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CIHM/ICMH
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1
2
3
1
2
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4
5
6
DEPAKTMKNT OF THE INTERIOR
D. 8. GEOLOGICAL AND GKOOIIAPHICAL SURVEY OF THE TEBaiTOBIBS
F. V. IIAYDEN, Oroi.oowt-in-Ohabo*
MISCELLANEOUS FUBLICATIONB-No. 12
HISTORY
OK
ISrORTH AMERIOA-ISr
PINNIPEDS
A MONOfiKAIMI or THE
WALRUSES, SEA-LIONS, SEA-BEARS
SEALS
OF
NORTH AMERICA.
Bv JOEL ASAPH ALLEN
AisMantin lln Mimeiim of Comparative Zoology at Cambridgi
Special Collaborator of the Survey
WASlMNCiTON
GOVERNMENT I'llINTINO OFFICE
18S(»
<• ^
:• :
PIIEFA'I'ORV NOTE.
United States Geological and
Geograpuioal Survey of the Teuritories,
Washimiton, D. C, 'fitly 1, 1880.
Tlii' picsciit si'iies ol' iDonofiiaplis of the North Anit'ricHii
Piiiinji(
nl>li-
cation of the part already i)repared relatinji' to the Peiniipeds,
As nearly all of the K])ecies belonjiinj^' to this {:ronp foinid in
the nortlunii lienMsjdiere are nuMubers of the North American
fauini, the ]n'esent treatise is virtually a mono}.';raph ol all the
species occurring;' north of the ecpiator, anrinter, it i»roves to be, to only a small
degree, a rei>etitioii of the account given by ]Mr. Elliott, also
rejuoduced at length. The history Captain liryant gives of
the changes in the numbers and relations of the different
classes of these animals at the rookeries, under the present
system of management of tl>e Fur Seal business, forms a valu-
able l)asis for generalization in regard to the future regulation
of this industry, and is aiso an important contxibution to the
life-history of the species.
The cuts, some thirty in munber, illustrating the cranial char-
uctt'rs of the Walruses, were drawn for the present work by Mr. J.
H. Blake, of Cambridge, and engraved by ^lessrs. Bussell and
Eichardson, of Boston. The Survey is indebted to Professor
Baird, Secretary of the Smithsonian Institution, for aseries of six-
teen original tigui'es, engraved «y Mr. H. H. Nichols, of Washing-
ton, from photographs on y «M,d. vllustrating the skulls of CaWoWti-
nus ?( rsin MS, Peale's ^'■Haiivhwmaantarcticuii,''^ (Jystophora crLstata,
^-^.»f-^^f-:rf*f-,^
(»f London for electros of (7 ray's ^^HalUyon richardsi," and of a
seri''H yf historic tignres of the walrus i>ublished in the Society's
»' l»roceedinj;s," Ity tiie late Dr. Gray, and to the proprietors of
"Science (Jossii),"for electios of the full-lenj;th figures of seals.
These were reci'ived throuyh Dr. Cones, who also furnished the
full-length \iews of EnmetopiaN .stelleri and CaUorhinus ursinns.
Mr. Allen desires nie to express, in tiiis connection, acknowl-
edgnients of liis indebtedness to IMof. Si)encer F. Bdrd, Secre-
tary of the Smithsonian Institution, and to Prol". Alexander
Agassiz, Director of the iNluseum of Comparative Zooh)gy, for
the liberality with which they have placed at his service the
rich material relating to this gTon]) of aninuds ( )lt.K.\ 1 1 >.K— Wiilnmcs 5-180
Sjuniiviiiy .I
tii'Uriiil olMcrvatiiuis iuid cliiinicteM of the j^roiip 5-12
(Jfiicia 12-14
S.Miii|i»i.-( III till- iii'iina 14
til'IIUS (•IHIII.EM > 14
S.viniiiyiiiv anil lii.stmv 14-17
Species 17-23
( M)Oiia;m s KosM Alii s— Atlantic Wi'lrus 23-147
Syuoiiviuy anil liililiii!;iai>liii'al ict'iTcnces 23-20
Kxtcrnal iliarartcis 2()-;)8
Srxnal ilill.Ti iiics 3S-43
Inii'.iilual viiiiat inns anil vaiiatiouH dcprmlL'Ut npou age 43-45
McaHiilrnirnts 111 .sliiills 40
Unutitii)n 47-37
l''o«sil ri'iiiains ."i7-fl5
(rfogiapliiral ilistrilmliim. pivsrnt ami pUHt 05-79
f '(last lit Ndi til Ainriira Oi5-71
("oast lit' Kniopc 71-79
NoinciiclatuiT 80
EtyuKdoiiy bO-82
Litcratiiiv . , 82-107
( icnt ral history 82-92
Fi)j,nivs 92-107
Habits and tlu« cliaBe 107-133
Products 133-134
Food 134-137
Fiiui'tiiins of the timks 137-1""
Knciiiics 13H-
Diiincstiiation 140-
Oiioha;m K oiiKsif—l'acitlc Walrus 147-.'
Synonymy and liildiiijiiaphical references 1
Kxlenial iliaraeters and skeleton 147-1
Measurements of ' deton 14!»-1
Measinenielits ot . ..tills 15
Uill'ereiitial eliaracters 150-171'
Nomenclature . .. 170-171
General history 171-172
Figures 172-174
< teou;rapliical distribution 174-178
llaliits, food, commercial products, and the chase 178-180
Family tVl'AKI ID.K— Fared Seals 187-411
Synonymy and characters of the group 187
Technical history 188-207
Higher gi-oups 188-190
(leuera 190-193
Species ia3-207
IX
■j.iii
•j;i7-j:is
J44
X TABLE OF CONTENTS.
Page.
t'li.aiirliT.H 111' I he I'lNMl'KIH \--('()ntuiiir(l.
KmiiiUv OTxVIUIDJ:— KmitiI ScalH.
S\ iiiipsiH of till, ^iiiiia unci .spiicics 2U8-ai3
M.vlhic.'il :iu(i uiiilitiiiiiiMiibli' spcciea 2U->l(i
tliDm^liliical ilistiiliiilion 216
f'o»,sil ( Itiiiics ''l?.-!'"!
Milk (Iciititidii 2''l-2"4
•Infj,'iiliiiilips of (Iciititimi ^24
I'lLsition of (h,. 1,181 upper piriiiiincnt mou.r ^iri
(rfui rill oliaervationn "ja.'i-'J''"
Habits ""' .„"-
J J 1
Products .,.,g
Pestrurlioii of Fur Si iils for tiirir peltries i;'Ji*-J31
Genus Ml MKTOIMA.S ._i;j1
Ei;Mi:roi'iA.s biklleui— StfUci s Sea-Lion 2;i'j-274
S.vuou.viiiy iiml l)il)lio-raphi(Ml n iVjicncos L':)2
Kxtt'ina! cliaraclir.s
Extoriial niia.surt'uicnts
Skull ' '"
.McaauriTiiiiits of skulls ..
TlM'l ll ^rji,
Sk(li't(Mi 24(1-241
ili'a.siiiriUfUt.s of skrlt'ton 212-244
Sexiuil, ailolfsifut, ami iuilividual variation 244
Gpofir.ipliiial \ aiiatiou
Comparison with allied .speeies 244-24
.McasiiiTiiii ills of skulks of OTAiilA JliDATA 247
Geouiapliical iKsiiiliiiliuu 248
General bistm y and nonienelature 248-254
Habits 254-274
Genus ZAUii'iirs 275
Zai.01'hi-,s ( Ai.itniiMANrs— Califcuiiiau Sea-Lion 27ii-;il2
Syuouyniv ami bibliosiiapbieal references 270
External eliaraelrrs ]'_' 27ii-27^
Younu .,7^
•'''l"!i'- 27b
Si^«; 27>^-28;i
External measurements 2711-280
Measuieiiients of skeleton (d' female 281-283
^"^"11 283-285
ileasnieiuenls of skulls 285
Dentition .jjjg
Sexual dirt'erenees 287
Variation with age 287-289
Coraparison with allied speiiies 289
Geographiealdistiibiition 280-291
General liistory and nomenclature 291-290
H"l"'« 296-312
Ge.nusOAi.LoimiNis 312-4l(i
UALi.oHHiiM s imsLNUb— Northern Fur Seal 3i;j
Synonymy and bibliographical references 313-314
External eharaetiTS ;j14
Color jjl^
Pelage ;j|,-,
Size 31»-319
External measurementa ... 319
Ears ;j2u
Fore limbs ;j2i)
Hind lindis 320
TAIJLK OF CONTENTS.
XI
Page.
Chiira«tci sol' till- I'i.vmikiha— ('(intinind.
raniily OTAIillD.K— Kiiicd S.uls.
Gi'nuH('ALi.ij|;iii.Niis.
CAi.LOiiiil.Ni'8 UU8I.NLS — Northern Kur Seal.
Skull 820-323
Moasuvcmonts of skulls 323
Tc-etli 3i;4
Skclctdii 324-32t)
^Mtasiin nil Ills of ski'letun 325
Si'XUiil (lill'.iiniiH 325-327
Difl'iicnccH iiHuhiii;; frtiiii age 327
Inilividiiiil variiitioii 'S'JS
Comiiaiiwin with alliccl Hpocies 329-331
Mcasurciiu uts nf skulls of AliCTOCEPHALUB AUfiTIIALIS 331
Oi'Diiiniiliiral ilistriliution and niifiration 332-335
GeiH'i'al liistmy and ncninuclature 335-33'J
Fijiuii's 330-3 U
Ilalnts 341-371
Till' cliasi! 371-3Tr
Miidr lit Caiiluri- STl'-:;"-
llistiiiy aud pKisjiicts iif the Kur Seal business at the Prybilov
Islands 37H-3«1
EufUiifS (if llic I'm- Seals Ilj^l
History of tlie Kiir Seal Fishery at thi; I'rybllov Islauda, Alaska.
iVimi 186!) 1(1 1«T7, liy Ciiaui.hs Huyani' :)i?J-tll
I'reliiniuary aud acueral oliservatioiis ;ihj-:j:<.-
Kecelit ehauuis in the lialiits and relative uiimbers (d' the
ditlerellt classes (if Seals :il?>?-;i!(5
Cause (if tile eliani;es in I lie lialiits (if the Seals, &e 3ilh-4U2
Albinos and sexually almoriual individuals 4(13
Descriiition of the youni; ; vaiiatiou in eohn' with ago, &c. . . . 403
M.iltin.i.' 404
Sexual orj^aus, &e 405
I'ower (if siispendiug respiration 40G
Natural enemies 400
Eflbet of elimatie iutluences 407
Number of Seals reiiulred for the subsistence of tlie natives. . 409
Winter resorts aud habits of the Seals 410
Family PUUCIDiE—Karless Seals 412-7.10
Characters of the tirmiii 412
Technical liistory 412-4«li
Highet groups 412-414
Genera 414-421
Species 421^00
Classilication 400-407
Synopsis of sub-families and genera 401-463
Synon\ niatic list of the species 463-407
Geographical distribution 407-409
Fossil remains 409-481
North America 409-470
Europe 47(i-481
Milk dentition 481-484
General habits and instincts 484-486
Food 486
Enemies 487
Mitiratious 487-491
Locomo.^ion on land 481-490
Seal huutin); 490-540
XII
TAMLK OF CONTliNTH.
Chiini' HiH (il llic I'lNsirKiifA— Colli iiiiiiil.
F .iiiil.v I'llOCID.K- KiiiliHH SciiIh.
Si-aliiif; iliHii'iriN
WimI I in I'll III mi I
Nrw rdiiriilliiiiil
illin MllMTl 111 "(Mrl'lllllllll ' SlIlM
Niivii /I'liililii ami Kiini Sni
Wllltr Sill
('aH|iiiiii Sisi
Niirlli I'acilii
Son III rarilir ami A iilanl ii' Siiim
MiIIioiIh of i:i|iliiii , ,V 1
Slioif liiiiiliii^
l')Hii|iiniaii\ iiiiilioilH
Hv tlliMlls ilT IlrlH. .
'Jill' HI -a I I io\ , ..... ...
'I'll!' .sfal lioiik
'I'llr ■ SKiiiikla"
Ici' liiiiil ill'.:
In llii' liiiHol' liolliiiia
(Ill I 111' 1 oa.sl 111 Si'uli. Ilami
In I III' .Ian Mavi II Slim
|laii;:i'i K anil iiinri tajnlir.s ol' \ir liiiiiliiii;
.SpcrirH liniilril
Alililiilaiii 1 III' Si alH III |iai I Iriilai liM'iilllirH
I'rodinlH . .
I'li'iiiiralion 111' llii' |iioiliirlM
WiiHli Till ili'Hli III linn 111' Siiils
Di'i'i'i'iiHi' rriiiii in.jiiilii'i'iii.s liiinUiit;
Si'iii.s anil Sral liiliilili); in llirnlilrii llinr in llir ( iiilt' ill' St. laiwri'iici'
Sub-Ciiinily I'liiii in.i,
fii'nim I'liiii ,\
I'jioi A VII I i.iNA— Ilailinr Si'iil
S\iionwii> anil liililii>nia|iliiriil 11 Iii'iihth
K\ 111 nil I I'liarai Ins
HlHlini'livi' rllllliU'll TH
Iliiliviiluiil iiml HI' X mil vaiialioii ... .
Mt'iiMiii'1'tnrnln iif (lie HkiillH
-r)4»
.')41l-,'^^,0l
:.::.{
.■i.'.:!-.",.-.7
.-,.''.7
.V)7-.'i,-.!)
.'"i.''ili-.'"i(i7
.")i!.*>-ri7 1
liTI-.W!
.-.74
r>7.''>-.'>H4
.'H4'-.'iHH
,MI7-»K»
.SIt7-tlOI)
i>iHi-(iu:i
IID.'I-DU.'')
U(I6
01)7-1114
Ulilt-lllll
•11 -.Mil;!
IIM-Iiltl
uii) rdu
(illl-tlL'il
iYM-im
tw()-(i;rj
o:)'j-ii:i7
ii;i7
urn
ii;iu
TAIlLi: OK ('ONTKNTS.
CIlllliK I' IH III lll> riSMI'I'.IllA -('(llltlllUl'll.
I'.uuil.v I'llOClK.lv
Siili liiiiiil\ I'iKii l\.i:.
(Ic'lillH rilni A.
I'lKlc A ( I'Ai.cil'llll.l I l.l|c|„NI.ANI)|l'A— Illllp HiMtl.
tii> ;iiiil liililiii;;rapliiital icfi^iciKtoM
Kxiriiinl rliiiiai'lciM
SUiill and mLcIi lull
MraHiirriiii'iitH 1)1' I hi- Hki'li'tnn
MiaNiiii'iiii'iilH III' III! sKiill
(ii'lii'ial IiIhIiii V and iihiih nrlaliirr
• li'ii:;ia|ilil< al ili^l li lull inn
II aliil-i, iHiiiliirlM. anil liiiiilln;:
(JllllH lll^l ninriKir \
IIimHIIiiIIKh A I'Ahi I M A — Itiiiliiin Sial
Syiiiiii.Nins anil liililliij,'ia|iliiral 1 1 I'liinrrM
KxliTiial I liaiaili IN
Si/i-
tii'iii'ial liiHlni \
Iirii;:ra|iliiral iIInI i iliiillnii
lialiiK
(irllll^ II Mil IIH.Ill H -
< ii-ni-ral IiInIiii \ anil iliHriisMJun nl' the ''(ii-nim IN'ha " iit' Sropoli .
IlAI,|rl|i|;UI - i.in I'l ,■^— (ila\ Si al
S\ 111 my in \ anil liilillnurapliiral irrrri'iirnH
• lOsIri nal iliaiarli IH
MiaNlllrllH lit H III' mU nils
(li'iiV'iapliiral ilisli iliiil ion
(inirial li ill I II V anil niniii inlatiiK^
Ilaliils
(iriin« Mll.NArill n ..
Mi.NAi iii.s iiail'irAl.lH— WihI Inillan Siiil ...
(Miaiai Ills
I>anipli'i\s arriiiiiil
1 1 ills anil (iiissc .s ai riainlH. iKi:), I HP)!
(iiav H ai riinnlN. IXI'.I, |h7I
(■ill nil (llr W'rnl Inillall Sial.s. I Hid
Aiial\Mi« anil ili.siii.ssinii nl' I In Intiiiiniiii^
Alllnilli n nl' I III' .laniaiiaii oi I'liliii Siiil
( li'nL'l;i|illlr.ll ilisit ilillljiill
Sail rainih CVSl'orillilMllN.K
(il'llllrt ('Vhl'll'IMKA
(!VKHiriiiii!A I iii>i Ai \— llonilcd Sial .
S\ 111 iiiv Illy and lillilin;.'iaphi('al ii riTriir.tm
Kxiri nal rliarai lii.s
Skill Inn and sUiill
.Mrasiiii nnnl.-i 111' HiiiillK
MiiLsiininiiilti 111' till' Hliili'tiin
liin^iapliii m1 iIimI I IIiiiI inn and nii mil nniiii'iirlatilKt
llalillM
Iliinliliu and jiiiidinls
XIII
Pa(jc<.
(14(1
(m-(M7
(M7 llal
(i.'il
(I,'.'J
flr.u-ii.'M
ll.-lj
(ir.,-i-(i7r.
(irir>-(i.';7
():.7
B,-.*<-(Itll
(ItiO
litii
(Kl'J lilili
lillll (170
(17n-(i7"i
117:1 -ii7ii
(17(MWJ
Ii7()
(I7(M17'<
tYjH
(17H-(iM|
(iKI-liK-.-
()«:;
(W'J-(i«!t
liH.'!- i;«:i
(lH!l-7lMi
(in'.l^li'.tll
(iiHi li'.i::
(IIM
(m.'i-(i!Hi
lillli-li'Jn
(i!lil-7il(i
707-7nx
7iiK.7'j:t
7llH
7(IH 710
7|ip 7l"i
17.". 7 M
-\H
7IK-7'JI)
(Un-TJl
7'J 1-723
7'j;i
7-j:i
7:'i-7»::
72.1 72(i
7'J(i 721*
7:111 7:1:1
7:12
7:i:t
7:1:1-7:17
7;w-71o
7ln-7H
741-712
XIV
TAIU.K OF CONTENTS.
Chara4'U)i'M of tin- I'inmi'I hia— Cciiitiiiiiril.
Family riKMID.K.
Sub- family— ('vi-i(pi'iii iiiiii.N a;.
.i: 761-704
I'liuiij viliiliiia 761
Plioca fotiila 702
Plioca fiiii'iilaliillcu 701!
Eritfiiatlms liarlmlus .. 703
m.stiio|ilioia fasi'iata .. 703
Hulicliiiiiis ;;iy|iiiH 704
ryHli>|ilioia ri i.slata 764
.Mariorhiiiiisaiicim' 'mtris "04
B. Adtiitidiin iiiiU I'lncrtii. n 765-774
Family OlMiii.i,M!>.v, 705-7IW
(Moha'iiiiK riixiniirun — .\tlaiitlo WaJruB 765-708
.Ailditioiial iiliiiiiciH 705
Sizi anil rstiiiial ainicanilKM! 705
(ii'oniaiiliiciil ilistiihuliou 700-707
Nova Zi'inbla
Kraiiz-.Iomf Laud
Aliiiiiilaii' (' ill Wolatenhobne Sound
SpitzlMljirll, &c
lirlaiid
SiipiMiHi'il pi'cHi'Mi'i^ of Walraflos in the Autarotio Soaa .
'I'lw 'Wall iirt a foriiiiilalilt> nnta^oniHt
Ciiiiosiiv ami liailcHsncsH of thr Walrus
I/oi iiniotioii : iiHC of tin) UukH ill climbing
KiHuriH of till' Walrus
Odobcmiig obemin — I'aciQo Walrus ,
Disliiliiiiioii
Family OtaiiiidjE
( )tai iiH at t he (iolapagos Tslnnda
FoHHil Olarii'S
('apt nil- of Sea Lions for menafjerles
Zalophun cal{f.
41.
Fi};. 2, p.
4J.
Fie. 'M'-
4;).
Vig- 4.1>-
u:t.
Fi^:. r.,p
ii:i.
Fiji. O.p.
1)4.
Fif.'. T,p.
'.»4.
Fi«. H,p.
!t4.
Fin. l»,p.
!l."i.
Fiti. 1", p
'.Hi.
Fi«. ll.p.
Kill.
Fi};- 1-. !'•
101.
Fi-. r!,l>.
l.-.:i.
Fili. M, p.
I-'Im I."i II
i.-.(i.
1 ."i7 .
1 IL. !•'. P-
Fi;;. Ii-i. p.
i:.s.
Fi;;. 17. p.
i."i«.
Fi-. 1.S, p
l.-.!t.
Fi^;. l!".p
I.V.I.
Fi;;. Jii. p
ICO.
Fi^'.'Jl.p
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FiL'. -JJ, P
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Fi^'.-J:i,i)
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Fi.-. •-•4, P
i(i:i.
Fi;.. -J.-., p
IGli.
Fi«. -jr,. p
104.
Fii; -7, p
Km.
Vi'A--Jx.\<
II 10.
Fi-.21l, p
100.
Imk- :•
107.
Fit'.iil.p
. 107.
Fi-. .'l^, p
lOK.
Fi-. ;)3, 1
. IIW.
Fi-. :;i, 1
. 101).
Fi;;. ;t.->, 1
. lO'.t.
Fi-. :ili. I
. i7;i.
Fi,;;. ;i7, |
. ;;,■>».
Fi-. -.iX, 1
.:ii7.
Fi;;. :i!l, 1
. :i'.'i.
Fi-. -li), 1
.:i'.'i.
Fi-. 41,1
:rj^'.
Fi;;. 4J. i
. :cj;;.
Fi-. lit, 1
. .')0:i.
Fi;;.4J. 1
. .V().
Fis.4.'-..|
. "iHO.
Fiu. 4(i, I
.581.
Fi-. 47, 1
. .182.
Fig.4H,|). '8:1
Fi(f. 4«, p. Udl .
Odvbimiwi rimmarut. skull nl' ((oinilf in pnilili' anil lowiT.juw from above.
OdAibniPuim rnnmariui, skull nl' fiiiiiilf fiDiii iilinvf.
Odob-iiiuH riiKiiKinin. .•^KuU nl !• miilr tVniii liclcw.
OlauH M;i-iiiis s " Udsuianis srii .Mnrsiis \iiivf;;icuH."
(•laiis MaLiiiiis's I'linus alniistniMis Ocriiui (li'nniiuici."
(H'Siii'i'.s '■ KiisMianis."
tiisiici s •■ Varca iiiaiiiia " (.\ililii)(ia tii Icoiifs .\iiiiiial).
(ii'sui r's •■ KoHiiianiM" (Innics Aniiiiill.. l.'iOli/.
l)r Vri'is 'Sia IIkisc," 100!I.
HisHil (Ji'iaiil's " \ValiU8s," lOia.
Martin's '• Wall-niss," 17ti.">.
l!llll'(ill's ■' I.r Mdl'Sl'.' 17*i."i.
OiliihiriiiiK iiIk^hhii, llilri- \ ii WH i)f licail.
(Mliihitiynim iilieiiit'<. sliiiU in imililc.
Odiilxi'iiiis riiKiiiiiniK, skull in pmliJi-.
Oil'ihifiniH iiJHiiinntn, skull liniii I'riiiit.
Oiliilni'iDiH ubi'siiK, skull rriiiii liniit.
(J(tiib(riiUK niHiiKirim. m ripilal viiw of xkuU.
(JduhiviiiiK iibexiin. iMii|iii.il \ ii-w nl' skull.
(Miihii'itiiH roHmaruH, skull IVom almvf.
(Jdiil'd'UUX (ibf.Kiis, m1;u11 I'l nil almvi-.
OdiihifiiiiH iibfKtiu. .Minn- skull fimii almvi'.
Odiibdniw nmiiKinin, Mnni- skull I'lnni above.
(Jddbunian obebitu, .miiiu- skull I'miu I'riiut.
Odobwnnn rumiMnui, .viniii.i; skull f'lniu t'niut.
Odobii'iiu.i riiKiiioruii, .skull I'lcmi liclovv.
OdiibiviiHH ohemix. skull riimi lirl.iw.
Odiihii'nnn rnKKinriiK. |ci«ir.j,,« ridiii aliiivi'.
Oddbiiuun (ibentin, lowri jaw I'linu abuvo.
Odobimius rimmaniH. Inwcijaw t'rnni side.
(JdnhiiiniH tilnKim, liiwci- jau I'lnni Bitlc.
nddbiiiitm riiHiitdiKu, liiwii .jaw nl' young from above*
Odvbd'uiiH (ihcKiiK, Inwi-r jaw nl \ nuug from nbovo.
Odulnriiiin riiniiiariDi, lower, j.iw nl' young from Hide.
OilnhirnuH iibi'xu)!, lower jaw nl' ynuiig I'rom Biilc
Odobd'iiiin ubt'sim. Cnnk'.s fl-ure of tlif aiiironl.
KumeUijniiH stelUri, li-uriB of animal
VaUi/rliiniin nrninun, li-urcM of animal.
(JaUurlii'ii^KK iiitiintin. skull nf fiinalr iu protilc.
('allurliiniig urtiiiniii, skull of I'l luiiln from ubovi-.
Cnlliirlnniin iimintiii, skull nf fiinali lower jaw.
('ulliiihuiiin iirniniiK. skull of finialr from below.
J'liiicd litiiliitii. auiinal.
. " llalichwriiM ant'irrliCHH," I'liiln, Hkull iu prollle.
" llulii'lidriin <(iiliir<:t.icuK," I'rali', okall from above.
" niilirliwrus iintdrctiKng," I'lule, Bkiill lioin below.
" Ilalichiernii antnrclicut" I'ealc, lower juw.
" tlalicyun richtirdgi" (iray. skull iu profile.
Fhiica firtida, uuinuil.
XVI
IJST OK IIJ.l'riTltAI lONH.
I'iu ■'", l>
n:i:i
Ki«.r.i,|.
)ll)l
Kili.V.M.
7'.' (
Klu.r.:i,|.
7-JH
V\H. M, |.
7211
Ki«. r,r., p
Tlsn
KiK. .'>«, I,
7;ii
KiK ■■•T, |.
711
Ki-. r.K, |,
71:.
KiU'. . ■■,!), p
7 Hi
Hlmio, p
717
t'Imrti ^fruliinflif-ii, ;ilMiii;il.
llntirliirniH 'iri^jiiiK. ;iiijiiiiil
('ilKliiiilinrii frinlii>ii. :iiiiiti;il
I'llHli'iiliiifii ••iiHldld mIuiII ill piiillli'.
I'liKfiiiiliinii irinldlii ^^liiill hciiii iiliiiviv
('jltiliiithdrii rrinldlii. sKiill ficiiij ImIuvv.
(hlKhi/iliiirii ftisliilii. lipHi r |ii»
Mnrriifliiiiiiii(iiiiiiiiiliriiHlr\H, hIiiiII in piiilll«.
MiirrnrliiniiH tiii;iiiiilini/ihin. hKiiII limii :iIm)V«.
Miifriirhiiiiix iiiiiiiiKlirdHhix. hUiiII ('miii lii'low.
Mdvriirhiddn dii'iUKtim/ilriii. lnwi r.jiiw.
HISTORY OV Till'] NORTH AMKRICAN
riNNIPKDS.
'i'lic l'iiiiii|>iMls, <)!• I'iiuilixtliti, niiliracin^ llic Seals iiiid Will-
riis<'s, iuv (■oiiiiiioiily r<'<'(><^iii/«'d by r«M'«Mi(, sysleriiiitiit wiitcrs
as coiislitiilinjf a siilxndcr ol" Mic order hWtv, or Oariiivoioiis
Matiiiiials. Tliey are, in slioit, ti'iie Cortniutni, iiiodiiied lor an
a<|iiatie existence, and have <-onse<|iieiitly l»eeti sometimes
termed '■'•Amphihinus ('antirnra.''^ Theii- whole I'orm is modilied
for life in the water, whieh element is their true liom«-. Ileie
they display extreme activity, lint on lan transi'ormed
into swimmin;;' orj;ans. As tiieir riiime impli(^s, they are Jin-
footed, (lencially speakin;^, the body may be compared to two
cones ioined basally. I'niikethe other niaiine mannnals, the
I'imiipeds arc all well clothed with hair, while several of them
ha\e, underneath the exterior coars<-r hair, a thick, soft, silky
nndcr liM. In contrasting' thetn with the ordinary or terrestrial
mammals, we note that the l)o(ly is only ex<'eptionally laiscd,
and the limbs are contined within the common internment to
beyond the knees ami elbows, and \\\v. hence to only a slight
dejirce sciviceabh^ for terrestrial locomotion.^ The first di^it of
the manns is generally lenj^thened and enlarj^cd, as arc both
the outer dibits of the pes. As compared with other Fcm\
they present, in osteolo;^ical characters, many ob\ioiis poinisof
dilVercnce, especially in relation to the strnctnrc of the sknii.
limbs, and pelvis, and in (h-ntition. The sknil is ,s hct wcfii tli<- piilatiiic iiiid Iron .ntition, the canines are j;i'catly de\eIoi)ed,
soinetiines enoiMKHisly so; the ](»\ver incisois aie never more
than tour in nnndter, an-' sometimes only two; the iippt-r incisors
usually nund>er six, hut sometimes only Ibui', or even two; the
jrrindin^ teeth (premolars and molars) are j;enerally simjile in
structure, and usually didei' from each other merely in resi)ect
to size, or the number of roots by which they are inserted. The
pelvis dilVei's from that of the teirestrial /'V/vr in the sliortness
of the iliac portion and the eversion of its anterioi- border; the
ischiac bones Itaiely meet tbi' a short distance in the male, and
arc usually widely separated in the female, the i»elvic ai'ch thus
remaining' in the latter permanentl,\ open \entrally.
