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DUTCH N.W. NEW GUINEA.
A CONTRIBUTION
TO THE
PHYTOGEOGRAPHY AND FLORA
OF THE
ARFAK MOUNTAINS, &c.
BY
L. S. GIBBS, F.L.S., F.R.M.S.
With Four Plates and Sixteen Text-figures.
(Read before the Newcastle Meeting of the British Association
for the Advancement of Science, September 1916.)
LONDON:
TAYLOR AND FRANCIS, RED LION COURT, FLEET STREET.
JULY 1017.
Price 12s. 6
3U
CONTENTS.
PAGE
INTRODUCTION 1
1. HISTORICAL ... ... ... ... ... ... ... ... 1
2. SUMMARY OF PREVIOUS WORK 2
3. SCOPE AND CONDITIONS OP PRESENT WORK ... ... ... ... 5
GENERAL CHARACTERS OF THE COUNTRY 7
1. TOPOGRAPIIY ... ... ... ... ... ... ... ... 7
a. The Immediate Shore-Line ... ... ... ... ... ... 8
1. " Korang " or Coral-Limestone Zone ... ... ... ... 9
c. Inhabited Zone of Foot-hills and Lower Ranges ... ... ... 11
d. Crests of Main Range and Lake Basins ... ... ... ... 11
2. METEOROLOGY 13
3. PlIYTOGEOGRAPHY 14
ITINERARY AND DESCRIPTIVE ACCOUNT OF VEGETATION ... 15
a. BEACH FORMATION : Immediate Shore-Line ... ... 15
1. INUNDATION FOREST BELT : " Korang " or Coral- Limestone Zone ... 16
c. SECONDARY ASSOCIATIONS : Inhabited Zone of Foot-hills and Lower
Ranges 18
d. Low MOUNTAIN FOREST FORMATION ABOVE 7000' (to which syste-
matic collection was limited) : Crests of Main Range and Lake
Basins ... ... ... ... ... 21
PLANT ASSOCIATIONS OF THE LOW MOUNTAIN FOREST
FORMATION- 27
A. FOREST ASSOCIATIONS ... ... ... ... ... 27
1 a. Mossless Forest.
1 ft. Aruucaria Forest.
2 a. Intermediate Mossy Forest.
2 /3. Dwarfed Intermediate Mossy Forest.
2 y. Shrubberies.
3. Mossy Forest.
PAGE
a. S.W. RIDGE .' 27
la. Mossless Forest, 7000-7500' (trees 40-50') 27
2 a, ft. Intermediate Mossy Forest, 7500-8500' (trees 20-40')... 28
3. Mossy Forest, 8000' (trees 10-20') 29
1. LAKE BASIN, 7000' 29
2 a. Intermediate Mossy Forest (trees 40-50') 29
1/3. Araucaria Forest ... ... ... ... ... ... 31
3. Mossy Forest 32
2 j3. Dwarfed Intermediate Mossy Forest (trees 10') 33
c. SLOPES OF KOEBRE MOUNTAIN ... ... ... ... ... 33
2 a. Intermediate Mossy Forest, 7000-7500' (trees 30-40') ... 33
2 /3. Dwarfed Intermediate Mossy Forest, 7500-8500' (trees
10-30') 33
2 y. Shrubberies bordering Crest of Mountain, 8500-9000' ... 34
B. OPEN "OPPORTUNITY" ASSOCIATIONS ... ... ... ... ... 34
a. S.W. EIDGE, 7000-8500' ' 34
Papuan Rest- and Camping-places ... ... ... ... 34
Small Landslips 35
b. MARSH BY $ LAKE, 7000' 35
Sand Pans with running water ... ... ... ... ... 35
Denser Sedge Growth in Boggy Areas ... ... 36
Shallow Standing Water 37
Grass-jungle on Edge of Forest ... ... ... ... ... 37
c. OPEN SUMMIT OF KOEBRE MOUNTAIN, 9030' ... ... ... 37
Cladonia Association ... ... ... ... ... ... 37
PHYTOGEOGRAPHICAL CONCLUSIONS 39
SOME PLANT ASSOCIATIONS OF THE N.W. COAST 49
SYSTEMATIC RESULTS 50
PRINCIPAL BIBLIOGRAPHY 58
SYSTEMATIC ACCOUNT OF THE PLANTS COLLECTED 59
1. ARFAK MOUNTAINS ... ... ... ... ... 59
2. DOREI BAT, ISLANDS TO HUMBOLDT BAY ... ... 185
EXPLANATION OF THE PLATES 223
\_The altitudes given are approximate^
A CONTRIBUTION
TO THE
PHYTOGEOGRAPHY AND FLORA
OF
DUTCH N.W. NEW GUINEA.
INTRODUCTION.
1. HISTORICAL.
THE history of the navigation and exploration of Now Guinea from the
earliest times has been most exhaustively summarised by Dr. Wichmann in
the two volumes of history included in " Nova Guinea." The following
account is, therefore, strictly limited to records of botanical collections and
observations in N.W. New Guinea alone.
From historical times the N.W. coast of Dutch New Guinea paid tribute
to the Sultans of Tidor, and was subsequently included in the Residency of
Termite, which now administers the whole of the Sultan's dominions.
Termite, the capital, has always been the centre of the N. New Guinea
trade in Bird*-of-Paradise, the " Passaros del Sol " of the old Portuguese
navigators, who widely distributed these highly prized objects both east and
west, Oriental potentates and Moorish sultans with their courtiers vying in
the possession of such treasured symbols of royal power and magnificence,
which adorned the headgear on ceremonious occasions.
Subsequent to this the golden returns from the spice trade in the
Moluccas, a monopoly most stringently held by the Dutch East India Co.,
attracted the attention of other European nations. In the search both for
wild varieties of the precious trees and suitable areas for possible exploitation,
N. New Guinea offered the most promising field, the Dutch, from their base
in the Moluccas, again holding most of the trumps. All navigation or
detailed survey of the N.W. coasts of the country, with investigation into
the numbers and condition of the inhabitants, dates from this period about
the 18th century.
In j-ecent years, thanks to the dictates of fashion and enhanced value, the
volume of the trade in Birds-of-Paradise has again enormously increased,
regulated, however, in the Dutch Possessions by most adequate measures,
stringently enforced, for the protection of these beautiful creatures.
All intercourse in N.W. New Guinea during the open season is dependent
on this trade. Thanks to the enterprise of Tidorese, Malay, Arab, and
Chinese traders, in whose hands it is chiefly concentrated, the coast Papuans
have been brought more or less into contact with the outside world. This
has resulted in a certain amount of intermarriage, as a ready means of con-
solidating and extending trade relations in the chief centres of distribution,
and also in spreading a knowledge of Malay, even in remoter communities.
In the season the chief trading-stations swarm with the miscellaneous agents
engaged in this lucrative business, whilst the Papuans are occupied in hunting
in the interior on their own account, or for the various Ternate traders or
agents, so that it is next to impossible to obtain quarters or procure native
hunters or carriers. It is advisable to take this fact into consideration in
connection with biological work in this part of the country.
2. SUMMARY OF PREVIOUS WORK.
Geelvink Bay was first discovered and mapped out in 1705 (21, i.
138-152) by Jacob Weyland, who commanded the ships 'Geelvink' and
' Nova Guinea/ In the course of his work he touched at Dorei Bay,
situated to the N.W. of Geelvink Bay.
In 1775 Forrest (1, 95-114) in the 'Tartar,' a 10-ton ship, in which
he had sailed from Balambangan to obtain nutmeg and clove trees for
the purpose of introducing them into that island, l spent part of January
and February at Dorei, enjoying fine weather all the time, and his
excellent straightforward account is quite one of the best relating to this
place. The 'Tartar' lay off Wousi, where Forrest describes the whaleback
Papuan houses spreading over the water, as they are still to be seen at the
present day, and also the Arfak Mountains rising below Dorei. He even
mentions Oranswari and Wariap as two inhabited places along the coast
to the south, and notes the correct time to reach them by fast sailing in a
favourable wind. He states that the people had many prnus, and were said
1 Off N. Borneo.
to deal honestly with the Chinese who traded with them, conditions which
still prevail.
In July 182-1, Duperrey (21, i. 316) in the 'Coquille' spent over a
fortnight at Dorei, and the first collection of plants from there was made
by Lesson, the surgeon accompanying the expedition (2, ii. 534).
Dumont d'Urville (3, iv. 578-612) in the Voyage of the ' Astrolabe ' gives
a verv good general account of Dorei Bay, where he stayed in August 1827,
after the discovery of Humboldt Bay, which was named, but not touched ;it.
He mentions taking in water " au limpide ruisseau de Wirsi," and refers to
the very rainy weather. The local Papuans are described as living in terror
of the Arfakis, some mountain tribes established above Wousi on what is
now the site of Manokoeari, who were always treacherously attacking the
coast people, one such attack occurring duriug the stay of the ' Astrolabe/
Lesson, who accompanied him, made another collection of plants.
Wallace in 1858 (5, ii. 298-326) spent three months and a half, from
March to July, at Dorei Bay collecting zoological specimens, and was there
when the S.S. 'Etna' (8, 78), fitted out by the Dutch Netherlands Indian
Government for ethnological and natural history investigations, put into
the bay in June of that year. Some of the 'Etna's' company attempted to
ascend the Arfak (8, 73) from the N. side, but turned back after reaching
1500'.
In September 1872, D'Albertis (9, 67-71), having spent three months at
Andai with Beccari making zoological collections, succeeded in reaching
Hatam, an Alfuero village, situated at 5000', three days' journey from Andai,
where he spent a month, enduring great privations. This intrepid explorer
was the first European to penetrate into these mountains and reveal their
ornithological treasures, for, of the magnificent collection of Birds-of-
Paradise and other types obtained by him, nearly all proved new to
science.
In June 1875, Hatam was again visited, this time by Beccari (11, 35-38),
who spent a month in this inhospitable place, much hindered by rains and
swollen torrents. He attained 2040 in. in exploring the surrounding heights,
and brought back, in addition to much valuable zoological material, a large
collection of plants unfortunately not yet fully worked out.
Rosenberg in 1869 and 1870 (10) made several journeys to Dorei and
the adjacent islands of the N.W. coast, of which he gives an interesting
description. Accompanied by Malay hunters he spent three months at
Andai, staying with the Missionary VVoelders, with the intention of attaining
Hatam (10, 99). Several abortive attempts were made, and he claims this
achievement for his hunters, a claim since disproved (21, ii. 142). A
vocabulary of the Noemfoer (coast) and the Alfuero (mountain) languages,
with some excellent original drawings, are included in his book.
32
Teysmann, the well-known Dutch botanist, visited Dorei in 1871 (14,
61-95), and also touched at Andai and Wariap in coasting up Geelvink
Bay, making large collections of plants.
In 1891 one of Veitch's orchid collectors, David Burke (21, ii. 539),
a gardener, made his way to Small Hatam, a remarkable feat, referred to
by E. St. Vraz, who in 1896 (15, 232-235) spent several weeks in Great
Hatiim, some distance beyond Small Hatam, which he states was D'Albertis'
and Beccari's objective. Though much hampered by rain and the usual
difficulties with carriers and hunters, the standing barrier to all work in the
mountains of this country, St. Vraz's account is detailed and interesting.
In 1898 Manokoeari (21, ii. 677), the old Alfuero "campong" 1 on
Dorei Bay, was raised to the status of a Government Station, to which
Mr. van Oosterzee was appointed Assistant Resident, an appointment held
until 1913, when he was obliged to retire through ill-health.
In April 1904 (17, 998-1021) van Qosterzee visited the Angi lakes,
the first European to explore the more southern portion of the Arf'ak. On
arriving at Ternate, on the way to Manokoeari, I heard with great regret of
the serious illness of this very able administrator, who during his 16 years'
residence at Manokoeari had made himself familiar with the Papuans and
their language, his sympathetic rule giving him unusual influence with the
chiefs, or "Koranos" as they are called. Manokoeari also bears witness
to his initiative in the beautifully situated Residency and " Pasangrahan," 2
in avenues of Casuarinas shading the level white roads, in the efficient water
supply, substantial pier with well-arranged godowns, and the broad street of
Chinese " tokos " 3 .
In 1908 Pratt, the well-known collector, with his two sons, spent some
time at the Angi lakes. They were accompanied in the first instance by
van Oosterzee, who, however, returned immediately to the const.
In 1912 (25, 77-78) Gjellerup, attached ;is Officer of Health to the
Exploration Detachment 1911-12, proceeded to the lakes, accompanied by
a mining engineer named Hubrecht. They spent April and May in that
region, where the former made important botanical collections, now in course
of publication in " Nova Guinea/' Unfortunately, many specimens were lost
owing to the desertion of his carriers.
I had the pleasure of meeting Dr. Gjellerup in Java soon after his return
from New Guinea, and acknowledge with keen appreciation his generous
kindness in giving me the fullest benefit of his experiences, not only
in relation to valuable details in organization and the areas to which he
had devoted most attention, but also for most welcome data on Papuan
1 Campong, a native settlement or village.
a Rest-house.
3 Shops.
idiosyncrasies, with special reference to the mountain tribes and the best
method of dealing with them.
On my return from Wariap in 19 L3 I was fortunate in finding Mr. Pratt
and his sons at Manokoeari, where they had arrived to organize another
expedition to the Angi lakes. I gladly welcome this opportunity of thanking
them for permission to reproduce the successful photographs subsequently
taken in that region, including several purely botanical subjects selected
especially at my request. My own results there had not proved satisfactory.
3. SCOPE AND CONDITIONS OF PRESENT WORK.
The chief objective of the present work was a direct outcome of results
obtained and observations made on Mt. Kinabaln in British North Borneo
in 1910.
The great interest in the higher regions of that, mountain lies in the extra-
Malayan facies of the vegetation and elements in the systematic composition
of the flora. The predominance there of species in Orchidacese, Myrtacea3,
Ericaceae, chiefly Rhodoilemlrons and Vacciniums, is such a striking feature,
that one felt it must be derived from some more continental type of
vegetation than the limited occurrence on an isolated mountain could
account for.
Celebes and the Moluccas did not suggest sufficient scope for the proposed
key to the solution ; but the mountains of New Guinea with their huge
area, approximate rainfall, and great altitude offered a perfect basis of
comparison ; while Dr. Beccari's description in "Malesia" of his work and
observations on the plants found by him on the Arfak mountains showed a
very close approach to the phytogeographical facies of the Kinahnlu " Massiv."
This impression was further confirmed by the account of Mr. Pratt's first
expedition to the more southern region of the range, and a few plants collected
by him in an open marsh by the larger of the two Angi lakes, kindly
shown to me at Kevv by Dr. Stapf, finally led me to decide on that locality
for comparative work.
Through the broad-minded interest and influence of my friend the late
Sir Kenelm Digby, permission for the expedition was accorded, through the
kind offices of the Foreign Office, by the Government, of the Netherlands.
I must express my deep gratitude to His Excellency Mr. A. W. F. Idenburg,
then Governor-General of the Netherlands-India, for the extreme courtesy
of his reception and the interest he evinced in the scientific objects of the
projected work, the success of which was assured, thanks to his generous
assistance and detailed instructions.
My thanks are also due to Mr. 0. Lulofs and Mr. J. C. Kielstra for
much kind attention during my stay at'Buitenzorg.
At Ternate the Resident, Mr. Gh. L. J. Palmer van den Broek, was
again most kind in forwarding my plans, and from Mr. L. J. J. M. (now
Captain) Tabbers, Acting Assistant-Resident at Manokoeari, I met with
every sissistance which military training in grasp of the situation, judgment
of men, and appreciation of fact and detail could do to ensure success.
To Captain J. .F. E. ten Kloosten, who most kindly procured me
Dr. Gjellerup's sketch-map of the Angi lakes from the military carto-
graphical office at the Wousi Bivouac, with permission to reproduce the
same in this work ; to Mr. J. W. Langeler, of the Dutch Navy, attached
to the Mamberamo expedition ; and to Mr. F. J. F. van Hasselt, Chief
Missionary at Manokoeari, I am further indebted for much help and
thoughtful consideration.
Any difficulty in reaching Wariap, about 60 miles south of Manokoeari,
the best point of departure for the mountains, was obviated through the
kind offices of Captain J. V. L. Opperman, Commander of the Mamberamo
expedition 1913-1914. He gave me, with the escort kindly provided by the
Dutch Government, a passage in the Government boat the ' Valk,' which
was conveying his expedition to its destination. The ' Valk ' landed us at
Waren, 10 miles south of Wariap.
As there had been recent fighting between the coast and hill people,
Mr. Tabbers judged it advisable to send an unusually strong escort, com-
prising five Amboinese armed policemen, " Pradjoerit," and ten convicts,
" Orang rante," attached to them, to help in the carriage of provisions should
the Papuans desert, according to their usual custom. It was no doubt partly
owing to these two circumstances that all my carriers remained with me on
this occasion ; I noticed great reluctance to stray far from the protection
of the guns.
The sergeant in charge, always called " Serzan," a Timorese, who had
served for 20 years through the Achin campaigns, was a. most capable and
tireless little man, a thorough soldier, not only very clever in handling his
own men, who did him credit, but also in dealing with the Papuans, not
such an easy matter.
The " Orang rante," all enormous men, who made a great impression on
the Papuans, turned out very handy and willing. One of them acted as
my cook, both in the mountains and until my departure from Manokoeari,
while several became quite expert in carrying out botanical processes.
It is a great tribute to the humanity of the Dutch rule that men like these
Malays, to whom confinement is death, should be sent to expiate their
offences, chiefly crimes due to jealousy and gambling, in outlying stations,
where they have congenial work under easy and almost independent
conditions. Escape is out of the question, as the Papuans shoot at sight
any stragglers in the forest so it is not even necessary to guard them, while
the best behaved are allowed to earn money by gardening or acting as
" boys " to the civil and military officers.
GENERAL CHARACTERS OF THE COUNTRY.
1. TOPOGRAPHY.
The north-west coast of New Guinea is bleak, mountainous, and sparsely
inhabited. The mountains, from 5000-7000', rise abruptly from the sea,
there being little sloping foreground and few small bays to shelter
FIG. 1.
N.W. NEW GUINEA.
the schooners and Papuan praus, which only trade during the favourable
inonsun. There is no good anchorage tor large boats between Sorong, an
island on the extreme north-west, the first point of call, and Dorei Bay,
a fine harbour, consisting of two bays, sheltered by the islands of Mansinam
and the much smaller Meoswar. A low mangrove spit, on which a Papuan
" cainpong " is situated, separates the two bays, the further one being much
smaller, very shallow, and probably rapidly silting up.
Manokoeari lies in the centre, on the lower slopes of a very recent
u korang '' or coral limestone range about 500' in height, while to the
east the coast runs out into a low peninsula, where most of the Papuan
plantations are situated, the soil there being much better than that of the
sterile " korang " of the range.
To the south rise the Arfak Mountains, 9000' high, which run in
parallel ranges down the western shore of Greelvink Bay, at about forty
to fifty miles inland, except at one point Tanaroeboe, one day's journey
along the beach from Andai, where spurs from the mountains abut steeply
on to the sea. How far south the range extends has not been determined,
nor whether it is continuous with the mountain chain of the north-west coast,
as some writers have inferred.
The main buttress of the Arfak rises above Andai, a Papuan " campong "
at the mouth of the Andai River, about four to five hours by native prau
across the bay from Manokoeari. From Andai there is a tract to Amberbaki
on the north-west coast, which passes through Hatam, a four days' journey
according to Rosenberg (10, 79).
The two small Angi lakes, to the vicinity of which this collection was
limited, lie at an altitude of 7000' and 9000' respectively, on the southern
portion of the range. The lakes are accessible from several points on the
coast, of which the immediate shore-line only is sparsely inhabited, the
few inhabitants being restricted to the beach, hence their name, " Orang
pantai " or " beach people/'
A huge intervening low-lying belt of coral limestone or " korang "
extends from the beach to the loot-hills of the Arfak. This tract of country,
intersected by the alluvial terraces and large inundation-areas of the rivers,
which pour down from the mountains in the rainy season, is devoid of
inhabitants and suggests very recent elevation.
The main topographical features of this part of the W. coast of Geelvink
Bay may therefore be roughly divided into four zones, which also corre-
spond to the general plant-formations : (a) The Immediate Shore-Line ;
(b) "Korang" or Coral-Limestone Zone; (c) Inhabited Zone of Foot-hills
and lower Ranges; (d) Crests of main Range and Lake Basins from 7000'.
a. THE IMMEDIATE SHORE-LINE.
The immediate shore-line is sparsely inhabited, though many fine rivers
and torrents sweep down from the mountains. With the exception of some
mangrove formation, between Dorei Bay and Andai, and Oranswari and
Wariap, the beach sweeps southward, consisting of big stones, shingle, or
9
loose sand. Large blocks of coral, as it' just broken off from the reef, lie at
the water's edge, where the surf beats ceaselessly and the dip of the beach
is general!)' too steep for native praus to land without running the risk of
being smashed to pieces. It is the presence of landing-places, with good
water, whicli limits the stages in the tedious journey along the beach from
Andai to Sjari.
The rivers in flowing into the sea either break up into many shifting
mouths or are barred by banks of sand or shingle into semi-lagoons, with
only a small exit to the sea.
Most of the trees fringing the beach are prostrate or semi-prostrate, as if
torn up by the force of the waves. Stagnant lagoons, impenetrable bog, and
shallow standing water occur in parts, while the undergrowth is covered
and the ground strewn with seaweed, evidence of the retreating swirl of
great waves. The natives told me that when the north wind blows the
sea washes all over this belt of country, fish being often found stranded
on bushes far inland.
The few small native ; 'campongs" are placed just above the beach,
whore sand-banks have accumulated, on which some Casuarinas mark the
permanence, as at Wariap and Waren.
I returned from Wariap by the beach in December 1913, the first time
this journey had been made by a European, to be followed by Mr. Pratt in
April 1914, at the height of the north monsun. He described the whole
region as then more or less under water, the rivers, pouring down from the
mountains in floods, being beaten back over the land by the huge surf raised
by the north wind, which bars the exit of their waters to the sea.
This interesting observation accounts for the shifting river mouths, and
also explains why the native habitations are generally so far from water and
so few in number.
1. " KORANG " Ott CORAL-LIMESTONE ZONE.
Behind the beach the low-lying belt of " korang," covered with forest,
stretches uniformly from the coast to the foot-hills of the Arfak, a sterile
porous formation showing so little depth of soil that it gives the impression
of walking over a reef.
Rosenberg (10, 80) in 1870 refers to the recent appearance of this coral-
limestone area, which he concluded must be still rising, and he quotes the
older inhabitants of Andai as saying that they remembered low scrub where
the forest now stands.
Van Gelder (20, 94) considers that a gradual rising of the whole of the
north coast of New Guinea is taking place, or a lowering of the sea-level,
which amounts to the same thing. He found evidence of this fact at
10
Manokoeari, in the presence along the beach, at slight elevation above the
sea, of an undoubtedly very recent " Schelp Gonglomeraat " (shell-limestone)
that must have been formed in the surf zone.
The rivers cut their way through alluvial deposits of mud or sand, or
form great open spaces their present inundation-areas, all sand and stones
in the dry season, but which, in the wet, are lakes of standing water.
MAP OF ANOI LAKES
(reduced from Dr, K. G. Gjellerup's sketch-map).
Dr. Gjellerup's route.
At Wanessi, the head of such an area on the Momi River, a day's journey
from Wariap, the remains of an old " campong " can still be seen, where
some of the hill people tried to establish themselves, but were forced to retire
to Wariap on account of the floods.
This belt is uninhabited.
11
c. INHABITED ZONE OF FOOT-HILLS AND LOWER RANGES.
Once on the foot-hills the character of the country changes : the ground
is broken by rocky outcrops and boulders, and the streams are now mountain
torrents, plunging over great boulders which form their beds.
Crossing the Soedomi River, a rocky stream which falls into the Momi,
one mounts steeply up the flanks of the spurs to the crests of the ridges,
which are cleared. Native houses are first seen at about 1000', and from
thence upwards, on the crests and slopes of the ridges, where the soil is
deep and easy to work, there is evidence of extensive cultivation, past
and present.
All the region of the subsidiary spurs and lower ranges seems to be
inhabited. From commanding views it is easy to trace the line of the ridge,
by which the approach to the lakes is made from Sjari, by the smoke rising
from the houses, which are always perched in a conspicuous position,
probably for the double purpose of observation and defence. Only in the
case of one or two head-hunting tribes, who seem to inspire the whole
neighbourhood with terror, were the houses situated on the lower slopes.
On the third, or Sjari ridge, from the Soedomi River, one looks down on
to the Momi again, which rises in these mountains at right angles to its
coast course. From below Soekoem the course of the Momi River (fig. 2)
is erroneously indicated ; it should be that suggested for the Runsiki River
(fig- 2).
Van Oosterzee (17, 999), who went up from Sjari, 20 miles along the coast
from Wariap and 10 miles south o Warcn, mentions crossing the rivers
Waren and Waidiri, both flowing into the Momi, but the Waren enters the
sea at Waren, halfway between Sjari and Wariap. The Soedomi was the
only river we crossed, coming up from Wariap, till the Momi was again met
with as a rocky torrent. Crossing to the west bank of the Momi we climbed
another long spur in a westerly direction, till the latter joined the crest of
the main range at 7000', called the S.W. ridge in this work, where the
inhabited zone suddenly ceases, a fact probably determined by the steeper
gradients and narrower crests, and the exposed and poor gravelly soil.
d. CRESTS OF MAIN RANGE AND LAKE BASINS.
The route to the lakes lies along the narrow crest of this ridge, marked
as " Bonjas Gebebergte " on fig. 2, which runs S. t S.W. from 5000-8500' in
height, bounded on the west by a valley, said by the Papuans to be that
of the river issuing from the largest of the two lakes (Tilaan of fig. 2), and
on the east by short spurs which run down to the Momi valley.
The Angi lakes are situated at about 7000' and 8000' respectively,
separated on the west by the long Koebre Mountain, 9000' high, and
12
bounded to the east and the south by the S.W. ridge. The largest lake is
called the Wannasin Bean or " Woman " Lake by the Papuans, in contra-
distinction to the higher and smaller one, the Warmasin Snoon or "Man"
Lake. These names are so well recognised that the Malays with me
invariably referred to each lake as "Prempuan" or "Anak," the Malay
translation. According to Gjellerup (25, 77), the greatest length of the
lowest or " Woman " lake is 9 km. south-west to north-east, and the greatest
breadth is 4 km. on the south side. This lake is so deep that van Oosterzee
(17, 1010) found that, on the south side, at a few hundred in. from the
edge, he could no longer touch the bottom after 90 m. Grjellerup (25, 77)
states that eels are the only living fish in it. On the north side the con-
verging hills gradually slope into a marsh, the exit of the Tilaan River,
which, according to both van Oosterzee (17, 1018) and Gjellerup (25, 77)
joins the Ransiki River, which is stated by the former to fall into the sea
at Wariap, obviously a mistake for the Momi River.
The western shore of this lake is inhabited by a small Alfuero tribe, their
communal houses being in four groups on the extreme edge of the water,
in separate bays, the " kebuns " stretching up the slopes behind. Further on,
the sides of Koebre rise steeply from the water, while to the south a low
ridge, about a couple of hundred feet high, connects with the S.W. ridge,
which forms the boundary to the east.
The smaller or " Man " lake is reckoned by Gjellerup (25, 77) as roughly
about 4 km. from the " Woman " lake, and he gives its north and west sides
as inhabited. Van Oosterzee (17, 1013), who spent most of his time on
the upper lake, estimated it to be 7 km. long, and 2 broad in the southern
portion and he counted some thirty houses on the western shore with
about 1000 souls ; the houses seemed to me more numerous to the south-
west, and I hardly think there can be so many inhabitants at the present
time. He mentions the morass at the north end of the lake, also rafts
on its surface, the cultivation of potatoes on the shores, and that eels
are the only fish found in its waters. St. Vraz (15, 234), while at Great
Hatam, heard from the natives that at four days' journey south there was a
large lake called " Tschemti," with many fish and crocodiles, on whose banks
Manikianos lived.
Inquiring of some of the Papuans with me, who had been up at the lakes
before, if it were possible to reach the north coast and Manokoeari from the
upper lake, they answered " Yes," and that it took twenty days, but, of course,
in their estimate of time they would lose count beyond a certain number.
Van Oosterzee met some Papuans (17, 1010) by the lower lake, who told
him they had come from the north, and Gjellerup (25, 77) also mentions a
way to the north coast, facts which agree with St. Vraz's information.
As I heard it was possible to return to Wariap in two days by keeping
to the S.W. ridge, I decided at once on such a welcome alternative to the
13
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five days spent in coining up. Our return
route therefore followed the S.W. ridge,
leaving the spur by which we had ap-
proached it from the Momi to the right.
Continuing due north we gradually de-
scended to about 5000', then bearing east
along a lateral spur which finally dipped
very steeply to the rocky bed of a tribu-
tary of the Moini. After crossing the
latter, we emerged on to the north bank
of the Momi River, whore we bivouacked
for that night, returning to Wariap the
next day.
The chief difficulty of this somewhat
strenuous route is the want of water,
there being none between the vicinity of
the lakes and the tributary of the Momi.
According to fig. 2 this route would
follow the " Bonyas Gebebergte," which
join the spurs of the Arfak indicated on
the sketch-map to the north of the sup-
positional position of the Ransiki River.
The altitudes in the sketch-map,
judging by the character of the vege-
tation, seem somewhat underestimated.
2. METEOROLOGY.
The following facts referring to the
north-west coast are taken from Braak's
(24, 210) paper on the climate of New
Guinea, which summarizes all the infor-
mation at present available.
On the north coast the east or dry
monsun prevails from May and June to
September and October, though even
then much rain may fall. In Mano-
koeari the east monsun is the driest time
(24, 219). It is probable that in the
interior the difference between the cha-
racter of the E. and W. monsun is much
less pronounced.
In N. New Guinea it is at present
not possible to give actual figures, though
14
it may be accepted that the temperature throughout the whole year is very
constant, and that the mean variation is probably between 26 and 27 C.
(24,223).
Braak's table (24, 221) quotes the average rainfall for several years for
Sorong, Manokoeari, Djende on Roon Island, and Windesi to the south
of Geelvink Bay. In both the latter stations the rainfall is much higher
than on the more exposed N. const at Sorong and Manokoeari.
In the historical summary the prevailing weather has been quoted
whenever recorded, and it will be seen to vary considerably. At Mano-
koeari, January and June were considered the wettest months. During
a stay there in November 1913 the weather was very fine; jbut on my
return in December the rainy season had set in and there was rain every
day, generally in the afternoon, the temperature being markedly cooler.
In the intervening period, spent at Waren and Wariap, on the coast of
Geelvink Bay, in the Arfak, and on returning- to Manokoeari along the
coast, only three half-days of rain were experienced, and these occurred on
the way up and while at the lakes, where the rain was also accompanied
by strong \A'ind. Otherwise the fine still weather was a constant source of
amazement to the Papuans, the rainy season being well overdue, and they
attributed this lucky chance to the fact that sometimes at the ch;mge of the
monsun a short halcyon period of fine weather sets in.
At Waren and Wariap a very strong N.W. breeze invariably sprang up
between 3 and 4 P.M., causing a sudden fall of temperature.
Unfortunately my only available calendar was lost at Waren. Con-
sequently, no readings were taken either there or at Wariap, and only one or
two in the mountains, which were as follows :
On the Momi River, 3500'. 75 F. 6 P.M.
On the crest of the S.W. ridge, 8500'. 68 F. 4 P.M.
On the "Woman" lake, 7000'. 60 F. 7 A.M. ; and on the following
day at the same hour, 55 F.
Up at the lakes it was always cool in the early morning, the sun being
very hot in the middle of the day, but cooling down in the afternoon and at
night, though warmer thau at the same altitude on Kinabalu. Both Pratt
and Gjellerup reported very wet weather during their several stays at the
3. PHYTOGEOGRAPHY.
The general plant-formations, as is shown below, agree in broad outline
with the topographical zones already described. They are :
(a) Beach Formation : Immediate Shore-Line.
(6) Inundation Forest Belt : " Koran g " or Coral-Limestone Zone,
15
(c) Secondary Associations : Inhabited Zone of Foot-hills or Lower
Ranges.
(d) Low Mountain Forest above 7000' : Crests of Main Range and
Lake Basins, 7000-9000'.
All systematic collection was limited to the last formation.
ITINERARY AND DESCRIPTIVE ACCOUNT OF
VEGETATION.
(a) BEACH FORMATION.
Permanent sand-spits, Waren and Wariap.
The beach at Waren forms a long sweep on ench side as far the eye can
reach. A plantation belonging to a Japanese, who with his son had permission
to accompany me to the lakes, was situated at the mouth of the Waren River,
which, dammed up by a sand-bank, formed a green lagoon, with only a very
narrow outlet to the sea. A clnmp of Casuarina equisetifolia proved a certain
stability, but the river-exit, with the dip of the beach, must be always in a
state of flux with each N. monsun season. The great accumulation of s;md
to which Waren and also Wariap owe their security from the inroads of the
surf, must be due to the amount brought down and deposited by the rivers
at their mouths.
Where the beach broadens out in the immediate vicinity of Waren,
a Pes-caprcB association with Tacca pinnatifida is formed, to be succeeded by
typical beach-shrubs, like Tliespesia populnea, Canavalia obtusifolia, Sccevola
Koenigii, Vitex trifolia, Clerodendrun inerme, Premna nitida, and a Gmelina,
probably villosa, which must successively bank up the sand against the wash
of the surf, as the Japanese had cleared behind them and planted coco-nuts
on the pure sand, with cotton and pineapples asundercrops. This is the only
spot along the coast besides Wariap where such a risk could be taken. Where
the beach was lower and narrower, the surf w:ithed through beach-jungle or
under Casuarinas to the overhanging fringe of forest trees.
Sub-emerged Beach.
It is several hours from Waren to Wariap along the coast to the north,
by what may be styled a sub-emerged beach. A little beyond Waren the
sand decreases in volume, strand plants disappear, and the beach narrows
considerably. Huge trees of Barringtonia speciosa lie prostrate to semi-
prostrate over the sea. Young plants of Pandamis &\>.,Draca>na angustifolia,
an immense Crinum, probably C. macrantheruin, with giant stools of
Asplenium Nidus, no doubt displaced from the branches of trees as they
fell, crowd the ground, all dusted over and growing plentifully amongst
much water-washed debris and plant detritus, both terrestrial and marine,
16
which, with myriad prostrate Barringtonia-seedVmgs, attached by the one
anchor-root, all pointing seawards, bear witness to the force of great waves
retreating from their rush inland.
In many places this undergrowth was so thick, or the prostrate trunks so
numerous, that it was easiest to walk through the surf, outside the branches
of the fringing trees. The rivers, where there is no sand to bar their exit,
form small mangrove swamps at their mouths, which have to be waded
through. In one of these small swamps a tree covered with a vine of
Mucuna Kratkei,a\so known from the N.E. and S.W., whose numerous huge
racemes formed a dome of brilliant red flowers, was a magnificent sight.
Wariap.
Here, where the people all remembered Mr. van Oosterzee and the
Pratts, I was welcomed as an old friend the "korano," a very fine
man physically and quite a personality, and the " guru " (teaclier), to whom
I had a letter from Mr. van Hasselt, having already paid their respects at
Waren. It was arranged that the "korano," Manao, should act as guide to
my party to the Likes, and the Wariap people of themselves offered to
accompany me as carriers, promising to remain as long as I stayed there
a promise sealed on " Pinang " and " Zabacco," as they call the latter, and
faithfully kept. " Pinang " replaces betel-nut on the coast of N. New
Guinea, being obtained from the wild Areca macrocalyv Zipp. (12, i. 18)
and eaten with lime and the fruit of Piper Siriboa (14, 69).
Wariap, situated on a sand-spit through which the Momi has cut its
broad way on one side, forming a good harbour for praus, while on the
other Casuarinas are massed, is quite a large and busy " campong," where
much prau-building and making of Pandanus mats (nokes) and sago-holders
is carried on.
The long whale-backed houses are built above the beach, on a level spit
of very fine sand, which, overgrown with grass and Pes-Caprce, is broken by
shallow green lagoons shadowed by a jungle upgrowth of Thespesia populnea,
Abrus precatorius, Ccesalpinia Nuga, Wedelia tiflora> etc.
(V) INUNDATION FOREST BELT.
Just behind the beach formation this forest forms a huge unbroken green
wall, in which the pyramidal branching of Terminalia Catappa is easily
distinguished from the outside, whilst most of the trees are covered with the
heavy green curtains, falling straight from the crowns, of Zanonia macro-
carpa, a Cucurbitaceous liane. In this forest Ficus, Macaranya, and
Artocarpus sp., the latter with enormous leaves often about 1 m. long
and m. or more broad, mostly predominate their trunks screened with
immense fronds of climbing ferns, spreading radially all the way up,
or Epipremnopsis Hugeliana, fiaphidophora Peepla, other Philodendron spp.,
17
Piper Forstenii, Potlios, and various large-leaved root-climbers. Piles of
these huge leaves accumulate under the trees, to be dispersed by the floods
of the rainy season. A giant Korthalsia, its interlaced stems scrambling up
and down the trees or spreading in tangled mass over the ground, was the
only " rotan " seen. Asplenium Nidus was abundant ; but epiphytic orchids
were lew and far between, and the absence of flowers or fruits was most
striking.
In this forest there is little undergrowth. Sodden leaves mostly fill up
the interstices of the porous korang. Where light shoots through, some
thin grass or patches of the creeping Geoplnla renifornris and Ilemigraphis
re plans, or single specimens of the small semi-herbaceous shrub Amarocarpus
Wichmannii, with horizontal dorsi-ventral branches, appear ; but the most
conspicuous objects are the huge Z&ntaua capsules, the size of large
pumpkins, in all stages of decay, scattered over the " korang," which is so
porous that surface-water soon drains through; but where there is standing
water, groups of Sago Palms occur, often forming swamps covering large
areas.
A track from Waren to the Arfak runs for a couple of days through
Sago swamps.
Native Plantations.
These are dotted through this forest where little islands of soil accumulate,
as at Waren, or alluvial deposits have been formed by the rivers, as at
Wariap. Here the well-stocked " kebuns " 1 surprised me by their extent,
many kinds of bananas and plantains, coco-nuts, papaya, cassava, " kladi," 2
" ubi," 3 "labu," 4 egg-fruit, and various " sayur" 5 etc. being grown, with
clumps of bamboo, probably planted.
On passing through this " korang " zone we followed a new route,
keeping to the south bank of the Momi, so obviating the necessity of
crossing the river, which is rather deep at its mouth. This track passes
through the Wariap plantations and the sterile "korang" forest beyond,
cutting off a great angle of the river. Then it skirts the shifting banks of
the river as the latter cuts through alluvial deposits of mud and sand,
or crosses wide sand-banks, the splash of crocodiles heralding our approach.
These sand-banks afforded a fine view over the Momi, showing Camarinas
and the symmetrical Terminalia, the latter veiled in all-obliterating Zanonia,
backed by the distant mountains. Thence the track alternates between the
dome-like " korang " forest and the thicker undergrowth of alluvial flats, or,
when nearer the river, over oozy slime which spreads over the stems and
leaves of a small Licuala palm and the giant KorlJuilsia, the chief under-
growth in such areas.
1 Gardens. a Colocasia antiquorum Schott. ' Sweet potatoes.
4 Gourds. ' Vegetables.
C
18
Momi Inundation Area.
From the forest one emerges into the blazing sunshine, on an open plain,
which it takes several hours to cross, all stones, gravel, sand, and " lalang,"
dotted with small trees of Casuarina equisetifolia. This open space forms an
inundation-area of the river, and is under water during the rains. Amongst
the stones Geodorum pictum, a pink-flowered orchid, was characteristic, and
Peristylus goodyeroides was found among the " lalang " which covered the
sandy areas. The most remarkable feature was abundant clumps of a new
Pteris, P. bamliisoides, with erect rhachises resembling stems rising from an
underground rhizome, each rhachis about 2 m. in height, clothed \vith
segments from the base to the apex, originally bilateral, becoming spiral
later through the twisting of the cortex. The species was also seen near
Wariap, in passing through the " kebuns." Professor Bower, to whom
I submitted this interesting new fern, suggests that " the general habit
might possibly compare with that of Pteris grandiflora, in so far that both
are probably creeping rhizomatous."
Pandanus Trees.
The second day, while still in the " korang " forest, we passed through a
striking group of old Pandanus trees, about 30 in. high, each rising out of
the forest on numberless grotesque stilt roots for about a quarter of the
height, succeeded by a straight stem with much-branched top ; the old
leaves hung in limp masses from every possible resting-place and strewed
the wet ground underneath. These weird trees gave a pregnant impression
of the scenic possibilities of this genus under primitive conditions. Even
the Malays were impressed ; but the Papuans spoke of similar isolated
groups scattered through this forest. These groups possibly represent
the first vegetative covering of the " korang/' displaced later by more
rapidly growing dicotyledonous forest trees, as only those plants which
could respond by equal vertical growth and so maintain the same level
as their competitors would have a chance of survival under such enveloping
conditions. No fruit was seen on the trees or underneath them.
(c) SECONDARY ASSOCIATIONS.
Once on the foot-hills the character of the forest changes, the sodden
effect of the "korang'* belt works out, and it loses the mud-washed look
and steaminess suggestive of constant inundation. The ground, rocky and
broken, is strewn with Sapindaceous and Anacardiaceous fruits, red Pometia,
and other brightly-coloured kinds, and undergrowth in the shape of plants
and shrubs appears, Musscenda frondosa being general. The foliage of both
trees and lianes is less monstrous and more varied in form, while a graceful
epiphytic flora puts in an appearance. From time to time small cleared
19
spaces were reached, evidently known rest camps, where we halted ten
minutes to rest the carriers.
After a tedious climb up the flanks of the spurs, clothed in high forest,
one emerges on to cleared narrow ridges, covered with long grass, showing
the first signs of cultivation, where a beautiful view opens out, on the
one side to the blue island-dotted waters of Geelvink Bay over the
foot-hills and the flat inundation-belt just passed through, which spreads
out like a green table, and on the other side across the Momi valley to the
central mountain range with its many outlying spurs. After more climbing,
bamboo thickets evidently planted, with the magic plant of Malaya, Justida
Gandarussa, never known to set seed, further confirm the impression of man's
vicinity. Fine forest succeeded these abandoned gardens, from which we
emerged on to old " kebuns " on the broad crest of the lower Serao range.
Here were the Serao houses, surrounded by present cultivation, where
we were very well received by the korano and his charming wife, the
prettiest Papuan woman seen, though many are nice-looking.
The Serao people great friends of Manao's cordially invited me to
sleep in their house ; but as all the Papuans with the Japanese streamed in,
to say nothing of the original inhabitants, I decided to camp outside in a
newly made " kebun," with the " Pradjoerit" and " Orang ranteV'
The korano's house was very large, with split-bamboo flooring and a few
small partitions, while against each side a narrow strip was thickly sanded
over for fires. Opposite the entrance a second door opened on to a balcony,
commanding a lovely view over the dip of the ridge to the immediate Momi
valley and the further spurs of the Arfak. A house inhabited by a head-
hunting tribe was pointed out on the slopes below.
Native Plantations.
In the " kebuns " the luxuriance of the crops and method in cultivation
is surprising. Sweet potatoes of very fine quality, gourds, plantains etc.,
and papayas, the latter replaced by maize and tobacco as the altitude
increases, with some of the finest sugar-cane I have seen, are planted,
the standing crops in diagonals alternately, with sweet potatoes and gourds,
chiefly Lagenaria hispida, as undercrops.
Some of these plantations were situated on the steepest slopes, where,
toiling up in the pitiless sun, one sinks to one's kneos in fine deep soil.
Fortunately there are always many logs lying in succession as they were
felled, which facilitate the ascent.
The plantations or gardens are surrounded by a strong double stockade
against wild pig, with notched poles slanting both ways at certain points for
ingress and egress. One or two communal houses are generally built in the
middle of the plantation, each on a maze of criss-crossed poles, about 20
from the ground, with a veranda back and front, approached by a notched
C2
20
pole from the front only. Where the crest is narrow the houses are built at
the edge, so that, entering on the level in front, the elevation at the hack
accords with the slope of the hill.
Strict etiquette demands that your name and business be shouted out by
the most sonorous voiced Papuan at the point of ingress into the plantation,
before intrusion on their domain, for the information of the owners. You
are then received by the inhabitants, both men and women, all standing or
sitting unarmed on the balcony, and after friendly greeting and distribution
of tobacco one passes on in peace. This custom is described in the account
of the ' Etna' expedition for the northern part of the Arfak (8, 74), and by
van Oosterzee (17, 1002 & 1004) on the occasion of his expedition to the
Angi lakes, in the Sjari region.
From the third ridge on leaving the Soedomi River, we looked down on
to the Momi again, and descended to it over secondary slopes overgrown with
Rnbus roscefolius, the fruit dirty-red in colour, hard, and like a small rasp-
berry in shape, just as insipid but very different in appearance to the large
scarlet, strawberry-like fruit of the same species in the uplands of British
N. Borneo. A boggy bamboo-thicket lined the bed of the river, which we
crossed, to camp for the third night on the other side in an old "kebun."
Whole families of the hill people came down to visit us, even with babies in
arms, each party, after wandering round gloating over the various sights of
the camp, building its own shelter, to which they retired to cook their meal
and spend the night. I distributed tobacco to the men, women, and bigger'
children, and rice to the babies, of which the very tiniest demanded its quota.
Some of these people came on with us to the lakes, as others had done from
Serao a source of considerable relief to the coast carriers, many of whom
were getting tired from the steady climbing. These mountain people are
splendid carriers, but it is next to impossible to get them to go down to the
coast.
