\g New York State Cullege of Agriculture At Cornell University Ithaca, N. YB. Bibrary Cornell University Library ' QK 160.F5 m | ii 4 001 301 237 Manual of il 192 | | woe MANUAL FLOWERING PLANTS IOWA PART 1. POLYPETALAE. MANUAL OF THE FLOWERING PLANTS OF IOWA BY T. J. FITZPATRICK. Fellow of the Iowa Academy of Sciences A THESIS. PRIVATELY PUBLISHED 1899. Copyright Applied For. PREFACE. Tus volume is the result of continued study of the flowering plants of lowa during a period of seven years. The author has collected in more than thirty counties and in most portions of the state. To the collections thus ob- tained the author’s many friends have made generous additions either by donation or by exchange. So far as possible all the literature relating to Iowa botany has been canvassed and the collections in the State University have been examined. All previous treatises have been in the nature of lists; many annotated, some not; and were mostly confined to the flora of restricted loealities. In this volume all the scattered information is gathered and the author endeavors to correct previous errors. to confirm the observations already made. and to add new information obtained by fersonal effort. This work is founded primarily upon the private herbarium of T. J. and M. F. L. Fitzpatrick and in this connection the author wishes to state that he is under obligations to the followiug parties who have sent many rare or infre- quent and interesting specimens: I'red Reppert of Muscatine county. Prof. B. Fink of Fayette county, E. W. D. Holway and Herbert Goddard of Winne- shiek county. R. I. Cratty of Emmet county. Prof. Pammel, C. R. Ball and Wilmon Newell of Iowa State College, J. P. Anderson of Decatur county, J. fi. Mills of Henry county, and Profs. T. tl. Macbride and B. Shimek of the State University, and to many others who have lent aid in a limited amount. In the determination of difficult material the author has received aid from Prof. Wm. Trelease and Prof. J. B.S. Norton of the Missouri Botanical Gar- den, F. Lamson-Seribner of the U.S. Department of Agriculture, and R. I. Cratty. The question of nomenclature is as yet unsettled. For some reason the new nomenclature has not been received with much favor by Iowa botanists. Be- cause of this the author has thought it best to pursue a conservative course. All needful changes have been adopted and by the use of synonyms both the old and new systems are presented. The author shall be pleased at all times to receive information and material so that he may be able to extend his knowledge of a subject the pursuit of which has been a source of much pleasure. T. J. Firzpatrick. LAMONI, Iowa, July 39, 1899 : MANUAL OF THE FLOWERING PLANTS OF IOWA. ee ae ae ANALYTICAL KEY TO THE FAMILIES. Series 1. PHANEROGAMS or FLOWERING PLantTs: those producing true flowers and seeds. Class 1. ANGlosPERMS: plants which have the ovules contained in a closed ovary. Subclass 1. DicoryLepons: stems differentiated into bark, wood, and pith; leaves netted-veined; embryo with a pair of opposite cotyle- dons; flowers usually 4 or 5-merous. Division 1. PoLyprraLas: calyx and carolla present, petals separate. Apetalous forms occur as will be noted farther on. The following wholly artificial key is taken from various governmental publications and adapted to the Iowa flora. A. Stamens ten to many and more than twice the number of sepats. * Calyx entirely frec from the ovary. Pistils many, distinct, enclosed in a hollow receptacle; leaves alternate, stipulate. Rosa, in RosacEak, 49. Pistils several, contained in pits on the upper surface of a large convex receptacle. Nelumbo, in NyMPHAEACEAE, 7. Pistils more than one, distinct, not enclosed in the receptacle. Stamens inserted on the edge of a disk which lines the calyx-tube, dis- tinct; anthers 2-celled. RosacEAk, 44. Stamens monodelphons, united with the base of the petals; anthers 1- celled. MALVACEAE, 23. Stamens inserted on the receptacle. Small trees; filaments shorter than the anthers. ANONACEAE, 6. Herbs or woody climbers; filaments longer than the anthers. Flowers dicecious; twiners; leaves alternate, palmately lobed. MENISPERMACEAR, 7 Flowers perfect, if climbers the leaves are opposite. Leaves peltate; petals persistent. Srasenia, in NYMPHAEACEAE, 7. Leaves not peltate; petals deciduous. RANUNCULACEAE, 1. Pistils several-lobed, the ovaries united below the middle. RESED4CEAE, 16. Pistils several, their ovaries cohering in a ring around an axis. MALVACEAE, 23. Pistil solitary as to the ovary but styles or stigmas may be several Leaves punctate with pellucid or black dots. HYPERICACEaE, 22. Leaves not punctate with pellucid or black dots. Ovary simple, 1-celled, 2-ovuled; fruit a drupe. Drupaceag, 43. Ovary simple, 1-celled, several-ovuled; fruit a legume. MIMOSACEAE, 43. Ovary compound, 1-celled; placenta central. PoRTULACcACcEag, 21. Ovary simple, 1-celled; placenta parietal, many-ovuled. Leaves 2-3-ternately compound or dissected. RANUNCULACEAE, 1. Leaves peltate, palmately lobed. Podophyllum, in BERBERIDACEAE, 7. Ovary compound, 1-celled; placentae 2 or more, parietal. Sepals caducous; juice milky or colored. PAPAVERACEAE, 9. Sepals deciduous, 4. CAPPARIDACEAE, 16. Sepals persistent, 3 or 5. CISTACEAE, 17. Ovary compound, several-celled. Calyx valvate in the bud, and persistent; stamens monodel- phous; anthers 1-celled. MALVACEAE, 23. Deciduous; anthers 2-celled. TILIACEAE, 25. Calyx imbricated in the bud, persistent. Ovaries on many partitions; aquatics. NYMPHAEACEAE, 7. Ovaries on 5 placentae in the axis. SARRACENIACEAE, 9. ** Calyx more or less cohercnt with the surface of the compound ovary. Ovary 8-30-celled; ovules many, on partitions, aquatic. NYMPHAEACEAE, 7. Ovary apparently 10-celled, each division 1-ovuled. Amelanchier, in POMACEAE, 49. Ovary 2-5-celled. Leaves alternate, stipulate. PoMACEAE, 49. Leaves opposite, exstipulate. SAXIFRAGACEAE, 51. Ovary 1-celled; ovules parietal. Fleshy jointed prickly plants; petals many. CACTACEAE, 49. Rough-leaved plants; petals 5 or 10. LOacgag, 59. Ovary 1-celled; placenta free, central; pericarp opening by a lid. PorRTULACACEAR, 21. B. Stamens of the same number as the petals and opposite them. Pistil solitary; flowers mostly perfect; herbs, shrubs, or woody vines. Ovary 1-celled; anthers opening by valves. BERBERIDACEASE, 7. Ovary t-celled; anthers longitudinally dehiscent. PORTULACACEAE, 21 Ovary 2-4-celled. Calyx-lobes small or wanting; petals valvate. VITACEAR, 29. Calyx 4-5-cleft valvate in the bud; petals involute. RuAMNACEAE, 29. Pistils 3-6, separate; flowers dicecious; woody vines. MENISPERMACEAE, 7. C. Stamens of the same number as the petals «nd alternate with them or not more than twice us many. * Ovary superior, the calyx entirely free. + Ovarics 2 or more, separate. Stamens hypogynous, distinct. Leaves pellucid-functate. RUTACEAE, 27. Leaves not pellucid-punctate. ‘ Tree; leaves odd-pinnate; flowers paniculate. SIMARUBACEAE, 28. Herbs; leaves thick, succulent; flowers cymose. CRASSULACEAE, 53. Herbs; leaves not fleshy; inflorescence various. RANUNCLACEAE, 1. Stamens inserted on the calyx, distinct. Stamens twice as many as the pistils. CRASSULACEAE, 53, Stamens neither the same nor twice the number of the pistils. Leaves stipulate. Rosacear, 44. Leaves exstipulate. SAXIFRAGACEAE, 51. + + Ovaries 2-5, separate above, mvure or less united below, Leaves pellucid-punctate. , RUTACEAE, 27. Leaves not pellucid-punctate. Trees or shrubs; leaves opposite, palmately lobed or pinnate; fruit a samara. ACKERACEAE, 30. A small shrub; leaves opposite, 3-foliolate. STAPIYLEACBAB, 31. + + + Ovary 5-lobed, 5-celled; style compound. JERANIACEAE, 26. tt + + Ovary simple, 1-celled with one parietal placenta. Flowers irregular, the upper petal enclosing the others in the bud. PAPILIONACEAR, 33. Flowers mostly regular, the upper petal enclosed by the lateral in the bud. CAESALPINACEAE, 42. ++4+ + Ovary 1, compound, as shown by the number of cells, placentae, styles, or stigmas. Ovary 1-celled. Corolla irregular, spurred; petals 4; stamens 6. FumARIACEAE, 9. Corolla irregular, lower petal spurred; petals and stamens 5. VIOLACEAR, 17. Corolla regular or nearly so. Ovule 1; stigmas 3; small trees or shrubs with resinous or milky acrid juice. ANACARDIACEAE, 32. Ovules 1 or 2; herbs. A few of the CRUCIFERAE, 10. Ovules 2 or more, central or basal. Petals inserted on the throat of the calyx. LYTHRACEAE, 55. Petals not inserted on the throat of the calyx. CARYOPHYLLACEAE, 19 Ovules several or many, on two or more parietal placentae. Leaves pellucid-punctate or black-dotted. HYPERICACEAE. 22. Leaves non-punctate. Sepals 5, unequal or only 3. CISTACEAE, 17. Sepals and petals 4; stamens 6. CRUCIFERAE, 10. Sepals and petals 5: stamens 5 or 10. SAXIFRAGACEAE, 51. Ovary 2-several-celled; flowers irregular. Stamens 6 or 8; anthers I-celled, opening at the top. PoLYGALACEAR, 21. Stamens 11 or 12, dehiscence longitudinal; petals inserted on the throat of the gibbous or spurred calyx. Cuphea, in Ly THRACEAR, 55. Stamens 5 to 8 or 10; petals mostly hypogynous. Ovary 3-celled; leaves opposite, digitate; trees. HIPPOCASTANACEAE, 31. Ovary 5-celled; leaves alternate, simple; herbs. BALSAMINACEAE, 27. Ovary 2-several-celled; flowers regular. Stamens neither the same nor twice the number of the petals. Petals 5; stamens collected into 3 clusters. HYPERICACEAB, 22. Petals 4; stamens 6, 2 shorter, rarely 2 or 4. CRUCIFERAE, 10. Petals 5; stamens more, distinct. ACERACEAE, 30, Stamens of the same or twice the number of the petals. Ovules and seeds 1 or 2 in each cell. Herbs, with perfect symmetrical flowers. Ovary deeply lobed, 5-celled; sepals 5. GERANIACEAR, 26. Ovary globose, breaking into 5 2-celled carpels, sepals 5. : LINACEAE, 26. Shrubs or trees. Leaves 3-foliolate, pellucid-punctate. RUTACEAE, Leaves palmately lobed and veined; fruit a 2-winged samara. ACERACEAE, 30. io zt Leaves simple, pinnately veined, non punctate. Fruit a globose or lobed pod; seeds arillate. CELASTRACEAE, 28. Fruit a berry-like drupe. ILICINEAE, 29. Ovules and seeds several to many in each cell. Stipules caducous; leaves opposite, compound. STAPHYLEACEAE, 31 Stipules wanting when the leaves are opposite. Stamens 10, monodelphous below; leaflets 3, obcordate. OXALIDACEAR, 27. Stamens distinct, hypogynous. , CARYOPHYLLACEAE, 19. Stamens distinct, perigynous. Styles usually 2, simple. sometimes cohering. SAXIFRAGACEAE, 51. Style 1; capsule often 1-2elled. LYTHURACUHAE, 55, * * Calyz-tube m ore or less adherent to the ovury. Ours climbing annuals, with °-forked tendrils. . CUCURBITACEAE, 59. Mostly erect herbs, not tendri) bearing. Ovules and seeds two to many in each cell of the ovary. Ovary l-celled; ovules many, basal. PORTULACACBAR, 21. Ovary 1-celled; placentae 2 or 3, parietal. SAXIFRAGACEAE, 51. Ovary 2-several-celled. Anthers dehiscing by apical pores; style 1. MELASTOMACEAE, 55. Anthers not dehiscing by apical pores. Stamens inserted on a disk which fills the bottom of the calyx. CELASTRACEAE, 28. Stamens inserted on the calyx. Stamens 4 or 8, rarely 5; style 1. ONAGRACEAR, 56. Stamens 5 or 10; styles 2 or 3, distinct. SAXIFRAGACBAE, 51. Ovules and seeds solitary in each cell of the ovary. Stamens 2 or 8; styles 1; stigma 2-4-lobed; herbs. ONAGRACEAE, 56. Stamens 4 or 8; styles or stigmas 4; aquatics. HALORAGIDACEAR, 54. Stamens 8, but 4 of them sterile, scale-like, styles 2; small tree. HaMAMELIDACEAE, 54, Stamens 4; style and stigma 1; small shrubs. CoRNACEAE, 65. Stamens 5; flowers umbellate, rarely capitate. Fruit dry, consisting of 2 coherent carpels; styles 2. UMBELLIFERAE, 60. Fruit a 2-5-celled drupe; styles 2-5. ARALIACEAE, 64. Gamopetalous forms in polypetalous families. Forms which have their petals more or less united into one piece. * Stamens more numerous than the lobes of the corolla. Ovary 1-celled, with one parietal placenta. The upper petal enclosing the others in the bud. PApILIoNAcCEAR, 33. The upper petal enclosed by the lateral in the bud. CARSALPINACEAB, 42. Ovary 2-celled, 2-ovuled. POLYGALACEAE, 32. Ovary 3-many-celled. Stamens 10, hypogynous, united below; styles5. ONXALIDACEAE, 27. . Stamens many, monodelphous; styles many. MALVACEAE, 23. **® Stamens as iany as the lobes of the ovary. Annuals, climbing, tendril bearing. CUCURBITACEAE, 59, Shrub, with alternate simple |. aves. ILIcINEAE, 29, Apetalous forms in polypetalous families. Forms having the corolla and sometimes the calyx wanting; calyx often petaloid. * Ovary or its cells with many ovules. Ovary inferior, 4-celled; stamens 4. Ludwigia, in ONAGRACEAE, 56, Ovary inferior, 1-celled; stamens usually 8. Chrysosplenium, in SAXIFRAGACEAE, 52. Ovary superior. Pod 5-celled, 5-beaked, the beaks falling away at maturity; stamens 10. Penthorum, in CRAssULACEAE, 53. Pod 3-celled, 3-valved, many-seeded. AIZOACEAE, 60. Pod 1 or 2-celled; placentae central. Stamens inserted on the calyx. LYTHRACEAE, 55. Stamens hypogynous or inserted at the base of the calyx. CARYOPHYLLACEAE, 19. Pod 1-celled, with one parietal placenta. RANUNCULACEAE, 1. Pods 2 or more, separate, simple. RANUNCULACEAE, 1. * * Ovary or its cells usually with 1 or 2 ovules. Pistils 2 or more, distinct or nearly so. Stamens hypogynous; leaves punctate. RUTACEAE, 27. Stamens hypogynous; leaves not punctate; calyx usually petaloid. RANUNCULACEAE, 1. Pistil 1, simple or compound. Ovary inferior, 3 or 4-celled; aquatics. HALORAGIDACEAE, 54. Ovary naked; calyx none. Callitriche, in HALORAGIDACEAE, 55. Ovary free from the calyx; trees or shrubs. Ovary 2-celled; fruit a 2-winged samara. ACERACEAE, 30. Ovary 3-celled; fruit drupaceous or dry. RHAMNACEAE, 29. , RANUNCULACEAE Juss. Crowroot FAmity. Annual or perennial herbs, with colorless usually acrid juice; and poly- petalous or apetalous, regular or irregular flowers. Leaves simple diverging by the beaks into nearly separate follicles. Moist meadows and woods; May-June; fre- quent; Winneshiek, Allamakee, Fayette, Scott, Linn, Johnson, and Cerro Gordo counties. SULLIVANTIA T & G. Perennials, with lobed or eut-toothed petioled rounded leaves, and small white flowers in a loose cymose-panicle. Calyx campanulate, adherent to the base of the ovary, 5-cleft. Petals 5, oblanceo- late, twice the length of the calyx. Stamens 5, included. Capsule 2-celled, 2-beaked, with many wing-margined seeds. S.ohionis T. & G. Stem low, 4-10 inches high, weak, nearly leafless; leaves palmately veined. Limestone cliffs; May-August; infrequent; Jones, Delaware, Jackson, and Cedarcounties. (Sullivantia sullivantia (T. & G.) Brit.) MITELLA lL. Perennials, with alternate radical leaves and opposite cau- line leaves, and small flowers in a slender spike. Calyx 5-cleft, short, adnate to the base of the ovary. Petals5, pinnatifid. Stamens 10, included. Styles 2. short. Capsule 1-celled, 2-beaked, opening by two equal valves. M. diphylla L. Bishop’s cap or mitre-wort. Stem hairy, 8-16 inches high; leaves cordate acute, dentate, sublobate; radical ledves long-petioled, cauline leaves 2; very Short-petioled; flowers white. Rich woods; May-June; frequent; Winneshiek. Fayette, Scott. Muscatine, Henry, and Johnson counties. HEUCHERA L. Perennials, with long-petioled radical leaves, and green- ish or purplish flowers in small clusters, paniculate. Calyx campanulate, 5- cleft, segments obtuse, tube adnate to the base of the ovary. Petals 5, small, spatulate, entire, inserted with the 5 stamenson the throat of the calyx. Styles 2. Capsule !-celled, 2-beaked, opening between the beaks. H. hispida Pursh. Stem 2-4 feet high; stem, petioles, and veins hirsute with long spreading hairs; leaves rounded, cordate at the base, 5-9-lobed, toothed; flowers greenish, oblique; stamens exserted. Prairies; May-June; common. . CHRYSOSPLENIUM L. Stems low. Calyx 4~5-lobed, yellow inside, ad- nate to the ovary. Petals wanting. Stamens 8-10, short. Styles2. Capsule obecordate, 1-celled, 2-valved, many-seeded. C. alternifolium L. Golden Suxifrage. Found ina deep ravine among moss; Winneshiek county, the only locality in the state. PARNASSIA L. Scape-like perennials. with solitary terminal flowers. Calyx deeply 5-parted, imbricated in the bud, persistent. Petals 5, spreading, white, imbricated in the bud. Tertile stamens 5, alternate with the petals, sterile stamens in clusters at the base of the petals. Stigmas 4, sessile. Ova- ry 1-celled, with 4 parietal placenta. Seeds winged. P.carcliniana Mx. Scapes glabrous, 10-20 inches high, with one sessile leaf near the base; radical leaves petioled, round-oval; sterile stamens in each cluster 3, gland-tipped; petals ovate, obtuse, greenish veined, 3-4 times the length of the sepals. Wet meadows and river banks; June-August; infre- quent; Emmet, Fayette, Muscatine, Johnson, and Story counties. HYDRANGEA L. Shrubs, with opposite petioled exstipulate leaves, and corymbose often apetalous flowers. Calyx-tube hemispheric, adnate to the ovary, 8-10-ribbed, 4-5-lobed. Petals 4-5. Stamens 8-10, inserted on the disk. Styles 2, diverging. Ovary 2-4-celled. Capsule membranous, 2-celled, many- sceded, about 15-ribbed. GOOSEBERRY FAMILY. 53 H. arborescens L. Wild Hydrangea. A shrub, 4-10 feet high, with ovate pointed serrate leaves, and mostly fertile flowers in cymes. Reported from Lee county. GROSSULARIACEAE Pumort. GoosEBERRY FAMILY. Low shrubs, with alternate palmately-lobed single or clustered leaves, and solitary or racemose flowers. Calyx 5-lobed, adherent to the ovary, withering persistent. Petals 5, small. Stamens 5, alternate with the petals, inserted with the petals on the top of the calyx-tube. Styles 2, distinct or united. Ovary 1-celled, with 2-parietal placente. Fruit a berry. Represented by the genus Rises L. * Pedurnecles axillary, solitary, 1-3-flowered. R. gracile Mx. Cummon Wild Gooseberry. Stems recurved, often with long and stout red thorns; leaves rounded; lobed and incised; flowers whitish; calyx-tube short and narrow; calyx-lobes oblong-linear, filaments slender, connivent or parallel, much exserted; berry smooth. Woods; May; common. R. cynosbati L. Prickly G. Ueaves similar to the preceding, pubescent; calyx campanulate, tube longer than the lobes, stamens and style not ex- serted, styles united; berry armed with long spines. Woods; May; frequent; Winneshiek, Clayton, Dubuque; Fayette, Delaware, Scott, Muscatine, Henry, Johnson, Story, Dallas, Webster, and Cerro Gordo counties. R. oxyacanthoides L. Flowers greenish or purplish, peduncles short; stamens not exceeding the oblong calyx-lobes. Hillsides; April-May; re- ported as frequent from Delaware and Henry counties. ** Flowers in drooping racemes. R. floridum L’Her. Wild Black Currant. Stems without thorns or prickles; leaves heart-shaped, 3-5-lobed, toothed; flowers large, whitish; bracts longer than the pedicels; stamens and style short; berry smooth, black. Woods; May; frequent; Winneshiek, Allamakee, Clayton, Delaware, Seott, Muscatine, Fayette, Johnsoa, Emmett, Story, Winnebago, Floyd, Cerro Gordo, Lyon, Woodbury, and Taylor counties. R. rubrum L. Red Currant. Fruit globose, red. This species is the com- mon garden currant, occasionally an escape. R. aureum Pursh. Golden, Buffalo, or Missouri Currant. Shrub 5-12 feet high, with 3-5-lobed, leaves, and showy golden-yellow flowers in short racemes. Frequent in cultivation, occasionally spontaneous; April-May. CRASSULACEAE DC. Orpine FamILy. Herbs with cymose flowers. Sepals and pistals of the same number. Petals as many as the sepals or wanting. Stamens twice as many as the sepals. Fruit.capsular, dehiscent. PENTHORUM. Plants not succulent; carpels united. SrepuM. Plants very succulent; carpels distinct. PENTHORUM L. Perennials, with leafy stems, and yellowish green apetalous flowers. Sepals and pistils 5. Stamens 10. Fruit of 5 one-celled carpels, united at the base, each horned above. Seeds small, many. P. sedoides L. Ditch Stonccrop. Stem 1-3 feet high; leaves scattered, sessile, lanceolate, tapering both ways, serrate. Common in wet soil; July— September. SEDUM L. Leaves thick, succulent, sepals, petals, and pistils 4-5. Stamens 8-10. Carpels distinct, many-seeded. S. telephium L. Orpine.’ Live-forever. A glabrous somewhat glaucous od HALORAGIDACEAE perennial with ovate to obovate, obtuse, coarsely dentate leaves, and purple flowers in a dense cyme. A rare garden escape observed in Muscatine and Johnson counties. HAMAMELIDACEAE Z/udl. Witon Haze, FaMizy. Small trees, with alternate simple petioled pinnately-veined leaves, deciduous stipules, and axillary clustered or capitate flowers. Calyx superior, imbricated in the bud, deciduous Petals inserted on the calyx, valvate in the bud, deciduous. Ovary 2-celled. Styles 2, distinct. Repre- sented with us by the genus HaMAMELIs L. H. virginiana L. Witch-huzel. Tree 10-20 feet high; leaves oval or obovate, wayvy-toothed; flowers yellow; calyx 4-parted, with 3 scale-like leaves im- mediately below; petals 4, long-linear; stamens 8, 4 fertile and alternate with the petals, the remainder sterile; fruit a 2-celled woody capsule, opening at the top, cells 1-seeded; seeds pendulous. Wooded ravines; flowers appearing in October and the fruit the following July; infrequent; Clayton, Dubuque, and Delaware counties. HALORAGIDACEAE Al. & Garcke. WATER-MILLFOIL FAMILY. Marsh or aquatic plants. Leaves alternate, opposite or whorled, entire or toothed; submerged leaves pinnately divided into narrow segments. Flowers solitary, fascicled or whorled, sessile. Calyx su- perior; tube coherent with the ovary. Petals small, inserted on the calyx, alternate with its segments. Calyx and corolla sometimes want- ing. Stamens 1-8, inserted with the petals. Ovary inferior, with 2-4 1-ovuled cells. MYRIOPHYLLUM. Leaves whorled; flowers moncecious or polygamous, the parts in fours. PROSERPINACA. Leaves alternate; flowers perfect, the parts in threes. CALLITRICA. Leaves opposite; flowers monvecious; sepals and petals wanting. MYRIOPHYLLUM L. Waver Miuirort. Perennial aquatics. Leaves crowded or whorled, the submerged dissected into capillary divisions. Flow- ers moncecious or polygamous in the axils of the upper leaves; the upper stam- inate. Stamens 4-8. Fruit nut-like, 4-lobed, 4-celled. M. scabratum Mx. Stem slender; leaves in whorls of + or 5, lower pin- nately parted, floral linear, toothed or serrate; stamens 4. Shallow lakes; July; frequent; Muscatine, Appanoose, Ringgold, and Fremont counties. (M. pinnatum (Walt.) B. 8. P.) M. heterophylium Mx. Stem stout; leaves in 4’s and 5's: floral leaves ovate or lanceolate, crowded, serrate, stamens 4. Lakes; July; Winnebago, Cerro Gordo, and O’Brien counties. M. spicatum L. Leaves mostly pinnately-parted, capillary; floral leaves small, shorter than the flowers, ovate, entire or toothed; flowers in an inter- rupted spike; petals deciduous; stamens 8. Ponds; July; Muscatine,- Cerro Gordo, Story, and Dickinson counties. PROSERPINACA L. Perennial aquatics, with alternate lanceolate leaves, and perfect small sessile axillary Howers. Calyx 3-sided, adnate to the ovary, limb 3-parted, Petals wanting. Stamens 3-4. Stigmas 3, cylindrical. Fruit bony, 3-4-angled, 3-4-celled, 43~4-seeded. P, palustris L. Jlermaid weed, Glabrous, mostly simple; emersed leaves oblong or linear-lanceolate, sharply serrate; the immersed leaves pectinate or pectinate-pinnatifid, the segments linear, stiff, acute, often serrulate, flowers LoosESTRIFE FAMILY. 