The existing rinnipeds constitute three xcry distinct minor
iiroups or families, ditl'criii;^' ipiitc widely from each other in
importaid characfeis: these arc the Walruses, or Otlolxniithr, the
Eared Seals, or Ohirilthv, and the Marless Seals, or /'hociila: The
tirsi two arc far more nearly allied than arc cither of these with
the third, so that the (hlolxcnidic and (Htnilthr ma\ be together
contrasted with the I'hochUv. Tiie last named is the lowest oi
mo>l .i;cncralizcd yroup. while the others a[)pcar to stand on
nearly the same plane, and about e([ually iemot< from the Pho-
viiJd. The AValruscs arc really little more than thick, clumsy,
obese Ibrms of the Otarian type, Avith the canines «'normously
develo]>cd. and the whole skull i-orri'lat ively modilied. The
lind)-structiire. tin' mode of life, and the whole economy are
essentially the same in the two fiioups, and, aside iromthe cran-
ial modifications ])rescnted by the (hlohdiiuhv, wldeh ar<^ obvi
ously related t(» the development of thecauines as huj;(! tusks,
the Walruses are merely elephantiueOtariids, the absence or
presenee of an external ear beiuo- ju reality a feature of minor
importance.
The characters of the suborder and its three families amy be
more foriually stated as follows : *
"The (.■liiUiicf'.r.'s Iktc j;ivun uio in part tlioso collated !•>• 1 >r. I'licdddrc ».
Gill ill 187:'. (• Ariim;;('iii(Mit of tho rainilifs .>t' llir Mai iiiials.' Sniillisniiiaii
Mi<('('lliiiiciiiis ('(illcct ions. \o. ■.>:!(•. )i|i. ."iCi. (H, li",M. l'\ wlioiii I lir dist inctivc
t'latnicN ol' llirx uiiiii|is well' lirst loriiiiilati'i!. 'I'licv Jiav; , liowcvcr. been
ansncjaiiil.
CIIAIfACTKHS (»l' IMNMITlilA.
Limbs i>iniiil'iii'iii, cir iiinilHii'il intn >.\viiiiiiiiu;;' nruiins, ;iiiil ciirlnsril li> ni
Ix'yitiiil III!' I'lhiiws ,'tii(l ktii't's witliiii I lie I'lHiiiiiiiii hiti'^iiiiiriit.
Dij^ilsiil' till' liiiillM.s tlcircii.siii;; ill Iclljilll illltl si/c iVmii t Ik' I'iinI
to tilt' lil'lli; III' lliiisi' III' tilt' |M's, the lii'Nt, iiiiil lit'tli l;ir;;i'xt iiliil
lolly•('^*l , llir llircc iiiiililli' mil's .slioitiT iiiiil Miliri|ii;il. I't'lvis
with till' ilijir |iiiitiiiii MTV ^limt, ami tin' .iiili'iioi- Ipnriji'r iniiili
cvri'lt'il; isilijii liairl.N iiii'i'l iiiji l(\ a sliiirl >viii|(li\.siN (ncvrr
liucliylosrd) anil in tlir IViiiali' iisiiall.v wiilrly .sr|iaiali'il. Skull
P'lit'rally ;;ii'atly cDiiipirssi'il iiili'iitrliitally : I'arial |Miitii(ii
iiHiiiilly sliiirt anil ratliiT liri>ail, ami llir luain-iaM' aliiiiptly t'X-
pandi'il. liarliryiiial liuiir iiii|M'i'tbi'iitr ami Jiiiiii'il In tlir maxil-
lary, cncl'isi'il wliidly williin tlir nlliit. ralatiiirs usually si'pa-
ratril liya variiil.v, kIIiii nrnuisiilcralili' si/c rriuii llir tViuiials.
Tyiii|uinic lioni's si'parali'il alsoliya varuityrrom tlii'i'XDccipil-
als. Di'iititinii sini|ilr. jii'iH'i'ally unsprriali/ril, till' imdais all
.similar in striii'liirr. Itii'liliiuus dcnliiiiin riiiliim-nlary, iii'vcr
truly luni'tiiHial, and ^riirrally not jit'isisti'iit ltr,\iiiid tin- ln'tal
slaj,'r id' llir animal. I'rrniani'iit im'isius usually !; lu |. snmi-
times .] {('fiKUti'liiint and Miifi'inliiiiiis). nr rviii > ( Diliilm iiiik);
raniiics .; ; nndars- ;, ' , nr '; I 'INNiri",! )rA
A. llimi li';^sr;i>)ialdr id' Idillii I llltird tiU'waid and llsiil in li'l irstri.'ll jdi'o-
miiliiin. Nick lrii;;l liriH'd (isjii riallv in laiuilv ,1 1 i. .^kullwilli
till' masliiid proii's>i's lar^i- and salient ^espeeially in I In- mult.'S),
ami with disiinel .'liispliennid eaiials. Anteiinr I'eei nearly as
larjie as the )iipsteriiu'. tlieir di,i;its rajiidly deeicasiii'^ in lenntli
I'riim tile liisi Id the lillli, wiilniut ilistinet ela.\s, and with a
hroad earl i la;; i noils liorder ex I end in;; heyoiid I he di;;il-. Hind
Icct .sileepi ible of Mical expansion, I he I liiee iiiiddii' di^ils only
willi e laws, and all I he di;;ifs terminat in;;' in lon^i, narrow, i ar-
tilai;iiioiis tlaps. united li.'isally. I'eiiinr wiili the Irocliaiiter
ininiir well developed < !lii;,«SK;nAUA.
I. Witlioiit external ears. I'orm Ihiek and heavv. .\iiterior por-
tion of the skull <>reatly swollen, ffivin;:" support to the enor-
mously developed eaiiiiies, wliieh i'orm li>n>^, protrudiiiff tu.slroee.s.ses, ami tlie surface of the mastoid
processes contiiiuons with the auditory Imlhc Odolnvnldcc.
II. With small oxtonial ears. I'-orm slender and clon'^ated. Ante-
rior portion of the skull not uiin.siially swollen, anc' tlio canines
not highly sj)i'ci;ilized. Incisors of deciduous dentition -}, only
the outer on either side cuttinj^ the j^um ; of permanent denti-
tion 4, the two central pairs of tlus npi)cr with a transverse
irroove. I'ostorhital ]ii'()ce.sses strongly developed. Siirfteo
of the ma.^^i/id jiroeesses not eonfimious with the anditorj''
hnlhe Otarilda:
B. Hind li';;s not ca|>;ildi' of heiuf; tiirneil forward, and not sorvicoablo
for terrestrial locomotion. Neck short. Slcull with the mastoid
•Jr view of the uncertainty respecting the proper notation of the grind-
ing teeth, they will in the jiresent work be designated simply as molars,
"with noiiltemiit at distingnishiiig "premolars" from "molars.''
CIIAI.'ACTKKS or I'INNII'KDIA.
IH'iM'i'.s.si's swiillrii, lull IK 1 1 smIu'IiI, ,\\\i\ w it liiir' ilistiiicl ali,H|ilu'-
iioiii 1','iiiiils. Anli'iior iimli.s siiiiiiji'i' ijiim t hi' iioHtriltii', Ihr first.
ilit;it litllc, il'iiii\, liiiij;cr tliiiii tlir iirxl Niicccciiiii;; diics, all
iU'int'il with Nti'<)ii<; rhiw.s, \\ hicli jiii' tri'iiiiiiiil. liiiiil iVi'i I'a-
]itllil(' of iiioiii I'jitr <'\|iMiiMi()ii, shiil'l : (limits (iiHiiaJl.v) all aiiiinl
witii Niroii;;- claws, anil witJKMit icrmlnal laitilayiimiis llaps.
I'riiiiii with nil tiarr i>t' the ti'iM'haiilt'r niiimr. . . Iii:i'i iiiijAPA.*
III. Without cxtci'iial cat's, rostoi'liital ju'ori'SHcs watititi;;, m' very
small. Itn'isiits vaiialiji' (';, :|. i>i'}). lirridnoiis ili'titil ion not
|M'i'sislrnl licvotiil I'd'lal life I'limidii.
The riiiniptulH prcs'iit ii lii;;li drorcc of cci'dtriil ilcNclop-
ment, iiiul arc easily «loiiH'sti<'al('(l under liiv(»ial)le e<»iirs to
certain favorite hreedino' oromids, and diirino' \]w .season of re
])rodnction !ea\<' the water, and pass a coiisideraltle period upon
land. The ICarle.ss Seals, on the other hand, with the exception
of the Sea I'Mcphants, do not so nniforiiily resort to |»articnlar
lireedin;; oronnds on land, and leave the water (»idy for very
short intervals. They nsiially hrinj;' forth their yonn;; on the
ice, most (»f the sp"cies l»«'inj;' conlined t]>ears to
be strictly tropical, and very few of them rauj^e into troi>ical
waters. As a i^ronp, the ['innipeds ari^ distinctively character-
istic- (tf the Arctic, Antarctic, and Temperate ])ortiortions of the Northern iremisidiere. The
Otariidw and riiocida', on tlie other hand, are abumlantly
represented on both sides of the equator, as will be noticed
more in detail later.
*l''or the .suKj;osti()ii
to my friciid l>r. Kllio.
ho terms Grcsmgrada and UcpUgrada I am indiOiteil
^011f.S.
FAMILY ODOHyl^.NlDiK.
Wl»!r:j;;j:;.
'• 'J'rkhnidii, (iJt.w, l.niiiliMi Mfd. K'riius., If^-JI, ItiKl" (I'iirnil.v ). .lj(«(i Gray,
Jri(7»((7/((/ri, (iit.w, Ann. ol' I'liilos.. l^•,>^), :tl(i; Ann. and .M.ij;. Niil. Hint., ^d
H (laniily).
Tiivhiciiia, (iu.VY, London's .Mai;. Nat. lli.st., i, 1K!7, r>:{H ; "Zotil. Krthiis and
Tt'n<)i',;r' (.sulil'aniily). Jn pait only, oi('xt. ii, 1850, 'ill; Cat. Srals and
WIialcK. H(i(i, :i:5 (snlifaniily). In jiart only _- Trichicina (jiny, 1837.
" Triihtrliiild stn Cuiiipodoiiliti, IfiiooKKs, Cat. Anal, and Zoiil. Muh. 18a8, 37."
Trirhnlioiilni, (iiKlii:!,. Fanna . '27. (ill, 7(1 ( -.= " TtiihivhUhv MrooUcs, (irrvais").— Ai.i.r.x, Hnll.
,Mns. Comp. Zoiil.. ii. 1^70, *,M.
J!iiniiiiili33; Diet. ."^iLNal., lix, l^'JO,
Iti.'i (family').
(|i;m;i;ai. ohsi'-uvations.
.Viiioiij; llic disliiictiv*' li'adin'.s ol" (lie (hlolnvnUkv iiio the
('iu»iiii()ii.s(l('v»'l(»|)iiM'iit of I he iij)i)cr('imiii('s, aiul tlie <'on.s»'quont
gic'iit rnliirjicinciit of tlu' aiilcrior ])(>rti<)n of tlic skull for their
reception jiihI sii])j)oit, tiie early losis of all the iiu'isors excej)t
the outer i)air of the upper jaw, the; caducous character of the
posterior molars, auar vertebra' iciiiaiiiiny' of proj)ortioualely
the same lenj;th. \n consecpu'uce of their obesity, the ribs and
tile proximal s(>o|iients <»f lhelind)s are lonjicr in the Walni.ses ''
than ill tile iOared Seals, wliiU' tlie distal sej^ineiits (d the li.Mbs
are iclatively sliortei. The scapula is long and narrow, instead
of short and broad, a.s in the Otariida', iwid it.s ciest is ithiced
(i FAMILY tiDoll/KNIlM;.
irioif iiiilnidilv. Acnii'diii^lN , ill rcspccl to L; iiiid liiiilis in the one r(»litrastC(I
with ;^rr;il tliicKiicss ol' IxMly. iiiid distall.v ii dispi »])()rti()iiiito
K'diitiidii (>r I lie cNti'diiitics in llic (»tlicr. TIic iiKid stiikin;;"
tlirrticiiccs^ liowt'Vci', exist ill llic ciaiiial riniriiclcrs. roiiitinj;'
Ikhii the unreal devt'lopincnlor tln' iippci- caiiiiics in ihr W ;il-
I'liscs, ;iiid tlic ('oiis('(|ii('iil iiiodilit'iilions ol' (lie facial portion of
tlic skull. Ill llic (}hir!!(l(i\ tin- j;<'iit'ial contoiir of tlic skull is
stroii,L;l,\ I isint'; in llic (hlolKiiiida, \\ is iiiii(|iic. owiii;; toils
;iicat expansion antcri(»il,v. In respect toolliereianial features,
the WalinsesdilVeilioin the Eaved Se:ils in liavinjiiio post-ovb-
ital ]»i'oee.sHos, and in the niastoiil [troeesses lieinj;' not . separated
from the auditory hiilhe. Tiie teeth are all sin^ile rooted, and
iiave in the periiianent dentition no distinct crowns.
On coiiipariiiji' the Odolxviiidiv with the I'lioriihv, the ditVer-
enee.s in j^oneral .stnieture are Ibiuul to be far ••reater than ob-
tain between the Wabnses and I'iared Seals, espe<'iall,v in rej^ard
to the hind exti'eiiiities; these in the J'liocidd- beinji directed
ba<'kward. and useless as organs of teirestiial Ioconioti(»n.
Hence, in s(. .ar as tL<' (hlohwnUUv and ()t(irii(l(v n'^vvv in linib-
aiid skiiU-strneture, they both similarly depart from the Phocine
tyi»e. As already indicated in the synopsis of the sidxtrdei' Z'/«-
nijx'ilid, the I'Ikh'kUv dill'er far more from either the OdolxvnUlw
and OtariUUv than do these latter from each other. This ditier-
onee is esi»ecially emphasized in the' skull; for whih; tlie Odo-
h(vni(l(v and Otariiilw aji'ree in all important cranial characters,
aside from the special features correlatf'd with tlu; immense
enlar;;<'ment (»f the u])per canines in the former, they widely
differ from the riiocida: This is es])ecially seen in the absence
in the latter of an alis]dienoid canal, in the greatly swollen andi-
toiy bulla', the position of the carotid foramen, and the non-
salient character of the mastoid ]m>cesses.
The few points in which the Walruses dilfer in myology from
other Pimiipeds, Dr. 3Iurie statv's to be ''the presence of a co-
I'aco-brachialis, a llexor brevi.s manus, a pronator quadratus, an
op])onens jtoUieis, and a palmaris brevis," in the ])Ossession of
Avhich it differs both from Otarid and J'Iiock, but tliat in other
resj)ects they " muscularlyju'esent general a jireement."' ''('om-
]>ared with the Seals [I'liora .^] there are two extra peionei and
a llexor Imca ishallucis." "•Though deficient in concha, the auri-
cular muscles aic remarkably large."*
I'roi-. Zoai. Soc. 1,(111(1.. 1*370, 1). .')4.').
OMM'.l.'AI. (tllSl-.KVATlONS.
i
••( 'nlisi<|»'rill,n tin' Nt'fV (litTcifllt ;lttitll liltic (l(!viii(iii of tlii- iin sclcs of tlir liiiid Icj;', aixl in this rcsiioct
himmIc iVdiii tlic Sell, vet Imt slijilitly."*
Ill n's]icct to fill' iMisitjdii iuiil cliiiriictcr of till' visc«!ra, a yen-
fill! a;ii'tM'iiifiit lias Itcccii iiutcd with lliosc of tlic other IMiiiii-
pcds, and tlicv i»icsciit iiolliin;; that calls lor special notice ill
tlic present connection. As Dr. ^Inrie lias stated, tlu-re is littlo
api>recial»le dilferciice exiiihited tlironj^lioiit the IMnnipeds in
the construction of the alinitMitary canal. " It is .siiiij)ly that of
a Carnivore, with. lio\v«'ver, a moderate-sized ca'ciini. The great
ylaiidnlar snperticics and correlated large lyini>hatie.s point to
means of sjicedy and freipient digestion; and in the Walrns
these apparatus are extraordinarily develoi)ed.''t
In a<*cor(lance with the characters already given (p. .'i), if any
snbdivision of the Pinnipeds into groups of higher rank than
families is to be made, it seems evident that the OdohaunJcc
and Otarlidn' are to be collectively contrasted with the PhocUlw;
in other words, that to nnite the Otarildw and Phocidw as a
group of co-ordinate rank with the Odohwnuhc is to lose sight
of the Avide ditterences that sei)arate the two first-named fami-
lies, as well as of the many important features shared in com-
mon by the (hlolxviiidw and Otarndtv, by Avhieh both are trench-
antly se])arated from the Phocidw.
Although the Walruses are now \ery generally recogniz«'d as
constituting a natural faunly of the Pinnipeds, ranking co-ordi-
nately with the I'^ared Seals on the one hand and with tho Earless
Seals on the other, the affinities of few groui)s have been more
diversely intcrju-eted. As early as the thirteenth c«'ntnry. the
author of the"kSi)eeidnm Regale'', — one of the earliest works re-
lating to natural history, in which the Walrns is mentioned, —
stated distinctly that the Walrus Avas an animal closely related
to the .Seals: and wetin^7'i. p. 4()I.
8 FAMILY ODOH/EXlDyi:.
llicm with forms with whicli they hiul no lohitionshii*. Tn the
ii'fuiR'v of scioncc, nothhig was perhaps more natural tliau that
an'inals sliouhl be chissitiecl in accordance Avith their mo(h' of
]ife, their liabitat, or their external form, and w<' are heiu-c not
surprised to find ihat llondelet, (lesner, Ahh'ovaiuhis, Joustou,
and other pre-Linna?an writers, arranged tlie Pinnipeds, as well
as the Sirenians and Cetaceans, with the iishes. or tiiat other
early Avriters should term all four-footed creatures "Qiiadin
peds,*' and divide them into "Land Quadrupeds" and "Quadru-
peds of tlu' Sea." While all marine animals were by some early
writers classified as ''fishes,'"* the Pinnipeds were much sooner
«lisassociated from the true fishes than wer«» the Cetaceans and
Sirenians, the mannnalian aflinities of which Avere not at first
lecognized by even the great Linne himself, who, as late as the
tenth edition of his "Systema datura'" fl75S). still left them
in the class •• J'isrcs."
In view of the several exceUeut descri])tions and very credit-
,d)le figures ol' the Atlantic Walrus that ap])earcd as early as
the sixteentli and seventeenth centuries (a detailed account of
which will be given later), it is sur])rising iii;'t the early sys-
t^ematic writcis sliould display such romi>lete ignorance of souu-
of the most olnious external characters of this animal, as was
Jiotably the case with Linne, Klein. Brissou, ICrxleben, and
Gmelin, who strangely associated th<' AVidriis and the ]\binatee
as niend)er> of the same (/eiiKs, ami groujied them with such
diverse creatures as Sloths and Elci)hants. Linne, it is true, in
the earlier editions of the '' Systema Xatura>,'' placed the Wal-
rus with the Seals in the genus Phoro, in his order Fem\ — a near
hit at their true affinities. Later, however, foil wing probably
Klein and Brisson, he fell into the grave error of r.'nioviiig tluMii
to nearly the most unnatural association possible. In this con-
nection, it may prove not uninteresting to sketch, in brief out-
line, the strange history of the classification of this singula)"
grou]) of fin-footed Carnivores.
.\.s already stated, l^inne's first allocation of the grou]» was
the natural one. Brisson,t in IToO, led in the long role of error
by forming his third "'order" of mammals of the lOIcphant, the
*Mosf modfi'ii l!ii)<'iiii!L{c8 still ivtiiin nlics of (liis iincii'iii cnstoiii, as
«'\iiict'ii, for ('\;iiiiiilf. ill siicli I'hi^'ii.sli words ;is s7((7/-//.s//, crdii-jisli. irlidh-
Jlslitri/. >ic(il-ji>flu'rji, etc.. wliilc hrdljifili (Swcilisli), ividrixclt (Djiiiisli), irallJlKk
(GiTiiiiiii), ftc.. iirc coimiioii Ncrnariiljir iiiiini's jiiipiii'd to Cclaccuns.
tRi'^nr Aiiiiiial. I7.")(i, p. t-'.
(!i;m:i{Ai. ousKiaATioNs.
1)
NValius, and llic .Manatee, tlie two h.s\ iiaiiKMl eoiistitiitiiiu Iiis
•• j;euus Odohnni.s." This was a iiiaiki'd i<'ti(K'es.si(»ii IVoin even
the system of Ivleiii,* of a lew years' eailier date, w1h» l)ioiitilit
to<;etlier as one family tlic Seals. ( )ttei's, Meaver, NN'alrns, and
Manatee. Linne, in 17(»<1.+ not <»niy removroui)s than ;;('nera. plaeed the Waliuses and Seals tojiether im-
uiediately after the ('avidvores. Schrel)ei',§ at about th«? same
date (1777), ad»»j>t<'d a siudlar ehi.s.sitication, t!u' Walrus .staud-
iuy next aftei' the Hlejthant fiid lU'.'cedinfi' the Seals. Scliro-
l)Oi*s ;;enus 'I'ricliirliKs contained also the Dufionj;' and tlio
Manatee. ris.son to
(rinelin. standiu;;' next to the l']le[»hant, and as.sociated yeueric-
all\ with the Siiciuans. r>lunu'nl)acli.'| from 17HStillas late
even as ISi',"). still arranged the Walrus and the Sireniaus in
the ucnus 'rriclii'clnis. in other res|)('cts. the Walius appears
with new associal"s. the ji^cnus 1', IciiccIiks Iteinji' uinted with
Oniitliiirln/iirlnis to form a "faniily"(I) of his "(»rder" I'abiintd.
The order I'dliiiiiia. as the name implies, was composed of the
web-footed manunals, and divided int(> three •• families," namely,
"A. (Hires" (eonsistinjiof the yenus Castor); -']>. Fcrw"' {I'lioco
and Liitra); and "C. Jlrnfii" {OrnHhorlii/iicJiii.s and Trichechm).
Tills is essentially also the ariauyvmeid j»ro]«)S(;d by Klein in
1751.
The lirst step toward disinendteriiij^- ;.! uunatural conj^lo-
Jiieratiou known previously under the names Tiicheclitis and
Odoheniis was made by Het/ius** in 1704, wlu> divided the
gcuu-s Tricluchiis of former autluu-s into throe yenera, namely.
ManutKN, for the Manatee; JIifdioiiKtlis. for Steller's Sea-Cow
{= lihi/tiiiH Illij-er, ISll); an. lit.
tSyst. \li"^. Aiiiiii.. 1T77. p. .".!»:!.
vNSiiiip'di.. ii. [I77t'..'l. 1.. v'lio.
II Sy.';!. \!i(.,i, .-.!).
^Hiindlt. <1. Xiiliiriifscli.. I7"<-, p. II-.', and lalcr iililimis.
"Koiigl. NrinivK. Acad. ii,\,i llaiidliii;;,. \\. I7'.M, iip. •J-r)-;!0(l.
10
lA.Mii.v (ii»(»r..i;Mii.K.
I [
\\\ii Iiotli llif \\';iliii> iiikI llic 1 )ii,niiii,u. NVliilc lliis \v;is in
tilt' iiiiiiii a iiiosi iiiipdi taiil ami iiro.mc -^'x e iiiiio\ati»iii. Kc;
zius sociiis t(i \i;\\{' laltdi'i'd, liUt- >('\('ral still carlici- wiitns.
iiiidcr tlic iiiipicssioM lliaf tlic Walrus, like ilic iMi.^oiiu. /""/ /"<
hind fat. ( )/.('i('tsK()\ sk_\ . * alioiit a year later. aiil\'
iuiKiiaiit nf li'ct/ius's [>a|M'|-. also placfd. as cuiioiisly liapiiriird.
ilic Walrus and the hii^diiu to.iActliff in tlic ufims Triilucliits,
iMM-ausc lie siii)ji(>scd the Dw^owj: lii((l hliid Jrct.Wki' tlif Walrus I
These euiious aiit it lift ieal mistakes iiitlieate how little was
known Ity systematic wiiteis ai»(>ut tlic stiiictuic ol' these ani-
mals as late as the close of the last .s'/M«r«.v"; '•!'. TricluTiis i and the Manatee, to which he
gave the generic names lespeetively of Phiti/sfonnis { = ll(ili<-uri .
niiger, 1811) and OxystoiiiKs { = M((nafii,s. Ifetziiis. 17!»4). lea\ in;.;
only the Walrus in TriclurhKs. The genus TrirlKTlins. however,
as first instituted by Artedi (IT.'W) and Liniic (ITriS). as will lie
shown later, did not rehite in any way to the Walrus, being
applied exelnsively to the ^Fanatee. It was not till 17«>(i that
the term was lirst made to cover both the then known Sirenians
and the "Walrus, althongh the iMubroilment of the two groups
began with Urisson, ten years eailier.
The l*innipeds and Sirenians, collectively considered, were
lirst separated as disthict groups by illiger§ inl.Sll, who raised
them to the rank <»f orders, they forming respectively his orders
Piiiiiipcdut and \(it(iiitii».. xiii. KDti. jij). :{:i-:i7ri.
t'l'illll. l^I.'IM.Mlt., \>. IT','.
t Diifi Natioiiiii-.Ntiiscmii (Irr Xatiirj^i'scjiiciih', li. IS(i;!, [vp. iMl-lt.'jS.
V^ riixhiiiiuisSvstciiiatis.Maiimialiuiint Avium. l?'lt,iip. Uri. V.VJ; Aliliamll
tier Akacl. Wi'^scnscli. /ii Itcrliii. ISOl-l-'JI. (IKJ.'.V i>lt. :!H !:">!•. imsnim.
CENEUAL OIJSEHVATIONS.
11
(•(iui\ iilfiil ill extent with the order Pinniimlla. The i»i'0[niety
of the eliaiiyes introducetl by Illigcr was not speedily leeognizcd
by con temporary writers; (Javier, and many subsequent syste-
matists tor hall' a eentury, ])lacin,L: tlie Pinnipeds amonn' the
(jtinirora and tiie ttirenians amon.n' the CtUicca, with the rank
respectively ol" I'anulics, the lanuly P//oc/(/ft' einbrachiy all the
Pinnipeds. Dr. .1. 1'. Gray, in ISL'l,* and af:;ain in 182"),! widely
separated the Walruses from the Seals as a laniily, Trichcchida',
which he most stranjicly placed (together with the Sirenians) in
the order ('t lier I 'inniiieds. Wagler,** in 18.'{0, made the Walruses merely
a genus of his order Crsi. Nilssou,tt in 1837, divided the Pin-
nipeds into two sections, the second of which embracd not only
TriclurliUN, but also JlalichaiKs^ ('!iNto2>hora, aiul OtarUt. Tur-
ner, |f in 1848, from a study of tiie skulls, separated the Piiiiii-
l)eds into three natural groups, considered by him to hold the
rank of stdifamilies, namely: Arctoccphalina. embra<'ing Otdria
and An'toccphuluN ; Trichcciua, consisting (»f the genus '• Trivht-
c«6'"; and ritovina, embracing all the other Seals. Gill, §§ in 1800^
* " London Mi'd. JJcpos., 1821. p. :?0'2," upiul (Jniy.
t Annals of riiilosopliy. 'Jd sit., vol. x. ia.'.'>, p. niO.
t Eoiidoii's Miifj. Nat. Hist., vol. i. p. '>"':!.
v> I'ani. Rt'g. Anini.. \). .M.
!| Diet. Sci. Nat., t. lix, p. 307.
*'. "t'al. of Ills Anatoni. and Zoiil. Mns.. ]i. ;{(!." aiiud (tint.
**\aiiiil. S.\(st. Ainpli., p. '27.
ttVrtcnsk. Akiid. llandl.. 18117, !i:{i'>; WiP};mann's Aiob. f. Natnrg., 1811,
jt. :i(lO(transl.)
ttl'mc. Zoiil. Soc. I.oik:,, 1848, pp. 85, 88.
VH^ I'ldc. I".ssc\ Ins|if!ii('. \ol. V, 11. 7.
rJ (iKMI.'A oi rill. lA.Mll.V.