The next day, proceeding through secondary forest up a lateral spur, old
plantations opened out at 5000'. At about 6000' a couple of bushes of the
copper-coloured Rhododendron Icetum, red when older, one of the glories of
the Arfak, heralded the approach of the mountain collecting-grounds, which
I alone intended to work. The korano of Wariap and his grandson Waspiri
pointed out, on the southern flanks of the spur near the valley below, two
houses of the " Orang Jatoe," or bad people, noted head-hunters according
to my informants, Waspiri adding that the victims were decapitated at the
house observed from the Serao ridge, the resulting trophies being brought up
here. With glasses the people could be seen standing on a rise near by, while
in front of the house, in a cleared space, twelve men were sitting, in two
rows of six each, singing some barbaric chant, accompanying their song of
defiance with reiterative movements of arms and legs, in Polynesian fashion.
Certainly, unlike all the other mountain people, they did not attempt to
21
approach our party, which was, of course, much too large to invite any form
of attack.
Higher up, a large solitary house inhabited by very friendly people marked
the limit of the inhabited zone. Immediately above this the path enters
true " rimbu," l then strikes the track along part of the main range, which,
running approximately N. to S.W., is clothed in low mountain forest, the
plant-formation common to the crest of the main range and the lake basins.
(d) Low MOUNTAIN FOREST FORMATION ABOVE 7000'.
From 7000' low mountain forest prevails up to the lakes, and systematic
collection was limited to this formation.
Advancing due south, with increase of altitude the crest of theS.W. ridge
soon becomes narrower and the trees smaller, showing a gradual transition
from mossless forest to an intermediate mossy forest with a fine variety of
mosses and hepatics ; while on higher points true mossy forest prevailed,
but very limited in incidence and species. Coniferous trees, Phyllocladus,
Podocarpus, Uacrydium, and Libocedrus, were conspicuous on this ridge, the
facies of the vegetation being strongly reminiscent of that of the ridges of
Kinabalu at the same altitude, but suggesting a wider and more continental
origin.
Small open rest- and camping-places from time to time gave vantage
points of observation, where the ground was always bright with clumps of a
brilliant orange Dendrobium and the pretty mauve Burmannia disticha :
otherwise, white and yellow predominated amongst the shrubs and trees of
the mountain forest.
Other open spaces were caused by small landslips of the loose granite-
gravel soil, on which no rock was seen exposed. In one place quite a large
part of the crest had slipped away, leaving a steep wall about 100' high,
difficult to scramble up, as the gravel gave beneath the feet. Probably the
higher points of the ridge, now overgrown with roots of trees, are due to this
agency. These open spaces afforded beautiful views over the Sjari ridge
with its tiny trails of smoke, to the blue waters of the Bay in the distance on
the east, and limited on the west by the densely-wooded slopes of the Tilaan
valley.
That night, the fourth after leaving Wariap, we camped on the highest
part of this ridge, on an open space above a mountain-torrent. I was
awakened in the night by a wild stamping of feet, accompanied by a
stentorian chant of powerful voices in unison, taken up by each Papuan
party in turn. The " Papuas," as they are always called, rigidly kept to
their separate clans, each putting up its own shelter. This somewhat
alarming incident proved to be a primitive method of keeping warm.
1 Virgin forest.
fiven on the march when resting for meals the Papuans invariably sorted
themselves into their own family groups. Notwithstanding the heterogeneous
nature of the party over forty carriers, men, women, and children, the men
in most cases accompanied by their wives and other children, with the hill
people joined on, I never heard any complaint from the Serzan nor a
discordant note or quarrel.
No doubt the presence of my staunch friends Manao and Waspiri, both
very fine men, and that of old Basi, the korano of another "campong," a
fund of quaint humour and good temper, who had been up several times
before, had a great deal lo do with this result. There were, however, many
outside elements over which their control \>as not acknowledged; but the
absence of bad temper and quarrelsomeness is always a distinguishing
feature of primitive people who have been spared contact with our so-called
civilization.
The next morning, after some hours' progress along the crest, we emerged
from the small forest into a lower scrub, to look down on to the brilliant blue
waters of a lovely lake, surrounded by slopes and ridges wooded to the water's
edge as they ran into the lake, intersected by the white beaches of many bays
of varying size and outline. This lake lies in a trough between the ridge we
were on and Koebre Mountain, of which the bare summit with a few scattered
trees limits the sky-line west, as it rises straight from the water's edge, except
for a little cultivated land where the lew Alfuero houses were dotted along
the shores. One or two moving spots on the water showed that these people
were observing us from the frail rafts made of three palm-stems tied together,
which are their only means of transport (PL 1. fig. 2).
On the eastern side at one's feet the prevailing forest runs down to the
water's edge where the banks are steep (PL 2. fig. 4), but where the slopes
are gradual it is replaced by an open marsh bordering this side of the lake, on
which some long spinneys and an isolated forest-patch intrude (PL 2. fig. 3).
Intersected about the centre by a few forest-clad lateral spurs from the
S.W. ridge, the marsh sweeps round to the north, where a break in the hills
marks the exit of the Tilaan River. This northern portion was not touched
by me in the course of this work.
Angi Lakes.
Turning abruptly to the right, we descended steeply over roots and
trunks of trees through a sheltered mossy forest, then splashing over logs
and bog on to the marsh. Here it was open and easy walking where water
streams over the coarse quartz sand, bright with Riedelicu, Dendrobiums,
and Rhododendrons, but impassible where boggy and covered with fern and
sedge.
The marsh is not a natural association, but it is kept open by the Alfueros,
who were busy burning it off during our stay ; a practice which no doubt
23
facilitates drainage, as the small rivulets, issuing from the slopes, which
stream over the marsh to drain into the lake, would form a water-logged
area under forest conditions, impeding access to and from the lake. This
is no doubt what the Papuans imply when they explain the burning by
saying that it keeps the ground u panas" or dry. But more important still
is the fact that the houses on the opposite side command the whole of this
cleared area, so that the arrival of strangers can be controlled and warning
received in case of hostile attack.
The practice of burning areas for observation or otherwise must always
have been prevalent in the Arfak, as Forrest, in 1705 (1, 108), whose stay at
Dorei Bay coincided with a very dry January, saw from there " many great
fires on the mountains of Arfak."
Passing over the central portion of the marsh, we made our way to some
rising ground, where the camp was pitched in front of an isolated forest
patch facing the lake, from which it was separated by a muddy tract, where
Juncus lampocarpus predominated. This tract soon became a morass from
the constant Papuan traffic to and from the lake, where a long white beach
formed the landing-stage of the native rafts.
The day after our arrival the Alfueros streamed over to visit us, accom-
panied as usual by wives, children, and babies. Wearing no clothes, many
were plastered over with some black pigment, possibly for greater warmth.
They brought to trade "ubi," corn-cobs, tobacco, and splendid potatoes. The
latter, grown on the upper lake, were a most welcome delicacy after unlimited
sweet potatoes, and as much appreciated by the Malays and Papuans as by
myself. My people traded everything that could be scraped together for
them and for the tobacco, which was said to be of very good quality. The
" Pradjoerit" and " Orang rante" exchanged their salt rations and matches,
relying with touching faith on my supplies, even parting with the buttons on
their uniforms. The Papuans traded their rice and sago rations, to return
to the coast exhausted as a result of an " ubi " diet. St. Vrsiz (15, 235)
mentions potatoes as doing well at Hatam, where they had been introduced
for twenty years through the Missionary Woelders from Andai. The
tobacco was carried at the top of long poles, rolled into large pointed ellipses,
which looked like clubs.
In 1857 (8, 75), when the northern part of the Arfak was visited by
members of the * Etna ' Expedition, they were told that tobacco was not
grown on the north side but on the east, and that it was distributed from
Hatam to Amberbaki, Dorei, and the south-west coast of Geelvink Bay.
That it should be easiest to bring this appreciated article three days' journey
down from Hatam to Andai, and then by " prau " along the coast, in prefer-
ence to the two to five days' journey from the Angi lakes, proves how little
intercourse there is between the Hill and Coast tribes, and also what a
natural boundary the inundation "korang" belt forms.
24
The korano of Koebre was quite a superior man, a blood-brother of
Manao's, who brought him up to me, when he presented me with splendid
potatoes and corn-cobs, and I gave him knives and a "kain" J in return.
It was interesting to note how the character of the surrounding forest,
even in such a small area as this lake-basin, varied with the exposure. The
eastern slopes were characterized by mossy forest, while to the south-e;ist
Araucaria Beccarii predominate.!, gregarious and in groups, to near the
water's edge (PL 1. fig. 1). To the north and north-east the forest was not so
homogeneous, older Libocedrus arfakensis and Podocarpus papuanus, with the
graceful palm Kentia Gibbsiana, standing out above the mass level, both on
the slopes and the ridge. On the western slopes of Koebre it was much drier
in type.
The most fertile part was the isolated patch of intermediate mossy forest
behind our camp, which reminded me of Fiji in its beautiful moss-flora and
wealth of creeping orchids. The possibilities of this patch, though continually
worked through, seemed inexhaustible.
Accompanied by two of the " Pradjoerit," Manao and his friend the
Alfuero korano, with the latter's two delightful boys, most keen to help in
collecting and looking for plants, I spent a day on Koebre. We crossed the
lake on two of the rafts tied together, following Dr. Gjellerup's advice. It
was a very tedious journey, taking about an hour and a half; while coining
back in the dark, with stormy gusts of wind and rain, we spent about two
hours in crossing. The two rafts, attached by a rotan-tie at each end, worked
against each other as the waves splashed up between.
The summit of Koebre is a bare, open, lichen-covered plateau, of which
the wind-swept character is revealed in the shrnbs, either prostrate and
spreading on the ground or of clipped, erect, and compact habit. A few
single trees which have survived the fires to which the open character of
this summit is due, dot the surface, while in gullies and depressions small
trees are crowded into shrubberies surrounded by a ring of burnt wood.
It was amazing to see solitary grotesque Myrmedomas, over a metre high,
also recorded by van Ooster/ee (17, 1008) plants of such size, to say nothing
of the terrestrial habit, being quite unknown to me (PL 3. fig. G). The
same may be said of an extraordinary Hydnophytutn^ just like a collection of
pipes standing upright on the ground, each pipe representing a hollow stem,
about one dm. across, bearing flowering branches round the rim. A couple
of small isolated trees of Dacrydium novo-gumeense bore an abundance of
small red cones.
From the summit there is a splendid view over the smaller, or " Man "
(<$ ), lake, beautiful in outline, with much cultivation round its shores, of
which the upper slopes are much more densely wooded and the lower more
thickly inhabited than the "Woman" (?), especially towards the south,
1 Cloth.
25
where the excellent potatoes are grown. To the north the surrounding hills
slope on both sides to a marshy area, which marks the exit of a river, as on
the lower lake.
To the south-west the houses of another head-hunting tribe were pointed
out, who seem to dominate this part of the country, as those before men-
tioned cause a reign of terror at lower altitudes. These people possibly
represent the tribe called " Hiraj," about which St. Vraz (15, 234) was told
at Hatam thnt they lived beyond the lake Tschemti, and were spoken of
as cannibals ; but I never heard any of these " Orang jatoe " referred to as
anything but head-hunters by my informants, Manao and Waspiri.
Our stay at the <$ lake, much as I should have liked to prolong it,
was limited to six days. The camp, never very dry at the best of times,
became sodden and under water from so much trampling, and many of the
Wariap people, who had faithfully kept their promise to stay with me, were
suffering from bad colds and rheumatism, such a sudden change of tempera-
ture necessarily telling on people accustomed to tropical heat, when warm
clothing cannot be provided.
The Malays of my escort were also becoming depressed, as they always
do when out of their accustomed environment, so the order to break camp
was hailed as a happy deliverance by everybody but myself. The beauty of
these surroundings, their extraordinary variety, afforded such a wealth of
material for work and reflection, which, with crisp air, cool temperature, and
splendid weather, made one long to spend more time in this lovely spot.
Our return was along the S.W. ridge again, leaving on the right
the spur by which we had ascended. Beyond this point a fine high forest
developed as the altitude decreased, the huge climbing ferns still wreathing
the trunks of the much finer trees, the undergrowth showing less herbaceous
variety with more sub-staging of shrubs and young trees. A small clump
of Corsia arfakensis, a new species, grew on some dead wood, and at
about 5000' a group of two or three magnificent Agatliis Dammara, with
towering white steins, too large to climb, and very small crowns, occurred.
Great lumps of white dammar stood out on the trunks, which the
" Pradjoerit," to whom these trees were familiar in Amboina, immediately
wanted to fire, so like the wasteful Fijians with their beautiful " Dakua "
trees, which I sternly forbade. This practice, I was glad to see, seemed
unknown to the Papuans.
Many young trees showing the fastigiate youth form were seen, and one
of the Papuans found me a young seedling, about 2 m. high, from which I
took the foliage, but had to content myself, so far as fruiting material went,
with some old scales found beneath the old trees ; they were sufficient,
however, to determine the species.
Continuing down a lateral spur, running in an easterly direction, we
26
passed through a somewhat dense undergrowth of a small Licuala sp., where
the pretty white Medinilla arfakensis, almost a small tree, a very handsome
Bulbophyllum covering a prostrate log, and the climbing Diclirotricliium
brevipes, another of Beccari's Hatam records, were also growing. From an
open space we caught a glimpse of the buttress of the ridge we had descended
from, with the glorious Agathis trees towering far above the rest of the
forest.
Farther on, at another unusually large cleared resting-space. vaulted
over by trees, we came upon some horizontal sticks, resting on a couple of
forked supports stuck in the ground, on which a number of little forked
prongs were arranged in groups. Waspiri exclaimed when he saw this
peculiar arrangement, explaining that it was the practice of the Coast people
to have these places, which show what parties have recently been in the
mountains, the arrangement of tho little prongs indicating how many and
whether women or children, if the parties had returned, or if any of their
members had been killed. On this occasion it was m:ide out that a man and
woman, missing from Wariap, had been killed in the hills.
Finally, always working east, we passed through the bamboo clumps,
which herald cultivation, on to a large plantation, with a solitary house in
the centre. Here an old man showed us the way down to the Momi River,
an impossibly steep track, up which, considering the way it was worn, the
people of the house must fetch all their water every day.
From this plantation a view up to the Serao ridge showed the " kebun,"
in which we had camped on the second night, and also the house of the
head-hunters on the lower slopes.
Camping by the river that night we reached Wariap next day through
the " korang " forest, joining our old tract later, just before the Pandanus
group.
At Wariap my first inquiry was for the ' Valk/ as the Commander had
most kindly promised to call in there on his return from the Mamberamo
River, on the chance of our being there. As there was no news I decided
not to wait in tliat sand-fly stricken spot, but, giving the men two days' rest,
to return by the beach to Manokoeari, which everybody at Manokoeari, even
Mr. van Oosterzee, had told me was quite impossible. The iSerzan, however,
after searching inquiries, found that this route was feasible and well-known
to the Wariap people, taking four days. Two " praus" from Wariap carried
kit and provisions, landing at night at the native camping-places, to which it
is always wisest to keep, though somewhat long, as they are determined by
good water and landing-stages.
I had hoped to be in time for the December boat to Java, but on crossing
in u praus " from Andai to Manokoeari, once past the mouth of the estuary,
we could see the smoke of the steamer across the bay. Watching intently
which direction she took, the Serzan exclaimed, " Poelang ! " (Home). This
27
entailed a month's wait at Manokoeari for the next boat ; I put in the time
working about Dorei Bay, as Dr. Grjellerup told me it had not been collected
over. As the next boat called at Humboldt Bay, the limit of the Dutch
possessions, which is only visited every other month, I was able to take that
trip as well, collecting at each stopping-place, with very good results.
The coast-collections proved very interesting, but phytogeographically
so distinct from the Arfak plants, no two species proving common to both
regions, that they have been separately enumerated.
PLANT ASSOCIATIONS OF LOW MOUNTAIN FOREST
FORMATION.
A. FOREST ASSOCIATIONS.
[Endemic species are marked , and those of wider distribution *.]
a. S.W. RIDGE.
1 a. Mossless Forest.
On the main range, or S.W. ridge, at 7000', a mossless forest asso-
ciation prevails, of slender straight trees about 13-] 6 m. high, with a very
open undergrowth of chiefly herbaceous plants.
Undergrowth. Alpinia domatifera, 1-2 m., always in appreciable colonies
of one height, the flowers varying from white to red with red fruit, and
A. arfakensis var. subsessilis, with pink flowers and white fruit, were
dominant, more or less covering the open ground.
Liancs. * Gleichenia linearis, spread over supports up to 7 m., while
the trunks of the trees were wreathed in the climbing ferns *Nephrolepis
acuminata and P^lybotrya arfakensis, from base to branches, the long fronds
standing out radially from the stems. Freycinetia Gibbsece, with very hand-
some red sheaths, hung bunched from the trees or spread in thick masses
underneath, and F. flaviceps, with yellow fruit, was more slender in habit.
Trees. A group of Quercus Lauterbachii, the ground underneath strewn
with magnificent acorns of all sizes, some of those collected having proved
the largest known, represented a family recorded by D'Albertis (9, 69),
Beccari (12, i. 177), and St. Vraz (15, 33) from Hatam. *Podocarpus
Rumphii, recorded by Beccari, but not seen in fruit, was abundant ; likewise
Phyllodadus kypophyttvSj the Kinabalu species, and Podocarpus papuanus,
recorded as P. imbricatus (which it very much resembles, the seedling form
being indistinguishable) by Beccari from Hatam, and since found by Kloss
on Mt. Carstensz. Advancing due south, as the crest of the ridge narrows,
a gradual transition to an intermediate mossy forest of smaller trees with
branched stems and denser crowns, the trunks and buses covered with small
hepatics and mosses, takes place as the altitude increases.
2 a, /3. Intermediate Mossy Forest.
Undergrowth. Sphagnum novo-guineense, with *Rhacopilum spectabile
and the magnificent and abundant Dawsonia gigantea, ull in fruit, with the
creeping *Lycopodiutn cernuum, formed part of the prevailing moss-carpet,
from which rose the orchids Platantliera elliptica, vars. longicalcarata and
elatior, green in colour and varying in size, gregarious and general in this
association ; Cryptostylis arfakensis, with red lip and green perianth, and
Bulbophyllum muricatum, with large red-spotted yellow flowers, were found
in single examples. Clumps of Alpinia domatifera (dwarfed), *Gahnia
psittacorum, previously known only from Australia and Tasmania, * Histiopteris
incisa, *Dipteris conjugata, gregarious as usual where more open, with the
small shrubs Diplycosia Lilianea?, about "25 m. high, with rigid branches and
striking white-tipped red flowers, Vaccinium pilosifiorum with pretty, very
hairy rose-pink corollas, also seen as an epiphyte, and V. leptospermoides,
with red-pink flowers, were often grouped together at the base and between
the dwarfed trees.
Tiny tufts of Gentiana Vanderwateri, with large white flowers, showed
up in damp places and, where drier and the small trees opened out, the
minute Lobelia arfakensis spread its large patches on the ground, dotted with
white flowers, while innumerable seedlings of all the conifers previously
mentioned, with Dacrydium and Libocedrus, formed the most general and
conspicuous part of the undergrowth, including the slender tree-ferns
Alsopldla arfakensis, with stems 1 dm. through and about 1 m. high, the
fronds 1 m. long, and Cyathea arfakensis not much larger.
Epiphytes. The small ferns collected on the moss-grown trunks of
the trees were * Trichomanes palmattfidum, Hymenopliyllum cincinnatum,
*Lindsaya hymenopliylloides, Poly podium remigerum, *P. stenopliyllum,
*P. clavifer, and the minute yellow orchids, Octarrhena cylindrica, with
Dendrobium glauco-viride (magenta), Phreatia spatlmlata (white), and Piper
arfakianum.
Lianes. * Gleiclienia linearis and *G. volubile abounded, with Freycinetia
GibbsecK, much less luxuriant, F. ftaviceps, and Calamus arjakianus ; a slender
bamboo, identical in habit and appearance with the Kinabalu plant, but not
seen in flower, and Lyonsia albifiora were also pretty general.
Trees. Dacrydium novo-guineense with Libocedrus arfakensis, Podo-
carpus papuanus, *P. Rumphii, and * Phyllodadus hypopliyllus wore dominant,
as the profusion of seedlings testified ; but a great variety of other small
trees were associated, especially towards the southern portion of the ridge,
where it broadens out again and is consequently more sheltered Urimys
arfakensis, with a dense round crown, bore its small, white, later pink flowers
on pendent pedicels ; Spirceanthemum bullatum, *13ceckea frutescens ; Back-
housia arfakensis, flat-topped like a Leptospermum, with very small coriaceous
leaves, and smothered in golden-orange flowers. A Psyclwtria sp., resembling
P. sarmentosa, with white panicles of flowers, Idenburyia arfakensis repre-
senting a new natural order, Myrtus flavida var. glabrescens, a glabrous form
of the Kinabalu specie?, Jambosa arfakensis, with small thick leaves and
white flowers, Palmervandenbroefcia papuana, an interesting new genus in
Araliacese, and Timonius filipes, were in full flower.
3. Mossy Forest.
The occurrence of this sterile type of mossy forest, as distinct from the
intermediate form, was limited to the highest points of the ridge, where
the prostrate and erect trunks of the small stunted trees with the ground
between were swathed in long moss, which, as usual in this type of associa-
tion, stands out straight from its supports, rigidly turgid and generally in
vegetative condition.
*Schizcea malaccana, *Gahnia psittacorum, 1 m. high, Halorrhagis suf-
fruticosa, Nepenthes maxima v. l>revi folia of the exposed peaks of the serpentine ridges of Kinabalu.
* Lycopodium cernuum, * Gleichenia vulcanica, * Dip/ten* conjugata, grew
densely, also * Spathoglottis aurea and *Dianella ccerulea.
At another more recent slip, Gleichenia vulcanica with *Lycopodium
cernuum were associated with *Gahnia psittacorum, 1'50 in., * Burmannia
disticha, the woody herbaceous llalorrhagis suffruticosa, the single shoots with
verticillate spinous leaves terminated by a panicle of red flowers, Oldenlandia
nutans, of similar habit, but quite herbaceous, with white flowers ; Myrtus
arfakensis, as a small spreading shrub, and the prostrate M. koebrensis,
with reddish foliage and stems, both with yellow flowers, spread over the
easier gradients.
b. MARSH BY ? LAKE.
Sand Pans with running water.
In certain areas of the marsh, more towards the centre, where coarse
quartz-sand formed a solid surface, intersected by shallow streams or bathed
in films of streaming water, many small herbaceous plants grew spaced
between the larger clumps of more showy species.
Herbaceous Plants. The little *Schizcea malaccana witli the cosmopolitan
* Lycopodium carolinianum, creeping tightly to the ground, and *L. cernuum,
P2
36
the minute *Bulbostylis capillaris var. trifida, Centrolepis novo-guineensis in
tufts, the first record of the genus in New Guinea, *Xyris pauciftora,
Eriocaulon leucogenes, very variable in size, * llurmannia disticha, up to 1 m.,
one or two examples of the little white Spiranthes papuana ?, *Polygonum
alatum, the small yellow *Utricularia bifida, and the mauve *U.racemosa
from a few cm. to 2 dm. in height, both new records for New Guinea,
dotted the surface ; while abundant patches of the yellow and red RiedeHa
montana var. puberula, also var. arfakensis, cream and dark red, both about
50 m. high, with the beautiful terrestrial orchids, Dendrobium fruticicola,
the foliage '50 m., above which the brilliant orange flowers, red when older,
rose to '30 m., D. latifrons, also yellow, and the splendid D. rhomboglossum,
1 m. high with large convex flowers, magenta-pink outside, white inside, borne
the whole length of the rhachis, made glorious patches of colour, blended
with small slender plants of Halorrhagis su/ruticosa, Oldenlandia nutans, and
Coleus Gibbsece, white to mauve, all of similar habit and dominant on the
marsh in all situations ; young plants of Nepenthes maxima var. nana, the
mature trailing over any support, with *Lycopodium divaricatum abounded.
Shrubs. As the above plants eased oft' and a ranker growth inhibited
close burning, shrubs occurred in small groups, chiefly *Bceckea frutescens,
Rhododendron Devriesianum, of which the magnificent white flowers, turning-
pink later and fragrant at night, are about 1 dm. across, the whole inflor-
escence being 10 dm. in diameter, R. Icetum, the little profuse flowering
R. Vonroemeri, and the bright red R. Gibbsece, all further emphasize the
wonderful colour-scheme of the more open portions of this interesting marsh
association.
Denser Sedge Growth in Boggy Areas.
Boggy areas with deep ditches, standing water, or larger streams bore an
impenetrable sedge-growth which included *Dryopteris Beddomei, *Lyco-
podium divaricatum, * Cladium falcatum and *C. germanicum, *Sdrpus
setaeeus, Trisetum latifolium, Ischcvmum aristatum var. arfakensis, *Juncus
lampocarpus, Phaius Tankervillece var. papuanus, the flowers brown, white
inside, Halorrhagis suffruticosa, Oldenlandia nutans, Coleus Gibbsece, and
* Emilia sonchifolia, all drawn up to the level of the surrounding sedge.
In other boggy areas sloping down to the lake, generally under water,
*Juncus lampocarpus predominated, easing off with * ' Carex Gaudichaudiana
to the sand-pan association.
Edging the lake, limiting the white coarse quartz-sand beach, a higher
and drier sandy bank, well bound together by the roots of the sedges, and
broken in parts where small streams ran into the lake, carried *Marchantia
polymorpha, * Lycopodium divaricatum and *L. complanatum, Cladium arfak-
ense and *C. germanicum, * Carex Gaudichaudiana, *Gahnia psittacorum,
* Rhynchospora aurea and *R. glauca, Trisetwn latifolium, * Dianella
37
ccerulea, Riedelia orchioides, '75 in. high with red perianth-segments and
red-tipped lip, replaced, where the sandy ground was more exposed, by
the creeping *Isachne miliacea, * Centella asiatica, *Gna sorbifolia, *Gleichenia
vulcanica, and Oleandra cuspidata were common all round, the scrambling
*Gleichenia Icevigata abundant in the open to under the trees, *G. glauca up
to 6 m. \\ith *G. linearis, *Cladium falcatum, * Gahnia psittacorum up to
2 m., Halorrhagis sujfruticosa, Uldenlandia nutans, and Coleus Gibbsece, all
drawn up to the level of their environment, with the shrubs Medinilla
Forbesii, * Mclast oma malabathricum var. adpressum, the lanky single shoots
of young plants of Poikilogyne arfakensis, Rhododendron Devriesianum,
It. Ict'tum, and the pink R. undulaticalyx. The gregarious Sccevola Lauter-
hachU, with long semi-scandent shoots, shining leaves, and for the genus
large yellow flowers, known hitherto from N.E. New Guinea, dominated in
the tangled mass of fern and sedge.
c. OPEN SUMMIT OF KOEBRK MOUNTAIN, 9000'.
Cladonia Association.
The open summit of Koebre forms a flat plateau with a hard surface of
disintegrated quartz-granite on which quartz-gravel lies so thickly that it
gives quite a white effect. Where the rock is exposed it snows as large a
proportion of quartz-veins as of granite.
Herbaceous plants. On this open plateau, for which systematic burning
is again responsible, a remarkable association of Cladonia spp. prevailed,
spreading over the whole area as a uniform grey carpet, about 3 cm. high,
composed of * Cladonia verticillata, *C. didyma, and *C. cocci/era, displaced
38
in certain parts by stretches of short *Pteridium aquiliniini var. lanuginosum
or *Gleiclienia vulcanica, and in damper places dense mats of C&ilrolepis
novo-guineensis.
Associated with the Cladonla were clumps of Riedelia montana var.
arfakeusis, and colonies of * Burmannia disticha and * Gahnia psittacorum,
the latter dwarfed to '50 in. high. Patersonia novo-guineensis in full flower,
the gleaming white or pale mauve corollas well exposed, was dotted in tufts
all along the summit, a most interesting first record of a genus known from
Kinabalu, Mt. Halcon in the Philippines, and Australia.
A small example of the gi'eenPlatanthera elliptica was a remarkable find,
the species alone being previously known from S.W. New Guine;i, while
the two new varieties, so abundant in the moss-grown forest of the S.W.
ridge and the small high mossy forest-patch by the lower l:ike, were not seen
on the slopes of Koebre. Patches of the familiar D. rhomboglossum and
D. fruticicola, both as conspicuous as on the marsh, with the magenta
D. infractum, represented the terrestrial orchids in flower, with Glomera
Gibbsece epiphytic on the upper branches of a wind-swept shrub.
Didiscus koebrensis with many radiating stolons, near a species already
collected on Mt. Scratch ley in the S.E. supposed to bo a variety of
I), sanicitlce/olius of Kinabalu and Mt. Halcon. Oldenlandia nutans var.
alpinum was dwarfed to '50 m. ]\]yrmedoi>ia arfakiana (PI. 3- fig. 6) with
its slate-blue flowers the size of a shilling, showing all round the formless
bristly steins, of which the fleshy consistency no doubt resists the fires, grew
in solitary examples, one of which, about a metre in height, sent out a metre-
long shoot from the apex at right angles to the main stem.
Shrubs. The exposed and wind-swept character of this summit plateau
was shown in the shrubs, which either spread prostrate over the surface or
were clipped back into small compact shapes, Hibbertia novo-guineensis, a
plant with large flowers and spreading habit and very near to H. scandens
of E. Australia, is the first species in this genus, hitherto known from
A-Ustralia and New Caledonia, to be described for Malaya or New Guinea.
The ever-present *J3ceckea frutescens took on a prostrate form, Acronyclna
arfakensis, with insignificant white flowers and reddish fruit, was either pros-
trate or erect, while Myrtus prostrata, with reddish stems and small coriaceous
leaves, had developed quite dorsiventral shoots, resting on the ground.
Small erect shrubs were *Bensloivia umbellata with yellow foliage,
Diplycosia Liltanete, and Vaccinium villosiftorum, while the dark green Tel-
mmt/iodia rotundifolia, the typical Styirtielia Gjellerupii, with white flowers
and pink berries, and another Styplielia sp., of which the material was not
sufficiently complete for identification, Rhododendron angiense, Sericolea
novo-guineensis with Psycliotria vaccinioides, were of clipped compact habit.
A couple of small trees of Dacrydium novo-guineense dwarfed to about
4 111., with stiff, Araucaroid, ascending branches, bore many tiny ripe cones,
39
glinting red through the dark green foliage, a fortunate fact, deciding the
genus of a very prevalent species of conifer, of which it was certain that the
abundant seedlings in various growth-forms could only represent a new
Dacrydium sp., but the search for fruiting specimens at lower altitudes
proved unsuccessful.
In reference to this exposed lichen-covered plateau a quotation from
Lorenz (22, viii. (1909) 178) in "Nova Guinea" suggests the presence of a
similar association on the Charles Louis Mountains, while Mr. Stroeve of the
Dutch Navy, whom I had the pleasure of meeting at Manokoeari after my
return from the Arfak, described what seemed a very similar formation as
occurring on the mountains to the south of Geelvink Bay at about the same
altitude, where not only the practice of burning obtains but potatoes are also
grown. I heard later with great regret of the untimely fate of this promising
young officer, who, after most successful exploration and surveying work
on the Ruffaier River, was treacherously shot by arrow by Papuans on the
Waroza River, near the coast to the east of Geelvink Bay (26, 782).
PHYTOGEOGRAPHICAL CONCLUSIONS.
These may be summarized as follows :
(1) New Guinea, the centre of distribution for many so-called
Polynesian, Australian, and to a lesser extent Malayan types, of
which the Papuan species are not only older in type, but also
show extraordinarily pronounced specific differentiation.
(2) Wide distribution in New Guinea of endemic Mountain types.
(3) Low Mountain Forest formation approximates to the Ridge forma-
tion of Mt. Kinabalu and the Philippines.
(4) Mountain Forest Flora of the Arfak Mountains possibly represents
the oldest Papuan type.
(5) Pteridophytic and Bryophytic Floras are more Polynesian than
Malayan in incidence and luxuriance.
(6) Open " Opportunity " Plant Associations offer further proof of the
autochthonous character of the Papuan Flora.
1. NEW GUINEA THE CENTRE OF DISTRIBUTION FOR MANY SO-CALLED
MALAYAN, POLYNESIAN, AND AUSTRALIAN TYPES.
Evidence of New Guinea as the centre of distribution for many plant-
types, so far considered Polynesian or Australian, has been strikingly
confirmed on the present occasion. Emphasized by the well-known botanists
who have worked out the results, such evidence has been a marked feature
of recent Dutch and German exploration, and was also shown in Kloss's
Mt. Oartensz collections 1 .
1 Ridley, II. N., " Report on the Botany of the Wollaston Expedition to Dutch New
Guinea," Trans. Linn. Soc., ser. 2, Bot. ix. (1916) 1-269, pis. i.-vi.
40
Geological support for this theory is quoted by Wallace (7, 443) in
' Island Life,' who points out that the " 1000 fathom line, which indicates
the land area which would be produced if the sea-bottom were elevated 6000',
extends in a broad mass westwards, then sending out two great arms, one
reaching to beyond Lord Howe's Island, while the other stretches over
Norfolk Island to the great barrier reef, thus forming a connection between
Tropical Australia and New Guinea."
Sieberg 1 explains the same configuration in greater detail as follows :
" Neuguinea und der Bismarckarchipel gehoren nach E. Suess dem inner-
sten unter den Bogen junger Faltengebirge an. die vom Osten her sich
gegen die alte starre Festlandstafel Australien anlegen und die in Neu-
seeland zusatnmentreffen. Dieser innerste australishe Bogen setzt sich aus
zwei annahernd parallelen Teilstiicken zu?ammen, die beide in die Nord-
nordwesthalbinsel Neuseelands miiriden. Die Nordwestlich streichenden
Ketten holier Faltengebirge, die das innere Neuguineas der Lange nach
durchziehen (Bismarck und Kratkegebirge, Viktor-Emanuel, (Charles-Louis
Gebirge) reichen untermeerisch noch weithin und ragen in ihren hochsten
Spitzen als die Inselgruppe der Louisiaden und Neukaledonien ernpor ; der
ostliohe parallelbogen setzt sich zusammen .aus Neumecklenburg, den
Salomonen und Neuen Hebriden."
Of the preponderating influence of the New Guinea Flora, when suffi-
ciently well-known, we have an instance in the Orchidacese, a family which
in the N.E. has received the expert attention of Schlechter (23), the
well-known authority and collector. Dr. J. J. Smith, of Buitenzorg, an
equally keen expert in this family, has not only described nearly all the
orchids of the N.W. and S.W. so far known, but has also consistently
enjoined on collectors in the Dutch possessions to devote particular atten-
tion to these plants, so that from both these sources we can form some idea
of the general incidence and distribution of tin's one order throughout the
country.
Such is the wealth of material in a single instance in this little-explored
island that Schlechter, whose unrivalled experience in N.E. New Guinea
has spread over a number of years, summarizes his results in orchids alone
as 116 genera with 1450 species, of which 1102 are new (23, i. 14, xiii).
He considers that nowhere in the world is the Orchid flora so rich, quoting
2600 species as being known from there. Orchids known from Australia
and common to New Guinea he (23, i. 14, xix) looks upon as branches of
the Malayan-Papuan flora in Australia, the Orchid flora of tropical Australia
having developed under strong Malayan-Papuan influence, but it has had
little influence on the Papuan flora. This remark, to those who have
1 Sieberg, A., " Die Ei'dbebentatigkeit in Deutsch-Neuguinea (Kaiser- Wilhelms-land und
Bismarckarchipel)," Peterm. Mitth. Ivi. pt. 2 (1910) 118.
41
worked over the two areas, succinctly summarizes the whole relation of
both floras.
This Malay-Papuan influence is the determining factor extending to the
South Sea Islands, while the Moluccas show Malay with Philippine ground-
types and a strong Papuan influence (23, i. 14, xx), and Schlechter describes
New Guinea as the centre of distribution of an endemic Orchid flora (" Aus-
gangscentrum einer eigener Orchideenflora ") (23, i. 14, xx) as evidenced
by Corysantltes, supposed to be Australian, but now truly Papuan, 13 species
being known from N.E. New Guinea alone. The presence of this genus in
Australia, New Caledonia, Samoa, Java, Philippines, and the Himalayas
suggests radiation from a Papuan centre of development (23, 14, xxiii).
This overwhelming Papuan influence is amply demonstrated in many
other families, which not only show an actual numerical predominance in
species, but also a greater range of specific differentiation than is known
from elsewhere : for example, Libocedrus (4), Drimys (19), Myrtus (6) ; of
Pandanus and Freycinetia, I have never seen such a wide range in form ;
while in Rhododendron, Vaccinium, Styplielia, the numerical predominance
along with the great morphological range of form is unsurpassed in any
other region, except perhaps, in the case of the first, Central China.
Further interesting proof of the soundness of this point of view is afforded
by comparison of the Australian and New Guinea representatives of the
same genera, the Papuan forms showing decidedly the oldest types.
In the present collection perhaps Trimenia is the most striking case in
point. T. weinmannicefolia Seem., described in 1852 from Fiji, a dioecious
plant, remained the type of a supposed monotypic Polynesian genus, closely
allied to Piptocalyx Moorei, also dicecious, ranking as a monotypic Australian
one, till Ridley described T. papuana from Mt. Carstensz in 1916. T. arfak-
ensis is included in the present paper.
Two species of Trimenia are now known from New Guinea, both
hermaphrodite, while two new species in a closely allied new genus, Iden-
burgia, show a syncarpous bilocular ovary, proving not only that the dioecious
habit of the two isolated outliers of this order is probably derived, but also
that the systematic position of Trimenia and Piptocalyx in the apocarpous
Monimiacese is untenable, necessitating the new order Trimeniacese.
An equally convincing example is that of Pullea, a genus established by
Schlechter in Saxifragacese to include two plants with inferior ovary from
N.E. and N.W. New Guinea respectively, to which P. papuana is now
added, with a N. Queensland species, P. Stutzeri = CaUicoma Stutzeri
F. Muell., first distributed as Stutzeria by him, but afterwards included in
Callicoma. Pullea, therefore, now includes three distinct Papuan species
and one N. Australian, but, had the latter plant first been accorded proper
generic position, the former would have been cited as evidence of a wave of
Australian immigration into New Guinea.
42
Of the two Papuan species in BackJiousia, another of the so-called
Australian endemic genera, one was described by Ridley from Mt. Carstensz,
to which a second, one of the commonest trees in the Arfak, is now added.
Both have normal calyx-lobes, while the Australian species show a secondary
petaloid development of the lobes, evidently derived in character, though
previously considered a generic distinction.
The same case may be quoted for Didiscus, where the older types,
according to the Monographer, 1 showing a normal calyx, occur in Malaya
and New Guinea, with one species in N. Australia, whereas those with
aborted calyx-lobes, by far the most numerous, are known from the rest of
that continent, with one species from some Polynesian Islands.
Further evidence is also forthcoming in interesting new records of
genera not previously known from New Guinea, namely, Hibbertict, hitherto
considered Australian and New Caledonian ; Centrolejns, abundant at 7000'
and 9000', connects New Guinea with the open summit of Kinabalu in
N. Borneo at 13,000', Mt. Halcon in the Philippines at 7000', and S. China
on the one hand, and Australia and New Zealand on the other ; while the
genus Patersonia, supposed on inadequate knowledge to be endemic Australian,
shows the same distribution, with the exception of S. China, and now in-
cludes three very distinct Mahiyan mountain types.
In the case of species, Gahnia psittacorum, abundant everywhere in the
Arfak from 7000', is the first record for Malaya of a plant widely distributed
in E. Australia to Tasmania.
2. WIDE DISTRIBUTION IN NEW GUINEA OF ENDEMIC
MOUNTAIN TYPES.
In considering questions bearing on the phytogeography of New Guinea,
as a whole, it is important to take into consideration the fact that it is a
country of 786,000 km. in area (23, i. 14, i), most of which is mountainous in
character and undisturbed in condition. A huge region of vast ranges, in
which all intercourse between the relatively few and scattered inhabitants
is not only restricted by natural barriers, but also by the many different
languages, no two tribes having a common speech, even when living in
apparent proximity on the same range of mountains. Both these facts have
effectually debarred outside penetration into the mysterious back-country.
Though our knowledge of the plant-covering of these mountains is
extremely limited, such phytogeographical exploration so far accomplished
points, as would be expected under such virgin conditions, to the homogeneity
and stability of the flora as a whole. This fact is evinced in the pronounced
endemism and wide distribution throughout the whole country, further
1 Domin, K., " Monographie der Gattung Didiscus (DC.)." Sitz. Kon. bohm. Ges. d.
Wise. ii. Cl. (1908) 21.
43
convincing proofs of which are afforded in the present collection. The
following instances may he quoted.
All the mountains of New Guinea show a preponderating number of
species belonging to Rhododendron and Vaccinium in Ericaceae and Styphelia
in Epacridacese, though but few identical species have been so far recorded ;
but that this point is only a question of further investigation is proved by the
distribution of the very distinct R. Vonroemeri, which, most abundant in
the Arf'ak, is recorded for the ( Cyclops Mountains in the north, and from
the Hellwig Mountains and Mt. Carstensz in the south-west ( = 72. calceo-
larioides Wernham). Medinilla Forbesii, collected in the south-east and the
south-west, is now established for the north-west as well, while Timonius
filipes and Podocarpus papuanus, first collected on Mt. Carstensz, are
common on the Arfak at higher altitudes. Quercus Lauterbachii 1 and
/ SccBvola Lauterbachii,^ generally collected in the north-east, were not pre-
viously known from Dutch N.W. New Guinea.
In small herbaceous plants two new species of Didiscus link up the
Arfak with the Owen Stanley range in the south-east, while Eriocaulon
leucogenes, Trisetum latifolium, and Gentiana Vandencateri are common to
the former and Mt. Carstensz. Spirantlies papuana is now known from the
north-east and north-west, Platanthera elliptica from north and south-west.
New species in genera first recorded from New Guinea through Kloss's
Carstensz collections, such as Trimenia and Backhousia, are now proved to
be common to the Arfak as well ; also the genera Pullea and Sericolea,
the latter represented by six species in the north-east, one in the south-east,
three on Mt. Carstensz, and now by two in the Arfak; while Libocedrus,
known from the north-east, south-east, and south-west, and Dacrydium from
the north-east, south-west, and south-east, are abundantly represented by
one very distinct species each in the Arfak.
3. THE LOW MOUNTAIN FOREST FORMATION APPROXIMATES TO THE
RIDGE FORMATION OF MT. KIN ABA LU AND THE PHILIPPINES.
The low forest formation approximates very closely to that of the
serpentine ridges of Kinabalu from 7000', but with a larger proportion of
what I would have previously described as southern hemisphere types, but
prefer now to refer to as Papuan.
Identical plants so far recorded are Phyllocludus liypophyllus, Myrtus
jiavida var. ylabrescens, a glabrous variety of the Kinabalu plant, while
closely allied species occur in Dawsonia, Podocarpus, Ducrydium, Centrolepis,
Patersonia, Didiscus, Qentiana, and others.
1 Schumann, K., and Lauterbacli, K., ' Flora der deulachen Schutegebiete in der Siidsee,'
Leipzig, 1901. Naclitrage, 1905.
44
The prominence of orchids, Myrtacese, Rhododendrons, Styphelias, and
Vacciniums, in relation to other plants, is equally emphasized, but members
of the Araliacese, an old type of plant so prominent in the Arfak of New
Guinea, are, so far as we know, absent on the N. Bornean mountain.
From descriptions of the Philippine mountain-ridge vegetation it would
appear that the Papuan facies again predominates, thus approximating it to
that of the Arfak and Mt. Kinabalu, the absence of araliaceous types being
less marked, though of the typical P;ipuan genera Anomopanax is represented
by one species, and Kissodendron and Mackinlaya are absent. Phyllocladu*
hypophyllus is common to all, with allied Dacrydium and Podocarpus spp.
Glochidion Merriltii is also common to the Arfak, but a thinning out in the
number of Styphelia and Vaeciniuni spp. is apparent, while the Rhododendrons
show a great reduction in variety of type.
4. THE MOUNTAIN FOREST FLORA OF THE ARFAK MOUNTAINS
POSSIBLY REPRESENTS OLDEST PAPUAN TYPE.
It is in the Arfak alone, of all the Papuan mountain ranges of similar or
greater altitude, that two different localities have been worked over on three
separate occasions. This fact gives a wider field of comparison in relation
to this range. On comparing the general type of vegetation and the
generic and specific distinctness of the plants so far collected on this range
with those known from similar localities, one feels inclined to look upon
these granite mountains as carrying an older part of the Papuan flora.
Where so little is known, however, of the mountain flora of the country as
a whole, and collections remain limited to vertical sequence, it is impossible
to base such an impression on any actual fact, and the hypothesis is only
worth advancing as a possible consideration for future workers.
The contrast is very striking between the recent " korang " belt, from
which this range on the east rises almost without transition as abruptly as
the mountains on the north from the sea, and the well-defined limits and
deep soil of the foot-hills and lower ranges. The extreme homogeneity of
the mountain forest zone, the extraordinarily small incidence of outside or
immigration plants, combined with the relatively broad crests of the ridges
and the tremendously weathered condition of the main range, are all facts
which point to general stability in condition, spreading over a considerable
epoch of time.
The climate of the Arfak would also seem to be more favourable than on
other ranges, and the nature and structure of the plants are not so limited
by the edaphic and atmospheric factors which play such a large part in the
restriction of floral conditions on Kinabalu.
For instance, all the Arfak plants show coriaceous to very small leaves,
and the hairy covering or tomentum, common to many of the Kinabalu
45
plants, is quite absent a fact possibly to be attributed to the warm well-
drained soil of disintegrated granite and the even conditions of illumination
with less direct exposure to wind. The " Nebelwald " of the German
botanists, described as predominating on the mountains of the north-east,
with trees swathed in lichen, is absent, also the " Krlippelwald" association,
while the turgid vegetative form of mossy forest is limited in incidence.
But, of course, so far as the two former associations are concerned, the
lower altitude is a possible and limiting factor.
6. PTERIDOPHYTIC AND BRYOPHYTIC FLORAS ARE MORE POLYNESIAN
THAN MALAYAN IN INCIDENCE AND LUXURIANCE.
