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Les cartes, planches, tableaux, etc., peuvent dtre film6s A des taux de reduction diff^rents. Lorsque Ie document est trop grand pour dtre reproduit en un soul clich6, il est film6 A partir de I'angle supArieur gauche, de gauche A droite, et de haut en bas, en prer" all. CWerilyitisweUknoUZ 'i^hUJltiS^^^^^^ "^"*^'* ''' of the salte of sUver, hydn)eyanic acid &c »nT^ «f^«'o™P08ing several leav.ofpIan^carbo;ie'ac7a^ammo;i^So Z^^T "?.*'.^ ^^- forms of physical force, but to no purpose aa thl eZ Jf -n T"' «' "*^^ iUustrate my meaning and I am IZ^ examples mil be sufficient to phy^ foL. ^' "°* °°^ '''''^'^ ''*•> '^' correlation of the And first of plants. water, ammonia, carbonic acid &c and 7.^ • 'ts e„ret.ons or decay, such as ar ' 'rom them it reform 7eoZi^oTZ 1^ 1 ^*^'' ^^' '^^' ^" -^. they were derived. Tl^t ^S rw:Ser^^^^^ '"°"^- eff^ted by L veget^bk li^rfo^Ts ^^ ^ ^'^^if *^« ^''-^^ whence it is derived : at pre^nt weTalf ^!^ '^ . ^^ "^^ ^^ ^" «'^^' ««« namely, germination whirh^SrjenLrl''^ ''.t '*^ ''^ ^**««°' growth of the plant, 'and closelj ^t rnrm^irl^ '''^'^" ''""" *^« ^'■ hea?rm;i: :^r3ol"fhe"\^^^^^^ '^^''' ^ ^''^ -«"- of imposition; some^^^e ISrlrric^rralt" tT ^"*" '^■ which the starch is converted first into dextrine aud*^ T ^^ '"''"'' "^ solved. A part however does not «flnl, t ?' '''° '"^^ '"8"' *°<* ^'8- passes downLdslTreTa; ItL J ^"*/^'"°g oxygen f„>m the air stable conditionTt can atS^rndrr. ''^''"*'°" ^^ '^' '"^^^^ -<» '"ost meantime in the pla^^ onhe ^ we Tavl" "*"' ''?'""'°"'° ^''-^ '^ '^^ ^^=;ruit^3^^ -..somrs^zf^^rv::"^^^^ cont;i^Tni"^jrt:pr Ti ''' '""^^''^^ '^^ ^'- *^« -*»«' let us weigh the pl^t tJi^ forlT "^'^ """^""^ '''*'"°- ^« -^««*««<1 this shall find-putli!! 1 o^/l 1' '"*^ '"PP'^'^S we had weighed the seed we the young pCShatso-^^J^ '"Z"'"*"" that may have been absorbed by way 8 Structural An^*- "^ '.^^■: -^fi.-^ .: 11 from complex bodies such as malamide, (C» H • • i\» O') is of little consequence in the consideration of the question I am now concerned with. However the process be looked upon we have here again the deconirosiUon of one body, and the formation of another, but in an inverse order, as it were to that observed in the former case ; for whereas in that the body which was formed had more stability than that which was decomposed, and there was in conse quence a surplus of force; in this case the body formed is in a much lower state of combination than that decomposed, therefore (Law IV) force has been required, and this force we have seen is supplied in the form of light. There is another question, in connection with this supply of force to plants of great interest and importance, but which I think may be best consider^ when I come to speak of the influence of light and heat on growth and develop ment, where they do not seem to act so directly through the chemical force If we pass now to the consideration of animal life, wo shall iind it to bo under the influence of the same laws as those which govern the life of vegetable^ but to be m one sense diametrically opposed to the latter; for firsUy, as tomattlsr it requires for its maintenance to be supplied with organic compounds, animals not possessing the power of appropriating to themselves mineral matter* and secondly, as to force; it is supplied with no dynamic agency JVom any external sourrce, at least not in the same direct manner that plants are. This diflference may be briefly expressed by saying that through plants the other physical forces are converted into chemical force; and in animals this chemical force is reconverted into the ordinaiy forms of physical force and so returned to its former state. In each case, by its passage through particular forms of structure, taking on, in some part of its course, the various forms of force caUed vital. And did this hold good throughout it would afford an ex- ceedingly beautiful and useful line of demarcation between the two kingdoms but unfortunately as with all other distinctions, the exceptions to it are numer- ous, though with them we need not now concern ourselves, as they have not the least effect upon tne theory for which I am contending. Now these two facts are easUy seen to be intimately connected with one another ; for as the food of animals is organic matter, that is, matter in a state of weak union, of highly complex chemical constUution, the elements of which readily, upon the least provocation enter into more inUmate combination at the same time (Law III) evolving force, so animals may be said to appropriate force along with the matter they eat. Further, plants receiving force from without, do not