\\;is tin- IH\( :iiillior \\Im» r(r();^iii/('(l I Ik- \\ iilriisc.s iis loiiiiiii;; a
disliiict lUiiiilN , w iiicli lie h riiicil lioxnmriihi . lit I liis step, lie \\;i,s
iiiiiiir(li;itrl\ lulluwcd l>,\ ro ordiimlc in niiik with his
'■ /'Af/r«/Wr«," roiisistin;; of tlic I'Iiiu'kUv :iiiiI i>l((t'iiilties of hotli tiie I'iiinipeds aiil., :',. ('(Plllp. /cic'il., \ip|. ji. |). '.'1.
I I'.'IIIIIM . •>'.•.
||\:iii I'lrmijiii, Ami. Mm-, illli^l. NmI . ili iii'lt;ii|iic. i, I-T7, |p. Till.
J.
(iKNKIUI, OUSKUVATIONS.
caiiiiH's were roiiiiil :it . \ii\ ns, l»iit \';iii Knu'dcii i.s .s|i(»ii;;l,\ (»!'
the ii|iiiiioii tliiit IIh' It'clli (l(',s( rilx-d l»\ l{ii.\ liiiiikcslcr,* Irmn
the Ii'rd < 'ni;; <»(" I'lii^iliiiid, ill IHJJ."*^ iiiid iiiiiiwd Triclircliothm li".r
//■///, ;ir<' I hose of his ,\li((lliiit ramus. The other
hones hi'lieved to represent it are nine \erlehra', |»art of a pel
\ is, a hinnerus, a femur, several metatarsal, meta<-ai'pal, and
phalangeal hones, etc., ainl part of a tusk. Says Van Meneden:
'• I lie lnanchede max ilia ire est tout ce < pie in ills possedoiis de la
tele. liCS dents maiiipieiit, liiais le h(»id esl asse/ coiiijdel,
pour tpi'oii piiisse hieii iii;.;er de leiirs caracleies par les ah i-oles.
Nous poiiMins, (111 reste, fori hieii aiissi apprt'cier la forme dt;
cet OS, dislinj^iier sa symphyse et sa hrievcte.
•' l/os est luise a son c\tn''iiiile anti'-iieiire, la syin|(liyse est,
fori cniirteet Tos ii'a pas plus dV-paisseiir siir la, li;;ne mediaiie
que siir le coh'-. Lesalveolessoiil compaiat i\ emeiit (hit ;;randes:
h's trois dernieres sont. a pen ]>re semlilahles, ranti'-iieiire est, la
plus petite. ( "est Till virse dans Ic Morse, liaca nine (leva if «'tre
tort ;;raiide. II iTs a ipi'iiiie settle alvi'ole |iiiiir ane dent iiiei
si\e.
'* \a' i(ii]>s (111 maxillaire esl remai<|iiahle jioiir sa coiirhiire.
Toui(!la ]iarlie jiost(''rieiire, (pii const Hue, la hranrhcdn maxillairi^
maiKpie. On xoil siir la face e\tern(; trois Irons mentonniers.
-> Mil coiiiparanl ce masillaire a celiii dii M(M'se vivaiit, (Ui Noif
((Ue la SMiiphyse est loiite ditferelile, (jii'il e\ist(^ line ;;ran(le
ahcole pour la dent caniiie el (les traces d'line petite al\(''oUi
pour line iiicisi\c ipii restait prohahleineiit (•a(!li(''(! sous les j;(,'U-
civcs. Dans le. iMors(^ vivaiit, il try a pas de place jioiir line
canine |yraiide| an maxillaire, infc'iriciir."
• Sec hiVDlid. p. arts of AUuthcfiuui.
Van Hencden's descriptions and liyuri's of the lower Jaw
Iraguu'ut indicate features widely ditferent from those of the
coiTespondiny part in the Walrus, especially in the shortness ol
the symphysis and in the curvature of the i)art represented, but
above all in the nuMd)er, rcLitiv*' size, and foriu of the alveoli,
and particularly in the larjjic size of that of the canine, which
must have been almost as hiyhly specialized as in the Sea Lions.
That the tusks referred to it by Van IJencden (those described
by Lankester especially, as well as the frajiuient he himself fig-
ures) beloiiji' here, there seems to be at leasi ro(»m for reason-
able doubt.* The dilfcrences presented by the Jaw fragment
of Trichechotlou as compannl with the <'orrcs])onding part ol
Aladheriitm arc c\cn still Juorc marked.
The more obvious characters distincti\c tif the three genera
oi \\vii OiUthanidn . a^aX picscnt Iciiowii, may Ite luietly indicated
as follows :
Slliiiijiniy (if //(( ivi'ly to the
Walrus in a generic sense). — IJr.issON, Kegnc Anim.. 17.">*i, 43 (used
strictly in a generic sense, hut emhracing ■•!. t.a \'aelie marine —
OiloheHHn" = \\;i\v\\s: "•,'. Le Lamaulinc — Mdinilii"." The characters
given apply almost exclusively to the Walrus).
*Van Beneden himself says: --M, U'ay I.ankaster avait vu en Angleterre
diffdrcutes grandes dents, i)rovenaul duii Inu'hiii h'ay I.ankaster" as a syno-
nym of Trkliiihodoii liOiiiitncliii. dc.scrilpcil l.y himself much later! In view
of the iiueertaiuties of (he ease, it is to he regretted that Ik; did not ])ropo5e
il new generic as well as specific name lor his 'iv'uhivhiuhtii hnnihirlii.
♦ With refereiKe only Jo the lower Jaw, tlw only ktiown )iart, in case of
the extinct tyjies, readily susceptilile ol' eomiiaiison.
SYNONYMY AND N< )Mi:NCI,ATri{K.
1;
;>
Diluhiiiiiis. MAi.M(ii;i;N,<"»rv(rs. K. N'.t. AU;iil. Forli. IHti:i, (iHlUj. i;!ii.
]:,>s,iiitni.-<. Ki.i:i\,(^ii;nl. Dis-i. I{icv. Hist. Niil., IT.M, M\'M (;iii]>lir(l in a ;;cii-
fiic sense exeiiisivcly to llie \Yiilnis), — "S ui'oi.i, IntiKil. llisl.
Nat., 1777. —." — (iii.i, ("ex Stdpoli "), I'loe. Kssex Inst., v. If^titl. 7.
I'lioni. I.inm';, ."^vst. Nat.,i. 17.> (in pail uniy).
Trichi 'litis, I.iNNi';, .^yst. Nat., 170li, 4i>(iniiarl (ini\ ; ni(| of Linne, 17riH, nm
Artedi, I7rw ; liaso
of the )iresent eentiiry.
(idiihiimlhiriinii. (in.vTiOLlCT, Ihill. Soe. (ieol. (h> I'ranee. •," ser.. xv, 18")?.
(I'J4 ( = ■■ Tiirliirlnin ri>siiiiirnx'' anet. — fonnded on a supposed fossilV
Oilmitobinins, Sfxi»i;\ ai.i„ Ofvers. K. Vet. Akad. Forh., 1-.V.», 441.
.' I'll ell I'll oilim, l..v\Ki;.srKU, (Quarter. .loiuu. Cicol. .'^oe. Loud., xxi, Ifftio, 'J-iti,
pi. X. xi (hased on I'ossil tusks from tlie K'ed Crajj-, i'.n^iland).
Tlic iiiiiiH- 'rriclin-liiis, lur so Imiii' :i tiiiu- in m ficnil use loi
tlif \V;iIiiiscs. ]»>ov<'.s iMd, iis loiij; ;|o(> sliuwii l>v \Vi»'o||i;uiii,* von
liiicr. .Miillcr. Stiiiiiiiiis. ;m«l liitcf lt\ idlu r "viiti'is. to Itt'loiij^' at
.ill to llicsc iiiiiiimls. lull to tilt' .M;iiiiit<'('. The iiiiiiic Trli'ltirlnix
oi'i;.;iii;itiMl with .\rtt'tli in IT^IS. insi posilniiuoiis worl^ t clitcd
ii\ li:siiiii iiniM'iinc. Fistiihi . . . . " 'I'lic cita
lions nntlcr TriilurliKs fiiilnacf no allusion to the W'aliiis. luii
i«'Iatt' wholly to Siicnians, of to tln' .Maiiatcf. a.s the latr«'i' was
tlnii Uiiown.T Artt'. llC.
tlehthyolo-ria, 1738, pars i, ]). 74; pars iii, \>. 71>: pars i\, ]>. Itlil. In Arie-
di's work tile name is twice written Trkhrcliiis ami twii'c Tliricliirlimt, On p.
71 iif pars i. wliere it lirst occurs, its deri\alioii is given, namely : •• Trhln-
r'nin ;i \^iti^ rriiiii .J'' i \;fl<)c piicis ipda solus inter pisces fere hirsulus sit."
: The references in a eenerid way ajipcarto include .'ill the ."sircnians then
known.
v^']. g.. "Deniiuin duo utrini|iie I'mimiil. louuit udine :>pii liani;e crassii u
poUicis."
m
u;
'llll. (il.M S Mixili.lAI S.
<<(iii\iil(iil lu I he T/Vr ul Linin- (S\.s|. N;il ., •il. \, I 7.">Sj. I.iiiiif,
ill I7.")H. liisl iiitindiirt-il AltcdiVs ;;fiiiis Tritli)flnis, ;il wliicli
lime Ik- plmc*! in il uiilvtiic Miiiuitcf, I Mi;;()iiu, iiikI Slflh-i's
Sea ('(»\\, I<-:i\ in;; t III' Waliiiscs still in /'linen. Mi,, (lia^iimsis
of till' ;;ciilis* ciiiliiactMl mtiic ul' the ilisi ilirt i\ c rliaiarti) > nj
llic Waliiis. Ill I7«I«; (iLMli «■l\
aMcrilii'd to tlic Walrus in loiiiH-r fditifuis, when I In- Walinst ;
wi'ii- plan-il iiniii'i' I'linm. Ilnicc, to uliatcvei tli<- ;;fii(iic
naiiif I'lichvclms iiia\ he ic|'rral>l<', it r»itaiiil,\ is not prrtimnt
to tlic Walrus. 'I'liis liciii;^ scltltMl, tin- ipn-stion arises. What
jfi'iH'i'ic iiaiiic is oCiiiHpiestioiiaMc appliraiiilit.N to the Walruses .'
Here the real iliMienlt.N in the tase he^^ins, tor aiilhors who
ailiiiil the inapplii-ahilitN ot' 'I'rifli clnis to this ;,iioiip are not
agreed as to what shall lie snitst it iite<| lor it. .Seandinav ian
Avriters, as .Maliii,i;ren (Isiilj and lalljehorj,^ (|s7l;, ;iiid I'eitrs
( JSOI j aiiionj;(Ierman ant horil ies. Iia\ e lor some \ ears eiM|iIo,\ ed
(hlolxniiis, a name apparent l\ ori;:inat in;^ with l-inne (as Oiluht
iiiis) in 17."!.'», and adopl<'d in a j^eiieiie sense li\ Itiisson in I7.'i(i.
A iiKdlilied I'orm ol" it ((hhnihihitniis) w as als(» emplox ed li\ Sun
de\aliin I.S,"i't. (lill,in isiKi, and other recent Aineiiean wrileis.
have l»roii;ih( into some pidiniiieiiee t he name 1,'iisiiiiirii.s. \\\si
ii.sed in a ;4eneiie isense li\ Klein in I7."»|, li\ Seo|M»l: in 1777, h.N
l*allas| in I.S.'JI, and In liamont i in Isilj ; while the ;:;reat mass
of Mn^iiish and ('onlineiital w riti-rs still elin;.; to '/'riclicilnis.
'Ilie ;;eiiera (hl(tl)iiH)lli( rlinii and 'I'riclicrliixluH, based on fossil
ri^iiiaiiis of file Walrus, iiavealso heeii reeeiillv introduced into
1 lie literal lire of the sill ijecl, the former li\ (Iratiolet in ls."»,s. and
the. lafter li.\ l,aiilvesler in ISIm; Iml these (espe tlielirst)
*"l)riillil:iiii. nmlaii-s r\ o-^c
lii;ji;iiM> III riii(|iii' iiilriiiiN (Inn. l.iiliJM ^iiiiiiil.'i. I'cilis ikoIci inns ((lailiiniil i
ill |iiiiii:iiii." \(/«/. .V((/., 1(1. \. i, 17.".'', p. lit.
t 'I'lic. sec I III (I (liM^iinsis dl' I'rii llll liiiK is, in In II, :is IfilldU > : •• ilcnii s |ii lino
rcH niilli nljiiii|ni . I.aniiiiii .sn|ii rinics snlil.irii. .Mnlaics i \ ns.sc iii^nsn
lilliminc : iiili rins duo. I.iiliia ^cniiiiala. i'lilis |in.sl( rinics (:niii|i('ii( .s cn-
iidiiiiali ill piiiiiaiii.'" — Ai/f/. .\V(^, ed. sii. ITiK'i. i, |i. If".
;y,(«il. l{:iNS()-.\hial., \nl. i. •Jfi'.l.
ii\S('ilsn!i.s willi llic Scjl linrscs. |i|i. Id. IIm.
■«!
'nil; si'KciKs OF tiik gknijm.
17
ii]t|M-iii' to Im* ictnTiil)!*' lo the «-\istiii;; \V:ili'iis<-s, and of ciiiiisc,
IXTOIIM' lllcr«>l,V HVIIOII.VIIIM of (Ml'licl- IIIIIIICS. ( 'oll.SC(|IM'llt l,V t Im)
clKticc, rvidciilly lies Ih'Iwccii (hli)h(viiits:\\\\\ h'nsitianis. ihhha:-
viis liiis sixlrcii \<'iirs' piiorilv over liosiinttiis, if we ^o Itjirk to
till' riii'lifst, iiitrodiictioii of IIk-sc iiaiii<-s into svstfniiitic nonicn-
flii.lMi'f.* It is tin*- tliiil liiisiiiiirii.s \\i\s{\u' *':ii\U's\ liiilin nanio
a|i|)li<-i'(;lian;;cal)l,v witit
Mots and Mnrnii.s l»y Olans Ma^^nns, (h'simt, Ih-ilMTstain, anil
{){\\v\n^ hill onlji ill (I rcniitvnlar sciisr. Altlion;;li used l»\ Kh^in
systrniatiraliy in ITr*!, , as did also i>riss(»n in I7ri(j. 'I'lie. wliolo
«pU'stion tnrns on what siiall Im- considerrd as the proper start-
in;4 |ioint lor ;;enerii- nonienejatnre, ahoni which opinion is still
dixidrd. ir the early ;^i-iieric names of Artedi, Klein, llrissori,
ami liinne (prior to J7r)(',cics
Oiliiliiiiniit, Liiiiic, "l)i;;ili ;iiil., |Mist. .'>, )i;iliili|M'S. \iiis>i Mormin, Di'iitOS
iiilciiiicilii Hii|)rriiiiiH Iminissiitii.''— .s'//«/. AVt/., \7'.i7) (cd. fir), f/.t. — Jiosmarua,
Kli ill, (^iiii.l. I)i:.ii. r.ri\. Ilisl. Nat., 17r>l, 40, 1)'^'.
till :icr(inl;iii( T willi (ii.sldiii ill Kiiiiiliir riisi'H, llin iiiiiiD'. i>l' I In-, riiiiiily 1)0-
coiiii's thlohii iiiild', — Ml illur HiiHiiiariiUv, nor Trlchcchiilw lining tciiiiltio.
Misc. Tub. No. IL' li
18
Tin; OKM'S (>IK)H/KMS.
I ,.1
with those ol' th(! (iiilph ol' St. Lmiirmr. The tusks (if those
of the I'l'o/eii iSea arc iiiuch loii^^ci'. uhmc sh'ii(h'r. iiikI liavc a.
twist and inward curvatiiiv. '* iShaw, a few years later, tlioii^iht
that tlie W'ahiis (h'seiihed ami liuiiied in the aeeonnt of Cap-
tain Cooli's last \(»ya;;(', Ihonnh perhaps n(»l speeilieallv distinct.
I'loni lh(»se of tlie Arctie shores of llnrojie, slmuld he regarded
as l)eh)nj;in;;' to a dilferent \aiiet,\.t lie appears, li(»wever. to
Lave based his opinion wholly on li^^uresof the animals, and par-
tienlarly on those j^iv-en l»y ("ook and .lonston (the latter a copy
of lierrard's, at second-hand fi'oia I)e l.ai't). Illiucr, in isll,
formally reco;inized two s]»ecies in his '• rehcrltliek der Siin;;;-
thiorc iiach ihrer \'ertheihiii,i;- iiher die \\'elttheih>,"| namely,
Tnvhcchux >v>,s;/ho'».s', occurring' on the northein shoresof (West-
'.ni ;') Asia, Europe, and North Ameiiea, and 7'. o/>r.s7/,v, occur-
riufi' on the northwestern shores of North America anil the ad-
joinin*;' iiortheasteru sbor<'s of .Vsia. While I do n()t lind that
he bus anywhere j'iven the distinctive eliaraet<'rs of those two
species, he, in the abov»'-cited ]»aper, also nauu'd the ainnial
described and li<;iired by Cook, T. (livvtujiUN. V. Cuvier, in ISiT),
ill describinji' the dent it ion of the ''Morses," says: "Ces dents out
ete decrites d'apres plusieurs tetes qui senddent avoir ai)i)artenu
11 deux especes, a en Juj;er du nioius par les proportions de cpiel-
ques uues de lours parties, et noii seulenient par lY'tendue de
lours defenses, caraetore (pi: iiv.dt d(^ji\ fait soup^ouuor a SUaw
* Arctic Zoology, vol. i, 1792, pp. 170, 171.
nF<> says: "An excellent represeutalioii is also given in pi. 52 of the
Inst voyage of our illnstrions navigator, Captain Cook. It is easy to sec,
liowtjvcr, a reinarkalile ditlerence lietween the tusks of this last, and those
of the former kind Hgured in Jonston, and it clearly appears, thiit though
this (lilTorenco is not such as to justify our considering them as two distinct
species, yet it ohliges us to remark them as v.arieties; and it should seem,
that, in the regions tlien visited liy Captain Cool<, viz. tlie iey coasts of the
American ctrntinent, in lat. 7(», the Walrus is found with tusks nnicli Icuigcr,
thinner, and iar more sharp-jiointed, in proportion, tlian the eiiiii(!i'iiK, and 7'. cndh-i.
Tile llrst (V. roKinarus) was juineipally eliMi'acteiized l>y lia\ inj,'
di\ cijiini;' tnsks, idtont as !un,u' as the lenu'lli tit'tlie whole he;id,
tiiiMll\ grooved on the outside, and with two distinct ^iroovcs on
the inside: by the ])()ssession of live Itat'k teeth, the last iwo
very small; l»y t lie lower ed^c of the uasal openin;;' lieinn but
little produced ; lt,\ the occipital crest heinn' stionyiy developed ;
and iiy the yicat specilic i^ravity of the Itoiiesof the skull. The
second ( '/'. UtnijUhnx) was piiucii)ally characterized hy the tusks
equallinii'or ex('eores, p. 23.">.
tWicf^mann, in coiniuonting upon Fremcry'.s supposed spccifu' tlitlcvonces,
observes as follows respecting probal>l(» sexual and individual ditl'erenceft
in the tusks any Fn'iiiciy as distinctive of several
species, iiiid alter nieiitioiiiiif;' at leiinth otlar features of varia-
tion observed by biin in a eonsiderable series of skidls, describ-
inn several of Ids sj>ecirnens in detail, and arriving;' at< the con-
clusion that up to that time all th(^ supposed sjiecies of Walrus
constituted really l)ut a sinjile sjiecies, a(hled another, under
the ajtpropriate uaine TrichechuH lUibim. This with sid)se(|uent
autintrs has shared the fate of Freniery's si)eeies,* beinj;- (fonsid-
ered as based merely on individual variation.
As will be more fully noticed later, two nonunal species have
been foiuuled on the fossil remains of the NN'alrus, namely, Tri-
chirhiiH viniiniduits, DeKay, ISlli, and Ihlobvnoihn'ium larte-
tiaiiiii.. of Gratiolet, the former based on remains from Accomac
County, Virginia, and the latter on n^mains from near Paris,
France. LanUester, in 180"), added still another, based on
tusivs I'rom the. lied Cray of Knyland, under the na/ne Trichc-
chodon hnxhj/i.
Dr. Leidy, in ISOO, in a pajx-r on fossil remains of the Wal-
nis from the, eastern coast of the United tStates, a<;ain noticed
the dill'erences in the s'v/.v, leuj^th, and curvature of the tusks
in specimens fronx the northwest coast of Xorlh .Vuu'rica and
the ((unmon Walrus of the ZS'orth Atlantic, lie says : ''In the
course of the precedin;;' iuvestiji'atious |referiin,t>' to previous
portions of his i)aper], I was led to examine ;• specimen, in the
cabinet of tile Academy of Natural Sciences [of IMiiladelpliia],
consisting;' of llie sluifed skin of a j)ortion of the head envelop-
ing' the Jaws of a species of Walrus apparently ditferiu";' from
tl'e true 'I'rIclHTKN ro.siiianis. of whii "i, as characteristic, 1 have
viesved tin- ti;;'ures of the skull and skeletons as ^iven by Dau-
benton, Cuvier, and De Blainville. The specimen was pre-
sented by Sandwith Drinker, Esq., of Canton, China, and was
I)robably derived from the Asiatic shore of the Arctic Ocean.
From the worn condition of the upper incisor and molars, it
appears to have l)elon!;'ed to an old individual ; and in the case
of tlie lower jaw, the teeth appear to have been entirely worn
out. The tusks are verv nuu-li larger and arc narrower tlKin in
11-
*^Giubcl, ill ld55, refcirctl to I remoiy's and St.iimius'.s species as still need-
ing cf)niirniati()u: "Die von ri/iiicrv nacli (lev licsclial't'ciilieit dcr Ziilme
uuterscLiedenen Ai'teii, 'Ti: longidcim mid Tr. Cooli; siiid liingst als unlialtbar
erkanut wovdeii luul audi die von Staiiuius uiil" SuliiideldiD'crenzen begriiu-
detc Art, 2V. dubiiiH, entbelirt noch der wcitoru Bestiitigung." — Siiugcthicrc,
p. ViS, footnote.
t-.'l
Tin: si'Kf'irs nv the oenus.
21
the 7'. ro.siiKirii.s, niid llicy ciu-vc dowiiwiird, outward, and in-
wiinl, instead of contiiiiially divi'i'j;iii<;' as in this species. At
tlieir eMiei'.ncnce tVoiii '.lie aheoli the tusks are Iwo and threo-
«|U;utei' inelies ajtart, near (lie middle livc^ and ii (juarter ini'hos,
and at their tips only one inch. Their len;Lith is twenty-two
inelies and their diameter at the alveolar hord. r autero-posteri-
orly \\\o and a (piarter inches, and transversel\ one and a half
inches. Towards their lower part tliey are twisted from within,
forwards and outwardly." After (piotiny Pennant's remark
(already ju'iven, see [i. 17) about siuular ditlerencos noted by him,
he adds that "the superior incisor and molar teeth are also very
nuieh smaller than in the fossils of T. ronmarKs,''^ and he i^ives
measurement sshowinjit his diU'erence. He then says: "The hairs
of tlu? upper lip of the T. romiKintti are stated by Shaw, to be about
three inches lonj^', and almost ecpud to a straw in diameter.* lu
tlio specimen innler consideration, the hairs of the moustache
are stiff-pointed spines, not more than one line long- at the upi)er
l)art of the li[>, and they j;radually increase in size until at the
lower and outer part of the lip they are about one inch in lengtli."
lie further achls, in the same connection : " Since presenting tlie
above comnnniication to the Society, the Academy has received
from ]\Ir. Drhdcer, of Canton, an entire specimen of the AVah'us
from Xorthern Asia. In this individual, Avliicli measures in a
straight line eight feet from the noso to the tail, the tusks aro
ten inches long, and diverge from their alveoli to the tips, where
tliey are five and a half inches apart, but they are slender, as in
the stuflfedhead above mentioned, and appear as if they would
ultimately have obtained the same length and direction. Per-
haps the pecuharities noticed may prove to be of a sexual ehar-
acter."t
As will be shown later, we have hero the more prominent ex-
ternal differences characterizing the two species of AValrus for
the first time expUcitly stated from direct observation of speci-
mens. If Dr. Leidy had had at that time good skulls of the
two species for comparison, the other important cranial ditfer-
euces (noted beyond) could not have escaped him, and he per-
haps would have been led to formally recognize the Pacific Wal-
rus as a species distinct from the Atlantic Walrus.
I have met with nothing further tonching this subje(;t prior
to Mr. U. W. EUiott's report on the Seal Islands of Alaska,
* "Shaw's Zoology, vol. i, pt. i, p. 2'M."
tTraiis. Amer. Phil. Soc. Phila., vol. x.i; i)p. d"), 8G.
90
THE GKNUS ODOli.EXrS.
piildislied in l.STo, in Miiicli, iiiidcr the li('ii(lin,n' "The Walrus
of IJcriii*;' 8('a, (liosiniirns Jircticus)" lie says: — "I write Mlio
Walrus of IkriiKj >Sva\ bewuisc; (liis auiiaal is (jaitc distinct
from the AValrus oi'llie North Atlantic and (Irccnland, dilTciiny
from it specilieall.v in a very .strilduj;- manner, l»y lis <4r('alcr size
and semi-liairlcss skin."* Tliis is all lie says, lio\v(i\ cr, icsjx'ct-
mtX tlieir dilTcroiiCL'S, no reference Ix'in;^' made to the I'eally dis-
tinctive features. Tluis (he matter rested lill, in J.S7(), (lili Ibr-
mally lecoji'uized two si)ecic.s in his "List of the I'rincipal Use-
ful or Injurious .Mamnuds,"t in a eatalo,i;ue of a "CoHei-tion to
Illustrate th(; Animal Resources o''lhe Uidted States" in tli«' ex-
hibit of the National ^Museum at tiie International l']xhibi!ion
of 187(5, hohl in Piiiladelphia. This is inerely a Jiominal list, in
which ap])ears, under ^•Eofimarlda'," the foUowinj^, which I hero
fully and literally transcribe :
KasMAitcs (iiiKSis, (Illijrcr,) Gill.
TIic [Atl;iii(ic] Wiilnis.
Athnif ic (.'nasi.
i
]i()S.MAia;.s CooKii, (I'lvnicry,) Cij!.
Till' [Piicific] V'alrii.s.
I'aciilr ( 'i:|isi .
Here is simply a no.uinal recognition of two species without
ex])ressed I'easons tlierefoi'. in an article on the UoHinnridiC,
published in 1877, Dr. (lill a.^aiu says: "Two siK'cies ii]>])earto
exist — one {It. ohesiin) inliabiliuin' the northern Atlantic, and the
other (/i. t'ooldi) the n(n'tlieru l'acilie."|
Van !>ene(h'n, on the other hand, in 1877, distinctly afUrms
his disbelief in tin; existence of two spe(;ies. In referring to the
subject he says : "Xous ne croyons jtas (|ue h.'S IMorses du d6-
troit de Lehrinj;' diil'erent specih(|uement de eeux de la mer de
Batiiii ou de la Nouvelle-Zendde, et c'est a tout, a notre avis,
que Fremery a essaye de les repartir en especes distinctcs
d'a])res les modifications de leurs dents." He further adds the
festinuniy of von liaer as follows: "Von ]>aer s'est occnp6 en
lS;ir> de cett(! qnestion s\ FAcademitj de St. r(5tersbourg ct I'il-
lustre naturaliste m'eerivait, ])eu de temps avant sa uiort, an
sujet de la difference legere des Morses, a I'Est et a I'Ouest de
*lii']iortoii the Pry'iilov Groin) or Seal iMlandH').' Alaska, 187:5 (not paged).
Also, Keport on Ihe Condition of Ad'airs in Alaska, 187.'), p. lO).
tTliis "List" is aiioiiyuions, and is licnce, perhaps, not properly qnotablo
in tins eomieclion, althonyh its anthorsliip Is known to tlio present Awiter.
tJcdnison'sNew Universal Cyclopsedia, vol. iii, 1877, p. 1725.
1:^
SYNONYMY.
23
lamer (llMciiilc, (|ii"il ••c^iinliiit Ics (lil'liTcnccs coiimic (U's iiio-
(liliciitioiis loc'iik's*. ('" ii'csl piis r;i\ is dc Henry \V. IClliot, qui
coiisi(lei'(! ]<' ^lorsc dii iiniia]ili.
(,)iiad. lie. 1777. :i:!0.
jyjuiiK )iii(riinis (I nippiijtohimiiK J'lilso diihin, Mornc or S< a Jlornf. 1>av, Syii..
hvx,, iin.
7r(T//co.w, JMauti.xs, Sjiii/li,, 1(J7.'), 7'-^, jd. J*, tij;-. I). — V.c.r.in:, licsilir. mid Xatiir-
(jlcs( li. (J Win la lid, 171^, ijl; 17(>;>, lOli; ]Jc.sorip. el: J list. Nat. dii Gio'iil.,
17()."), 5;
En.— Goi-niK, ".Moi-jdioL, 1, 1817, ^11"; Act. Acad.