The Pteridophytes and Bryophytes of the Arfak proved exceedingly
varied and luxuriant in incidence, more Polynesian than Malayan in facies,
but endemic Papuan in type. In mosses Spiridens, with a limited distribu-
tion in Polynesia, and since recorded from the north-east and north-west of
New Guinea and Borneo, is probably a Papuan generic type.
6. OPEN " OPPORTUNITY " PLANT ASSOCIATION.
In a mountainous forest country like New Guinea, sparsely inhabited,
with little or no intercommunication between the different tribes, and no
migratory herds of grazing animals, there is everything to conserve and
nothing to modify natural conditions.
Lying in what may be called the centre of the monsun region, the whole
country is subjected to a more or less regular rhythm of alternating air
currents. Beccari (12, i. 216) has emphasized that the north-east monsun,
blowing regularly from November to April, must affect the general distribu-
tion of plants in the regions which come under the immediate influence of
these prevailing winds. This opinion has been already advanced by myself, 1
and later 2 from observations on the granite core of the exposed summit of
Kinabalu, before I had seen Beccari's convincing remarks in relation to his
own observations throughout Malaya.
Beccari aptly remarks that seeds are lighter than grains of sand from
volcanoes, and are adapted to remain longer in the air. Authenticated
instances of the possibilities of long-distance transport of grains of sand and
volcanic ash by the agency of wind are quoted by him (12, i. 216-220).
Warming 3 advances the same theory, for which extraordinary proof is
forthcoming in recent work of the Geological Survey of India. La Touche 4
1 L. S. Gibbs, " A Contribution to the Montane Flora of Fiji, with Ecological Notes,"
Journ. Linn. Soc., Bot. xxxix. (1909) 137.
2 . " A Contribution to the Flora and Plant Formations of Mt. Kinabalu and the
Highlands of Brit. N. Borneo," I. c. xlii. (1914) 47.
3 T. Warming, ' History of the Flora of the Faeroes,' Botany of the Faeroes II. Copen-
hagen, 1903.
4 T. H. D. La Touche, Mem. Geol. Surv. Ind. xxxv. (1902) 42.
46
discovered small undamaged fora minif era in the desert sand of Barmar and
Bikanir, which must have reached the heart of the desert by wind trans-
portation over a distance of 500 miles from the coast of Cutch.
This interesting observation gave the first clue to the remarkable work
organized by T. H. Holland, 1 to explain the origin of several intermittent
saline lakes in the Rajputana desert, in which the quantity of salt stored
is in excess of the amount that could be accumulated by normal freshwater
rivers acting within any reasonable geological period under present physio-
graphic conditions. The Rann of (.hitch dries up in the hot dry season, to
be covered with a thin incrustation of salt. This salt is transported by
strong winds from the south-west, which blow regularly from April to June,
to be followed by the rainy season, when the salt, deposited on the surface
of the desert, is washed in solution into convenient hollows, forming small
lakes. It was found, as a result of this investigation, which should be widely
known amongst botanists, that during four months of the hot season of 1908
the amount of wind-borne salt passing a front of 300 km. broad and 100 m.
high must have been something of the order of 1HO,000 tons.
Recently our airmen 2 operating in Mesopotamia have found the " dust
chokes the engines and the sand above blows as high as 4000'." Further
interesting proof of the constant direction of air-currents is afforded in the
first report on upper air research in Australia, 3 when difficulties were
encountered owing to the fact that Melbourne is on the south coast of
Australia, and the prevailing winter winds, as well as the upper currents in
advance of cyclonic disturbances, are from a northerly direction, and thus
carry the balloons out to sea. Material carried by wind is deposited on
reaching contrary currents, when should the seeds carried be precipitated on
to a suitable habitat germination takes place (12, i. 220).
Cross-currents would be most likely met with on the summits of high
mountains, where conditions in the tropics would alone be favourable to
ombrophobous plants requiring temperate conditions. We have now sufficient
evidence to prove that the area of high open country on the immense chains
of mountains in New Guinea must carry an enormous number of species of
this type of plant. The summits of Mt. Kiuabalu in Borneo, Mt. Halcon
and others in the Philippines, and Bonthain Peak in Celebes, would offer
the only suitable habitat for such plants between New Guinea and the
Himalayas in the west monsun region of distribution.
In those cases where not only the suitable area but also the fauna is much
restricted, though identical conditions of temperature and rainfall prevail,
1 T. H. Holland : successive Annual Reports of the Geological Survey of India published
in Recordt G.S.I, during 1904-09; "Discussion on the Physiography of Arid ' Lands,"
Report Brit. Asaoc. Adv. Sci. for 1914, 363 (1915).
a Candler, Edmund, ' Daily Mail,' Dec. 19, 1916.
Commonwealth Bureau of Meteorol., Bull. 13 (Melbourne, 1915).
47
possibilities in the evolution of new species are practically inhibited, and we
get the same or very closely sillied representatives.
But when cross-currents from the Antarctic (24, 222) cause precipitation
in Australia, the opportunity habitat is that of a large and open arid plateau
with little or no elevation, low rainfall, and a different and unlimited insect
fauna. Where constant conditions in illumination, temperature, and limited
rainfall prevail, there is practically nothing to limit the multiplication of
species able to survive the widely prevailing nrid conditions of this continent,
of which the rainfall is less than 10" over 100,000,000 sq. miles. 1 A
multitude of plants, so similar in appearance that it is difficult on casual
acquaintance to separate the different species, are evolved, especially as
annuals or ephemerals of the eremaea. To quote the genus Didiscus again,
which well exemplifies this theory : according to Domin 2 the Calycina
section represents the oldest form, limited to four species, of which two are
Papuan, one extending to N. Borneo and the Philippines ; the others occur
in Celebes and N.E. Australia respectively. Pseudo-calycina, considered
atavistic, is represented by one Papuan species and two in N.E. Australia.
The rest, or Eudidiscus, are all mostly annuals, younger types, represented
largely in W. Australia, with one in the Polynesian Islands, where restricted
conditions wouid .limit the evolution of the many closely allied species, so
marked a feature of Australian xerophytic types.
Open plant associations at the Angi lakes belong to the " Opportunity "
category, as under normal conditions these areas would be in forest, being
only kept open by the means of artificial burning at regular intervals. It is
extremely interesting in this respect that it should be possible to compare
two distinct associations of this type, developed under absolutely opposed
conditions, viz. :
1. The open marsh at 7000', sheltered in position, showing varying
conditions of soil and constant conditions of moisture.
2. The Cladonia association of Koebre at 9000', where constant con-
ditions of exposure and drainage prevail with a hard surface soil, sterilised
by the constant burning.
No more certain proof is afforded of the derived character of a flora
than that offered in a forest country by open spaces due to artificial circum-
stances, providing conditions, rainfall, and temperature are favourable.
This fact is well exemplified in New Zealand and the Pacific Islands, where
both rainfall and original plant covering once approached present Papuan
conditions, but now, under the stress of competition with ombrophobous
immigrant plants, not a single native species will be found in such areas.
1 Griffith Taylor, " Discussion on the 1 hysiography of Arid Lands," Report Brit. Assoc.
Adv. Sci. for 1914, 303 (1916).
2 Domin, K., " Monographic der Gattung Didiscus (DC.)." Sitz. Ktin. bo'hni. Ges. d.
Wiss. ii. Cl. (1908) 13, 20-23.
48
So much is this the case that the history and country of origin of immigrant
man, to whose agency the presence of this " Opportunity flora " is indirectly
due, can be pretty accurately determined by the nature of the invading plants.
It is the absence of aliens in the " Opportunity " associations of the Arfak
which form the engrossingly interesting feature in the phytogeography
of this region.
1. Open Marsh.
Endemism is the dominant note in the plant-covering of this marsh.
Most of the dominant plants found there are now described for the first time,
or were first collected by Beccari and Gjellerup. One or two have been
previously described from New Guinea, as Erwcaulon leucoyenes and Trisetum
latifolium ; others, to quote new Papuan records alone, are incidental wind
immigrants from the Himalayas, as Xyris pauciflora, Polygonum strigosum,
and Viola distans. The latter is unknown in Malaya with the exception of
the Philippines, while the others reach N.E. Australia, and therefore their
incidence in New Guinea was to be taken for granted. In Utricularia
racemosa and 7. bifida, the former shows the Himalayan range and the
latter is limited to Malaya, while Gahnia psittacorum, abundant on both
these open areas and also at home in the forest, reaches E. Australia and
Tasmania. Most of the other Cyperacese are cosmopolitan tempei-ate types,
of incidental wind distribution, like the cryptogams, of which, in relation to
the freshwater Alga?, Professor West writes " that all the species observed
are ubiquitous, few of the tropical ascending to 7000', the one exception
being Closterium Bacillum, known only from Burma." The lichens, most
mosses and ferns, including the Lycopodiums of these areas, are also cosmo-
politan, while the only Selaginella collected is endemic.
The one plant to suggest man's agency was J)esmodium Scalpe, an
unexpected representative of a genus that, may almost be described as alien
to the Papuan mountain flora, but, growing on the site of van Oosterzee's
and the Pratts' camps, it may be considered the one relic of alien intrusion.
On Kinabalu this plant has so far only been found at Lobang, on the
invariable camping-site.
2. Cladonia A ssociation of Koelre.
Nonb of the ombrophobous herbaceous plants with the exception of the
Riedelias, a feature of the open spaces of the S.W. ridge and of this summit
plateau, were collected in the surrounding forest. They were all plants
requiring constant illumination and low temperature for their development,
of which the germination of the seeds would be inhibited under shade
conditions.
Some of these plants, such as the Dendroliums and Centrolepis, are
common to the open spaces of the S.W. ridge and to the marsh ; of the
49
others, Didiscus has been recorded from Mt. Scrutchley in the south-east,
and the Platanthera from the south-west.
The presence of the other plants must be due to wind-incidence, and they
would be derived from natural exposed areas above the tree-level on the
mountains of greater altitude to the east and south of the country.
The cryptogams, again, as is the case on the marsh, are all cosmopolitan.
This remarkable ridge association of Koebre combines some of the most
peculiar elements of what have been considered the Malayan, Polynesian,
and Australian floras.
The plants found there show roughly what the systematic enumeration
of the species collected proves in detail, that the flora of the mountains of
New Guinea, almost unknown outside the last ten years, must now be
considered the axle of a wheel of distribution, of which the spokes alone have
so far been familiar to us. This is in agreement with all recent work at
similar or greater altitudes. Had that axle, even now barely investigated,
been worked out first, we would, as a matter of course, speak of the
dominance of Papuan elements in neighbouring floras as the German and
Dutch botanists have already rightly suggested.
SOME PLANT ASSOCIATIONS OF THE N.W. COAST.
Dorei Bay.
The chief plant association of Dorei Bay is that of the " korang " forest
clothing the low coral-limestone range which rises immediately behind Mano-
koeari to the height of about 500', in a gradual slope from the sea-shore.
This forest is still in its pristine condition, as all the surface-water drains
through the sterile and porous subsoil, to a certain level line, about 200'
above the beach, which marks the issue of the small streams representing the
drainage of the ridge. This line also limits possible cultivation, as below it
the " korang " is covered with sufficient depth of soil, due mostly to the
erosive action of thse streams, to allow of necessary but not luxuriant
cultivation.
The old "pisang" 1 plantations of the Alfueros, now run to seed, with
secondary jungle upgrowth, abut on to the natural forest at this level, on
which both the reservoirs collecting for the water-supply of Manokoeari are
sit.u.-ited at different points.
The peculiarities of this " korang " forest were noted by Forrest in 1750
(1, 111), who wrote "there being no underwood it is easy travelling under
the lofty trees "; and Dumont d'Urville in 1827 (3, iv. 581) estimated the trees
in the forest as 80-200' high, writing of a " sol degage, arbrisseaux clairsemes,
1 Banana.
50
fongeres de petite taille, et fort peu de plantes herbacees," and further on
(3, iv. 602), " Tous les environs du Havre proprement ditsont occupes par des
tbrets a 1'etat de nature, situees sur un sol entierement madreporique, qui
s'eleve en pente tres douce " ; and, finally, Wallace, in 1852 (6, 173), describes
" the Dorey promontory is a raised coral reef, and, geologically speaking, a
very recent one. The beach is a mass of dead and broken coral, not yet
ground into sand, quite impracticable for walking, and from this beach up
into the jungle, and even on to the hill, to the height of 200' or 300', there is
scarcely a perceptible change in the coral rock, and the masses of coral and
shells that everywhere strew the surface. In some of the gulleys, however,
I found traces of a core of stratified rock."
I did not work over any of the gullies, as once off the "korang" range
the conditions are all secondary, every inch of ground having been under
present or past cultivation.
With regard to the beach, the coral mentioned by Wallace had possibly
been washed up by a heavy N.W. monsun, as our own beaches are often
covered with shingle during the winter gales, to be dispersed again later.
At the period of my stay it was certainly not an apparent factor.
On the "korang" the soil is so thin that the coral is always visible,
mostly covered with dead leaves. The most interesting portion is along the
flat-topped summit where the surface is more even and advantageous to
plants, and in parts small soak-areas hold shallow standing water. The
immediate flanks proved barren of results, being very dry with great over-
hanging outcrops of pure " korang "-like cliffs, too porous to offer much hold
for plants.
On what may perhaps be referred to as the drainage-line of the streams
issuing from the range, quite a different type of undergrowth prevailed,
almost luxuriant in character, comprising chiefly ferns, Zingiberaceous and
Araceous plants.
Trees. Most conspicuous were fine isolated examples of that magnificent
palm *Pigafetta pilaris. Too beautiful to cut down, I only took some old
fruit and the measurements of the immense leaves, those shed, with the
old flowering rhachises, remaining piled around each tree, which in con-
sequence form isolated spots in the forest. Dr. Beccari, however, with his
personal knowledge of this locality and expert interest, had no difficulty in
determining this splendid species. *Ficus myriocarpa, *F. celebica, *F. botryo-
carpa, and F. Irachiata, the two latter with green receptacles which all
contained water, were very general, with the large-leaved c Macaranga riparia,
*Mallotus tilicvfolia, Aglaia Gibbsece, with large branching white racemes,
* Euonymus javanicus, and *Albizzi.a moluccana.
Climbing plants. All the trunks of the trees were covered with root-
climbing epiphytes, as in the "korang" forest of the inundation-zone of
51
Geelvink Bay Pothos sp. not seen in flower or fruit, *Piper Forstenii
with liuge leaves, also sterile. The scandent fern Thysanosoria dimorpho-
phylla (PI. 4. fig. 7) with fertile fronds at the apex of the shoots, the type
of a new genus, was very common with *Lygodium digitatum, Freycinetia
lanceolata, *Drac,tvna angustifolia, *Flagellaria indica in huge examj)les
running up the tallest trees in the forest, and *Zanonia macrocarpa enveloping
most of them in its heavy curtains.
Epiphytes. As in the littoral korang forest these were not numerous and
not a single orchid uas collected. Possibly the thick swathing growth of
climbing epiphytes and stem-clasping lianes may, to a certain extent, account
for their absence, also perhaps the porous nature of the subsoil, which
reduces the constant evaporation so essential to the support of the large
epiphytic flora common to the primary high forest of the foot-hills. The
moss *Pelekium trachypodum on dead wood, the fern *Antrophyum reticu-
latum, with the white-flowered Mymecodia puloinata, were collected.
Undergrowth. The hepatic *I)iimortiera velutina was found in patches,
and the terns * Stenosemia aurita always in colonies, while *Asplenium laser-
pitiifolium, *Aspidium Lenzeanum, with *Dryopteris truncata and *I)iplazium
proliferum, the two latter like small tree-ferns in habit, were dotted about.
Small colonies of the creeping * Hemiyraphis reptans, *Geophila reniformis,
with the orchids, always grouped, Microstylis Gibbsete and the larger orange-
green M. xanthocheila, \\ ith Liparis maboroensis var. bistriata. Of larger
plants, *Centotheca lappacea and *Schleria margaritifera, the small white
Draccena novo-guineensis with very screwed leaves, were abundant; Pellionia
Vanhasseltii massed on a prostrate trunk and on the ground beneath, the
stinging Laportea armata about 1-2 in. high, gregarious on a small soak-
area, and the shrubby Arnarocarpus Wichmannii, about 1 m. high with
dorsiventral branches, were scattered over the surface, which showed no
understaging of shrubs and little even of young trees.
In the denser undergrowth at the base of the forest, on the drainage-line,
abounded *Aspidium pachyphyllum with fertile and sterile fronds, Aglaionema
novo-guineensis about 1 in. high, the leaves crowded towards the top and the
flowers with green spathes and white spadix and red fruit, and Cyrtosperma
macrot urn ; Alocasia acuta, a peculiar plant with large fleshy leaves on long
petioles, crowded at the apex of a stem about 2 m. high, with the flowers
bunched in the axils of the leaves, was common, with the white *Peristrophe
ialappcefolia, not previously recorded outside Java. Schismatoglottis dorensis
spread in large colonies in more open places where the white translucent
Clavaria Gibbsece and * Dictyophora phalloidea, always yellow, were plentiful,
the presence of the latter being invariably revealed by the smell, with
*G easier Jim hiatus und the smaller G. mirabilis var. tricliifer.
52
Clearings at Edge of Forest and Secondary Jungle.
This form o tropical upgrowth was not so rampant as is usually the case,
on account of the poor " korang " subsoil, but it proved more than usually
interesting in character in showing such a large proportion of endemic
Papuan and Moluccan species.
In clearings Gigantochloa novo-guineensis, apparently cultivated, was in
flower, the fine Pandanus Tabbersianus bearing fruits almost 1 m. long,
*Mallotus tilicefolia, *MellocTiia arborea, *Kleinhofia hospita, *Tamarindus
indica, *Rubus moluccanus, * Callicarpa erioclona, white, Premna nitida, also
white-flowered with black fruit, and the ubiquitous * Wedelia biflora, with the
scandent * Allopliyllus Cobbe, *Flagellaria indica, *Rhyssopteris timorensis,
the bright yellow Sccevola novo-guineensis, *Merremia nymphceifolia, with the
epiphytic -Loranthus Versteegii, of which the flowering, vertically hanging
shoots bear dense red racemes, standing out at right angles to the stems for
half their length, were all general. There has been some question lately
about the fertilization of LorantJius sp. by butterflies. I can only say that, on
vigorously pulling one of the long shoots in the first excitement at seeing
such a peculiar Loranthus form, I was covered by showers of very large red
ants. The denseness and position of the racemes would certainly favour
fertilization by these insects.
Just below the drainage-level of the forest-clad range, a damp area of
some extent, shaded by a secondary unidentified upgrowth, proved a very
good collecting-ground for fungi. In fact, almost the whole total of the
species collected were found on this area, the " korang " forest itself being
too dry in character to encourage this form of growth.
Cultivation.
In the grounds of the Residency, bread-fruit, Avocado pears, pumiloes,
custard-apples, mangoes, Canarium nuts, jambu, limes and lemons, pine-
apples, with small water-melons and hill-rice, have been successfully grown.
Roses were always in flower in the garden, of which the chief interest was a
well-grown young Araucaria Beccarii about 5 m. high, brought down from
the Angi lakes by Mr. van Oosterzee, who had laid out and planted these
gardens. The old convict gardener who looked after them lo.st no oppor-
tunity of denouncing the " korang," which certainly outcrops in most
inopportune places, and the shallow soil. Surrounding the " Pasangrahan "
and the quarters of the " Pradjoerit," all the usual " sayur " were grown by
the " Orang rante " in the well-kept gardens, and also by the Chinese, who all
have their own plots of cultivated ground.
53
Mangrove Association at Langg&n.
On the spit of land between the two bays a Papuan " campong " was
built among the mangroves, where *^Egiceras floridum flourished as a round
shrub about 1 m. high, covered with the white flowers and quaint fruit. On
a tree overhanging the sea-water, * Lycopodium phlegmariodes with Dendro-
bium pseudo-ealceolum in flower and various Dischidia spp. abounded.
*Pandanus polycephalus with small red fruit bunched at the apex of the
peduncle, common through the Moluccas, with *Excacaria Agallocha and
the climbers * Tristellateia australasica, a mass of yellow flowers, *Derris uligi-
nosa and Sarcolobus retusus, marked the land-edge of the mangrove-spit ;
while Freycinetia Beccarii, Erythrospermu>n candid^,, the yellow-flowered
*JDurandea parvifolia, a Gardenia sp., with *Pollia sorzogonensis as under-
growth, were found where the ground was more consolidated, and young
colonies of *Pigafetta pilaris were quite abundant towards the shores of the
second bay.
Wousi and Genbela.
Wousi, in earlier times the watering-place for all the boats calling at
Dorei Bay, where the Papuan " campong," with a fringe of houses built over
the sea, still stands, is a tiny valley cut out of the " korang " range by the
action of the stream, the range from this point gradually sloping to the level
ground. At the time of my stay Wousi was also the site of the Military
Bivouac, and the base for the work of the Exploration detachments so suc-
cessfully organized from 1907 by the military authorities at Amboina. The
survey having been completed by the successful results of Captain Opper-
man's Expedition (27, 542-3) the bivouac has now been closed.
In the military cartographical office hung a huge map of Dutch New
Guinea, originally blank, filled in by degrees with tracings of the work
of each Exploration detachment in turn. When I was shown this map
only one blank space remained, viz. the source of the Mamberamo River.
With the successful results of Captain Opperman and Mr. Langeler that
blank space has also disappeared, and with it this admirable era of organized
exploration is closed, the whole of the Dutch possessions in North New
Guinea being now mapped out.
At Wousi, under the shade of the overhanging trees, the ideal and
classical anchorage was reserved for naval and military needs, and all the
ground underneath the beautiful century-old trees behind the beach was
cleared and grass sown, forming a fine green sward, where the quarters of
the military and naval officers were built. Farther up the valley were the
open barracks for the native troops, carefully arranged with intersecting
white paths, beyond which again excellent gardens had been made up the
bu.iki of the stream, where all the vegetables for the needs of the forces were
54
grown, a herd of the beautiful little Bali cattle being also kept for regular
killing ; on these occasions the requirements of the official residents of
Manokoeari were also taken into account.
Genbela.
Beyond Wousi secondary forest with Malay houses at intervals bounded
the beach to Genbela, where fine sands run out to the cape that limits the
bay to the east. In the. forest the fungus *Favolus scaber, the lianes
* Entada scandens and Mucuna Krdtkei were collected. *Peristroplie
jcdappcefolia, *Hemi(/r aphis reptans and H. dorensis, with *Geopldla reniformis,
formed constant undergrowth. On the edge of the sandy beach *Pandanus
dubius grew in clumps, the young plants unbranched with thin leaves about
3 m. long; the old trees about 8 m. high, with many branched crowns and
shorter leaves of much stiffer consistency, bore large round heads of glaucous
mericarps. Behind this beach, on a level stretch of country with intermittent
sago-swamps, the native plantations were mostly situated.
Island of Roon.
We stopped a day at Djende, the chief place on the island, of which the
gneiss J formation carries a mainland type of vegetation. Djende lies in a
deep sheltered bay, the native houses being all built over the water, like
a miniature Brunei. It is surrounded by hills about 500' high, which, from
the little seen, seemed to carry a very interesting association of plants.
Along the road bordering the bay, Anthoceros bullato-spongiosus associated
with the minute mosses *Garckea phascoidea and *Wilsoniella pellucida and
young plants of * Lycopodium cernuum grew in the shade, while *Sclileria
margaritifera and 0tanthera novo-gmneensis were found in the open.
On the forest slopes Cyailiea runensis grew as undergrowth, where the
orchid Vrydagzynea elongata was growing sporadically with Centotlieca
lappacea ; * Trichomanes bipunctatum was collected as an epiphyte, also
* Piper Forstenii in flower, with hanging yellow ? spikes, about 4 dm. long.
The forest was in a very dry condition, rather a surprising fact, taking the
heavy rainfall into account (p. 13), also considering it was the rainy
season ; this fact again proves the fallacy of the all-embracing term rain-
forest applied in general to all and sundry tropical forest formations. The
actual rainfall is by no means the dominant factor, as it is the limiting
characters, no matter how small their incidence, which have to be taken into
account ; that is to say exposure, soil, and drainage, while should drier
A. Wichmann's Uerichte in Jiull. nos. 43, 44 & 40 v/d Maatscli. ter bev. v.in het Natuurk.
Onderzoek d. Nederl. Kolon. (N. Guinea Exped. 1903, Bull. nos. 3, 4 & 6). 80. Leiden.
55
conditions obtain for only one week, that week will limit the conditions of
the forest formation.
In N. New Guinea as in Brit. N. Borneo I saw no forest that answered
to the description of rain-forest. 1
Humboldt Bay.
This bay forms a most beautiful inlet, broken in outline, surrounded by
mountains and protected by a small island, on which a very picturesque
Papuan pile-village with an imposing " spirit house " is situated. As the
Tidorese never penetrated so far (8, 87) the natives here are still unspoilt in
primitive condition. The Government Station is at the head of the bay, just
behind the beach. On a marshy spot near the " campong " * Tlioracostachyum
hypolytroides, a sedge with white upper leaves, bracts, and inflorescence, grew
abundantly. On the banks of the river Gigantocldoa novo-guineensis was in
flower, as at Manokoeari, also *I)raccena angustifolia. We entered the
" rimbu/' the Controleur having most kindly found me two Malays who knew
the forest, on the edge of which a handsome Casuarina with a spreading crown,
most distinct in habit, was abundant. Ascending the ridge, clothed in fine
high forest on deep brown soil, we found as undergrowth * Trichomanes
javanicum var. rhomboideum, *Diplaziuin maximum, with *Selaginella plumosa,
creeping, the palms Licuala montana with red fruit and *Arenga microcarpa,
the aroid Q Holochlamys Seccarii, like a Caladium in habit, and the under-
trees *Pipturus argenteus and Clerodendron Lindawianum var. glabrescens,
the latter with handsome white flowers and black fruit, also the climbing
*Polypodium normale and Calamus humboldtianus.
Over the crest of the ridge, where great mounds of dead leaves testified
to Megapode activity in building their nests, we descended by a stream into
quite a different type of vegetation, one of those sudden changes in
comparatively small areas, which so constantly characterize the mixed
tropical forest.
Along the course of the stream, in which I picked up pure alabaster,
much fancied by the Papuans for nose-ornaments, * Trichomanes humile was
found on rocks with * Vittaria elongata, the hanging fronds to 2 m. in length.
The fungi *Polyporus arcularius and *Hirneola polytriclia grew on dead
wood.
As undergrowth the handsome Pteris torricelliana with fronds 1*50 m
long, and //i/j><>lt;j>is grandifrons on a rhachis about 3 cm. through, bearing
magnificent single deltoid fronds about 5 m. long, the petiole about 3 m.,
and lamina 2 in. long, rising at intervals from an underground rhizome, \vas
1 L. S. Gibbs, " A Contribution to the Flora and Plant Formations of Mt. Kinabalu and
the Highlands of Brit. N. Borneo," Journ. Linn. Soc., Bot. xlii. (1914) 8.
56
quite distinct from anything in ferns so far familiar. *Angiopteris evecta was
abundant, and Alsophila straminea well represented. Begonia humboldtiana
with mottled leaves and pink flowers grew all along the banks in large
clumps, but it was impossible to find any normal fruit, the capsules being
all swollen through the action of some insect. Ficus conocephalifolia, most
aptly named, with enormous leaves and bearing red receptacles, was a
common tree.
Bosnik, on the Island of Wiak.
On the return to Manokoeari, half a day spent on Bosnik, on ne of the
Schouten Islands, was interesting, it being the first time the steamer had
called there, this new Government station having been built as an alternative
to Mosmer as a point of call. The island is coral-limestone, a high ridge
rising behind Bosnik, and low-lying flats in front of the latter are rapidly
being cleared for the " campong " plantations.
Under the strand trees bordering this area * Cycas ctrcinalit, the ferns
* Dryopteris stenobasis, *Diplazium polypodioides, with *Fleuria ruderalis,
Ophiorrhiza insularis, Piper bipunctatum, and P. bosnicanum, the latter
epiphytic on a tree, were found.
As this was quite a new station, the strand trees were not yet stripped
of their epiphytic treasures, and one splendid example, a veritable garden of
various plants, yielded *Lycopodium phlegmaroides, *Psilotum flaccidum, the
orchids Hippeopliyllum alboviride, Dendrobium inconspicuum and D. potomo-
philum, Eria rigida var. papuana, and Sarcanthns bicornis, all with incon-
spicuous flowers, with the handsome Medinilla rhodorhachis with pink
flowers.
All the streams drain through the " korang " into the sea at the beach
level, so that it is possible to drink fresh water from the salt.
At all the other stopping-places the time was too short to do more than
collect a few plants, or, the islands being entirely under cultivation, possibilities
in collection were limited to the beach.
SYSTEMATIC RESULTS.
Over 330 plants were collected in the Arfak, of which 100 have proved
new to science, with one new natural order and five very distinct new genera.
Of. the new species perhaps the most interesting are a Dacrydium, the first
species in fruit to be described from New Guinea, a Libocedrus, a genus new
for Dutch N.W. New Guinea, and a Kentia. A new species each in Trimenia,
Telminthodia, and Backhousia, establish Papuan preponderance in those
genera, while a new species in Patersonia, Centrolepis, and Hibbertia represent
new generic records for New Guinea.
I/- ,-2/A
^e'*<.* . He. Y* V -
57
The few widely distributed plants included comprise about 30 new
records for New Guinea, of which the most interesting are Xyris pauci/ora,
Bulbostylis capillaris var. trifida, Spathoglottis aurea, Polygonum strigosum,
Viola distans, and Hydrocotyle rotundifolia.
Several of Beccari's Hatam plants have been re-collected, viz. Riedelia
orchioides, Palmeria arfakiana, in a J example, of which the ? is so far de-
scribed, Styphelia trochocarpoides,and Dicrotrichium brevipes. Of Gjellerup's
plants, as would be expected in working over the same ground, the dupli-
cate examples are more numerous, especially in the orchids, but even in
that order Smith has described 20 new species with four new varieties, and
in Ericaceae two Rhododendrons, four Vacciniums, and one Diplycosia prove
new, with one variety in Styphelia in Epacridacese.
Collections made subsequently round Manokoeari (Dorei Bay) on some
of the islands along the coast, and at Humboldt Bay, have been separately
enumerated, no two species proving common to both the mountain and coast
flora. About 150 plants are comprised in this list, which includes one new
genus and several new species in ferns, with 27 new species in other genera,
and interesting new records, of which the larger portion is more Malayan in
type. Wide distribution of endemic plants is again a very marked feature.
In working out these collections I must express my thanks to the expert
botanists who have so kindly determined those orders in which they were
interested. To Dr. J. J. Smith of Buitenzorg I am under especial obligation,
not only for undertaking the Orchidacese, Ericacese, and Epacridacese, but
also for many valuable details concerning work already done in the Arfak,
and in arranging for the services of one of his trained native collectors
who accompanied me to New Guinea. I am indebted to Dr. Odoardo Beccari
for working out the Palmse ; Dr. Valeton for the Ziugiberaceae and
Rubiacea? ; and M. Casimir de Candolle and Professor J. Macfarlane for the
Piperacere, Meliaceajj and the Nepenthacese. In London my thanks are due
to Drs. Stapf and Rendle and the staffs of Kew and the British Museum for
much kind help ; and in particular to Professor G. S. West for the deter-
mination of the Freshwater Algoe ; Miss A. Lorraiu Smith for the Lichens ;
Dr. A. B. Rendle for Pandanacese, Cyperacese, and Gramineje, and a new
genus in Urticacese ; Dr. 0. Stapf for Utriculariaceae and valued criticism ;
Messrs. J. Ramsbottom, A. Gepp, L. Hutchinson, E. G. Baker, H. N. Ridley,
and J. R. Drummond for Fungi, Bryophytes and Pteridophytes, Euphor-
biacese, Melastomacese, Moracea3, and Solanacese ; and to Mr. S. Moore for
the Labiata?, Acanthacese, and Composites, and his great kindness in revising
the proofs.
The plants may be consulted at the British Museum, Kew, Leiden, and
Buitenzorg, and in my own collection (on loan to the British Museum).
r
58
PRINCIPAL BIBLIOGRAPHY FOR DUTCH N.W. NEW GUINEA.
1. FORREST, T. A Voyage to New Guinea and the Moluccas from Balambangan.
London, 1779. (2nd ed. 1780.)
2. LESSON, P. Voyage autourdu Monde sur la Corvette La Coquille. Paris, 1838.
3. DUMONT D'URVILLE, J. S. C. Voyage de V Astrolabe, 1826-29. iv. Paris, 1832.
4. DE BRUIJN KOPS, G. F. Bijdrage tot de kennis der Noord- en Oostkusten van
Nieuw-Guinea. Natk. Tijdschr. Niederl. Indie, Deel i. (1850) 163-222.
5. WALLACE, A. R. Malay Archipelago. London, 1869.
6. . Notes on a Voyage to New Guinea. Journ. R. Geogr. Soc. xxx. (1860)
172-177.
7. . Island Life. London, 1880.
8. GOES, H. D. A. VAN DER. Nieuw Guinea, ethnographisch et natuurkundig onder-
zocht in 1858. Bijdr. t. de Taal-, Land- en Volkenkunde, (2) v. (1862) 72.
9. D'ALBERTIS, L. M. Una mesa fra i Papuani del Monte Arfak. Boll. Soc. Geog.
Ital. x. 2 (1873) 67-71.
10. ROSENBERG, H. VON. Reistochten naar de Geelvinkbai. 's Gravenhage, 1875.
11. BECCARI, O. Esplorazione dei Monte Arfak. Boll. Soc. Geog. Ital. i. (1876)
35-38.
12. . Malesia : i., ii., iii. 1877-1890.
13. SCHEFFER, R. H. Plantes de la Nouvelle Guinee. Ann. Jard. Buit. i. (1876)
1-60.
14. TETSMANN, M. J. E. Voyage a la Nouvelle Guinee. Loc. cit. 61-95.
15. ST. VRAZ, E. Reise nach Neu-Guinea. Petermann's Mitth. xliv. (1898)
232-35.
16. WARBURG, O. Monsunia, Beitrage zur Kenntniss der Vegetation des Slid- und
Ostasiatischen Monsungebietes. i. Leipzig, 1900.
17. OOSTERZEE, L. A. VAN. Tijdschr. Kon. Ned. Aardr. Gen. (2) xxi. (1904)
998-1021.
18. VALETON, Tir. Plant* Papuanae. Bull. Soc. Dep. Agric. Indes Neerl. x.
(1907) 1-70.
19. HARTMANN, A. Repertorium op de Literatuur betreffende de Nederlandsche
Kolonien. 1866-1893. 1894-1905. 1906-10. Published by Murtinius
Nijhoff.
20. GELDER, J. K. VAN.^ Verslag omtrennt eene geologische verkenning van de
Mambei-amo-rivier op Nieuw-Guinea. Jaarb. Mijnw. Ned. Oost-Ind. Bat.
xxxix. (1910) 87-112.
21. WICHMANN, C. E. A. Nova Guinea. History, i.-ii. (1909-12).
22. NOVA GUINEA, Botany, viii. (1907-13), (1913-15) ; xii. (1913-15).
23. SCHLECHTER, R. Die Orchidaceen von Deutsch-Neu-Guinea. (1911-1914.)
24. BRAAK, C. Het Klimaat van Nieuw-Guinea. Natk. Tijdschr. Nederl. Indie,
Deel Ixxiii. (1914) 179-266, with Map.
25. SMITH, J. J. De Exploratie van Nieuw-Guinea. in Tijdschr. Kon. Ned. Aardr.
Gen. (2) xxx. (1913) 77-8.
26. . Ibid. xxxi. (1914) 782.
27- . Ibid, xxxii. (1915) 542-3.
[References to papers not reftrring to Dutch N. If. Jfeu- Guinea
are girtn in the text.']
PLATE I.
FIG. 1.
FIG. 2.
PLATE II.
FIG. 3.
Fio. 4.
PLATE III.
FIG. 6.
PLATE IV.
P. Highley, photo.
THYSANOSORIA DIMORPHOPHYLLA, Gepp.
59
SYSTEMATIC ACCOUNT OF THE PLANTS COLLECTED AT THE
ANGI LAKES, 7000-9000', IN THE ARFAK MOUNTAINS, IN
DUTCH N.W. NEW GUINEA, IN DECEMBER 1913.
[An asterisk denotes new records for New Guinea. Collectors' names and
localities are given for Dutch New Guinea only.~^
THALLOPHYTA.
FRESHWATER ALG^l. (G. S. WEST.)
The geographical distribution has not been included, since all the species
observed are ubiquitous except Closterium Bacillum Joshua, known only
Erom Burma. The reason for this general ubiquity is the altitude,
few of the tropical types ascending to 7000'.
MYXOPHYCE^:.
CHROOCOCCUS MINOR Nag. Gatt. einzell. Alg. 1849, 47, t. 1 A, f. 4 ; Rabenh.
Flor. Europ. Alg. ii. 30.
No. 5971.
CHROOCOCCUS TURGIDUS Nag. Gatt. einzell. Alg. 1849, 46 ; Rabenh. Flor.
Europ. Alg. ii. 32.
No. 5971.
MERISMOPEDIA GLAUCA (Ehrenb.) Nag. Gatt. einzell. Alg. 1849, 55, t. 1 D,
f. 1.
No. 5971.
OSCILLATORIA AMPHIBIA Ag. in Flora, x. 1827, 632.
No. 5971.
OSCILLATORIA ANGUSTISSIMA W. & G. S. West in Journ. Bot. 1897, 300.
No. 5723.
OSCILLATORIA LIMOSA Ag. Dispos. Algar. Suec. 1812, 35.
No. 5723.
OSCILLATORIA TENUIS Ag. Algarum Decades, ii. 1813, 25.
No. 5723.
OSCILLATORIA TEREBRIFORMIS Ag. in Flora, x. 1827, 634.
No. 5971.
LYNGBYA VERSICOLOR (Wartm.) Gom. in Ann. Sc-i. Nat. 7 e serie, Bot. xvi.
1892, 147, t. 4, f. 4-5.
No. 5971.
STIGONEMA OCELLATUM Thuret, Essai de classification des Nostochinees, Ann.
Sci. Nat. 6 e serie, Bot. i. 1875, 380.
No. 5971.
SCYTONEMA MIRABILE (Dillw.) Thuret. \_ = S. figuratum Ag. Syst. Algar.
1824, 38.]
No. 5971.
BACILLARIE^E.
FRAGILARIA PARASITICA (W. Sin.) Grun. Van Heurck, Synops. Diat. Belg.
1885, t. 45, . 30. Odontidium parasiticum W. Sin.
No. 5978. Epiphytic on Surirella robusta var. splendida.
EUNOTIA BICAPITATA Grun. Van Heurck, 1. c. 1885, t. 35, . 11.
No. 5971.
EUNOTIA LUNARIS (Ehrenb.) Grun. in Van Heurck, 1. c. 1885, 144, t. 35,
f. 3, 4, ot 6.
No. 5971.
EUNOTIA ROBOSTA Ralfs. Van Heurck, 1. c. 1885, 144, t. 33, f. 11-13.
No. 5971. Both forms originally described by Ehrenberg as E, octodon
and E. polyodon were plentiful.
NAVICDLA APPENDICULATA Kiitz. Bacill. 1844, 93, t. 3, f. 28.
Nos. 5723 and 5971.
NAVICULA BICAPITATA Lagerstedt, Spitsberg. Diat. 1873, 23, t. 1, f. 5.
No. 5971.
NAVICULA EXILIS Kiitz. Bacill. 1844. Van Heurck, 1. c. 101, t. 12,
f. 11, 12.
No. 5723.
NAVICULA MAJOR Kiitz. Bacill. 1844, 97, t. 4, f. 19.
No. 5971.
NAVICDLA PAR VA (Greg.), jy.gibba var. brevistriata Van Heurck, 1. c. 78,
t. 6, f. 5.
No. 5971.
NAVICULA VIRIDIS Ktttz. Bacill. 1844, 97, t. 4, f. 18.
No. 5971.
COCCONEMA LiEVE (Nag.) G. S. West in Journ. Linn. Soc., Bot. xxxviii.
1907, 159. Cymbella laivis Nag. in Kiitz. Spec. Alg. 1849, 59.
No. 5971.
NITZSCHIA COMMUNIS Rabenh. Alg. no. 949 ; Flora Europ. Alg. i. 1864,
159. Van Heurck, 1. c. 184, t. 69, f. 32.
No. 5723.
61
NITZSCHIA CONSTRICTA (Kutz.) Pritch. Infus. 1861, 780. N. dulia W. Sm.
Brit. Diatom, i. 1853, 41, t. 13, f. 112.
No. 5978.
NITZSCHIA PALEA (Kutz.) W. Sm. Brit. Diatom, ii. 1856, 89. Synedra
Palea Kiitz. Bacill. 1844, t. 3, L 27 ; t. 4, . 2.
No. 5978.
NITZSCHIA FRUSTULUM (Kiitz.) Grun., var. PERMINUTA (Grun.). Van Heurck,
1. c. t. 69, f. 4.
No. 5723.
HANTZSCHIA AMPHIOXYS (Ehrenb.) Grun. in Kongl. ST. Vet.-Akad. Handl.
xvii. no. 2, 1880, 103.
No. 5723.
SURIRELLA ROBUSTA Ehrenb. in Ber. Akad. Berlin, 1840, 215. S. nobilis
W. Sm. Brit. Diatom, i. 1853, 32, t. 7, f. 63.
Var. SPLENDIDA (Ehrenb.) Van Heurck, 1. c. 185, t. 72, f. 4.
No. 5978.
All the specimens were covered with the small epiphyte Fragilaria
parasitica (W. Sm.) Grun.
CHLOROPHYCE.E.
SCENEDESMUS ACUTiFORMis Schroder in Forschungsb. Biol. Stat. Plon, v.
1897, 17, t. 2, . 4.
No. 5971.
ULOTHRIX SUBTILIS Kutz. Phyc. Germ. 1845, 197 ; Tab. Phyc. ii. 1852,
t. 85, f. 1.
No. 5971.
MIOROSPORA PACHYDERMA (Wille) Lagerh. Conferva pachyderma Wille in
Ofvers. a K. Vet.-Akad. Forh. 1881, no. 8, 20, t. 1, f. 28-35.
No. 5971.
SPIROGYRA sp. (sterile).
No. 5978. This species was undeterminable, but may possibly be one of
the forms of Sp. decimina (Mull.) Kiitz.
NETRIUM OBLONGUM (De Bary) Liitkem. in Cohn's Beitrage zur Biol. der
Pflanz. viii. 1902,407."
Var. CYLINDRICUM W. & G. S. West in Journ. Bot. 1903, 8 (sep.),
t. 446, f. 10.
No. 5971.
62
PENITJM sp.
No. 5971.
Several specimens were observed of a small Penium, but they were not in
good condition for identification. It is possible that it is a form of P. pliy-
matosporam Nordst.
CLOSTERIUM BACILLUM Joshua in Journ. Linn. Soc., Bot. xxi. 1885, 652,
f. 4-6.
Long. 224 p ; lat. 34 /t.
No. 5971.
Known previously only from Burma.
MICRASTKRIAS DECEMDENTATA (Nag-.) Archer in Pritch. Infus. 1861,
726.
Long. 68 ft ; lat. 68 fi ; lat. isthm. 15 /A ; crass. 24 /*.
No. 5971.
COSMARIUM L^VE Rabenh. Flor. Europ. Alg. iii. 1868, 161 ; G. S. West
in Journ. Linn. Soc., Bot. xxvii. 1899, 386, t. 10, f. 1-6.
No. 5971.
COSMARIUM PSEUDOPYRAMIDATUM Lund in Acta R. Soc. Scient. Upstila,
ser. 3, viii. 1871, 41, t. 2, f. 18.
No. 5971.
COSMARIUM PUNCTULATUM Breb. W. & G. S. West, Brit. Desm. iii. 1908,
206, t. 84, f . 13, 14 ; t. 102, f. 22.
No. 5971.
HYALOTHEOA DISSILIENS (Sin.) Breb. in Ralfs' Brit. Desm. 1848, 51, t. 1,
f. 1.
No. 5971.
LICHENES. (A. LORRAIN SMITH.)
*CLADONIA COCCIFERA (L.) Willd. Fl. Berol. 1787, 361.
Arfak Mts., Koebre ridge, growing thickly on burnt open summit, 9000'.
Dec. 5598.
Distrib. Cosmopolitan.
*CLADONIA DIDYMA (F^e) Wain., var. MUSCIGENA Wain., Monogr. Clad. i.
1887, 141.
Arfak Mts., Koebre ridge, 9000', growing thickly on burnt open summit
Dec. 5720.
Distrib. New Caledonia; West Indies; Central America j Brazil; Chile j
Peru.
63
*CLADONIA VERTICILLATA (Floerk.) Schaer. Lich. Helv. Spic. 1831, 31.
Arfak Mts., Koebre ridge, 9000', carpeting on burnt open summit.
Dec. 5739.
Distrib. Cosmopolitan.
STICTA VARIABILIS Achar. Lichenogr. 445. Schum. & Laut. N. 30.
Arfak Mts., Angi lakes, epiphytic in forest by ? lake, 7000'. Dec.
5905.
Distrib. N.E. New Guinea. Polynesia to E. Africa.
FUNGI. (J. RAMSBOTTOM.)
PYRENOMYCETES.
H YPOCREACE^E.
PODOCREA COHNU-DAM^: (Pat.) Lind. in Engl. & Prantl, Naturfam. i. 365
(1897).
Lower foot-hills, Arfak Mts., 500', terrestrial in forest. Jan. 5703.
This species was described from China by Patouillard, who placed it in
the genus Hypocrea. The stromata of typical species of this genus are
crustaceous, cushion-shaped or hemispherical, and those s| ecies which have
an elongated or vertical stroma are better separated off as a distinct genus.
