PHYSIOLOGY
BRIEFER COURSE
GOLTON
BIOLOGY
LIBRARY
G
.wt
X-RAY PHOTOGRAPH OF HAND
SHOWING SHOT CARRIED FOR TWENTY YEARS
(From Recreation, by permission of G. O. Shields.)
BRIEFER COURSE
PHYSIOLOGY
ILLUSTRATED BY EXPERIMENT
BY
BUEL P. COLTON, A.M.
AUTHOR OF " PHYSIOLOGY, EXPERIMENTAL AND DESCRIPTIVE
"PRACTICAL ZOOLOGY"; AND PROFESSOR OF NATURAL
SCIENCE IN THE ILLINOIS STATE NORMAL
UNIVERSITY
BOSTON, U.S.A.
D. C. HEATH & CO., PUBLISHERS
1900
BIOLOGY
LIBRARY
G
COPYRIGHT, 1899,
BY BUEL P. COLTON.
Nortooob
J. S. Gushing & Co. - Berwick & Smith
Norwood Mass. U.S.A.
PREFACE.
THE author's " Experimental and Descriptive Physiol-
ogy" has been adopted by a large number of schools and
colleges. But there are many schools in which, owing to
the youth of the pupils, the shortness of the time allotted to
the subject, or the meagerness of laboratory facilities, such
a rigorous course cannot be taken. For such schools this
simpler book is written. While it contains considerably
less experiment and dissection than the larger book, it is
still based upon experimental work. No teaching of physi-
. ology is worthy of the name unless it rests upon experi-
ment, observation, and dissection. The ridiculous answers
of the pupil who has learned mere "book physiology"
furnish the standard jest of the educational journal. Try-
ing to teach physiology without experiment is not only in
opposition to modern views of pedagogy and psychology,
but it is equally at variance with the common sense of the
business man's view. Such teaching is a mere mummery
of words it teaches neither how to know nor to do.
In fitting this work for the less mature mind, special
attention has been paid to conciseness and brevity of state-
ment and to clearness of exposition. Sentences and para-
iii
735021
iv PR El- ACE.
graphs have been made short; chapters are short, with
definite summaries appended.
Function, rather than structure, has been made promi-
nent. Only so much of anatomy as is really needed to
understand the working of the organs has been introduced.
The experimental work and directions for dissection, as
well as some of the more difficult points, have been put in
smaller type, so they may more readily be omitted where
it is not possible to complete all the work in the given
time.
Although hygiene has been given a prominent place,
yet it may be claimed that when the pupil is well grounded
in the functions of the different organs, from observational
and experimental work, many of the rules of hygiene will
readily occur to him as natural inferences. When other
rules for the preservation of health, which might not occur
to him, are suggested, he will see their significance because
he understands the underlying principle ; and he not only
can, but will, obey the rule better because he sees reason
in it, and does not follow it blindly as an arbitrary law
thrust upon him. Questions are given, at the end of each
chapter, to test the pupil's knowledge of principles by
application to new cases.
Some of the more desirable reforms in nomenclature have
been adopted ; among these are the use of the terms ante-
rior, meaning toward the head ; posterior, in the opposite
direction ; dorsal, toward the back ; ventral, toward the
region of the belly. These terms, used instead of " up "
PREFACE. V
and "down," "front" and "back," will do away with
much confusion, especially since we are obliged to use the
structure of horizontal-bodied animals to illustrate human
anatomy. Many Latin terms, such as "vena cava" should
be replaced by English, as caval vein. Postcaval vein and
precaval vein are easier and better than " vena cava infe-
rior " and "vena cava superior." In many cases the
English plural may well replace the Greek or Latin form,
as ganglion, plural ganglions. Blood tube is better than
"bloodvessel." The best authorities say spinal bulb in-
stead of the long "medulla oblongata." Food tube is
simpler than " alimentary canal," especially as the tube is
not canal-like. The rib-bearing vertebras are thoracic, and
are no more "dorsal" than the other parts of the spinal
column.
Effort has been made to lay stress on the more impor-
tant topics, and the skeleton is relegated to a subsidiary
place, as a knowledge of it has so little to do with practical
hygiene. The heart and the stomach receive full treat-
ment, while matters of such slight importance as the hair
and nails are briefly dismissed.
The order of topics is the result of long experience.
For many years the author has sought to find the most
natural sequence of subjects, so that, as the work pro-
gressed, the pupil would find the way best prepared for
him. Without claiming that this is the best sequence, the
writer is sure that it is the " path of least resistance."
The subject of Alcohol has been treated in full com-
vi PREFACE.
pliance with the law. Copious quotations have been taken
from the best authorities on this subject. The same
high-grade illustrations have been used that brought such
favorable comment on the earlier work.
This briefer edition has, too, the full benefit of the
criticism of the eminent authorities whose names are listed
in the larger work.
TO THE TEACHER.
For any practical work in physiology it is very desirable
to have a room furnished with tables and supplied with
water.
Each pupil should make full notes and drawings of the
work done and the organs studied and dissected. Only by
so doing will he firmly fix and retain what he gathers from
day to day.
In the larger work by the author are many experiments
and dissections given in full which are here omitted in
order to present a briefer course. In the larger work there
is also given a list of books which are most helpful in
teaching physiology.
CONTENTS.
CHAPTER I.
PAGE
INTRODUCTION l
Health. Care of the Body. Hygiene. Physiology. Organ. Func-
tion. Anatomy. Tissues. Cells. Physiological Division of Labor.
CHAPTER II.
MOTION 7
Motion Necessary to Life. Experiments with our Muscles. Action
of Muscle. Structure of Muscle. Connective Tissue. Laws of
Muscle Action. Flexors and Extensors. Symmetry. Muscles and
Bones. Levers. Locomotion. Uses of Bones.
CHAPTER III.
THE GENERAL FUNCTIONS OF THE NERVOUS SYSTEM SENSATION
AND. MOTION 2 4
Muscles controlled by Nerves. Voluntary and Involuntary Motion.
The Spinal Cord. The Spinal Nerves. Structure of Nerves.
Function of Nerves. Structure of the Spinal Cord. Ganglions.
Reflex Action of the Spinal Cord. Reflex Action of the Spinal
Cord in the Frog. Function of the Nerve Roots. Importance of
Reflex Action. Inhibition. Nature of a Nervous Impulse. Har-
mony in Muscle Action. Nerves depend on Blood Supply.
CHAPTER IV.
CIRCULATION OF THE BLOOD 39
The Blood and its Work. The Rate of the Heart Beat. Position
and Size of the Heart. The Valves of the Heart. The Blood
Tubes Connected with the Heart. The Action of the Heart. Work
and Rest of the Heart. Action of the Large Arteries. Variation
in Blood Supply. Plain Muscle Fibers in the Walls of the Arteries.
vii
viii CONTENTS.
I'AGK
Circulation of Blood in the \Yel> of the Frog's Foot. Blood Mow in
the Capillaries. The Veins. The Valves in the Veins. F.t'fect of
Pressure on the Veins. Rate of Blood Flow. Nourishment of the
Walls of the Heart. Effect of Gravity on Circulation.
CHAPTER V.
CONTROL OF CIRCULATION. THE BLOOD AND THE LYMPH .... 64
The Effect of Emotions on Circulation. Rhythmic Action of the
Heart. Nerve Control of the Heart. Sympathetic Nervous System.
The Vagus Nerve. Inhibition. Vaso-motor Nerves. Blushing.
Regulation of the Size of the Arteries. Effect of Exercise on the
Size of the Arteries. Effects of Alcohol on Circulation. The
Bloocl. The Corpuscles of the Blood. The Plasma. Hemoglobin.
Coagulation of Blood. Fibrin. Amount of Blood. Distribution of
Blood. The Lymph Spaces. Lymph Tubes. Lymphatic Glands.
Flow of Lymph. Massage. Transfusion of Blood.
CHAPTER VI.
RESPIRATION 84
The Close Relation between Circulation and Respiration. Organs
of Respiration. Structure of the Lungs. The Windpipe. Cilia.
The Pleura. The Diaphragm. Action of the Diaphragm. Move-
ments of Respiration. Forces of Inspiration. Resistances to Inspi-
ration. Elastic Reactions of Expiration. Forced Respiration. Rate
of Respiration. Modifications of Respiration. Lung Capacity.
Hygiene of Respiration. Breathing through the Mouth. Control
of Respiration. Chemistry of Respiration. Composition of the
Air. Exchanges between the Air and the Blood in the Lungs.
Oxyhemoglobin. The Gases in the Blood. Production of Heat
and Motion in the Body. Oxidation of Live Tissues. Body and
Locomotive Compared. Storage of Oxygen in the Tissues. Re-
breathing Air.
CHAPTER VII.
VENTILATION AND HEATING. DIST AND BACTERIA 114
Need of Ventilation. Grates. Principles of Ventilation. Stoves.
Furnaces. Foul-air Shafts and Fans. Steam and Hot-water
Heating. Direct and Indirect Heating. Dead Dust. Sources of
Dust. Live Dust. Consumption. Disease Germs. Bacteria. How
to avoid Dust. Sweeping. Contagious Diseases. Putrefaction.
Preservation of Foods. Need of Removal of Waste.
CONTENTS. ix
CHAPTER VIII.
PAGE
EXCRETION 130
The Skin throws off Perspiration. The Structure of the Skin. The
Epidermis. Color of the Skin. The Dermis. Sweat Glands. Es-
sentials of a Gland. Blood Supply of Glands. Oil Glands.
Composition of Sweat. Amount of Perspiration. Functions of the
Skin. Regulation of Bodily Temperature by the Skin. Distribu-
tion of Heat in the Body. Regulation of Bodily Temperature by
Food and Clothing. The Kidneys. Work of the Kidneys. Rela-
tion of the Skin and Kidneys.
CHAPTER IX.
Foons AND COOKING 144
Necessity of Food. Food Defined. Foodstuffs. Proteids. Impor-
tance of Proteids. Meat. Fish. Eggs. Milk. Cheese. Vege-
table Proteids. Carbohydrates. Grains. Wheat. Corn. Rice.
Oats. Potatoes. Vegetables. Fruits. Water. Impurities in
Water. Typhoid Fever. Ice-water. Boiling Water. Salts.
Necessity of a Mixed Diet. Effects of Cold on Appetite for Fats.
Vegetarians. Tea. Coffee. Beef Tea. Cooking. Soups.
CHAPTER X.
THE DIGESTIVE SYSTEM 159
The Object of Food. The Digestive Tube. Organs of Digestion.
The Mouth. The Teeth Kinds, Structure, Arrangement. Care
of the Teeth. Salivary Glands. Action of Salivary Glands. Saliva
and its Uses. Mucus and Mucous Glands. The Pharynx. Swal-
lowing. The Gullet. The Structure of the Stomach. Gastric
Glands. Blood Supply of the Stomach. Stomach Digestion.
Churning Action of the Stomach. Time of Stomach Digestion.
Chyme. Absorption from the Stomach. The Intestine. The
Liver. The Pancreas. Bile. Pancreatic Juice. The Portal Circu-
lation. Functions of the Bile. Work of the Pancreatic Juice.
Intestinal Juice.
CHAPTER XL
ABSORPTION DIGESTION COMPLETED 181
Absorption. Villi. Routes of Different Foods after Absorption. Dif-
fusion and Osmosis. Absorption a Vital Process. The Lacteals
and the Lymphatics. Outline of Digestion. The Colon. Work
of the Large Intestine. Constipation. Laxative and Constipating
X CONTENTS.
PAGE
Foods. Hygiene of Digestion. Deliberation in Eating. Thorough
Mastication. Fleets of Repose on Digestion. Conversation at
Meals. Value of Soups and Desserts. Hot Drink at Meals. Time
of Meals. Eating between Meals. Amount of Food Needed.
Errors in Diet.
CHAPTER XII.
NUTRITION 195
Ledger Account of the Body and its Organs. Blood a Mixture of
Good and Bad. Action of Diseased Kidneys. Blood Stream like
Water Pipes and Sewer Combined. A Living Eddy. Importance
of Renewal of Blood and Lymph. Fat as a Tissue. Hibernation.
Respiration and Oxidation of Candle. Glycogen. Muscular Exer-
tion and Excretion of Urea. Metabolism. Indestructibility of
Matter. Indestructibility of Force. Utilization of Energy in the
Body and in Machines. Correlation and Conservation of Energy.
CHAPTER XIII.
ALCOHOL 208
Alcohol and Crime. Alcohol and Energy. Alcohol and Heat. Alco-
hol and Muscular Energy. Arctic Explorers. Alcohol and Train-
ing. Stimulants. Narcotics. Temperance Drinks. Testimony of
Physiologists. Testimony of Army Officers. Testimony of a Natu-
ralist. Physiological Effects of Alcohol.
CHAPTER XIV.
EXERCISE AND BATHING 226
How Exercise is Beneficial. Exercise for General Health. Nature's
Rewards and Punishments. Exercise prolongs Life. Choice of
Exercise. Games of School Children. Tennis. Baseball and
Football. Boxing. Bicycling. Exercise for Middle-aged Men.
Taking Cold. Diarrhea. Bathing. Cold Baths. Bath Mits. Time
for Bathing. Warm Baths vs. Cold Baths. Exercise Arterial Mus-
cles. Habit of Cold Bathing acquired gradually.
CHAPTER XV.
THE BRAIN 235
The Coverings of the Brain. Parts of the Brain. The Cerebrum.
The Cerebellum. ' The Spinal Bulb. Brain of a Cat or Rabbit.
Cranial Nerves and their Functions. Hemispheres of the Cerebrum.
Brain Convolutions and Intelligence. Gray and White Matter of
the Brain. Neuroglia. Functions of the Cerebrum. Pigeon
with Cerebrum Removed. Functions of Cerebral Cortex. Center
CONTENTS. xi
PAGE
of Sensations itself Insensible. Crossed Control of the Body. Lo-
cation of Brain Functions. Left Hemisphere better Developed.
Location of Sensation Centers. Brain Work and Brain Rest.
Sleeplessness. Fatigue. Control of Mind. Habit of Resting the
Brain. Nervous Tissues least Affected by Starvation. Blood Supply
of the Brain. Fainting. Apoplexy. Meningitis. The Water
Cushion of the Brain. Relative Activity of Gray and White Matter.
CHAPTER XVI.
EFFECTS OF ALCOHOL ON THE NERVOUS SYSTEM 250
The Chief Effect of Alcohol is on the Nervous System. Inebriety re-"
garded as a Disease. Moral Deterioration produced by Alcohol.
Narcotics. Opium. Cocaine. Chloral Hydrate. Chloroform.
Tobacco. Cigarette Smoking.
CHAPTER XVII.
GENERAL CONSIDERATIONS CONCERNING THE NERVOUS SYSTEM . . . 261
Nerve Stimuli. Kinds of Nerve Stimuli. Essential Similarity of All
Nerve Fibers. Relation of Stimulus and Sensation. Reaction
Time. Reflex Action. Connection of Brain 'Centers. Nature of
Sensation. Subjective Sensations. The Relative Nature of Sensa-
tion. Induction Currents used in Physiological Experiments.
Dreams. Lingering Effect of Sensations. Habits are Acquired
Reflex Actions. Usefulness of Resting. Nervous System vs. Tele-
graph System. Efferent Currents. Afferent Currents.
CHAPTER XVIII.
THE GENERAL SENSES 271
The Body a Collection of Organs. Influence from the External World.
Classification of the Senses. General Sensations and Special Senses.
The Muscular Sense. Importance of Muscular Sense. Dependence
of Sight on Muscular Sense and Touch. Pain. Pain a General
Sense. Extent of Pain. Use of Pain. Hunger and Thirst.
CHAPTER XIX.
THE SPECIAL SENSES TOUCH AND TEMPERATURE SENSE .... 278
What we learn by Touching Objects. Cutaneous Sensations. Nerve
Endings in the Skin. Touch Corpuscles. Touch the most General
of the Senses. Pressure Sense. Local Sign. Test by Compass
Points. Reference of Sensation to the Region of the Nerve End-
ings. Temperature Sense.
xil CONTEWS.
CHAPTKR XX.
PAGE
THE SENSE OF SIGHT 285
Protection of the Eye. The Lacrymal Secretion. External Parts of
the Eye. The Conjunctiva. Muscles of the Eyeball. Movements
of the Eye. Coats of the Eye. The Sclerotic Coat. The Choroid
Coat. The Retina. The Cornea. The Iris. The Pupil. Regula-
tion of the Amount of Light admitted into the Eye. The Refract-
ing Media of the Eye. The Aqueous Humor. The Vitreous Humor.
The Crystalline Lens. The Lens Capsule. The Hyaloid Mem-
brane. The Ciliary Muscle. Inversion of the Image. Adjustment
for Distance. Action of the Ciliary Muscle. Defects of Eyesight.
Structure of the Retina. Importance of the Retina. The Blind
Spot. The Optic Nerve not Sensitive to Light. Sympathy between
the Eyes. Pain in the Eyes. Color Sensations. Color Blindness.
Stereoscopic Vision. After-images. Care of the Eyes.
CHAPTER XXI.
TASTE, SMELL, AND HEARING ^01
Uses of the Sense of Taste. The Papilla;. Nerve Supply of the
Tongue. Solution Necessary for Tasting. Flavors. Effect of
Temperature on Taste. The Sense of Smell. Why we Sniff. The
Sense of Bearing. The External Ear. The Tympanum. The
Middle Ear. The Eustachian Tube. The Internal Ear. The Pro-
duction of Sound. The Equilibrium Sense. The Care of the Far.
The Use of the Ears.
CHAPTER XXII.
THE VOICE 309
The Ear and the Voice. What we can learn from our own Throats.
The Vocal Cords. Reinforcement of Sound. Pitch and Voice.
Voice and Speech. Vowels and Consonants. Difference between
Voices. Hoarseness. Whispering. Culture of the Voice.
CHAPTER XXIII.
ACCIDENTS. WHAT TO DO TILL THE DOCTOR COMES 314
How to stop Flow of Blood from Arteries. Bleeding from the Upper
Arm. Bleeding from the Neck. Wounds in the Thigh. Bleeding
from Veins. Hemorrhage of the Lungs or Stomach. Bleeding from
the Nose. Treatment of Burns. Danger from Burning Clothing.
CONTENTS. xiii
Treatment of Fainting. Broken Bones. Sunstroke. Treatment of
the Drowned. Swimming. Suffoeation in Wells. Bites of Cats,
Dogs, etc. Wounds from Thorns, Rusty Nails, etc. Snake Bites.
Poisons and their Antidotes. Poison Ivy. The Sick-room. Qual-
ities of the Nurse. Care of the Sick.
CHAPTER XXIV.
THE SKELETON 330
Axial Skeleton. Appendicular Skeleton. Uses of the Bones. Study
of a Vertebra. Table of the Bones. The Spinal Column. Articu-
lations of a Vertebra. The Cervical Vertebras. Atlas and Axis.
The Thoracic Vertebras. The Lumbar Vertebras. The Sacrum and
the Coccyx. Flexibility of the Spinal Column. Curves of the Spinal
Column. Cavities of the Skeleton. Pronation and Supination.
Weight of the Bones. Microscopic Structure of Bone. Classifica-
tion of Joints. Sprains and Dislocations.
CHAPTER XXV.
THE MUSCLES 341
The Number of Muscles. The Arrangement of Muscles. Forms of
Muscles. Names of Muscles. Peculiar Muscles. Heart Muscle.
Three kinds of Muscular Fiber Compared. Each Fiber a Cell.
Muscles of Expression. Muscles and Fat. Convulsions. Rigor
Mortis. Some Prominent Muscles. Sculpture and Anatomy.
APPENDIX . . 347
Antidotes. Disinfectants. Vital Statistics.
GLOSSARY 360
INDEX .... 371
PHYSIOLOGY.
CMA.P,TER I. ,'.::
Health. Is it not a splendid thing to be well and
strong ? To be full of bounding health ? To " feel one's
life in every limb " ?
Who does not desire to prolong, so far as possible, this
condition characteristic of youth ?
Natural and Artificial Modes of Life. An animal
living in a state of nature may keep well and live its
natural period of life without knowing anything about the
laws of health. But as students or indoor workers, many
of us lead a sedentary life ; we are not natural, but often
highly artificial, in our mode of living. We move about
but little, whereas the animal abounds in motion. We
concentrate energy upon mental effort, thus diverting a
large share of our sum total of energy away from the pro-
cess of nutrition. We often shut ourselves in rooms nearly
air-tight. We eat poorly chosen and ill-prepared food.
We devour it hastily, often when we are not in fit con-
dition to take food. In short, we frequently disobey the
laws of Nature. Now, Nature punishes every violation of
her laws. She never forgives, never forgets.
1
Value of Knowledge. The out-of-door worker may
not suffer so much from ignorance in these matters.
From the character of his occupation, he is, to a certain
extent, obliged to obey Nature. He gets enough fresh
air. His bodily exertion generally brings a hearty appe-
tite, vigorous digestion, active circulation of the blood.
Still, he would greatly profit by knowing something of the
nature of his food, its wholesomeness or unwholesomeness.
The fact that he has fair health is no proof that he always
does the best thing. His natural mode of life may keep
him in tolerably good condition in spite of his violation of
certain laws V b#t ^(f J-coldd :uXidpubtedly learn how to
economize in the purchase, preparation, and proper com-
bination of foods.
Importance of the Care of the Body. Any machine
of man's invention must be kept in good running order if
we would have it do good work, or last long. We must
keep a machine clean, well oiled, and not overtax it. Are
not our bodies worth equal care ? If some part of a ma-
chine is broken, we may replace it at moderate expense ;
but none of the vital organs can be replaced. We may
get a new mainspring for a watch, but we cannot obtain a
new stomach or lungs.
Its Admirable Mechanism. Aside from the above
considerations the human body is worthy of study for its
own sake. Viewed simply as a mechanism, it is wonder-
ful. Each organ is so well adapted to its work, and all
the organs work so harmoniously through their connection
and control by the nervous system, that we never cease to
admire it. We admire a doll, or other toy, so ingeniously
constructed that it can move its eyes or walk a short time
after being wound up. But this live mechanism, which is
INTRODUCTION. 3
self-winding, self-regulating, self-repairing, self-directing,
amazes us.
Hygiene. We take up the study of the human body
mainly that we may learn how to preserve health ; the
science of health is hygiene.
Physiology. In order to keep the various organs in
good order we must know what their natural work is, and
how they do it ; the science of the action of the body and
its parts is physiology.
Organ. Any part, or member, of the body, which has
a special work to do, is called an organ, as the hand, the
eye, or the stomach.
Function. -- The work, or action, of each organ is
its function.
Anatomy. In order to understand the working of
each organ it is usually necessary to know something
of its construction ; the science of structure is anatomy.
We do not need to go far into anatomy to obtain a fail-
knowledge of the manner in which our organs do their
work. The surgeon, of course, must be able to locate
accurately the various blood tubes, nerves, muscles, etc.
We need to know only the general structure of the body
and, more in detail, some of the more important organs,
such as the heart, the lungs, the larynx, the eye, etc.
It is fortunate for us that these organs in the sheep, pig,
and cow are so nearly like our own that they serve
admirably to enable us to understand ourselves.
Tissues. Every organ is composed of several different
kinds of material. For instance, in a slice across a ham
we see the skin on the outside, then fat, lean, and bone.
4 I'HYSIOLOGY.
These "primary building materials" of the body we call
tissues. A tissue may be defined as an aggregation of
similar cells devoted to a common work.
Cells. The whole body is made up of small parts
called cells, comparable to the bricks in a house. These
cells are of various shapes in the different tissues.
In the more active tissues the cells are alive, and each
cell may be compared to the ameba, a little mass of living
jelly-like substance called protoplasm. The ameba is a
protozoan often found in the slime at bottom of stagnant
water. Within this is a small, rounded part called the
nucleus. Most of the cells of the body differ from
the ameba in having a distinct
outer covering or cell wall. A grape
serves very well to show what a cell
is like. The whole body is built up
Nucleus. f cells, few of them large enough
Fig. i. Epithelial Ceils from to be seen by the naked eye.