Cius. Lcoi>. (Jarol., xv, i, 1"^:'>1, H, pi. iv (dentition, etc.).— Vox Baku,
M(;ni. Acad. «t. Petersb. Math, etc., vi'= Kcr., ii, lH:ir», 199 (blood-
vessels of limbs).— Jakgkr, Miiller's Aieh. llir An.'it., 1844, 70 (den-
til ion — Labrador Hi)ecinien; .
Walross, Mauticx.s, Zocdogischc Garten, xi, 1870, 283 (etymology).
Wallrun sen Mors, RUYSCU, Theatr. Animal., 1718, 159, pi. xliv (liguro same
as .lonstou'i)).
Walrus, WuUM, Mus. Worm., 1G55, 289 (fig. from Do Lact).— Wymax, Proc.
IJost. Soc. Nat. Hist., iii, 1850, 242 (relation to l*acliyderma). — Lea,
Proc. Acad. Nat. Sci. Pliila., 1854, 2C5 (nsc of the skin). — Wheat-
LAXi>, Proc. Essex Institute, 1, 1854, G2 (remarks on a sknll). —
SoxxTAG, Nar. GrinncU Exj)!. Exp. 1857, 113 (woodcnt — group of
Walruses). — Murray, Geogr. Distr. Mam., 180(5, 128, map, xxviii*
(distribution; in part).— Hayes, Open Polar Sea, 186t, 404 (hunt-
ing).— Packard, Lull. Essex Institute, i, 18G9, 137 (fm-mer exist-
ence in Gulf of St. Lawrence). — Atwood, Proc. Bost. Soc. Nat.
* " Lcs Morses des cOtos d(^ Sibdrio ou do I'est do I'AhIo out les dents cani-
nes phis lortes quo les Morses do Spitssbcrg ot do GroDuland, nics tlisait-il
dans une, lettro "
t Ann. du Mus. Roy. d'llist. Nat. Bclgique, pt. 1, 1877, 45.
tibid., p. 17.
I
A
24
OD015ii:\i:S UOSMAIM'S — ATLANTIC WALRUS.
H
Hist., xiii, L~7(l, ii'JO (icnuaks on a wkiill IVuiii IIk; (iiill' of St. Law-
rence). — TuiiNEK, .Toiini. Aiiat. iiihI I'liys., v, 1^70, lir)(rcliifions of
luTicardiiim). — L'ink'. l»inii'.li (ircciiland, Ij-T", 1'.'() ((listiil)nliun),
!J1>', -'.V,', -.;:•,' (fliiisc).
Arctic Wtilnix, I'kxxant, .'^yimii. (^'ikkI., IT"!, :s;i,">; Aiftic /in".]., v,'(l cil., i,
ITU--', 1(W (ill jiart).
Fossil It'iilnix, I'.AiM'dN, Lfiiicloii I'hil. Mil;;., wxii, 1-(C>, '.)> (iii> locality). —
MnciiKi.i,, .Smith, iV (.'(h)1m;i!, Ann. Now York L.vc Nat. Hist., ii,
li^'i^, 'J71 (Ibssil, Aciiiac (-'o., \'a. — doiibf I'iiIIn' ict'rircil to tlu' cxist-
iiij; species) ; IMinli. New I'liil. .Jiiiiiii., y, 1~'J~. :!-.") (alisiiact of tlio
last). — ![a!;i,.\\, Kdiiil). New I'iiil. .loiirn., xvii, ISM, ;>ti(l; Tiaiis.
(Jeol. Soe. reini., i, l-^li.") 7.">; Med. and I'livs. Ke.caiclics, Itfll.i, 277
(same si»eeinieli). — Lyki.F, iV OWI'.N, I'loe. Lond. (ieid. Soe., iv, I'ri;],
;W (fossil. Mavtiia'a ^'illeya^d, Mass.): Aiiur. .loiiiii. Sci. and Arts,
xlvi. If};'., :U'.» (same).
La Viichc nuiritic. IJiiis.sox, lliii. Anini.. 17.">(i, i^.
"Morscli, (i.mi:m\. L'eisc dureli liusslaiid, iii, 17;')!. It!.').
Morse ml \ (tvlic iiiuniK. lU rrox. Hist. Nat., xiii, ]7t).'), :C)f', pi. liv (animal). —
l>Ai:iii:xr(»x, Million's Hist. Nat., xiii, I7(i.">, tl.'i. i>l. Iv (sknll). — HoL-
i.AXKiii;. Alire^ed'llist. Nat. des Quail. Vivip.. i, 17'.l(l, pi. xii, ti;;-. :!. —
1'. C'rviKi;, Diet, des Sci. Nat., xxxiii, Ir^K!, '..'7; Dents des .Mam..
lft>5, )i^:i, 1)1. xcv.
Morse, HUET, Coll. de Mam. dii Mus. dllist. Nat., l^Ot*, .V.), pi. liii (ti^-. from
IJullou).
Sea Cow, Siiuldhasi, Phil. Trans., Ixv, 1775, 249.
PAoca, IJoxXAXio, KenimNat. Hist., i, (no date), !.'>'.), ]d. xxxix, tii;. •i7 {a i)oor
veiji'esentiition of J)e Laefs ii;;ine, witli the yonn;; one omitted).
P/joca rosmaritx, LlNXl5, Sy.st. Nat., i, l?")!^, 38.
2Vich<:chHs rosmariis, LiXiXi':, Syst. Nat., i, 17110, 411. — M(r,t,i;i{, I'l'od. Zool.
Dan., i. 177ti, 1.— Sciii!i:iJi:i!, Sihii;etli., ii, 177."). -'(W, j)!. Ixxix (from
Ballon). — Zl.MMKUMAXX, Geogr. (Jesehielite, i, 1778, iiDD; ii. 17H),
4t>l.— FAitiMcns. Taiina (inenl., l7Kt, 4.— Li{Xi.i;iii;x, Syst. He;,'.
Aniin., 17."'7, .">'.»:{.— (;.mi:i.ix, Syst. Nat., i, 17."'-*, :")'.•. — SllAW. Nat.
Miscel,, 171)1, pi. e( Ixxvi ; (Jen. Zoill., i, lH(t(», '2:U (in part), lig. (W,
(from Jonston). — nu:Mi;xitA<'i[, Handh. derNatnrjjeseli., 178s. ij-^;
18-Jl, l:S(); It^-J.'). 11-J; Abl.ild. natnr. (;e, U71.— I?AUT()X, I'liil. Ma^^, xxxii, 180.-., 98 (fos-
sil; locality not stated),— (i. Cuvji:i!, Tableau el<'!menl., 171)8, 17'i;
Lemons 1."), .''.0, 01, 04, 0"^, 7.") (dis-
tribution). — l)i:.sMAi!i:sT, Noiiv. Diet, d'llist. Nat., xxi, l-^l>, ;!90;
Mam., 1820, 2.'j:{,— ScouicsDY, Account Ant . Kcfxions, i, 1820, 002 (K«^n-
oral history). — •' Ki:i;sri;i!\. CapitisTrielieelii IJosmari Descrip.Ost,,
1821,— ."■— ScAinii, Kdinb. I'liil, .Jonrn., ii, 182."), 28',! (Orkney): .»ar-
diuc'.sNat. l.ilirary, .Mam., vii, 1838, 219, pi, .xx (orij,'inal li,u;uro
ifa
SYNONYMY.
25
of animal).— Harlan, I'auii. Anier., 1^'2'i, 111 ; Ediul). Now Pliil.
Jouni., xvii, WM, ;?G0 (fossil); T.van.s. (ieol. Soc. renii., i, lf*\'>:>, 72
(eaiiic); M('*l. and Pliys. IJcpos., 1835, L'77 (same).— GoDJLVN, Anicr.
Nat. Hist., i, 18-^0, a.Vt.— Sciiix/, Nairn-. il.-r.Siiii-.-tli., 1>'-J7, 1(;'.>, pi.
Ixv (two liyiiics — " AbbilduM!^ nacli BliiiUL'ubaLli imd .SL'lniiid'"). —
Lkssox, Man. do Mam., 1827, 'J08.— Ross, App. Puitv'.s Fourth Voj-.,
1828, ID'2; Api>. Koss'.s Second Voy., I8l!5,xxi.— Fi.kminc, I'.iit.Anini.,
1828, 18.— liAi I", Natuiw. Aldiandl. Wiirtonib., ii, l-^2-\ lii7 (denti-
tion) ; "Bull. Sei.Nat., xvii, 1821), 280" (abstract).- Fisciiicit, Synop.
Mam., 1821), 213. — GuicKiN-Mi'iNT.vn.i.K, Icon, du I'ei^ne Auim. do
(i.Cuvi-r. ilani., 18211-18:58, It), pi. xix, li^'. .") (animal).— FuKMr.iiV,
lji,jdra;;t()t de natuurk. Wctcnscii., vii, 18:?1, 384.— Dkloxgciiami'S,
Mom. Soc. I>inn. do Norm.'iudio, v, 131. 101 (doutitiou). — WlL-
SOX, Nat. llisl. Quad, and AVIiales, 1837, 14.", pi. cccxxxiv, lig.2
(iininuil); Kncycl. Brit., 7th ed., xiv, 12"). — BKr.r., Brit. Quad.,
1837, 2r)8 (animal and skull; original ligures). — Vox Baku, Mem.
Acad. .St. I'etersb. Math., etc., (5" s('V., iv, 1838, it7, pi. xlvii (distribn-
tion). — WllXi.MAXX, Arch, llir Natingesch.. 1838, 11:5 (dentition). —
IIamii.tun, Jaid. Nat. Libr., Mam., viii, 1831), 103, pi. i (animal, and
woodcut of skull, — original ligure). — IJiCltAUD.SOX, Zool. Boeciiey's
Voy., 1831), ().— Br.Aixvir.r.i;, Ostoojiraphie, I)esl'ho«|ues, 1840-.'>1, 19,
pi. i (skeleton), pi. iv (skull).— Dkaay, Nat. Hist. Now York,
ZoiJl., i, 1842, 50.- ZiMMKKMANX, Jahrb. fiir Mineral., 1845, 73. —
WA(iNr,i{, Schrcber's S,'iuy.(tli., vii, l<|i;. , V'!^; Odoutog.,
1855,(^2, pi. xxxvi, fig. 5 (dentition). — Nii.ssox, Skand. Faini., 1847,
318.— (iKKVAis, Zool. et I'al. Fraiicais, i, 1^4-'-.'>2, 14(1.— Uuay. Cat.
Seals in Brit. Mus., l.-.'.O, :;2; I'roe. Zo.il. .^oc. Loud., 1S53, 112 (ou
iittitiules.'ind ligures): fat. .S'alsand AVluiles. 1K!1!, 3(>, 307.— OWKX,
Proc. Zool. Soe. Loud., 1>'.")3. I(i3(anat. and dentition) ; Ann. ami
Mag. Nat. Hist., xv, 185.'>, 22ti (from th • foregoing); Cat. Osteol.
Coll. Mus. Collego Surg., 1853, 031 (skeleton); Fucycl. Brit., xvi,
1854, 403, iig. 112 (.skull); Odontography, 1'^.>I. 510, pl.exxxii. tig. 8
(dentition); Orr's Circhuif the Sciences, Zoiil., i, li-54, 230, tig. 27
(skeleton); Comp. Anat. an, 4".t(». 498,
507 ; iii, 1808, 338, 524, 780.— Blasiu.S, Faun. Wirb. Deutschl.. i, l-i57,
202, ligs. 148-150 (skull).— Van dkij Hokvicx, Ilandb. Zo,.|. Kugl.
Ed. ii, 18,">8, 007.— Vo\ SciiUKNCK, h'eiseu im Aiuur-Lande, i, 1-59,
179 (in part only).— Lkidy, Trans. Amer. Phil. Soe. Phila., xi, I'OO,
pis. iv, v (in part); .lourn. Acad. Nat. Sci. Phila., 2d ser., vii, l-ti9,
410.— Woi.i' A- Sc[.ATi:i!. Zool. Sketches, i, 1801, No. 10.— Gkijuai'.d,
Cut. Bones .M;iui. Brit. .Mus., 1^02, 14."). — Nkwtox, Proc. Zo.ll. .8oc.
Loiul., IftU, 499.— .<< i.ATiMt & Bautu-.TT, Proc. Zool. Soe. Lond.,
1HI')7, Hl-i, ^^p.t.— Vox .MiDDr.NDOitri', SiliirisehoReise, iv, 1S(;7, 934 (in
l)art only).— Buown. I'roe. Zoiil, Soe-. Loud., L-*lH, ;!35, 427 (liaMts
and disiribulion) ; Man. Nat. llisl. (Jreeuland, 1-T."), :!,'>.— .Mii;ii;,
Proc. Zoid. Soe. I, Olid.. I-'(W, t;7 (rejiori on cause of death of sprci-
meu in Zoiil. (laid., I.oiid.); H70, 5"'l ; Trans. ZoiU. Soe. I.nud., \ii,
pt. vi, 1H71,411. pis, li-lv (anatomy), — (Jii.ri.v. I'roe. and Trans, Nova
Scotia Inst. "at.. Sci.. ii. p(.3. l.-'70, 123(witli ,i plate).- I.'ki;;;' . /j id-
20
ODOILKNUS :.>lSMAUrS ATLANTIC WALlfUS.
ay, XcwtiiuiiillMiid). — lli:c<;i.iv, Rei-
MMi iiacli (IriiiXoiditdliiiiiiccr, iii, 1>74, ■11! (I ml "its anil ]., :!'• stT., ii, 1^74, 1G4 (IoksII. Fraiici.').—
(Iri.i,i\i;i;, I'loc. '/.oiW. Sue. Loiiil.. 1-74, ">"^!l (size 1)1' liliMiil-coi'jiii.s-
ck's). — 1'i:ii.1)i:n, Zuiild^iisi, ;!(! scr., i. l-^TT, ;!(!(> ((lisiriliiiiinii and
t'lKHl).— ^'A^■ lirNKDiix. Ann. Mns. l.'oy. li ibnlion, ^rncral lialiits. and liissil remains). — IviXK,
Hanisli (iifinland. its l'c(i|)l(' and its I'lodncts, 1~77, VHi.
Trichicluin loiKjitli iin. l'i:i:Mi;i!Y. l>ijdra;j; lot dc Xalunil;. Wchnsidi., \ i, l^'M,
:W4.
Trirluclnix vlr, j)l.
\ix, lijjs. I, a, h (I'ds.sil. Acconiai' C'i>., Va.).
t Tricliivliiis (Inhius, 8i.\N'Nirs. MiiUcr's Aicli. iur Aunt., l.';4"J, 4(t7 (witliout
locality).
lioxiiittfUfi (ircliciiK, LiLLlKiic )!!(;. Fanna rd'vcr.s Svcrij^rs ocli Xorgos Uyj^gr.,
1874, »i74.
Eobiiitiniii iriclicdnix, T..\.M()NT, Seasons w if litho Sea-borHes, 1801, 141, 1(57 (two
]ilatcs). — (in. I., Johnson's Xcw Univ. Cycio])., iii. 1^77. (i:!li.
liOgniurus ohcmis, GiLi., I'roc. llsscx Inst., v, ISGd, 11! (in i)ait only); Interna-
tional Exliil). 1871), Anini. Resources U. S., No. x!, l'^7(), 4 (Aflantic
Waliiis; no deseiii)tion) ; JoImisou'.h X^(^w Univ. C'ycl.. iii, 1877,
17-,'.">.— r.\cK,\i:i), I'loc. Host. Soe. Xal. Hist., \. \>i\i'<. v!71 : .Mem.
Dost. Soe. Xal. llisl.. i, 18()7,'J4(i (Ib.ssil).— Lkidv. .louni. Aead. X'at.
.Sei. Pliila., viii, 1-77, "JU, ]il. xxx, fig. (i (I'o.ssil. .Sontli ('ar(din;i).
Odohiiiotlivriiiiii hirtiliiuiinn, (.ii;ATiiH.i;T, Hull. Sue. (li'ol. de I' ranee, •."' si'r..
XV, l.-3."^. I'l'.' 1, 111. V (fossil, near I'aris, Trance).
OdoHldbiniiix ro.s/, -f//(s, SrXDKVALl., (il'ver. K. Vet. Akad. I'orli.. 1>.V.». 441 ;
Zcit.x li. (Ji'sammt. Xatnrw. Hallo, xv, 18()0, '2711.
Odobainix ronmanix, Mai,mi;iii:x, (JlVer. K. Yet. Akad. I'orli, 18il;!. (18(14), 1"0
(food and liabi(s), 5(1."), ]il. vii (dentition) ; AViegniann's .Arcliiv f.
Xatnrgcschichte, ]8i)4. Cw (tran.slated from (UVers. K. Vet. Akad.
Forli., l-r,;!. i:iOet sell.). — I'KiT.Hs, Monatsli. Akad. Wiss. Jicrlin, 18(;4,
ti8,"), jd. (tlcntition) ; Ann. and JIag. Xat. llisl., (;J), xv, lS()o, 3u5
(abstract).— Rink, Danish lireeulaud, 1877, 4;>0.
t Tr'nhicodon huxli\j\, Laxkicstkh, Quarter. Journ. Gcol. ,Soc. Loud., xxi,
18(ir), yjC), ids. X, xi (fossil; Rc8.
trhixii iiraitia, Fahricius, Fauna Grusnl., not Callorhinus uraintta ; seeBKOWN,
Proc. ZoiJl. Soc. Lend., 18G8, 357, 348.
Morse; Vuvhc marine ; Chvral marine ; B6ic d, la grande dent (French).
Bon hiitriniis, RuY.scil, 1. e.
Hvalroxs (Swedish and Danish).
Harhcst; Ilrahuns (Norwegian).
Morsk (Lapp).
Wallroiis; MtrrjiJ'rrd (Gennan).
Walrus; Sea Cow; Sea Home (Fnglisli).
External Char acters.— As regards general form, the head,
ill conipiirison witli the size of tlie body, is rather small, squar-
EXTKUNAF- CIIAHAfTKlIS.
'21
isli ill (Hitliiif, Itiil iiiiicli l()U,u('r lliiin lii'(i;i(l. witii llic iiiii/.zle
iihiiipllv iniiiciilcd :iiul •soiiicwliiii l)i!oltc(l i>y llic llfs. Tin- nnsiiils arc soaicwliat crcsccntic
ill --h.ipi', placi'd \ I'iiicaliv. \\ilii tlif uppiT i)ai'l iiimc ('\piUirly. The tail is scarcely, if
at all, \ isible, bein^i;' enclosed within tlie te.iiUineiitsof the body.
The fore limbs are free only from tlieelltow; as in the J*iniiii>eds
generally, (hey are jireatl\ ex])aiided, llai, and somewhai fin-
like, !)ut wilii miK'li more t'reedom of motion than is the case in
tJK' PltDvidiv. They are armed with live small llat nails, i>laeed
at considerable distance from the end of the cartilaginous toe-
tlap. The first or inner diyit is sli;^litly the htni^'est, the others
beiii;;- each stu-eessively a little sliorter till the fifth, w hich nearly
equals the first. The hind liinl) is enclosed within the te;;u-
nieuts of the body nearly to the heel; the free portion when
expanded is lan-sha])e(l, but w hen (!losed the sides are nearly
parallel. The first and fifth digits are eousiderably longer and
larger than the middle ones, the lifth being also rather larger
than the lir.st. They are all ])r(n'ided with snndl nails, placed
at some tlistanee from the Knu\ of the toe-Hap. The soh's of both
lore and hind extremities are l)are, rough, and '' warty,"' and
the dor.sal surf aee of the digits as far as tin; jn'oximal jdialanges
is also devoid of liair. In the young and middle-aged, the body
is rather thiekly eovere \\ lii're
more or less wriiiUled and thrown into Idhls, es|ireially o\erlIie
slionldei's, where t lie folds are dee|i ami liea\y. 'IMie axerau'e
Icii^i'lli of lour adiill males e\amined is aliont 10.', Icei. \ar\in,u'
from!).] lo 11 i'eet. .\iltliors, however, eoiiiiiionl.\ .ui\e rather
lar.u'er dimensions, and a length ot'lwehc feet is said to lie not
inrrei|iieiitly attained. The lar,!;vst Itristles \ary in length iVoin
'J.L'."» lo LI.Tl inches.
Vnn>> Dr. Miirie's paper on the ;4('iieral anatomy ol' a yoiinji
individual I add a few riirtiicr details. Dr. Miiiie descrihes the
muzzle as capahli' of ureal niohilily, and the mxstaeial hristlesas
eiuviii;;' in dilfereiil diicctioiis aei-ordiiiu to the imiseiilar tension
of the ]»arts to which they are alt ached. •'When the nostrils are
rt'lax<'d they drop fmwards and the hristles inwards. At such
tiiiu'.s tiie n arcs are apart liilly ].\ inch; Itnt when they are <'oii-
trii. Il'.t.
KXTKk.NAL CIIARAOTKHS,
2\)
III respect lo llie iiiystacial l)ii.stles, Dr. Millie's li^^iii* s of tiie
lieail iiiid iiiii/zle of tln^ .Voiiiijf s|)e<-iiiieii desciilted \ty liini
(drawn lioiii |)lioti<;;i'apii,s, sniiie tVoiii the liviii;; animal) rejt-
resent tlieiii as (|iiite Ion;;', tlie loii;L;<'st lieiii^' sailles lu'come slioilcr in adult life, heiiii;- |)erliai)s worn oil
hy coiistaiil friclioii. Tlie itrislles in the specimens I have sfteii
JMiic no rcsciiildaacc lo l!ie lon.i;' curving' luistles i'i;;iired and
dcsciilied hy Dr. Miiiir as existing' in the yoiiiiu animal. They
were coiisider.ihly (oiiethirdj longer, however, in tin; ycainu'est
of four specimriis in I'rofessor Ward's collection than in the
oldcsi. uiviiiu siipjioit to llic oiiiiiion already slated that llie\
itccome shorter as the animal advances in a^'ct
.\s already noted, the fore feel are formed much as in other
IMniiipeds, more nearly aurecin;^', however, with those of the
()liirli(l(f {h:i\\ with those of the t'kocitliv, esjtecially with res])ect
to freedom of movement, having' the power of pronation and
supination to a considerable decree. "In the Walrus," says Dr.
Millie, '•ihe hiimeriis, radius, and ulna can he so placed that
liiey meel at ail acute aii^le, the lower limb of which is in a
uivat measure fre<'. '{"lie dibits, on the other hand, can to.i'ellier
he t iiriied backwards at a sliari» alible with the radius and ulna,
>o that the bones of the limbs altouellier forai an S ■shaped
liUiire. In the Heal tlii^ anlibraciiiiiiii and dibits bend on each
i''
f:
Sfc l';isMir;i(''.H GcniiMii tiimsliilidii, p. i:i'J.
t hi J',ill;is's li;; I 111 ■fill lliH "Icoili's'") of ;i \ nini;; i\;nii|il<' oC llir I'lnilic Wnl-
nis, llic iiiysliU'iiil luisllcH arc irin'csculcd as very luiii;-, as in llic vnmi^ of
11k: Allaiilic species.
30
ODOILENTS }{()SMAl{rs — ATLANTIC WAI.KTS.
I'. V,
■ r'\
other more iiiiniiliiily, tliiis < In tlic net of swiniiniiiii
the Wiilnis cv itlciilly ciiii use its lore limb iis \\\v jis tlic cUiow.
with ;i kind of rotiiry movi'iiiciit (>; the maims and iiiitiliracliimii;
Init in the Seal liic lotaiy a<'tii)n takes phice only at the wrist,
and al»o\(' that a sort ol'^inylynioid of hack and I'oiwaid mo\c-
nient."
••The pahnar snrfaee or sole of {lie nianns is not r.iiiiKe a par-
hji' sliovel in li;iiire. 'I'heic is a .!;reat eallons. ron,i;hei;»'d and
warty pad at the ]>roximal eml or l»all of tlie hand: an, while. La-
nioiit says a lull ;; row ii old male will wei;,di at least. .'5,000
])oiiiids.f Aside, t'loiii Dr. .Miirie's iiieasiireiiieiits ol'a .voiiiit^spe-
ciiiieii, I have iii(;t, witli iiodetaihid liieasiii-eiiieids «)!' the, Atlan-
tic Walrus, exeepi thosi! j,'iv(^n by I)i'. iii,|: whieli are, as fol-
lows :
I'-t. III.
JlxiniiK! length 12 3
j,iii-iii <.r ii.aii I r,
111 cm I 111 of iiiii/,/lc 1
J li--l,iiir<' I'lniii ii(>s<^ tr> vyi' H
l)i^I;ilii(j hcl Wren ryi'M lii
J", \lrliMion (if t ll-.k lir\(ili(l 1 lie llionlll I II
] li,->liilicc 111' IiisUn ;i|i:i|'I ;it Iiusc J
] lisl.inii' of lilsUs :i|i;irl :it (i|is II
Ij(Ii;;||i (if fciiv ||i|i|i(f U U
ISlvjiillli ,,f foil llipiirr I I
Li'iiy;lli of hi Ml I lli|i|pir 1 In
Urcadlli of liiiiil lli|i|ii T, ilislimliil 2 I'eet, with a
ciieiiiiit'erenee at. the shoulders of Id feet ; and the leiiii'th of the
tiisUs as L'tl iiH-hes. Ilaniiltoii I savs an iiiilix idiial Uilled inOrlc-
lie\ . ill ISLM, whieli he saw, '• was alioiil ten feet in leiio(li," with
the head \'.'>S inches in lenj^lli. From the si/e of the lusk.s (ex-
serted S.j inches) it ajipears to lia\c been far from fidly ;(rowii.
Daiilieiitoii j^i\es the leiiulli of the specimen he described as
lU feel, with a eirci I inference at t In-shoiilders of S I'eet. Ijuniont
'Acromil of the .\irl. I.'c;^., vol. i, |i. .'ifi-,'.
>.Mr. I.aiiioiil, ill his •'Sca.soiis willi the Sca-hor.scs" (p. — i, ;;ivc.s llic
vii;;iil of an old male as il.noii |ioiiinN. Imf in his " Yachfiii^; in I Ik; Arctic
Seas" (p. "'.I), he says, "A fiill-si/eil ohl hull Walrus imist weiyli iit least.
GjCiMi Ijis.. anil ,-.iich a Walrus, if \eiy fat, will inoiluee Ci.'iO Ihs. of Iilnbber,
bul sehloiii niore I hail 7MI) Ihs., whieli is I think the iinnuic :inioiint
yiclileil li\ till 1I10-.I ohese of our \ ill iiiis." lie speaks, luiwever, ill aiioflicu
place (p. !-;(), of one that ''yiehleil hetwccn 70it ami -0(1 ]ii)un .IS II I'tTl ."» iiiclics.* Mvnuri iiM-a^iiic-
iiK'iils III' lliii'i- ihIiiII iii:ilrs rruiii iilisl iin'iil (Milltilj sKiii-i :il'i- :ih
I'liHows: (I) li'lljilli (I'lnni liosi' ti» lilil), 10 I'ril ■"» imliivs; (J)!» I'l'i-I,
i'l iiirlii's; (;;) 10 li'it III iiiclics; (l)S fi-cl o inilics. 'i'lii- liist,
lliirr Will" I'lilly inliill, w liili- uiii- ■if I Ill-Ill. Id jilili^r I'imiii il^
hroKiii, wmii liisks;iii«l |>iiilly ii;ik«Ml, sciincd skin. Wiis \<'r\ nlil ;
tlic oIIkt \v;is iiiiI iiiori' tli;iii Iwu lliiids ;;r"\\"- 'llii'si' iii;i\ :ill
li;i\r lii'cii s|i('riiiH'li.s ol' less tliiili I he ;i\ riilj^i' sizi-. AiMilij;,
liowiMT, l."» lo IH iiu'lu's lor I he l<'n;;lli ol" llic liiiHlliml* (in>' iii'ii-
ilirlllilfil), wiiiilil ;^i\i';i irii;4tli ol' ;il>i»iit IL' I'lrt lor lln- l;ii;;»'r
iliili\ iilliills.^
Most ol' t lie old w lilcis wcir coiilctil wit li stilt in;;' it to lie iis
hiip'iis iiii o\ :iiiil ;is thick iis ;i Iio^^sIkmiI. 'I'Iic iiccoiiiits of lli<^
color iirr iilso ilisci'c|Kiiit ; I'liiiriciiis's stiitciiiciit Hint llic color
\;iiics V ''li ii;^!', Ilic voilli;,'^ liciii;; liliick, tiicji (|iisk.\, liitcr |»;ilcr,
iiiid liii.ill.\ ill old ;i,iic white, luiviiij;' Itccii (|iiot(d it.v most suit-
sc(|iiciit coiii|)ilcis. Writers who liiivc ;;i\cii the color I'loiii
iictiiiil oliscr\!itioii li;i\c never, however, coidiriiied l''id»riciiis's
;iccoiiiit, they iisiiiilly descrilMii;;' tln' c'olor of IIm^ hair iis "yel-
lowish hrowii," "yellowish jiijiy," " tauiiy," ''very Ii<;ht yellow-
ish-;;ray," etc., some of whom explicitly state that after (;xteiid<'(l
oliservatioiis tliey have never met with the chaii;^e,s of color
with a;;(' noted hy l-'aliriciiis. TlmH, Mr. Ivohert, l»rown say.s
that althonjili he has seen Walruses of all staj^os, from Itiitli
until nearly mature aj;e, he never saw any of a l)la<;k color, all
hf'inj;' of" the ordinary hrown color, though, like mo.st animals,
they ;,'et lijilifer as tln^y {i;v()\v old."| Scorcsby says that the
skin of the Walrus is coverod "with a short 'llowish-brown
coloied liair."§
Dr. (lilpin stat<'H that Ids Jiahrador siHtcinuMi was thinly cov-
ered with " aresse,d li};ht yellowish-f^reen hair," about an inch
in len^'th. lie jwlds that the, siirfju!»! of the. whole skin was
mmm
34
()Doba:m:s uosmaruh — atlanthj walrus.
