UC-NRLI B 2 flSM MSO ___ " VESTIGES OF THE NATURAL HISTORY CKEATION. Bqmnt of Start) lEtrition. LONDON: JOHN CHURCHILL, PRINCES STREET, SOHO. MDCCCXLVII. LOAN STACK Cr IW CONTENTS. PACK THE BODIES OF SPACE THEIR ARRANGEMENTS AND FORMATION L CONSTITUENT MATERIALS OF THE EARTH, AND OF THE OTHER BODIES OF SPACE 16 THE EARTH FORMED GEOLOGICAL CHANGES 25 LOWER AND UPPER SILURIAN FORMATIONS FIB8T FORMS OF LIFE 29 DEVONIAN ERA FISHES ABUNDANT 35 CARBONIGENOUS ERA COMMENCEMENT OF LAND PLANTS . . 42 PERMIAN ERA FIRST TRACES OF REPTILES 50 ERA OF THE TRIAS AND OOLITE REPTILES ABUNDANT FIRST TRACES OF BIRDS AND MAMMALIA 54 CRETACEOUS ERA 69 ERA OF THE TERTIARY FORMATION MAMMALIA ABUNDANT . 74 ERA OF THE SUPERFICIAL FORMATIONS EXISTING SPECIFIC FORMS ABUNDANT 81 GENERAL CONSIDERATIONS RESPECTING THE ORIGIN OF THE ANIMATED TRIBES . . 8tf PARTICULAR CONSIDERATIONS RESPECTING THE ORIGIN OF THE ANIMATED TRIBES . 99 551 IV CONTENTS. PACE HYPOTHESIS OF THE DEVELOPMENT OF THE VEGETABLE AND ANIMAL KINGDOMS 112 AFFINITIES AND GEOGRAPHICAL DISTRIBUTION OF ORGANISMS 147 EARLY HISTORY OF MANKIND 211 MENTAL CONSTITUTION OF ANIMALS 236 PURPOSE AND GENERAL CONDITION OF THE ANIMATED CREA- TION 255 NOTE CONCLUSORY 274 NOTES . , 279 ERRATUM. Page 56, last line for " four," read " fore." THE BODIES OF SPACE, THEIR ARRANGEMENTS AND FORMATION. IT is familiar knowledge that the earth which we inhabit is a globe of somewhat less than 8000 miles in diameter, being one of a series of eleven which revolve at different distances around the sun, and some of which have satellites in like manner revolving around them. The sun, planets, and satel- lites, with the less intelligible orbs termed comets, are com- prehensively called the solar system ; and if we take as the uttermost bounds of this system the orbit of Uranus (though the comets actually have a wider range), we shall find that it occupies a portion of space not less than three thousand six hundred millions of miles in diameter. The mind fails to form an exact notion of a portion of space so immense ; but some faint idea of it may be obtained from the fact, that, if the swiftest race-horse ever known had begun to traverse it, at full speed, at the time of the birth of Moses, he would as yet have accomplished only half his journey. It has long been concluded amongst astronomers, that the stars, though they appear to our eyes only as brilliant points, are all to be considered as suns, representing so many solar systems, each bearing a general resemblance to our own. The stars have a brilliancy and apparent magnitude which we may safely presume to be in proportion to their actual size and the distance at w r hich they are placed from us. Attempts have been made to ascertain the distance in some 2 THE BODIES OF SPACE, instances by calculations founded on parallax; that is, the change of relative situation produced on a heavenly object by our planet passing from one part of its orbit to another ex- actly opposite ; it being previously understood that if, upon this base of nearly two hundred millions of miles, an angle of so much as one second, or the 3600th part of a degree, could be raised, the distance might be assumed in that instance as not less than 19,200,000 millions of miles! In the case of the most brilliant star, Sirius, even this minute parallax could not be found ; from which, of course, it was to be inferred that the distance of that star is something beyond the vast distance which has been stated. In some others, on which the experiment has been tried, no sensible parallax could be detected ; from which the same inference was to be made in their case. We seemed thus to be left in a hope- less state of ignorance regarding the measurements of the sidereal unijyerse, as if it were such a question as man was not destined ever to answer ; but at length, in our own time, responses came from several points almost at once. By Pro- fessor Henderson, it was ascertained that the star a of the constellation of the Centaur, the third in brightness in our heavens, but in reality a double star, and believed for various reasons to be among those nearest to us, had a parallax of a full second, from which it was inferred that the distance was the vast sum of miles which has been stated. Afterwards, Bessel assigned a parallax of thirty one hundredths of a second to the double star 61 Cygni, placing it at a distance nearly 670,000 times that of the sun.(') Such are but the first steps we take in imagination amongst the hosts of orbs by which we are surrounded. If we suppose that similar intervals exist between all the stars, we shall readily see that the space occupied by even the comparatively small number visible to the naked eye must be vast beyond all powers of human conception. The number visible to the eye is about three thousand ; but when a telescope of small power is directed to the heavens, a great number more come into view, and the number is ever increased in proportion to the increased power THEIR ARRANGEMENTS AND FORMATION. d of the instrument. In one place, where they are more thickly sown than elsewhere, Sir William Herschel reckoned that fifty thousand passed over a field of view two degrees in hreadth in a single hour. It was first surmised by the ancient philosopher, Democritus, that the faintly white zone which spans the sky under the name of the Milky Way, might be only a dense collection of stars too remote to be distinguished. This conjecture has been verified by the instruments of mo- dern astronomers, and some speculations of a most remark- able kind have been formed in connexion with it. By the joint labours of the two Herschels, the sky has been "gauged" in all directions by the telescope, so as to ascertain the condi- tions of different parts with respect to the frequency of stars. The result has been a conviction that, as the planets are parts of solar systems, so are solar systems parts of what may be called Astral Systems that is, systems composed of a multi- tude of stars, bearing a certain relation to eaclT other. The astral system to which we belong 1 , is conceived to be of an ob- long, flattish form, with a space wholly or comparatively vacant in the centre, while the extremity in one direction parts into two. The stars are most thickly sown in the outer parts of this vast ring, and these constitute the Milky Way. Our sun is believed to be placed in the southern portion of the ring, near its inner edge, so that we are presented with many more stars, and see the Milky Way much more clearly, in that direction, than towards the north, in which line our eye has to traverse the vacant central space. Nor is this all. A motion of our solar system with respect to the stars, first suggested by Sir William Herschel, in 1783, has since been verified by the exact calculations of M. Argelander, late director of the Observatory at Abo. The sun is proceeding towards a point in the constellation Hercules. It is, therefore, receding from the inner edge of the ring. Motions of this kind, through such vast regions of space, must be long in producing any change sensible to the inhabitants of our planet, and it is not easy to grasp their general character ; but grounds have nevertheless been found for supposing that not only our sun, but the other suns of the system, pursue a wavy B2 4 THE BODIES OF SPACE, course round the ring, from west to east, crossing and recross- ing the middle of the annular circle. " Some stars will de- part more, others less, from either side of the circumference of equilibrium, according to the places in which they are situ- ated, and according to the direction and the velocity with which they are put in motion. Our sun is probably one of those which depart furthest from it, and descend furthest into the empty space within the ring."( 8 ) According to this view, a time may come when we shall be much more in the thick of the stars of our astral system than we are now, and have of course much more brilliant nocturnal skies ; but it may be countless ages before the eyes which are to see this added resplendence shall exist. The evidence of the existence of other astral systems is much more decided than might be expected, when we consider that the nearest of them must needs be placed at a mighty interval beyond our own. The elder Herschel, di- recting his wonderful tube towards the sides of our system, where stars are planted most rarely, and raising the powers of the instrument to the required pitch, was enabled with awe-struck mind to see suspended in the vast empyrean astral systems, or, as he called them, firmaments, resembling our own. Like light cloudlets to a certain power of the telescope, they resolved themselves, under a greater power, into stars, though these generally seemed no larger than the finest particles of diamond dust. The general forms of these systems (nebulce) are various. So also are the distances, as proved by the different degrees of telescopic power necessary to bring them into view. The furthest observed by the astro- nomer were estimated by him as thirty-five thousand times more remote than Sirius, supposing its distance to be about twenty millions of millions of miles. More recently, the Earl of Bosse has brought his superb instruments to bear upon these distant objects, and thus re- vealed them in more wondrous forms than before. Many which Herschel saw only as filmy matter, spread in patches over the sky, are now found to be vast aggregations of stars. Many which to the elder philosopher seemed round and well THEIR ARRANGEMENTS AND FORMATION. O defined, are seen by his successor to have branches starting out in different directions filaments, as he calls them, the language applicable, to the smallest of objects examined in our hands, being thus found applicable to the promontories of those great continents, each atom of which may be said to be millions of miles removed from another. Such is the universe, as developed to the perceptions of the modern philosopher different indeed from that of our fore- fathers, who did not know the bounds even of this little world, and beheld in the sun, moon, and stars only a set of menial lights ordained, usefully or not, to attend it. And to such con- templations are we raised by modern science, if we choose to leave for them the strifes and self-seekings of our social scene. Thinking of such acquisitions of knowledge, one cannot but go warmly along with the living Herschel when he speaks of the discoveries of Struve, Bessel, and Henderson, as among the fairest flowers of civilization. They surely justify, as he says, " the vast expenditure of time and talent which have led up to them," and show that " there are yet behind not only secrets of nature which shall increase the wealth and power of man, but TRUTHS which shall ennoble the age and the country in which they are divulged, and by dilating the intellect, re-act on the moral character of mankind." ( 3 ) Where our perceptive faculties are baffled, we dream ; where they compass their object, w r e inquire after cause. Such is a law of our minds, which cannot have been bestowed upon us without being designed for a good end. And, indeed, it is by experience placed beyond all doubt, that to yield to this impulse is to use a direct means of improving our condition on earth, and to advance in the scale of moral as well as in- tellectual being. Nor are we left to doubt that extensions of knowledge, either in simple fact or in cause and relation, are not to be estimated by their immediate and apparent effects ; for both are there often good results of the most tangible kind where no such thing was expected as from Napier's disco- very of the logarithms, or, to take an opposite instance, from Smith's ascertainment of the supra-position of rocks and it is utterly impossible in any way to reckon the benefits which 6 THE BODIES OF SPACE, light confers upon mind wherever it is allowed to enter. As- suming, then, the legitimacy of such inquiries, and yet hold- ing by the reverence which Created owes to Creating, we may without fear yield to the instinct which sends us to ask after cause with regard to this vast and beauteous scene. How has it been that these orby myriads have taken the places in which we find them ? To what authorship are we to ascribe the whole ? In philosophising, the prime difficulty is to bring down the mind to sufficiently simple conceptions. Many can soar and mystify, and come to nothing ; to few is it given to find truth where it usually lies, amongst the things most familiar. The ideas which the ancients formed of the movements of the heavenly bodies were lofty, but utterly false. It was re- served for the geometers of the last two centuries, by pursuing truth on more solid grounds, to establish the simplicity which is now known to extend through the physical constitution of the universe. It has been fully ascertained that the planets have obtained their forms, keep their places with regard to the sun and to each other, and pursue all their various mo- tions, in obedience to certain laws which are to be every day seen acting on the humblest scale in our very presence. Thus, the earth is a globe for the same reason that a dew-drop is so. It is slightly flattened at the poles, as a consequence of rotation on an axis when in a soft state, for the same reason that a mass of clay whirled rapidly round will become of a similar shape. The sun and earth are mutually attracted in proportion to their respective masses, and inversely as the square of the distance, which is a law prevailing with not less certainty upon two rose leaves floating on the summer lake into which they have fallen. The revolution of the planet or satellite in an orbit round a central mass is, again, the result of a composition of two opposite forces one of them this attraction of gravity in its proper proportions, the other a primitive motion of the one mass away from the other in a straight line ; and this phenomenon is exemplified when we see a stone which has been thrown from a boy's hand, brought in a curve to the ground. All THEIR ARRANGEMENTS AND FORMATION. 7 these marvels rest on mathematical calculations of the nicest exactness, insomuch that, taking one as an example, astrono- mers have computed ten years beforehand, the time at which the planet Jupiter would pass our meridian, and the predicted time was correct within half a second. Since Xewton stated the laws of gravity and of the pla- netary motions, there have been some important additions to his philosophy. It has been shown, that certain perturbations in the planetary movements, which appeared to him as denoting a necessary end to the system, observe periods, and are only further proofs of the stability of the whole arrangement. It has also been discovered that the laws of motion extend beyond the solar system. Amongst the serene orbs, which seem so still to our ordinary perceptions, we now know that there is no such thing as rest. Stars are ascertained to have proper motions, of the same nature with that found in our own sun. Many are seen to be, in reality, double or triple that is, com- posed of a plurality of suns, which perform regular revolution- ary motions around each other in ellipses. The periods of some of these movements and revolutions are of such brevity, that their elements are already in the book of the astronomer ; others are seen to be of such vastness, that the times which have determinated the youth and death of our oldest empires, would be, in the comparison, but as a little spoke in some enormous wheel. Yet of all of them no doubt can be entertained that they depend upon those simple physical laws which preside over every particle of tangible matter in our own sphere. Here it is right to advert to some general features of the solar system, most of which have also been discovered since the days of Xewton. It is, in the first place, remarkable, that the planets all move nearly in one plane, corresponding with the centre of the sun's body. Xext, it is not less worthy of attention, that the motion of the sun on its axis, those of the planets around the sun, and the satellites around their primaries, ( 4 ) and the motions of all on their axes, are in one direction namely, from w r est to east. Had all these matters been left to accident, the chances against the uniformity would have been, though calculable, inconceivably great. 8 THE BODIES OF SPACE, Of the forty-three motions ascertained in the early part of this century, it was found by Laplace, that the adverse chances were as upwards of four millions of millions to one. It is thus powerfully impressed on us that the uniformity of the motions, as well as their general adjustment to one plane, must have been a consequence of a single cause acting through- out the whole system. Some of the other relations of the bodies are not less re- markable. It is, perhaps, of little consequence that the larger planets are towards the outside of the system, since there is an absence of regularity in the gradation in this respect. In the series of comparative densities we find an approach to a regular gradation : they stand thus in decimals, the Earth being considered as 1 Mercury, 2*95 ; Venus, '99 ; Earth 1 ; Mars, -79 ; Jupiter, -23 ; Saturn, -11 ; Uranus, -26 ; the last being the only very decided violation of the rule. Then the distances are curiously relative. It has been found that, if we place the following line of numbers, 03 6 12 24 48 96 192 and add 4 to each, we shall have a series denoting the re- spective distances of the planets from the sun. It will stand thus 4 7 10 16 28 52 100 196 Merc. Venus. Earth. Mars. Jupiter. Saturn. Uranus. It will be observed that the first row of figures goes on from the second on the left hand in a succession of duplications, or multiplications by 2. Surely there is here a most surprising proof of unity in the solar system. It was remarked, when this relation was first detected, that there was the want of a planet corresponding to 28 ; the difficulty was afterwards considered as overcome, by the discovery of four small planets revolving at nearly one mean distance from the sun, between Mars and Jupiter.( 5 ) The distances bear an equally interest- ing mathematical relation to the times of the revolutions round the sun. With respect to any two planets, the squares of the times of revolutions are to each other in the same propor- tion as the cubes of their mean distances, a most surprising result, for the discovery of which the world was indebted to THEIR ARRANGEMENTS AND FORMATION. 9 the illustrious Kepler. Sir John Herschel truly observes " When we contemplate the constituents of the planetary system from the point of view which this relation affords us, it is no longer mere analogy which strikes us, no longer a general resemblance among them, as individuals independent of each other, and circulating about the sun, each according to its own peculiar nature, and connected with it by its own peculiar tie. The resemblance is now perceived to be a true family likeness ; they are bound up in one chain interwoven in one web of mutual relation and harmonious agreement, subjected to one pervading influence, which extends from the centre to the furthest limits of that great system, of which all of them, the Earth included, must henceforth be regarded as members." ( 6 ) The tendency of all the later discoveries has been to deepen the conviction arising from the first, that the physical affairs of the universe are under the regulation of laws ; the forms, the distances, the movements, the inter- dependencies of the bodies of space, are determined in this, and in no other more arbitrary manner. And what does a law imply ? It is an arrange- ment in which we see invariable uniformity and self-consis- tency. In the case of these physical laws, we can bring it to mathematical elements, and see that numbers, in the expres- sion of space or of time, form, as it were, its basis. We thus trace in law, Intelligence often we can see that it has a bene- ficial object, still more strongly speaking of mind as con- cerned in it. There cannot, however, be an inherent intelli- gence in these laws ; we cannot conceive of mind actually working in the agglomeration of a dew-drop or the orbitual revolution of the moon. The intelligence appears external to the laws ; something of which the laws are but as the expres- sion of the Will and Power. If this be admitted, the laws cannot be regarded as primary or independent causes of the phenomena of the physical world. We come, in short, to a Being beyond nature its author, its God ; infinite, in- conceivable, it may be, and yet one whom these very laws present to us with attributes showing that our nature is in some way a faint and far-cast shadow of His, while all the 10 THE BODIES OP SPACE, gentlest and beautifullest of our emotions lead us to believe that we are as children in his care, and as vessels in his hand. We must consequently understand and this is for the reader's special attention that when we speak of natural law, we only speak of the mode in which the Divine Power is exercised. It is but another phrase for the action of the ever present and sustaining God. Viewing Nature in this light, the pursuit of science is but the seeking of a deeper acquaintance with the Infinite. The endeavour to explain any events in her history, however grand or mysterious these may be, is only to sit like a child at a mother's knee, and fondly ask of the things which passed before we were born. In modesty and reverence, in the spirit of the love of truth, and that craving of an innate help- lessness which seems as if it could never be satisfied till it knew all, we may even inquire if there be any trace of the origin of that arrangement of the universe which is presented to our notice. In this inquiry, we start with the clear fact of the orbs being determined in their forms by law. That law neces- sarily infers a previous form of matter, one in which the mole- cules were separately moveable fluid or gasiform just as the law by which the dew-drop is spherified, implies that the constituent particles were in such a condition before it took effect. We thus see the Will which constitutes law acting in a non-material manner in that portion of what we are ac- customed to call Creation. In the places and relations of the orbs, there is equal proof, though of a less popularly tangible kind, that law was concerned. The work was done by the will of God, expressed in the form of the law of gravitation. When we corne to consider the motions, and regard them as necessarily containing results of an impulse, we are apt to sup- pose some immediate and more direct application of divine power necessary ; but this cannot stand a second considera- tion. We see the motions inextricably wrought up in rela- tion with the magnitudes, as well as the arrangements ; a totally different mode of their origin is therefore inconceiv- able. Having, moreover, in gravitation a general source of THEIR ARRANGEMENTS AND FORMATION. 11 motion, and knowing in what various ways a motive power may be applied, see, for a familiar example, the wheels of a clock revolving under the influence of a weight, all difficulty in supposing an actual origin of a natural kind for the motions of the heavenly bodies vanishes, however obscure our notions may remain as to the process concerned in the case. Thus everything leads us to the belief that there was a previous form of matter, the alteration of which into the present was brought about in the manner of, though certainly not by any self-dependent efficacy in, Natural Law. At this point we might rest, for in the general conclusion that the orbs were formed and arranged in such a manner, enough has been gained for the present object. It is worth while, however, to touch slightly on the ideas which have passed through certain great minds with respect to the births of these bodies. The first idea of what has been called the nebular cos- mogony arose with Sir William Herschel, in consequence of the observations which he made regarding a class of heavenly bodies, to which the appellation of nebula had been applied, in reference to their cloud-like appearance. Some of these bodies were ascertained, by a high telescopic power, to be only astral systems like our own, placed at such a vast dis- tance, that the individuality of the stars composing them was lost to ordinary perceptions. Others resisted the highest telescopic power which the astronomer applied, and, from various considerations, he came to regard them as masses of diffused luminous matter. In these he further discovered a variety of appearances, marking what seemed a gradation of characters, as if they had been in various degrees of conden- sation ; and hence he was led to surmise that they were solar systems in the process of being formed out of a previous con- dition of matter. Laplace now stepped forward to show that, if such a luminous matter existed, and if nuclei were esta- blished in it, these might become centres of aggregation for the neighbouring diffused matter ; on such centres a rotatory motion would be established, wherever, as was the most likely case, there was any obliquity in the lines of direction in 12 THE BODIES OF SPACE, which the opposing currents met each other: this motion would increase as the agglomeration proceeded : at certain intervals, the centrifugal force acting in the exterior of the rotating mass would overcome the agglomerating force, and a series of rings would thus be left apart, each possessing the motion proper to itself at the crisis of separation. These, again, could only continue in their annular form if uniform in constitution. There being many chances against this, they would probably break up, and be agglomerated into either one or several masses, which would then become representa- tives of the primary mass, and perhaps give rise to a similar progeny of inferior masses. All this Laplace showed to be possible under the physical laws of the universe, and he con- ceived that such might be the actual history of all such systems as ours, the four small planets between Mars and Jupiter being an example of a ring which agglomerated into distinct parts, and the rings of Saturn instances of satellites which have not yet attained, if they ever will attain, the ultimate form assigned to such bodies in general. This hypothesis, it will be observed, only comes to the point at which we must needs arrive under a consideration of the " web of relation" traceable in the constituents of the solar system namely, that they have had a common origin in a soft and diffused form of matter. Such a form of matter may now, as is alleged, be no longer actually seen in the heavens ; and yet there may remain good reasons for believing that it once existed. One of these will afterwards be presented in the facts connected with the density of the planets and the internal heat of the earth. As another, I may point to the curious phenomenon called the zodiacal light, an oblate luminosity surrounding the sun, and very conspicuous in the twilights of tropical climes ; a remnant, as has been supposed, of the dif- fused solar atmosphere of the nebular cosmogony. There is even a support to the hypothesis in what would seem at first to be an anomaly and an objection the existence of the many binary and ternary solar systems. It may be supposed that, at a certain point in the confluence of the matter of these regions of space, the solar nuclei would become involved in THEIR ARRANGEMENTS AND FORMATION. 13 a common revolutionary motion, linked inextricably with each other, though it might be at sufficient distances to allow of each body having afterwards its attendant planets. Such a phenomenon is occasionally realized to us on the surface of a river flowing between irregular banks. There we not only see single dimples rotating and passing onward, results of that obliquity in the meeting of currents which is thought to have set solar systems in motion, but often two or more of these dynamic microcosms will come within a range of mutual influence, and go on wheeling around each other. These fantastic eddies, which the musing poet will sometimes watch abstractedly for an hour, little thinking of the laws which produce and connect them, form an illustration of the mechanism of binary and ternary stars, and bring an unex- pected aid to a hypothesis of the history of the heavenly spaces. A remarkable approximation has also been made to what may be called an experimental verification of this cosmogony, by a living professor, M. Plateau, of Ghent. Divested of technical terms, the experiment was nearly as follows : Placing a mixture of water and alcohol in a glass box, and therein a small quantity of olive oil of density precisely equal to the mixture, we have in the latter a liquid mass relieved from the operation of gravity, and free to take the exterior form given by the forces which may act upon it. In point of fact, the oil, by virtue of the law of molecular attraction, in- stantly takes a globular form. A vertical axis being intro- duced through the box, with a small disc upon it, so arranged that its centre is coincident with the centre of the globe of oil, we turn the axis at a slow rate, and thus set the oil-sphere in rotation. " We then presently see the sphere flatten at its poles and swell out at its equator, and thus realize on a small scale an effect which is admitted to have taken place in the planets." The spherifying forces are of different natures, that of molecular attraction in the case of the oil, and of universal attraction in that of the planet ; but the results are analogous, if not identical. Quickening the rotation makes the figure more oblately spheroidal. When it comes to be so quick as 14 THE BODIES OF SPACE, two or three turns in a second, " the liquid sphere first takes rapidly its maximum of flattening, then becomes hollow above and below around the axis of rotation, stretching out continually in a horizontal direction, and finally, abandoning the disc, is transformed into a perfectly regular ring" At first, this remains connected with the disc by a thin pellicle of oil ; which, however, on the disc being stopped, breaks and disappears, and the ring then becomes completely disengaged. The only observable difference between this ring and that of Saturn, is that it is rounded, instead of being flattened ; but this is accounted for by the learned professor in a satisfactory way. A little after the stoppage of the rotatory motion of the disc, the ring of oil, losing its own motion, gathers once more into a sphere. If, however, a smaller disc be used, and its rotation continued after the separation of the ring, rotatory motion and centrifugal force will be generated in the alcoholic fluid, and the oil-ring, thus prevented from returning into the globular form, divides itself into several isolated masses, each of which immediately takes the globular form. These " are almost always seen to assume, at the instant of their forma- tion, a movement of rotation upon themselves, a movement which constantly takes place in the same direction as that of the ring. Moreover, as the ring, at the instant of its rupture, had still a remainder of velocity, the spheres to which it has given birth tend to fly off at a tangent ; but, as on the other hand, the disc, turning in the alcoholic fluid, has impressed on this a movement of rotation, the spheres are especially carried along by this last movement, and revolve for some time round the disc. Those which revolve at the same time upon them- selves, consequently then present the curious spectacle of planets revolving at the same time upon themselves and in their orbits. Finally besides three or four large spheres into which the ring resolves itself, there are almost always pro- duced one or two very small ones, which may thus be compared to satellites. The experiment presents, as we see, an image in miniature of the formation of the planets, according to the hypothesis of Laplace, by the rupture THEIR ARRANGEMENTS AND FORMATION. 15 of the cosmical rings attributable to the condensation of the solar atmosphere." ( 7 ) It must of course be admitted that the process of the experiment was of a reverse kind, and at- tended, as far as M. Plateau's description informs us, by slightly various effects ; but the general reflection which it gives of Laplace's cosmogony is certainly such as to confer upon that hypothesis a strong probability. To conclude this section of the history. What we see is a boundless multitude of bodies with vast empty spaces between. We know of certain motions amongst these bodies ; of other and grander translations we are only beginning to get some knowledge. Beside this idea of locality and move- ment, we have the equally certain one of a former soft and more diffused state of the materials of these bodies ; also a tolerably clear one as to gravitation having been the deter- mining cause of both locality and movement. To no other conclusion, as it appears to me, can these various ideas lead, than to that of universal space being formerly occupied with gasiform matter ; this, however, of irregular constitution, so that gravitation caused it to break up and gather into patches, producing at once the relative localities of astral and solar systems, and the movements which they have since observed, in themselves and with regard to each other, from the daily spinning of single bodies on ideal axles, to the mazy dances of vast families of orbs, which come to periods only in millions of years. How grand, yet how simple the whole of this pro- cess for a God only to conceive and do, and yet for man, after all, to trace out and ponder upon. Oh, truly must we be in some way immediate to the august Father, who can think all this, and so come into his presence and council, albeit only to fall prostrate and mutely adore ! 16 CONSTITUENT MATERIALS OF THE EARTH, AND OP THE OTHER BODIES OF SPACE. THE orbs being all inextricably connected in the manner which has been described, are we also to presume that the constitution of the whole is uniform ? that is to say, do the whole consist of the same chemical elements ? What are elements ? This is a term applied by the chemist to a limited number of substances, (fifty-five are ascer- tained,) which, in their combinations, form all the matters present in and about our globe. They are called elements, or simple substances, because it has hitherto been found impossible to reduce them into others, wherefore they are presumed to be the primary bases of all matters. It has, indeed, been surmised that these so-called elements are only modifications of a primordial form of matter, brought about under certain conditions ; but if this should prove to be the case, it would little affect the present speculations. Analogy would lead us to conclude that the modifications of the primordial matter, forming our so-called elements, are as universal, or as liable to take place everywhere, as are the laws of gravitation and centrifugal force. It therefore ap- pears likely that the gases, the metals, the earths, and other simple substances, (besides whatever more of which we have no acquaintance,) exist under proper conditions, as well in the astral system which is thirty-five thousand times more CONSTITUENT MATERIALS OF THE EARTH. 17 distant than Sirius, as within the bounds of our own solar system or our own globe. Matter, whether it consists of about fifty-five ingredients, or only one, is liable to infinite varieties of condition under different influences. As a familiar illustration, water, when subjected to a temperature under 32 Fahrenheit, becomes ice ; raise the temperature to 212, and it becomes steam, oc- cupying a vast deal more space than it formerly did. The gases, when subjected to pressure, become liquids ; for ex- ample, carbonic acid gas, when subjected to weight equal to a column of water 1230 feet high, at a temperature of 32, takes this form : the other gases require various amounts of pressure for this transformation, but all appear to be liable to it when the pressure proper in each case is administered. Heat is a power greatly concerned in regulating the volume and other conditions of matter. The chemist will probably yet tell us what additional amount of heat would be required to vaporize all the water of our globe ; how much more to disengage the oxygen which is diffused in nearly a proportion of one-half throughout its solids ; and, finally, how much more would be required to cause the whole to become vapori- form, which we may consider as equivalent to its being re- stored to its supposed original nebulous state. He may calculate with equal certainty, what would be the effect of a considerable diminution of the earth's temperature what changes would take place in each of its component sub- stances, and how much the whole would shrink in bulk. The earth and all its various substances have at present a certain volume in consequence of the temperature which actually exists. If, then, we admit that its matter and that of the associate planets was at one time diffused throughout the whole space now circumscribed by the orbit of Uranus, it follows, after what we know of the power of heat, that the nebulous form of matter was attended by the condition of a very high temperature. The nebulous matter of space, pre- viously to the formation of stellar and planetary bodies, must have been a universal Fire Mist, an idea which we can scarcely comprehend. The formation of systems out of this 18 CONSTITUENT MATERIALS OF THE EARTH matter implies a change of some kind with regard to the con- dition of the heat. Had this power continued to act with its full original repulsive energy, the process of agglomeration by attraction could not have gone on. We do not know enough of the laws of heat to enable us to surmise how the necessary change in this respect was brought about ; but we can trace some of the steps and consequences of the process. Uranus would be formed at the time when the heat of our system's matter was at the greatest, Saturn at the next, and so on. Now this tallies with the exceeding diifuseness of the matter of those elder planets, Saturn being not more dense or heavy than the substance cork. It may be that a sufficiency of heat still remains in those planets to make up for their distance from the sun, and the consequent smallness of the heat which they derive from his rays. And it may equally be, since Mercury is nearly thrice the density of the earth, that its matter exists under a degree of cold for which that planet's large enjoyment of the sun's rays is no more than a compensation. Thus there may be upon the whole a nearly equal experience of heat amongst all these children of the sun. Where, meanwhile, is the heat once diffused through the system, over and above what remains in the planets ? May we not rationally presume it to have gone to constitute that luminous envelope of the sun, in which his warmth- giving power is now held to reside ? It may have simply been reserved to constitute, at the last, a means of sustaining the many operations of which the planets were destined to be the theatre. The tendency of the preceding considerations is to impress the notion that our globe is a specimen of all the similarly- placed bodies of space, as respects its constituent matter and the physical and chemical laws governing it, with only this qualification, that there are possibly shades of variation with respect to the component materials, and undoubtedly with respect to the conditions under which the laws operate, and consequently the effects which they produce. Thus, there may be substances here which are not in some of the other bodies, and substances here solid may be elsewhere liquid or AND OF THE OTHER BODIES OF SPACE. 19 vaporiform. We are the more entitled to draw such conclu- sions, seeing that there is nothing at all singular or special in the astronomical situation of the earth. It takes its place third in a series of planets, which series is only one of num- berless other systems forming one group. It is strikingly if such an expression may be used a member of a democracy. Hence, we cannot suppose that there is any peculiarity about it which does not attach to multitudes of other bodies ; in fact, to all that are analogous to it in respect of cosmical ar- rangements. It therefore becomes a point of great interest what are the materials of this specimen ? What is the constitutional cha- racter of this object, which may be said to be a sample, presented to our immediate observation, of those crowds of worlds which seem to us as the particles of the desert sand- cloud in number, and to whose diffusion there are no con- ceivable local limits ? The solids, liquids, and aeriform fluids of our globe are all, as has been stated, reducible into fifty-five substances hitherto called elementary. Of these, forty are well- characterized metals, twelve non -metallic bodies, and the remaining three solid substances of intermediate character, which form a con- necting link between the two great groups. Among the non- metallic elements, four viz., oxygen, hydrogen, nitrogen, and chlorine, are permanently gaseous ; bromine is fluid at common temperatures ; and the remainder (with the ex- ception of fluorine, which has never been isolated, and whose physical characters are consequently unknown) are solid. The body oxygen is considered as by far the most abun- dant substance in our globe. It constitutes a fifth part of our atmosphere, eight-ninths of the weight of water, and a large proportion of every kind of rock in the crust of the earth. Hydrogen, which forms the remaining ^part of water, and enters into some mineral substance, is perhaps next. Nitrogen, of which the atmosphere is four-fifths composed, must be considered as an abundant substance. The metal silicium, which unites with oxygen in nearly equal parts to form silica, the basis of about a half of the rocks in the c2 20 CONSTITUENT MATERIALS OF THE EARTH earth's crust, is, of course, an important ingredient. Alumi- nium, the metallic basis of alumina, a material which enters largely into many rocks, is another abundant elementary sub- stance. So, also, is carbon, a small ingredient in the atmo- sphere, but the chief constituent of animal and vegetable substances, and of all fossils which ever were in the latter condition, amongst which coal takes a conspicuous place. The familiarly -known metals, as iron, tin, lead, silver, gold, are elements of comparatively small magnitude in that ex- terior part of the earth's body which we are able to inves- tigate. It is remarkable of the elementary substances that they generally exist in combination. Thus, oxygen and nitrogen, though in mixture they form the aerial envelope of the globe, are never found separate in nature. Carbon is pure only in the diamond. And the metallic bases of the earths, though the chemist can disengage them, may well be supposed unlikely to remain long uncombined, seeing that contact with moisture makes them burn. Combination and re- combination are principles largely pervading nature. There are few rocks, for example, that are not composed of at least two varieties of matter, each of which is again a compound of elementary substances. What is still more wonderful with respect to this principle of combination, all the elementary substances observe certain mathematical proportions in their unions. When in the gaseous state, one volume of them unites with one, two, three, or more volumes of another, any extra quantity being sure to be left over, if such there should be. Combinations by weight are also governed by fixed and un- changing laws, of the greatest beauty and simplicity. It has hence been surmised that matter is composed of infinitely minute particles or atoms, each of which belonging to any one substance can only associate with a certain number of the atoms of any other. There are also strange predilections amongst substances for each other's company. One will re- main combined in solution with another, till a third is added, when it will abandon the former and attach itself to the AND OF THE OTHER BODIES OF SPACE. 