Saccardo placed them in the subgenus Podocrea, which Lindau afterwards
(/. c.) raised to generic rank. Atkinson (Bot. Gaz. xl. 401 (1905)) showed
that the generic name Podophyllum had meanwhile been proposed by Karsten
(Hedw. xxxi. 294 (1892)) and therefore had priority. As Podophyllum was
a name used by Linnaeus (1735 and Sp. PI. i. 723 (1753)) for a genus of
Berberidaceae it cannot be duplicated amongst the fungi, and Podocrea must
stand.
XYLABIACE^E.
XYLARIA POLYMORPHA (Pers.) Grev. Flor. Edin. 355 (1824).
On dead trunk in forest. Lower foot-hills by Momi River, Arfak Mts.
500'. Dec. 6151.
Distrib. World-wide.
XYLARIA DOMINGENSIS (Berk.) Sacc. Syll. i. 315 (1882).
On dead wood in forest, foot-hills by Momi River, Arfak Mts., 500'.
6136.
Distrib. West Indies, etc.
04
BASIDIOMYOETES.
AGARIC AC E^:.
OMPHALIA ARFAKENSIS Ramsbottom, sp. nov.
Tota alba. Pileo carnuloso, ex umbilicato infundibuliforme, margine primum
inflexo, crenato, 1-2 cm. lato ; stipite fistuloso, 1 cm. longo, 2 mm. crasso ; lamellis
decurrentibus, valde distantibus, postice latissimis (hinc triangulis), interdum
dichotomis, venoso-connexis ; sporis suballantoideis, 3-guttulatis, 6-7 p X 3-4 p ;
basidiis c. 25 p. X 5 p.
Caespitosa ad lignum putridum.
Bab. In forest, foot-hills by Momi River, ArfakMts., 500'. Dec. 6148.
XEROTUS CINNAMOMEUS Ramsbottom, sp. nov.
Mesopus ; cinnamomeus, coriaceus ; pileo convexo, centra papillate, c. 2 cm.
diam., ad centrum plicato sulcato, margine acuto, integro ; stipite subfistuloso,
deorsum cylindrico, sursum compresso sulcato, basi strigoso ; lamellis parcis (c. 10),
valde distantibus, non decurrentibus. latis, intermixtis brevioribus, interdum venae-
formibus ; sporis ellipsoideis, subgranulosis, 6-7 p X 4-5 p ; basidiis c. 50 p x 8 p.
Ad lignum putridum.
The radiating depressions on the upper surface of the pileus mark the
position of the gills below. A trsmsverse section of the stipes shows a pseudo-
sclerenchymatous tissue, which greatly simulates the sclerenchyma seen in
Gramineae and certain other glumiferous Monocotyledons.
Hab. In forest, lower foot-hills by Momi River, Arfak Mts., 500'. Dec.
6150.
P O L Y P O R A C E JR.
HKXAGOKA APIARIA (Pers.) Fr. Ii.picr. Syst, Myc. 497 (1838).
Foot-liills by Momi River, Arfsik Mts., plentiful, 400'. Dec. 6149.
Distrib New Guinea. India, Ceylon, Java, Philippines, etc.
Fungi Imperfecti.
LEPTOTHYRELLA SERICOLE/E Ramsbottom, sp. nov.
Pycnidiis ampbigenis, sparsis, dimidiat-o scutellatis, radiato-cellulosis, atris
c. 500 p diam., zona purpurascenti cinctis, sporis fusoideo-ellipsoideis vel clavulatis,
rectis vel curvulis, hyalinis, multiguttulatis, continuis, dein medio uniseptatis. non
constrictis, 25-35 p x 4-5 p.
In foliis Sericoleae.
On Sericolea arfakensls Gibbs (p. 147).
65
BRYOPHYTA. (A. GEPP.)
The number of species brought home is but small, and obviously fails
to represent adequately the richness of the moss-flora in the districts visited.
Attention was exclusively directed to the collection of fruiting plants.
HEPATICJE.
*RICCAHDIA MAXIMA Schiffn. He[>at. Flor. von Buitenzorg, i. 57 (1900).
Arfak Mts., Angi lakes, forest patch by ? lake, on dead wood in forest,
7000'. Dec. 5680.
Distrib. Java ; Sumatra.
*MARCHANTIA POLYMORPHA Linn. Spec. Plant, ed. i. 1603 (1753).
Arfak Mts., Angi lakes, on open gravel-bank by ? lake, 7000'. Dec.
5906.
Distrib. Cosmopolitan.
MUSCI.
SPHAGNUM JUNGHUHNIANUM Doz. et jMolk. Bryologia Javanic-a, i. 27, tab. 18
(1855).
Arfak Mts., Angi lakes, in open marsh by ? lake, 7000'. Dec. 5967.
Distrib. New Guinea. Malay Islands ; Formosa ; Japan ; India.
SPHAGNUM NOVO-GUINEENSE Fleisch. et Warnst. in Engler, Pflanzenreich,
Heft 51, Sphagnales, p. 520 (1911); Nova Guinea, xii. (1914) 127,
t. xxxiv. B.
Arfak Mts., ridge running up to Angi lakes, terrestrial in moss-grown
forest, 8000'. Dec. 6006.
Distrib. New Guinea (D.S.W., Goliath-Gebirge, de Kock}.
*FUNARIA CALVESCENS Schwaegr. Spec. Muscorum, Suppl. i. sect. 2, 77,
tab. 65 (1816).
Arfak Mts., Angi lakes, on ground amongst bracken, "here burnt, on
bank of ? lake, 7000'. Dec. 5938.
Distrib. Warmer regions of the world.
*RHODOBRYUM GIGANTEUM Paris, Index Bryolog. 1116 (1898).
Arfak Mts., Angi lakes, abundant on ground in isolated t'orrst patch by
? lake, 7000'. Dec. 5892. " Very handsome moss."
Distrib. Malay Islands ; India ; Bourbon ; Hawaii.
RHIZOGONIUM SPINIFORME Bruch in Flora, 1846, 134.
Arfak Mts., Angi lakes, on living tree in isolated forest patch by ? lake,
7000'. Dec. 5979. Also, on dead wood in same forest patch, 7000'. Dec.
5894.
Distrib. New Guinea (N.E.). Throughout the tropics.
DAWSONIA GIGANTEA C. Mull, ex Geheeb in Bibliotheca Botanica, Heft 44.
13 (1898).
Arfak Mts., abundant in moss-grown forest of ridge running up to Angi
Likes, in mossy forest sloping down to ? lake and in the open by same,
7000-8500'. Dec. 5523. Angi lakes on open banks by edge of ? lake and
in forest, 7000'. Dec. 5935.
Distrib. New Guinea (D.N.W., Mt. Arfak, Beccari ; D.S.W., Went-
Gebirge, von Roemer ; Hubrecht-Gebirge, van Noulmys ; Siriwo-Fluss,
Janowsky) .
DAWSONIA BECCARII Brotherus et Geheeb in Bibliotheca Botanica, Het't 44,
13 (1898).
Arfak Mts., ridge running up to Angi lakes, steep open slopes of gravel,
8000'. Dec. <$ & ? together. 5521. Ridge running up to Angi lakes
in open gravelly spaces, 8500'. Dec. 6005.
Distrib. New Guinea (D.N.W., Mt. Arfak, Beccari).
RHACOPILUM SPECTABILE Reinw. et Hornsch. in Nov. Act. Acad. Cses. Leop.
xiv. 721, tab. 40 (1828).
Arfak Mts., ridge running up to Angi lakes, in forest, 8000'. Dec.
6121, 6122.
Distrib. New Guinea (D.N.W., Mt. Arfak, Beccari ; D.S.W., Went-
Gebirge, von Roemer ; N.E.). Malay Islands to Fiji and New Caledonia.
SPIRIDENS REINWARDTI Nees ab Es. in Nov. Act Acad. Cses. Leop. xi. 143,
tab. 17 (1823).
Arfak Mts., Angi lakes, in isolated forest patch by $ lake, 7000'. Dec.
5591.
Distrib. New Guinea (D.N.W., Mt. Arfak, Beccari ; S.E., Armit,
MacGregor, Miclwlitz ; N.E.). Malay Islands.
ENDOTRICHELLA ARF^KIANA C. Mull, ex Geheeb in Bibliotheca Botanica,
Heft 44, 16, tab. 14 (1898).
Arfak Mts., Angi lakes, with Rliizogonium spiniforme on living tree
in isolated forest patch by ? lake, 7000'. Dec. 5979 pro parte.
Distrib. New Guinea (D.N.W., Mt. Arfak, Beccari}.
TAXITHELIUM SUBSTIGMOSUM Broth, in Engl. & Prantl, Nat. Pflanz. i. Abt. 3,
1092 (1908).
Arfak Mts., Angi lakes, small forest by ? lake, on dead wood, 7000'.
Dec. 5965.
Distrib. New Guinea (D.N.W., MacCluer Bay, Naumann ; N.E.).
6?
ECTROPOTHECIUM ARFAKENSE Broth, et Geheeb in Bibliotheca Botanica,
Heft 44, 24 (1898).
Arfak Mts., Angi lakes, on dead wood, in isolated forest patch by
? lake, 7000'. Dec. 5904.
Distrib. New Guinea (D.N.W., Mt. Arfak, Beccari).
HYPNODKNDRON DIVEKSIFOLIUM Broth, et Geheeb in Ofvers. Finsk. Vet. Soc.
Forh. xl. 191 (1898).
Arfak Mts., Angi lakes, forest patch by ? lake, carpeting in forest, 7000'.
Dec. 5667.
Distrib. New Guinea (S.E., Mt. Dayman, Artnii).
PTERIDOPHYTA. (A. GEPP.)
FILICALES.
The references to the descriptions of the following ferns are to be found
in Christensen's ' Index Filicum,' 1905-13.
*TRICHOMANES DIGITATUM Sw.
Arfak Mts., S.W. ridge running up to Angi hikes, 9000'. epiphytic in
moss-grown forest. Dec. 6000.
Distrib. Malay Islands to the Mascarenes and New South Wales.
*TRICHOMANES PALMATIFIDUM K. Mull.
Arfak Mts., S.W. ridge, Angi lakes, 8000', epiphytic in moss-grown
forest, Dec. 5520:
Distrib. Java.
TRICHOMANES PALLIDUM Bl.
Arfak Mts., Koebre Mt,, 8000-9000', epiphytic in forest. Dec. 5728.
Distrib. New Guinea (D.N.W., Arfak Mts., Beccari ; D.S.W., Hellwig-
Gebirge, von Roemer ; Mt. Carstensz, Kloss ; S.E.). Trop. Asia. Polynesia.
TRICHOMANES APHLEBIOIDES Christ.
Arfak Mts., Angi lakes, 7000', epiphytic in forest by ? lake, 5947 ;
forest patch by ? lake, epiphytic, 6137. Dec.
Distrib. New Guinea (D.N.W., R. Begowri, Gjellerup ; D.S.W., Noord
Rivier, Versteeg).
TRICHOMANES MEIFOLIUM Bory.
Arfak Mts., Angi lakes, 7000', epiphytic in forest patch by ? lake. Dec.
5678.
Distrib. New Guinea (D.N.W., Arfak Mts., Beccari ; D.S.W., Mt.
Carstensz, Kloss}. Malay Islands to Polynesia and Reunion.
68
HYMENOPHYLLDM AUSTRALE Willd.
Arfak Mts., Angi lakes, 7000', epiphytic in forest patch by ? lake.
Dec. 5893.
Distrib. New Guinea (D.N.W., Arfak Mts., Beccari- N.E.). India.
Malay Islands. Australasia.
*HYMENOPHYLLUM PANICULIFLORUM Presl.
Arfak Mts., Angi lakes, 7000', epiphytic in forest patch by $ lake.
Dec. 6141.
Distrib. Malay Islands and Japan.
*HYMENOPHYLLUM SALAKENSE Racib.
Arfak Mts., Angi lakes, 7000', epiphytic in forest patch by ? lake.
Dec. 6138.
Distrih. Java.
*HYMENOPHYLLUM KUHZII Prantl.
A rials. Mts., Angi lakes, 7000', epiphytic in forest patch by ? lake.
Dec. 5679.
Distrib. Java.
HYMKNOPHYLLUM (LEPTOCIONIUM) CEKNUUM Gepp, sp. nov.
Rhizoma longe repens pilosum; stipites remoti erecti pilosi obsolete alati, ad
10 cm. longi, 1 mm. crassi. Frons 20-22'5 cm. longa. 5-7'5 cm. lata, lanceolata,
rhachi omnino sed anguste alata pilosa ; pinnis alternis 20-jugatis contiguis ovato-
lanceolatis pinnatifidis, costa alata pilosa ; pinnulis 1-2-dichotome lobatis ; segmentis
ultimis planis serrulatis 2 inin. long., 0'5 nun. lat., costula anguste et dentate cristata.
tSori in lobis brevibus pinnarum superiorum terminales, 1-8. Indusium vix ad
medium divisum, valvis rotundatis integris, basi obovatum, longitudinaliter
cristatum. Receptaculum exserturn.
Hab. Arfak Mts., Angi lakes, 7000', terrestrial in spinneys by ? lake.
Dec. 5964.
The tall narrowed frond is in general habit somewhat like IF. Zol-
lingerianum as figured by Van den Bosch (Hymen. Javan. t. 50), and bears
about 20 pinna) on each side. In the dried specimens the frond-apex is
cernuons and the pinnae complicato-decurved. The narrowly winged stipes
and rhachis, tlie flat serrulate ultimate segments, the narrowly dentato-cristate
costules, and the hairiness of stipes, rhachis, costfe, and costulesare characters
to be noted.
HYMENOPHYLLUM (LEPTOCIONIUM) CINCINNATUM Gepp, sp. nov.
Rhizoma longe repens pilosum ; stipites remoti erecti pilosi obsolete alati,
3-5 cm. longi, 0'5 mm. crassi. Frons circ. 5 cm. longa, 2 cm. lata, lanceolata.
rhachi alata pilosa ; pinnis alternis 10-12-jugatis contiguis (circ. 1 cm. longis,
0'5 cm. latis) pinnatifidis ; pinnulis 1-2-dichotome lobatis ; segmentis ultimis planis
paucidentatis, 2-3 mm. longis, 0'5 cm. latis. Sori solitarii in lobis brevibus pin-
narum terminates. JiH/itsimn ad medium divisum, valvis obtusis truncatisve integris,
basi obovatum, parce longitudinaliter cristatum. JReceptaculum exsertum.
Hab. Arfak Mts., S.W. ridge, Angi lakes, 8500', epiphytic in moss-grown
forest. Dec. 5989.
This species has sori much as in H. cernuum (No. 5964), but is not one-
quarter the size, and is not cristate on the costules of the segments. It
approaches //. holochilum Van den Bosch (Hymen. Javan. t. 34), hut has a
stouter, more hairy rhizome and stipes (the latter winged throughout its
length), and much narrower ultimate segments. The dried plants are very
convolute.
OYATHEA ARFAKENSIS Gepp, sp. nov.
Stipes circ. 18 cm. longus muricatus inferne fuscus supeme griseo-purpurascens ;
rhachis purpurea baud nitens supra ferrugineo-pubescens glande juxta cujusque pinnae
basin instructa. Frons lineari-lanceolata, circa 80 cm. longa, 13 cm. lata, bipinnata ;
pinnae alternae 35-jugatae stipitatae lanceolatae, 8 cm. long*, 2 cm. lata?. obtusae,
pinnnlis 15-20-jugatis, stipitatis 1 cm. longis, 0'4 cm. latis, imbricatis oblongis
obtusis, inferioribus paucipinnatis (segmentis rotundatis), superioribus lobatis vel
crenatis vel integris (versus apicem) ; rhachibus pinnarum supra pubescentibus, infra
squamulis albidis sparse instructis. Sori costales 4-5-jugati ; venuke pinnularum
5-6 parum conspicuae simplices f urcatuave. Textura coriacea ; frons supra griseo-
purpurea, infra pallide brunnea. Indusium persistens hemisphericum.
Hab. Arl'ak Mts., S.W. ridge, Angi lakes, 8000', undergrowth in moss-
grown forest. Dec. 6008. " Tree-fern."
. Tlie pinnae are not contiguous, but are attached to the rhachis at intervals
of about 3 cm. In the dried specimen the shortly-stalked opposite rows of
pinnulse are appressed to one another and directed upwards (apically),
displaying the sori and concealing the upper surface.
CYATHEA FUSCA Baker.
Arfak Mts., Angi lakes, 7000', undergrowth in forest by ? lake. Dec.
5932. "Tree-fern, 3 in. in height (pinnae)."
Distrlb. New Guinea (S.E.).
ALSOPHILA ANGIENSIS Gepp, sp. nov.
Stipes (?). Frons tripinnatifida ; rhachis purpureo-fusca sparse aculeata impolita
sparse et breviter atro-hispida ; pinna brevi-stipitata, 37 cm. longa, 19 cm. lata,
oblongo-lanceolata ; rhachis pinnse supra atro-tomentosa purpureo-fusca, infra sulcata
furfuraceo-squamulata ; pinnula? suboppositse 30-jugatai contiguae imbricataeve
sessiles horizontales lineari-lanceolata} acuminatse 1'5 cm. lataa, fere ad costam
pinnatisecta? ; costa pinnulae supra fulvo-tomentosa, infra sulcata furfuraceo-squamu-
lata ; segmenta 2G-jugata lineari-oblonga falcatula integra 3 mm. lata, supra
f usco-purpurea glabra, infra glauca ; costula segmenti supra glabrum, infra pauci-
pilosa et squamulis flavidis timbriatis versus basin vestita ; venulae inconspicuae
tenues 14-jugat, 1-2-furcata?, pauci-pilosa?. Sori (?). Textura coriacea.
70
[Jab. Arfak Mts., Angi lakes, 7000', spinney by ? hike, undergrowth in
forest. Dec. 5968. " Tree-fern, 3 m. in height, brown paleze on young
stein and fronds. Brown and with thorns on old."
This plant is represented by two pinnae which are entirely sterile. It is
tentatively referred to Alsophila. In some respects it recalls A. glauca, but
differs in having the pinnules crowded together and imbricated, and the
rhachises dull and unpolished. The pinnules are inserted on the pinna-
rhachis at intervals of about T25 cm. and overlap ; the middle pinnules are
horizontal, tlie lower are deflexed.
ALSOPHILA ARFAKENSIS Gepp, sp. nov.
Stipes (?). Frons tripinnatifida, pinnis remotis alternis stipitatis. Rhachis
(versus apicem) pallida, supra breviter tomentosa, infra glabra ruguloso-aspera ;
rhachis pinnulae similis est. Pinnae infimae 28 cm. longse, 9-10 cm. latse. lineari-
lanceolat* breviter acuminatae ad apicem pinnatifidae. Pinnulae rt 12-jugatae remotse
alternae stipitatse, usque ad 5 cm. longse, 1 cm. latse, lineari-lanceolatai obtuse
acuminatse, ad | pinuatifidse ; costa pinnulae supra tomentosula, sed versus apicem
glabra, infra rugulosa squamulis paucis lauceolatis rubris ornata ; segmenta 12-
jugata (apice excluso), 3'5 mm. lata, oblonga obtusissima crenata margine
recurvata ; costula segmenti supra glabra, infra squamulata ; venulse 5-jugatae
furcata? simplicesve. Sori 4-5-jugati prope costain dispositi, segmenti latitudinem
haud obtegentes. Textura coriacea.
Hal. Arfak Mts., S.\V. ridge, Angi lakes, 7000-8500', common under-
growth in moss-grown forest. Dec. 5990. Also common about ? lake,
in forest. " Slender tree-fern, stem 1 dm. in diameter and 1 m. in height.
Fronds 1 m. long."
The material consists of the two .lowest pinnoe and the top 37 cm. of a
frond. A small dark gland is present at the base of the stalks of the
pinnules.
*DRYOPTERIS (LASTHEA) BEDHOMEI 0. Kuntze.
Arfak Mts., Angi lakes, 7000', common in open marsh by ? lake. Dec.
5939. " Creeping rhizome."
Distrib. Philippine Islands ; Ceylon ; South India ; South China.
DKYOPTEHIS (LASTRKA) VILLOSIPES Gepp, sp. nov.
Rhizoma erectum squamulis lineari-lanceolatis ferrugineis dentatis vel subintegris
vestitum. Stipites 12'5 cm. longi, c. 1 mm. crassi, atro-pui-purei, inferne squamulis
angustis capillaribus c. 1 mm. longis ferrugineis vestiti, superne breviter et sparse
pubescentes et squamulis paucis lineari-lanceolatis ornati (ut etiam rhachis). Frons
lanceolata 10 cm. longa, 3 cm. lata; pinnae subcontiguae alteruae plerumque
brevi-stipitatae, versus frondis apicem sessiles, elliptico-lauceolatse 30-jugatae,
horizontales, inferne 1'3-1'7 cm. longae, 0'4-0 g 5 cm. latae, fere ad if pinnatiticlae,
ad apices crenatae vel subintegrae obtusss; segmenta 0-8-jugata approximata sub-
falcata obtusa, segmento infimo superiori oblongo quain reliquis majori interdum
71
libero ; venulse segment! paucae pinnate dispositae inconspicuae. Sort singuli
plerumque ad media segmenta, prope pinnae costam dispositi, indusio persistente.
Textura coriacea ; lamina superne fusca, inferne griseo-viridis.
Hal. Arfak Mts., Koebre Mt., 7000-8000', epiphytic in forest. Dec.
5627.
The lower pinnules are slightly deflexed, and the lowest pair are a trifle
shorter than the pair above them. The plant differs in every respect from
I), viscosa.
POLYBOTRYA ARFAKENSIS Gepp, Sp. nOV.
Stipes paleis linearibus brunneis deciduis 0'5 cm. longis vestitus et earum
cicatricibus muricatus, 15 cm. longus, atro-purpureus. Frons circa 1 m. longa,
40 cm. lata, bipinnata obovato-lanceolata ; rhachis atro-purpurea paleacea muricata
(velut stipes) ad apicem haud evoluta, guperne breviter ferrugineo-tomentosa, inferne
parum furfuracea, pinnas circa 9 alternas remotas dimorphas (inferiores 6 fertiles,
superiores 3 steriles) gerens; pinnae brevi-stipitatae ad apicem abrupte (haud
evolutse), steriles circa 30 cm. longse, 13 cm. latae, lanceolatae, fertiles circa 25 cm.
longse, 7'5 cm. latae ; rhachis pinnae superne ferrugineo-tomentosa ; pinnulse alternae
circa 12-jugatae stipitatae ; pinnulae steriles 6'5 cm. longae, 1'2 cm. latae, oblongo-
lanceolatae acuminatae apice serrato, usque ad \ pinnatifidae, lobis (fere 20-jugatis)
rotundatis crenulatis, venulis in lobis pinnatae dispositis c. 3-jugatis; pinnulae fertiles
3'5 cm. longae, 4-8 mm. latse, lineares usque ad costam pinnatisectae, lobis parvis
oblongis sessilibus propter soros copiosos omnino obtectis. Textura subcoriacea.
Lamina superne purpureo-brunnea, inferne pallidior glabra.
Hal. Arfak Mts., S.W. ridge, Angi lakes, 7000', climbing in forest,
clasping trunk to top. Dec. 5984. " Fronds 1 in. long, sterile like barren
portion of fertile frond ; fertile frond distinct."
This fern is remarkable for its dimorphous fronds and pinnae and for the
arrested growth of its frond-apices. The material consists of one frond,
the six lower pinnse of which are fertile, and the three upper are sterile.
The apex of the main rhachis has failed to attain its full development, as also
have the apices of the secondary rhachises (pinnse both fertile and sterile).
Hence, the proper apices are absent. Whether the apical growth has been
temporarily interrupted or permanently nrrested, and whether the arrest of
growth is due to injury or is of normal occurrence in the life-history of the
plant is uncertain. The lower pinnse are shorter than the others. The
sterile pinnules are inserted at intervals of about 2 cm., the fertile at about
1*5 cm. The fertile pinnules recall those of Osmunda javanica in shape, but
not in arrangement. Dimorphism of frond and pinnse occurs in the tropical
American P. osmundacea.
P. arfakensis climbs by means of its rhizome and puts out its fronds at
right angles to the axis of the tree up which it climbs,
72
DIPTEKIS CONJUGATA Reinw.
ArFak Mts., Angi lakes, 7000', growing on bank by edge of ? lake, in
open. Dec. 5913. " Also abundant on open steep gravel slopes on S.W.
ridge, running up to lakes from 8000-8500'."
Distrib. New Guinea (D.N.W., Arfak Mts., Andai, Beccari ; D.S.W.,
Noord Rivier, Versteeg : submontane region, von Roemer : Mt. Carstensz,
Kloss ; S.E.). Asia. Polynesia.
OLEANDRA CUSPIDATA Baker.
Arfak Mts., Angi lakes, 7000', common on edge of forest patch by
? lake, growing in clumps. Dec. 5559. " Up to 2 in. in height, leaves
in interrupted whorls up the stem Also seen on S.W. ridge."
Distrib. New Guinea (D.N.W., Arfak, Bcccari ; D.S.W., Noord Rivier,
Versteeg ; Mt. Carstensx, Kloss}.
NEPHROLBPIS ACUMINATA Kuhn.
Arfak Mts., S.W. ridge, Angi lakes, 7000', climbing in forest. Dec.
6123. "Fronds with apical fertile portion or with entire fertile fronds."
Distrib. New Guinea (D.S.W., Mt. Carstensz, Kloss). Malay Islands.
Perak.
HUMATA PUSILLA Carr.
Arfak Mts., Angi lakes, 7000', epiphytic in forest, spinneys by ? lake.
Dec. 5960.
Distrib. New Guinea (N.E.). Melanesia.
HUMATA NEOGUINENSIS C. Chr.
Arfak Mts., Angi lakes, 7000', epiphytic in forest or creeping on edge.
Dec. 5588. " Sterile and fertile frond."
Distrib. New Guinea (D.S.W., Hellwig-Gebirge, von Roemer).
HUMATA ALPINA Moore.
Arfak Mts., Angi lakes, 7000', epiphytic in forest patch by ? lake.
Dec. 5674.
Distrib. New Guinea (D.S.W., Low country and Hellwig-Gebirge,
von Roemer; N.E.). Malay Islands. Polynesia.
*DAVALLIA DISSECTA J. Sm.
Arfak Mts., Angi lakes, 7000', epiphytic in forest patch by ? lake.
Dec. 5592.
Distrib. Java ; Sumatra.
DAVALLIA (PROSAPTIA) SCHLECHTERI C. Chr.
Arfak. Mts., Koebre Mt., 8000-9000', epiphytic in forest slopes. Dec.
5634, 5640, 5625.
Distrib. New Guinea (D.S.W., Mt. Carstensz, Kloss ; N.E.).
73
DAVALLIA (PROSAPTIA) CONTIGUA Spr.
Art'ak Mts., Angi lakes, 7000', epiphytic in isolated Forest patch by ?
lake. Dec. 5895.
Distrib. New Guinea (D.S.W., Noon! Rivier, Virsteey : Mt. Carstensz,
Klo948.
Distrib. New Guinea (D.S.W., Hellwig-Gebirge, von Roemer; S.E.).
Malay Islands. Polynesia.
R
76
DRYOSTACHYUM SPLENDENS J. Smith.
Arfak Mts., Angi lakes, terrestrial on edge of spinneys by ? lake, 7000'.
Dec. 5970.
Distrib. New Guinea (D.S.W., coastal lowlands, von Roemer}. Malay
Islands.
CHEIROPLEURIA BICUSPIS Presl.
Arfak Mts., lower part of S.W. ridge, terrestrial in high forest, 5000-
6000'. Dec. 6134. Fertile frond only.
Distrib. New Guinea (S.E.). Java ; Philippine Islands ; Formosa ;
Liu-kiu Islands.
GLEICHENIA VULCANIOA Bl.
Arfak Mts., Koebre ridge, terrestrial on burnt open summit, common,
9000'. Dec. 5611. Angi lakes, abundant at edge of forest and spinneys
by ? lake, 7000'. 5727. S.W. ridge, terrestrial on open steep gravel slopes,
8000-9000'. 5996.
Distrib. New Guinea (S.B.). Malay Islands.
GLEICHENIA GLAUCA Hook.
Arfak Mts.. Angi lakes, abundant on edge of forest by ? lake, 7000'.
Dec. 5668. " One frond climbing up to 7 m."
Distrib. New Guinea (D.S.W., Mt. Carstensz, Kloss). Asia. Australia.
Polynesia.
*GLEICHENIA L^VIGATA Hook.
Arfak Mts., Angi lakes, abundant on edge of forest patch by ? lake,
7000'. Dec. 5577. "3 in. in height, almost arboreal in habit. Leaves
glaucous underneath."
Distrib. Malay Islands.
GLEICHENIA LINEARIS Clarke.
Arfak Mts., Angi lakes, running up trees at edge of forest patch by
? lake, 7000'. Dec. 5575. " Massed/' S.W. ridge, common in forest,
where more open, 7000-8000'. 5991. "Scrambling up to 7 m. ; half
frond/'
Distrib. New Guinea (D.N.W., Arfak Mts., Beccari ; D.S.W., Noord
Rivier, Verstteg ; S.E.). Tropics and subtropics.
*ScHiz^A MALACCANA Baker.
Arfak Mts., Angi lakes, common on open drier and gravelly parts of
marsh by ? lake, 7000'. Dec. 5945. S.W. ridge, 850^ in moss in
forest undergrowth. 6011.
Distrib. Malay Islands. Burma.
77
LYCOPODIALES.
LYCOPODIUM SERRATUM Tlmnb.
Arfak Mts., Angi lakes, forest patch by ? lake, terrestrial in humus,
7000'. Dec. 5724.
Distrib. New Guinea (N.E.). Asia. Polynesia. Mexico. Bourbon.
LYCOPODIUM SQUARROSUM Forst.
Arfak Mts., Angi lakes, spinneys by ? lake, 7000', epiphytic in torest.
Dec. 5725.
Distrib. New Guinea (N.E. ; S.E.). Asia. Polynesia. Mascarene
Islands.
LYCOPODIUM PINIFOLIUM Bl.
Arfak Mts., Angi lakes, epiphytic under Araucaria forest by ? lake,
7000'. Dec. 5936.
Distrib. New Guinea (D.S.W., Mt. Carstensz, Kloss ; N.E.). Java ;
Borneo.
LYCOPODIUM CERNUUM L.
Arfak Mts., Angi lakes, common on open marsh by ? lake, 7000'. Dec.
5658, 5925. S.W. ridge, creeping in forest and on open steep slopes, 8000-
9000'. 5995.
Distrib. New Guinea (D.N.W., Arfak Mts., Beccari ; D.S.W., Mt.
Carstensz, Kloss ; N.E. ; S.E.). Tropics and some subtropics.
LYCOPODIUM CASUARINOIDES Spring.
Arfak Mts., Angi lakes, scrambling on edge of forest by ? lake, 7000'.
Dec. 5944.
Distrib. New Guinea (D.S.W., Mt. Carstensz, Kloss). Trop. Asia.
LYCOPODIUM CLAVATUM L., var. WALLICHIANUM Spring.
Arfak Mts., Angi lakes, open gravel bank by ? lake, 7000'. Dec.
5911. Open marsh by ? lake. 5556.
Distrib. The species is recorded for New Guinea (D.N.W.*, Arfak Mts.,
Beccari ; S.E.). The species is cosmopolitan in temperate regions. The
variety occurs in Java and India.
LYCOPODIUM COMPLANATUM L.
Arfak Mts.. Angi lakes, by open marsh and bank of ? lake, 7000'. Dec.
5912.
Distrib. New Guinea (D.N.W., Arfak Mts., Beccari). Asia. Polynesia.
Africa, America; but mostly in the north temperate zones.
H2
78
LYCOPODIUM VOLUBILE Forst.
Arfak Mts., Angi lakes, scrambling up to 10 in. in forest spinneys by
? lake, 7000'. Dec. 5954. S.W. ridge, scrambling- in moss-grown forest,
7000-8500'. 5982.
Distrib. New Guinea (D.N.W., Arfak Mts., Beccari ; S.E.). Malay
Islands. Australasia. Polynesia.
PsiLOTUM FLACCIDUM Wall.
Arfak Mts., Koebre Mt., epiphytic in forest, 7500'. Dec. 5633.
Distril. New Guinea (D.N.W., Arfak Mts., Beccari ; D.S.W., Noord
Rivier, Versteeg ; Mt. Carstensz, Kloss ; N.E.). Tropics.
SELAGINELLA ANGUSTIRAMEA F. Muell. & Baker.
Arfak Mts., Angi lakes, terrestrial on edge of forest by ? lake, 7000'.
Dec. 5726. " Growing in patches."
Distrib. New Guinea (N.E. ; S.E.).
SPERMATOPHYTA.
CONIFERS.
TA x ACE^E.
DACRYDIUM NOVO-GUINEENSE Gibbs, sp. nov.
Arbor parva, dioica, in statu juvenili erecta ; ramulis foliis longioribus 5-seriatim
imbricatis pncditis, postea ramulis gracilioribus folia breviora gignentibus. Folia
in statu plantae juvenili laxiuscula, squarrosa, acicularia, incurvato-pungentia, sectione
triangulare ; folia in statu adultiori parva, dense imbricata, 5-seriata squami-
formia, triangularia, apice pungentia, facie plana, dorso carinata ; folia ramoruin
fertilium arete applicata, rhombica, acuta, dorso carinata, sectione sub-tetragona.
Strobili masculi ignoti. Strobili feminei ad apicem ramulorum brevissimorum
axillarium erecti, parvi ; bractese 24, lineares, apice apiculata?, incurvat^, carinata?,
margine membranaceae, basi incrassata?, demum carnosse, rubrse, bractea fertilis
unica, terminalis. Squama ovuligera fere usque ad basin libera, sub anthesi teres,
cucullata. Ovulum in statu pollinifero liberura, squama ovuligera adhuc immatura
circumdatum. Seminn erecta vel obliqua, bracteam summum sterilem superantia,
ovoideo-angulata, viridia, nitida, basi ad | vel i squama ovuligera incrassato-
cupuliforme cincta.
Hob. Arfak Mts., crest of ridges and forest by ? lake, 7000-9000'.
Seedling, ? lake, 7000'. Dec. 5508. Koebre" ridge, open summit, 9000'.
? . Fr. with mature foliage. 5648.
Foliage of seedling branches 1 cm. across, the leaves 4 mm. by '5 mm. ;
tbat of more adult branches 1*5 mm. across, the leaves 1*5 by *4 mm. ;
foliage of fruiting branches 2 mm. across, the adpressed lozenge-shaped leaves
1*7 mm. by 1 mm. Strobilus 7 mm. by 3 mm., imbricating, with bracts
79
FIG. 3.
Dacrydium noro-fluitieense Gibbs. A. Youth foliage ; B. Mature $, nat. size; C. Ovule
in pollination-stage, ovuliferous scale appearing; D. a. Lateral, b. ventral view;
E, F. Stages in development of ovule; H, I, J, K. Stages in development of
strobilus; L. Seed with ovuliferous scale still attached; M. Strobilus, showing
swollen bases of sterile bracts forming strobilus (seed shed) ; M.c. Ovuliferous scale.
80
gradually increasing in length till at the apex they are 3 mm. by - 5 mm., of
which the apical bract only is fertile ; the mature strobilus, with swollen
bract-bases, is 7 mm. by 4 mm., not including the seed. Ovuliferous scale
is 2 mm. in height and 2 mm. broad, persisting in the axil of the fertile
bract after the seed is shed. Seed 5 by 2 mm. All measurements of the
fruiting-stages and drawings are from material in formalin. Pollination
drops were seen on the young ovules.
The mature fruiting-form of this species was only seen in one or two
isolated examples on the open summit of Koebre* ridge, small trees +10 m.
high, with short trunks and rounded crowns of rigid ascending branches,
bearing numerous small red and fleshy cones. In younger stages this plant
was very abundant on the crest of the S.W. ridge and in the forest round
the ? lake. The foliage of the seedlings is plumose in character to '50 in.
in height, when the small adpressed scaly leaves begin to appear.
In the peculiar habit and the strap-shaped laminae of the bracts composing
the strobilus, this species is quite distinct from known members of the genus.
PODOCARPUS PAPUANUS Ridl. in Trans. Linn. Soc. ser. 2, Bot. ix. 158, quoad
spec. Klossianum de Gamp III apportatum, non aliorum.
Arfak Mts , Angi lakes, common on the surrounding ridges and in
spinneys by the ? lake, 7000-9000'. Fl. ? ., Fr. (yg.). Dec. 5540.
Diittrib. New Guinea (D.N.W., Arfak Mts., Hatam, Beccari ; D.S.W.,
Mt. Carstensz, Kloss).
A fine tree, 30 m. in height, very like P. imbricatus Bl. in habit and in
the dimorphic foliage, both seedling and youth form of foliage being identical
tn both species in fact, like Beccari, I took the young plant for that species
in the field, but the foliage of the mature tree is more spreading and distinct,
the scales of the $ cones differ in shape, while the ? cones are larger and
very glaucous in appearance. The fusion of the lamina of the fertile bract
with the ovuliferous scale is also less complete than is the case in P. imbri-
catus and the position of the seed is more oblique. Dr. Beccari most kindly
sent me some of his Arfak material, published as P. imbricatus^ which, on
comparison, proved identical with the above.
The description of this plant being limited to the mature foliage
and one <$ cone, I append what is necessarily wanting in the original
diagnosis :
Arbor alta ; truncus teres, erectus ; rami fere penduli, copiose ramulosi ; ramuli
flexuosi, graciles. Folia dimorpha, juvenilia 1 cm. longa, 2 mm. lata, biseriatim
expansa, plana, anguste linearia, mucronato-pungentia, decurrentia, apicem et basin
versus decrescentia. Strobili feminei 1*2 cin. longi, '5 mm. lati, ad apicem ramu-
lorum brevium erecti. Bractece 2 ve!3, 2 mm., glaucescentes, laminae teretiusculae,
apice obtusae, 2 mm. longse, '8 mm. latse (inter se connatse), basi incrassata, verru-
culosa 4-5 mm. longa, 1 vel 2 superioribus fertilia, lamina bracteis fertilioribus cum
FIG. 4.
Podocarjnts pupuanus llidl. A, B. Youth foliage ; C. Mature foliage, sterile ; D. Branch,
bearing 2 $ strobili, one allowing two fertile bracts (pollination-stage) ; E. $ strobilus
(fertilization-stage) : all uat. size. G, H, I. Same stages : J. Strobilus with oblique
ovule ; K. Mature foliage : all X 4.
82"
squama ovuligera tota longitudine connata. Squama ovuligera cum ovulo connata,
viridis, nitida. Ovula juvenilia in statu archegoniale, oblique erecta, bracteas
superantia, 7 mm. longa, 5 mm. lata.
All the measurements given are from formalin material. On a mature
tree in fruit small branches of the young foliage occurred up the stem. The
? strobili were in the pollination to the archegonial stage, and may be com-
pared with similar stages in P. imbricalus (Gibbs, in Ann. Bot. xxvi. (1912)
pi. xlix. figs. 1-6). The terminology given on p. 518, /. c., is that followed on
the present occasion.
There seems little doubt that Giulianetti's specimen, included by
Mr. Ridley in his description of P. papuanus, represents P. imbricatns Bl.,
as Dr. Stapf had already nnmed it on the Herbarium sheets at Kew ; the two
? cones on the specimen prove the correctness of this determination. These
cones were not seen by Reorders (Nova Guinea, viii. (1911) 615).
PODOGARPUS RUMPHII Bl. Rumphia, iii. (1847) 214 ; Becc. Malesia, i. 179.
Arfak Mts., S.W. ridge, in forest from 7000-9000'. Veg. Dec. 5985.
Distrib. New Guinea (D.N. W., Arfak, Beccari ; D.S.W., Lobo, Zippelius ;
N.E.). Moluccas, Amboina, Celebes.
A tree, + 16 m. high, very common on the crest of the ridge, but not seen
in fruit ; therefore the determination must remain uncertain. Leaves over
2 dm. long.
PHYLLOCLADUS HYPOPHYLLUS Hook. f. Ic. PI. 889 ; F. Muell. in Trans. Roy.
Soc. Viet. i. (1888) 32.
Arfak Mts., common on ridges. Koebre ridge, open summit, 9000'.
? (very yg.). Dec. 5657. Seedling, 5657 a. S.W. ridge, foliage glaucous.
? (yg.). 5992.
Distrib. New Guinea (S.E.). N. Borneo, Philippines.
Very plentiful on ridges and in the forest, showing glaucous and non-
glaucous foliage as on Kinabalu. Only the remains of some mature cones
were collected. I fail to distinguish any difference between the above species
and P. protractus Pilg. It is a very variable plant like other Phyllocladus
spp., differing according to the age of the plant and whether the fertile
branches occur on the old or the young wood. The series of variations
obtained by me on Kinabalu are duplicated in the Arfak material, and appear
also marked in the large amount of material from the Philippines available
at Kew for comparison.
P I N A C E M.
AGATHIS DAMMAKA (Lum) ; A. Rich. Conif. Ixxxiii. t. 19 ; Rumph. Herb.
Amboina, ii. 174, t. 57 ; Becc. Malesia, i. 180 ; Warb. Monsunia, i.
182 ; K. Laut. in Engl. Bot. Jahrb. 1. (1913) 48.
Arfak Mts., S.W. ridge, 5000', in high forest. ? . Dec. 6127.
Veg. (yg. plant). 5747.
Distrib. New Guinea (D.N.W., Waigiou, La Billardiere ; Ramoi, Jobi
Island, Ansus, Beccari}. Auiboina, (Celebes, Borneo, Java, Philippines,
Malay Peninsula.
A beautiful tree, about 40 in. high, with straight white shaft branching
at the very top into a small, not very spreading crown with yellowish-
green foliage.
The distinct habit of these trees, of which three to four were growing
near together, the crowns rising above the surrounding forest, was especially
noted, to be confirmed by the excellent description given by Rumphius above,
quoted by Parlatore (DC. Prod. xvi. 2, 374). Great lumps of white resin
stood out on the straight white trunks.
The trees were all too big to climb, so it was only possible to collect the
old scales underneath, still in sound condition and in some cases attached
to the axes of the cones. Many young trees, + 20 in. high, showed the
fastigiate youth form characteristic of the genus (16, t. ix. f. 1). The leaves
in the above collection were from a young plant 2 m. high, with one whorl
of single branches + 2'75 in. long. As the leaves vary in size and texture
with the age of the plant, it is impossible to base a species on vegetative
characters alone, and A. Labillardieri Warb. is no doubt synonymous with
the above, as already suggested by Lauterbach, I. c.
ARAUCAKIA BECCARII Warb. Monsunia, i. 187. A. Cunningliamii Becc. in
Malesia, i. 180 (non Ait.). A. Cunning kamii Ait., var. papuana Laut.
in Engl. Bot. Jahrb. 1. (1918) 51.
Arfak Mts., gregarious in parts in forest by ? lake. Seedlings. Dec.
5934. Young plant (1 m.). 5748. Old foliage and cones (pro-embryo
stage). 5749.
Distrib. New Guinea (D.N.W., Arfak Mts., Hatam, Beccari).
These handsome trees were quite abundant on the eastern bank of the
lake. They were about 25 in. in height and mostly in fruit. The large
erect cones are borne on the horizontal uppermost branches of the trees.
Many of the mature cones seen on the largest trees must be double the size
given by Beccari, but the indurated bases of old leaves densely investing the
trunks rendered climbing out of the question, and, having no axes with us,
they could not be felled. A smaller tree, felled by " parang, v yielded two
? cones, one of which approximated to Beccari's measurements, while the
other was smaller. Both the cones were in pro-embryo stage, with the seed-
coat already quite indurated. The ovuliferous scale, not shown in A. Cun-
ninghamii, is very noticeable in this species, as Beccari has described, and in
this character it approximates to J. JIunsteinii K. Schum. (Fl. Kais.
Wilhelmsland, 12 (1889)). The cone-scales a re more elongate than in A. Cun-
nuiijkamii, with narrower base, more swollen apophysis, and a more pungent
apex. The leaves of the fertile branch are more spreading, 1 cm. long and
84
3 mm. broad at the dilated base, with pungent apex more or less reflexed.
In habit these trees differ from A. Cunningliamii, for, though fastigiate-
pyramidal in growth, the branching is not so defined or symmetrical, showing
little trace of the candelabra-like habit so familiar in the latter. There is
also a difference in the size and shape of the leaves, in their stomatal
markings, and in the much larger size of the ? cones. I must thank
Dr. Beccari for material of his species, which he most kindly sent me for
purposes of comparison ; also Dr. Stapf for carefully considering the points
of difference between the two plants. (Pis. 1, 3, figs. 1, 5.)
In both the available cones an apparent orifice (ori., JB) is visible on the
swollen pulvinus of most of the bracts, behind the apex of the ovuliferous
scale, possibly due to rapture of tissue.
FIG. 5.
Araucaria Beccarii Warb. A. Sporophyll; o.s., ovuliferous scale, w., wing.
B. Lateral view ; ori., apparent orifice. C. Dorsal view.
LlBOCEDRUS ARFAKENSIS Gibbs, Sp. nov.
Arbor alta, in diversis ramis monoica ; rami teretes, cortice fusco obducti, ramuli
oppositi, distichi. Folia decussatim opposita, in statu juvenili omnia conformia,
linearia, in statu adulta adpresse quadrifariatim imbricata, diffonnia, marginalia
navicularia, subacuta, 'coriacea, maxima ex parte adnata, complicata, apice solum
libera, facialia plana, squamiformia, triangularia, carinata, acuminata. Strobili
masculi in ramulis lateralibus solitarii, terminales, cylindracei. Antheroe GO -seriatse,
spiraliter dispositse, stipite breve, connectivi appendicula squamiformi, late ovata,
leviter peltata, chartacea, loculi 3-6, globosi, deorsum 2-valves. Strobili feminei in
ramulis brevibus erecti ; bractese 4, elongato-ovatae, demum lignosse, appendicem
magnum late ovatum obtusum antice proferentes. Nucula elliptica, subacuta, alata,
altera subobsoleta, altera elongato-ovata.
Hob. Arfak Mts., on ridges and in the forest by ? lake, 7000-8000'.