1
I i
covcivd by "sciU's iiiul bald warty patches," and tbat tbo skin
itsi'b" was tbrowii hito " welts and folds'' <»ii tlu' neck and slioul-
deis.
Mv. Ibown Ihrther says that " the, very circumstantial account
of tlie number of mystacial bristles given in some accounts is
ni(»st erroneous; they vary in the lunnber of rows and in the
numlM'i' in each row in almost every specimen. They are ele-
vatt'd on a minute tubercle, and tiie spaces between these bris-
tles are covered with downy whitish hairs."*
Many other writers also note the scars and warty i)atches and
partial absence of hair n'ferred to above by l>r. (rili)in. Mr.
Brown, in speakinj; of those he met with in Davis .Straits, says:
^' I iiavc seen an old Walrus <|uite spotted with leprous-lookinji'
marks consisting of irregular tubercular-looking whiter carti-
laginous hau'less blotches; they api»eart d to be the cicatrices
of woumls inliicted at dilfercnt times by ice, tlu' claws of the
Polar Uear, or nu't witli in the wear and tear »>f tiie rough-and-
tnndde life a Sea-horse must lead in X. hit. 7 P."* ^Ir. Lamont
further adds that in the Spitsbergen seas the "old bulls are
always very light-colored, from being nearly devoid of liair:
their skins ai'e rough and rugose, like tliat of a Hidnoceros.
and they are gt-nerally quite covered with scars and wounds,
inlbcted l)y harpoons, hinces, aiul bullets wiiich they have
escaped from, as well as by tlie tusks of one another in tights
.init.ug themsehes.''t From these reports, especially tiiat of
Mr. IJrowM, Dr. 'riei ]ias iid'erred that the Walrus is subject
to skin diseas hat the; ''glandular spots*' thus produced
are mista' liealed cutaneous \\ounds.'' However this
may be. etty well established that many of these marks
are reall^> ars of wounds.
Respecting other external characters, especially the tusks,
and their variations with age, sex, and accidental causes, I
transcribe the following from Mr. Lamont's entertaining book,
which will be found s(» freely ([noted in subsequent ])ages:
"Old bulls," he obser\-es, •<■ very fre(iuently have oue or both
of their tusks broken, whi(!h may arise from using them to assist
in chunbering up the i(,'e and rocks The calf has n(»
tirsks the lirst year, but the second year, when he has attained
to about the size of a large Seal, he has a i)air al)out as large as
*l*ioc. ZoiJl. .Soc. Loud., IciiJd, p. l-'cr.
t Seasons with tlio .Smi-Uoi'ses, p. i;57.
ITraiiy. Zool. Soc. Loud., vol. vii, 167'.i, p. 122.
44
KXTF.HNAL CIIAIIACTKRS.
35
I
tlu' fiiiiiiieteethofa lion ; tlu' tliinl year tln'y lire about .six inches
lon^.
"Tnsks vary very ""K'h i" >^^'/'^' i"'^/-()l' hnll's tasks may be stated
as twenty-lour inehes lon<;',* and lour jxmnds apiece in weight;
but we obtained several pairs above tliese dimensions, and in
particular one pair, which measured thirty-one inehes in length
when taken out of tlie head, and woi;;hed eight pounds each.
► 'uch a pair of tusks, however, is extremely rare, and I never,
ti» the best of my belief, saw a pair nearly eciualto them among
more than one thousand Walruses, although wc took the utmost
pains to secure the best, and always inspected the tusks care-
fully with a glass before we tired a shot or threw a harpoon.
•' ( "ows' tusks will anraiic fully as long as bulls', from being less
liabh^ to b(! broken, but tlu\y are seldom wo/r than twenty inches
long and three ])onnds each in weight. Tiiey are generally sot
much ('loser together than the bull's tusks, sometimes overlap-
ping one another at the points, as in the case with the stuft'ed
specimen at the British .Mus■
^1^
iK
m
!^-''
3G
ODOBvENUS KOSMAKUS — ATLANTIC WALRUS.
suspected by AVieji'inaiiu luitl Staniiius (see (intca, p. 11>), who
believed that the Jeiiiale liad longer, sk'udeier, and more eoji-
veryiny tusks thau the male. Tlu.'re is also a speeimeu in the
collection of the Museum of Comparative Zoiilogy, Cambridge,
ill which the tusks are very long and slender, and eoii^^-ge
to such a degree that their points actually o\erlai).
Ill concluding this rat!ierrand)ling notice of the external cliar-
acteis and aspect of the Atlantic Walrus, I append the (juaint
and very correct description of tins animal, Avritti'U by the mis-
siouary Egede as early as 1740. I give it from Kriinitz's Ger-
man translation from the original Danish :
"Der Walh'oss, oder das Meerpferd, ist eine Art \on Fisch,
desseu Gestalt einem Seehunde gleicliLjmmt: jed .-h ist es
Aveit grosser uiul stiirker. Seine Pfoten sind mit fii f Klaueu
verseheu, wie die I'foteu des Seehuudes ; doch kiir/er von Xil-
geln ; uud der Kopf is dicker, ruuder und stiirker. Die Uaut
diescis Thieres ist, vornehmlich am liaise, einen Daumen dick,
imd aller Orten faltig, uud runzlig. Es luit eiu dickes und
bratmes Haar. In dem obern Kiuubacken sitzeu zwey ki-umme
Ziibne, welche aus dem Muiide liber der Unterlipi)e hervorrageu ;
und eiueu oder zwey Fuss lang, uud bisweileu aueh wohl noch
liinger siud. Die WaUi'osszalme sind in eben solcliem Werth,
als die Elephanteuziihue. luwendig siud sie dicht iiud fest, au
dor Wurzel aber hold. Seiu Maul ist wie eiu Ochsenmaul;
uuton iind oben mit staclilichten Borsten, in der Dicke eines
Strohhalms, besetzt, uud dieso dieuen ihm anstatt eines Bartes.
Oberhalb des Mundes sind zwey Naselocher, wie bey dem See-
hunde. Seine rothe xUigen seheu ganz feurig aus ; und weil
sein Hals ganz ausserordentlich dick ist, kanu er nicht leicht
um sicli herum selien ; und dieserhalb dreht er die Augeu im
Kopfe horum, waun er etwas ansehen will. Er hat, gleich dem
Seehunde, einen sehi* kurzen Schwauz. Seiu Fleisch hat eine
Aehnlichkeit mit dem Schweinenlleische. Es pflegt sich tlieses
Thier mehrentheils auf dem Else aufzuhalten. Indesseu kann es
so lange auf dem Lande bleiben, bis es der Hunger niithigt, in
die See zu gehen ; iudem es sich von deuen Fischen und Meer-
Insokteu uiiter} ilt. Wdun es im Zorne ist, briillt es wie ein
Ochs. Die Meerpferde smd beherzt, und stehen sich eiuander
bis in dea Tod bey. Sic leben in bestiindigem Kriege mit denen
Biiren, denen sie mit ihreii grossen und starken Ziihnen genug
zu schaffen machen. Oefters tragen sie den Sieg davon; und
.il
KXTKHX/ T> (FrARACTKUS.
'■> 7
■woniji'stciis kiiiiii>l<'ii sic so hiiiuc, I»is si(> todt ziir lOnlc iiicdcr-
faiU'ii."*
AiHitlicr iiftiMiiit of the Wiilnis, lioiii its hciiii;' one of tiic
Oiirlicst extant, is also of espcciiil iiilcrest in tlie present cou-
iieetioii. 'riionj»li repeatedly eo})ie(l, in part or wlioll.v, l»y tlio
earliei' authors, and also l»y von J':u'r, I tliink it deseivin.n' of
reprodnetion here. It was written by Irof. A. E. Yorst, and was
based on the yonnj; specimen taken to llolhuitt in lOl.'J. It is
liere c()pi<'d from I)e Laet (l)escrip. [ndiic Occident.), by wlioni it
was pnbli.slied in HJ.'>;>:
"BeUuain lian<; niaiinani vidi, inagnitndlne vituli, aut eanis
Britannici majoris, Plioc.T non diasimilem; capite rotnndo, ocu-
lis bovillis, naribns depcessis ac patulis, (luos modo contralie-
bat, modo dicbicebat, auriiim loco ntrinquc foramina; rictns
oris rotnndo nee ita \asto, snperiori parte aut labro mystaca
gestabat setis cartilaj>ineis, erassis ac rilane carebat. I'ostica parte
repebat magis quam incedebat. Cute (^rassa, coreacea, pilisque
brevibus ac temiisibus obsita vestiebatur, colore cinereo. Grun-
nitum apri instar edebat, sen crocitabat voce gravi et valida.
Kepebat per aream extia aiutiendo, herumque gestantem cibuni ac offerentem
magno nisu ac gruuuitu accedebat, sequebaturque, nidore ejus
allectus. Lardum ejus gustantibus baud insuave visum est.
* Hfvni tIauM Egi'rtc, Missioniirs mid liischolV's in Groulaml, Ijescliioibung
uutl Xatuv-Gcsfhu'lito von Gninland, iibcrsctzet von D. Joh. Co. Kriinitz.
Jlir Ivii]>ri 111. ]{( rlin, vorlogrs August Mylius, 17G3. i)}). l()(i-10f^.— Since
transrvil'iiig the above I have met with an early (I7(i8) Englitsli transhitiou
of this Avork, in whieh nn English rondoring o ' the above description may
be found at p. 12.'>.
11
ns
()1)( )B.'E\US I?OS.MAHUS — ATLANTIC WALllUS.
■1 ';
(.:'
Coiispii-icbiiiilur il»i(Uiii duo iiuijoiimi ciipitM, ilciitilms duobus
exertis ICl('j)liaiit<>iiiiii iiist;ii'. loii^is ;if cmssis ct iilbiciiiitilni.s
iinmitii. fjtii (Icoi'siiiii \<'rsiis |»fctiis s]»('('t;il>;iiit. iMinuii.coria
cere mil IC ixnulo iicudissc I'ci'cliaiit .\ii,i:li (jiii attidciiiiit.
Ilisct' dt'iitiltii.s rui)('s asccndci'c siMino snstiiicrc ;ij('l»;iiit. rt pro-
dfilii! ill (•(iiitiliclitclii sell tfiTMlii \\\ soimiiiiiii il;i r;i|ii;ilil .^li'j^a-
tiiii. i'iil>iiliii!i ajcliam illis esse lolia nltl()i::^a ac iiiauiia, lu'rhtc
ciijusdaiii t' I'liiidoiiiaii.s nascent is. NVc piscihiis \ iNcrcaut ra.i'ni-
vormii <'ss('. \idi iltidcm pcncin i-Jusdcni aiiiiiialis osst'iini, ro-
tunduiii, i'ultitiiiii ct aiiiplins loii^iini. ciassinii, |)ondoi-(i,sinn ac
solidniii, in tine ]n'(>i)e j^landcMi lon.uc cras.sioieni ac jotundioreni.
llujns pulvcre ad calcidiini jtcllcndinn .Moscovita' rctnntur."*
A still earlier desciij)ti()ii ol'tlie Walrus is juivcn hy Purcliast
in liis account of the first voyage ''into llie North Seas," by
^^'illialll Uari'nts, a Dutch iiavij;atoi', \vli(» met with AN'abnses
oil Oraii.iic Island, in l.?,tt, Iraiislaled from the Dutch b; \y.
Philip. The account says they "'^ went to one «if those Islands
[of OraiigeJ, where th<^y I'ouiid about two hundred Walriisheu,
or Sea-horses, lyinjj,- upon tlie shore to bast tlieiuselves in the
Suune. This Sea-horse is a Avoiulerful strou<;' ]Mouster of the
Sea, uuich biji<>er than an Oxe, which keeps continually in tlu^
Seas, haviiiji' a skin like a Sea-ealfe or Seale, with very short
hayre, mouthed like a Lion, and many times they lye u]»on the
Ice; they are hardly killed unlesse you strike them , just upon
the forehead, it hath foure Feet, but no Eares, and commonly
it hath one or two youiif>- ones at a time. And when the Fisher-
men chance to find them upon a Hake of Ice with their young
ones, sliee easteth her younj;' ones before her into the water, and
then takes them in her Amies and so i)luiiji('th up and downo
■with them, and when slice will revenge her-selfe upon the IJoates,
or make resistance against them, then shee casts her young ones
from her agaiiie, anil with all her force goeth towards \\u' Doate
thinking to overthrow if They have two
teeth sticking out of tluiir mouthes, on each side one, each being
about half an Ell long, and are esteemed to bee good as any
Ivory or EleiHiants teeth, especially in Mil scon via, Tiirtiiria,
and thcreal i •; where they are knowne, for they are as white,
hard, and c\eii as Ivorie."
Skxial Dii'KK1m:>;ci;s. — The subject of sexual dilfciences ni
the Walruses has received very little attention at the hands of
' Nov lis Oil lis sell l)cs( riplid Jmliii' Occident:! lis, |)]i. lis, '.V,), 1(i;i;),
tlli< I'll'^riiiics, \(il, iii, p. ITii.
SKxr.M. ini'i r.i;i:\rKS.
ao
sysl('iiiiiti<' wiilt'i's, wlio li;i\c, imlft'd, no jjositivc iiilni'iiiatidii to
olVci', ;mm1 \<'r\ little can he yh-aiH'd IVoiii other .soiiircs. All that
1 have met \vith,atfeii»retlye\leiisive researt'h.1iasalrea(l\ Iteeii
iiieiileiilally ix'ivvu in the loreyoiii.u' aeeoiiiit of t'le extenial char
Meters. All that (an hi' .^atheredis that in the lemale the tusks
are smaller and thinner, and th<^ ji'enenil size oi' i!ie animal may
lie inleni'd ti> he somewhat smaller than in the mal<-. In tact,
the external eharaeters in the adult animal of t lie speeies under
eonsideiation have never as yet heen j;iven with much detail,
the few naturalists who have met with it in life seemin-; to take
it lor .granted that an aniiaal so long- known, and so familiar to
tliom, nuist he well known, thereby ronderinj;- a careful and d*;-
tailed descript ion unnecessary. The very good descrijttion j;ivon
by Ur. (iilpin (see (intri), ]»]>. '"U, 32, ;?.">) of an adult is about all
that I have met with in the way of detailed descriptions of the
adults of either sex.
The lijiures and descri|)tions given of the young, especially
those rce<'ntlyi»ublished by Dr. J. 3[urie,* leave lit tie to be desired
as regards the external eharaeters in early life. The absence of
references to any strongly marked sexual dift'erenees in the
adult might ])erhaits be taken as negative evidence that none
exist; but on the basis of analogy Avitli the other IMnnipeds,
especially with the OtnriUl(V, we shonI;1 hardly exi»ect their
absence. Even in the case of the skulls, leT sexed specimens
appear to have come under the observation of specialists. Wo
" re and there, however, meet with references to supposed sex-
ual difl'erences in the size and character of the tusks, and also
in respect to the size of the skull and the density and weight of
the bones in those of supposed females as comi)ared with tlK)So
of supposed males. Thus, Wiegmann, in 1832, in referring to
the species described by Fremery, in 1831, says, in remarking
ujwn Fremery's " Trlchcchm Cooliii,''^ that he remeud)ers having
heard from a Greenland traveller that the female Walrus has
longer and slenderer tusks than the male, and states, on the au-
thority of Fremery, that a young specinuMi in the IJoyal Museum
of JloUand, having long, slender tusks, was reganU'd by Tem-
minckas a fenude. IJe als(» considers, (ui the ground of analogy,
that the greater or less develo]>ment of the occipital and other
crests of the skull, as well as the relative weight of the bones,
* " Kfst:nxli(s iqioii tlic Anatomy of tlio riuniitedia.— I'fiit I. < »ii I lie Wal-
rus ('/Vi./Kc/iH.f ivmiittniH, liiim.)." — Trans. Zi)t')l. Soc. Loud., vol. \ii, 1>7'.*. pp.
411-1(11, witji wdoilciits, and iilatus li-lv.
■M
i
iiE
M
If*
)■
40
(•DOIS.KNIS IJOSMAKUS ATLANTIC WAMJIS.
'i ■
( i.
1(» be only (litlrrciiccs of a scvual cliaractcr.* Staimiiis.f icii
years later, citcil the views >.. reiiiiniiick ami \Vi(L;iiiaiiii (as
above .uiveii) resi»ee(iiij;' sexual «lil]'ereiu'es iu Walruses, but adds
nothing' u«'\v to the subject. Laniont (see (inlci'i. \>. ."!.">) states
that the ''tusks vary very nuu-h in si/e and slia|H' aeeoidiiiii to
the a^'e and sex of the animal." "(Jows' tusks,'' he says, ••will
areraije fully as lonji' as bulls', Irom beiuj^' less liable to l»e broken,
but they are seldom more than twenty inches lonu and three
pounds each in weight. They are j^i'nerally set nuich closer
together than the bull's tusks, sometime o\('rhii)i»inji- at tlio
points, us in the case with the stuftcd specimen at the I>ritish
Museum." lie gives the UMigth of tusks in the male as 2t inches,
aud the weight as i jjounds each.
A skekton, marked as that of a female, in the Museum of
Comparative Zoology, coHeeted in the Greenland seas by ])r.
Kane, has the bones very light, soft, and i)orous, as coni]»ared
with those of male s]>eelmens. The skull (see tigg. l-;») is nuich
smaller, with the (;rests and ri'('lnod(')ilicit sciii hoIIIc. mil Aiialofricii beU'gt werden." — Arch, fur
Xafiirficsfh., 18:]2, pii. l'^-', I'J!*.
tMiillcr'sArcli. fiir. Aiiat., 1^11, j.. :«>•,'.
Sr.XIAI. DIFl'KRKNCKS.
41
ttisks (see li:r. 1) is iloiibllcss iihiioriiiiil. nml is doiilttl<'ss owiiii;-
to their iiiisyiiiiiictviciil ared imnnrous specimens of lioth sexes, whicii were lost v.iili the ill-
fated vessel.
till theXalioual Miiseinu there is also a female skull of (he I'aciii'' \\ alms
that ])rese)its corresiiondin}; diHerences as compared with male skulls of the
same Hpceies.
I
VARIATIONS DEPENDENT UPON AGE, ETC.
43
coiijecturo.s of Wk'giiuuiu ami Tcumiinck. What othor dilier-
cucc'S obtain, especially in external (^Isaiactois, can as yet bo
only conjectured. It i.s to be hoped, lio\v<'ver, that wc shall
not have lony to wait for detailed accounts of the external
characters of the adults of both sexes.
Fio, 'X — Odoha'uiifi ronmarus, 9 .
Individual Vaeiatio>>!, and ^'Al{IA^IONs dependent
UPON Age. — That a wide rauye of individual variation obtains
in this si)ecios is sufflcicntly evident from an examination of
even a limited series of skulls. These dilierences have been
noted in (.'onsiderable dcitail by Fremcry, Wicgniann, Stannius,
and Jaeger, as will be presently noticed more in detail iu ju'e-
senting the j^enend histoiy of the species. Still greater difler-
ences, of course, result from dilierences of age. These collect-
ively, as A\ill be noted later, have formed the basis of several
nominal species. All the Pinnipeds appear to be subject to a
wide range of variations of thischaraitter, and none more so than
the AValruses. Tliese atVect to a considerable extent tlie general
l)roportions of the skull, and especially tlie form and relative
development of different I)ones. Tliese latter dilierences are
best seen in eom})arativfly young skulls, since most of the
sutures close at a ratliei" early age. Among tliese variations
are especially noteworthy those of the nasal bones, the inter-
maxillaries, and the frontals, and to a les.-. degree those of the
base of the skull. Tlu' crests and ridges for nuis. 20.
\.VliIATK>N.S DKl'ENDKNT UPON ACJE, KTC.
45
TIm! iVoiitiils Viiry * Z 0^ <>*
1 '<
1-
o
il « : S f2
•qjSnoi '.ttiifao.ttoi
•)JV(Ib 09aC|8Il> 'BJ0ilI3U| J,)dd£],
•j«ioni yiv\ IB o)BiBd JO mPIAV
O rt g -T • O M
?1 ?i CI H ■ Si cI
s s s s s
•8ojao8 iiiaojt.iBium .loddn jo i(^noi
I*; f5 C5 XI 1^
« ci .- S »
•sdj} IB ?,iB(Iu ojun^Bip 'KjuinBO
•oBtiq )B K.iSpa jBn
aojxa uooAvj"! ojubjbii) 'souiub,')
S 3 fi S fS ?? S
HI
MSBq IB ^i.nMi.i|iMii.).ii.) 'K.mniiQ | J,'
-(SJCIOOI
JO onc[)Bll!lI JO pu) .loi.wjsod
O) Q!ii!XBnu.>(nt jd.i.ip.ioq .wi.wjuv'
Tt h. ja e = X t-
S ?1 H fi ? S
^5 s fS X 1^ M I-
I- !• is {I ri I- »
*H -H r-< fj ffl 1^ M
•.\I.iou.itm! ipi'lAv 's niiH| n:ni:.>v
•.\'t.xoiaojS()4l uipiAV 'Kouofj iiisBX
•i[iSaoi '8.>no(i iiisBX
I vussoj
I iBiodmo} uoD.ttjoq
CO fS
•XOg Of O O \3 \3 O "to
r', a O O O o O
^ .3 -3 -a '53 -a "TS
•jaqninn ouSoxbjbo
'?5 -I
« « «
to©
5 ^ 5 t-< •«
rt » O
lias
DKNTITION.
47
I)i",.NriTif»N. — The (It'iititioii of Wiilniscs, for \iirioii.s rcu-sons,
lias hccii :i |M'i|ilt'.\iii;; siilijcci, and lias cii^a^ii'd tlif attoiitioii
of iiiaii\ niiiiifiit wiitcrs. In tlic adult stajnc it pivsi'iits many
altiioniialitics, and, ln'sidcs, issuhject to nmcli individual varia-
tion. Iiotli in tin- tnnporarv and adult series. For a Ion;;' time
its deviations from tli«' normal type were not well untler.stood,
and even now leadin;;' authorities do not appear to be roi)er notation. A.s previously
staled, iIm' incisors of both Jaws, except the outer i)air in the
upper, disa]>pear soon after birth, and before middle life is
reached the last tttotli of the molar series on each sule iu both
jaws also usually disapjiears. A brief history of the princii)al
iiiv<'sti^'ations. and opinions held at different times resi)ectiug
thetlentition of llie Atlantic \Valrus (for the investigations
resjicctin;;' the dentition of this ^roup appear to have been based
almost wholly uiton this species), is herewith appended as form-
ing a lii;.thly interesting chajjter in the technical history of the
species. Ill this historical sketch will lie found noted many
facts relating to the general subject, given by the authors whose
papers ai'<' here biiclly summarized.
The dentition of tlie Atlantic Walru.; has bt'cii discussed in
greater or less detail by iJai)p, von l.aer, Wiegmaun, Frcmery,
k>tannius, .lacgcr, ( )weii. Mahiigreii, Peters, and various other
writers. NViegmanii, in IS.'JS, pretty fully presented the early
history of the subject, noting the almost total lack of informa-
tion respecting the matter shown by Linne, who evidently paid
little attention t(> the references to the subject made by previ-
ous writers. The credit of first giving any definite statement
respecting the nnnd)er of the teeth and their character is due
to Anderson, who, in 1734, gave the niunber of molars as four
above and three below. Brisson, in IToG, gave the number as
four both above and below; while Crantz, in 170,'), again gives
foiu' above and three below, and ipiite fairly describes the nor-
mal dentition of the adult.* In the same year, Daubenton gave
also again four below on each side as well as four above.
' I iii>pt'n(l iu lull Ci'uutz's description:
'• It liail no sliiiip («ci\s<)/T« in its mouth, anil noiio at all liifoiv, Init only
four tcotli on I'adi side ; on tlierii^lit siilcol" tlii" nntltT-jaw three pretty hroad
eoncave jrrindors The two lonj^ t usks or horns grn\viii out of
its laee aliove the nose, and liendiuft down over its mouth, so as almost to
barricade it nil. sei'nr to h(> more an imiiodinient than a liel]) to It
Tlierijht tusk is aliont an inch lon<;er than the left, and its wliole lengtli
' -HI
m\\
• 'I
>
C. f
I 1' :,
''..
:M
4H (Mum.KMS I.'OSMAIMS— Ari.AM'lC WALias.
No ;intli<>r ]niof to Sclirchci- (177"i) iippcars to li;i\(' iiicr •vith
(Iccidiioiis iiici.voi's. \\Im» I'omul l\V(» such ii|»|»i'i' iiicistus on ciu'li
si.s'.S7( iiifcrnKt.rilhirid (W^ Wiiliiosses mid (h'r Sehlieide/.iihiie
i || ' dessellu'ii liiit keiiiieii ieiiieii, iiideiii er iiiir cine vortiefllielie
Abhiindhiiiji' iiiit sehruieii Zeiehiimi.u'eii dieser Kiioeheii \fr-
sc'hiediMier Thieve zii^escliickt li;itte.'' Caiiipev, in eritieisin.i;'
Limio's errors rejiiirdiiijn' the Wiilnis,t K'ves four imisois (,^^,)'
aud four iiiohirs altove and livt^ lielow (-,Z-,) ^*'^' soiueliiiies only
four below). Tlu^ observations of Sehreber, (loethe. and Taiii-
j , per ap])ear to luive been ucneiaily oxcilooked by siibsiMineiit
writers, so that it was h'ft for (r. Ciivier to iliseover anew the
liresenee of deeiibious ineisors in the yoniifi" Wahiis. lietweeu
the canines he recoj;nized two incisors similar t.> the molars,
which he says the majority of observers had overlooked, because
they are not lixed in the interiiiaxillaiy, and between these
again two i)ointen alleni Anselien
naeh auserhalb dem Zahnlieische nicht zn bemerkeu seyn, /umal da sie nicht
am Rande der obern Kinnlade, sondern mehr hineinwiirts stehen. Ich linthi
sie an einem zur Natnraliensanindnng hiesiger Universitilt gehorigen Wall-
rossschiidel ; xmd da derselbe, besage seiner Grosse, vcm einem jungen
Thierc ist ; so glaube ich beynalie gar, dass si(^ bey zunehiiiendem Alter des
Thieres ausfallen nnd nicht wieder wachsen. Sie konimen also hier in keine
weitero Betrachtung, als d.iss sie dem Systeniatiker (;inen Wink geben, dis
Thier nicht zu weit von dem Robbengeschleehte zn entfenien." — Saugcthiere,
Th. ii, p. 2C0.
tl qtiote the French edition of C.'miper's works ((Eu^Tes, torn, ii, p. 480,
Paris, 1S0I>), the only one accessible to me.
4M
KHNTITION.
40
Inwcrjiiw.* I''. (Juvicr jiiivc latt'i'tilsu the siiiiic dciitiil loninila.
IIcdcciiM'd tliiit tlic peculiar or imoiiiiilons deiititioii of the Wal-
ruses indicated that tiiey were ait isohited ^inaip, lutviii;^ atliui-
ties. on tlu' (»ne iiaud, with the Carnivura, and, on the other,
witli the Ikiuninantslt
Accordin.i; to \Vie;uniann, I'lidolpliil (in ISOL*) reco;;ni/ed
liie lirst ofliie series ui' loircr ^rindin^' teetii as a canine. 5
Thus, as \Viej;niann lon;^' sin<'e oliserved, liu' suhject reniairu-d
till Ifapp was so I'ortuuate, iu ISiiS, as to Inive opportunity to
exanuue a tuMal specinu'ii. in this example, he Ibund six inci-
sors in the upper Jaw and live in the lower (ij^^)- He also
expressed it as his heliet' that the lirst low«'r molar should he
rej;arded as a canine, hecause (1) it was somewhat further
removed from the rest than tlu; others wore frcuu each <»t her;
because (li) of its j;reater length and thickness in the adult ani-
mal: hccause (.'{) it stands close to the temporary ov nnlk inci-
sors, and shuts aj>ainst the outernu)st of the ui)per incis(»rs;
and hecause ( t) it lacks the transverse de])ressiou seen on the
inu'-r side of the crown of the back-teeth. The dental '')rmula
recojiuized hv him for the Walrus may be considered as
1.