21 latter. A fourth being added, the third will perhaps leave the first, and join the new comer. Such is an outline of the information which chemistry gives us regarding the constituent materials of our globe, and their combinations. How infinitely is the knowledge in- creased in interest, when we consider the probability of such being the materials of the whole of the bodies of space, and the laws under which these everywhere combine, subject only to local and accidental variations ! In considering the cosmogenic arrangements of our globe, our attention is called in a special degree to the moon. In Laplace's hypothesis, satellites are considered as masses thrown off from their primaries, exactly as the primaries had previously been from the sun. The orbit of any satellite is also to be regarded as marking the bounds of the mass of the primary at the time when that satellite was thrown off; its speed likewise denotes the rapidity of the rotatory motion of the primary at that particular juncture. For example, the outermost of the four satellites of Jupiter revolves round his body at the distance of 1,180,582 miles ; hence, according to the hypothesis, the planet was once about 3,675,501 miles in circumference, instead of being, as now, only 89,170 miles in diameter. This large mass would take rather more than six- teen days six hours and a half (the present revolutionary period of the outermost satellite) to rotate on its axis. The innermost satellite would be formed when the planet was re- duced to a circumference of 309,075 miles, and rotated in about forty- two hours and a half. From similar inferences, it would result that the mass of the earth, at a certain point of time after it was thrown off from the sun, was no less than 482,000 miles in diameter, being sixty times what it has since shrunk to. At that time, the mass must have taken rather more than twenty-nine and a half days to rotate, (being the revolutionary period of the inoon,) instead of, as now, rather less than twenty-four hours. The time intervening between the formation of the moon, and the earth's diminution to its present size, was probably 22 CONSTITUENT MATERIALS OF THE EARTH like one of those vast sums in which astronomy deals so largely, but which the mind altogether fails to grasp. The observation made upon the surface of the moon by telescopes tends strongly to support the hypothesis as to all the bodies of space being composed of similar matters subject to certain variations. It does not appear that our satellite is provided with an atmosphere of the kind found upon earth ; neither is there any appearance of water upon the surface. Yet that surface is, like the face of our globe, marked by inequalities and the appearance of volcanic operations. These inequalities and volcanic operations are upon a scale far greater than any which now exist upon the earth's sur- face. The mountains are, in many instances, equal in height to nearly the highest of our Andes. They are generally of extreme steepness, and sharp of outline, peculiarities which might be looked for in a planet deficient in meteoric agen- cies such as those which operate so powerfully in wearing down ruggedness on the surface of our earth. The vol- canic operations are on a stupendous scale. They are the cause of the bright spots of the moon, while the want of them is what distinguishes the duller portions, usually but erroneously called seas. In some parts, bright volcanic matter, besides covering one large patch, radiates out in long streams, which appear studded with subordinate foci of the same kind of energy. A large portion of the surface is covered with circular eminences, called Ring Mountains, of various dia- meters, from a quarter of a mile to several hundred miles, and in some places as close together as the circles on the surface of a boiling pot, which they in no small degree resemble. Some even intrude upon and obliterate portions of the neigh- bouring circles, thus leading to the idea of date, or a succes- sion of events on the moon's surface. Generally, in the centre, there is a mount, which appears to be connected, in the way of cause, with the annular eminence, beyond which again vast boulder-like masses are in some instances seen scattered. What, however, most strikes the senses of an ob- server, is the vast profundity of some of the pits between the AND OF THE OTHER BODIES OF SPACE. 23 ring and the inner mount ; in one case, this is reckoned to be not less than 22,000 feet, or twice the height of .ZEtna. These characteristics of the moon forbid the idea that it can be at present a theatre of life like the earth, and almost seem to declare that it never can become so. But it is far from un- likely that the elements which seem wanting may be only in combinations different from those which exist here, and may yet be developed as we here find them. Seas may yet fill the profound hollows of the surface ; an atmosphere may spread over the whole. Should these events take place, meteorolo- gical phenomena, and all the phenomena of organic life, will commence, and the moon, like the earth, will become a green and inhabited world. ( 8 ) It is unavoidably held as a strong proof in favour of any hypothesis, when all the relative phenomena are in harmony with it. This is eminently the case with the Laplacian cos- mogony, for here the associated facts cannot be explained on any other supposition. AVe have seen reason to believe that the primary condition of matter was that of a diffused mass, in which the component molecules were kept apart through the efficacy of heat ; that portions of this matter agglome- rated into suns, which threw off planets ; that these planets were at first very much diffused, but gradually contracted by cooling to their present dimensions. Now, as to our own globe, there is a remarkably distinct memorial of the supposed high temperature of the materials, in the store of heat which still exists in the interior. The immediate sur- face of the earth, be it observed, exhibits only the temperature which might be expected to be imparted to such materials by the heat of the sun. There is a point a very short way down, but varying in different climes, where all effect from the sun's rays ceases. Then commences a temperature from an entirely different cause, one which evidently has its source in the interior of the earth, and which regularly increases as we descend to greater and greater depths, the rate of increment being, in general, about one degree Fahrenheit for every fifty feet ; and of this high temperature there are other evidences 24 CONSTITUENT MATERIALS OF THE EARTH. in the phenomena of volcanoes and thermal springs, as well as in what is ascertained with regard to the density of the entire mass of the earth. This approximates five and a half times the weight of water; hut the actual weight of the prin- cipal solid substances composing the outer crust is as two and a half times the weight of water ; and this, we know, if the globe were solid and cold, should increase greatly towards the centre, water acquiring the density of quicksilver at 362 miles below the surface, and other things in proportion, and these densities becoming much greater at greater depths ; so that the entire mass of a cool globe should be of a gravity infinitely exceeding five and a half times the weight of water. The only alternative supposition is, that the central materials are greatly expanded or diffused by some means ; and by what means could they be so expanded but by heat ? Indeed, the existence of this central heat, a residuum of that which kept all matter in a vaporiform chaos at first, is amongst the most solid discoveries of modern science,( 9 ) and the support which it gives to the nebular hypothesis is highly important. We shall heratefter see what have been supposed by some to be traces of an operation of this heat upon the surface of the earth in very remote times ; an effect, however, which has long passed entirely away. 25 THE EARTH FORMED GEOLOGICAL CHANGES. IN our version of the romance of nature, we now descend from the consideration of orb- filled space and the character of the universal elements, to trace the- history of our own globe. We shall see that it falls into connexion in an interesting manner with the primary order of things indicated by La- place's Hypothesis. The nature of the materials of the externe or crust of our globe, is known to a greater depth than might be supposed, in consequence of the relation of position of its various masses. C'onfused as these at first appear, an order of ar- rangement, connected with time, has been detected in them by the labours of modern geologists. It is found that a cer- tain kind of rock, below which there is never, in ordinary circumstances, any other kind, is of crystaline character. Sometimes elevated in naked mountain masses, sometimes found only at great depths below other rocks of a different kind, Granite (for such is its name) appears as the basis rock of the earth's crust ; the form into which the once fluid matter of our planet was primarily resolved, although, in many instances, subjected, under heat, to new move- ments at times' long subsequent. The crystals of granite are of distinct substances quartz, felspar, mica, and horn- blende (each of which is, again, a combination of a cer- tain number of the simple or elementary substances) : two 26 THE EARTH FORMED : of these, sometimes three, associated in various proportions, compose the rock, which thus appears in many varieties, passing under different names. Where granite does not appear upon the surface, or else some other igneous rock, such as will presently be adverted to, we find that great flooring overlaid with rocks of a diffe- rent character and history namely, what are called Aqueous or Sedimentary Rocks. These are in the form of strata or beds, and have evidently been for the most part produced as a sediment of sand, clay, or other materials, at the bottom of seas, the matter being hardened by heat and pressure after its deposition. Whence the materials of these rocks ? With some peculiar exceptions, each group of them has been derived from the substance of such rocks as were previously in exist- ence, the earliest from the original granite, and so on in suc- cession ; and this, by means or processes which continue in operation at the present day. That is to say, the atmosphere, by the chemical action of its materials, and the vapours with which it is charged, wears down whatever rocks are exposed to it ; rivers carry the particles into the sea, the sea also erodes the rocks against which it impinges, and strews the matter along its bottom; thus are sediments laid down, to be in time formed into rocks. Many of the earliest or lowest strata are obviously composed of material but slightly changed from the original granite ; such are all the rocks bearing the name of Gneiss. Others present the component materials in different combinations ; as, for instance, where, with clay de- rived from the felspar and the addition of mica, Micaceous Schist has been formed. Sometimes the quartz forms a se- dimentary rock by itself. For such elections of materials, as they may be called, we see natural means of accounting, when we reflect that the lighter particles of any substance suspended in moving water are liable to be carried most ra- pidly, and to the greatest distance. ( 10 ) It is also to be re- marked of all these early rocks, that they have evidently been subjected to an extraordinary degree of heat, insomuch that they generally have acquired a new crystaline texture, are strangely waved and contorted, and often cannot be GEOLOGICAL CHANGES. 27 distinguished from the underlying granite, the adjacent parts of which may sometimes be detected as having been placed there after the deposition of the aqueous rocks. Now the lowest stratified rocks are sometimes found lying in a nearly horizontal position, as they would be originally formed ; but more generally they are tilted up in high in- clinations, with the broken edges directed towards granitic mountains ; indicating that the rise of these mountains from below was the cause of the change of position in the stratified rocks. Thus the earliest strata were in their turn exposed to the wearing influences of sea and atmosphere, and the mate- rials appropriated to form new rocks. And, precisely as might be expected, these new rocks are laid down uncon- formably to the old ; that is, their verges rest at an angle against the sides of the senior formation. These new rocks are again, in their turn, broken up and placed in high inclina- tions by new and similar upbursts of igneous rock ; so as to become liable, of course, to similar disintegration. Such a repetition of wearings down and raisings up, implying fre- quent changes of land and sea, has been in reality the history of our globe since it took its present shape. A granitic crust, containing vast and profound oceans, as is proved by the ex- tent and thickness of the earliest strata, was the infant condi- tion of the earth. Points of unconformableness in the over- lying aqueous rocks, connected with protrusions of granites, and other similar presentments of the internal igneous mass, such as trap and basalt, mark the conclusions of subsequent sections in this grand tale. Dates, such as chronologists never dreamed of compared with which those of Egypt's dynasties are as the latter to a child's reckoning of its birth- days have thus been presented to the now living generation, in connexion with the history of our planet. The aqueous rocks, taken in their details, are a vast num- ber. Geologists, however, group them in formations or sys- tems, partly with reference to their lithological characters and the breaks in stratific arrangement above described, and partly with regard to an entirely different class of particulars. It is now time to say that, from an early portion of the sedimen- 28 THE EARTH FORMED, ETC. tary rock series to its close, the mineral masses are found to enclose remains of the organic beings (plants and animals) which flourished upon earth during the time when the various strata were forming ; and these organisms, or such parts of them as were of sufficient solidity, have, in many instances, been preserved with the utmost fidelity, although for the most part converted into the substance of the enclosing mineral. Now, as we pass along through the series of strata, we find a cessation of certain specific forms of plants and animals, while others come into view ; at some points, the change is almost complete at others, it is very considerable. Such demarca- tions are taken into account by geologists in the grouping of the rock series. They speak of a Palceozoic Period, compre- hending a large early section, terminating at a point where the specific forms are for the first time almost wholly changed ; a Secondary Period, and a Tertiary Period, these also giving groups of species all but distinct. Under each period are reckoned certain systems, more or less organically distinct, and these we shall now proceed to treat separately. 29 LOWER AND UPPER SILURIAN FORMA- TIONSFIRST FORMS OF LIFE. THE first leaves of the Stone Book have been damaged by fire. It is acknowledged by all geologists that the rocks of what has been called the Gneiss and Mica Slate System, were, subsequently to their deposition, exposed to so high a tempera- ture, that, though organic remains had been entombed in them, we should have had no chance of seeing these preserved. ( n ) If it be a fact, therefore, that no distinct traces of plants and animals are presented in these rocks, it is no conclusive proof that such did not then exist. But no such fact can be said to have been ascertained. Certain fragments resembling the cases of infusoria (shelled animalcules) have been observed.( 12 ) A few patches of limestone (primary limestone) indicate organic life as having been present where they were formed ;( 13 ) and a similar inference has been drawn from certain experi- ments of M. Braconnot, in which he detected ammoniacal products from masses of this formation. From analogy, moreover, seeing that all other sedimentary formations con- tain remains of living beings it appears, a priori, unlikely that this should be, any otherwise than apparently, devoid of them. ( 14 ) However all this may be, it is certain that the next formation the Silurian is the first in which we find incontestable monuments of the early life of our planet. The Silurian System is so named, because a district in western England, where it is largely presented at the surface, and where it was first thoroughly examined, was that occu- 30 LOWER AND UPPER SILURIAN FORMATIONS: pied in the time of the Romans by a people called the Silures. It is in reality developed much more extensively in Scandi- navia and Russia, as well as other parts of the world. With us it- is a series of clayey and arenaceous beds, of hard con- sistence ; but the characters are different in other countries. And what were the vessels of the mystery of life upon our earth in the era of the Silurian formation, as far as these rocks can inform us ? One would imagine that, if our present amount of geolo- gical knowledge had come to us by some sudden revelation, it would have been with a kind of awe that its first recipients would have waited for this portion of it. But had they done so, they would quickly have had to admit that nature is simpler than man's wit would make her, for, behold, the in- terrogation only brings before us the unpretending forms of a few humble sea-plants, certain zoophytes and polypiaria, and a variety of shelled marine animals ! To descend to particulars Fucoids, or markings produced by fuci, a tribe of sea-plants, appear in the Lower Silurians of Russia, below any ascertained animal remains, thus sup- porting the obvious conclusion that vegetation must have started fully as early as animal life, since the one thing is necessary to the support of the other. In America, the same vegetable remains are presented in the very first ascertained fossiliferous strata ; but in England they are not as yet found quite so early. In the Lower Silurians of Sweden, not only are there distinct impressions of such plants, but Professor Forchhammer speaks of courses of true coal, composed, as he thinks, of sea-weed, and gives an opinion that the alum-slate of that country owes its combustible character to the carbon, sulphur, and potash, derived from marine vegetation. ( 15 ) Of the animals, the first we are called upon to notice are Polypiaria, the creatures to which we are indebted for those vast coral reefs by which the course of the mariner is so often obstructed in tropical seas, beings thus productive of great results, and yet in themselves extremely humble, as is partly indicated by the composite form in which they exist. Next to them may be ranked certain humble animals (graptolites) FIRST FORMS OF LIFE. 31 allied to the sea-pens of modern seas, a family usually inha- biting mud and slimy sediment in deep water. We come to creatures comparatively well organized, and yet still within the lowest division of the Animal Kingdom, when we speak of Crinoidea, which might be described as a lowly kind of star-fish, fixed on the top of a flexible stalk arising from the sea-bottom. Numberless calcareous plates enter into the composition of the stalk, body, and multitudinous tentacula or arms of the crinoid, forming altogether a wonderful example of the elaborateness of pattern on which nature sometimes works ; and yet it is a very humble animal, only, indeed, a stomach of one aperture, with arms wherewith to supply itself with food. The echinodermata, however, to which order it belongs, are the destructives of their grade ; and thus soon, it therefore appears, were animals set up as a police over the rest, to effect the great providential object of con- trolling the numbers of living beings. We have spoken of the Animal Kingdom and one of its divi- sions. It may be well here to state that these divisions have a reference to the grade and general character of the animals. An animal is said to be low, when its organization is of a simple kind, subservient to a comparatively narrow range of functions, and suited to a comparatively narrow field of exist- ence, if, like the polypes, for instance, and crinoids, it be fixed in situation, and consist mainly of an alimentary recep- tacle, with means of filling that with food. Elevation is marked in the scale, by an animal ceasing to be compound (which is the case of the coralline polypes), assuming a power of locomotion, having sex assigned to separate indivi- duals, exchanging a multiplicity of parts serving one end (as the many pairs of feet in the centipede) for a smaller number ; attaining, in short, at once a more complex and more concen- trated organization. On such grounds, the animal kingdom may be primarily divided into Vertelrated and Invertelrated ; animals with a back-bone and the superior nervous system which that structure implies, and animals devoid of that structure and possessing a humbler nervous system. In the latter are placed, first, the rayed animals {Radiata) ; then, on 32 LOWER AND UPPER SILURIAN FORMATIONS: one higher platform, the Articulata or jointed animals, as Crustacea, insects, spiders, and the Mollusca, or pulpy animals, of which oysters, cowries, and cuttle-fish are ex- amples. To all these, the Vertebrata are as a great and beau- tiful superstructure upon a rustic basement, in the four great classes of ascending rank Fishes, Reptiles, Birds, Mammalia. To resume our view of the Lower Silurian fossils : Of Articulata, we have first a few examples of its lowest class, the Annelides or sea worms, a group ill adapted for pre- servation ; otherwise, they might have been seen in greater abundance. Some which were found in a building stone at Lampeter, far down in the Lower Silurians, lie in a long coil, as they might be supposed to have laid themselves down to die upon the sea-bottom ; they are considered as allied to the nereis of the present seas. But by far the most conspi- cuous articulate of the early rocks is a member of the class Crustacea. This is the Trilobite, a marine animal to which the wood- louse makes the nearest approach in general form, but marked by two groovings along the length of the body, so as to pre- sent a tri-lobed form ; hence the name. The animal of our present seas most resembling it, is the limulus, or king- crab. It is surprising how many species and even genera of the trilobite lived in the Silurian seas, and in what quantities their remains are found in various parts of the earth. The means of locomotion possessed by the animal are not known ; but from its form we must suppose it to have lived at the bottom of the sea, having its eyes directed upwards and side- ways. The latter organs presenting facets such as are seen upon similar animals of our era, we may be said to have a proof furnished to us by this humble creature, " that the water of those oceans and the supernatant atmosphere, was as transparent a medium at that time as it is at the present day, and that therefore no material permanent alteration can have resulted in either during the thousands of years which have elapsed since the creation of the animal world on this earth." ( 1 ). The trilobites stand low in the Crustacea, nor FIRST FORMS OF LIFE. 33 were any higher animals of that order (such as crabs, lobsters, &c.,) yet in existence. Of the division Mollusca, the predominant form in point of numbers was the bivalve order Brackiopoda, which is now but slenderly represented upon earth. So numerous are the specimens in the Silurian rocks, that an eminent geologist calls this emphatically the Age of Brachiopods. ( n ) The ani- mal is a humble one, having two shells, not connected by a hinge, as is usual in superior bivalves, but kept together by a bundle of fibres. Its destiny is to remain fixed at the bottom of deep seas, and live upon nutritive particles, which it col- lects by means of two spiral-shaped arms, extending from the margin of its mouth, and from which the order has received its name. The superior abundance of life in the depths of the ocean, far from land, may be inferred, as a fact of this period, from the comparative number of the brachiopod fossils. Of univalves, which, generally speaking, rank above the bivalves, there are remains of all the three classes. The first and humblest, Pteropoda, most of which are naked, and therefore incapable of preservation in the fossil state, appear only in a few slight conical shells, indicating an animal allied to the genus Criseis, still common in the Mediterranean. Of the next, Gasteropoda, there are many fossil species. There are also representatives of the last class, Cephalopoda, amongst which are now found some of the highest of the invertebrate animals, as the nautilus, cuttle-fish, and poulp. The cepha- lopods (orthoceratites, &c.,) pursuing a free-swimming life, and highly organized for the catching and destroying of the weaker marine animals, were the lords of the organic world in their day. Such are the organisms of the Lower Silurian era, the first age of organization upon earth of which any very distinct memories have been left to us. There was as yet no fish nor any other kind of vertebrated animal, nor any creature which lived upon dry land. The zoology of the Upper Silurians is only different in as far as it presents, for the most part, new species of the same families, and a greater abundance of speci- mens ; one rock (Wenlock limestone) is a mere mass of the D 34 LOWER AND UPPER SILURIAN FORMATIONS. remains of corals, crinoids, and trilobites, held together by- shale, and gives us, in England,* a few obscure traces of fish. It is most remarkable how uniform has been the Fauna of the earth in those primitive ages. Silurian rocks have now been examined in England, Russia, Germany, and North America, with great care ; also in South America, the south- ern part of Africa, and even at the Falkland islands, the very antipodes of Britain ; yet in no place has any essential differ- ence of fossils been detected. Brachiopods, orthoceratites, trilobites, are almost everywhere characteristic fossils. In the Alleghany mountains, in the hills of Herefordshire, on the slopes of the Ural chain, which divides Europe from Asia, we have remains of the same animal tribes. ( 18 ) There are differences of species that is to say, the fossils of different regions present certain minor peculiarities but even this is only partial, and does not materially interfere with the general fact that there has been a remarkable uniformity of life in the primeval seas. In the present era, it is hardly necessary to say, the case is very different. Even seas so near as the Red Sea and Mediterranean, present wholly different genera of mollusks. It has been thought that there might be a cause for the greater uniformity of life in those ages, in the greater uniformity of temperature, resulting from the as yet unspent heat of the surface, arising from the internal incandescence ; but perhaps the more probable cause was simply the compa- rative newness of life upon earth, and its little experience of those external agencies by which it is liable to be affected, and which, we shall see reason to believe, have operated in producing the many shades of variation which now mark the organic kingdoms. 35 DEVONIAN ERA. FISHES ABUNDANT. WE now advance to a new chapter in this marvellous history that of the Devonian era. The term Devonian System is applied to an important and conspicuous group of strata, overlying the Silurian, and largely developed first in the South of Devonshire (whence the name), and in Cornwall, South Wales, Herefordshire, Shropshire, and Worcestershire ; also in Scotland, the valley of the Rhine, Russia, and the west- ern states of America ; besides, in all probability, many parts of the world as yet unexplored. In Scotland, the great Gram- pian elevation, composed of granite and gneissic rock, is skirted by a dense formation of conglomerate and red sand- stone, extending in a sweep from Dunbarton to Stonehaven, and so on to Morayshire, Ross, and Caithness. This passes by the general name of the Old Red Sandstone, which was at first used as an appellative for the system ; but it has latterly been abandoned, as redness is not found to be a prevailing peculiarity of the strata in other countries. In Russia, a surface as large as the whole of Great Britain is occupied by this formation. It reaches a thickness of ten thousand feet in England. The general forms of life prevalent in the Silurian era are continued in the Devonian, with the remarkable addition of a large development of the humblest vertebrate class Fishes. There is here, as there was in the Silurians, an abundance of zoophytes, corallines, crinoids, crustaceans, and mollusks, but mostly presenting those inferior variations which natural- ists regard as constituting distinct species : speaking strictly, D2 36 DEVONIAN ERA. out of about eight hundred so-called species of the Silurian epoch, one hundred pass into the Devonian formation, where, however, they gradually disappear, while new ones as gradu- ally take their place. For such changes of species, adopting this word in the sense usually attached to it, geologists suggest causes in physical changes, as the rise of a sea-bottom by gradual filling up, or the intrusion of a new mineral material into the ocean, or one of a more decisive kind proceeding from such revolutions as are indicated by unconformableness in the strata. But on this point much obscurity at present rests, for, as our survey is extended into other countries, it is found that extensive changes of species occur without any apparent dependence on at least some of these causes ; so that, in these instances, some other explanation remains to be sought for. Corallines (favosites, cyathophylla, stromatopora) are amongst those genera which pass from the Silurians to the Devonians ; they are so abundant, as in some places to con- stitute entire strata, (Devonshire marbles.) The crinoids and trilobites are also continued as families throughout this era. Of the latter we have a new species (brontes), marked by several new features, including a set of claws resembling those of the common lobster, and the whole length of which is judged to have been not less than four feet. Some of the new brachio- pods are of very peculiar shape ; amongst the gasteropods are some which approach existing forms. The lordly cephalo- poda continue to be largely represented, but in a considerable change of form ; for while the chief animals of this class in the Silurians (orthoceratites) had a simple, straight, or slightly curved shell, those new to the present era (as clymenia) had one forming a complete spiral. The most remarkable circumstance connected with the Devonian formation, is its presenting us with fish. A few faint traces of this class had, as we have seen, been presented in the Upper Silurians of our own country, though, it may be remarked, wanting in the corresponding rocks of Russia. We are now to see such memorials of them in the Devonian formation as show that the seas of that era had in many FISHES ABUNDANT. 37 places swarmed with such inhabitants. M. Agassiz, of Xeuf- chatel, to whom the investigation of the subject has chiefly been committed, has ascertained upwards of a hundred species of Devonian fish, to which number it is to be expected that many additions will yet be made. The predominating fishes of this system, and the only ones which (as far as fossils show) existed for some ages, are arranged by M. Agassiz in two orders, with a regard to their external covering, which that naturalist holds to be, in fishes, a reflection of the internal organization. Both orders, it is to be remarked at the very first, are manifestly of an inferior character to the two other orders which afterwards came into existence, and still are the principal fishes of our seas, these being covered by true scales, and respectively named ctenoid and cycloid, from the forms of that part of their organization. The two orders of early fish are covered with integuments considerably different in character ; the one (placoids) with irregular enamelled plates, the other (ganoids} with regular enamelled scales, the first being not placed over each other, as scales are, but laid edge to edge, in the manner of a pave- ment. These characters, according to M. Agassiz, were accompanied by a rudimentary or cartilaginous skeleton, while the ctenoids and cycloids possess an osseous struc- ture. The cephalaspis has a longish tail-like body inserted within the cusp of a large crescent-shaped head, some- what like a saddler's cutting-knife. The body is covered with strong plates of bone, enamelled, and the head was protected on the upper side with one large plate, as with a buckler hence the name, implying buckler-head. A range of small fins conveys the idea of its having been as weak in motion as it is strong in structure. In the coccosteus, the outline of the body is of the form of a short thick coffin, rounded, covered with strong bony plates, and terminating in a long tail, which seems to have been the sole organ of motion. While the tail establishes this creature among the vertebrata and the fishes, its teeth, chiselled, as it were, out of the solid bone of the jaw, like the nippers of a lobster, 38 DEVONIAN ERA. suggest its propinquity to the invertebrate part of creation . The pterichthys has also strong bony plates over its body, arranged much like those of a tortoise, and has a long tail; but its most remarkable feature, and that which has suggested its name, is a pair of narrow wing-like appen- dages attached to the shoulders, which the creature is supposed to have erected for its defence when attacked by an enemy. A group of ganoids seem to have been the police of their day, possessing a powerful development of sharp conical teeth situated on the margin of the jaws. One genus, the holoptychius, introduced near the close of the Old Red era, and passing up into the next, presents a flat oval form, mea- suring in one specimen thirty inches by twelve, with a cover- ing of strong plates, wavily grooved and overlapping each other, the head forming only a slight rounded projection from the general figure. We here find another early and startling example, in addition to the brontes, of animals which may be called large. In the strata of this formation at Db'rpat, there are gigantic bones, which were at first thought to belong to reptiles, but have since been ascertained to be remains of fishes, leading to the conjecture that the animals to which they appertained could not be less than thirty- six feet long (i9). M. Agassiz has lately announced nine genera of sharks of the division Cestraceon in the Devonians of Russia. It is in this voracious family that we see the placoids represented in modern seas ; the ganoids are all but unrepresented in our time. Of both classes, one invariable peculiarity has at- tracted much attention. " In all recent fish, with the excep- tion of the shark family, the sturgeon, and the bony pike, the vertebral column terminates at the point where the caudal fin is given off, and this fin is expanded above and below the body, forming what is called a homocercal tail. In all those, without exception, which have been found in strata of the Palaeozoic period, [placoids and ganoids,] the caudal fin is he- terocercal) being formed of two unequal branches, the upper FISHES ABUNDANT. 39 one expanded immediately from the vertebral column, while the lower one is given off at a point some distance from the extremity." ( 20 ) Xow it is a remarkable fact, that this one- sided tail is a peculiarity in the more perfect fishes (as the salmon) at a certain stage in their embryonic history ; as is also the inferior position of the mouth, peculiar to the early fishes. More than this : in the earlier periods of embryonic life, there is no vertebral column. This organ is represented in embryos by a gelatinous cord, called the dorsal cord, which in maturity disappears as the vertebrae are formed upon it. M. Agassiz has satisfied himself that this was the nature of the organization of the early fishes, as it is that of the sturgeon of the present seas. It is not premature to re- mark how broadly these facts hint at the parity of law affect- ing the progress of general creation, and the progress of an individual foetus of one of the more perfect animals. ( 21 ) An- other feature of the placoids, bringing them down towards the level of an inferior portion of the animal kingdom, is the distinct marks which the dermal plates bear, in many speci- mens, t)f processes for muscular attachments. This suggests a peculiarity of articulate animals, and powerfully hints that the cartilaginous skeleton had not been, as in higher verte- brata, the grand support of the frame, and the basis of its strength. An eminent geologist is of opinion that the species of this era vary locally, as far as might be expected from what we see of the distribution of animal life in the present times. Nevertheless, throughout the distant parts of the earth where Devonian strata are found, the general characters of animals and also vegetable life are nearly the same. It is further ob- served, that whatever particular family is continued with little change through a succession of strata, is also amongst those most widely extended over the world. It is the opinion of M. Brongniart, who has distinguished himself by his in- vestigation of vegetable fossils, that the fuci of these early seas indicate a higher temperature than now prevails at many of the places where they are found. He regards this as a 40 DEVONIAN ERA. proof of the more equable diffusion of a tropical climate in ancient times, and distinctly attributes it to the action of the internal heat of the earth. The early animals are not so uni- form over large geographical areas as the plants. M. Agassiz surmises, from an examination of the fishes of the ancient seas, that the ocean did not at first contain much salt, but gradually acquired its present infusion of that material ; a theory, it may be remarked, which derives support from the suggestion, that the salt of the sea has been mainly brought thither, in the course of time, by rivers, washing it in particles out of the land, in common with other detritus, while it is obvious that rain does not restore it.( 22 ) It is easy to suppose a compara- tive absence of salt in the early ocean affecting animal and vegetable marine life in different ways and degrees. As yet overlooking possible exceptions of a narrow and dubious kind ( 23 ) we meet with no traces of land plants : remains of terrestrial animals have not even been suspected. This exclusively marine character of the flora and fauna of the early ages is usually thought to betoken the non-existence of dry land. But there are reasons apart from the fossil history for believing that great masses had been exposed to the atmo- sphere in those ages. The earliest strata give token of vast disintegration. In our time, this process is usually seen tak- ing place chiefly in the atmosphere, and at the point where land and water meet ; in a much less activity below the sur- face of the ocean. It would thus appear likely that there was dry land in the eras of the earliest stratified formations, though, from whatever cause, it bore no vegetation and sus- tained no animals, or was only a scene of life in certain rare and favourably situated places. The ages of mountains, from which this inference is derived, form one of the most curious as well, as trustworthy chapters in geological science. It is as certain that the Grampian mountains of Scotland are older than the Alps and Apennines, as it is that civilization had visited Italy, and had enabled her to subdue the world, while Scotland was the residence of " roving barbarians." The Pyrenees, Carpathians, and other ranges of continental FISHES ABUNDANT. 41 Europe, are all younger than the Grampians, or even the in- significant Mendip Hills of southern England. Stratification tells this tale as plainly as Livy tells the history of the Roman republic. It tells us to use the words of Professor Phillips that at the time when the Grampians sent streams and detritus to straits where now the valleys of the Forth and Clyde meet, the greater part of Europe was a wide ocean. 