again while they continue to hve, or even perhaps till long after Uieir deatfi, give out that force but hoard It up; whereas animals taking in this force with their food, give it out in the performance of all tiieir functions, such as innervation, muscular action, seer*, tion, &c., and after tfie animal body has arrived at its ftdl siie, so that it requires no force or matter to be used in building it up which is not again given out in • I do not speak here of the water used by animals, because although they injeat and m a sense assimilate it, it eridenUy can take no active part in the economy, at do the organic articles in their food, and though it is extensively used, it is so as a purely pas- •ive agent. 12 its breaking down, it stands in the same relation to force that it does to matter not retaining any but giving out in some form or other aU that it veoeiveB. Bui It wUl of course be observed that this is not the case as long as any processes of growth or development remain to be accomplished. ^^ .,, ^'t^ T "°f '^' *''' ""'"'^ '''^"'^"^ '" *t« «^ ''"y as we have done the vegetable, and to do so let us begin with the egg The egg being subjected to a certain degree of heat, prebably required torive TrtS L^ :fT'^ ''^"'* '* " '"^"p"^' """^ °«' itself iTi^r It^tT 7 ' "^ '"*"" '"*° *''*' ^»* sortof decomposition as didT^ eed in the former ease, oxygen is absorbed, and carbonic acid and perhaps wa ter IS given out, through the minute pores of the sheU or sac as the Vase mayle ll^'l^ t^ r '"t"^-. ^ " consequence of the formation of these simpk gethr with free oxygen foree is evolved. (Laws I and III). At the same ime gradually, from a shapeless mass of albumen, a living being is built up. To effect this force is required ; on the other hand force is known to be given out Moreover we know that on the one hand it is gradually Uberated; and on the u! f^'ST/ * ~*«'f V"'^ ^«'«- Surely the conclusion foreed upon us ,8 that the force here evolved is applied when foree is known to be used- rather than say, here foree is annihilated; there ereated-.,r here foree becomes atent ; there it is reused from its dormant state. For is it not unphi'.osophical to suppose a cause which is more than adequate to produce an observed effect? shell with It, It will be found to contain considerably less matter than did the egg from which it preceeded j we have seen where that matter has gone And Jf the tissues of the young bird be analysed they will be found to contain at least some of them, molecules such as hemato-fibrine = C » « H^'N** S" O") more complex, less stable, and held together by a weaker affinity, than were those of the ongmal constituente of the egg, thus displaying (besiL the moiThological and hictological transformations that have tVen plStS. re fo,^ wa! deS ''''"" ^'''" '""-^ ""' '^"^ ^'^ -- -J^-- ^^ In another place I stated that my idea of the agency of cells, as such, is that they represent the form of matter threugh which the physicalf rees pas Sthd conversion into the vital. Now the form of vital foree mnifested by^lls vS with their structure; and for any one kind of action we must have an apprenri ate form of cell. But contained in aU seeds and eggs capable of life w!Zve ^Z'l^l^f;.'^''' T'^f'^^-^^^^y^^^ chemical ZZ^Z ;^n,r- n ?• f "P *^' P^""^* " '"'^"'' "'^ ^^'^- i^ "^b^nt makes the material link incomplete ; and the chemical foree instead of passing to the vital, IS given out in other forms, chiefly as heat. But whether thTegg or seed llT ?w r ""*' ''^ '° '"^ deoompoBition it must result, fremihe C l^d down that the same amount offeree will be given out by the time that its elmentfl, m either case, have reached the same chemical level Now after the young bird or mammal, as the case may be, has used the food la^^ up for, or supplied to it by the female parent and has t(; shift for it^ h 13 receives no force from vithout, as does tLe plant in a similar situation, and here, and not till we arrive at this point does the difference above mentioned begin, but its food consisting of such matters as being in a state of loose com- bination readily admit of the evolution of power by entering into greater inti- macy of union. (Law III). At the same time, as if this were not sufficient, a free element of pre-eminently strong affinities is taken in by the animal to com- bine with them, as they run down to form the simple compounds. But here it may be said that the young animal has not only to furnish energy for the performance of its organic and animal functions — respiration, circulation, locomotion, innervation, &c., — but also it has to grow ; increase in size. Its tissues continually breaking down in the