55 solitary cr several together; fruit sharply angled. Ponds; July; rather rare; Museatine, Story, and Boone counties. CALLITRICHE L_ Leaves spatulate or linear, opposite. Flowers monc- cious, 1-3 together, axillary, sessile. Sepals and petals wanting. Sterile flowers with one stamen; anther cordate, 4-celled, becoming 1-celled. Ovary 4celled. Styles 2, distinct. Fruit nut-like, 4-lobed, 4-celled, membranous, indehiscent. C. vernal. Perennial; floating leaves tufted, obovate, 3-nerved, narrowed into a petiole; submerged leaves linear; flowers usually between a pair of bracts; fruit ovate, styles shorter than the fruit. In a slough near Cedar riv- er, Museatine county; June-September. (C. palustris L.) C. hetercphylla Pursh. Similar to the preceding species; floating leaves broadly spatulate, often retuse, narrowed into a petiole; fruit obovate; styles longer than the fruit. Wet slough; July; infrequent; Decatur county. MELASTOMACEAE FR. Br. MEADow-BEauvty FAMILY. Herbs with opposite parallel-veined leaves and perfect regular cymose flowers. Petals inserted on the calyx-throat, alternate with the calyx- lobes, stamens inserted with the petals, double their number; authers opening by pores at the apex. Represented with us by the genus RExIA L. R. virginica L. Meadow-Beruty. Stem 1-2 feet high, square, angles some- what winged, rootstock tuberiferous: leaves with 3-5 nerves, sessile, acute, oval-lanceolate, nerves and margin more of less bristly; flowers peduncled, bright, purple; calyx persistent 4-cleft, lower portion of the tube coherent with the ovary, the upper free; petals 4; stamens 8; anthers long-linear, 1- celled, with a short spur near the filament; fruit a capsule, 4-celled, with 4 placentze from the central axis, many-seeded. In swampy places along Cedar river; July; Muscatine county. LYTHRACEAE JZ/ndl. LoosestRire FaMiy. Herbs, with mostly opposite entire exstipulate leaves and axillary or whorled flowers. Calyx free from the ovary but enclosing it, persistent, 4-7-toothed. Petals inserted on the top of the calyx-throat, equal in number to the calyx-teeth, deciduous. Stamens 4-14, inserted on the calyx-tube. Style 1; stigma capitate. Capsule 2-4-celled, with axial placente, many-seeded. * Flowers regular. Lyrarum. Flowers rarely solitary ;.calyx tubular; petals usually 6. AMMANNIA. Flowers solitary; calyx globular; petals and stamens usually 4. RotaLa. Flowers solitary; sinuses of the calyx appendaged; petals and stamens 4. DipipLis. Flowers solitary; petals none; stamens 4. ** Flowers irregular. CupHEa. Calyx more or less spurred; petals 6; stamens H or 12. LYTHRUM L. Leaves opposite or scattered, sessile. Flowers purple, | solitary, axillary. Calyx tubular, striated, 5-toothed, with appendages in the sinuses. Petals 5-7. Stamens 5-14. Capsule oblong, 2-celled. L. alatum Pursh. Wing-cngled Loosestrife. Perennial, smooth; stem 1-3 feet high, with many long slender branches; leaves oblong-ovate to linear- lanceolate, acute, base rounded or cordate, the upper mostly alternate; petals deep purple; stamens of the short styled flowers exserted. Rich soil; July- August; common. AMMANNIA L. Low smooth herbs, with opposite narrow leaves, and sub- 56 ONAGRACEAE. sessile cymose axillary flowers. Calyx globular, 4-angled, 4-toothed; sinuses with horn-like appendages. Petals 4, purplish, deciduous. Stamens 4-8. Capsule globular 2-4-celled. A. coccinea Rottb. Stem 4-8 inches high, branched from below; leaves linear-lanceolate; stamens more or less exserted; style slender, elongated. Swampy places, shores; July-September; infrequent; Muscatine, Louisa, Johnson, Decatur, Story, and Lyon counties. = : ROTALA L. Low herbs. Calyx semiglobose or campanulate; sinuses with tooth-like appendages. Petals and stamens 4. Capsule globular, 4- celled. R. ramosior (L.) Koehne. Stem 5-6 inches high, smooth, simple; leaves linear-oblanceolate, opposite, tapering to a petiole; flowers solitary, axillary, sessile; appendages shorter or as long as the calyx-teeth. Low wet places; July-September; infrequent; Muscatine, Henry, Benton, and Lyon counties. DIDIPLIS Raf. Aquatic or marsh plants, with opposite leaves, 4-angled stems, and mostly solitary axillary flowers. Calyx 4-lobed, appendages none. Petals wanting. Stamens usually 4, inserted on the calyx-tube. Capsule globose, 2-celled. D. linearis Raf. A low glabrous much branched plant, 2-6 inches long; leaves long-linear or lanceolate, mostly sessile; flowers small, greenish; calyx- lobes broad, triangular; style short; capsule small. Mud flats, shallow water; June-August; infrequent; Devatur and Ringgold counties. (D. diandra (Nutt.) Wood.) CUPHEA P. Br. Herbs, with opposite or verticillate leaves, and axillary irregular unsymmetrical flowers. Calyx tubular, gibbous or spurred at the ‘base, apex 6-toothed, with 6 appendages, mouth oblique. Petals 6, unequal. Stamens 11 or 12, inserted on the throat of the calyx, unequal. Capsule ob- long, included, 1-celled, dehiscent. C. viscosissima Jacq. An erect viscid-pubescent annual, 6-20 inches high, branched; leaves ovate-lanceolate, slender petioled, apex bluntish, base rounded; flowers purple, axillary, short-peduncled; petals ovate, clawed. Dry soil; July-September; reported as belonging to the flora of Iowa by Arthur. (Parsonia petiolata (L.) Rusby.) ONAGRACEAE Dumort. EVENING-PRIMROSE FAMILY. Herbs, with alternate or opposite leaves, and perfect symmetrical flowers. Calyx herbaceous or colored, of 4 lobes, rarely less, valvate in the bud, tube cohering with the ovary. Stamens as many or twice as many as the petals and inserted with them on the top of the calyx- throat; anthers 2-celled, introrse; pollen grains often cohering by threads. Ovary inferior, 4-celled. Fruit a 2-4-celled many-seeced capsule. * Parts of the flower in fours. + Culyx-tube not prolonged beyond the summit of the ovury; fruit dehisecnt. Lupwiacia. Leaves opposite or alternate, stamens 4; capsule short. + + Calyx-tube prolonged beyond the summit of the ovary; fruit dehiscent. EPLLOBIUM. Leaves often opposite; seeds silky tufted. OENOTHERA. Leaves alternate; seeds naked. t+ + + Fruit indehiscent. GauRa. Filaments appendaged at the base; fruit indehiscent. ** Parts of the flower in twos. CiRcaEA, Sepals, petals, and stamens 2; fruit bristly. EVENING-PRIMROSE FAMILY. 57 LUDWIGIA L. Perennial herbs, with opposite or alternate leaves, and solitary axillary flowers. Calyx-lobes, petals, and stamens 4, or petals want- ing. Capsule short, many-seeded. L. alternifolia L. Secd-box. Stem 2-3 feet high, branched, smoothish slightly wing-angled; leaves lanceolate to linear-lanceolate, alternate, ses- sile; flowers short peduncled; calyx-lobes ovate, acute, reddish; petals yel- low; capsule cubical, wing-angled. Damp soil; June-August; rare; Johnson, and Decatur counties. L. polycarpa Short & Peter. Stem 2-4 feet high, glabrous; leaves nar- rowly lanceolate, alternate, sessile, tapering both ways, those of the runners oblong spatulate; flowers small, sessile; capsule 4-sided, longer than the calyx-lobes, ,;with 2-linear awl-shaped bractlets from the base. Borders of ponds aud in shallow water; July-September; frequent; Fayette, Scott, Muscatine, Johnson, Story, Appanoose, Decatur, Ringgold, and Taylor coun- ties. L. palustris (L.) Ell. Marsh or Water Purslane. Stem smooth, creeping or floating; leaves ovate or oval, petioled, opposite; calyx-lobes short; petals small or wanting, reddish in terrestrial forms; capsule oblong, 4-sided, ses- sile. Wet places; July-September; infrequent; Fayette, Muscatine, Johnson, Benton, and Emmet counties. (Isnardia palustris L.) EPILOBIUM L. Mostly perennials. (Leaves nearly sessile, opposite or seattered. Flowers violet, purple, or white. Calyx-tube scarcely prolonged beyond the ovary; limb 4-5-lobed, deciduous. Petals 4. Stamens 8, with short anthers. Capsule linear, many-seeded. Seeds with a tuft of long hairs at the end, the coma. ; E. angustifolium L. Great Willow-herb. Fire-weed. Stem simple, 4-6 feet, high, glabrous of finely pubescent; leaves scattered, mostly entire, narrowly lanceolate, tapering both ways; flowers large, purple, in a long raceme; calyx-limb deeply parted; petals entire; lobes of the stigma long. Rich woods; June-September; frequent; Dubuque, Scott, Johnson, Winnehago, and Shelby counties. (Chamaenerion angustifolium (L.) Seop.) E. coloratum Muhl. Stem 1-3 feet high, much branched, 4-angled, hoary pubescent; leaves lanceolate, contracted to slender petioles or sessile, acuminate, sharply serrulate or denticulate, lower opposite; flowers small, pale, somewhat nodding, short-peduncled; coma reddish-brown. Low grounds; July-September: common. E. adenocaulon Haussk. This species resembles the preceding; leaves lanceolate or ovate-lanceolate, somewhat serrulate or denticulate; inflores- cence and capsules glandular-pubescent; coma white. Wet places; July- September; frequent; Johnson, Story, Hamilton, Floyd, Hancock, Dickinson, and Emmet counties. E. lineare Muhl. Stems somewhat branched, mintely hoary-pubescent; Jeaves numerous, linear or linear-lanceolate, short-petioled; fowers many, small, pale; capsules hoary; coma dingy. Wet places; July-September; iu- frequent; Fayette, Scott, Story, Hamilton, and Emmet counties. OENOTHERA L. Evenine Primrose. Leaves alternate. Calyx-tube pro- longed beyond the ovary, deciduous, limb of 4 reflexed lobes. Petals 4, yel- low in our species. Stamens 8; anthers linear, versatile. OE. biennis L. Stem 1-5 feet high, sometimes branched, pubescent or hairy; leaves lanceolate or ovate, tapering both ways, acuminate, remotely toothed, the lowest petioled; flowers in a leafy spike; calyx-tube long; cap- 58 ONAGRACEAE. sules oblong, pubescent or hirsute, sessile, seeds prismatic-angled. Fields and waste places; July-September; common. (Onagru bicnnis (L.) Seop.) OE. rhombipetala Nutt. Stem 1-4 feet high; simple, appressed pubes- cent; leaves narrowly lanceolate to linear, entire or denticulate, the lowest ‘tapering to a petiole; flowers in a leafy or bracted spike; calyx silky canes- cent; petals rhombiec-ovate; capsules linear; seeds obovoid, not angled. Sandy soil, prairies; July-September; Muscatine county, reported from Fayette, Story, and other counties, often confused with the preceding. OE. fruticosa L Sundrops. Biennial or perennial; stem 1-3 feet high, branched, villous to nearly glabrous; leaves oblong-lanceolate, entire or den- ticulate: flowers in a loose corymb; capsule short-pedicelled, ribbed and ‘winged. Low grounds; June~August; infrequent; Muscatine, Johnson, and Henry counties. (Kneiffia fruticosa (L.) Raimann.) OE. serrulata Nutt. Stem 6-16 inches high, perennial, woody below, canescent or glabrous; leaves mostly linear or lanceolate, irregularily and ‘sharply denticulate; flowers axillary; calyx-tube funnelform, nerved; petals obovate, crenulate. Dry soil, pastures and waysides; June-July; frequent; Winneshiek; Allamakee, Fayette, Story, Shelby, Fremont, Cerro Gordo, Han- cock, Diekinson, Ida, Emmet, Sioux, Woodbury. and Lyon counties. (Meriolix serrulata (Nutt.) Walp.) GAURA L. Leaves alternate, sessile. Flowers white or rose-color, in spikes or racemes. Calyx-tube much prolonged beyond the oyary, deciduous; calyx-lobes 4, reflexed. Petals 4, unequal, clawed. Stamens usually 8. Stigma 4-lobed. Fruit nut-like, indehiscent, 1-celled, 1-4-seeded. G. biennis L. Stem 3-6 feet high, much branched, villous or downy-pube- scent; leaves oblong-lanceolate or lanceolate, remotely denticulate; flowers sessile, white, turning pink, in numerous slender spikes; fruit sessile, 4- ribbed, oval or oblong, villous pubescent. Fields and waste places; July— September; common. G. coccinea Pursh. Stem 6-20 inches high, erect or ascending, much branched, canescent, pubescent or glabrate; leaves oblong-lanceolate to linear-oblong, entire, repand or denticulate; flowers sessile, red, turning scarlet; fruit sessile, terete below, 4-sided above, canescent. Dry soil, ‘prairies; May—August; frequent; Woodbury, Harrison, Munona, and Fremont counties. G. parviflora Dougl. Stem 2-6 feet high, villous-pubescent; leaves ovate- lanceolate, acuminate, repand-denticulate, base narrowed, softly pubescent: flowers many, pink, sessile, in an elongated spike; fruit sessile, 4-nerved, obtusely 4-angled, glabrous or nearly so. Meadows and fields; May-August; frequent; Woodbury, Harrison, Pottawattomie. and Fremont counties. CIRCAEA L. Perennials, with opposite petioled thin leaves, and small whitish racemose flowers. Calyx-tube slightly prolonged, the end filled by a cup-shaped disk, deciduous. Petals 2, obeordate. Stamens 2. Fruit small, bur-like, indehiscent, clothed with white hooked hairs. Cc. lutetiana L. Stem simple, 1-2 feet high; leaves ovate, long-petioled, base rounded, acuminate. entire or slightly toothed; bracts none: fruit 2- celled, slender pedicelled, bristly. Rieh woods; June-August; common. C. alpina L. Stem weak, 5-0 inches high; leaves cordate, thin, shining, coarsely toothed; bracts minute; fruit 1-celled, obovate-oblong, soft hairy. Rich woods; June-August; infrequent; Winneshiek, Clayton, and Muscatine ‘counties. Gourp FAMILY. 59 LOASACEAE Peichen) Loasa FAMIny. Erect herbs, with stinging or viscid pubescence, alternate or opposite exstipulate leaves, and regular perfect solitary or cymose flowers. Calyx- tube adnate to the ovary, limb 4-5-lobed, persistent. Petals 4-10, inserted on the throat of the calyx. Stamens many, inserted with the petals. Ovary 1-celled, with 1-3 parietal placentas. Capsule 1-celled. Represented in our flora by the genus MENTZELIA L. M. ornata T. & G. Stem stout 1-2 feet high, roughish-pubescent; leaves sessile, or lower petioled, oval to lanceolate or oblong, sinuate-pinnatifid, flowers usually solitary, terminal, yellowish white, opening in the evening, petals 10, twice as long as the calyx-lobes; capsules oblong. Sandy and rocky bluffs along the Big Sioux river, Cedar Bluffs; June-September; com- mon; Woodbury county. (Pammel) (Af. decupetalu (Pursh.) Urban & Gilg. } CUCURBITACEAE B. Juss. Gourp Famtry. Climbing or trailing succulent herbs with tendrils. Leaves alternate, petioled, palmately lobed, palmately veined. Flowers moncecious or dioecious, axillary, sometimes gamopetalous. Calyx usually campanulate, 5-toothed or 5-lobed, imbricated in the bud. Corolla mouopetalous, rotate or campanulate. Stamens 5 or 3, of which one is usually 1-celled. Stigmas 2 or 3. Ovary inferior, l-cclled, with 3 parietal placentas. Rep- resented in cultivation by the gourd, pumpkin, melon, squash, cucum- ber, etc. EcuHINocysTis. Fruit echinate, 4-seeded. Sicyos. Fruit prickly, 1-seeded. ECHINOCYSTIS T. & G. Annuals with 3 forked tendrils, climbing high. Flowers moncecious, small, greenish white; the sterile in a long compound raceme, the fertile solitary or in small clusters; petals 6; anthers 3, more or less united; stigmas 2; fruit fleshy, inflated, becoming dry, bursting above, clothed with weak prickles, 4-seeded. E. lobata T& G. Wild Cucumber. Leaves broad, thin, sharply and deeply 5-lobed; fruit oval, spines nearly half an inch long; seeds black. Low grounds along waterways: July-September; common. (Micrampelis lobata (Mx.) Greene.) SICYOS L. Anouals with 3 forked tendrils, climbing high. Flowers moncecious, from the same axil, small, whitish. Petals 5, united below. Anthers cohering ina mass. Style slender, with 3 stigmas. Ovary 1-celled, with a single suspended ovule. [Fruit ovate, dry, indehiscent, filled by the seed, spiny. S. angulatus L. One-sceded Bur-Cucumber. Clammy hairy; leaves roundish cordate, with 5 pointed angles or lobes. Rich soil, along rivers and in waste places; July-Septeraber; frequent; Scott, Muscatine, Louisa, Des Moines, and Dallas counties. CACTACEAE JZindi. Cactus Famity. Ours green and fleshy plants, with flattened and joined stems armed with spines and prickles, and solitary sessile flowers. Sepals, petals, and stamens numerous, their bases cohering and adnate to the inferior 1- celled many-ovuled ovary. Fruit a l-celled berry. Represented in our flora by the genus Opuntia Mill. 60 UMBELLIFERAE. O. rafinesquii Engelm. Prickly Pear. Prostrate, green; joints obovate to suborbicular or oval; leaves spreading, subulate; spines axillary, 1-5 together; prickles tufted, axillary, reddish brown; flowers large, yellow, with a reddish center; stamens shorter than the petals; stigmas 4-10; fruit fieshy, edible. Sandy soil; June-July; infrequent; Muscatine and Lyon counties. (O. humi- fusa Raf.) O. fragilis (Nutt.) Haw. Prostrate or decumbent; joints ovate, nearly terete or flattish; leaves small, reddish; flowers yellow, smaller than those of the preceding; fruit dry at maturity. Dry soil; July-August; common; Lyon county. AIZOACEAE A. Br. CARpPEt-weEED FAMILY. Mostly prostrate and branching herbs, with opposite or verticillate leaves, and small regular perfect flowers, either solitary, cymose or glomerate. Calyx of 5 sepals, free from the ovary. Petals wanting. Represented in our flora by the genus Mo.iueo L. M. verticillata L. Carpet-weed. Prostrate, much branched, annual, form- ing patches; leaves spatulate, whorled at the joints; pedicels slender, 1-flow- ered, clustered with the leaves; stamens mostly 3; stigmas 3; capsule 3-celled, 3-valved; seeds reniform. Sandy shores, waste places; July-September; common. This species is included by different botanists under FicoIpEAE or CARYOPHYLLACEAE. UMBELLIFERAE B. Juss. Carnot FAMILy. Herbs, with usually hollow stems, alternate compound or simple leaves, sheathing petioles, and small flowers in compound umbels, rarely in heads. Calyx adherent to the ovary, limb obsolete or 5-toothed. Petals and stamens 5, inserted on a fleshy disk which covers the ovary. Styles 2, distinct or united at their thickened bases. Ovary 2-celled, 2-ovuled. Fruit dry, consisting of 2 coherent carpels. A well defined but difficult order. * Flowers white. + Fruit flattened dorsally. Daucus. Leaves pinnately decompound; calyx 5-toothed; fruit prickly along the ribs. CONIOSELINUM. Leaves 2-3-pinnately decompound; fruit winged on the lateral lobes. TIEDEMANNIa. Leaves simply pinnate. HERACLEUM. Leaves large, ternately compound; flowers large. + + Fruit flattened luteratly. Sium. Leaves once pinnate; ribs prominent, equal, corky. Carum. Leaves decompound; fruit ovoid or oval; ribs filiform, inconspicuous. Cicuta, Leaves pinnately compound, the veins running to the notches; ribs flattish, corky. CONIuM. Stems spotted; leaves tripinnately dissected; ribs wavy. : ‘++ + Fruit linear-oblong, slightly flattened laterally. CRYPTOTAENIA. Leaves 3-foliolate; fruit glabrous; ribs obtuse, equal. BrEruLa. Aquatics; leaves simply pinnate; fruit globrous, nearly galbose; ribs incon spicuous. CHAEROPHYLLUM. Leaves ternately decompound; fruit glabrous. OSMORRHIZA. Leaves biternately divided; fruit bristly. + + + + Fruit ovate or obovate. ERYNGIUM. Leaves parallel-veined, coriaceous, bristly-toothed; flowers in dense bracted globose heads, . ** Flowers yellow. + Fruit ylobular, armed. with hooked prickles. SANICULA. Leaves palmately divided; flowers greenish yellow. Cartnor FAMILY. 