UNIVERSITY OF CALIFORNIA COLLEGE OF AGRICULTURE AGRICULTURAL EXPERIMENT STATION BERKELEY, CALIFORNIA YIELD, STAND AND VOLUME TABLES FOR DOUGLAS FIR IN CALIFORNIA FRANCIS X. SCHUMACHER BULLETIN 491 APRIL, 1930 UNIVERSITY OF CALIFORNIA PRINTING OFFICE BERKELEY, CALIFORNIA 1930 YIELD, STAND AND VOLUME TABLES FOR DOUGLAS FIR IN CALIFORNIA FEANCIS X. SCHUMACHER! INTRODUCTION The United States Forest Service has reecntly completed a study of the yields of Douglas fir (Pseudotsuga taxifolia Britt.) for even-aged stands of Oregon and Washington. 2 The work was not extended to stands south of the Willamette-Umqua divide in Oregon because from observation it is believed that this line roughly divides the Douglas fir forest into two types of decided difference in stand characteristics. But the commercial range of the species on the Pacific slope extends into California about as far south as Yosemite National Park in the Sierra and about San Francisco Bay along the coast. To report the yields of well-stocked, even-aged stands of the species in California is the object of this bulletin. That there are significant differences in certain stand character- istics between the two general regions seems established from the work presented herein. GROWTH OF DOUGLAS FIR STANDS IN CALIFORNIA The growth of the species is shown by tables which state the yield of even-aged stands over a period of years. Age, timber productive quality of the area, and stand density are the most important growth- determining factors of a stand. As there is no satisfactory way of expressing stand density in absolute terms, normal-yield tables based on the ideal density which produces maximum volume are presented. Basic Data The normal-yield tables for Douglas fir are based on 159 sample plots scattered through the geographical range of the species in California. i Assistant Professor of Forestry and Assistant Forester in the Experiment Station. 2 McArdle, R. E. Rates of growth of Douglas fir forests. West Coast Lumberman, 54:90-95, 1928. This article summarizes the results of the study. The complete work is to be published soon as a bulletin of the United States Department of Agriculture. 4 University of California — Experiment Station Plot Selection. — Within even-aged stands plots were established so as to enclose a comparatively complete crown canopy by excluding the larger openings which follow failure of reproduction or accident and at the same time to include within boundaries the area equivalent to that which seemed to be used by the enclosed timber. Plots were surveyed with staff compass and chain. Age Determination. — The age of each plot was determined by counting the annual rings on cores extracted (with Swedish increment borers) from near the base of several trees. By the age of the tree is understood the number of rings on the core plus the necessary cor- rection for height growth to the point of boring. The age of the oldest tree was taken as the plot age although the difference between the ages of the youngest and oldest tree examined was seldom more than two or three years. Field Measurements. — Diameter breast high of every tree was measured with diameter tape and tallied by species and crown class (dominant, codominant, intermediate, or suppressed). The heights of fifteen to twenty-five trees were measured with the Forest Service hypsometer, from horizontal distances measured with the Leitz Fardi Range Finder of 20-centimeter base. Heights were plotted over diameter on cross-section paper in the field, the number of measurements necessary being judged at the time by the range of diameters present and their dispersion around the free-hand curve. A short description of physiographic features completed the field work on each plot. Office Computations. — The computational work necessary for each plot is evident from following paragraphs. The yield tables were constructed by correlating dependent growth variables with age and site quality by the method described by Bruce and Reineke, 3 and the stand tables are based on CharlierV method of calculating theoretical frequencies. Normal Yield Tables Tables 1 to 11 and figures 1 to 11 indicate the growth of Douglas fir in fully-stocked stands in California, for age and site index. 5 Site index is herein defined as the height that the average dominant Douglas fir will attain, or has attained at 50 years of age. Average '■> Druce, D., and L. H. Reineke. Multiple curvilinear correlation in forest investigative work. Unpublished contribution of the United States Forest Service. 1927. < Charlier, C. V. L. Die Grundzuge der mathematischen Statistik. p. 3-125. Lutke und Wulff, Hamburg. 1920. : - Before constructing these tables the sample plot data were compared to the yield tables for Douglas fir in Oregon and Washington. See p. 27. Bul, 491] Yield, Stand, and Volume Tables for Douglas Fir TABLE l Height of the Average Dominant Tree* Age, Site index — height of average dominant at 50 years years 60 80 100 120 140 30 feet • 39 feet 54 feet 67 feet 81 feet 95 40 50 68 85 102 120 50 60 80 100 120 140 60 68 89 112 135 156 70 74 98 122 147 170 80 79 104 131 158 182 90 83 110 138 166 192 100 86 114 146 173 201 110 89 118 152 179 209 120 92 122 156 185 216 130 96 125 159 189 220 140 98 128 162 193 224 150 99 130 164 196 228 160 100 132 165 198 232 * The height from average ground level to tip of the dominant tree of average basal area for the dominant class. SO 60 70 80 ^20 /JO /-90 /so Too Fig. 1. — Height of the average dominant tree for age and site index. These curves were used in site classification of the plots. University of California — Experiment Station TABLE 2 Height of Average Tree* Age, Site index — height of average dominant at 50 years years 60 80 100 120 140 30 feet feet 41 feet 58 feet 72 * feet 85 40 58 77 94 110 50 47 71 92 110 131 60 57 81 104 127 148 70 65 89 114 140 163 80 70 96 123 152 176 90 75 102 132 160 187 100 78 107 139 168 196 110 82 112 145 176 120 85 117 149 180 130 88 121 154 184 140 90 124 157 188 150 91 126 159 192 160 92 127 161 194 The height from average ground level to tip of the tree of average basal area. /30 /-K) /5Q Age /r? c/ears Fig- 2. — Height of the average tree for age and site index. Bul. 491] Yield, Stand, and Volume Tables for Douglas Fir TABLE 3 Number of Trees to the Acre* Site index — height of average dominant at 50 years Age, 60 80 100 120 140 years Number of trees to the acre 30 1060 672 485 394 40 780 497 364 297 50 1033 601 386 278 230 60 790 475 302 220 182 70 643 382 241 176 147 80 530 313 200 148 121 90 445 260 168 125 100 100 378 225 143 104 85 110 324 193 122 91 120 282 170 107 80 130 254 152 95 70 140 230 138 87 62 150 212 124 79 58 160 198 113 75 54 The number of trees that have reached a height of at least 4.5 feet (breast height). Fig. 3. — Number of trees to the acre for age and site index. University of California — Experiment Station TABLE 4 Basal, Area to the Acre* Age, Site index— height of average dominant at 50 years years 60 80 100 120 140 30 sq.ft. sq. ft. 198 sq. ft. 217 sq. ft. 230 sq.ft. 243 40 223 243 267 285 50 205 237 264 290 305 60 214 249 281 305 319 70 222 260 295 316 328 80 228 271 305 323 334 90 233 280 313 329 339 100 238 288 318 333 342 110 242 294 322 336 120 245 298 326 338 130 248 302 328 340 140 250 305 330 341 150 251 308 331 342 160 252 309 332 343 The sum of the cross-sectional areas at breast height, in square feet. 60 70 SO 90 Age /n years Fig. 4. — Growth in basal area to the I /20 J30 1<K) for age and site index. Bul. 491] Yield, Stand, and Volume Tables for Douglas Fir TABLE 5 Average Diameter, Breast 1 High* Age, Site index — height of average dominant at 50 years years 60 80 100 120 140 inches inches inches inches inches 30 5.9 7.7 9.3 10 6 40 7.2 9.5 11.6 13.3 50 6.0 8.5 11 2 13 8 15 6 60 7 1 9.8 13.1 15.9 17.9 70 8 11.2 15 18.1 20.3 80 8 9 12 6 16.7 20 22.5 90 9 8 14.0 18 5 22 25 100 10 7 15.3 20.2 24.2 27 110 11 7 16 7 22.0 26.0 120 12 6 17 9 23.6 27.2 130 13 4 19 1 25.2 29.8 140 14 1 20 2 26.3 31.8 150 14 7 21 3 27.7 32.9 160 15 3 22.4 28.5 34 1 The diameter in inches of the tree of average basal Fig. 5. — Average diameter breast high for age and site index — the diameter of the circle of average basal area. 10 University of California — Experiment Station TABLE 6 Mean Diameter, Breast