I-'renu'ry. in 1S,")1, also made reference to the dentition of the
Walruses; hut his paper bears mainly upon the question of
whether there are one or more species of these animals, and
will be further noticed in another connection. He notes partic-
ularly the presence, in some of his skulls, of two small molars
abo\ e, behind the largo ones.
Wieginann,|| in 1838, contributed facts additional to those
already recorded, but his memoir is largely devoted to a discus-
sion of the obsen-ations of preceding Avriters. He assents to
Rudolphi's and Kapp's interpretation of the honiologieal rela-
tion of the lirst large tooth of the lower jaw; refers to finding
' Ki'jfiio Aniiual, toiii. i. 1817. p. 1G8.
t Dents (li'sMiiiii., p. -23^.
tAiiiitoini;>cli-itliysiolojfis(li(> Abliaiulluugi'ii, p. 145.
vNVVifjfiiiaimsiiy.s: ". . . . UelJcnVu's ist liiiddlplii dcrorstr, dordic iintorca
1"< lli of six incisors l)<)(ii iihovf^ iiiid below, iuul to
tlic (-.irly (Iccidiioiis ciiiiractcr of tin; last (liltli) upper molar,
and I lie rreipicnt disappearancui t/f Mic loiirMi. lie ('(/ncludes
thill I lie norniiil number of the back-teeth is ^~-' and lliat in
4 — 'I'
earl.N lile the dent it ion of the Wall lis is not \vi(h'ly ditl'eicnt from
that of other Pinnipeds.
'I'he same year (l.S.'{.S), Mae;;illivray* eonsi(h'red the normal
dentition of the Walrus to l)e \.'^.', <'• ',1^; TJii. + M. ^-^'
= j^ = ;;:;. ills conclusion was based on th(^ examination of a
])er Jaw there are <>n each side tliiee imisors, the first
oi- inner extremely small, the seciond a little lari^cr, the third or
oiiltr disj>ro|>or(i()nately iar;;e, beiii;; e((iial to the laij^cr .yiind-
eis. The socket of this tooth is placed in tiie iiiterniaxillar\
bone, but towaids its mouth it is|»ai'lly i'oniied Ity the maxillaiy.
The small incisors ha\e deej) conical sock<'ts. The canine"
tooth is disi)Iaccd, beiiij;' thrust outwards licyoiid the line of the
other teeth, and caiisin.in' the peculiar l)ulj;in;;' of tlic head. The
lateral iiuMsor is on llic IcncMiI' its aiitcrioi' liiai.niii. and the
first urindcr is o|»posite to its middle, 'i'liere arc ti\e j^rindcrs.
]ia\iii,u ccoiid, and third nearly c(pial.
the latter beinu' a little larger, the fourth much smaller, and
the IllVli vciy small, flic tusks, or enormoiisls de\cl(»ped
canine teeth <»f the upper jaw, arc comiu'csscd, <"oiiicai. a little
curved backward, directed dow nwards and a little forwards, and
somewhat ili\cr;4in,i;-, but in some iiidi\ idiials, when \ery loiri;.
they ajiain coii\(r;.:e towards the points. In adults, lhcincisor>
are obliteiatetl, exceptiii!.; the lateral pair of the up|)er jaw; the
fifth jj,rlndcr in liotli Jaws has al-o disappeared, and sometimes
the fourth in one or both Jaws."
■ iiriti.'dniiuidniiicds, JsiJH, j.p. -^Jl i, -J-J 1 .
if
;(;;!
DKNTITION.
51
StiUiiiiii.s. ill ISl'J, riiiUicr coiitrilmti'd to llic siilijrct hy
suldiii;; <»i).s(a'v;ili<)ii.s icspccliii^ Viirinlioii i'l the iiiuiiIh'I' of
t\H\ l(M>tli icsiiltiii^' IVoiii ;i;;(', ' in di'tail llic incisive
driitilioii ol" ii sciics of four skulls of (liU'erciit, )i|?«'S. In two
youii^' skulls, tln! outer tcin|)onii',v incisor of the iipiHT Jaw on
either si(l(5 icmainod; tlu5 alveoli of IIk; s;'cond pair were still
distinct, while the alveoli of tiie iiddd' : paii' weit^ nearly oblit-
erated. In another, the alveoli of the, inin'r pair of ineisorH were
wholly obliterated; those of the second pair wen; baiely re(!Og-
nizalih', while those; of the outer j>air wer»} distinct, the teeth
havin^i' fallen later. He was also able lo i'eeoj:;nize the alveoli
of six incisors in the lowei- Jaws of tin; skulls Just mentioned,
and states that Ik; lhouj;ht Kapp's view of the homology of the
first lower bac k-tooth (considered as a canine) was jtrobably
correct. He liirthei- takes exceptions lo the value of tin; char-
acters assumed by I'^remery as the l»asis of several species of
\Valriis(\s.* lie adds, in respect to the lusks, that in old
a.nc they become \vlioll,\ solid to the base. In regard to the
ii|)pcr molars, lie notes thepicseiice of live in sev<'ral instaiicciS,
and liiids tiiat, as a rule, the fourth disappears before the fifth,
or, at least, that, its alveolus becomes s(»oner obiilerate(l. llo
also ctudirms tiie siMtemeiit jucviously made by Wiegmann,
that th(! alveoli becoiin! lilled by depositions of bony iimtler in
concentric layers at the bottom and on tin; sid( s.
''I'lirsc ;illc)j;c(l sprcilic, cliiO'dclrrs ln>, not ices in dotail, jinil coiKsititTH thorn
ii.s (l('|icii(li'iii, iijiim ji;;c. lie, says: " iJiiH Mis.siiclMi (liener Clianiktcro <,'ilu'lll
KclioM iiiiK(l('iiilJiiis)aii(lt',tiawH(ii('.s('lln'ii iiiir riirv(illi;r aiiHyrwachseiic'J'liicre,
iiiciit abcr I'iir Jniij^c! uiiwciidliai' Hind, dcnn das Htiiik.sto Waclistliiim der
Eckziilnio fall), erst, (>ti'cid)ar in cino Hpiitero Lcbciispdriodi) in widclier
naniciitlicli dii', Ix'idcn iiincrstcn Sclincidcziilinti nnddio bcidcn lolztcn iJack-
/.iibno j(!diT HciU; drx oljcrcn ICinnlado schon j^escliwundcii Hind. Uiorzu
kouuiit noch dor Unintand, dass aiuh din ICok/illino Ixii sehr alton 'i'liioreu
an del- Spitzo ItodcnfcnJ al);;('nnizt; .sii.il, dcnniacli in siiiittTon Lclx-nsHta-
tlicn an Jiiin;^o waLrschcinlich wicdor abnclinicn. Endlidi Hclicint cs ja
scllist, alsol) dio Liiiifjo dioscr Ziihno jo iiach don Gcsolilrclitorn vorscliioden
wiiri'.
" i^lioii so wcni;;- (i('\vi(lii nuiclito ich util" di(i l"urclinn;,;on dicscr Ziihno
Ic^iin. llinr Zaid, wic iiinr fStiirko uach Hind Hio ))oi vorschiodonon iibri-
Hiiis niijif \()ii cinaMiloi' aliU(:icliii<;, lie notes lliiit tlie l:ist m|i|mt iiioliii' Itiid
already fallen tVoin the ri^lit side, hut still remained on tlio
left, belli lid wliieli, as well as heliind the ah et tins of the fourth
iiiohir on the ri<;ht side, was a little shallow pit, in wliieli, diir-
irij; fd'tal life, a tooth hiid perhaps stood. In front :A' these
heforeiiientioiied teeth were three molars on eaeli side, and in
front of these a eonieal iiieisoi, and the alxcoli of the others
were traceable, altlioii;4h alieiid.v filled with a spongy substance.
f the lower jaw, there were li\(' teeth on ea<'li side, with traces
of three already lalleii ludal incisors on one side and of two on
tli(! other, '{'he second skull (!>•/ inches by Trj) was somewhat
older, the canines bein^' about live inches lony. There were
present in this skull three ii|>per molars cui e:icli side, and a
lilled-wp alveolus behind them. Of tlu'se te<'tli fas also in tin;
other skull), tlu^ middle one wu.s the lar^^est and most worn.t
In front of these, and somewhat distant from them (')'"), was an
incisoron (;ach side, and in addition to these another jiair of small
c — .'{; .lae^icr thus
recoj;ni/in^, as had Ifudolphi, Uapp, \Viej;iiiann, Fj'emery, and
Stannius, the lirst of the; low«'r-Jaw series as a canine. Iteliind
these were 1 races of lli«s alveoli of the fourth jiair of molars. In
the third skidl (leii«rth 12.\ inches, breadth lOA), still older, with
tusks a foot lonj;', wen^ three u]»[»er "baek-liMith " on each side,
close to;;('tlier, the middh; one beinj; the lar;;est, and in frontof
these a eultinj;-tooiii. The low<'r Jaw had also lour teeth on
each side, homiil\ in IJH' iipprr liiiiii I lie lowci-Jiiw : ;iii(l
lliiil. iiiiHc i;ii<'j\ , ;i siii;ill InoIJi \\;is siiiM'imlflcdbcliiiKl I lie nor
mill loiii-, ill llic upper jiiw, iiiid still inure rniely in Hie lower
j;i\\ ; llie roriniilii -A' tlie ileiitilion of siieli \iirielies, in exeess,
iH.in-.— i.r,-:, <■•
'ni.
;;, M,
I —I
1—1
= LM»." < hveli liereinnkes.
no relereiiee totlie lileinl lire of llic, sii'oject, aiid evldcntl.y ^avc-
a very erroneous inlerprelnlion ol' tiic; denial loriiiida. In Ids
■atei' relereiiees lo llie siihjecl lie, ^ives an enliiely diHereiit in-
terpretation, and one more, nearly af^reeiiij;' with lliat now <;oin-
monly a(M;ei>led. In his lalesi i«!leicn(U', lo llie, suhjecl,* iitj
writes: " In Hie W a!iiis(7V/(7/a7i».s' rfmnariin) \hv normal incisive
roriiiiila is I ransitorily represented in Hie veiy yoiin;;' animal,
wliieli lias three teeth in each i)rema\illary and I \v(» on each
side of the lore part of the lower jaw; Iheysoon disappeai'
except the outer pair above, which rcmaiiM'htse, to the maxil-
lary suture, on the inner side of Hie sockets of Hie eiiormoii.-,
canines, and commence the series of small and simple, niolus
which they resemhie in size and form. In the adult. Hien^ iMe,
usually three such molars on c k h side, hehiiid the permaiienf
incisor, and four similar teeth on each side, of the lowei' jaw;
the anterior one passing' into the intcrsiiace. between the npi»er
incisor and the lirst molar Tin; canii'es iii'e of enormous
si/e Their hom()lyp(i below I'clain.s Hicsizoan.l sliajte of
tile siiceeedin;;' molars," Tlit^ formulii of Hie normal dentition
a])parciitly hove re('o;;nize Ihe lower Jaw, the lirst permanent looHi is
I'c^iardeil as a canine, because it is thicker and rounder than tluj
jtostciior leelli, and lacks tiie eross-furrow that marks the oth-
.\ii,ilinii\ 1(1" Vi'i'ti^hriitcs, vol. iii, y. ',V.W.
tOilniil,.-., 1,. M-J, |,1. :!(;, lijr. 7; Siill}irl ll., p. !v!;>.
^riirs iiic, iiowcvi!!', US mIiiiwii liy Miiliiigrcii (kim^ licyoiid), incccdcd in tho
riiii)iy() liy (cinporary tccili.
;l
li-
hn
■ V
54
( MM tU.KXrs K( )S.MAUrS ATI-ANTIC WALUUS.
; !
ors. The \(miij;aiiiiiia! \^i^ live iipiici iii()liiiil\»i'iii teeth (•' llaclc-
zslline''), tlie lii.st t\v'll dentition: 1. J^J, C. \-\, M. :-;. While
(iieliel ae('e]»ts the first ])ernianent tooth of the in<>lariforiM s<'rie.s
(if the Mi»i>er jaw as an ineisor, and the lirst in the lower Jav as
a canine, he recoj-nizes only two iiersisteid nioLirs on each side,
above and below.
Malm.uren,* in ISG4, ti;.>ured the dentition from a Iti'tal s]K»ci-
men, and i»nblished an elaborate i»a]»er on the deidition of the
Walrns, in which he leviewed at some lenjith the history of the
snbject, notieinjj;- (^nite fully the writings of the early authors,
from ( 'rautz to the Ciiviors, and the papers of liapii, Owen,
Wiegmaun, Xilsson, and other later writers. The fonnula he
I)resonts as that of the peraumcnt dentition is : I. ^-^J, C. ~\,
M. !|5J|=^=18; and for the deciduous dentition: 1. 1~^, 0. ]^\,
M,
4—4 Hi
J" 'JO +
4 — 4~Iii
The specimen figured shows both the permanent and deeidu-
ons dentition. The deciduous teeth are most of them separately
lignred, of natural size, as minnte, slender, spindle-i'ooted teeth,
with short, thi(;kened crowns. The permanent teeth are already
in place, altlumgh even the n]»per <'anines liad inobably not
pierced the gum. The middle i»air of incisors of both jaws had
ah'eady disappeared, leading only their distinctly recogiuzablo
alveoli. His specimen api)ears to have liad bnt a single cadn-
cous molarbehind the permanent series, Iron Avhich he assumes
the nnniber of upper molars to be 4 — 4 insteid of 5 — .").
The following year, Peters f referred to Malmgren's paper,
jiublishing a jdate illustrating the dentition as existing in a some-
what older skull (received from Labrador) than that tignred by
Malmgren. I'eters here takes e.\ce])tion to Malmgren's assumed
numberofba'.'k-teeth, which, in accordance with the views of Kapp
and Wiegmaun, Veters believed shoidd be I'—.', instead of j~i
>^ . i) — u' 4 — 4
•M»rvcrsiy;t ill' Kiiii.ul. \'(t.-Ak;iil. I'orlmmll., It^O.i, i)]). r)(lo-r)-J:.', jil. vii.
■ tTln' ii;i|M T liiMim |>iililislifit ill Swcdisli. I am iiiuihli' to I'ollnw liiiii in his
llisCllSsioll (if lIlC SlllljlMl .
t M(iiial>l). I\. P. Akail., l-^'i."), \)]>. (!t!r)-(i87, id. faciny p. Cc^.").
u
■■■i
k
DENTITION.
;JD
t'ii
According to lA-U-rs, .M;il!ii,i,'i\'ii, froui not liuditi.^' niorc than
I'oiu' iii>[t('r hiick Ici'tli ill any of I lit' many skulls of \arious aj;os
lie liiid cxaniiiicd, concluded lliat wjicn a lifth is present it is
aliiioriaal. TIk' youn}^' skull li,uui'ed and desciihed l)y P»;ters,
JHtwcvcr, lias in the ui)]»ei' Jaw tlie I'onrth and liltli hack-teeth
still in place on tiie lij^lit sidi', and the t'ourtli on the left side,
with an aheolusot'a liltli. This Peters considered as allording
new proof of the correctness of Wiogniainfs formula. As
already noticed, live molars have been recognized l»y I'remery,
l^app, (iiehel, and Owen, and, though perhaps not always pres-
ent, are frequently to be met with.
The dental formuhi of the Wabus, as determhied by liapp
and AVieginauu, hivs l)eeu adopted by Van der Iloeveu * and
Blasius, f as well as by Peters, and essentially by (liebel. Gie-
bel, however, gives oidy four deciduous lower incisors, instead
of six. Owen, in his later works, agrees in this point with Gie-
bcl, but takes apparently no cognizance of the deciduous fourth
and fifth molars, to which he refers, however, in Ids earlier
papers.
Gray, | in 18G(>, althouuh quothig the formida given by
Eapp, 'vlo ^s the folk-vlug: "Cutting teeth ^ in young, I in
adult; grniders —^ in adult, truncated, all single-rooted; ca-
nines, upper very large, exserted." He, however, quotes Iiapi)'s
fonnula, and also that given by Owen in his " Catalogue of the
Osteological Series of the IMuseum of the Eoyal College of Sur-
geons" (1853, p. 030).
Professor Flower, § in 18G0, gave a diagraui of the dentition of
the Walrus based on many observations made l>y himself and
on "those of others, especially' Professor Malmgren,'' in which
both the temporary and permanent dentition is indi, p. 738, Eutjlisli cd.
\ Siiiiy;i'tlu(i'e Di'iUst'lilaiidw, l>i7u, iip. 2G1, :i0'2,
I Cat. Seals and Wliaks, p. :35.
vN .IiiMin. Aiiat. and Pliys., iii, ]). 27'2.
.1
■mm
m
' .")<) oltoli.KNl S HOSMAHIS AI'LANTIC WAI.IM'S.
,,' , ; UIifi'cqiK'iitly pfl'sistciil to cxd'ciiic old iiyf, iilllioii.uli ('(Hiiliiolily
,, ' losi ill inacciMtcd skulls. TIk'si- iii(liiiiciiliii',v tcctli iirc iisiuilly
' ii dosciilicd as ' iiiilk-tcctli '; (ivcn llic postciior ones arc sonic-
times so called, luit it apjsoars to iiic an open (lucstion wlicllicr
tlio\ rcscnt pennaiiciit teeth in a nidiinciitary
I or aborted condition."
, J Huxley, ill liis "AnatoHiN of \'ertcl>rated Animals" (pp. I'A'A),
j . i 301), piiltlislied in l.sTl', ado])ts tlie f'ollowinji' as the dental for-
;; ' ; mulaofthe Walruses: "I. ,'-^, ('. J-[, i>.ni. in. :;-;!+ ;::^V' lie
I C ! says: >• Tlu^ dentiTioii of the Walrus is extremely peculiar. In
\\.< the adult, there is one simple conical tooth in the out«'r part of
I : the i)remaxilla, followed by a huge tusk-like canine, and three,
short, simple-fanji'od teeth. Sometimes, t\V(» other teeth, which
' ' i soon fall out, lie behind these, on each sid«' of the upper Jaw. In
, the mandible theie are no incisors, but a sin;;le slntrt canine is
' 1 ; followed by three similar, simple teeth, and l)y one other, which
I is caducous." * Txdh here and in the formula no reference is
made to the deciduous incisors, althouj;li the caducous molars
are recognized.
In the fore^oiiiii risitmr, we have seen how va^tui Avas the
information bcaiiii.u' on this .viibj;'t possessed l)y all writers
I, l)i-ior to altout tile Ix'^iiniiiiiu' of the present ceiitiii\ : how the
jji ' ' earlier notices of the existence of iiicisois in i lie noiiiii:' were
overlooked and rediscoxcrcd l»y hiter writers, as well as how
' slowly the tirst periiianeii', tooth of the molariform series of the
ui>per Jaw came to be ^('neially reco^iiixed as a true incisor and
not a molar; how, later, the numi)er of incisors in theyonn^was
fouiul to be six in tlii' upper Jaw and six in the lower Jaw, with,
as a rule, two small caducous molars on each side in the upper
jaw, and one on each side in the lower behind the i>ermanent
' . grindinj;' teeth ; that the tirst permanent molariform tooth of
the lower Jaw was a canine and not a molar; and that l>y dif-
ferent writers the inimber of incisors i-eco^nized in the lower
jaw has been sometimes four and sometimes six, and the cadu-
cous upper mohirs re<;arded sonK'tiii:"s a.s one and sometimes
as two. Finally, that the true formula of the full dentition of
the Walrus is I. i!^!*; C. \-\; I'm. M. ]-;;=l;?=;U. It lience
appears that the dentition of the Walruses is peculiar and some-
what abnormal in four featui'cs, namely, (1) the early disap-
pearance of all the incisors except the outer pair of the upper
* AiiMt. N'crti'ln-. Aiiiiii., \)\k ;?(jO, ;?G1.
Hfc^ffl
FOSSIL IIKMAINS.
i)l
jaw, (L') llic ('iionnoiis (l('V<'lo]iiin'iit of fluMiiipcr caiiincs. (.'») tlic
sli.uiil spcriali/.alinii of l lie liiwt'i' caiiiiics, aiitl (I) t he <'a(!iic()!is
ciiaiiictci' III' till' two postciiur |iaiis nl' molars ol' 1 hi' iipiu r jaw
anil till' itostcrioi' |)air in tlic litwi r jaw. 'I'lir ciirly (Iciititidii of
tln' Walrus (lillcrs mainly iVom tli.il ol' iiiost ullici' riiiniprils in
liaviH,u six lower incisors instead of lonr. I lit' incisi\»' Ibi'Miiila ol'
otJMT Pinnipuls, as^ciicially I't'coj^uizcd, Itcinj;' usually v^ !. IVc-
qucnlly i;"!;, and sonu'tiMics (as in Mocrorli i mis iuu\ t'l/stopliura)
•-^-, — never, at least in the i>einiaiient dentition, ^-~;|, but I am
far Iroiii sure this number may not sometimes appear in the
deciduous dentition, in the Sea Otter {Knlii/dris), there are
said to be six lower incisors in the yonn^i', wliile only four are
present in adult lile. The middle i)air of h)wer incisors so early
disai>]tear that e\cn in very yonny specimens they are some-
f'lnes wanting;. Ifapj) found in a f(etal s])e('imen tliree on one
side and only two on the other, and ([uite a inimber of ]>romi-
nenl writers on the subject have reco^iiii/ed two j>aii's of lower
incisois as the normal number. In many specimens, the alveoli
of three pairs have been found, and, in addition to tin? instances
already jiiven, 1 may ad '. that theic is a younu' skull in the
Museum of ('om])arative /o(lloi;y lliat shows decided traces of
thri'c paiis. the outer incisor on one side bein^■ still in j>lace.
In view of all that is at jtresent known respectinj^' the sid)-
ject, 1 a(h)pt the followinji formula; .is beini;- well-established, —
premisinji', however, that they are substantially in accord with
the view of the ease presented by Professor Flower in 18(I!» : —
Temporary dentition: 1. i|i;|; <'.JeI; M. i5^ = ||; = ;w.
Permanent dentition : I. i-;^-; t\}5-|; M. •^^^=JJ = 2G; the
last two upper molars and the last lower one on each side
being rudimentary and often absent.
Fossil Re3IAIx\S.— llemahis of the Atlantic Walrus, in a fossil
state, have been fonnd at various points along the Atlantic
coast li'om Elaine to South Carolina, and in Europe as far south
as l-lngland and France. The first noticed front American locali-
ties was thus mentioned by Barton in 1S05, but the locality
is not given, lie says: "The bones of one of these large
animals have been foun»l. Tines*? ap])ear to have belonged to
as]»ecies <»f trichcchns; j)erhaps to the triclicchun rosinarus or
?Ho/-.sc."* Messrs. Mitchill, Smith, and Cooper described, hi
*Liiii(l(.ii Phil. Mill;., vol. .\xxii, !->()'>, p. Dd
Mm
»aJU,>J*-
I
i:'
oS olxiB.EMS IJOSMAIiCs — ATLANTIC WALKIS.
1tfMllv fcftTicil it: hut hitcr it wiis rc^nrdcd 1>\ l)fi\ii.\
Jl-S r('i»i'('S('ntiii;n' ii distinct species, to wliicli lie ii;i\e liie n;i:iie
Trlcli cell IIS ririiiiiiiiiiiis.\ In iSlI, l,,veil described a tusk ul»
tainod from the Tertiary Clays of (iardiner. Maine, wliicli
Owon reii'ardod as pi'ol»al»l\ belonijiHiU' to an extinct s|»ecies.§
Lyellll also refers to a skidl lie ohtained at .Martha's X'ineyard,
Massaeliiisetts. Ilcdcseiihes tliis skull as >• lilferin.L;fr(Mn skulls
of the cxistiiiij;' species {Tricliivliii.s rosmiini.s, IJnn.). with wliich
it was eoiiipaicd hy Professor Owen, in lia\ in,u' only six molars
and tw(» tusks, whereas those of the recent lia\ ti four molars on
each side, besides occasionally a rudimentary one. The front
' I tusk is rounder than that of the recent walrns."<|
In 1S."»7, Dr. Leidy** described and li^iired a skidl found on
the sea-beach at Lonj;' l>ranch, 3Iouuiouth Couidy, New .Jersey,
where it was (»btain»'d by Prof. .1. V. l''razer in IS."),'), and refers
to another specimen (consistin;j;' of the facial portion of a skull)
J discovered at the same locality by Prof. (Jeo. II. Cook. The
first-nanK'd specimen, says Dr. Leidy, ''has lost a portion of the
craniuui i>roi)er, and the exserted ixution of one tusk, but other-
Avise, except beiny a little water-worn, is in a ^ood state of
preservation. It is luichan^^ed in texture, and nearly so in
colour; and it helonyed to an old individual, as all the sutures
are com])letely obliterated. The form of the facial ]>ortion of
this specimen corresponds with that of the specimen from Vir-
ginia, [described by JJeKay and precediuji' Avriters,J abox'e men-
tioned ; and the entire skull closely resembles that of the recent
Walrus, TrichccliiiN rosmai'iis, as represented in Ihi^ li<;ures of
Danbenton, (.'iivier, and De lilainviUe ; and its measurements
are also sufllciently near those j^iven by the lirst-named auth(»r
to recoj^nize it as the same s]iecies.
* Anil. bye. \:it. Hist. New Ycuk, vol. ii, Is^S. j.. -JTl.
tEdiuli. New I'iiil. .loiirii., vol. xvii, l-^IM. p. :!C.(l.
tNat. Hist. Xfw Yoik, Zoiilof^ty, \>t. i, t>'l'J, [p. aCi, |il. \ix, (ins. t, I'lH-kiiitl. .A[. 'ilC.
IIAincr. .(.lurii. Sci. and Arts. vol. xlvi, IsM, )>. oil*.
^1 As is well known, t lie cxistiiij;' Wah'is lias occasionally only I lie imiiilni
oftoctli foiiiid in tlic .Marllia"s Vini'yard siu'cinicn.
**Traiis. .Via. I'liil. Soc, \(d. xl. \. -:>. ids. i\, \, li^-. 1.
m4il
"I,
FOSSIL REMAINS.
50
"Tlif tusks in tliu fos.sil ciirvod (lowmvai'dly in a e "^he remains of the sanu' species which prol>ably during
the glacial period extended its habitation very far south of the
latitude in which it has been found in the historic iieriod."! In
view of the now well-known former extension of the habitat of
the Moose, (Jaribou, lieindeer, Musk Ox, and other northern
manuiials, southward to Kentucky, the latter hypothesis seems
the more probable one, and that the species in glacial times
inhabited the eastern coast of the United States southward to
Virginia, if not even l)eyond this point.
3[ore recently. Dr. Leidy has announced the occurrence of
AValrus remains in the phosphate beds of Ashley Jtiver, South
Carolina, and has described aud figured a tusk from that locality.
* Trans Aiiii'f. IMiil. Soi'., vol. xi, jiji. f^l!, 84.
tlliid.. p. .-^l.
^ii, 'i;
\^
GO
< »1 >( )nA:NIJS 1{0SMAHL.S — ATLANTIC WAMJl'S.
( ■':!'
''Til is spcciiiicii," lie siiys, " is its Idacli as cboiiy, dense, licstvy,
and Itiittlc, and is iicaily coiiiplctt', cxccpl a( llic tliin border oi"
the |»idp eavil.v. 'rii(MMu\atiiri' is ,«;li>;lit, and it indii-ates the
tootli li> he oi' the h'l't si(U'." He <;iveH its .
FOSSIL li'K.MAlNS.
01
'• ]| \v;i,s])iirli;ill,v imbedded in :i liiyt-r t»l" blue el;iy ii tu(»t in fliiek-
iiess, (tveiliiid by i) lii.verofli-iliterela.v l\\<> I'eel two iiielies (liick,
('()iil;iiiiiii,n (lists iiiid f^\n'\ls iA' Mi/d (iriiKiria, Maroniii iinhiilBull. Soe. Gdol. de Frauce, 2-= «6r., xv, 1858, p- C24.
i
i
; • \'S'
*S J.«
,:i*^aj^
fi2
(tltlill.r.MS K'OS.MAKMS — ATLANTIC WALKI'S.
}|.
1 H'" t
sont (I'liillcins »|ii(' ties piiiticiiliMitf's relatives ])oiir lii plnpart i\
IVi;;*' ct ill! scxr, iiiiisi <|ni> Til <'tiil)li M. (icivais."*
Vail r.('ii»' oceiiju'et ipie (Iratiolct u
(leeiit sons le nom iVOdohnmtlurv. Lai'tet I'avait reinis s\ (Jratio-
let. Xous avons examine cette tete avee tout le soin mlcessairc
et lions partapjeons conipletenient I'avis que M. I'anl (lervais a
exj>i'iin6 t\ son snjet dans la Zoolof/ie et la PnU'ontoUxjic frani;ai8es
(p. SS), c'est-a-(liie, (pie ee erane IVaittnre et .pit a subi I'actiou
du fen, n'est antie (!hose (pi'iui erane de >rorse vivant qui 6ii(t
rapporte da Xord.