Although the cells are closely
packed together, each cell leads, in one sense, an inde-
pendent life. But all work together to maintain the life
of the body. The cell is like the individual in a com-
munity. Each lives primarily for itself, yet all work
together for the good of the whole.
Epithelial Cells from the Inside of the Cheek. With the blade of
a very dull knife, or the handle of a scalpel, gently scrape the inside of
the cheek. Place a little of the white scraping on a slide in a drop
of water, cover with a cover slip, and examine under a quarter-inch
objective. Many cells will be seen, some of them showing nuclei.
Compare these cells with the accompanying figure.
The Physiological Division of Labor. We are aware
of the advantages of division of labor in a community. If
INTRODUCTION. 5
each person learns to do one thing well, all together work
economically for the common good, time is saved, and
better goods are produced. In the body there is a division
of labor similar to that of a community. Each organ has
its own work to do, and all work together for the common
welfare. The cells of each tissue have certain properties
and peculiarities of form differing from the form and
properties of the cells of any other tissue. While the
general structure of all cells is essentially the same, and
while they all have certain properties in common, each has
some one kind of work that it can do well, and to which
work it devotes itself. The nerve cells receive impressions
from the outer world, carry nervous impulses, and control
the various activities of the body. The muscle cells have
as their work the production of motion. All the cells
must take food for themselves and grow. Each has a
birth, life, and death, as each individual in a community
of men ; and as the community endures, while the indi-
vidual members are continually changing, so, in the body,
while the form remains about the same from year to year,
the cells are continually changing, some dying, and others
taking their places. .
In an animal of a single cell, like the ameba, the one
cell must do everything for itself. The higher animals all
begin their individual life as an egg, which is, in fact, a
single minute cell. This grows and divides, forming two
cells. By repeated division there accumulates a mass
of cells. These take on the arrangement peculiar to the
kind of animal from which the egg came. But as the cells
increase in number one group of cells takes up one part
of the work of .the body, other cells another part of the
work, and so on.
In studying history (sociology) we have to deal with the
6 J'HYSIOLOGY.
individual, the community, the state, and the nation. The
cell is an individual, the community is a tissue, the state is
an organ, and the nation is one body.
Let us proceed to study the nature of the individual cell,
and the combined actions of these individuals in that com-
munity called the human body.
Summary. i. Health is essential to comfort and efficiency in
work.
2. Our artificial mode of life is at variance with nature's laws.
3. Only by obeying the laws of nature can we preserve health.
4. We should learn these laws of nature from the advice and ex-
perience of others, and not by the expensive process of suffering from
disobedience.
5. Anatomy is the science of structure. Human anatomy is the
science of the structure of the human body.
6. Physiology is the science of function.
7. Hygiene is the art of preserving health.
8. Cells are the units of structure in the body.
9. A tissue is a group of similar cells having a single function.
10. An organ is a part having a special work or function. The
organs work together for the common good of the whole organism.
This working together results in
1 1 . The physiological division of labor, in which each organ works
for all the others, and is dependent on all the other organs.
Questions. i. What are some of the ways in which we most fre-
quently violate the laws of health ?
2. Name the more important organs of the body and their functions.
3. Name the different tissues of one of these organs.
CHAPTER II.
MOTION.
Motion and Life. Motion is the most manifest sign of
life. While we are sitting still, as we say, there are fre-
quent slight motions of the head, body, and limbs. Even
during sleep the movements of breathing may be seen ;
the hand laid upon the chest may feel .the beating of the
heart, and the finger detect the pulse in a number of
places.
We must move to get our food, or at least to eat and
digest it. Motion is necessary for breathing, for circu-
lating the blood, for getting rid of wastes. We often move
to avoid injury.
Motion is necessary for seeing : we must turn the face
toward the object; we move the eyeballs; within the eye
are motions to regulate the amount of light admitted, and
to adapt the eye for seeing at different distances.
In feeling, we put forth the hand to touch the object.
In tasting, we touch the tongue to the object. In smelling,
we sniff ; and sniffing is a respiratory motion. In hearing
and in speech there is also motion.
How are all these motions produced ?
Experiments with the Muscles in our own Bodies. i. Clasp
the front of the right upper arm ; draw up the forearm strongly
and as far as possible. Note what changes are felt in the biceps
muscle.
2. Repeat the experiment, and with the thumb and finger feel the
cord, or tendon, at the lower end of the muscle, just within the angle of
the elbow.
7
8 PHYSIOLOGY.
3. Place a weight in the hand, and repeat the act, noting the con-
dition of the muscle during the experiment ; also note the condition
of the tendon.
4. Span the muscle, placing the tips of the fingers in the angle of
the elbow, and the tip of the thumb as far as you can up the arm ; again
bend the arm. What change in the muscle does this show ? Any
muscle that bends a limb, as does the biceps, is called a flexor muscle.
5. Clasp the back of the right upper arm ; forcibly straighten the
arm. The muscle lying along the back of the arm is the triceps muscle.
It is called an extensor muscle because it extends, or straightens, the
arm.
6. Clasp the upper side of the right forearm near the elbow ;
clench the right hand quickly and forcibly ; repeat rapidly.
7. Notice the thick mass of muscle at the base of the thumb ;
pinch the forefinger and thumb strongly together. What changes can
be seen and felt ?
8. Place the hand on the outside of the shoulder ; raise the arm to
a horizontal position; repeat with a weight in the hand.
Fig. 2. The Shortening and Thickening of the Biceps Muscle in raising the
Forearm.
9. Stand erect with the heels close to each other, but not quite
touching; let the arms hang freely by the sides; rise on tiptoes,
without moving otherwise ; repeat ten times.
10. Place the tips of the fingers on the angles of the lower jaw;
shut the teeth firmly on a piece of rubber, and note the bulging of the
masseter muscles.
MOTION. , 9
11. Press the fingers on the temples ; again shut the jaw firmly, and
feel the action of the temporal muscles.
12. Make a narrow band of paper that will snugly fit the forearm
when the hand is open ; now clench the fist strongly.
13. With a tape measure take the circumference of the upper arm
when the arm hangs free ; again when the forearm is strongly flexed.
14. In the same way measure the forearm when the hand is open,
and when the hand is clenched.
By these experiments we learn that when a muscle works it becomes
shorter, thicker, and harder.
Nerves and Muscles of a Rabbit's Leg. In the hind leg of a
rabbit the sciatic nerve may be found by separating two large
muscles on the sides of the thigh, beginning behind the knee joint.
The shape and connections of the muscles may be learned, and also
the distribution of the nerve.
The Action of Muscle. The action of muscle is always
a "pull." The muscle shortens, at the same time thick-
ening and hardening. These changes in muscle are
roughly shown in the preceding experiments of feeling
the arm during its action. But the isolated calf muscle of
the frog may be made to prove the characteristic changes
with great clearness.
Action of Frog's Muscle. A frog may be killed painlessly by put-
ting a teaspoonful of ether into a fruit jar of water, immersing the frog
and capping the jar. When the frog becomes motionless, its head
should be cut off and a wire run down the spinal column to destroy the
spinal cord. After cutting the skin around the base of one thigh the
skin may easily be stripped from the whole hind limb. If the muscles
on the back of the thigh be gently separated there will be found a white
thread running lengthwise, the sciatic nerve. It should be severed
near the hip and carefully turned down upon the calf muscle. It should
not be pinched or dragged. The muscles of the thigh should now all
be cut away, being careful not to sever the nerve near the knee. The
hip joint should be unjointed. With the handle of the scalpel the calf
muscle should be separated from the shin bone, and just below the
knee the shin bone and all the muscles except the calf muscle severed
10
PHYSIOLOGY.
It now the heel cord be cut off below the heel there will remain such a
preparation as is represented in the accompanying figure, consisting of
the thigh bone with the calf muscle hanging from it, and the sciatic
Femur
Origin ~
Belly of Muscle -
Insertion
Weight
SHORTENED
ELONGATED
Fig. 3. Action of the Calf Muscle of the Frog, showing the Relations of the
Sciatic Nerve.
Origin
Bundle of Muscle Fibers
nerve still connected with the calf muscle. This may be supported by
holding the end of the thigh bone in a clamp on a retort stand. A
light weight should be attached to
the heel cord. The muscle and
nerve should be moistened with
water containing a little salt. On
pinching the free end of the nerve,
or cutting off the least bit with
scissors, the muscle will be made
to act. The shortening and thick-
ening will be plainly seen, and
by taking it between the thumb
and finger the hardening may be
felt.
Muscle Sheath
CROSS SECTION
Tendon
Insertion
LONGITUDINAL SECTION
Fig. 4. The Structure of Muscle.
Structure of Muscle.
Chipped beef shows well
the structure of muscle. The
MOTION.
II
white network is the connective tissue. In the meshes
is the red muscular tissue. The partitions which run all
through the muscle are continuous with the muscle sheath,
and both are continuous with the tendons at the ends of
the muscle. In fresh muscle the sheath and the parti-
tions are nearly transparent, and are not easily seen.
When the meat is cooked or salted the connective tissue
becomes white and opaque.
Microscopic Structure of Muscle. In frog's or rabbit's muscle
observe the thin, transparent membrane covering the muscle, the muscle
sheath. With forceps tear away part of the
muscle sheath. Tear the muscle to pieces,
and note its fibrous structure. A shred of
muscle may be mounted in a drop of nor-
mal saline solution on a slide, and exam-
ined with low power of the microscope. If
examined with a higher power the cross-
markings, or striations, will be seen. Such
muscle is called striated or striped muscle.
All of the muscles used in ordinary motions
are of this kind.
Fig. 5. Two Muscular Fibers
showing the Terminations of
the Nerves.
Effects of Cooking Muscle. In
well-cooked corned beef the connec-
tive tissue is thoroughly softened,
and the muscle fibers are easily separated. Thorough
cooking, especially slow boiling, will soften the connective
tissue, and may render palatable meat that, cooked other-
wise, would be exceedingly tough on account of the large
amount of connective tissue.
Imitation of Structure of Muscle. A good way to
represent the structure of muscle is to take a number of
pieces of red cord to represent the muscle fibers. Wrap
each in white tissue paper ; this represents the individual
12
fiber sheath. Lay a number of these side by side ; wrap
all in a common sheath ; let the tissue paper project be-
yond the threads, and here compress it into a compact
cylinder ; this last corresponds to the tendon.
Connective Tissue the Skeleton of Muscle. If all
the muscular tissue were removed from a muscle, the
sheaths and partitions would remain, just as they do in a
squeezed lemon or orange. The connective tissue forms
a framework for all the soft tissues of the body, and if
their working cells were removed, the connective tissue
would remain, and show more or less completely the form
of the part. Connective tissue, therefore, may be called
the skeleton of the soft tissues. Muscle consists, then,
essentially of a collection of soft, transparent tubes, filled
with the semi-fluid muscle substance. By scraping the
surface of a steak with a dull knife the muscle substance
may be obtained, leaving the connective tissue. This is a
good way to get the nutritious part of beef for an invalid.
Importance of Muscles. The different materials of
which the body is built up are called tissues. Thus we
find muscular tissue, bony tissue, nervous tissue, etc. The
muscles make up nearly half of the weight of the body.
This fact of itself should lead us to consider the muscles
of high importance. Add to this the facts above noted,
that the muscles are so largely concerned in the nutrition
of the body, the chief agents for its protection, essential
for the reception of ideas, and absolutely indispensable for
the expression of ideas, and we can see the reason for
beginning the study of physiology with the examination
of the muscles and their action.
Laws of Muscle Action. The chief characteristic of
muscle is its ability to shorten ; incidentally, it at the
MOTION'. 1 3
same time thickens and hardens. But it does its work by
shortening, pulling on the bones by means of the strong,
inelastic tendons, thus producing motion. The action of
the muscle as a whole is the result of the characteristics
of the cells of which it is composed. The individual cells
and fibers shorten, and their combined action is seen in
the muscular movement.
Extent of Muscle Shortening. A muscle may be
made to shorten one third of its length, but probably
never shortens that much in the living body.
Duration of Muscle Shortening. A muscle cannot be
kept shortened for any great length of time. If one holds
his arm out horizontally as long as possible he soon feels '
fatigue, later pain, and he may feel soreness in the muscle
for several days. The law of muscle action is to alternate
periods of rest with periods of action. In many exercises,
as in walking, the limbs act alternately, one resting or
recovering position while the other works.
Alternate Action of Flexors and Extensors. If we
consider the biceps and triceps of the arm, we see that
they are compelled to act alternately if they would do
effective work. They might both shorten at the same
time, and are made to do so in such an attempt as that
of holding the arm rigidly bent at a right angle ; as, for
instance, in wrestling " square hold," in which case one
wishes to prevent his opponent from either pushing or
pulling him. But while the two muscles act, no motion is
produced. When the flexor shortens, the extensor length-
ens, and vice versa.
Normal Condition of Muscle. The muscles are always
slightly stretched, as shown by the fact that when a cut
is made into a muscle the wound gapes open ; the tension
14 PHYSIOLOGY.
of the muscle is further shown by the fact that when a
bone is broken, as in the upper arm or thigh, the ends
of the bones slip by each other, and the limb has to be
strongly stretched to bring the ends back together. Mus-
cles act better when slightly stretched, and probably need
a slight resistant action of the opponent muscle.
Symmetrical Development of the Muscles; The mus-
cles of the two sides of the body are the same in number
and arrangement. At birth they are probably about equal
in size, weight, and strength. Most persons early become
right-handed, and the greater use of the right hand and
shoulder makes the muscles of this side larger and heavier.
The muscles pulling on the bones slightly modify them
in shape. The whole body may become noticeably un-
symmetrical. Most persons step harder on one foot than
the other, as shown by the sound of the footstep, and as
shown by the constant wearing of one shoe sole or heel
faster than the other. In many persons one shoulder is
habitually carried higher than the other. Symmetrical
development should be carefully sought, and any tendency
to a one-sided development should, so far as possible, be
avoided. We should use the left hand more. There are
many advantages in being able to use either hand. In
carving, in shaving, in bandaging, in administering medicine,
it may be necessary to use the left hand skillfully. The
pianist and the harpist use the two hands about equally,
while the violinist puts much more skill into his left hand.
Trainers of athletes often begin by developing the left
side of the body till it equals the right in size and strength.
Muscles the Source of Strength. Our strength de-
pends on our muscles. It is a fine thing to have strong,
well-developed muscles, not only because they give beauty
MOTION, 1 5
of form, but because extra strength and endurance may
be needed in case of accident, to save one's own life or that
of others. In a case of fire the ability to climb, to go up
or down a rope "hand over hand," may be all important.
Any one's life may depend on his ability to run far and
swiftly, to swim, to jump, or to lift a heavy weight.
Skeletal Muscles. When we look at the skinned car-
cass of an animal in the market, we observe that the mus-
cles almost completely cover the bones. Those which are
attached to the bones are called skeletal muscles. They act
upon them as levers, giving to motion strength, quickness,
and precision. Without bones our motions would be like
those of an earthworm or slug, slow and uncertain. The
muscles, acting through the bones, can lift a weight that
would crush the muscles if laid directly upon them, while
a bone, able to support a heavy weight without being
crushed, has no power in itself. The muscles have active
strength, the bones have passive strength.
Relation of the Muscles and the Bones. Suspend the skeleton
from the ceiling in the most open space in the room. Let the pupils
study it ; not to learn the names of all the bones, but to get a general
idea of the forms and relations of the parts. It is well to have the
skeleton constantly at hand, to show the location of the various organs
as they are taken up. If possible, supply the class with separate bones
from another skeleton, and let the pupils place each separate bone
alongside the corresponding one in the complete skeleton.
Pass to the skeleton, and locate the biceps muscle. After examining
Fig. 2, show the points of its origin and insertion. Feel the biceps
of your arm. Note that its thickest part is opposite the most slender
part of the bone. But at the enlarged end of the bone the muscle has
narrowed to a slender tendon, which passes over the joint to be attached
to the next bone, thus giving more slenderness, flexibility, and freedom
of motion to the joint. The muscle which closes the mouth, as in
pursing up the lips, is not attached to any bone, but in shortening
reduces the aperture.
1 6 /'//}'S/OUhi}'.
Flexion of the Forearm. Take the bones of the arm that are
articulated (if there is not an artificial hinge at the elbow, one can
readily be made of wire) ; put a strong rubber band in place of the
biceps muscle ; fasten this to the head of the humerus by cords, and
by the lower end to the radius, where the rough place, an inch or so
from the elbow joint, shows the insertion of the tendon. Have the
rubber stretched so that when not held it will flex the forearm. This
will serve to show the action of the biceps, though we must be careful
to bear in mind that the muscle does not pull the arm up because it has
been stretched, as is the case with the rubber. In the case of the
muscle, we know that the live muscle has the power of shortening when
stimulated, and in this respect is totally unlike the rubber. The live
cells, or units, act in concert.
Levers. The essentials of a lever are the point about
which the lever turns, called the fulcrum, the place where
the power is applied, called the power, and the part to be
moved, called the weight. In the body, the fulcrum is
some joint, the power is the place where the muscle is
attached, and the weight is the part to be moved.
Kinds of Levers. In flexing the forearm, the weight
is the hand or the hand and what is in it ; the fulcrum is
the elbow joint ; and the power is the point where the
tendon of the biceps is attached to the radius. This kind
of a lever is what the books call a lever of the third class.
The triceps, on the back of the arm, pulls on the projection
of the ulna (the inner bone of the forearm when the palm
is up), back of the elbow. The elbow is here, also, the
fulcrum, and the hand (or the object to be pushed by the
hand) is the weight. This kind of lever, where the fulcrum
is between the power and the weight, is called a lever of the
first class. In raising the weight of the body, by stand-
ing on tiptoe, we use a lever of the second class. Here
the ball of the foot is the fulcrum. The weight is the
weight of the whole body, resting on the ankle joint, while
MOTION. I/
the power is the calf muscle. We may find many exam-
ples of levers in the body if we look for them.
(1) Tapping on Floor. (2) Rising on Toe. (3) Lifting Weight.
Fig. 6. Three Kinds of Levers as shown by the Foot.
P Power. W Weight. F Fulcrum.
Kinds of Levers shown by the Foot. The different
classes of levers may be further illustrated by different
motions of the foot. In tapping the toes on the floor
while the heel is lifted, or in pressing down the ball of the
foot while running the treadle of a sewing machine, we
have an example of a first-class lever. In raising the
weight of the body on tiptoes, or as the foot is used in
taking each step, the foot is used as a lever of the second
class. When one lifts a weight with the toes, the foot is
used as a lever of the third class. These three classes of
levers are illustrated in the accompanying figures.
Advantages and Disadvantages of Levers in the Body. The
action of the bones of the forearm as a lever may perhaps be better
understood by the following considerations : If the arm consisted
merely of the biceps, suspended from the shoulder, it is evident that
its only action would be a straight pull. Suppose the biceps, thus
hanging alone from the shoulder, had a hook at its lower end, it could,
when it shortened, lift a weight just as far as it shortened, and no
i8
PHYSIOLOGY.
farther. It could not swing the wi:ight outward, or push it upward.
But from the way in which the biceps is attached to the forearm, when
the muscle shortens an inch it may move the hand a foot. Of course
the hand moves much faster, and we have a great gain in speed by
reason of this lever arrangement. But we cannot lift so heavy a weight
at this faster rate, as we could at the
elbow. For instance, suppose one were
to carry a heavy basket with a bail
handle by slipping the arm through
the bail up to the elbow. Now. it is
evident that the biceps is supporting
the weight. If it is as heavy as can be
held here, we know that we could not
hold the same weight in the hand with
the elbow bent at a right angle.
Ball
Articular Extremity
Medullary Cavity
Hard Bone
Study of One of the Long Bones.
For this, take, preferably, a femur
or a humerus. Let us suppose we ha\
a femur.
1. Observe its shape, cylindrical,
somewhat curved, enlarged at the ends.
2. The ends have smooth places,
where they fitted other bones.
3. Along the sides, especially near
the ends, are ridges and projections,
where the muscles were attached.
4. There are small holes in the
bone, where blood tubes passed in and
out.
5. Saw a femur in two, lengthwise,
and make a drawing showing :
(#) The central marrow cavity.
() The spongy extremities, noting especially the directions of the
bony plates and fibers.
6. Observe the width of the lower end of the femur, where it rests
on the tibia. Suppose these two bones were as narrow at their ends,
where they meet to form the knee joint, as they are at their centers,
what kind of a joint would they make ? Illustrate by piling up a num-
Spongy Bone
Articular Extremity
r. 7. Longitudinal Section of
Femur.
MOTION.
her of spools on end ; the column is more lightened than it is weak-
ened by the hollowing out of the sides of each spool. And the central
hollow of the spool does not greatly weaken it. A given weight of
material has more strength when in the form of a hollow cylinder. The
bones combine well two very desirable quali-
ties, lightness and strength. If in our col-
umn of spools we place a wide rubber band
around the junction of two spools, we have
something very similar to the capsular liga-
ment, which surrounds the joints.
Joints. The ends of the bones, where
they fit together in the joints, are covered
with a layer of smooth, elastic, whitish or
transparent cartilage. The motion in the
joints is made still more easy by the synovia,
resembling white of egg. The ends of the
bones are held together by tough bands and
cords of ligament, a form of connective
tissue very much like tendon. Bones are
closely covered by a tough coat of connective
tissue called the periosteum.
All these structures can easily be found
by dissecting a sheep shank gotten from the
butcher, or in the hind leg of a rabbit.
Locomotion. Locomotion is mov-
ing from place to place and should
be distinguished from mere motion.
By continuing such observations as
we made when we began to study
our motions, we can analyze and
understand many of the common
movements which we habitually
make.
Standing. Although we are not ordinarily conscious of
the fact, when we are standing still we are using many
muscles. The accompanying figure illustrates how some
Action of the Muscles
in Standing.
20 PHYSIOLOGY.
of the muscles act in keeping the body upright. Our
weight, or, we would better say, the force of gravity, is
continually trying to pull us down to the ground. The
joints are all freely movable, and hence as soon as the
muscles cease to act properly, in balancing against each
other, we lose our equilibrium, and fall if we do not
quickly regain it.
Walking. In walking, we lean forward, and if we take
no further action we fall. But we keep one foot on the
ground, pushing the body forward, while the other leg is
flexed and carried forward to save us from the fall. We
catch the body on this foot, and repeat the action. To
show how we are really repeatedly falling and catching
ourselves, recall how likely one is to fall if some obstacle
is placed in the way of the foot as it moves forward to
catch the weight of the body.
Running. In running, the action is more vigorous.
The propulsion by the rear leg is now greater. It gives
such a push as to make the body clear the ground, whereas
in walking, the rear foot is not lifted till the front foot
touches the ground. But in running there is a time when
both feet are off the ground.
Locomotion by Reaction. Take two broomsticks and place them
crosswise under the ends of a board. Run along the board. This
shows that the direct effort in running is to push one's support from
under him. When a horse plunges forward in the mud, he only thrusts
his feet farther into the mud. Our effort in progression is primarily to
push the earth out from under us, and it is by reaction that we go
forward. It is the same problem with the fish swimming forward by
striking backward and sideways against the water, and with the bird
beating downward and backward upon the air.
Bones combine Lightness and Strength. The mus-
cles, then, make use of the bones as levers. We carry
MOTION. 21
these levers with us all the time. Hence the desirability
of having them as light as is consistent with the requisite
degree of strength. The body follows the same law of
mechanics that we use outside of the body. A hollow
pillar or hollow tube has a greater strength than the same
amount of material in the form of a solid cylinder. The
long bones of the limbs are hollow, and near their ends,
where we have found that they need to be enlarged, we
find a spongy structure, where lightness and strength are
secured by the interlacing fibers and plates of bony
material.
Uses of Bones. The part that the bones play is of a
passive nature ; they support the tissues, protect some
parts, and serve as levers on which the muscles act. We
may not call the bones dead tissues, for they receive blood
and grow. But the active muscles use them as a man uses
a crowbar, as a mere tool. It will therefore be more
interesting to return to the muscles, and learn the causes
and conditions of their activity.