High* Age, Site index — height of average dominant at 50 years years 60 80 100 120 140 inches inches inches inches inches 30 5 6 7 8.4 9 6 40 6 3 8.5 10 7 12 2 50 5 1 7.6 10 2 12 8 14 5 60 6.1 8.9 12 14 8 16.7 70 7.0 10.3 13.8 16 9 19.0 80 8.0 11.6 15.7 18 9 21.3 90 8.9 12 9 17.3 20 9 23.7 100 9.8 14 2 18 22 9 25 6 110 107 15.6 20.6 24 8 120 11.6 16.8 22 26 6 130 12 4 17.9 23.4 28 4 140 13.0 19.0 24.8 30 2 150 13 6 20.1 26.2 31 9 160 14 .2 21.2 27.6 33 5 * The mean of all diameters on an average acre. s £ v. s & > y y 1 /, > s 1" A ^ «5j «? s> 90' J ^ /O 20 30 <K> 50 CO 70 60 90 /OO //O /20 /30 tfO (SO /60 Age /n years Fig. 6. — Mean diameter breast high for age and site index — the average of all diameters in the stand. Bul, 491] Yield, Stand, and Volume Tables for Douglas Fir 11 TABLE 7 Cubic Volume to the Acre* Age, Site index — height of average dominant at 50 years years 60 80 100 120 140 cu. ft. cu.ft. cu. ft. cu.ft. cu. ft. 30 3,300 4,900 6,500 7,700 40 2,300 5,000 7,200 9,350 10,900 50 3,650 6,400 9,000 11,700 13,100 60 4,800 7,600 10,500 13,200 14,800 70 5,700 8,550 11,750 14,500 16,200 80 6,400 9,350 12,750 15,500 17,400 90 6,950 10,000 13,550 16,400 18,400 100 7,400 10,500 14,300 17,200 19,200 110 7,700 11,000 14,900 17,950 120 7,950 8,150 11,400 11,700 15,400 15,950 18,600 19,200 130 140 8,350 8,500 12,000 12,300 16,400 16,800 19,800 20,300 150 160 8,600 12,500 17,200 20,800 * The cubic volume of the entire stem of all trees from ground to tip but without limbs or bark. The volume table used is given following p. 22. Fig. 7. — Growth in cubic volume to the acre for age and site index. 12 University of California — Experiment Station TABLE 8 Mean Annual. Growth in Cubic Volume to the Acre* Age, Site index — height of average dominant at 50 years years 60 80 100 120 140 30 cu. ft. cu. ft. 110 cu. ft. 163 cu.ft. 217 cu.ft. 257 40 58 125 180 234 270 50 73 128 180 234 262 60 80 127 175 220 247 70 82 122 168 207 232 80 80 117 159 194 218 90 77 110 151 182 205 100 74 105 143 172 192 110 70 100 135 163 120 66 95 128 155 130 63 90 123 m 140 60 86 117 141 150 57 82 112 135 160 54 78 107 130 The cubic volume on the acre divided by the age 60 70 80 90 Age tn years /OO //O /20 /30 MO /SO /GO Fig. 8. — Mean annual growth in cubic volume to the acre for age and site index. Bui* 491] Yield, Stand, and Volume Tables for Douglas Fir 13 TABLE 9 Number of Trees Eight Inches and Over,, to the Acre Site index — height of average dominant at 50 years Age, 60 80 100 120 140 years Number of trees eight inches and over 30 185 265 258 252 40 252 278 251 230 50 191 279 258 221 198 60 250 277 230 190 170 70 266 260 203 165 143 80 269 234 179 144 118 90 260 210 158 124 98 100 243 190 139 195 85 110 225 174 122 SI 120 210 159 106 80 130 199 146 94 70 140 187 135 85 63 150 178 124 79 58 160 167 114 75 54 O JO 20 30 <K> 50 CO 70 80 90 /OO //O /20 /30 /fO /SO /60 Age /n c/eors Fig. 9. — Number of merchantable trees to the acre for age and site index. 14 University of California — Experiment Station TABLE 10 Volume Board Measure to the Acre* Site index — height of average dominant at 50 years Age, years 60 80 100 120 140 bd. ft. bd. ft. bd. ft. bd. ft. bd. ft. 30 7,760 17,050 27,900 37,000 40 16,000 31,700 47,700 59,400 50 8,940 25,200 45,000 64,800 76,200 60 15,060 34,300 56,900 77,400 90,600 70 21,000 42,700 67,300 89,000 103,500 80 26,500 49,650 76,200 98,400 114,800 90 31,400 55,700 83,800 107,400 124,100 100 35,900 60,600 91,000 115,300 131,500 110 39,400 42,200 65,650 68,200 73,200 76,400 97,600 102,700 122,200 120 127,600 130 44,600 46,750 107,800 133 , 700 140 111,800 139,000 150 48,300 79,700 115,700 142,900 160 49,600 82,400 119,000 146,600 * The board foot contents of the trees by the International log rule of J^-inch kerf between a stump of one foot and a top diameter inside bark of 5 inches scaled in 16-foot logs with 0.3-foot trimming allot- ment to each. Gross volumes are presented, no account being taken of cull factors. The volume table used is given following p. 22. MQ/QOL S HO' I30.0OG / /20,OOC ,/ / / /OQOOO / 9QOO0 80.000 70,00C / / eopoo // / / 5O.0OC / It ■9O.000 30.000 20,000 /QOOO O LL ' . f / / ' r / ■20<Q •oo- § ! C 10 20 30 fO 30 00^ 70 80 90 /OO I/O 120 130 1*0 ISO 160 Age. //? years Fig. 10. — Growth in volume board measure to the acre for age and site index. Bul. 491] Yield, Stand, and Volume Tables for Douglas Fir 15 TABLE ll Mean Annual Growth m Board Feet to the Acre* Age, Site index— height of average dominant at 50 years years 60 80 100 120 140 30 bd. ft. bd. ft. 259 bd. ft. 568 bd. ft. 930 bd. ft. 1,234 40 400 793 1,193 1,485 50 179 504 900 1,296 1,525 60 251 565 948 1,290 1,510 70 297 610 962 1,270 1,480 80 331 620 952 1,230 1,436 90 349 619 931 1,193 1,380 100 359 606 910 1,153 1,315 110 359 597 888 1,112 120 352 568 859 1,065 130 343 553 830 1,028 140 334 546 799 993 150 325 531 771 953 160 310 515 744 916 The board foot volume on the acre divided by the age. Fig. 11. — Mean annual growth in volume board measure to the acre for age and site index. 16 University of California — Experiment Station height of the dominant, or of the dominant and codominant stand is generally accepted as the most accurate and readily measurable factor of timber-productive quality of an area, because it bears a very close relationship to volume production within the limits of normal stocking. Although the yield tables for Douglas fir in Oregon and Washing- ton define site index as the height of the average dominant and codominant at 100 years, the height of the average dominant at 50 years is used here in order to conform with site index as defined in other California yield studies. 6 ' 7 Height curves used in determining the site-index of each plot are shown in figure 1. Check of Basic Data Against the Yield Tables Table 12 shows the check of the values of the 159 sample plots against the yield tables interpolated to nearest year of age and nearest foot of site index. TABLE 12 Check of Basic Data Against 1 Yield Tables Basal area All trees per acre Average d. b. h Volume in cubic feet Volume in board measure. Aggregate difference, per cent* -0.0 -0.2 +0 9 -0.0 +0.8 Mean difference, per cent** Standard error of estimate, per centf 16 4 27.0 15 .7 16 3 20 4 Standard error of yield table value, per centj ±1 30 ±2 14 ±1.24 ±1.29 ±1.67 * The aggregate difference is the sum of the plot values expressed as a percentage difference from the sum of corresponding tabular values. ** The mean difference is the mean of the per cent deviations of the plot values from corresponding tabular values. t Standard error of estimate (<r es <) = — in which x = deviation of each plot from its tabular value N in per cent, 2=the sum, and N = number of plots. % Standard error of yield table value is the same as that ordinarily understood as standard error of the mean, the mean here being tabular value for age and site index. It is expressed thus: a™ =— % =L . Stand Tables Although yield tables are basic to the solution of many forest management problems, they are not complete without stand tables as problems of valuation and utilization require knowledge of such stem distribution. Stand ta bles for Douglas fir are given in table 13. 8 6 Schumacher, Francis X. Yield, stand and volume tables for white fir in the California pine region. California Agr. Exp. Sta. Bui. 407:1-26. 1926. 7 Schumacher, Francis X. Yield, stand and volume tables for red fir in Cali- fornia. California Agr. Exp. Sta. Bui. 456:1-32, 1928. 8 The analysis of stem distribution and construction of stand tables is explained on pp. 32. Buu 491] Yield, Stand, and Volume Tables for Douglas Fir 17 TABLE 13 Normal Stand Table, for Douglas Fir Including all Trees Age of stand in years D. b. h. class, inches 30 40 50 60 70 80 90 100 110 120 130 140 150 160 Number of trees by diameter classes Site index 60 feet at 5C years 0.0- 2.0 2 0-40 173 233 251 184 107 52 22 8 97 135 177 159 110 65 32 11 2 58 90 127 132 105 72 36 16 6 28 60 89 102 91 70 46 27 16 1 15 36 62 79 80 67 49 31 24 2 13 22 44 59 65 60 45 33 32 5 8 14 31 43 52 51 45 36 36 9 7 9 21 33 41 43 40 34 40 12 2 2 7 16 26 33 37 36 32 42 16 4 2 5 12 21 27 32 33 30 44 19 5 2 4 9 17 23 28 29 27 43 21 6 1 3 4.0- 6.0 8 6.0- 8.0 15 8.0-10.0 21 10.0-12.0 12.0-14.0 25 27 14.0-16.0 26 16.0-20 42 20.0-24.0 24 0-28 . . 