; GRAY'S SCHOOL AND FIELD BOOK OP BOTANY CONSISTING OF "LESSONS IN BOTANY" AND "FIELD, FOREST, AND GARDEN BOTANY" BOUND IN ONE VOLUME REVISED EDITION PUBLISHERS' PREFACE TO GRAY'S SCHOOL AND FIELD BOOK OF BOTANY. THIS work consists of the "LESSONS IN BOTANY" and the "FIELD, FOREST, AND GARDEN BOTANY," bound together in one complete volume, forming a most popular and comprehensive SCHOOL BOTANY, adapted to beginners and advanced classes, to Agricultural Colleges and Schools, as well as to all other grades in which the science is taught. It is also adapted for use as a handbook to assist in analyzing plants and flowers in field study of botany either by classes or individuals. The book is intended to furnish Botanical Classes and beginners with an easier introduction to the Plants of this country, and a much more compre- hensive work, than the MANUAL. Beginning with the first principles, it progresses by easy stages until the student who is at all diligent is enabled to master the intricacies of the science. It is a Grammar and Dictionai-y of Botany, and comprises the common Herbs, Shrubs, and Trees of the Southern as well as the Northern and Middle States, including the commonly cultivated as well as the native species in fields, gardens, pleasure grounds, or house culture, and even the conservatory plants ordinarily met with. GKKAY'S LESSONS IN BOTANY REVISED EDITION THE ELEMENTS OF BOTANY FOR BEGINNERS AND FOR SCHOOLS BY ASA GRAY NEW YOKK : CINCINNATI : CHICAGO AMERICAN BOOK COMPANY GRAY'S BOTANICAL SERIES Gray's How Plants Grow Gray's How Plants Behave *Gray's Lessons in Botany Gray's Field, Forest, and Garden Botany (Flora only) *Gray's School and Field Book of Botany (Lessons and Flora) Gray's Manual of Botany. (Flora only) *Gray's Lessons and Manual of Botany Gray's Botanical Text-Book I. Gray's Structural Botany II. Goodale's Physiological Botany Coulter's Manual of Botany of the Rocky Mountains Gray and Coulter's Text-Book of Western Botany EDITIONS OF 1901 *Leavitt's Outlines of Botany (Based on Gray's Lessons) *Leavitt's Outlines of Botany with Flora (Outlines and Gray's Field, Forest, and Garden Botany) *Leavitt's Outlines and Gray's Manual Copyright, 1887, by ASA GRAY REV. LESSONS W. P. II PREFACE. THIS volume takes the place of the author's LESSONS IN BOTANY AND VEGETABLE PHYSIOLOGY, published over a quarter of a cen- tury ago. It is constructed on the same lines, and is a kind of new and much revised edition of that successful work. While in some respects more extended, it is also more concise and terse than its predecessor. This should the hetter fit it for its purpose now that competent teachers are common. They may in many cases develop paragraphs into lectures, and fully illustrate points which are barely, but it is hoped clearly, stated. Indeed, even for those without a teacher, it may be that a condensed is better than a diffuse exposition. The book is adapted to the higher schools, " How Plants Grow and Behave " being the " Botany for Young People and Common Schools." It is intended to ground beginners in Structural Botany and the principles of vegetable life, mainly as concerns Flowering or Phanerogamous plants, with which botanical instruction should always begin ; also to be a companion and interpreter to the Man- uals and Floras by which the student threads his flowery way to a clear knowledge of the surrounding vegetable creation. Such a book, like a grammar, must needs abound in technical words, which thus arrayed may seem formidable ; nevertheless, if rightly apprehended, this treatise should teach that the study of bot- any is not the learning of names and terms, but the acquisition of knowledge and ideas. No eifort should be made to com- mit technical terms to memory. Any term used in describing a plant or explaining its structure can be looked up when it is wanted, and that should suffice. On -the other hand, plans of iv PREFACE. structure, types, adaptations, and modifications, once understood, are not readily forgotten ; and they give meaning and interest to the technical terms used in explaining them. In these "Elements" naturally no mention has been made of certain terms and names which recent cryptogam ically-minded botanists, with lack of proportion and just perspective, are en- deavoring to introduce into phanerogamous botany, and which are not needed nor appropriate, even in more advanced works, for the adequate recognition of the ascertained