42 CARBONIGENOUS ERA. COMMENCEMENT OF LAND PLANTS. THE next group of rocks is called the Carboniferous For- mation, from the remarkable feature of its numerous inter- spersed beds of coal. It commences with the beds of the mountain limestone, which, in some situations, as in Derby- shire and Ireland, are of great thickness, being alternated with chert, (a siliceous sandstone,) sandstones, shales, and beds of coal, generally of the harder and less bituminous kind, (anthracite,} the whole being covered in some places by the millstone grit, a siliceous conglomerate, composed of the de- tritus of the earliest formation. The mountain limestone, attaining in England to a depth of eight hundred yards, greatly exceeds in volume any of the primary limestone beds, and shows an enormous addition of power to the causes con- nected with animal life, by which this substance is supposed to have been produced. In fact, distinct remains of corals, crinoidea, and shells, are so abundant in it, as to compose three-fourths of the mass in some parts. Above the mountain limestone commence the more conspi- cuous coal beds, alternating with sandstones, shales, beds of limestone, and ironstone. Coal is altogether composed of the matter of a terrestrial vegetation, transmuted by putrefaction of a peculiar kind, beneath the surface of water and in the absence of air. Some estuary shells have been found in it, but few of pelagic origin, and no remains of those zoophytes and crinoidea so abundant in the mountain limestone and COMMENCEMENT OF LAND PLANTS. 43 other rocks. Coal beds exist in Europe, Asia, and America, and have hitherto been esteemed as the most valuable of mineral productions, from the important services which the substance renders in manufactures and in domestic economy. It is to be remarked, that there are some local variations in the arrangement of coal beds. In France, they rest imme- diately on the granite and other primary rocks, the interme- diate strata not having been found at those places. In other countries, traces of coal are found in the Devonian formation. These last circumstances may only show that different parts of the earth's surface did not all witness the same events of a certain fixed series exactly at the same time. Some features of the condition of the earth during the de- position of the carboniferous group, are made out with a clear- ness which must satisfy most minds. First we are told of a time when carbonate of lime was formed in vast abundance along the shores and islands of the ocean, accompanied by an unusually large population of corals and encrinites ; while in some parts of the earth there were pieces of dry land covered with a luxuriant vegetation. Xext we have a comparatively brief period of volcanic disturbance, (when the conglomerate was formed.) Then the causes favourable to the so abundant production of limestone, and the large population of marine radiata, decline, and we find the masses of dry land increase in number and extent, and begin to bear an amount of forest vegetation, far exceeding that of the most sheltered tropical spots of the present surface. The climate, even in the lati- tude of Baffin's Bay, was torrid ; and the atmosphere has been supposed by some to have contained a larger charge of car- bonic acid gas (the material of vegetation) than it now does. The forests or thickets of the period included no plants speci- fically the same with those now known upon earth. They mainly consisted of gigantic vegetables, many of which are not represented by any existing types, while others are akin to kinds which, in temperate climes at least, are now only found in small and lowly forms. That these forests grew upon a Polynesia, or multitude of small islands, is considered 44 CARBONIGENOUS ERA. probable, from similar vegetation being now found in such situations within the tropics. With regard to the circumstances under which the masses of vegetable matter were transformed into successive coal strata, geologists are divided. From examples seen at the present day, at the mouths of such rivers as the Mississippi, which traverse extensive sylvan regions, and from other cir- cumstances to be adverted to, it is held likely by some that the vegetable matter, the rubbish of decayed forests, was carried by rivers into estuaries, and there accumulated in vast natural rafts, until it sunk to the bottom, where an overlay of sand or mud would prepare it for becoming a stratum of coal. Others conceive that the vegetation first passed into the con- dition of a peat moss, that a subsidence then exposed it to be overrun by the sea, and covered with a layer of sand or mud ; that a subsequent uprise made the mud dry land, and fitted it to bear a new forest, w*hich afterwards, like its predecessors, became a bed of peat ; that, in short, by repetitions of this process, the alternate layers of coal, sandstone, and shale, con- stituting the carboniferous group, were formed. It is favour- able to this last view that marine fossils are rarely found in the body of the coal itself, though abundant in the shale layers above and below it ; also that in several places erect stems of trees are found with their roots still fixed in the shale beds, and crossing the sandstone beds at almost right angles, show- ing that these, at least, had not been drifted from their ori- ginal situations. On the other hand, it is not easy to admit such repeated risings and sinkings of surface as would be required, on this hypothesis, to form a series of coal strata. Perhaps we may most safely rest at present with the suppo- sition that coal has been formed under both classes of circum- stances, though in the latter only as an exception to the former. The plants of the carbonigenous period have been investi- gated with great care by several able naturalists, and above eight hundred species have been ascertained. The living plants of our own era are at least 80,000, and it is difficult to suppose the flora of that remote age to have been so much more COMMENCEMENT OF LAND PLANTS. 45 limited. It must, however, be observed, that there are many conceivable circumstances to account for the non-preserva- tion or transmission of many of the plants of this era. The numerous fungi, and other lowly forms, could scarcely have left clear memorials of themselves in the rocks, or in the masses of coal ; and it has even been ascertained by experi- ment, that some of the highest forms of vegetation perish with surprising quickness in water. If we might assume, never- theless, that the plants actually ascertained, form in any degree a representation of the flora of this period, they would imply that the early terrestrial botany of our globe was greatly less varied than the present, and composed chiefly of plants of comparatively simple form and structure. ( 24 J In the ranks of the vegetable kingdom, the lowest place is taken by plants of cellular tissue, and which have no flowers, (cryptogamia,) as sea-weeds, lichens, mosses, fungi, ferns. Above these stand plants with vascular tissue, and bearing flowers, in which again there are two great subdivisions ; first, plants having one seed-lobe, (monocotyledons,) and in which the new matter is added within, (endogenous the cane and palm are examples ;) second, plants having two seed-lobes, (dicotyledons,) and in which the new matter is added on the outside under the bark, (exogenous the pine, elm, oak, and all the British forest-trees are examples ;) these subdivisions also ranking in the order in which they are here stated. Now it is found that the predominant plants of the coal era are of the cellular and cryptogamic kind, while the dicotyledons are comparatively rare. There is, indeed, one exogenous family, which occurs in considerable numbers, and, perhaps, figured more conspicuously in the living woods than in the dead coal beds namely, the conifers ; but this, again, is held as the lowest family of its class. That many trees of higher families now existed, seems unlikely, when we learn that such trees occur in considerable numbers in subsequent formations, showing that there was nothing positively to forbid their being preserved in the coal measures, if they had then existed. The master-form or type of the era was the fern, or breckan, of which about one hundred and thirty species have 46 CARBONIGENOUS ERA. been ascertained as entering into the composition of coal. The ferns are plants which thrive best in warm, shaded, and moist situations. In tropical countries, where these condi- tions abound, there are many more species than in temperate climes, and some of these are arborescent, or of a tree-like size and luxuriance. ( 25 ) The ferns of the coal strata have been of this magnitude, and that without regard to the regions of the earth where they are found. In the coal of Baffin's Bay, of Newcastle, and of the torrid zone, alike, are the fossil ferns arborescent, showing that, in that era, the present tropical temperature, or one even higher, existed in very high latitudes. In the swamps and ditches of England there grows a plant called the horse-tail, (cquisetum^) having a succulent, erect, jointed stem, with slender leaves, and a scaly catkin at the top. A second large section of the plants of the carboniferous era were of this kind, (equisetacece,} but, like the ferns, reach- ing the magnitudes of trees. While existing equiseta rarely exceed three feet in height, and the stems are generally under half an inch in diameter, their kindred, entombed in the coal beds, seem to have been generally fourteen or fifteen feet high, with stems from six inches to a foot in thickness. It is to be remarked that plants of this kind (forming two genera, the most abundant of which is the calamites) are only represented on the present surface by plants of the same family : the species which flourished at this era gradually lessen in number as we advance upwards in the series of rocks, and disappear before we arrive at the tertiary formation. The club-moss family (lycopodiacece) are other plants of the present surface, usually seen in a lowly and creeping form in temperate latitudes, but presenting species which rise to a greater magnitude within the tropics. Many specimens of this family are found in the coal beds ; it is thought they have contributed more to the substance of the coal than any other family. But, like the ferns and equisetaceae, they rise to a prodigious magnitude. The lepidodrendon (so the fossil genus is called, from the scaly exterior) has probably been from sixty-five to eighty feet in height, having at the base a COMMENCEMENT OF LAND PLANTS. 47 diameter of about three feet, while the leaves measured twenty inches in length. In the forests of the coal era, the lepidodendra would enjoy the rank of firs in our forests, affording shade to the only less stately ferns and calamites. The internal structure of the stem, and the character of the seed-vessels, show them to have been a link between single- lobed and double-lobed plants a fact worthy of note, as it favours the idea of a progress in vegetable creation, in the line of an improved organization. It is also curious to find a missing link of so much importance in a genus of plants which has long ceased to have a living place upon earth. The other leading plants of the coal era are without re- presentatives on the present surface, and their characters are in general less clearly ascertained. Amongst the most re- markable are the sigillaria, of which large stems are very abundant, showing that the interior has been soft, and the exterior fluted, with separate leaves inserted in vertical rows along the flutings and the stigmaria, a plant apparently cal- culated to flourish in marshes, or pools, having a short, thick, fleshy stem, with a dome-shaped top, from which sprung branches of from twenty to thirty feet long. Amongst mono- cotyledons were some palms, (flabellaria and nceggerathia?) besides a few not distinctly assignable to any class. The conifers of the coal are comparatively rare, and are only as yet found in isolated cases, and in sandstone beds. One discovered in the Craigleith quarry, near Edinburgh, consisted of a stem about two feet thick, and forty-seven feet in length. Others were afterwards found, both in the same situation, and at Newcastle. Leaves and fruit, being wanting, an ingenious mode of detecting the nature of these trees was de- . vised by some naturalists residing in the northern capital.( 26 ) Taking thin polished cross slices of the stem, and sub- jecting them to the microscope, they detected the structure of the wood to be that of a cone-bearing tree, by the presence of certain " reticulations" which distinguish that family, in addi- tion to the usual radiating and concentric lines. That par- ticular tree was concluded to be an araucaria, a species now found in Norfolk Island, in the South Sea, and in a few other 48 CARBONIGENOUS ERA. remote situations. The conifers of this era may be said to form the dawn of dicotyledonous trees, to which, it has already been noticed, the lepidodendra are a link from the monocotyledons. The concentric rings of the Craigleith and other conifers of this era have been mentioned. It is inte- resting to find in these a record of the changing seasons of those early ages, when as yet there were no human beings to observe time or tide. The rings are clearly traced ; but it is observed that they are more slightly marked than is the case with their family at the present day, as if the changes of temperature had been within a narrower range. Such (if we are to be allowed to rest with positive evi- dence) was the vegetation of the carbonigenous era, com- posed of forms low in the botanical scale, mostly flowerless and fruitless, but luxuriant and abundant beyond what the most favoured spots on earth can now show. The rigidity of the leaves of its plants, and the absence of fleshy fruits and farinaceous seeds, unfitted it to afford nutriment to animals ; and, monotonous in its forms, and destitute of brilliant colour- ing, its sward probably unenlivened by any of the smaller flowering herbs, its shades uncheered by the music of birds, it must have been a sombre scene to a human visitant. But neither man nor any other animals were then in existence to look for such uses or such beauties in this vegetation. It was serving other and equally important ends, clearing perhaps the atmosphere of matter noxious to animal life, and certainly storing up mineral masses which were in long subsequent ages to prove of the greatest service to the human race, even to the extent of favouring the progress of its civilization. Traces of land plants previous to the Carboniferous era are isolated at the best, and, till we know more about them, they cannot be allowed greatly to affect our views of the botanical history of the globe. Geologists speak of a fern leaf in the Silurians of Wales ; in those of America, a plant apparently allied to the lepidodendron ; in the American lower Devonian, some allied to ferns. These phenomena, if fully established, would not interfere with general deductions from the mass of early land vegetation found in the coal COMMENCEMENT OF LAND PLANTS. 49 era. There might be patches of vegetation long before the time of the great coal flora ; and from such pieces of land might those early specimens have been wafted. The Carboniferous formation exhibits a scanty zoology compared with either those which go before, or those which come after. The mountain limestone, indeed, deposited at the commencement of it, abounds unusually in polypiaria, crinoidea, and mollusca ; but when we ascend to the coal- beds themselves, the case is altered. We have then only a limited variety of shell mollusks, with fragments of a few species of fishes, and these are rarely or never found in the coal seams, but in the shales alternating with them. Among the fishes, the conspicuous form is the Sauroid family, which receives its name in consequence of a character of teeth, scales, and even osteology, resembling that of the Sauria, and evidently leading on to that section of reptiles. ( 27 ) One of the most noted species is the Megalichthys Hibbertii, discovered by Dr. Hibbert Ware, in a limestone bed at Burdiehouse, near Edinburgh, and of which other specimens have been found in the coal measures of Yorkshire, and low coal shales of New- castle. The enormous size of the animal is inferred from teeth belonging to it, not less than four inches long. At this point we find the first traces of land animals, in the fossil re- mains of terrestrial insects ( 28 ) and the foot-prints of reptiles, the first in England, the latter in America. ( ?9 ) Coal strata are nearly confined to the group termed the carboniferous formation. Thin beds are not unknown after- wards, but they occur only as a rare exception. It is there- fore thought that the most important of the conditions which allowed of so abundant a terrestrial vegetation whatever these were had ceased about the time when this formation was completed. The termination of the carboniferous formation is marked in some regions by symptoms of great disturbance. Coal- beds generally lie in basins, as if following the curve of the bottom of seas. There is no such basin which is not broken up into pieces, some of which have been tossed up on edge, others allowed to sink, causing the ends of strata to be in 50 PERMIAN ERA. some instances many yards, and in a few, several hundred feet, removed from the corresponding ends of neighbouring fragments. These are held to be results of volcanic move- ments below, the operation of which is further seen in nume- rous upbursts and intrusions of fire-born rock, (trap.) That these disturbances took place about the close of the formation, and not later, is shown in the fact of the next higher group of strata being comparatively undisturbed. Other symptoms of this time of violence are seen in the beds of conglomerate which occur amongst the first strata above the coal. These, as usual, consist of fragments of the elder rocks, more or less worn from being tumbled about in agitated water, and laid down in a mud paste, afterwards hardened. ( 30 ) It is to be admitted for strict truth, that in some parts of Europe the carboniferous formation is followed by superior deposits, without the appearance of such disturbances between their respective periods; but such cases apparently are exceptive. PERMIAN ERA. FIRST TRACES OF REPTILES. IN this subordinate manner, may be noticed a short series of strata, following, whether conformably or otherwise, upon the carboniferous formation, and to which a general name has been applied, from its being unusually well developed in the portion of Russia which formed the ancient kingdom of Permia. This sub-formation comprehending in ascending order a group of sandstones, called with us the Lower New Red Sandstones, and amongst the Germans Rothe-todte- liegende ( 31 ) a thick calcareous bed called with us the Magnesian Limestone, by the Germans Zechstein, and some other strata is, in respect of fossils, a continuation of the carboniferous system. With it, however, ends a range of animal forms which first appeared in the Silurians, and passed, with the changes which have been indicated, through the Devonian and Carbonigenous eras. FIRST TRACES OF REPTILES. 51 The total number of specific forms, which had heen dimi- nishing in the carbonigenous era, is in this still further reduced ; one recent author says, from about a thousand to a hundred and sixty-six, of which only eighteen are common to the inferior strata. ( 32 ) It appears as if, while some new species continued to present themselves, the animal kingdom were now generally undergoing a decay, for even specimens of particular families are less abundant than formerly. Instead, for example, of the hundred species of corals of the carbonifer- ous formation, there were now only fifteen, and of these but three or four abundant. Of the numerous crinoidea of the past, but one now remained, and this is rarely found. The trilobite has now vanished, to appear no more. For hundreds of brachiopods, there were now only thirty, ten of them old. The cephalopods almost disappear at the very commencement of the Permian era. It cannot at present be determined whether this diminution of fossils is owing to an actual reduction of the amount of life in the ancient seas, or only to some such simple cause as the occurrence of deposits which were not favourable to the pre- servation of animal remains. It may even be that the prin- cipal cemeteries of the age have not yet been hit upon by research ; for certainly this is neither the most extensively nor the most rigidly examined of the various formations, and we are made the more suspicious by finding that, at this part of the rock series, several important fossiliferous strata are present in one region and not in others. It has been ascer- tained, however, by Permian researches, that extensive changes of specific forms in the ancient seas were not, as has been supposed, necessarily and essentially connected with great physical disturbances ; for both do we find that the un- conformability of strata or memorials of disturbance between the carboniferous and Permian do not affect the fossils, and that a conformable succession of strata over the Permian is attended by a great usually called a complete change of species. At this termination of the Permian, modern geolo- gists close what they call the Palaeozoic Period, on a suppo- sit'on that an ancient creation had now passed away, to givo B2 52 PERMIAN ERA. place to one entirely new. And this view is eagerly embraced by those who argue for repeated interferences of creative power. But not only is such a notion discountenanced by the nature of the subsequent organisms, an advance to higher species of particular classes, and to a new class the next in the animal scale, but it is utterly overthrown by the recent discovery of plants in the higher formations, (Trias of France and certain Liassic beds in the Alps,) identical with carboni- ferous species. Where such changes of fossils occur, the more reasonable supposition is, that notwithstanding conformable- ness of strata, a local suspension of deposits for a considerable time is indicated, a time during which the usual changes of species were proceeding, probably at their usual rate, and which was sufficient to present something like a complete change of forms when the deposits were re-commenced. ( 33 ) In the Permian formation, besides the principal orders of animals which previously existed, we have the first undoubted traces of another, succeeding fish in the animal scale, namely, Reptiles- This is a most important event in our history, for it gives us, for the first time, a class of vertebrate animals capable of breathing the atmosphere and walking upon the land. We shall presently see that it was a class destined for a long suc- cession of ages to flourish over the soil, in many various and some most formidable shapes, and without any superiors to keep them in check. As yet, but a few bones of reptiles have been discovered in the Zechstein of Thuringia in Upper Saxony, and in quarries near Bristol. By Professor Owen, who has carefully examined them, they are said to be of the lacertilian or lizard order (specifically called by him, palseosaurs, thecodonts, monitors, &c.), but for the most part of gigantic size, and differing from modern lizards in very re- markable characters of the vertebrae, teeth, and dermal plates. To them, as to all the reptiles of this and several subsequent great periods, belonged a fish-like form of the vertebral column, in as far as its component bones were biconcave, or shaped like a double egg-cup, a peculiarity regarded by this eminent anatomist as probably fitting the animal for partially FIRST TRACES OF REPTILES. 53 marine habits. And that the full importance of this pecu- liarity of the early reptiles may be appreciated, the reader must be made aware that modern reptiles have a ball-and- socket form of the vertebrae that is, a convexity at the one side fitting into the hollow of the adjacent bone ; but this form only when they are mature animals, for in the embryotic state of the crocodile and of the frog the form has been ascer- tained to be biconcave, which gradually changes as the animal approaches perfection. The teeth of the thecodonts and palaeosaurs were fixed in distinct sockets, like those of the modern crocodiles. In this respect, they were superior to the modern varanians, the nearest living tribes, which have the teeth imbedded in comparatively shallow cavities along the bottom of a groove in the jaw. 54 ERA OF THE TRIAS AND OOLITE. REPTILES ABUNDANT. FIRST TRACES OF BIRDS AND MAMMALIA. GEOLOGISTS now apply the term Secondary Formation (once of wider application) to those intervening between the end of the Permian or close of what they call the Palaeozoic Period, and the termination of the Chalk Series, afterwards to be described, at which place there is another and almost total change of specific forms. The first of these formations is called with us the Upper New Red Sandstone ; it consists, in England, of only a group of strata of that kind, surmounted by some variegated marls. But on the Continent, below a stratum equivalent to these marls, there is one of limestone, bearing the name of the Muschelkalk, and full of shells. The system is there called Trias, on account of its thus consisting of a triple group of strata. TRIAS. THE organic relics of this system are most abundant in the Muschelkalk. There we are presented with a great number of crinoids and shells, all differing in specific character from their predecessors of the same orders. The crinoid called, from its elegant lily-like shape, Encrinites Moniliformis, is a conspicuous fossil. The brachiopods, here almost extinct, are REPTILES ABUNDANT. 55 replaced by ostraceae of various genera a change from the animals of deep to those of shallow seas. The univalve mol- lusks also indicate a condition of the sea advancing towards that which exists near the present shores. In the new forms of cephalopoda were some marking their advanced character by their non-possession of a shell or stony skeleton. In this case, the existence of the animal is only betrayed by its horny mandible, constituting the fossils called rhynchulites. We find in this system further traces, but still obscure and local, of the reptilian class. Before proceeding to speak of them, it is necessary to remark that the ingredients and ar- rangements of rocks, with fossil remains, do not form the sole materials of the history compiled by the geologist. He is equally contented when he can find an intelligible fact told by what may be called a writing of nature upon these stone tablets. So low as the bottom of the carboniferous system, slabs are found marked over a great extent of surface with that peculiar corrugation or wrinkling which the receding tide leaves upon a sandy beach when the sea is but slightly agitated ; and not only are these ripple-marks, as they are called, found on the surfaces, but casts of them appear on the under sides of slabs lying above. The phenomena suggest the time when the sand, ultimately formed into these stone slabs, was part of the beach of a sea of the carbonigenous era ; when, left wavy by one tide, it was covered over with athin layer of fresh sand by the next, and so on, precisely as such circumstances might be expected to take place at the present day. Sandstone surfaces, ripple-marked, present themselves throughout the sub- sequent formations : in those of the New Red, at more than one place in England, they further bear impressions of rain-drops which have fallen upon them the rain, of course, of the incon- ceivably remote age in which the sandstones were formed. In the Greensill sandstone, near Shrewsbury, it has even been possible to tell from what direction the shower came which impressed the sandy surface, the rims of the marks being somewhat raised on one side, exactly as might be expected from a slanting shower falling at this day upon one of our beaches. These facts have the same kind of interest as the 56 THE TRIAS AND OOLITE. season rings of the Craigleith conifers, speaking of the identity of the familiar processes of nature in those early ages with those of our own. Hearing of memorials of this kind will prepare the reader to learn that the earliest intelligence we have respecting land- walking animals consists, in great part, of their mere foot- steps, impressed on the wet sand or mud, which afterwards became rock. Let no one undervalue such testimony. The fidelity of an impression from a foot, as certifying by what or whose foot the impression was made, is acknowledged in judicial procedure ; and often has this kind of evidence fixed the opinion of judge and jury, when every other has failed. So much being premised, we proceed to examine the Triassic reptiles. In the lower beds of the upper new red sandstone, near Shrewsbury, we are introduced to a new lacertilian, presenting some remarkable characters, and named the Hhynchosaurus. From the few fragments of the animal which have been discovered, it would appear to have had a toothless head, resembling that of a bird, and en- closed in a bony sheath ; also a hinder toe directed back- wards, in which feature we also see an assimilation to the next higher vertebrate class. Footmarks, impressed in the way which has been described, and attributed to this animal, confirm the appearances presented by the extraordinary ar- rangement of its locomotive organs. In the same beds occur a few bones, and a great number of footsteps, which Professor Owen has fixed as the double me- morials of a group of animals, to which he has given (from the structure of their teeth) the name of Labyrinthodonts, and which he classes with the JBatrachia, that order of rep- tiles to which the frog and toad belong. Those who are ac- customed to regard this as a group of generally small and insignificant animals, will be surprised to learn that the laby- rinthodonts were of the size of a large hog. Their footmarks, discovered alike in America and the elder continent, " bear a singular resemblance to the impression that would be made by the palm and expanded fingers and thumb of the human hand." But it is evident that the four extremities of the animal had REPTILES ABUNDANT. 57 been, like those of the kangaroo and some other genera, much smaller than the hinder, some specimens of which measure eight inches by five. These batrachia present affinities to the fish class in their biconcave vertebrae and the formation and arrangement of the teeth. Their nostrils being also, like those of the Sauria, placed near the extremity of the head, indicate a partially marine habitat, such an arrangement being required to enable the animal to breathe while nearly altogether sunk in the water. Quarries of the red sandstone of this system also present an abundance of footmarks attributed to tortoises, thus point- ing to the contemporaneous existence of a third order of rep- tiles, the Chelonia. The first examples were discovered by the Rev. Dr. Duncan in the quarry of Corncockle Muir, Dum- friesshire, where the slabs incline at an angle of thirty-eight degrees, and the footmarks are distinctly traced up and down the slope, as if, when the surfaces w r ere those of a beach at, however, a lower inclination the animal had had occasion to pass only in that direction, in its daily visits to the sea. Some slabs similarly impressed, in the Stourton quarries, Cheshire, are further marked with a shower of rain which we know to have fallen afterwards, for its little hollows are impressed in the footmarks also, though more slightly than on the rest of the surface, the comparative hardness of a trodden place having apparently prevented so deep an impression being made. It is in the celebrated Muschelkalk that, for the first time, we find examples of a group of reptiles which have excited more attention than perhaps any other fossil animals. The same group, it may be remarked, occurs in the English lias and subsequent formations ; but the mere fact of writing in England should not make us postpone to that place an order of beings which we find earlier in another portion of what, geologically, may be regarded as but one great zoological province. These animals, called collectively Enaliosauria, or Marine Saurians, abounded throughout a long period of the earth's history, while mammalian life was yet hardly de- veloped ; but they disappeared in what we shall have to speak of as the Cretaceous Era. The Ichthyosaur, of which ten 58 THE TRIAS AND OOLITE. species have been distinguished, was an animal of marine habits and great bulk, (reaching about thirty feet in length,) in which to the form of the fish there were united, in a re- markable way, characters of animals higher in the scale. A body, framed upon a purely piscine vertebral column, con- taining a huge voracious stomach, and terminating in a ver- tically expanded tail, in which respect it also preserved the fish character, was furnished with the head of a crocodile, and four fins approximating to the character of the paddles of the whale, but composed of a greater number of bones, and thus showing an affinity to the fins of fishes. Over all was a skin resembling that of the cetaceous animals. Nor should it be omitted that the sternum or breast-bone presents a structure resembling that of the ornithorhynchus or duck-rat of Aus- tralia. The vast jaws of this animal, having a stretch of seven feet ; its eye resting in a socket eighteen inches in diameter, and defended by an apparatus of bony plates, like that of a bird of prey ; the powerful range of teeth, and the position of the breathing apertures near the extremity of the snout ; all speak to the naturalist of ferocious habits like those of the modern crocodile, to which the ichthyosaur may be considered as a link from the predaceous fish. A curious light has been thrown upon these habits by the pellets voided by the animal, which have been found in great quantities in a fossilized state, and bear the name of coprolites. In these we find fragments not only of fish, but of reptiles, arguing that the animal must have been a destructive creature both to its own class and to that below it. The genus next in importance is the Plesiosaurus, so called as being near to the saurian character. This animal was under eighteen feet long, and altogether a feebler creature than the Ichthyosaur, which seems to have made it a prey. Yet it was itself one of the destructive potentates of the early seas. A body, generally fish-like, though framed on vertebrae presenting less concave ends, and which terminated in a short tail, serving only as a rudder, was furnished with a long neck and small head, together with four slender paddles, more cetacean than those of the Ichthyosaur. Moving, like that REPTILES ABUNDANT. 59 animal, quickly in the water, by means of the special organs designed for the purpose, the Plesiosaur would have a further advantage in its long, flexible, serpent-like neck ; but the small size of the head, though there we find the same superior arrangement of teeth seen in the thecodonts, must have rendered it a much less formidable creature than that last described. Professor Owen regards it as fitted to live near shores and to ascend estuaries. The attention of the geologists of the United States has been called to certain footmarks in the sandstones of the valley of Connecticut, indicative, as they think, of birds of the orders Grallatores (waders) and Rasores (scrapers.) " The footsteps appear in regular succession on the continuous track of an animal, in the act of walking or running, with the right and left foot always in their relative places. The dis- tance of the intervals between each footstep on the same track is occasionaHy varied, but to no greater amount than may be ex- plained by the bird having altered its pace. Many tracks of dif- ferent individuals and different species are often found crossing each other, and crowded, like impressions of feet upon the shores of a muddy stream, where ducks and geese resort." ( 34 ) Some of these prints indicate small animals, but others denote birds of what would now be an unusually large size, one having a foot fifteen inches in length, and a stride of from four to six feet. There are anomalies in the forms of some of the feet ; but their being the vestiges of birds has for some years been generally admitted. There is, however, some uncertainty re- garding the date of the rocks which present these memorials, for the phenomena of superposition only denote their being between the carboniferous and cretaceous formations, and an exact place is assigned them, merely upon the strength of the discovery that they present fish of certain genera never found above the Triassic series. Along with distinctly ornithic footmarks are those of the Labyrinthodont. Altogether, above thirty species of Triassic birds are made out from these ves- tiges by American geologists. 60 THE TRIAS AND OOLITE. THE chronicles of this period consist of a series of beds, mostly calcareous, taking their general name (Oolite system) from a conspicuous member of them the oolite a limestone composed of an aggregation of small round grains or spherules, and so called from its fancied resemblance to a cluster of eggs, or the roe of a fish. This texture of stone is novel and striking. It is supposed to be of chemical origin, each spherule being an aggregation of particles round a central nucleus. The oolite system is largely developed in England, France, Westphalia, and Northern Italy ; it appears in Northern India and Africa, and patches of it exist in Scotland, and in the vale of the Mississippi. It may of course be yet discovered in many other parts of the world. The series, as shown in the neighbourhood of Bath, is (beginning with the lowest) as follows: 1. Lias, a set of strata variously composed of limestone, clay, marl, and shale, clay being predominant; 2. Lower oolitic formation, in- cluding, besides the great oolitic bed of central England, fullers'-earth beds, forest marble, and cornbrash ; 3. Middle oolitic formation, composed of two sub-groups, the Oxford clay and coral rag, the latter being a mere layer of the works of the coral polype ; 4. Upper oolitic formation, including what are called Kimmeridge clay and Portland oolite. In Yorkshire there is an additional group above the lias, and in Sutherlandshire there is another group above that again. In the wealds (moorlands) of Kent and Sussex, there is, in like manner, above the fourth of the Bath series, another addi- tional group, to which the name of the Wealden has been given, from its topographical situation, and which, composed of sandstones and clays, is subdivided into Purbeck beds, Hastings sand, and Weald clay. There are no particular appearances of disturbance between the close of the Trias, and the beginning of the Lias and Oolite REPTILES ABUNDANT. 61 system, as far as has been observed in England. Yet there is a great change in the materials of the rocks of the two forma- tions, showing that, while the bottoms of the seas of the one period had been chiefly arenaceous, those of the other were chiefly clayey and limy. And there is an equal difference between the two periods in respect of both botany and zoology. While the Permian and Triassic systems, with the single exception of the Muschelkalk, show comparatively scanty traces of life, those in the lias and oolite are extremely abundant, particularly in the department of animals, and more particularly still of sea mollusca. The distinguishing characters of the zoology appear to be uniform over a great space. " In the equivalent deposits in the Himalayan Moun- tains, at Fernando Po, in the region north of the Cape of Good Hope, and in the Run of Cutch, and other parts of Hindostan, fossils have been discovered, which, as far as English naturalists who have seen them can determine, are undistinguishable from certain oolite and lias fossils of Europe. ( 33 ) The dry land of this age presented cycadese, " a beautiful class of plants between the palms and conifers, having a tall, straight trunk, terminating in a magnificent crown of foliage." ( 36 ) There were tree ferns, but in smaller pro- portion than in former ages ; also equisetaceae, lilia, and coniferse. The vegetation was generally analogous to that of the Cape of Good Hope and Australia, which seems to argue a climate between the tropical and temperate. It was, how- ever, sufficiently luxuriant, in some instances, to produce thin seams of coal, for there are such in the oolite formation of both Yorkshire and Sutherland. The sea, as for ages before, contained algae, of which, however, only a few species have been preserved to our day. The lower marine animals present themselves in great abundance, and in some interesting varieties of form. Corals, absent in the lias, reappear in the oolite in quantity sufficient, at some places, as we have seen, to constitute entire strata. The crinoids are also numerous, and amongst these are new genera showing an advance of organization from those of 62 THE TRIAS AND OOLITE. preceding systems. The pentacrinite, instead of a round, has a five-angled stalk, with an increased profusion of tentacula ; it had also the superior character of a power to float about, and attach itself where it pleased. To this fossil of the lias succeed others of the same family in the oolite comatula arid ophiura which are entirely free-swimming, thus supporting the general appearances of an advance of animal characters as we proceed from lower to higher formations. Here also appear other examples of the order to which the crinoidea belong (echinodermata) ; namely, the echinus, or sea-urchin, and the goniaster, which last is regarded as a link between the echinus and star-fish. Among the crustaceans of the oolite, a conspicuous place is due to the limulus, or king-crab, of which several species occur in this formation. This animal is remarkable as the genus of our time to which the trilobite makes the nearest approach ; and the appearance of the limulus at the time when the trilobite vanishes, (the carbonigenous era,) is spoken of by a distinguished geologist as " one of those beautiful links in natural history, of which the strata forming the earth's crust have afforded so many proofs." ( 37 ) Here also we have, in the eryon, an early example of the highest crustacean order, (decapoda,) and one to which the modern lobster and cray-fish belong. Insects resembling the dragon- fly have been found in the oolite. The deeper oolitic seas were occupied by various species of terebratula, a brachiopodous mollusk remarkable as having lived in one form or another from the earliest to the present time. In the shallower seas were other bivalves. There was also abundance of all the univalve classes, Pteropoda, Gastero- poda, and Cephalopoda. Of the last we see an advance of characters in the ammonites and belemnites, which now appear in many varieties. The belemnite, which belonged to the higher order of the class, those having only two branchiae, calls fur some particular notice. It is an elongated, conical shell, terminating in a point, and having, at the larger end, a cavity for the residence of the animal, with a series of air-chambers below. The animal, placed in the upper cavity, REPTILES ABUNDANT. 