61 t + Fruit flattened dorsally; stightly in Thaspium. PasTINACA. Leaves pinnately compound; dorsal ribs fililorm, the lateral extending into wings, POLYTAENIA. Leaves 2-pinnate; fruit with the lateral ribs thick corky, dorsal ribs ob- scure. THasPiuM. Leaves 2 3-ternate; fruit oblong, with 5-7 wings, slightly flattened. t+ + Fruit flattened laterally. FOENICULUM. Leaves dissected into filiform segments; ribs prominent. PIMPINELLA. Leaves ternately compound; ribs slender. Zizis. Leaves 2-8-ternate; fruit oblong, with filiform ribs; stylopodium wanting. DAUCUS L. Biennials, with bristly stems, and pinnately decompound leaves. Involucre of foliaceous pinnately parted bracts. Involucels of many entire or toothed bracts. Flowers white. umbels compound, concave. Calyx ‘teeth obsolete. Petals 5, the 2 outer often larger and deeply 2-cleft. Fruit oblong, dorsally, flattened, with 5 bristly primary ribs and 4 winged second- ary ones. Stylopodium depressed. D. carrota L. The common cultivated carrot, frequently found wild in cultivated fields and waste places; July-September; Dubuque, Fayette, Story, Appanoose, and Decatur counties. CONIOSELINUM Hoffm. An erect glabrous perennial, with pinnately decompound leaves, involucre none or of few bracts, involucels of narrow linear bracts. and compound umbels of white flowers. Calyx-teeth none. Fruit oval or oblong, flattened, glabrous; dorsal ribs large, lateral broadly winged; oil-tubes usually’? or 3 in the intervals. Stylopodium slightly coni- eal. C. canadense T. & G. Stem striate, 2-5 feet high; leaves decompound into linear oblong segments. Reported from Scott and Story counties. TIEDEMANNIA DC. Glabrous perennials. Involucre and involucels present. Flowers white. Calyx 5-toothed. Fruit elliptic-ovate, dorsally compressed, glabrous; ribs equidistant, obtuse, the lateral ones broadly winged; oil-tubes solitary in the intervals and 2-6 on the inner face. Stylo- podium short, thick, conical. T. rigida (L.) Coult. & Rose. Whole plant smooth; stem 2-6 feet high, terete, striate, hollow, simple or sparingly branched; leaves all simply pseudo-pinnate; common petiole channeled, somewhat margined; leaflets lanceolate to linear, often slightly falcate, sessile, entire or remotely toothed toward the apex; umbels terminal and lateral. Sloughs and uplands; August; common; Johnson, Benton, Jasper, Story, Decatur, and Ringgold eounties. (Oxypolis rigidus (L.) Britton ) HERACLEUM L. Perennials, with ternately compound leaves, and white flowers in large umbels. Involucre of few bracts, deciduous. Inyolucels of small bracts. Calyx-teeth smal] or obsolete. Petals obcordate, the 2 outer often larger and 2-cleft. Fruit obovate, flattened, somewhat pubescent; dorsal ribs 4, obtuse, the lateral broadly winged. Stylopodium thick, conic. H. lanatum Mx. Cow-Pursnip. Pubescent or woolly, 4-8 feet high; petioles much dilated; leaflets 3, petiolulate, round cordate, about 3-lobed; irregularly cut-toothed. Moist woods; June-July; frequent and generally distributed; Winneshiek, Allamakee, Fayette, Story, Winnebago, Webster, Calhoun, Cerro Gordo, Decatur, Union, Pottawattomie, Shelby, Woodbury, and Emmet counties. , SIUM L. Smooth perennials, with simply pinnate leaves, the lower often pinnatisected, and compound umbels of white flowers. Involucre and involu- cels of numerous uarrow bracts. Calyx-teeth minute. Fruit glabrous, ovate 62 UMBELLIFERAE. to oblong; ribs prominent, corky, nearly equal, oil-tubes 1-3 in the intervals. Stylopodium depressed. S. cicutaefolium Gmel. Stem 2-6 feet high, striate; leaflets 3-8 pairs, linear to lanceolate, sharply serrate, the submerged leaves finely dissected; rays numerous. Sloughs and swamps; July-September; frequent. CARUM L. Ours a glabrous biennial or perennial, with pinnatisected leaves, divisions filiform, and small white flowers in terminal compound um- bels. Calyx-teeth minute. Petals inflexed at the apex. Fruit ovate or ob- long, glabrous, slightly compressed. Stylopodium conical. C. carui L Curaway. Stem erect, 1-2 feet high, branching; lower leaves long-petioled, the upper sessile or nearly so, bases of the petioles dilated. Introduced, waste places; May-July; Emmet county, reported from Fayette county. CICUTA L. Smooth poisonous perennials. Leaves pinnately compound; leaflets serrate. Involucre usually wanting. Involucels of many slender bractlets. Flowers white. Calyx-teeth 5, broad. Fruit subglobose, glabrous, with 5 equal flattish corky ribs; oil-tubes solitary in the intervals. Stylopo- dium depressed. C. maculata L. Water Hemlock. Stem 2-6 feet high, spotted or streaked: leaves bipinnate or tripinnate; leaflets lanceolate, mucronately serrate; veins running to the notches; fruit nearly terete. Sloughs and marshes; June— August; common. ‘ C. bulbifera L. Stem 1-3 feet high; leaves 2-3 pinnate; leaflets linear, remotely toothed; upper axils bearing clustered bulblets. Swamps; July— September; infrequent; Muscatine, Linn, and Emmet counties. CONIUM L. Biennials, with tripionately dissected leaves, and white flow- ers. Involucre and involucels of narrow bracts. Calyx-teeth obsolete, fruit ovate, compressed; ribs equal, prominent, wavy when immature; oil-tubes wanting. Stylopodium depressed. C. maculatum L. Poison Hemlock. Stem 2-6 feet high, spotted, glabrous, striate, sometimes glaucous; roots fusiform, fleshy; petioles very much dilated. Waste places; June-July; Winneshiek and Scott counties. CRYPTOTAENIA DC. Glabrous perennials, with 3-foliclate leaves, and white flowers. Involucre and involucels none. Calyx-teeth obsolete. Fruit glabrous, linear-oblong, with 5 obtuse equal ribs; oil-tubes solitary, beneath each rib and in the intervals. Stylopodium slender-conical. C. canadensis DC. Stem smooth, 1-3 feet high; leaflets large, pointed, ovate, doubly serrate, often lobed; umbels irregular, unequally few-rayed; pedicels unequal; fruit often curved. Moist woods; June-September; com- mon. (Deringa canadensis (L.) Kuntze.) BERULA Hoffm. A glabrous aquatic or marsh perennial, with simple pinnate leaves, leaflets serrate and variously cut, involucre and involucels of narrow bracts, and terminal umbels of white flowers. Calyx-teeth minute. Fruit glabrous; ribs slender; pericarp thick, corky; oil-tubes numerous. Stylopodium conic; styles short. B. angustifolia (L) Mert. & Koch. Stem 6-30 inches high; leaflets 7-19, ovate to linear-oblong, serrate, laciniate, or lobed. Reported from Sioux eounty. (B. crectu (Huds.) Coville.) CHAEROPHYLLUM L. Annuals, with ternately decompound leaves, pin- natifid leaflets, and white flowers. Involucre usually none. Involucels of many bractlets. [Fruit narrowly oblong to linear, notched at the base, with Carnot FAMILY. 63 5 equal ribs: oil tubes solitary in the intervals. Stylopodium conical. C. procumbens (L.) Crantz. Stems weak, spreading, smoothish or some- what hairy, 4-16 inches high; umbels féw-rayed; leaflets trifid or pinnatifid; fruit acute. Low woods; April-May: infrequent; Johnson, Henry, Story, and Decatur counties. : OSMORRHIZA Raf. Glabrous or hirsute perennials. with large aromatic roots, and biternately divided leaves. Involucre and involucels few-leaved. Flowers white. Calyx-teeth obsolete. Fruit long, linear, bristly, attenuate atthe base. Stylopodium small, conic. O. longistylis DC. (abrous or somewhat pubescent: stem 1-2 feet high; style nearly as long as the cvary. Rich woods; May-June: common. (Wash- inytonia longistylis (Torr.) Britton.) O. brevistylis DC. Villous: stylopodium and style about as long as the width of the ovary. With the preceding. (WW. cluytoni (Mx.) Britton.) ERYNGIUM L. Perennials, with coriaceous leaves, and bracted sessile flowers in dense heads. Calyx-lobes prominent, rigid, persistent. Petals in- flexed. Fruit ovate or obovate, scaly; ribs obsolete; oil-tubes usually 5. Stylopodium wanting; styles filiform. E. yuccaefolium Mx. Button Snakeront. Stem 2-4 feet high, glabrous, striate; leaves often 1-2 feet long, rigid, grass-like. parallel-veined, tapering toa point, remotely bristly; heads ovate-globose; flowers white. Prairies; July-August; common. (E. aquaticum L.) SANICULA L. Smooth oiennials or perennials, with few palmately divided leaves, and greenish yellow flowers. Umbels irregular. few-rayed. Involucre and involucels few-leaved. Calyx-teeth prominent and persistent. [Fruit globular, ribless, with 5 oil-tubes, densely clothed with hooked bristles. S. marylandica L. Stem 1-3 feet high, but little branched; leaves 3-7- parted, root-leaves long-petioled; segments irregularly serrate or dentate, often incised; staminate flowers many, long-pedicelled; calyx-segments lan- ceolate, cuspidate; petals and anthers greenish white; pistillate flowers ses- sile; styles slender, recurved, longer than the bristles. Low woods; May- July; common. S. gregaria Bicknell. Stems usually clustered, weak. erect, 1-3 feet high; leaves 5-divided; segments doubly serrate, teeth bristle-tipped, incised; calyx lobes ovate, obtuse; petals and anthers yellow: styles longer than the bris- tles. This species probably belongs to our flora. S. canadensis LL. Biennial, stem 1-4 feet high, branched: leaves 3-5 divided; segments cuneate-obovate to oblong, serrate and incised; staminate flowers never in separate heads; styles shorter than the bristles. Low woods; May-August; common. PASTINACA L. Biennial, with pinnately compound leaves, involucre and involucels mostly wanting. and yellow flowers. Calyx-teeth obsolete. Fruit flat, oval, the dorsal ribs filiform, the lateral broad and nerved. Stylopodium depressed. P. sativa L. Stem 3-+ feet high. grooved, glabrous; root fusiform: leaves downy beneath; leaflets ovate to oblong, toothed. terminal 3-lobed, all more or less lobed or incised. Fields and waste places; July-September; frequent. POLYTAENIA DC. Perennial, glabrous herb, with pinnately decompound leaves, upper opposite, lower alternate and long-petioled, invrolucre wanting, involucels of narrow bractlets. and bright yellow flowers in compound um- bels. Calyx 5-toothed. Fruit flattened, obovate to oval, glabrous, dorsal ribs small, obscure, lateral thick and corky. Stylopodium wanting. 64 UMBELLIFERAE. P. nuttallii DC. Stem 2-+ feet high; pedicels and involucels pubescent; leaf-segments narrow, entire or lobed. Prairies and upland woods; May- July; rather rare; Scott, Muscatine, Johnson, Story, Appanoose, Decatur, Ringgold, and Montgomery counties. THASPIUM Nutt. Perennials, with ternately divided leaves, and yellow flowers. Involucre wanting. Involucels 3-leaved. Calyx 5-toothed. Stylo- podium wanting; styles slender, exserted. Fruit oblong, ovoid, with promi- nent wings; oil-tubes solitary in the intervals, 2 on the inner face of the car- pels. T. barbinode Nutt. Stem 2-3 feet high, somewhat angular, striate, loosely branched, somewhat pubescent on the joints or in the umbels; leaves 1-3-ter- nate, mostly biternate: leaflets ovate, with cuneate base, coarsely serrate, often cleft or parted; flowers light yellow; fruit broadly oblong, 5-7 winged. Rich woods; May-June; probably common. FOENICULUM Adans. Biennials or perennials, with striate stems, leaves biternately dissected into numerous filiform segments, and yellow flowers. Calyx-teeth obsolete. Involucre and involucels wanting. Fruit elliptic-ob- long, subterete; ribs 5, prominent; oil-tubes solitary. Stylopodium large, conic. F. officinale All. Whole plant aromatic; stem 4-6 feet high, purplish green, somewhat glaucous. Waste places; July-September; rarely escaped from cultivation; Johnson and Story counties. (F. foeniculwm (L.) Karst.) PIMPINELLA L. Glabrous perennial, with ternately compound leaves, involucre and involucels scanty or wanting, and yellow fowers. Calyx-teeth obsolete. Fruit ovate to oblong, laterally compressed; ribs slender, equal; oil-tubes numerous, 2-6 in the intervals. Stylopodium thick, broadly conic. P. integerrima (L.) A. Gray. Stem smooth, glaucous, 1-3 feet high; leaves 2-3-ternate, leaflets ovate to lanceolate, entire; fruit broadly oblong; stylopodium small or wanting. Rocky woods; May-June; common. ZIZIA Koch. Glabrous perennials, with undivided or ternately divided leaves, and yellow flowers. Involucre wanting. Involucels of small bractlets. Calyx-teeth prominent. Fruit glabrous, with slender ribs; oil-tubes large, solitary, in the intervals and one in each rib. Stylopodium wanting. A genus distinguished from Thaspium by the wingless fruit. Z. aurea (L.) Koch. Stem 2-3 feet high, radical leaves long-petioled, all except the uppermost 2-3-ternate; leaflets ovate to lanceolate, sharply serrate, sometimes incisely lobed or parted; rays 15-20; fruit oblong. Low grounds, prairies and meadows; May-June; common. Z. cordata (Walt.) DC. Stem 2-3 feet high; lower leaves long-petioled, ovate to orbicular, crenate, base deeply cordate, stem leaves short-petioled, ternate or quinate, segments ovate or oval, crenate or lobed; rays 7-16; fruit ovate or oval. Woods; May-June; Lyon county. ARALIACEAE Vent, Ginsena FAMILY. Ours perennial herbs or shrubs, with alternate pinnate leaves, and um- bellate flowers. Petals not inflexed, valvate in the bud. Styles usually 2-5. Ovary 2-5-celled; cells l-ovuled. Fruit a 2-5-celled drupe. ARALIA. Leaves alternate, decompound; styles 5. PANaX. Leaves verticillate, palmately 3-7-foliolate; styles 2-3, ARALIA L. Herbs or shrubs from thick perennial roots. Flowers, white or greenish, more or less polygamous. Calyx-tube adherent to the ovary, limb 5-toothed. Petals 5, oblong, spreading, deciduous. Stamens 5, alter- Doag-Woov Famtny. 65 nate with the petals. Styles slender, recurved above, persistent, Cells of the ovary with a single suspended ovule. A. racemosa lL. Spikenard. Stem smooth, branched; leaves large, biter- nate-pinnate; leaflets ovate, base cordate, doubly serrate, taper-pointed; um- bels numerous, racemose; styles 5, united; fruit 5-celled. Rich woods; July; frequent. A. naudicaulis L. Wild Sarsaperilla. Nearly stemless; roots long, hori- zontal; leaf long-stalked, ternate, divisions 3-5-foliolate; leaflets ovate, taper- pointed. serrate; umbels 2-5, on a scape shorter than the leaf-stalk; fruit 5- celled. Rich woods; May-June; infrequent; Winneshielk, Allamakee, Fayette, Scott, Muscatine, Henry, Johnson, Story, and Winnebago counties. A. spinosa L. Hercules Club. A shrub or low tree, 8-30 feet high, stem and stalks prickly; leaves long-petioled, bipinnate; leaflets ovate, pointed, serrate, glaucous and more or less pubescent beneath; umbels in terminal compound panicles, peduncles and pedicels pubescent; flowers white. Rich soil; June-August; an escape from cultivaticn; Johnson county. PANAX L. Ours a perennial herb, froma fusiform, simple or branched root. Leaves at the summit of the stem, verticillate,.digitately compound. Umbel simple, solitary, terminal. Calyx-limb obscurely 5-toothed. Petals 5, spread- ing. Stamens 5, alternate with the petals. Styles 2-3. P. quinquefclia L. Ginseny. Stem slender, 1-foot higk; leaves palmately 3-7-foliolate; leaflets mostly 5, stalked, obovate-oblong, taper-pointed, coarsely serrate; flowers greenish yellow; styles 2: fruit 2-celled, bright red. Rich woods; July-August; becoming infrequent; Winneshiek, Fayette, Jack- son, Delaware, Muscatine, Henry, Johnson, Story, and Cerro Gordo counties. (A. quinquefolia Dee. & P1.) CORNACEAE Link. Dog-Woop Pamity. Ours shrubs, with simple opposite or alternate exstipulate leaves, and cymose flowers. Calyx-tnbe adherent to the 1-2 celled ovary; limb minute, toothed or lobed. Petals valvate in the bud, distinct, alternate with the calyx-teeth, epigynous, sometimes wanting. Stamens as many as the petals, and alternate with them. Ovules pendulous. Fruit a 1-2- seeded drupe, crowned with the calyx. CORNUS L. Leaves with one exception opposite, entire. Flowers per- fect, white in our species. Sepals, petals, and stamens 4. Style 1, slender; stigma terminal, flat or capitate. Drupe with a 2-celled, 2-seeded stone. * Lerves opposite. + Lentves glabrate or minutely pubescent beneath. C. paniculata L’Her. Panicled Cornel. Dogwood. A shrub 3-10 feet high, branches erect, grayish, smooth; leaves lanceolate or ovate-lanceolate, long- acuminate, base acute, minutely appressed-pubescent on both sides, pale be- neath; flowers cymose-paniculate, white; fruit globose, white: stone subglo- bose, slightly furrowed. Rich soil, woods and thickets; May-June; common. (C. candidissima Marsh.) C. stolonifera Mx. Red-osier Dogwood. Shrub 3-12 feet high, wultiplies freely from stolons thus forming clumps; the branches bright red-purple, glabrous or slightly, appressed pubescent; leaves oval, ovate, or ovate-lanceo- late, acute or short acuminate, base rounded or narrowed, minutely pubescent on both sides, whitish beneath; flowers in flat-topped cymes, white; fruit ” 66 CORNACEAE. whitish or white, globose; stone variable. Moist soil; June-July; ,a species frequently reported but is confused with the others t + Leaves downy-pubescent beneath, sometimes becoming glabrate. C. sericea L. Silky Cornel. Kinnikinnik. A shrub 3-10 feet high; branches mostly purplish; branchlets and inflorescence silky-downy; leaves lanceolate, ovate or elliptical, base rounded or acute, glabrous above, white or silky- downy often rusty beneath; cymes broad, flat, compact; calyx-teeth lanceo- late, conspicuous; fruit globose, pale blue; stone oblique, base somewhat pointed. Moist woods; May-July; common. ((. amonum Mill.) C. asperifolia Mx. Shrub 4-8 feet high; branches reddish brown, mostly pubescent; branchlets and inflorescence rough-pubescent; leaves narrowly ovate to round-ovate or oblong, more or less acuminate, base acute or obtuse, rough pubescent above, whitish and roughish woolly beneath; cymes loose, broad, often paniculate; calyx-teeth small; petals oblong-lanceolate, white; fruit globose, white; stone slightly furrowed. Low woods; May-June; com- mon. C. circinata L’Her. A shrub 4-10 feet high; branches greenish, warty, glabrous; leaves broadly ovat, acuminate, white; tomentose beneath; cymes depressed, dense; fruit globose, light blue; stone globose, ridged. Rich woods, May-June; rare; Winneshiek, Fayette, and Cerro Gordo counties, re- . ported from Story and Benton counties. ** Leaves alternate. Cc. alternifolia L. f. Shrub 8-20 feet high; leaves often clustered at the ends of the branches, oval, acuminate, base narrowed or rounded, hoary beneath; cymes pubescent; fruit globular, blue; stone obovoid. Rocky woods; May-July; rather rare; Fayette, Jackson, Delaware, Muscatine, Henry, and Johnson counties. Acacia ACERACEAE Actaea Agrimonia Agrostemma Ailanthus Family Ailanthus AIZOACEA Alfalfa Alyssum -Amelanchier Ammannia Ampelopsis Amphicarphaea Amorpha Amygdalus ANACARDIACEAE Anemone Anemonella Apios Apple Family Apple ANONACEAE Aquilegia Arabis ARALIACEAE Aralia Arenaria Argemone Aruncus Asimina Astragalus Baneberry Baptisia Barbarea Barberry Family Basswood BERBERIDACEAE Berula Bicuculla Bishop’s Cap Bitter Sweet Blackberry Bladder Campion Bladder-nut Family Bladder-nut Bloodroot Bouncing Bet Box-Elder Brasenia Brassica Buckeye Family Buckeye Buckthorn Family Buckthorn Burning Bush Bush Clover Buttercup CACTACEAE Cactus Family CAESALPINACEAE Callirrhoe 37 | 30 48 20 28 28 60 35 15 51 55 30 42 ~ 32 28 59 59 42 24 INDEX. Callitriche Caltha Camelina Capnoides Caper Family CAPPARIDACEAE Capsella Caraway Cardamine Carret-weed Family Carpet-weed Carrot Family Carum CARYOPHYLLACEAE Cassia Castalte Catehfly Catgut Caulophyllum Ceanothus CELASTRACEAE Celastrus Cerastium Cercis Chaerophyllum Chamaenerion Cherry Chick weed Chinquefoil Chrysospleniura Cicuta , Circaea Cissus CISTACEZ Claytonia Clematis Cleome Clover Cohosh Blue Columbine Comarum Conioselinum Conium Corn Cockle CORNACEAE Cornus Cornel Corydalis Cow-herb Crab Apple Cracca Crane’s-bill CRASSULACEAE Crataegus Cress Crotalaria Crowfoot Family Crowfoot CRUCIFERAE Cryptotaenia Cucumber, wild One-seeded 54 16 10 16 16 11 62 15 60 60 60 62 19 42 20 37 © 29 28 28 21 42 62 57 44 21 47 59° 62 58 30 17 22 16 34 48 61 62 20 65 65 65 10 19 50 37 26 53 50 15 34 10 62 59 59 CUCURBITACEAE Cuphea Currant Custard Apple Family 6 Dalea Dame’s Violet Daucus Delphinium Dentaria Deringa Desmanthus Desmodium Dewberry Dianthus Dicentra Didiplis Dog-Wood Family Dogwood Draba DRUPACEAE Dutchman's Breeches Echinocystis Eglantine Elodes Epilobium Eryngium Erysimum Evening-Primrose Family Evening Primrose Evuonymus Faleata False Indigo False Flax Fire-weed Flax Family Flax Floerkea Foeniculum Fragaria FUMARIACEAE Fumitory 34, Gaura GERANIACEAE Geranium Geum Ginseng Family Ginseng Glade Mallow Gleditschia Goat’s Beard Goat’s Rue Golden Saxifrage Golden Seal Gooseberry Family Gooseberry Gourd Family Grape Family GROSSULARIACEAE Ground Plum 36 1 61 6 15 62 43 38 46 19 10 56 65 65 15 43 10 59 49 23 57 63 12 56 bv 28 42 36 16 57 26 26 at 64 47 9 9 58 26 26 46 64 65 25 43 45 37 52 6 53 53 59 29 53 37 68 Gymnocladus HaALORAGIDACEAE HaAaMAMELIDACEAE Hamamelis Heart’s-ease Hemlock Hepatica Heracleum Hercules Club Hesperis Heuchera Hibiseus HipeocasTANACEAE Holly Family Hop-tree Horseradish Hosackia Hydrangea Hydrastis HYPERICACEAE Hypericum Tlex ILICINEAE Impatiens Todanthus Isnardia Isopyrum June-berry Jewel-weed Family Ketmia Kinnikinnik Kneiffia AKuhnistera Lake Cress Larkspur Lathyrus Lead-plant Leather-flower Lechea LEGUMINOSAE Lepidium Lespedeza LIMNANTHACEAE LINACEAE Linden Family Linden Liquorice, Wild- Live-forever LOACEAE Loasa Family Loco-weed Locust, Honey Loosestrife Loosestrife Family Lucerne Ludwigia Lupinus Lychnis LYTHRACEAE Lytbrum ig to Win om DO wis ig INDEX. Mallow Family Mallow Malva MALVACEAE Malvastrum Maple Maple Family Marsh Cress Marsh Marigold May Apple Meadow-Beauty Family Meadow-Rue Meadow-Sweet Medicago Meibomia MELASTOMACEAEB Melilot Melilotus MENISPERMACEAE Venispermum Mentzelia Mermaid weed Meriolix Micrampelis Milkwort Family Millfoil, water MIMOSACEAE Mitella Miter-wort Mochringia Mollugo Moonseed Family Moonseed Mouse-tail Mustard Family Mustard Mustard, Worm- seed Myriophyllum Myosurus Napaea Nasturtium Nelumbo ' New Jersey Tea Nine Bark Nuphar NyMPHAEACEAE Nymphaea Oenothera ONAGRACEAE Opulaster Opuntia Orange-root Orpine Family Orpine Osmorrhiza OXALIDACEAE Oxalis Oxygraphis ) Oxrypolis Oxytropis wwuwok WMD MR WW Wk WO HPO Wd Or or ar @W wet or 23 24 24 25 30 30 11 wt Oworobuwoons Ho io we ov eet 56 59 6 53 52 63 on 10 61 a8 Panax Pansy Parnassia Paroselit PAPAVERACEAE Papaw PAPILIONACEAE Parsonia Partridge Pea Pasque-flower - Pastinaca Pea Family Peach Penny-cress Penthorum Pepper-root Petalcstemon Phaca Physocarpus Pimpinella Pink Pink Family Plum Family Plum, wild Podophyllum Poison oak Poison ivy Polanisia POLYGALACEAE Polygala Polytaenia POMACEAE Poppy Family PORTULAC4CEAE Portulaca Potentilla Prairie Apple Prickly Ash Prickly Poppy Proserpinaca Prunus Psoralea Ptelea Pulsatilla Purslare Family Purslane Pyrus Radish RANUNCULACEAE Ranunculus Raphanus Raspberry Rattle-box Reseda RHAMNACKAE Rhamnus Rhus Ribes Robinia Rock Cress Queen-of-the-Prairie 33 56 +2 2 63 33 44 il 53 15 a7 38 45 64 19 19 Pitcher-Plant Family 43 44 us, 32 a 16 a am 63 wwe to we oF CW WPA khe OMG -2 gris to INDEX. 