What makes Complex Muscular Action Harmoni-
ous. Have you ever seen two persons, each using the
right hand, try to sew, one holding the cloth, the other
using the needle ? Would they get along well ? Suppose
one were to hold the needle, and the other were to try to
thread it, each using one hand ? Why is it that the right
hands of two persons cannot work so well together as the
right and left hands of one person ? What connection is
there between the two, that one knows just what the other
is doing and when it does it ? Why can two individuals
never, with any amount of practice, work so in unity as the
parts of the individual ?
22 PHYSIOLOGY.
Let us seek the answers to these questions in the follow-
ing lessons.
READING. How to Get Strong and How to Stay So^
Blaikie ; Sound Bodies for Onr Boys and Girls, Blaikie ;
Physiology of Bodily Exercise, Lagrange.
Summary. i . Motion is involved in nearly every activity of the
body.
2. The action of muscle is a shortening, accompanied by a thick-
ening and hardening.
3. Muscle consists of fibers with a connective tissue sheath for
each fiber, bundle of fibers, and for the muscle as a whole.
4. The skeletal muscle fibers are striated.
5. The muscles make about half the body's weight.
6. Muscles may shorten one third their length.
7. They cannot remain shortened long at a time.
8. The muscles should be developed symmetrically.
9. In the limbs the muscles are fusiform and have their greatest
diameter opposite the central, or narrower, portions of the bones, con-
cealing the fact that the bones are largest at the ends, as is so manifest
in the skeleton.
10. The bones serve as levers by which the muscles exert their
force.
1 1 . The bones of the limbs are hollow cylinders combining lightness
-and strength.
12. The joints have a smooth motion due to the cartilage and
synovia.
13. Locomotion is brought about by reaction.
Questions. i . What effect is produced by carrying a heavy satchel
for a long distance without resting?
2. Which is more tiresome, standing still or walking? Why?
3. When the boy, who thinks he can strike a hard blow, says,
Feel my muscle,' 1 does he usually call attention to the muscle used in
striking?
4. Find other examples of levers in the body.
5. Find examples of the three kinds of levers, not in the body,
which we use often.
MOTION. 23
6. Why is it easier to sit with one leg crossed over the other?
7. What is the effect on muscles of light clothing?
8. How may the arms be used to illustrate the three kinds of
levers ?
9. Analyze and explain jumping, hopping, etc.
10. What is "curvature of the spine"? How caused and how
avoided ?
1 1 . What makes people bow-legged ?
12. Why are the sides of the body often sore after walking on icy
pavements ?
CHAPTER III.
THE GENERAL FUNCTIONS OF THE NERVOUS SYSTEM.
SENSATION AND MOTION.
What makes Muscles Shorten ? We have seen that
the muscles have the power of shortening ; that in shorten-
ing they act on the bones as levers to produce our varied
motions. What makes the muscles shorten ?
Voluntary and Involuntary Motions. Some motions
we will to make. We will to sit, to stand, to walk, to run,
or to stretch out the hand. Such motions, originating in
a brain activity, are called Voluntary. Other motions are
Involuntary. The will does not control the heart beat.
Most persons cannot keep from winking when a quick
motion is made toward the face, even if they know they
will not be hit. But all of these motions, whether volun-
tary or involuntary, are dependent upon the nervous
system.
The Cerebro-spinal Nervous System. This consists
of the brain, the spinal cord, and the spinal nerves. The
brain will be described later.
The Spinal Cord. The spinal cord is a cylindrical
body extending from the brain along the cavity of the
spinal column. Its diameter is not uniform throughout.
Between the shoulders is an enlargement called the cer-
vical enlargement, where the large nerves are given off to
24
NERVOUS SYSTEM.
Fig. 9. Diagram showing Arrangement of Nervous System.
26 PHYSIOLOGY.
the arms. In the region of the loins is the lumbar enlarge-
ment, where the nerves are given off to supply the poste-
rior limbs. The cord is not so long as the cavity of the
spinal column, and the space posterior to the cord is occu-
pied by the nerves extending to the posterior limbs, and
these nerves are given off at a very sharp angle, and con-
tinue backward for some distance before they emerge
from the cavity of the spinal column. But in the region
of the shoulders the nerves spring off at about a right
angle with the cord. The outside of the cord is white, but
the central portion consists of what is called gray matter.
The white portion is made up of fibers, but the gray matter
consists of nerve cells as well.
The Spinal Nerves. These are given off in pairs from
the sides of the spinal cord, passing out between the suc-
cessive vertebrae. In the regions of the shoulders and
loins the spinal nerves are large, as they supply the large
muscles of the limbs ; but in the middle of the back, where
only the muscles of the body wall are supplied, the nerves
are small. We have thirty-one pairs of spinal nerves.
The Roots of the Spinal Nerves. Each spinal nerve
arises by two roots, one nearer the back, called the dorsal
root, the other nearer the ventral surface, the ventral root.
These two roots soon unite to form one spinal nerve.
The Ganglion of the Dorsal Root. On the dorsal
root, just before it unites with the ventral root, is a swell-
ing, the ganglion of the dorsal root. Like all ganglions, it
is largely made up of nerve cells, being a center of con-
trol rather than a means of communication.
The Cerebro-spinal Nervous System of the Rabbit or Cat. It
will prove helpful at this point to examine the nervous system of a
NERVOUS SYSTEM. 2*]
rabbit. The animal may be killed painlessly by shutting it in a tight
box with a sponge holding a teaspoonful of chloroform or ether. The
smaller the box, the less anesthetic necessary. A large glass jar inverted
over the animal is convenient.
For a support, nail a foot of two-by-four scantling edgewise to a base
board. Lay the animal on this and tack out the feet. Slit and pull
aside the skin from the nose along the back to the base of the tail.
Remove the muscle along the sides of the back and neck. Between
the skull and the first vertebra is a space covered by a thin membrane.
This may be cut through with scissors. Then bone forceps may be
employed by inserting the point of a blade on each side of the cord and
cutting through the bone. In this way the whole of the dorsal portion
of the spinal column may be removed, exposing the spinal cord through-
out its entire length. With care the nerves and their roots may be
found. The nerves extending to the anterior limbs are easily traced,
but for the nerves to the posterior limbs more work is needed.
Structure of Nerves. When we trace the sciatic nerve
outward, we find that it is continually subdividing. This
division continues until the branches are too small to be
seen by the naked eye. Microscopic examination shows
that a nerve is made up of a great number of fibers bound
together in a common sheath of connective tissue, as is
the case with muscle. When the nerve divides there is
ordinarily no true branching or forking, but certain of
the fibers simply separate from the rest, as in the separa-
tion of the fibers in floss silk.
Structure of a Nerve Fiber. A single nerve fiber is
too small to be seen by the naked eye, being only about
one two-thousandth of an inch in diameter. It consists of
the following parts :
1. The Axis Cylinder, a central strand, or core, of semi-
transparent, gray material.
2. The Medullary Sheath is a layer of white, oily
material around the axis cylinder.
28 rnvsjoLOGY.
3. The Nerve Fiber Sheath is a thin, transparent outer
sheath of cdnnective tissue.
Function of Nerve Fibers. -The sole function of the
nerve fiber is to convey nerve impulses. The nerve im-
pulse passes along the axis cylinder as an electric current
passes along an insulated wire.
Nerve Fiber Sheath
Axis Cylinder
Medullary Sheath
Fig. 1 0. Structure of a Nerve Fiber.
Gray Nerve Fibers. In the sympathetic nerves there
are many fibers which have no medullary sheath, but con-
sist simply of the axis cylinder and the nerve-fiber sheath.
These are called gray nerve fibers.
Cross-section of the Spinal Cord. If a thin slice of
the spinal cord be made as shown in Fig. 11, it will be
seen that the central part is darker in color than the outer
part. The central part is known as the gray matter, in
distinction from the rest, which is called the white matter.
The white matter of the nervous system is made up of
nerve fibers whose structure and use we have just con-
sidered. But the gray matter has a different structure and
a different function. Instead of being made up mainly of
fibers it is composed of cells, one of the forms of which is
represented in Fig. 12. Some of the branches of these
cells are continued, and become the axis cylinders of nerves,
and it is believed that every nerve fiber begins as a branch
of some nerve cell. One of the best places to see these
nerve cells is in the gray matter of the spinal cord, near
NERVOUS SYSTEM. 29
the place where the ventral root of the spinal nerve arises.
This part of the gray matter is called the ventral horn of
the gray matter. If this portion be examined under a
moderately high power of the microscope, there may be
seen a number of cells with radiating branches.
Dorsal or Sensor
Root
Ganglion
i
Ventral or Motor Root
Fig. 1 1 . Cross-section of Spinal Cord.
Functions of the Spinal Cord. The spinal cord has
two main functions :
1. Its conducting power, by means of the white fibers
which make up the outer part of the cord. These fibers
may be regarded as connecting the gray matter of the
brain with all parts of the body.
2. The gray matter is the center of the reflex actions of
the cord.
Ganglia. Masses of nerve cells make up nerve centers,
or ganglia, such as are on the dorsal roots of the spinal
nerves. These also would show under the microscope
that their chief constituent is a collection of nerve cells
which give off one or more branches.
30 /V/KVA >/.(;<;r.
The gray matter of the spinal cord is considered LI col-
lection of ganglions. We see that the outer layer of the
brain is grayish in color. Within is white matter, consist-
ing of nerve fibers that connect the cells of the gray layer
Fig. 12. A Large Nerve Cell from the Gray Matter of the Spinal Cord.
with the various parts of the body through the base of the
brain, the spinal cord, and spinal nerves.
No Sensation without the Brain. After a fowl's head is cut off it
" flops " around for some time, and it may even jump clear from the
ground. If one takes hold of its feet to pick it up, it may begin to
struggle as if it were trying to escape.
Now, we know that the bird cannot feel anything after its head is
cut off, because the body is completely separated from the brain, which
is the center of sensation. So with the frog. After its head is cut
off, it cannot feel anything.
Reflex Action of the Spinal Cord of the Frog. A frog may be
killed as directed on p. 9. Cut off its head and suspend the body
from any convenient support, such as the ring of a retort stand.
1. On pinching the toes the foot will be drawn up.
2. The sciatic nerve should now be severed as before directed
(p. 9). At the instant of cutting the nerve the muscles below will
twitch, because the nerve fibers running to them are stimulated.
NERVOUS SYSTEM. 31
3. If the toes are again pinched, it is found that the uninjured leg
will draw up, but not the one whose sciatic nerve has been severed.
4. If a wire be run down the spinal cavity, the spinal cord will- be
destroyed, and during the operation the uninjured leg will act spas-
modically, because the nerve fibers going to its muscles from the cord
are stimulated.
5. Pinching the toes no longer gives response, because the cord,
which acted as the center of this reflex action, is destroyed.
The Gray Matter of the Cord the Center of Reflex
Action. In simple sensation of touch, pressure on the
Afferent Dorsal Root
Sensor Fiber
Skin
Motor Fiber v v
Efferent Ventrg| Root
Fig. 1 3. Diagram of Reflex Action of the Spinal Cord.
(After Landois and Stirling.)
toes starts a nerve current or nerve impulse which runs up
to the brain. The sensation is in the brain, but is referred
to the foot. Hence we should be careful not to speak of
a sensation being carried. In voluntary muscular action
the impulse starts from the brain, goes to the muscles, and
makes them shorten or relax.
But in reflex action the current runs up the nerve to the
spinal cord. The gray matter of the central part of the
cord receives the message, and sends back a nerve impulse
to the muscles to make them shorten and pull the foot
away from the source of injury.
32 PHYSIOLOGY.
The Parts Essential to Reflex Action of the Spinal
Cord : -
1. A sensitive surface (the skin, for instance).
2. Afferent nerve fibers.
3. A nerve cell, or cells, in the center of the spinal cord.
4. Efferent nerve fibers.
5. Working organ, as muscle or gland.
Phases of Reflex Action. In the above experiment
on the frog the steps in order were :
1. Stimulation of the nerve endings in the skin of the
toe.
2. Passage of a nerve impulse up the afferent fibers to
the spinal cord.
Nerve-Cell
Afferent Fiber ^ \ ,_ _.,
x Efferent Fiber
Skin !,
&-- Muscle
Fig. 14. Scheme of Reflex Arc.
3. Reception of the impulse by a cell, or cells, of the
gray matter in the cord.
4. Sending back a nerve impulse
5. Along an efferent fiber, or fibers, to
6. Muscles which shorten and move the foot.
Importance of Reflex Action. It is important that
we understand the nature of reflex action, for very many
of the processes of the body are regulated by it. Not
only the more manifest motions, such as winking when
NERl/OUS SYSTEM. 33
anything comes quickly toward the eye, dodging, jumping
when suddenly touched by anything hot or when pricked
by a pin, but also the adjustments of the essential processes
of life, circulation, respiration, and digestion, are brought
about through reflex action.
Destination of Nerve Fibers. The sciatic nerve is
composed of many fibers. If this nerve is traced outward,
it is found to be continually subdividing, and sending small
branches to the muscles, and finally in the muscles one
fine nerve fiber goes to each muscle fiber. (See Fig. 13.)
Many fibers go on past the muscles to the skin. We can
feel in any part of the skin, and we can tell just where we
are touched. These fibers from the skin, then, carry
nerve impulses inward, as those going to the muscles
carry impulses outward.
Nerve Roots and their Functions. Experiments on
the lower animals, and accidents in the case of man, show
that all the fibers of the nerves that carry currents to the
muscles pass out from the spinal cord into the ventral
root, and that all the fibers that carry currents inward
enter the spinal cord through the dorsal root. Hence, the
dorsal root is often called the afferent root, and the ventral
the efferent root. Since ingoing impulses produce sensa-
tion, the dorsal root is called the sensory root, while the
ventral root, carrying currents outward to produce motion,
is called the motor root.
Effect of Stimulating a Spinal Nerve. Experiments
have shown that if, in an uninjured animal, a nerve, or
more properly a nerve trunk, as the sciatic nerve, be
stimulated, for instance, by a suitable electric shock, two
effects are produced : first, motion in the parts whose
34 PHYSIOLOGY.
muscles are supplied by the nerve ; second, sensation,
which is referred to the parts of the skin supplied by the
branches of the nerve.
Effect of Severing a Spinal Nerve. If. instead of simply stimu-
lating the nerve, the nerve is severed, the same two effects will he pro-
duced. After severing the nerve, if we stimulate the end of the nerve
still connected with the limb, we get action of the muscles in that limb.
If we stimulate the end of the nerve connected with the body, a sensa-
tion will be produced, and this sensation will be referred to the parts
from which the nerve fibers arise, probably in the skin of the limb.
Effect of Stimulating the Ends of Severed Nerve Roots. If we
now turn to the roots of the nerve, and make similar experiments, we
obtain the following results : Stimulating the dorsal root causes sensa-
tion referred to some outer surface, and no other effect is noticed.
Cutting the dorsal root also causes sensation. Stimulating the end of
this root still connected with the spinal cord causes sensation: but
stimulating the end of the root connected with the nerve gives no
appreciable result.
Stimulating or cutting the ventral root causes motion in the parts
whose muscles are supplied by fibers from this root. After severing
this root, if the end connected with trie spinal cord be stimulated, no
effect is noticed ; but stimulating the end still connected with the nerve
is followed by shortening of the muscles supplied.
Effect of Severing All the Spinal Nerves. Severing
all the spinal nerves destroys all power of sensation and
voluntary motion in all parts of the body except the head.
After severing all the dorsal roots, no sensation would be
produced by stimulating any part of the body, and after
severing all the ventral roots no act of the will can cause
any of the muscles of the body to act. Severing all the
nerves, or severing all the roots, cuts off all communication
of the brain with the body, and so far as motion and sensa-
tion in the body generally are concerned, has the same
effect as severing the spinal cord below the head.
NERVOUS SYSTEM. 35
Cramp. Cramp is a spasmodic shortening of the
muscles, attended with pain.
Tetanus. Tetanus (or locked jaw) is a spasmodic and
continuous shortening of the muscles, causing rigidity of
the parts they supply. It is due to the disordered and
excessive stimulation of the muscles through the nerves.
Crossing of the Fibers from the Brain to the Spinal
Cord. Both the brain and the spinal cord consist of two
lateral halves connected by cross fibers. Each half of the
brain is connected with the opposite half of the body.
This is accomplished by the crossing of the fibers. The
fibers that carry nerve impulses outward are now known
to cross as they leave the brain, at the very beginning of
the spinal cord, in the part known as the spinal bulb.
The sensations arising from touching anything with the
right hand, therefore, are in the left half of the brain, and
the right half of the brain controls the left hand.
Voluntary Interference with Reflex Actions. We
have seen that the jerking of the hand away from a hot
object is due to reflex action of the spinal cord. One
might, by a powerful effort of the will, keep the hand on
an object that is hot enough to burn the skin. One may
command the foot to remain quiet when it is tickled ; but
as soon as the person is asleep, the same stimulations
would be followed by the reflex actions such as we have
considered.
In these cases of interference it is understood that the
brain sends a nerve impulse down to the centers of the
reflex action, and stops or diminishes their operation.
This retarding influence of a group of cells is called inhi-
bition. It is not always due to voluntary interference,
but may be due to reflex interference, as we may see later.
36 rHYSIOLOGY.
The Nature of a Nervous Impulse. Of the nature
of a nerve impulse we know but little. It is convenient
to compare the nervous system, with its conducting fibers
and central ganglia, to a telegraph system. And electric-
ity is the most convenient stimulus for exciting nerve im-
pulses. Yet a nerve impulse is very different from an
electric current. A nerve fiber is a poor conductor of
electricity. An electric current may travel along a copper
wire at the rate of between 100,000 and 200,000 miles a
second, while a nerve impulse in a motor nerve travels
only 1 70 feet in a second.
Transmission of Motor Impulses. When a motor fiber is stimu-
lated in the middle of its course we observe only one effect, the
shortening of the muscle at its lower end. But there is every reason
to believe that the nerve current, or impulse, runs along the nerve in
both directions from its starting point. But while the action of the
muscle at the peripheral extremity manifests the existence of the cur-
rent, there is nothing at the central extremity to give such evidence.
Transmission of Sensory Impulses. Similarly, when a sensor
nerve fiber is stimulated at some intermediate point, we have a sensa-
tion in the brain due to the current brought by the afferent fiber, and
which we refer to the outer end of the nerve fiber. Probably a nerve
impulse passed from the point of stimulation to the outer end of the
fiber ; but as there is nothing at the outer end of the nerve fiber to
interpret it, we get no evidence of such impulse except, by refined
physiological tests.
Harmony in Muscle Action. In throwing a stone a
number of muscles are used. Each one of these must
shorten in the right way and at the right time or the throw
will not be accurate. Each muscle shortens under the
influence of a nerve impulse started by the brain and
brought by a motor nerve. If any muscle shortens an
instant too soon, or a little too strongly, the stone goes to
NERVOUS SYSTEM. 37
one side. In a tune on a piano we know that the right
keys must be struck ; that each must be struck at the right
time, with the proper degree of force, and held for the
right length of time, or we have discord instead of har-
mony. What the player is to the instrument, the brain
is to the body.
Temporary Loss of Muscular Power. It may have
happened to you that after sitting long in one position you
attempted to stand, but found that you could not do so.
One leg failed to act at the bidding of your will. When
the foot is " asleep " we get little sensation from it ; we
hardly know whether it is touching the floor or not. Press-
ing on it with the other foot causes no pain.
We try to stand when the foot is asleep, but we are
unable to do so. The brain starts the nerve currents, and
they run along the nerve as far as the compressed part ;
here they stop. They cannot reach the muscles of the
leg below. Hence the muscles do not shorten, and we
do not rise, no matter how strongly we will to do so.
Why is it that the nerves and muscles thus sometimes
lose their ability to perform their natural activities ?
Dependence of Nerves and Muscles. This has been
explained by saying that owing to external pressure, the
nerve has temporarily lost its power of conducting nerve
currents. But what beside the nerve has been com-
pressed ? What process in the limb has been interfered
with by the pressure due to the position in which one has
been sitting or lying ? What is the temperature of the
benumbed limb ?
On what are the nerves and muscles so dependent for
the maintenance of their activity ?
38 niYSioi.OGY.
READING. Power through Repose, Call ; The Technique
of Rest, Brackett; Muscles ami Xerres^ Rosenthal.
Summary. I . Motions are voluntary or involuntary, but all are
under control of the nervous system.
2. The cerebro-spinal nervous system consists of the brain, the
spinal cord, and the spinal nerves.
3. Each spinal nerve has two roots : the dorsal, which is afferent
and sensory ; the ventral, which is efferent and motor.
4. A ganglion is a nerve center largely composed of nerve cells.
5. Nerves are made up of nerve fibers.
6. A nerve fiber consists, of the central core (or axis cylinder),
which conducts the nerve impulse, the medullary sheath, and, outside,
the nerve-fiber sheath.
7. The spinal cord has in its outer part white nerve fibers, in its
center gray nerve cells.
8. These cells are branched, and at least one branch becomes the
axis cylinder of a nerve fiber.
9. The gray matter of the cord is the center of the reflex action.
10. The nerve fibers from each half of the brain connect with the
opposite half of the body.
11. The nervous system is comparable to a telegraph system.
Questions. i. Name as many involuntary motions as you know.
2. What other cases of reflex action do you know ?
3. The story is told of a young Roman (Mucius Scaevola) that to
show his fortitude he thrust his hand into the fire and held it there
until it was destroyed. What physiological action does this illustrate ?
4. Why is a man partially paralyzed when he has broken his neck
or back ?
5. How does the nervous system differ from a telegraph system?
CHAPTER IV.
CIRCULATION OF THE BLOOD.
The Blood and its Work. We know that if any animal
is bled freely, it soon becomes weak, then unconscious, and
soon dies, if the escape of blood be not stopped.
We observe the natural difference in color of different
parts of our bodies ; for instance, the lips and cheeks.
We often note varying color, as in blushing and pallor.
We wish to understand these differences and changes ;
also to know what to do in case of fainting or bleeding
from wounds. We may prolong and make more useful
our own lives and those of others by knowing, in a practical
way, something about the causes, prevention, and remedies
of the colds, congestions, and inflammations to which we
are subject.
Nearly every part of the body bleeds when cut. There
is no bleeding when we trim the nails or cut the hair, and
the outer skin has no blood in it. But the inner skin, and
almost every tissue within it, if pierced even by the finest
needle, yields blood. We see a little blood oozing from
the surface of a fresh steak or roast.
What kind of a substance is the blood ? Is it uniformly
distributed through the tissues, like water soaked up into a
cloth, or is it in distinct cavities ? Why is it so essential to
life ? How does it do its work ?
The Rate of the Heart Beat. The heart beats about
seventy-two times a minute in men ; in women, about
39
4O PHYSIOLOGY.
eighty. At birth the rate is from one hundred and thirty
to one hundred and forty, and gradually decreases till
about the age of twenty, when the average of seventy-two
is reached. This rate holds till old age, when it increases.
The rate is increased by muscular activity, food, external
heat, internal heat (fever), pain, and mental excitement.
Music accelerates the pulse rate. The pulse rate varies
during the twenty-four hours, being lowest during the
night, and highest about 1 1 A.M. Certain diseases increase
the frequency of the pulse. Some drugs quicken the
pulse rate, and others dimmish it.
The Heart Beat and the Pulse. i . The heart beat, felt at the left
of the breast bone.
2. The pulse, felt at the wrist and at various parts of the body.
Perhaps the most convenient place to study it is at the temple. Lay
the forefinger lightly along the cheek just in front of the ear. Count
the pulsations for a minute.
Let one or two pupils who are quick at figures step to the blackboard
and put down the number of pulsations of each pupil, and divide by the
number thus reporting, to get the average.
1 . Let all in the class count the pulse while sitting. Probably it
will be best to discard the first trial, as there are likely to be several
failures from one cause or another. Then, too, there is usually a slight
excitement at the beginning of a wholly new experiment. Get the aver-
age of the class.
2. Find the pulse while sitting; rise quickly, and immediately begin
to count the pulse. Compare with the pulse as taken while sitting.
3. Compare the pulse before and after meals.
4. With the thumb and finger lightly clasp the windpipe, well back.
The pulse in the carotid arteries will be felt.