performance of their respective func- tions, it has to restore them continually, and not only this, has to add steadily to their bulk. Whence comes the supply offeree that shall be adequate to these wants ? And here we see the purpose of the vegetable kingdom in its relation to the animal, perhaps more beautifully displayed than from any other point of view. For the vegetable having received force from without, holds it in trust, as it were, for the animal kingdom ; and in the use of vegetable food it is known that the animal does not raise the compounds contained therein to any higher state of combination than that in which it receives them — except in the case of hemato-fibrinc, and perhaps a few other instances. To build up its nitrogenijsed tissues it receives albumen, fibrine, and caseine; for its adipose tissue it takes in fat, though it can also form this last from staioh or sugar ; its gelatinous and cartilaginous tissues are supplied from the albuminous bodies by a process of diminishing complexity, and so on. But this is not of much importance, after all, for the result would be the same, namely, that on the one hand animals receive compounds of great com- plexity and of loose chemical construction j and on the other hand their excreta consist of chemical forms of great comparative, and some of great actual simpli- city, between the elements of which there exists great strength of affinity, there- by furnishing the conditions (Law III.) for the evolution of vast quantities ot force, which being directed into the proper channels by means of the various forms of cells, through which it acts, performs all the functions of the body. But here, if this be granted, another difficulty will arise in most minds, which at first sight might seem almost insurmountable, fcr, it will be asked, is it possible that such an immense amount of power of various kinds, put forth by animals, can be derived from the decomposiiion of what seems to be the compa- ratively small quantities of food they digest? To illustrate this forcibly, I was once for five days and four nights exposed to a temperature of from zero or below that point to a few degrees above the freezing point : during this time I was sup- plied with no food, no artificial heat, and travelled every day on foot through deep snow from morning till night. Now, I ask, could the muscular force employed, the heat evolved, and the vis nervosa put forth (without speaking of other forms of force liberated in less amount), have been derived from the decomposition cf the tissues lost during that time ? I make no doubt that the reply must be in the affirmative. The answer to this difficulty resolves itself into two distinct parts. 1. The >» »"' : fii 14 consideration of the quantitative ivlaHnna ^r »i *> cconomiepowei.ofanin>asco„p^^^^^^^^^^^^^ «»d 2. The contrivance. ^ ^ *^ *^°'' °^ " '"""''i^c of merely human one hand, and the vaLstrm/of vl „ the 'Ji^ ' *'r'?'"' '<'"'' «" »^« resultant of any vital force wh"n i Jol V ,. ' ^"* '^ ^^ ''""'^ ?«* 'he this being bettfr unde/srod could morf. ,1'"'° """^ '^^ ^'^^ P'^^^ ^°«««. 1. we can do in the oJ:'i::ZZZS ."^^^ITV'^' '"'"""" -and I think it may safely be cranted fl,„f f ^^""ot'on we can measure; offby the body takes^hisf^rmrreJt!^^^^^^^^ 'f ''' '"'«^ S-" •nation to the result we are s ekin. F„ °? tl roT . \ ^ T""*^ 'P^''^^'- twecn motion and heat be recolleSlthat Ll 'f^'T'^'^ ^^^^ exists be- 772 pounds one foot is only el!l 1 1 . ! '"°^' ^°''° *'^P^^'<^ "^ '«'"«? wa Jthrough one dele Fal' aid T 1 '''^'"'^ ^^ ™^° °"^ P"""^ of of chemical action ifLessanfi^tl 1.7 ^'^ T'^'' ^'^* ^ ^"'^^ »'"°""* heat-how much'eat forSsttce a 1^7. " ^"^''"^''^ •^"■'""^^ o*' while, at tl. same tim^irgivroHaJ^^n^^^^^^^^ ? lamp wi„ yi,„, disproportion tltrmrdteSL'tr^^^^^^^ 2. That the animal body should beTsutrirr " ^'' F""" '' ''"^'"S- a^d Hand soitlilyt^wr;: S^X^t I^Tf ^ ^^"^ us. And it must be regarded as a result of f^! ■■ '''''^ '^""S proportion of the force given off tekes the fori f .""""^"S '^'' «« g^^at a is a much ehea^r ^o Jof fo^Lt l^t™ IT^^^^^^^ '^-^ Part IT. 15 cessity alive, in the ordinary acceptation of that term, yet it must be in a stat<> closely allied to vitality, as recently dead tissue, or as in the case of the seed vrhieh is said to be in a state of " dormant vitality." And here we have as it seems to me, an almost, if not quite, insuperable bar to the doctrine of equivo- cal generation ; which can never be received till some exception be pointed out to the law I have laid down. It may be considered as proved by the following facts that nerve force and elec- tricity are not identical. For (1) no electric current can be detected in a nerve along which nerve force is known to be passing. (2) By ligaturing a nerve its conducting power for electricity is not in the least impaired ; while for nerve force it h destroyed. And (3) if a piece of nerve be removed and the ends thus left be connected by means of a conductor, electricity will still pass along it, but nerve force will not.