23 7 28.0-32.0 Total 1030 788 642 530 445 378 325 282 251 230 210 198 Site index 80 feet at 5C years 0.0- 2.0 189 240 259 183 105 53 21 5 72 134 175 157 122 69 36 14 4 42 71 104 119 101 77 48 26 12 19 39 67 85 85 70 52 33 23 2 10 21 40 55 65 60 50 37 37 7 6 13 26 37 46 48 43 37 43 13 2 4 6 15 25 32 36 37 32 47 22 5 4 5 9 16 23 28 31 29 46 25 8 1 2.0- 4.0 3 6 11 17 21 24 25 43 28 13 3 2 4 7 13 16 19 21 39 29 15 6 1 1 3 5 9 12 15 17 35 28 18 8 1 1 2 4 6 10 12 14 30 27 19 10 3 4.0- 6.0 6 0-80 1 3 5 7 9 11 26 25 19 12 5 1 2 8.0-10.0 10.0-12.0 4 6 12 0-14 g 14.0-16.0 9 16.0-20.0 23 20 0-24 23 24 0-28 19 28 0-32 12 32 0-36 6 36 0-40 1 Total 1055 783 600 475 382 314 261 225 194 172 152 138 123 114 Site index 100 feet at 50 years 0.0- 2.0 65 101 139 136 106 65 36 17 5 18 47 73 92 88 75 51 30 19 2 10 20 41 56 64 61 51 37 37 7 3 10 22 33 41 45 43 37 49 16 3 2 5 11 20 27 32 34 31 49 26 7 1 1 3 6 11 17 21 25 26 45 30 14 3 2.0- 4.0 1 3 6 11 14 18 20 39 31 18 6 1 1 2 4 7 10 12 14 32 28 20 10 3 1 1 2 4 7 9 10 26 25 20 12 5 1 4.0- 6.0 1 2 3 4 6 8 19 20 19 14 8 3 6.0- 8.0 1 2 3 5 6 15 18 17 13 9 5 1 2 2 3 5 12 15 15 13 10 5 3 8 0-10.0 1 2 3 4 10 13 14 12 9 7 3 1 1 10.0-12.0 2 12.0-14.0 2 14.0-16.0 3 16 0-20.0 9 20 0-24.0 11 24.0-28.0 28 0-32 12 12 32 0-36 10 7 4 2 36 0-40 40 0-44 44.0-48.0 48 0-52.0 Total 670 495 384 302 245 202 168 143 123 107 95 86 79 75 18 University of California — Experiment Station Table 13 — (Concluded) Age of stand in years D. b. h. class, inches 30 40 50 60 70 80 90 100 110 120 130 140 150 160 Number of trees by diameter classes Site index 1W feet at 50 years 0-20 26 51 77 92 86 67 45 26 16 14 20 39 52 60 56 47 35 33 7 2 8 19 29 37 41 40 35 48 18 3 1 4 8 16 21 26 29 28 48 28 10 1 2 0-40 1 4 8 13 17 20 21 40 31 16 5 1 2 4 7 10 13 15 32 28 21 10 3 1 1 2 4 6 8 10 24 24 20 14 6 2 4 0- 6.0 1 1 2 4 6 7 17 20 19 14 9 4 1 1 2 3 4 5 13 16 16 14 10 5 3 1 1 2 3 3 9 13 14 13 11 7 3 1 6 0- 8.0 8 0-10 10 0-12.0 1 2 3 7 10 12 12 10 8 5 2 1 2 6 8 10 10 10 8 5 3 1 1 1 1 5 6 8 8 8 8 6 4 2 1 12.0-14.0 14.0-16.0 1 1 16.0-20.0 4 20 0-24 5 24 0-28 7 28.0-32.0 32 0-36 8 g 36.0-40.0 40 0-44 7 6 44.0-18.0 48 0-52 4 2 52 0-56 1 105 72 Total 486 363 280 220 177 146 122 92 81 65 59 54 Site index 140 feet at 50 years 0.0- 2.0 12 26 48 63 69 62 49 34 29 3 10 10 22 33 42 45 43 37 44 13 1 2 3 8 16 22 28 31 30 49 29 10 1 2.0- 4.0 1 4 7 13 17 20 23 44 31 18 6 1 2 4 7 10 12 14 32 29 21 11 3 1 4.0- 6.0 6.0- 8.0 1 2 3 6 8 9 23 24 21 14 7 3 1 1 2 3 5 6 16 18 18 15 10 5 1 1 1 2 3 4 11 14 15 13 11 7 3 1 8.0-10.0 , 10.0-12.0 12.0-14.0 14.0-16.0 16.0-20.0 20.0-24.0 24.0-28.0 28.0-32.0 32.0-36.0 36.0-40.0 40.0-44.0 44.0-48.0 48.0-52.0 Total 395 300 229 183 147 121 101 86 Bul, 491] Yield, Stand, and Volume Tables for Douglas Fir 19 VOLUME TABLES Preliminary to the study of yields in cubic and board feet, volume tables in these units were prepared. 9 The basic tree data of the tables presented are from measurements taken by the Division of Forestry from eight, previously measured, even-aged sample plots in Mendo- cino and Trinity counties. The ages of the trees measured were from 33 to 111 years. Table 14 is the volume in cubic feet, and states the entire volume of the stem, including stump and top, but without bark. It was pre- pared by correlating cylindrical form factor with diameter, height and site index. As no significant relationship was discovered with site index, the table may be used for any site class. TABLE 16 Check of Basic Tree Data Against Volume Tables Aggregate difference, per cent Mean difference, per cent Standard error of estimate, per cent Standard error of volume table value, per cent Cubic foot volume -0 -0.7 -0.7 -0.4 11.7 12.1 ±0.71 ±0.81 Table 15 is the volume in board measure. It includes the board- foot contents of the trees between a one-foot stump and top diameter inside bark of five inches. It was prepared by correlating the number of board feet to a cubic foot with the diameter and height of the trees. Table 16 shows the check of the basic tree data with the volume tables. DISCUSSION The generic name of Douglas fir, Pseudotsuga, implies that its common name is a misnomer in that the tree is not a true fir of the Abies genus, such as red and white fir. One of the outstanding differences in characteristic growth between Douglas fir and the California true firs already studied 10 is the fact that the crown of the former becomes rather widespread when not confined by neighboring trees. Now diameter breast high bears a 9 The check of the volumes of the basic tree data against the volume tables for immature Douglas fir in Oregon and Washington is explained on pp. 35, io See Bul. 407 and Bul. 456 previously referred to. 20 University of California — Experiment Station noticeably constant ratio to crown width in any one timber species; hence the net result of widespread Douglas fir crowns with their asso- ciated greater trunk diameters at breast-height — when the stand is deficient in number of trees — is the tendency to form complete crown canopies and therefore to approach normal stocking by basal area. Figure 12 indicates this within the limits of the data presented. The regression of average diameter breast high on number of tree is Average d.b.h. in per cent of the tabular value ^ 1,000,000 Number of trees in per cent of the tabular value Now basal area in square feet is .00545 times the number of trees times the square of average diameter breast high in inches. But 30 -90 SO 60 70 GO 90 JOO //O /SO /30 tfO /50 /CO /70 /80 /SO 200 A/umber of trees on p/ofs /n per cenf of c//e/d tob/e. Fig. 12. — Relation between average diameter and number of trees. within a given site-age class by the above equation, the number of trees times the square of the average diameter is constant ; that is, basal area tends to be independent of the number of trees as long as there are at least sufficient trees to allow a complete crown canopy. The true firs, on the other hand, have characteristically narrow crowns even when growing in the open; hence they have not the ability to form complete crown canopies when deficient in number of trees. Deficiency in number of trees within a site-age class results in deficiency in basal area, because of the narrower crowns and the crown diameter— diameter breast high ratio. Therefore, average diameter is proportional to the number of trees and not to the square Bul. 491] Yield, Stand, and Volume Tables for Douglas Fir 21 L •^Z5C t § 20C Si Co//fom/a -x /^Oregon drd Wos/i/ng/on ^/ou ^6 <^0 ^0 6<? ao /tfO /.a? /w /<^? Age /n years /ZOO /ooc a) S 60C ^ 50C ^300 \ \^rOreana nns/ hSas/7/rafnr, Co/zforn/'o ^ ^ ao \ 6C I 20 40 60 80 /0O Age //? yeans /20 W /60 Fig. 13. — Comparison of California stands by basal area and by number of trees with those of Oregon and Washington for site index 140' feet — height of the average dominant and codominant tree at 100 years. 22 University of California — Experiment Station root of the number; while the effect on basal area of increasing the number of trees through normality to an overstocked condition is that it rises to a maximum and then falls off. 11 The differences in growth of Douglas fir between the central and southern parts of its range are evident from figure 13 which shows the comparison of yield values in basal area and in number of trees with age for average site class. One must infer that the stand in California breaks up earlier in life than it does farther north for the following reasons : (1) It has fewer trees to the acre throughout and these decrease at a greater rate. (2) It grows faster in basal area when young, but after about 100 years this growth practically stops though in the north it is still vigorous. Such differences are not unknown in other species which have a wide latitudinal range. In taking part in a recent discussion as to the relative merits of timber producing regions in the United States, Zon 12 compares the yield of two Russian species — Scotch pine and birch — in northern and southern provinces of that country and notes the same tendencies. 11 See figures 6 and 7, Bui. 456. 12 Zon, E, Forestry versus climate. Jour. Forestry. 26:711-713. 