69 Rori rea 11 | Soapwort 19 | Touch-me-not 27 a 49 | Spesia 38 | Triadena Rosacka pi ame es z 44 | Spikenard 65 | Tree-of-H 9 Hi Ben w Rose Family 44 | Spiraea 45 | Vidichiom 3d Rose Mallow oy Spring Beauty 99 Turni mm Rotala 56 | Squirrel Corn 10 e a oe 45 | Staff-tree Family 23 | UMBELLIFERAE 60 ue Anemone 3 | STAPHYLEACEAB 31 | Ulnaria 45 Rue Family 27 | Staphylea 31 ce RurackaE 27 | Starry Campion 19 oe 40 = Be Sad 3 i Tiola ene 9 oy VIOLACEAR nt i Snorer 63 | St. John’s-wort 99 | Violet Family 17 a 19 7 Vi inia C SiceRGan St. John’s- Wort irginia Creeper 30 NIACEAE 9 . Virgin’s-Bow 17 Sarsaparilla 65 | 3 Family 22 ieee ee 3 SAXIFRAGACEAE pil, |: oPerODy Tnteh 53 | Viti fi Sasi Strawberry 47 itis 30 aifrags SL | Strophosty! Saxifrage, Golden 52 Sai te es = Wall-flower, Western 12 Saxifrage Family 5] : oon ao ‘ te Washingtonia 63 Seed-box 87 | sa y 5 | Water Cress Is 12 Sedum 53 Scett : 32 | Water Lily 8 Senaca Snakeroot 32 Ss Be ae 49 | Water Lily Family 7 Senna Family a2 ous 35 | Water Millfoil Senna, Wild 42 oan TODS 58 Family 54 Senna, Low 43.) Pulcesmon 3 Whur Grass 1A Service-berry 51 | Talinum 99 illow Herb 36 Shepherd's purse 11 | Tephrosia 37 Willow-herb, Great 57 Sickle-pod 14 | Thalietrum 3 Wind-flower 1 Sicyos 59 | Thaspium 64 | Witch Hazel Family 54 Sida 25-1 Tedemannia 61 | Witeh-hazel 54 Side-saddle flower 9 | Thelypodium 13 | Wood-sorrel a0 Silene 19 | Thiaspi 31 | Wood-sorrel Family 27 SIMARUBACEAE 28 | TMLIACEAE 2 = ‘ Sisymbrium 13 | Tilin ba Xanthoxylum 28 Sium 61 | Toothache tree og | Zizia 64 Snakeroot 83 | Toothwort 15 ERRATA. Errors are an especial nuisance. Page 3 line 2 for petiolate read petiolulate. Page 3 line 37 for preceeding read preceding. Page 3 line 46 for Emmett read Emmet. Page 4 line 28 for petiolate read petiolulate. Page 16 line 38 for preceeding read preceding. Page 16 line 39 for foliate read foliolate. Page 27 line 6 for trifoliate read trifoliolate. Page 28 line 1 for Celistraceae read Celastraceae. Page 31 line 8 for preceeding read preceding. Page 31 line 14 for mountaim read mountain. Page 31 lines 27 and 44 for foliate read foliolate. Page 33 line 7 coralla read corolla. Page 33 lines 26, 32, 37,41, and 45 for foliate read foliolate. and so on through the Pea Family. Page 53 line 29 for Emmett read Emmet. Page 60 line 44 for globrous read glabrous. Page 60 line 44 for galbous read globose. All other mistakes are so obvious that the reader may correct, overlook or eondemn at his pleasure. < MANUAL OF THE FLOWERING PLANTS OF IOWA PART 2. GAMOPETALAE. MANUAL OF THE FLOWERING PLANTS OF IOWA BY T. J. FITZPATRICK. Fellow of the lowaAcademy of Sciences A TNESIS. PRIVATELY PUBLISHED 1899: Copyright Applied For. ANALYTICAL KEY TO THE FAMILIES. Division 2. GAMOPETALAE: calyx and corolla usually present, the petals more or less united. Polypetalous forms occur in the Asclepiada- ceae, Oleaceae, Pyrolaceae, and the Monotropaceae, and apetalous forms occur in the Oleaceae as noted at the end of the key. A. Stamens more numerous than the lobes of the corolla. Stamens free from the corolla or merely adnate to its base. Ovary superior, 4-5-celled; fruit a cupsule. Low evergreen perennials. PYROLACEAE, 102. Waxy-white parasitic plants. MONCTROPACEAE, 103. Ovary inferior, advate to the calyx; fruit a many-seeded berry. VACCINIACEAE, 103. Stamens borne on the tube or at the base of the corolla. Leaves alternate. EBENACEAR, 105, Leaves opposite and basal. ADOXACEAE, 69. B. Stamens as many as the lobes of the corolla und inserted on the tube opposite the lobes. PRIMULACEAE, 104. C. Stamens ux many as the lobes of the corolla and inserted alternate with them, or ewer, - : * Ovary inferior, adherent to the ealyx-tube. + Anthers united, ercept in Kuhnia of the Compositae, the Ambrosiaceac, und mony of the Campanulaceae, Flowers in involucrate heads. Flowers all ligulate; juice milky. CICHORIACBAE, 97. Flowers tubular or the outer ligulate; juice rarely milky. Stamens distinct or nearly so. AMBROSIACEAR, 95. Stamens united by their anthers around the style. COMPOSITAE, 72. Flowers not in involucrate heads. CAMPANULACEAE, 101. + fF Stamens inserted on the corolla; anthers separate. Stamens in ours 3, fewer than the corolla-lobes. VALERIANACEAE, 71. Stamens as many as the corolla-lobes, one lessin Linnuew of the Caprifoliaceae. Leaves opposite or verticillate, usually stipulate. RUBIACEAE, 70, Leaves opposite, exstipulate. CAPRIFOLIACEAE, 67. ** OQoury superior. + Corolla Inregulir, more or lexs 2-lipped; stamens 4 and didynaumeus, or only .2. t Carpels 1—/?-xeeded. Ovary 4-lobed, becoming 4 1-seeded nutlets. LABIATAE, 131. Ovary not lobed; carpels separating into l-seeded nutlets. VERBENACEAE, 130, Lt Carpels 2—many-seeded, Ovary 1-celled; stamens 2; marsh or aquatic herbs. LENTIBULARIACEAE, 128. Ovary 1-celled; stamens 4; brownish or whitish parasitic herbs. OROBANCHACEAR, 127. Ovary 1 celled or becoming partially 4-celled; fruit with 2 long recurved MARTYNIACEAE, 129. beaks. Ovary and capsule Z-celled; placentae parietal; seeds winged in our species; small trees or woody vines. BIGNONIACEAE, 128, Ovary and capsule 2-celled; placentae axile. Seeds usually many. SCROPHULARIACEAE, 120, Seeds few, borne on curved projections. ACANTHACEAB, 129, +7 Corolla somewhat irregular; stamens 5, Ovary deeply 4-lobed. Echium, in BORRAGINACEAB, 112. Ovary notlobed; filaments bearded. Verbascum, in SCROPHULARIACEAE, 121, TTT Corolla trreygular. { Stamens of the same number us the lobes of the corolla, Ovaries 2, separate; stigmas and somvtimes the styles united. Filaments distinct; pollen granular. APOCYNACEAE, 106. Filaments monodelphous; pollen in waxy-masses. ASCLEPIADACEAE, 107. Ovary deeply 4-lobed. Leaves alternate. BORRAGINACEAE, 112. Leaves opposite. Mentha, in LABIATAE, 133. Ovary 1, not deeply lobed, 1-celled, 1-ovuled. PLANTAGINACEAB, 139. Ovary 1-celled, with the ovules parietal or in 2 parietal placentae. Leaves opposite. _ GENTIANACEAE, 109. Leaves basal or alternate, 3-foliolate. MENYANTHACEAE, 110. Leaves basal or alternate, toothed, lobed, or compound. HYDROPHYLLACEAE, 111. Ovary 2—10 celled. Leafless white or yellowish parasitic vines. CUSCUTACEAE, 116. Leaves ulternate and stipulate or opposite and exstipulate; stamens in- serted on the tube of the corolla. Stamens 4; capsule 2-celled, circumscissile. PLANTAGINACEAE, 139. Stamens 4; ovary 2—4 celled, becoming 2—4 1-seeded nutlets. VERBENACEAE, 130. Stamens usually 5. / ‘Fruit of 4 1-seeded outlets, or of 2 2-seeded carpels. BORRAGINACEAE, 112. Fruit a few-seeded capsule. d Calyx 5-cleft; style 3-lobed or 3-cleft. POLEMONIACEAB, 110. Calyx of 5 distinct sepals; styles 1 or 2, 2-cleft or entire; mostly twin- ing vines. Leaves normal. CONVOLVULACEAE, 115. Leaves reduced to minute scales. CUSCUTACEAER, 116. Fruit a many-seeded capsule or berry; style solitary. SOLANACEAE, 117. Lt Stamens less in number than the lobes of the corolla, Stamens 4, didynamous. Ovary 2-celled; cells several-ovuled. ACANTHACEAE, 129. Ovary 2—4-celled; cells 1-ovuled. VERBENACEAE, 130. Stamens with anthers 2; ovary 4-lobed. Lycopus, in LABIATAE, 133. Stamens 2; ovary 2-celled. ° Herbs. Veronica, in SCROPHULARIACEAE, 124. Trees or shrubs. OLEACEAE, 106. Polypetalous forms in gamopetalous families. Ovaries 2, separate. ASCLEPIADACEAE, 107, Ovary 1, compound. Stamens 2. rarely 3 or 4; petals 4. OLEACEAE, 106. Stamens in ours 8 or 10; petals 4 or 5, > Low evergreen perennials. PYROLACEAR, 102. Waxy-white parasitic plants. MONOTROPACEAB, 103. Apetalous forms in gamopetalous families. Trees or shrubs; fruit a l-celled 1-seeded samara or drupe. OLEACEAE, 106. HOoNEYSUCKLE FAMILY. 67 GAMOPETALAE: those with both calyx and corolla present, the latter having its petals more or less united, rarely separate or wanting. CAPRIFOLIACEAE Vent. HonrysuckiE FAMity. Shrubs or herbs, with opposite exstipulate leaves, and perfect mostly 5-parted regular or irregular flowers. Calyx-tube adherent to the ovary. Corolla monopetalous, tubular, or rotate. Stamens inserted on the calyx- tube, usually of the same number as the lobes, one less in Linnaea. Style 1; stigmas 3-5. Fruit a berry, drupe, or capsule, 1-several- seeded. Tribe Lonicereae. Corolla tubular, limb regular or irregular; style fili- form. * Herbs; jflowcrs axillary; corolla 5-lobed. LINNAEA. Stamens 4. TRIOSTEUM. Stamens 5. . * * Shrubs, erect or climbing. SYMPHORICARPOS. Stamens 4-5; berry 4-celled. 2-seeded. LONICERA. Corolla tubular; stamens 5; berry 2-3-celled. DIERVILLA Stameas 5: fruit a 2-celled, 2-valved, many-seeded capsule. -Tribe Sambuceae. Corolla regular, deeply 5-lobed, rotate; stigmas 3-5, mostly sessile. Sambucus. Leaves pinnate; fruit a 3-seeded berry. VipukNnom. Leaves simple; fruit a i-celled, 1-seeded drupe. LINNAEA L. A single species, creeping, evergreen, perennial. Leaves roundish, slighly ecrenate, petiolate. Peduncle slender, erect, 2-flowered. Calyx small, 5-parted, deciduous. Corolla campanulate, lobes 5, about equal. Stamens 4, 2 longer, inserted toward the base of the corolla. Style slightly exserted; stigma capitate. L. borealis L. Somewhat pubescent; stems 6-30 inches long; pedicels from the axils of small branchlets; flowers nodding; corolla roseate, fragrant, about a half inch long. Moist woods; June; infrequent; Winneshiek county. TRIOSTEUM L. Coarse perennials, with opposite ample more or less con- nate entire leaves, and sessile axillary flowers. Calyx-tube ovoid, limb of 5 nearly linear segments. Corolla tubular, but little exceeding the calyx, gibbous at the base, limb of 5 nearly equal lobes. Stamens 5, included. Style 1, exserted; stigma capitate, lobed. Ovary 3-celled, forming a drupe. Seeds 3, bony, ribbed. T. perfoliatum L. Fever-wort. Horse Gentian. Stems 2-4 feet high, sim- ple, soft-pubescent to hirsute; leaves oval, acuminate, contracted towurd the base; flowers usually clustered, brownish purple, drupe orange color. Up- Jand woods; May—June; common. T. angustifolium L. Stem 2-6 feet high, hairy; leaves lanceolate, acumi- nate, tapering to the base. hardly connate; flowers mostly solitary, yellow- ish. Woods: May-July; infrequent; Johnson and Lee counties. SYMPHORICARPOS Juss. Low branching shrubs, with opposite oval short-petioled entire leaves and small roseate axillary flowers. Calyx-tube globose, teeth 4-5, short, persistent. Corolla campanulate, limb regular, 4—5- lobed. Stamens 4-5, short, inserted in the throat of the corolla. Stigmas capitate. Fruit a globose berry, 4-celled, 2 of the cells with a single seed, the others abortive. S. vulgaris Mx. Indian Currant. Coral-berry. Stems slender, 2-4 feet high; leaves nearly sessile, downy beneath; flowers in short thick clusters; 68 CAPRIFOLIACEAE. stamens and bearded style incluted; berriessmall, red. Along streams and bordering uplands; July; common but confined mostly to the southern half of the state; Lee, Henry, Jefferson, Van Buren, Wapello, Appanoose, Monroe, Decatur, Lucas, Ringgold, Union, Clark, Adams, Taylor, Page, Montgomery, Fremont, Pottawattomie, Harrison, Monona, Shelby, Ida, Woodbury, Sioux, Lyon, and Story counties. (S. symphoricarpos (L.) MaeM.) S. occidentalis Hook. Wolfberry Leaves petioled; flowers larger than those of the preceding, spicate, axillary, terminal; corolla bearded within; stamens and glabrous style exserted; berries white. Upland woods; June- July; frequent; Emmet, Lyon, Dickinson, Shelby, Pottawattomie, Fremont, Page, Winnebago, Story, Fayette, and Winneshiek counties. LONICERA L. Bushy or climbing shrubs, with opposite entire leaves. and usually showy flowers. Calyx-tube subglobose; teeth 5, very short. Corolla tubular or funnel-form; limbin 4 or 5 often unequal lubes. Stamens 5, ex- serted. Stigma capitate. Ovary 2-3-celled, becoming a few-seeded berry. L. sullivantii Gray. Honeysuckle. Stems climbing, 3-6 feet high; leaves glabrous, often much glaucous, sessile, niostly connate-perfoliate, oval or ob- ovate, the upper pair fortaing an orbicular disk; corolla pale yellow, hairy within, tube half inch long, limb shorter; filaments nearly glabrous; berries red. Woods; June; common; Winneshiek, Allamakee, Clayton, Dubuque, Fayette, Jackson, Delaware, Muscatine, Johnson, Henry, Fremont, Pottawat- tomie, Winnebago, Chickasaw, Cerro Gordo, and Emmet counties. ~ L. glauca Hill. Stems 3-4 feet high, usually bushy; leaves glabrous. less glaucous, oblong, upper pairs connate; flowers greenish yellow or purplish, tube short, limb nearly as long, hirsute within: style and base of the fila- ments hirsute; berries yellow. Woods: May-June; common; Winneshiek, Allamakee, Fayette. Story, and Emmet counties, specitnens referred to this species have been collected in Delaware, Johnson, Cerro Gordo, Uenry, and Calhoun counties. (L. dioicr L.) L. tartarica L. Bush Honeysuckle. An erect glabrous shrub, 5-10 feet high. with ovate short petioled leaves, and pinkish or whitish flowers on axil- lary bracted peduncles. Frequently cultivated; has been collected along the Volga river, Fayette county, probably an escape. DIERVILLA Moench. (Leaves petioled, ovate, acuminate, serrate. Flow- ers in axillary cymes. Calyk-tube oblong, the limb of 5 linear segments. Corolla funnel-form, nearly regular, 5-lobed, twice the length of the calyx. Stamens 5, exserted. Fruit a capsule, 2-celled, 2-valved, many-seeded. D. trifida Moench. A small shrub, 4-6 feet high, with taper-pointed leaves, and pale yellow flowers; capsule oblong, with a slender beak, crowned with the persistent calyx-lobes. Bluff thickets; June-July; infrequent; -Winneshiek, Fayette, and Muscatine counties. (D. diervilla (L ) MacM.) SAMBUCUS L. Small shrubs, with simple pinnate leaves. and small cymose white flowers. Calyx-lobes small, 5 or none. Corolla-limb 5-cleft, spreading. segments obtuse. Stamens 5. Stigmas3. Fruita pulpy berry, 3- seeded. S. canadensis L. Common Elder. Stems more or less woody, 4-10 feet high, with a large white pith, leaflets 5-11, ovate-oblong, acuminate, smooth; fruit dark purple. Rich soil, fields and open woods; June-J uly; common. S. racemosa L. Reid-berrie?d Elder. Stems woody, 3-10 feet high, with a reddish-brown pith; leaflets 5-7, ovate-lanceolate, acuminate, slightly downy beneath; cymes panicled; fruit red. Woods; flowers in May, fruit ripe in MoscuaAtTeL FaMIny. 69 June; infrequent; Winneshiek, Clayton, Fayette, Delaware, and Scott coun- ties. (S. puhens Mx.) VIBURNUM L. Small trees or shrubs, with simple leaves, and white flowers in compound cymes. Calyx small, persistent, 5-toothed. Corolla rctate, deeply 5-lobed, segments obtuse. Stamens 5. Stig:mas 1-3, sessile. Fruit a 1-celled, 1-seeded drupe. Fetioles frequently minutely stipulate. V. lentago L. Swect Viburnum. Sheep-berry. Shrub or small tree, 8-30 feet high; leaves ovate, long-acuminate, closely and sharply serrate, with long margiped petioles; cymes 3-5 rayed, sessile; drupes oval, half inch long, ripe in autumn, edible. Woods; May-June; frequent throughout the state. V. prunifolium L. Black Haw. Similar to the preceding; leaves oval, ob- tuse or acutish. finely serrulate. Low woods; May-June; rare; Johnson county, reported from Huinbolt county. V. pubescens (Ait.) Pursh. Downy-lectwved Arrow-wood. Shrub 2-3 feet high; leaves ovate to oblong-ovate, short acuminate, broadly dentate, pin- nately veined, downy when young, base cordate, short-petioled or nearly ses- sile; drupe dark purple, oval; stone flattish, slightly 2-grooved on both sur- faces. Rocky woods; May-June; infrequent; Emmet, Fayette. Johnson, Henry, Decatur, Story, and Cerro Gordo counties. A form of this species with the characteristic fruit but the petioles a half inch or more long is found in Johnson county. The forms referred to the next species may belong here. V. dentatum L. Arrow-wood. A shrub, 4-10 feet high; branches slender, glabrous, gray; leaves ovate, oval or orbivular, acute: or short-acuminate, coarsely dentate, both sides glabrous or pubescent in the axils of the veins beneath, base rounded or slightly cordate, petioles one-fourth of an inch to an inch long, glabrous; drupe globose-ovoid, blue, becoming blackish; stone deeply grooved on one side, rounded on the other. Woods; May-June; infre- quent; forms have been referred to this species from Jackson, Delaware, Johnsun, Henry, Winnebago, and Dallas counties. : V. opulus L. Cranberry-tree. Wild Guelder-rose. Shrub 5-12 feet high; leaves deeply 3-lobed, 3-ribbed, broadly ovate, base rounded or truucate, lobes acuminate, dentate, entire in the sinuses; petioles stipuled. and with 2 glands above; drupes globose or oval, red, acidic. stone orbicular, flat, not grooved. Woods; June-July; rather rare; Allamakee and Delaware counties, reported from Fayette county. ADOXACEAE Fritsch. MoscuaTEL FAMILY. Dwarf perennial herbs, with scaly or tuberous rootstocks, basal and opposite ternately divided leaves, and capitate cluster of greenish flow- ers. Calyx-limb of 3 or more teeth. Corolla rotate, 4-6-lobed. Stamens of 4-6 pairs, in the sinuses of the corolla, separate or partly united; anthers peltate, l-celled. Style 3-5-parted. Fruit a greenish drupe, with 3-5 cartilaginous nutlets. Represented by the monotypic genus Apoxa L. A. moschatellina L. Musk-root. Moschatel. Smooth, 3-5 inches high, radical leaves 1-4, ternately compound, long-petioled; cauline leaves a single pair, 3-parted or 3-cleft; flowers greenish or yellowish, 4-5 in a slender pedun- culate glomerule. Rocky woods; May; rather rare; Winneshiek county, re- ported from Fayette and Floyd counties. 70 RUBIACEAE. RUBIACEAE B. Juss. MADDER FAMILY. Shrubs or herbs, with opposite or verticillate entire leaves. Stipules leaf-like, conuecting the bases of the petioles or none. Flowers perfect but frequently of two forms in Houstonia. Calvx-tube adherent to the ovary; limb 4-5-cleft. Corolla regular, inserted upon the calyx-tube; lobes 4-5. Stamens 4-5, inserted on the corolla-tube, alternate with its lobes. Styles 1-2. Ovary 2-4-celled. Fruit a capsule, berry, or a drupe. . Houstonia. Low herbs. with opposite leaves, and solitary or cymose flowers. CEPHaLsNTaus. Leaves opposite or in 3’s; flowers in dense globular peduncled heads. GaLiumM. Leaves verticillate; stipules wanting; flowers cymose. HOUSTONIA L. Low herbs. Stipules small, entire, connate with the petioles. Flowers solitary or cymose, white or bluish. Calyx 4-toothed or 4- lobed, persistent, the lobes becoming distinct in fruit. Corolla tubular, with 4 spreading lobes, longer than the calyx. Stamens 4, inserted on the throat of the corolla. Style 1; stigmas2. Ovary 2-celled. Fruit capsular, few-several- seeded. Flowers often dimorphous, that is some flowers with exserted style and included stamens, others with included style and exserted stamens. H. minima Beck. Least Bluets. Annual; stem 1-4 inches high, glabrous; radical leaves spatulate or ovate, cauline leaves ovate to linear; peduncles axillary, the earlier ones usually 1 inch long, later ones shorter; flowers vio- let or purple; calyx-lobes conspicuous; corolla-lobes about equalling the tube. Fields and prairies; March-May; common; Johnson county. H. angustifolia Mx. Perennial; stems tufted, from a woody root, 4-20 inches high; leaves often fascicled, narrowly linear, 1-nerved; flowers numer- cus, in compact terminal cymes, white or purplish, short-pedicelled; lobes of the corulla bearded inside; capsule obovoid. Prairies; June-July; frequent; Taylor, Fremont, and Shelby counties; reported from Pottawattamie, Harri- son, and Woodbury counties. CEPHALANTHUS L. Shrubs, with white flowers in dense globose pedun- ecled heads. Calyx-limb 4-toothed. Corolla tubular, 4-cleft. Stamens 4. Style 1, filiform, much exserted; stigma capitate; fruit dry, 2-4-celled. C. occidentalis L. Button-bush. Shrub 2-10 feet high; leaves opposite or in 3's, ovate to lance-oblong, pointed, petioled; stipules small, triangular. Low grounds, along streams and swamps; July-August; frequent in the southeast quarter of the state but rarer elsewhere; Allamakee, Clayton, Clin- ton, Jackson, Scott, Muscatine, Louisa, Lee, Henry, Johnson, Appanoose, Story, Decatur, Page, Fremont, and Humbolt counties. GALIUM L. Herbs, with angled stems, verticillate leaves, and cymose flowers. Calyx-limb of four small teeth. Corolla rotate, 4-cleft, valvate in the bud. Stamens 4, short. Styles 2. Fruit globular, dry or fleshy, separat- ing into 2, 1-seeded, indehiscent carpels. G. aparine L. Annual, stems 2-4 feet long, weak, reclining, retrorsely prickly; leaves usually in 8's, sometimes in 6’s or 7's, oblanceolate to linear, mucronate, peduncles axillary, 1-2-flowered; fruit densely covered with hooked prickles. Low woods; May—August; common. G. circaezans Mx. Wild Liquorice. Perennial, stem 1-2 feet high, branched, downy or smooth; leaves large, in 4’s, oval or ovate-lanceolate, 3- nerved, obtuse; peduncles few-flowered; fruit nearly sessile, hispid. Rich woods; May-July; frequent; Johnson, Henry, Muscatine, Lee, Van Buren, Decatur, Fremont, and Pottawattamic counties. VALERIAN FAMILY. 