63 could raise or depress itself in the water at pleasure by a pneumatic operation upon the air tube pervading its shell. Its tentacula, sent abroad over the summit of the shell, searched the sea for prey. The creature had an ink bag with which it could muddle the water around it, to protect itself from more powerful animals, and strange to say, this has been found so well preserved, that an artist has used it in one instance as a pigment, wherewith to delineate the belemnite itself. There are many fishes, some of which (acrodus, psammodus, &c.) are presumed, from remains of their palatal bones, to have been of the gigantic cartilaginous class, (placoidean,) now represented by such as the cestraceon. It has been con- sidered by Professor Owen as worthy of notice, that, the cestraceori being an inhabitant of the Australian seas, we have, in both the botany and ichthyology of this period, an analogy to that Continent. The pycnodontes, (thick toothed,) and lepidoides, (having thick scales,) are other families de- scribed by M. Agassiz as extensively prevalent. In the English lias there is a vast abundance of the enalio- sauria which we have seen commence in the foreign Mus- chelkalk, and, in addition to these, specimens of Pterosauria or Winged Saurians, a type of being, the most new, perhaps, of all which the geological record has presented to us. The Pterodactyls, as the animals of this order have been called, were saurians of small size compared with their associates, being not larger than a modern cormorant ; but the marvel in their case consists in bat-Lke wings extended upon the fore- finger, by which the animal was enabled to pursue its way in the air. This order became extinct in the time of the chalk formation. The only existing animal of which it may even remind us is the draco volans or flying lizard, which has a membrane by which to support itself in leaping from tree to tree. In the proper oolite, there is added an enaliosaurian (the Pliosaur} in which there is a very close approach made to the crocodilian order, but upon a scale of enormous mag- nitude, the animd having apparently been as large as the ex- 64 THE TRIAS AND OOLITE. isting whales. Here, too, we find the true crocodilia largely developed, and five genera have been described, (Teleosaurus, Steneosaurus, Cetiosaurus, &c.) The two first are like croco- diles of our own time in all respects, except a somewhat greater bulk, and certain peculiarities, indicating more aquatic habits. The last derives its name from the approximation to the whale tribes seen in the form of its vertebrae. In this group there is a genus presenting ball-and-socket vertebrae, and thus proving its advanced character ; but, strange to say, the concavity is in this case directed backwards, instead of forwards, which is the universal arrangement in similar cases, in our era. The first glimpse of the highest class of the vertebrate sub- kingdom Mammalia is obtained from the Stonesfield slate, where there have been found several specimens of the lower jaw-bone of a quadruped evidently insectivorous, and inferred, from peculiarities of structure, to have belonged to the marsu- pial family, (pouched animals.) ( 3S ) It may be observed, although no specimens of so high a class of animals as mam- malia are found earlier, such may nevertheless have existed : the defect may be in our not having found them ; but, other things considered, the probability is that heretofore there were no mammifers. It is an interesting circumstance that the first mammifers found should have belonged to the mar- supialia, when the place of that order in the scale of creation is considered. In the imperfect structure of their brain, deficient in the organs connecting the two hemispheres and in the mode of gestation, which is only in small part uterine this family is usually regarded as only a little advanced above the character of the bird. The highest part of the oolitic formation presents some phenomena of an unusual and interesting character, which demand special notice. Immediately above the upper oolitic group in Buckinghamshire, in the vicinity of Weymouth, and other situations, there is a thin stratum, usually called by workmen the dirt-bed, which appears, from incontestable evi- dence, to have been a soil formed, like soils of the present day, in the course of time, upon a surface which had previously REPTILES ABUNDANT. 65 been the bottom of the sea. The dirt-bed contains exuviae of tropical trees, accumulated through time, as the forest shed its honours on the spot where it grew, and became itself decayed. Near Weymouth there is a piece of this stratum, in which stumps of trees remain rooted, mostly erect or slightly inclined, and from one to three feet high ; while trunks of the same forest, also silicified, lie imbedded on the surface of the soil in which they grew. Above this bed lie those which have been called the Wealden, from their full development in the Weald of Sussex ; and these as incontestable argue that the dry land forming the dirt-bed had next afterwards become the area of brackish estuaries or lakes partially connected with the sea ; for the Wealden strata contain exuviae of fresh- water tribes, besides those of the great saurians and chelonia. The area of this estuary comprehends the whole south-east province of Eng- land. A geologist thus confidently narrates the subsequent events : " Much calcareous matter was first deposited [in this estuary], and in it were entombed myriads of shells, appa- rently analogous to those of the vivipara. Then came a thick envelope of sand, sometimes interstratified with mud ; and, finally, muddy matter prevailed. The solid surface beneath the waters would appear to have suffered a long continued and gradual depression, which was as gradually filled, or nearly so, with transported matter ; in the end, however, after a depression of several hundred feet, the sea again entered upon the area, not suddenly or violently for the Wealden rocks pass gradually into the superincumbent cretaceous series but so quietly, that the mud containing the remains of terrestrial and fresh-water creatures was tranquilly covered up by sands replete with marine exuviae." ( 39 ) A subsequent depression of the same area, to the depth of at least three hundred fathoms, is believed to have taken place, to admit of the deposition of the cretaceous beds lying above. From the scattered way in which remains of the larger terrestrial animals occur in the Wealden, and the intermix- ture of pebbles of the special appearance of those worn in rivers, it is also inferred that the estuary which once covered r 66 THE TRIAS AND OOLITE. the south-east part of England was the mouth of a river of that far-descending class of which the Mississippi and Amazon are examples. What part of the earth's surface presented the dry land through which that and other similar rivers flowed, no one can tell. It has been surmised, that the par- ticular one here spoken of may have flowed from a point not nearer than the site of the present Newfoundland. Professor Phillips has suggested, from the analogy of the mineral com- position, that anciently elevated coal strata may have com- posed the dry land from which the sandy matters of these strata were washed. Such a deposit as the Wealden almost necessarily implies a local, not a general condition ; yet it has been thought that similar strata and remains exist in the Pays de Bray, near Beauvais. This leads to the supposition that there may have been, in that age, a series of river-receiving estuaries along the border of some such great ocean as the Atlantic, of which that of modern Sussex is only an example. The zoology of the Wealden is chiefly remarkable for the additions which it makes to the list of reptiles presented in previous formations. Besides some new crocodilia (Sucho- saurus and Goniopholis), and several chelonia (Tetrosternon, etc.), we have here the principal constituents of a group, which Professor Owen has described as a distinct order, under the name of Dinosauria, the remaining form being the Mega- losaurm of the oolite. These were terrestrial crocodile-like animals, with some features of organization recalling the lacertilia, and also such a massive and stately form of the ex- tremities, as to remind us of the large land pachyderms. The animal last named, from twenty-five to thirty feet long, with an enormous muzzle furnished with strong teeth, must have been by far the most formidable land creature of its age. The very opposite habits of the Iguanodon, an equally huge herbi- vorous reptile, lead me to suspect an error in the classifica- tion : but passing from this its size and stately limbs are such as equally to excite our surprise. From the scapula or blade-bone of the remaining genus, the Hylceosaurus, the ap- proximation of the whole of the dinosaurs to the mammalian type of structure has been inferred. REPTILES ABUNDANT. 67 The imagination eagerly aspires to picture the world of the Oolitic Era, when there were scarcely any living creatures of more exalted character than reptiles. There were then vast tracts of dry land, as now ; their surface bore a luxuriant vegetation of no mean kind. The meteoric agencies, the rise and fall of tides, were common phenomena of that time, as of the present. Day after day, through long drawn ages, the sun passed on his course. Night after night, the sparkling garniture of the sky looked down on this green world. But a being of superhuman intelligence, coming to examine our globe, would have seen all this existing only for fishes and still humbler creatures in the sea, and for reptiles, insects, and perhaps a few birds, and still fewer opossums, upon land. He would have beheld the tyrant sauria pursuing their car- nivorous instincts upon the wave, upon the shore, and even in the air ; huge turtles creeping along the muddy coasts ; still more huge megalosaurs traversing the plain ; frog-like ani- mals of the bulk of modern boars, croaking in the marshes ; and with all this, the air filled with multitudes of insects. But no flocks would have met his eye upon the mountains, no herds quietly roaming in the valleys. He would encounter no tiger or elephant in the jungle. None of the smaller mam- malian quadrupeds, as the dog, the genet, the hedgehog, the hare, the mole, would have presented themselves. And not only were no human beings to be seen, but our supernatural visitant would know that this scene must lie spread out in perfect capability for their reception, during ages upon ages, before such beings were to exist; the stream flowing and glittering in the sun, but not to cheer the eye of man ; the season passing, but not to yield its fruit to him ; the whole jocund earth spread out in unenjoyed beauty, as yet unwitting of the glory and the gloom which human impulses were to bring upon it. How strange to reflect on the contemplations of the supposed visitant. What a vast void ! What a stretch of time before there was to be even a commencement to its proper filling ! And yet the certainty that in good time, in the ripeness of the plans of the mighty Author, the higher animals were to come, and among the last the Creature of r2 68 THE TRIAS AND OOLITE. Creatures who, in his infinity of device, was to turn it all to his use the historical being of the world ! It has been sup- posed by some geologists, that there was a special adaptation of the earth at this time to its predominating tenants, as if it presented only low muddy coasts and marshes fit for the residence of reptiles. And it has been thought that this state of the earth is what led to the existence of so many reptiles. But all such speculations rest on insecure grounds. When we consider that the Age of Reptiles, as it has been called, is interposed between an age of fishes and an age of mammals, reptiles being also intermediate to these in the animal scale, we cannot but surmise that the fact depends on some organic law, rather than upon one in physical geography. An obser- vation of some importance to this question is made by Mr. Darwin in his Journal. Describing the Galapagos islands in the Pacific Ocean, where turtles and lizards replace the her- bivorous mammalia, and are the predominating forms of life, he says " The geologist, on hearing this, will probably refer back his mind to the secondary epochs, when lizards, some herbivorous, some carnivorous, and of dimensions comparable only with our existing whales, swarmed on the land and in the sea. It is, therefore, worthy of his observation, that this archipelago, instead of possessing a humid climate and rank vegetation, cannot be considered otherwise than extremely arid, and, for an equatorial region, remarkably temperate." CRETACEOUS ERA. THE record of this period consists of a series of strata, in which chalk beds make a conspicuous appearance, and which is therefore called the Cretaceous System or formation. In England, a long stripe, extending from Yorkshire to Kent, presents the cretaceous beds upon the surface, generally lying conformably upon the oolite, and in many instances rising into bold escarpments towards the west. The celebrated cliffs of Dover are of this formation. It extends into Northern France, and thence north-westward into Germany, whence it is traced into Scandinavia and Russia. The same system exists in Xorth America, and probably in other parts of the earth not yet geologically investigated. Being a marine deposit, it establishes that seas existed at the time of its formation on the tracts occupied by it, while some of its organic remains prove that, in the neighbourhood of those seas, there were tracts of dry land. The cretaceous formation in England presents beds chiefly sandy in the lowest part, chiefly clayey in the middle, and chiefly of chalk in the upper part, the chalk beds being never absent, which some of the lower are in several places. In the vale of the Mississippi, again, the true chalk is wholly, or all but wholly absent. In the south of England, the lower beds are (reckoning from the lowest upwards), 1. Shankland or greensand, " a triple alternation of sands and sandstones with clay ;" 2. Gait, " a stiff blue or black clay, abounding in shells, which frequently possess a pearly lustre ;" 3. Hard 70 CRETACEOUS ERA. chalk ; 4. Chalk with flints ; these two last being generally white, hut in some districts red, and in others yellow. The whole are, in England, about 1200 feet thick, showing the considerable depths of the ocean in which the deposits were made. Chalk is a carbonate of lime, and the manner of its pro- duction in such vast quantities was long a subject of specula- tion among geologists. Some light seemed to be thrown upon the subject a few years ago, when it was observed, that the detritus of coral reefs in the present tropical seas gave a powder, undistinguishable, when dried, from ordinary chalk. It then appeared likely that the chalk beds were the detritus of the corals which lived in the oceans of that era. Mr. Dar- win, who made some curious inquiries on this point, further suggested, that the matter might have intermediately passed through the bodies of worms and fish, such as feed on the corals of the present day, and in whose stomachs he has found impure chalk. This, however, cannot be a full explanation of the production of chalk, if we admit some more recent dis- coveries of Professor Ehrenberg. That master of microscopic investigation announces, that chalk is composed partly of " inorganic particles of irregular elliptical structure and gra- nular slaty disposition," and partly of shells of inconceivable minuteness, " varying from the one-twelfth to the two hun- dred and eighty- eighth part of a line" a cubic inch of the substance containing above ten millions of them ! The chalk of the north of Europe contains, he says, a large proportion of the inorganic matter ; that of the south, a larger proportion of the organic matter, being in some instances almost entirely composed of it. He has been able to classify many of these creatures, some of them being allied to the nautili, nummuli, cyprides, &c. The shells of some are calcareous, of others siliceous. M. Ehrenberg has likewise detected microscopic sea-plants in the chalk. The distinctive feature of the uppermost chalk beds in England is the presence of flint nodules. These are generally disposed in layers parallel to each other. It was readily pre- CRETACEOUS ERA. 71 sumed by geologists that these masses were formed by a chemical aggregation of particles of silica, originally held in solution in the mass of the chalk. But whence the silica in a substance so different from it ? Ehrenberg suggests that it is composed of the siliceous coverings of a portion of the microscopic creatures, whose shells he has in other instances detected in their original condition. It is remarkable that the chalk with flint abounds in the north of Europe ; that without flints in the south ; while in the northern chalk siliceous animalcules are wanting, and in the southern present in great quantities. The conclusion seems natural, that in the one case the siliceous exuviae have been left in their original form ; in the other, dissolved chemically, and aggre- gated on the common principle of chemical affinity into nodules of flint, probably concentrating, in every instance, upon a piece of decaying organic matter, as has been the case with the nodules of ironstone in the earlier rocks, and the spherules of the oolite. What is more remarkable, M. Ehrenberg has ascertained that at least fifty-seven species of the microscopic animals of the chalk, being infusoria and calcareous-shelled polythalamia, are still found living in various parts of the earth. These species are the most abundant in the rock. Singly they are the most unimportant of all animals, but in the mass, forming as they do such enormous strata over a large part of the earth's surface, they have an importance greatly exceeding that of the largest and noblest of the beasts of the field. Moreover, these species have a peculiar interest, as the only specific types of that early age which have survived to the present day. While the specific features of all higher ani- mals have been again and again changed since that period, these humble creatures have preserved the characters they then possessed shall we say, through a continuing unifor- mity in the conditions under which they have lived, while all other animals have been exposed to circumstances productive of change ? All the ordinary and more observable orders of the inhabi- 72 CRETACEOUS ERA. tants of the sea, except the cetacea, have been found in the cretaceous formation zoophytes, radiaria, mollusks, crus- tacea (in great variety of species), and fishes in smaller variety. Down to this period, the placoid and ganoid fishes had, as far as we have evidence, flourished alone ; now they decline, and we begin to find in their place fishes of two orders of superior organization, those which predominate in the present creation. These are osseous in internal structure, with corneous scales. The enaliosaurians disappear in this formation, while the land reptiles, so numerous in the two preceding periods, become much diminished in numbers. Of the latter, one of the most remarkable was the mosaesaurus, which seems to have held an intermediate place between the monitor and iguana, and to have been about twenty-five feet long, with a tail calculated to assist it powerfully in swim- ming. Fuci abounded in the cretaceous seas, and confervas are found enclosed in flints. Of terrestrial vegetation, as of ter- restrial" animals, the specimens in the European area are com- paratively rare, rendering it probable that there was little dry land near. The remains are chiefly of ferns, conifers, and cycadese, but in the two former cases we have only cones and leaves. There have been discovered many pieces of wood containing holes drilled by the teredo, and thus showing that they had been long drifted about in the ocean before being entombed at the bottom. The series in America corresponding to this, entitled the Ferruginous Sand formation, presents fossils generally iden- tical with those of Europe, not excepting the fragments of drilled wood ; showing that, in this, as in earlier ages, there was a parity of conditions for animal life over a vast tract of the earth's surface. To European reptiles, the American formation adds a gigantic one, styled the Saurodon, from the lizard-like character of its teeth. We have seen that footsteps of birds have been announced from America, in the new red sandstone. Some similar isolated phenomena occur in the subsequent formations. In CRETACEOUS ERA. 73 the slate of Glaris, in Switzerland, corresponding to the Eng- lish gait, in the chalk formation, the remains of a bird have been found. From a chalk bed, near Maidstone, have like- wise been extracted some remains of a bird, supposed to have been of the long- winged swimmer family, and equal in size to the albatross. 74 ERA OF THE TERTIARY FORMATION- MAMMALIA ABUNDANT. THE chalk-beds are the highest which extend over a consi- derable space; but in hollows of these beds, comparatively limited in extent, there have been formed series of strata clays, limestones, marls, alternating to which the name of the Tertiary Formation has been applied. London and Paris alike rest on basins of this formation, and another such basin extends from near Winchester, under Southampton, and re- appears in the Isle of Wight. A stripe of it extends along the east coast of North America, from Massachusetts to Florida. It is also found in Sicily and Italy, insensibly blended with formations still in progress. Though compara- tively a local formation, it is not of the less importance as a record of the condition of the earth during a certain period. The hollows filled by the tertiary formation must be consi- dered as the beds of estuaries and gulfs, left at the conclusion of the cretaceous period. We have seen that an estuary, either by the drifting up of its mouth, or a change of level in that quarter, may be supposed to have become an inland sheet of water, and that, by another change of the reverse kind, it may be supposed to have become an estuary again. Such changes the Paris basin appears to have undergone oftener than once, for, first, we have there a fresh- water formation of clay and limestone beds ; then, a marine-limestone formation ; MAMMALIA ABUNDANT. 75 next, a second fresh- water formation, in which the material of the celebrated plaster of Paris (gypsum) is included ; then a second marine formation of sandy and limy beds ; and finally, a third series of fresh-water strata. Such alternations occur in other examples of the tertiary formation likewise. The end of the Secondary Formation, which we have just seen take place, presents in some respects a remarkable re- semblance to the close of what is called the Palaeozoic period in the Permian strata. Looking broadly at the specific forms of the next higher strata, they appear to have undergone a total change. Again do we now witness a difference of the shelly cephalopoda. There is also a gradual reduction and finally a disappearance of the specific forms of gasteropods, formerly abundant. It has heretofore been a belief of geolo- gists, that at this point, as at the former, there was an entire renewal of life upon our planet ; but several considerations forbid such a conclusion in the second as well as in the first instance. First, the specific forms are not wholly changed, for a few do pass into the next higher strata. Second, there is, in the higher formation, an apparent following of an order applicable to the whole palaontological history, as something under one law, seeing that birds and mam- malia, the next classes in the vertebrate scale, are then added. In the words of Sir R. Murchison, who believes that a true geological passage may be found between the two for- mations, the upper secondary rocks judging from many of their generic forms " seem to have prepared the way for the sequence of the tertiary strata." For these reasons, the idea of an entire renovation of life at this time what is commonly called a new creation is not now maintained anywhere with confidence. The more rational explanation of the appearances is one suggested by actual facts observed in the strata ; that the final cretaceous beds were deposited in seas more than usually deep, and which were therefore no proper habitat for the animals previously existing ; that an interval of time afterwards took place, which is not represented by any strata which have been discovered ; and that, by the time the ter- tiary formation commenced, the usual modifying influences 76 ERA OF THE TERTIARY FORMATION. having never ceased, the fauna had undergone such an amount of change as naturalists are accustomed to describe (their language being wholly arbitrary) as a renewal of species. It is in perfect harmony with this view, that from the com- mencement of the Tertiaries, and as we ascend in the series, we find more and more specific forms identical with those still existing upon earth, as if we had now reached the dawn of the present state of the zoology of our planet. By the study of the shells alone, Mr. Lyell has formed a division of the whole term into four sub-periods, to which he has given names with reference to the proportions which they respec- tively present of surviving species first, eocene; second, miocene ; third, older pliocene ; fourth, newer pliocene. ( 40 ) This division, however, is to be regarded as not safely appli- cable to the Tertiaries generally, except as a convenient means of indicating various portions of the series. The eocene period presents, in three continental groups, 1238 species of shells, of which forty-two, or 3*5 per cent, yet flourish unchanged. Some of these are remarkable enough ; but they all sink into insignificance beside the mam- malian remains which the lower eocene deposits of the Paris basin present to us, showing that the land had now become the theatre of an extensive creation of the highest class of animals. Cuvier ascertained about fifty species of these, all of them long since extinct. About four-fifths are of the order Pachydermata, thick-skinned animals, to which our modern elephant, rhinoceros, horse, and pig belong. Nearly the whole of these, however, belong to a family which is now confined to South America and Sumatra, namely, the tapirs, an animal of squat figure, and possessing a short pro- boscis, an inhabitant of the woods, and. an herbivore, but of unsocial habits. It is curious to find that a family now so limited in its range, had formerly been distributed over France, England, and other parts of the earth. Naturalists have conferred the names, Palaeotherium, Lophiodon, Cory- phodon, &c., upon the ancient extinct tapirs, which seem chiefly to differ from modern species in a few peculiarities of the constitution of the teeth, and in having three, instead of MAMMALIA ABUNDANT. 77 four toes upon the fore feet. One British specimen seems to have been about a third larger than the modern animal. Another section of the Paris eocene remains have served to reconstruct a family to which the general name Anoplo- therium has been given, from a regard to its deficiency of all offensive or defensive weapons. These are the first examples of bi-hooved animals as yet discovered upon earth ; they were strictly herbivorous, and make a slight approach to the cervine or deer tribes. The common anoplothere was about the size of an ass, but less elevated from the ground, and with a tail of above three feet in length ; it is supposed to have been of aquatic habits, and an expert swimmer and diver, but also given to browsing upon land. Associated with these we find the first example (choeropotamus) of an animal ap- proaching to the hog tribe, being nearest to the peccary of South America. We learn from the remainder of the Paris fossils, and from others found in the eocene, that the earth now possessed fresh- water reptiles ; serpents of the size of the boa ; natatorial, wading, and rapacious birds ; rodents (dormouse and squirrel) ; species allied to the racoon, the genet, and fox ; also bats and monkeys. Lastly, the oldest tertiaries of America present us with the Zeuglodon, a herbivorous whale resembling the dugong, having a stinted development of the extremities, but an enormous tail, and reaching altogether the length of a hundred feet. In the miocene sub-period, the shells give eighteen per cent, of existing species, showing a considerable advance from the preceding era with regard to the inhabitants of the sea. The advance in land animals is less marked, but yet considerable. The predominating forms are still pachyderms, and the tapiroid animals continue to be conspicuous. Here occur remains of the Dinotherium, a creature said to exhibit an affinity to the cetacea in the form of its head, and to the tapir in the character of its teeth. It is most distinguished by its huge size, being not less than eighteen feet long ; it had a mole-like form of the shoulder-blade, conferring the power of digging for food, and a couple of tusks turning down from 78 ERA OF THE TERTIARY FORMATION. the lower jaw, by which it could have attached itself, like the walrus, to a shore or bank, while its body floated in the water. Dr. Buckland considers this and some similar miocene animals, as adapted to a semi-aquatic life, in a region where lakes abounded. Besides the tapirs, we have in this era animals allied to the glutton, the bear, the dog, the horse, the hog, and lastly, several felinse (creatures of which the lion is the type) ; all of which are new forms, as far as we know. There was also an abundance of marine mammalia, seals, dolphins, lamantins, walruses, and whales. The shells of the older pliocene give from thirty-five to fifty ; those of the newer, from ninety to ninety-five per cent, of ex- isting species. The pachydermata of the preceding era now dis- appear; but others enter upon the scene elephantoid animals, the hippopotamus, rhinoceros, and horse. All of these bear a striking resemblance to pachyderms of the same families still existing. We have, in the mastodon and mammoth, which succeed each other in the strata, elephants variously distin- guished from the present by peculiarities in their dentition, and hence considered as of different species, though this is a purely arbitrary distinction. What is remarkable of these ancient animals is their having lived in countries so far beyond the present range of their family, namely, throughout the whole temperate region of Asia and Europe, (England not being excepted,) and even so far north as the seventieth degree of latitude. The mammoth also inhabited North America. Its chief external peculiarity was a pair of long curved tusks extending forwards and upwards from the upper jaw. The numerous remains of the animal in the most super- ficial strata, and the discovery (in 1801) of a specimen with its flesh and hide entire in a mass of ice at the mouth of the Lena in Siberia, show that it must have lived down to compa- ratively modern times. The pliocene gives many other new families. From remains which have been found, however fragmentary in many cases, there cannot be a doubt that all the principal mamma- lian forms, except the highest and a few others, now existed throughout the earth, and in species which only differed from MAMMALIA ABUNDANT. 79 those now living in slight peculiarities, chiefly of dentition. Bears, badgers, hyaenas, and feline animals ; moles and other insectivores ; otters and weasels ; the wolf and dog, then roamed for prey as now ; besides an extinct felina, the machairodus, possessing teeth like curved saws. England had beavers and bears, little different from living species; only, one of the former family was of huge bulk. We also had the hippopotamus and rhinoceros. Oxen, deer, camels, etc., inhabited the great zoological province with which we are connected ; and monkeys and apes passed far beyond the tropical regions to which they are now confined. In India, besides the pachyderms of the European eocene, there were ruminants in abundance (including an extraordinary one, of huge bulk, named the Sivatherium), carnivores, rodents, and insectivores. Here also were monkeys, of unusual bulk ; but the most wonderful animal as yet discovered in this region was a tortoise, not distinguishable in any point of structure from a land species now living, but reaching the surprising length of eighteen feet. The discoveries among the tertiaries of South America have been of a not less interesting cha- racter, in as far as they equally show an approach to the existing zoological characters of that region. Dr. Lund, a Danish naturalist, presents us with a monkey, indicating the features of the platyrrhine or New World group ; and the edentate order, which is still more peculiar to that region, is there preceded by examples of vast size. In the megatherium, megalonyx, scelidotherium, and mylodon, we have a family of sloths, of elephantine magnitude, which lived by breaking down and eating trees. The toxodon surprises us not less, being an equally huge member of the rodent order, that order which now includes most of the smallest quadrupeds. ( 41 ) One remarkable circumstance connected with the tertiary formation remains to be noticed, the prevalence of volcanic action at that era. In Auvergne, in Catalonia, near Venice, and in the vicinity of Rome and Xaples, lavas exactly re- sembling the produce of existing volcanoes are associated and intermixed with the lacustrine as well as marine tertiaries. The superficies of tertiaries in England is disturbed by two 80 ERA OF THE TERTIARY FORMATION. great swells, forming what are called anticlinal axes, one of which divides the London from the Hampshire basin, while the other passes through the Isle of Wight, both throwing the strata down at a violent inclination towards the north, as if the subterranean disturbing force had waved forward in that direction. The Pyrenees, too, and Alps, have both un- dergone elevation since the deposition of the tertiaries; and in Sicily there are mountains which have risen three thousand feet since the deposition of some of the most recent of these rocks. The general effect of these operations was of course to extend the land surface, and to increase the variety of its features, thus improving the natural drainage, and generally adapting the earth for the reception of higher classes of animals. 81 ERA OF THE SUPERFICIAL FORMATIONS EXISTING SPECIFIC FORMS ABUNDANT. WE have now completed our survey of the series of stratified rocks, and traced in their fossils the progress of organic crea- tion down to a time which seems not long antecedent to the appearance of man. There are, nevertheless, memorials of still another era or space of time which it is all but certain did also precede that event. The first that calls for notice is the phenomenon to which geologists have applied the term denudation. Great hitches and slips are detected in superficial strata, such as, if left in their original state, must have caused considerable inequalities on the face of the country ; yet all is found as smooth the joinings are all as much reduced to a common level as if some gigantic artificial force had been used for the purpose. Again, a great valley has been scooped out in the midst of sedimentary strata, leaving the edges of these facing each other from the opposite sides, with perhaps here and there an isolated mass starting up to the height of the two sides, being composed of matter which has resisted the agency by which the adjoining matter was removed. Here, it is thought, we see incontestable traces of the operation of moving water. The second fact we are called to notice is, that over the rock formations of all eras, in various parts of the globe, but con- fined in general to situations not very elevated, there is a layer of stiff clay, mostly of a blue colour, mingled with fragments of rock of all sizes, travel-worn, and otherwise, and to which geologists give the name of Diluvium, as being 82 ERA OF THE SUPERFICIAL FORMATIONS. apparently the produce of some vast flood, or of the sea thrown into an unusual agitation. It seems to indicate that, at the time when it was laid down, much of the present dry land was under the ocean a supposition which we shall see supported by other evidence. The included masses of rock have been carefully inspected in many places, and traced to particular parent beds at considerable distances. Connected with these phenomena are certain rock surfaces on the slopes of hills and elsewhere, which exhibit groovings and scratchings, such as we might suppose would be produced by a quantity of loose blocks hurried along over them by a flood. Another associated phenomenon is that called crag and tail, which exists in many places, namely, a rocky mountain, or lesser elevation, presenting on one side the naked rock in a more or less abrupt form, and on the other a gentle slope ; the sites of Windsor, Edinburgh, and Stirling, with their respective castles, are specimens of crag and tail. Finally, I may advert to certain long ridges of clay and gravel which arrest the attention of travellers on the surface of Sweden and Finland, and which are also found in the United States, where, indeed, the whole of these phenomena have been observed over a large surface, as well as in Europe. It is very remarkable that the direction from which the diluvial blocks have generally come, the lines of the grooved rock surfaces, the direction of the crag and tail eminences, and that of the clay and gravel ridges phenomena, be it ob- served, extending over the northern parts of both Europe and America are all from the north and north-west towards the south-east. We thus acquire the idea of a powerful current moving in a direction from north-west to south-east, carrying, besides mud, masses of rock which furrowed the solid surfaces as they passed along, abrading the north-west faces of many hills, but leaving the slopes in the opposite direction unin- jured, and in some instances forming long ridges of detritus along the surface. These are curious considerations ; and it has become a question of much interest, by what means, and under what circumstances, such a current was produced. But in the present state of our knowledge, all that can be EXISTING SPECIFIC FORMS ABUNDANT. 83 legitimately inferred from the diluvium is, that many portions of the northern nations of Europe and America were then under the sea, and that a strong current set over them. Connected with the Diluvium is the history of Ossiferous Cavprns, of which specimens singly exist at Kirkdale in Yorkshire, Gailenreuth in Franconia, and other places. They occur in the calcareous strata, as the great caverns generally do, but have in all instances been naturally closed up till the recent period of their discovery. The floors are covered with what appears to be a bed of the diluvial clay, over which rests a crust of stalagmite, the result of the drop- pings from the roof since the time when the clay bed was laid down. In the instances above specified, and several others, there have been found, under the clay bed, assem- blages of the bones of animals, of many various kinds. At Kirkdale, for example, the remains of twenty-four species were ascertained namely, pigeon, lark, raven, duck, and par- tridge ; mouse, water-rat, rabbit, hare, hippopotamus, rhino- ceros, elephant, weasel, fox, wolf, deer (three species), ox, horse, bear, tiger, hyena. From many of the bones of the gentler of these animals being found in a broken state, it is supposed that the cave was a haunt of hyenas and other pre- daceous creatures, by which the smaller ones were here con- sumed. This must have been at a time antecedent to the floodings which produced the diluvium, since the bones are covered by a bed of that formation. It is impossible not to see here a very natural series of incidents. First, the cave is frequented by wild beasts, who make it a kind of charnel- house. Then, submerged in the current which has been spoken of, it receives a clay flooring from the waters contain- ing that matter in suspension. Finally, raised from the water, but with no mouth to the open air, it remains unin- truded on for a long series of ages, during which the clay flooring receives a new calcareous covering, from the drop- pings of the roof. Our attention is next drawn to the erratic blocks or boulders, which in many parts of the earth are thickly strewn over the surface, particularly in the north of Europe. Some G2 84 ERA OF THE SUPERFICIAL FORMATIONS. of these blocks are many tons in weight, yet are clearly as- certained to have belonged originally to situations at a great distance. Fragments, for example, of the granite of Shap Fell are found in every direction around to the distance of fifty miles, one piece being placed high upon Criffel Moun- tain, on the opposite side of the Solway estuary ; so also are fragments of the Alps found far up the slopes of the Jura. There are even blocks on the east coast of England, supposed to have travelled from Norway. The only rational conjec- ture which can be formed as to the transport of such masses from so great a distance, is one which presumes them to have been carried and dropped by icebergs, while seas existed upon the space between their original and final sites. Icebergs do even now carry off such masses from the polar coasts, which, falling when the retaining ice melts, must take up situations at the bottom of the sea, similar to those in which we find the erratic blocks of the present dry land. While the diluvium and erratic blocks clearly suppose a part at least of the present land to have at one time been sunk to a considerable depth in the sea, there is another set of ap- pearances which as manifestly show the steps by which the land was made afterwards to re-emerge from that element. These consist of terraces, which have been detected near, and at some distance inland from, the coast lines of Scandinavia, Britain, America, and other regions ; being evidently ancient beaches, or platforms, on which the margin of the sea at one time rested. They have been observed at different heights above the present sea-level, from twenty to above twelve hundred feet ; and in many places they are seen rising above each other in succession, to the number of three, four, and even more. The smooth flatness of these terraces, with ge- nerally a slight inclination towards the sea, the sandy compo- sition of many of them, and, in some instances, the preser- vation of marine shells in the ground, identify them perfectly with existing sea-beaches, notwithstanding the cuts and scoopings which have at frequent intervals been effected in them by water-courses. The irresistible inference from the phenomena is, that the highest was first the coast line ; then EXISTING SPECIFIC FORMS ABUNDANT. 