1928. Douglas I Diameter breast height, inches Basis, No of trees... Th< Diameter breast height, inches 6 7 8 9 : 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 , 40 41 42 43 44 1 Basis, number of trees 70 80 10 20 30 40 49 59 70 84 97 111 126 140 157 174 190 206 224 240 261 277 296 316 335 355 374 396 415 437 457 480 502 525 546 570 593 617 640 680 87 107 124 142 160 182 203 226 250 273 299 323 350 378 405 334 463 492 526 553 583 613 642 684 707 738 770 810 836 872 910 945 985 20 115 140 162 189 214 240 268 298 320 348 375 420 454 480 523 560 598 634 670 715 754 792 830 878 912 955 997 1030 1080 1120 1180 1210 1270 29 33 50 73 97 120 144 170 198 225 254 285 317 350 384 418 452 490 530 573 617 666 700 741 792 833 880 928 975 1020 1070 1110 1180 1220 1280 1340 1390 1450 1500 50 100 60 84 112 138 165 193 223 256 291 328 365 403 445 482 525 573 617 664 705 760 815 910 967 1020 1080 1130 1190 1250 1300 1380 1420 1490 1550 1600 1680 1720 36 Ag( tr_. Stump height, 1 foot. Trees scaled in 16-foot logs with 0.3-foot trimming allowance to 5 i Basis, 215 trees, measured by the Division of Forestry, 1927, in ev Heavy lines in the tables show limits of basic data. 22 University of California — Experiment Station root of the number; while the effect on basal area of increasing the number of trees through normality to an overstocked condition is that it rises to a maximum and then falls off. 11 The differences in growth of Douglas fir between the central and southern parts of its range are evident from figure 13 which shows the comparison of yield values in basal area and in number of trees with age for average site class. One must infer that the stand in California breaks up earlier in life than it does farther north for the following reasons : (1) It has fewer trees to the acre throughout and these decrease at a greater rate. (2) It grows faster in basal area when young, but after about 100 years this growth practically stops though in the north it is still vigorous. Such differences are not unknown in other species which have a wide latitudinal range. In taking part in a recent discussion as to the relative merits of timber producing regions in the United States, Zon 12 compares the yield of two Russian species — Scotch pine and birch — in northern and southern provinces of that country and notes the same tendencies. ii See figures 6 and 7, Bui. 456. 12 Zon, R. Forestry versus climate. Jour. Forestry. 26:711-713. 1928. TABLE 14— Douolas Pra— Volume in Cubic Feet Tola, hri.ht i„ ,..t s -l-l-l-l " 80 90 - | 1, | ,29 ,80 ,40 | ,60 | ,80 | 170 | ,80 | ,90 200 n"m- .'»" 15 9 63.2 81.8 8 09 30.0 69 5 76 3 83.1 900 108 123 21 30 3 68.7 75.8 128 6S.7 148 158 88.3 98.1 ,08 118 127 ,70 26 1 10 27.9 55.5 m 180 13 9 68 108 130 101 81 5 138 150 202 2,9 231 247 ,86 29, 90 8 ,97 21, 326 36, 510 580 35 8.66 10.7 180 36.1 \Z 1 29 2 - :;; 11 11 30 22 29 1 553 69.1 66 8 81 6 86.7 90 8 17 4 200 29 6 24 30.9 161 169 171 179 187 200 , 4 31.9 36.10 160 168 176 182 200 208 214 223 ,3 I 62 66 61 65 70 79 84 89 3 3 2 3 6 160 ,70 \ 26 197 205 213 221 232 242 2M 261 I » 239 260 271 160 186 218 230 240 324 338 318 179 236 246 260 270 283 349 361 202 267 320 333 319 361 32 91 8 104 109 120 -.: Jlu 343 300 377 1 38 328 346 359 375 390 200 349 385 380 307 413 3O0 310 370 390 424 Unsis, No. 2 .7 19 20 „ 50 36 ,3 .6 u • „ 4 9 267 TABLE 15 DovauiS Fir-Volume in Board Feet Total hoight in Tort br ^B"' 00 70 . J»oJj»J_™Jjlo| "0 I ,40 1 ,50 1 160 1 170 1 ,80 | ,90 1 200 Ba«i«, Volume in board feet ; 4 : 17 " Jl_ 47 34 62 :: ■ ■ : 107 » 126 ,: 8 32 42 50 60 — !il_ 79 91 100 113 ,24 139 160 161 175 10 » 30 44 01 73 95 no 125 140 152 ,67 185 200 216 230 19 10 40 58 81 07 1,2 128 143 162 ,8, 198 2,5 ' 238 254 271 295 21 59 115 M !« 187 m 200 220 287 260 2 ' 332 391 m 12 87 u 13 Si" 162 ,98 223 259 290 Hir 306 Z 366 427 Z 600 570 " 15 97 142 189 225 258 294 330 367 106 118 483 526 570 602 626 16 111 160 214 254 29, 332 370 415 410 500 543 595 642 692 735 7 17 140 103 268 285 " «6 m 483 515 604 680 708 z z 5 10 157 128 291 350 403 458 617 572 634 605 751 8,7 883 950 ,000 8 20 171 250 320 384 445 805 559 628 700 788 827 900 075 1010 1110 21 190 273 348 418 482 550 613 690 764 817 900 080 1070 1140 1210 , 22 209 299 375 452 526 600 662 750 836 007 978 ,070 1150 1230 1300 s 23 224 323 420 i-iii 573 656 718 812 900 985 1080 1210 1320 1410 5 21 240 350 454 _530_ 617 705 774 880 075 ,060 1140 ,240 1310 1420 1510 8 25 261 >78 480 573 664 735 837 950 ,060 1,10 ,230 ,320 1150 1530 1610 Z 2 2 666 70S 1210 1400 1510 1510 1630 !™ ;™ 27 ■...7 051 inv, ILW 2 28 310 463 598 700 815 918 1010 1150 1270 1390 1500 1620 1870 2000 „ 29 335 482 634 741 860 078 1070 1210 1340 1470 1800 1720 1880 1000 2100 30 355 626 670 702 010 1040 1140 1300 1430 1560 1700 1830 1990 2,00 2200 31 374 553 715 833 967 1090 1200 1380 1510 1650 1800 1030 2000 3230 2380 „ 32 415 613 792 2 z un 1350 1M0 1590 irni ,<«] 2300 16 3 » 2» , 31 437 642 830 975 1130 1280 1100 1000 1760 1910 2100 2160 2130 26,0 2780 36 457 684 878 1020 1,90 1340 1500 1690 1850 2040 2200 2380 2680 2740 2900 „ ,070 37 502 738 955 1110 ,300 1480 1620 1830 2000 2230 MOO 2600 2800 3000 3200 „ 38 525 770 097 ,180 ,380 1530 1700 1910 2100 2320 2530 2710 2930 3160 3310 39 546 810 1939 1220 ,420 1600 1760 2000 2200 2430 2640 2830 3060 3190 3400 o 40 570 836 1080 ,280 ,490 1680 1890 2090 2300 2550 1730 2020 3200 3420 3630 n 11 593 872 1120 1340 ,550 1730 1920 2180 2390 2640 2880 3080 3330 3850 3760 42 617 910 1180 1390 1600 1800 2000 2240 2490 1710 < 3200 3180 j;i«i 3920 " 640 945 1210 1450 1680 1890 2090 2310 1580 2830 310O 3340 3600 3810 4050 14 m ,,:, 1170 1500 1720 1950 2170 2440 2680 2950 3830 3180 3760 3980 1 B„i.. number ol tree, 1 20 19 50 36 j 33 ,6 " 1 11 4 • ■1 • o 215 ■ [i.trri,,,!,.,,,!,! rulf i»,-mrli k TABLE 15 22 i IB — Volume in Board Feet rot Total height in feet nu 110 120 130 140 150 160 170 180 190 200 Basis, Number it ] 1 of trees Volume in board feet sou 30 34 39 45 52 58 66 73 79 87 5 the 47 53 62 69 77 86 96 107 117 126 16 wit 68 79 110 91 125 100 140 113 152 124 167 139 185 150 200 161 216 175 230 10 CaJ 95 19 fol] 128 143 162 181 198 215 * 238 254 274 295 24 155 177 200 220 240 260 286 309 332 353 24 at i 187 220 259 294 212 238 279 261 306 354 287 338 390 448 312 366 427 483 340 400 460 526 368 430 500 570 394 454 535 602 421 493 570 626 14 250 290 330 14 320 367 13 yea 406 11 vigi 332 370 415 460 500 543 595 642 692 735 7 373 415 415 461 463 518 515 575 564 613 680 665 740 718 798 778 860 824 915 5 wid 627 10 458 517 572 634 695 751 817 883 950 1000 8 the Zon birc 505 559 628 700 768 827 900 975 1040 1110 3 550 613 690 764 817 900 980 1070 1140 1210 6 600 662 750 836 907 978 1070 1150 1230 1300 5 the 656 705 718 774 812 880 900 975 985 1060 1080 1150 1240 1240 1340 1320 1420 1410 1520 5 i 1140 8 l: 735 837 950 1060 1140 1230 1320 1450 1530 1640 807 890 1010 1120 1210 1300 1310 1400 1410 1510 1540 1630 1640 1770 1750 1880 3 867 951 1080 1200 2 918 1010 1150 1270 1390 1500 1620 1760 1870 2000 976 1070 1210 1340 1470 1600 1720 1880 2000 2100 1040 1140 1300 1430 1560 1700 1830 1990 2100 2200 1090 1160 1210 1200 1270 1350 1380 1440 1510 1510 1590 1690 1650 1740 1800 1900 1930 2090 2220 2380 2500 2380 2500 2620 2030 2200 2300 1 1850 2000 2150 1280 1400 1600 1750 1920 2100 2260 2430 2610 2780 1340 1500 1690 1850 2040 2200 2380 2580 2740 2900 1400 1560 1760 1920 2130 2300 2500 2680 2890 3040 1 1480 1620 1830 2000 2230 2400 2600 2800 3000 3200 1530 1700 1910 2100 2320 2530 2720 2930 3160 3340 ' 1600 1790 2000 2200 2430 2640 2830 3060 3290 3490 , 1680 1890 2090 2300 2550 2730 2920 3200 3420 3630 1730 1920 2180 2390 2640 2880 3080 3330 3850 3760 1800 2000 2240 2490 2720 3000 3200 3480 3700 3920 1890 2090 2310 2580 2830 3100 3340 3600 3810 4050 i 1950 2170 2440 2690 2950 3330 3480 3750 3980 4200 1 i 33 16 11 9 11 4 2 215 ach< i d. i. b. in top by International rule (H-inch kerf). m-aged stands in Mendocino and Trinity Counties. APPENDIX Bul. 491] Yield, Stand, and Volume Tables for Douglas Fir 25 YIELD AND STAND TABLES Basic Data The sample plots on which the yield and stand tables are based were measured by the Division of Forestry in 1927. Out of the 175 plots originally measured, 16 were discarded (see table 20). The 159 actually used are from the following- watersheds given in table 17. TABLE 17 Distribution of Plots by Principal, Watersheds Region and watershed Number of plots Coast Range: 3 3 2 5 Big River Eel River Van Duzen River 3 38 6 Mad River 12 23 Trinity River 38 Klamath River Sierra Nevada Mountains: 5 14 Yuba River Feather River 5 2 Total 159 The composition of the plots by basal areas of the various species included is shown in table 18. TABLE 18 Composition op Basal Area of the Plots Used Species Douglas fir Western yellow pine Oak, laurel and madrone Redwood White fir Sugar pine Incense cedar Grand fir Total Basal area in percentage of total 94.99 1.48 1 04 98 57 45 37 12 100.00 26 University of California — Experiment Station The distribution of the plots by site and age classes is given in table 19. In this table, site index is defined as the height of the average dominant and codominant at 100 years, as the tables were first constructed on site index so denned for purposes of comparison with yields of Douglas fir in Oregon and Washington. TABLE 19 Distribution of Plots by Site and Age Classes Age in Site index —height in feet of the average dominant and codominant tree at 100 years years 75-84 85-94 95- 104 105- 114 115- 124 125- 134 135- 144 145- 154 155- 161 165- 174 175- 184 185- 194 195- 204 205- 214 Total 25- 34 35- 44 1 1 3 10 5 1 2 8 2 1 4 2 10 2 1 5 6 7 2 2 1 8 8 45- 54 1 1 4 12 55- 64 . 