85
Seedling, ? lake, 7000'. Dec. 555(1 Youth form, S.W. ridge, 8000'.
5500. Koebre ridge, 8000'.
formibus, subteretibus ; spicis parvis densifloris.
Scandent, and of moderate size. Sheathed stem apparently 15-20 mm.
in diam. Leaf-sheaths (in the small portion seen by me) closely armed with
short small prickles. Ocrea extraordinarily large and long, as much as
50 cm. in length, very gradually long-acuminate, slightly inflated, enfolding
the younger part of the stem, covered with thin fuscous-furfuraceous scurf,
chartaceous, exsuccous, rigid, later splitting longitudinally on the outer side
and not dissolving into fibres, fugaciously furfuraceous outside and orna-
mented all over, and rather closely, with oblique, interrupted, slightly raised,
spinulit'erous lines or ridges. Leaf-slieatli flagella slender and very long,
armed, but not very regularly, with ternate or quinate claws, intermingled
with smaller scattered prickles. Leaves non-cirriferous ; in one specimen
about 1 m. long on the whole ; the petiole alone 35 cm. long, almost
biconvex, slightly prickly, and with obtuse margins ; rhachis smooth,
bifaced, with an acute salient angle above, and armed beneath, at first, with
quinate and higher up with ternate claws. Leaflets very conspicuously
approximate into very few distant groups (3 in the specimen at hand)
with long vacant spaces of rhachis interposed ; in one specimen the leaflets
are 7 on each side of the rhachis, of which 8 (4 on each side) form a
basal group, and 7 (3 on each side with a deeply bilobed apical one) form
the terminal group ; between these two main groups is another intermediate
12
94
formed of 2 opposite leaflets onlv ; the leaflets are lanceolato-ensiform, or
oblanceolate, and taper almost equally to both ends, have the base rather
acute, and are shortly and suddenly acuminate in a slightly bristly-spinulous
tip ; are papyraceous, rather firm, glossy on both surfaces, but especially
above, very slightly paler beneath, with an acute mid-costa and a few slender
side-nerves, all quite naked on both surfaces, though at times the mid-costa
is slightly spinulous above near the apex ; margins remotely and minutely
ciliate-spinulous, more closely near the apex ; transverse veinlets very
crowded, fine and sharp ; intermediate leaflets 40-50 cm. long, 3'5-4 cm.
broad, the lowest smallest, those of the terminal group shorter, but not
narrower. Male spadix somewhat shorter than the leaves (70 cm. long in
one specimen), erect, strict, having an elongate pedicellar part bearing only
one dense and narrow cupressiform panicle, about 20 cm. long (perhaps at
times the spadix is longer, and with more than one panicle) ; the primary
spathe is very elongate, and at first enfolds the spadix ; it is tubular,
flattened with acute edges, closely sheathed in its lower part, and is produced
above into an open, thinly membranous, lacerate, lanceolate-acuminate limb,
it is sprinkled all over outside with minute tuberculiform prickles ; the
panicle is composed of several gradually diminishing, very approximate, very
densely flowered, short spikelet-bearing branchlets, inserted at an angle of
45, subtended by secondary thinly membranous, dry, lanceolate-acuminate,
more or less lacerated, secondary spathes, only a little shorter than their
respective branchlets ; the branchlets have the appearance of being small,
simple, cylindrical spikes, 5-6 cm. long in the basal part of the panicle,
gradually a little shorter above ; in fact, however, they are composed of
several, very short, very closely drawn together, alternate, distichous
spikelets, the lowest of which are 8-10 mm. long, with 7-8 very approximate
flowers on each side ; the upper spikelets gradually diminish in length and
number of flowers ; spathels bracteiform, membranous, concave with a
triangular acute apex, surpassing the involucre; this is deeply cupular,
or subcampanulate, truncate, not distinctly two-keeled, and bidentate on the
side next to the axis. Male flowers very closely packed together, oblong,
6 mm. long; the calyx deeply 3-lobed; the corolla twice as long as the
calyx, narrowing a little above to a bluntish apex. Female spadix and fruit
unknown.
Hab. Humboldt Bay, ridge behind the village, 700', scrambling in high
forest. (?. Jan. 6267.
A very distinct species, but with very marked affinities with C. macro-
chamis Becc. It belongs to Group V. of my monograph, characterized
mainly by non-cirriferous leaves, and by the leaf-sheaths furnished with
a long clawed flagellum ; in that group it falls into the division which
contains C.macrocldamys and other Papuan species, all having very large and
elongate chartaceous ocrese. Even among these species C. humboldtiantti
95
is easily distinguishable by its relatively short leaves, having few, lanceolate,
grouped leaflets, glossy and smooth on both surfaces ; by the sheaths
prolonged into an excessively long, slightly inflate ocrea, which is orna-
mented with oblique spinuliferous slightly raised ridges ; by the strict
long-pedicelled spadix with a very densely flowered panicle ; and by the
primary spathe having an elongate, thinly-membranous, lacerate, externally
prickly limb.
CALAMUS ARFAKIANUS Becc., sp nov.
Gracilis, scandens, caudice 5-10 mm. diarn., vaginis flagelliferis, spinis gracilibus,
interdum brevissimis, parce armatis, in ore et in ocrea brevi barbatis ; foliis non
cirriferis, brevibus, petiolo brevissimo, segmentis paucis per greges paucos inter se
remotos approximatis, lineari-lanceolatis, subulato-acuminatis, tenuiter papyraceis,
unicostulatis, nervis secundariis utrinque 1-2 tenuibus, costa media et nervis omnibus
levibus; segmentis intermediis circiter 20 cm. longis, 15-20 mm. latis, duobus
terminalibus basi breviter connatis vel fere liberis; spadicibus valde elongatis et
flagelliformibus ; spathis elongatis arete vaginantibus ; spadicis 3 ramis prinuiriis
paucis, remotis, spicas numerosas detiexas f erentibus ; spicis majoribus 15-18 mm.
longis, floribus utrinque 13-15 fere contiguis ; spatbellis concavis, bracteiformis ;
spadicis $ ramis primariis paucis, remotis ; spicis majoribus 4-5 cm. longis, floribus
utrinque 8-10 ; fructibus late ovato-ellipticis, abrupte mucrouatis, 15 mm. diam. ;
scininLs albumin e homogeneo.
IScandent and very slender. Sheathed stem 5-6 or at most 8-10 mm. in
diam. Leaf-sheaths flagelliferous, not or only slightly gibbous above, at
times almost, smooth, and only slightly (and very minutely) tubercled-
spinulous in their upper part; but in some specimens rather densely covered
with scattered or slightly confluent bristle-like spines, which become closer,
considerably longer, and more hair-like, near and on the margins of the
mouths of the same leaf-sheaths, and of the ocrea, which because of them
looks bearded. Leaf-sheath flagella very slender and long, and armed
irregularly with very minute scattered claws. Leaves non-cirriferous, short,
40-60 cm. long; petiole very short, 2-3 cm. long at most; rhachis fuga-
ciously rusty-furfuraceous like the petiole, armed beneath with unequal,
rather long-tipped, scattered or ternate claws. Leaflets few, 15-19 in all,
very irregularly set, usually approximate in three groups separated by long
vacant spaces of rhachis; they are narrowly lanceolate or linear-lanceolate,
gradually subulately acuminate, thinly papyraceous, almost shining and about
the same colour on both surfaces ; have the mid-costa very sharp above, and
one, or at times two slender secondary nerves on each side of it ; mid-costa
and nerves quite naked and smooth on both surfaces ; transverse veinlets
rather sharp and not very crowded ; the margins have only a few distant,
almost inconspicuous, appressed spinules, closer and more distinct in the
apical part ; the leaflets of the interme iiute and lower groups are about
96
20 cm. long and 15-20 mm. broad ; those of the terminal group somewhat
smaller ; the two terminal are the smallest, free, or more or less connate,
at the base. Male and female spadices similar, flagelliform, very long and
slender, with a very few distant partial inflorescences ; primary spathes very
narrowly tubular and elongate, very closely sheathing, more or less armed
with scattered small claws ; the lowest slightly compressed, the others
cylindraceous. Male partial inflorescences (primary branches) zigzag
sinuous, spreading, inserted outside their respective spathes with a distinct
axillary callus ; the lowest 18-20 cm. long, with 7-8 rather distant spikelets
on each side; the upper ones gradually shorter, and with fewer spikelets;
secondary spathes 8-10 mm. long, smooth or at times very slightly spinulous,
tubular -infundibuliform, or with a narrow flattened base, and slightly
enlarged above, very closely sheathing, obliquely truncate and glabrous at
the mouth, where acute at one side ; spikelets flattened, 6 mm. broad, comb-
like with very approximate or contiguous bifarious flowers ; spikelets
strongly deflexed, inserted just at the mouths of their respective spathes ;
the lower spikelets 15-18 mm. long, bearing 13-15 flowers on each side; the
others gradually shorter; spathels bracteiform, broad, concave, acute, strongly
striately-veined; involucre cupular, obsoletely two-keeled and two-toothed
on the side next to the axis. Male /lowers very close together, ovate, sub-
acute, 2 - 5 mm. long; the calyx deeply 3-lobed, strongly striately-veined ;
the corolla twice as long as the calyx. Female spadix has shorter partial
inflorescences and fewer but larger spikelets than the male spadix ; the
largest (lowest) spikelets are 4-5 cm. long, and carry 8-10 flowers on each
side; spathels broadly infundibuliform, truncate; involucrophorum obliquely
campanulate ; involucre deeply cupular, truncate, as long as the involucro-
phorum ; areola of the neuter flower rather conspicuous, subcupular. Female
flowers ovate, inserted at an angle of about 45. Fruiting perianth almost
explanate, not pedicelliform. Fruit broadly ovoid-elliptical, 17-18 mm. long
(not taking into account the beak), 15 mm. broad, equally rounded at both
ends, suddenly topped by a beak 2'5 mm. long ; scales in 21 longitudinal
series, rather deeply grooved along the centre, straw-yellowish with a very
narrow dark brown marginal line; tip bluntish. Seed irregularly globose,
with homogeneous albumen.
Hob. Arfak Mts., Angi lakes, common all through the forest round ? lake
at 7000-8000'. "A small rotang, excellent for tying." 5977 (? type-
specimens) with the sheathed stem 8-10 mm. in diam. and rather densely
prickly leaf-sheaths. The male plant (6144, Dec., Mt. Arfak, 5000' on
the S.W. ridge, scrambling in high forest) : very slender; sheathed stem
8-10 mm. in diam. ; leaf-sheaths nearly smooth.
It is evidently related to the Australian C. Muelleri, from which it mainly
differs in the leaflets not being distinctly 3-costulate, and in the female spadix
97
having the flowers inserted at an angle of about 45 (not horizontal), and in
its larger fruit.
C. arfakianus is characterized by its slender stem, by the cirriferous leaf-
sheaths, and short non-cirriferous leaves, with a few narrowly lanceolate
leaflets approximated into a few (three) distant groups; the two terminal
leaflets free, or but slightly united at their bases ; by the very slender and
long filiform spadices, not very dissimilar in the two sexes, and by the ovoid-
elliptical, suddenly-beaked fruit, having the seed with equable albumen.
CALAMUS ARFAKIANUS var. IMBERBIS Becc.
Vaginis inermibus, in ore ininiine barbatis ; ocrea brevi truncata ; segmentis
interdum obsolete 3-costulatis.
Differs from the type in having the leaf-sheaths unarmed and strongly
striate longitudinally, and not bearded at their mouths ; in the ocrea being
short, truncate, and marcescent, and also without bristles ; the leaflets in
this variety have the side nerves more distinct, and at times appear almost
3-costulate.
Hob. Arfak Mts., S.W. ridge, leading to Angi lakes ; scrambling in
forest at 7000'. ? . Dec. 6143.
CALAMUS PRATTIANUS Becc., sp. nov.
Gracilis, scandens, caudice circ. 1 cm. diam. ; vaginis flagelliferis, spinis graci-
libus subsetiformibus densiuscule obsitis, in ore dense longiuscule hispido-barbatis ;
foliis non cirriferis, brevibus ; petiolo brevissimo ; segmentis paucis, inaequidistantibus,
saepe utrinque geminatis (non per greges remotos approxiinatis), anguste lanceolatis,
subulato-acuminatis, chartaceis, fimiis, unicostatis, utrinque nudis vel superne in
costa media et in nervis secundariis duobus inconspicue remoteque spinulosis ;
segmentis intermediis 15-18 cm. longis, 20-25 mm. latis, superioribus sensim
minoribus, duobus terminalibus basi liberis vel breviter unitis ; spadice c? gracillimo,
elongato ; inflorescentiis partialibus paucis, brevibus, remotis, spicas paucas erecto-
patulas ferentibus ; spicis majoribus 3 cm. longis, flores 13-15 utrinque ferentibus,
remotiusculis, spathellis breviter lateque infundibularibus, truncatis. Caetera desunt.
Scandent and very slender. Sheathed stem about 1 cm. in diam. Leaf-
sheaths flagelliferous, not or but slightly gibbous above, rather densely
covered with scattered or slightly confluent bristle-like spines, becoming
closer and considerably longer and more hair-like near the mouths of the
same leaf-shaths and their ocrese, which latter are very short and on account
of these bristles look densely bearded. Leaf-sheath flagella very slender,
long, and armed irregularly with very minute scattered claws. Leaves non-
cirriferous, short (the few seen by me are 35 cm. long); petiole very short,
or almost obsolete ; rhachis fugaciously rusty-t'urfuraceous, armed with a
line of solitary, rather long-tipped claws. Leaflets few, about 16 in all ;
very irregularly set, usually approximate in pairs on each side of the rhachis,
with irregular vacant spaces of rhachis interposed ; they are narrowly
98
lanceolate, gradually tubulately acuminate, papyraceous, rather rigid, almost
shiny and of about the same colour on both surfaces, have the mid-costa very
sharp above, and one or two slender secondary nerves on each side of it ;
underneath they are quite smooth, but above have occasionally a few very
minute spinules on the mid-costa, and on a secondary nerve on each side
of it ; transverse veinlets much interrupted and not very sharp ; the margins
almost smooth, or very inconspicuously spinulous, more distinctly so at the
apex; the intermediate leaflets are 15-18 cm. long and 20-25 mm. wide,
the lowermost and the upper ones gradually smaller, the two terminal being
the smallest, free, or more or less connate at the base. Male spadix
flagelliform, very long and slender, with very few and distant partial
inflorescences ; primary spathes very narrowly tubular and elongate, very
closely sheathing, more or less armed with scattered small claws, the lowest
compressed with rather acute margins ; the others cylindraceous, somewhat
produced and lacerated at apex; partial inflorescences ascendent, rather
short, the lowest and largest about 10 cm. long with 5-6 spikelets on each
side ; 'secondary spathes very narrowly tubular-infundibuliform, or with a
flattened base, and slightly enlarged above, obliquely truncate at their
mouths, and there slightly produced at one side into a triangular point,
which embraces the bases of their respective spikelets ; spikelets spreading,
the largest (lowest) about 3 cm. long, with 13-15 flowers on each side; the
upper are shorter, and have fewer flowers ; spathels shallowly and broadly
infundibuliform, truncate, strongly striately- veined ; involucre cupular,
obsoletely two-keeled and slightly two-toothed on the side next to the axis.
Male flowers ovoid, inserted at an angle of about 45, not in contact with
each other, but separated by the blades of their respective spathels. Female
spadix and fruit unknown.
Hob. Arfak Mts., near the Monswoon Snoon (cT lake) at about 8000'.
Collected by Mr. A. E. Pratt.
It is closely related to C. arfikianus, from which it differs in the leaves
having unequidistant, but not distinctly grouped leaflets, on that account
resembling C. Muelleri more than C. arfakianus. From C. Muelleri it differs
also in having larger spikelets, and having ascendant non-contiguous flowers.
C. Prattianus is characterized by its slender stem with cirriferous leaf-
sheaths and short non-cirriferous leaves ; by the ocreai and the mouths of
the leaf-sheaths being furnished with long bristles ; by the lanceolate
unequidistant (but not distinctly grouped) and almost smooth leaflets ; by
the very slender and long-flagelliform male spadix having only a few partial
inflorescences furnished with but few spreading spikelets ; these have briefly
infundibular spathels and bear the flowers inserted at an angle of about 45,
a little apart one from the other.
99
CENTKOLEPIDACEiE.
CENTROLEPIS NOVO-GUINEENSIS Gibbs, sp. nov.
Planta perennis, densissiine csespitosa, multifoliata. Folia basi vaginata, vagina
hyalina, margine longe pilosa, lamina lineare, canaliculata, leviterscabriuscula, obtusa.
Pedunculus gracilis, scabriusculus ; gliunis 2, inaBqualibus. calyptratis, margine
hyalinis, unitioris. Flares bractea unica hyalina stipati. Stamen 1. Ovarium
2-loculare, loculis 2, superpositis. Styli 2.
Hah. Arfak Mts., Angi lakes. Abundant in marsh by ? lake where
open on sand, 7000'. FJ., Fr. Dec. 5566. Koebre ridge, abundant on
open burnt summit, where damp, 9000'. Fl., Fr. 5646.
Plants 2'4 cm. high, with branched stems, all densely matted with white
hairs at the base. Leaves 8 mm. long, with vagina 3 mm. long and 1 mm.
broad, the lamina 4 mm. by 5 mm. Peduncle 1*5-1'8 cm. in length, much
longer exported in 5646, each bearing one spikelet, 4 mm. long. Glumes 2,
the largest 3'5 mm. long, and the smaller 3 mm. Hyaline bract 4 mm. long,
o>ate acute. Stamen with versatile anther 1 mm. in length; filament
3'5 mm. long. Ovary 1*5 mm. long. Styles 3 mm.
Very near C. pliilippinensis Merr., but is a smaller plant with shorter,
radially arranged, spreading leaves, hairy at the base, with, as far as seen,
one flower in each glume.
This interesting record affords further striking proof of the wider
distribution of a so-called Australian genus. One species alone was previously
known from Asia, viz. C. cambodiana Hance, till Merrill in 1906 found
C. pliilippinensis on Mt. Halcon at 2400 in. In 1910 1 C. kinabaluensis was
found by me in N. Borneo, on Mt. Kinabalu at 13,000', which now, wich
the above, gives 4 well-defined Asiatic species, indeed 5, if an undescribcd
plant in the Kew Herbarium should belong to this genus ; and no doubt
further exploration of the magnificent mountain ranges of New Guinea with
their unlimited possibilities will yield many more.
1 CKNTBOLEPIS KINABALUENSIS Gibbs, = C". philijipinemis W. B. Turrell, ex Gilbs
in Journ. Linn. Soc., Bot. xlii. (1914) 172 (non Merr.).
Planta perennis, 2-3 cm. alta, caespitosa. Folia disticha, imbricata, + 1'2 cm. longa, vagina
6 mm. longa, 2 mm. lata, hyalina, glabra, lamina 8 mm. longa, 5 mm. lata, obtusa, sctosa.
Pedunculiis 1'2 cm. longus; glumis 2, inaequalibus, inferne latis concavisque, superne angustatis,
apice obtusis, dorso carinatis, 4*5 mm. long. Flares 6-8, in quaque gloma 34, flos qnisque
bractea hyalina, 3'5 mm. longa. Filamentum 3 mm. longum, anthera 1'2 mm. longa. Ovarium
1'5 mm. longum, stylis 2 mm.
Hab. Kinabalu, granite core, summit, cracks in granite, forming mats. Fl.,Fr. Feb. 4207.
A note of Turrell'* on the herbarium sheet at Kew states, " The number of flowers in a
ppikelet varies in this plant. I have found 4, 6 or 8 flowers in one spikelet. The Philippine
type is described as having 4 flowers and all the flowers of Merrill 6160 that I have
dissected have this number in each spikelet." The plant also differs in the glabrous
distichous leaves and the shape of the glumes, which have bluut apices.
100
X YllID A C E M.
*XYRIS PAUCIFLORA Wildm. Phytogr. i. 2, t. i. f. 1.
Arfak Mts., Angi lakes, common in open marsh by ? lake. FL, Fr.
Dec. 5928.
Distrib. India (" Foot-hills of the Himalayas, in marshes, from Nepal
eastwards, N. Bengal to Burma," Hook. f. Fl. Br. Ind. vi. 365), Ceylon,
Malay Peninsula, N. Borneo, Celebes, Philippines, and China. N.E. Australia.
ERIOCAULACE^E. (A. B. RENDLE.)
ERIOCAULON LEUCOGENES 1 Ridl. in Trans. Linn. Soc. ser. 2, Bot. ix. (1916)
240.
Arfak Mts., Angi lakes, common on marsh by $ lake, where open and
sandy, 7000'. Fl. Dec. 5567.
Distrib. New Guinea (D.S.W., Mt. Carstensz, Kloss).
" Flower heads mauve." The plants show a great range in size from
6 cm. to 20 cm., the heads varying in diameter from '5 to 1 cm.
JUNCACE^E. (A. B. RENDLE.)
JUNO us LAMPOCARPUS Ebrh. Calam. n. 126 ; Ridl. in Trans. Linn. Soc. ser. 2,
Bot. ix. (1916) 231.
Arfak Mts., Angi lakes, open marsh by ? lake, 7000'. Fl. Dec. 5927.
Distrib. Temperate Europe and Asia, N. and E. Africa, Eastern N.
America, S.E. Australia and New Zealand.
LILIACE^E.
DIANELLA CGERULEA Sims, Bot. Mag. (1801) t. 505 ; Schum. & Laut. 219 ;
Nova Guinea, viii. (1914) 996 ; Ridl. in Trans. Linn. iSoc. ser. 2, Bot.
ix. (1916) 230.
Arfak Mts., in high forest below, and in open spaces on the S.W. ridg",
running up to Angi lakes, also in open marsh by ? lake, 7000-8000'. Fl.
Dec. 5519.
Distrib. New Guinea (D.N.W., Arfak Mts., Gjellerup; D.S.W., G.-lieb,
Brandenliorst; Mt. Carstensz, Kloss; N.E.). Philippines, New Caledonia,
and Fiji ; N.E. Australia to Tasmania.
Acaulescent, and with light blue flowers.
LUZURIAGA ASPERICAULIS Hall. f. in Nova Guinea, viii. (1914) 991,
t. clxxxi.
Arfak Mts., twining in mossy forest on S.W. ridge running up to Angi
1 A species of Eriocanlon was also collected by Miss Gibbs on Mt. Kinabalu at 12,000'
(no. 4209) by Kadamaiau torrent, on the granite core near the summit of the mountain ;
it was mixed with Centrolepis kinabaluensis Gibbs (no. 4209) (see p. 99). .It is a csespitose
plant, forming small cushions 2 - 5 cm. high, with glabrous leaves 2-2'5 cm. long, jhl mm.
wide in the middle, linear-tapering from a broad membranous base. The specimens are all
Bterile. No Eriocaulon has hitherto been recorded from the grauitd core of the mountain.
101
lakes, 8000-0000'. Fl. Dec. 5536. Common in forest slopes by ? lake,
7000'. Fl., Fr. 5744.
Distrib. New Guinea (D.N.W., Arfak Mts., Gjellerup).
A common and very pretty twiner with white flowers. The fruit, which
was not available in Dr. Gjellerup's specimens, is a roundish berry, 8 mm. loi'g
by 6 mm. broad, 1-2-seeded, with persistent style and remains of perianth-
tube at the base.
IRIDACE.E.
PATERSONIA NOVO-GUINEENSIS Gibbs, sp. nov.
Planta perennis ; caulis brevissimus. Folia circiter 5-8, scapum sequantia vel
superantia, disticha, linearia, rigida, acutissima, in marginibus dense rufo-pilosa,
tenuiter inultistriata. Spathce subsequales, lineari-oblongae, acutissirnse, carinatie
(carina ad f rufo-pilosa vel glabra) striatae, scarioso-marginatse. Bracteoe spathis
similes nisi magis membranacese necnon angustiores brevioresque, ad apicem piloso-
marginatae vel glabrse. Flores in spatha 2-3, sessiles; tubus gracillimus, vix exsertus;
lobi exteriores obovati, patentes, in carina dorsali ad apicem dense barbati, interiores
nulli. Filamenta in tubum integrum connata. Stylus superne, leviter incrassatus ;
lobi stigmatici lineari-oblongi, papillosi.
Hob. Arfak Mts., Koebre ridge, plentiful on open burnt summit, 9000'.
Fl., Fr. Dec. 5600.
A small rigid plant, with gleaming white or pale mauve flowers, + 1-3-5 dm.
in height. Leaves from 1-2-3-5 dm. by 5-6 mm., characterized by a line of
matted branched hairs, containing brown colouring-matter, up the dorsal
keel, and from the vagina to the apex on the adaxial margin of the leaf.
Sheathing scales at the base of the leaves from 3'5 cm. long, lanceolate acute,
brown, the old bases persisting round the stem. Scape 7'3 cm. long, more
or less exserted from the sheathing leaves, which it exceeds or equals, or it
may be shorter. Spathes 3'8 cm. by 5 mm. Perigonium tube 2 cm. long,
lobes 8 mm. long, including apical tuft of hairs 1 mm. long. Anthers 3 mm.
Stigma-lobes 2 mm.
This species is near P. Lowii Stapf. It differs in the somewhat broader
leaves with marginal line of brown hairs and the generally white corollas
with an apical tuft of hairs to each lobe. The transverse section of the leaf
shows many more fibro-vascular strands and is more attenuated at the margin
than in the above species. There is also no trace of an inner perianth, as
Stapf also found for the Kinabalu plant. The marginal line of matted
branched hairs is common to some Australian species, viz. P. sericea R. Br.
and P. pygmcca Lindl., but in every case apparently the hairs are hyaline
and much finer in texture.
This genus, for many years supposed to be endemic in Australia, now
includes three Malayan mountain types, viz. P. borneensis Stapf and P. Lowii
from Mt. Kinabalu in N. Borneo, the latter also common to Mt, Halcon in
the Philippines, and the above.
102
ZlNGIBEEACE^. (TH. VALETON.)
ALPINIA ARFAKENSIS K. Sch. in Engl. Bot. Jahrb. xxvii. (1899) 296.
Var. SUBSESSJLIS Val., var. nov.
Folia subsessilia minora, glaberrima, siccando valde convoluta coriacea. Ligula
20 mm. longa vel longior, valde macerata. Anther cm. longum,
112
clinandrio parvo, concavo cum costula longitudinal! in rostellum decurrente, auriculis
obtusis. Anfhera cucullata, longe ultra thecas producta, ambitu subquadrangula
vel melius 7-angula, apice abrupte subulato-acuminata, basi truncata, c. 0'14 cm.
longa, connective in costulam longitudinalem incrassato, thecis suborbicularibus.
Pollinia 4, geminata, lateraliter compressa, triangula, extus convexa. Ovarium
pedicellatum 6-costatum, c. O9 cm. longum. Gapsula oblongo-obovoidea, basi
acuta, c. 1-1 cm. longa, pedicello c. I'l cm. longo.
Hob. Arfak Mts., Angi lakes, forest patch by $ lake, edge of forest in
humus, terrestrial, 7000'. Fl., Fr. Dec. 5G89.
The first species of the section Rhachidilulbon recorded from New Guinea.
It much resembles L. brgvistylis J. J. S. (L. montana Lindl. var., brevistylis
J. J. S.).
Flowers green with a brown labellum.
LIPARIS ( PLATYCHILUS ?) GIBBSIJR J. J. S., sp. nov.
Rliizoma repens, radicans, teres, vaginis magnis ad c. 3'25 cm. longis tectuin.
Pseudobulbi c. 2'5 cm. distantes, cum rhizomate angulum anguste acutum facientes,
sicco tenues, teretes, c. 2'5 cm. longi, 1-folii, basi nonnullis vaginis magnis acute
acuminatis ad c. 4 cm. longis cincti. Folium inarticulatum, erectum, lanceolatum,
acuminatum. acutum, basi sensini in petiolum angustatum, nervis 5 majoribus subtus
prominentibus, sicco membranaceum, c. 9'75-10 - 5 cm. longum, 2'l-2'25 cm. latum ;
petiolus canaliculatus, nervosus, c. 2'5-4 cm. longus. Inflorescentia in pseudobulbo
mature terminalis, folio brevior, satis multiflora, pedunculo c. 4'5 cm. longo, vaginula
angusta acuta c. 2'75 cm. longa ad basin, rhachide fractiflexa, c. 6 cm. longa. Bractcce
adpressse, lanceolato-trian guise, acutissimae, concavse, 1-nervise, ad c. 0'55 cm. longse.
Flores c. 16, quaquaversi, patentes, sepalis patentissimis, convexis. Sepalum dorsale
oblongum, superne angustatum, obtusum, 3-nervium, costa media dorso apicem versus
prominente, c. 0*65 cm. longum, 0'25 cm. latum. Sepala lateralia divergentia,
oblique oblonga, f'alcatula, obtusa, vix apiculata, 3-nervia, costa media dorso pro-
minente, c. - 67 cm. longa, 0'3 cm. lata. Petala reflexa, linearia, obtusa, basi
leviter oblique dilatata, leviter convexa, 1-nervia, c. 0'57 cm. longa, medio O'OG cm.
lata. Labellum porrectum, cum gynostemio angulum acutum faciens, basi gyno-
stemio adnatum, in utraque parte canaliculse latse longitudinalis convexum, subtus
convexum, e basi contracta quadrangula sensim dilatatum, 2-lobum, obcordatum,
lobis remotis obtusis grosse crenatis cum lobulo brevi obtuso in sinu, basi contracta
excepta ciliolatum, 3-nervium, nervis exterioribus ramosis, callo majusculo antice
dentes 2 erectos parallelos falcato-triangulos acutiusculos utrosque basi antice in
costulam brevem exeuntes gerente in basi, explanatum c. 0'67 cm. longum, antice
0'45 cm. latum, parte contracta c. 0'2 cm. longa, 0'275 cm. lata. Qynostemium
superne valde hamato-incurvum, dorso convexum, basi valde dilatatum, medio labello
adnatum, ovario multo latius, c. O32 cm. longum, basi 0'17 cm. latum, clinandrio
concavo, intus ad basin rostelli dente acuto instructo, auriculis subquadrangulis
Anthera cucullata, transverse quadrangulo-ovalis, apice truncata, basi emarginat?
bidentata. Stigma, obverse rotundato-triangulum, margine elevatum. Ovariuii
6-sulcatum, cum pedicello c. 0'4 cm. longum, curvatum.
Hob. Arfak Mts., Angi lakes, forest patch by ? lake, epiphyte in forest
7000'. Dec. 5Gi*trib. New Guinea (D.N.W., Arfak Mts., Gjellervp).
116
CERATOSTYLIS ( EUCERATOSTYLIS) ANGTENSIS J. J. S., sp. nov.
Caules approximati, teretes, c. 12'5-15 cm. longi, macerati 0'17 cm. diam., basi
vaginis tubulosis superne accrescentibus sese amplectentibus sicco membranaceis laxe
reticulato-venosis brunneis ad c. 2*25 cm. longis tecti. Folium erectum, subulato-
teres, acutiusculum, antice leviter longitudinaliter costatum cum sulco tenuissimo,
c. 2'7 cm. longum, maceratum c. 0*125 cm. diam. ; vagina tubulosa, antice rumpens,
c. 0'3 cm. longa. Inflorescentia fascicularis, pluriflora, squamata, pedunculis
partialibus tenuibus, 1-floris, c. 0'3 cm. longis, basi vaginulam latam obtusam
membranaceam c. 0'26 cm. longam gerentibus. Bractea ample cucullata, obtusa,
ovario multo brevior, membranacea, c. 0'14 cm. longa. Flores parvi, bene 0'4 cm.
longi, sepalis dorso parce pubescentibus. Sepalum dorsale oblongum, obtusum,
3-nervium, c. 0'25 cm. longum, bene O'l cm. latum. Sepala lateralia lacinia
oblique oblonga concava pedem gynostemii bene superante decurrentia, mentum
reversum ovario parallelum oblongum apice subinflatum obtusum bene O'l cm.
longum formantia, oblique oblonga, apice angustata, obtusa, 3-nervia, c. 0'24 cm.
tota fere 0'4 cm. longa, 0'125 cm. lata. Petala oblique oblonga, acutiuscula,
falcatula, 1-nervia, c. 0'2 cm. longa, fere 0'07 cm. lata. Lalellum cum pedi
gynostemii angulum fere rectum faciens, gynostemio parallelum, ima basi ultra
apicem pedis gynostemii obsolete productum, supra basin leviter recurvum, con-
cavum, gynostemium bene superans, basi unguiculato-angustatum, -| parte superiore
contractum carnoso-incrassatum et utrinque convexum, obtusum, parte mediana
marginibus incurvum et ciliolatum, 3-nervium, lineis 2 elevatis puberulis parallelis
intus usque ad medium, explanation c. 0'35 cm. longum, medio 0'14 cm. latum, apice
incrassato bene O'l cm. longo, fere O'l cm. lato. Gynostemium ultra medium
bifidum, c. 0'14 cm. longum, brachiis parallelis, oblongis, obtusis, extus convexis,
intus concavis. AntTiera cucullata, transverse suboblongo-elliptica, apice truncata,
c. 0'0(5 cm. lata. Pes gynostemii reversus, cum ovario angulum acutum faciens,
apice a sepalis lateralibus libero obtusangule incurvus, c. O'l cm. longus. Ovarium
pubescens, c. 0'375 cm. longum.
Hob. Arfak Mts., Augi lakes, isolated forest patch by ? lake, epiphytic,
7000'. Dec. 5718.
A very inconspicuous species.
The colour of the flowers is white.
DENDKOBIUM ( CADETIA) SUBRADIATUM J. J. S. in Fedde Rep. xii. (1913)
27 ; in Nova Guinea, xii. (1915) 273, t. c. 171.
Arfak Mts., Angi lakes, terrestrial in forest by ? lake, 7000'. Dec.
5542 and 5908.
Distrib. New Guinea (D.N.W., Arfak Mts., Gjellerup).
DENDROBIUM ( LATOURIA) RHOMBOGLOSSUM J. J. S. in Bull. Jard. Boh Buit.
2 e ser. n. ii. (1911) 9 j in Nova Guinea, xii. (1913) 44, t. xiii. 34.
Arfak Mts., Angi lakes, marsh by ? lake, terrestial, abundant, 7000'.
Dec. 5511. Koebre ridge, between c? an d ? lake, burnt open summit
plateau, 9000'. Dec. 5009.
Distrib. New Guinea (D.N.W., Goliath Mts., De Kock).
117
" Plant up to 1 ra., perianth segments magenta outside and white inside ;
abundant."
DENDROBIUM ( LATOURIA) CURVIMENTUM J. J. S., sp. nov.
Caules (1 adest) tenuiter clavati, c. 6-nodes (absque internodiis summis foliiferis
brevissimis), c. 19'5 cm. longi, apice 3-folii, internodio maximo incrassato infra folia
c. 6-3 cm. longo. Folia erecto-patentia, ovato-elliptico-lanceolata, apicem versus
angustata, acuta, .basi breviter contracta, sicco rigide coriacea, c. 5-67 cm. longa,
1-2-1'G cm. lata. Infloresceiitice e nodis summis inter folia, folia superantes,
c. lo-florae, pedunculo tenui, c. 5'2 cm. longo, vaginulis c. 4 tubulosis acutis donato,
rhachide sicco angulata, c. 4 cm. longa. Bractece late triangulae, acute, concava?,
c. 0'3 cm. longae. Flores parvi, incurvi, carnosuli, glabri. Sepalum dorsale cuin
ovario anguluin obtusum fornians, oblongo-triangulurn, dimidio superiore angustutnin,
obtusum, concavum, 5-nervium, c. 0'54 cm. longurn, bene 0'3 cm. latum. Sepala
latcralia ad pedem gynostemii decurrentia, mentum ovario parallelum valde incurvum
breve crassum obtusum basi subconstrictum bene 0'3 cm. longum formantia, late oblique
triangula, subobtusa, basi antice in laciniarn brevem triangulam dilatata, margine
superiore valde rotundata fere quadrangula, concava, 5-nervia, fere 0'5 cm. longa,
basi 0'5 cm. lata. Petala oblique anguste oblonga, falcatula, obtusa, basi leviter
contracta, apice erosula, concava, 1-nervia, c. 0'46 cm. longa, 0'14 cm. lata.
Lalelluiii pedi gynostemii insertum, unguiculatum, 3-lobum, concavum, carnosum,
subexplanatum c. 0'4o cm. longum, 0'6 cm., bene explanatum 0'65 cm. latum, ungue
pedi gynostemii parallelo valde recurvo, canaliculato, fascia longitudinal! bicostata
ad basin laminae dilatata et in dentes 3 conicos exeunte instructo, dentibus lateralibus
membra na angusta labello adnatis, callo rotundato verruculoso ante dentem inter-
medium, carinula transversa utrinque ad basin lobi intennedii, lamina transversa,
c. 0'3 cm. longa ; lobi laterales inexplanati porrecti, paralleli, gynostemium et lobum
intermedium superantes, explanati lobum intermedium aequar.tes vel vix superantes,
late oblique ovati, rotundati, concavi ; lobus intermedius transversus, bipartitus,
concavus, subtus convexus et crasse carinatus, carina infra sinum in apiculum
crassum conicum exeunte, lobulis oblique triangulis, margine antico rotundatis,
obtusis, explanatis divergentibus, explanatus c. 0'14 cm. longus, 0'325 cm. latus.
Gynostemium a dorso compressum, in utraque stigmatis parte obtusangule dilatatum,
absque anthera c. 0-175 cm. longum, clinandrio concavo, margine denticulate, fila-
mento incurvulo, oblongo, rotundato, auriculis paulo brevioribus, obtusis, leviter
denticulatis. Anthera cucullata, transverse ovali-reniformis, basi rotunda to-biloba,
apice breviter rotundato-producta, c. 0-16 cm. lata. Pollinia 4, in corpuscula
2 oblique obovata supra convexa subtus plana unita, lateraliter compressa, interioni
qua m exteriora angustiora, c. 0'07 cm. longa. Stigma obverse triangulum, basi
rotundatum, apice truncatum. Pes gynostemii cum ovario angulum acutum faciens,
reversus, inedio fere rectangule incurvus, oblongus, obtusus, valde excavato-concavus,
iiifxplunatus c. 0'3o cm. longus. Orariutn valde incurvum, 6-sulcatum, c. 0'33 cm.
longum ; pedii-i'llus I'l cm. longus.
Hub. Arfak Mts., Koebre ridge, forest, 8000-9000'. Fl. (green). Dec.
5647.
118
This Dendrolium belongs to the small-flowered species of the section
Latouria. The shape of the labellum calls to mind that of I), dendrocolloides
J. J. S. and D. latifrons J. J. S., but the habit is very different.
DENDROBIUM ( TRACHYRHIZUM) LATIFRONS J. J. S., sp. nov.
Rhizoma breve, vaginis brevibus tubulosis sese amplectentibus omnino obtectum,
radicibus dense verrucosis. Caules simplices, elongati, supenie flexuosi, c. 47 cm.
longi, internodiis c. 3'7-l'3 cm. longis. Folia patentia, oblonga, oblique bidentata,
dentibus acutiusculis, costa media supra sulcata subtus prominente, marginibus in
sicco recurvis, sicco tenuiter coriacea, ad c. 5'5 cm. longa, 1*8 cm. lata, superiora
et inferiora decrescentia ; vaginae tubulosse, apice truncate, internodiis breviores.
InJlorescenticB in caulibus foliatis axillares, vaginam dorso ad basin perforantes,
arcuatae, laxe 3-5-florse, pedunculo cauli adpresso, c. 3-4 cm. longo, nonnullis
vaginulis tubulosis ad basin, rhachide c. l'5-2 cm. longa. JBractets adpressse,
triangulse, concavse, c. 0%3 cm. longae. Flores quaquaversi. Sepalum dorsale late
triangulum, marginibus curvatum et base utrinque angulatum, obtusum, concavum,
7-nervium, c. O8 cm. longum et latum. Sepala lateralia cum pede gynostemii
inentum obtusum formantia, antice omnino libera, late oblique triangula, obtusa,
breviter crasse obtusa apiculata, margine postico basi angulata, 7-nervia, costa
media dorso obtuse prominente, c. O95 cm. longa, basi 1'15 cm. lata. Petala
oblique subovalia, basi lata, breviter obtusiuscule triangulo-acuminata, concava,
minutissime erosulo-ciliolata, basi 3-nervia, c. 0'77 cm. longa, O53 cm. lata.
Labellum ungue brevi lato 5-nervio pedi gynostemii insertum, 3-lobum, curvatum,
subtus supra basin concavum cum nervis 5 obtuse prominentibus, intus fascia
latissima longitudinali carnoso-incrassata subquinquecostata cum costis 3 mediants
alte elevatis basi in appendicem reversam rotundato-trapeziformem obsolete retusam
concavam carnosam unguem sequantem, c. 0'2 cm. longam, O35 cm. latam, producta
in basi lobi intermedii in lamellam transversam porrectam excavationem anticam
obtegentem verrucosam antice dentes c. 5 porrectis iiTegulariter conicis gerentem
utrinque nonnullas costulas transversas emittentem terminante, intus papillosum,
explariatum c. 1'13 cm., usque ad apicem loborum lateralium O93 cm. longum, ad
lobos laterales c. 1'43 cm. latum, ungue c. O2 cm. longo ; lobi laterales erecti,
patentes, oblique ovati, rotundati, basi lati, concavi, margine postico manifesto
incurvi ; lobus intermedius latissimus, concavus, 2-lobus, medio plica supra concava
subtus convexa et in sinu dente brevi triangulo donatus, lobis transverse oblique
rotundato-quadrangulis, apice truncatis, irregulariter crenatis, angulum exteriorem
versus repandulis, expknatus c. 0'43 cm. longus, 1'3 cm. latus. Gynostemium
(foecundatum) bene 0'3 cm. longum, apice porrecto utrinque dente donato, auriculis
triangulis. Pes gynostemii cum ovario angulum obtusum fere rectum formans,
late oblongus, intus pubescens, basi dorso convexus, apice altius excavatus et dorso
roturidato-gibbosus, c. 0'7 cm. longus. Ovarium (foecundatum) obconicum, 6-
sulcatum, c. 0'4 cm. longum ; pedicellus c. 1*3 cm- longus.
Hob. Arfak Mts., Angi lakes, terrestrial in marsh by ? lake, 7000'.
Dec. 5558.
This species is nearly allied to D. appendiculoides J. J. S., from which it
differs, so far as the single specimens allow comparison, by its more robust
119
stems, less flowered racemes with the peduncle pressed against the stem,
broader petals, the lip with different-shaped keels, shorter, blunter, not
falcate lateral lobes, a shorter and much broader mid-lobe and a broader basal
appendix, whereas the column-foot is much less excavated. Moreover, the
plant was collected in a much higher altitude above the sea.
The species recalls I), prosteriglossum Schltr. var. oltusilolum Schltr. to
mind ; it is possible that the two plants will prove to be identical.
The flowers are described as yellow.
DENDROBIUM ( OXYGLOSSUM) TRIFOLIUM J. J. S., sp. nov.
Caules approximati, c. 4-5 cm. longi, inferne vaginis mox fatlscentibus, apice
c. 3-foliatL Folia erecto-patentia, lanceolata, oblique obtusa, breviter apiculata,
omnino sed praesertim apice minutissime denticulata, costa media subtus prominente,
sicco rigide coriacea, c. 2'6-4 cm. longa, 0'67-0'83 cm. lata ; vaginae inferne tubu-
losse, superne canaliculato-conduplicatae. Injlorescentia pseudoterminalis, brevissima,
c. 3-flora. Uractete rachidem amplectentes, late triangula3, acute acuminatae, ad
c. 07 cm. longse. Flores erecti, majusculi, c. 3'1 cm. longi, sepalis sicco dorso parco
nigro-punctatis. Sepalum dorsale ellipticum, obtusum, apice minutissime denticu-
latum, 5-nervium, c. 1'25 cm. longum, 0'65 cm. latum. Sepala lateralia kcink
elongata partim anguste lineari ad pedem gynostemii decurrentia, inentum rectum
ovario adpressum c. 1'7 cm. longum, superne marginibus auticis longitudine c. 0'6 cm.
connatis anguste calcariforme apice retusum formantia, parte anteriore oblique
trkngulum, marginibus curvatis, subacuta, acute apiculata, apice minutissime
denticulate, 7-nervia, costa media dorso prominente, c. 1-2 cm. longa, basi 2 cm.
margin e antico 2*9 cm. metientia. Petala spathulato-rhombea, vix acuminata,
acuta, superne papilloso-ciliolate, 3-nervia, c. 1'23 cm. longa, 0'57 cm. late.
Labellum pedi gynostemii et gynostemio pamllelum et adpressum, pedi gynostemii
longitudine c. 0'95 cm. adnatum, gynostemium superans, anguste lineare, apicem
versus cliktetum, apice 3-lobuin, concavum, infra apicem subventricosum, ecallosum,
inferne 3-, superne 7-nervium, expknatum c. 2'6 cm. longum, ad lobos kterales
fere 0'4 cm., basi partis liberse 0'175 cm. ktum; lobi kterales breves, rotundati,
papillosi ; lobus intermedius revolutus, triangulus, acutus, papilloso-cilioktus,
c. 0-25 cm. longus, basi O16 cm. ktus. Gynostemium a dorso compressum, in
utraque stigmatis parte obtusangulum, absque anthera c. 0'2 cm. longum, clinanclrio
in utraque fikmenti parte kcinukto, filamento subukto, incurvo. Anthera cucullata,
antice visa quinquangukris, apicem versus angustete, truncate et parte superiore
adpresse puberuk, basi 2-loba, c. 0'2 cm. late. Pollinia 4, in corpuscuk 2 oblique
obovate supra convexa subtus concava unite, oblique oblonga, 2 interiora quam
cxtci'iora paulo angustiora. Stigma suborbicukre. Pes gynostemii lineans, canalicu-
ktus c. 1'7 cm. longus. Ovarium pedicelktum ckvatum, suj^erne 3-alatum et
3-costatum, ak superiore quam cetera? longiore et in dentcm libenHD trkngulum
producte, sparsa nigro-punctetum, c. 3 cm. longum.
llab. Arfak Mts., Angi lakes, on bank by ? lake. 7000', terrestrial.