'• Nons avons etndie eett snr nil t'rajiinent de crane (hmt la
eavite een'-hiale a (''t('' ntiUstJe pour iiii iisa;;'e (piehMtiapie et (pii
anra ('id apportc dans eet etat ])ar (piehpie i»("('lieiir dii Nord.
C'est h; e('»t«!i droit et iioii le e(»t('' jianehe (pii est eoiiservtJ.
"(-ehii qui se Ironve devaiit ee rra<;iiieiit de crane et (ini a
devaiit Ini nii choix de seel ions dcs diverses le^ioiis s.
He ennmerates no less than twelve or tifteen spechneiis of these
remains, mostly I'ragments, collected from varions hicalities, all
from the so-called "IJed ("rag" forniati(ni of England, or its
eqni\'alent. The principal hxtalities are Sntton, Felixstow, and
Bawdsey, hi England, hnt he refers also ty tlnMr o(!cnrrence at
diilerent points in Jjelginin. The iiiaj(nity of the specimens
of the tusk obtain(!(l, Avrites .Mr. Lankester, "are its ])ointcd
'Dull. Soc. G<;ol. de rranec, I^' st^r., ii. 1H7-I, pp. ICO, 170.
tDcsL'iip. (tcsOsHcineutsFossilcsdcslJivironsd'Aiivcvs, Aim. Mtis. d'llist.
Nat. (le I]clKi,i,'iu\ i, 18T7, pj). 10. 11.
t'vt.i; lid's, I.'iiiniiiiK fds^ih, l';ti7, p. 1)1)4. — Id., Jlouitininl dca mora . , . , ,
p. 48, 1^70."
FO.SSIL KKMAINS.
<;;;
toniiiiiiitioiis; hill otiifi- s|icciiiii'iis, of tlu^ bust' aiiil iiitcnncili-
iitc |»orti()iis, liu\«' collie lo li;;lil. 'riirtdif-lioiil its U'iij;tli," Mr.
I.;iiilv«'»ti'i- cuiiliiiiu's, •• w iiirli in soiih' cxamplrs must Iiaxc Ih'uii
I'lillv liiivc I'l't't, llic liisU is sli;^lilly curved; but in those wliicli
apptar In 1»<' I'lilly j;ro\vii tlic curve, is (loiisiilerably ;;i'e;ilei'
lowanls the leriiiiiial point, the (lirccttioii of the ciii\c, probabl^N
<>i\iii^ ihe tiislv, it' its l'iiini;;raih^ atllnities Ixt established, a
j'elrollccted position, as in tlic Diiiotluiriiun. Tiie Cia;;' tusk
is \eiy iiiiuih compressed hiterally, so liiat. its transverse sec-
tion lias an elUptical outline, whilst that of th»i Dinothrriuin-
tusk is nearly cireidar. 'I'he amount of lateral compression
is, however, extremely variable, as it is also in tin; iixiii^^- VV'al-
nises; the amount also of the lat«!ral as well as the antcro-
j)osieii(»r llection <»f the tusk appears to vary, as in the recent
Tfivlnriis, the xariability of which in the si/e and form of its
ni>ks is well known. A single lar^e furrow on the outer siir-
I'acc. two on the inner, and one on tin; inner curved margin, ex-
tend aloiij; the wiiole length of the tusk ill many specimens,
exactly similar to those oil some liisks of Walrus; but in both
the recent and fossil specimens llie_\ are subject to much varia-
tion, i.i I heir major or minor dcM-lopmci it. No appearance of aii\
weaiili;;' of Ihe point of the tusks b_\ use diirill.y life is obsiTVa-
l)!r: and indeed the j;reater backward cui'vatnre of that part
sei'iiis to result from its freedom from iisan'c, since in the Walrus
the jioint of the tusk is rapidly worn away, which of course
checks any tendency to curvature which mi,i;lit become. a|)par-
cut if the tusk were not used against such hard substances us
rocks and blocks of ice.
'• I'loiii ail cxaniinatiun of the yeuoral contour and form of the
tusks, without reyard to their substance or structure, one would
unquestionably be led to reyard tlieui us belonj^in;;- to an animal
siiiiihu' to the existiu,';' Walrus, inasmuch as it is in this animal
alone that this form of tusk, \\*ith its longitudinal furrows, j;reat
h.'iijitli, and j;entle curvature, is found."'
.Vfter describin}^' ill detail the structure of ihese fossil tusks,
as shown in seittiyns and as revealed by (he microscope, ^Mr.
Laiikester further observes: -'In its microscopical striutlnre,
the dentine of the fossil tusks presents a complete resemhlauee
lo that of the Walrus,* .... The dentinal tubes are verv
*Tli('ir inicii)s('(>])it'!il siructiu'i', iis wot! ;ih cxttTiKiI \'tn\\\, tar illustrated
liy uuiiu'i'iius liL;iii'.'s, loniiin;^' jilatcs x and \i, afcoiniiaux iiii; Mr. t.aidvf.s-
ii'r's jiaprr.
rM
04
(»I)()IL1>NLJ.S K'OMMAIMS ATI.ANI'IC WAMJl'-S.
I
'I-
'■'1
^i(^^-:1!^l
nearly of the Siiiiic : izc, and ('(inally closclv paclccd, and ate
connected with stellate lacnna' in some nnnilteis near the jjcri-
pliery of the tooth. This Vtinclnic, which is not pc ^n liar to the
Walrns, is, nevei'lheh'ss, a l«'st of alllnitv, inasnnicli as the form
of the lacuna- vaiies in dillercnt animals, 'i'liev are not .net
with in the tnsks of the I'roboscidea or tin' liippopotannis, init
occnr in the cnrions incisors of the l)n,n(»n,ii. i'he ■dentinal
cells' of the < 'ra,n' tnsks also reseiiihle those <»f the W'ali'iis
In Htrucfiin^ the cement exactly resembles thatof the Walrus,
displayin,:;' vascnlar canals, honc-lacnna', and canalicnli. of the
.siune form and disj)osition ; hnt the |>rop(»rtion which it hears
to tln! tlii(;kiiess » I" the othei' tooth tissues appears t(t he lar;>«T
in the Walrus than in the; lossil.
" I'^'rom the foicn'oini;' remarks it will he apparent that w;- have
in thesivfossil tusks characteis which ally them most closely to the
lai'ii'c canines of the ;j,'einis lVo.S(; to establish the ^enus Trichccodoa to
r<'ceiv(( the animal tinis iiidicated. The Jusu(i(;ation of a e,en-
erio .sepiirutioii mu.st bes(m<.dit iuthefaetof the yreat anticpiity
of the 14od Crag, and the consequent probability of the associ-
ation of other and more distinctive attributes with those of the
tusks."
As re^'urds its ji'eolo^^ical posithni ami associations. Mi". Lau
ke.ster adds: " It a])i)ears that the Trichaothm IlKxlciji, like the
Cetice."n remains of the Cray and hw^v. Sharks' teeth,
is a ;, 1>.\ the ;^(.'(>l(»;4i(.'al «'\ idciM'c ol' llirir ;iiiti(i-
iiit\ i!s Itv tli<' ;ictii!ill,\ ■»l).s('i'V('(l and inliriittcdlN s!ij;lit dilVci-
ciicrs of roiMi and striifl inc. Mr. liaiiUcslcr docs not iiiloiiii iis
rcsi»;'(i iii.u I Ik- locality wIicik-c <'aiiic liis spcciiiicns oi' t lie t iisks
ol' ilir li\iii.';' Walrus with wiiicii lie coinitais'il tin- fossil liisks.
Ill I his coll licet ion it iiia.v l»<' added (see further on this point the
Mccoiint nl' 0(l()l)(r'tiis<)h(Niis'j;]\<'n\t{'y{m(\){hn{ (he disks of the i'a-
cilic species (^/(/o/yrr/*»f.s- o/>r.s».s) are not only !oii;;ciand slenderer
than I hose of liie Atlantic species {(). rosiiHiriis), hut are shaiper-
j)oiiilcd and more inciiiNcd, and do not present the worn and
hntkeii appearance so often (indeed, iisiially) seen in the tasks
of lid individuals of the latter. Whet her oi- not tlic\ present
dilfercnces of striK'tiirc has not, so far as known ,0 me. hceii
iiiicroscopieally determined. The tusks of the l*acilic species,
flirt herm(»re, sometiiiK's atfain the si/(^ indicated for the tusks
of" TriclKcodon hnxhuiW For the jircscnt I niiist consider Laii-
k(!Stci"s Tricltcwdon hiudcjil as certainly not j;«'iierically separ-
able from the exislin^' Walruses, although it may have; differed
from the exisliny Atlantic species in larger si/*; and jiossihly in
other cl'.aracters, as so often ha[>pen,s among' the iiniiiediate [»ro-
genitorsof existing speeios in other gToui>s of inainmals.
\'an IJeiieden has recently reviewed iit considerable length
the hi loryof the supposed and actual fossil remains of tin;
Walrus,* sliowing that most of those reported as found in dill'er-
eiit ]»arts of l^'ranc • aiuKjcrmany were really those of d.iHereut
>speci(!s of extinct tSireniaus or other animals than the Walrus.
Van Ucneden, liowever, describes and iigures a dorsal vertebra
Ik; considers as that of the Walrus, found near Deurne, and a sea-
]»hoid hone from Anvers.
(iKiKiKAl'IIlCAI. DlSTUnU'TlON, PKKSJINT AND PAST. — 1.
(Uxtst oJ'Nortk America. — As already shown (^/H/m,pp.'~>7-<»l ), the
Walrus, like the Musk Ox, the (Jaribou, and the Moose, langed
(hiiiiig the great lee Period nn.ch beyond the southern limit of
its Itoiindary at the time the eastern coast of North Ainericii
was first visited by lOiirop 'aiis. AVhile its icmains have been
found as far south :;s Xew .Jersey, Vii'ginia, and even South
< -arolina, thert; is no (;vi.
.Misc. I'ul>. No. IL' r»
fid
nl>0|',,i;M S IJOSMARIS All-AN'I H' WAMMIS.
I
No\ ;i S(i»ti;i, ;i.^ ucll ;i - t lit- sliorc-i mid isliiiids to I In- iioiHiWiird;
hill llii^ ;i|i|M-:irs lo liiivi- \u-i\i :il lliiil liiiic llicji' soiif licrii liinit
Ol l|i>ll illlllidll.
Ill M;i\, I."».'I1, II1C3 well' iiicl uilli l»\ .hiiiics ( iniicr, nhoiit
llic i>l;iiiil ol' '" K':ilii<-;i"' (piol^iMv Siihic I.s|;iii(lj, w li(» t liiis irlcis
1(» IImiii: "Ahoiil Hh- siiid Isliiiid {li:iiii(;i| jiic v»'r\ j^icmU;
Ix'iislcs ;is ;ii(';it iis oxen, wliicli li;i\c luo ^^ifiit tcctli in tlicir
fiioiitlis Iil I'reiyhted
with these lislies." Another writer sa_\s tiiat he had seen
;i "drs Hal lidl al oiiee" ol' llieir teeth, ••uliieh ale a i'oole
and -iiiiielinies more in len'^IJi."" 'riie\ al^o. al ahoiil I li< ,-ame
time, rieipieiited the >o called •• l;ir "' oil' < 'ape Kiel on.
Sa.\ s < hai'les lau ;iieate store of Morsses or Sea Oxen, wlii<'h were a
sleepc upon liic rock<'S: Imt when we apinnachcd nere unto
them uilli our lutale tlie,\ east liieiir-el\e> into tiie >ea and
j)iirsiied ii.> with such fiirie as that ue were ;^lad to lice from
lliein." It is later said that the nnmlMr of these "Sea Oxen'"
was "ahoiil thirt\ or fortv.":j l''rom Ihe accounts of e -'Sea < )xen " u ere a<'custoiiied to
resort to these \arioiis islands during Apiil, .Miiv and .liine.
loi' Ihe. purpose of luin;;ili;; forth t heir \oiiii;:. 'I'liiis, ■■'rhoinas
.lames ol' iJristoll,"' in sjieakin^ of t he " Isle of l.'amea.'" sa.\ s it
was situated "in 17 de;irees, some lift ie lea;4ues from thedrand
IJas. lie. re New foniidlaiid : and i-^alioiil I w eiit ie lea.uiu s aiioiit .
and soiiH' pail of the Island is Hal Sands and shoiilds: and IIk
list I commeih on liaiike do do ilieir kiiidcj in Ajiril, Mav i\: ■liiiie.
by iiiiiiihers of I liou>aiids. w hicli lish is verv hi;;: ami halh tun
^I'eat tcelh; and I he skiniie of I lu'iii is lik< Iliiffi'S l« al her : ami
l apjiiaiaiice ol' rain, tlie,\ immediately
iclicil Id the water with ;4real preeipilal ion. The\ are, when
oh! of I he w aler. \ er\ iiii w ield \ , aiioiin;.; about nine,
moiitli<. Tl.>e> never ha ve more than two at a time, and seldom
more t hail one.
'•'I'lie eehoiiri<'s are t'ornieil principally by nat me. bein;4a j^rad-
iial slope ol'sol'l rock, with w hich I he .Ma;;(lalen Islands abound;
aiioiil so to 100 yards wide at the ualer side, and spreadiii;^ so
;i> lo contain, near Hie siimmil, a very considerable number.
I b II- t lie\ are siiirered lo coiae and amuse t liemseh cs lor a eon
•idi lable time, till t lie_\ acipiire a boldness, bein;^at their lirsl,
laiidin;^ so e.\ceediii;;|\ timid as lo make, il impossible lor any
peiMiii lo ajipioach theiii. In a few weeks t hey asscmlle in
•^leal numbers; roriiierl,\, wlich iindisi iirbed by tiie AmericanM,
!'• Ill'' a moil III of .e\ en or <'i;.;hl ( I ion sand ; and the foi iii ol' the
ecjioin ie not allow inj; 1 hem !oremainconlij4iioiis lo the water, llie.
ion most ones ine, insiMisibly piisiicd abov(! Ilie slojie. vVlicii
! ■'.3
iij
n
J fl
'i"r
IImI;IiivI, Vi)y(i;4cs, Mil. iii,|i.y;!7.
M'li.'iilivoix, viil. \, 11. 'iMi.
y-A'l
SJti&kiM
68
ODOIi/KM S l.'(tS.M.\i;rs ATI.AN IK UAMM'S.
:|i:-'^
'>' '■.
■' f
llicy lire Miiivcd to a cuiivciiiciit (lisliiiicc the lislicriiicii, liii\ iiij;'
'|ir»»\i(Ic(l tlic iicccssiny apparaliis, kiUc tlic adsaiila.nc <»!' a
sea wIihI, tlie slope of the cclioiivlc. the darkness of the nij^lit de-
prives tlicm of e\cry direction to the water, so that they stray
al»oiil and are killed at leisnre, those that arc nearest the shore
bein;; tiie lirst \iclims. In this manner there has Iteen killed
(iftecn or sixteen hundred af one cut. They then skin them,
and take oil' a coat of fat that always surrounds them, which
they dissolve by heal into oil. 1'he skin is cut into slices of
two or tlax'c inches wide, and e\]>ortcd to Ameiica for <'arriaye
traces, and lo iaiuland for ^^liu'. The teeth is an iid'erior sort
of ivory, and is manufactured ibr the same jturposcs, but soon
luins yellow."*
Accoidin.!.;' to Dj. A. ►S. I'ackard, Jr., its bones are still ibund
at th(^ localities mentioned by Hhuldham. "According' to tra-
dition," he further says, "it also iidiabited sonu; of the harbors
of Cape Breton ; audi have l)een informed by a lisherman in
Maine, whose word 1 d(» not doubt, that on an islet near Cape
.Sal)le, Nova Scotia [probably tin- '' isle of liamea" of the early
voyagers abcady (|uotedj, its bones are found abundantly
on the sandy shore, litteen to twenty feet abov<; the sea. In
the St. Lawrence (lulf they were extciininated during;' the
jniddle of the last ceidury. The last one seen (tr hca.rin, wilting a few years later (in bSOD), in
referring to the former oocurrence of the Walrus on the shores
TJiil. Trims., yoI. Ixv, p. '^41).
tProc. Bust. .Soc, Nat. Ili.sl., v(.l. x, ]«)(>, p. >J71.
(UKMIKAI'IIK AL DISIIMIll 'IfoX.
no
;iii(l i'-l;iii\iil ('itiii|i;iiiv of .Mis( on", roiiiided (iiiiinj;' tlie e;iilier
piiil oT tlie .■.e\eiitee:illi eeiililiA. 1>\ lll<' l\ill,u iil' I'^nilice, ;ind
\vli(i>e epIieiiM I'lil eily of New l.'oclielle. iiiliiiiierilii^' ;it one time
some llioiisiiiids, li;is jiiissed i!\v;iv le;i\iiiji iio siyn. 'I'lie miir
(lered Se;i liorses iinxc lel'l :i more eiidiirini;' monument timii
tlie miirdeiers." lie liirtlier iidds: '■'l"lioii,i;li we li;i\e no
iiecoiints Inter timn the sevenleentli and ei.uliteentli eeiitiiries of
tlieir inliahitin;:' Sable Island,-^*! it is very proltable tliat tliey
continued to icsort tliere until tliey entirely left, tliese, latitudes.
Its dillieiilty of access, its heiiif^' nninliahitcd, and its sandy
liars frinj^cd Avilli a ceaseless snrf, ])oint it out as tlieir last
hold."*
Di'. (iilpin also r(M'(mls the ('a])tiire of a Walrus in tin; Stiaits
(if lielle Isle, Lalirador, in IMarch, ISO!*, which was diaimcd on
liie ice lor li\(' miles, and then taken by ship to S!. .lohii's.
New I'onndlaiid. aiid thence to Ilaliiax. No\ii Scotia, where it
was desciilied and li'^iired Ity Dr. (''lusion."§
*!' . <'»J»
■ I'ror. .'iiiil 'I'raiis. Xova Si^oria JumI. Nat. Si'i., vol. ii, pt. I!, ])p. lv''M27.
t lliid.. 11)1. l-.':!-1"^7, witli a plate.
i/.o.'iinMi-i, 1871, -rjoO.
^Vv'H'. /,oul. Si)c. liond., 1808, p. 4',V.\.
'()
!iii(l:iiil ;il)(iiit Tori iMJiillic, on llic wcslciii coiist
ol' (licchl.'iinl. ill liililiidr Til', but tlicyscciii Id Innr, >iiic(f
I li;il ihilc. nical ly tlrcrcwsrd in niiiiilicis iiloiiu I lie w liolr nf tlic
( 1 l((ill;l!|(| en; I si. ClI pill ill i'ciN Icll. Ill ili- | i:i piT oil t lie •• .Mlllli
II 111 11; I (li Noilli ( Irccnliiml niid ( IiiiiimH I-iiiid," oIiscia cd in is;,')
h,\ llic r.iilisli Arctic I'Apcdil ion, iil'lcr iilliidiii.L; to tlicir roiMici'
;iliiiiid;iiicc .ihoiil I'ort I'onllvc. ns ol»scr\ fd hy l\;iiic;iiid lliiycs,
Miys: •• < 'iiiioiisly ciioii;;ii. wcdid not sec one ol I licsc iiniiiiiils
ill llic \ iciiiily of Toil i'oiilkc nor in Sinitli Soi !«•, until wt;
rciiclicd l''i;iiil3 and (tS'5 >;. lat., where it oc<'iirs pretty numerously ;it times,
'^riie darin,n' task of enteiin;^' into c(»iitest with this animal from
the kayak on the open sea forms a re,L,uIar sport to the natives
(d' Kati.i;amiut in (>i'P X. lat. Tiie num'ier yi'urly killed has
not been separatel\ calciilaled, . . . but they can hardl.\
ex(i<."|:
'I'lie westeininost point at which it has been oliserxcd is .--aid
to be t he western shore of lliidsop's liay. Mr. -I. ( '. itoss stales
it to bean iiihabitanl of tlie wi'Si coast of I'atlin's May and
"'I'lir /(ii'i|(i;;h t, :',(! ^c|■., \n\. i, |i. ol)!!, Sc|)l .•llllicr, l'"^/".
( III MSS. IKllrs lie JKIS liilldJS' pJMCld ill IMV ilis|ios;il.
I Diiliisli < ;i((iil.niil, ils i'((p]p|i' :iii(i its I'inilncls. pp. I'JIi-JiT, tH77.
n
fJI'.ocIv'AI'lIK ,M, DISTKIItTITION.
71
: ri'
hi m
i;r|(lll>i' I!;IV- mill lo lie ((r!';isi(tli;ill\ liH'l willl ill I ln' IMilllicni
|;iil (,r riiiicf IJci^cnrs Inlci, lull siiys it is iiiikiiow ii lo llic
ii;ili\ (--ol Itoiilliiii.* Dr. Kicliaidsoii s;i\s: ''Tiic \\ iiliiiscs wcrii
\ci\ iiiiiiicroiis ill I.L;I(»(»lik iiiid uit Ilic oilier |t;iits uT I lie <'(iii,sl
Id iIm' cMstwiird of lilt' l''iiry siimI llccia's Sirait. Tlirs arc not
loiiiKi. Iiowcxcr. al IIh' iiioiiIIi of llic Copper Aline Ijiver,
;il!li(i!i,'^li liie hiaek w hail' liati lieeii soiiKjIiiiies (Irilled I liillier."t
lie also rel'ers lo lis heiiin iiiikiiown lo tlie Ivskiiiio.' ol' the
('opperiiiiiie and Mat-ken/ie ifivers. ;j; No species of Walrus
;ippeais lo lia\t' excr Itceii seen on tin' Arctic c(tasl of America
helwceii llic !l7lli and l."),Slli iiierialnied luiiong th('ir Jolk-lore, may
he th(^ Waliiis."H I''!eming- states that one was killed in the
Sound of Stockness, on the east (!oast of Iluiris, in December,
1817, 1[ while another, accoidin;;- to Maegillivray and others, was
killed in Oikney in .Fiine, 1S2."».** Mr. \i. Brown adds that onci
was seen in Orkney in IS57, and another in IjTor' Isles abont tlio
siune time. ft It appeals tohave !iever ocenrred in Iceland, ex-
cept as a li' (; straggler. Many years ago tliey are said to have
lived on the shonss of I'^inmark, and at a innch lat< <• date to
haveabonndedonsonujof the i.slands oil" this coast. Mi'. Lamout
says: " We learn from the voyag(! of OhthcMv, wiiich was per-
formed aboid a tlnaisand yeais ago, that tlie AN'alriis then
abounded on the coast of I'inmarken itself; tiieyhave. however,
abandoned 'hat coast for some eentnries, although indi\idnal
stragglers lia\e been occasionally cajttnred there ii|» to within
■' Ivd.ss's ■.'(! \()y., All]!., is;!."), p. wi.
tSiiii|)l. I';iri-.v".s ','(1 Voy,, |i. ;!:if.
t Ziii'ilo^y of IJccclii'y's V^ty;i;j;c, Mam., !-•:'>'.', p. <>.
VI lli'ctor lidccc's History of ScotliiiHl. .■is(|iiot('cl liy I'rili.sli /(iiilni;i>its.
1! I'loc Zoill. S(ir. I.ohil.. HiW, |i. \:\\\.
• isiiiisii \iiiiiKii>, p. i:i.
■'lidiiili. New I'liil. .(oiini., \ (il. ii, p. !!'■'!•; I5ri)isli (,!ii;iut still in the southeast <'orii<>i'|, and
Jiyk Yse Island, which in their turn aic now veiy inleiior
lniiitin;^-j;round to the Itanks and sUeiiies lyinj;' to the north of
Sititzherficn.
" J'oilninit«'ly Cor the persecuted Walruses, liowever,tliese lat-
tei' disti'icts are only accessilile, in open seasons, or ])erliaps once
in three or loui' summers, so that, tliey }ivt a little breathinji'
time there to i)reed and re])lenisli their inimhcrs, oi' undoubt-
edly the next twenty or Ihiity years would witness the total
extinction of Hosmtirns Irirlicrns on the coasts of the islands of
Northern I'^urope.
"Th(^ Walrus is also loundail round the (!oasls (»f No\a Zeni-
l>la, but not in such ninubejs as at Spitzberj^en; and he inider-
g'oes, if |)ossible, more jx'rsecution in those islands frctin some
colonies of Itussians or Samo'icdc's, who, I am told, reuularly
winter in Xova Zembia ibr tlu' purpose of hunt inj^' and lish-
in,yv'f
''The war of extermination,'' sa\s Mr. Kamoiil, in his later
Avork. •• which has been cariied on Ibr many years in Spitzi)er-
j;en and N(»vaya Zendya has driven all the Arctic faiuia jmain-
iiialsj from their old haunts, and. in seekinj;' I'ctreats more inac-
cessible to man, it is piobable that they have had in some
defiTce to alter their habits. I'or exaMi]>le, up to about twenty
years aj^o it was eustomai'y for all ^\'alrus hunteis to entertain
a reasonable lio|»e that by waitinj^' till lat(> in the season all Ibr-
mer ill-luck miyht be com])ensated in a few fortunate hours by
Ifillinj;' some hundreds on shore; in fact, favorite haunts were
AV( II known to the lishers, and were visited successively before
linally lea\ in;; the liuntinj;j;i'ounds. >iow, althouj>h the Arctic
seas ai'c ex])l(U'e(l by steamers and visited anraially by as l)o]d
and enlerprisinj'' hunteis as foimerly, siu-h a windfall as a herd
of Walruses ashore is seldom heard of,
" Each year better found \ essels and mor(^ elaborate weai)ons
* Jb\ LiiiiKiJil liassiiMT rcpoilcd the iiipl iiic of :i Ijuj^o IhiII " in Mm^^cii")
Soiuid near Ihc, NortJi ('.i|ic aliunl iHCiri." — Vdrhl'nuj in ihc Arctic licua, \\. 58,
foodioic.
t Hi'ii.sdii.s with the (Sra-lior.scs, pp. 1(17, Kib.
(;i;(»(ii;Ai'iii(\i. distimisi ikin.
I ■(
jUf sent "iiil In li;iri\ tin- Wiilriis; ;is ii n»iisc(|iifii<'(' cn civ mm-
.■soii tlicrc is niiMicr «lir(i('iill v in olttiiiiiiii;;- ;i (miuh — lor two rc;i-
sniis. I iKisriiliiliKiis wliirli li;i\r \i ■mi I led iliti) w li;il w iis siifc Irr*!
iiiy ;;!()iiml liisl ,vc;ir iiict'l liiciiciH-iuv, ;iii«l liiiil' iiic killed, u liilc
the (itlicr liiiir csciiiiiii,'^ will he IoiiimI iirxl ,\ riir ;i sIc)) r;nl Ikt
ii\\;i\. Tliis iiilclliuciil rcli'ciitiii^' ul' the Wiiliiis hciorc ;i siipc
lior nicDiN will, 1 hclicxc, pi'csrrNr I lie species iii'iei' its sciii'eily
ill aecessihle w;ilers reinlers it no loiter ;iii oliject of sport niiiiiiiierce. Tliiit tli( \\ ;ilriis, ... is heiiij^ (Iriveii IVoiii every
(iistrict where llie li:iii(l of iiiiiii is lelt, is eerlaiii."*
Mr. Alfreil Newton, wriliiij;' in lS(>t, iMjsjx'ctiii^' llicir loniier
])reseiiee on the eoiists of J<''iiMii:irk, iiiid theii'distriliiitioii at tliiit
.serves : '• I se«' no icason to doiild (he a.sseition, oi jier-
liaps it would he safer to say the inference, that in former days
Walruses hahitiially frecpieiited the coasts of i'lniiiark'. fn tlie
sixteenth and seAcnteeiith ceiitiirios t hey were certainly ahniid-
aiit altoiit l!ear I.-laiid; llie,\ are spoken of there as ' iyin;^' Iik(^
houses upon heaps, '|t| . . .: ye) for the last thirty years proli-
alily not one has lieeii seen there. Now they are lieninied in l»y (he
packed ice of the i'olarSea on the one side and (heir nicrciless
eiieiiiies on the other, 'i'he result cannot admit of any r);;-l."».i|, we r<'ad: '' 'I'o the North part of that Conn
trey are the places wliere they have their i''uircs. as .Sahles,
Martenis, <;re('se iievci's, Foxt's white, hiacke, and icd, Minkiis,
]']riiiiiies, Minivers, ami Harts, Therc^ are als(» a lisiios te(3th,
wliicli fish is called a ."Morsse. The, takers fiicreof dwell in a
place called i'ostesoia, which hrin;; them njxni Harts to Liim-
pas to sell, and from LitiujxiN carrie tiieiii to a place calh.'d Col-
inoyro, wiien^ the hij^li Market is holden on Saint Nicolas day,"§
On Ilondins's iiia|> of I'nssia a(;(;onipanyin|). 2i:i, 'Z\\.
