«»-vJ ■ i,«. IMAGE EVALUATION TEST TARGET (MT-3) !^- A^ L^.< fc 4^^ :/. 'm "W 1.0 I.I 1.25 150 •i^ IIIIU ^ 1^ 1^ M 1.8 JA III 1.6 V <^ .A ^r()diolio of the coal-measures are still un- certain as to their affinities. Thoy do noi come within the characters of the genera Cai'diiiia, Anthracosia, itc, to which fossils occurring in similar situations in the Hritish coal-fields have boon referred. They aro all thin shells, marked with growth lines, but destitute of other ornamenta- tion, and, so far as can bo observed, without tooth. In so far as external foi'm is concerned they may all bo referred to the genei'a Modiolu and Anodon. J?ut mere form may- bo a very fallacious guide, and 1 shall notice what seem to me to be the distinct specific foi'ms under the jirovisional name Naiadites, intending thereby to expicss my belief that tboy aro probably allied to the Unionida\ They aro cer- tainly distinct from any of the shells nf the marine carboni- fermu limestones, and ai-o never associated with marino fossils. It is possible that their neai'ost living analogue is the Bi/s'iO-anodonfa of D'Orbigny, from the River Parana." At the same time five species wore described, and indica- tions were given as to their local and stratigraphical distribution. A sixth species was subsequently discovered, Vol, X, p. 39. r I r Bivalve SkelU of Nova Scotia. 8 and another roforrcd to the Hamo ,i,n'oiip has since l)Oori found to belong' to tho <,'crius Anthraco^ia or Carhonicohi. Ileforo the ))iil)li(ation of the second o lit ion of " Acuilian Geoloijiy" in IHIJS, I iuid sent spociinons to my friend, the late Mv. Salter, of lae (ieologicul Survey of (Ireat Britain, who was at the time studyint^ the Hritisji species, and he described tliena with some other fossils from Xova Scotia which I had placed in his hands, in a paper in the Journal of the Geological Society' with figures of three of the species, which he referred to his two new genera Anthracoptera and Anthracomya, then recently established for the British species. Ho thu- diopped my genus '• Xaiadites " aiul substituted two other names of later date. I might have objecied to this, but I have made it a rule never to raise (lucstioiis of priority or of mere nomenclature, and I I'elt quite sure that Sailer was not a man to do any injustice, while I fully recogni/od his superiority as an authority on fossils of this Uind. There was, however, a more important point involved, having relation to the whole question of the conditions of accumulation of coal. Salter held the shells to be probably marine, and on this ground my name Naiadites was objectionable to him, while one of his names, Anthracomya, implied the idea of burrowing creatures allied to the ATya or sand clai Now, throughout the whole thiclcnes.s of the coal-formation of Nova Scotia, there is an entire absence of the species of marine moUusks found in the underlying marine limestones, while the bivalve shells in question occur almost exclusively in the coal measures and are not found in the ailmittedly marine beds. The question was an imjjortanl one with reference to the mode of accumulation of coal, a subject then engaging my atten- tion ; for though the occurrence of a i'ow e.vceptional bods holding marine shells might be explicable as the result of occasional invasions of the sea on beds usu;illy beyond its roach, the association of those shells with the Ijeds of coal was so constant and intimate that if they could be proved to be marine, a similar conclusion might naturally bo 1 Vol. XIX, p. 80, 1863. 4 Notes on Sperimenx. roachod rcs|)ocling tho coul itself, utui even Horiu' of tho plaiitH associiitwl willi it. 1 ihorcforo submittod to Mr. Sailor and pulilishcd in my new edition the foilowinii; tiictH, teiulin^f to hIiow that niy so-called Nuiuditos were froah- water or ostuaiine shells. 1. Under the microscope the thicker shells, even those of the Anthracoptera type which most resemble marine species, present an internal lamellar and subnacreoiis layer and a thin layer of vertical pi-ismutic fibres, covered with a well developed epidei-mis in the manner of the shells of the Unionidieor freshwater mushcls. 2. Th'j liii;ament unitint; tho valves was external, and there seem to iiave been no hinge teeth. The shells wore closed or very slii(htly o])en posteriorly, and in s()mes])ecies there are indications of a byssus or '• beard " for attach- ment. The jfenoral aspect is in some species that of mussels, in others that of Unios or Anodons. 3. I know of no instance of the occurrence of these shells in the marine limestones or in association with species known to be marine. 4. The mode of their oe-cui-renco precludes the idea that they were burrowors, and favors the supposition that they may have been attached by a byssus to floating timber and to one another. 