85 an elevation took place, and the second highest became so, the first being now raised into the air and thrown inland. Then, upon another elevation, the sea began to form, at its new point of contact with the land, the third highest beach, and so on down to the platform nearest to the present sea- beach. Phenomena of this kind become comparatively familiar to us, when we hear of evidence that the last sixty feet of the eleva- tion of Sweden, and the last eighty-five of that of Chili, have taken place since man first dwelt in those countries ; nay, that the elevation of the former country goes on at this time at the rate of about forty-five inches in a century, and that a thousand miles of the Chilian coast rose four feet in one night, under the influence of a powerful earthquake, so lately as 1822. Subterranean forces, of the kind then exemplified in Chili, supply a ready explanation of the whole phenomena, though some other operating causes have been suggested. In an inquiry on this point, it becomes of consequence to learn some particulars respecting the levels. Taking a parti- cular beach, it is generally observed that the level continues the same along a considerable number of miles, and nothing like breaks or hitches has as yet been detected in any case. A second and a third beach are also observed to be exactly parallel to the first. These facts would seem to indicate quiet elevating movements, uniform over a large tract. It must, however, be remarked that the raised beaches at one part of a coast rarely coincide with those at another part forty or fifty miles off. We might suppose this to indicate a limit in that extent of the uniformity of the elevating cause ; but it would be rash to conclude positively that such is the case. In the present sea, as is well known, there are different levels at different places, owing to the operation of peculiar local causes, as currents, evaporation, and the influx of large rivers into narrow-mouthed estuaries. The differences of level in the ancient beaches might be occasioned by some such causes. But, whatever doubt may rest on this minor point, enough has been ascertained to settle the main one, that we have in these platforms indubitable monuments of an elevation of the 86 ERA OF THE SUPERFICIAL FORMATIONS. land from the sea, and the concluding great event of the geo- logical history. The idea of such a deep immersion of the land unavoidably suggests some considerations as to the effect which it might have upon terrestrial animal life. Some, regarding it as a complete submersion, argue that terrestrial life would be, oh such an occasion, extensively, if not universally, destroyed. Nor was the idea of its universal destruction the less plau- sible, when it was believed that the present land animals are an entirely new set of species, introduced since the conclusion of the Tertiary Formation. It must now be owned that there are insurmountable objections to such hypothesis. First, it is not true that the specific forms of the tertiary epoch have all of them disappeared. There are several for example, a badger of the Miocene which are not in the slightest degree distinguished from living species. Many reptiles, now living in India, have been proved to be coeval with the Himalayan Anoplothere, Mastodon, and Hippopotamus. Second, the specific distinctions alleged in a great number of cases be- tween tertiary and existing animals are extremely slight, and such as we have no fixed principle by which to be assured that they mark new species, in the sense of a new creation. Finally, the tertiary animals of America indicate an approxi- mation to the character of existing animals in that region, and tertiary animals of the other great continent, equally ap- proximate to those at present occupying it ; showing that the demarcations of the present great zoological provinces had been already marked out, and have never been obliterated. There is therefore enough to justify us in believing that no entire submergence of the earth took place at the time of the Diluvium, though how nearly it might approach completeness we cannot say. There are some other superficial formations, of less conse- quence on the present occasion than the diluvium namely, lacustrine deposits, or filled-up lakes ; alluvium, or the de- posits of rivers beside their margins ; deltas, the deposits made by great ones at their efflux into the sea ; peat mosses ; EXISTING SPECIFIC FORMS ABUNDANT. 87 and the vegetable soil. The animal remains found in these generally testify to a zoology on the verge of that now pre- vailing, or melting into it, there being included many species which still exist. In a lacustrine deposit at Market- Weighton, in the Vale of York, there have been found bones of the elephant, rhinoceros, bison, wolf, horse, felis, deer, birds, all or nearly all presenting peculiarities different from existing species, associated with thirteen species of land and fresh- water shells, " exactly identical with types now living in the vicinity." In similar deposits in North America, are remains of the mammoth, mastodon, buffalo, and other animals of extinct and living types. In short, these superfi- cial deposits show precisely such remains as might be ex- pected from a time at which the present forms of the animal world had been generally assumed, but yet so far remote in chronology as to allow of the dropping of many species, through familiar causes perhaps we should only say the obliteration of many peculiarities called specific in the in- terval. Still, however, several of the most important living species have left no record of themselves in any formation beyond what are, comparatively speaking, modern. Such are the sheep and goat, and such, above all, is our own species. We thus learn that, compared with many humbler animals, man is a being, as it were, of yesterday. 88 GENERAL CONSIDERATIONS RESPECTING THE ORIGIN OF THE ANIMATED TRIBES. THUS concludes the wondrous section of the earth's history which is told by geology. It takes up our globe at the period when its original incandescent state had nearly ceased ; conducts it through what we have every reason to believe were vast spaces of time, in the course of which many super- ficial changes took place, and vegetable and animal life was gradually evolved ; and drops it just at the point when man was apparently about to enter on the scene. The compilation of such a history, from materials of so extraordinary a cha- racter, and the powerful nature of the evidence which these materials afford, are calculated to excite our admiration, and the result must be allowed to exalt the dignity of science, as a product of man's industry and his reason. It is now to be remarked, that the whole series of opera- tions displayed in inorganic geology is concluded upon as having taken place under the agency of natural laws. Those movements of subterranean force which thrust up mountain ranges and upheaved continents, stand in inextricable connec- tion, on the one hand, with the volcanoes which are yet belch- ing forth lavas and shaking large tracts of ground, as, on the other, with the primitive incandescent state of the earth. ORIGIN OF THE ANIMATED TRIBES. 89 Those forces which disintegrated the early rocks and of the detritus formed new beds at the bottoms of seas, are still seen at work to the same effect in every part of the globe. To bring these truths the more clearly before us, it is possible to make a substance resembling basalt in a furnace ; lime- stone and sandstone have both been formed from suitable materials in appropriate receptacles ; the phenomena of cleavage have, with the aid of electricity, been similated on a small scale, and by the same agent crystals are formed. In short, the remark which was made regard- ing the indifference of the cosmical laws to the scale on which they operated, is to be repeated regarding the geo- logical. A common furnace will sometimes exemplify the operation of laws which have been concerned in the produc- tion of a Giant's Causeway ; and in a sloping ploughed field after rain, we may often observe, at the lower end of a furrow, a handful of washed and neatly deposited mud or sand, capable of serving as an illustration of the way in which Nature has produced the deltas of the Nile and Ganges. In the ripple-mark on sandy beaches of the present day, we see Nature's exact repetition of the operations by which she im- pressed similar features on the sandstones of the carboni- genous era. Even such marks as wind-slanted rain would in our day produce on tide-deserted sands, have been read upon tablets of the ancient strata. It is the same Nature working every where and in all time, causing the wind to blow and the rain to fall, and the tide to ebb and flow, inconceivable ages before the birth of our race, as now. So also we learn from the conifers of those old ages, that there were winter and summer upon earth, before any of us lived to liken the one to all that is genial in our own nature, or to say that the other breathed no airs so unkind as man's ingratitude. Let no one suppose there is any necessary disrespect for the Creator in thus tracing his laws in their minute and familiar operations. There is in reality no true great and small, grand and familiar, in nature. Such only appear, when we thrust ourselves in as a point from which to start in judging. Let us pass, if possible, beyond immediate impressions, and 90 GENERAL CONSIDERATIONS RESPECTING THE see all in relation to Cause, and we shall chastenedly admit that the whole is alike worshipful. We have, then, in this history, a planet formed, and a long and complicated series of superficial changes effected upon it, all through the efficacy of simply natural laws, which we can see at work at this day in numberless familiar ways. But mixed up with these geognostic changes, and apparently as a final object connected with the formation of the globe itself, there is another set of phenomena presented in the course of our history the coming into existence, namely, of a long suite of living things, vegetable and animal, terminating in the families which we still see occupying the surface. The question arises In what manner has this set of phenomena originated ? Can we touch at and rest for a moment on the possibility of plants and animals having likewise been pro- duced in the way of natural law ; thus assigning but one class of causes for everything revealed to our sensual obser- vation ; or are we at once to reject this idea, and remain con- tent, either to suppose that creative power here acted in a different way, or to believe unexaminingly that the inquiry is one beyond our powers ? Taking the last part of the question first, I would reply, that I am extremely loath to imagine that there is anything in nature which we should, for any reason, refrain from exa- mining. If we can infer aught from the past history of science, it is, that the whole of nature is a legitimate field for the exercise of our intellectual faculties ; that there is a con- nexion between this knowledge and our well-being ; and that if we can judge from things once despaired of by our inquiring reason, but now made clear and simple, there is none of Nature's mysteries which we may not hopefully attempt to penetrate. To remain idly content to presume a various class of immediate causes for organic nature, seems to me, on this ground, equally objectionable. Granting, then, that the inquiry should be entered upon, it may be right to insist, in the first place, upon certain general considerations, which, supposing we enter upon it in a scientific ORIGIN OF THE ANIMATED TRIBES. 91 spirit, appear to throw the balance of likelihood on the side of ordinary natural causes. The production of the organic world is, we see, mixed up with the production of the physical. It is mixed in the sense of actual connexion and dependence, and it is mixed in regard to time, for the one class of phenomena commenced whenever the other had arrived at a point which favoured or admitted of it ; life, as it were, pressed in when- ever and wherever there were suitable conditions, and, once it had commenced, the two classes of phenomena went on, hand in hand, together. It is surely very unlikely, d. priori, that in such a complex mass of phenomena there should have been two totally distinct modes of the exercise of the divine power. Were such the case, it would form a most extraordinary, and to philosophic consideration ought to be a most startling ex- ception, from what we otherwise observe of the character of the divine procedure in the universe. Further, let us con- sider the comparative character of the two classes of phenomena, for comparison may of course be legitimate where the natural system is not admitted. The absurdities into which we should thus be led must strike every reflecting mind. The Eternal Sovereign arranges a solar or an astral system by dispositions imparted primordially to matter; he causes, by the same majestic means, vast oceans to form and continents to rise, and all the grand meteoric agencies to proceed in ceaseless alternation, so as to fit the earth for a residence of organic beings. But when, in the course of these operations, fuci and corals are to be for the first time placed in those oceans, a par- ticular interference of the divine power is required : in the belief of the ignorant, the very hand of Deity is necessary ; in that of the sage as sages are amongst us only a " creative fiat" is demanded ; but, in either way, special attention to the object, such as a human being has to pay in the progress of his affairs, is presumed. And not only on this one occasion, but all along the stretch of geological time, this special atten- tion is needed whenever a new family of organisms is to be introduced : a new fiat for fishes, another for reptiles, a third for birds ; nay, taking up the present views of geologists as to species, such an event as the commencement of a certain 92 GENERAL CONSIDERATIONS RESPECTING THE cephalopod, one with a few new nodulosities and corrugations upon its shell, would, on this theory, require the particular care of that same Almighty who willed at once the whole means by which infinity was replenished with its worlds ! I have here contemplated the question as one remaining to be settled by science. We must not, however, overlook that the decision is exposed to great prejudice, in consequence of our minds being prepossessed by a more or less distinct con- clusion in favour of organic creation by some kind of special exercise of divine power. This is the idea which first rose in the human family, being that which the unassisted mind is apt to form out of the appearances presented to it ; precisely as, with regard to the heavenly motions, the geocentric theory was that which the appearances first suggested, and therefore was first embraced by man. This idea of the organic creation has rested almost undisturbed to the present day, because, till a recent period, science came little near it, and means for testing its soundness scarcely existed. It is different now, when, besides the cosmical arrangements being seen to have been brought about under natural law, the same influence is traced through the whole series of geognostic changes since the beginning of our planet's existence. But yet, this know- ledge being recent, the ancient idea of the creation of organisms continues to have that hold upon our minds which early im- pressions and long-continued habits tend to give to even the most unphilosophical convictions. It is necessary to keep this in view, if we would enter upon the inquiry in a philosophical spirit, and with the pure desire to arrive only at such a con- clusion as the balance of evidence may justify. In intimate connexion with the ancient idea, is the supposi- tion found resting in many minds, that to presume a creation of living beings by the intervention of law, is equivalent to superseding the whole doctrine of the divine authorship of organic nature. Were this true, it would form a most im- portant objection to the law theory ; but I think it is not only not true, but the reverse of the truth. As formerly stated, the whole idea of law relates only to the mode in which the Deity is pleased to manifest his power in the natural world. ORIGIN OP THE ANIMATED TRIBES. 93 It leaves the absolute fact of his authorship of and supremacy over nature, precisely where it was ; and only tells us that, instead of dealing with the natural world as a human being traffics with his own affairs, adjusting each circumstance to a relation with other circumstances as they emerge, he has originally conceived, and since sustained arrangements fitted to serve in a general sufficiency for all contingencies ; himself, of course, necessarily living in all such arrangements, as the only means by which they could be, even for a moment, upheld. Were the question to be settled upon a consideration of the respective moral merits of the two theories, I would say that the latter is greatly the preferable, as it implies a far grander view of the divine power and dignity than the other. For one thing, it places the leading divine attribute of fore- sight in a much more sublime position. " If," says Dr. Buckland, contemplating the possible establishment of this doctrine " if the properties adopted by the elements at the moment of their creation adapted them beforehand to the infinity of complicated useful purposes which they have already answered, and may have still further to answer, under many dispensations of the material world, such an aboriginal constitution, so far from superseding an intelligent agent, would only exalt our conceptions of the consummate skill and power that could comprehend such an infinity of future uses under future systems, in the original groundwork of his crea- tion'' It is also to be observed, though almost needlessly, that any objection of this kind would lie equally well against other doctrines which the enlightened part of mankind have long admitted. The whole purport of science is to ascertain law ; it has, in the course of time, transferred one set of phenomena after another out of the region of marvel into that of law, thus showing a true divine regulation in them. Suppose, then, that there be a balance of probability from actual evidence in favour of an organic creation by law, we should only, in so settling the matter, be concluding upon one department of the great system of things, as we have been concluding upon others throughout all the ages of philosophy. 94 GENERAL CONSIDERATIONS RESPECTING THE To return to a consideration of the positive arguments for an organic creation by law, we have seen that this stands in harmony with our conclusions as to the cosmical arrangements, and also those regarding the geognostic changes. We are now to observe that it consorts equally well with what we know of the actual history of organic beings upon the earth. These came not at once, as they might have been expected to do if produced by some special act, or even some special inter- position of will, on the part of the Deity. They came in a long-extending succession, in the order, as we shall afterwards see more convincingly, of progressive organization; grade following grade, till, from a humble starting-point in both kingdoms, the highest forms were realized. Time, we see, was an element in the evolution of Being, as it is in the repro- duction of an individual at the present day. At the beginning of geological investigation, it was thought that some imme- diate external conditions ruled the appearance of particular classes of animals at particular times : as that the absence of dry land was the cause of the late commencement of terrestrial animals; that there being for a long time only reptilian land vertebrata was owing to an overcharge of the atmosphere with carbonic acid the store from which came the chief material of the abundant vegetation of the carbonigenous age ; and so forth. But it is now seen that the progress of the animal world was, in its main features, independent of such circum- stances. There was dry land unreckonable ages before there were any land animals. The sea abounded in invertebrate animals before there were any fish, though the conditions required for the existence of both are the same. The oolitic continents where only reptiles roamed could have equally supported mammalia, for which the atmosphere was then fully fitted, even upon the admission of the carbonic acid theory, as the coal was by that time formed; yet mammalia came not. It was also a dream at the dawn of true geology, that fresh creations of animals were connected with great physical revolutions of the surface ; as if, at particular times, all had perished in storms of volcanic violence, and been replaced with a wholly new fauna. But this idea is likewise passing away. ORIGIN OF THE ANIMATED TRIBES. 95 It is now seen that changes in specific forms took place quietly in the course of time, while no volcanic disturbances are traceable. In short, it is always becoming more and more clear that organic progress both the specific changes in classes formerly existing, and the accession of new and higher classes depended, not by any means wholly or immediately upon ex- ternal circumstances, but in great part upon time. All this looks very unlike either special working or special willing on the part of the Creator, but, on the contrary, very like the simply natural procedure of things in the world of our own day. There are some other facts in the history of fossils, which it is difficult to reconcile with the idea of special creative effort, but which perfectly harmonize with that of a creation through the medium or in the manner of law. It is admitted, for instance, that " the differences which exist between extinct faunas and the animals now living are so much greater in proportion as these faunas are most ancient" Passing down- ward in the formations and backward in time, we first find species identical with the present ; next, only genera ; after- wards, only families or orders. These are the words of naturalists ; but the truth simply is, that in early formations, animals resembled the present in broad general characters ; afterwards, they resembled them in characters more parti- cular ; finally, they become identical. Always as we advance, the total mass of the animal creation puts on more and more of the appearances which it now bears. It may be asked if this does not seem to imply that the present system of things is essentially connected with the past ; in which case, if the present is a natural system, we have an additional proof that the past was a natural system also. So also it is admitted that, however nearly the specific forms may experience an entire change from one formation to another, there are always resemblances and approximations between each two which are adjacent to each other. " If," says M. Pictet, an opponent of the views here advocated, "we compare two successive creations of one and the same epoch, such as the faunas of the five divisions of the cretaceous formation, we cannot fail to be struck with the intimate connexion they 96 GENERAL CONSIDERATIONS RESPECTING THE have with each other. The greater part of the genera are the same : a great part of the species are very closely allied and easily confounded. [Referring to two of these sub-forma- tions,] is it probable that the albian fauna had been completely annihilated, and then, by a new and independent creation, replaced by a fauna altogether new, and so similar to it ? I am aware that these facts may be referred to the general plan of creation [that is, a supposed plan, according to which the divine power had operated in its special successive creative operations] ; but is the mind entirely satisfied with this expla- nation ?" I cannot but echo the last question. Can we be content to assume for, after all, it is assumption that a series of miraculous creations was invariably to be in the manner of a piecing on and blending from one to another, when we have the alternative of presuming (grant it were to be left to presumption alone) that these connexions are only memorials of a natural law presiding over the development of the whole organic creation, and making it one and not many things ? I can only wonder that a man learned in the subject can see such a difficulty as he has here stated, and find it more easily passed over than the bare fact that certain mam- malia have not changed for three thousand years, for such is the only difficulty he states on the other side. It must further be recollected that we are not only to account for the origination of organic being upon this little planet, third of a series which is but one of hundreds of thousands of series, the whole of which again form but one portion of an apparently infinite globe-peopled space, where all seems analogous. We have to suppose, that every one of these numberless globes is either a theatre of organic being, or in the way of becoming so. This is a conclusion which every addition to our knowledge makes only the more irresistible. Is it conceivable, as a fitting mode of exercise for creative intelligence, that it should be constantly paying a special attention to the creation of species, as they may be required in each situation throughout those worlds, at parti- cular times? Is such an idea accordant with our general conception of the dignity, not to speak of the power, of the ORIGIN OF THE ANIMATED TRIBES. 97 Great Author ? Yet such is the notion which we must form, if we adhere to the doctrine of special exercise. Let us see, on the other hand, how the doctrine of a creation in the manner of law agrees with this expanded view of the organic world. Unprepared as most men may be for such an announce- ment, there can be no doubt that we are able, in this limited sphere, to form some satisfactory conclusions as to the plants and animals of those other spheres which move at such im- mense distances from us. Suppose that the first persons of an early nation who made a ship and ventured to sea in it, observed, as they sailed along, a set of objects which they had never before seen namely, a fleet of other ships would they not have been justified in supposing that those ships were occupied, like their own, by human beings, possessing hands to row and steer, eyes to watch the signs of the weather, intelligence to guide them from one place to an- other in short, beings in all respects like themselves, or only showing such differences as they knew to be producible by difference of climate and habits of life ? Precisely in this manner we can speculate on the inhabitants of remote spheres. We see that matter has originally been diffused in one mass, of which the spheres are portions. Consequently, inorganic matter must be presumed to be everywhere the same, although probably with differences in the proportions of ingredients in different globes, and also some difference of conditions. Out of a certain number of the elements of inor- ganic matter are composed organic bodies, both vegetable and animal : such must be the rule in Jupiter and in Sirius, as it is here. We, therefore, are all but certain that herbaceous and ligneous fibre, that flesh and blood, are the constituents of the organic beings of all those spheres which are as yet seats of life. Gravitation we see to be an all-pervading prin- ciple : therefore there must be a relation between the spheres and their respective organic occupants, by virtue of which they are fixed, as far as necessary, on the surface. Such a relation, of course, involves details as to the density and elas- ticity of structure, as well as size of the organic tenants, in proportion to the gravity of the respective planets pecu- 98 ORIGIN OF THE ANIMATED TRIBES. liarities, however, which may quite well consist with the idea of a universality of certain types, such as we see exemplified upon earth. We come to comparatively matter of detail, when we advert to heat and light ; yet it is important to con- sider that these are universal agents, and that, as they bear marked relations to organic life and structure on earth, they may be presumed to do so in other spheres also. The con- siderations as to light are particularly interesting, for, on our globe, the structure of one important organ, almost univer- sally distributed in the animal kingdom, is in direct and pre- cise relation to it. Where there is light there will be eyes, and these, in other spheres, will be the same in all respects as the eyes of tellurian animals, with only such differences as may be necessary to accord with minor peculiarities of con- dition and of situation. It is but a small stretch of the argu- ment to suppose that, one conspicuous organ of a large por- tion of our animal kingdom being thus universal, a parity in all the other organs species for species, class for class, king- dom for kingdom is highly likely, and that thus the inha- bitants of all the other globes of space bear not only a general, but a particular resemblance to those of our own. It must be obvious, that, if organic beings are thus uni- versally distributed, the idea of their having all come into existence through the immediate agency of laws everywhere applicable, is strictly conformable to the principle laid down for our own limited sphere. As a set of laws produced all orbs, their motions and geognostic arrangements, so a set of laws overspread them all with life. The whole productive or creative arrangements thus appear in unity. 99 PARTICULAR CONSIDERATIONS RESPECTING THE ORIGIN OF THE ANIMATED TRIBES. IT being thus shown that there is a general probability in favour of organic creation as a natural event, it becomes necessary to inquire whether this idea be supported or con- tradicted by special facts connected with the constitution of organic bodies Here the patrons of opposite views appear at first sight to have an advantage. " Yes," they say, " it may appear that the bodies of space have been formed and arranged in the manner of natural law ; it may be admitted that all the geognostic phenomena have had a similar history ; but when you come to treat of life and organization, you find yourself in a totally different field. These are mysteries, in the con- sideration of which physical laws fail you, and you are forced to make reference to causes of a different order. Before the origin of life upon our planet could be supposed to have taken place as a simply natural phenomenon, it would be necessary to show that it can be produced at this moment out of inor- ganic elements. But this cannot be shown ; and we must therefore conclude that a special interference of deity was re- quired at the commencement of every species." It is nevertheless true, that much of this is mere assump- H2 100 PARTICULAR CONSIDERATIONS ON THE tion, contrary to the actual state, and much more contrary to the tendency, of science. First, with regard to the constituents of organic bodies, it is found that they are merely a selection of the simple sub- stances which form the inorganic or non-vitalized world. Carbon, oxygen, hydrogen, and nitrogen, are the chief. The first combinations of these in animals are into what are called proximate principles, as albumen, fibrin, etc., out of which the animal body is composed. Now, so far from there being any- thing peculiar or mysterious in these combinations, it is acknowledged that they are simply chemical. "It is now certain," says Dr. Daubeny, of Oxford, " that the same simple laws of composition pervade the whole creation ; and that if the organic chemist only takes the requisite precautions to avoid resolving into their ultimate elements the proximate principles upon which he operates, the result of his analysis will show that they are combined precisely according to the same plan as the elements of mineral bodies are known to be."( 42 ) A particular fact is here worthy of attention : " The conversion of fecula into sugar, as one of the ordinary pro- cesses of vegetable economy, is effected by the production of a secretion termed diastase, which occasions both the rupture of the starch vesicles, and the change of their contained gum into sugar. This diastase may be separately obtained by the chemist, and it acts as effectually in his laboratory as in the vegetable organization. He can also imitate its effects by other chemical agents." ( 43 ) The same writer elsewhere adds, " No reasonable ground has yet been adduced for supposing that, if we had the power of bringing together the elements of any organic compound, in their requisite states and pro- portions, the result would be any other than that which is found in the living body. Every fresh discovery," he says, " is tending to break down the barrier between the two classes of organic and inorganic bodies, as far as regards their chemical combination." ( 44 ) It is much to know the elements of organic bodies, and that the first combinations of these are simply chemical. The powers by which these combinations take place are next to ORIGIN OF THE ANIMATED TRIBES. 101 be inquired into. The predominant idea hitherto has been, that the vital affinities are of a totally distinct nature, de- pending upon a mystic something, to which the term vital principle was applied. But this idea is now on the decline. Admitting the vital affinities, as powers superseding and counteracting ordinary chemical affinities, it is seen that the idea of a distinct inscrutable principle on which they depend, is " both unsupported by evidence and useless in the explanation of facts." ( 45 ) It is becoming evident that living structures result from the action of a multitude of natural forces in combination "gravity, cohesion, elasticity, the agency of the imponderables, and all other powers which operate both on masses and atoms." Professor Draper, of New York, in making this statement, says " It is astonishing that in our days the ancient system which excludes all connexion with natural philosophy and chemistry, and depends on the ficti- tious aid of a visionary force, should continue to exist ; a system which at the outset ought to have broken down by the most common considerations, such as those connected with the mechanical principles involved in the bony skeleton, the optical principles in the construction of the eye, or the hydraulic action of the valves of the heart."( 46 ) So much for the combinations concerned in living bodies ; but how shall we hope to see their forms brought under any relation to physical laws ? On this point we have some illustrations in the phenomena attending the production of crystals, a class of bodies which has been said to stand be- tween the inorganic and the organic. From the agency which has been employed by Mr. Crosse in making crystals formerly supposed to be of Nature's production alone, it is now incon- testable that crystallization is dependent on electric agency, the special forms being the result of the peculiar nature of the constituent substance and the conditions under which the imponderable is applied. Here are obviously natural means of producing forms almost as various as those of living beings, and equally determinate and regular. A certain community of cause in the two instances is indicated by the surprising resemblance which some examples of crystallization bear to 102 PARTICULAR CONSIDERATIONS ON THE vegetable forms. In some, the mimicry is beautiful and com- plete ; for example, in the well-known one called the Arbor Diance. An amalgam of four parts of silver and two of mercury being dissolved in nitric acid, and water equal to thirty weights of the metals being added, a small piece of soft amalgam of silver, suspended in the solution, quickly gathers to itself the particles of the silver of the amalgam, which form upon it a crystallization precisely resembling a shrub. Vege- table figures are also presented in some of the most ordinary appearances of the electric fluid. In the marks caused by positive electricity, or which it leaves in its passage, we see the ramifications of a tree, as well as of its individual leaves ; those of the negative, recal the bulbous or the spreading root, according as they are clumped or divergent. These phenomena seem to indicate that the electric energies have had something to do in determining the forms of plants. That they are intimately connected with vegetable life is indubitable, for germination will not proceed in water charged with negative electricity, while water charged positively greatly favours it ; and a garden sensibly increases in luxuriance when a number of conducting rods are made to terminate in branches over its beds. With regard to the resemblance of the ramifications of the branches and leaves of plants to the traces of the positive electri- city, and that of the roots to the negative, it is a circumstance calling for especial remark, that the atmosphere, particularly its lower strata, is generally charged positively, while the earth is always charged negatively. The correspondence here is curious. A plant thus appears as a thing formed on the basis of a natural electrical operation the brush realized. We can thus suppose the various forms of plants as, imme- diately, the result of a law in electricity, variously affecting them according to their organic character, or respective germinal constituents. In the poplar, the brush is unusually vertical, and little divergent ; the reverse in the beech : in the palm, a pencil has proceeded straight up for a certain distance, radiates there, and turns outwards and downwards ; and so on. We can here see at least traces of secondary means by ORIGIN OF THE ANIMATED TRIBES. 103 which the Almighty Deviser might establish all the vegetable forms with which the earth is overspread. ( 47 ) We turn to the minutiae of organic structure and embry- ology, as affording us some further illustrations of an instruc- tive kind. It is now ascertained by microscopic research, that the basis of all vegetable and animal substances consists of nucleated cells ; that is, cells having granules within them. Xutriment is converted into these before being assimilated by the system. The tissues are formed from them. The ovum destined to become a new creature, is originally only a cell with a contained granule. We see it acting this repro- ductive part in the simplest manner in the cryptogamic plants. " The parent cell, arrived at maturity by the exercise of its organic functions, bursts, and liberates its contained granules. These, at once thrown upon their own resources, and entirely dependent for their nutrition on the surrounding elements, develop themselves into new cells, which repeat the life of their original. Amongst the higher tribes of the crypto- gamia, the reproductive cell does not burst, but the first cells of the new structure are developed within it, and these gradually extend, by a similar process of multiplication, into that primary leaf-like expansion which is the first formed structure in all plants." (* 8 ) Here the little cell becomes directly a plant, the full formed living being. It is also worthy of remark that, in the sponges, (an animal form,) a gemmule detached from the body of the parent, and trusting for sus- tentation only to the fluid into which it has been cast, becomes, without further process, the new creature. Further, it has been recently discovered by means of the microscope, that there is, as far as can be judged, a perfect resemblance between the ovum of the mammal tribes, during that early stage when it is passing through the oviduct, and the young of the infusory animalcules. One of the most remarkable of these, the volvox globator, can hardly be distinguished from the germ which } after passing through a long foetal progress, becomes a com- plete mammifer, an animal of the highest class. It has even been found that both are alike provided with those cilia, which) 104 PARTICULAR CONSIDERATIONS ON THE producing an appearance of revolving motion, is partly the cause of the name given to this animalcule. These resem- blances are the more entitled to notice, that they were made by various observers, distant from each other at the time. ( 49 ) It has likewise been noticed that the globules of the blood are reproduced by the expansion of contained granules ; they are, in short, distinct organisms multiplied by the same fissiparous generation. So that all animated nature may be said to be based on this mode of origin ; the fundamental form of organic being is a cell, having new cells forming within itself, by which it is in time discharged, and which are again followed by others and others, in endless succession. It is of course obvious that, if these cells could be produced by any process from inorganic elements, we should be entitled to say that the fact of a transit from the inorganic into the organic had been witnessed in that instance ; the possibility of the commence- ment of animated creation by the ordinary laws of nature might be considered as established. Now it was announced some years ago by Prevost and Dumas, that globules could be produced in albumen by electricity. If, therefore, these globules be identical with the cells which are now held to be repro- ductive, it might be said that the production of albumen by artificial means is the only step in the process wanting. This has not yet been effected ; but it is known to be only a che- mical process, the mode of which may be any day discovered in the laboratory. ( 50 ) Admitting, however, all these views regarding life and or- ganization, the advocates of interference have still to say that a transition from the inorganic to the organic, such as we must suppose to have taken place in the early geological ages, is no ordinary cognizable fact of the present time upon earth : structure, form, life, are never seen to be imparted to the in- sensate elements ; the production of the humblest plant or animacule, otherwise than as a repetition of some parental form, is not one of the possibilities of science ; if, then, we trace back the generations of organisms to the Silurian or any earlier epoch, and acknowledge the world of that time to have been one in which the present order of natural events ORIGIN OP THE ANIMATED TRIBES. 