4 1 5 1 7 2 5 2 7 4 5 2 2 59 65- 74 39 75- 84 . 2 85- 94 95-104 1 3 105-114 1 5 6 1 12 115-124 125-134 7 3 1 11 1 135-144 145-154 2 2 155-164 165-174 11 1 1 12 9 4 1 Total 5 6 6 8 24 24 25 24 159 20 30 /OO //O 720 /JO /<?0 /50 /60 /70 JO 60 70 60 90 Age /n c/eors Fig. 14. — Comparison of heights of average dominant and codominant tree in the California plots with the height curve for Oregon and Washington stands of the same average site index. Bul, 491] Yield, Stand, and Volume Tables for Douglas Fir 27 Comparison of the California Sample Plots with Yield Tables for Douglas Fir in Oregon and Washington Yield tables for Douglas fir in Oregon and Washington define site index as the height of the average dominant and codominant tree at 100 years. In order, therefore, to compare the values of the California sample plots with the Oregon- Washington tables, each California plot was assigned a site index number as defined for the tables of the northern material. That the latter 's height growth curve for the average dominant and codominant, on which site index is based, fits the California data is shown in figure 14. Then the values of each California plot were compared with the Oregon-Washington yield tables and the percentages of the former to the latter were arranged O 20 'fO 60 80 /OO /20 /<K) /60 /60 200 220 ' 210 260 280 300 Qosot ere& //? per cent of Oregon- IVosn/ngton g/e/ct tables 20 10 60 60 /OO /20 /1V /60 Z&O 20O 220 210 260 260 300 Number of trees /n p?r cent of Oregon -IVosh/hgton g/e/ct tob/es Fig. 15. — Frequency distribution of the California sample plots in per cent of the Oregon-Washington yield tables by basal area and by number of trees. 28 University of California — Experiment Station in a frequency array by basal areas and by numbers of trees. Figure 15 shows these dispersions graphically. The comparison of the means for the original 175 plots are as follows : By basal area, + 33.2% ± 2.65% By number of trees to the acre, — 4.3% ± 2.49% Were the means of the California plots by both basal area and number of trees either higher or lower than the Oregon-Washington tables by about the same amount, one might doubt the validity of the comparison, as the differences might be due to different conceptions as to what constitutes normal stocking, on the part of those who originally laid out and measured the plots in the two regions. But as the basal area of the California material is 33 per cent higher, and the number of trees 4 per cent lower, this can hardly be the case. Rejection of Abnormal Plots The rejection of abnormal plots is based on the above comparison. Those which deviated by about two standard deviations from the mean difference of the California plots were checked over for explanation of their abnormal values. As the explanation was seldom evident from the measurements taken or from the plot description, nearly all were rejected. Table 20 summarizes the rejected plots. TABLE 20 California Plots Rejected as Abnormal Age, years Site index Per cent of difference from corresponding values in Oregon- Washington yield tables Basis for rejection By basal area By number of trees 63 71 + 55 + 62 Too many incense cedar trees 67 98 + 45 + 61 Too many incense cedar trees 72 109 + 84 +107 Basal area and number of trees too high 67 122 + 95 + 80 Basal area and number of trees too high 111 125 +147 + 76 Basal area and number of trees too high 27 130 +126 - 8 Basal area too high 27 132 +152 + 9 Basal area too high 168 146 + 44 - 26 Basal area too high 91 148 + 74 + 24 Basal area too high 27 151 +125 - 20 Basal area too high 45 158 +173 + 53 Basal area and number trees too high 45 156 +139 + 45 Basal area and number trees too high 168 150 + 80 + 11 Basal area too high 45 171 + 83 + 88 Too many redwood sprouts and tan oak trees 45 178 + 53 + 139 Basal area and number trees too high 33 200 + 99 +125 Too many redwood sprouts Bul. 491] Yield, Stand, and Volume Tables for Douglas Fir 29 The means of the remaining 159 plots are as follows : Basal area: + 25.9 ± 1.97% Number of trees: — 9.6 ± 1.97% Obviously these figures cannot be accepted as due to chance fluctua- tion. There must be differences in Douglas fir stand characteristics between the southern and central part of its range on the Pacific slope. Construction of the Yield Tables on the 100- Year Site Index Rather than correlate the percentage deviations of basal area, num- ber of trees, and of other growth units with age and site, the original units are correlated directly with age and site on the 100-year site index and later transferred to the 50-year site index to conform with site as defined for other California species. Basal Area, Number of Trees, and Cubic Volume. — Plot values on the acre basis for these variables were correlated with age and site by comparing them with the multiple linear regression equation, and, by a series of successive estimates, converting the net regression lines for age and for site index as well as the relationship between actual and estimated values, to curvilinear forms where necessary. The calculation of the correlation, measured by the correlation index, is analogous to the Pearsonian correlation ratio : CI ^m in which CI = correlation index <j est = the standard error of estimate; the standard deviation of the de- pendent (y) variable measured from the regression line or curve. o- J/ = the standard deviation of the dependent variable. The term-^ measures the percentage dispersion of the dependent ffy variable due to factors other than the independent variables — in this case, age and site index — considered ; that is, it measures the extent of the independence of the relationship. The numerical value of the correlation index and of the standard error of estimate give the best idea of the association of a particular dependent variable with age and site index. These are : For basal area : <j est = 34.8 sq. ft. ; CI = .845 For no. of trees: cr es< = .116 log trees; CI = .909 For cu. volume : <r e8t = 1930 cu. ft. ; CI = .880 30 University of California — Experiment Station Average Diameter Breast High. — This is the diameter in inches of the tree of average basal area. It varies as the square root of the total basal area divided by the number of trees. If the curves for these variables are accurate, it may be calculated directly from them. This was accordingly tried, giving the average diameter breast high of the yield tables. A check on the work is afforded by the relationship, »(l-:)=™[f; ■(§-:)'] in which BA =■ the total basal area, T = the number of trees, D s= average diameter breast high, and subscripts a and t refer to actual and tabular values respectively. The basal area of each plot in per cent of its tabular basal area was subjected to this equation with the following results: Mean = 100.38% ; standard deviation = 1.72% showing a satisfactory check. Height of Average Tree. — This was arrived at through the relation- ship of the ratio of height of average tree to height of average dominant and codominant with average diameter (fig. 16). •k £- too •^ 90 /s 3' 12 "t% 14 -*® ^r \' -3 1 tl— /H i — H ioy ? + ^0 X4C fi: O 2 4 6 8 10 /2 /4 /& t8 20 22 24 26 28 30 32 34 36 Z Aver&oe- cf /a meter breosf high Fig. 16. — Ratio of height of average tree to height of average dominant and codominant tree for average diameter. Volume in Board Feet. — This is based on the correlation of the ratio of board feet to a cubic foot, with the average diameter (fig. 17). The curved ratio applied to cubic volume gives board foot volume. Bul. 491] Yield, Stand, and Volume Tables for Douglas Fir 31 -0 * / MS ^JZ , 15 ' \ 20 "T'r ^^T 'zZ- ^ s ** >< **» ^ 1 ®Jf /^3I // t V 7 /' 10 i aa a<? ad ae jo 3a 3<? 