The Position of the Heart. The base of the heart is
in the center of the chest, just back of the breast bone, but
the apex points downward and to the left.
The Covering of the Heart. The heart is inclosed in
a loosely fitting membranous bag, the pericardium. Within
CIRCULATION OF THE BLOOD. 41
the pericardium and around the heart ivS a small quantity
of liquid, called the pericardial fluid.
The Size of the Heart. A person's heart is about the
size of his clenched hand.
The External Features of the Heart. The heart is
cone-shaped and the bulk of it is made up of the ventricles,
the auricles being two ear-like flaps at the base, one on
each side. There is a deep notch between the auricles
and the ventricles. The line of division between the two
ventricles is marked by a groove, which runs obliquely
along the ventral surface. In this groove are blood tubes
and usually considerable fat.
The Internal Structure of the Heart. The two halves
of the heart are completely separated from one another
by a partition. Each half, in turn, has valves which,
part of the time, separate the cavity of each auricle (at
the base) from the cavity of the corresponding ventricle
(at the apex).
The Valves of the Heart. Between the auricles and
the ventricles are curtain-like valves, whose upper edges
are attached to the inner surface of the walls at the upper
margin of the ventricle. These flaps are somewhat tri-
angular, and have strong white, tendinous cords extending
from their edges and under surfaces to the walls of the
ventricle below. In the right half of the heart there are
three flaps, and this valve is called the tricuspid valve. In
the left side there are two flaps, which, together, constitute
the mitral valve. In the resting heart these flaps hang
down along the walls of the ventricles so that on opening
the heart one would see only a single cavity in each half
of the heart.
42
The Semilunar Valves. From the base of the right
ventricle arises the pulmonary artery. Within its base,
just as it leaves the ventricle, are three pocket-like valves,
like " patch-pockets." They are in a circle, with their edges
touching, and thus surround the opening, with their
mouths opening away from the heart. A similar set of
valves are within the base of the aorta, which arises from
the left ventricle. Both these sets of valves are called
semilunar valves.
Dissection of the Heart. No description (nor even figures) can
give a clear idea of the heart. A good model will be of some assist-
ance. But the heart itself should be examined carefully and then dis-
sected. The heart and lungs of a sheep should be obtained (ask the
butcher to save the "pluck," i.e. the heart and lungs taken out together).
The relations of the heart to the lungs and other organs should first
be studied, and then the pericardium opened. Observe the outside of
the heart, and then cut the heart open to see the points given in the
above description. After the heart is severed from the lungs the auri-
cles may be cut off; then, by pouring water into the ventricle, the
action of the valves between the auricles and the ventricles will be
seen. Pressing on the outer surface of the right ventricle will make
the water escape through the pulmonary artery. If this be split open,
the semilunar valves at its base may be found.
The Blood Tubes connecting the Heart with Other
Organs. The aorta (the largest artery in the body)
arises from the base of the left ventricle, and supplies
with blood every organ of the body except the lungs.
The pulmonary artery springs from the base of the right
ventricle and sends blood to the right and left lungs.
Two large veins enter the right auricle, the precaval vein
from the anterior regions of the body and the postcaval
vein brings blood from all the organs of the posterior por-
tions of the body. The pulmonary veins return the blood
from the lungs to the left auricle, two from each lung.
CIRCULATION OF THE BLOOD.
43
44
PHYSIOLOGY.
fL External Jugular Vein
Internal Jugular Vein
2 Subclavian Artery
b Subclavian Vein
1 Carotid Artery
i Aorta
in Precaval Vein
IV Postcaval Vein
Gastric Artery
Splenic Artery
Hepatic Artery
Pancreatic Artery
ff Renal Veins
5 Renal Arteries
T Iliac Arteries
i Iliac Veins
Fig. 16. Distribution of Arteries and Veins.
CIRCULATION OF THE BLOOD.
45
The Distribution of the Arteries and Veins. The
organs of the body receive a supply of blood in propor-
tion to their size and activity. The artery supplying the
blood and the vein which returns it usually lie side by side
(see Fig. 16). The larger arteries are usually deep-seated
and in protected places.
Demonstration of the Action of the Heart. The heart may be
mounted as shown in Fig. 17, and its action illustrated by compressing
the ventricles with both hands. Instead of the apparatus here shown
two retort stands may be used, though not so convenient.
Capillaries
of the Lungs
Capillaries
of the Body
Fig. 17. Demonstration of the Action of the Heart (Heart Diagrammatic).
The Action of the Heart. - - The -heart consists of
muscle fibers so arranged that they form a thick-walled
bag, which stands expanded when the muscles relax. But
when the fibers shorten the whole heart contracts, and the
46 PHYSIOLOGY.
cavity is much reduced in size, if not entirely obliterated,
and the blood is forced out.
The complete action of the heart consists of three parts,
the contraction of the auricles, the contraction of the
ventricles, and the pause.
The Pause. During the pause the blood is steadily
pouring into the auricles ; into the right auricle from the
caval veins, into the left auricle from the pulmonary veins.
At this time the curtain-like valves between the auricles
and the ventricles are open, and their flaps hang loosely
beside the walls of the ventricles. The blood, therefore,
as it passes into the auricles, passes on into the ventricles.
As the ventricle fills, the valves float up, as seen in the
experiment of pouring water into the ventricle.
The Contraction of the Auricle. When the ventricle
is full, but not stretched, and the auricle partly full, the
auricle suddenly contracts, thus forcing more blood into
the ventricle, and distending it. At the same time the
valves, which were already nearly closed, are tightly closed
by the pressure of the blood which is forced up behind
them. The flaps of the valves are kept from going up too
far by the tendinous cords and by the papillary muscles to
which the cords are attached.
The Contraction of the Ventricle. Next comes the
contraction of the ventricle, slower, but more powerful
than that of the auricle. As the walls of the ventricle are
drawn together, the blood is subjected to pressure. It
cannot go back into the auricles, for the more it presses
against the valves, the more tightly they are closed. The
semilunar valves are closed by back pressure in the aorta
and pulmonary artery. But the pressure of the blood in
the ventricles is so much greater that the semilunar valves
CIRCULATION OF THE BLOOD,
47
are forced open, and nearly all the blood is driven out of
the ventricles ; from the right ventricle into the pulmonary
artery, and from the left ventricle into the aorta.
While the ventricles are contracting and forcing their
blood out, the auricles are slowly filling by the steady
inflow through the veins.
Systole and Diastole. The contraction of the heart is
called the systole, and its dilation the diastole.
Dilation of the Ventricle. As soon as the ventricle
has completed its contraction it dilates, and most of the
blood that has accumulated in the auricle simply falls into
the ventricle. The dilating ventricle exerts a slight suc-
tion, so the blood is in part drawn into the ventricle. Dur-
ing the remainder of the pause the blood accumulates in
Auricle
Fig. 1 8. Diagram of the Heart, showing the Action of the Valves.
the auricle and ventricle till the auricle again contracts and
the cycle is repeated. This is true of both halves of the
heart, which work simultaneously, the right heart pumping
dark blood while the left heart pumps bright blood. The
left ventricle is thicker walled and stronger than the right.
48 PHYSIOLOGY.
Work and Rest of the Heart. The time taken by the
different parts of the heart beat are about as follows : The
auricle contracts about one eighth of the time and rests
the other seven eighths. The ventricle contracts about
three eighths of the time and dilates during about five
eighths. If we suppose the whole period of the heart beat
to be twenty-four hours (instead of eight tenths of a sec-
ond), we can more easily see how much of the time the
heart is actually at work, and how much of the time the
heart is resting.
Auricle contracting (working) \ of the time 3 h., resting 21 h.
Ventricle contracting (working) f of the time 9 h., resting 15 h.
No part of the heart, therefore, is working longer than a
man would who only works nine hours a day. Some ob-
servers state that the resting period is even greater than
these figures would show.
Since the contraction of the ventricles immediately fol-
lows that of the auricles, one half of the time is occupied
by the whole contraction of the heart, and during half the
time the whole heart is resting. This is different from our
usual statements regarding the work of the heart. We
hear it said that the heart never rests. Its work and rest
follow each other at such short intervals that we do not
appreciate the interval of rest that comes between the suc-
cessive impulses that we feel. Suppose a policeman had
the power of sleeping at will, and that he slept thirty min-
utes of each hour, and that in the remaining thirty minutes
he made the rounds of a block. If we saw him passing
regularly once an hour, every hour of the twenty-four, we
might suppose that he did not sleep at all during the entire
time. This ratio of work and rest is fairly constant in
the varying rates of heart beat.
CIRCULATION OF THE BLOOD.
49
The Beat of the Heart. The apex of the heart is
always in contact with the chest wall. Consequently, it
never strikes it. At each beat it pushes hard against the
chest wall. This push may be felt and seen, and is called
the heart beat.
The Sounds of the Heart. There are two sounds of
the heart :
1. A short, sharp sound made by the closing of the semi-
lunar valves.
2. Just preceding this sound a longer, duller sound may
be heard during the contraction of the ventricles. This is
supposed to be due to the vibrations of the walls of the
ventricles and of the large valves.
As soon as
Nucleus
Isolated Fibers
Fibers Joined
Action of the Large Arteries. The large arteries
have in their walls a yellow elastic tissue. When the
blood is forced into them, they are stretched,
the ventricle ceases to contract,
and sends no more blood into
the arteries, they "stretch back."
We should not say contract, for
it is simply an elastic reaction.
As the artery reacts it presses on
the blood, and hence the blood
tries to escape in every possible
way. It cannot go back, for it
fills the pockets of the semilunar
valves, and closes them with a
click. A rapid wave is sent for-
ward that gives the pulse, and a slower but still rapid
stream flows along the arteries, through the pulmonary
artery to the lungs, and through the aorta and its branches
to all the other parts of the body.
Fig. 1 9. Plain (Unstriated) Muscu-
lar Fibers from the Bladder.
50 PHYSIOLOGY.
The elastic reaction of the arteries thus helps to make
steady the flow of blood, which is intermittent as it leaves
the heart. The medium-sized arteries also have elastic
tissue in their walls, and regulate the blood flow in the
same way.
Variation of the Amount of Blood Needed. Each
organ requires a supply of blood in proportion to its
activity. An actively working organ, like the brain, de-
mands much more blood than bone, practically inactive.
Further, working tissues, such as the brain and muscles,
need a great deal more blood while they are at work than
when they are resting. An organ needing a constant large
supply of blood might secure this by having a large artery.
But how can the supply be regulated so that an organ
may receive, now more, now less, according to its needs ?
Plain Muscle Fibers in the Walls of the Arteries.
This is regulated by the medium-sized and small arteries
Connective Tissue
Endothelium
Muscle Fiber
Fig. 20. Plain Muscle Fiber. Isolated and in Wall of Artery.
leading to the parts. In the walls of these arteries are
muscle fibers of a different kind from those of the skele-
CIRCULATION OF THE BLOOD. 51
ton. These fibers are spindle-shaped cells, as shown in
Fig. 19, with a nucleus near the center, and do not have
the cross-markings of the fibers of the skeletal muscles ;
they are in consequence called nonstriated, smooth, or
plain muscle fibers. They are arranged circularly in the
walls of the arteries. These fibers have, in common with
all muscle fibers, the power of shortening. When they
shorten they reduce the size of the artery, and, there-
fore, for the time, less blood can flow through the
artery. When the muscle fibers cease to shorten, the
artery widens, and allows more blood to pass through it.
Illustration of the Action of Muscles in Arterial Walls. To
illustrate the action of the muscles in the walls of an artery, let the
water run through a hose or large
rubber tube. Now, if a row of per-
sons take hold of this tube, the grip Endotheiium
of their hands is like that of the
muscles. When the hands tighten internal Elastic
their grip, the caliber of the hose La y er
or tube is diminished, and less water
is allowed to flow through it. When Circular Mus-
cle Fibers
the hands relax, the tube, being
elastic, allows more liquid to flow
through it.
Illustration of a Small Artery.
To represent a small artery, take
a small, thin-walled rubber tube and
wind a red thread around it. Now,
if the thread could be made to Fl * 21 - Coats of a Small Artery.
shorten, it would diminish the cali-
ber of the tube. The representation would be more exact if the thread
were cut into many short pieces, and if each piece were thicker in
the middle, and were then glued to the tube. If the whole were
covered by a layer of tissue paper, the structure of the artery would be
roughly represented.
52 PHYSIOLOGY.
Plain and Striated Muscle Fibers Compared. These plain mus-
cle fibers are further like those of the skeletal*muscles in that they are
under the control of the nerves, but they are involuntary in their action.
PIGMENT
CELLS
ARTERY
Fig. 22. Part of Frog's Web (low magnifying power).
We cannot interfere with the action of these muscles, no matter how
strongly we may will to do so. Without our thinking about it, more
CIRCULATION OF THE BLOOD.
53
blood goes to the muscles of the legs when we walk, more to the brain
when we are studying, to the digestive organs after eating, etc. The
Walls of Capillari
Tissues of Web
\
Fig. 23. Part of Frog's Web (highly magnified).
plain muscle fibers shorten at a much slower rate than the striated
fibers. They are also slower in relaxing. Since the plain muscles are
usually found in the walls of hollow organs such as the heart, blood
54
PHYSIOLOGY.
tubes, digestive tube, etc., they are sometimes called visceral muscles in
distinction from the skeletal muscles.
The Circulation of Blood in the Web of a Frog's Foot. For this
get a frog with a pale web. Take a piece of shingle six inches long
and three inches wide. Cut a round hole, half an inch in diameter,
near one end of it. Wrap the frog in a wet cloth, with one leg project-
ing, and tie it, thus wrapped, to the shingle. Tie threads around two
of the toes, and stretch the web, but not too tightly, over the hole.
Keep the web moist. Place the shingle firmly on the stage of a micro-
scope. Examine first with a low power. The large tubes which grow
smaller by subdivision are arteries. The large tubes which are
formed by the union of smaller ones are
the veins. The finer tubes, forming a net-
work in every direction, are the capillaries.
They receive the blood from the arteries
and pass it on to the veins.
Put on a higher power, a one-fifth or
one-sixth objective. It may now be seen
that the colored corpuscles float more in
the center of the stream, and with a steady
motion, while the colorless corpuscles keep
close to the walls of the capillary, and seem
to adhere to them, advancing with a hesi-
tant motion, seeming to roll along against
the wall of the capillary.
Close your eyes for a moment, and re-
flect that in all the active tissues of your
body for example, the muscles, brain, and
digestive organs there is a similar net-
work of fine tubes with a current of blood
running through them. The current is not so rapid as it seems, for the
microscope magnifies the rate of flow as well as the size of the cor-
puscles. The blood really is moving slowly in the capillaries, and it is
very important that it should be so, for in the capillaries the work of
the blood is done. Part of the liquid of the blood soaks through the
thin walls of the capillaries, and nourishes the surrounding tissues. All
the other parts of the circulatory system exist for the purpose of send-
ing a continuous, slow, and steady stream of blood through the
capillaries. (See pages 72 and 73.)
Fig. 24- Capillary Blood
Tubes of Muscle.
CIRCULATION OF THE BLOOD.
55
Connective
Tissuf
Artery
Fig. 25.
Cross-section of Small Artery
and Vein.
The Blood Flow in the Capillaries. The arteries
divide and subdivide, and become capillaries, which have
connecting branches, form-
ing a close network of tiny
thin-walled tubes. These
penetrate nearly every tis-
sue of the body. The blood
cannot do its full work till it
is in the tissues/and to reach
the tissues it must soak
through the walls of the
capillaries. The work of
the heart and arteries is to
keep a steady flow of blood
through the capillaries, that the tissues may be constantly
supplied.
How is it that the jerky action of the heart, at each
contraction sending a jet of
blood into the arteries,
shown by a spurt when an
artery is severed, and also
indicated by the intermit-
tent pulse, how is this
intermittent flow converted
into the steady, uniform
current that we have seen
in the capillaries ?
Experiments illustrating the
Blood Flow in the Capillaries.
A few experiments may make this Fig. 26.
Surface View
Longitudinal Section
matter more clear.
Capillaries, composed of a single
layer of cells.
Material : i . A common rubber syringe.
2. A glass tube three feet long and seven sixteenths of an inch
outside diameter.
56 PHYSIOLOGY.
3. Four inches of the same size glass tubing, for making connec-
tions.
4. Several nozzles, made of the same size glass tubing, all fine, but
of varying degrees of fineness.
5. India-rubber tubing, twelve feet, three eighths of an inch inside
diameter. This should be black, pure gum, rubber which is more
highly elastic than the other kinds.
6. Three feet of rubber tubing, same size as above.
7. Four inches of white rubber tubing, same size as above, for
making connections.
In all the experiments, have one of the students assist by holding
the outlet tube, so that (i) all the members of the class may see the
stream, and (2) that the stream may be suitably directed, as into a pail
or sink.
Count aloud, to mark the exact time of each compression of the
bulb, so the students can compare this with. the time and duration of
the jets of water.
Be very careful to use perfectly clean water, as any fine particles of
sediment drawn into the tube are likely to clog the fine outlet of the
nozzle. And it is well to take the further precaution not to let the
supply tube touch the bottom of the water-supply dish, as some fine
sediment may get in in spite of previous care.
EXPERIMENT i. Remove the nozzle of the syringe, and put in its
place the long glass tube. Work the syringe, and note that the jet is
jerky, following each contraction of the bulb.
EXPERIMENT 2. Substitute the rubber tube, three feet long, for
the glass tube. On working the bulb the stream will be found inter-
mittent.
EXPERIMENT 3. Take off the rubber tube and replace the glass
tube, adding the nozzle. Here the pressure will be so great that it is
likely to push off the nozzle unless the assistant holds it firmly. It
could be tied on, but this takes more time. On working the bulb,
greater effort must be made on account of the resistance caused by the
narrower outlet.
EXPERIMENT 4. Once more substitute the rubber tube, this time
with a glass nozzle in its end. Now, on working the bulb, resistance
will be felt, and the stream will be constant, or nearly so, and will con-
tinue for some time when the bulb is no longer worked. This is be-
cause the rubber has been stretched, chiefly laterally, and is now
CIRCULATION OF THE BLOOD.
57
" stretching back." That is, by the elastic reaction of the rubber tube
the jerky action of the bulb is converted into the steady flow that we
see. In the first experiment we had a rigid tube and practically no
resistance. In the second, although the tube was elastic, there was no
resistance, so the elasticity was not brought into play. In the third,
there was resistance, but the tube was inelastic. In the fourth, the
resistance brought into play the elasticity of the rubber tube, and the
elastic reaction of the tube continues (so to speak) the action of the bulb
between two successive strokes. In this experiment the pulse can be
felt in the tube.
The Veins. The capillaries, after penetrating the tis-
sues, reunite to form small veins, which in turn reunite
to form larger ones, till finally two great veins, the caval
veins, precaval and postcaval, return the blood to the
heart. The veins, like the arteries, are smooth inside and
elastic (though less elastic than the arteries). They are
thinner than the arteries, and, in consequence, collapse
when the blood flows out of them, whereas the larger
arteries stand open, after they are emptied of blood.
The Valves in the Veins. The only valves in the arte-
ries are those which we have seen at the beginning of the
aorta and pulmonary artery.
Many of the veins have
similar pocket-like valves,
though less strong than
those of the arteries. They
are usually in pairs, but some-
times single or in threes. It
is important to note that they
all have the mouths of the
pockets toward the heart, so
that the blood flows freely
toward the heart, but is prevented from flowing the other
way on account of the filling of the valves by the reflow
Open Shut
Fig. 27. Venous Valves.
58 PHYSIOLOGY.
of the blood stream. When the blood is flowing through
the veins toward the heart the valves lie against the walls
of the veins.
The valves are most numerous in the medium-sized veins,
and especially in the veins of the extremities ; more abun-
dant in the leg than in the arm. Valves are absent from the
caval and some other veins, and from the very small veins.
Illustration of Venous Valves. Make a cloth tube (or take the
lining of a boy's coat sleeve) and sew three patch-pockets on the in-
side, in a circle, i.e. with edges touching each other. Make the pockets
a little " full." Pour sand, shot, or grain through the sleeve first in
one direction and then in the other.
Evidences of Valves in our Veins. With the forefinger stroke
one of the veins on the hand or wrist toward the tips of the fingers.
The veins swell out. The blood meets resistance in the valves of the
vein. Their location may be determined by their bulging out during
the experiment.
Stroke a vein toward the body, and the blood is pushed along with-
out resistance.
Let the left hand hang by the side. Note the large vein along the
thumb side of the wrist. Place the tip of the second finger on this
vein just above the base of the thumb. Now, while pressing firmly
with the tip of the second finger, let the forefinger, with moderate
pressure, stroke the vein up the wrist. It may be seen that the blood
is pushed on freely, but comes back only part way. It stops where it
reaches the valves, filling the vein full to this point, but leaving it col-
lapsed beyond, as shown by the groove. Remove the second finger,
and the vein immediately fills from the side nearer the tip of the fingers.
These experiments show that the blood in the veins moves freely
toward the body, but cannot flow outward to the extremities.
Dissection of the Valves in a Vein. The valves may be seen by
dissecting out the jugular vein (or any other large superficial vein) of a
cat, dog, or rabbit. Split the vein and pin it out on a board.
Effect of Pressure on the Veins. Since the valves in
the veins open toward the heart, any intermittent pressure
on the veins helps to push the blood on toward the heart.
CIRCULATION OF THE BLOOD. 59
The valves are most numerous in the superficial veins and
those of the muscles. The pressure of the muscles during
their action (thickening while shortening) produces pres-
sure on the veins ; and as the muscles act for a short time
only, and then relax, this alternate compression and release
aids very considerably in moving the blood on toward the
heart. It is worthy of remark that this effect is more
pronounced at the time the muscles need the most active
circulation ; namely when they are in action, and are using
the most blood. The heart has power enough to pump
the blood clear around from each ventricle to the auricle
of the other side of the heart ; but this outside aid comes
in good play to relieve the heart at a time when it has an
unusual amount of work to do, as when one is using a
large number of muscles vigorously.
" Every active muscle is a throbbing heart, squeezing
its blood tubes empty while in motion, and relaxing so
as to allow them to fill up anew."
Rate of Blood Flow in the Arteries, Capillaries, and
Veins. The blood flows most rapidly in the arteries,
slowest in the capillaries. Why is this ?
When an artery divides, the two branches taken together
are larger than the one artery that divided to form them.
Stated more exactly, the sum of the areas of the cross-
sections of the branches is greater than the area of the
cross-section before branching. Hence as the blood flows
on it is continually entering wider and wider channels ;
and we are told that the united cross-section of all the
capillaries fed by the aorta is several hundred times that
of the aorta itself.
The Flow of the Blood compared with the Current of
a Stream. If we walk along a stream, we see that the
6o
PHYSIOLOGY.
channel varies considerably in width and depth. Where
the channel is large, whether from increased width or depth,
there the current is slower, but wherever the channel is
reduced, the current is more rapid. So the stream in
the relatively narrow artery is swift. In the capillaries,
Pulmonary Vein -
Left Auricle ..
Left Ventricle
Aorta
Digestive Tube
Pulmonary Artery
Lymph Vein
Right Auricle
- Right Ventricle
Caval Vein
- Liver
Fig. 28. Plan of Circulation. (Dorsal View.)
although any individual channel is small, these channels
all together are wide ; the result is the same whether a
river widens out into a single lake, or divides into a great
number of channels running past innumerable islands.
CIRCULATION OF THE BLOOD. 6l
All the tissues of the body may be regarded as so many
islands lying between the capillary streams.
The Blood Flow in the Veins. When the blood re-
collects in the veins it is entering narrower channels, and
its rate is quickened ; but as the veins are wider than the
arteries, the stream does not enter the heart with the veloc-
ity with which it left that organ. The veins hold more
blood than the arteries, and in dissecting the cat or rabbit
it will be noticed that the arteries are emptied of blood;
that the tissues of most of the organs are fairly free from
blood ; but -that the great veins, such as the caval veins,
are full.
Blood Tubes compared to Two Funnels. If the blood
tubes leaving the heart could all be united, they would be
best represented by a funnel with its tube connected with
the heart. If another funnel were placed with its mouth
to the mouth of the first, their point of union, the widest
point, would represent the capillaries; and if the second
funnel had a wider tube than the first, it would fairly rep-
resent the veins which return the blood to the heart.