105 was prevalent, we necessarily can see no natural origin for species, and a miraculous one must be admitted. Here we have undoubtedly the strongest of the argu- ments usually adduced against a natural origin of life. Yet it is one which may easily be replied to. In the first place, there is no reason to suppose that, although life had been imparted by natural means after the first cooling of the surface to a suitable temperature, it would have continued there- after to be capable of being imparted in like manner. The great work of the peopling of this globe with living species is mainly a fact accomplished: the highest known species came as a crowning effect thousands of years ago. The work being thus, to all appearance, finished, we are not necessa- rily to expect that the origination of life and of species should be conspicuously exemplified in the present day. We are rather to expect that the vital phenomena presented to our eyes should mainly, if not entirely, be limited to a regular and unvarying succession of races by the ordinary means of generation. This, however, is no more an argument against a time when phenomena of the first kind prevailed, than it would be a proof against the fact of a mature man having once been a growing youth, that he is now seen growing no longer. We might consider the primitive production of species either as one phenomenon of the nature of the develop- ment of an individual embryo, and that phenomenon as past, just as the individual creation is perfected at birth, or as ex- pressly and wholly a consequence of conditions, which being temporary, the results were temporary also. From the occu- pation of all the great geographical provinces with a more or less full suite of the forms of life, a new development may have hardly any chance of being now drawn forth, and none of being advanced to any extent, even though the same life- creating laws be still in force. Or the operations of these laws might be observant of times, and those of rare occur- rence, so that hundreds of human generations may pass without an opportunity of witnessing such effects. However it may actually have been, assuredly the most rigid disproof of primitive creation as a fact of our time could be no con- 106 PARTICULAR CONSIDERATIONS ON THE elusive argument against a natural creation at a time when the earth was vacant of all organic tenantry, if for such a creation any positive arguments can be adduced. Secondly, it is far from being certain that the primitive im- parting of life and form to inorganic elements is not a fact of our times. Such a doctrine is not generally received in the scientific world ; but the reasons for rejecting it may at least admit of criticism. The leading one is, that, in a great num- ber of instances where the superficial observers of former times assumed a n on -generative origin for life, (as in the ce- lebrated case in Virgil's fourth Georgic,) either the direct contrary has been ascertained, or exhaustive experiments have left no alternative from the conclusion that ordinary genera- tion did take place, albeit in a manner which escapes observa- tion. Finding that an erroneous assumption has been formed in many cases, modern inquirers have not hesitated to assume that there can be no case in which generation is not concerned ; which is certainly far from being allowable. There are several persons eminent in science who profess at least to find great difficulties in accepting the doctrine of invariable gene- ration. Dr. Allen Thomson, one of the professors in the Edinburgh University, has stated several considerations aris- ing from analogical reasoning, which appear to him to throw the balance of evidence in favour of the primitive production of infusoria, the vegetation called mould, and the like. One seems to be of great force ; namely, that the animalcules, which are supposed (altogether hypothetically) to be produced by ova, are afterwards found increasing their numbers, not by that mode at all, but by division of their bodies. If it be the nature of these creatures to propagate in this splitting or fissiparous manner, how could they be communicated to a vegetable infusion ?( 51 ) It has been shown by the opponents of this theory, that when a vegetable infusion is debarred from the contact of the atmosphere, by being closely sealed up or covered with a layer of oil, or only receives oxygen which has passed through sulphuric acid, whereby all animal admixtures have been destroyed, no animalcules are produced ; but can we be sure, in such circumstances, that we have not ORIGIN OF THE ANIMATED TRIBES. 107 set aside some other simple condition requisite for a non-ex- ovo generation ? Who can tell what effect such exclusion of air or such mode of admitting oxygen may have upon the operation of the imponderables in the case ? To this I do not believe that any satisfactory answer could be given. Perhaps the fashionable doctrine is in nothing placed in greater difficulties than it is with regard to the entozoa, or creatures which live within the bodies of others. These ani- mals do, and apparently can, live nowhere else than in the interior of other living bodies, where they generally take up their abode in the viscera, but also sometimes in the cham- bers of the eye, the interior of the brain, the serous sacs, and other places having no communication from without. Some are viviparous, others oviparous. Of the latter it cannot reasonably be supposed that the ova ever pass through the medium of the air, or through the blood-vessels, for they are too heavy for the one transit, and too large for the other. Of the former, it cannot be conceived how they pass into young animals certainly not by communication from the parent, for it has often been found that entozoa do not appear in certain generations of a human family, and some of pecu- liar and noted character have only appeared at rare intervals, and in very extraordinary circumstances. A candid view of the less popular doctrine, as to the origin of this humble form of life, is taken by a distinguished living naturalist. " To explain the beginning of these worms within the human body, on the common doctrine that all created beings proceed from their likes, or a primordial egg, is so difficult, that the mo- derns have been driven to speculate, as our fathers did, on their spontaneous birth; but they have received the hypo- thesis with some modification. Thus it is not from putrefac- tion or fermentation that the entozoa are born, for both of these processes are rather fatal to their existence, but from the aggregation and fit apposition of matter which is already or- ganized, or has been thrown from organized surfaces. * * Their origin in this manner is not more wonderful or more inexplicable than that of many of the inferior animals from sections of themselves. * * Particles of matter fitted by 108 PARTICULAR CONSIDERATIONS ON THE digestion, and their transmission through a living body, for immediate assimilation with it, or flakes of lymph detached from surfaces already organized, seem neither to exceed nor fall below that simplicity of structure which favours this won- derful development ; and the supposition that, like morsels of a planaria, they may also, when retained in contact with living parts, and in other favourable circumstances, continue to live and be gradually changed into creatures of analogous conformation, is surely not so absurd as to be brought into comparison with the Metamorphoses of Ovid. * * We think the hypothesis is also supported in some degree by the fact, that the origin of the entozoa is favoured by all causes which tend to disturb the equality between the secerning and absorbent sy stems." ( 52 ) Here particles of organized matter are suggested as the germinal original of distinct and fully or- ganized animals, many of which have a highly developed re- productive system. How near such particles must be to the inorganic form of matter may be j udged from what has been said within the last few pages. While these appear as good general arguments for primi- tive life-production as a common occurrence in nature, there is a series of facts which goes a fur way to prove that such occurrences must have taken place in comparatively modern times. The pig, in its domestic state, is subject to the attacks of a hydatid, from which the wild animal is free ; hence the disease called measles in pork. The domestica- tion of the pig is of course an event subsequent to the origin of man ; indeed, comparatively speaking, a recent event. Whence, then, the first progenitor of this hydatid ? So also there is a tinea which attacks dressed wool, but never touches it in its unwashed state. A particular insect disdains all food but chocolate, and the larva of the oinopota cellaris lives no- where but in wine and beer, all of these being articles manufac- tured by man. There is likewise a fish called the pymelodes cyclopum, which is only found in subterranean cavities con- nected with certain specimens of the volcanic formation in South America, dating from a time posterior to the arrange- ments of the earth for our species. Whence the first pyme- ORIGIN OF THE ANIMATED TRIBES. 109 lodes cyclopum ? To have produced all these various animals, there must have been means in force at a time long subse- quent to that at which the production of life by miracle is assumed to have taken place. And what is this but to con- nect the ancient events, assumed to have come by miracle, with the modern cases of doubted primitive generation ? Does it not tend to show that the ancient and modern events are of one character, both alike results of a silent immutable energy imparted to nature by her Divine Author, and in the working of which there is no regard to great or small ? Seeing such reasons for believing the general dictum of the philosophical world on primitive generation to be inconclu- sive, we may be prepared to review without surprise or incre- dulity the well-known experiments of Mr. Crosse, which seemed to result in the production of a small species of insect in considerable numbers. This gentleman was pursuing some experiments in crystallization, causing a powerful voltaic battery to operate upon a saturated solution of sili- cate of potash, when the insects unexpectedly made their ap- pearance. He afterwards tried nitrate of copper, which is a deadly poison, and from that fluid also did live insects emerge. Discouraged by the reception of his experiments, Mr. Crosse soon discontinued them; but they were some years after pursued by Mr. Weekes, of Sandwich, with pre- cisely the same results. This gentleman, besides trying the first of the above substances, employed ferro-cyanate of potas- sium, on account of its containing a larger proportion of carbon, the principal element of organic bodies ; and from this substance the insects were produced in increased numbers. A few weeks sufficed for this experiment, with the powerful battery of Mr. Crosse : but the first attempts of Mr. Weekes required about eleven months, a ground of presumption in itself that the electricity was chiefly concerned in the pheno- menon. The changes undergone by the fluid operated upon, were in both cases remarkable, and nearly alike. In Mr. Weekes' s apparatus, the silicate of potash became first turbid, then of a milky appearance ; round the negative wire of the battery, dipped into the fluid, there gathered a quantity of 110 PARTICULAR CONSIDERATIONS ON THE gelatinous matter. From this Mr. Weekes observed one of the insects in the very act of emerging, immediately after which it ascended to the surface of the fluid, and sought concealment in an obscure corner of the apparatus. The insects produced by both experimentalists seem to have been the same, a species of acarus, minute and semi-transparent, and furnished with long bristles, which can only be seen by the aid of the microscope. It is worthy of remark, that some of these insects, soon after their existence had commenced, were found to be likely to extend their species. They were sometimes observed to go back to the fluid to feed, and occa- sionally they devoured each other. ( 53 ) The reception of novelties in science must ever be regu- lated very much by the amount of kindred or relative pheno- mena which the public mind already possesses and acknow- ledges, to which the new can be assimilated. A novelty, however true, if there be no received truths with which it can be shown in harmonious relation, has little chance of a favourable hearing. In fact, as has been often observed, there is a measure of incredulity from our ignorance as well as from our knowledge, and if the most distinguished philosopher three hundred years ago had ventured to develop any striking new fact which only could harmonize with the as yet un- known Copernican solar system, we cannot doubt that it would have been universally scoffed at in the scientific world, such as it then was, or, at the best, interpreted in a thousand wrong ways in conformity with ideas already familiar. The experiments above described, finding a public mind which had never discovered a fact or conceived an idea at all analo- gous, were of course ungraciously received. It was held to be impious even to surmise that animals could have been formed through any instrumentality of an apparatus devised by human skill. The more likely account of the phenomena was said to be, that the insects were only developed from ova, resting either in the fluid, or in the wooden frame on which the experiments took place. On these objections the following remarks may be made. The supposition of impiety arises from an entire misconception of what is ORIGIN OF THE ANIMATED TRIBES. 1 1 1 implied by an aboriginal creation of insects. The experi- mentalist could never be considered as the author of the existence of these creatures, except by the most unreasoning ignorance. The utmost that can be claimed for, or imputed to him is, that he arranged the natural conditions under which the true creative energy that flowing from the primordial appointment of the Divine Author of all things was pleased to work in that instance. On the hypothesis here brought forward, the acarus Crossii was a type of being ordained from the beginning, and destined to be realized under certain physical conditions. When a human hand brought these conditions into the proper arrangement, it did an act akin to hundreds of familiar ones which we execute every day, and which are followed by natural results ; but it did nothing more. The production of the insect, if it did take place as assumed, was as clearly an act of the Almighty himself, as if he had fashioned it with hands. For the pre- sumption that an act of aboriginal creation did take place, there is this to be said, that, in Mr. Weekes's experiment, every care that ingenuity could devise was taken to exclude the possibility of a development of the insects from ova. The wood of the frame was baked in a powerful heat ; a bell- shaped glass covered the apparatus, and from this the atmo- sphere was excluded by the fumes constantly rising from the liquid, for the emission of which there was an aperture so arranged at the top of the glass, that only these fumes could pass. The water was distilled, and the substance of the silicate had been subjected to white heat. Thus every source of fallacy seemed to be shut up. In such circumstances, a candid mind, which sees nothing either impious or unphiloso- phical in the idea of a new creation, will be disposed to ihink that there is less difficulty in believing in such a creation having actually taken place, than in believing that, in two instances, separated in place and time, exactly the same insects should have chanced to arise from concealed ova.( 54 ) 112 HYPOTHESIS OF THE DEVELOPMENT OF THE VEGETABLE AND ANIMAL KINGDOMS. WE have now seen arguments, of both a general and par- ticular kind, for the simply natural origin of life upon our planet. But, whatever force may be allowed to these argu- ments, no attempt has as yet been made to show how, even if life originated in its first and humblest forms in this manner, it passed on, otherwise than by a series of interferences, through that double series of higher forms terminating in the dicotyledons and mammalia, which we have seen rising throughout the geological ages, and leaving the earth occupied by its present organisms. In now proposing to make such an attempt, I deem it ne- cessary, for the sake of simplicity, to confine attention mainly to the animal kingdom, the vegetable department of nature, which starts from a common, or at least, contiguous basis, being sure to fall into any system which may be found appli- cable to the other. It has already been intimated that the succession of animals throughout the geological eras is generally conformable to the gradation of forms in the animal scale, taking these in broad masses, and allowing for such imperfections in the series as geology itself at once leads us to expect, and partly THE VEGETABLE AND ANIMAL KINGDOMS. 113 accounts for. We have seen that, overlooking an era with doubtful vestiges of life, there was first one in which only sea plants and invertebrate marine animals flourished ; after- wards one presenting the meaner (cartilaginous) fishes ; and that higher (osseous) fishes, reptiles, birds, and mammals, came at long intervals throughout the subsequent ages. Even when we pass into details, we find the succession to be in so many instances conformable to the gradation of the special groups of animals, that no doubt can remain that such is the case in all. Thus, for example, there is, between the Silurians and the oolite, a clear advance from humble to more highly organized cephalopoda and echinodermata. In the same time, the trilobite is exchanged for the superior but con- nected form, limulus, and the brachiopoda sink beneath the new and superior class of bivalves, the lamellibranchiata. We see, in an order of fishes of the carbonigenous era, an approximation to the reptile class. By-and-by, come ichthy- osauri, half-fish half-crocodile; afterwards, a succession of forms ending in true crocodiles. Some difficulties have, indeed, been brought forward ; but they are, on a just view of the sciences concerned, of no real importance, and I only deem them worthy of notice in a subordinate place. ( 55 ) Leaving for a future section the particulars of the animal scale, which will there lend us further illustration, it may now be observed that, while the external features of the various creatures are so different, there has been traced, throughout large groups of them, a fundamental unity of organization, as implying, with respect to these groups, that all were constructed upon one plan, though in a series of im- provements and variations giving rise to the special forms, and bearing reference to the conditions in which each animal lives. Starting from the primitive germ, which, as we have seen, is the representative of a particular order of full-grown animals, we find all others to be merely advances from that type, with the extension of endowments and modification of forms which are required in each particular case ; each form, also, retaining a strong affinity to that which precedes it, 114 HYPOTHESIS OF THE DEVELOPMENT OF and tending to impress its own features on that which suc- ceeds. This principle is partly matter of familiar observation. It is obvious to all, that an ordinary mammalian quadruped has a strong analogy of form to the human being ; its head, its forelegs, its hinder extremities, have each their representative parts in our frame. But the ordinary observer is surprised to learn how much further the principle is carried. For example, the hind leg of the horse looks very different from one of our limbs, in as far as it seems to have a knee presented backwards, and possesses no toes. In reality, the part of the horse corresponding to our knee is high up near the body of the animal, and the hock corresponds to our heel. It has toes, moreover ; but they are sunk in the hollow of a hoof, which serves as a shield to that part. The horse, the dog, and many other quadrupeds, may be said to walk upon their toes (hence called digitigrade) : others, as the bear and badger, present the whole foot to the ground, as man does (hence called plantigrade). Thus, too, the wing of the bird contains bones representing those of our arm, though modified for so different a purpose. The paddles of the whale tribes and seals are other curious modifications of a member substantially the same. The bat, again, has the bones of its hand developed to an unusual extent, so as to become a frame for the membrane by which it flies : in the extinct pterodactyle, the same pur- pose was chiefly served by a development of the forefinger alone. The fundamental resemblance which lurks below various appearances is often startling. Thus, the giraffe, with its long neck, has, in that part, no more bones than are to be found in the neck of the elephant or pig, which hardly seem to have any neck at all. The cervical vertebrae are but seven in every one of the mammalian animals. Sometimes, an organ appears entirely wanting in one family, as feet in the serpent tribes, a pelvic region in the whale, the wing in the bird called the apteryx ; and yet it is not truly wanting. Usually, some rudiment of it appears, as if nature had been willing to give it, but had kept it back from a com- THE VEGETABLE AND ANIMAL KINGDOMS. 115 plete development, as knowing it to be not needed in that instance. On this ground, the notion of a much ridiculed philosopher of the last century, respecting a human tail, may be said to be not quite without foundation. Between the fifth and sixth week, a tail exists in the human embryo ; it then goes back; but still in the mature subject its elements are seen clumped up in the bone at the bottom of the spine, the os coccygis. Unity of organization becomes the more remarkable when we observe that the corresponding organs of animals, while preserving a resemblance, are sometimes put to different uses. For example, the ribs become, in the serpent, organs of loco- motion, and the snout is extended, in the elephant, into an instrument serving all the usual purposes of an arm and hand. It is- equally remarkable, that there should be, in the ori- ginal plan of the animal structure, a double set of organs, one or other of which is selected for development according to the needs of the particular animal. Thus, there are in the plan both gills and lungs, two wholly distinct kinds of respiratory apparatus, the one being designed for a watery and the other for an atmospheric medium. The mammalia, as creatures destined to breathe the air, are furnished with lungs ; but, at an early stage of the foetal progress, this is not the case. They have at that time a branchial apparatus. Afterwards, this goes back, and the lungs are developed from a different portion of the organism. Lungs, on the other hand, are possessed by certain fishes in a rudimental form ; it is the well-known air-bladder of those fishes, which are understood to profit by it, as an additional means of floating. So, also, the baleen of the whale and the teeth of the land mammifer are different organs. The whale, in embryo,' shows the rudi- ments of teeth ; but, not being wanted, they are not deve- loped, and baleen is brought forward instead. But the most remarkable circumstance attending the law of unity of organization is, that an organ will sometimes be seen developed to a certain extent, but wholly without use. i2 116 HYPOTHESIS OF THE DEVELOPMENT OP This organ will, perhaps, be seen serving a purpose in a par- ticular family of animals ; but we advance into an adjoining or kindred family, and there find a rudiment of the same organ, which, owing to the different conditions of this new set of creatures, is of no kind of service. Thus, some of the serpent tribes possess rudimentary limbs. In other instances, a portion of organization necessary in one sex is also pre- sented in the other, where it is not necessary. For example, the mammae of the human female, by whom these organs are obviously required, also exist in the male, who has no occasion for them. It might be supposed that in this case there was a regard to uniformity for mere appearance' sake ; but that no such principle is concerned, appears from a much more re- markable instance connected with the marsupial animals. The female of that tribe has a process of bone advancing from the pubes for the support of her pouch ; and this also appears in the male marsupial, who has no pouch, and re- quires none. The same law of unity presides over the vegetable king- dom. Amongst phanerogamous plants, a certain number of organs are always present, either in a developed or rudi- mentary state ; and those which are rudimentary can be developed by cultivation. The flowers which bear stamens on one stalk and pistils on another, can be caused to produce both, or to become perfect flowers, by having a sufficiency of nourishment supplied to them. So, also, where a special function is required for particular circumstances, nature provides for it, not by a new organ, but by a modification of a common one. Thus, for instance, some plants destined to live in arid situations, require to have a store of water which they may slowly absorb. The need is arranged for by a cup- like expansion round the stalk, in which water remains after a shower. Now the pitcher, as this is called, is not a new organ, but simply the metamorphosis of a leaf. It is thus proved, with regard to the constituent beings of large sections of the animal kingdom, that they are bound up in a fundamental unity, however various in degree of endow- THE VEGETABLE AND ANIMAL KINGDOMS. 117 ment and in the purposes which they serve in the world. They may be said to stand in a connexion analogous to that in which the planets are placed by the third law of Kepler. And the inference with regard to their origin is the same. Precisely as it is impossible to suppose a distinct exertion or fiat of Almighty Power for the formation of the earth, wrought up as it is in a complex dynamical connexion, first with Yenus on the one hand and Mars on the other, and secondly with all the other members of the system, so is it impossible to conceive the same power using particular means for the production of a particular animal species, an indivi- dualized fraction, as it now appears, in a vast system which would not be complete without it, and into whose adjacent parts it melts by the finest shadings. Supposing, for a moment, that each species had been distinct in its origin, these shadings would have been unnecessary; and there would at least have been a strong probability against a unity of organization being adopted as part of the plan. In that case, abortive or rudimentary organs must have been con- sidered as a kind of blemish the thing of all others most irreconcilable with that idea of perfection which a general view of nature irresistibly attributes to its author. If, on the other hand, we admit that the animal kingdom took its rise in a general law, we see in the shadings and the organic unity something not only harmonious with, but essential to the system. Rudimentary organs, too, appear but as harm- less peculiarities of development, and interesting evidences of the manner in which the Divine Author has been pleased to work. It must be easy to see how this class of facts bears on the great question. Organisms we know to have been produced, not at once, but in the course of a vast series of ages ; here we now see that they are not a group of individually entire things accidentally associated, but parts of great masses, nicely connected, and integral in their respective totalities. Time, and a succession of forms in gradation and affinity, become elements in the idea of organic creation. It must be seen that the whole phenomena thus pass into strong analogy 118 HYPOTHESIS OF THE DEVELOPMENT OF with those attending the production of individual organisms. But it becomes something more than analogy when we have learned the facts attending the embryonic development of animals. First surmised by the illustrious Harvey, after- wards illustrated by Hunter in his wondrous collection at the Royal College of Surgeons, finally advanced to mature con- clusions by Tiedemann, St. Hilaire, and Serres, embryonic development is now a science. Its primary positions are 1. that the embryos of all animals are not distinguishably different from each other ; and, 2. that those of all animals pass through a series of phases of development, each of which is the type or analogue of the permanent configuration of tribes inferior to it in the scale. With regard to the latter proposition, it is to be remarked that, while it is generally true of the whole forms of animal being, it is more particu- larly true of departments of the organization, as the nutritive system, the vascular system, the nervous system, &c., each of which is destined for a peculiar degree of development in dif- ferent groups of animals, according to their needs. Speaking, however, roundly, it is undoubted, respecting nearly all animals, that they pass in embryo through phases resembling the general as well as the particular characters of others of lower grade. For example, the comatula, a free-swimming star-fish, is, at one stage of its early progress, a crinoid that is, a star-fish fixed upon a stalk at the bottom of the sea. It advances from the form of one of the lower to that of one of the higher echinodermata. The animals of its first form were, as we have seen, among the most abundant in the earliest fossiliferous rocks : they began to decline in the New Red Sandstone era, and they were succeeded in the Oolitic age by animals of the form of the mature comatula. Thus, too, the insect, standing near the head of the articulated animals, is, in the larva state, an annelid or worm, the anne- lides being the lowest in the same class. The higher crus- tacea, as the crab and lobster, at their escape from the ovum, resemble the perfect animal of the inferior order entomo- straca, and pass through the forms of transition which charac- terize the intermediate tribes of Crustacea. The salmon, a THE VEGETABLE AND ANIMAL KINGDOMS. 119 highly organized fish, exhibits, in its early stages, as has been remarked, the gelatinous dorsal cord, the heterocercal tail, and inferior position of the mouth, which mark the mature example of the cartilaginous fishes. The frog, again, for some time after its birth, is a fish with external gills, and other organs fitting it for an aquatic life, all of which are changed as it advances to maturity and becomes a land animal. The mammifer only passes through still more stages, according to its higher place in the scale. Nor is man him- self exempt from this law. His first form is that which is permanent in the animalcule. His organization gradually passes through conditions generally resembling a worm, a fish, a reptile, a bird, and the lower mammalia, before it attains its specific maturity. At one of the last stages of his foetal career, he exhibits an intermaxillary bone, which is charac- teristic of the perfect ape ; this is suppressed, and he may then be said to take leave of the simial type, and become a true human creature. Even as we shall find, the varieties of his race are represented in the progressive development of an in- dividual of the highest, before we see the adult Caucasian, the highest point yet attained in the animal scale. To come to particular points of the organization. The brain of man, which exceeds that of all other animals in com- plexity of organization and fulness of development, is, at one early period, only " a simple fold of nervous matter, with difficulty distinguishable into three parts, while a little tail- like prolongation towards the higher parts, and which had been the first to appear, is the only representation of a spinal marrow. Now, in this state it perfectly resembles the brain of an adult fish, thus assuming in transitu the form that in the fish is permanent. In a short time, however, the structure is become more complex, the parts more distinct, and the spinal marrow better marked ; it is now the brain of a reptile. The change continues ; by a singular motion, certain parts (cor- pora quadrigemind) which had hitherto appeared on the upper surface, now pass towards the lower ; the former is their per- manent situation in fishes and reptiles, the latter in birds and 120 HYPOTHESIS OF THE DEVELOPMENT OP mammalia. This is another advance in the scale, but more remains yet to be done. The complication of the organ in- creases; cavities termed ventricles are formed, which do not exist in fishes, reptiles, or birds ; curiously organized parts, such as the corpora striata, are added ; it is now the brain of the mammalia. Its last and final change alone seems want- ing, that which shall render it the brain of MAN. ( 5G ) And this change in time takes place. So also with the heart. This organ, in the mammalia, con- sists of four cavities, but in the reptiles of only three, and in fishes of two only, while in the articulated animals it is merely a prolonged tube. Now in the mammal foetus, at a certain early stage, the organ has the form of a prolonged tube ; and a human being may be said to have then the heart of an insect. Subsequently, it is shortened and widened, and becomes divided by a contraction into two parts, a ventricle and an auricle ; it is now the heart of a fish. A subdivision of the auricle afterwards makes a triple-chambered form, as in the heart of the reptile tribes ; lastly, the ventricle being also subdivided, it becomes a full mammal heart. We have now to remember that, corresponding generally to these progressive forms in the development of individuals, has been the succession of animal forms in the course of time. Our earth bore crinoidea before it bore the higher echinoder- mata. It presented crinoidea, annelides, and mollusca,, before it bore fishes, and when fishes came, the first forms were those cartilaginous types which correspond with the early fo3tal condition of higher orders. Afterwards there were reptiles, then mammifers, and finally, as we know, came man. Was it, then, too much to say that, when we learned the facts of embryonic development, we should see something more than analogy between the progress of species upon the earth and the production of an individual organism ? The tendency of all the illustrations is undoubtedly to make us look to development as the principle which has been immediately and mainly concerned in the peopling of this globe, a process extending over a vast space of time, but which THE VEGETABLE AND ANIMAL KINGDOMS. 121 is nevertheless connected in character with the briefer process by which an individual being is evoked from a simple germ. What mystery is there here and how shall I proceed to enun- ciate the conception which I have ventured to form of what may prove to be its proper solution ! It is an idea by no means calculated to impress by its greatness, or to puzzle by its profoundness. It is as much marked by simplicity as perhaps any other of those which have explained the great secrets of nature. But here, again, it may be said, lies one of its strongest claims to our faith. My proposition is that the several series of animated beings, from the simplest and oldest up to the highest and most recent, are the results, first, of an inherent impulse in the forms of life to advance, in definite times, by generation, through grades of organization terminating in the highest dicotyledons and vertebrata, these grades being few in num- ber, and generally marked by intervals of organic character which we find to be a practical difficulty in ascertaining affinities ; second, of another inherent impulse connected with the vital forces, tending, in the course of generations, to modify organic structures in accordance with external circumstances, as food, the nature of the habitat, and the meteoric agencies, these being the " adaptations" of the natural theologian. We may contemplate these phenomena as ordained to take place in every situation, and at every time, where and when the requisite materials and conditions are presented in other orbs as well as in this in any geographical area of this globe which may at any time arise observing only the variations due to difference of materials and of conditions. The nucleated vesicle is the fundamental form of all or- ganization, the meeting-point between the inorganic and the organic the end of the mineral and beginning of the vegetable and animal kingdoms, which thence start in dif- ferent directions, but in a general parallelism and analogy. This nucleated vesicle is itself a type of mature and inde- pendent being, as well as the starting point of the foetal progress of every higher individual in creation, both animal 122 HYPOTHESIS OF THE DEVELOPMENT OF and vegetable. We have seen that it is a form of being which there is some reason to believe electric agency will produce though not perhaps usher into full life in albumen, one of those component materials of animal bodies, in whose com- binations it is believed there is no chemical peculiarity for- bidding their being any day realized in the laboratory. Remembering these things, it seems, after all, an obvious idea that a chemico- electric operation, "by which germinal vesicles . were produced, was the first phenomenon in organic creation, and that the second was an advance of these through a suc- cession of higher grades, and a variety of modifications, in accordance with laws of the same absolute nature as those by which the Almighty rules the physical department of nature. Leaving the first of these supposed processes to rest upon the arguments which have been adduced with regard to a possible transition from the inorganic to the organic, as a natural fact, we have two things to be accounted for first, grade ; and, second, external peculiarities. We have to con- vince ourselves, both that a fish may advance to be a reptile, and a reptile to*be a bird being a distinct step onward in complexity of organization and that particular organs are capable of being modified, so as to suit external conditions, for example, the bill of a bird to the picking up of food in shallow waters, or the throat of the foetal marsupial to the reception of the mother's milk without a danger of choking. With regard to grade, it may be admitted at once that, in Nature's government, there is no observable appearance of such promotions. But it may be asked, if, supposing such events to be within the scope of nature, we are necessarily to expect to see them take place, or even to hear of them having been recorded. To settle this question, let us first inquire into the proportion of the number of these grades to the space of time believed to be represented in the fossiliferous series of rocks. Mr. Lyell tells us that the space between our sun and some of the remote star- clusters, of which the distance to Sirius (not less than nineteen millions of millions of miles) is but a fraction, may no more than compare with the space of THE VEGETABLE AND ANIMAL KINGDOMS. 