71 G. boreale L. Northern Bedstraw. Perennial, stem 1-2 feet high, rigid, smooth and glabrous; leaves in 4's, linear-lanceolste, acute, 3-nerved; flow- ers white, uumerous, ia terminal panicles; fruit hispid or smooth. Low prairies and rocky woods; May—August; common; Winneshiek, Allamakee. Fayette, Jackson, Jones, Floyd, Cerro Gordo, Hancock, Emmet, Dickinson, and Lyon counties. G. trifidum L. Small Bedstraw. Perennial, stem weak, decumbent or sub- erect, diffusely branched, retrorsely roughish on the angles; leaves mostly in 4's, sometimes in 6's, linear or oblanceolate, obtuse, margins and midrib rough; peduncles 1-7-flowered; flowers white, frequently 3-parted, pedicels capillary; fruit smooth. ‘Wet woods; June-July; common. This species varies much. Variety pusiLLuM Gray is a small form with-narrow linear leaves, in 4’s, and peduncles 1-2-flowered. Variety LatIFolium Torr., usually less branched; leaves in 4’s, ovate, obtuse or oblanceolate; peduncles about 3-flowered; flowers 4-parted. These are the varieties given in Gray's Manual. They are not now recognized. G. concinnum T. & G. Similar to the preceding, perennial; stem decum- bent, much branched, angles scabrous; leaves all in 6's, linear, slightly pointed, i-nerved, margins scabrous upward, peduncles filiform, 2-3 times forked; flowers small, numerous, cymose, white; fruit smooth. Dry woods; June-July; common. G. triflorum Mx. Sweet-scented Bedstraw. Perennial, diffuse, procumbent or ascending; stem 1-3 feet long, weak, rough on the angles; leaves in 6’s, narrowly oval or elliptical-lanceolate, cuspidate, l-nerved, margins rough; peduncles 3-flowered; flowers greenish, pedicelled; fruit clothed with hooked bristles. Rich woods; June-July; common; Jackson, Jones, Muscatine, Johnson, Decatur, Story, Winnebago, and Emmet counties. G. asprellum Mx. Rouyh Bedstraw. Perennial; stem diffuse, much branched, retrorsely hispid; ieaves in 6’s or in 4's and 5’s on the branches, Janceolate, acuminate or cuspidate, margins and midrib retrorsely prickly; peduncles 2-3 times forked; flowers white; fruit smooth and glabrous. Allu- vial soil; June-August; reported from Jones, Lee, Johnson, and Cerro Gordo counties, perhaps frequent. VALERIANACEAE Batsch. VALERIAN FaAmtity. Herbs, with opposite exstipulate leaves, and cymose-clustered flowers. Calyx-tube adherent to the ovary; limb sometimes pappus-like. Corolla tubular or funnel-form, 4- 5-lobed. Stamens inserted on the corolla tube, alternate with the lobes and usually fewer in number. Ovary inferior, 3-celled, but one cell fertile and l-ovuled. Fruit indehiscent, witha sin- vile suspended ovule. VALERIANA. Fruit 1-celled; calyx-lobes becoming pappus-like. VALERLANELLA. Fruit 3-celled; calyx-lobes minute or obsolete. VALERIANA L. Perennial herbs, with strong-scented roots, mostly pin- nate leaves, and dicecious or dimorphous flowers. Calyx-limb small ‘at first but evolving a plumose pappus as the seed develops. Corolla limb 5-lobed, nearly regular. Stamens 3. Fruit 1-celled, 1-seeded. V. edulis Nutt. Edible Valerian. Stem 1-4 feet high, thick, smooth; root ample, spindle-shaped, edible; leaves densely ciliate, radical spatulate, cau- line pinnately parted into 3-7 long and narrow divisions; flowers whitish, 72 COMPOSITAE polygamo-dicecious, in cymose clusters, disposed in a long narrow interrupted panicle. Low prairies; May-June; common; Winneshiek and Fayette counties. VALERIANELLA Poll. Annual branched herbs, with tufted basal leaves, and terminal capitate, corymbed or panicled flowers. Calyx-limb short and toothed or obsolete. Corolla small, 5-lobed, nearly regular. Stamens 3. Fruit 3-celled, one cell 1-seeded, the others empty. V. chenopodifolia (Pursh) DC. Stem glabrous, 1-2 feet high; lower and basal leaves spatulate, obtuse, sometimes repand, the upper oblong or lance- olate, entire, sessile; flowers in dense cymes; corolla white; fruit triangular- pyramidal, glabrous or pubescent. Moist soil; May-July; reported from Lee county by Arthur. COMPOSITAE Adans. TuHisTLE FAMILY. Curs herbs, with watery or resinous sap, and opposite alternate or ba- sal exstipulate leaves. Flowers perfect, pistillate or neutral, infrequent- ly moneecious or dicecious, few or many borne ona common receptacle which is subtended by a common involucre composed of few or many bracts arranged in oneormoreseries. Receptacle naked or chaffy, smooth, pit- ted or honeycombed. Calyx-tube adherent to the ovary, thelimb wanting or of bristles, teeth, scales, awns, or crown-like or cup-like andis known as the pappus. Corolla tubular, 5-lobed or 5-cleft or the maryinal flowers with astrap-shaped or ligulate corolla; the heads are radiate when the ray- flowers are present, discoid when absent, the tubular are the disk“flowers. Stamens usually 5, borne on the corolla and alternate with its lobes, the an- thers united into a tube which sheaths the style, syngenesious, distinct or nearly so in Kuhnia, sometimes appendaged at the apex or sagittate or tailed at the base, the pollen grains globose, rough or echinate. Ovary 1- celled, 1-ovuled; style of the fertile flowers 2-cleft, the branches various and naked or variously appendaged; stigmas marginal; styleof sterile tiowers usually undivided. Fruitanachene. A large family and well represented in Iowa. Some writers include the two following families with this one. Tribe Vernonieae. Heads discoid; flowers tubular, perfect, never yellow; style-branches filiform or subulate; leaves alternate or scattered. VERNONIA. With us the only genus. Tribe Eupatorieae. Heads discoid; flowers tubular. perfect, never yel- low; style- -branches thickened upward, obtuse, minutely pubescent; leaves alternate, opposite or whorled. EupPATORIuM. Achenes 5-angled; pappus of numerous capillary bristles; maNEENEEE bracts more than 4. Kuanta. Achenes 10-ribbed; involucral bracts striate-nerved; pappus strongly ae mose. Lisrris, Achenes 10-ribbed; involucral bracts faintly striate; pappus plumose or bar- bellate; heads spicate or racemose. Tribe Astereae. Heads radiate or discoid; style-branches of the perfect flowers flattened, with triangular appendages above; flowers often yellow; anthers not tailed at the base; leaves alternate. GRINDELIA. Heads large; flowers fertile, many; pappus of caducous awns. CHRysoPsis, Heads many-flowered; pappus double, the outer of chaffy scales, the inner of capillary bristles. APLOPAPPUS. Heads many-flowered, radiate; pappus persistent, of many unequal bristles; achenes white-tomentose. SoLipaco. Heads small, radiate, tew-many-flowered, yellow, rarely white; pappus of numerous nearly equal capillary bristles. THISTLE FAMILY. 73 BOLToNLA. Heads many-flowered; rays white or purp ish, disk yellow; pappus of a few short bristles and 2-4 awns. AstTER. Heads usually small, many-flowered. on leaf; peduncles; rays white, blue or purple, disk yellow; involucre wellimbricated. paypus simple, copious, of capillary bristles; achenes, flattish. Erigeron. Heads many-flowered, on naked peduncles; invotucre bt little imbricated; pappus a single ruw of capillary bristles with intermixed sma2ier o1es or with an outer pappus of scales or little bristles; achenes flattened. Tribe Inuleae. Heads usually small, discoid, except in Inula where the ravs are large and yellow; anthers sagittate, unappendaged ahove, tailed be- low; style-branches obtuse or lruncate, naked: pappus capillary or none. Our species have the receptacle not chaffy. ANTENNARIA. Heads dicecious;: pappus of the staminate flowers club-shaped, of the pistillate flowers united at the base and falling away together. ANAPHaLIS. Heads dicecious or polygamo-dicecious; pappus not club-shaped nor united at the base, . GNAPHALIUM. Heads with all the flowers fertile; pappus capillary. InuLA. Heads large, many-flowered, with yellow rays; pappus of rough capillary bristles. Tribe Heliantheae. Heads radiate or discoid; involuere herbaceous or foliaceous; receptacle chaffy; pappusa cup or crown, or awns, bristles or scales, never capillary, sometimes none; style-branches truncate or hairy-append- aged. * Disk flowers perfect, but sterile, the rays fertile. Potymnta. Involucral scales in 2 rows, the 5 outer foliaceous; achenes thick, short, not flattened; pappus none SILPHITM. Involucral scales thick, imbricated in several rows: achenes flattened, wing- margined, in several rows; rays in 2 or 3 series; pappus none or 2 teeth. PARTHENIUM. Involucral scales short. roundish. in 2 rows: achenes flattened, keeled and margined: rays 5, very short, the heads appearing discoid; pappus of 2 small scales. * * Disk-flowers fertile; rays deciduous except in Hcliopsis, or none. Ecuiera. Rays white. short; involucral scales in 2 rows; receptacle flat; chaff awn- like or bristle-like. HEtropPsis. Rays yellow, fertile; achenes 4-angled. ECHINACEA. Rays rose-colored. pistillate, sterile: achenes 4-sided. RupsBeck1a. Rays yellow, neutral: achenes 4-sided, marginless. LEPACHYS. Rays yellow. neutral: achenes flattened laterally and wargined. HELIANTHUS. Receptacle flat or convex: achenes flattish, neither margined nor winged; pappus of 2 deciduous scales or awns. ACTINOMERIS. Receptacle convex; achenes laterally flattened, 2-awned; leaves decur- rent. CorREopPsIs. Receptacle flat; achenes flat; pappus of 2 short teeth or awns, or a mere border, or none. : BIDENS. Receptacle flat; achenes flat: pappus of 2-6awns or teeth which are hispid or barbed. : a Tribe Helenieae. Similar to the preceding tribe: receptacle not chaffy; pappus a row of chaffy scales, bristly-dissected in Dysodia; style-branches truncate or with hairy tips. HELENtwM, Receptacle oblong; rays 3-5-cleft; involucral scales distinct; leaves decur- rent. Dysupia. Receptacle flat; rays few, short; involucral scales united into a cup; ill- scented plants. Tribe Anthemidece. Rays mostly white or none; involucral scales im- bricated, usually dry and scarious; pappus a short crown or none. Strong- scented herbs with alternate leaves. * Receptacle chaffy. ANTHEMIS. Heads large; achenes terete. ACHILLEA. Heads small; achenes flattened. 74 COMPOSITAE * * Receptacle naked, sometimes huiry. CHRYSANTHEMUM. Heads large, many-rayed; pappus none. TANACETUM. Heads small, corymbed, rayless; flowers yellow; pappus a short crown. ARTEMISIA. Heads small, racemose, spicate or panicled, rayless; pappus none. Tribe Senecioneae. Leads radiate or discoid; involucral scales little if any imbricated, not scarious; receptacle naked; pappus of soft capillary bristles. SENECLO. Flowers yellow; heads usually radiate CACALIA. Flowers white or cream-colored, all perfect; heads discoid; sap often milky. ERECATITES. Flowers whitish. the marginal pistillate, the central perfect; heads dis- coid. Tribe Cynareae. Heads large, discoid; flowers tubular, perfect; involu- eral scales much imbricated; anthers long-tailed at the base, elongated-ap- pendaged at the tip; style-branches short or united, smooth, often witha pubescent ring below. ARCT1uM. Leaves not prickly; involucral scales hooked; pappus of rough bristles. Cnicus. Leaves prickly; pappus-bristles plumose. VERNONIA Schreb. Ivon-weed. Perennials. with leafy stems, and corym- bose cymes. Leaves alternate, acuminate Heads discoid, 15-40-flowered: flowers purple, tubular, perfect. Invulucre shorter than the flowers, of many imbricated scales, the inner longest. Pappus double, onter seale-like. inner copious. capillary. Receptacle naked. Achenes cylindrical, ribbed. Species are with difficulty determined. V. fasciculata Mx. Stem 3~7 feet high, glabrous; leaves narrowly lanceo- late to linear, sharply denticulate; heads many, cyme close, involucre bell- shaped, half as long as the showy flowers; scales obtuse, closely appressed. Low grounds; August-September; common. V. noveboracensis (L.) Willd. Stem 3-6 feet high, pubescent or glabrate, leaves lanceolate, serrulate; involucre purplish, scales ovate or lance-ovate, with filiform or subulate spreading tips. Moist svil; July-September; re- ported from Scott, Union, Pottawattamie, and Harrison counties. Specimens from Des Moines, Lee, and Cass counties referred to this species. V. glauca (L.) Britton. Stem 2-5 feet high, glabrous, or finely pubescent; leaves thin, the lower oval or obovate, the upper ovate-langeolate, acute or acuminate, pale beneath, sharply serrate; inflorescence loosely branched; heads few, slender-pedunclea, 10-20-flowered; involucral scales acute or acu- minate, appressed. ‘Woods, August-September; reported, but questionable. (VY. noaveboracensis latifolia Gray.) V. baldwinii Torr. Stout, 2-5 feet high, densely tomentose-pubescent; leaves lanceolute or oblong-lanceolate, acute or acuminate, sharply serrate, scabrous above, tomentulose beneath; heads stout-peduneled; involucral scales acute or acuminate, squarrose. Dry soil; July-September. Forms from Van Buren, Appanoose, Decatur, and Ringgold counties referred to this species. EUPATORIUM L. Perennials. Flowers in corymbose heads, white, blue, or red, never yellow, all tubular, 4-cleft. Involucre imbricate. Receptacle flat, naked. Pappus a single row of capillary scabrous bristles. Style ex- serted, deeply cleft. Achenes 5-angled. E. purpureum L. Jue-Pye Weed. Stem simple, 8-8 feet high, often pur- plish, pubescent above, mostly glabrous below; leaves 3-6 ina whorl, short- petioled, large, ovate-lanceolate to lanceolate, coarsely serrate, pointed; flow- ers flesh-color; involucral scales numerous, imbricated in several rows, Low gounds; August-September; common. ‘ THISTLE FAMILy. 75 E. altissimum L. Stem simple, 2-6 feet high, soft-pubescent, leafy; leaves opposite, sessile. lanceolate, tapering both ways, conspicuously 3-nerved, en- tire or toothed beyond the middle; involucral scales oblong, shorter than the heads, in about two rows, outer shorter; heads 1-5-flowered. Dry ground; August-September; frequent; eastern and southeastern counties, sparingly westward; Winneshiek, Benton, Iayette, Muscatine, Des Moines, Henry, Johnson, Story, Decatur, and Harrison counties. E. serotinum Mx. Stem 3-6 feev high, usually branched, soft-puberulent, leaves mostly opposite, a few upper alternate, long-petioled, ovate-lanceolate, 3-nerved, coarsely serrate; heads 12-15-flowered, in compound corymbs; ‘in- volucre pubescent; scales obtuse. Low ground; August-September, infre- quent; Fayette, Scott, Muscatine, and Woodbury counties. E. perfoliatum LL. Boneset. Stem 2-4 feet high, hairy, corymbosely branched; leaves opposite or in 3's, connate-perfoliate, lanceolate, pointed, crenate-serrate, downy beneath; heads 10-15-flowered, in a dense corymb, scales narrowly lanceolate, shorter than the flower. Low grounds; August; common. The number of flowers in the head and the connate character of the leaves vary greatly. E. ageratoides L. White Snake-root. Stem smooth, simple or branched, 1- 3 feet high; leaves opposite, petioled, thin, broadly ovate, pointed, 3-nerved, large toothed; heads 10-20-flowered, in a compound corymb, involucral scales in one row, oblong, obtuse. Woods; July-September; common. E. coelestinum L. Mist-flower. Plant pubescent or puberulent, 1-3 feet high; leaves opposite, ovate, petioled, acute or obtuse, crenate-dentate, base truncate or slightly cordate; heads cymose-corymbose; flowers blue or violet. Moist soil; August-September; reported from Lee county by Arthur. KUHNIA L. Perennials. Heads paniculate-corymbose, cream colored, dis- eoid, 10-20-flowered. Involucral scales narrowly linear, loosely imbricated, parallel nerved. Pappus a row of white plumose capillary bristles. Achenes cylindrical, 10-striate. K. eupatorioides L. Stem 2-3 feet high, erect, usually simple, minutely pubescent,-leaves alternate, the upper sessile, the lower short-petioled, lanceo- late or linear-lanceolate, entire or sparingly dentate, acuminate. Dry soil, August-September; common. ‘ K. glutinosa Ell. Usually taller and stouter than the preceding, pubes- cent or tomentulose; leaves all sessile, lanceolate to ovate-lanccolate, sharply serrate, those of the branches linear-lanceolate and entire; heads numerous, densely clustered in cymes. ‘Dry soil; August-September; reported from Fayette and Story counties; probably common. (K. eupatorioides corymbulosa T. & G.) LIATRIS Schreb. BurTron SNAkgeRooT. Buiazine-STaR. Perennials, from a roundish corm, erect, simple, leafy. Leaves long-linear, entire, rigid, ses- sile, or lower oblong-lanceolate or oblong-ovate, petioled. Heads discoid, few to many-flowered. Involucral scales well imbricated in many rows. Recep- tacle naked. Pappus of from 15-40 capillary bristles. Achenes slender, tap- ering, 10-ribbed. : * Tnwolucre cylindric or turbinate, 15—16-flowered. L. squarrosa (L.) Willd. Stem 1-3 feet high; frequently hairy; leaves rigid, long-linear; heads few, racemose, about an inch in length; involueral scales tapering to an acute point, leaf-like, spreading; corolla-lobes hairy in- side, pappus plumose. Prairie soil; August-September; Scott, Wapello, 76 COMPOSITAE. Decatur, Taylor, Page, and Cass counties; common in southern Iowa. (La- einaria squarrosa (L.) Hill.) L. cylindracea Mx. Stem 6-18 inches high; heads few, oblong-eylindrical, 1; inch long; scales ovate, with appressed mucronate tips; pappus plumose; corolla-lobes hairy inside. Prairies; Adgust-September; frequent, but appar- ently local; Fayette, Jones, Muscatine, and Story counties. (Lacinuria cylin- dracea (Mx. ) Kuntze ). : ** Involuere hemispheric; 15—45-flowered. L. scariosa (L.) Willd. Stem 3-5 feet high, scabrous-pubescert; leaves lanceolate, lower oblong-lanceolate and long-petioled; heads large, short, re- motely racemose, few or many, 25-40-flowered; involucre hemispherical; seales obovate, obtuse, margins scarious, often colored; pappus barbellate. Prairie: August-September; common. (Lacinwria scariosa ( L.) Hill.) L. pycnostochya Mx. Stem 3-5 feet high, hairy or smoothish; upper leaves linear-lanceolate; heads small, many, few-flowered, ina dense spike, 6-20 inches long; seales oblong or lanceolate with recurved or spreading pet- al-like tips. Low prairies; August-September; common. (Laeinaria pycro- stachya (Mx. ) Kuntze). L. punctata Hook. Stem 4-30 inches high, mostly glabrous; leaves nar- rowly linear, punctate, rigid; heads 3-6-flowered, usually many ina dense spike; scales oblong, acuminate, punctate; pappus very plumose; corolla-lobes naked. Dry sandy knolls; August-September; Fremont, Harrison, Woodbury, Lyon, Emmet, Hancock, and Dickinson counties. ( Lucinaria punctata ( Hook.) Kuntze ). GRINDELIA Willd. Coarse perennial glabrous herbs. Leaves alternate, sessile or clasping. Heads large, terminating the branches, radiate or dis- coid, many-flowered; rays pistillate. Involucral seales closely imbricated in several series, narrow-acuminate, green-tipped. Pappus of 2 or more awns. Achenes short, thick. glabrous. G. squarresa (Pursh) Dunal. Stem 1-3 feet high, leafy; leaves alternate, sessile, linear or spatulate-oblong, serrate; leaves and heads balsamic-viscid; rays pistillate, wanting in var. NUDA Gray. Waste places; August; infre- quent; Woodbury, Sioux, Lyon, Ida, Palo Alto, Muscatine, Henry, Lee, Ma- baska, Boone, and Fremont counties. CHRYSOPSIS Nutt. Perennials, with hairy stems, and large heads ter- minatiug the branches. Rays many, yellow. pistillate. Involucral scaleslin- ear, imbricated. Receptacle flat. Pappus double, outer of short racher ehaft- fy bristles, inner of long capillary bristles. Avhenes linear, flat, obovate or linear-oblong. C. villosa (Pursh) Nutt. Stem about 1 foot high, hairy, corymbosely branched; leaves alternate, oblong-spatulate, entire, hairy. Dry soil; Au- gust; frequent; Lyon county. APLOPAPPUS Cass. Mostly perennials. Leaves alternate. Involucre hemispherical, of many closely imbricated scales. Receptacle flat. Heads many-flowered, radiate; rays pistillate, yellow. Disk flowers yellow. Pap- pus of simple bristles. Achenes short, top-shaped or linear. A. spinulosus DC. Perennial; stem 4-12 inches high, usually spreading; leaves pinnately or bi-pinnately parted, lobes bristle-tipped; achenes ptibes- cent. Loess mounds, forming dense patches; April-September; Harrison, Monona, and Woodbury counties. (Hriocarpum spinulosian (Nutt.) Greene.) re THISTLE FaMiLy. (7 SOLIDAGO L. GoLpEN-Rop. Perennials, with slender erect stems, near- ly sessile, entire or serrate, alternate cauline leaves, and racemose or cluster- ed, yellow, rarely white, heads. Involucral scales appressed, oblong, rarely with green tips. Receptacle small, not chaffy, alveolate, fimbrillate in the last. Rays pistillate, few to many. Disk flowers perfect, yellow. Pappus simple, capillary, seabrous. Achenes teretish, ribbed. Flowers appear in August and September. * Inflorescence in ihe axils of feather-veined leaves. S. latifolia L. Stem 2-3 feet high, angular, ziz-zag, smooth or downy a- bove; leaves broadly oval, thin, deeply serrate, acuminate. base tapering into «w margined-petiole, pubescent beneath; flowers in short axillary clusters, the terminal racemose; scales appressed; rays 3-4. Rich woods; July-September; frequent; AlNlamakee, Fayette, Delaware, Jackson, Johnson, Henry, Des Moines, Story, Decatur, Winnebago, and Emmet counties. (S. flexicaitlis L. S. bicolor L. Stem 1-2 feet high, simple, hairy; leaves elliptical to oblong, acute both ways, the lower slightly serrate, tapering into a petiole; flowers in sinell axillary clusters, rays 6-12. white; involucral scales obtuse. Woods; in- frequent; Johnson and Winneshiek counties. * ® Tnjlorescence a terminal panicle. + Leaves not triple-ribbed; rucemes not secund, S. speciosa Nutt. Stem 3-6 feet high, simple, smooth; leaves thickish, the upper oval or oblong, smooth, rough-margined, lower ovate or broadly o- val, sub-serrate, petioles margined; panicle ample, thyrsoid, rough-pubescent, seules oblong; rays 5-8. Upland borders; common in eastern Iowa, rarer westward. S. rigidiuscula (T. & G.) Porter. Stem rather strict, 24 feet high, gla- hrous below, rough-pubescent above; leaves lanceolate or oblong-lanceolate, entire or the basal crenate, margins ciliolate, sessile-or the lower petioled. Prairies; common throughout western Iowa, but the range is not well known as the species has been confused with the preceding. (8. speciosa angustata Gr.) + + Leaves not triple-ribbed; raccines secund,. S. patula Muhl. Stem 4-5 feet high, smooth, angular-striate; leaves am- ple, ovate, acute, serrate, very rough above. smooth and veiny beneath; heads rather large, in short racemes, on spreading branches; scalesobtuse. Swamps, Cedar river region; Muscatine county. S. ulmifolia Muh]. Stem 2-4 feet high, glabrous; branches pubescent; leaves elliptic-ovate or oblong-lanceolate, tapering both ways, thin, veiny and soft-hairy beneath, sharply serrate or the upper entire; branches of the pan- icle long, slender, recurved; involucral scales lanceolate-oblong; ray and disk flowers 3-4. Woods; common; variable; Winneshiek, Jackson, Johnson, Hen- ry, Des Moines, Decatur, and Taylor counties; reported from Fayette, Scott, aud Story counties. t + + Leanes more or less triple-ribbed; stem glabrous. panicle sometimes pithescent. S. missouriensis Nutt. Stem 2-3 feet high. simple, smooth throughout; leaves linear-lanceolate, tapering both ways, entire or sparingly denticulate, rough-margined, the lower frequently lanceolate, serrate; racemes short, dense: involucral scales obtuse, thick. Dry woods and prairies; common; flowering earlier than the other species; Muscatine, Johnson, Story, Decatur, Ringgold, Page, Fremont, Cerro Gordo, Dickinson, and Lyon counties; repor- ted from Fayette, Monona, and Harrison counties. 