Nourishment of the Walls of the Heart and Blood
Tubes. The cardiac (coronary) arteries spring from the
aorta just above the semilunar valves, and send blood into
the muscular walls of the heart ; and these arteries, like
others, divide, forming capillaries, through which the heart
muscle is nourished, The cardiac veins return the blood
to the right auricle.
Influence of Gravity on Circulation. Although the
heart pumps the blood around through the body inde-
pendent of the force of gravity, yet the circulation is influ-
enced by this force. For instance, a person who has
62 PHYSIOLOGY.
fainted should be laid flat on his back, that the heart may
more easily drive blood to the brain. Many persons go to
sleep more readily while sitting than while lying down.
A sore hand feels less pain if held up, as in a sling, than
when hanging by the side, and a sprained ankle does
better rested on a chair, as less blood flows to it. Nearly
every one has noted the pain following the pressure of
blood when a sore hand, or foot, is suddenly lowered.
Experiments illustrating the Effect of Gravity on Circulation.
Let all the pupils in the class stand. Let one arm hang freely by the
side. Hold the other arm straight up as far as the clothing will readily
permit. Observe :
1. The difference in the color of the two hands.
2. The difference in fullness, both in the feeling of fullness and in
the prominence of the veins.
3. The difference in temperature ; place the backs of the hands
against the cheeks.
The position largely determines the amount of blood in the hand,
and the amount of blood determines the temperature, the size, and the
color.
Summary. i. The heart beats about seventy-two times a minute.
2. The pulse is a wave running along an artery.
3. The pulse varies with age, health, food, etc.
4. The heart has two main cavities, one in each half of the heart,
and two independent streams are flowing through it.
5. Valves allow the blood to flow through the heart in one direc-
tion, but prevent a reversal of the current.
6. The heart is a hollow muscle, and. by contraction forces the
blood out into the arteries.
7. The heart works rather less than half the time.
8. The large arteries, by elastic reaction, push the blood on while
the heart is resting.
9. Circular muscle fibers in the walls of the medium-sized arteries
regulate the blood supply to the organs.
10. In the arteries the blood flow is rapid and intermittent, in the
capillaries slow and constant.
CIRCULATION" OF THE BLOOD. 63
11. The thin walls of the capillaries allow the liquid part of the
blood to soak out and nourish the tissues, and to soak back into the
capillaries bearing waste matter.
12. The veins are thin walled, and collapse when empty, while the
arteries are thick walled, and stand open when empty of blood.
13. Arteries carry blood from the heart, while veins carry it toward
the heart.
14. The veins have valves which allow the blood to pass toward
the heart, but not away from it.
15. Any intermittent pressure on the veins aids the blood flow.
1 6. The blood flow is most rapid in the arteries, slower in the veins,
slowest in the capillaries.
17. Gravity influences circulation.
Questions. i. Why do the large arteries lie deep?
2. In which direction should the limbs be stroked to promote circu-
lation?
3. How does slapping the hands around the body warm the ringers?
4. How can a horse or a cow be comfortable with the head down
for a long time?
5. Why are the walls of the left ventricle thicker than those of the
right ?
CHAPTER V.
CONTROL OF CIRCULATION. THE BLOOD AND
THE LYMPH.
The Effect of the Emotions on Circulation. In our
every-day experience we have evidence of the control of
the heart and blood tubes by the nervous system. We know
that certain emotions affect the circulation of the blood ;
for instance, blushing and pallor. Certain emotions may
also quicken or retard the action of the heart. Excessive
grief or joy has produced sudden death by stopping the
beat of the heart.
Let us look a little more closely at that part of the
nervous system that has such intimate relation to the
blood system.
The Rhythmic Action of the Heart. In the first
place, the action of the heart is automatic. The heart of
the frog continues to beat a long time after it is removed
from the body. This is regarded by many as due to the
action of certain ganglia imbedded in the walls of the
heart, especially in the auricles ; while others say that
since the ventricle, in which no ganglia have been found,
may beat independently of the auricles, rhythmic contrac-
tion is characteristic of heart muscle, and that we are, at
present, unable to explain it.
But while the impulses that originate the action of the
heart arise within the heart itself, still the beat of the heart
is constantly modified by nerve impulses reaching it from
without.
6 4
Carotid Plexus
Superior Cervical Ganglion
Middle Cervical Ganglion
Pharyngeal Branches
Cardiac Branches
Deep Cardiac Plexus
Superficial Cardiac Plexus
Solar Plexus
Aortic Plexus
Lumbar Ganglia
Fig. 29. Vertical Section of Body, showing Sympathetic Nerves and Ganglia of Right
Sid,e and their Connection with the Cerebro-spinal Nerves.
66
PHYSIOLOGY.
Sympathetic Nerve Chains
GRAY
Sources of the Heart's Nerve Supply. The heart re-
ceives its nerves from two sources, the sympathetic system
and the vagus (or pneumogastric) nerves.
The Sympathetic Nervous System. The sympathetic
nervous system consists of two rows of ganglia in the body
cavity, one along each side
of the spinal column, re-
ceiving branches from the
spinal nerves, and sending
branches to all the
internal organs of
the body, the
heart and lungs in .
the thorax, and the
stomach, intestines, and the
other organs of the abdomi-
Ganglion of
Dorsal Root
Sympathetic
Ganglion
Fig. 30. Relation of Spinal Cord and
Sympathetic Nervous System (Diagram).
nal cavity. In many places
these nerves form a thick
network called a plexus.
One very large plexus is on the dorsal surface of the
stomach, and is called
spinal cord the solar plexus.
Sympathetic
Ganglion
The Vagus Nerves.
The vagus nerves
are a pair of the cranial
nerves arising from the
sides of the spinal bulb ;
and passing downward,
they give branches to
the pharynx, the gullet,
the stomach, the larynx, the windpipe, the lungs, and the
heart. Now, whatever other function the vagus nerves
Fig- 31. Ideal Cross-section of the Nervous
System. (After Landois and Stirling.)
CONTROL OF THE CIRCULATION.
6 7
may have, they seem to have the power of retarding, or
stopping altogether, the beat of the heart ; and stimulation
of the vagus nerves may make the heart pause in a relaxed
condition. Other nerves may quicken the heart beat, but
the vagi are regarded as a break on the heart's action.
Inhibition. This is a case of inhibition. It is well
known that a severe blow over the stomach may cause one
to faint by stopping the heart. This
is due to reflex inhibition of the heart.
The blow sends a nerve impulse by
fibers of the sympathetic system to
the center in the spinal bulb, and
thence an impulse is taken by the
vagus nerves to stop the heart.
Vaso-constrictor Nerves. In an
experiment with the rabbit's
ear it has been shown that
stimulating the sympathetic
nerve in the neck causes the
ear to become pale. This is
due to the constriction of the
arteries of the ear, because
the nerves have made the
muscle fibers of these arteries
shorten. Such nerve fibers
are called constrictors, or vaso-con-
strictors. They run in the sympa-
thetic nerve, but have their origin
and center in the spinal bulb.
Vaso-dilator Nerves. Other fibers
may cause the opposite effect, namely, dilation, and are
therefore called vaso-dilators. Examples of these may be
Lungs -
Heart
Stomach
Fig. 32.
Diagram of vagus
Nerve.
68 PHYSIOLOGY.
found running to the arteries of the limbs. When the
muscles of any organ, say the legs, act, they need a greater
supply of blood. Now, at the same time that nerve im-
pulses are sent to the muscles of the legs to make the
muscles shorten, impulses are sent along other fibers of
the same nerves to make the arteries dilate, and allow
more blood to flow to these muscles.
Vaso-motor Nerves. The vaso-constrictor and the
vaso-dilator nerves taken together are called vaso-motor
nerves.
Centers of Control of Circulation. The centers of
control of the blood tubes are in the cerebro-spinal nervous
system. There is no evidence that the sympathetic gan-
glia are centers of reflex action.
Blushing. How is it that the face sometimes flushes
so suddenly ? Because of some emotion, you say. But
how does the emotion bring this about ? We have already
learned about the muscles in the wall of the arteries. We
are now prepared to understand that in the normal condi-
tion nervous impulses are acting on these muscles, keeping
them partly shortened, and so keeping the arteries of a
moderate size. Under the influence of certain emotions,
the caliber of the arteries is suddenly enlarged, and hence
the change in color.
The Regulation of the Size of the Arteries. Through
the sympathetic system the blood supply of all the organs
of the body is regulated. Any organ needing more blood
sends a message (nerve impulse) to some nerve center, and
in response nerve impulses are sent to the muscle fibers of
the supplying artery, and the amount of blood sent to that
organ is regulated. For instance, a piece of ice is laid
CONTROL OF THE C/RCULATfOJV.
6 9
upon the skin of the hand. The part becomes pale, as
the arteries have become narrowed. If this action be con-
tinued, there may set in a decided reaction, and the part
become more red than usual, when the reaction has
widened the artery more than it was
before the constriction.
Effect of Exercise on the Size of the
Arteries. As there is only a certain
amount of blood in the body, it is evi-
dent that if one organ receives
an extra supply, some other sympathetic
organ or organs must, for the Ganglions
time, receive less. For in-
stance, one begins to walk vigorously.
The large muscles of the lower limbs
and trunk become active, and they need
more blood. They therefore send mes-
sages to some nerve center (probably in
the spinal cord), and by reflex action the
arteries supplying the lower limbs are
widened, and these muscles receive more
blood. But these muscles make up a
very considerable part of the weight and
bulk of the body. While in action they
take the lion's share of the blood. The
brain, at such a time, would receive less, Fig . 33 . Ventral Vie
and it would be folly to expect the brain
to work at its full capacity while the
blood was called away to other organs.
of Spinal Cord with
Sympathetic Gang-
lions of One Side.
Regulation of the Effects of Exercise. When we ex-
ercise vigorously, the heart beats faster, and this of itself
would tend to increase the blood supply to all organs.
70 PHYSIOLOGY.
But this mechanism for widening the channel leading to
the working organs, while the arteries to the other organs
are made smaller, or at least are not enlarged, solves the
problem of supplying each part according to a greatly
varying need, while not sending too much to a part not
needing it.
EFFECTS OF ALCOHOL ON THE CIRCULATION.
" Alcohol stimulates the heart, producing increased force
and rapidity of the cardiac beat. It thus tends to increase
the blood pressure by acting on the heart, and to increase
the flow of blood from the arteries into the veins. The
effect on the blood pressure is, however, partly counter-
acted by a coincident dilatation of the blood vessels of the
skin, which thus become flushed, and tends to produce
more sensible perspiration." Treatise on Hygiene, STE-
VENSON and MURPHY.
"The warm and flushed condition of the skin which
follows the drinking of alcoholic fluids is probably, in a
similar manner, the result of an inhibition of that part
of the vaso-motor center which governs the cutaneous
arteries." FOSTER.
The control of the muscles in the walls of the arteries
being thus interfered with, the circular muscles are no
longer made to shorten, and the artery dilates, thus allow-
ing more blood to flow into it.
We may thus account for the flushing of the skin of the
face, which in many individuals quickly betrays indulgence
in alcoholic drink. If this flushing is too often repeated,
the arteries gradually "lose tone," and the condition be-
comes permanent. The circulation in the whites of the
eyes may be affected, making them "bloodshot."
CONTROL OF THE CIRCULATION. ?1
Similar congestion occurs in the mucous membrane of
the stomach from the presence of alcohol, which may
become a permanent inflammation followed in time by
very extensive changes in appearance and function. It
is said that most of the alcohol swallowed is absorbed
directly from the stomach, and hence the intestines are
not so directly affected.
Good authorities state that alcohol arrests the develop-
ment of the corpuscles. It diminishes the size, alters the
form, and reduces the number of the corpuscles. Since
the work of the blood corpuscles is so important this
reduction in their number and efficiency must very
appreciably affect the nutrition of the body as a whole.
When the blood is " out of order" the body is out of order.
The Blood, The blood is composed of a clear liquid,
the plasma, and the blood cells, or corpuscles. In a drop
of blood under the microscope the plasma occupies the
clear spaces between the corpuscles. The corpuscles
make up one third of the bulk of the blood, and the
plasma two thirds.
Microscopic Examination of the Blood. To get a drop of blood
from the finger, wind a cord around the finger, beginning at the base,
drawing the cord moderately tight, until the last joint is reached. By
this time the end of the finger is usually well distended with blood.
With a clean needle make a quick, sharp, light puncture near the base
of the nail ; this ordinarily brings a small amount of blood. Put a
small drop on each of several slides and quickly cover with coverslips.
Examine with a high power.
The Colored Corpuscles. These are often called the
red corpuscles. But while in the mass they give the
blood a red appearance, individually they are faint yellow-
ish red. In shape they are seen to be circular disks, hol-
lowed on each side like a sunken biscuit. As they are
72 PHYSIOLOGY.
hollowed on both sides they are more accurately described
as biconcave. These corpuscles tend to gather side by
side, in rolls, like coins. They are cells without nuclei.
The Colorless Corpuscles. In the open spaces be-
tween the rolls of colored corpuscles may occasionally
be found some spherical corpuscles. They are usually
White Corpuscles
HIGHLY MAGNIFIED
White Corpuscle
Red Corpuscles
in Rolls
MODERATELY MAGNIFIED
Fig. 34. Red and White Corpuscles of the Blood.
called the white corpuscles, but are better designated as
the colorless corpuscles, since the others have only a slight
color, and these have none. They usually have a dotted
appearance. It is not so easy to distinguish the two kinds
of corpuscles as it is in the case of the frog's blood, for the
two kinds are more nearly of the same size in the human
CONTROL OF THE CIRCULATION. 73
blood ; and, further, when the colored corpuscles of human
blood are seen flatwise they present a circular outline, while
the frog's colored corpuscles are elliptical. But with a
little study the two may be distinguished. As in the frog's
blood, the colorless corpuscles have ameboid movements,
though they are not very marked unless the blood be
warmed to about the temperature of the human body.
Flexibility and Elasticity of the Corpuscles. It will
be well here to examine again the frog's web. (See p. 54.)
It will occasionally be seen that when one of the colored
corpuscles is pressed against an angle at the forking of
the blood stream, it is sometimes bent, and that as soon as
the pressure is discontinued the corpuscle springs back to
its former shape, showing that it is elastic.
Frog's Blood. A drop of frog's blood, mounted as the human blood
was, will be helpful, as there is a very decided difference in the size and
shape of the colored and colorless corpuscles. Further, the colorless
corpuscles of the frog will show ameboid movements, i.e. slow changes
of form, if watched a while.
The Plasma. The plasma consists chiefly of water,
having in solution various salts, including common salt;
it also contains the nourishing materials for the tissues.
These nourishing materials, obtained from the food by
digestion, consist chiefly of proteids, fats, and sugar. The
plasma also contains waste matters, from the working
tissues, on their way out of the body. How the food is pre-
pared for the building of tissue, and how the waste matter
is removed from the body, we shall study a little later.
The Color of Blood. The difference in color of an in-
dividual corpuscle and the blood in the mass may be better
understood by comparing it with something that we see
more frequently. A tumbler of currant jelly has a rich,
74 PHYSIOLOGY.
red color, but a thin layer of the same jelly, as when one
takes a spoonful on a plate, has a pale color, more yellow-
ish. The colorless plasma with the colored bodies in it
may be compared to a glass dish filled with cranberries
and water.
Hemoglobin. The coloring matter in the blood, then,
is wholly in the colored corpuscles. Examination of these
corpuscles shows that their color is due to a substance
called hemoglobin. There is a small amount of iron in
the hemoglobin, and the presence of this small quantity
of iron appears to be essential to give the blood its color.
When we come to the study of respiration we shall see
that the hemoglobin in the corpuscles is the chief agent in
picking up the oxygen from the air in the lungs and carry-
ing it to the tissues in the body.
The Coagulation of Blood. When the blood escapes
from its natural channels it usually changes from a liquid
to a jelly-like condition. This is known as coagulation.
It is due to the formation of threads of fibrin from the
plasma. These threads of fibrin entangle and inclose the
corpuscles, and the two constitute the clot, or coagulum, as
it is more technically termed. The liquid that afterward
separates from the clot is the serum, and differs from the
plasma only in the removal of the fibrin, which is exceed-
ingly small in quantity, though of great importance in its
action. Many experiments have been made, and much
has been written about the coagulation of the blood, and
perhaps its real cause is not yet clear. But we know that
the coagulation often serves to stop the flow of blood from
wounds, and this is its main use.
Fibrin. If freshly drawn blood be stirred rapidly with
a bundle of wires (perhaps the most convenient stirrer is
CONTROL OF THE CIRCULATION. ?$
a little roll of wire screen), there will soon collect on the
wires a stringy substance. Thorough washing will soon
leave this colorless. It is fibrin. If the stirring has been
done thoroughly, the blood will no longer clot, no matter
how long it may stand.
Liquid Blood and Coagulated Blood. The following
scheme shows the difference between the liquid blood and
the coagulated blood :
( Serum
Liquid Blood
Plasma.
( Fibrin
Clot
Coagulated Blood.
Corpuscles
Amount of Blood. The blood constitutes about one
thirteenth of the weight of the body. In a body weighing
one hundred and fifty pounds this would be about six
quarts.
Chemical Reaction of Blood. Blood is alkaline.
Specific Gravity of Blood. Blood is somewhat heavier
than water, owing to the salts and other matters dissolved
in it.
Quantity of Blood in Different Organs (approximately).
i. One fourth is in the heart and the larger arteries
and veins (including those of the lungs).
2. One fourth in the liver.
3. One fourth in the skeletal muscles.
4. One fourth in the other organs.
The Lymph Spaces. We have seen that the capillaries
have very thin walls. Through their walls part of the
plasma of the blood soaks out, and is then called lymph.
It passes into irregular cavities in the tissue called lymph
spaces. Most of these lymph spaces are minute chinks or
76 PHYSIOLOGY.
crevices in the connective tissues of the different parts of
the body.
The Lymph Tubes. Opening out of the lymph spaces
are irregular passage ways called lymph capillaries, and
these lymph capillaries are continuous with thin-walled
tubes, the lymph tubes. These lymph tubes might be
called the lymph veins, since they join still larger tubes,
closely set with valves, similar to those of the veins. But,
unlike the blood veins, the lymph veins do not gradually
increase in size by confluence. They suddenly form a
large tube, the receptacle of the chyle, beginning in the
upper part of the abdomen. This tube soon narrows and
passes through the diaphragm, close to the spinal column,
and up along the column near the aorta, and empties into
the veins of the neck at the junction of the left jugular
and left subclavian veins. This tube is the thoracic duct,
or the main lymph duct. It has numerous valves, and,
like some of the smaller lymph veins, it presents a beaded
appearance, due to the filling and bulging out of the valves.
In the right side of the neck is a short right lymph duct
which receives lymph from the right side of the head,
neck, and thorax, and from the right arm. The lymph
tubes, as a whole, are usually called the "lymphatics."
Lymph Spaces in the Frog. In dissecting the frog, the looseness
of the skin is very noticeable. The large spaces under the skin are
lymph spaces. Sometimes considerable lymph is found here, so that
in holding up a frog the sagging of the skin from the weight of the
lymph may be easily seen.
Valves at the Mouth of the Lymph Tubes. There
are valves where these lymph ducts empty into the veins
which prevent any reflow of liquid into the ducts, but allow
the lymph to pass freely into the veins.
CONTROL OF THE CIRCULATION. 77
Muscle Fibers in the Walls of the Lymph Tubes. -
There are plain muscle fibers in the walls of the lymph
ducts.
Lymphatic Glands. In its course the lymph passes
through many kernel-like masses, the lymphatic glands.
Lymph contains corpuscles which are considered identical
with the colorless blood corpuscles. It is tho'ught that
these corpuscles are formed in the lymphatic glands.
The Flow of Lymph. The flow of lymph is partly
due to the blood pressure in the capillaries ; this pressure
is caused by the heart. (In the frog there are two small
hearts, not, however, near the blood-pumping heart,
and these pump the lymph along.) In our bodies the flow
of lymph is largely aided by any pressure that may be
brought to bear on the lymph veins ; for, on account of
the valves, as in the blood veins, any pressure must push
the liquid toward the heart. Thus the action of the mus-
cles in the limbs, in the chest, in the abdomen, in the
movements of breathing, and in the bending of the body,
etc., all help in this flow, which is always, probably, very
much slower than that in the blood veins.
Relations of Blood Flow and Lymph Flow. It will
now be seen that while the blood leaves the left ventricle
by one tube, the aorta, it returns to the right auricle, not
merely by the two caval veins, but that a part of the blood
(i.e. of the liquid part of it) does not return by blood veins,
but having left the blood system proper through the thin
walls of the capillaries, it is brought back to the heart by
the lymph veins, which, however, join the blood veins just
before they empty into the heart. There is, in other
words, only one set of distributing tubes, but there are two
sets of collecting or returning tubes.
PHYSIOLOGY.
Left Jugular Vein
Mouth of Lymph ...
Vein
Right Lymph Vein
Right Subclavian
Vein
Precaval Vein
Postcaval Vein
Main Lymph Vein
(Thoracic Duct)
Lymph Capillaries
Blood Capillaries
Fig. 35. Diagram of the Circulation of Blood and Lymph (Dorsal View).
CONTROL OF THE CIRCULATION.
79
Lymph
Capillary
The Lymph. Lymph is a clear liquid. (Chyle and
the lacteals will be considered when we study digestion.)
It is more watery than the blood plasma, but contains a
share of all its nutritious substances. Lymph may be
defined as "diluted blood minus red corpuscles." The
blood proper never reaches the tissues.
The Cells of the Body live in Lymph. The cells
of the tissues are bathed in the lymph which fills the
spaces in the connective
tissue (and we have seen
that the connective tissue
pervades nearly all the tis-
sues of the body), as water
may fill the spaces left
between stones built into
a wall. The cells get all
their nourishment from the
lymph, and into the lymph
they throw all their waste
matter. Each cell may be
compared to an individual
ameba, which lives in
water, and takes all its
nourishment from that
water, and throws all its
waste product into the
same water. As water is
the medium in which the
ameba lives, so we may say lymph is the medium in
which the cells of the body live.
Cells of the Body Aquatic. The cells of the body,
i.e. all the active, working cells, may, therefore, be said
Oxygen
Food
Water
Other
Wastes
Fig. 36. Relation of Blood and Muscle.
(Lymph being Middleman.)
80 PHYSIOLOGY,
to live an aquatic life, and only dead cells, as of hair,
epidermis, etc., live in air. We might also say that not
only the human body, but all animal life is aquatic.
Importance of Lymph. We can see that the move-
ment and renewal of lymph are as necessary as the circu-
lation of the blood itself ; is, in fact, the most important
part of it.
Lymph Cavities or Serous Cavities. We have noticed
the pericardial liquid. There is also a small quantity of
similar liquid around the lungs in the pleural cavities, and
in the abdominal or peritoneal cavity, around the digestive
organs ; also in the cavities of the brain. The liquid in
each case is lymph, and these cavities, often called serous
cavities, are lymph cavities. They communicate with the
lymph tubes.
Dropsy. In health the amount of the liquid in these
cavities is small, but in certain disorders it may accumu-
late. In general, such affections are called "dropsy."
The lymph may also accumulate in the tissues of the
extremities, causing swelling of the limbs.
Variation in the Composition of Lymph. It is evi-
dent that the materials needed by the cells of the different
tissues are not the same. So, as one tissue takes certain
materials and another tissue others, it is clear that the
lymph will not be of quite the same composition in the
different parts of the body. This difference is further
due to the difference in the waste products thrown out
by the different cells. Hence the composition of the
blood varies considerably in different regions. But the
lymph from all the tissues unites with the blood from all
the tissues in the right heart, and on their way to it in the
CONTROL OF THE CIRCULATION. 8 1
larger veins. So the constant slight differences in com-
position of the blood and lymph in the various tissues are
counterbalanced by the mingling of the currents from
these various parts in the large arteries and veins.