* But it must equally be allowed that some exceedingly clnso relationship does exist between them, for if a current of electricity be passed along a motor nerve, even for a part of its course, contraction of the muscles supplied by that nerve is the result. ^ If an afiFerent nerve be experimented upon the same way, whether a nerve of common sensation, the optic, auditory, gustatory, or olfactory, a prick- ing sensation is expeilenced, flashes of light seen, sounds heard, a peculiar taste, or a phosphoric odor perceived, in accordance with the function of the nerve operated upon.f Now if in the case of the magnetization of a bar of iron by the passage of a cur- rent of electricity round it, a conversion of the electric force into the magnetic be conceded, such connection can scarcely be denied in these cases in which a perfect parallelism to that of the magnetization seems to exist. But to follow out the analogy — for magnetism will under the proper conditions produce elec- tricity — the converse should hold, namely, that the nerve force will produce the electric ; and this is seen in the most remarkable manner in the case of the electric fishes of which the Torpedo, the Gymnotus clectriciis, and the SUui-us clec- tricus are best known. In them a special division of the nervous system is set apart for the production of nerve force, which by means of a particular form of apparatus — supplying the special material substratum required in this case — is converted into electricity. That in this case the nerve force is in fact converted into electricity, or bears some very analogous relation to it, is as well capable of proof as that in other cases it excites the contraction of muscles, for from the electric lobe proceeds a large nerve trunk, which when it reaches its destina- tion in the electric apparatus, divides into minute branches which ramify profusely in all directions. Now if this nerve be divided, the apparatus fails to evolve electricity ; if partly divided or injured, the discharge is weakened ; if the elec- tric lobe be removed, destroyed, partially taken away, or injured, similar results follow ; and if the lobe be irritated the discharge is increased. If now the nerve be divided, and the cut end belonging to the peripheral extremity be acted upon • Lectures on Physiology by Prof. Fraser. t For the facts in this paragraph as well as for several others of the same kind fur- ther on, I am indebted to Dr. Carpenter's article in the Phil. Tran. for 1830 on " The mutual relations of the Vital and Physical forces." f't > ^ ''f i 16 t^t::^^;rrr^Jt„r^^^^^^^ o. He., .e result wiU eUctri^ity from the apiaratus " *^' "^*'' «"'' "» ^volutioa of J^'it^tn/Lteio^rr^ -^ -sole nomena connected' Z^TJZ neToeT"'" "' '''^'''''^' "^ ^« P^O" -usclejn their relaUon to orither^rla; "kT"'"- "''" *"""'' «"^ ^ cription. It iB, however, probaWe tha. ^ " ^''° '" '^' ^^^'^ d««- relation that nerve force Cs to .."• 5 • '? ^'* ^^''''''' »^*'««° the produced in the other For^hL Tthf " ^'^.^ °""' ''"'^ *« «^« -«"»» certainly not converted direc7lyZ^ 1^ T! t^' """^^ *^^ "^'^^ ^^'^^ ^^ «hip to it, as we shall ^e further o;. T„ the nt ^T ^ ""^ •^^*^"* ''^^'- trie fish it is highly probable thLtL 1 . ^"'^ '" *^« ^"^ °^ »!>« elec- nerve force by VcCttnt^ alT^^^ the tion and even death of the animal after l.fl? ' 1"'* *^" '^^'''^^ ^^^u^" and the fact that no other orS Ifl^L" u' ^ "''"^i^' "'^'^ ^'' ««"« ti°»«.=*^ de^, to justify us in com^i^fTofh;: ^H^r' °"*' -"'^ -"> ^^ - there :nrn:Tou^t:tratr^:;^^^^^^^^^ converaion of heat into nerve-force ^y22L ?. ""'^ "°'^ ^'^""^ "!»» '^ the nerves which transmit suerSpttl'X'he ''"'^"' -tremities of n>ent of this, and which would annfyTn' II l" J '^"'"* ^°'' '^' ^^'^^'^h" of this form. Given a force appH K ho. ^r^^ T' "°"'' "^^^ ^""^^^'^-^ of a different chamcter passfn. alol thif n!^^ 'l\^''''> «» *^^ «"^-^««"t or (2) it must have been roused from a h!^ ^TT^ " "°"''-'- '- "««ted ; by eonverslo. of the forceTwctTas in '*"''' "' ^'^ ^* '^ *^« '^"^tan; «een reasons above for rejecting tte two I^ T -"""^^ ''' ^"^ ^^ ^ave therefore to accept the iL eXati- "' '* '"^ "'"'^"^ *« »« by a t^^ltlLTZlir ^^^^^^^ .^^.^ --^- ^^- i3 caused the optic nerve, flashes of light Ts^rtt! "^^^ ''^"" '^^"^' '»PP"^'» to appUed to the course of a motorTem it' .