78 COMPOSITAE. S. serctina Ait. Stem 3-6 feet high, glabrous, often glaucous, branched above; leaves lanceolate, taper-pointed, sharply serrate above the tapering base, smooth on both sides, rough ciliate; panicle ample, pubescent: involu- cral scales linear, obtuse; rays long. The var. GiGANT#zA Gray has the leaves more or less pubescent beneath. Open woods and borders; common; Winne- shiek, Clayton, Dubuque, Muscatine, Johnson, Henry, Des Moines,.Decatur, Taylor, Winnebago, Dickinson, and Emmet counties; reported from Fayette, Story, Hamilton, and Woodbury counties. S. rupestris Raf. Stem slender, glabrous, 258 feet high; leaves thin, linear-lanceolate, tapering both ways, entire or sparingly serrate; heads small; rays short, 4-6. Loess mounds; Woodbury county, reported by Pam-. mel, also reported from Seott county by Nagel and Haupt. ++ + + Leaves more or less triple-ribbed; stem pibescent. S. canadensis L. Stem 3-5 feet high, pubescent; leaves lanceolate, acu- minate, sharply serrate or nearly entire, pubescent beneath, rough above; heads numerous, small; rays short. Borders and fields; common; varies greatly; Winneshiek, Clayton, Dubuque, Floyd, Delaware, Johnson, Henry, Decatur, Calhoun, Winnebago, and Emmet counties; reported from Fayette, Scott, Story, Hamilton, Dickinson, Monona, and Woodbury counties. ‘ S. nemoralis L. Stem 6-24 inches high, cinereous-pubescent, simple or corymbed above; leaves spatulate or oblanceolate, entire or the lower some- what serrate and tapering into a petiole; heads small, in numerous usually recurved racemes; involucral scales linear-oblong, appressed. Dry soil; com- mon; Winneshiek, Dubuque, Floyd, Delaware. Johnson, Henry, Des Moines, Decatur, Story, Calhoun, Winnebago, Dickinson and Emuinet counties; re- ported from Fayette and Hamilton counties. *** Heads in compound corymbs or in. cymose clusters. +t Stem densely pubescent. S. rigida L. Stem 2-4 feet high, stout, leafy, softly pubescent or scabrous; Jeaves oval or oblong, thick, rigid. entire or obscurely serrate, feather-veined; lower leaves oblanceolate, tapering into slender petioles; heads numerous, large, many-flowered; involucral scales very obtuse; rays 7-10. Prairies; common; Winneshiek, Floyd, Johnson, Decatur, Fremont, Jasper, Winne- .bago, Emmet, and Lyon counties; reported from Fayette, Seott, Story, Hamil- ton, Monona, and Woodbury counties. + + Stem glabrous or nearly so. S. riddellii Frank. Stem 1-3 feet high, glabrous, stout; leaves long and narrow, entire, froma sheathing base, mostly recurved, radical leaves very long, tapering into long petioles; heads numerous, many-flowered, in a dense corymb, the branches and pedicels rough-pubescent; rays 5-7. Low prairies, sloughs, often growing in water; infrequent; Muscatine, Floyd, Hancock, and Emmet counties; reported from Story and Hamilton counties. S. lanceolata L. Stem from a long and slender rootstock, 1-3 feet high, much branched, usually smooth; leaves narrowly lanceolate, sessile, 3-nerved, heads small, in sessile glomerate clusters disposed in flat-topped corymbs; rays small, 15-20;'late flowering. Sandy or wet soil, pastures and copses common; Johnson, Henry, Decatur, Dickinson, and Emmet counties; reported from Fayette, Scott, and Story counties. (Huthamia greminifotia (L.) Nutt.) BOLTONIA L’Her. Bushy-branched glabrous perennials, with striate angled stems, entire leaves, and Aster-like flowers. Heads mauy-flowered, radiate; rays many, pistillate. Scales in two series, appressed, with scarious TrmisTLE FAMILY. 79 margins, Receptacle convex, punctate. Pappus of minut2 bristles and ofteu 2-3 awns. Achenes flat, obovate, 2-3-winged. B. asteroides (L.) L'Her. Stem 1-6 feet high; leaves long-lanceolate, en- tire; involueral scales acuminate. Movist soil, sloughs and low places; July- September; frequent; Muscatine, Johnson, Ilenry, Lee, Decatur, Ringgold, Taylor, Winnebago, and Emmet counties; reported from Story, Harrison, Mo- nona, and Woodbury counties. ASTER L. Aster. Perennials, with alternate leaves, and panicled, cor- yinbed, or racemose heads. Disk-flowers perfect, yellow to purplish; rays pistillute, 3-toothed at the apex; white, blue, or purple, never yellow. In- volucral scales louse, imbricuted. often with green tips. Pappus simple, or double in the last, capillary. Receptacle flat, alveolute. Achenes compressed. Common in late summer and autuinn. : A. Pappus simple, of capillary bristles. * Lower and bos.el leeves cord tie and slendcr-petioled, the stem leaves wot cordate- clauspiing. + Rays white, vivlet, or rese. A. corymbosus Ait. Stem 1-2 feet high. nearly smooth, somewhut flex- uous: leaves umple. ovate, acuminate, thin, coarsely serrate, all but the up- permost with a heart-shaped base. and on long wingless petioles; heads ob- long, in small open curymnbs; rays 6-9, white. Rich woods; Muscatine county. (A. dimericutus 1. ) . A. macrophyllus lL. Stem 2-3 feet high, rigid, rough-pubescent: leaves thickish, rough, serrate. pubescent beneath, the upper ovate or oblong, ses- sile or on margined petioles, the lower cordate and long-petioled; heads one- half inch high, larger than the preceding; scales obtuse, the inner longer and thioner. Lilly woodlands, not rare; Muscatine county. ++ Rays blue or purple, rerety white. t Leaves entire or nearly so, firm, or thick. A. azureus Lindl. Stem 1-3 feet high. rigid, roughish, panicalately branched; leaves rough, the radical and lower cauline leaves lanceolate, more or less with corcate bases, on long petioles. the upper lanceolate to linear or subulate, wing-petioled or with attenuated base to sessile; rays violet blue; involucre obeonic; scales with abrupt green tips, appressed. Prairies and thickets; frequent; Johnson, Story, Des Moines, Appanoose. and Decatur counties. A. shortii look. Stem 2-4 feet high, rough-pubescent; leaves lanceolate or ovate-lanceolate, long-acuminate, entire, on slender naked petioles, base deeply cordate, smooth above, minutely pubescent beneath, the upper leaves sessile; racemes paniculate; involucre campanulate; scales with green tips. Cliffs and banks; infrequent; Winneshiel, Muscatine, Johnson, and Lee coun- ties, reported from Fayette county. t { Leaves mostly sharply scrrate. thin. A. cordifolius L. Stem 1-3 feet high, smooth or slightly pubescent, rarely hairy, paniculately branched above; leaves ovate-cordate. sharply serrate, on long slightly margined petioles, or upper lanceolate and sessile; heads pan- icled; rays pale blue or white; involucre obconie, scales appressed, tipped with short green points. W oods; common; Winneshiek, Johnson, and Decatur counties, reported from Fayette, Story, and Des Moines counties. A. ‘sagittifolius Willd. Stem 2- 4 feet high, rigid; leaves ovate-lanceolate, pointed, serrate, pubescent beneath, the lower with the base truncate or cor- 80 COMPOSITAE. date and wing-petioled, the upper linear or lanceolate, sessile; heads race- mose, scales loose, linear-subulate. Woods; common; Fayette, Delaware. and Johnson counties; reported from Winneshiek, Allamakee, Clayton, Dubuque, Des Moines, Henry, Plymouth, and Monona counties. A. drummondii Linal. Stem stout, 2-4 feet high, pale, grayish pubescent; leaves ovate or ovate-lanceolate, acuminate, rough above, canescent beneath the lower and basal cordate, with slender naked petioles, the upper with cor- date or rounded bases and margired petioles, those of the branches sessile. Dry soil, open woods; infrequent; \Vinoeshiek, Delaware, Muscatine, Henry, Des Moines, Johnson, Jasper, Winnebago and Dickinson counties, reported from Story and Floyd counties. ** Leaves not cordate or petinled, some or all of tre cuuline with corde or auriculute clasping biwses. + Stem rouyh, or hirsute-pubescent. t Leaves entire, oblong, linear, or Lancevlate. @ Leaves sessile, stronyly cordate-clasping. A. patens Ait. Stem 2-4 feet high, rough-pubescent, loosely paniculate above; leaves ovate-oblong to oblong-lanceolate, cordate-clasping; heads large, termiral; rays many, violet-blue; scales slender, tips acute, spreading. Forms from Delaware county have been referred to this species. A. phicgifolius Muhl. Stem 2-4 feet high, rough-pubescent; leaves lan- ceolate to oblong-lanceolate, eutire, acuminate, auriculate-clasping, much contracted below the middle, thin, rongh above, pubescent beneath; rays many, purple blue. Woods; Winnebago county. (A. patens var. phloyifolius Nees.) A. novae-angliae L. Stem 3-7 feet high, stout, hairy, leafy, corymbosely branched; leaves lanceolate, auriculate-clasping, entire, acute; rays many, violet-purple, rarely varying to rose-purple, or white; scales nearly equal, green throughout, narrowly linear. Sloughs and low places; common; Win- neshiek, Floyd, Delaware. Johnson, Jefferson, Des Moines, Appanoose. De- cetur, Story, Winnebago, Lyon, and Emmet counties; reported from Fayette and Monona counties. / 2% Leaves slightly cordate claspiny. A. oblongifelius Nutt. Stem 1-2 feet high, much branched, hirsute-pu- bescent, leafy; leaves narrowly oblong to oblong-lanceolate, entire, clasping, rough or hispidulous on both sides, rough-margined; rays violet purple; invo- lucre hemispheric, the bracts glandular. Prairies and rocky bluffs; infre- quent; Fayette, Muscatine, Des Moines, Johnson, Pottawattam‘> Calhoun, and Lyon counties; reported from Story, Monona, and Woodbury counties. A. amethystinus Nutt. Resembles the preceding, stem taller, 2-5 feet high, hoary puberulent or somewhat hirsute; leaves linear-lanceolate, rough on both sides, partly clasping; involucral scales erect, the green tips merely spreading, involucre turbinate, the bracts hispid. Moist places, searee; re- ported from Floyd and Stcry counties. ¢ t The lower and must of the upper leaves serrate. A. puniceus L. Stem stout, 3-6 feet high, hispid; leaves oblong-lanceo- late, auriculate-clasping, acuminate, coarsely serrate to sparingly denticulate or sometimes entire; heads rather jurge, sparsely paniculate or thyrsoid- erowded; involuere of loose, thin, equal, linear seales, arranged in tivo rows; rays ¥% inch long, showy, violet, varying to purple or white. Swampy grounds; frequent; Fayette, and Museatine counties, reported from Story county. THISTLE FaMILy. 81 + + Stem glabrous or sparingly pubescent above. Se t Leave sharply serrate, ee & A. laevis. A. prenanthoides Muhl. Stem 1-3 feet high, slehder, nearly glabrous or hairy above; leaves thin, ovate-lanceolate, taper-pointed, sharply serrate in the middle, tapering into a slender, elongated, entire portiun towards the base, upper surface rough, lower sinooth, base auriculate-clasping: heads about 4 ineh high, on short divergent peduncles, involueral scales narrow, somewhat spreading; rays light blue or whitish. Rich woods; frequent; Jackson, Delaware. Muscatine, and Johnson counties, reported fram Vayette and Story counties. A. laevis L. Stem 2-1 feet high. smooth, often glaucous; leaves lanceolate or oblong-laneeolate, entire or sub-serrate, thickish, radical ard lower leaves large, wing-petioled, the upper partly clasping by an auriculate base; heads chiefly solitary on the branchlets; involucre hemispherical. scales with broad, acute, appresse., green tips. Prairies and rocky woods; frequent and widely distributed; Winneshiek, Fayette, Floyd, Delaware, Johnson, Henry. Deca- tur, Winnebago, [uyon, Dickinson, and Emmet counties, reported from Story and Hamilton counties. t £ Leaves entire or nearly so. A. longifolius Lam. Stem 1-3 feet high, glabrous.or somewhat pubes- cent, branched; leaves lung-lanceolate to linear-lanceolate. entire or nearly so, acuminate, base sessile, clasping; heads numerous, about an inch broad, the scales narrow, green, acute, in 1 or 2 rows. Moist soil; specimens from Delaware and Henry counties referred to this species *** Leaves petioled or sessi’e, not claxping or scarcely so. + Leaves entire, silvery, silky or canescent or both sides. A. sericeus Vent. Stem 1-3 feet high. smooth, very rigil, brauched; leaves lanceolate or oblong, sessile, entire, silvery white on both sides; heads large, mostly terminal, on slender branches; rays many, violet-purp'’e; scales leaf-like, tips spreading. Common on the prairies in western and southern Jowa, infrequent eastward; Winneshiek, Delaware, Muscatine,, Johnson, Decatur, Ringgold, Taylor, Page, Fremont, Montgomery, Pottawattamie, Winnebago, Lyon, and Emmet counties; reported from Fayette, Story, Buena Vista, Monona, Harrison, and Woodbury counties. t ¢ Leaves often toothed, neither silvery, silky nor canescent. t Heads corymbose. A. ptarmicoides-(Nees) T. & G. Stems simple, tufted, 8-20 inches high, smocth or sparingly pubescent; leaves linear-lanceolate, 1-3-nerved, acute, entire, rough-margined, lower tapering to a short petiole; heads in a flat corymb; involucral scales imbricated in 3 or 4 rows, appressed, without green tips; rays white. Sandy soil; Muscatine, Emmet, and Lyon counties. t ¢ Heads solitary at the ends of the branchlets. A. dumosus L. Stem 1-3 feet high, glabrous, paniculately branched; leaves linear or linear-lanceolate, entire. acute or obtusish, rough-margined, those of the branches small and bract-like: heads many, terminating the branches and branchlets; involucre campanulate, the scales in + rows, linear- subulate, obtuse or acutish, appressed, green-tipped; rays 15-30, white to pale violet. Sandy soil; reported from Benton county by Arthur; specimens from Lee county have been referred to this species. ttt Heads paniculate, not in 1-sided racemes. A. salicifolius Lam. Stem 2-5 feet high, slender, paniculately branched, leafy, glabrous or slightly pubescent above; leaves lanceolate or linear-lan- 82 COMPOSITAE. ceolate, sparingly dentate or entire, rough-margined, acute or acuminate. sessile or somewhat clasping, the lowest sometimes petioled; beads pumer- ous, rather large; involucre turbinate, its braets linear-oblong, green-tipped, acute or obtusish, appressed, in 4 or 5 series; rays many, violet, purplish, or white. Low moist soil; common; Johnson, Henry, and Deeatur counties. A. paniculatus Lam. Stem 2-8 feet high, glabrous. paniculately branched; leaves lanceolate or oblong-lanceolate, acuminate, base narrowed and sessile or clasping, sparingly serrate to nearly entire; heads many, rather broad; involucre hemispheric, the bracts linear-lanceolate, acute or acumi- nate, appressed, green-tipped, in 4 or 5 rows; rays many, white or purplish. Moist soil; Fayette county, reported from Story, Woodbury, Lyon, Lee, and Henry counties. A. tradescanti L. Stem 2-5 feet high, slender, panieulately branched, branches ascending and sometimes pubescent in lines; eauline Jeaves Jancev- Jate cr linear-lanceolate, acuminate, narrowed to the sessile base. entire or more commonly sharply serrate in the middle; heads numerous, racemose; in- volucre hemispheric to broadly turbinate. its braets linear, aeute., green- tipped, appressed. in 4 or 5 rows; rays white or whitish, short and narrow. Fields and low places; specimens from Delaware and Johsun county have been referred to this species. A. ericoides L. Stem 1-3 feet high, bushy, paniculately branched, gla- brous; leaves firm or rigid, the basal spatulate, obtuse, dentate. with mar- gined-petioles; cauline Jeaves linear, entire, acute, the uppermost linear-subu- late; heads many; involucre hemispherie to campanuiate, the braets green- tipped, linear-lanceolate or lanceolate, in about 3 rows; rays 15-25, white or rosy tinged. Dry soil; Henry county, reported from Lee and Monona coun- ties. ttt + Heads racemose and usually t-sided on the branches. ‘A, diffusus Ait. Stem 1-4 feet high, much branched. pubescent or nearly glabrous; leaves lanceolate or oblong-lanceolate. tapering both ways, sharply serrate in the middle or the smaller entire, sessile, thinnish; heads disposed singly or in clusters along the flowering branches; rays short, white or some- times purplish; involuecral scales linear, obtuse or acutish. Low woods; eom- mon; Story, Fayette, Delaware, Johnson, Henry, Appanoose, and Decatur counties. (A. leteriflorus (L.) Britton). A. multifforus Ait. Stem about a foot high, bushy-braneched, grayish, pubescent; leaves small, linear, entire, sessile or partly clasping; heads small,crowded in numerous racemes; herbaceous tips of the invulucral seales spreading; rays white, rarely bluish. Sandy soil, woods and prairies: eom- mon; Winneshiek, Fayette, Floyd, Delaware, Scott, Muscatine, Des Moines, Henry, Jefferson, Johnson, Story, Hamilton, Appanoose, Decatur, Ringgold, Taylor, Harrison, Woodbury. and Emmet counties. B. Leuves petivled or sessile, entire; pappus double, the inner bristles long, the outer shorter. ; A. umbellatus Mill. Stem 2-8 feet high, glabrous or pubescent above. striate, corymbosely branched; leaves lanceolate to oblong-lanceolate, taper pointed, base narrowed into short petioles. upper sessile, usually pubescent on the veins beneath, entire, margin hispid; heads many, in compound flat corymbs; involucre campanulate or hemispheric, the scales acutish or obtus- ish, in 8 or 4 rows. Low grounds; Johnson and Story counties, reported from THISTLE FAMILY. 83 Fayette county. (Doellingeria wmbelluta (Mill.) Nees.) The variety ruBENS Gray has the lower surfaces of the leaves pubescent and is reported from Story county. (D. wmbellate pubens (Gray) Britton). ERIGERON L. Frranane. Herbs, with alternate simple leaves, and soli- tary or corymbose pedunculate heads. Ray-flowers many, pistillate, white or purple; disk-flowers yellow. Involucral scales narrow, equal, scarcely im- bricated. Receptacle flat or convex. Pappus of capillary bristles, mostly simple. Achenes compressed,:?-nerved, frequently pubescent. * Hewds smull, rays short, iaconspicuous, in several rows. E. canadensis L. Horse-weed. Stem 1-5 feet high, erect, hairy; leaves linear to linear-lanceolate, usually entire. radical cut-lobed; heads small, numerous, paniculate; rays short, white. Waste places; July—October; com- mon. (Leptilon canadense (L.) Britton). E. divaricatus Mx. Stems low, 3-15 inches high, diffuse and decumbent, hairy; leaves linear, entire; heads small, corymbose; rays purple. Prairie or o- pen woods; June-August; common; Emmet, Story, Johnson. Muscatine, Appa- noose, Decatur, Ringgold, and Fremont counties. (L. divuricatum (Mx.) Raf.) ** Heads larger, rays elongated, conspicuous, in one or more rows. E. annuus (L.) Pers. Daisy Fleabune. Stem 2-4 feet high, branching, hirsute; leaves ovate to lanceolate, sessile or lower un margined petioles, coarsely and sharply toothed; rays numerous, purplish. Fields and waste places; June-August: common. E. strigosus Muhl. Daisy Fleahune. Stem 2-3 feet high. branched, ap- pressed pubescent or nearly smooth; upper leaves lanceolate. lower oblong or spatulate, 3-nerved, tapering into a petiole. all mostly entire; rays white. Fields; June-August; common. (E£. ramosus (Walt.) B.S. P.) E. philadelphicus L. Commun Fleabane. Stem about 2 feet high, hairy; leaves clasping by a heart-shaped base, ovate, or lower spatulate, entire or remotely toothed, thin; heads few, corymbed; rays rose-purple. Upland woods; May-July; common. E. bellidifolius Muhl. Robin's Plantain. Stem 1-2 feet high, hairy, sim- ple; radical leaves obovate or spatulate, sparingly tootred, stem leaves re- mote, lance-oblong, clasping, entire: heads few, large, on slender peduncles; rays 50 or more, linear-spatulate, rather broad, purple. Woods; May-June; infrequent. Winneshiek. Iayette,. Muscatine, Delaware, Johnson, Henry and Jackson counties. (E. ptilchellus Mx.} ANTENNARIA Gaertn. Perennial white-woolly herbs, with alternate en- tire leaves, and corymbed heads of dicecious or polygamo-dicecious flowers. Heads many-flowered. discoid. Scales of the involucre imbricated, white or colored, dry and scarious. Receptacle sub-convex, foveolate, not chaffy. Pappus a single row of bristles, in the pistillate flowers capillary and united at the base, in the staminate flowers thickened and barbellate above. Fertile corollas slender. Achenes terete or flattish. A. plantaginifolia (U.) Richards. Stem 4-18 inches high, simple, with rupning stolons, forming patches, radical leaves spatulate to obovate or oval, 3-nerved, petioled, cauline leaves sessile, oblong or lanceolate to linear, the upper small, bract-like; heads in a close terminal corymb. Vields and upland woods; April-May; common. ANAPHALIS DG. White-woolly perennials, with many of the characters of the preceding genus. Fowers dicecious, usually the pistillate heads with a few perfect but sterile flowers in the center. Pappus in the pistillate flowers BE COMPOSITAE. not united at the base, of copious capillary: bristles, in me staminate flowers not thickened above. , NS A. margaritacea (L.) Benth. & Hook. Stem 1-2 feet bigh. simple, corym- bosely branched above, leafy; leaves nearly linear to broadly Janeeolate, aeu- minate, entire, sessile, becoming green above, the larger obscurely 3-nerved; involueral seales pearly-white, numerous, unequal, imbriecated, obtuse. Fields and open woods; July-August; frequent; Delaware, Johuson, and Deeatur counties; perhaps generally distributed. This species is confused with the following, the general appearance of which it resembles. GNAPHALIUM L. Woolly or pubescent herbs, with sessile alternate leaves, and variously clustered heads of fertile flowers. The pistillate flow- ers in several series, the perfect flowers, central, few. Pappus of capillary bristles. , G. polycephalum Mx. Annual or winter-annual; stem 1-3 feet high, woolly or totnentose; leaves lanceolate or linear-lahiceolate, sessile. glabrate above, white-woolly beneath, margids undulate; heads in corymbose or pani- culate clustets; pappus-bristles distinct. This species is frequently reported. (G. obtusifolium L.) INULA’ L. ‘Coarse paces laa with alteraate | simple Jeaves, and large many-flowered heads. '' Rays'pistillate, yellow. Disk flowers perfect, yellow. Involueré hemispherical, of imbricated'foliaceous sedles. Receptacle naked. Pappus simple, of capillary bristles. Anthers sagittate at the base. the auri- eles caudate. Achenes somewhat 4 or 5-ribbed. I. Hhelenium L. ‘Elecumpune. ‘Stem 2-6 feet high, from w# large thick root: leaves large, oblong or'ovate, woolly beneath,’ entire or deuticulate, radical petioled, cauline clasping; scales ovate. Open, woods;- July-August; loeal; Johnson ‘county; rarely i in waste’ places. Decatur county. POLYMNIA L.