123 time which has probably elapsed since the origin of the coral- line limestone over which the Niagara is precipitated at the Falls. Now, the number of grades of what may be called the first degree (transitions from class to class) passed through by the vertebrata since their origin in the Devonian rocks is, at the utmost, three. Such a leap in organic progress has, therefore, only taken place once in many millions of millions of years. If such be the case, all chance of such grade transi- tions being witnessed within the four thousand years of his- torical humanity vanishes. As to the possible occurrence of such unusual events in the midst of a series which appear fixed and regular, let us call forward an illustration from the Ninth Bridgewaier Treatise of Mr. Babbage. The reader is requested to suppose himself seated before the calculating machine, and observing it. It is moved by a weight, and there is a wheel which revolves through a small angle round its axis, at short intervals, presenting to the eye successively, a series of numbers engraved on its divided circumference. Let the figures thus seen be the series, 1, 2, 3, 4, 5, &c., of natural numbers, each of which exceeds its immediate ante- cedent by unity. " Now, reader," says Mr. Babbage, " Let me ask you how long you will have counted before you are firmly convinced that the engine has been so adjusted, that it will continue, while its motion is maintained, to produce the same series of natural numbers ? Some minds are so constituted, that after passing the first hundred terms, they will be satisfied that they are acquainted with the law. After seeing five hundred terms few will doubt, and after the fifty thousandth term the propensity to believe that the succeeding term will be fifty thousand and one, will be almost irresistible. That term will be fifty thousand and one ; and the same regular succession will continue ; the five millionth and the fifty millionth term will still appear in their expected order, and one unbroken chain of natural numbers will pass before your eyes, from one up to one hundred million. " True to the vast induction which has been made, the next 124 HYPOTHESIS OF THE DEVELOPMENT OF succeeding term will be one hundred million and one ; but the next number presented by the rim of the wheel, instead of being one hundred million and two, is one hundred million ten thousand and two. The whole series from the commence- ment being thus, 1 2 3 4 5 99,999,999 100,000,000 regularly as far as 100,000,001 100,010,002 the law changes. 100,030,003 100,060,004 100,100,005 100,150,006 100,210,007 100,280,008 " The law which seemed at first to govern this series failed at the hundred million and second term. This term is larger than we expected by 10,000. The next term is larger than was anticipated by 30,000, and the excess of each term above what we had expected forms the following table : 10,000 30,000 60,000 100,000 150,000 THE VEGETABLE AND ANIMAL KINGDOMS. 125 being, in fact, the series of triangular number s,( 57 ) each mul- tiplied by 10,000. " If we now continue to observe the numbers presented by the wheel, we shall find, that for a hundred, or even for a thousand terms, they continue to follow the new law relating to the triangular numbers ; but after watching them for 2761 terms, we find that this law fails in the case of the 2762nd term. " If we continue to observe, we shall discover another law then coming into action, which also is dependent, but in a different manner, on triangular numbers. This will continue through about 1430 terms, when a new law is again intro- duced which extends over about 950 terms, and this, too, like all its predecessors, fails, and gives place to other laws, which appear at different intervals. " Kow it must be observed that the law that each number presented by the engine is greater by unity than the preceding number, which law the observer had deduced from an induc- tion of a hundred million instances, was not the true law that regulated its action, and that the occurrence of the number 100,010,002 at the 100,000,002nd term was as necessary a consequence of the original adjustment, and might have been as fully foreknown at the commencement, as was the regular suc- cession of any one of the intermediate numbers to its immediate antecedent. The same remark applies to the next apparent deviation from the new law, which was founded on an induc- tion of 2761 terms, and also to the succeeding law, with this limitation only that whilst their consecutive introduction at various definite intervals, is a necessary consequence of the mechanical structure of the engine, our knowledge of analysis does not enable us to predict the periods themselves at which the more distant laws will be introduced." It is not difficult to apply the philosophy of this passage to the question under consideration. Let us remember that the gestation of a single organism is the work of but a few days, weeks, or months ; but the gestation (so to speak) of a whole creation is a matter involving the enormous spaces of 126 HYPOTHESIS OF THE DEVELOPMENT OF time which have been described. Suppose that an ephe- meron, hovering over a pool for its one April day of life, were capable of observing the fry of the frog in the water below. In its aged afternoon, having seen no change upon them for such a long time, it would be little qualified to con- ceive that the external branchiae of these creatures were to decay, and be replaced by internal lungs, that feet were to be developed, the tail erased, and the animal then to become a denizen of the land. Precisely such may be our difficulty in conceiving that plants and animals are capable of advancing by generation to a higher type of being. Granting that, during the whole time which we call the historical era, there have been no movements of this kind, nor even any of the less rare transitions in which only specific modifications are concerned, we know the historical era to be only an infinitesi- mal portion of the entire age of our globe. We do not know what may have happened during the ages which preceded its commencement, as we do not know what may happen in ages yet in the distant future. All, therefore, that we can properly infer from the apparent fixity of organic forms is, that such is the ordinary procedure of nature in the time im- mediately passing before our eyes. Mr. Babbage's illustra- tion enables us to understand how this ordinary procedure may be subordinate to a higher law which in proper season interrupts and changes it. It has been seen that, in the reproduction of the higher animals, the new being passes through stages in which it is successively fish-like and reptile-like. But the resemblance is not to the adult fish or the adult reptile, but to the fish and reptile at a certain point in their fetal progress ; this holds true with regard to the vascular, nervous, and other systems alike. It seems as if gestation consisted of two distinct and independent stages one devoted to the developme/it of the new being through the conditions of the inferior types, or, rather through the corresponding first stages of their develop- ment; another perfecting and bringing the new being to a healthy maturity, on the basis of the point of development THE VEGETABLE AND ANIMAL KINGDOMS. 127 reached. This may be illustrated by a simple diagram. ( 3S ) The foetus of all the four classes may be supposed to advance in an identical condition to the point A. The fish there diverges and passes along a line apart, and peculiar to itself, to its mature state at F. The reptile, bird, and D mammal, go on together to C, where the reptile diverges in like manner, and ad- vances by itself to R. The bird diverges atD, and goes on to B. Here it is appa- rent that the only thing required for an advance from one grade to another in the generative process is that, for example, the fish embryo should not diverge at A, but go on to C before it diverges, in which case the progeny will be, not a fish, but a reptile. To protract the straightfor- ward part of the gestation over a small space is all that is ne- cessary. Xow we may never see an example of the working of the actual law which is supposed to be capable of producing such an advance of grade ; but something approaching to it in effect has been observed. Sex is fully ascertained to be a matter of development. All beings are, at one stage of the embryotic progress, female ; a certain number of them are afterwards advanced to be of the more powerful sex. From this it will be understood that no absolute distinction exists ; all such are merely apparent. The ingenious Huber first made us aware of an instance, in a humble department of the animal world, of arrangements being made by the animals themselves for adjusting the law of development to the production of a par- ticular sex. Amongst bees, as amongst several other insect tribes, there is in each community but one true female, the queen bee, the workers being false females or neuters ; that is to say, sex is carried on in them to a point intermediate between the female and male, where it is attended by sterility. The preparatory states of the queen bee occupy sixteen days ; those of the neuters, twenty ; and those of males, twenty-four. Now it is a fact, settled by innumerable observations and ex- 128 HYPOTHESIS OF THE DEVELOPMENT OF periments, that the bees can so modify a larva, which other- wise would result in a worker, that, when the perfect insect emerges from the pupa, it is found to be a queen or true fe- male. For this purpose they enlarge its cell, make a pyra- midal hollow to allow of its assuming a vertical instead of a horizontal position, keep it warmer than other larvae are kept, and feed it with a peculiar kind of food. From these simple circumstances, leading to a shortening of the embryotic con- dition, results a creature different in form, and also in dispo- sitions, from what would have otherwise been produced. Some of the organs possessed by the worker are here wanting, We have a creature " destined to enjoy love, to burn with jealousy and anger, to be incited to vengeance, and to pass her time without labour," instead of one " zealous for the good of the community, a defender of the public rights, enjoying an immunity from the stimulus of sexual appetite and the pains of parturition ; laborious, industrious, patient, ingenious, skilful ; incessantly engaged in the nurture of the young, in collecting honey and pollen, in elaborating wax, in con- structing cells and the like ! paying the most respectful and assiduous attention to objects which, had its ovaries been de- veloped, it would have hated and pursued with the most vin- dictive fury till it had destroyed them!"( 59 ) All these changes may be produced by a mere modification of the em- bryotic progress, which is within the power of the adult animals to effect. By the arrangements made and the food given, the embryo becomes sooner fit for being ushered forth in its imago or perfect state. Development may be said to be v *thus arrested at a particular stage that early one at which the female sex is complete. In the other circumstances, it is allowed to go on four days longer, and a stage is then reached between the two sexes, which in this species is destined to be the perfect condition of a large portion of the community. Four days more make it a perfect male. It may be observed that there is, from the period of oviposition, a destined dis- tinction between the sexes of the young bees. The queen lays the whole of the eggs which are designed to become workers, before she begins to lay those which become males. THE VEGETABLE AND ANIMAL KINGDOMS. 129 But the condition of her reproductive system evidently governs the matter of sex, for it is remarked that when her impregnation is delayed beyond the twenty-eighth day of her entire existence, she lays only eggs which become males. We have here, it will be admitted, a most remarkable illustration of the principle of development, although in an operation limited to the production of sex only. Let it not be said that the phenomena concerned in the generation of bees may be very different from those concerned in the repro- duction of the higher animals. There is a unity throughout nature which makes the one case an instructive reflection of the other. () We shall now see an instance of development operating within the production of what approaches to the character of variety of species. It is fully established that a human family, tribe, or nation, is liable, in the course of generations, to be either advanced from a mean form to a higher one, or degraded from a higher to a lower, by the influence of the physical conditions in which it lives. The coarse features, and other structural peculiarities of the negro race only continue while these people live amidst the circumstances usually associated with barbarism. In a more temperate clime, and higher social state, the face and figure become greatly refined. The few African nations which possess any civilization exhibit forms approaching the European ; and when the same people in the United States of America have enjoyed a within-door life for several generations, they assimilate to the whites amongst whom they live. On the other hand, there are authentic instances of a people originally well-formed and good-looking, being brought, by imperfect diet and a variety of physical hardships, to a meaner form. It is remarkable that prominence of the jaws, a recession and diminution of the cranium, and an elongation and attenuation of the limbs, are peculiarities always produced by these miserable condi- tions, for they indicate an unequivocal retrogression towards the type of the lower animals. Thus we see nature alike willing to go back and to go forward. Both effects are simply 130 HYPOTHESIS OF THE DEVELOPMENT OF the result of the operation of the law of development in the generative system. Let us trace this law also in the production of certain classes of monstrosities. A human foetus is often left with one of the most important parts of its frame imperfectly developed ; the heart, for instance, goes no further than the three- chambered form, so that it is the heart of a reptile. There are even instances of this organ being left in the two- chambered or fish- form. Here we have apparently a realiza- tion of the converse of advance of grade, so far, at least, as one organ is concerned. Seeing a complete specific retrogression in one point, how easy it is to suppose a simply natural pro- cess, reversing the phenomenon, and making a fish mother develop a reptile heart, or a reptile mother develop a mammal one. It is no great boldness to surmise that a super-adequacy of force in the measure of this under-adequacy (and the one thing seems as natural an occurrence as the other) would suffice in a natatorial bird to give it as a progeny the orni- thorhynchus, or might give the progeny of an ornithorhynchus the mouth and feet of a true mammalian, and thus complete at two stages a passage from one class to another. Perhaps, with the bulk of men, even those devoted to science, the great difficulty is, after all, in conceiving the par- ticulars of such a process as would be required to advance a fish into a reptile. And yet no difficulty could well be less substantial, seeing that the metamorphosis of the tadpole into the frog a phenomenon presented to our observation in count- less instances every spring is, in part at least, as thoroughly a transmutation of the fish organization into that of the reptile, as the supposable change of sauroid fishes into saurian reptiles could ever be. It is different, as being only a process in ordi- nary generation ; but it realizes, as far as the necessary organic changes are concerned, the hypothetic view of an advance of one grade of animal forms into another. There is another fact connected with the reproduction of the batrachian order of reptiles, that, when the young are enclosed in a dark box sunk in a river, with holes through which the water may flow, the animals grow, but never undergo their destined change : THE VEGETABLE AND ANIMAL KINGDOMS. 131 they became gigantic tadpoles, and the reptile characters are not developed. Here the progeny of a reptile literally be- comes a fish, and transition of species is thoroughly realized, although in retrogression. And this is an instance in which the whole animal is concerned. Now surely no one will deny that that which we see nature undo she is able to do, and might be seen doing, if the proper occasion were to occur, or were the requisite attendant conditions realized. So much with regard to grade. Let us now consider the principle of modifidbility that part of the hypothesis to which we are to look for an account of the external variations and adaptation of animals. Here we are directly opposed by the prevalent doctrine among naturalists, that species is intransible, and has so continued during all the time that scientific observation has existed. There is a certain volubility, they admit, in organisms, throughout successive generations, and for this variability external conditions may account ; but such varia- tions show a disposition to give way, when the original con- ditions are resumed, or when the changed individual is mingled in alliance with the original stock. There is there- fore a fixed and immutable character which we call species, and which can only be traced to an origin differing as an event from the procedure of nature in our own time. I trust to be able to show that this doctrine is in such a condition with regard to facts, and has of late been obliged by facts to make so many shifts in its assumptions, that it is not entitled to the respect usually assigned to it. The doctrine theoretically attaches the term species to every organism which manifests the same peculiarities throughout a series of generations. Practically, without waiting to watch successive generations, or where, from the extinction of the organism, this may be impossible, natura- lists give the appellation to every organism which presents a plurality of individuals similarly marked. Very small pecu- liarities suffice. A particular spot on the wing of a butterfly constitutes it a distinct species. The Golden Plover of Aus- tralia was so reckoned, and got the name of Xanthocheilus, K2 132 HYPOTHESIS OF THE DEVELOPMENT OF from only having a small portion of yellow in the commissure of its bill. At the same time, in palaeontology, such a peculiarity as an extra-plication in the enamel of a fossil pachiderm's tooth, is sufficient to obtain a specific name for that animal, and constitute its origin a separate miracle. With equal facility, naturalists of this predominant order make up groups of species into genera, and groups of genera into families and tribes. Suppose the doctrine were to be taken according to the practice, we should possess a fact speaking strongly against fixity of species. It has been pointed out by an eminent botanist that, amongst recent fossil plants, are poplars, pines, birches, and hornbeams, like those now existing, but not the same. Thus one species has replaced another in even com- paratively recent times. It may be asked, if the same change of species has not been going on since. The vague descrip- tions of ancient botanists forbid our speaking confidently of the intermediate ages. But look to the present time. In districts examined narrowly at no distant day, new species are continually being found by new investigators. It will be said, that these additions are owing to the acuteness of modern observers. But this is begging the whole question. " We do not know," says our author, " that we are entitled to assert that botanists were so mole-eyed thirty years ago, that their quick-sighted successors have been able to add twenty-five per cent, to the number of ascertained species growing at their own doors." ( 6I ) Grant, then, that the pecu- liar plants in question really are species, the probability un- doubtedly is, that they are new species, true examples of that very phenomenon which the superstition of science would hold to be a supernatural event. Still take the doctrine according to the practice, and let us see how it stands with regard to certain facts recently ascer- tained. Amidst all the dogmatism which has been indulged in on this subject, the assumed distinction of species has given way in numberless instances, both in the vegetable and animal kingdoms. In botany, the wider distinction of genus, and even that of whole tribes, has proved in some cases falla- THE VEGETABLE AND ANIMAL KINGDOMS. 133 cious. According to Dr. Lindley, " So entirely in the simplest forms of Thallogens [an assemblage embracing sea- weeds, fungi, and lichens] is all trace of series missing, that in some of them their reproductive matter has been regarded by certain writers as altogether of an ambiguous nature. In their opinion, it is even uncertain whether this matter will reproduce its like, and whether it is not a mere repre- sentation of the vital principle of vegetation, capable of being called into action either as a Fungus, an Alga, or a Lichen, according to the particular conditions of heat, light, mois- ture, and medium, in which it is placed ; producing Fungi upon dead or putrid organic beings; Lichens upon living vegetables, earth, or stones ; and Algae where water is the medium in which they are developed. Kiitzing endeavours to maintain the following propositions connected with this sub- ject: 1st, the formation of organic matter can only take place by means of the previously dissolved elements of other organic principles ; 2nd, simple globules, such as Crypto- coccus, Palmella, and Protococcus, can give birth to different formations, according to the influence of light, air, and tem- perature ; 3rd, we must regard all the forms of lower algae as vegetations of a very simple structure, and distinguish them from each other, notwithstanding that in certain circum- stances they maj^ raise themselves to vegetations of a higher form ; for, in other circumstances, they can exist and mul- tiply independently ; 4th, the same formation may be pro- duced by primitive formations of altogether different kinds." " It has been said," adds Dr. Lindley, " that Algae are aquatics, while Lichens and Fungi are terrestrial ; but Fungi will develop in water, when they assume the form ofAlgce"(^) Undoubtedly, eight so-called genera of fungi are now set down as only variations of one plant (Telephora sulphurea), arising from peculiar conditions of culture. Even in higher departments of the vegetable kingdom, the revolutions have been very remarkable. Six so-called species of pine are wound up into one in a recent memoir on the Coniferae. The cowslip, primrose, oxlip, and polyanthus, which were always regarded as distinct species, are now 134 HYPOTHESIS OF THE DEVELOPMENT OF found to be producible from one set of seeds, under various conditions ; they are radically one plant. So also " the clove, pink, and carnation are only varieties of a flower growing among the ruins of some of our old castles, the Dianthus caryophyllus." The artichoke of the garden and the cardoon (a kind of thistle) of the South American wild, are held as distinct species in all botanical works ; yet the artichoke, in neglect, degenerates into the cardoon. ( 63 ) The ranunculus aquatilis and the ranunculus hederaceus are, in like manner, set down as distinct species; but behold the secret of their difference ! While the former plant remains in the water, its leaves are all finely cut and have their divi- sions hairy ; but when the stems reach the surface, the leaves developed in the atmosphere are widened, rounded, and simply lobed. Should the seeds of this water plant fall upon a soil merely moist without being inundated, the result is the ranun- culus hederaceus the presumed distinct species with short stalks, and none of the leaves divided into hairy cut work ! ( 4 ) To come to a more familiar instance. It is now fully ascer- tained that the various bread-forming grains, wheat, barley, oats, rye, are resolvable into one. If wheat be sown in June, and mown down so as not to be allowed to come to ear till the next season, the product will be found to consist partly of rye or some other of the cereals. Oats have in like manner been transformed into rye, barley, and even wheat. Till a recent period, this phenomenon was doubted ; but it has been tested by experiment, and reported on by so many credible persons, that it can no longer be rejected. And it appears that poor- ness of soil has the same effect as mowing down. One observer states that, in a field of wheat near Lucerne, he saw ears resembling barley, but with grains similar to rye, growing from the same stem with ears of wheat. ( 65 ) Dr. Lindley, who publishes these facts, acknowledges there being no theoretical improbability in such transformations, seeing that, " in orchidaceous plants, forms just as different as wheat, barley, rye, and oats, have been proved by the most rigorous evidence to be accidental variations of one common form, brought about no one knows how, but before our eyes, and THE VEGETABLE AND ANIMAL KINGDOMS. 135 rendered permanent by equally mysterious agency." It is more than probable that the greater number of what may be called the domesticated plants, are unsuspected variations of others, which, growing wild, are recognised as different species. One noted instance of such transition has been detected within the last few years, in our different kinds of cabbage, savoy, brocoli, and cauliflower. They are all common descendants of a plant which is sometimes found growing wild upon our sea-shores, the brassica oleracea a transition which no one can appreciate till he has compared the tough slender stem and small glaucous leaf of the original, with the stout fleshy stem and large succulent leaves, sometimes gathered into a heart several feet in circumference, which he will find in the most familiar of the cabbages. What respect, it may be asked, can we attach to the doc- trine of intransibility of species, when we find its adherents wrong in so many instances ? Admit their explanation, that a mere mistake has been made in calling that species which was only variety, what guarantee can we have for the fixity of any so-called species, when it has given way in such in- stances ? What t* species, if it cannot be fixed upon such a vast assemblage as the Thallogens, or even the progeny of the Telephora sulphurea ? Apart from all theorising about the absolute characters of species, do not these facts show a transibility and intercommunion of forms totally at variance with those general opinions as to fixity which now reign in the scientific world ? In the animal kingdom, we have fewer illustrations of modifiability or transition ; but they tend to exactly the same effect. We shall here pass over the succession of forms which appears in common infusions. Neither shall we enter into the particulars of a late curious investigation by a Danish naturalist, which results in showing alternative forms in the succession of certain animals low in the scale, in- cluding the medusa; that is, as it were, A giving birth to B, B to C, and C to A again. C* 5 ) Such matters are as yet obscure, however highly they may promise in time to illus- trate this question. Let us rather look to departments of this 136 HYPOTHESIS OF THE DEVELOPMENT OP kingdom which come broadly under the observation of natu- ralists. In the mollusca there occurs a modifiability of a most remarkable nature. Fresh- water species of these, exposed to brackish water, assume, where able to survive the change, characters in the exterior form of the shell proper to their marine congeners, and involving differences from the original animal much greater than is usually sufficient with naturalists to constitute a distinction of species, if not of tribe or family. Many years ago, Pennant remarked the singular modification of stomach which the common trout appears to have under- gone in the lakes of the county of Galway, in consequence of feeding on shell-fish. The integument has become as thick as the gizzard of a bird, manifestly in consequence of an effort of nature to accommodate herself to the peculiar food of the animal. So also, when a common gull was fed upon corn, the parietes of the stomach were found, on examination after death, to be thickened. ( 67 ) The peculiar forms of the mandibles of birds are grounds of specific distinction ; yet it is now ascertained that these are variable under particular conditions as to food. It has been tried with confined birds ; and even in a wild state there occur individuals strangely modified in this respect, the magpie, woodpecker, and rook, having all been found with the crossed mandibles of the loxia.( 68 ) Look also at the changes from the wild animals to those domesticated ones which are known to be descended from them. " When the eggs of the wild goose," says Professor Low, " are taken, and the young are supplied with food in unlimited quantity, the result is remarkable. The intestines, and with them the abdomen, become so much enlarged, that the animal nearly loses the power of flight, and the powerful muscles which enabled him in a wild state to take such flights, become feeble from disease, and his long wings are rendered unserviceable. The beautiful bird that outstripped the flight of the eagle, is now a captive without a chain." Another change is the transition from grey to white plumage. In the domestication of the pig, the author last quoted admits that there are reduc- tions of the number of teeth, and variations of the number of the dorsal, lumbar, sacral, and caudal vertebrae, producing THE VEGETABLE AND ANIMAL KINGDOMS. 137 differences greater than what are usually regarded as suffi- cient to constitute species. But the most striking observations on this subject are those of M. Roulin, made during a resi- dence of several years in Columbia, relative to the races which had been introduced there in a domesticated state by the early voyagers, and allowed to run wild during the three centuries which have since elapsed. As an example, the hog : " Wandering all day in the woods, this animal has lost nearly all marks of servitude ; its ears have become erect, its head broadened, and raised at the upper part, and its colour has been rendered permanent." It has, in short, returned to a strict resemblance to the wild boar of France. The cow, also, from the cessation of the practice of milking, has lost the abundant flow of milk which is found in her species in Europe : to get milk from her at all, it is necessary that her calf should be left with her. M. Roulin arrived at the fol- lowing conclusions : that animals naturalized in new countries undergo durable changes, bringing their organization into accordance with the climates in which they are destined to live ; and that habits of independence soon make the domestic species resume the characters of the wild species from which they have sprung. We have here, it will be owned, equal proof that the tuskless hog of our farm-yards is the same animal which roams the forest in formidable state and arma- ture, as that the wild boar is the same with the domestic pig. It is difficult, after what we have now seen, to regard the idea of species or specific distinction as descriptive of a fact in nature ; it must be held as merely representing certain appearances presented, perhaps transiently, to our notice. The history of the question seems to be this. Naturalists, starting with a limited fund of observation, mainly, indeed, consisting of the remark which the most superficial observa- tion supplies, that like usually produces like, lay it down as an axiom that species is a determinate thing. In a little time, certain modifiabilities are observed. To maintain the axiom intact, these are called varieties. Afterwards, much greater variabilities are witnessed, even to the dissolution of genera among the cryptogams and cereals, and the community of 138 HYPOTHESIS OF THE DEVELOPMENT OF algae and fungi water and land plants. Still, to keep the axiom whole, these are held in doubt, or relegated to a place in the elastic region of the varieties. Such is the stage which we have now attained. But this is a process the reverse of philosophical : it is to start with a theory, and then make facts succumb to it. Were the process reversed and the facts taken first, we should see that a great modifiability exists in organic nature, especially in the humbler departments of the two kingdoms. And seeing that this modifiability presents itself within the scope of a very limited experience, it might safely be inferred that something much greater would be de- tected if our range of experience were extended, especially since the world presents us with results which can only be naturally accounted for in this manner. It is here a fact to be specially remarked, that the greatest variability, the most striking instances of transition or intercommunion of forms, are offered in the lower grades of being. In these depart- ments of nature, generation is rapid and abundant in compa- rison with the reproduction of the higher forms. What requires perhaps a century in the one case (say a series of three generations) will be accomplished in a day in the other. Nothing, therefore, seems more natural than that phe- nomena connected with the reproduction of the higher animals should require a much longer time to be evolved than those connected with the lower. The time may be, in the one case, such as to fall within our range of observation (and this range, as far as scientific accuracy is concerned, is but a day), while in the other case it may be, and indeed, on a just comparison, we should expect it to be, beyond even the whole space of what is called the historical era. Such is precisely the point to which the present theory would lead us. We see that permanency of specific distinctions in the higher or- ganisms would sink, as it has done in so many of the lower, if we had as long a time to observe their reproductive history as would, in embryology, be equivalent to the space of time during which we have observed the humbler creatures. We see this persistency and think it fixed, exactly as men have hitherto seen the solar position in the universe. We advance among THE VEGETABLE AND ANIMAL KINGDOMS. 139 the stars at the rate of two millions of millions of miles a year ; but astronomers tell us that it would take ninety millions of years to enable us to pass through the whole, even at this rapid rate. Well, therefore, might the unassisted eye and unexamining intellect presume the place of the solar system to be fixed, for it is evident that no human tradition could record changes indicating the translation. Yet we do pass on to Hercules, although forty centuries failed to remark the circumstance. So may specific distinctions in the higher animals have been changed in the course of the vast periods which geology shows to have elapsed since the commence- ment of organization upon earth, although, during that inap- preciable segment of the great cycle which has passed since man woke to the mysteries of nature, no single transition of the kind might have been observed. The whole case reminds us greatly of the objection which stood against the earth's motion from the days of Aristarchus downwards, that there ought in that case to be an observable parallax. As there was no observable parallax, because the earth's orbit is an in- significant space in comparison with the distance of the stars, so is our observation of animal changes insufficient to show transitions of species in the higher grades of the kingdom, be- cause it is a mere span in comparison with the vast ages actually concerned in the phenomenon. A similar principle of explanation applies to the alleged tendency of variety to be obliterated. While it is only to be expected that a single animal showing an originality of form will fail to impress it on its posterity, if it be absorbed in alli- ance with animals possessing no such peculiarities, there is no reason to believe that a variety uniting with a creature like itself will not have descendants of its own character. We judge on this question in the midst of a fully-peopled world; but we must cast back our minds to a time when it was only in the course of being filled with living things. We must think of a time when, for example, over large portions of the surface mountain tracts were rising, perhaps beside low and marshy grounds, or when forests began to spread over exten- sive regions. Here a new field of existence is presented. 140 HYPOTHESIS OF THE DEVELOPMENT OF The fecundity of nature has ordained that her creatures shall ever be pressing upon the verge of the local means of subsist-. ence. A colonizing principle accordingly comes into play. On such an occasion, it might be that individual wading birds began to advance into dry grounds and woods, elected to the new life perhaps by some of those varieties of appe- tency which occur in all tribes ; thus exposing themselves to new influences, and ceasing to experience those formerly ope- rating, until, by slow degrees, in the course of a vast space of time, the characters of the pheasant tribes were evoked. ( 69 ) Here, it will be at once perceived, re-absorption of peculiarities was not likely to occur, for the field of colonization, so to speak, was sufficiently wide to allow of the new families wandering farther and farther away from the original grounds and the ancestral tribes, while return was prevented by the full popu- lation continually pressing behind. Altogether, this presents a very different view of varieties from that which is commonly presented, when we see a single peculiar individual standing in the midst of, and necessarily allying itself to, the original stock. The process of variation as a consequence of changed conditions and appetencies being left unchecked, and that for a vast space of time, we obtain at length creatures fixedly peculiar ; that is, however, merely creatures which appear so, because there is no replacing them in the former conditions in this densely-peopled globe, and, though there were, the retrogression to the anterior forms would require a space of time beyond the range of human observation. It may now be remarked, that, in this hypothetic variabi- lity, the possibility of re-union may, and in all probability does, depend upon the degree of similarity which still exists in the different individuals, supposing them to be members of the same stirps or line of being, for I believe that no others are capable of intermixture. As has been remarked by a venerable naturalist " Many bulbous roots that have been increased during a long succession of years by offsets, become absolutely incapable of bearing seed ; 'and it is not more strange that plants which in different soils and climates have diverged from the original form of the first created individual, THE VEGETABLE AND ANIMAL KINGDOMS. 141 should refuse to bear seed by the one which has departed most widely, and yet produce it readily by another, which still agrees with it in some important points." ( ;0 ) Admit this, and the grand basis of specific distinction, the possibility of intermixture, can no longer be laid hold of. Plants and animals of one line are only to be expected to unite, which, being of one grade of organization, are also sufficiently near to each other in those peculiarities liable to modification from external causes, on which the so-called distinctions of species are grounded. The illustrations of our hypothesis are now closed. We have seen that, even judging from short spaces of time, there is a great and incontestable modifiability of organic forms, so great as to have absorbed the presumed distinctions of species in many noted instances. We have seen that this modifi- ability, by some hidden law, immediately obeys external con- ditions. It has also been seen that, though no transition from grade to grade was ever observed to take place, the means and mode by which it could naturally happen are not concealed from us ; they are pictured before our eyes in the metamorphosis of the tadpole, and even practically exem- plified in a narrow degree in the natural history of the bee. It has been shown that no organism is independent, but all stand in a web of intimate relation, undeniably indicating that their origin is one connected phenomenon. It has been seen that the higher animals, when their organization is ex- amined, are only improvements upon the lower advanced forms of the same beings ; and the same holds good regarding plants. In conformity, too, with this gradation of forms, is the succession of the actual animals throughout the geological ages ; a fact most important not merely one calling to be explained, as is at the utmost allowed by men of science of the present day, but one which helps to explain, a piece of actual tangible evidence, and bearing wholly, when taken in connexion with proofs of other kinds, in favour of the natural origin of species. Surely when, in addition to all this, we learn that life is believed by many men of science to spring occasionally, even now, from inorganic elements 142 HYPOTHESIS OF THE DEVELOPMENT OF when we find that, moreover, it is generally admitted by that class of men to be in itself a simply natural phenomenon, we cannot but say that at least VESTIGES have been seen of the natural ordinances or arrangements, by which the Al- mighty Father caused this globe (and probably others within our ken) to be overspread with the many creatures whose per- fection is his praise. Rigid proof is not, indeed, attained ; but we have all the evidence which is attainable in the case. It is evidence from various quarters, all perfectly homogeneous ; it harmonizes with everything else which science tells us of the history of the universe ; it supplants a mean with an exalted idea of the Deity, and has nothing opposed to it but the prejudices formed in the nonage of our race. For these reasons, I must, till disproof is offered, regard the theory of Progressive Development as the true explanation of the origin of organic nature. The simplest and most primitive types of being, under a law to which that of like production is subordinate, gave birth to a type superior to it in compo- siteness of organization and endowment of faculties; this again produced the next higher, and so on to the highest. There has been, in short, a universal gestation of nature, analogous to that of the individual being, and attended as little by circumstances of a startling or miraculous kind as the silent advance of an ordinary mother from one week to another of her pregnancy. We see but the chronicle of one or two great areas, within which the development has reached the highest forms. In some others, as Australia and the islands of the Pacific, development has not yet passed through the whole of its stages, because, owing to the comparatively late uprise of the land, the terrestrial portion of the develop- ment was there commenced more recently. It would com- mence and proceed in any new appropriate area, on this or any other sphere, exactly as it commenced upon our area in the time of the earliest fossiliferous rocks, whichever these are. Nay, it starts every hour with common infusions, and in similar humble theatres, and would there proceed through all the subsequent stages, granting suitable space and con- ditions. Thus simple after ages of marvelling is Organic THE VEGETABLE AND ANIMAL KINGDOMS. 