5. Tlie attachment of shells of spirorbis to tho outer sur- face of many specimens seems to show that they wore free in clear water wlicn living, wliile the dense piling together of these f^hells in some beds almo^tt unmixed with othei- material, and their occasional occurrence in patches asso- sociated with fossil wood, points to the same conclusion. (I. They are associated witli tine sediments, vegetable de- bris, the crusts of minute crustaceans and remains of fishes more likely to have been iidiabitants of fresh or bi'ackish water than of the sea. On these grounds, and being unable from the specimens in mypossession to make out evidence of generic distinction,! continued to use the name Naiadites; using however, Salter's names as subgeneric, so as to keep our species in harmony with those of England as described by the Geological Bi'Hi/ve Shel/s of Nom Scotia. 6 Survey. The miitlor vvhk \vAl in this form in my cilitioa of 18(;s. It Heoms, hovvevor, tlmt in Hiibstitiitin^' a ti^'uro not poriiaps very uccuratoly druwii fi-.>m a flutloneil Hpoci- mon, for llio fi^nire which Suitor hml ^tjivoii fi-om an ani^uiar and com I trussed oxam|)lo, I caiHcd Homo misnndoi-standiiiLf as to one of the specios, loadiiii,' to tlic HUpposition tiiat one of thoHo named by SaltcM- wan dirtoront from that which I roco^'ni/,od by the same name. Tho ditforonco was really in state of pi-esorvation witli some inaccuraev in drawing' in both canes, I Mhall <,mvo below copies of these imperfect fi^'ures, which however, represent actual appearances wliich may mislead collectors, alon« with a ti^nire carefully copied from a younawson. 1 2 ss^^- l''i>:ures 1 ^o'^. — N(ti• «-Small specimen .natural size and enlarged. F,g. 7.-Large specimen, natur^size. Fig. 8.- SJSSr'EnTar'ged Tlf'' ""'^^^^ ""^ anterior end sl^tly * !( I Bivalve Shells of Nova Scotia. U Supplement to Acadian Geology, 1800. p. 4:5 (as xVaiadites) ; Salter Journal of Geological Society. Vd. XIX, 1868, p. 79; Acadian Geology, second edition. I8(i8, p. 20;; Wheelton Hind, Journal of Geological Society, Vol. J., 18 ^pper Coal-formation, Pictou. Supplement to Acadian Geology, last edition; Salter, Journal of G.olog.ca Society, second edition, p. 205; Wheelton Hind Journal of Geological Society, 1. c. ' Shell elliptical, smooth or with very fine concentric lines. I'.pidermis tliin, in many specimens absent. More than twice as long as wide. Anterior margin narrowed in front of beak. Beaks about one-sixth of the length from the an- terior end. Posterior end somewhat narrowed at extremity This species is usually found in gray arenaceous beds of the upper coal-formation and the millstone grit. It is com- paratively rare in the middle coal-formation. All our museum specimens are from Pictou and Sydney. The species may be compared with A. lanceolata of Great Britain. Bivalve Shells of Nova Scotia. 13 4. Anthracomya nvnlis, Daw son. 11 BnlaSd^T'^""'""""''" "'""■'' ^^"^^'"'" ^'arboniferous, Pamboro. Supplement to Acadian Geology, I8.i0; Salter, Journal of Geo- r'4r. 'wf ';,'■"; r'Tv '''''''^" ^-ology. second edition.. p. -05 , W heelton Ilnul, Journal of Geological Society, 1. c. Ih.sspeces has the general form of the .smaller speci- meuH ofelongafa, but i. broa.ler bohiud and more tumid in tront, 80 as to be at once distiniruishable by the eye It occurs .paringly in bedH from the millstone grit and lower Carboniferous to the middle coal-formation. Oiu- specime,:s are from the South Joggins, Riversdale and Parrsboro. It may be compai-ed with A. dolabrata of Jingland, but is always much smallei-. 5. Anthracomya obtenta, Dawson. 12 Cape^'lSeto/al'tumUiT ''''""' ^'''^'^^' ^^oal-fonnation, Mabou. Acadian Geology, second edition, p. 205, (as A. ohiusa, a name Which 1 hnd was pre-occupied for a species now included in tins genus.) ];] *• -^j*^- J'''""'^"/'"""^""'"'^^'''^"'"' <^'Oal-forniation Pictou restora tionof a Hattened and imperfect specimen, enlarged x 2 ' M Notes on Specimens. (Jonoriil foiin rounded, and piobahly when not chan,'?ed by prcHHiii'c tumid. Anterior end l)i'0{id and ubi-uptly i-ounded ; hinge line Hti-aiijht. Hcaks raisoii and Komewhat near the front; lower and po.sterior inari>;inH broadly i-ounded. shell thin, wrinkled when flattened, strongly marked with growth-lines. This species resembles sjmewhat A. Adamsii var. erpansa, England. It is rare. Our only specimens are from McLel- lan's Brook, IMctou, and .Mabou, in Cape Breton, and are mostly rtattoned, except some very young examples from llie latter ))lace. In addition to fragments of plants and comminuted debris of vegetable 'natter, the beds holding Naiadite.s, contain a number of other animal remains, constituting a peculiar fauna altogether different from that of the lower carl)oniferous marine limestone, and also in many re8|)ects distinct fi'om tiiat of the sandstones of the millstone grit and upper coal formation. This fauna, though not that