The Spleen. The function, or functions, of the spleen are not well
understood. It is believed to have something to do with the renova-
tion of the blood, perhaps forming colorless corpuscles and destroying
colored corpuscles. At any rate, the physiologists generally call it a
blood gland. It is unlike true glands in that it has no duct, and forms
no secretion to be poured into any cavity, like the glands of excretion
and secretion. It has been found, in the case of accidents to man, and
by experiment on the lower animals, that life may continue after this
organ has been removed.
Massage. A system of pressing, rubbing, and knead-
ing the muscles is known as massage. It helps the flow
of the blood and lymph, thus aiding in washing out the
waste products from the muscles and other parts of the
body that are to be reached by pressure. We have seen
that one of the benefits of exercise is to promote the cir-
culation of the blood and of the lymph, and so to help
get rid of the waste matters that are produced by the
activity of the various organs. Many invalids cannot take
active exercise. So this passive exercise may very fairly
take its place, and assist in the nutrition of the tissue by
accelerating the flow of blood and lymph, bringing new
nourishment and carrying away wastes. For students
who do not take sufficient exercise it is a good thing to
rub the body thoroughly and briskly, not only after a
bath, but often with the hands or with a dry towel.
Transfusion of Blood. Transfusion of blood is the transfer of
blood from the blood vessels of one animal to those of another. Trans-
fusion may be direct or immediate, as when the blood vessels of the
two animals are connected by tubing so that the blood passes from one
to the other without exposure to the air ; in indirect or mediate trans-
82 PHYSIOLOGY.
fusion the blood is first drawn into a receptacle. In indirect transfusion
the blood is often defibrinated before transference. The blood may be
introduced either into an artery or a vein; if -into a vein it is sent in
the direction of the natural flow, i.e. toward the heart ; if into an artery,
in either direction. Soon after the discovery of the circulation of the
blood the operation of transfusion began to be practiced, and high
hopes were indulged in as to its value. But it was soon found to be
attended by so much danger that it is now seldom used. It is resorted
to (i) after great loss of blood, (2) after some forms of poisoning part
of the blood is withdrawn and replaced by fresh blood, and (3) in
certain disordered conditions of the blood. The chief dangers are (i)
the introduction of air which forms minute bubbles and stops the blood-
flow in the capillaries, (2) the introduction sometimes causes coagula-
tion within the blood vessels, and (3) the serum of the introduced
blood sometimes destroys the corpuscles of the blood to which it is
added. In the earlier practice lamb's blood was employed, but now
when transfusion is practiced on man only human blood is used. It
has been found safer and better after great loss of blood from hemor-
rhage, to introduce a salt solution of about the natural degree of salt-
ness of the blood ; this restores the normal volume of circulating liquid,
and avoids most of the dangers except that of introducing air. The
numerous fatal results of this operation have shown that it should not
be resorted to except in cases of extreme necessity.
For directions about stopping the flow of blood from
wounds see Chapter XXIII. and the books named below.
READING. Prompt Aid to the Injured, Doty ; Emer-
gencies, Dulles; Emergencies, Howe; First Aid to the
Injured, Lawless; First Aid to the Injured, Morton; First
Aid in Illness and Injury, Pilcher ; Sickness and Accidents,
Curran.
What other process keeps pace with the coursing of the
blood through the body, being its running mate, so to
speak ?
Summary. i . Blushing, and other variations in blood supply, are
under the control of the sympathetic nervous system.
2. The sympathetic nervous system consists of two rows of ganglia
CONTROL OF THE CIRCULATION". 83
in the body cavity near the spinal column, with fibers running to the
internal organs. It is also connected with the cerebro-spinal nervous
system.
3. The heart beat is automatic and rhythmic.
4. The heart beat is regulated by the sympathetic nervous system
and by the vagus nerves.
5. The blood consists of a liquid, the plasma, in which float the
colored and colorless corpuscles.
6. When blood is shed it coagulates, tending to check its own
escape.
7. Lymph is like the blood diluted and lacking the colored cor-
puscles.
8. A set of lymph tubes conveys the lymph into the veins to join
the flow toward the heart.
9. In its course the lymph passes through the lymphatic glands.
Questions. i. What makes the hands grow red and puff up on
sitting in a warm room after snow balling?
2. How is a mustard plaster effective?
3. Why does light exercise before retiring promote sleep?
4. Why are the feet often cold after studying?
5. How does the application of ice, or cold water, relieve head-
ache?
6. Why should the clothing be changed after getting wet?
7. What is the meaning of humor, in the expressions "good-
humored," " bad-humored " ? Have these expressions a real physio-
logical significance?
CHAPTER VI.
RESPIRATION.
The Close Relation between Circulation and Respira-
tion. Is it not a very striking fact that we take one
breath for every four heart beats ? That whatever quick-
ens the breathing also quickens the heart, so that the two
always keep in al-
most the same ratio ?
Let us learn what
are the many inti-
mate relations of
the blood pump and
the air pump, the
blood system and
the air system, of
Circulation and Res-
piration.
The Organs of
Respiration.
1 . The lungs and
air tubes.
2. The structures
which increase and diminish the size of the chest, princi-
pally the diaphragm, and the muscles acting on the ribs.
The Parts of the Lungs. i. The Air Vesicles, an
immense number of small sacs, which communicate with
8 4
Fig. 37. The Trachea and Bronchial Tubes, showing
Two Clusters (Alveoli)) of Air Vesicles.
RESPIRATION.
1. Pulmonary Orifice
2. Aortic Orifice
3. Left Auriculo-Ventricular Orifice
4. Right Auriculo-Ventricular Orifice
The heavy black line between the heart and the liver represents the diaphragm.
Fig. 38. Front View of the Thorax. The Ribs and Sternum are represented in
Relation to the Lungs, Heart, and other Internal Organs.
86
PHYSIOLOGY.
the outer air by the bronchial twigs, the bronchi, and the
trachea.
2. The Pulmonary Capillaries, forming a thick network
around and between the air sacs. These capillaries receive
their blood from the pulmonary artery, and return it to the
heart by the pulmonary veins.
Elastic Tissue in the Lungs. The air vesicles, with
their supplying air tubes and their surrounding blood tubes,
are bound together by elastic tissue, which fills up most of
the intervening space.
The Windpipe or Trachea. The windpipe has in its
walls C-shaped cartilages, with the open part of the C
on the dorsal surface. These cartilages continue in the
bronchi, and so on until in the smaller twigs they finally
disappear. The cartilages are held together, and the
dorsal gap of the cartilages (the gap would be like that of
a series of horseshoes piled one on top of another) bridged,
by tough fibrous tissue, with much elastic tissue, and
with plain muscle fibers ; the plain muscle fibers are very
abundant in the smaller air tubes.
The Mucous Membrane. The lining of the trachea
is a mucous mem-
brane. It pours
out on its surface
a substance some-
what like white of
egg, called mucus.
This keeps the air
Fig. 39. Ciliated Cells lining the Air Tubes (x 300).
particles of dust that are in the inspired air. There is a
constant slow current of mucus toward the throat, whence
it is, from time to time, hawked up.
RESI'IRATION.
Cilia. This current of mucus is caused by the cilia
projecting from the lining cells of the trachea. They are
little hairlike projections, in countless numbers, like a field
of grass, each stalk having the power of bending back and
forth, making a quick stroke toward the throat, then a
slower recover stroke. Thus the united wavelike action
of the myriads of lashing cilia paddles the mucus head-
ward. It is a very common error to suppose that the cilia
produce air currents. This is not their function, and it
can readily be seen that they cannot create currents of air,
as they are wholly submerged, like grass growing on the
bottom of a shallow pond of slimy water.
Location of Mucous Membrane. All the cavities and
passages in the body to which the air has access, such as
the digestive and respiratory passages, etc., are lined by
mucous membrane (not all
Ciliated). Trachea
The Pleura. The out-
side of each
lung is cov-
ered by a thin
adherent mem-
brane, the pleu-
ra, which com-
pletely invests
it, except at the root of
the lung, where the bron-
chus and blood tubes
enter. Here the pleura
turns toward and adheres to the inner wall of the chest,
forming its lining (still called the pleura), and below passes
over the anterior surface of the diaphragm. The lung is
Pleural Space
(Exaggerated)
Chest Wall- -
~~~ Pleura
Fig. 40. Diagram of the Lungs and Pleurae.
88 PHYSIOLOGY.
thus free, except at its root, where the air and blood tubes
enter. A very small quantity of liquid moistens the con-
tiguous surfaces of the pleurae on the outside of the lung
and the inside of the chest wall, so they move easily one
upon the other during respiration. As the lungs are
always distended enough to fill the chest cavity, these two
surfaces are always in contact. In pleurisy (inflammation
of the pleurae) pain is felt in breathing from friction or
adhesion of these surfaces.
Important Facts concerning Respiration. In study-
ing respiration, let us constantly keep in mind these
facts :
1. The lungs are highly elastic, and
2. Highly porous, each air vesicle being in direct com-
munication with the outer air by means of
3. Air tubes that always stand open
4. And are always moist internally.
5. The pulmonary capillaries closely invest each air
vesicle.
6. The lungs are always expanded enough to fill all
the space in the chest not occupied by other organs, and
7. Freely movable, except at the place of entrance of
the bronchi and blood tubes.
8. The smooth, moist pleurae.
The Diaphragm. The diaphragm is a thin muscle
making a complete partition between the abdominal cavity
and the chest cavity. It is convex anteriorly, concave pos-
teriorly ; its ventral border is attached to the inside of the
chest wall about opposite the lower end of the breast bone,
thence obliquely along the border of the ribs (as felt in
front), and the dorsal attachment is posterior to the ventral
RESPIRATION'.
8 9
attachment. Its general position is shown in Figs. 38, 40,
and 43.
To show the Action of the Diaphragm and Lungs. MATERIAL.
Bell jar with stopper, sheet of rubber large enough to cover the mouth
of the jar, toy rubber balloon, cork (rubber preferred), glass tube, strong
rubber band (such as boys use for slung shots), marble.
Triangularis Sterni
Internal Mammary Vessels v.
Left Phrenic
Nerve
Pleura
Puimonalis
Pleura Costalis
Mediastinum \ Sympathetic Nerve
( Thoracic Duct
Vena Azygos Major I p osterior
, Pneumogastric Nerves )
Fig. 41. A Transverse Section of the Thorax, showing the Relative Position of the
Viscera and Reflections of the Pleurae.
PREPARATION. Lay the marble on the center of the sheet of rub-
ber, double the rubber over it, stretching the rubber strongly over the
marble, and tie the marble firmly in its place. Stretch the sheet of
rubber over the mouth of the jar with the projection made by the marble
on the outside, and fasten with rubber band. Bore a hole in the cork,
90 PHYSIOLOGY.
and fix the glass tube snugly in it, so that the lower end of the tube will
extend about half-way down the jar. Tie the balloon on the lower end
of the glass tube.
EXPERIMENT i. Inflate the balloon. Consider that it requires
some expenditure of energy to do this. When the mouth is taken away
from the tube the balloon immediately collapses.
EXPERIMENT 2. Insert the balloon and tube into the jar, but do
not cork, and repeat Experiment i. The same results as before are
noticed, and it will further be seen, or rather heard and felt, that when
the balloon is inflated some air comes out of the jar around the tube,
and when the balloon collapses air again enters the jar.
EXPERIMENT 3. Again inflate the balloon, and while it is inflated
tightly cork the jar. If all the parts fit well, the balloon should now
remain inflated. This may at first seem strange, as the mouth is taken
away from the tube, and the tube left entirely open to the air. But it
will be seen that to just the extent that the balloon contracts, so much
more space is left in the jar outside the balloon. This means diminished
pressure, and the pressure of the outer air presses the diaphragm up,
and keeps the balloon partly distended, maintaining equilibrium.
EXPERIMENT 4. Pull the diaphragm down, using the marble as a
handle. This shows the expansion of the lung by the pressure of the
external air when more space is given by the depression of the dia-
phragm. On releasing the diaphragm, it springs upward, and the
balloon becomes reduced in size, driving out part of the air that was in
it. This shows how expiration is accomplished, so far as the diaphragm
is concerned.
If a bell jar be not at hand, a lamp chimney or a quart bottle may be
used, after cutting off the bottom, as follows : File a deep notch across
near the bottom ; heat an iron rod, and apply the end of it to one end
of the notch, and slowly draw the rod around to the other end of the
notch (the rod may need to be reheated) . After cracking off the bot-
tom of the jar, file the edges so they will not cut the rubber.
Let each pupil make a drawing, showing the position of the parts in
inspiration and in expiration.
Illustration of the Minute Anatomy of the Lung. To illustrate
the minute anatomy of the lung, take a rubber balloon, a glass tube,
two rubber tubes, one dyed red, the other blue, a bag of netting, with
one side dyed red and the other side blue. Tie the balloon on the end
of the glass tube, slip the bag of netting over the balloon and tie it,
RESPrRATION. 91
with the ends of the rubber tubes on the corresponding sides of the
bag. Slip a short piece of the rubber tube on the end of the glass
tube, and when the balloon is inflated shut the air in by means of a
CILIA I i.BRONCHIAL TUBE.
Fig. 42. Minute Structure of the Lungs, showing Air Vesicles
and Capillaries.
pinchcock. The balloon represents an air vesicle, the glass tube a
bronchial twig, the blue tube a subdivision of the pulmonary artery,
the netting the capillaries around the vesicle, and the red tube one of
the branches of the pulmonary veins.
The Movements of Respiration. The process of res-
piration consists of two acts, inspiration and expiration.
Two Active Forces in Inspiration. In inspiration
the principal active forces in the body are, first, the dia-
phragm ; and, second, the muscles which elevate the ribs.
Work of the Diaphragm in Inspiration. The dia-
phragm is a muscle, and when its fibers shorten, the dia-
phragm is pulled down. In moving down it presses on
the abdominal organs, and makes the abdomen protrude
laterally and ventrally. This lowering of the diaphragm
increases the space in the chest ; the air already in the
PHYSIOLOGY.
chest expands to fill this greater space. When expanded
it exerts less pressure than before, and the air outside,
having greater pressure, enters till equilibrium is produced.
The air enters through the trachea, presses on the inside
of the elastic lungs, and makes their bases extend, follow-
ing the diaphragm in its descent. The bases of the lungs
remain in contact with the upper surface of the diaphragm
all the time.
. . . . Increased Air
Space
Inspiration Expiration
Fig. 43. Diagrammatic Sections of the Body in Inspiration and Expiration.
Work of the Chest Walls in Inspiration. Certain
muscles of the chest wall elevate the, ribs and breast bone.
This act widens the chest, and the air, as before, presses
in through the open trachea, and keeps the sides of the
lungs in contact with the inner surfaces of the chest walls.
Effort required in Depressing the Diaphragm. -
Inspiration requires considerable effort, because the dia-
RESPIRATION. 93
phragm in its descent presses upon the elastic organs of
the abdomen (stomach, liver, etc.), and these organs, in
turn, are pressed against the elastic walls of the abdomen.
It is somewhat like pressing a pillow down into a rubber
bag; the pillow springs up as soon as the pressure is
stopped, because of its own elasticity as well as that of the
bag. Therefore, as soon as the diaphragm relaxes, the
elastic walls of the abdomen retreat, and the abdominal
organs rise to their former place.
Effort Required in raising the Ribs. When the ribs
are elevated, the cartilages which connect the ventral ends
of the. bony parts of the ribs with the breast bone are
slightly bent. When the muscles relax, the elasticity of
the rib cartilages helps to bring the ribs back to their
former position, thus reducing the chest to its former
width.
Expiration Easy. Thus we see why expiration is easy ;
in fact, " does itself " (in ordinary respiration) by elastic
reactions. But inspiration is harder than it would be if it
were not for the fact that the descent of the diaphragm
meets resistance, and the ribs, in rising, have to overcome
resistance in bending the costal cartilages, and in raising
the weight of the chest walls and shoulders.
Potential Energy stored in a Door Spring. When
one opens a door that has a spring to shut it, he has to
expend more energy to open the door than he would if he
did not have to bend (twist or compress) the spring at the
same time. But no effort is needed to shut the door. The
door was opened and shut at the same time ; i.e. when
the door was opened force was stored in the spring (in the
form of what is called potential energy), and this stored
energy shuts the door while we pass on. We can better
94 PHYSIOLOGY.
afford to employ more energy while opening the door than
to take the extra time to shut it. If, then, a door with such
spring were fastened open, it might remain open for a long
time. When released it flies shut. If one, in this case,
asks, "Who shut the door?" the answer is, "The person
who opened it."
The Storing of Energy during Inspiration. So in
the act of inspiration we perform a double work in storing
energy by which the expiration is performed without active
muscular effort.
Review of Forces of Respiration :
FORCES OF INSPIRATION.
1. Depression of the diaphragm.
2. Muscles elevating the ribs.
3. Pressure of the external air.
RESISTANCES TO INSPIRATION.
1. Compression of the abdominal organs and stretching
abdominal walls.
2. Bending the rib cartilages and lifting the chest.
3. Stretching the lungs.
ELASTIC REACTIONS OF EXPIRATION.
1. Elastic reaction of the abdominal walls and contents.
2. Elastic reaction of the rib cartilages.
3. Elastic reaction of the lungs.
Forced Respiration. Thus far we have been speaking
of ordinary respiration. In forced respiration, as in shout-
ing, many muscles are brought into play to expel the air
rapidly and forcibly. In such an act as coughing there is
vigorous action of the abdominal muscles.
RESPIRATION. 95
Abdominal and Thoracic Respiration. The main part
of respiration is performed by the diaphragm, and the more
common mode of respiration is therefore called abdominal
or diaphragmatic respiration. In women of the civilized
races respiration is more largely accomplished by the action
of the thoracic muscles, and is called thoracic or costal res-
piration. In children the respiration is of the abdominal
type.
The Rate of Respiration. The rate of respiration in
the adult varies from sixteen to twenty-four per minute,
the average being about seventeen times a minute ; about
one respiration for every four heart beats. Light is favor-
able to respiratory activity. The rate is affected by the
position of the body, state of activity, temperature, diges-
tion, emotions, age, disease, etc. Ordinary inspiration
takes slightly less time than expiration.
Modifications of Respiration. Coughing is a forcible expiration,
usually directed through the mouth, and for the purpose of getting rid
of some foreign substance, or caused by irritation. In sneezing there is
first a deep inspiration, and then the current of air is forced out, chiefly
through the nose. Sneezing may be prevented by pressing firmly on
the upper lip. Crying, laughing, sobbing, are modifications of respira-
tion connected with certain emotions. Yawning and sighing are deeper
breathings, caused by etijiui, depressing emotions, or a deficient ventila-
tion. Hiccuping is sudden inspiration, produced by spasmodic action
of the diaphragm, accompanied by sudden closure of the glottis, and is
often caused by some disorder of stomach digestion. Snoring is caused
by breathing through the mouth and setting the soft palate into vibra-
tion. Sniffing is sudden inspiration : the diaphragm is suddenly pulled
down, the air in the nasal cavity is thus drawn downward, and the air
we wish to test, or the odor we wish to inhale, is thus drawn into the
upper nasal cavities ; whereas in ordinary inspiration most of the air
passes along the lower part of the nasal passage. In hawking, the air
is forced out through the narrowed passage between the root of the
tongue and the soft palate to remove mucus. Gargling is forcing air up
96 PHYSIOLOGY.
through liquid held between the tongue and the soft palate. Panting,
whistling, blowing, spitting, sucking, and drinking are also modifica-
tions of respiration. In case of choking it is well to hold the head for-
Jl
II
COMPLEMENTAL AIR.
120 CUBIC INCHES.
AIR THAT CAN BE BUT SELDOM IS TAKEN IN.
TIDAL AIR. 20 to 30 Cubic Inches Air Taken in
and Sent out at Each Breath.
RESERVE AIR.
100 CUBIC INCHES.
m
^ AIR THAT CAN BE BUT IS SELDOM DRIVEN OUT.
RESIDUAL AIR.
100 CUBIC INCHES.
AIR THAT CANNOT BE DRIVEN OUT.
II
t>
-S -S
-
QU
Figr. 44. Diagram of Lung Capacity.
ward, and perhaps downward. A smart slap between the shoulders
sometimes helps dislodge anything stuck in the throat, and it may be
necessary, in addition, to hold a child with its head downward.
RESPIRATION. 97
Capacity of the Lungs. Have the class stand, and each pupil raise
his right hand.
1 . Tidal Air. Let all breathe together, at the ordinary rate and
depth, and let the hand rise about three inches during inspiration, and
fall again during expiration. The amount of air taken in at an ordinary
breath is from 20 to 30 cubic inches, or about a pint. This is called
tidal air.
2. Complemental Air. As before, let the hand go up and down
with the breathing, but at the end of the third inspiration, instead of
stopping with the usual amount, keep on breathing in as much as pos-
sible, letting the hand rise accordingly. This air that can be taken in
above the ordinary breath is called the complemental air, and it is
estimated to be, on the average, about 120 cubic inches.
3. Reserve Air. Begin as before, and at what would be the end
of the third expiration continue to drive out as much air as possible,
indicating the degree by correspondingly lowering the hand. This air
that can be breathed out beyond the ordinary expiration is called the
reserve air, and is reckoned at about 100 cubic inches.
4. Residual Air. The air cannot all be breathed out. The re-
mainder is called the residual air, and is computed to be about 100
cubic inches.
The Vital Capacity. All the air that can be breathed out after a
full inspiration, i.e. the sum of the complemental, tidal, and reserve
air, would be about 240 to 250 cubic inches, and is called the vital
capacity. Of course these figures represent only the average of cer-
tain experiments and observations. By practice any one can con-
siderably increase his vital capacity.
A Test of the Capacity of the Lungs. A simple method of
measuring these stages of respiration is to take a gallon bottle and
first carefully graduate it to pints by pouring in water and marking on
the outside with a file. Then invert the bottle in a trough of water,
and inhale from it by means of a rubber tube. Or fill the bottle, in-
vert in water, and exhale into it.
Hygiene of Breathing. Those persons who take con-
stant exercise in the open air are likely not to suffer much
from deficient respiration. But persons following seden-
98 PHYSIOLOGY.
tary occupations, such as that of the student, not calling
for deep breathing (and often the air taken in is of poor
quality), need to pay especial attention to the matter.
Breathing through the Mouth. We should breathe
through the nose, and not through the mouth. The nasal
passages are fitted for the introduction of the air (i) by
being narrow, but of large area; (2) by having their lining
membranes richly supplied with blood ; (3) by the abun-
dant secretion of mucus by this membrane. The air,
coming through this narrow channel, is warmed, and a
large part of any dust it may contain is caught by the
sticky mucus that covers all the walls of this passageway.
If we breathe through the mouth (especially out of doors
in cold weather), the air may not be sufficiently warmed
before entering the lungs, and much more dust would be
carried into the lungs. Then, too, the air has a drying
effect on the throat, whereas the mucus of the nasal pas-
sages will moisten the air as it enters. The cilia, which
extend from most of the cells lining the respiratory pas-
sages, are constantly causing the mucus to slowly flow
toward the external opening, so a good share of the dust
is gotten rid of. A further advantage of breathing through
the nose is that we detect odors, and can thus judge of the
quality of the air.
Breathing and Circulation. The fact has been noted
that breathing directly aids the circulation of the blood.
This is due to the way air pressure is made to affect the
large veins. Breathing also may very considerably aid
the flow of lymph. Every deep inspiration brings pres-
sure to bear on the main lymph duct as the diaphragm
descends. Every forced expiration has the same effect.
We must keep in mind that the tissues are fed directly by
RESPIRATION. 99
the lymph that surrounds them ; that while the lymph is
continually fed by the blood, there is not a great pressure
given in this way. The lymph stream is largely depend-
ent on the pressure of the surrounding organs. When
one takes a good deal of muscular exercise the lymph is
renewed with rapidity enough to supply the tissues with
food, and to carry away their wastes. But in those who
sit quiet a large share of the day, taking no more exercise
than is necessary to take them to and from their places
of business, the lymph becomes too nearly stagnant, the
tissues are not well nourished, and the whole body suffers.