36 Average diameter breasf high Fig. 17. — Relation of the number of board feet per cubic foot to average diameter. Site Index Translated to Height of Average Dominant at 50 Years In order to change the basis of the yield tables from height of average dominant and codominant at 100 years to height of average dominant at 50 years, the latter site index was plotted over the former (fig. 18) and the final tables re-read accordingly. ML **f GjS 4f- -I S /2 ° ft Co S*Z9 £ £ p? 6x/ -7 +, 4 O 20 <?0 60 SO /OO /20 i<90 /60 /80 200 22.0 2<90 Site index ~ height of average c/om/nanf ond codom/nonh of /oo years Fig. 18. — Relation of site index based on the height of the average dominant tree at 50 years to site index based on the height of the average dominant and codominant tree at 100 years. 32 University of California — Experiment Station Construction of the Stand Tables The distribution of trees by diameter class in a stand forms a fre- quency series which may be analyzed and graduated into a frequency curve when four constants are known — (1) the mean diameter, (2) the standard deviation, (3) the coefficient of asymmetry, (4) the coeffi- cient of excess. These were computed for each of the 159 sample plots. Average Diameter, Mean Diameter and Standard Deviation. — Average diameter, mean diameter and standard deviation are tied together in any one stand by the relationship, o* = A dhl ?-M dbh * in which o- = the standard deviation of diameter distribution, Adbh = the diameter of average basal area, Mdbh = the mean of the diameters breast high. As these three constants were computed independently for each plot, their relationship was checked as follows: / * / / <0 ■£ V / r> ■§ .^ J / r A- ? ome fer A y>+s F a / J™ / < < \ r -/ ^az °- 5j r o 8 °°u lo ^Stor ic/aro ' ' c/ev iaf/o/ 7 sy &+ p& p-r* ■O/O 06 ""^5 ~~**35 p < j 1 / i 5 z J 2 f 3 -< j r— J tf o ' <*5 Averoqe dto meter- breast high Fig*. 19. — Relation of mean diameter and standard deviation to average diameter. Bul,. 491] Yield, Stand, and Volume Tables for Douglas Fir 33 1. The plots were sorted into classes according to the squares of their average diameters using class intervals of 50 square inches. 2. For each plot within the respective classes, were tallied the squares of its average diameter, of its mean diameter, and of its standard deviation. Adding the sums of the squares of mean diameter and of standard deviation, and subtracting this total from the sums of squares of average diameter left an aggregate difference of 15 hundredths of one per cent. •hlU r § y 2 ,17. i, • t3+^ -rj>* ">A3 -" *fc* **<& / -+'// > A **?26 Vj^ , 4s 2 ? 6 8/0/2/^/6 /6 20 22 2f- 26 28 30 32 3* 36 36 Fig. 20. — Relation of the coefficient of asymmetry to mean diameter. 3. Within each class interval were plotted the square root of the average of the mean diameters squared, and of the average of the standard deviations squared, over the square root of the average of the average diameters squared (fig. 19). Straight lines were fitted to these points so that M dbt ?+o* = A dbt ? Asymmetry and Excess. — The coefficient of asymmetry (/?..) and the coefficient of excess (/? 4 ) of the plots were correlated with mean diameter (figs. 20 and 21). Starting with average diameter of a site-age class from table 5, its mean diameter and standard deviation were read from figure 19, and, for the indicated mean diameter, its coefficient of asymmetry and 34 University of California — Experiment Station 5" \ y \ —01 \ \ 2 3 ^ 02 /6 ~ ^ 1 \ 1 1 / N / ' . if / V 1 ' 1 //V" ■-la _ +'„ \ 1 1 \ 1 \J v N 1 1 •s / o a -f 6 e to /2 ./# /6 /e ao 22 a<f ^<5 ae jo sa 1 j-? jc js ffeon d/'ornef-er- breosf /?/gh Fig. 21. — Relation of the coefficient of excess to mean diameter. of excess taken from figures 20 and 21 ; from these parameters, with the aid of Charlier's Tables, 13 the theoretical frequencies were calculated. 13 Charlier's "Type A" frequency curve has the form AT r in which F(x) = frequency of x (in this case frequency per unit of one-half standard deviation measured from mean diameter). N = total frequency. <r = standard deviation. 1 -x 2 <Mz)=- 03 (x) V2T 2 <* 3 0o dx* 4 (z)=^0o dx* These are tabulated for unit frequency with x in terms of standard deviation in Charlier. Coefficient of asymmetry, &= -— 3 („ 3 = the 3rd moment measured from the mean). Coefficient of excess, fo^f ~ 4 - 3 ) ("4 = the 4th moment measured from the mean, Bul. 491] Yield, Stand, and Volume Tables for Douglas Fir 35 VOLUME TABLES Basic Data From 10 to 50 taper measurements were taken on each of eight of the yield study sample plots, two of the plots on cut-over lands of the Union Lumber Company, Mendocino County, two on holdings of the Casper Lumber Company, Mendocino County, and four on the Trinity National Forest in Trinity County. Diameters were measured along the stem of each felled tree out- side and inside bark at breast-height, at each tenth of length above breast-height, at each fifth of length from the lowest tenth downwards and at 1 per cent of total height from the ground. TABLE 21 Basic Data op the Volume Tables Plot County Trees measured Plot age Site index* Ft. Bragg No. 1 42 10 25 42 25 25 48 50 33 33 45 45 68 68 72 111 200 Ft. Bragg No. 2 210 Casper No. 2 178 Casper No. 3 171 Minersville No. 3 Trinity 93 Trinity 90 Minersville No. 14 Trinity 109 South Fork Trinity River No. 16 Trinity 143 * Height of average dominant and codominant at 100 years. Table 21 shows the number of trees by plots and the range in age and site of the data. Each tree was plotted on cross-section paper and its cubic volume computed as the sum of the sectional volumes, each by the Smalian formula. The section lengths were in per cent of total height starting with the stump of 1 per cent, the second section of 3 per cent, the third of 6 per cent, and the remaining nine sections each having length of 10 per cent of tree 's total height. Comparison with Douglas Fir Volume Tables for Oregon and Washington It would only make for confusion to construct volume tables for a particular region when tables for the same species based on data of another region may apply. As there is no readily observable difference 36 University of California — Experiment Station between the forms of Douglas fir in California as against Oregon and Washington, the volumes of the California data were checked against the cubic volume table for immature Douglas fir in Oregon and Washington. 14 The volumes of the tree data basic to the latter table were, how- ever, computed as of different sectional lengths than those noted above for the California trees. Stumps of 1% feet were used and all other sections, regardless of tree's size, were cubed in 10-foot lengths. In order to ascertain what differences in volume result from the two methods of calculation, the trees of Minersville Plot No. 14 were cubed by both methods. It was found that for constant height, both methods gave the same results independent of diameter; but for con- stant diameter, volume of trees less than about 50 feet in total height averaged 6 per cent higher when cubed by the method used for the California data, though the calculated volumes of taller trees were independent of the method of computation. However, only 19 out of the 267, or 7 per cent of the trees of all the plots are less than 55 feet tall ; so that the difference in method should carry little weight in explaining any difference between the actual volumes and those tabulated for the species in Oregon and Washington. Following are the results of the check of the California trees against the Oregon-Washington volume tables : Number of trees 267 Aggregate difference — 2.4 per cent Mean difference — 6.2 per cent Standard error of estimate 13.4 per cent Now if the California trees of all sizes have consistently greater or less taper than the Oregon-Washington trees, there should be no cor- relation between the per cent deviation and tree size. In other words, a blanket correction factor might be applied to the table to arrive at true average volume. This, however, is not the case. The multiple correlation coefficient between per cent deviation and diameter and height was found to be r 123 = .485 ± .034 in which subscript , = per cent deviation of the California volumes, 2 = diameter at breast-height, „ = Total height. ] 4McArdle, R. E. A set of volume tables for second-growth Douglas fir in western Oregon and Washington. Issued in mimeographed form by the Pacific Northwest Forest Experiment Station, June 10, 1926. Bul,. 