Dec. 5907.
This species, the flowers of which agree in colour with I), pe
120
Svhltr., differs from the latter in the larger flowers, a blunt dorsal sepal,
differently shaped petals and lip, lacinulate clinandrium, not 5-winged ovary.
The flowers are described as dull yellow, with a red tip to the labellum.
DENDROBIUM AGATHOD^MONIS J. J. S. in Bull. Dep. Agr. Ind. Neerl. n. xxxix.
(1910) 7 ; etc.
Arfak Mts., ridge running up to Angi lakes, terrestrial in open gravel
spaces, 8000'. Dec. 5530 and 5596.
Disifib. New Guinea (D.N.W., Cyclops Mts., Gjdlerup ; D.S.W., Aga-
thodiimonsberg, von Roemer, Hell wig Mts., van Noukuys).
Perhaps it would be better to keep the typical I), agatliodcemonis apart
as a species and not to unite with it the different forms I successively added.
However, these forms are, in the material at hand, almost all represented by
single specimens, and therefore it is very difficult to make a decision.
The flowers of the Arfak specimens are described as magenta.
DENDROBIUM ( CALYPTROCHILUS) PAPUANUM J. J. S., sp. nov.
Caules approximati, elongati, superne flexuosi, sicco inferne teretes superne alto
sulcati, c. 46 cm. longi, internodiis ad c. 2'6 cm. longis. Folia basi semitbrta, ovato-
lanceolata, longe acuminata, acutissima, longe mucronata, acumine denticulata, costa
media subtus prominente, sicco rigidiuscula, c. 6-7'5 cm. longa, 1-2-1-9 cm. lata ;
vaginae tubulosse, internodia paulum superantes, minute verrucosse, novelise sicco
fusee punctatse. Inflorescentiae ad nodos caulium defoliatorum, abbreviate,
c. 4-flora3, pedunculo nonnullis vaginulis brevibus tubulosis tecto. Bractece valde
approximatse, patentes, ovato-trian guise, acute acuminataa, concava^, sicco c. 0'85 cm.
longae. Flores parvi, macerati c. 1*4 cm. longi. Sepalum dorsale subovale, apice
brevissime productum, obtusum, valde concavum, 3-nervium, c. 0'65 cm. longum,
0'4 cm. latum. Sepala lateralia lacinia oblique triangula ad pedem gynostemii
decurrentia, marginibus anticis longitudine c. O65 cm. connata, men turn magnum
conicum ovario parallelum suberectum anguste obtusum c. 0'9 cm. longum formantia,
parte libera oblique ovato-triangula, pra^sertim margine postico valde curvata, obtusa,
apiculata, concava, c. 4-nervia, c. 0'625 cm. longa, margine antico c. 1'35 cm., basi
1 cm. metientia. Petala oblique subelliptico-oblonga, subulato-apiculata, concava,
superne erosula, 3-nervia, c. 0'65 cm. longa, 0'27 cm. lata. Labellum gynostemium
sequans, basi gynostemii longitudine c. 0'4 cm. adnatum, concavum, apice cucullatum,
ecallosum, c. 5-nervium, nervis exterioribus ramosis, cucullo plicato margine apicali
truncate dense et breviter subulato-lacinulato, explanatum cuneatum, late obverse
rhombeum, fere I'l cm. longum, 0'84 cm. latum, ab apice cuculli usque ad marginem
apicalem c. 0'37 cm. metiens. Gynostemium recurvuluin, a dorso compressum,
apicem versvis dilatatura, in utraque stigmatis parte obtusangule dilatatum,
c. 0'2G cm. longum, filarnento curvulo, lineari, obtuso, dorso convexo, auriculis
magnis, quadrangulis, obtusissimis, dimidio superiore apice leviter productis et
denticulatis, dimidio inferiore rotundatis. Antliera cucullata, arnbitu transverse
subquadi-angula, leviter 6-angulata, basi biloba, apice producta, truncata et papillosa,
c. 0'22-j cm. lata. Pollinia 4, oblonga, in corpuscula 2 oblique subovalia supra
convexa subtus concava conglutinata. PCS gynostemii linearis, canaliculatus,
121
c. 0'9 cm. longus. Ovarium pedicellatum clavatum, minutissime furfuraceo-
puncticulatum, c. l - 25 cm. longum.
Hal. Arfak Mts., Angi lakes, epiphytic in forest patch by ? lake, 7000'.
Dec. 5712.
Amongst the small-flowered purple or pink coloured species of the section
Cali/ptrochilus this one differs from D. roseum Schltr. in its lonf acuminate
leaves, smaller flowers, and much broader lip without callus.
The flowers are pink.
DKNDROBIUM ( CALYPTROCHILUS) INFRACTUM J. J. S. in Fedde Rep. xii.
(1913) 118 ; in Nova Guinea, xii. 340, t. cxxiii. 225.
Arfak Mts., Koebre' ridge between <$ and ? lake, terrestrial on burnt
open summit plateau, 9000'. Fl. Dec. 5655.
Distrib. New Guinea (D.N.W., Arfak Mts., Gjellerup).
DENDROBIUM ( CALYPTROCHILUS) RIPARIUM J. J. S. in Fedde Rep. xii.
(1913) 117 ; in Nova Guinea, xii. 343, t. cxxiv. 227.
Arfak Mts., Angi lakes, forest patch by ? lake, epiphytic in forest, 7000'.
Fl. Dec. 5896.
Distrib. New Guinea (D.N.W., Arfak Mts., Gjellerup).
DENDROBIUM ( CALYPTROCHILUS) FRUTICICOLA J. J. S. in Fedde Rep. xii.
(1913) 116 ; in Nova Guinea, xii. t. cxxv. 229.
Arfak Mts., Angi lakes, terrestrial in open marsh by ? lake, growing ia
patches, abundant, 7000'. Fl. Dec. 5510. Forest patch by ? lake, 7000'.
Fl. (orange). Dec. 5553. Koebre ridge between and ? lake, terres-
trial on open burnt summit plateau, 9000'. Dec. 5608.
Distrib. New Guinea (D.N.W., Arfak Mts., Gjellerup).
The flowers show the same slight differences as in the original specimens.
DENDROBIUM ( CALYPTROCHILUS) PARVIFOLIUM J. J. S., sp. nov.
Planta parva (?). Caulis ramosus, valde flexuosus, sicco longitudinaliter alte
suleatus, pars adest 13'5 cm. longus, internodiis c. O'G-O'S cm. longis. Folia parva,
patentissiina, oblongo-ovata, apicem versus angustata, breviter acuta, apiculata,
prsesertim superne erosula, costa media sicco supra sulcata subtus cum nervis sequen-
tibus prominente, rigida, sicco ad c. 1*1 cm. longa, fere 0.4 cm. lata; vagina?
tubulosse, verruculosae, internodia aequantes. Ltflorescentice (1 adest) in flexubus
caulium foliatorum, brevissimse, 1-florae, pedunculo abbreviate, nonnnllis vaginulis
tubulosis verruculosis obtecto. Bractea cucullata, apiculata, c. 0-85 cm. longa.
Flos c. 1-8 cm. longus. Sepalum dorsale ovatum. apiceru versus angustatum,
anguste obtusum, breviter obtuse apiculatum, basi latum, concavum, 5-nervium,
c. U'7o cm. longum, 0'4 cm. latum. Sepala lateralia lacinia falcatulo-trian^nla a(tlum dorsale horizontal, oblongum, obtusum, breviter obtuse conico-apiculatum,
concavum, superne vix erosulum, 3-nervium, c. 0'37 cm. longum, 0'15 cm. latum.
Sepala lateralia porrecta, cum pede gynostemii mentum subrectanguluin cum ovario
angulum obtusum faciens obtusum postice c. 0'13 cm. longum fonnantia, margiue
proximo ultra medium conglutinata, oblique triangula, falcata, margine extoriorc
rotundata, obtusa, breviter obtuse conico-apiculata, concava, 3-nn-via, tola c. ();}.") cm.
longa, 0'17 cm. lata. Pffnln gyoostemio :i(l])rcssa, parallela, oblique oblonga, suj>erne
denticuluta, longe suhulalo-aruininata, l-ncrvia, c. 0'3 cm. longa, O'l cm. latu,
L
124
acumine O'l cm. longo. Labellum membrana tenui 3-nervia pecli gynostemii
insertum, mobile, erectum, sigmoideum, e basi contracta dentibus brevibus reversis
falcato-triangulis et intus callo erecto quadrate ornata dilatatum, fascia mediana
longittidinali convexo-incrassatum et subtus convexum, lateribus supra convexum
subtus concavum, 3-nervium, \ parte superiore recurva et utrinque plica parva sed
distincta supra convexa a parte inferiore separata, itaque subtrilobum, medio utrinque
minute ciliolatum, inexplanatum c. 0'15 cm. longum, explanatum spathulato-
obovatum, c. 0'17 cm. longum, O075 cm. latum, lobo intermedio rotundato, semi-
orbiculari, convexo, subtus concavo, carnosulo, eciliato, c. O05 cm. longo, 0'075 cm.
lato. Gynostemium cum ovario angulum obtusum faciens, curvatum, totum
c. 0-12 cm. longum, clinandrio concavo, filamento elongate, subulate, auriculis
filamentum bene superantibus, subulatis, falcato-incurvis, margine inferiore lobulo
subobsoleto obtuso munitis. Anthera cucullata, semiglobosa, apice producta incurva
truncata, connectivo gibboso-incrassato papilloso, c. 0"05 cm. alta. Pollinia in cor-
pusculum semiglobosum unita. Rostellum triangulum, obtusum. Stigma magnum,
fere tetam faciem inferiore.m gynostemii occupans, profunde excavatum, obovato-
triangulum. Pes gynostemii cum ovario angulum obtusum faciens, subrectus, basi
dorso convexus, apice vix recurvulus, oblongus, obtusus, canaliculatus, crassiusculus.
Ovarium 6-sulcatum, c. 0'075 cm. longum.
Arfak Mts., Angi lakes, forest patch by ? lake, epiphyte in forest, 7000'.
Fl. Dec. 5662.
Differs from B. myrtillus Schltr., probably its nearest ally, by the
mentum, the broader slightly three-lobed lip biauricled at the base and
only ciliolate in the middle beneath the semiorbicular mid-lobe, and the
papillate connective.
The flowers are described as yellow ; in the dried plant the petals are
distinctly red.
BULBOPHYLLUM ( MONOSEPALUM) MURICATUM J. J. S. in Bull. Dep. Agr.
Ind. Need. n. xlv. (1911) 9; in Nova Guinea, viii. (1911) t. C, B ;
xii. 374. Monosepalum muricatum Schltr. Orch. D. Neu-Guinea,
(1912) 682.
Arfak Mts., ridge running up to Angi lakes, terrestrial in mossy forest,
8000'. Fl. Dec. 5997.
Distrib. N.E. New Guinea.
Flowers yellow with red spots.
BULBOPHYLLUM ( NEMATORHIZIS) OVALITEPALUM J. J. S., sp. nov.
Eliizoma elongatum, repens, filiforme, longinode, ad nodos plerumque 1-subpseudo-
bulbis pauces radices emittens, pars adest 25 cm. longa, internodiis c. O5-1 cm. longis,
vaginis tubulosis, quarn internodia multo brevioribus. Pseudotulbi c. 1 '5-3-2 cm.
distantes, erecti, oblique ovoidei, sicco valde rugosi, c. 0'5-0'65 cm. longi, 1-folii.
Folium erectum, breviter petiolatum, oblongum ad ligulatum, obtusum, apice leviter
insequaliter vel subsequaliter breviter obtuse bilobulatum, basi acutum, costa media
subtus prominente, c. rS-3'1 cm. longa, sicco 0-45-O55 cm. lata ; petiolus cana-
liculatus, c. G'l-0-2 cm. longus. Inftorescentice ad basin pseudobulborum et ad
125
nodos rhizomatis solitariae, pedunculo 1-floro filiformi, c. 1'3 cm. longo, mferne
c. 2-vaginulis tubulosis superne leviter arnpliatis obtusis c. 0'125 cm. longis donato.
Bractea brevis, cupuliformis, obtusa, c. 0'08 cm. longa. Flos parvus. Sepalum
dorsale ovatum, obtusum, concavum, 3-nervium, c. 0'34 cm. longum, 0*225 cm.
latum. Sepala kteralia oblique ovalia, obtusissima, breviter obtuse apiculata,
3-nervia, c. 0'35 cm. longa, 0'22 cm. lata. Petala vix obovato-ovalia, obtusissima,
3-nervia. Labellum mobile, curvum, carnosum, ambitu subquadrangulo-ovale, basi
truncatum, apice rotunda turn, dimidio inferiore excavato-concavum cum marginibus
erectis, antice convexum, glabrum, c. 0'2 cm. longum, 0'12 cm. latum. Gynostemium
breve, apice obtuso, auriculis brevibus. Anthera cucullata, explanata reniformis,
obtusa, c. 0-075 cm. lata. Pollinia 4, lateraliter compressa, a latere visa obovata,
exteriora extus convexa, interiora manifeste tenuiora. Stigma majusculum, trans-
versuna. Pes gynostemii cum ovario angulum obtusum faciens, quadrangulus, apicem
versus paulo angustatus, truncatus, c. 0'075 cm. longus et fere sequilatus. Ovarium
G-sulcatum, parce punctatum, c. 0'25 cm. longum ; pedicellus tenuis, supra basin
articulatus, c. 0'45 cm. longus.
Hab. Arfak Mts., Angi lakes, forest patch by ? lake, epiphytic in forest,
7000'. Dec. 5683.
This seems to be the first known species of the section NematorJuzis from
the Arfak Mts.
The column was not in a very good state, hence the description of it
wants completion.
The flowers are green.
BlILBOPHYLLUM ( PELTOPUS) OCTARRHENIPETALUM J. J. S. in Fedde Rep.
xii. (1913) 400 ; in Nova Guinea, xii. 400, t. cxlriii. 277.
Angi lakes, forest patch by ? lake, epiphytic, 7000'. Fl. (white).
Dec. 5504.
Distrib. New Guinea (D.N.W., Arfak Mts., Gjellerup).
BULBOPHYLLUM ( DiALEiPANTHE) FRiSTis J. J. S. in Fedde Rep. xii. (1913)
399 ; in Nova Guinea, xii. 419, t. clviii. 296.
Arfak Mts., Angi lakes, slopes by " campong," ? lake, terrestrial in
forest, 7500'. Dec. 5638.
Distrib. New Guinea (D.N.W., Arfak Mts., Gjellerup).
Only an inflorescence.
BULBOPHYLLUM ( HYALOSEMA) TRICANALIFERDM J. J. S. in Fetfde Rep. xii.
(1913) 398 ; in Nova Guinea, xii. 426, t. clx. 302.
Arfak Mts., Angi lakes, epiphytic in forest patch by ? lake, 7000'.
Dec. 5548.
hi ft nb. New Guinea (D.N.W., Arfak Mts., Angi lakes, Gjellerup).
Flowers yellow with brown spots.
L2
126
PHREATIA ( BULBOPHREATTA) SPATHULATA J. J. S., sp. nov.
Pusilla. RMzoma elongatum, repens, ramosum, radicans, vaginis tubulosis
tectum. Pseuclobulbi parvi, remoti, omnino vaginis obtecti, 2-folii. Folia erecta,
insequimagna, lanceolato-spathulata, obtusa, retusa, apiculata, sicco ad c. 2'3 cm.
longa, 0'35 cm. lata. Inflorescentia folia bene superans, laxe c. 13-flora, pedunculo
tenui, c. 4'5 cm. longo, vaginulis c. 6-tubulosis apiculatis inferne magis approximatis
ad c. 0-35 cm. longis donate, rhachide tenui, c. 3 cm. longa. Bractece ovato-triangula?,
acuminatse, .acutissima?, ad c. 0'225 cm. longse. Flores parvi. Sepalum dorsale
ovatum, obtusum, concavum, 1-nervium, c. 0'2 cm. longum, O14 cm. latum. Sepala
lateralia divergentia, late oblique ovata, acuta, oblique concava, 1-nervia, c. 0-2 cm.
tota 0'25 cm. longa, 0'2 cm. lata. Petala oblique ovata, subacuta, concava, 1-nervia,
c. 0-16 cm. longa, Ol cm. kta. Labellum leviter sigmoideum, unguiculatum, con-
cavum, explanaturn c. 0'14 cm. longum, ungue quadrangulo, basi utrinque in lobulum
parvum rotundatum dilatato, probabiliter 2-glanduloso, c. 0'07 cm. longo, basi
O06 cm. lato, abrupte in laniinam dilatato, lamina semilunato-reniformi, apice
latissime rotundata, medio leviter retusa cum lobulo distincto obtuso in sinu, lobis
lateralibus obtusa, incrassationibus 2 convexis intus ad basin, intus papillosa,
c. 0-06 cm. longa, O175 cm. lata. Gynostemium breve, cum ovario angulum
obtusum faciens, dorso convexum, papillosum, absque anthera c. O075 cm. longum,
clinandrio alte excavato. Bostellum latum. Stigma transversum, margine elevatum.
Pes gynostemii reversus, c. Ol cm. longus, basi ovario adpressus et intus convexus,
parte superiore incurva, quam partem inferiorem angustiore, truncata. Ovaritim cum
pedicello trigono torto clavatum, c. 0'35 cm. longum.
Hob. Arfak Mts., ridge running up to Angi lakes, epiphytic in forest,
8000'. Dec. 6004.
A small plant remarkable for the two convex thickenings at the base of
the blade of the lip.
Flowers white.
PHREATIA ( RHIZOPHYLLUM) DENSISSIMA J. J. S. in Fedde Rep. xii. (1913)
26 ; in Nova Guinea, xii. 438, t. clviii. 314.
Arfak Mts., Angi lakes, epiphytic in forest patch by ? lake, 7000'.
Fl. (green). Dec. 5550.
Distrib. New Guinea (D.N.W., Arfak Mts., Gjellerup).
OCTARRHENA CYLINDRICA J. J. S., Sp. DOV.
Caules approximati, compressi, simplices, basi valde radicantes, c. 5-5 cm. longi,
cum vaginis c. 0'375-0'4 cm. lati, internodiis c. O5-O7 cm. longis. Folia equitantia,
erecto-patentia, articulata, lateraliter compressa, linearia, interdum falcatula, acuta,
basi valde obliqua, dorso c. 2-3'5 cm. longa, medio 0'275-0'35 cm. lata; vaginse sese
amplectentes, tubulosse, lateraliter cornpressse, antice alte fissa3, apice valde obliqua?,
internodia superantes. Inflorescentia axillares, erectae, dense multiflora?, cylindricse,
pedunculo tenui, c. 1-75-2 cm. longo, inferne nonnullis vaginulis tubulosis longe
acuminates, superne numerosis vaginulis bracteiformibus donate, rachide angukta,
c. 2'4-2'S cm. longa. Bractece e basi lata longe lineari-acuminatse, concave, iiTe-
gulariter marginatae, c. 0'14 cm. longae. Flores vagi, minimi c. 0'17 cm. lono-i.
127
Sepalum dorsale erectum, triangulum, obtusiusculura, convexum, c. 0'06 cm. longura
et latuin. Sepala lateralia oblique ovato-triangula, apice interdum plus ininusve
contracta, anguste obtusa, c. 0-075 cm. longa, 0'07-0'075 cm. lata. Petala diver-
gentia, anguste oblique triangula, subfalcatula, acuta, convexa, 1-nervia, c. 0'07 cm.
longa. Labellum simplex, decurvuin cum ovario angulum obtusum faciens, medio
fere obtusangule incurvurn, subsigmoideum, manifeste concavum, 1-nervium, basi
medio tantum affixum, callis 2 longitudinalibus parallelis approximatis oblongis in f
partibus inferioribus labelli, explanatum ovato-oblongum, anguste obtusum, c. Ol cm.
longum, fere 0'05 cm. latum. Gynostemium cum anthera 0. O'OG cm. longum,
clinandrio cum dorso gynostemii angulum rectum faciente, concavo. Anthera
cucullata, ovato-triangula, basi leviter emarginata, apice anguste truncata, c. 0'04 cm.
longa. Pollinia 8, clavato-pyriformia. Bostellum productum. Stigma trans-
versum. Ovarium obconicum, cum pedicello angulum obtusum faciens, c. O'OG cm.
longum ; pedicellus crassus, tortus, c. 0'06 cm. longus.
Hob. Arfak Mts., ridge running up to Angi lakes, epiphytic in forest,
8000'. Dec. 5993.
From the other species this one is readily distinguished by its moderately
dense and broad leaves and dense spikes.
The flowers are said to be yellow.
Var. MAJOR J. J. S., var. nov.
Gaules elongati, inferne defoliati cum radicibus adpressis, ad c. 21 cm. longi.
Folia 2-4-3 cm. longa, 0-37-0-475 cm. lata. Pedunculus c. 2*25-25 cm. longus ;
rhachis 4-5 cm. longa. Flores majores. Sepalum dorsale c. O'l cm. longum,
0-075 cm. latum, lateralia 0'12 cm. longa, 0'8 cm. lata.
Hob. Arfak Mts., Angi lakes, mossy slopes by ? lake, 7000-8000'.
Fl. Dec. 5536.
This differs from the type-specimen only by its longer stems and larger,
differently coloured flowers.
The flowers are white.
In the dried specimens the flowers of the type as well as those of the
variety are greenish with a large white anther.
DICOTYLEDONE.E.
P I P B R A C E M. (0. DB CANDOLLE.)
PIPER ARFAKIANUM C. DC., sp. nov.
Kamulis glabris, junioribus in nodis hirtellis; foliis modice petiolatis glabris,
lirnbo ovato-acmninato basi ima sequilatera acuto apice longe et acute acuminate,
5-nervio, petiolo basi ima vaginante ; pedunculo glabro quam petiolus breviore,
spica quam limbus pluries breviore cylindrica et apice obtusa, rhachi pilosa,
bractese glabrae, pelta transverse elliptica centre late et brevissime pedicelkta,
ovario inferne rhachi at baud profunde iinmerso superue libero ovato et glabro,
stigmatibus rotundato-ovatis.
128
Hob. Arfak Mts., S.W. ridge running up to Angi lakes, in forest, 8000'.
Fl. Dec. 5525.
Dioicum, epiphytum. Ramuli spiciferi 1 mm. crassi, collenchyma libriforme in
fascicules discretes dispositum, fasciculi intramedullare, 1-seriati, canalis lysigenus
unicus centralis. Limbi in sicco firmi fuscescentes et pellucido-punctulati, superi
usque ad 5 cm. longi et 2'5 cm. lati. Petioli usque ad 5 mm., pedunculi usque
ad 3 mm. longi. Spica matura 7 mm. longa, 4 mm. crassa, in sicco nigra.
PlPEK PILOSULINODUM C. DC., sp. nOV.
Ramulis tantum in nodis pilosulis, primum Isevibus postea lineatim lenticellatis ;
foliis parvis modice petiolatis, limbo ovato-acuminato basi obtuso apice obtusiuscule
et sat longe acuminate supra glabro subtus baud dense piloso, 5-nervio, petiolo
piloso basi ima vaginante ; pedunculo glabro quam petiolus breviore, spica florente
quam limbus paullo breviore, rhachi hirsuta, bractese glabrse, pelta rotunda,
staminibus 2, antheris tetragonis 4-valvatis.
Hab. Arfak Mts.. slopes of Koebre ridge between Angi lakes, 8000'. FL
Dec. 5624.
Dioicum, epiphytum. Ramuli spiciferi 1 mm. crassi, collenchyma libriforme
in fascicules discretes dispositum, fasciculi intramedullares 1-seriati, canalis lysi-
genus unicus centralis, in ramulis 2 mm. crassis lineatum lenticellatis cellulae
sclerosse circum collenchyma creberrimse. Limbi in sicco subcoriacei fuscescentes
et creberrime pellucido-punctulati usque ad 4'2 cm. longi et 2 cm. lati. Petioli
circiter 5 mm., pedunculi 3 mm. longi. Spicse florentes 3 cm. longse, 1 mm.
crassse, bracteae pelta 0'75 mm. diam.
F AGACE^I.
QUERCUS LAUTERBACHII Seemen in Engl. Bot. Jabrb. xxiii. Beitr. Ivii. 54 ;
Si:hum. & Laut. 264, t. iv. figs. A-E.
Arfak Mts., S.W. ridge, 6000'. Fr. Dec. 6124.
Distrib. New Guinea (D.S.W., Arfak Mts., 6000', A. E. Pratt ; (Herb.
Brit.)N.E.).
A tall tree with slender stem. Pratt's collection consists of a single
acorn, very striking in size, 6'5 cm. long, with cupule 6 cm. by 5 cm. and
nut 4'5 cm. long and about the same in breadth, enclosed in the cupule for
I'D cm. from the base. I came across the same group of trees on the exact
spot kindly described to me by Mr. Pratt ; the ground was strewn with
acorns of all sizes, of which the one originally collected would represent the
largest. The leaves are larger than in Seemen's measurements, with the
veins pilose on the under surface, but they were collected from young
plants.
URTICACEJE.
PIPTURUS PAPUANUS Gibbs, sp. nov.
Arbuscula vel frutex : ramulis internodiis brevibus, cortice rugoso-lenticelloso ;
innovationes tomentosae. Folia parva, petiolata, lanceolata, sensim angustata,
acuminata, basi obtusa, integerrima, leviter revoluta, rigide membranacea, supra
129
hirsuta, pilis albidis adpressis dense obtecta, demum asperrima, subtus subtiliter
brunneo-velutina, trinervia, costa media prominente nervis 2 lateralibus, arcuato-
anastomosantibus, reticulo inter venas conspicuo. Glomeruli feminei axillares,
sessiles, pisifonnes, densiflori. Perigonium tomentosum, fructiferurn albo-carnosum.
Hab. Arfak Mts., Angi lakes, edge of forest by ? lake, 7000'. Fl. ? ,
Fr. Dec. 5955.
Leaves +6 cm. by 1*5 cm., green, but drying a dusty brown, densely
covered on the upper surface with punctiform cystoliths. Petioles 1*3 cm.,
slender, densely puberulous. Stipules 4 mm., lanceolate, acute, tomentose
on the outside. Glomerules 4 mm. across, in fruit, forming a white swollen
receptacle. Flower ? 3 mm. long ; style filiform, hirsute, 2'5 mm. Achene
+ 1 mm. long, covered with short stiff white hairs.
This plant is distinct in the small narrow leaves with scabrid upper
surface and the very well-marked network of minor veins, forming minute
interstices, covered with dense brown tomentum on the under surface. In
the shape of the leaves it is nearest to P.mindartensis Elm., but differs in the
small size, more rigid texture and entire margins, the prominent venation
with fewer lateral veins, and the brown coloration when dry.
GIBBSIA Rendle, gen. nov. (A. B. RENDLE.)
Flores inonoici, unisexuales : HCasculi alabastro depresso-globosi, mucronati ;
perianthio fere ad basin 5-partito, segmentis valvatis, ovatis. Stamina 5, antberis
subrotundis, dorsifixis, in alabastro inflexis. Ovarii rudimentum lanuginosum.
Feminei perianthio brevi, late cupuliforme, persistente et basin fructus cupula
carnosula adnata tegente. Ovarium ovoideum valde obliquum ; stigma apicale,
sessile, discoideum, papillosum, margine setulis fimbriato; ovulum a basi erectum,
micropyle elongata, superne ampliata et fimbriata. Fructus subdrupaceus, parvus,
valde obliquus, exocarpium tenue, carnosulum, et super cupulam demum separabile,
endocarpium crustaceuin. Semen conforme, testa membranacea ; albumen carnosum ;
cotyledones parvae, ellipticae ; radicula superior.
Frutices, foliis alternis, petiolatis, crenato-serratis, 3-nerviis, subtus canescentibus.
Stipiilae membranaceae, in unam intrapetiolarem alte bifidam connatae, deciduie.
Inflorescentice axillares, solitariae vel binae, foliis multo breviores, dichotomse.
Flores in cymulis parvulis androgynis apice ramulorum ultiinorum breviter pedi-
cellati. Bracteee minutae, ovatse, scariosae.
Recalls the genus Delregeasia in its leaf-characters, but in characters of
flower and fruit is nearest the Malayan and Pacific Island genus Leucosyke,
which, however, has a penicillate stigma and the fruits crowded on a fleshy
receptacle ; there is no trace of a fleshy receptacle in Gibbsia. The fruit is
very characteristic ; the upper portion of the thinly succulent exocarp
separates like a cap, and the endocarp enclosing the seed is then readily
separable from the cup formed by the union of the somewhat fleshy perianth
with the lower part of the exocarp.
130
GIBBSIA INSIGHTS Rendle, sp. nov.
Fruteoc ramulis foliiferis colore cinerascente et pilis brevibus appressis hispidis.
Folia breviter petiolata, lanceolata, acuminata, margine crenato-dentato recurvato,
in facie superiore eleganter reticulato-impressa, hispidula, subtus inter nervos rubros
conspicuos dense albo-tomentosa ; petiolus ut in ramulo hispidus. Stipules infra
medium bifidse, triangulari-acutse, in margine superiore ciliolatse, uninerviaB. In-
florescentice geminatse, ssepius bis vel ter dicbotoma?, pedunculo ramisque tenuibus,
foliis 4-2-plo breviores. Cymulce ssepe floribus evolutis 7. Antherae connective
umbonato ; filamenta perianthium aequantia. Exocarpium laete-brunneum.
Leafy branchlets 2-2'5 mm. thick. Leaves 5-8 cm. long, 1'2-1*7 cm.
wide ; petiole 3-7 mm. long. The leaves have a striking appearance ; the
three main nerves are impressed on the upper face, which is beautifully
embossed with small chequer-like areas representing the transverse and
small connecting veins ; the red main nerves on the lower face stand out
conspicuously on the intensely white tomentum covering the rest of the
surface. Stipules 3'5 mm. long, united in the lower third, each with a
strong median nerve. Inflorescences generally in pairs in the leaf-axils, to
2'5 cm. long ; the small dense cymes are crowded at the ends of the short
ultimate branches, the number of flowers in each varies a typical one
is represented in fig. C, of which a diagrammatic analysis is given in fig. D.
Bracts about *5 mm. long. Flowers jointed on very short pedicels, which
are less than 1 mm. long. Bud of male flower 1'5 mm. in diameter, perianth-
segments 1'6-1'7 mm. long. Ovary slightly compressed, ovoid, with a
narrow rounded keel running along the back and bearing a row of short
forwardly-pointing setse, at the base of each of which is a black dot ;
similar but slightly larger setse surround the stigmatic disc, covering it
when young but ultimately becoming reflexed (figs. I, J). Fruit about
1 mm. long ; crowned with the withered stigmatic disc ; exocarp bright
chestnut-brown.
Hob. Arfak Mts., Angi lakes, edge of forest by ? lake, 7000'. Fl., Fr.
Dec. 5961.
A second species of this genus was collected by Mr. Kloss on Dr.
Wollaston's Dutch New Guinea Expedition. The material is scanty, but
there is no doubt that it represents a plant congeneric with the Arfak
specimen and a distinct species. A description is appended :
GIBBSIA CARSTENSZBNSIS Rendle, sp. nov.
Frutex ramulis foliiferis veluti in G. insignis hispidis. Folia longius petiolata,
elliptica, acuminata, margine crenato-dentato, recurvato, in facie superiore reti-
culato-impressa, hispidula, subtus nervis exceptis albido-tomentosa ; petiolo tenui,
hispido. Stipulce supra medium bifida3, acutse. Inflorescentia et fruclus velut in
G. insignis.
Leaves 4'5-7 cm. long, 2-2'5 cm. wide ; petiole l-2'5 cm. long; bracts
75 mm. long.
131
Hab. Dutch New Guinea :i Mt. Carstensz, 5500-6700'. Collected by
0. B. Kloss. Fr. Jan. 1913.
Distinguished from G. insignis by its relatively much broader elliptical
leaves and larger petioles ; the upper face is less conspicuously chequered.
The bracts are also somewhat larger.
FIG. 8.
Gibbsia insignis, Rendle. A. Branch, bearing leaves and inflorescences ;
B-L. Description in text. A. B. Rendle, anal. ; P. Highley, del.
132
Description of Fig. 8 (B-L) (p. 131).
B. Cymules clustered at the ends of branchlets ; X 4.
C. A typical cymule with its subtending bract, B ; the terminal female flower was
incompletely developed, a, b, lateral bracteoles, each subtending- a female flower
the lateral bracteoles of which (a', b' in fig. D) subtend each a male flower the
four male flowers indicated; the stalk of each of these male flowers bears a pair of
minute bracteoles, each of which subtends an undeveloped male flower with a
lateral bracteole. The apex of the inflorescence and the bracteoles a and b are
pushed forward away from the axis, x 15.
D. Diagram of same.
E. Male flower, with dehiscent anthers ; X 4.
F. Stamen ; X 15. G. A stamen taken from a bud, showing the umbo-like dorsal
connective ; X 15.
H. Female flower ; x 15.
I. Young stigma, covered by the ring of inflexed setae.
J. Mature stigma. I and J, x 40.
K. Fruit, showing detached upper part of exocarp (a), endocarp containing seed (b),
and cupule (c) ; X 8.
L. Ovule : X 20.
SANTALACE^S.
EXOCAEPUS sp. ?
Arfak Mts., Koebre ridge, common in forest undergrowth from 8000-
8500'. Veg. 5617.
A leafless parasite, with dark green, rigid, flattened, much branched
shoot?, about 1 m. high. The cladodes are 5 mm. broad, showing swollen
round scars alternately up the nodes, on which the flowers are evidently
borne in fascicles. I am indebted to Mr. S. Moore for this determination,
but, as he points out, in the absence of flowers or fruit the genus must
remain uncertain.
HENSLOWIA UMBELLATA Bl. Mus. Bot. Lugd.-Bat. i. (1850) 243 = //. Rein-
wardtiana Bl. cf. Koords, Excurr-ionsfl. Java, ii. 168 ; Schum. &
Laut. 300; Nova Guinea, viii. (1910) 287 ; Ridl. in Trans. Linn.
Soc. ser. 2, Bot. ix. (1916) 146.
Arfak Mts., Koebre ridge, open summit, 9000'. FL, ) 100). It differs in the
smaller leaves with much shorter tomentum underneath, the few-flowered
racemes, and the truncate to 3-lobed petals.
SERICOLEA ARFAKENSIS Gibbs, sp. nov.
Fruticulus epiphyticus, sparsim ramosus. Kami teretiusculi, innovationes fulvo-
sericei. Folia opposita, ovata, acuminata, basi rotundata, minute dentata, incurvata,
superne glabra, subtus fulvo-sericea, multinervia, reticulationibus conspicuis ; petiolo
brevi, sericeo. Raeemi abbreviati, pedunculati; floribus parvis, oppositis vel sub-
umbellatis, pedicellis gracilibus fultis. Bractece knceolatae, acuminatse. Sepala 4,
lanceolata, acuta, extus sericea, carinata. Petala 4. cuneato-obovata, apice truncata
et brevissime trilobata, basi ciliata. Stamina 1/-2, puberula. Discus 5-lobatus.
Ovarium conicum, glabrum, biloculare ; stylus subulatus ; stigma truncatum ;
ovula in loculo utroque 2, pendula, anatropa.
Hob. Arfak Mts., Angi lakes, ridge above ? lake, 8000'. Fl. Dec.
6009.
Epiphyte with hanging branches + *50 m. long, in moss-grown small
forest. Leaves 3'5 cm. including hair-like acumen + 5 cm. long by 1'9 cm.
broad. Petiole 2 mm. long. Racemes +3 cm. long, including peduncle
+ 1'4 cm. and pedicels 1*1 cm., both slightly sericeous ; flowers 4 mm., pink.
Sepals 2 - 5 mm. long. Petals 4 mm. Stamens 1'5 mm. long ; anthers shorter
than the filaments, with apical dehiscence. Ovary and style +3 mm. long.
This delicate little plant is, so far, distinct in Sencolea in the more remote
pairs of leaves and the 4-merous pink flowers.
The leaves were infested with Leptotliyrella sericolece Ramsbottom (p. 04).
Pyrsonota Ridl. and Mischopleura Wern\mm = Sericolea. In the generic
diagnosis of the latter 10 stamens are given, but in one bud of S. ovalifolia
examined 15 stamens were seen.
DlLLENIACEJS.
HlBBERTIA (SUBSESSILES) NOVO-GUINEENSIS Gibbs, Sp. HOV.
Frutex prostratus ; rami volubiles, teretes, junioribus pubescentibus, demum glab-
rati. Folia lineari-lanceolata, acuminata, in petiolum basi dilatatum longe attenuata,
Integra, chartacea, supeme glabra, inferne villosula. Flores solitarii, terminales,
subsessiles, flavi. Sepala longe ovata, acuminata, coriacea, concava, subtus villosa.
Petala 5, obovato-obtusa, integra. Stamina calyce breviora, antberis linearibus,
filamentis filiformibus. Ovaria 5 ; styli flexuosi ; stigmata simplicia, apice concava.
Hab. Arfak Mts., Koebre ridge, spreading on burnt open summit, 9000'.
Fl. Dec. 5619.
A twining woody plant with large yellow flowers. Largest leaves +8'5
by 1 cm., the midrib much impressed on the upper surface and prominent on
the lower. Flowers +4 cm. across. Sep;ils unequal, l'j-1'8 cm. by 9 mm.
Petals 1'5 by l'2-3 cm., equalling the sepals. Stamens unequal, red-brown
in colour (dried), the longest 6 mm. long; anthers 2 mm. long, with apical
149
pores ; filaments 4 mm. long. Ovaries 4 mm. and styles 6 mm. long. The
material is insufficient to determine the number of ovules in each ovary.
A species which resembles in habit and flowers //. scandens (Willd.) Gilg,
from Queensland and New South Wales, but is at once distinguished by the
more linear leaves and smaller flowers, and the longer stamens with shorter
anthers and longer filaments. This is the first record of the genus from
New Guinea and adds another to the long list of genera so far considered
limited to Australia. The genus is also common to New Caledonia, and
further investigation will no doubt prove it to be widely spread in New
Guinea.
GUTTIFER^E.
HYPERICUM MUTILUM L. Sp. PI. ed. 1, 787 ; cfr. Maximowicz in Bull. Ac.
Sc. St. Petersburg, xi. (1881) 171.
Arfak Mts., Angi lakes, 7000', open marsh by ? lake. Fl. (yellow),
Fr. Dec. 5963.
Distrib. New Guinea (D.S.W., Mt. Carstensz, Kloss; N.E.). Sumatra,
Java, N. Borneo, Philippines, Central and S. China; New Zealand and
Tasmania. Madagascar. N. and S. America.
VIOLACE.E.
*ViOLA DISTANS Wall. Cat. n. 4022.
Arfak Mts., Angi lakes, open marsh by ? lake, 7000', creeping amongst
grass. Fl., Fr. Dec. 5962.
Distrib. Temperate Himalayas, Nilghiri Mts. to Ceylon. S. China.
Flowers white, striped violet on lower lip. Drawings of the style on
herbarium sheets at Kew and the British Museum, including some excellent
dissections by the late C. B. Clarke, show complete identity with the above
specimens, which only differ in the so far recorded colour of the flowers
(violet and blue).
BEGONIACE^E.
SYMBEGONIA ARFAKENSIS Gibbs, sp. nov.
Herba, caule erecto in sicco rufo, pilis paucis longis suberectis pubescentibus
onusto. Folia sessilia, oblique lanceolata vel ovata, basi uniauriculata, apice angus-
tata acuta, grosse serrato-dentata, supra glabra, subtus albo-punctata, in nervis
rufo-pubescentia. Inflorescentia terminalis, a basi ramosa, bracteis multis, latis,
apice obtusis, albidis. Flares d 1 sepalis 2 ; staminibus haud multis in columnain
gracilem connatis, supra liberis ; antheris globosis. Fructus solitarii vel bini,
pedicellati, 3-alati, pedunculo fructifero puberulo, alis apice divaricatis, protractis*
acutis, sparsim serratis. Placenta bitida.
Hob. Arfak Mts., Angi lakes, in forest by ? lake. Fl. h
papery cortex, very glandular on the young wood. Leaves' + 1 cm. by
8 mm., with midrib slightly impressed on the upper surface and indistinct
on the lower; petioles 4 mm. long, glandular. Peduncles sparsely villous
when young, 2-3 mm. long. Flowers subtended by two villous, narrow-
linear bracts 3 mm. long. Calyx-lobes villous when young, 2 mm. lung,
narrowing from a broad base. Petals glandular, 4 mm. by 1 mm. Stamens
as long as or just exceeding the petals ; anthers globose. Style thick,
152
5 mm. long. Ovary 1'5 mm. long. Fruit gland-pitted, 2'G mm. by 3 mm.
Seeds large, 2 mm. long.
This plant is nearest to M. rufo-punctata Panch., a New Caledonian
species, but differs in tbe prostrate habit, ovate leaves, and pedunculate
flowers.
MYRTUS (AUSTROMYRTUS) ARFAKENSIS Gibbs, sp. nov.
Frutex prostratus, valde ramosus, glaberrimus et glanduligerus. Folia opposita,
petiolata, late elliptica, basi subattenuata, apice obtusa vel emarginata, margine
revoluta, coriacea, supra in sicco fuscescentia, subtus pallidiora, pagina utraque
glandulis rufis prominentibus conspersa. Flores cymosi, pedicellati, bibracteolati,
cymis axillaribus, subpedunculatis, 3-floris vel solitariis. Calyx 4-lobatus, lobi
triangulares, obtusi. Petala 4, flavida, unguiculata. Stamina oo , biseriata. Stylus
filiformis. Ovarium biloculare, ovulis biseriatis. Bacca carnosa.
Hob. Arfak Mts., Angi lakes, S.W. ridge, on open steep slopes, 8000'.
Fl., Fr. Dec. 5994.
In this plant the stems are terete, with the older wood dark brown in
colour, the younger reddish and rugulose with larger glands ; the small
leaves are closely arranged up the branches, the younger shoots being
reddish in colour. The leaves are +7 mm. by 5 mm., but with the incurved
margins appear 3 mm. across ; the midrib is thick, somewhat raised on the
u-iderside but not conspicuous. Petiole thick and glandular, 1'3 mm. long.
The flowers form little subsessile cymes, or are on solitary pedicels + 1-3 mm.
long, with small, triangular, translucent, gland-dotted bracts at the base of
the calyx. Calyx-tube 2 mm. long, with obtuse lobes 1'5 mm. long and the
same in breadth at the base. Petals suborbicular, +2 mm. by 1'5 mm.
Stamens 4'5 mm. long, with minute globose anthers. Style 4 mm. long.
Berry black, crowned by persistent calyx-lobes, 5 mm. by 5 mm.
This plant is nearest to M. prostrata, described above, and in the field
was very similar in habit and appearance. It is easily distinguished by
the entire glabrous and more glandular leaves, tbe longer stamens, and
bilocular ovary.
MYRTUS (AUSTROMYRTUS) KOEBRENSIS Gibbs, sp. nov.
Ffutex vel arbor parva, ramosissima ; ramis cortice cinerascente obductis, tota
planta utrmque glaberrima. .Folia parva, ovata, obtusa; basi in petiolum breveiu
attenuata, margine leviter recurvata, glandulosa, coriacea. Flores 4-meri, axillares,
solitarii vel bini, pedicellis foliis minoribus. Calyce tubo turbinato, basi bibracteolato.
Sepala late obtusa, glandulosa. Petala flava, reflexa, basi unguiculata. Stamina
biseriata. Ovarium biloculare. Ovula biseriata.
Hob. Arfak Mts., in shrubberies on open summit of Koebre Mt., 9000'.
Fl. Dec, 5614.
A compact shrub to small tree, with very small leaves +'9 mm. by
5 mm., smooth and shining above, with the midrib indistinct on both
surfaces ; petiole 1 mm. long. The flowers arise in the axils of the leaves
153
all along the branches, on pedicels 4 mm. long, with two small hracts at the
base. Calyx-lobes triangular, 1 mm. long and 1 mm. broad at the base,
obtuse to acute and erect in flower. Petals 2*5 by 1'5 mm., the lamina
reflexed in flower. Stamens with filaments 3 mm. long j anthers minute.
Style 3*5 mm. long. Ovary 2 mm. long by 1 mm. Pedicels, calyx, and
ovary thickly gland-dotted.
This plant approaches some forms of M. tenuifolia (Sm.), Mez, but is
distinguished by the glabrous glandular habit, much smaller leaves, and
the yellow 4-merous flowers.
JAMBOSA (CLAVIMYRTUS) ARFAKKNSIS G-ibbs, sp. nov.
Arbor parva ; ramuli teretes, saturate brunnei. Folia coriacea, opposita, petiolata,
oLovata, spathulata, rigida, verisimiliter impellucida, glabra, leviter recurvata,
basi attenuato-cuneata, apice obtusa vel emarginata, supra crebre punctata, subtus
]jarit, fronds 1 m., naked stipes 50 m., young plantlets make roots on
parent plant."
Distrib. New Guinea (D.N.W., Ramoi, Beccari-, D.S.W., Hellwig-Gebirge,
ron Roemer ; N.E. ; S.E.). Malay Islands. Polynesia. Trop. Australia.
HYPOLEPIS GRANDIFRONS Gepp, sp. nov.
Frons maxima subdeltoidea (pinna basalis c. 72 cm., pinna media c. 92 cm. longa)
tripinnata ; pinna? primariae lanceolate acuminate 22-25 cm. latae ; pinna? secundaria}
alternse c. 40-jugate breviter stipitatae, intervallis c. 2'5 cm. insert*, lineari-lanceolate
acuminate usque ad 12*5 cm. longe 2'5 cm. late ; pinnule alternae 20-22 -jugate
breviter stipitate oblongse obtusissime pinnatifide, lobis anticis 5 et posticis 4
instructs, apice flabellatae ; lobi oblongi paucicrenati vel integri, venulis 3 anticis
1 postico percursi ; lobus anticus inferior duplo major, venulis 3-jugatis furcatis
percursus. Sori in lobis solitarii, in sinu marginis antici positi, lobulo (indusio)
parvo luteo-fusco involute obtecti ; in lobo antico inferiore duo sori adsunt.