Deep Breathing. It is a grateful relief to the whole
system to stand, stretch, inhale deeply and slowly several
times, and to repeat this every hour or so. Every one en-
gaged in office work or studying should form this habit,
especially if he does not give an hour daily to exercise in
a gymnasium, or otherwise.
Respiratory Sounds. During respiration sounds are
produced by which the skilled physician can tell much as
to the condition of the respiratory organs.
The Control of Respiration. Breathing is an involun-
tary act. Still we can modify it. We can hold the breath
for a time ; but it is stated that one cannot hold the breath
long enough to produce death by suffocation.
The muscles of respiration are under the control of
nerves. The center of respiratory control is believed to
be in the lower portion of the spinal bulb. This respira-
tory center is one of the most vital points in the body, for
if it is destroyed, breathing is completely stopped, and
death ensues. This center is affected by the condition of
the blood. For instance, if the blood going to this center
has not enough oxygen, the center hastens the process
100 PHYSIOLOGY.
of breathing by nerve impulses sent to the muscles of
respiration.
The Control of the Diaphragm. The diaphragm is
under the control of the phrenic nerves, which arise from
the third, fourth, and fifth cervical nerves. If the neck is
broken above the point where these nerves are given off,
death almost always immediately follows, because the con-
nection of the respiratory center and the diaphragm is
broken.
Composition of Dry Air (by volume) :
Oxygen . . . . . . .... . 2 1 .00
Nitrogen . . . . . v . . . . . 79.00 .
Carbon Dioxid .04
100.04
Experiments illustrating the Chemistry of Respiration. EX-
PERIMENT i . If a piece of phosphorus be burned under a fruit jar
inverted and with the mouth under water (for directions consult any
chemistry), the oxygen will be consumed and water will enter part way
to take its place. The remainder is nitrogen.
EXPERIMENT 2. If a burning taper be lowered into this nitrogen,
the flame will be extinguished.
EXPERIMENT 3. If a chemical laboratory is at hand, some carbon
dioxid should be generated and tested to -show that it extinguishes
flame.
EXPERIMENT 4. Lime water is the test of carbon dioxid, and may
easily be prepared by putting a piece of quicklime the size of a hen's
egg into a quart of water.
EXPERIMENT 5. Pour a little clear lime water into a jar contain-
ing carbon dioxid, and on shaking the contents the lime water will be
rendered milky.
EXPERIMENT 6. By means of a tube (a straw will serve) breathe
through a small quantity of lime water to show that there is carbon
dioxid in the expired breath.
EXPERIMENT 7. If a jar be inverted over a lighted taper, the flame
will soon be extinguished. Test the gas with lime water to see that
carbon dioxid is produced by a burning candle.
RESPIRATION. IOI
EXPERIMENT 8. By holding a clean, cold tumbler over a burning
taper it will be seen that water vapor is produced by the burning.
EXPERIMENT 9. Breathing into a clean, cold tumbler shows that
water is produced also in the process of respiration.
EXPERIMENT 10. A very brilliant experiment and one that is very
instructive at this point is to burn a watch spring in oxygen. In this
process the oxygen unites with the iron, forming iron oxid.
EXPERIMENT n. If a piece of watch spring be placed in water, it
will soon rust. Rust is also an iron oxid, only the process is slow,
instead of rapid as in the case of combustion, and just as much heat is
given off, but not much at any given instant.
EXPERIMENT 12. If a short piece of magnesium ribbon can be
obtained, it may be burned in the presence of the class, though it is not
well to look long at the excessively strong while light.
EXPERIMENT 13. ManMslimi;wild also 'ru^'in water, forming a
white rust, or magnesium oxid, as in burning. , , , ,
EXPERIMENT 14. If a ) In privy vaults :
1. Mercuric chlorid in solution, I : 500.
2. Carbolic acid in solution, five per cent.
(c) For the disinfection and deodori/alion of the surface of masses of
organic material in privy vaults, etc. :
Chlorid of lime in powder.
For Clothing, Bedding, etc. (a) Soiled underclothing, bed linen, etc.
1. Destruction by fire, if of little value.
2. Boiling at least half an hour.
ANTISEPTICS AND DISINFECTANTS. 349
3. Immersion in a solution of mercuric chlorid of the strength of I : 2,000
for four hours.
4. Immersion in a two per cent solution of carbolic acid for four hours.
(b} Outer garments of wool or silk, and similar articles, which would be
injured by immersion in boiling water or in a disinfecting solution :
1. Exposure in a suitable apparatus to a current of steam for ten minutes.
2. Exposure to dry heat at a temperature of no degrees C. (230 de-
grees F.) for two hours.
(V) Mattresses and blankets soiled by the discharge of the sick :
1. Destruction by fire. *
2. Exposure to superheated steam, 105 degrees C. (221 degrees F.), for
ten minutes. (Mattresses to have the cover removed or freely exposed.)
3. Immersion in boiling water for half an hour. .
Furniture and Articles of Wood, Leather, and Porcelain. Washing,
several times repeated, with :
I. Solution of carbolic acid, two per cent.
For the Person. The hands and general surface of the body of attend-
ants of the sick, and of convalescents, should be washed with :
1. Solution of chlorinated soda diluted with nine parts of water, i : 10.
2. Carbolic acid ; two per cent solution.
3. Mercuric chlorid, i : l,ooo.
For the Dead. Envelop the body in a sheet thoroughly saturated
with :
1. Chlorid of lime in solution, four per cent.
2. Mercuric chlorid in solution, i : 500.
3. Carbolic acid in solution, five per cent.
For the Sick Room. (a) While occupied, wash all surfaces with :
1. Mercuric chlorid in solution, I : 1,000.
2. Carbolic acid in solution, two per cent.
(l>) When vacated, fumigate with sulphur dioxid for twelve hours, burning
at least three pounds of sulphur for every thousand cubic feet of- air space in
the room ; then wash all surfaces with one of the above-mentioned solutions,
and afterward with soap and hot water ; finally throw open doors and win-
dows, and ventilate freely."
350
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POISONS AND ANTIDOTES.
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352
LVD ANTIDOTES.
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PO/SONS AND ANTIDOTES.
353
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354 PHYSIOLOGY.
Daily Excretions. Sweat, from 1.5 Ibs. to 4.5 Ibs. ; urea, about
1 oz. ; organic matter exhaled, :! grains ; urine, 53 oz.
"Of the entire excreta, 32 per cent pass off by the breath ; 17 per-
cent by the skin ; 4C.5 per cent by the kidneys ; 4.5 per cent by the
alimentary canal." C'TTTKII.
Number of Sweat Glands. The number of sweat glands may
be as high as 3,500 in a square inch, and the average is estimated at.
2,800 per square inch ; as there are about 2,500 square inches of body
surface, it is readily computed that there are several millions of sweat,
glands.
Number of Hairs on the Human Head. The average number
of hairs on the head is 120,000. They are set obliquely, and are con-
trolled by muscles so that they may be made to stand erect, or nearly so,
under the influence of certain emotions, as fear, anger, etc.
Huxley and others have classified the races of men according to the
hair, into the Ulotrichi, or crisp or woolly haired division, including
the negroes, bushmen, etc. ; and Leiotrichi, or smooth-haired, sub-
divided into the Australioid, the Mongoloid, the Xanthochroic, and the
Melanochroic.
In Europeans the hair is oval in cross-section ; in the Japanese
and Chinese it is circular.
Circulation. Rate of blood flow : in the large arteries, from 12 to
16 inches a second ; in the caval veins, about 4 inches a second ; in the
capillaries, from 1 inch to 1.5 inches a minute. A portion of the blood
makes the complete circulation (in a horse) in less than half a minute.
This is found by putting some readily detected chemical into one jugular
vein, and noting how soon it appears in the other jugular vein. The
time necessary for all the blood to pass through the heart is estimated
as follows : Each ventricle pumps about six ounces of blood at each
stroke. At this rate thirty strokes, 25 to 50 seconds (or less), would
have pumped all the blood in the body. Still, some of the blood (from
the shorter circuits) may have been pumped twice, and some (from the
longer routes) may not yet have been around once. And since the
total amount of blood has been only approximately determined, these
figures are not very accurate.
Number of blood corpuscles to the cubic inch, about 83,000,000.
Dr. Tanner's Forty Days' Fast (Newspaper Account). No
Food but "Water Taken. When Dr. Tanner came to New York
from Minnesota he weighed 184 pounds. He was six weeks making ar-
VITAL STATISTICS.
355
rangernents for his fast; and when lie began his experiment his weight
was 157 pounds. He weighed 121 pounds on the day his fast ended.
He had therefore lost 62 \ pounds since he came to the city, and 3(3
pounds since he began his fast. Dr. Hammond, the well-known New
York physician whose assertion that a forty days' fast was a physical
impossibility led Dr. Tanner to make the attempt, came out in "a card
in the New York papers declaring that he believed the fast had been
fairly conducted.
On each day of his fast Dr. Tanner weighed as follows :
DAY.
1st
POUNDS.
DAY.
25th
POUNDS.
1314,
3d
... 153
20th
. " . . . . I3ld
5th
7th
. . . 147
. 1434
27th .....
28th . .
129!
llth
13th
14th
. . . 139|
133
29th .....
30th
31st
130
16th
. . . 132
32d
1274
17th (8.30 P.M.) . .
17th (HAM)
135^
33d
34th
i2e|
18th
19th
136
35th
36th . ...
20th (4 P.M.) . . .
20th (5 A.M.) . . .
21st .
. . . . 135
. . . . 1 35
37th
38th
39th
1224
22d
1334
40th
1214
26th ......
Cavities of the Body. 1. Mucous cavities (open to the external
air). Digestive tube, respiratory passages, genito-urinary passages, ex-
ternal and middle ear, etc.
2. Serous cavities (closed). They may all be said to be lymph cav-
ities. They are the lymph spaces throughout the body, and the large
spaces, called the pleural cavity around the lungs, the pericardial cavity
around the heart, the peritoneal cavity in the abdomen, the arachnoid
cavity around the brain, and a similar one along the spinal cord.
3. Synovial cavities in the joints.
4. Blood cavities, the inside of the heart and blood tubes.
5. Secretion cavities, the cavities and tubes from the glands ; for
example, the bile sac and its duct.
6. Bone cavities.
356
PHYSIOLOGY.
LOSSES OF THE TISSUES DURING STARVATION*.
(FROM KXPKRIMEXT ON A CAT.)
Fat los's:'. per i-Mit.
Blood .... ;.->
Spleen .... "71 "
Pancreas ... "64 "
Stomach ... "39 "
Pharynx, gullet . " 34 "
Skin " 33 "
Kidneys. ... "31 "
Liver . " 52 "
Heart ....
Intestines . . .
Muscles of locomo-
tion ....
Respiratory appa-
ratus . . .
Bones ....
Eyes
Nervous system .
loses 44 per cent.
" 42
" 42 "
" 22 "
" 16
" 10 "
" 2 "
QUANTITY OF WATER IN 1,000 PARTS.
Teeth 100 Bile 880
Bones 130 Milk H87
Cartilage 550 Pancreatic juice 900
Muscles 750 Urine 936
Ligament 768 Lymph 960
Brain * . . 789 Gastric juice 975
Blood 795 Sweat 986
Synovia 805 Saliva W">
THE LOSS OF WATER FROM THE BODY.
From the Alimentary canal (feces) 4 per cent.
" Lungs 20
" " Skin (perspiration) 30 "
" " Kidneys (urine) 46 "
ELEMENTS IN THE HUMAN BODY.
Oxygen 72.0 Chlorin .085
Carbon 13.5 Fluorin 08
Hydrogen 9.1 Potassium .026
Nitrogen 2.5 Iron 01
Calcium 1.3 Magnesium 0012
Fosforus 1.15 Silicon 0002
Sulfur 147 Copper, lead, aluminum . (traces)
Sodium 1 100.
DAILY RATION OF A U. S. SOLDIER DURING THE LATE WAR.
Bread or flour 22 oz.
Fresh or salt beef (or pork or bacon 12 oz.) 20
Potatoes (three times a week) 16 "
VITAL STATISTICS.
357
Rice
Coffee (or tea 0-.24 oz.) . .
. ... 1.6
... 1.6
. . . . 2.4
64
Vinegar
Salt
COMPOSITION OF F<
WATKU. PROTEIDS.
32
30DS.
FATS.
16
CARBO-
HYDRATES.
Beef, lean
72
19.3
3.6
Beef, fat
51
14.8
29.8
Mutton, lean ....
72
18.3
4.9
Mutton, fat ....
53
12.4
31.1
Veal
63
16.5
15.3
Pork, fat
39
9.8
48.9
Poultry
74
21
3.8
Whitefish
78
18.1
2.9
Salmon
77
16.1
5.5
Eels (rich in fat) . .
75
9.9
13.8
Oysters
75.7
11.7
2.4
SUGAR.
Milk
86
4.1
3.9
5.2
Buttermilk ....
88
4.1
.7
6.4
Cream
66
2.7
26.7
2.8
Cheese, full ....
36
28.4
31.1
. . .
Cheese, skim ....
44
44.8
6.3
Eggs, white . . . .
78
20.4
Eggs, yelk . . ...
52
16
30.7
. . .
STARCH.
Bread
37
8.1
1.6
51
Flour
15
10.8
2
70.8
COMPOSITION
OF THE
BLOOD.
Water
Solids
Corpuscles ....
. 130
Proteids (of serum)
. 70
Fibrin (of clot) . .
2.2
Fatty matters (of serum)
1.4
Inorganic salts . .
6.0
Gases, urea, kreatin,
etc. . .
6.4
gill.
5.1
4.4
4.8
4.7
2.3
1.2
1.0
1.4
2.7
2.7
.8
.8
4.9
4.5
4.9
1.6
1.3
2.3
1.7
784
216
1000
358 PHYSIOLOGY.
COMPOSITION OF CASTKK' ,11 K'K.
Water 99.44
Solids
Pepsin 319
Salts 218
Hydrochloric acid 02
.557
100
Fluids of the Body (FORD). 1. Circulating fluids, chyle,
lymph, blood.
2. Fluids for digestion, saliva, gastric juice, pancreatic juice, bile,
intestinal juice.
3. Fluids of closed cavities, of the arachnoid, pleura!, pericardia!,
and peritoneal sacs, of joints, of the eye and ear, and of cells.
4. Secretions for protection, cerumen or wax, tears, fluid of mucous
membranes, oily fluids on the surface of the body.
5. Fluids for discharge, intestinal secretion, renal or kidney se-
cretion, perspiration, vapor from the lungs, etc.
Acids and Alkalies of the Body. Acids, gastric juice, mu-
cus, chyme, contents of large intestine.
Alkalies, saliva (or neutral), pancreatic juice, intestinal juice,
bile (or neutral), contents of small intestine, sweat.
Amount of Digestive Liquids. The amount of saliva secreted
daily is estimated at from 1 to 3 pints, of gastric juice from 10 to 20 pints,
of bile from 2 to 3 pints. The amount of intestinal and other juices is
difficult to estimate. But it is readily seen that a very large amount of
liquid is daily separated from the blood to be used in the preparation of
the food for absorption into the blood. This is to be looked upon as an
investment. It is supposed to be reabsorbed with large returns in addi-
tion to the prepared food ; and if anything interferes with the absorp-
tion of the food material, especially if the secretion goes on, it is plain
that bankruptcy will follow as surely as in the business world whenever
there is a continual expenditure without corresponding returns. The
condition known as "diarrhea" illustrates this condition, perhaps, as
well as any well-known condition of the body.
Specific Gravity of the Liquids of the Body. As all the
liquids of the body have dissolved and suspended in them various salts
and other matters, they are all heavier than water.
VITAL STATISTICS. 359
Alcohol and Longevity. Investigation by Baer has shown that the
average expectation of life among users and dealers in alcoholic liquors
is very much shortened. The following table gives a comparative view of
the expectation of life in those who abstained from and those who used
alcohol :
EXPECTATION OF LIFE.
AGE. ABSTAINERS. ALCOHOL USERS
At 25, 32.08 years, 26.23 years.
"' 35, 25.92 " 20.01 "
" 45 J 9.92 " * i$.i9 "
" 55' '4-45 " "- 16 "
" 65, 9.62 " 8.04 "
TABLE SHOWING THE INFLUENCE OF ALCOHOL UPON THE
MORTALITY FROM VARIOUS DISEASES.
GENERAL MALE POPULATION. ALCOHOL VENDERS.
Brain disease, 11.77 P er cent - r 4-43 per cent.
Tuberculosis, 30.36 " 36.57 "
Pneumonia and pleuritis, 9.63 " H-44 "
Heart disease, 1.46 " 3.29 "
Kidney disease, 1.40 " 2.11 "
Suicide, 2.99 " 4.02 "
Cancer, 2.49 " 3.70 "
Old age, 22.49 " 7-5 "
GLOS BA RY.
Albumen (al-bu'-meri). The white of an egg.
Albumin (iil-hi'i'-min). A proteid substance, the chief constituent of
the body. Its molecule is highly complex, and varies widely within
certain limits in different organs and in different conditions.
Albuminuria (al-bu'-mi-nu'-ri-a}. The presence of albumin in tin- urine,
indicating changes in the blood or in the kidneys.
Amylopsin (am-i-lop'-xin}. A ferment said to exist in pancreatin.
Anabolism (an-ab'-o-lizm}. Synthetic or constructive metabolism.
Activity and repair of function ; opposed to katabolism.
Arbor Vitae (ar'-6or vl'-te). A term applied to the branched appear-
ance of a section of the cerebellum.
Argon (ar'-gron). A newly discovered element similar to nitrogen
(found in the air).
Arytenoid (ar-i-te'-noid}. Resembling the mouth of a pitcher, as the
arytenoid cartilages of the larynx.
Atlas (at'-las). The uppermost of the cervical vertebrae (from the
mythical Atlas who supported the Earth).
Auricle (aw'-ri-kl). The auricles of the heart are the two cavities be-
tween the veins and the ventricles. Also, the pinna and external
meatus of the ear.
Axis (ak'-sis). The second cervical vertebra, on which the head, with
the atlas, turns.
Bacterium (bak-te'-ri-um}, pi. bacteria. A genus of microscopic fungi
characterized by short, linear, inflexible, rod-like forms without
tendency to unite into chains or filaments.
Biceps (bl'-seps}. Biceps brachii, the flexor of the arm.
Bicuspid (bi-kus'-pid}. Having two points ; the bicuspid or premolar
teeth; the bicuspid valve, between the left auricle and the left ven-
tricle.
Brachial (bra'-ke-al or brak'-i-al}. Pertaining to the arm.
360
GLOSSARY. 361
Bronchus (brong' -kits'), pi. bronchi. The two tubes into which the tra-
chea divides opposite the third thoracic vertebra, called respectively
the right and left bronchus.
Caffein (kaf'-e-in'). An alkaloid that occurs in the leaves and beans of
the coffee-tree, in Paraguay tea, etc.
Canaliculus (kan-a-Uk'-u-lus^), pi. canaliculi. The crevices extending
from lacunae, through which nutrition is conveyed to all parts of
the bone.
Canine (ka-nl)i' or ka'-nin*). The conical teeth between the incisors
and the premolars.
Capillary (kap'-i-la-ri or ka-pil'-a-ri). A minute blood-tube connecting
the smallest ramification of the arteries with those of the veins.
Capsule (kap'-auV). A tunic or bag that incloses a part of the body or
an organ.
Carbohydrate (kar-bo-hi'-draf). An organic substance containing six
carbon atoms or some multiple of six, and hydrogen and oxygen in
the proportion in which they form water; that is, twice as many
hydrogen as oxygen atoms. Starches, sugars, and gums are carbo-
hydrates.
Cardiac (kar'-di-ak}. Pertaining to the heart.
Carotid (ka-rot'-id}. The principal right and left arteries of the neck.
Carpus (kar'-pus). Belonging to the wrist; as the carpal bones.
Cartilage (kar'-ti-luj}. Gristle of various kinds, articular, etc.
Casein (ka'-se-m). A derived albumin, the chief proteid of milk, pre-
cipitated by acids and by rennet at 40C.
Cecum (se'-kum). The large blind pouch or cul-de-sac, in which the
large intestine begins.
Centrum (sen'-trum). The center or middle part ; the body of a verte-
bra, exclusive of the bases of the neural arches.
Cerebellum (ser-e-bel'-um*). The inferior part of the brain, lying below
the cerebrum.
Cerebrum (ser'-e-brum'). The chief portion of the brain, occupying the
whole upper part of the cranium.
Cervical (ser'-m-kal}. Pertaining to the neck, as cervical vertebrae.
Chordae tendineae (kor'-de}. The tendinous cords connecting the
fleshy columns of the heart with the auriculo-ventricular valves.
Choroid (ko'-roid}. The second or vascular coat of the eye, continu-
ous with the iris in front, and lying between the sclerotic and the
retina.
362
Chyle (kll). The milk-white fluid absorbed by tin- lacteals during di-
gestion.
Chyme (kirn'). Food that has undergone gastric digestion, and has not
yet been acted upon hy the biliary, pancreatic, and intestinal
secretions.
Cilium (stT-i-um), pi. cilia. The eyelashes; also the hair-like appen-
dages of certain epithelial cells, whose function is to propel fluid
or particles along the passages that they line.
Ciliary (sil'-i-a-ri). Pertaining to the eyelid or eyelash ; also by ex-
tension to the ciliary apparatus or the structure related to the
mechanism of accommodation. Pertaining to the cilia.
Circumvallate (sir-kum-val'-at). Surrounded by a wall or prominence,
as the circumvallate papillae on the tongue.
Clavicle (klav'-i-kl). The collar-bone.
Coccyx (kok'-siks). The last bone of the spinal column, formed by the
union of four rudimentary vertebrae.
Cochlea (kok'-le-a). A cavity of the internal ear, resembling a snail-
shell.
Conjunctiva (kon-junyk-ti'-va). The mucous membrane covering the
anterior portion of the globe of the eye, reflected on, and extending
to, the free edge of the lids.
Corpus Arantii (kor'-pus). The tubercles, one in the center of each
segment of the semilunar valves.
Corpuscle (kor'-pus-l). A name loosely applied to almost any small,
rounded or oval body, as the blood corpuscles.
Cortex (kor'-teks}. Bark. The outer layer of gray matter of the brain ;
the outer layer, cortical substance, of the kidney.
Cricoid (krl'-koid}. Ring-shaped, as the cricoid cartilage of the
larynx.
Dentine (den'-tin}. The ivory-like substance constituting the bulk of
the tooth, lying under the enamel of the crown and the cement
of the root.
Diabetes (di-a-be'-tez*}. The name of two different affections, diuln-t<-x
mellitn*, or persistent glycosuria, and diabetes insipidus, or polyu-
ria, both characterized, in ordinary cases, by an abnormally large
discharge of urine. The former is distinguished by the presence
of an excessive quantity of sugar in the urine.
Dialysis (di-ar-i-*is\ The operation of separating crystalline from
colloid substances by means of a porous diaphragm, the former
GLOSSARY. 363
passing through the diaphragm into the pure water upon which the
dialyzer rests.
Digastric (di-yas'-trik). Having two bellies, as the digastric muscle,
enlarged near each end and with a tendon in the middle.
Duodenum (du-d-de'-num). The first part of the small intestine, begin-
ning with the pylorus.
Emulsion (e-inul'-shun}. Water or other liquid in which oil, in minute
subdivision of its particles, is suspended.
Enamel (eM-am'~el). The hard covering of the crown of a tooth.
Endothelium (en-do-the'-li-um). The internal lining membrane of
serous, synovial, and other internal surfaces, the homolog of epi-
thelium.
Enzyme (en'-zim}. Any chemic or hydrolytic ferment, as distinguished
from organized ferments such as yeast; unorganized ferment.
Epiglottis (ep-i-ylot'-is). A thin fibro-cartilaginous valve that aids in
preventing food and drink from passing into the larynx.
Esophagus (e-8Q/ y -a-grt*). The musculo-membranous tube extending
from the pharynx to the stomach.
Eustachian (u-sta'-ki-an). Eustachian tube, the tube leading from the
middle ear to the pharynx.
Facet (fas'-et}. A small plane surface. The articulating surface of a
bone.
Femur (/e'-mer). The thigh-bone.