491] Yield, Stand, and Volume Tables for Douglas Fir 37 /05 - ■2o +40 +/ +40 - -28 yJ1 ^ II IS' Si" 23* ' +2 +3 ^+< 4 3/^/6 20 24 28 32 36 40 44 48 D/a/7?efer breosf /?/gn /r? //7cf?es /30 \ %)/20 ki +2- ¥ & 1 »+/7 +/S 27 ^4 ^c *2 Cr <V 7 +<= 20 40 60 80 /OO /20 /40 /60 /80 200 Tofo/ fre/Qhf /n feef Fig. 22. — Comparison of the California tree volumes with the Oregon-Washington cubic foot volume table by diameter and by height. 38 University of California — Experiment Station Site index was also included as an independent variable in the trial correlation, but the coefficient was not materially increased thereby. It seems likely that site quality does not affect taper in comparatively young timber. Differences in taper due to site prob- ably become significant in mature timber only, for several volume tables for mature timber in which site quality is one of the important variables are now in use. The volumes of California immature Douglas fir compared to the cubic volume table for Oregon and Washington vary with diameter and with height, as shown in figure 22, in which the deviations of the data from the multiple regression equation are compared with the net regression lines for diameter with average height and for height with average diameter. The differences must be due to one or more of the following mensurational factors which make for systematic differences in volume when the latter is based on diameter at breast-height out- side bark and on total height of tree : a Differences in bark thickness. b Differences in taper near the base of the tree. c Differences in taper in the upper part of the bole. In order to compare taper of the species between the two regions, it is necessary that the basic data be analyzed and compared. For this purpose, the original field data from Oregon and Washington were loaned by the United States Forest Service. 15 Lower Taper and Bark Thickness. — These factors were analyzed in one operation rather than separately because their effect on volume is dependent upon their sums. The taper of a typical timber tree is concave towards its axis from the tip downward until a point is reached, usually within the first tenth of its length from the ground, below which it becomes convex toward its axis. The importance of the lower taper from a volume- determining standpoint lies in the fact that the diameter of the tree is nearly always taken at 4% feet from the ground (breast-height), which may or may not be above the point of taper inflection, depend- ing partly upon the size of the tree and partly upon many other factors difficult of measurement and analysis, and too involved for ready application. It thus happens that the diameter at breast-height is The writer is deeply indebted to Director T. T. Munger of the Pacific Northwest Forest Experiment Station, United States Department of Agricul- ture, for the use of 1600 taper measurements — over 80 per cent of the basic data of the Oregon-Washington volume tables. Bul. 491] Yield, Stand, and Volume Tables for Douglas Fir 39 is not satisfactory for accurate volume determination in conjunction with a volume table. But as it comes at such a handy point, prac- tically all volume tables are based upon it. If the taper inside bark of Douglas fir were the same throughout the upper nine-tenths of its length in its entire range on the Pacific slope, it is evident that trees of the same total height and diameter inside bark at one-tenth height would have the same volume. But if the lower taper and bark thickness differ with latitude, while the upper taper remains the same, their volumes may differ significantly if based on a diameter, outside bark, below the point of inflection, because in one case the diameter measured will be greater than in the other. The following method was used to analyze the effect of bark thick- ness and lower taper of the California Douglas fir on cubic volume as tabulated in the Oregon-Washington volume table : (1) Using the northern tree data, diameter breast high outside bark was correlated with total height, site index and diameter inside bark at one-tenth of total height. The effect of site index was found to be negligible, and was dropped as a variable. (2) The regression which was found to be linear, was put up in the form of an alignment chart, and a new diameter outside bark at breast-height read for the 267 California trees according to their total height and diameters inside bark at one-tenth height, by referring these measurements to the chart. (3) Having assigned to each California tree the diameter at breast-height outside bark which it would have had, had bark thickness and lower taper been the same as that of the northern data, its cubic volume was again checked against the volume table, on the new diameter and total height. The multiple correlation coefficient between per cent deviation of the tree volumes from the tabular for diameter breast high and height was computed to be r 123 = .173 ± .060 a much less significant figure than the correlation based on the original check; but the mean of the per cent deviation = + 4.3% ±0.8%, which is approximately 10 per cent higher than the original check. This indicates that in the lowest tenth of length, the California trees have greater taper, greater bark thickness, or both, than the northern trees, for the greater the ratio of diameter at breast-height outside bark to an upper diameter inside bark, the less becomes volume for a given diameter at breast-height, other factors remaining constant. 40 University of California — Experiment Station 4- 8/2/6 20 2<? 26 32 36 <?0 -?"? Diameter /r?3/o / e horfc of //o heiohf in inches 90 ( 1o//fi or/7/t 7~^ 85 ySVrego ^ Was/?// no/7 ?q/o/ 7 . 60 /D >T0 J> /o Fc >r A\ f era ge L hamt sfer ->C 20 <K) CO SO /OO /20 fi?Q Tbfo/ he/ghf /n feef /60 /SO 200 220 Fig. 23. — Comparison of form quotations of the California trees with the form quotients of the Oregon- Washington trees by diameter inside bark at one-tenth height and by total height. Bui,. 491] Yield, Stand, and Volume Tables for Douglas Fir 41 Upper Taper. — The upper taper of the Douglas fir material was analyzed by comparing form quotients of the trees from the two regions. For this purpose form quotient is defined as the diameter inside bark at one-half total height divided by diameter inside bark at one-tenth total height. It is evident that the higher the form quotient the closer does the bole approach a cylinder in form except near the tip. Figure 23 shows the effect of tree size on form quotient for both the northern and the California trees. It must be concluded that the volume tables for immature Douglas fir in Oregon and Washington — in which the northern foresters have full confidence, as they are based on nearly 2000 trees — do not apply to immature trees of the same species in California, because (1) The California trees have greater basal flare, the tendency of which is less volume for a given diameter at breast-height, (2) The California trees have higher average form quotients with consequent tendency to greater volume. This is in general, however, more than offset by the loss in volume due to basal flare. Therefore, since the average taper of immature Douglas fir in California differs from that of the northern states, tables 14 and 15 were prepared. STATION PUBLICATIONS AVAILABLE FOE FKEE DISTRIBUTION BULLETINS No. 253. 263. 277. 279. 283. 304. 310. 313. 331. 335. 343. 344. 346. 347. 348. 349. 353. 354. 357. 361. 362. 363. 364. 366. 367. 368. 369. 370. 371. 373. 374. 380. 