Sporangia circum penicillum paraphysium articulatarum disposita. Textura
membranacea ; color valde viridis. Rhachis primaria (? 1 cm. crassa) strain im-a ;
rliachis secundaria et tertiaria atque pinnularum coste supenie sulco pilis articulatis
brevibus villoso exaratae ; costa et venule pilis paucis hinc illinc instructae.
Hab. Humboldt Bay, ridge behind " campong," 500', terrestrial by stream
in high forest. Jan. 6258. "Magnificent single fronds from underground
creeping rhizome, 5 m. long, petiole 3 m., lamina 2 in. deltoid."
The huge fronds, the stipitate pinnules, and the few sori distinguish this
species from the rest. The material consists of two pinnae with fragments of
the main rhachis attached.
PTERIS (EUPTERIS) BAMBUSOIDES Gepp, sp. nov.
Rhachis erecta crassa atro-purpurea subnitens minute pubescens. Pinnae inter-
vallis circa 4 cm. insert*, erecto-patentes, alternse (?) brevi-stipitatae, ad basin imam
1-2 -dichotomy, segmentis usque ad 40 cm. longis e basi cuneata linearibus sensim
attenuatis, fertilibus integris, sterilibus serrulatis, versus apicem spinuloso-serratis,
margine incrassata venuliformi, lamina nitente, costa inferne plerumque rubella et ad
latera saepe pubescenti, venulis plurimis (circa 33 in centimetri spatio) conspicuis
simplicibus furcatisve. In Just urn angustum brunneolum membranaceum.
Ildb. Arfak Mts., inundation area of Momi River, and common down to
Wariap. Dec. 5732. "Each shoot up to 3 m. tapering; growing in
clumps. Pinnre on young fronds plane, but on rhachis twisting in growth,
finally arranged spirally."
196
The material collected consists of a longitudinal half of a fragment of
rhachis, about 38 cm. long, with pinnae attached. The pinna-segments
closely resemble the pinnse of P. moluccana, but differ, of course, in being
dichotomously disposed at base.
The remarkable fronds of this fern, rising to a height of 3 m. from the
ground and growing in clumps, must form a conspicuous feature in
the savannah landscape. It is surprising that so well developed a species
should not have been recorded previously, unless, indeed, its distribution is
extremely limited.
PTERIS TORRICELLIANA Christ.
Humboldt Bay, ridge behind " campong/' 400', by stream in high forest,
undergrowth. Jan. 6252. " Petiole 1'50 m., frond 1 m. long."
Distrib. New Guinea (N.E.).
VlTTARIA ELONGATA Sw.
Humboldt Bay, ridge behind "campong," 200', epiphytic by stream in
high forest. Jan. 6254. " Pendent from rock, fronds 2'80 m. long."
Distrib. New Guinea (D.N.W., Soron, Ramoi, Beccari ; D.S.W., coastal
lowlands, Versteeg ; N.E. ; S.E.). Trop. Asia. Polynesia. N.E. Australia.
ANTROPHYUM RETICULATUM Kaulf.
Manokoeari, track to Ambani, 700', epiphytic in high forest. Jan.
6193.
Distrib. New Guinea (D.S.W., coastal lowlands, Versteeg ; N.E. ; S.E.).
Madagascar to Polynesia and tropical Australia.
POLYPODIDM (PLEOPELTIS) NORMALE Don.
Humboldt Bay, ridge behind " campong/' 500', common, climbing in high
forest. Jan. 6271.
Distrib. New Guinea (N.E.). Trop. Asia. China. Madagascar.
*LYGODIUM DIGITATUM Presl.
Manokoeari, track to Ambani, 500', common in forest and clearings.
Jan. 6208.
Distrib. Philippine Islands. Malacca.
ANGIOPTERIS EVECTA Hoffm.
Humboldt Bay, ridge behind " campong," 500', very common by stream
in high forest. Jan. 6259.
Distrib. New Guinea (D.N.W., Arfak Mts., Beccari; D.S.W., Mt. Car-
stensz, Kloss ; S.E.). Tahiti. Trop. Asia. Africa.
OPHIOGLOSSUM PEDUNCTJLOSUM Desv.
'Dammar Island, in open " kebun " near seashore. Jan. 6289.
Distrib. New Guinea (N.E.). Trop. Asia. Australasia.
197
LYCOPODIALES.
LYCOPODIUM PHLEGMARIOIDES Gaud.
Manokoeari, Langgen, epiphytic on tree hanging over sea. Dec. 6221.
Distrib. New Guinea (N.E.). Malay Islands. Polynesia.
LYCOPODIUM CERNUUM L.
Island of Roon, young plants, spreading in the open, in shade by road
round bay. Jan. 6225.
Distrib. New Guinea (D.N.W., Arfak Mts., Beccari ; D.S.W., Mt. Car-
stensz, Kloss ; N.E. ; S.E.). Tropics and some subtropics.
PsiLOTUM FLACCIDUM Wall.
Schouten Islands, Bosnik, Wiak, epiphytic on strand trees. Jan.
6273.
Distrib. New Guinea (D.N.W., Arfak Mts., Beccari ; D.S.W., Noord
River, Versteeg ; Mt. Carstensz, Kloss ; N.E.), Tropics.
SELAGINELLA PLUMOSA Baker.
Humboldt Bay, ridge behind " campong," creeping in high forest, 500'.
Jan. 6265.
Distrib. New Guinea (N.E.). Solomon Islands. Trop. Asia.
SPERMATOPHYTA.
CYCADALES.
CYCADACE^E.
CYCAS CIRCINALIS L. Sp. PI. ed. 1, 1188 ; F. Muell. Pap. PI. ii. 71 ;
Schum. & Laut. 153 ; Nova Guinea, viii. (1910) 343.
Schouten Islands, Wiak, Bosnik, under trees on seashore. Veg. Jan.
6278.
Distrib. New Guinea (D.S/W., Noord River, Versteeg ; S.E. : N.E. and
adjacent islands). S. Asia to Polynesia.
MONOCOTYLEDONS*;.
P A N D A N A C E M. (A. B. RENDLE.)
FREYCINETIA OBLANCEOLATA Martelli in Webbia, iii. 176.
Manokoeari, "korang" forest, 500'. Fl., . Jan. 6158. Spathes
light green. Fl., ? . 6157. Spathes pink-green.
Distrib. New Guinea (D.N.W., Dore, Teysmann).
The specimens agree with Martelli's description, except that the number
of the stigmas varies from 2-4 in the above specimens.
198
FREYCINETIA BECCARII Solms-Lanb. in Ann. Jard. Buit. (1883) iii. 100.
Manokoeari, Langgen, scandent on land-edge of mangrove association.
Fr. Jan. 6219.
Distrib. New Guinea.
Fruit red-brown, 2'5 mm. long, not quite ripe.
PANDANUS DUBIUS Spreng. Syst. Veg. iii. (182G) 897 ; Schum. & Laut. 159.
Manokoeari, Genbela, plentiful by cape, on edge of sandy beach. Jan.
6215. Wakde Island, gregarious on edge of beach. Fr. Jan. 6248.
Distrib. New Guinea and adjacent islands. Java, Borneo, Moluccas,
Celebes, Philippines, Carolines, Marianne Islands, and New Hebrides.
*PANDANUS POLTCEPHALUS Lam. Encycl. i. (1785) 372.
Manokoeari, Langgen, common in mangrove association. Fr. Jan.
6230.
Distrib. Amboina, Batjan, Ceram, Timor, Sumatra.
" Plant + 10 m. high, with branched head. Fruit red."
PANDANUS TABBERSIANUS Rendle, sp. nov.
Arbor parva. Folia valde spiraliter ordinata, e basi dilatata linearia, superne
gradatim attenuata, basin versus late canaliculata cseterum plana ; marginibus in
parte basilari nudis caeterum dentibus minutis aeutis crebris munitis ; costa media
in facie inferiore prominente, velut in margine denticulata. Syncarpium magnum,
pendulum, pendunculo longo suffultum, anguste lanceolato-ellipsoideum, axe fibroso-
lignoso, spathis plurimis indutum. Spathce lineari-lanceolatse vel lanceolatse, sub-
yequilongaB, coriaceae, extus leves, carinatse, marginibus et carina (basi excepta) velut
in foliis denticulate. Drupce mature brunnea?, numerosissime, confertse, irregu-
lariter 5-7-gonse, parte apicale libera et infra pileum depresso-pyramidatum angulosum
constricta ; stigma laterale, horizontale, plus minus compressum superne autem rotun-
datum vel interdum bilobatum ; endocarpium osseum, anguste ellipsoideum, basi
angustatum, mesocarpium inferne fibrosum.
Hab. Manokoeari, forest edge, track to A mbani, 100'. Fr. Jan. 6213.
Plant 5 m. high. Leaves 3 m. long, 13 cm. wide at the base, 6 cm.
wide about the middle ; teeth l'5-2 mm. long. Syncarp 4 dm. long,
1 dm. in greatest diameter, core up to 5 cm. in diameter ; peduncle 3'7 dm.
long. Drupes 1'8-2'1 cm. long, 4-5 mm. thick, upper free part about
*5 cm., cap 22*75 mm. long ; endocarp about 1 cm. long.
From the description evidently near P. Englerianum Martelli from Neu-
Mecklenburg, which, however, has a much larger trigonous syncarp, scarlet
drupes, and a discoid stigma. The syncarp of our species is described as of
a uniform chocohite-brown colour.
This plant has been named in honour of Mr. (now Capt.) Tabbers, Acting
Assistant Resident at Manokoeari, to whose ready help, practical experience,
and judgment Miss Gibbs was much indebted during her stay.
199
GR AMINES. (A. B. RENDLE.)
CENTOTHECA LAPPACEA Desv. in Journ. de Bot. (1813) 70 ; Schum. & Laut.
185 ; Ridl. in Trans. Linn. Soc. ser. 2, Bot. ix. (1916) 250.
Island of Roon, ridge beyond Djende, common undergrowth in forest,
400'. FJ. Jan. 6234.
Distrib. New Guinea (D.S.W., Utakwa River, Kloss; N.E. and adjacent
islands). West Africa, through India, Malesia, and Polynesia.
GIGANTOCHLOA NOVO-GUiNEENSis Rendle, sp. nov.
Culmus arborescens, ramis teretibus glabris. Folia subsessilia, lineari-oblonga,
apice subabrupte acuminata, ad basin valde obliquam et inaequaliter cordatam paullo
angustata, multinervia sed haud conspicue, baud tessellata, utrinque glabra, margine
scabridula ; vaginae tubuliformes, persistentes ; ore auriculatae, ciliats ; ligula angusta,
ciliata. Inflorescentiae terminales et axillares, rbachi elongata, ramis multo brevi-
oribus, suberectis, spiculis capituliforme fasciculatis, capitulis distantibus, ad apicem
rami interdum glorneratis. Spicula numerosissimte, dense confertae, lanceolate,
compressse, pluriflorse ; glumce inferiores, saepe 2, steriles, fertilibus minores, late
ovatae, apice mucronate, rnultinerviae, in margine superiore sparse ciliolatse ; glumce
ft- 1- tiles 2-5 distichae, 1 vel 2 superiores imperfects?, late ovatae, breviter acuminatie,
apice pungente, in margine superiore ciliolatae, inultinervise ; palea glumse subsequalis
vel paullo brevior, bi-carinata, carinae minute ciliolatae, lamina inter carinam et inar-
ginem 3- vel 2-nervata, pars inter carinas haud nervata. Lodiculee 0. Stamina 0,
exserta, fikmenta pro majore parte longitudinis in tubum tenuiter membranaceum
demuin valde fragilem connata ; antherse anguste lineares, obsolete apiculatie.
Oniriiim breviter pedicellaturn, anguste pyramidatuni, glabrum, stylo elongate,
filiforme, superne in stigmata plumosa 3- vel 2-diviso.
Culms about 16 ft. high ; the young culm about 1 cm. thick ; sheaths
membranous, tubular, 11-12 cm. long, with margins ciliate above, and a
truncate auricled apex bearing a fringe ot long erect stiff cilia, 1*8 cm. long,
small leaf-blade, subsessile, ovate, acuminate, with shallowly cordate base.
Fully formed leaves on (flowering shoots 4-4'5 dm. long, 7-9 cm. wide,
on the young leafy shoots up to 6 dm. long, 9'5 cm. wide ; apex acuminate,
ending in a twisted scabrous point; ligule a ridge hearing stiffish ciKse
which are about 1 cm. long. Rhachis and branches of inflorescence smooth,
glabrous; the rhachis up to 6 dm. long; spikelets crowded in somewhat
distant heads, l'2-2 cm. long, which are generally 3-5 cm. apart, but closer
toward the end of the axis, where they sometimes unite to form a spherical
compound head. Branches of inflorescence and heads subtended by the dry,
In-own, persistent bract-sheath, which is about equal in length to the head.
Spikelets about 1 cm. long, 4 mm. broad. Lower barren glume 4 mm. long,
9-nerved, upper 6 mm. long, 11-nerved ; flowering glumes 8-9 mm. long.
A very distinct species, suggesting Oxytenantkera in habit of in-
florescence.
Hab. Manokoeari, open cultivated slopes above "campong," 100'. Jan.
6270 a & b. Humboldt Bay, beyond " campong " in the open and along
stream. Jan. 6266 a & b.
200
CYPERAOE^. (A. B. RENDLE.)
THORACOSTAOHYUM HYPOLYTROIDES C. B. Clarke in Hook. . Fl. Brit. Ind. vi.
(1894) 680.
Humboldt Bay, by "campong," swamp behind beach, abundant. Fl.
Jan. 6249.
Distrib. New Guinea. Asia and tropical Australia.
*SCLERIA MARGARITIFERA Willd. Sp. PI. iv. 312.
Manokoeari, edge of forest, 200'. Fl., Fr. Jan. 6166. Island of
Roon, near Djende, in the open, by road round the bay. Fr. Jan. 6239.
Distrib. Polynesia. Australia.
PALM.E. (0. BECCARI.)
ARENQA MTCROCARPA Becc. in Schum. et Hollr. Fl. von Kaiser Wilhelms
Land (1889) 16. A. gradlicaulis Bailey in Queensl. Agric. Journ. iii.
pt. 3 (1898) 202. A. microsperma (sphalmate pro A. microc.arpa)
Becc. in Rechinger Bot. u. Zool. Ergebnisse, etc. v. (1913) 64, et in
Schum. et Laut. Fl. deut. Schutzg. in d. Siidsee, 204. Didymosperma
microcarpa Warb. in Mons. ined. ex Schum. et Laut. 1. c. 204. D. novo-
guineensis Warb. in Mons. ined. ex Schum. et Laut. 1. c. D. humile
Laut. et K. Schum. in Schum. et Laut. 1. c. 204. ? Sayuerus austra-
lasicus Wendl. et Dr. in Linnsea, xxxix. (1875) 219.
Humboldt Bay, by river near " campong." FL, Fr. Jan. 6268.
Distrib. N.E. New Guinea and N.E. Australia.
CARYOTA RUMPHIANA Mart. Hist. nat. Palm. iii. 195 ; Blume, Rumphia, ii.
140 ; Becc. Malesia, i. 70, 74.
Dammar Island. Fl., Fr. Feb. 6288.
Distrib. Moluccas.
LICUALA MONTANA Dammer et K. Sch. in Schum. et Laut. Fl. deut. Schutzg.
in d. Siidsee, 200 ; Becc. in Webbia, i. 291 (in note).
Humboldt Bay, ridge behind the "campong." Fl. Jan. 6262.
Distrib. N.E. New Guinea.
PIGAFETTA FiLARis Becc. Malesia, i. 89 (Pigafettia). Metroxylon filare Mart.
Hist. nat. Palm. iii. 216 & 343. Sagus Jilaris Bl. Rumphia, ii. 154 &
128. Pigafettia papuana Becc. Malesia, i. 89.
Manokoeari, ridge behind, single specimens in forest, 500'. Jan. 6180.
Distrib. New Guinea (D.N.W., near Andai, Beccari}. Moluccas.
201
A RACEME.
HOLOCHLAMYS BECCARii Engl. in Malesia, i. 265 ; Schum. & Laut. 212 ;
Nova Guinea, viii. (1912) 806.
Humboldt Bay, ridge behind "campong," 500', undergrowth in high
forest. Fl. Jan. 6261.
Distrib. New Guinea (D.N.W., Ramoi, Andai, Beccari ; D.S.W., Hellwig
Mountains, von Roemer [fl. Nov.] ; N.E.). Aru Islands (Mostley,
' Challenger' Exp. Herb. Kew.).
Spadix green when collected with only remains of spathe.
CYRTOSPERMA MACROTUM Becc. (MS.) ex Engl. in Bull. Soc. Tosc. Ort. iv.
(1879) 295; Malesia, i. 279, t. xxiv. figs. 1-6; Nova Guinea, viii.
(1910) 249 ; Ridl. in Trans. Linn. Soc., ser. 2, Bot. ix. (1916) 24.
Manokoeari, track to Ambani, undergrowth in high forest, on "korang."
Fl. (yg.). Jan. 6211.
Distrib. New Guinea (D.N.W., Batanta, Beccari ; D.S.W., Noord River,
Versteeg, Brandenhorst, marsh land, von JKoemer ; Mt. Carstensz, Kloss ;
S.E.).
Plant 1 m. high. Spathe white.
SCHISMATOGLOTTIS DORENSIS Gibbs, Sp. DOV.
Herba parva ; caudiculus hypogaeus. Cataphylla lineari-lanceolata, mem-
branacea. Folia oblongo-lanceolata, acuininata, basi breviter late cordata, supenie
Mavo-viridia, subtus pallida, nervis 16, vagina lata. Spatha solitaria, albida, basi
constricta, lamina convoluta, dilatata, cuspidata. Spadicis pars feminea triente
inferior! dorso spathae adnata, superne laxiflora, inflorescentia mascula a feminea
interstitio nudo separata. Staminum filarnenta quam antherse paullo longiora ;
thecae obovoidese, concavo-rufescentes. Staminodia truncata, ruf o-punctata. Ovaria
oblonga, ovoidea, stigmate parvi sessili coronata, ovula plurima.
Hob. Manokoeari, terrestrial in secondary jungle, forming large patches.
200'. FL, Fr. Jan. 6167.
Plant +3 dm. high. Largest leaf +1*5 dm. by 5 cm., acumen 1 cm.
Petiole 1'8 dm. long, vagina 3'5 cm. Peduncle 7 cm. long ; cataphylls
+ 2'5 cm. long. Spathe white, 3'6 cm. long, basal portion 1*5 cm. by
4 mm. Spadix white, 2 cm. long ; ? portion cylindric, 1'2 cm. by
2'5 mm., constricted above ; upper part club-shaped, 1 cm. by 4'5 mm., with
stamens below and staminodes at the apex. Fruit green (when collected),
2 cm. by '7 mm.
Near S. Klossii Ridl., but differs in the longer petioles, larger and broader
leaves, longer peduncles, and the naked interval between the upper and lower
portion of the spadix. Mr. N. E. Brown was good enough to give me big
opinion on this plant as a new species.
202
AGLAONEMA NOVO-GUINEENSIS Engl. in Bot. Jahrb. xxv. (1898) 22 ; Sohum.
& Laut. 214.
Manokoeari, undergrowth in damp secondary jungle, 100'. Fl., Fr.
Jan. 6164.
Distrib. N.E. New Guinea.
Stem 1 m. high, with leaves aggregated at the top. Spathes green,
sometimes bipartite ; spadix white ; fruiting spathes green, fruit red.
ALOCASIA ACUTA (Engl.) Hall. f. in Bull. Herb. Bois. vi. (1898) 605 ;
Malesia, i. 294 ; Schum. & Laut. 214; Nova Guinea, viii. (1910) 251.
Manokoeari, common on edge of forest, where damp, 200'. FL, Fr.
6190.
Distrib. New Guinea (D.N.W., Andai, Beccari; D.S.W., Noord River,
Versteeg ; N.E.). Skru and Aru Islands, Batyan (Herb. Kew.).
A very striking plant with stem 2 m. high, the large fleshy leaves aggre-
gated at the top, bearing numerous flowers, on short peduncles, in their axils.
The greenish spathes enclose the spadix, which is white at the top (<$}, then
red (sterile), and green at the base ( ? ). The fruit contains 2-6 bright red
seeds, with fruiting spathes red at the base and green above.
FLAGELLARIACE^E.
FLAGELLARIA INDICA L. Sp. PI. ed. 1, 333; F. Muell. Pap. PI. i. 73;
Ri.il. in Journ. Bot. xxiv. (1886) 358 ; Schum. & Laut. 215 ; Ridl.
in Trans. Linn. Soc. ser. 2, Bot, ix. (1916) 231.
Manokoeari, scandent on the highest trees in the forest and in open
clearings. Fl. Jan. 6210.
Distrib. New Guinea (D.S.W., Mt. Carstensz, Kloss ; N.E. ; S.E. and
adjacent islands). Tropical Asia, Africa, and Australia.
C M M E L I N A C E M.
POLLIA SORZOGONENSIS (E. Mey.) Endl. Gen. PL (1841) 368 ; Schum. & Laut.
216; Nova Guinea, viii. (1913) 907; Ridl. in Trans. Linn. Soc. ser. 2,
Bot. ix. (1916) 231.
Manokoeari, Langgcn, undergrowth in secondary jungle. Fl. (white).
Jan. 6222.
Distrib. New Guinea 'D.N.W., Salt Spring, Begowri River, Gjellehtp ;
D.S.W., Mt. Carstensz, Kloss; N.E. and adjacent islands). India and Ceylon;
Indo-China; Formosa ; Malaya, N.E. Australia, and New Caledonia.
Li LI ACE^E.
DRACAENA ANGUSTIFOLIA Roxb. Fl. Ind. ii. 155 ; Ridl. in Journ. Bot. xxiv.
(1886) 357 ; Schum. & Laut. 220 ; Nova Guinea, viii. (1914) 1002.
Humboldt Bay, edge of forest. FL, Fr. Jan. 6250.
Distrib. New Guinea (D.N.W., Humboldt Bay, Bogowri and Tor rivers,
203
Gjellerup; D.S.W., Merauke, Versteeg ; Okaba, Brandenhorst ; N.E. ; S.E.).
India. Indo-China, Philippines and Formosa, Malay Archipelago and
N.E. Australia.
Common, but not in flower about Manokoeari.
DRAOENA NOVO-GUINEENSIS Gibbs, sp. nov.
Caulis simplex, fmticosus. Folia petiolata, ascendentia, spiraliter arete imbri-
cata, lineari-lanceolata vel oblanceolata, acuta vel acuminata, membranacea, ^iridia,
oblique parallele costata. Panicula longe pedunculata, quam folia brevior. Flores
solitarii vel bini, pedicellis graciliter dispositis. Bractece membranacese, deltoideo-
acuminatae. Perianthium albo-viridulum, lobis ultra medium liberis, obtusis, apicem
incurvatis, tubo basin versus angustato. Stamina gracilia, filamentis complanatis,
antheris basi divaricatis. Ovarium sessile : stigma 3-lobatum. Bacca disperma.
llab. Manokoeari, common undergrowth in high forest on " korang."
500'. Fl., Fr. Jan. 6195.
A very characteristic little plant, '25--50 m. in height, well marked by
the spirally twisted leaves, like a screw pine in appearance. Leaves 3-4*6 dm.
in length, including linear petiole + 1 dm. long, slightly broader and sheathing
at the base, by 3-3'5 cm. broad, the shorter leaves being the broadest. In-
florescence l'3-2'7 dm. long, including peduncle 1-1*4 dm. long and 6 cm.
broad, generally branching three times, but in the single fruiting example
unbranched and epedunculate. Bracts +3 cm. long, those subtending
flowers 3 mm. long ; bracteoles 2 mm. Perianth 1'3 cm. long, with
segments 7 mm. long and 2 mm. broad. Stamens 5 mm. long ; anthers
3 mm. ; filaments 2'5 mm. by 1 mm. Style 1*1 cm. long, with minute stigma.
Ovary 2 mm. long Fruit 6 by 7 mm..
This species seems distinct from the Malayan Dracaenas, so far known, in
the lengthily petiolate spirally arranged leaves, with very straight linear
petioles, and the strap-shaped filaments to the anthers. This is the first
member of the genus to be described from New Guinea.
R C H I D A C E JE. (J. J. SMITH.)
VRYDAGZYNEA ELONGATA Bl. Fl. Jav. n. ser. 1, 61, t. 28, f. 1, etc.
Island of Roon, ridge beyond Djende, terrestrial in forest. Fl. Jan.
6240.
Distrib. N.E. New Guinea.
HIPPEOPHYLLUM ALBOViEiDE J. J. S. in Fedde Rep. xi. (1912) 135 ; in
Nova Guinea, xii. (1915) 217, t. Ixxi. 123.
Schouten Islands, Island of Wiak, Bosnik, epiphytic on strand tree.
Fl. (white). Jan. 6283.
Distrib. New Guinea (D.N.W., Tor River, Gjellerup).
Q
204
LIPARIS ( EOLIPARIS) MABOROENSIS Schltr. Orch. D. Neu-Gruinea, (1911) 18G.
Var. BISTRFATA J. J. S., var. nov.
Planta minor. Folia 4-5'4 cm. longa, 2'3-3'2 cm. lata. Inflorescentia 1-flora.
Sepala et petala c. 0'475-0'44 cm. longa. Labellum 0'56 cm. longurn. Ovariuin
pedicelktum c. 0'6 cm. longum.
Manokoeari, tract to Ambani, terrestrial in " korang " forest, 800'
above the sea. Fl. (green). Jan. 6187.
Distrib. N.E. New Guinea.
I think this plant is L. maboroensis Schltr. I have kept it apart because
of the smaller flowers and the two purple lines on the labellum.
MICROSTYLIS ( CREPIDIUM) XANTHOCHILA Schltr. in Schum. et Laut. Nachtr.
Fl. d. Schutzgeb. Siidsee, (1905) 102, etc.
Manokoeari, tract to Ambani, terrestrial in high forest on " korang/'
800'. Fl. (orange-green). Jan. 6188.
Distrib. N.E. New Guinea.
MICROSTYLIS ( CREPIDIUM) GTIBBSLE J. J. S., sp. nov.
Caulis e basi decumbente radicante adscendens, c. 11-folius. Folia oblique
subovato-oblonga, plus minus falcata, subacuta, apiculata, minute undulata, nervis
conspicuis c. 3 subtus prominentibus, c. 4'5-5 cm. longa, 1'6-1'7 cm. lata;
petiolus canaliculatus, cum vagina c. l'G-1'7 cm. longus. Inflorescentia erecta, laxe
multiflora, pedunculo c. 5'25 cm. longo, vaginulam bracteiformem 1 gerente, rhacbide
6 cm. superante. Bractecs parvae, reflexae, oblongo- ad lanceolato-triangulse, aeutae,
ad c. 0*27 cm. longa?. Flores patentissimi. Sepalum dorsale subovato-ovale,
obtusum, convexum, apice concavum, 3-nervium, fere 0'3 cm. longum, 0*175 cm.
latum. Sepala lateralia reflexa, oblique orbiculari-ovata, obtusa, basi convexa,
superne concava, 3-nervia, c. 0'26 cm. longa, fere 0'2 cm. lata. Petala oblique
ligulata, obtusa, 1-nervia, c. 0'25 cm. longa, 0'08 cm. lata. Labellum hippocrepidi-
forme, fovea triangula concava incrassatione triangula lata convexa circumdata,
incrassatione parva ad basin, explanatum totum c. 0'43 cm. longum, lobis lateralibus
obsoletis vel subobsoletis, obtusis, lobo intermedio late triangulo, in lobos 2 oblique
triangulos acutos margine interiore repandulos bifido, laciniis 4 elongatis subulatis
praeserfcim interioribus falcatis et lobum intermedium superantibus ad bene O'l cm.
longis exterius descrescentibus utrinque, auriculis majusculis, anguste oblique trian-
gulis. acutis vel acutiusculis, fere 0'2 cm. longis, basi c. 0'08 cm. latis. Gynostcmnim
a dorso compressum, oblongo-quadrangulum, apice dilatatum, dorso convexum, bene
O'l cm. longum, clinandrio concavo, auriculis divergentibus, antheram superantibus,
oblongis, truncato-obtusis, intus convexis, carnosulis. Anthera cucullata, transverse
quinquangulari-reniformis, obtusissima, 0'06 cm. lata. Bostellum latum, retusum.
Ovarium pedicellatum c. l-l'l cm. longum.
Hob. Manokoeari, tract to Arnbani, terrestrial in " korang " forest,
700'. Dec. 6192.
The nearest allies of this species are M. pedicellaris Rchb. f., M. pectinata
J. J. S., and M. wariana Schltr., which all are characterized by the elongate
pedicel.
205
It differs from the insufficiently described M. peduellaris in the inflor-
escence showing a distinct naked portion at the base, in the pedicels which
have 3-4 times the length of the sepals, and in the teeth of the lip ; from
M. pectinata by the more numerous and smaller leaves, smaller flowers,
longer auricles of the lip, etc.; and from M. wariarta by the more numerous
smaller leaves, the differently coloured flowers, longer auricles to the lip, etc.
The leaves appear to have been coloured.
The flowers are said to be purple.
DENDROBIUM ( CADETIA) POTAMOPHILUM J. J. S. in Bull. Jard. Buit. 2 e ser.
n. viii. (1912) 18. Cadetia potamophila Schltr. Orch. D. Neu-Guinea,
(1912) 438.
Schouten Islands, Wiak, Bosnik, epiphytic on strand tree. Jan. 6282.
Distrib. N.E. New Guinea.
Schl center's sketch and description suits this plant very well ; the
length of the spathe, however, is 1 cm., whereas Schlechter gives 0'5 cm.
Flower white, labellum veined, with purple tip.
DENDROBIUM ( APORUM) PSEUDOCALCEOLUM J. J. S. in Bull. Dep. Agric. Ind.
Neerl. n. v. (1907) 34, etc.
Manokoeari, Lauggen, epiphytic on prostrate tree, overhanging sea.
Jan. 6227.
Distrib. N.E. New Guinea.
Flowers mauve.
DENDROBIUM ( APORUM) INCONSPICUUM J. J. S., sp. nov.
Gaules elongati, ramosi, valcle compressi, laxe flexuosi, nitidi, laxe foliati, superne
valde attenuati et folia rudimentaria gerentes, parte adest 74 cm. longa, internodiis e
basi apicem versus paulo dilatatis, ad c. 4 cm. longis et 0'47 cm. ktis. Folia erecto-
patentia, lateraliter compressa, lanceolato-linearia, acuta, sicco margine superiore
c. 4'7 cm., margine inferiore 5'2 cm. longa, 0'4 cna. lata ; vagina? tubulosae, valde
compressse, ensiformes, internodia a^quantes. Inflorescentice e nodis caulis partis
superioris, fasciculatae, squamatte, pedunculis brevibus, 1-floris. Flares parvi. Sepa-
lum dorsale triangulum, obtusum, 3-nervium, c. 0'25 cm. longum, 0'225 cin. latum.
Sepala lateralia lacinia oblique triangula ad pedem gynostemii decurrentia, mentum
obtusum cum ovario angulum fere rectum faciens medio fere valde obtusangule
incurvum fere 0'25 cm. longum formantia, marginibus anticis libera, oblique triangula,
obtusiuscula, 4-nervia, c. 0'25 cm. longa, basi 0'36-0'37 cm. lata. Petala oblique
lanceolata, acuta, 1-nervia, c. 0'26 cm. longa, 0'07 cm. lata. Labellum cum pede
gynostemii angulum acutum faciens, supra basin et in | partibus supra basin
obtusangule recurvum, unguiculatum, valde dilatatuin, 3-lobum, basi 3- supra basin
5-nervium, fascia lata longitudinal! incrassato apice 3-loba, lobulo intermedio
laterales bene superante basin lobi intermedii vis attingente, explanation, c. 0'4 cm.
longum, ad lobos laterales 0'35 cm. latum, ungue oblongo-quadrangulo ; lobi laterales
sinibus parvis a lobo intermedia separati, late trianguli, fere quadranguli, obtusi,
irregulariter marginati ; lobus intermedius laterales superans, transversus, late
206
retuso-subbilobulus, cum lobulo parvo fere aequilongo triangulo obtuso in sinu,
c. 0'06 cm. longus, 0'15 cm. latus. Qynosteminm breve, c. O075 cm. longum,
filamento subulate, auriculis subquadrangulis, 2-lobulis. Anthera cucullata, ainbitu
antice 6-angulata, apice truncata, basi 2-lobula, c. - 07 cm. lata. Pollinia 4,
oblonga, lateraliter compressa, interiora quam exteriora tenuiora, exteriora extus
convexa, c. O'OG cm. longa. Stigma profunde excavatum, quadrangulum, margine
inferiore rotundatum. Pes gynostemii cum ovario angulum subrectum faciens,
medio fere obtusangule incurvus, parte superiore quam partem inferiorem bene latiore
concava cum incrassatione longitudinali verruculosa, truncatus inexplanatus
c. 0'25 cm. longus. Ovarium 6-sulcatum, c. 0'23 cm. longum, cum pedicello
c. O4 cm. longo, tenuiter clavatum.
Hob. Schouten Islands, Wiak, Bosnik, epiphytic on strand tree, common.
Jan. 6272.
The description of D. nycteridoglossum Rchb. f., which is said to be of
Papuan origin, more or less fits this plant. As, however, the description
is very poor and the exactness of the information about the origin is very
uncertain, further comparison with that species is for the present useless.
The material was very poor, and especially the description of the lip
wants further confirmation.
The flowers are said to be yellow.
ERIA ( CYLINDROLOBUS) RIGIDA Bl. Mus. Bot. Lugd.-Bat. ii. 183, etc.
Var. PAPUANA J. J. S. in Nova Guinea, xii. (1913) 76, t. xx. 59.
Schouten Islands, Wiak, Bosnik, epiphytic on strand trees. Jan. 6276.
Though I now think that this plant is not a form of Eria rigida Bl., I
quote it under the above name because I do not yet wholly understand
Schlechter's Papuan species of this group.
Distrib. New Guinea (D.N.W., Humboldt Bay, Gjellerup}.
Shoots, 50 m. long. Flowers with outer perianth-segments white, the
inner pink.
POMATOCALPA ORIENTALS J. J. S. in Nova Guinea, xii. (1913) 101, t. xxviii.
85, etc.
Boeroe, Tifoe harbour by road to signal point, forest. Fl. (orange),
Jan. 6296.
Distrib. New Guinea (D.N.W., Humboldt Bay, Gjellerup ; D.S.W.,
Merauke River, nat. coll.). Kei Islands, Amboina, Obi.
SARCANTHUS BICORNIS J. J. S. in Bull. Dep. Agric. Ind. Neerl. n. xix. (1908)
35, etc.
Schouten Islands, Wiak, Bosnik, epiphytic on strand trees. Fl. Jan.
6274.
Distrib. New Guinea (D.S.W., Digul River, Brandenliorst ; Noord River,
Verxteeg; N.E.).
Flowers with white petals and yellow labellum.
207
DICOTYLEDONEJE.
CASUARINACE.E.
CASUARINA EQUISETIFOLIA Forst. Char. Gen. PI. Aust. 103, fig. 52.
VVariap, near, on a stony and open inundation area of the Momi River.
Fr. Dec. 6126.
Jjistrib. N.E. & S.E. New Guinea and adjacent islands. Solomon and
Marianne Islands, Polynesia, N.E. Australia, Malaya, India to E. Africa.
PIPERACE.E. (C. DE CANDOLLE.)
PIPER BOSNICANUM C. DC., sp. nov.
liamulis glabris laevibus ; foliis sat longe petiolatis glabris, limbo ovato-rotundato
basi ima aequilatera acuto apice breviter et obtusiuscule acuminate, o-nervio, petiolo
basi ima vaginante ; pedunculo glabro quam petiolus breviore, spica inatura quani
limbus pluries breviore, rhachi hirsuta, bracteae glabra? pelta rotunda centre pedi-
cellata, ovario libero rotundato glabro, stigniatibus ovatis brevibus, bacca ovatae
glabrse stipite duo pluries longiore.
Hab. Schouten Islands, Wiak, Bosnik. Fl., Fr. 6277.
Dioicum, in arboribus scandens. Kamuli spiciferi 2 mm. crassi, collenchyma in
fascicules discretes dispositum et haud libriforme, fasciculi intramedullares 1-seriati,
canalis lysigenus ventralis perphericique plures. Lirnbi in sicco formi pellucido-
punctulati, usqua ad 10 cm. longi et 7 cm. lati. Petioli 1'5 cm., pedunculi 1 cm.
longi. Spica matura 2 cm. longa, bractese pelta 1 mm. diain. Stigmata 3 sessilia,
bacca in vivo aurantiaca in sicco nigra 7 mm. longa et 5 mm. crassa. Species primo
udspectu P. buruanum, Miq. referens.
PIPER FORBTENII C. DC. in Prodr. xvi. i. 348 ; Scheffer in Ann. Jard. Buit.
i. (1876) 50 ; Nova Guinea, viii. (1914) 1007.
Roon Island, ridge beyond Djende, 300', scandent in forest. Fl., ? . Jan.
6233.
Distrib. New Guinea (D.N.W., Dore, Teysmann ; D.S.W., near Alkmaar,
van Roemer). Amboina, Halmaheira, Philippines.
Liane, climbing up trunks of trees, with huge leaves and pendent yellow
spikes, 4 dm. long and 3 cm. thick. Also abundant in the high forest
round Manokoeari, but only seen there in sterile condition.
PIPER BIPUNCTATDM C. DC., sp. nov.
Kamulis glabris ; foliis breviter petiolatis, limbo elliptico-lanceolato basi aequi-
latera acuto apice acuta, et sat longe acuminato supra glabro subtus puberulo,
5-plinervio nervo centrali nervum adscendentem utrinque opposita vel alterne mittente
quorum supremus a 2-2'5 cm. supra basin solutus, nervo laterali adscendente utrinque
a basi soluto, petiolo puberulo basi ima vaginante ; pedunculo fere glabro petiolum
superante, spica subflorente quam limbus pluries hreviore, rhacbi hirsuta, bracteae
glabrse pelta rotunda centre pedicellata, starninibus 2, antheris ovatis 4-valvatis
quam filamenta oblonga pluries brevioribus.
Hab. Schouten Islands, Wiak, Bosnik (coral), in clearings. Fl. Jan.
5709.
208
Dioicum, epiphytum. Eainuli spiciferi 1 mm. crassi, collenchyma in fascicules
discretes a latere valde productos dispositum et baud libriforine, fasciculi intra-
medullares 1-seriati, canalis lysigenus unicus centralis. Limbi in sicco inembranacei
usque ad 12'5 cm. longi et 3'5 cm. lati, creberrime pellucido-punctulati punctatique
punctis rotundis vel oblongis. Petioli 6 mm., pedunculi 10 mm. longi. Spica sub-
florens 3 cm. longa et 2'5 mm. crassa, bracteae pelta 1 mm. diam.
HORACES. (H. N. RIDLEY.)
Ficus INFECTORIA Roxb. Fl. Ind. iii. 550 ; Sebum. & Laut. 273.
Wakde Island, on beach. Fl. Jan. 6244.
Distrib. N.E. New Guinea. India, Burma, Malaya, and Philippines.
" A large tree with white latex/'
Ficus MYRIOCARPA. Miq. in Ann. Mus. Lugd.-Bat. iii. 230-296 ; Schum. &
Laut. 281.
Manokoeari, in high forest, on " korang," 200-500' Fr. Jan. 6171.
Distrib. New Guinea (N.E., and adjacent islands). Amboina.
" Large forest tree, the leaves with stipules 4-5 cm. long and red recep-
tacles. The fruiting branches are cauline, long, and are also borne on the
roots."
*Ficus CELEBICA Bl. Bijdr. 461.
Manokoeari, tract to Ainbani, in high forest on " korang," 200'. Fr.
Jan. 6209.
Distrib. Celebes, Malay Peninsula, and Philippines.
" A forest tree with orange-red receptacles."
*Ficus BOTRYOCARPA Miq. Ann. Mus. Lugd.-Bat. iii. 233,
Manokoeari, in high forest on "korang." Fr. Jan. 6179.
Distrib. Celebes.
"A tree, with green receptacles borne on hollow cauline branches.
When cut water came out of the figs."
Ficus BRACHIATA King in Ann. Bot. Gard. Calc. i. 106, pi. 136.
Manokoeari, common in high forests on " korang," 200-500'. Fr. Jan.
6172.
Distrib. Sumatra.
"A tree with green receptacles, 3*7 cm. across, borne on long, cauline,
branching and pendent, fruiting shoots. On cutting figs open, water
came out."
FlCUS CONOCEPHALIFOLIA Ridl., Sp. nOV.
Arbor, rainis crassiusculis cortice cervino. Folia oblonga breviter obtuse cus-
pidata base rotundato cordato margine integro, superne glabra subcoriacea, subtus in
nervis 7 paribus hirtis, nervis intra margines anastomosantibus, subtus minute albo-
200
punctata scabra, 40 cm. longa, 18 cm. lata, petiolo 12 cm. longo. Stipules persis-
tentes lanceolatse acuminatae hirtse, 2 cm. longie vel ultra. Syncarpia rubra globosa
2 cm. crassa (in sicco) parce hirta, bracteis 2 lanceolatis acutis dissitis in dorso.
Bracteee basales nullae. Pedunculus '5-1 cm. longus bracteatus. Bractece ad
ostiolum lanceolatse acuminatse hirtse, plures erectae. Flores masculi ad ostiolum
monandri, pedicellati, sepala 4 lanceolata subacuta, anthera reniformi magna trail s-
versiin biloba. Feminei plerumque longe pedicellati, pistillo oblongo, stylo lateral!.
Hab. Humboldt Bay, ridge behind " kampong." Fr. Jan. 6260.
A specimen of what appears also to be this species is in Herb. Brit. Mus.
labelled " Java, Straits of Sunda J(oseph) B(anks)."
I cannot match this fig with anything yet described. It seems in some
points to be most nearly allied to F. Beccarii, but the fruits in this and its
allies are all borne on separate branches. The persistent and large stipules
are peculiar. The leaf, which is large, has just the texture and scabrid feel,
as well as the whitish dottings, of Conocephalus. The only figs seen contained
male flowers and gall flowers, some of which have, like the male flowers,
long stalks, while others are almost sessile.
URTICACE.E.
LAPOKTEA ARMATA Warb. in Engl. Bot. Jahrb. xiii. (1891) 293 ; Schum.
& Laut. 289.
Manokoeari, undergrowth, in high forest, on "korang," gregarious in
damp places, 500'. FL, <$ ? . 6186.
Distrib. N.E. New Guinea.
These plants, with herbaceous slender stems, 1-2 m. high, very stinging
leaves, grey in colour, and green flowers, formed a colony in a small soak
area on the top of the forest ridge which rises behind Manokoeari. As
Warburg's description only covers the ? plant, that of the c? is appended.
Inflorescence 2'5 cm. long, branched, with peduncle 7 cm. long. Flowers
arranged in small sessile glomerules, 2 mm. across, on the secondary branches
of the rhachis, composed of more numerous flowers than the $ , which
are mostly single in my specimens. Perianth 4-partite, 1'5 mm. across,
with lanceolate-acute segments, 1'5 mm. in length, covered with a few scat-
tered white hairs.
FLEURYA RUDERALIS (Forst.) Gaud. Voy. Uranie, 497 ; Schum. & Laut.
291 ; Scheffer in Ann. Jard. Buit. i. (1876) 48 ; F,Muell. Pap. PI.
41,59.
Schouten Islands, Wiak, Bosnik, under trees by sea-shore. Fr. Jan.
6279.
Distrib. New Guinea (D.N.W., Teysmann ; N.E. ; S.E.). Java, Marshall
and Solomon Islands, Ellice and Union Group ; Tahiti.
Herbaceous, with pink stems and white flowers.
210
PELLIONIA VANHASSELTII Gibbs, sp. nov.
Herba decumbens, glaberrima ; caulis simplex, plurisulcatus. Folia alterna
petiolata vel subsessilia, elliptico-lanceolata, basi uno latere subrotundata, altero
cuneata, apice in acumen longum acutissimum sensim attenuata, infra medium
integerrima, superne grosse insequaliter obtuse -den tata, penninervia, fere sequilatera
(basi excepta), viridia, carnosa, supra cystolithis linearibus baud densis, subtus palli-
diora, cystolithis minutis densissime obtecta. Flores feminei albidi, pedicellati,
laxiuscule glomerati. Perianthium 5-merum, persistens, segmentis apice leviter
retusis, longe aristatis, setis perianthio duplo longioribus. Staminodia 5, parva,
segmentis opposita, inflexa. Achcenium compressum.
Hob. Manokoeari, epiphytic or terrestrial in high forest, 500'. Fr. Jan.
6168.
This plant, up to 5-4 dm. in height, more or less decumbent in habit,
was found growing massed on the dead prostrate trunk of a tree, and also
beneath on the ground. Leaves dark green when dried, fleshy when
growing, +18 cm. long, including the acumen +3 cm. long, 4'5 mm. broad.
Flower fascicles l'5-2 cm. across. Pedicels +3 mm. long. Perianth-
segments 2 mm. long, with mucro 3 mm. long. IStaminodes +'5 mm. long,
when expanded +1 mm. long. Achene 1*5 mm. by 1 mm., light brown in
colour.
Near P. nigrescens Warb., but differs in the deeply dentate leaves, with
fewer veins, drying dark green in colour.
This species is named in honour of Mr. R. F. J. van Hasselt, Chief
Missionary at Manokoeari, who, from long residence and perfect knowledge
of Papuan dialects, enjoys the confidence of these people, and to whom I
was much indebted for valuable information and help, most kindly given.