143 Creation, while yet the whole phenomena are, in another point of view, wonders of the highest kind, being the un- doubted results of ordinances arguing the highest attributes of foresight, skill, and goodness on the part of their Divine Author. Early in this century, M. Lamarck, one of the most dis- tinguished of modern naturalists, suggested that the gradation of animals depended upon some general law which it was important for us to discover. So far he was right ; but the theory which he consequently formed with regard to the causes of the varieties of animated being was so far from being adequate to account for the facts, that it has had scarcely a single adherent. What M. Lamarck chiefly grounded upon was the well-known physiological fact, that use or exercise strengthens and enlarges an organ, while disuse equally atrophies it. He conceived that, an animal being brought into new circumstances, and called upon to accommodate itself to these, the exertions which it consequently made to that effect caused the rise of new parts : on the contrary, when new circumstances left certain existing parts unused, these parts gradually ceased to exist. Something analogous was, he thought, produced in vegetables, by changes in their nutrition, in their absorptions and transpirations, and in the quantity of caloric, light, air, and moisture which they received. This principle, with time, he deemed sufficient to have produced the advance from the monad to the mammal. His illustrations were chiefly of the following nature. The bird which is attracted to the water by the necessity of seek- ing there its food, wishes to move about on the surface of the flood, and for this purpose strikes out its toes. Through the consequent repeated separations of the toes, the skin uniting them at the roots is extended and at length becomes webbed. In like manner, the shore bird which has no desire to swim, but has to approach the water for food, is constantly subject to sink in the mud. The bird, disliking this, exerts all its efforts to lengthen its legs ; the result is, that, by continual habit for many generations, the legs of this order do at length become long and bare, as we see them. The error of the 144 HYPOTHESIS OP THE DEVELOPMENT OF theory is in giving this adaptive principle too much to do. What undoubtedly is effectual in modifying the exterior peculiarities of animals was obviously insufficient to account for the great grades of organization. In the present day, we have superior light from geology and physiology, and hence arises my suggestion of a process analogous to ordinary ges- tation for advancing organic life through its grades in the course of a long but definite space of time, with only a recourse to external conditions as a means of producing the exterior characters. It must nevertheless be acknowledged that the germ of this natural view of the history of the world is presented in the work of Lamarck. But the idea that any of the lower animals have been con- cerned in any way with the origin of man is not this de- grading ? Degrading is a term expressive of a notion of the human mind, and the human mind is liable to prejudices which prevent its notions from being invariably correct. Were we acquainted for the first time with the circumstances attending the production of an individual of our race, we might equally think them degrading, and be eager to deny them, and exclude them from the admitted truths of nature. Knowing this fact familiarly and beyond contradiction, a healthy and natural mind finds no difficulty in regarding it complacently. So also, on becoming aware of the genetic history of our species, we might expect a rational and well- ordered mind to receive the idea with submission, as a view of the manner in which Divine Providence has been pleased in this instance to work. One source of the prejudice here to be contended with rests in our associations with the word ancestry. From seeing our immediate seniors possessed of venerable qualities, we naturally incline to venerate an an- cestry ; we presume its constituent elements to be something superior to ourselves. When called upon, therefore, to place any of the inferior orders of Being in this relation, a shock unavoidably follows. But here the error lies in transferring our idea of the qualities of a sire or grandsire to a collective ancestry. The elder people of the earth are in reality its children, and we are its true senate. The feeling due to THE VEGETABLE AND ANIMAL KINGDOMS. 145 early generations is the half-pitying benevolence which we daily bestow upon childhood. It follows that the still earlier generations antecedent to the perfection of the human type, ought to be regarded with an extension of this same feeling the modification of it which humane natures daily exemplify in their treatment of the inferior animals. Our children, it may be said, are the representatives of the first simple and impulsive men of the earth : the lower animals represent the earlier pre-human stages of life. The right conception of the case is, that in these stages we are not to look for what is venerable, but, on the contrary, for what is humble and ele- mentary. We are to expect but the primitice of man's mas- terful life something not even ascending to the dignity of " the infant mewling in its nurse's arms." If thus prepared, we should experience no shock on hearing that the human form was preceded genealogically by others of humbler aspect, no more than we are on learning that every indivi- dual amongst us passes through the characters of the inverte- brate, fish, and reptile, before he is permitted to breathe the breath of life. A deep moral principle seems involved in the history of the origin of man. He is the undoubted chief of all creatures, and as such may well have a character and destiny in some respects peculiar and far exalted above the rest ; but it appears that his relation to them is, after all, one of kindred. Along with his authority over them, he bears from nature an obligation to abstain from wantonly injuring them, and as far as possible to cherish and protect them. Good men feel this duty, as if it were a command from a source above themselves. It seems to them, that if the help- lessness of childhood calls for kind and gentle treatment, much more does the essentially weaker character of the dumb creature. And if the innocence of infancy is touching, still more so is the even more harmless character which (over- looking carnivorous instincts implanted in certain families for a wise purpose) attaches to the lower animals. It is common, under the influence of prejudice, to do gross injustice to the characters of these denizens of nature's common. We do not sufficiently reflect on their respectable qualities. Yet we L 146 HYPOTHESIS OP THE DEVELOPMENT, ETC. must go to the dog for a type of the virtue of fidelity, and to the bee for that of industry. The parental affection of many animals is not below, if it is not considerably above, that of human mothers. Man nowhere exemplifies the virtue of patience in the practical perfection in which we see it in the horse and many other creatures which become the slaves of his convenience. Nowhere does he display that perfect mo- deration in wants. Alas for man's boasted superiority in how many respects does it fail beside the unassuming merits of the mere commonalty of nature ! 147 AFFINITIES AND GEOGRAPHICAL DIS- TRIBUTION OF ORGANISMS. ALL truth being self-consistent, we might expect that this view of the history of organic nature, if sound, would ac- cord with a just classification of plants and animals, supposing such to exist. It is certainly very desirable that our theory could have been subjected to this test ; but it cannot be, for naturalists are as yet only struggling towards true classifica- tions in both kingdoms. It becomes necessary, nevertheless, that we should make some inquiry into that order which has long been alleged to exist in animated nature, as, if any such thing truly exist, it either must agree with a genealogical system, or become its condemnation. The result of my own investigations is, that there is an order in animated nature, but that it has hitherto been much misunderstood both by those who incline to a theory of deve- lopment and others. The former naturally took hold of the idea of gradations, because it generally accorded with the notion of development. They pointed to that " chain of being," or series of ascending forms, which had long been supposed to extend between the animalcule and the human being. It was on the other hand successfully shown that beings did not form " a single and continuous series ;" that it was " impos- sible to place all living animals in such an order that we may L2 148 AFFINITIES AND GEOGRAPHICAL always pass from one species to another by following a de- crease in perfection." " On the one hand, there are classes of animals so insulated, that nothing connects them with others." " On the other, there are types of organization which are absolutely indivisible, and of which the most per- fect beings are superior to the mean of another type, while the most imperfect are inferior to it." All this is true : it remained unanswered by the advocates of the development theory ; and such was the position of the question when the earlier editions of the present work made their appearance. But the error actually lay in the original idea of a chain of being. The animal kingdom (and, by consideration of parity, we may presume the vegetable also) consists of a plurality of series going on side by side with each other, but not all to the same point in the scale. No wonder, accordingly, that some appear insulated, or that the highest of some types are superior to the meanest of others, while the most imperfect appear inferior. Nor is this merely a hypothetical view of the animal kingdom. It is clearly pointed to by some of the most interesting discoveries in embryology. It is supported by several important considerations regarding the general cha- racters of particular series. It likewise harmonizes with that order of fossils, which I have ventured to describe as not something calling in itself for explanation, but a fact which we may look to as one of the means of explaining something else the whole history of organization upon earth. Finally, such reformation as this new view calls for in our classifica- tions, is accordant in its general demands with all those re- cently effected by the greatest naturalists, by which external and comparatively accidental characters are overlooked, and only the more essential affinities regarded. If it goes beyond the march of living naturalists, it goes in the direction in which they are going, and over ground, to which I believe they must quickly come, whether they adopt a genealogical view of the organic world or not. The divisions of the animal kingdom, as we find it in Cuvier, are partly into grades, with a regard to dignity of organization first into Vertebrata (having an internal skele- DISTRIBUTION OF ORGANISMS. 149 ton) and Invertebrata, and afterwards into such divisions as these of the vertebrata, namely, Mammalia, Birds, Reptiles, Fishes. In these grades are comprehended animals of very various character, animals which only agree in this particular of a community of grade or rank. But other divisions in the common classifications are into groups or series of animals closely allied to each other in form and of one general cha- racter, as, for example, the cephalopoda, the echinodermata, the Crustacea. The one kind of division may be said to be transverse, the other longitudinal. Such a diversity gives rise to a suspicion that there is something wrong, something out of accordance with nature. And so it is. The true fundamental divisions are entirely of the latter kind longi- tudinal ; there only do we find persistence of characters ; the other so-called divisions are only the marks of stages which the true divisions, the Stirpes of being, have reached in their respective courses. It is nevertheless necessary, in the mean- time, to keep the existing classification in view, and to use its language, in order that my own views may be intelligible. Cuvier divided the Invertebrata into three great masses, the Radiata, the Articulata, and Mollusca. Of these the tw last appear as co-ordinate, though distinct from each other ; while the Radiata, again, may, excepting one class, be con- sidered as forming a kind of basis for the whole kingdom. The RADIATA are all of them animals of exceedingly simple structure, mostly inhabitants of the waters, many of them pro- pagating not by ova, but by division of their bodies, or by the throwing out of little bud-like excrescences. In this lower region are comprehended the Infusory animalcules, Internal Parasites (Entozoa), Sponges, Polyps, Sea-nettles (Acalephcz), and some other obscure classes. Some of these appear to be distinct and independent series, which advance no further ; such, in particular, are the internal parasites, which necessarily do not pass to any higher grade, because they have no sphere for further development. Others form the roots, as it were, of higher families. There are two admitted methods of investigating the affi- nities of beings. One is to observe the connexion between the 150 AFFINITIES AND GEOGRAPHICAL forms of the mature organisms ; another is to examine the embryotic progress, and watch the succession of forms there presented. It has for some time been ascertained that no animal, in the course of its development, passes through the forms of all the animals meaner than itself. For example, the sea-nettle is at one time like the monad, an infusory animal- cule, and then like the polyp; the mollusk is successively like the monad and polyp, but never like the sea-nettle. The articulate animal, again, is never like the polyp or sea-nettle, but proceeds at once from the monad form to that of the worm. This Professor Owen calls being " obedient to the law of unity of organization only in its monad stage." ( 7l ) The fact has been held as a difficulty in the way of the doctrine of unity ; but perhaps it is only one of the same nature with that intimated regarding the assumed scale of being. I see animals classed by their affinities in distinct lines, or series, which I regard as stirpes or races. I would therefore expect the unity of organization to be liable to some such limitation as Mr. Owen points out. Is it not, in reality, that each stirps has a unity of organization for itself, or, in other words, that there is such a unity only as far as each particular series of animals is concerned ? These breaks in unity and the breaks in the chain of being are but one thing : they are only dis- turbances to our preconceived ideas, not to a true view of nature drawn from its realities. I shall not attempt to place all these obscure animals in genealogical series. The state of zoological science demands that such an effort should be postponed for several years to come. Let us limit our attention to one class, the Echino- dermata, or star-fishes, which are perhaps improperly ranked with other Radiata, seeing that their character is so much superior. In general highly organized, and enjoying free movement at the bottom of the sea, these animals are signally destructive. Admitted to be in their lower forms intimately allied to the Polyps, they probably start in some portion of that extensive order. In their own class, however, as far as traceable backwards, they commence with the encrinus or stone-lily, a group of animals of which we have seen many DISTRIBUTION OF ORGANISMS. 151 varieties flourishing in the early seas, but which are now nearly extinct. The creature consisted of a stomach and arms, surrounded by long tentacles or arms, placed upon the top of a stalk fixed to the sea-bottom, the whole being com- posed of numberless minute calcareous plates, connected by gelatinous substance. In more advanced forms of the same order, (as the comatula and the extinct marsupite), the body and arms desert the stalk, and betake themselves to a free- swimming life ; but, as has been elsewhere mentioned, the young comatula lives for a time as an encrinus ; that is, upon a stalk. Seeing that the same animal, in an earlier embryotic stage, represents a polypidom, we conclude that in the poly- piaria is the origin of the echinodermatous line : it is first the polypidom, then the encrinus, then the free-swimming coma- tula, or feather star, the last being one of the most graceful animals in existence. In the higher genera of the latter family, the tentacles are shortened and reduced in number. In the Ophiurce, there are only five long and simple rays pro- jecting from the central body. Afterwards, in the Asteriada, or true star- fishes, the central part dilates step by step, until it fills up the interstices between the rays, and the form be- comes a pentagonal disk. From this there is a clear passage to the Echinus or sea urchin, which is merely a spheroidal animal in a calcareous case, through which numberless spines or tentacles project, for locomotion and the collection of food. This form again becomes elongated into the cylindrical soft- bodied Holothuria, with a circle of tentacles at the oral ex- tremity ; thence the transition is easy to the genus Fistu- laridcs, animals externally worm-like, and possessing the rudiment of a heart, with red blood in the arteries, so that, in this last echinoderm, we may be said to have come nearly, if not fully abreast with, the annelides, and to be approximating to some of the humbler fishes. ( 72 ) The reader cannot fail to have been struck by the great number of forms passed through in this line, in comparison with any other, before leaving the radiate sub-kingdom ; but, in reality, the echinodermata, though of radiated form, are much superior to the rest of that division in their organization, which is, if not complicated in 152 AFFINITIES AND GEOGRAPHICAL the usual sense of naturalists, full of extremely curious minute work. Their whole destiny seems to be of a high kind, for in the stone record their line of forms stands parallel with others in which the whole of the three lowest sub-kingdoms are passed through. Polypiarian animals and encrinites appear in the Silurian and many subsequent formations; at the commencement of the carbonigenous era, the latter are so abundant that we walk over large tracts of country, where the rocks beneath our feet are almost wholly composed of their remains. The Asteriadae appear in the upper Silurians, and are but faintly seen until the Lias, when they become conspicuous. In the Oolite, the Echinidae make their ap- pearance. These are the last which we could expect to be preserved in rocks, as the higher families possess no hard parts ; otherwise, we might perhaps have seen the succession of this class of fossils continued into the holothurise and fistularidae. It cannot fail to be noticed how well the pro- gression of forms agrees with the order of their appearance in the geological ages. The ground is now cleared for the two grand series of in- vertebrate animals, and first of the ARTICULATA. These are generally describable as animals " composed of a succession of rings, formed by the skin or outward integument, which from its hardness constitutes a kind of external skeleton ;" one class, however, the Annelides, have no hard investment. The pedigree of the Articulata is very brief. The embryo in most classes passes at once from the monad to the worm form, and then the articulate character is assumed. It can therefore scarcely be said that the radiate sub-kingdom comes before the articulate, though the one is lower in organization than the other. There is indeed reason to believe that the great classes of the Articulata are distinct stirpes, the commence- ment of each of which is little more than a step from the in- organic form of matter. This may seem inconsistent with the maxim Natura per saltum nihil agit; but maxims must be obedient to facts, not facts to maxims, and we may deem that a leap which in reality is none. DISTRIBUTION OF ORGANISMS. 153 The necessity of taking liberal views of the procedure of nature in the development of the organic world, is im- pressed upon us by a character found in the very first order of the articulata to which our attention is called. That the Annelides (worms) are the humblest of the articulate animals there is now no doubt ; yet, unlike their superiors, almost all of them have red blood, a feature of the highest sub-kingdom. Four leading forms in this class are described. Of the Tubicolida, or those inhabiting tubes, the serpula is an example. It forms for its habitation, usually upon some sea-immersed stone, an irregularly twisted calcareous tube, out of which it presents, floating in the water, a fan-like branchial apparatus of beautiful colours. The second order, Suctoria, is represented by the well-known leech ; the third by the earth-worm ; the fourth by the sea- mouse (aphrodita). In all of these groups, we see distinct advances in organiza- tion, and this is traceable in some in an interesting confor- mity with changes of scene and mode of life, from fixed situa- tions to free movement in the sea, from thence to the shore, and thence again to the land. From the iJfais, a simple marine worm which at the recess of tide burrows in the sand, there is a clear passage to the common earth-worm, which adopts a similar retreat on land, and comes to the surface when rain is falling. The fourth order, Dorsibranchiato, so called because of gill tufts ranged along the back, have an equally clear affinity, implying ancestral relationship to cer- tain land animals, which, however, naturalists at present regard as an independent class. The nereis, a well known dorsibranchiate, is an animal of great length, composed of a consecutive series of rings, each having a couple of processes at each side, which are used as oars for propelling the body through the water. One species is four feet long, and consists of several hundred segments. By conversion of the water- breathing apparatus into one fitted for aerial respiration, an increase of firmness and density to the external integument, and the development of a couple of limbs for each ring of the body, we see the nereis, as it were, transmuted into the 154 AFFINITIES AND GEOGRAPHICAL Myriapod.C 3 } Here, however, there may be more than one line of passage ; for the two great families of the myriapods, the Julidse and Scolopendridse, are diverse in character, the former being vegetable feeders, the latter carnivorous, and it appears as a rule in the genetic system, that true carnivores are always apart. Confining our view to the Scolopendridae, we see a remarkable continuity of character and habits transmitted to them from the presumed marine ancestor, (nereis,) allowing for the altered medium of existence. The scolopendra is an animal furnished with powerful destructive organs ; and, living under stones and the bark of trees, and in fissures generally, it is his custom to wind insidiously along, and dart upon any little animal which comes in his way. Of the nereides, on the other hand, we are told that they " usually live in the excavations of littoral rocks, in the hol- lows of sponges, in the interstices of the radicles of thalas- siophytes, under stones, and in general in all bodies which present fissures more or less profound . . . They all ap- pear to feed upon animal substances. . . M. Bosc tells us they live upon polypi and small worms, on which they throw themselves, by darting the anterior part of their body, which they have first contracted." The next articulate class demanding attention is the Crus- tacea, animals in which the annular sections are covered with a calcareous shell, and provided with jointed limbs, the respi- ratory apparatus being branchial ; all are aquatic, except some of the higher genera, which occasionally adventure upon the land. They are in two great groups, entomostraca and malacostraca, the former being the simpler, and exclusively marine. Emmerich considers the Trilobites which figure so conspicuously in the early rocks, as between the two divi- sions, but most nearly allied to the first ; whence it would appear that the Crustacea which make so early an appearance in the rock series, are humble animals, only preceded in their own sub-kingdom by a group, which, from their slight forms, might be ill-adapted for preservation in strata exposed after deposition to a high temperature. The .geological history of the Crustacea tallies in other points with their gradation. In DISTRIBUTION OF ORGANISMS. 155 the triassic epoch come the Macruri, which prevail to the present time ; afterwards, in the tertiary era, come the Brachyura. These are the fossil orders which have been best studied, and it is M. Agassiz who remarks " they succeed each other in the series of formations in the order of their organic gradation." The same naturalist remarks " the inti- mate analogy between these different types and the phases of the embryonic development of the Crustacea, which MM. Rathke and Erdl have afforded us the means of becoming acquainted with." As elsewhere remarked, the young of the decapoda are of the entomostracous form, and thus denote a passage of the one from the other. In one family of the Crustacea, there is a striking illustra- tion of what I regard as the true history of species. This is the family to which the well-known hermit-crab belong (Paguri,) distributed extensively in the tropical American islands, and upon our own coasts. Animals of this kind live in molluscan shells deserted by their proper tenants. They select one at the first for their residence, and afterwards, as they increase in size, they remove to larger ones. With the hind part of the body inserted in the hollow shell, they present the head and feet outwards. They move about in the shallow water, upon the shore, and even upon dry land, with great freedom, dragging their adopted mansion after them. A very slight examination of these animals shows that they are adapted by special peculiarities for this kind of life. In the common British hermit crab, the third and fourth pairs of locomotive limbs are of small size, being buried wholly within the shell, where they are applied to the columellar fold, as a means of fastening the animal in the recess. Farther in, and also employed in fastening the body to the shell, is the caudal part, with two holders developed for this express purpose, and as rough as a file. The hold is still further secured in some species by rows of suckers along the abdomen. Add to all this, that, for want of room at the mouth of the shell, only one of the pincer claws is well deve- loped, usually the right, while only the two front pairs of feet are used for locomotion, and we see that, whether we take 156 AFFINITIES AND GEOGRAPHICAL these crabs as a species, a genus, or a family, their ordinary form that thing which naturalists regard as immutable, and as originally the effect of a special creative effort is in direct relation to the existence and forms of turbinate shells for- merly possessed by a different class of animals, which must therefore have existed before the hermit crabs. Now mark the credulity to which the adherents of immutability must here be reduced. They must believe that the Creator, having a particular regard to the fact of molluscan shells lying use- less on the shore, formed, by special care or fiat, a family of crabs to occupy them. They must believe that the rough- ness of the caudal appendages, the development of suckers along the abdomen, the reduction of the two hind pairs of limbs, and the left pincer claw, were all subjects for this special care, and were beyond the power of what an eminent geologist calls " vulgar nature." Surely the Deus ex machind was never more remarkably exemplified. See, on the other hand, how these facts are accounted for on the development theory. According to this new light, the hermit crabs are simply a portion of some greater section of the crustacean class. Their peculiarities are modifications from the parent form, brought about in the course of genera- tions, in consequence of an appetency which had led these creatures to seek a kind of shelter in turbinate shells. They are as truly creatures of the Great God, as if they had been made in the manner of a human artist modelling a figure. But the means were inherent natural forces in the constitu- tion of the original tribe, tending, in generation, to accommo- date organic form to physical circumstances. The next class in general rank is the Insecta, a wonderfully varied group, yet all agreeing in having thirteen segments and three pairs of legs ; all, moreover, respiring by means of tracheae or tubes permeating the body, an arrangement having reference to their peculiar mode of locomotion, which, in the majority of species, is by flight through the air. The fact of the greater number of insect genera passing, in their larva state, through the annelidan or myriapodous form, points to these classes as their genetic origin ; yet this is a DISTRIBUTION OF ORGANISMS. 157 point on which the benefit of further investigation is desir- able. In the case of the Arachnida (mites and spiders), the highest articulate class, no humbler form is traceable in the embryo ; it is therefore impossible to assign them any pedi- gree. Can it be possible that the arachnida, or these with the insecta, have sprung almost or wholly at once from inor- ganic elements under the proper electric influences? On this subject, we are quite unprepared to make any positive affirmation ; but it certainly is remarkable that in no depart- ment of the animal kingdom, besides the infusoria and entozoa, have there been more frequent appearances of an aboriginal commencement of life than in the insecta. The acarus so often produced from certain solutions, where ova were rigidly excluded, is a lowly member of the arachnida. We now come to the MOLLUSCA, a portion of the animal kingdom, the importance of which, in point of numbers and the part they play in creation, none but students of zoology could fully appreciate. The infinite variety of bivalve and univalve shells presented upon our own coasts and brought from all parts of the world, will convey some idea of the mul- titude of forms comprehended under this sub-kingdom. The whole mass is after all resolvable into three divisions ; one of them comprising headless mollusks in bivalve shells ; the other two, headed mollusks in univalve shells (some, however, of all the three divisions being naked). The whole sub-kingdom appear to have a very brief genesis in the radiata, the only preceding forms in embryo being the infusorial and polypian. Here, too, as in the Articulata, we find that we must start at a point very near the fountain-head of organic existence. In the headless division, naturalists place three sub-divi- sions, called by them classes, in the following rank, according to ascending grade of organization Tunicata, Brachiopoda, and Lamellibranchiata. The two latter are the shell-fish of popular observation, headless, and mostly sessile, or destined to spend their lives in fixed positions. The Tunicata are similar in all essential respects, except in being of humbler organization, an inclosed, not in shells, but in a cartilaginous or coriaceous integument ; whence their name. It thus ap- 158 AFFINITIES AND GEOGRAPHICAL pears that the Brachiopoda, which are the predominant fossils of the Lower Silurian era, are the first animals we meet with in this line, having parts capable of commemorating their ex- istence. While the Brachiopoda are generally inhabitants of deep seas, the Lamellibranchiata, among which are included the oyster, muscle, and other testacea, affect the beds of shal- low seas, whence they spread in a variety of genera, towards shores, the mouths of rivers, and into fresh water. The Lamellibranchiates are higher than the preceding class ; they are the first bivalves which possess a true hinge. It is also remarkable that, with the decline of the brachiopods, at an early point in the secondary formation, rises the lamellibran- chiate class. There is here, therefore, an improvement in organization, an advance in habitat landward, and a succession of existence in the geological ages, all in harmonious connec- tion. NOT is this all. The lamellibranchiata are again di- visible into monomyaria arid dimyaria, the former having one adductor muscle, and the latter two ; the former, moreover, being intermediate between the brachiopods and dimyaria in respect of non-symmetrical form. Now the monomyaria succeed the brachiopods as an abundant and predominating form, and are succeeded again, in that respect, by the dimyaria. This beautiful harmony between the fossil history of the acephalous mollusks and their order in progressive organization is expressly declared by M. Agassiz. The three highest molluscan classes, univalved, possessing heads, and with hardly an exception destined for independent locomotion, stand apart from the bivalve orders; generally superior in organization, as beseems their higher destiny, but not on that account to be held as an advanced form in the same genealogy. The lowest univalve class called Pteropoda, from their mode of progression by a couple of wing-like membranes projecting from the neck may be described as marine slugs, generally of small size, many of them naked, others protected by a very delicate shell, which swim through the ocean in vast multitudes ; one species (clio) being in such abundance in the circumpolar ocean as to form the chief food DISTRIBUTION OF ORGANISMS. 159 of the whale. Professor Edward Forbes expresses his opinion that the larva of the pteropod will yet most likely be found to resemble an ascidian polype ; inferring a very brief descent from the starting-point of life in its class. The Gasteropoda & class of many families and genera, including limpets, whelks, cowries, snails, etc. have compa- ratively a high organization, the nervous system more con- centrated, the nutritive more elaborate, but yet are of slug- gish habits, usually moving by alternate contractions and ex- pansions of a fleshy disk placed upon their stomachs ; hence the name of the class. Many of the gasteropods are naked, others possessed of but slender protection. A large propor- tion are vegetable feeders, the marine species battening upon sea- weed, the terrestrial species upon herbage and fruit ; the rest are flesh-eaters, but the general character of the Gas- teropoda as a class is harmless, like that of the herbivorous mammalia. A clear gradation of forms passes through some of the families, from the simple cone of the limpet to the spiral of the snail. The descent of the class appears to be from some families of the preceding ; for " they all," says a minute observer of nature, ( 74 ) " commence life under the same form, both of shell and animal ; namely, a very simple spiral, helicoid shell, and an animal furnished with two ciliated wings or lobes, by w r hich it can swim freely through the fluid in which it is contained. At this stage of the animal's exist- ence, it corresponds to the permanent state of a Pteropod." In the univalve mollusks, as in the bivalves, it clearly appears that the humblest families are destined to a fixed place in the depths of the ocean. As we advance through the higher groups, we find, in parallel steps with an improvement in the organs of animal life (for example, the splitting of the sexes into different individuals), an advance in the sphere of existence, to a life on the surface of the ocean, to fresh water, and even to dry land. The humble Helicidce (snails) a family of the Gasteropoda, are the first animals which we en- counter as adventuring upon the firm surface of the globe. And it is interesting to remark, in this progression, the 160 AFFINITIES AND GEOGRAPHICAL requisite change in the mode of respiration namely, from branchiae, the apparatus necessary in aquatic life, to a vas- cular air-sac, the first form of lungs the proper breathing organ of terrestrial animals. In the peculiarly destructive Cephalopoda, we recognise the highest organization of which the molluscan form appears capable ; it includes the orthoceratites, ammonites, belemnites, etc. of the rock systems, and the nautilus and cuttlefish of the present era. Their descent is probably from the carnivorous families of the pteropoda ; for " the nucleus of their shells," says the naturalist last quoted, " is a spiral univalve, similar in form to the undeveloped shells above alluded to [those of the embryo gasteropods] ; and it is yet to be seen whether all cephalopoda do not commence their existence under a spiral-shelled f>teropodous form." It has also been remarked, that " the shells of two species [of pteropoda] afford indica- tions of a transition towards the cephalopoda ; one resembling in its straight conical form the belemnite and many other ex- tinct genera of that class, and the other having a partially formed chamber at the lower closed extremity ; and similar evidence is afforded by their internal structure." ( 75 ) This ge- nealogy, if it shall be affirmed, will afford an important illustration of the geological history, because it will show that cephalopoda might be expected to make their appearance as early in the rock series as any other mollusks possessing parts equally fitted to commemorate their existence. These ani- mals are to be supposed as an ultimate form, reached, not through the medium of all the lower molluscan orders, but only of one, and with respect to that one, it so happens that, though possessing hard parts of such delicacy as to have little chance of preservation, relics of it have been discovered as far down as any cephalopodous remains.( 76 ) This contemporaneity of the cephalopoda with the gasteropods and brachiopods, it may be remarked, would be in harmony with what we know of the economy of nature with respect to the destructive ani- mals. They seem to bear a relation to those upon which they are destined to prey, and to be a necessary accompani- ment to them. Hence they would require to be upon a diffe- DISTRIBUTION OF ORGANISMS. 161 rent genetic line which actually appears, in every advance of the animal kingdom, to be the case and developed con- temporaneously with the weaker tribes, the fertility of which would otherwise produce complete anarchy. Granting, then, this pedigree for the cephalopoda, it would be no anomaly in our theory, although remains of inferior mollusks should never be found lower down in any part of the earth. The cephalopods, though so highly organized in comparison with the gasteropods, do not advance, like these, to land forms, with apparatus for aerial respiration. They are, as a class, restricted to a pelagic life, admitting of occasional appear- ances on the surface of the ocean. Their respiratory system is accordingly branchiate, yet with marks of grade which are worthy of observation. It is, in the words of Professor Owen, a law determining animal rank, that " increased number [of parts] irrespective of correlative structure, in an organ of the animal body, is ever a mark of its inferiority." By this test, the nautilus, with its four branchiae, sinks below the belemnite and the cuttle-fish with only two ; and such is the basis of a divi- sion of the cephalopoda. In the whole of this order, however, there is a remarkable advance of the nervous system, though only to the effect of enabling the animal to supply itself with food by conquest over the inferior tribes. The nervous centres, which in lower mollusca were only protected by coverings which also served to cover the rest of the body, now become of sufficient importance to have a special protec- tion, in the form of cartilaginous plates, which naturalists interpret as the rudiment of an internal skeleton. In this way, the cephalopoda approach the borders of the vertebrate sub-kingdom. This remarkable class of animals affords in its details some evidences in favour of the development theory. The humble form of a straight or slightly curved shell prevails in the earlier ages. Curved shells increase afterwards. There are also tolerably distinct appearances of a transition of forms in the genera of clymenia, goniatite, and ceratite, which make their appearance in this succession in the rock formations. 162 AFFINITIES AND GEOGRAPHICAL The dibranchiate belemnites commence in the oolitic epoch, ushering in the jD sepise, the highest of all the cephalopo- ^ dous orders, and which have since con- o ^ tinued to exist. It is worthy of remark that, in the | succession of rocks, the forms of the -f. cephalopoda change much more abruptly than appears to be the case with other rt less organized mollusca ; that is, there jj are more Decided as well as more fre- .2 quent examples of what geologists call change of species in this class than in others. This is only one of the many =g ^ proofs of law in these phenomena. On e the theory of interferences, why should there be entire renewals of some sets w of animals and not of others? On | .2 the theory of law, we only see each -i line of organic being undergoing the modifications appropriate to its special * constitution, in connexion with the effi- g, | | | cacyupon that constitution of external | | !~ NATURAL RESOURCES LIBRARY 2101 Valley Life Sciences Bldg. 642-2531 LOAN PERIOD _** MONTH LOAN ALL BOOKS MAY BE RECALLED AFTER 7 DAYS DUE AS STAMPED BELOW. ttAu 2 9 ?n' SUBJECT TO REGAI i IMMED1AILLY ,-*n**? a F?fx fcysf-** 5 * PH^ uvxw PK FtB 1 8 2005 Don AM ? r W rxcJo. JUH * J " Moffi" >RM NO. DD8 M 11-02 UNIVERSITY OF CALIFORNIA, BERKEL Berkeley, California 94720-65 CS3C '' cc < < SSV t c <- CCC <^ '- <- f^ . < . <^S. b?or *e ' - : *' **f 5fo-.-;^ f< ?^^*gjr *L- *. ^ V '