which we would expect in fresli-water lakes or streams under ordinary conditions, seems of such a nature as to be appro])riate to bodies of shallow, fresh or brackish water loaded with vegetable matter, or to wide and sluggish creeks travoi-sing the g'eat swamps of the period, and occasioiuilly wiilening into lagoons, receiving much fresh water from the land, and having but little communication with the open sea. The beds supposed to be thus deposited are carbonaceous or bituminous shales and' laminated, impure limestones full of earthy matter, and blackened with bituminous and carbonaceous debris, In addition to the bivalve shells in question, they contain vast numbers of minute bivalve crustaceans. (Bairdia and Carbonia) ' Species of Eurypterus, JJiplosti/lus and Anthropalaemon, re])resenting crustaceans of higher types. Great numbers of the little Spirorbis carhonarius are also attached to many of the i)lants and other fossils. Numerous scales and teeth of ganoid fishes of the geneia PaUvonixcus, Mhizodus, &c., ' Rupert J Olios, London (Jeologiciil Miigaziiie, AuBusit, 18iM, p. 2t'.l), and .June, 1889, p. 356. Bivalve Shells of Nova Srolia. 15 alHo oci'iir, and teotli ol .lipnoid H.slu's (' tetwdns), uIno various HpecioH of sharks {Ctenopty chins. Psammodus, Diplodus, kv). Soino of tlieso sharks must liavt" attained to a considerablo wizo, and the}' no doubt found accoss to th^? inhuid waters by the outlets comrnunicatinii,' with the sen, and wore attracted to visit tiieso C'onij)arative!y impure higoon.s by the ubundunco of food which tlioy atfordod. ' Very i-arely there have been found in these beds bones of ain])hibianH and shells of pulmonatc sun\U,{Pupa vttmia. &c.). Animals of these kinds no doubt haunted the mai-gins of the lagoonw or creeks; but only occasionally left their remains in deposits accumulating in tiicse places. We ])erhaps obtain a glimpse of purer inland waters, similar to ihose of modern Canadian lakes, by means of u remarkable shell, discovered by Mr. Weston, of the Geo- logical Survey, at the .South Joggins in 1893, and which has been described by Mr. Whiteaves, K.d.S., under the name Asthenodonta Westoni. - It resembles in general form the large jiearl-uiussel of our modern lakes. (Manjaritana maryaritifera L.) and some sjiecimens are no less than nine inches in length, and of somewhat massive thickness anteriorly. It was found in a sandstone with drift trunks of trees, and may have come from some distance inland. Such a .-hell could scarcely have been a companion of our little Naiadites or Anthrocomyiv, and points to more favoi'- able conditions for fresh-water molluscan life in lakes or largo streams in the interior of the continent. Conditions tiivourable to such mollusks vvere probably, as I have elsewhere suggested, more i)i-evalent in the later Erian or Devonian than in the < 'arboniforous. Hence the occurrence of such large Anodon-like shells as Amniyenia CatUkillenm, Hall in New York, and Anndon Juke&u in the Kiltorcan beds in Ireland. The above discovery however now gives reason to believe in similar conditions as oxistinn' in higher grounds contemporaneously with the great coal swamps of tlie low plains of the carboniferous period. ' Notices of this fnuiiii will bu found in Aciidian geoloev, pp. 2(i" et sen nndsuppleinentH. * '' - Trans. Royal Society of Canada, Section iv, 18!^ 16 Notex on Specimen. Tli, Lepidodendra, Oalamitos and Cordaitos,and with dark bodies of impure water full of vegetable matter. The faunu' of these districts might bo equally ditforent. We know little as yot of the upland fauna; but may hope for more discoveries in this direction, especially in countries like Xova Scotia and Cape Breton' where there were elevated districts in the midst of the ai oas of coal accumulation. Appendix. Note on (Janus Carhonicola, Mcf'oy. (Anthraromi, Kinj;.) This genus, which occurs abundantly in the Coal Forma- tion of Great J3ritain, is represented, so far as known, in Nova Scotia by only two small species, both from the lower ])art of the Coal Formation, <,r possibly from the Lower Carl.oniferons. One of the.se is (\ atujidata {Naia- (Utea anipilata, Acadian GeoloMry, p, 204, tig. 4(;.) \t jg tVom Parrsboro, from beds holding tbssil plants and, so far as known, no marine shells. The other, 0. liradorica {Anihraco&ia Bradoriea, Ac. Ceol., j). 314, tig. lli'J b) is from a shale supposed to be Lower Carboniferous, at Haddeck, Cape r.retoii. The affinities of these shells are at present^ uncertain, but will probably be discussed by \)\: Wheelton Hind in u ibrthcoming paper, its associations would seem to iiKifcMc that the habitat of some of the species was simi- lar to that of the genus Anthraromya, which at Parrsboro are found in neighboring beds. The figure of C, Bradoriea is i-eproduced here to show the characteristic form. Carhonicola Bradoriea. I I