385. 386. 388. 389. 390. 391. 392. 393. 394. 395. 396. 397. 400. 405. 406. 407. Irrigation and Soil Conditions in the Sierra Nevada Foothills, California. Size Grades for Ripe Olives. Sudan Grass. Irrigation of Rice in California. The Olive Insects of California. A Study of the Effects of Freezes on Citrus in California. Plum Pollination. Pruning Young Deciduous Fruit Trees. Phylloxera-resistant stocks. Cocoanut Meal as a Feed for Dairy Cows and Other Livestock. Cheese Pests and Their Control. Cold Storage as an Aid to the Market- ing of Plums, a Progress Report. Almond Pollination. The Control of Red Spiders in Decid- uous Orchards Pruning Young Olive Trees. A Studv of Sidedraft and Tractor Hitches. Bovine Infectious Abortion, and Asso- ciated Diseases of Cattle and New- born Calves. Results of Rice Experiments in 1922. A Self-Mixing Dusting Machine for Applying Dry Insecticides and Fun- gicides. Preliminary Yield Tables for Second Growth Redwood. Dust and the Tractor Engine. The Pruning of Citrus Trees in Cali- fornia. Fungicidal Dusts for the Control of Bunt. Turkish Tobacco Culture, Curing, and Marketing. Methods of Harvesting and Irrigation in Relation to Moldy Walnuts. Bacterial Decomposition of Olives During Pickling. Comparison of Woods for Butter Boxes. Factors Influencing the Development of Internal Browning of the Yellow Newtown. Apple. The Relative Cost of Yarding Small and Large Timber. Pear Pollination. A Survey of Orchard Practices in the Citrus Industry of Southern Cali- fornia. Growth of Eucalyptus in California Plantations. Pollination of the Sweet Cherry. Pruning Bearing Deciduous Fruit Trees. The Principles and Practice of Sun- Drying Fruit. Berseem or Egyptian Clover. Harvesting and PaGking Grapes in California. Machines for Coating Seed Wheat with Copper Carbonate Dust. Fruit Juice Concentrates. Crop Sequences at Davis. I. Cereal Hay Production in California. II. Feeding Trials with Cereal Hays. Bark Diseases of Citrus Trees in Cali- fornia. The Mat Bean, Phaseolus Aconitifolius. Manufacture of Roquefort Type Cheese from Goat's Milk. The Utilization of Surplus Plums. Citrus Culture in Central California. Stationary Spray Plants in California. Yield, Stand, and Volume Tables for White Fir in the California Pine Region. No. 408. 409. 410. 412. 414. 415. 416. 418. 419. 420. 421. 423. 425. 426. 427. 428. 430. 431. 432. 433. 434. 435. 436. 438. 439. 440. 444. 445. 446. 447. 448. 449. 450. 451. 452. 453. 454. Alternaria Rot of Lemons. The Digestibility of Certain Fruit By- products as Determined for Rumi- nants. Part I. Dried Orange Pulp and Raisin Pulp. Factors Influencing the Quality of Fresh Asparagus After it is Harvested. A Study of the Relative Value of Cer- tain Root Crops and Salmon Oil as Sources of Vitamin A for Poultry. Planting and Thinning Distances for Deciduous Fruit Trees. The Tractor on California Farms. Culture of the Oriental Persimmon in California. A Study of Various Rations for Fin- ishing Range Calves as Baby Beeves. Economic Aspects of the Cantaloupe Industry. Rice and Rice By-Products as Feeds for Fattening Swine. Beef Cattle Feeding Trials, 1921-24. Apricots (Series on California Crops and Prices). Apnle Growing in California. Apple Pollination Studies in California. The Value of Orange Pulp for Milk Production. The Relation of Maturity of California Plums to Shipping and Dessert Quality. Range Grasses in California. Raisin By-Products and Bean Screen- ings as Feeds for Fattening Lambs. Some Economic Problems Involved in the Pooling of Fruit. Power Requirements of Electrically Driven Dairy Manufacturing Equip- ment. Investigations on the Use of Fruits in Ice Cream and Ices. The Problem of Securing Closer Rela- tionship between Agricultural Devel- opment and Irrigation Construction. I. The Kadota Fig. II. The Kadota Fig Products'. Grafting Affinities with Special Refer- ence to Plums. The Digestibility of Certain Fruit By- Products as Determined for Rumi- nants. II. Dried Pineapple Pulp, Dried Lemon Pulp, and Dried Olive Pulp. The Feeding Value of Raisins and Dairy By-Products for Growing and Fattening Swine. Series on California Crops and Prices: Beans. Economic Aspects of the Apple In- dustry. The Asparagus Industry in California. A Method of Determining the Clean Weights of Individual Fleeces of Wool. Farmers' Purchase Agreement for Deep Well Pumps. Economic Aspects of the Watermelon Industry. Irrigation Investigations with Field Crops at Davis, and at Delhi, Cali- fornia, 1909-1925. Studies Preliminary to the Establish- ment of a Series of Fertilizer Trials in a Bearing Citrus Grove. Economic Aspects of the Pear Industry. Series on California Crops and Prices: Almonds. Rice Experiments in Sacramento Val- ley, 1922-1927. BULLETINS— (Continued) No. 455. Reclamation of the Fresno Type of Black-Alkali Soil. 456. Yield, Stand and Volume Tables for Red Fir in California. 458. Factors Influencing Percentage Calf Crop in Range Herds. Economic Aspects of the Fresh Plum Industry. Series on California Crops and Prices : Lemons. 461. Series on California Crops and Prices: Economic Aspects of the Beef Cattle Industry. Prune Supply and Price Situation. Drainage in the Sacramento Valley Rice Fields. 459. 460. 462. 464. No. 465. 466. Curly Top Symptoms of the Sugar Beet. The Continuous Can Washer for Dairy Plants. 467. Oat Varieties in California. 468. Sterilization of Dairy Utensils with Humidified Hot Air. 469. The Solar Heater. 470. Maturity Standards for Harvesting Bartlett Pears for Eastern Shipment. 471. The Use of Sulfur Dioxide in Shipping Grapes. 474. Factors Affecting the Cost of Tractor Logging in the California Pine Region. 475. Walnut Supply and Price Situation. CIRCULARS No. 115. Grafting Vinifera Vineyards. 117. The Selection and Cost of a Small Pumping Plant. 127. House Fumigation. 129. The Control of Citrus Insects. 164. Small Fruit Culture in California. 166. The County Farm Bureau. 178. The Packing of Apples in California. 203. Peat as a Manure Substitute. 212. Salvaging Rain-Damaged Prunes. 230. Testing Milk, Cream, and Skim Milk for Butterfat. 232. Harvesting and Handling California Cherries for Eastern Shipment. 239. Harvesting and Handling Apricots and Plums for Eastern Shipment. 240. Harvesting and Handling California Pears for Eastern Shipment. 241. Harvesting and Handling California Peaches for Eastern Shipment. 243. Marmalade Juice and Jelly Juice from Citrus Fruits. 244. Central Wire Bracing for Fruit Trees. 245. Vine Pruning Systems. 248. Some Common Errors in Vine Pruning and Their Remedies. 249. Replacing Missing Vines. 250. Measurement of Irrigation Water on the Farm. 253. Vineyard Plans. 255. Leguminous Plants as Organic Ferti- lizers in California Agriculture. 257. The Small-Seeded Horse Bean (Vicia faba var. minor). 258. Thinning Deciduous Fruits. 259. Pear By-Products. 261. Sewing Grain Sacks. 262. Cabbage Production in California. 263. Tomato Production in California. 265. Plant Disease and Pest Control. 266. Analyzing the Citrus Orchard by Means of Simple Tree Records. No. 269. 270. 276. 277. 278. 279 282. 284. 287. 288. 289. 290. 292. 294. 295. 296. 298. 300. 301. 302. 304. 305. 307. 308. 309. 310. 311. 312. 313. 314. 315. An Orchard Brush Burner. A Farm Septic Tank. Home Canning. Head, Cane, and Cordon Pruning of Vines. Olive Pickling in Mediterranean Countries. The Preparation and Refining of Olive Oil in Southern Europe. Prevention of Insect Attack on Stored Grain. The Almond in California. Potato Production in California. Phylloxera Resistant Vineyards. Oak Fungus in Orchard Trees. The Tangier Pea. Alkali Soils. Propagation of Deciduous Fruits. Growing Head Lettuce in California. Control of the California Ground Squirrel. Possibilities and Limitations of Coop- erative Marketing. Coccidiosis of Chickens. Buckeye Poisoning of the Honey Bee. The Sugar Beet in California. Drainage on the Farm. Liming the Soil. American Foulbrood and Its Control. Cantaloupe Production in California. Fruit Tree and Orchard Judging. The Operation of the Bacteriological Laboratory for Dairy Plants. The Improvement of Quality in Figs. Principles Governing the Choice. Oper- ation and Care of Small Irrigation Pumping Plants. Fruit Juices and Fruit Juice Beverages. Termites and Termite Damage. The Mediterranean and Other Fruit Flies. 8m-4,'30