^o^ *'^' "'^^^ ^^"'^' ""^ «« °« i by it ; in all cases, causing a « of n T '"°'''° '" ^'^^ "^"^^'^ ^^^^^^ Which it is applied'as the LtTtcT^n Us Z^Z^ '"^ ^'"^" *'^ "^^ *^ »cal action, some of it may be dZllTf ^"^"^ ^^^ to rf«-ec« chem- I Carpenter, Phil. Tran. 1850. ' '^°- te-^.#^:*e»,.' 17 and the rapid cooling of bodies of ivhich tho nervous centres have been destroyed notwithstanding that respiration is artificially maintained, and tho circulation continues.*" Chemical re-agents applied to nerves in their course will produce all the effects which we have seen to follow the application of heat and electricity ;f while, as the converse of this, it is well known that nerve influence may change the chem- ical properties of the secretions in the most marked manner, and even probably produce chemical alteration in tho blood itself, or tho solid parts of the organ- ism ;| and that it excites chemical change in the muscles there can be no doubt. We have a striking instance of the conversion of light into nerve force in the phenomena of sight, the mode of which conversion being a matter of little im- portance in our present inquiry. Whether we ore inclined to accept Draper's|| very ingenious explanations of it or not, tho fact that such change does tako place can scarcely be denied ; tho argument for tho proof of this conversion would be similar to that used on a former occasion when speaking of heat. Conversely, although most cases of animal luminosity may be fairly referred to slow combustion, or phophoresence, in the part where such effect is manifested, yet this explanation does not seem adequate to account for all instances of this kind ; and it is believed that in some cases, more particularly in the cases of the marine Annelida, and some other of tho Articxilata, a conversion of nerve force into light takes place.§ The relation between nerve force and motion has been considered as being a more remarkable instance of conversion than any of those above specified,^ But this does not seem to me at all so clear as at first sight it might appear to be. For it is certain that in the relationship existing between motor nerves and their muscles, no conversion of nerve force into motion takes place, but, as wc shall hereafter see, a connection of an entirely different character obtains. How- ever this may be the converse of it holds good ; for motion in the form of me- chanical irritation applied to a nerve at ita periphery, or in its course, will be followed by a nervous current along that nerve and by excitation of its centre. That is to say, when applied to a nerve of common sensation it causes pain ; to the eyeball or optic nerve, flashes of light; to the auditory, sounds; by strikbg the tongue quickly and lightly with the tip of the finger, a distinct taste is de- veloped, sometimes saline and sometimes acid. t9° I have said that the motion produced by the contraction of a muscle cannot be r^arded as a continuation of the nerve force which called that muscle into action. It seems sufficient reason for this assertion that there is another, and distinct source known to which to refer for the proximate antecedent of the mo- • Carpenter, " Principles of comparative Physiology," p. 401 ; see also his " Human Physiology," pp. 417 et acq. t Carpenter, Phil. Tran. 1850. X Carpenter, " Human Physiology," pp. 740-746. II Draper's ''Human Physiology," pp. 392 et seq. § Carpenter, " Principles of Comparative Physiology," p. 447. Compare Todd and Bowman "Phys. Anatomy," pp. 224 et seq. ir Carpenter, Phil. Tran. 1860. V^ Baly's translation of Muller's Physiology, p. 1002. /i, xj Ik ..-AS 18 tion namely, the chemical change taking place in the muscle; the relation of which to the force put forth is bo well shown by the different amount of urea formed under the different circumstances of activity or rest of the muscle i= Still that there is an intimate relation between the current in the motor nerve and the muscular contraction is well known, and also that in a certain sense a quantitative relation exists between them, the degree of contraction in the muscle being entinjiy dependent, cwlcU paribus, Mpon the amount of stimulation or nerve force conveyed to it by the nerve supplying it. The relation then between nerve and muscular force, though intims >, is certainly not that of direct con- version, but seems to be extremely analogous to that which light bears to the force produced by the union of hydrogen and chlorine when their combination IS determined by the action of it« rays upon them ; for the amount of nerve iorce, as of light, supplied in any given time, other things being equal, detcr- »^TL^ amount of chemical