LOR AN TRACED.
LORANTHUS VERSTEEGII Laut. in Nova Guinea, viii. (1910) 289.
Manokoeari, epiphyte on trees in clearings. Fl. Jan. 6175.
Distrib. New Guinea (D.S.W., Noord River, Versteeg ; Bian River,
Brandenhorst ; S.E. Sogeri Region, Forbes (499, Herb. Mus. Brit.)).
A very distinct plant with many long hanging shoots, 2-3 m. in length,
bearing single, and sometimes binate, densely-flowered racemes in the axils
of the leaves on the old wood, for about halfway down each shoot. The
flowers are red at the base and yellow-green from about the middle. Some
o the racemes are longer than in the type, and some shorter, and the flowers
are somewhat longer ; but these are no doubt variable features. In Forbes's
plant the leaves are narrow, as in Brandenhorst's 278 (Herb. Kew.), acute,
and decurrent at the base, with corollas 5 cm. long, pubescent on the outer
surface.
211
PORTULACACE2E.
PORTULACCA QUADRIFIDA L. Sp. PL ed. 1, 445 ; Hemsl. CLall. Exp. Bot.
122 j Schum. & Laut. 309.
Island of Wakde, in " campong," creeping under trees. Fl. (yellow).
Jan. 6247.
Distrib. N.E. New Guinea and adjacent islands. Marshall and Solomon
Islands, Fiji and Vavau. Tropical Asia and Africa.
ROSACES.
RUBUS MOLUCCANUS L. Sp. PI. ed. 1, 1197 ; Scheffer in Ann. Jard. Buit. i.
(1876) 23; Schum. & Laut. 339; Nova Guinea, viii. (1910) 367,
(1912) 647.
Manokoeari, edge of forest. Fl., Fr. Jan. 6212.
Distrib. New Guinea (D.N.W., Dore, Teysmann ; D.S.W., Noord River,
Versteeg ; Hellwig Mts., von, Roemer ; N.E. and adjacent islands). India,
Indo-China ; Malay Peninsula, Sumatra, Amboina, Borneo, Philippines.
Berries red, sweet.
LEQUMINOS^.
*ALBIZZIA MOLUCCANA Miq. Fl. Ind. Bat. i. 26.
Manokoeari, edge of forest, on " korang," 100'. Fr. Jan. 5738.
Distrib. Moluccas.
A fine tree with white bole and spreading crown, pinnatisect leaves with
very small pinnules, and pendent white flowers, quite abundant on the edge
of the high forest behind Manokoeari. Only the old pods scattered on the
ground were collected, which, however, are very distinct, +1*4 dm. long
and 2 cm. broad, with a small acumen and a membranous wing, broader on
the placental suture. The small seeds are very close together, under 1 cm.
apart. These characters, with the fine dissected leaves, are well shown in
Koorder's 15643 (" cultivated at Buitenzorg ") as may be seen in Herb.
Kew.
ENTADA SCANDENS Benth. in Hook. Journ. iii. 332 ; Schum. & Laut. 346.
Manokoeari, track to Ambani, creeper in strand forest. Fl. (yellow).
Jan. 6191.
Distrib. N.E. New Guinea and adjacent islands. Tropics of both hemi-
spheres.
TAMARINDUS INDICA L. Sp. PI. 34.
Manokoeari, Wousi, in " kebun." Fl. Jan. 6217.
Distrib. Tropics of both hemispheres.
212
ALYSICAKPUS NTJMMULARIFOLIUS DO. Prod. ii. 353; Schum. & Lnuf, 277.
Island of Sorong, creeping under trees by sea-shore. Fl. (red-brown).
Jan. 6285.
Distrib. N.E. New Guinea. India, Indo-China; Malay Peninsula and
Andamans, Borneo, Philippines, N.E. and N.W. Australia, Fiji. Tropical
Africa and West Indies.
DERRIS ULIGINOSA Benth. in PI. Jungh. i. 252 ; Schum. & Laut. 360 ;
Valeton, Plant, pap. in Bull. Dep. Agric. Ind. Neerl. no. x. (1902) 19 ;
Nova Guinea, viii. (1910) 379.
Manokoeari, Langgeri, in mangrove association. Fl. Jan. 6229.
Distrib. New Guinea (D.N.W., Humboldt Bay; D.S.W., Etna Bay,
Wichmann ; Gelieb, Brandenlwrst ; N.E. and adjacent islands). South
Asia to N. Australia and Polynesia.
MUCUNA KEAETKEI Warb. in Engl. Bot. Jahrb. xiii. (1891) 329 ; Schum.
& Laut. 365 ; Nova Guinea, viii. (1910) 381, (1912) 652.
Manokoeari, Wousi, over trees by sea-shore. Fl. Jan. 6231.
Distrib. New Guinea (D.S.W., Alkmaar, on Noord River, Versteeg ;
N.E.).
A coast liane, with magnificent red flowers in huge interrupted in-
florescences.
In the absence of leaves and fruit, determination must be uncertain ; but
in the shape and size of calyx, approximate measurements, and form of floral
parts, with the number of ovules (5), my plant agrees with M. Kraetkei Warb.
Warburg does not describe the inflorescence, beyond stating that it occurs
often on the old wood ; in that case bunches of shortish racemes, 5-17 dm.
long, spring irregularly from the old wood, which is lenticellose. When
the inflorescence is on young wood the recemes are long and single, as in
Versteeg's 403 Herb. Kew.
LINAGES.
*DURANDEA PARVIFLORA Stapf in Hook. Icon. tab. 2822.
Manokoeari, Langgen, in open clearing. Fl. Jan. 6218.
Distrib. Solomon Islands.
A small tree with yellow flowers. The measurements of the flowers agree
with the above species ; also the size of the leaves and the inflorescence.
MELIACE^E. (C. DE CANDOLLE.)
AGLAIA GIBBSI^ 0. DC., sp. nov.
Kamulis adpresse et rufescente stellato-lepidotis ; foliis sat longe petiolatis 3-
jugis, foliolo terminal! sat longe petiolulato oblongo-obovato basi sequilatera acuto
apice brevissirne et obtuse acuminate, lateralibus oppositis modice petiolulatis
oblongo-ellipticis basi aequilatera acutis apice breviter et obtuse acuminatis, omnibus
213
supra glabris subtus ad nervos baud dense stellato-lepidotis ; panicula tenninali
folium fere sequante paullo supra basin trifida, ramis pyramidato-ramulosis, ramulis
adpresse et rufescente stellato-lepidotis spicatim cymub'geris, cymulis bifloris,
floribus pedicellatis ; calice 5-sepalo, sepalis rotundato-ovatis subtus stellato-lepidotis
supra glabris, ciliatis; petalis 5 obovatis glabris eciliatis, tubo stamineo glabro
urceolato integro et margine antherifero, antheris 5 glabris subsessilibus ovatis,
horizon taliter inflexis, connective ultra thecas obtuse producto ; ovario parce stellato-
lepidoto, stigmate globoso.
Bab. Manokoeari, in forest on " korang," 300'. Fl. Jan. 6204.
Arbor. Folia alterna 36 cm. longa. Foliola in vivo obscure virescentia, in
sicco firmule membranacea et creberrime pellucido-punctulata, terminale 16 cm.
longum et usque ad 6 cm. latum, lateralia supera 15 cm. longa et 5*5 cm. lata,
subsequentia gradatiin paullo minora, nervi secundarii subrecti utrinque 10-12.
Panicula} rami usque ad 13 cm. longi. Pedicelli 1 mm. longi. Flores in vivo
albis. Sepala 0*5 mm. longa. Petala membranacea, 1'5 mm. longa et usque ad
1 mm. lata. Anthene 0'75 mm. longse.
Species A. Roemeri C. DC. proxima, foliolis paucioribus et a nervo centrali
sequilateris, sepalis subtus stellato-lepidotis, petalis obovatis ab ilia discrepans.
MALPIGHIACE^E.
TRISTELLATEIA AUSTRALASICA A. Rich. Sert. Astrol. 38, t. 15; Scheffer
in Ann. Jard. Buit. i. (1875) 10 ; F. Muell. Pap. PI. 36 ; Schum. &
Laut. 387.
Manokoeari, Langgen, climber in mangrove association. Fl. (yellow),
Fr. Jan. 6223.
Distrib. New Guinea (D.N.W., Andai, Teysmann ; N.E. & S.E., and
adjacent islands). Singapore, Slum, Formosa, Malaya, Philippines ; to
E. Australia and New Caledonia.
RYSSOPTERYS TIMORENSIS Bl. ex A. Juss. in Deless. Ic. Sel. iii. 21, t. 350 ;
Scheffer in Ann. Jard. Buit. i. (1875) 10 ; F. Muell. Pap. Pi. 36 ;
Schum. & Laut. 387.
Manokoeari, tract to Ambani, 100', clearings, scrambling in rampant
upgrowth. Fl. (yellow). Jan. 6200.
Distrib. New Guinea (D.N.W., Sorong, Teysmann ; N.E. & S.E., to
adjacent islands). Malaya to Philippines and tropical Australia.
EFPHORBIACEJi. (J. HUTCHINSON.)
MALLOTDS TILI^EFOLIUS (Bl.) Miill.-Arg. in Linnsea, xxxiv. 190 ; Miq. in
Ann. Mus. Lugd.-Bat. iv. 123 ; Schum. & Laut. 396.
Manokoeari, track to Ambani, tree in clearings and forest. FL, < . Dec.
6202.
Distrib. New Guinea (D.S.W., Zippelius ; N.E. and adjacent islands).
Malay Archipelago ; Fiji.
214
MACARANGA RIPARIA Engl. in Bot, Jahrb. vii. (1886) 463 ; Schum. & Lanr.
397 ; Nova Guinea, viii. (1910) 235, (1912) 789 ; Kidl. iu Trans.
Linn. Soc. ser. 2, Bot. ix. (1916) 147.
Manokoeari, ridge behind " campong," common in high forest on
" korang," 300'. Fl., ? . Jan. 6177.
Distrib. New Guinea (D.N.W., Dore, Teysmann ; Humboldt Bay,
Gjellerup ; D.S.W., Noord River, Versteeg, von Roemer ; Utakwa River,
Kloss ; N.E. and adjacent islands). Kei Islands.
" A fine tree = 80 m. in height. Quite common."
EXCCECARIA AGALLOCHA L. Sp. PI. ed. 1, 1288; Schum. & Laut. 406 ; Nova
Guinea, viii. (1910) 241.
Manokoeari, Langgeu, in mangrove association. Fl., Fr., ? . 6216.
Distrib. New Guinea (D.N.W., Humboldt Bay, Wichmann ; D.S.W.,
Merauke, Brandenliorst ; N.E. and adjacent islands). Brit. India, Malaya,
Papuasia, Philippines, S. China, E. Australia, and Polynesia.
C EL ASTHACE.E.
EUONYMUS JAVANICUS Bl. Bijdr. 1146.
Manokoeari, 200', high forest on "korang." Fl. Jan. 6182.
Distrib. Indo-China, Timor, Philippines, Kei and Aru Islands, N.E.
Australia.
Tree, with greenish-white flowers.
SAPINDACE.E.
ALLOPHYLLUS COBBE Bl. Rumphia, iii. 131.
Manokoeari, 200', creeper in clearings and on edge of forest. Fl. (white),
Fr. Jan. 6203.
Distrib. India, Ceylon, Malay Peninsula, Andaman Islands, Java, Borneo,
Little Kei Islands, Philippines ; Samoa and Vavau.
GREWIA ACUMINATA Juss. in Ann. Mus. Paris, iv. (1801) 91, t. 48 ; r.f.
Hochreutiner's Cat. Bog. Nov. i. 47, in Bull. Inst. Bot. Buit. xix.
(1904) 46.
Dammar Island, on sea-shore. FL, Fr. Jan. 6291.
Distrib. Philippines, Borneo, Java, Malay Peninsula.
Shrub, with white flowers. This plant was found growing in a clump on
the edge of the beach, the forest having been cleared behind it. It agrees
well with the large series at Kew of the species included by Hochreutiner
with G. acuminata, though the leaves are larger than in the specimens seen.
215
FLACOURTIACE.E.
ERYTHROSPERMUM CANDIDA Becc. in litt.= Gestroa Candida Becc. Malesia, i.
184.
Manokoeari, Langgen, land edge of mangrove association. Fl. Jan.
6224.
Distrib. New Guinea (D.N.W., Ramoi, Beccari).
Tree with white flowers.
I am indebted to the kindness of Dr. Beccari for a specimen of his
Gestroa Candida, which proved identical with my plant. It also shows with
the above some flowers with undeveloped ovaries among the more numerous
$ ones. Dr. Beccari wrote that he now considered the plant to be an
Jirytlirospermum, and on investigation I am myself inclined to the same
view.
BEGONIACE.E.
BEGONIA (PETERMANNIA) HUMBOLDTIANA Gibbs, sp. nov.
Herba glaberrima ; caule erecto, ramosissimo. Folia petiolata, obovato-lanceolata,
basi valde inaequalia (uno latere cordato, altero cuneato), apice tenuiter anguste-
acurainata, grosse et regulariter inciso-serrata, chartacea, supra flavo-viridula, albido
punctulata, subtus dilute griseo-albida. Inflorescentia terminalis, quam folia
brevior. Flos tf sepalis 2, roseis, orbiculari-cordatis ; filamentis basi connatis.
Flores $ petalis 5, insequalibus ; stylis 3, connatis, apice spiraliter bifurcatis.
Fmctus 3-alatus, apice truncatus vel arcuato-obtusus.
Hob. Humboldt Bay, in high forest by stream, 300-500'. Fl. $ ? , Fr.
Jan. 6253.
A handsome plant, '75 m. high, growing abundantly in large clumps
along the banks of the stream. Leaves 12 cm. by 4'5 cm., sage-green in
colour, with 2-3 white spots in more or ess parallel lines between the lateral
veins, which are conspicuous, 4-5 on each side of the midrib. On the margin
the largest incisions vary from 1 cm. in depth and 1*5 cm. in breadth,
graduating insensibly to the extreme apex. Petiole 1 cm. long. Stipules
1'5 cm. long, often unequal in size, caducous, transparently membranous in
texture, and produced into long fine hair-like points. flower 6 mm. by
7 mm. (immature). ? flowers arising singly, opposite the upper leaves, on
pedicels 1'5-1*8 cm. long, the flowers being 2 cm. long and 3 cm. across ;
with unequal petals, of which the largest are 1'2 cm. by 6' 7 mm. ; the three
styles are bifid, spreading, 5 mm. long. Fruit I'l cm. long, 1'5 cm. across,
with equal membranous wings, 4 mm. broad in the centre and 5 mm. broad
at the apex of the fruit. The fruits were affected by insects and swollen out
of shape in nearly all the plants seen.
This species is distinct in the very regular and finely graduated serrate
incisions of the leaves, with their parallel rows of white markings.
216
LYTHRACE.E.
PEMPHIS ACIDULA Forst. Char. Gen. PI. 68, t. 34 ; F. Muell. Pap. PL 43-59 ;
Schum. & Laut. 460.
Wakde Island, growing thickly on coral-reef on coast, where washed by
sea. Fl. (white) , Fr. Jan. 6246.
Distrib. New Guinea (N.E. & S.E., and adjacent islands). Strand plant
distributed from Polynesia to E. Africa.
Shrub, prostrate, to 3 m, in height, where beyond the spray.
MELASTOMACE^E. (E. G. BAKER.)
OTANTHERA NOVO-GUINEENSIS Bak. f. in Trans. Linn. Soc. ser. 2, Bot. ix.
(1916) 50.
Island of Roon, near the road round the Bay, growing in the open. Fl.,
Fr. Jan. 6238.
Distrib. New Guinea (D.S.W., Utakwa, near to Mt. Carstensz, Kloss*).
" Small shrub, with white flowers and red fruit."
MEDINILLA RHODORHACHIS Bak. f., sp. nov.
Planta epiphytica, ramis 4-gonis vel levissime 4-alatis ad nodos barbatis ad
.3/". speciosam Blume accedens. Folia sessilia, opposita, oblonga vel elliptica, apice
breviter acuminata, basi cordata, 5-plinervia, costa subtus conspicua. Flores 4-meri
in paniculam dispositi. Paniculae folio breviores. Pedicelli stricti. Bractece
parvse. Galycis tubus campanulatus limbo subtruncato. Petala alba staminibus
paullo longiora. Staminum connections postice breviter calcaratus, antice bical-
caratus. Fructus ignotus.
Hob. Schouten Islands, Wiak, Bosnik, epiphytic on strand tree.
Flowers white ; rhachis pink. Fl. Dec. 6275.
Leaves (ex sice.) 2-3 dm. long and 7-5-13 cm. broad. Calyx +3'5 mm.
long. Petah 6 mm. long. Anthers 3-3'5 mm. long.
The distinguishing features of this epiphyte are the oblong or elliptical,
acuminate, 5-plinerved, sessile leaves, the panicle of small, white, 4-merous
flowers, the calyx being +3'5 mm. long. The anthers are shortly spurred
anteriorly and posteriorly.
ARALIACE^E.
POLYSCIAS SORONGENSIS Gibbs, sp. nov.
Frutex parvus, glaberrimus. Folia pedicellata, stipulata, imparipinnata, 5-G-
juga, rhaqhis cylindrica, longitudinaliter striata ; foliola opposita, graciliter pedicel-
lata, obovato-lanceolata vel oblonga, acuta, basi obliqua rotundata, apice sensim
acuminata, subintegerrima tenuiter membranacea. Panicula pedunculata nunc
bipedata, racemuli umbellati pedunculi secundarii verticelkti vel oppositi. Flores
parvi, breviter pedicellati, pedicelli sub flores articulati, apice in calyculum minimum
expansi. Calyx repando-5-dentatus. Petala 5, apice leviter cohserentia. Stamina 5,
217
filamentis brevibus. Discus planus. Styli 2 distinct!, elongati. Ovarium bilocukre.
Fructus ignotus.
Hal). Island of Sorong, under strand trees. Fl. Feb. 6287.
A small shrub +2 m. high. The petioles of the leaves are 1'2 dm. in
lengthj with adnate stipules 1*7 cm. long. The dried leaves are light green
in colour, the largest being 4'8 dm. by 3'1 dm. ; the largest foliole is 1*7 dm.
by 5 cm. ; the margins can hardly be described as serrate, but show subulate
teeth, +1 mm. long and 1*2 cm. apart; petiolules +1'3 cm. long. The
inflorescence is umbellate-racemose, +5'1 dm. by 4 dm., including peduncle
2'5 dm. long, which is subtended by 2 bracts; the secondary peduncles arise
in the axils of 2 bracts, and bear pairs of bracts up the axis, which bear single
flowers or small lateral umbels, or may dichotomise, bearing 2-4 umbels;
the umbels show an involucre of minute bracts and are +1 cm. across. The
flowers are green, on pedicels 2 mm. long ; petals broad at the base and
thickened at the apex, 2'5 mm. long ; anthers oblong, almost sessile, 2 mm.
long ; styles 1/2 mm. long ; ovary 1/5 mm. long.
MYRSINACE.E.
M.ESA RACEMOSA (K. Sch.) ; Mez in Pflanzenreich, iv. 236, Myrsin. (1902)
43 ; Schum. & Laut. 492.
Island of Wakde, on beach. FL, Fr. Jan. 6243.
Distrib. N.E. New Guinea.
Shrub or epiphyte, with white flowers and berries.
FLORIDUM Roem. et Schult. Syst. iv. (1819) 512 ; A. Rich. Toy.
Astrol. ii. 57, t. 21 ; Scheffer in Ann. Jard. Buit. i. (1876) 33.
Manokoeari, Langgen, mangrove association, on sea-edge. Fl., Fr.
Jan. 6228.
Distrib. New Guinea (D.N.W., Jobi Island, Barclay ; Gede Island, Dore
Bay, Teyftnann). Kei Islands, Amboina, Sumbawa, Philippines.
Small compact shrub with white flowers.
ASCLEPIADACEJ:.
SARCOLOBUS RETUSUS K. Sch. Fl. Kais. Willielmsl. 109 ; Schum. & Laut.
509 ; Valeton in Bull. Dep. Agric. Indes Neerl. no. x. (1907) 49.
Manokoeari, Langgen, twiner in mangrove association. Fl. Jan.
6220.
Distrib. New Guinea (D.N.W., Humboldt Bay, Wichmann ; N.E.).
Ternate, Timor.
Creeper on trees and on ground. Yellow-green flowers with ciliate
petals. Schumann notes that this plant is very poisonous.
218
C N V L V U L A C E J5. (A. B. RENDLE.)
IPOMCEA DENTICULATA Choisy in Mem. Soc. Phys. Geneve, vi. (1833) 447.
Island of Sorong, creeping under trees by beach, covering ground and
plants. Fl. (purple), Fr. Jan. 5740.
Distrib. N.E. New Guinea and adjacent islands. Seychelles to Polynesia
and Australia.
B O R AGIN AC E ;E.
TOURNEFORTIA ARGENTEA L. . Suppl. 133 ; F. Muell. Pap. PI. 59 ;
Sebum. & Laut. 519.
Wakde Island, a small tree along the sea-shore. FL, Fr. Jan. 6242.
Distrib. New Guinea (N.B. ; S.E. and adjacent islands). Tropical
Australia, Polynesia, Malaya and India to Mascarene Islands.
Small tree 10 m. high, leaves silvery from white pubescence, with white
flowers and green fruit.
VERBENACE.E.
CALLICARPA ERIOCLONA Schauer in DC. Prod. xi. 643 ; Miq. Fl. Ind. Bat.
ii. (1856) 889 ; Scheffer in Ann. Jard. Buit. i. (1876) 41.
Manokoeari, track to Ambani, common on the edge of forest and in
clearings, 200'. FL, Fr. Jan. 6205.
Distrib. New Guinea (D.N.W., Lesson ; Mansinam Island, Dore Bay,
Teyxmann). Philippines.
This plant is distinguished from C. cana L. by the large, more lanceolate,
irregularly serrate leaves, with very white pubescence underneath, and white
flowers with longer exserted stamens. C. repanda K. Sch. & Warb. is
possibly a synonym of this plant.
PREMNA NITIDA K. Sch. Fl. Kais. Wilhelmsl. 120 ; Schnm. & Laut. 523.
Manokoeari, common in clearings and round foot-hills. FL, Fr. 6163.
Distrib. N.E, New Guinea.
A tree with white flowers and black fruit. Also seen along the beach to
Wariap.
CLERODENDRON LINDAWIANUM Laut. in Schum. & Laut. Nachtr. Fl. d.
Schutzgeb. Siidsee, 372.
Var. GLABRIOR Gibbs. Tota planta manifesto glabrior.
Humboldt Bay, in high forest on range above river, 400', FL, Fr.
Jan. 6264.
Distrib. (of type). New Guinea (D.S.W., Mt. Carstensz, Kloss ; N.E. ;
S.E. [Forbes, Sogeri region, Brit. Mus.]).
A small tree with conspicuous white flowers and black fruit.
I cannot separate this plant from Lauterbach's species, which is evidently
very widely distributed. In my specimen the leaves are glabrous, with the
219
exception of the veins, which are pubescent on both surfaces ; the tomentum
on peduncles, pedicels and calyx is also much reduced.
VITEX TRIFOLIA L. f. Suppl. 293 j F. Muell. Pap. PI. 86; Schum. &
Laut. 524 ; Valeton in Bull. Dep. Agric. Indes Neerl. no. x. (11)07)
51.
Dammar Island, on seashore. Fl., Fr. 6290.
Distrib. New Guinea (D.S.W., Merauke, Koch ; N.E. ; S.E.nnd adjacent
islands). South Asia through Malaya and Japan, and through New Guinea
to New Caledonia.
A C A N T H A C E JE. (S. MOORE.)
HEMIGRAPHIS REPTANS T. And. ex Hemsl. Bot. Voy. ' Challenger,' i. iii. 173 ;
Schum. & Laut. 543 ; Valeton in Bull. Dep. Agric. Indes Ne'er!, no. x.
(1907) 58.
Manokoeari, common undergrowth in forest, up to 500'. Fl., Fr. Jan.
6160.
Distrib. New Guinea (D.N.W., Tobadi, Wichmann; N.E. and adjacent
islands). Aru Islands, Amboina, Philippines; New Hebrides and New
Caledonia.
Small creeping plant with white flowers and glossy fleshy leaves. Also
seen on the lower foot-hills of the Arfak range near Wariap.
HEMIGRAPHIS DORENSIS S. Moore, sp. nov.
Erecta, circa 25 cm. alt. Caule sparsim ramoso gkbrescente. Eamis pauci-
foliatis puberulis. Foliis petiolatis oblongo- vel ovato-lanceolatis, obtusis, basi
obliquis obtusisque vel rotundatis, margine dentatis vel dentato-crenatis, nonnun-
quam fere integris supra pilis brevibus strigillosis appressis inspersis subtus in nervis
puberulis. Spicis abbreviatis paucifloris, subsessilibus pedunculatisve. Bracteis
oblanceolatis obtusis ciliatis. Calycis bracteas suba?quantis segmentis juxta basin
solurnmodo connatis linearibus acutis ciliatis. Corolla calyce duplo longiore extus
glabra hujus tubo superne ampliticato lobis posticis quam antici plane latioribus.
Filament is inclusis glabris. Ovario oblongo, apice pilosulo. Stylo inferne pilosulo.
Ovulis pro loculo 6.
Hob. Track to Ambani in " korang " forest, 300'. Fl. 6189.
Leaves 2'5-8 cm. long, l'5-3 cm. broad, membranaceou?, on lower side
pale. Petioles 5-22 mm. long, slightly pilose. Spikes up to 2 cm. long,
without counting the filiform peduncle of equal length. Bracts +8 mm.
long. Calyx 9 mm. long. Corolla white, 14 mm. long ; tube in the lower
part l'25-2 mm., at the throat 5 mm. wide ; front lobes obovate, 5 mm. broad ;
hind lobes rotundate, 6'5 mm. broad. Filaments longer 3'5 mm. long,
shorter barely 1 mm. ; anthers 1'5 mm. long. Starninodium 0. Ovary
3 mm. long. Style 10 mm., stigma front lobe 2'5 mm. long.
This is nearest to H. caudigera S. Moore, which has larger caudate leave?,
longer bracts, &c.
220
*ASYSTASIA INTRUSA Bl. Bijdr. 79G.
Sorong Island, under trees by beach. Fl. Jan. 6286.
Distrib. Aru Islands, Little Kei Islands, Timor Laut, Java, and Malay
Peninsula.
'25-'50 m. high, with reddish-brown pink flowers.
*PERISTROPHE JALAPP.EFOLIA Nees in DC. Prod. xi. 494.
Manokoeari, near Genbela, common undergrowth in secondary strand
forest. Fl., Fr. Jan. 6216. Manokoeari, in high forest, common under-
growth on " korang," 200-500'. 6161.
Distrib. Java.
Herbaceous plant, "50 m. high, with white flowers. Also seen on the
foot-hills of the Arfak range.
RUBIACE.E. (Tn. VALETON.)
BIKKIA GRANDIFLORA Reinw., var. TENUIFLORA Val., var. nov.
Folia petiolata, elliptica, obtusa, basi acuta vel cuneata, tenuiter coriacea, nervis
lateralibus utrinque circa 5 tenuibus baud prominentibus. Pedunculi medio cupu-
latim bracteolati. Calycis lobi ensiformes acuti, calycem cylindricum sequantes.
Corollae tubus elongatus strictus, tenuis, versus limbum sensim ampliatus. CoroHce
lobi trigoni, basi dilatati, longiores quam lati. Capsula cylindrica, lineis elevatis
8 pertensa, calycis lobis accrescentibus coronata.
Leaves 135 mm. long, 60 mm. broad. Petiole 15-20 mm. long. Calyx-
lobes 15 mm. long, 3 mm. broad. Corolla-tube 17 mm. long, base 3 mm.,
apex 7 mm. broad. Corolla 105 mm. long ; lobes 20 mm. long, base 14 mm.
broad.
Island of Wakde, shrub, 8 m. high, growing in masses behind the coral
rocks of seashore. Fl. (white, sweet-scented). Jan. 6245.
Distrib. (of the species). Moluccas (type-specimen of Reinw. in Herb.
Lugd.-Bat.), Kei Islands, Aru Islands, S.W. New Guinea, Palau Islands,
Tahiti.
The species is characterized by the large, sword-like, curved calyx-lobes
and the triangular acute corolla-lobes, which are longer than broad. The
variety differs from the type in the very narrow corolla-tube.
OPHIORRHIZA INSULARIS Val., sp. nov.
Herha sublignosa, tomentosa puberula, in sicco fusca, caules incano-puberuli,
glabrescentes. Stipules abortivse vel califormis et puberse seca tricula nulla. Folia
petiolata, vulgo aequalia, summa interdum valde disparia, elliptica vel obovata vel
ovato-elliptica, apice obtusa vel obtusissima vel rotundata, basi cuneato-acuminata
ad longe in petiolum producta, membranacea, in sicco visca, supra dense punctulato-
scabrida, subtus parce tomentosa, nervi laterales utrinque 9-12, vene laxe obscure
reticulatse. Inflorescentia pedunculata folius multo brevior, incano-puberula semel
vel bis dichotoma vel subtrichotoma, ramis elongatis multifloris secundifloris.
221
Florrs brevi pedicellate Calyx 5-lobus, lobis dentifonnibus parvis. Corolla apice
5-cristata, crista brevi rotundata ; tubus brevis latus fauce dense hirsute lobi tubo
sequilongi. Ovarium dense tomentosmn. Stylus glaber. Capsula ad ramos secundae,
breviter pedicellate, scaberulse.
Leaves 75-100 mm. long or shorter, 40-45 mm. broad. Petiole 15 mm.
long. Peduncle 10-20 mm. long. Fruiting-branches 5'5 cm. long.
Hob. Schouten Islands, Bosnik, Wiak, under strand trees on " korang."
Small plant 2 dm. high. Fl. (white), Fr. Jan. 6284.
Collected also by Gjellerup in .1911.
GEOPBILA RENIFORMIS Don, Prodr. Fl. Nepal. 136 ; Schum. & Laut. 584 ;
Nova Guinea, viii. (1911) 485 ; Ridl. in Trans. Linn. Soc. ser. 2, Bot.
ix. (1916) 77.
Manokoeari, common creeping undergrowth in forest. Fl., Fr. 6181.
Distrib. New Guinea (D.S.W., Etna Bay, Koch-, Utakwa River, Kloss ;
N.E. & S.E. and adjacent islands). Cosmopolitan in Tropics.
AMARACARPUS WICHMANNI Val., sp. nov. (A. cuneifolius Val., var., in Nova
Guinea, viii. (1912) 769 et (1911) 502, p.p.]
Fruticulus ramis bilateralibus, horizontaliter expansis, ramulis haud regulariter
pinnatis, potius quasi dichotomis. Rami abbreviati, secus ramulos oppositi et alterni ;
oranes grosse rufo-villosi. Stipulce minutae, ovatae, longe 2-aristatae, rufo-hirsutae.
Folia subsessilia, obovato-cuneata, acuta vel obtusa, basi crasse membranacea,
siccando supra glaucescentia, subtus fusca, glabra, costa media prope basin cum
petiolo pulverulento-villosula, nervi laterales utrinque 4 cum costa prominuli, ante
marginem arcuatim conjuncti. Flares in apice ramorum abbreviatorum sessiles,
nunc 3 glomerati, bracteis et stipulis setaceis suffulti. Calyx cum ovario glabro,
campaniformis, lobis 4 ovatis, acutis, recurvis, ciliatis. Corolla breviter tubulosa,
limbo subpatulo, tubo circa aequilongo, lobis oblongis crassis, tubus circa antheras
fauce sessiles barbatus. Stylus corollae sequilongus. Drupa calyce parvo coronata,
pyrenas semiobovoideae, ventre piano, dorso profunde 3-4 sulcae tricostatae. Ramuli
nunc 50-300 mm. longi, brachyblastis 10-25 mm. longis dense obsessi. Folia secus
ramos subpersistentia, in apice ramorum abbreviatorum saepe 4-verticillata, 8-20 mm.
longa, 6-9 mm. lata. Flores 7 mm. longi; corolla 4'5 mm. longa. Drupaa 3-5 mm.
longse, pyrenae 2 '5 mm. latse.
Hab. Manokoeari, common undershrub in forest. FL, Fr. Jan. 6185.
Distrib. New Guinea (D.N.W., Wichmann).
" Shrub, 1-1-50 m. Branches dorsiventral, symmetrical ; white flowers.
Also abundant at Waren up to the foot-hills of the Arfak."
MYRMECODIA PDLVINATA Becc. Malesia, ii. 103.
Manokoeari, epiphytic in high forest, 300'. Fl. (white)., Fr. (red).
Jan. 6197.
Distrib. New Guinea (D.N.W., Andai, Beccart).
222
C U C U R B IT A C E M.
MACROCARPA (Bl.) Cogn. in Bull. Herb. Bois. i. 612 ; chum. &
Laut. 589.
Manokoeari, liane, everywhere in forest. Fr. Jan. 5746.
Distrib. New Guinea (N.E. and S.E.). Java, Borneo.
The most conspicuous plant seen along the coast in the lower forest. It
obliterates whole trees with dense walls of verdure, while the huge fallen
fruits, rotting on the ground, are the most striking objects in the forest.
The seeds, with transparent wings about 10 cm. across, often fill the air,
lazily borne on the breeze, like great butterflies, for which, indeed, I took
them at first in the distance.
GOODENIACE-E.
SC.EVOLA NOVO-GUINEENSIS K. Schum. in Bot. Jahrb. ix. (1887) 222;
Schum. & Laut. 594 ; Nova Guinea, viii. (1912) 693.
Manokoeari, common scrambler on edge of forest, clearings, and young
jungle. Fl. (yellow), Fr. Jan. 6199.
Distrib. New Guinea (D.N.W., Humboldt Bay, Gjellerup ; N.E.).
Apparently flowers all the year round.
COMPOSITE. (S. MOORE.)
WEDELIA BIFLORA DC. in Wight, Contrib. 18 ; Schum. & Laut. 600 ; Nova
Guinea, viii. (1910) 337.
Manokoeari, Genbela, common everywhere in open clearings and on
beaeh. Fl., Fr. Jan. 6201.
Distrib. New Guinea (D.S.W., Noord River, Biwak Island, Versteeg,
von Roemer ; N.E. and adjacent islands). Common strand plant in the
Tropics.
223
EXPLANATION OF THE PLATES.
PLATE 1.
Fig. 1. Araucaria Beccarii with Pteridium aquifolium in foreground ; vicinity of lake.
Fig. 2. Rafts made of three palm-trunks tied together, used by the Alfueros on both the
Angi lakes.
PLATE 2.
Fig. 3. Vegetation of marsh by $ lake, looking S.W. ; Koebre* Mt. in the background.
Baeckea fnttescens .
Fig. 4. Vegetation of forest by $ lake, looking N.W. ; Koebre" in background. Myrmedoma
arfakiana on extreme branches of tree *- , with Phyllocladus hypophyllus and
Libocedrus arfakensis * .
PLATE 3.
Fig. 5. Araucaria Beccarii * , with Podocarpus papuanus and Polysrias sp. in centre ;
Pteridium aquilinum in foreground ; vegetation of forest by $ lake.
Fig. 6. Vegetation of Koebre' ridge, showing Myrmedoma arfakiana, T50 m. high, flower-
ing , with tuber cut in longitudinal section * .
[Photographs are reproduced by kind permission of Mr. A. E. Pratt.'}
PLATE 4.
Fig. 7. Thysanosoria dimorphophylla Gepp. Apical portion of plant, \ nat. size.
INDEX OF NATURAL OEDERS, NEW GENERA
AND NEW SPECIES.
When more than one reference is given, the page on which the description
occurs is indicated by thickened numerals.
Acantbacese, 219.
Acronychia arfakensis Gibbs, 38, 144.
papuana Gibbs 29, 33, 144.
Agaricacese, 64.
Aglaia Gibbsise C. DC., 50, 212.
Alsophila angiensis Gepp, 32, 69.
arfakensis Gepp, 28, 70.
straminea Gepp, 56, 192.
Amaracarpus Wichmanni Val., 17, 51 }
221.
Anomopanax arfakensis Gibbs, 30, 163.
Anthoceroa bullato-spongiosus Gepp, 54 ;
191.
Apocynacese, 177.
Aracese, 201.
Araliaceze, 159, 216.
Asclepiadaceae, 217.
Auriculariacese, 186.
Bacillarieaj, 60.
Backhousia arfakensis Gilbs, 28, 34, 153.
Balsaminaceae, 145.
Begonia humboldtiana Gibbs, 56, 215.
Begoniacese, 149, 215.
Bignoniaceae, 179.
Boraginacese, 218.
Bulbophyllum arfakense J. J. S., 31, 122.
birugatum J. J. S., 31, 123.
ovalitepalum J. J. 8., 31, 124.
Burmanniacese, 103.
Calamus arfakianus Becc., 28, 32, 95.
humboldtianus Becc., 55, 93.
Prattianus Becc., 97.
Campanulacese, 183.
Casuarinacese, 207.
Celastracese, 214.
Centrolepidacese, 99.
Centrolepis novo-guineensis Gibbs, 36, 38,
99.
kinabaluensia Gibbs, 99.
Ceratostylis angiensis J. J. S., 31, 116.
Chlorophycese, 61.
Cladium arfakense Rend., 36, 90.
Clavaria Gibbsise Ramsb., 51, 187.
Clavariaceae, 187.
Coleus Gibbsise 8. Moore, 36, 37, 178.
Commelinaoeae, 202.
Compositse, 183, 222.
Convolvulacese, 218.
Corsia arfakensis Gibbs, 25, 104.
Corsiace89, 103.
Cucurbitacese, 222.
Cunoniacese, 142.
Cyathea arfakensis Gepp, 28, 69.
Cycadaceaj, 197.
Cylindrophora epitricha Ramsb., 190.
Cyperaceee, 89, 200.
Dacrydium novo-guineense Gibbg, 28, 30,
32, 38, 78.
Dacryomycetacese, 187.
Dendrobium curviruentum J. J. S., 33, 117.
inconspicuum J. J. S., 56, 205.
- latifrons J. J. &, 36, 118.
papnanum J. J. S., 32, 120.
parvifolium J. J. S., 121.
trifolium J. J. S., 32, 119.
Didiscus arfakensis Gibbs, 35, 166.
koebrensis Gibbs, 38, 165.
INDEX.
225
Dilleniacese, 148.
Diplycosia Lilian J. J. S., 28, 34, 35, 38,
170.
Dracaena novo-guineensis Gibbs, 51, 203.
Priinys arfakensis Gibbs, 28, 135.
Beccariana Gibbs, 34, 133.
Dryopteris villosipes Gepp, 33, 70.
Elaeocarpaceae, 146.
Elaeocarpus koebrensis Gibbs, 34, 146.
Epacridaceae, 167.
Ericaceae, 168.
Eriocaulaceae, 100.
Euphorbiaceae, 145, 213.
Fagaceae, 128.
Ficus conocephalifolia Ridl., 56, 208.
Flacourtiaceae, 215.
Flagellariaceae, 202.
Freycineda flaviceps Rend., 27, 28, 87.
Gibbseaj Rend., 27, 28, 31, 88.
Gentianaceae, 177.
Gesneriaceae, 180.
Gibbsia Rend., 129.
carstenzensis Rend., 130.
insignis Rend., 33, 130, 131.
Gigantochloa novo-guineensis Rend., 52,
55, 199.
Glomera angiensis J. J. S., 31, 115.
Gibbsiae J. J. S., 38, 114.
similis J. J. S., 31, 33, 113.
Goodeniacese, 183, 222.
Goodyera arfakensis J. J. S., 30, 108.
Gramineae, 89, 199.
Guepinia conferta Ramsb., 187.
Guttiferae, 149.
Halorrhagaceae, 159.
Halorrhagis suffruticosa Gills, 29, 35, 36,
37, 159.
Hemigraphis dorensis S. Moore, 54, 219.
Henslowia crassifolia Gibbt, 33, 132.
Hibbertia novo-guineensia Gibbs, 38, 148.
Homalanthus arfakiensis Hutch., 33, 145.
Hymenophyllum cernuum Gepp, 68, 69.
cincinnatum Gepp, 28, 29, 68.
Hypocreaceae, 63.
Hypolepis grandifrons Gepp, 55, 195.
Idenburgia Gilbs, 136.
Idenburgia arfakensis Gibbs, 29, 34, 139.
novo-guineensis Gibbs, 34, 137.
Iridacese, 101.
Jambosa arfakensis Gibbs, 29, 153.
Juncaceae, 100.
Kentia Gibbsiana face., 24, 30, 32, 91.
Kissodendron bipinnatuui Gibbs, 30, 161.
Labiataj, 178.
Lactuca prolixa S. Moore, 32, 184.
Leguminosae, 143, 211.
Lentibulariaceae, 180.
Leptothyrella sericoleae Ramsb., 64, 148.
Libocedrus arfakensis Gibbs, 24, 28, 29, 33,
84.
Liliacese, 100, 202.
Linaceae, 212.
Liparis Gibbsise J. J. S., 31, 112.
lacus J. J. S., 30, 111.
Lobelia arfakensis Gibbs, 28, ISi
Loranthaceae, 210.
Lucinaea reticulata VaL, 31, 180.
Lycoperdaceae, 190.
Lyonsia albiflora Gibbs, 28, 31, 177.
Lythraceae, 216.
Maesa frulicosa Gibbs, 30, 174.
Magnoliaceae, 133.
Malpighiaceae, 213.
Medinilla arfakensis Bak.f., 35, 33, 158.
rhodorhachis Bak.f., 56,216.
Melastomaceae, 157, 210.
Meliaceaj, 212.
Metrosideros arfakensis Gibbs, 29, 154.
Microstylis Gibbsiae J. J. S., 51, 204.
grandiflora J. J. S., 33, HO.
producta J. J. S., 30, 109.
Monimiaceae, 135.
Moraceae, 208.
Myrsinaceae, 174, 217.
Myrtaceae, 150.
Myrtus arfakensis Gibls,29, 35, 152.
koebrensis Gibbs, 34, 35, 152.
prostrata Gibbs, 38, 144, 151.
Myxophyceae, 59.
Nepenthaceae, 141.
Octarrhena cylindrica /. /. S., 28, 126.
226
INDEX.
Oldenlandia nutans Val, 35, 36, 37, 180.
Ornphalia arfakensis Ramsb., 64.
Ophiorrhiza insularis Val., 56, 220.
Orchidacese, 105, 203.
Palmse, 91, 200.
Palmervandenbroekia Gibbs, 162.
papuana Gibbs, 29, 162.
Pandauaceae, 88, 197.
Pandanus Tabbersianus Rend., 52, 198.
Patersonia noTO-guineensis Gibbs, 38, 101.
Pellionia Vanhasseltii Gibbs,- 51, 210.
Phalloidaceas, 188.
Phreatia spathulata J. J. S., 28, 126.
Pilocratera novo-guineensis Ramsb., 185.
Pinaceaa, 82.
Piper arfakianum C. DC., 28, 127.
bipunctatum A. DC., 56, 207.
bosnicanum C. DC., 56, 207.
pilosulinodum C. DC., 33, 128.
Piperaceae, 127, 207.
Pipturus papuanus Gibbs, 33, 128.
Pogonanthera hexamera Bak.f., 34, 158.
Poikilogyne Bak.f., 157.
arfakensis Bak.f., 29, 37, 157.
Polybotrya arfakensis Gepp, 27, 71.
Polygonaceae, 133.
Polyporaceae, 64, 186.
Polyscias sorongensis Gibbs, 216.
Poria Gibbsise Ramsb., 186.
Portulacacese, 211.
Psychotria vaccinioides Vol., 34, 38, 181.
Pteris baoibusoides Gepp, 18, 195.
Pullea papuana Gibbs, 29, 41, 141.
Rhododendron Gibbsise J. J. S., 36, 169.
undulaticalyx J. J. S., 37, 168.
Iliedelia exalata Val., 33, 102.
Rosacese, 143, 211.
Rubiacese, 180, 220.
Rutacese, 143.
Santalacese, 132.
Sapindacere, 145, 214.
Sarcosoina novo-guineensis Ramsb., 186,
190.
Saxifragaceae, 141.
Schisnmtoglottis dorensis Gibbs, 51, 201.
Sericolea arfakensis Gibbs, 32, 64, 148.
novo-guineensis Gibbs, 34, 38, 147.
Shefflera angiensis Gibbs, 31, 160.
arfakensis Gibbs, 30, 33, 159.
Solanaceae, 177.
Solanum Gibbsise J. R. Drum., 32, 177.
Sphaeropsidese, 64.
Spirseanthemum bullatum Gibbs, 28, 29,
142.
Syrabegonia arfakensis Gibbs, 32, 149.
parvifolia Gibbs, 33, 150.
Symplocacese, 175.
Symplocos arfakensis Gibbs, 30, 31, 175.
novo-guineensis Gibbs, 30, 176.
Taxacese, 78.
Tecomanthe volubilis Gibbs, 31, 34, 179.
Terminthodia rotundifolia Ridl., 38, 143.
Tbysanosuria, Gepp, 193.
dimorpbophylla Gepp, 51, 193.
TiliacesB, 214.
Trimenia arfakensis Gibbs, 29, 31, 41, 136.
Trimeniaceae Gibbs, 135.
Umbelliferse, 165.
Urticaceas, 128, 209.
Vacciniura ligustrifolium J. J. S., 174.
pilosiflorum J. J. S., 28, 34, 35, 172.
roseiflorum 7. J. S., 30, 173.
villosiflorum J. J. S., 38, 171.
Verbenacese, 218.
Violacese, 149.
Xerotus cinnamomeus Ramsb., 64.
Xylariacese, 03.
Xyridacese, 100.
Zingiberacea), 102.
PRINTED BY TAYLOR AND FRANCIS, RED LION COURT, FLEET STREET.
UNIVERSITY OF CALIFORNIA LIBRARY
Los Angeles
This book is DUE on the last date stamped below.
315
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