* Perennial branching herbs, with opposite or alternate thin lobed leaves,'ahd light yellow flowers. Heads radiate, many-flowered; rays few, pistillate; disk flowers perfect but sterile. Involucral scales in two rows; outer about 5, leaf-like, spreading; . inner smaller, membranaceous. Receptacle flat, chaffy. Pappus Wanting. P. canadensis’L: Stem 2-5'feet high, classe -hairy; leaves’ petioled, the lower pinnatifid, the upper deltoid-ovate or hastate, 3-5-lobed or entire: rays 5, in the typical form shorter than’the ‘involuere, ‘often minute or obortive: achenes 3-ribbed. The form represented in our flora is variety RapIATA Gray, which has’ the rays longer and 3-lobed. Wooded ravines; June-July; frequent; Winpneshiek, panes Clayton, Dubuque, Fayette, Museatine, and Des Moines counties. : SILPHIUM L. Rostn-weev. Coarse perennials, with alternate or opposite leaves, resinous juice, and large yellow-flowered heads. Ray-flowers mny, pistillate, fertile, in 2'or 3 rows, outer ligulate; disl-flowers perfeet, sterile: style entire. Involucral scales foliaceous, spreading, in several rows. Re- ceptacle flat, chaffy. Pappus wanting or of 2 teeth. Achenes broad. flat, dorsally compressed. * Stems terete; leaves allernate. S. laciniatum L. Compuss-plant. Whole plant covered with whitish hairs; stem 3-10 feet high, terete, simple; leaves alternate, pinnately parted; divi- sions linear, cut, sinuate-lobed or entire; petioles dilated at their bases; radical or lower leaves 1-2 feet long; heads large, several, sessile or short pedunéled, mostly alternately arranged along the upper part of the naked TwistLeE FAMILY. 85 stem: scales ovate, tapering into long and spreading points: achenes broadly winged. Prairies; July; frequent throughout the state. S. terebinthinaceum Jacq. Stem 4-10 feet high, branched, sealy. gla- brous or nearly so; leaves basal, large, ovate,"long-petioled, apex acute, base cordate,‘both sides rough, sharply ‘dentate; heads many; achenes obovate, narrowly winged. Prairies and'dry woods; reportéd from:Iowa. f * * Stems square; leaves or their petiole-bases conneate-perfoliate. S. perfoliatum L. Cup-plimt. Stem square, 4-7 feet high, often branched above; leaves large. thin, uvate,"rough, remotely-toothed, opposite or in 3°s, their bases connate, forming a cup around the ‘stem; lower leaves wing- petioled; bases of ' petioles dilated and connate;'heads corymbose.'on long peduncles; scales ovate, achenes ‘winged. “Moist soil; July-August; frequent throughout the state. © ° ge ‘ < ** * Stems tercte or slightly 4-angled; leaves opposite or whorled. S. integrifolium Mx. Stem'4-angled and grooved, 2-4 feet high, smooth or scabrous: leaves opposite or uppermost alternate, ovate-lanceolate. rough- pubescent or smooth, entire. rigid, sessile, - bases “cordate. partly’ clasping heads short-peduneled, in a’ close corymb:achenes broadly winged. Prairies, roadsides, and:open woods. July—Augist: frequent throughout the state. S. trifoliatum:>L. Stem 4-7\feet high, ‘glabrous, sometimes glaucous: leaves lanceolate‘or‘oblong-lancedlate, the 'middle‘ones in whorls of 3 or 4, the uppermost opposite. acuminate,’ narrowed at‘the base, rourh above, gla- brate beneath, short-petioled. entire or somewhat dénticulate; heads several or many; achenes rather broadly winged. Woods; reported from Winnebago and Clinton counties.’ © *: ; Ce PARTHENIUM L._ Perénnials, “with alternate leaves. and’ corymbose heads. Rays 5; ligules white, short; obcordate:' fertile; disk-flowers many, sterile. Involucre hemispherical;'scales in 2 series. ovate’ or roundish: Re- ceptacle conical, chaffy. Achenes compressed, crowned with 2 teeth ahd the remains of the corolla: °-- ) cei : P. integrifokium'L. Prairie Dock, Stem 1-4: feé¢t ‘high, rough-pubescent, mostly simple;" radical‘ or lower leaves long-petioled, ample. ovate. crenite- irene coriaceous, upper sessile:‘ heads many. Upland soil; Jane-Septem- ber: frequent; more common in the eastern and southern counties; Winne- aniale Allamakee, Fayette, Dubuque, ‘Muscatine, Johnson, Henry, eee Decatur: Ringgold, and’ Fremont counties. ~ ' ECLIPTA L.''A rough’ annual, with opposite leaves.'and' small sehen many-flowered heads.” Rays short, white, numerous, fertile; disk-flowers perfect, -4-toothed, fertile. Involucral scales in 2 rows. ovate-lanceolate, ° foliaceous. Receptacle flat. with bristle-like chaff Pappus wanting or’ of 2-4 teeth or short awns. Achenes thick. in the rays et 3-sided, in the disk- flowers compressed. more or less margined.'* ef E. alba L. Hasskarl. Stem procumbent or vaneenaiNE 1-3 feet high.ap- pressed-pubescent; leaves mostly sessile, lanceolate or oblong, tapering both ways, sparingly serrate; rays scareely exceeding the disk. River banks; July-August; infrequent; Johnson. Linn, Muscatine, and Lee counties. HELIOPSIS Pers. Perennials, with opposite petioled 3-nervéd leaves, and yellow flowers. Involucre short, of 2 or 3 rows of imbricated subequal scales. Rays 10 or more, fertile. - Receptacle conical, with linear chaff. Pappus cor- oniform,' of 2-3 teeth; or wanting. *‘ Achenes truncate. smooth, 4-sided. H. scabra Dunal. Rough Ox-cyc. Stem 2-4 feet high, somewhat pubescent “© 86 COMPOSITAE. scabrous; leaves broadly ovate to ovate-lanceolate, scabrous, serrate; rays oblong or linear, 2-3-toothed; achenes pubescent on the angles when young. Prairies and open woods; June-September; frequent throughout the state. H. laevis Pers. Similar to the preceding; stem 3-5 feet high, glabrous; leaves opposite, sometimes in 3's; ovate or ovate-lanceolate, rather thin, acv- minate, sharply dentate, smooth on both sides or roughish above; achenes glabrous, truncate. Woods; July-September; ee ee Muscatine, Deca- tur, and Fremont counties. (H. helianthoides (L.) B. 8. P.) ECHINACEA Moench. Perennials, with seco alternate 3-5-nerved leaves, and single terminal large heads. Involucral seales in 2 or 3 rows, °im- bricated, lanceolate, spreading. Receptacle conical, bristling with spiny-tipped chaff. Rays rose-purple. long, drooping, pistillate but sterile. Disk-tlowers purplish, shorter than the chaff. Pappusa few teeth. wenence thick, short, 4-sided. E. angustifolia DC. Pale Purple Cone-flower. Stem 1-3 feet ria bristly- hairy, simple; leaves lanceolate or linear-laneeolate, entire, tapering both ways, long-petioled, bristly hairy; rays 12-15, 2-3-toothed, 1-2 inches long. Prairies; June-September; common. (Brauneriu pallida (Nutt.) Britton ). E. purpurea Moench. Purple Cone-flower, Stem 2-4 feet high, rough- hairy to nearly smooth; lower leaves long-petioled, ovate, 5-nerved, upper usually 3-nerved, all rough and frequently serrate; involueral-seales in 3-5 rows, lanceolate; rays 15-20, dull purple.’ bifid, 1-2 inches long. Woods; July-August; infrequent; Appanoose, Decatur, and Louisa counties. (B. pur- purea (L.) Britton ). RUDBECKIA L. Biennialsor perennials, with alternate leaves, and termin- al showy many-flowered heads. Involucral scales about 12, in 2 rows, folia- ceous. Receptacle conical or columnar, ¢chaffy. Rays many, neutral, mostly yellow. Disk-flowers perfect. Pappusa toothed or cut margin, or wanting. Achenes 4-angled. R. triloba L. Brown-eyed Susan. Biennial; stem 2-5 feet high, with slender spreading branches; lower Jeaves 3-lobed or 3- “parted, serrate; ‘upper ovate- lanceolate, serrate, acuminate, subsessile; heads small; disk conical, dark purple; chaff awned, ays about 8, oval or oblong. Woods; August-Septem- ber; common. R. subtomentosa Pursh. ‘Stem 3-4 feet high, branching, downy and sca- brous; leaves mostly 3-lobed or 3-parted, downy, divisions ovate or ovate-lan- ceolate, serrate; heads corymbose, short-peduncled; disk globular, dull brown; chaff obtuse, bearded. Prairies and open woods; August-September; fre- quent; Muscatine, Johnson, and Decatur countiés; reported from Fayette and Story counties. ’R. hirta L. Black-eyed Susan. Biennial, bristly-hairy throughout; stem 1-2 feet high, mostly simple; upper leaves oblong ur lanceolate, mostly entire, sessile, lower long-petioled, spatulate, 3-nerved; rays 9-14; disk dull brown; chaff hairy at the apex. Woods and prairies; June-August; common’. R. laciniata L. Stem glabrous, 2-7 feet high; lower leaves 3-lobed or 5-7- parted, divisions usually cut or lobed, ovate-lanceolate, acuminate; heads on long slender peduncles; disk globular in flower, columnar in fruit; rays ob- lanceolate, drooping. Low grounds; August; common. LEPACHYS Raf. Perennial herbs, with alternate pinuately divided leaves, grooved stems, and terminal heads. | Involucral scales linear, in one row. Receptacle oblong or columnar, chaffy. Rays few, neutral, yellow. Disk- flowers perfect. Pappus of 2 teeth or wanting. Achenes compressed, 1-2- winged. TuistLe lAMiiy. ST L. pinnata T. & G. Stems 2-4 feet high, stigose-pubescent. slender. branched above; leaflets 3-7. narrowly lanceolate, entire or remotely serrate; disk ovoid: rays about 2 inches long, light yellow, drooping; uchenes ob- scurely 2-tipped. Prairies: Jaly-Augast; frequent and widely distributed. (Ratibida pinnate (Vent.) Barnhart.) a L. columnaris T. & G. Stem 1-? feet high. strigose-pubescent and scabrous. branched; leatlets 5-9, oblung to linear. entire or 2-3-cleft. heads on long peduneles: disk columnar, 1-2 inches long: rays usually shorter, ovate, vellow. Prairies; August: rare: Lenry, Page, Boone, and Lyon coun- ties. CR. colnueris (Siuas) D. Don). HELIANTEUS L. SunFLower. Annual or perennial herbs. with alternate or opposite leaves, and solitary or corymbed pedunculate heads which termi- nate the stem or branches. Involucre imbricated,, leaf-like. Receptacie plane or convex, with persistent chaff which embraces the achenes. Rays few to many, neutral, yellow. Pappus of » or 4 chaffy scales, wostly decidu- ous. Achenes compressed or 4-angled. * Disk purple. purplish or brown; rveeeptacle flat or conver, t Ours canals; leaves mostly alternate, the lower opposite. H. annuus LL. Common Sunflower, Stem 2-10 feet bigh, usually much branched, hispid or sexbrous: leaves mostly alternate. petioled. ovate or lower cordate, 3-nerved, serrate; involucral seales ovate or oblong, long- acuminate, ciliate; disk 1 inch or more across, Commonly cultivated anda common eseape fh waste places; infrequent in the wild state: July—Septem- ber. ; : H. petiolar:s Nutt, Stem simple or branched. 1-3 feet high, strigose-his- pidulous: leaves ovate-lanceolate or oblong, slender-petioled. usually entire; involucral scales lanceolate or oblong-lanceolate. sometimes ciliate; tips mu- cronate or attenuate: disk about half an inch broad. Dry soil; July-August: vare; Dubuque and Muscatine counties. + 4+-Perennial: leaves mostly opposite. the upper alternate. H. rigidus Desf. Stem 2-4 feet high, simple. sometimes branched, rigid, rough; leaves opposite. % nerved, nearly sessile, oblong-lanceolate or lower oval, entire or somewhat serrate. rigid, rough on both sides. tapering both ways: heads large, usually solitary. terwinal; scales ovate or oblong. acute. ov obtuse. ciliate, appressed; rays 15-25; disk dark. Prairies; August—Sep- tember: frequent; Floyd. Hangock, Johnson, Winnebago. Decatur, Ringgold, ‘taylor, Page. and Fremont counties; reported from Fayette, Seott. Louisa. Story. Woodbury, Sioux, Lyon, Dickinson, and Emmet counties. (H. seaber- rimus EM ) *%* Disk yellow or yellowish; receptacle conic or conver; perennial. + Leaves usally lanceolate, much longer thaw wide. ¢ Leaves mostly basal, the upper bract-lihe. H. occidentalis Riddell. Stem 2-3 feet high, somewhat hairy with run- nets from the base, simple, nearly leafless above; leaves oval: 3-nerved, lower on long huiry petioles, the upper small, nearly sessile and remote: heads small, 1-5. on long peduncles; scales lance-ovate. acute. often ciliate: disk yellow. Prairies; August-September; frequent; Winneshick, Jonnson, and Lee counties; reported from Clinton, Benton, Fayette, Henry, and Muscatine counties. ° : ‘ tt Leaves mostly eauline and alternate or opposite. @ Stem sedbrous, scabrate or hispid; leaves sessile or nearly so. KS COMPOSITAE. H. giganteus LL. [loots fleshy. rootstock creeping: stems hispid or scab- rous, 3-10 feet high, simple or branched above; leaves sessile or short- petioled, laneeolate, scabrous above. hirsute beneath, dentieulate or serrate, mostly alternate but sometimes opposite; heads several, long-peduneled; in- volucre hemispheric, the bracts lanceolate-subulate, ciliate or hirsute. squar- rose. Reported as rare from Fayette county. H. maximiliana Sechrad. Stem 2-8 feet high. stout, hispidulous-seabrous: leaves alternate, some opposite, narrowly lanceolate. tapering both ways, subsessile, entire or nearly so, very scabrous on both sides; heads over medium size, terminating the stem or branches: seales long-attennate; rays golden yellow. Prairies; August-September: infrequent; Emmet and Dnbuyne counties, reported from Ficyd, Story, Buena Vista. Monona, Woodbury, Sioux, Lyon, and Diekinson counties. 4 @ Stem glabrous, ghaucons, learcs petioled. H. grosse-serratus Martens. Stem smooth, glaucous, 3-8 feet high, mostly simple: leaves opposite; petioled, long-lanceolate, Jong-pointed, sharply serrulate or denticulate, whiter and pubescent beneath: scales subu- late; heads many, small, short-peduoeled; rays 15-20. Low grounds; Auguast- September; common. + } Leaves ovate, ovate-tanceotate, or oblong. + Leanes sessile, or nearly so. H. mollis Lam. Stem 2-3 feet high, simple. hirsute: leaves ovate to lan- ceolate, acute or acuminate. 3-nerved, base cordate-clasping; heads few or solitary; involucre hemispheric. the bracts lanceolate, acuminate. villous- canescent, slightly spreading. Dry soil; August-September; reported as be- longing to the flora of lowa. H. divaricatus L. Stem 2-7 feet high, glabrous or pubescent above; leaves usually opposite, divaricate, lanceolate or ovate-lanceolate, dentate or denticulate, rough above, pubescent beneath, apex long-acuminate, base truncate; scales narrowly lanceolate, hirsute or ciliate. Dry woodlands; July-September; specimens from Winneshiek, Fayette, and Henry counties have been referred to this species. t ¢ Leaves petioled, % Stems glabrous or prleberulent, H. decapetalus L. Stem 1-5 feet high, slender, glabrous or nearly so, branches puberulent; leaves thin, ovate or ovate-lanceolate. sharply serrate, roughish above. more or Jess finely pubescent beneath, the lower opposite, slender petioled, the upper alternate, apex acuminate. base rounded or trun- vate and decurrent on the petiole; scales linear-laneeolate. acuminate. hir- sute, ciliate, ‘spreading. Moist woods; August-Septeinber: specimens frum Floyd county have been referred to this species. H. tracheliifelius Mill. Stem 3-6 feet high, roughish-pubescent; leaves ovate-lanceolate, short-petioled, 3-nerved at the base, green on both sides, rougher above, lower sharply serrate: scales linear-lanceolate. long-acumi- nate, pubernlent, ciliate, larger than the disk, sometimes foliaceous. Bor- ders; August-September; infrequent. Specimens from Fayette, Johnson, and Story counties are referred to this species. H. strumosus lL. Rowtstoeck long and slender; stem 3-6 feet high, usually branching, sinvoth, often. glaucous, the branches rarely hispidulous: leaves ovate-lanceolate, ucuminate, abruptly contracted or tapering into mar- vined pevioles, entire or slightly serrate, rough above, whiter and often THistLE FAMILY. 80 downy beneath; scales ovate-lanceolate. acuminate or attenuate’ and spread- ing, ciliate. Borders ot woods, August-September; common. ‘The variety MOLiis ‘I. & G., has the leaves finely and densely canescent beneath and is reported frum Story county. ( A. strimcsus maerophylius (Willd. ) Britton’). 4 @ Stem scabrous, hispid, ov hirsute. H. hirsutus Raf. Stem 2-4 feet high. hirsute; leaves petiolate, ovate-lan- ceolate, base abrupt or roundish, acuminate. more or less serrate, very rough above, usually rough-hairy beneath; scales ovate-lanceolate, ucuminate. Often confounded with the preceding; Aagust-September; frequent: Fay- ette, Van Buren, Decatur, and Ringgold counties; reported from Scott, Musca- tine, Johnson, and Lee counties. ; 4 : H. laetiflorus Pers. Stem 3-5 feet high. branched above, scabrous or his- pid; leaves oval or oblong-lanceolate, acuminate both + ways. inore or less serrate, rough on both sides, thinner than: the preceding: heads rather short- peduncled: scales in ? or 3 rows, ovate to oblong-lancevlate, acuminate; rays numerous; disk yellow. Prairies; August-September; infrequent! Jackson. Muscatine, and Johnson counties: reported from Fayette and Story counties. H. tuberosus L. Jerusalem Artichoke. Stem branching above, 4-3 feet high, pubescent or hirsute: leaves of the branches aod upper part of the stem usually alternate. all oblong-lanceolate to ovate or subeordate, petioled, acu- minate. coarsely serrate, scabrous above, shortly pubescent or cinereous beneath; scales lanceolate. attenuate. The variety sUBCANISCENS Gray, has the leaves densely white canescent beneath and is fuund in Fayette county. Moist soil: September; frequent: Winneshiek, Floyd. Henry, Van Buren. Decatur. Dickinson. and Winnebago counties: reported from Fayette, Musca- tine. Lee. Story. Boone. Emmet.-Cass, Woodbury. and Sioux counties. ACTINOMERIS Nutt. Perennials. with simple or somewhat branched steins. and feather-veined decurrent leaves. Heads corymbed, many-tlowered: rays yellow, few, neutral. Involucral scales foliaceous, nearly equal. Ne- ceptacle chaffy. : A. squarrosg Nutt. Stem 4-9 feet high. winged; leaves alternate or lower opposite, oblong or ovate-lanceolate. tapering both ways. more or less serrate: rays 2-8. irregular: achenes flat, obovate. broadly winged; pappus of 2-3 awns. Low woods: July-Augest: frequent in the eastern, southeastern. southern. and central counties: Johnson, Appanoose, Decatur, Ringgold, Page, and Fremont counties; reported from Neott. Muscatibe, Des Moines, Jleury. and Story counties. (Verbesine alternifolia (L ) Britton). . COREOPSIS L. Tickserp. . neutral, or none. Pappus of 2-4 awns, roughened backward. Achenes slender. quadrangular or flattened parallel with the involucral scales. * Plants terrestrial; leaves lanceolate. serrate, undivided, rarcly lobed, + Rays present, conspicuous, sometimes wanting bev B. cernua. B. chrysanthemoides Mx. Stem 6-24 inches high. erect or reclining be- low; leaves sessile, lanceolate, tapering both ways, serrate, connate; heads erect; outer involucre shorter than the rays; achenes 2-f-awned. Wet places; August-September; common; Winneshiek. Johnson. Floyd, Story, Deeatur, Winnebago. and Emmet counties; reported from Fayette, Muscatine, and Woodbury counties. (B. levis (L,) B.S. P.) B. cernua L. Stem 6-24 inebes high, nearly smooth: leaves Janceolate. sessile, sometimes slightly connate, coarsely serrate; heads nodding, rays few or none; outer involueral seales longer than the head; achenes usually t-awned. Wet places; July-September; common: Johnson and Decatur counties, reported from Fayette, Scott, Ienry, Lee, Story, and Emmet coun- ties. ; , + + Rays rudimentary or none. B. comosa (Gray) Wiegand. Stem 1-3 feet high. straw-colored; leaves short-petioled, lanceolate, coarsely serrate, petioles broadly margined;, outer involucral scales foliaceous, 2-4 times as Jong as the heads; corollas pale greenish yellow, mostly 4-lobed; pappus awns usually 3. Mcist soil; August— September; probably common; Johnsva and Decatur counties; reported from Fayette county. (D. connate comosa Gray). Probably confused with the fol- lowing. ; B. connati Muhl. Swamp Beyyur-ticks. Stem 1-5 feet high, purple: leaves thicker and greener than in the preceding. lanceolate or oblong-lanceclate, coarsely serrate, petioled: involuecral bracts not foliaceous: disk-flowers orange; achenes with 2-1 awns. Wet places; August-September, common; reported from Fayette, Johnson, Heory, and Story counties. **® Plants terrestrial leaves mostly 1-3-parted or dissected, + Rays short, rudimentary, or nore. B. frondosa L. Beyyar-ticks Stem 2-4 feet high, branching. smooth or somewhat hairy; leaves petivled, 3-5-divided. divisions mostly stalked. lanceo- late, serrate; outer involucral seales slender, much longer than the head, ciliate below: rays wanting; achenes with 2 long retrosely barbed awns. Moist ground; July-September; common. + 4 Rays larye and couspicuons. B. aristosa (Mx.) Britton. Stem much branched, 1-3 feet high: leaves thin, pubescent beneath, slender petioled. the lower pinnately 5-7-divided. segments lanceolate, serrate, incised or pinnatifid, upper leaves less divided; invotucral bracts glabrous or ciliate, short, the outer linear or spatulate, usually ciliate, not exceeding the inner; awns slender. 2. barbed, as long as the achene. Low grounds; August-September; reported from Muscatine and Benton counties. (Coreopsis aristosa Mx.) B. involucrata (Nutt.) Britton. Stem 1-3 feet high. much branched, minutely pubescent; leaf-segments linear-lanceolate. long-acuminate, incised ov pionatifid; outer involucral bracts Hnear-lanceolate, hispid and ciliate, TristLE FAMILY. My much exceeding the inner ones: awns 2, short. Low grounds. waste places: July-September: coninon: Johnson and Decatur counties; cuntused with the preceding which it much resembles. (Coreapsix involucrata Nutt.) *** Plants cpuatiie, submersed leaves filiforuily disseetcd. B. beckii Torr. Water Merigold. Aquatic perennial This species was observed by Hitebeuvek in a pond in Stury county in 1883 but no specimens were collected. HELENIUM L. Erect branching perennials, with alternate decurrent leaves, and many-tlowered radiale heads. Rays several, cuneate, fertile, 3-5- eteft. Involucral scales linear. reflexed. Receptacle ‘convex to oblong, naked. Pappus of 5-9 one-nerved thin seales. Achenes turbinate. ribbed. H. autumnale LL. Sneezewecd. Stem 1-6 feet high, nearly smooth. nar- rowly wing-angled; leaves Javeeolate, sessile. tapering both ways, toothed; disk and ray-flowers yellow. Pastures, preferring alluvial soil: August— September; common; Winneshiek. Allamakeée, Clinton, Scott. Muscatine, Des Muines. Johnson, Floyd. Jefferson. Deeatur. Taylor. Fremont, [Lancock. Lyon, Dickinson. and Emmet counties: reported froin Fayette, Story. Harri- son, and Woodbury counties. : ‘ DYSODIA Cav. Annual or biennial herbs, dotted with large pellucid glands which exhale a strong odor, Leaves mostly opposite. Heads manv-flowered, usually radiate; rays pistillate. Involucral seales in one row. partially united. Pappus of one row of scales dissected into capillary rough bristles. Achenes 4-angled. eo : D. chrysanthemoides Lagasea. Fetid Marigold. Stem 6-15 inches high. much branched. smooth: leaves pinnately parted. the divisions narrow, bristle-toothed or out: heads with a few short rays. Roadsides and waste places. | August-Septela ber: common; an emigrant from the west. (D. prtpposa ( Vent.) A. S. Hiteheock ). ' ANTHEMS L. Ours annual branching herbs. with (-3-pinnately dissected leaves, and solitary terminal heads. Rays white, pistillate or in the first neutra?; disk-flowers yellow. Involucral seales siall. “searious, imbricated, shorter than the disk. Pappus wanting ora mete border. Achenes terete or ribbed. ; A. cotula L. Mayieed, Dog-fenncl. Stem about 1 foot high, nearly smooth. leaves finely 3-pinnately dissected; pappus none; an ill-seented herb. toad- sides and waste places: June-September; common. - : A. arvensis L. Corn Chamomile. Similar to ‘the preceding but not ill- svented; stem pubescent; pappus a mere border. This species has been found rarely in Story and Muscatine counties. : ACHILLEA L. Perennial herbs, with small corymbose heads. and alter- nute much divided leaves. Heads many-flowered; rays few. fertile. Involu- eral’seales with scarious margins, imbricated. Receptacle convex or flattish, chaffy. Pappus wanting.’ Achenes oblong. flattened. margined. A. millefolium L. Common. Varrcw or Millfoil. Stem simple. 1-2 feet high: leaves 2-pinnatel y-parted; divisions linear, crowded, 3-5-clefi: heads in a flat- topped corymb; rays 4-5. short, white or rose color. Prairies, fields, and open woods: June-September: common. CHRYSANTHEMUM L. Perennial herbs, with toothed. divided, or pin- natifid leaves, white fertile rays. and yellow disk flowers. Involucre broad, depressed, of many imbricated searious-margined scales. Receptacle fat or convex, naked. Pappus wanting. Achenes striate. similar. ye COMPOSITAE. C. feucanthemum L. Greve or White Daisy. Stem 1-2 feet high, simple or branched: lower leaves spatulate, petioled, more or Jess pinnatifid: upper Jeaves clasping. toothed: heads large, solitary, terminating stein or branches. Vields and waste places. June-September; infrequent but widely distributed: Allamakee. Dubuque, Johnson, Decatur. Page, and Cerro Gordo counties; reported from Muscatine. Story, Cass, Hardin, and Woodbury counties. TANACETUM L. Strong-scented perennials. with alternate 1-3-pinnately. disseeted leaves, and corymbose yellow discoid beads. Ipvoluere hemispher- ical: the scales small, imbricated. Receptaele convex. Pappusa mere bor- der. .\chenes angled or ribbed, terminated by a large flat disk. T. vulgare L. Common Tansy. Stem 2-3 feet high, smooth: leaflets oblony- lanceolate. cut-toothed or pinnatifid; pistillate flowers tubular, with an ob- Jique 3-toothed limb: pappus 5-lobed. Along roadsides and in waste places: August-September: infrequent: Winneshiek. Allamakee. Jobnson. and Deca- tur counties: reported from Fayette and Story counties. ARTEMISIA L. Wormwoop. Annual, biennial or perennial plants with en- tire, toothed. lobed, or dissected, alternate leaves. and nodding heads which are in panicled spikes or racemes. Involucre obovoid, imbrieated, of dry connivent scales. Receptacle small. naked. Ueads discoid: flowers few to many, tubular, perfect, or the marginal pistillate. Pappus wanting. Achenes obovoid. capped by a small disk. 4 * Maryinal flowers pistillate and fertile; dish-flowers perfeet but sterile. + Leaves pinnately dissected into narrowly linear lobes. A. caudata Ms. Biennial, stem smooth, simple, 2-4 feet high; leaves 1-3- pinnately divided: the segments alternate, filiform: heads small, globose. numerous, in an elongated panicle. Sandy soil, August-September; common locally; Winneshiek, Muscatine, Jounson. Cerro Gordo, Emmet. Winnebago, and Lyon counties; reported from Fayette and Story counties. A. canadensis Mx. Biennial or perennial: stem 1-2 feet high, pubesceut or glabrous; leaves usually pubescent, the lower petioled and 2-pinnately di- vided into linear Jobes, upper leaves sessile, Jess divided: heads short-pedun- cled, in narrow panicles, usually numerous. Dry soil: July-August; reported from Woodbury and Lyon counties. t+ 4 Leaves linear, cutive or the lower 3-cleft. A. dracunculoides Pursh. Perennial, stem 3-4 feet high, glabrous or can- escent when young, branched; leaves linear, entire or lower trifid: heads small, numerous, in an open panicle. Dry or prairie soil: August-September: frequent; Winneshiek, Floyd, Muscatine, Haneock, Dickiusop. and Wuimet counties; reported from Fayette and Story counties. ** Flowers all fertile, the inarginal pistitlute; receptacle hairy. A. frigida Willd. Whole plant silky-caneseent; stems in tufts, 6-15 inches high; leaves pinnatifid; segments linear, 3-5-cleft; heads small, globose, droup- ing, racemose. Sterile soil; July-September; infrequent: Lyon eounty. A. absinthium L. Conunon Wormwood. shrubby. finely caneseent: stems 1~4 feet high, much branched; leaves 1-3-pinnately divided into obovate or oblong obtuse lobes, lower long petioled, the upper short-petioled or sessile, the uppermost linear and entire; heads many. drooping, yellow. short-pedun- cled, racemose-paniculate. Reported from Fayette county as a rare escape. #*% Blowers all fertile, the marginal pistillate; receptacle smovih. + Leaves dissected. A. annual. Stem 1-4 feet high, branching, glabrous, leaves :-3-pin- TuistiE FAaMiny. a) nately dissected. segments oblung, deeply pinuatifid: heads sual, in an imple panicle. Waste places: July-September: infreguent. Muscatine and Deea- tur counties: reported from Keokuk county. A. biennis Willd. Biennial: stem simple. ereet. 1-4 feet high. glabrous: {eaves 1-2-pinnately divided: divisions linear. entire or cut-toothed: heads globular, erect, in axillary spikes, collectively in a narrow leafy panicle. Dry soil: August-September: frequent. t+ Leaves serrate ov entire, not purnatifid. A. serrata Nutt. Perennial: stem leafy. 5-9 feet high: leaves lanceolate. serrate, whitened beneath. the upper often linear: heads small, few-flowered, greenish Kottom lands: August-September: frequent: Winneshiek, Jack- son, and Museatine counties: reported from Fayette. Poweshiek. Stury. Win- nebhago, and Cerro Gordo counties. A. gnaphalodes Nutt. Perennial: whole plant woolly-canescent: stem 2 4 feet high, branched: leaves lanceolate. entire. or lower toothed or pinnati- fid, upper surface usually glabrate and green: heads ovoid, stuall, mostly ses- sile. in narrow leafy panicles. Sandy and prairie soil: \ugast-September: common. (dd. Ladoviciane gneplelodes TL & G.) SENECIOL. Herbs. with alternate leaves. solitary or corymbed heads. and mostly yellow fluwers. Heads many-flowered: flowers all tubular and perfect or else the marginal ligulate and pistillate. lovolucre of many equal erect-convivent seales, sometimes with a few bractlets at the base. Recepta- cle tlat. naked. Pappus of copious soft capillary bristles. S. aureus L. Golden Rayivort. Perennial: stems 1-3 feet high, glabrous or woolly when young: radical leaves thin, simple. orbicular or ovate-oval. crenate-toothed. jony-petivied: lower lyrate-pinpatifid: upper sessile or partly clasping, dentate or incised: rays 8-12. Wet woods and meadows: May- June; Johuson county: reported from Winoneshiek. Favette. Story. aud Wood- bury counties: perhaps frequent and widely distributed but has been confused with the following. S. balsamitae Muhl. Stems a few inches to two feet high: radical leaves oblong or spatulate. varying to lanceolate. tapering to the petiole. serrate. upper lanceolate. laciniate or pinnatitid. petioled or the uppermost sessile. Dry and rocky woods: May-July: common: Winbeshiek, Allamakee. Scott. Johnson, Decatur. and Shelby counties; reported from Fayette. Museitine. aud Story counties. (NS. aureus balsamitae T. & G.) S. obovatus Muhl. Stems 1-2-feet high: leaves rather thick, the basal obovate or sputulate, very obtuse, base cuneate. crevate-dentate. with short margined petioles. or the earliest nearly sessile in rosulate tufts: stem leaves few, sessile. oblong to spatulate. incised or pinnatifid. Wet prairies: April- June: reported from Story county. (S. aerets obovetus TO. & G) S. lugens Richards. Stem stout. 1-3 feet high: basal and Jower leaves ob- long. oval or slightly spatulate. denticulate. narrowed into margined petioles. upper leaves sessile, bract-like: heads cor ymbose; involucral seales lanceolate or oblong-lanceolate, with black tips. Moist soil: June-August: reported from Plymcuth county. S. palustris Hook. Annual or biennial: stem 6-20 inches high. woolly or villous. sometimes at length glabrate; leaves oblong-lanceolate. toothed or luciniate, the upper cordate-clasping; involucre naked below: rays mauy. pale yellow: pappus copious. Wet ground: June-August: Emmet, und Cerro Gordo counties; reported from Dickinson county. U4 COMPOSITAE. CACALIA L. Inpian PLanrain. Tall perennials. with alternate Ieaves, and disevid heads in flat corymbs. Flowers white or whitish, all tubular: corolla deeply 5-eleft. Involuere cylindric, the scales in ove row, erect-con- nivent, with a few bractlets at the base. Pappus of capillary bristles. * Heads 25-30-flowcered. C. suaveolens L. Stem 3-4 feet high. glabrous, grooved; leaves triangu- Jar-lanceolate, halberd-shaped., acuminate, serrate, wing-petioled. Woods; August-September; infrequent: Winneshiek, Fayette. and Muscatine coun- ties: reported from Jackson and Seott counties. (Synosmut suaveolens (Ls.) Raf.) = * Aeuds 5-flowercd; inavolucre of 5 scates. C. reniformis Muhl. Stem 3-8 feet high, angulate, grooved, not glaucous. leaves green on both sides, lower Jeaves reniform, very large, long-petioled, upper dilated fan-sbaped, toothed and angled, palimately-veined. petioled. Rich woods; July-September; infrequent: Winneshiek, Allamakee. and Joha- son counties; reported from Fayette county. (Mesudenia reniformis (Mubl.) Raf.) C. atriplicifolia L. Stem terete: 3-5 feet high, grooved. glaucous; leaves palmately- -veined and angulately-lobed, “toothed, petioled, glaucous beneath: lower triangular-renifor m, or slightly cordate. Rich woods; August-Septem- ber; frequent; Muscatine. Johnson, Jefferson. Henry, Des Moines, Van Buren, Decatur, Union, Ringgold, Taylor. Page. Fremont, and Pottawat- tamie counties. (Vesadenia atriplicifolii (L ) Raf.) C. tuberosa Nutt. Sena grooved and angled, 2-6 feet high, glabrous. from a tuberous root; ledves oval or oblong, entire or obscurely toothe |. thick, 5-7-perved; lower with long petioles, upper with, short margined petioles. Wet prairies; June-August; frequent: Allamakee. Fayette, John- son, Appanoose, Decatur, Union, Adams, Ringgold. Taylor. Page. Pottawat- tamie, ‘Shelby, Hancock, Winnebago, Cerro Gordo. Dallas. Webster. and Emmet counties: reported from Scott. Story, and Woodbury counties. (Mesadenia tuberosa (Nutt.) Britton), ERECHTITES Raf. lil-scented annuals, with alternate simple leaves, and paniculate-corymbed discoid heads of whitish flowers. Involucre cylindrical, of linear scales, in one row, with a few bractlets at the base. Flowers many. all tubular, fertile. the marginal pistillate. the central perfect. Pappus of soft capillary bristles. ; E. hieracifolia (L.) Raf. Firewced. Stem grooved. 1-4 fect high, often hairy; leaves lanceolate, or oblong, acute. unequally toothed, sessile, upper auricled. Rich open woods and burnt clearings; August-September; fre- quent; Winneshiek, Johnson. Des Moines, Lee. Decatur, and Calhoun coun- ties; reported from Fayette and Story counties. _ ARCTIUM L. Biennial herbs. with large petioled alternate leaves, and solitary or clustered discoid heads. Tnvolucral scales numerous, imbriecated, long-attenuated, with hooked tips. Heads globose. many-flowered: flowers tubular. perfect. Receptacle bristly. Pappus of numerous. rough, deciduous bristles. Achenes oblong, flattened, A. minus Schk. Burdock. Stem 2-4 feet high. branehed: leaves thin, broadly ovate, with a cordate base, or laneeolate with a cuneate base. smooth above, pale-canescent beneath; heads many. racemose on the branches, short- peduneled or sessile, about half an inch broad, inuer bracts not exceeding the RAGWEED FAMILY. 95 pink-purple flowers. Waste and cultivated grounds; July-October; common, (A. leppa minus Gray.) CNICUS L. Tinstie. Biennials or perennials, with sessile alternate leaves, and large terminal discoid heads. Involuere ovoid or spherical; scales numerous, spinous-tipped or unarmed, in many rows. Receptacle bristly. Flowers tubular, perfect. Style nearly undivided. Pappus eopious, plumose. deciduous. Achenes oblone. compressed, smooth. C. Ionceolatus (L.) Willd. Common LT. Stem 2-4 feet high; leaves sessile, decurrent, pinnatifid, hairy above, white-woolly beneath, lobes spiny; heads purple; scales lanceolate, spreading, all tipped with aspine. Waysides and pastures; July-September; common. (Cerdius lanceutatus 1.) C. altissimus (.) Willd. Stem 2-6 feet high; leaves oblong-ovate to lanee-oblong, rough-hairy above, white-woolly beneath, undivided to sinuate- toothed, or sinuate-pinnatifid, lobes or teeth prickly; heads Jarge; scales lance-ovuate, outer with spreading spines; flowers purple. Fields and open woods; August-September, common. (Curditus altissimus L.) The variety FILIPENDULUS Gray. is 2-3 feet high; roots tuberous; leaves deeply pinnatifid. Emmet county: reported from Cass, Cherokee. Palo Alto, and Woodbury counties. C. discolor Muhl. Similar to the preceding; leaves deeply pinnatifid into lanceolate or linear segments. [ields and borders; July-September; reported common. Variable and seems to pass inty the preceding. (Cricus altissimus discolor Gray. . Cardias discolor (Muhl.) Nutt.) : C. hillii Canby. Stem 1-2 feet high, simple or branched; root fusi- form; leaves green on both sides, lobed or pinnatifid; lobes rounded, dentate or prickly; lower leaves spatulate-oblong, narrowed to the base or petioled, the upper oblong, sessile and clasping; outer involucral scales ovate lanceo- late, tipped With short bristles, very glutinous on the back. inner narrowly laneeolate and acuminate. Fields: June-July; reported from Story county (Cardiaus hiltii (Canby) Porter). C. arvensis (L.) Hoffm. Canada T. Perennial, stem 1-2 feet high, with a long running root; leaves oblong-lanceolate, sinuate-pinnatifid, prickly. smooth or slightly woolly beneath; heads small; flowers rose-purple, imper- feetly dicecious; scales triangular, appressed, minute, prickly-pointed. Culti- vated fields; infrequent but widely distributed. (Cardius arvensis (LL.) Robs.) AMEROSIACEAE Reichenb. RaAGweep FAMILY. Annual or perennial herbs, mostly of a weedy aspect, with alternate leaves, or the lower opposite, and small heads of greenish or whitish. moncecious or dicecious flowers. Staminate and pistillate flowers in the same head, or in separate heads. Involucral scales few, separate or united. Receptacle chatfv, Corolla wanting in the pistillate flowers or reduced to a short tube or ring; the culyx-limb none or a mere border; style 2-cleft. Corolla in the staminate flowers tubular, funnelform, or obconic, 4-5-lobed; stamens usually 5, separate, or the authers merely connivent. A family often included in the ComposiTa£ from which it is mostly distinguished by the anthers not being truly syngenesious. * Staminate and pistillate flowers in the same heads. Iva. Represented in our flora by a single species. 96 AMBROSIACEAE. #8 Staminale and pistillaie flowers in separate heads. AMBROSIA. Involucral bracts of the staminate heads united. XANTHiuM, Involucral bracts of the staminate heads separate: involucre of the pistil- late heads closed, covered with hooked spines. IVA L. Herbaceous or shrubby plants. with opposite or alternate leaves, and small nodding greenish-white discoid heads. Fertile and sterile flowers in the same head. Anthers nearly separate. Involueral scales 3-9, roundish. - Receptacle small, chaffy. Achenes obovoid or lenticular, I. xanthiifolia (fresen.) Nutt. Warsh Elder. Annual, stem tall, 4-10 feet high. roughish; leaves mostly opposite, petioled, ovate. rhombic or cordate, lobed, doubly serrate or cut toothed, hoary: flowers in terminal or axillary panicled spikes, bracts not obvious; fertile fowers 5, corolla rudimentary or wanting. Waste grounds; July-September; common in the western coun- ties but less frequent eastward. : AMBROSIA L. Annual or perennial herbs, with opposite or alternate lobed or pinnatifid leaves, and inconspicuous greenish flowers in spike-like racemes. Flowers moncecions; fertile 1-3 together, axillary. near the base of the raceme; the involuere oblong, closed, achene-like; pistil with 2 long protruding styles; stamens wanting; sterile flowers many, surrounded by a depressed hemispherical cup formed from united scales. Anthers nearly separate. Pappus wanting. Achenes ovoid. * Leaves opposite, paliniately lobed or wudivided; receptacle naked. A. trifida L. Greet Rugweed. Annual; stem stout, 3-12 feet high, hairy: leaves opposite, petioled, hairy, deeply 3-lobed, lobes oval. long-pointed, ser- rate; petioles margined; fruit obovate, 5-6 ribbed: and tubercled. Low grounds; July-September; common. The variety INTEGRIFoLIA TT. & G., a smaller forin, has ovate undivided leaves. This and intermediate forms are frequently found. ** Leaves opposite and alternate, pimnatifid; receptacle chatfy. A. artemisiaefolia L. Annual; stem 1-3 feet high, much branched, hairy or rough-pubescent; leaves opposite or alternate, thin, mostly twice pinnati- fid, smooth above. hoary beneath; petioles ciliate; fruit obovoid or globular, armed with 6 short teeth. Fields and waste places: July-September; eom- mon. A. psilostachya DC. Perennial with slender running rootstocks; stem 2— 4 feet high, paniculately branched, whitish. hispid; leaves thickish, mostly once pinnatifid; fruit obovoid, tubercles small or wanting. “Low grounds and prairies; July-September; common in western Iowa, rare eastward; Fre- mont, Decatur, and Johnson counties; reported from Pottawattamie, Story. Fayette. and Muscatine counties. : XANTHIUM L. Annuals, with alternate petioled leaves. and moncecious heads of flowers. Fertile flowers in clusters, the sterile in racemes above. Sterile involucres of separate scales: receptacle cylindric. Fertile involuere closed, coriaceous, covered with hooked prickles, 2-celled, 2-flowered. Pappus wanting. Corolla thread-form, Anthers nearly separate. Achenes oblong, flat. X. canadense Mill. Cocklebur. Stem 1-3 feet high, frequently brown spotted; leaves ovate or cordate, 3-nerved, dentate. frequently lobed: fruit about l-inch long, densely prickly, beaks hooked or ineurved. [Fields and waste places: July-September; common. Cutcory FAMILY. 97 CICHORIACEAE PReicheub. Cutcory FAMIby. Herbs. usually with acrid or milky juice, alternate or basal leaves, and vellow, blue, purple, or white flowers in involucrate heads. Involucral bracts in one to several series. Receptacle tlat or flattish, scaly. smooth, pitted, ov alveolate. Flowers all alike, perfect. Calyx-tube udnate to the ovary, the limb of seales; or bristles. or both. or wanting. Corolla gamopetalous, ligulate; tube short or lone; ray usually 5-toothed. Anthers 5, united into a tube around the style, sagittate or auricled be- low, usually appendaged above. Ovary 1-celled. 1-ovuled; stvle 2-cleft or 2-lobed. Fruit an achene. This family is usually regarded as a tribe of the ComposiTAE and is also known as LIGULIFLORAE. * Pappus of seales, or of scales aad bristles, CLiCHORIUM. Flowers blue or white; involucre double: pappus of blunt scales. Iwkicta. Flowers yellow; involucre simple: pappus of chatf and bristles. ** Pappus pliumosc. TRAGOPOGON, Stems leafy: involucre simple; plume branches of the pappus interwebbed. * 8% Pappus of capillary bristles, not plumose, t+ vlchenes eylindric, prismatic or tercte. mot fluttenwed, } Achenes not beaked. HteERAcICM. Flowers yellow or orange, sometimes red; pappus tawny PRENANTHES. Flowers white. cream-color, or purple: pappus tawny or brown; heads often nodding. LYGODESMIA. Flowers pink or purple: pappus white: heads erect. tt Aehenes betked, sometimes bechkless th Troximon,; flowers yellow; heads solitary, SCAPOSE, TROXIMON. Involucre loosely imbricated; achenes 10- ribbed. TARAXACUM. Involucre calyculate: achenes t-A-ribbed. + , clehenes flat or flattish; pappus white; involucre imbricated; heads panicted. Lactuca. Flowers yellow or purplish; achenes more or less beaked. SoncHus. Flowers yeHow: achenes flattish, beakless, CICHORIUM L. Erect branching perennials. with deep roots. alter- nate and basal leaves. and axillary and terminal heads. Involucre of two series of herbaceous bracts. Receptacle flat, naked, or slightly fimbrillate. Achenes j-angled or 5-ribbed. truncate, beakless. Pappus of 2 or 3 short blunt scales, forming a crown. C. intybus L. Chicory. Basal leaves on the ground, runcinate-pinnatifid, narrowed to long petioles; stem-leaves lanceolate or oblong, lobed or entire. base clasping and auricled; heads many. Waste places; July-September: re- ported trom Fayette, Scott, Polk, Adams, and Woodbury counties. KRIGIA Schreber. Ours a perennial, with scape-like stems, mostly radical leaves, and several to many-flowered heads. Involucral scales in 2 rows. Pappus double, the outer of scales, the inner capillary. K. amplexicaulis Nutt. Roots fibrous; stem 1-2 feet high, smooth, with J—. leaves; cauline leaves sessile, clasping, oblong or oval, mostly entire; radical on winged-petioles, entire or toothed, rarely pinnatifid; heads 2-5, peduneled. Rich woods; May-June; frequent; Wioneshiek, Allamakee, . Jones, Muscatine, Johnson, Henry, Lee, and Cerro Gordo counties, reported from Fayette and Scott counties. (Trayopogon virginicum L. Adopogon virgini- cum (G.) Kuntze). K. dandelion (L.) Nutt. Seape slender, leafless, with a solitary head. 6- 18 inches high; roots tuberiferous; leaves basal, tufted. linear-lanceolate to 98 CICHORIACEAE. spatulate, entire, denticulate, sinuate, or pinnatifid. Moist soil; April-June: reported from Woodbury counbLy. TRAGOPOGON L. Glabrous biennials, rarely perennials, from long descending roots, with elongated-linear, nervose, entire. clasping leaves, and solitary heads of yellow or purple flowers. Involucre simple. of several Jarge equal lanceolate acuminate seales. Pappus plumose.