change in the muscle or in the mixed gases and consequently the amount of force (Laws I. and III.) that will be put forth' or evolved, but in neither case docs the determining agent supply the force thus yielded. But the nerve force as well as the light, (according to the theory here advocated) must have a resultant when it ceases to exist as such and I would suggest the possibility of that resultant being the heat, or part of it, that IS always produced during normal muscular contraction. If we consider now that on the one hand all the physical forces are mutually convertible into each other, and on the other that nerve force is considered as the highest form of power put forth by organised beings, besides being (as must be granted) probably correlated to all the rest; it must be allowed that the fore- going facts go far towards establishing the relation contended for here, between the vital and physical forces, for if each one of the two groups of force have its own forms of force convertible into each other, then it only requires one connect- ing link to establish the unity of all the forms of both groups. But because we cannot point out any one link that would fulfil this condition so as not to «UIow of any cavil, if is necessary that we should have a larce number of in- stances of conversion, each of which should be as reliable as possible under the ra?p!lTwr/'r\^^"'"\P™^''^"'"*'^^^ ?«'"*'"§•" »»« direction, we may establish that which cannot bo shown to be absolutely true by any one direct ^ Instances (or at least seeming instances) of conversion coming under this di- vision of the subject might be multipUed, but it would be tedious to do so and would serve no purpose, for if those instances already adduced are not received ^^^l «f r«l''«on, any others would hardly be so; and if they axe so looked upon tliere is no need of adding to them, and it must be obvious to everv one that If this view be the true one, every vital manifestation must be an example ot correlation, since every vital force, in its origin as such, must proceed from a physical one. I shall, however, brieBy consider muscular action^ for the 1 po^ of seeing whether this theory is capable of throwing any light upon it. Aach form of cell, as we have seen above, has its own pr oper fom of « cell IogyJ^7S," """""" Pbyio'ogy-" pp. 444 et seq. and Carpenter's " Human Physio- ««^. jTlfSj^^l^^Wn 19 force," which it evolves under certain determinate conditions; thi.s force hai for it« anteoedcnt that set iVco by the oheniioul chun^cs going on in the cell itaclf, the convention being effected by the particular form of matter (the cell) through which the force passcH. Now an the cell is the form of matter (jntr r.r- lelknce) throua;h the agency of which tlie physical iorccs are changed into the vital, HI) each form of cell has its own form of vital force wliicli must result from the fact of ite having its origin as vltnl /onr in that cell. As the proximate origin of the force liberated by each cell i.s to be found in ohcmical change, ho tlic stimulus that calls that cell into action is something that will determine the taking pluce of that chemical change, and may be itself exceedingly amall in quantity compared with the force which at first sight it might seem to produce. The cells* of muscle iiro eliPinically composed of cxeceilingly complex bodies ; the affinity exercised between the elements of which seems to be very slight and their stability remarkably feeble ; from which it results that their elements may easily be made to change their chemical condition (Law II.); and also that when they full into low forms of combination in which the affinity exercised by them will be great, a large quantity of force will be set Irec (Law IIL) ; and in this fact we have the reason for the great complexity in chemical constitution which obtains in tissues through which much force has to be evolved, that is, whose functional activity is great. f This being the state of affairs what is next required is, first, another force which t^hall so act upon the complex bodies as to cause the chemical change, and second, a form of matter, in the passage through which this force set free shall assume the form required ; these two conditions we have fulfilled in nerve force on the one hand, and muscular tissue on the other. But any other force may take the place of the nervous as when an isolated muscle is called into action by heat, clee- ctrioity, mehanical irritation, etc. ; and again the cell may be in some way so al- tered that it shall lose its property of directing the force set free into the normal vital channel, and then we shall have another form