Ferment (fer'-ment}. Any micro-organism, proteid, or other chemic
substance capable of producing fermentation, i.e., the oxidation
and disorganization of the carbohydrates.
Fibrin (fi'-brin}. A native albumen or proteid, a substance that, be-
coming solid in shed blood, plasma, and lymph, causes coagulation
of these fluids.
Fibula (fib'-u-la}. The smaller or splint bone in the outer part of the
leg, articulating above with the tibia, and below with the astraga-
lus and tibia.
Filiform (Jll'-i-forni). Thread-like, as the filiform papillae.
Frontal (fron'-tal). Belonging to the front, as the frontal bone.
Fungiform (fun'-ji-form). Having the form of a mushroom, as fungi-
form papillae.
Ganglion (yany'-gli-on\ pi. ganglions or ganglia. A separate and semi-
independent nervous center, communicating with other ganglia or
nerves, with the central nervous system, and peripheral organs.
364
Gastric (yas'-trik^). Pertaining to the stomach.
Gelatin (jel'-a-tin). An albuminoid substance of jelly-like consistence,
obtained by boiling skin, connective tissue, and bones of animals
in water. The glue of commerce is an impure variety.
Glosso-pharyngeal (glox'-o-fa-rin'-je-al). Pertaining to the tongue and
larynx.
Gluten (ylo'-ten). A substance resembling albumin, and with which it
is probably identified ; it occurs abundantly in the seeds of cereals.
Glycogen (yll'-ko-jen}. A white amorphous powder, tasteless and odor-
less, forming an opalescent solution with water, and insoluble in
alcohol. It is commonly known as animal starch. It occurs in the
blood and in the liver, by which it is elaborated, and is changed by
diastasic ferments into glucose.
Gustatory (gus'-ta-to-ri}. Pertaining to the special sense of taste and
its organs.
Hashish (JtaM'-etA). A preparation from Indian hemp, Cannabis in-
dica. It is a powerful narcotic.
Haversian (ha-ver'-zian*). Haversian canal, in bone, a central opening
for blood-tubes, surrounded by a number of concentric rings, or
lamellae, of bone.
Hemoglobin (hem-o-ylo'-bin}. A substance existing in the corpuscles of
the blood, and to which their red color is due.
Hepatic (he-pat' -ik). Pertaining or belonging to the liver.
Hilum (hl'-lum}. A small pit, scar, or opening in an organic structure ;
the notch on the internal or concave border of the kidney.
Humerus (hu'-me-rus}. The bone of the upper arm.
Humor (/m'-wor). Any liquid, or semi-liquid, part of the body.
Hyoid (hl'-oid}. Having the form of the letter U. The hyoid bone
situated between the root of the tongue and the larynx, supporting
the tongue and giving attachment to its muscles.
Hypo-glossal (hi-po-ylos'-al}. Under the tongue.
Iliac (tT-t-rtJfc). Pertaining to the ilium, or region of the flanks, as iliac
artery, vein, etc.
Incisor (m-sf-sor). The chisel-shaped front teeth.
Inhibition (in-hi-bish'-un). The act of checking, restraining, or sup-
pressing ; any influence that controls, retards, or restrains. Inhib-
itory nerves and centers are those intermediating a modification,
stoppage, or suppression of a motor or secretory act already in
progress.
GLOSSARY. 365
Innominate (i-nom'-i-nate). Nameless ; a term applied to several parts
of the body to which no other definite name has been given, as the
innominate bone, artery, vein, etc.
Invertin (in'-ver-tln\ A ferment found in the intestinal juice, and also
produced by several species of plants ; it converts cane-sugar in
solution into invert sugar.
Jugular (jo'-gu-lar^. Pertaining to the throat, as the jugular vein.
Katabolism (ka-tab'-o-lizm). Analytic or destructive metabolism ; a
physiologic disintegration ; opposed to anabolism.
Lacrymal (lak'-ri-mal}. Having relation to the organs of the secretion,
transfer, or excretion of tears.
Lacuna (la-ku'-na}. A little hollow space ; especially the microscopic
cavities in bone occupied by the bone corpuscles, and communicat-
ing with one another and with the haversian canals and the sur-
faces of the bone through the canaliculi.
Lamella (fti-mel'-a), pi. lamellae. A thin lamina, scale, or plate ; of
bone, the concentric rings surrounding the haversian canals.
Larynx (lar'-inyks). The upper part of the air passage between the
trachea and the base of the tongue ; the voice-box.
Legumin (le-grw'-rain). A proteid compound in the seeds of many plants
belonging to the natural order Leguminosae (peas, beans, lentils,
etc.).
Lumbar (lum'-bcir\ pertaining to the loins, especially to the region
about the loins.
Lymphatic (lim-fat'-ik}. Pertaining to lymph.
Lymphatics (lim-fat'-iks). The tubes that convey lymph.
Lymphatic glands. The glands intercalated in the pathway of the
lymphatic tubes, through which lymph is filtered.
Massage (ma-sazh'). A method of effecting changes in the local and
general nutrition, action and other functions of the body, by rub-
bing, kneading, and other manipulation of the superficial parts of
the body by the hand or an instrument.
Masseter (mas'-e-ter^). A chewing-muscle felt on the angle of the
jaw.
Medullary (med'-u-la-ri}. Pertaining to the medulla, or marrow ; re-
sembling marrow. Also pertaining to the white substance of the
brain contained within the cortical envelop of gray matter.
Mesenteric (mez-en-ter'-ik}. Pertaining to the mesentery, as artery,
vein, etc.
366 GLOSS. IK}:
Mesentery ^nez'-cn-lrr-i}. A fold of the peritoneum that connects cer-
tain portions of the intestine with the dorsal abdominal wall.
Metabolism (me-ta&'-d-Jtem). A change in the intimate condition of
cells ; (1) constructive or synthetic metabolism is called Anabo-
lism ; in anabolism, the substance is becoming more complex and
is accumulating force ; (2) destructive or analytic metabolism is
called Katabolism : in katabolism there is disintegration, the mate-
rial is becoming less complex, and there is loss or expenditure of
force.
Metacarpus (met-a-kirr'-imx). The bones of the palm of the hand.
Metatarsus (met-a-tar'-ms). The five bones of the arch of the foot,
situated between the tarsus and the phalanges.
Mitral (nil'-trtil). Resembling a miter; mitral valve, with two Hups,
between the left auricle and the left ventricle.
Molar (mo'-far). Mill; the grinding-teeth.
Mucous (wu'-fctw). A term applied to those tissues that secrete mucus.
Mucus (mu'-kus). A viscid liquid secretion of mucous membranes,
composed essentially of inucin, holding in suspension desquamated
epithelial cells, etc.
Myosin ( wi'-o-sm). A proteid of the globulin class, the chief proteid
of muscle. Its coagulation after death causes riyor mortis.
Narcosis (nar-ko'-sis) . The deadening of pain, or production of incom-
plete or complete anesthesia by the use of narcotic agents, such as
anesthetics, opium, and other drugs.
Narcotic (nar-kotf-ic}. A drug that produces narcosis.
Neural (nu'-ral). Pertaining to the nerves.
Neuroglia (nu-rog'-li-a*). The reticulated framework or skeleton-work
of the substance of the brain and spinal cord. The term is some-
times abbreviated to glia.
Nucleus (nu'-kle-us). The essential part of a typical cell, usually round
in outline, and situated in the center.
Occipital (ok-sip'-i-tal}. Pertaining to the occiput or back part of the
head, as the occipital bone.
Odontoid (o-don'-toid). Resembling a tooth ; the tooth-like process
(axis) of the second cervical vertebra, on which the atlas turns.
Olfactory (ol-fak'-to-ri}. Pertaining to the sense of smell.
Osmosis (os-wio'-sts)- That property by which liquids and crystalline
substances in solution pass through porous septa ; endosmosis and
exosmosis.
GLOSSARY. 367
Oxy-hemoglobin (ok-ai-hem-o-ylo'-bin). Hemoglobin united, molecule
for molecule, with oxygen. It is the characteristic constituent of
the red corpuscles to which the scarlet color of arterial blood is
due.
Pancreas (pan'-krc-as^. A large racemose gland lying transversely
across the dorsal wall of the abdomen. It secretes a clear liquid
for the digestion of proteids, fats, and carbohydrates. The sweet-
bread of animals, vulgarly called the "belly sweet-bread" in con-
tra-distinction to the thymus, or true sweet-bread.
Pancreatin (pan'-krv-a-t'm}. The active element of the pancreatic juice.
Papilla (pa-pil'-a}, pi. papillae. Any soft, conical elevation, as papillae
of the dermis, tongue, etc.
Papillary (pap'-i-la-ri}. Pertaining to a papilla; papillary muscles,
the conic muscular columns of the heart, to which the chordae
tendineae are attached.
Parietal (pa-ri'-e-tal). Pertaining to the walls, as the parietal bone.
Parotid (pa-rot'-id). Near the ear, as the parotid salivary glands.
Patella (pa-tel'-a}. The knee-pan.
Peptone (pep'-ton}. A proteid body produced by the action of peptic
and pancreatic digestion.
Pericardium (per-i-kar'-di-um}. The closed membranous sac or cover-
ing, that envelops the heart.
Periosteum (per-i-os'-te-um*). A fibrous membrane that invests the
surfaces of the bones, except at the points of tendinous and liga-
mentary attachments, and on the articular surfaces where cartilage
is substituted.
Peristaltic (per-i-stal'-tik}. The peculiar movement of the intestine
and other tubular organs, consisting in a vermicular shortening
and narrowing of the tube, thus propelling the contents onward.
It is due to the successive contractions of the bundles of longitudi-
nal and circular muscular fibers.
Peritoneal (per-i-to-ne'-al}. Pertaining to the peritoneum.
Peritoneum (per-l-to-ne'-um*). The serous membrane lining the interior
of the abdominal cavity, and surrounding the contained viscera.
The peritoneum forms a closed sac, but is rendered complex in its
arrangement by numerous foldings produced by its reflection upon
the viscera.
Phalanges (fa-lan'-jez), plural of phalanx (fa'-lanyks~). Any one of
the bones of the fingers or toes.
368 GLOSS. 1A'\ \
Pharynx ^/'-/m/^- The cavity back of the soft palate. It commu-
nicates anteriorly with the posterior nares, laterally with the eusta-
chian tubes, ventrally with the mouth, and posteriorly with the
gullet and larynx.
Plasma (plaz'-ma). The original undifferentiated substance of IUIM rut,
living matter. The fluid part of the blood and lymph.
Pleura (]>Hj'-rii). The serous membrane which envelops the lungs, and
which, being reflected back, lines the inner surface of the thorax.
Plexus (plek'-sus). An aggregation of vessels or nerves forming an
intricate net-work.
Pneumogastric (nu-md-gas'-trik). Pertaining conjointly to the lungs
and the stomach, or to the pneumogastric or vagus nerve.
Portal (por'-tal). Pertaining to the porta (gate) or hi him of an organ,
especially of the liver, as the portal vein.
Postcaval (post-ka'-ioal}. Pertaining to the postcava; the postcaval
vein, formerly called the inferior vena cava, or vena cava ascendens.
Precaval (pre-kaf-mil}. Pertaining to the precava; the anterior caval
vein, formerly called the superior vena cava, or vena cava de-
scendens.
Pronation (pro-na'-shun^). The turning of the palm downward.
Protoplasm (pro'-to-plazm). An albuminous substance, ordinarily re-
sembling the white of an egg, consisting of carbon, oxygen, nitro-
gen, and hydrogen in extremely complex and unstable molecular
combination, and capable, under proper conditions, of manifesting
certain vital phenomena, such as spontaneous motion, sensation,
assimilation, and reproduction, thus constituting the physical basis
of life of all plants and animals.
Ptyalin (tl'-a-lin}. An amylolytic or diastasic ferment found in saliva,
having the property of converting starch into dextrin and sugar.
Pulmonary (pul'-mo-na-ri). Pertaining to the lungs.
Pylorus (pi-lo'-rus). The opening of the stomach into the duodenum.
Radius (ra'-di-us). The outer of the bones of the forearm.
Renal (re'-naV). Pertaining to the kidneys.
Rennin (ren'-w). An enzyme, or ferment, to whose action is due the
curdling or clotting of milk produced upon the addition of ren-
net.
Retina (ret'-i-na}. The chief and essential peripheral organ of vision;
the third or internal coat or membrane of the eye, made up of the
end organs or expansion of the optic nerve within the globe.
GLOSSARY. 369
Sacrum (sa'-kruni). A curved triangular bone, composed of five con-
solidated vertebrae, wedged between the two iliac (pelvic) bones,
and forming the dorsal boundary of the pelvis.
Scapula (skap'-u-la). The shoulder-blade.
Sciatic (si-at'-ik}. Pertaining to the ischium; the sciatic nerve, the
main nerve of the thigh.
Sclerotic (skle-rot'-ik). Hard, indurated; pertaining to the outer coat
of the eye.
Semilunar (sem-i-lu'-nar}. Resembling a half-moon in shape; semilu-
nar valves, pocket-like valves at the beginning of the aorta and
pulmonary artery.
Serous (se'-rus). Pertaining to, characterized by, or having the nature
of, serum.
Serum (se'-rum*). The yellowish fluid separating from the blood after
the coagulation of the fibrin.
Solar plexus (so'-lar). Solar, with radiations resembling the sun.
Sphincter (sfinyk'-ter'), A muscle surrounding and closing an orifice.
Splenic (splen'-ik). Pertaining to the spleen.
Steapsin (sep'-,sm). A diastasic ferment which causes fats to combine
with an additional molecule of water and then split into glycerine
and their corresponding acids.
Sternum (ster'-num}. The breast-bone.
Subclavian (sub-kla'-vi-an). Situated under the collar-bone ; subcla-
vian artery and vein.
Sublingual (sub-ling 1 '-ywal). Lying beneath the tongue, as sublingual
gland.
Submaxillary (sub-mak '-si-la-ri}. Lying beneath the lower maxilla, as
submaxillary salivary gland.
Supination (su-pi-na'-shun}. The turning of the palm upward.
Synovia ($i-no'-vi-a). The lubricating liquid secreted by the synovial
membranes in the joints.
Tarsus (ar'-sws). The instep, consisting of seven bones.
Temporal (tem'-po-ral). Pertaining to the temples, as temporal artery,
vein, muscle, etc.
Tetanus (tet'-a-nus}. A spasmodic and continuous contraction of the
muscles, causing rigidity of the parts to which they are attached.
Thein (the'-in). An alkaloid found in tea.
Theobromin (Mr-o-bro'-mm). A feeble alkaloid obtained from cacao-
butter ; the essential substance found in cocoa and chocolate.
37O GLOSS A A'\'
Thyroid (thi'-roid). Shield-shaped, as the thyroid cart ilage of the
larynx.
Tibia (tib'-i-a). The larger (inner) of the two hones of tin- leg, com-
monly called the shinbone.
Trachea (tra-ke'-a or tra'-kc-it). The windpipe.
Triceps (tri'-seps). Triceps of the arm, the extensor of the arm. lyinu
along the back of the humerus.
Tricuspid (trl-kim'-piil). Having three cusps or points, as the trienspid
valve.
Trypsin (trip'-sin}. The proteolytic ferment of pancreatic juice.
Ulna (ul'-na). The larger (inner) of the two bones of the forearm.
Ureter (fi-re'-ter). The tube conveying the urine from the pelvis of the
kidney to the bladder.
Vaso-constrictor (rr/.x'-o-A-on-N/r/A-'-f <>/). Causing a constriction of the
blood-vessels.
Vaso-dilator (a*'-d-cK4a / -tor). Pertaining to the positive dilating mo-
tility of the non-striated muscles of the vascular system.
Vaso-motor (vas-o-ino'-tor). Serving to regulate the tension of the
blood-vessels, as vaso-motor nerves ; including vaso-dilator and
vaso-constrictor mechanisms.
Ventricle (ven'-tri-kl). Applied to certain structures having a bellied
appearance. The cavities of the heart from which the blood is
forced out through the arteries.
Vesicle (ves'-i-kl}. A small, membranous, bladder-like formation, as
air vesicle.
Villus (wT-its), pi. villi. One of the numerous minute vascular projec-
tions from the mucous membrane lining the small intestine, for ab-
sorbing digested food.
Vitreous O*T-re-w). Glass-like, as the clear, jelly-like, vitreous humor
of the eye.
INDEX.
Abdomen, cross section of, i6f.
Abdominal respiration, 95.
Absorption, 181.
Of fats, 182, 183.
From stomach, 175.
Accommodation, 291.
Acids, indigestion, 179.
Fatty, 179.
In poisoning, 323.
Tasting, 302.
Action of large arteries, 49.
Of gullet, 171.
Of heart, 45 ; rhythmic, 65.
Of diseased kidneys, 197.
Of muscle, 9.
Of ciliary muscle, 291.
Reflex, 30, 32, 263.
Adam's apple, 309.
Adjustment of lens, 290.
Afferent currents, 268.
Nerve fibers, 27, 28, 32.
Nerve roots, 33, 31.
After-images, 295.
Negative, 295 ; Positive, 295.
After-pressure, 281.
Air, complemental, 96, 97.
Composition of, 100.
Currents about stoves, 116.
Expired, 102.
Reserve, 96, 97.
Residual, 96, 97.
Sacs, 84, 91.
In the sickroom, 325.
Tidal, 96, 97.
Vesicles, 84, 91, 103.
Washed, 119.
Albinos, 288.
Albumen, 145.
Albuminuria, 199.
Alcohol, 208.
In the army, 216, 217.
Binz, 218.
Brunton, 250.
And circulation, 70.
Clum, 252.
And cold climates, 209.
And crime, 208.
Crothers, 251.
Effects of, 210, 223.
And energy, 208, 212.
And excesses, 252.
Greely, 216.
Halliburton, 222.
And heat, 209.
Hornaday, 221.
Howell, 212.
Luce, 260.
Martin, 253.
M'Kendrick, 222.
Miura, 213.
Moral deterioration, 253.
As a narcotic, 210.
And nerve centers, 251.
And nervous system, 250.
As a poison, 210.
Reichert, 213.
Rohe, 218.
Stanley, 216.
Stevenson and Murphy, 250.
As a stimulant, 210.
Thompson, 214.
And training, 209.
In the tropics, 221.
And water, 210.
Woodhull, 216.
Woodruff, 217.
371
372
INDEX.
Alcoholic beverages, 218.
Alkalies, in digestion, 179.
In poisoning, 323.
Alveoli, of the lungs, 84.
Ameba, 5.
Amount of blood, 75.
Of food needed, 193.
Of perspiration, 136.
Of saliva, 168.
Amylopsin, 178.
Anabolism, 203.
Anatomy defined, 3.
And sculpture, 346.
Anesthetics, 254.
Animal matter, 338, 339.
Protoplasm, 202.
Animals and plants, 205.
Antidotes to poisons, 322, 347.
Aorta, 44, 177.
Apex beat of heart, 49.
Apoplexy, 248.
Appendicular skeleton, 330.
Appendix, vermiform, 187.
Aqueous humor, 288, 289.
Arch, neural, 330, 331.
Aristotle's experiment, 266.
Arm, bleeding from, 314.
Arrangement of teeth, 164.
Of muscles, 341,
Arterial muscle, exercise of, 233.
Arteries, large, action of, 49.
Bleeding from, 314.
Distribution of, 44.
And exercise, 69.
Regulation of size, 68.
Structure of, 51.
Artery, carotid, 44, 314.
Gastric, 44.
Hepatic, 44, 177.
Iliac, 44.
Mesenteric, 177.
Pancreatic, 44.
Pulmonary, 42, 43.
Renal, 44.
Splenic, 44.
Subclavian, 44.
Articulating process, 331.
Articulations of vertebra, 335.
Artificial life, i.
Renewal of air, 116.
Auditory center, 244, 264.
Nerve, 239, 238, 305.
Auricles of heart, 41, 47.
Contraction of, 46.
Asiatic cholera, Bacillus of, 123.
Association fibers, 264.
Astigmatism, 292.
Atlas, 335.
Axial skeleton, 330.
Axis, 335 ; axis cylinder, 27, 28.
Bacilli, types of, 123.
Bacillus, of Asiatic cholera, 123.
Of diphtheria, 123.
Of hog cholera, 123.
Tuberculosis, 122, 123
Of typhoid fever, 123.
Bacteria, 124.
Of putrefaction, 127.
Baking meat, 156.
Powder bread, 189.
Ball and socket joint, 339.
Bandaging, 328.
Barley, 149.
Baseball, 229.
Bathing, 232.
The sick, 326; Time for, 233.
Bath mils. 232.
Baths, cold, 232; warm, 233.
Beans, dried, 189.
Bear, hibernation of, 201.
Bedding, changing in sickroom. 326.
Bee-stings, 324.
Beef extract, 155.
Tea, 155.
Beets, 189.
Beverages containing alcohol, 218.
Biceps, 8, 15.
Bicuspid teeth, 164.
Bicycling, 230.
Bile, 177.
Duct, 186, 177.
Functions of, 178.
Sac, 160, 186.
Bites of cats, 324; dogs, 324; snakes, 324.
Bitters, taste of, 302.
Blackberries, 189.
Bleeding from arm, 314; arteries, 314;
neck, 314; nose, 315; veins, 315.
Blind spot, 293.
INDEX.
373
Blindness, color, 295.
Blister, 132.
Blood, amount of, 75.
Changes in, 106.
Chemical reaction of, 75.
Coagulation of, 74.
Color of, 73.
Composition of, 71.
Of frog, 73.
Gases of, 104.
And glands, 134.
Mixture of good and bad, 196.
Quantity in different organs, 75.
Renewal of, 200.
And river, 195.
Specific gravity of, 75.
Transfusion of, 81.
Work of, 39.
Blood-flow, and exercise, 107.
And lymph-flow, 77, 78.
Rate of, 59.
Blood-pressure, of ventricle, 46.
Blood-stream and sewer, 199.
Blood-supply of brain, 247.
Of stomach, 173.
Blood-tubes joining heart, 42.
Blowing, 96.
Blushing, 68.
Boats upsetting, 321.
Body, care of, 2.
And locomotive, 109.
Temperature of, 108.
Boiled milk, 189.
Boiling meat, 156 ; boiling water, 152.
Bone, composition of, 338.
Corpuscles, 337.
Lamellae of, 337.
Bone, structure of, 18, 337.
Bones, broken, 317.
Of ear, 305, 306.
Hygiene of, 339.
Lightness and strength of, 20.
Relation to muscles, 15.
Table of, 333.
Uses of, 21, 330.
Weight of, 337.
Bow-legs, 339.
Boxing, 229.
Brain, 235.
Blood-supply of, 247.
Brain centers, connection of, 263.
Convolutions and intelligence, 240.
Ganglia of, 241.
Gray matter of, 241.
Hemispheres of, 240.
Location of functions, 244.
Parts of, 235.
Preservation of, 236.
Rest, 246, 247.
And sensation, 30, 243.
The water-cushion of, 248.
White matter of, 241.
Work, 246.
Bread, hot, 189.
Breathing, effect on circulation, 98.
Deep, 97, 98.
Hygiene of, 97.
Through mouth, 98.
Restoring, 320.
And swallowing, 170, 171.
Broiling meat, 156.
Broken bones, 317.
Bronchi, 43.
Bruises, 340.
Bulb, hair, 130.
Olfactory, 303.
Spinal, 245, 246.
Burning clothing, 316.
Burns, treatment of, 316.
Cabbage, 189.
Caffein, 155.
Cake, 189.
Calf muscle of frog, 9.
Camel's hump, 201.
Canaliculi, 338.
Canals, haversian, 337, 338.
Canals, semicircular, 305, 306.
Candle, heat of, 205.
And respiration, 201.
Cane sugar, 179.
Canine teeth, 164.
Capacity of lungs, 97, 96.
Vital, 97.
Capillaries, blood-flow in, 55 ; of frog's
web, 52, 53 ; of lung, 91 ; of
muscle, 54 ; pulmonary, 86.
Capsule of lens, 289.
Carbohydrate food, 147.
Carbohydrates, 147.
374
Carbon dioxid of air, 100, 102; in blood,
i