analogies and homologies. As this volume will be the grammar and dictionary to more than one or two Manuals, Floras, etc., the particular directions for pro- cedure which were given in the " First Lessons " are now relegated to those works themselves, which in their new editions will pro- vide the requisite explanations. On the other hand, in view of such extended use, the Glossary at the end of this book has been considerably enlarged. It will be found to include not merely the common terms of botanical description but also many which are unusual or obsolete ; yet any of them may now and then be encoun- tered. Moreover, no small number of the Latin and Greek words which form the whole or part of the commoner specific names are added to this Glossary, some in an Anglicized, others in their Latin form. This may be helpful to students with small Latin and less Greek, in catching the meaning of a botanical name or term. The illustrations in this volume are largely increased in number. They are mostly from the hand of Isaac Sprague. It happens that the title chosen for this book is that of the author's earliest publication, in the year 1836, of which copies are rarely seen ; so that no inconvenience is likely to arise from the present use of the name. ASA GRAY. CAMBRIDGE, MASSACHUSETTS, March, 1887- CONTENTS. PACE SECTION I. INTRODUCTORY 9 SECTION H. FLAX AS A PATTERN PLANT 11 Growth from the Seed, Organs of Vegetation 11 Blossoming, Flower, &c 14 SECTION m. MORPHOLOGY OF SEEDLINGS ....... 16 Germinating Maples 16 Cotyledons thickened, hypogaeous in germination 18 Store of Food external to the Embryo 20 Cotyledons as to number 22 Dicotyledonous and Polycotyledonons 23 Monocotyledonous 24 Simple-stemmed Plants 26 SECTION IV. GROWTH FROM BUDS; BRANCHING . ... 27 Buds, situation and kinds 27 Vigorous vegetation from strong Buds 28 Arrangement of Branches 29 Non-developed, Latent, and Accessory Buds 30 Enumeration of kinds of Buds 31 Definite and Indefinite growth ; Deliquescent and Excurrent . 31 SECTION V. ROOTS 33 Primary and Secondary. Contrast between Stem and Root . 34 Fibrous and Fleshy Roots ; names of kinds 34 Anomalous Roots. Epiphytic and Parasitic Plants .... 36 Duration : Annuals, Biennials, Perennials 37 SECTION VI. STEMS 38 Those above Ground : kinds and modifications .39 Subterranean Stems and Branches 42 Rootstock, 42. Tuber, 44. Corra, 46. Bulb and Bulblete . . 46 Consolidated Vegetation 47 SECTION VII. LEAVES 49 1. LEAVES AS FOLIAGE 49 Parts and Venation 60 Forms as to general outline 62 As to apex and particular outline 63 CONTENTS, As to lobing or division 56 Compound, Perfoliate, and Equitant Leaves 67 With no distinction of Petiole and Blade, Phyllodia, Ac. . . . 61 2. LEAVES OF SPECIAL CONFORMATION AND USB .... 32 Leaves for storage 62 Leaves as bud-scales, 63, Spines, 64, and for Climbing ... 64 Pitchers, 64, and Fly-traps .... 65 1 8. STIPULES 66 4. THE ARRANGEMENT OF LEAVES 67 Phyllotaxy, 67, of Alternate Leaves 60 Of Opposite and Whorled Leaves 71 Venation or Praefoliation 71 vra. FLOWERS 72 I 1. POSITION AND ARRANGEMENT, INFLORESCENCE .... 73 Raceme, 73, Corymb, Umbel, Spike, Head 74 Spadix, Catkin, or Ament 76 Panicle: Determinate Inflorescence 76 Cyme, Fascicle, Glomerule, Scorpioid or Helicoid Cymes . . 77 Mixed Inflorescence 78 2. PARTS OB ORGANS OF THE FLOWER 79 Floral Envelopes : Perianth, Calyx, Corolla 79 Essential Organs : Stamen, Pistil 80 Torus or Receptacle r 81 3. PLAN OF THE FLOWER 81 When perfect, complete, regular, or symmetrical 81 Numerical Plan and Alternation of Organs 82 Flowers are altered branches 83 4. MODIFICATIONS OF THE TYPE 86 Unisexual or diclinous 86 Incomplete, Irregular, and Ur.sym metrical 86 Flowers with Multiplication of Parts 88 Flowers with Union of Part* : Coalescence 88 Regular Forms, 89, Irregular Forms 90 Papilionaceous, 91, Labiate, 92, and Ligulate Corollas .... 93 Adnation or Consolidation 94 Position of Flower or of its Parts 96 6. ARRANGEMENT OF PARTS IN THE BUD 97 ^Estivation or Praefloration, its kinds 67 SECTION IX. STAMENS IN PARTICULAR 98 Androecium, 98, Insertion, Relation, &c 99 Anther and Filament. Pollen 101 SECTION X. PISTILS IN PARTICULAR 106 | 1. ANGIOSPERMOCS OR ORDINARY GYNOECIUM 106 Parts of a complete Pistil 106 Carpels. Simple Pistil 106 CONTENTS. Vii Compound Pistil with Cells and Axile Placenta 107 One-celled with Free Central Placenta 108 One-celled with Parietal Placenta 108 2. GTMNOSPERMOU8 GTN