of force evolved, which in ac- cordance with the rule, in cases of chemical combination, will be mostly or entirely heat. This aberration is seen during life in certain morbid states of the system as in pyrexia (?), and after death of course it always happens ; and if the cir- cumstances attending the death be such as to leave the elements in a more than usually unstable condition we shall have a rise in the temperature of the dead body, &» is oflen seen in cases of cholera, yellow fever, etc.J In health, because it would seem some little chemical change, probably connec- ted with nutrition, must always be going on, when the muscle is at rest and there- fore not liberating a ly of its proper force, electricity in small quantities, (for the * I follow Sharpie} and Carpenter (Prof. Fraser's lectures on Physiology,) in consi- <1ering muscles as ultimately composed of cells : in reality it does not appear to me to be of any consequence for our present purpose whether we call them such or fibrillar, t For it would seem to be a law, to which there are certainly seeming exceptions from the operation of other laws interfering with it, that the more complex a body is, the weaker is the chemical affinity exercised by its elements. t Carpenter's " Human Physiology" p. 410. #t«^ ^B^sEfeLii^'j '^'^^l HEl^'I 20 oSltl^ '"'''; '"^ ''''""' ^"* ■" «^" "" »'>° »"'^'" '« ^""^'l upon to give out ite proper form of oneiKy, Us evolution oui.- os * ^ *"^' mi of Ilfr ""T ^"""'^''"' '° "•"^^ ^ "»*'*' ^^«'P^'» '^'^ '•«*<" 'h*t irritable IfT.h r T '" ' '"^^ '*"^'" °«"^*'°" '»>»" ol-tainH in other pe p ) h„v more ot this free eleotrioity than others.f ^ ^ '' ^ Part III. And first of heat. within thorn h.? V '^""y ^''"" *'^*^ ^'^""•cal clianges jroing on by its pl^etou^^^^^ P"t°' *\^^'' ''""'^ ' ^«"» '^ *^« f-- -hic'>, form in cr^t lart tt • 5 '"^ ^'"^ "^''"^^"^ *« ^'^' f°'««' has to per- orm in grea part the assunilative and nutritive functions. I say in -.roLt oart lorn wat"''".r'' •" *'^ P'P^'^'''" '^ ^°™ -»-' -<1 where tea7^^^^^^ from water are they .n any degree separated ? that is has the affinit/ex eS bv ew b dilsT ?h " ""TJ 't^" ^'^^ "'^ ^""^^'"^^ -^'^ carbon'trZhes von^ the Wo ; . T I' "'""' ''*"' " ^°'«^ *° ^ff^^'t 'J^'^ decomposition be- yond the force that .sevolved in the formation of the new body (Law IV^ If then ^ "e on light, U 1. ci.vu- ;u^ certain amou.it of heat will be re- • Carpenter's " Human Phyiiology" p. 426 "^ t Carpenter's " Human PhyEiology" p. 429 quantity of iy wood 3d '1,7^ J''' " "^^ ° "'^ ''^"''''^' <"' ^ ^"PPO'e, agiven coal as there warcarblilr . ' " '" ''' combustion than would as much char- is not at al lerene^U Z,?:^^^^^^ .^ combustion j but if the affinity between them of heat evolved bv U,e 12! ^/. . ' u ""^ '^"""^^ '"''" *° ^"'""'<=t from the amount O-e ? n Ihe woS sTt noM"! ''""^^ f '"* "'^-^^ «=«•««"« between the H an^ 21 quiMd for II 'ly given amount of prowth and dc lopment oxhibitod by the plant ; bat IB, a quuntiUtivfi relation must exMt between ttto force supplied and the vi- tal forcfi pnt forth which depends upon it ; and thi« is seon to bo the cam in the moat striMng manner, for aeoordinj; to Bou8lnp?ault " the same annual plant in uniting at it« full development and Koing thnjugh all the proeeesof f flowering and maturation of ite seed, everywhere reooivea the sauio amount v,i ^.lar light and heat, whether it bo grown at the c(|uator or at th« temperate lono, it- rate of growth being in a preoiBcly direct ratio to the amount it roceivcH in any iron .irae. Very much the same thing is seen in the oaao of the lower oold-blw led anifc A though what ia the nature of t»-3 relation oxiHting here between the pr -^al afli vital force, I do not pretend to say ; it may bo that the former morel) Tniahep a neoessary condition for the evolution of the latter from other ources ; -t may be changed into it directly; or again the heat may alter its li rm and b oihin^ chemical force may so pass into the ■ ital ; be this as it may, the relation exists ;ind is well seen in the case of the ( -uttacea. For 1, the variety of thoir t '•m and organization (which may bo rc^r ,ded as so many varied manifestat rf the organising force) increase as we pwss from the polar seas towards the (;• or the number of species thus augmenting .'reatly as we go southward. 2. T, terenccs of form and organization are »< t only more numerous and more ohw »c- tcristic in the warm than in the cold rej; >n8 of the gloln) ; but they are also in important. 3. Not only are those Crusta, -n which are most elevated in the sc defieient in the polar regions, but their r(>iative number decreases rapidly as « pass from the equator towards the pole. 4 The average size of t he Cnutacea o; tropical r^ons is considerably greater than that of the tribes inhabiting frigid or temperate climes. 5. It is whore the temperature is most elevated that the peeuliarities of structure which characterize he several groups are most strongly manifested. And 6. There is a remarkable v oincidence between the temperature of different regions and the prevalence of certain forms of Cnutacea.* The rate of performance of their functionb in cold-blooded animals depends much upon the temperature in which they liv' Now as the respiratory process is an exponent of the rate of life of any animoj that is of the rate of chemical change taking place in the organism, it follows t^rom the above that should this be stopped, the length of life of the animal will be in the inverse ratio of the temperature to which it is exposed ; and so we nd it, for when firogs were con- fined in a limited quantity of water and not a owed to come to the surface to breathe They died in 12 to 32 minutes when the water was 90° " " 35 90 « <« '* 72 " 350 375 " '• « 50 '•' " 367 498 " " " 30 At the lowest temperature mentioned the prolongation of life was not due to torpidity, for all the functions of the animal were p- rformed, but slowly J ♦ Milne Edwards " Histoire des Cruataces iome iii pp. 555 et. seq. quoted by Car- penter in his article in Phil. Trsn. 1850. t Dr. F. W. Edwards " On the influence of physical agents on life." [■Is 'i. 22 In the production of larvae ft-om the eggs of insects, we see very much the same rektion between heat and the vital force as in the case of plants for he rate of development is in the direct ratio of the heat supplied, and Ihe fina transformation may be accelerated or retarded at pleasure, within certain irnft^ by regulating t^e amount of heat which they receive ; but i; every case-in eZ of hesameinsect-the same amount of heat is required, and must be supplSto effect the same transformation.* U rLT'f'^ ""^'""^ '" ^^' P*"*^ '^ S^^*^""" '» warm-blooded animals ^no doubt due in great part to the regularity of temperature that they are ca- paWe of sustaining under nearly all circumstances, and which is necessa^ to the continuan^ of their vitality, and I would be inclined to think (though I cannot anywhere find It so stated) that the temperature of warm-blooded animals de- creases as age comes on from the single fact (if it be a fact) that the period of gestation is prolonged in accordance with the advance of age Besides the influence exercised by light in the decomposition of carbonic acid and ammonia m contact with the green leaves of plants, there is no doubt that It IS a force which IS extensively used in the process of development, that in somecases at least, it determines the manner and direction of growth in a very remarkable degree A very curious example of this kind is furnished by the experiments of Mirbel upon thegemm* o{ Afarchantia poIymorj>Ju,. He found after thoroughly testing the matter, by repeated trials, that during the development of these little discs, stomata are formed upon the side exposed to the light, while root fibres grew from the under surface; and it is a matter of indif- ftrence which side of the disc is at first turned upwards, since each has the power of developing stomata or roots according to the influence it receives.t This division of my subject might be almost indefinitely extended, but I have not attempted to do more than notice some of the more salient poinL belonging to It, which IS all my space will admit of. ^ Higher in the scale of organization there are to be found such facts as the influence of light m the development of tadpoles into frogs ;-multitudes of the like instances will present themselves to the mind of every one Finally to test a theory we examine it in all its ramifications, and if it be found to be absolutely opposed to fact in any one case it cannot stand. So if any physical or vital force, however inconsiderable in amount, can be shown to 5" ^'f '!!!;? ?' "^^""^ '""'^ **^°''*'^' «« ^« «^««^« >ts operations, which do not proceed from some antecedent physical or vital force, such theoW can no longer be entitled to belief or consideration. But if, on the other hand, there are vital forces of which we do not know the antecedent force, or that thei have any except from analogy, it is the business of the holders of this view to en- deavour to clear up, and show the connection between such forces and their cor- IhfZ'tw" '° ' Tt' '' •" '^' '"'"^°'" '"'^^■' 0' »o «l>ow, if such be the case, that none exist, and so destroy a false hypothesis. •Carpenter, PhH.Tran. 1850. t Carpenter, Phil. Iran. 1850.