' 'm
><■!
:f Ife t
K;y
Miit^-*''
'I
,1' 'V ,
74
oi.oit.r.Ms i;nsM,\i;i s — \i i.amic \v am; rs.
' ■{' '
'Wf
;.:'
'I'lir Wiilrii.scs ii|i|i(;;!r lo lia\ (' Itccii 111^1 iiici wiili on ( 'liciii
Island in H>n.">, inid lolinxc iH-cnmc n,'arl\ r\|ci minalcil iIhtc
w il liin a \ civ I'l'W ,\cai' •. 'I'lic liislois nl' I hric dc-l i iicl ion 1 lnic
and a! S|iit/.lHi!^ pari oT ilic ■.(•\cnli lul li
cent ni.\ is ;^i\rn in tin- lollow in.i: cxcci pis : •• In I lie v( iiic hio.").
Sirjdicn llcnni't was iinplo\cd lt\ I lie roiapanic." in a ."-Wiip
called llic ^//vfrf. |o iIkisc parls X((il liuards ol' ijic <'aiic|--or
Ni!r\\a\ "|, ainl \\a^ a I ( 'lici ii' I hind :\\\i\ Ixillcd ;.ohic S( aiiorsc.s.
and ltroii;^lil home head Oaic I'loin tlicn<'c . . .
•■ 1 1 cere It is to lice mid erst ood, that t he ( 'on i panic !ia\ iii.^ I>\
(d'teii I'csorl and iiiiplo\ iiient to lliose parts. oiisei\c," says tliat as they approa'lie.
wc went alioard." 'I'lie next da,\ they went on shore ayain and
* liii( irpiii.it til ,.(iiiir I iiiic pridi' to 1 i.i' yciir iririti, uih1r('ak(! it, if it
lice not nil the heller phited. Tliey will also strike with their
Tectli ill him tliiit is next llieni: lint l)eeaiis(^ their Teeth <;row
downward, I heir str(>]ces are of small force and danj^er." They
took in ••ele\('n t mines of Oyle, and the teeth of all the beasts
;i(on's;ii(l."
'{"he I'ollowin;; year (KJOfI) they aj^ain set oid for Cherie Island,
arriving theic Jidy .".. They found Die ice still about tin; island,
and the Walruses not yet on slioic; '' I'or their nature is sucli,
thill tlie.\ will not come on land iis lon^j; as any lee is about the
liiiid." ( )n the Ilth they ])erceived on shore "of the beiists
sidlicieni to iiiiike our xoviijic, \ve(' ])repared to ;•■()(' killing.
Miister Wrhlcii and Miister /iV'?//(f7 appointed nice totidv'e eleven
Mien with mee, and to }^o(! l)eyond the beasts wlieic they lay;
tiiiit they iuid w(M' mi;4ht meet at the middest of them, and so
enclose them, thiit none of them should ;i('t into the Sea, . .
. . iind before six lionres were ended, we li;id slayne ;d)oiit
sexcii or eijiiit hundred Ueasts. . . . l''or ten diiyes spae(^
we jilved our businesse veiy hard, and brought it almost to nn
end." They took in "two and twentie tuns of the Oyle of the
IMorses, and three hogsheads of their Teeth."
in KiOS they :ig;iin reaehed Cherie island towaid the (Mid of
-bine, and on the 22d "came into a CoveAvherc the .Morses were,
and slew abont OOO. or ]()()(». of them in less than seven hoiires:
and then we plied our business untill the sceond of .Inly : at
wliiit time Ave Imd tidcon into our shi]) L'L'. tuniies and three hogs-
heads of Oyle." On their icliirn they look with theia two live
young Walruses, on(! of which lived till they reached London.*
The voyage in 100(1 w;is less successful. They slew at one
tiiiK! eighty, at iinotlier one hundred and liity, iind id siill iin-
"I'mcliii.s liis I'ilniiincs, vol. iii, |i|i. ri.">7-r)(j().
-. -wn
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IMAGE EVALUATION
TEST TARGET (MT-3)
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76
ODOB^NUS ROSMARTJS — ATLANTIC WALRUS.
other time forty-five; but they lost most of tliem in coiiseqiieuce
of bad weiitber. " In the yeore IGIO. the Coini>anie set out iwo
Ships, viz. the Lionesse for CJicrie Hand, Thonun IJdpe Com-
mander ; and the Amitie, for a Kortherne Discoverie, tlie Mas-
ter of which ship was Jonas Foole : who in the moneth of May
fell with a Land, and called it Greenland, this is the Land that
was discovered by Sir Hugh WiUonghby long before [Spctsberg
of the Hollanders], which Shij) Anitie continued upon the coast
of Greenland, discovering the Harbours and killing of Morces
[the first killed by the English on Spit>.bergen], untill the moneth
of August, and so returned for England, having gotten about
some twelve Tunnes of goods, and an Unicornes home.
"In the yeere 1611. the Companie set foorth two Ships, the
Mark Margaret Admirall, burthen one hundred and sixtie
tunnes, Thomas Edge Commander ; and the Elizabeth, burthen
sixtie tunnes, Jonas Poole Master, well manned and furnished
•with all necessarie Provisions, they departed from Blackwall
the twentieth of Aprill, and arrived at the Foreland in Green-
land in the Latitude of 79. degrees, the twentieth of May fol-
lowing, the Admirall had in her six Biskayners expert men for
the killing of the Whale : this was the first yeere the Compa-
nie set out for the killing of Whales in Greenland, and about
the twelfth of June the Biskayners killed a small Whale, which
jeelded twelve Tunnes of Oyle, being the first Oyle that ever
was made in Greenland. The Companies two Shalops looking
about the Harbour for Whales, about the five and twentieth of
June rowing into Sir Thomas Smith his Bay, on the East side
of the Sound saw on the shoare great store of Sea-horses: after
they had found the Morses they presently rowed unto the ship,
being in crosse Eoad seven leagues off, and acquainted the Cap-
tayne what they had found. The Captaync i nderstanding of
it, gave order to the Master, Stephen Bennet, that he should take
into his Ship flftie tunnes of emptie Caske, and set sayje with
the Ship to goe into Foule Sound. The Captayne went pres-
ently aw{iy in one Shallop with sixe men unto the Seamorse,
and tooke with him L&nces, and comming to them they set on
them and killed five hundred Morses, and kept one thousand
Morses living on shoare, because it was not profitable to kill
them all at one time. The next day the Ship being gone unto
the place & well mored where the Morse were killed, all the men
belonging to the Ship went on shoare, to worke and make Oyle
of the Morses ; and when they had wrought two or three dayes,
GEOGRAPHICAL DISTRIBUTION.
77
it fortuued that a small quantitie of Ice come out of Foule
Sound, and put the Ship from her Moriiig, . . . The Ship
being cast away without hope of recoverie, the Commander
Thomas Edge gave order, that all the Morse li^aug on shoare
shoold be let goe into the Sea, and so gave over making of
Oylc. . . ." Fitting np their boats as Avell as they could
they soon after abandoned the coast of Spitzbergen (" Green-
land "), and set sail for Cherie Island, ^\ here they found the
"Elizabeth" and returned to Spitzbergen "to take in such
Goods as the sayd Edge had left in Foule Sound, woorth flfteene
hundred pounds.'*"
As early as the year 1611, the previous persecutions of the
Walruses at Cherie Island had made them very wary. Thomas
Finch, in his account of a visit to this island by William Gom'-
don in August of that year, says: "At our comming to the
Hand, wee had three or foure dayes together very fine weather:
in which time came in reasonable store of Morses, . . . yet
by no meanes would they go on those beaches and places, that
formerly they have been killed on. But fortie or flftie of them
together, went into little holes within the Eocke, which were so
little, steepe and slipperie, that as soone as wee did approach
towards them, they would tumble all into the sea. The like
whereof by the Masters and William Oourdons leyort, was never
done."t
During the years 1612, 1613, 1614, and 1615, numerous vessels
were sent out from England to Spitzbergen for the products of
the Walruses and Whales, but generally met with indifferent
success, being much troubled with Spanish, Dutch, and Dan-
ish " interlopers." ' ' •' ' ■'
" In the yeere 1616, the Companie set out for Greenland eight
Sayle of great ships, and two Pinnasses under the command of
Thomas Edge, v, ho following his course, arrived in Greenland
about the fourtii of June, having formerly appointed all his
ships for their severall Harbours, for their making of their Voy-
age upon the Whale, and having in every Harbour a sufficient
number of expert men, and all provisions fitting for such a Voy-
age. This yeere it pleased God to blesse them by their labours,
that they full laded all their ships with Oyle, and left an over-
plus in the Countrey^ which their ships could not take in.
They imployed this yeere a small Pinnasse unto the East-ward
part of Greenland, Namely, the Hand called now Edges Hand,
* PurcluiH his Pilgrimes, vol. iii, pp. 4(54, 465
tlbid., p. 536.
78
ODOBiENUS ROSMARUS — ATLANTIC WALRUS.
and other Hands lying to the Korth-wards as farre as seventie
eight degrees, this Pinnasse was some twentie tunnes, and had
t\.elve men in her, who killed one thousand Sea-horses on
^flges Hand, and brought all their Teeth home for London^
In 1017, they " employed a ship of sixtie tunnes, with twenty
men in her, who discovered to the Eastward of Greenland, as
farre J^ or th- wards as seventy-nine degrees, and an Hand which
he named Witches Hand, and divers other Hands as by the
Map appearoth, and killed store of Sea-horses there . . ."*
The Dutch, Danes, and Spaniards began, in 1612, also to
visit Spitzbergen in pursuit of Whales and Sea-horses, but axe
reported by the EngUsli to have made indifferent voyages. The
company soon also had rivals in the " Hull-men," who, as well
as the Dutch, did them much "ill service."!
About the years 1611 and 1612, the Whale-fishery was found
to be more profitable than Walrus-hunting, and subsequently
became the main pursuit, not only by the English, but by the
Dutch and Danes. Yet the Walruses were by no means left
wh jily unmolested, having been constantly hunted, with more
or less persistency, down to the present day, and, as already
shown, were long since exterminated from Oherie Island and
other smaller islands more to the northward, and greatly re-
duced in numbers on the shores of Spitzbergen.
Walruses have been recently reported as occurring on the
outer or northwestern coast of Nova. Zembla, but as not exist-
ing on the inner or southeastern coast. Von Baer, on the au-
thority of S. G. Gmeliu and others, gave the eastern lunit of the
distribution of the Atlantic Walruses as the mouth of the Jene-
sei River, though very rarely single individuals wandered as
far eastward as the Piasina Kiver. He even regarded the Gulf
of Obi as almost beyond tlieir true home.| Von Middendorff,
however, considers von Baer's eastern limit as incorrect, and
cites old Eussian manuscript log-books ("handschriftliche
Schiftsbiicher") in proof of their occurrence in numbers in Au-
gust, 1736, as far east as the eastern Taimyr Peninsula, and of
their being met with in August, 1739, as far east as Chatauja
Bay. Still further eastward, in the vicinity of the mouth oi the
Lena Elver, he gives similar authority for their occurrence in
August, 1735, and says that Dr. Figurin attests tlieir presence
*Purclia8 his Pilgrimes, vol. iii, p. 467. tibid., pp. 472, 473.
tMdni. do I'Acad. des Sci. de St. P(5tersb.. vi° 8<5r., Sci. math., phys. etnat.,
tome iv, 2''" pars, pp. 174, 184.
GEOGRAPHICAL DISTRIBUTION.
n
-on the shores of the delta-islands of the Lena. Eespecting the
move easterly coast of the Siberian Ice Sea, he says it is cer-
tainly known that the Walruses of B iring's Sea extend west-
ward in great numbers to Koljutschiu Island. Only the males,
however, reach this limit, th'j females not extending beyond the
Aicinity of the mouth of the Kolyma Eiver.*
It lience appears that about 1735 to 1739 Walruses were
met with as far eastward as the mouth of the Lena Eiver; but
Wrangell, nearly a century later, explored quite thoroughly
this whole region without meeting with them, and I have found
only one reference to their existence on the Siberian coast be-
tween the Kolyma and Jenesei Elvers later than those cited by
von Middendorff.
According to a recent letter t from Professor iNordenskjold, of
the Swedish Northeast Passage Expedition, "two Walruses"
T\ere seen in August, 1878, a little to the eastward of the Jenesei
River, and that open water was found as far as the mouth of
the Lena. From this it would seem that there is nothing to
prevent, at least in favorable years, the W^alruses from passing
eastward to the mouth of the Lena. There still remains, how-
ever, a breadth of some thirty degrees of longitude (between
13(P and 3 60°) where as yet no Walruses have been seen. They
appear to have been only verj^ rarely met with to the eastward
of the Jenesei (longitude 82° E.), and to be uncommon east of
tlie Gulf of Obi.
At present the Atlantic Wiilrus ranges along the northeast-
eru coast of North America from Labrador northward to Ee-
pulse Bay and Prince Eegent's Inlet, and along the shores of
Greenland; in the Old World only about the islands and in
the icy seas to the northward of Eastern Europe and the neigh-
boring portions of Western Asia, where it rarely, if ever, now
visits the shores of the continent.
On the easttmi coast of North America, Walruses have been
met with as fiir north as explorers have penetrated, and as far
as the Esquimaux live. They winter as far north as they can
find open water, retiring southward in autumn before the ad-
vance of the unbroken ice-sheet. Kane speaks of their remain-
ing in Eenssellaer Harbor (latitude 78° 37') in 1853, till the sec-
ond week of September, when the temperature reached zero
of Pahreuheit. |
* Von MiddeiKloiiPs Sibiiischo Reise, Bd. iv, 1867, pp. 935, 936.
t Soo Nature, vol. xix, p. 102, December 5, 1878.
t Arctic Exploration, vol. i, p. 140.
80
ODOB^NUS ROSMARUS ATLANTIC WALRUS.
Nomenclature. — Several specific ntunes have been iii moro
or less current use for the Atlantic Walrus, or rather for the
Atlantic and Pacific species collectively. Accepting Odoba'nus
as the proper generic name of the group, there is nothing to
prevent the adoption of rosmarm for the specific name of the
Atlantic species. It Avas used for this species exclusively by
Linn<;, Erxleben, and other early systematic writers, the Pacific
Walrus being at that time unknown to the systematists. If lios-
marus be used as the geneiic name of the group, as it lias been
by a few late writers, as a substitute for the wholly untenable
one of Trichechtis, it will be, of course, necessary to adopt some
other name for the si)ecies. Dr. Gill has used obesus of Illiger ;
but as this was applied by Illiger exclusively to the Pacific Wal-
rus, it cannot properly be used for the Atlantic species. It
would be difl&cidt to select a subsequent name that would not
be open to objection, if one should stop short of trichechus, used
(inadvertently?) in a specific sense {^^ Rosmarus trichechus^') by
Lament in 1861. The name lonaidem of Fremery, 1831, was
based on what subsequent writers have considered as probably
the female, but the name is highly inai)propriate, inasmuch as
it is the Pacific species, and not the Atlantic, that has the longer
tusks. There are left virginianus of DeKay and dtMm of
Stannius : the first is objectionable on account of its geograph-
ical significance ; the other is only doubtfully referable to the
Atlantic species. Adopting Odohcenus for the genus, leaves
rosmartts available for the species, thus settling the whole diffi-
culty.
As aheady noticed [antea, p. 20), two species besides virgi-
nianus have been based on fossil remains, and have been made
the basis of new genera. The first of these is the Odobenothe-
rium lartetianum of Gratiolet, since referred by Defrance to the
existi":'ig species ; the other is the Tricheeodon huxleyi of Lan-
kester, which there is perhaps reason for regarding as the large
extinct progenitor of the existing Walruses.
Etymology. — The term rosmarm was originally used by
Olaus Magnus, about the middle of the sixteenth century, in a
vernacular sense, interchangeably with morstis, the Latinized
form of the Eussian word niorsz (or morss). It was used in tho
same way by Geaner a few years later, as well as by numerous
other pre-Linnaean authors. Eespecting the etymology of the
word, von Baer gives the following : " In dem historisch-topo-
ETYMOLOGY.
m
grapliischen Werke : Be gentium septentrionalium conditionibus
cet. Komae 1555 lieisst es : Normgium littns maximoa ac grandes
places elephantis hahet, qui morsi seu rosmari vocantur, forsitan
oh asperitate mordendi sic appellati, (Eine recht witzige Etymo-
logic !) quia, si quem hominem in maris littore viderint apprehen-
dereque poterint, in eum celerime insiliunt, ac dente lacerant et in
momento interimunty *
The same author also gives the following from Herbeistain
(1567): "Under andern ist auch ein thier, so grosse wie ein
ochs, und von den einwonem Mors oder der Tod geheissen
wird." t Hence, either from superstitious notions of the terri-
ble character of this animal, or from the resemblance of the
iiussian word morss to the Latin word mors, these terms be-
came early confounded, and rendered by the German word Tod,
or death, f
In the account of the exploits of the Norman Othere, where
the Walrus first finds its place in literature, it is termed Horse-
wael. As noted by Martens § and other writers, equivalent
words in other languages have become current for this animal,
as Walross or Wallross of the Germans, Wallrus of the Dutch,
*In an early (1658) English version of Olaus Magnus's work ("A Com-
pendious > ''story of the Goths, Swedes, &, Vandals and other Northern
Nations. .itten by Olaus Magnus, Arch-Bishop of Upsal, and Metropoli-
tan of Sweden", i». 231), this passage is rendered as follows: "The Xorway
Coast, toward the more Northern parts, hath huge great Fish as big as Ele-
phants, which are called Morn, or Rosmari, may be they aiie so from their
sharp biting ; for if they see any man on the Sea-shore, and can catch him,
they come suddenly upon him, and rend him with their Teeth, that they will
kill him in a trice." From this it would appear that Morsua, as used by
Olaus Magnus, might bo simply the Latin word morsus, from mordere, to bite,
tSee von Baer, M^m, de Acad, des Sci. do St. Pdtersb., vi« 8<5r., So.
math., phys. et nat., tome iv, 2^" pars, pp. 112, 113.
^ Von Baer quotes a passage from the "Rerum Moscoviticarum auctores
varii," originally published early in the sixteenth century, in which occurs
the phrase " scandut ex mari pisces morss nuncupati," which he regards as
the first introduction into Latin of the Sclavic name MopiKb. In Western
Europe it a little ^ater became current in the form oi Morse, whicii was soon
written Moras or Mors, from which Buflfon later forme*' the name Morse,
which has since been the common appellation of this animal among French
^vrite^9. Von Baer further observes that the accidental resemblance in
sound of this word to that of the Latin word for death (mors) appears to
have contributed not a little to the strange conception of the torribleness of
this animal whicli was early entertained and even still prevails in Western
Europe, although the Russian accounts do not speak of it.
«Zoolog, Garten, Jahrg. xi, 1870, p. 283, where the etymology of tho
numes of tho Walvus is briefly discussed.
Misc. Pub. No. 12 6
82
ODOBiENUS R08MARUS — ATLANTIC WALRUS.
and Walrm of the English. By the early Scandinavian writer*
it was termed Bosmhvzlr, which later became resolved into Boa-
mul, from which, perhaps, originated the Latin term Boamarm,
which has the same significance, introduced by Olaus Magnus
and Gesner, and the Norwegian word Boatungr. Gesner and
several subsequent writers also used the word Meerroaa, and we
have in English the equivalent term Sea-horae^ as one of the ap-
pellations of the Walrus, and also, but more rarely, Meerpferd
in German, and Clwval marin in French.
The current French term Morae appears, as already stated, to
have been introduced by Buffon as a modification of the Eus-
sian word moraa, used by Michow (1517) and Herberstain (1649).
Among other old vernacular names we find in English Sea
Coic, in French Vache marine, in Latin Boa marinm^ etc., while
by the early French settlers in America it was commonly termed
Bete f> la grande dent. ^
Literature. — 1. General Hiatory. — Passing over the by some
supposed allusions to the Walrus by Pliny as too vague and
uncertain for positive identification, * we meet, according to
von Baer, with ^^he flist positive reference to the present
species in the account of the exploits of the famous Norman ex-
plorer Othere, or Octher, who, about the year 871 (890 accord-
ing to some authorities), made a voyage to some point beyond
the North Cape, where he met with large herds of Walruses,
some of the tusks of which he is said to have taken to England
as a present to King Alfred, t Walruses appear to have been
* See K. E. von Baer, M6in, de I'Acad. Imp. des Sci. de St. Pdtersb., vi'""
s^r., Sci. math., pliys. et nat., tome iv, 3™« livr., 1836, (1837), pp. 101, 102. T ..
this admirable monograph I am greatly indebted for information respecting
the earlier publications bearing upon the history of the Walruses. To this
exhaustive memoir the reader is referred for a full exposition of this part of
the subject. The following short summary is based, so far as the early his-
tory of the subject is concerned, mainly upon von Baer's monograph, an
analysis of which will be presented at a subsequent page. (See posted,, p. 88,
footnote.)
t Hakluyt's rendering of this account is as follows: "The principall
purpose of his [Othere's] traveile this way, was to encrcase the knowledge
and discoverio of those coasts and countreyes, for the more comoditie of fish-
ing of horsowhalcH, which have in their teeth bones of great price and ex-
cellcncio : whereof lie brought some at his retume unto the king. Their
skinnes are also very {\;ood to make cables for shippcs, and so used. This
kind of whale is much lesse in quantitie then other kindes, having not in
length above seven elles." — Hakluyt's Voyages, vol. i, p. 5.
i
GENERAL HISTORY.
88
an object of chase on the coast of Finmark as early as 980, and
must have been met with by the Norsemen when they visited
Greenland about the end of the tenth century. Their tusks
were an article of commercial value among the Mongolian and
Tartar tribes as early as the twelfth to the fifteenth centuries.
Aside from the various notices by Scandinavian writers, the
earliest unmistakable reference to the Walrus, other than that
connected with Othere, as above mentioned, was, according to
von Baer (1. c, p. 108), by Albertuf^ Magnus, in the first half
of the thirteenth century.
Says this writer (as quoted by von Baer), whose account is here
paraphrased : The hairy Cetaceans have very long tusks, by
which they suspend themselves to the rocks in order to sleep.
Then comes the fisherman and separates near the tail as much
skin as he can from the underlying fat, and then attaches a
cord, which has at the other end a large ring, which he makes
fast to a post or tree. Then when the fish awakens (by all of
these operations he was not yet awakened), they cast a huge
sliug-stone upon his head. Being aroused, he attempts to get
away, and is held by the tail near to the place and captured,
either swimming in the water or half alive on the shore. This
ludicrous description von Baer believes had for its foundation
misunderstood reports of the Walruses' habit of reposing upon
the shore or upon ice-bergs, the use of their tusks in climbing
up to these places of rest, and their deep sleep, and that the
account of the mode of capture was based on an incorrect
knowledge of the use of the harpoon; and that the account
shows that as early as the thirteenth century the Walrus was
harpooned on the coast extending from the White Sea north-
wards. *
* This curious legend is quoted by Gesner in his Historia Animalia Aqua-
tilia, 1558, p. 254. The following rendering appears also in the above-cited
English version of Olaus Magnus : ' ' Therefore, these Fish called Boamari, or
MorA, have heads fashioned like to an Oxes, and a hairy Skin, and hair grow-
ing as thick as straw or corn-reeds, that lye loose very largely. They will raise
themselves with their Teeth as by Ladders to the very tops of Rocks, that
they may feed on the Dewie Grasse, or fresh 'Vater, and role themselves in
it, and then go to the Sea again, unless in the mean while they fall very fiat
asleep, and rest upon the Rocks, for then Fisher-men make all the has.te
they can, and begin at the Tail, and part the Skin from the Fat ; and into
this that is parted, they put most strong cords, and fasten them on the rug-
ged Rocks, or Trees, that are near; then they throw stones at his head, out
of a Sling, to raise him, and they compel him to descend, spoiled of the
greatest part of his Skin which is'fastened to the Ropes : ho being thereby
84
ODOB^NUS R08MARUS— ATLANTIC WALRUS.
The Walrus is also referred to by Hector Boethius in 1526, in
bis History of Scotland ; * by Herberstain (or Herberstein, as
also written) in 1549; by Par6 about the year 1600; and by Al-
drovandus in 1642.
Herberstain also very correctly indicates the habits of these
animals, which, he says, repair to the shore in large herds to
repose, and that while the herd sleeps one of their number
keeps watch. He compares their feet to those of the Beaver,
and refers to the value of their tusks tc the Russians, Turks,
and Tartars, and observes that they called them fish-teeth, t
Even before the middle of the sixteenth century. Walruses
had been met with on the eastern shore of North America. In
May, 1534, they were seen by Cartier, and later in the same
century by Fischer, Drake, and others, on the coast of Nova
Scotia and adjacent islands, and later still by other explorers on
the islands in the Gulf of Saint Lawrence (see antecl, p. 06), in
the accounts of whose voyages | occur interesting notices of
these animals.
In the year 1553, Edward VI of England sent an expedi-
tion under Willoughby and Chancellor to the White Sea, which
resulted in still further increasing our knowledge of the Wal-
ruses, especially of their distribution eastward along the Arctic
coast of Europe and Asia. Chancellor's short account § refers
especially to the uses made of the skins and tusks.
The earliest delineations of the Walrus appear to have been
made by Olaus Magnus in his " Tabula Terrarum Septentrio-
nalium" (1555), where he has portrayed many strange and fabu-
lous animal forms which there is reason to believe were based
upon this animal. 1 1 Gesner a few years later (1558), in his " His-
debilitated, fearful, and half dead, he is made a rich prey, especially for his
Teeth, that are very pretious amongst the Scythians, the Moscovites, Eua-
aiana, and Tartars, (as Ivory amongst the Indians) by reason of its hardness,
whiteness, and ponderousnesse. For which cause, by excellent industry of
Artificers, they are made fit for handles for Javelins : And this is also testi-
f'ed by Mechovita, an Historian of Poland, in his double Sarmatia, and
FuuIm Jovius after him, relates it by the Relation of one Demetrius, that
was sent from the great Duke of Moscovy, to Pope Clement the 7th." — Loc.
cit., pp. 231,232.
" "Scotorum Regni Descriptio, p. 90," as cit«d by various writers.
t Herberstain, as cited by von Baer, 1. c, p. 111.
tSee Hakluyt's Voyages, vol. iii, ed. 1810, pp. 237,238,242,249,254, etc.
\S See Hakluyt's Voyages, vol. i, ed. 1599, p. 237.
II Olaus Magnus's figures will be noticed later under the section devoted
to the fi};ure8 of the Walrus (posted, p. 92 et seq.).
genp:ral history.
85
toriuiii Aiiiinalium" (in the volnino devoted to the " Animalia
AquatiliJi "), tiiitlifully copied all ofOlaiis Magnus's figures under
the heading "De Cotis," and then presents, under the name
Ensmonis, the figure of the Walrus from Olaus Magnus. This
figure, however, he judiciously criticises, stating that the tusks
should be in the upper Jaw, and not in the lower, as they were
represented by Olaus Magnus. This last-named author, in the
later editions of his work " De Gentium Septentrionalium Con-
ditionibus," etc. (as in that of 15G3), rightly places, iiecording to
von Baer, the tusks in the upper jaw. Gesner (continues von
Baer) knew only the first edition of this work, and took his
figure from the above-mentioned " Tabula Terrarum Septen-
trionalium." Also were unknown to him the aecounts of the Wal-
rus given by "Herberstain, Chancellor, and Othere," so that he
made extracts from only Michovius and Albertus Magnus. He
also knew no better than to offer, as a figure of the Walrus, a
drawing he had received from Strassburg, representing, pretty
fairly, the head and tusks, while the rest was purely a fabrica-
tion. Some rhymes, which he further inflicts upou his readers,
show clearly how " awful" the conceptions of the Walrus then
were (or, as von Baer puts it, " Wie schauerlich nocli die Yor-
stellungen vom Wallrossewaren").* |, >, v ; .
In 1008, a young living Walrus was taken to England, having
been captured on Bear or Cherie Island off the coast of Nor-
way, t while four years later (1012) another yf ung Walrus, with
the stuffed skin of its mother, was taken to Holland. The first
appears to have been very intelligently described by ^lius Ever-
hard Vorstius, whose description is quoted by De Laet. J The
specimen taken to Holland was well figured by Hessel Gerard,
the young one doubtless from life, the figures being pu^^lished
by him in 1613, § and subsequently repeatedly copied (as will
be more fully noticed later).
In 1625, Purchas, in his history of the voyages of the English
to Cherie Island and Spitzbergen (then called "Greenland"),
gives much interesting information respecting the chase of the
* To ebow what these conceptions were, von Baer cites the passages
ahtady quoted (anted,, p. 81), in reference to the singular misinterpretations
given in Western Europe to the Russian name Moras. See von Baer, 1. c,
p. li:{. .
t Recuoil do Voy. au Nord, 2^ 6A., tome ii, p. 368.
t Nov. Orb. 8. Doscrip. Ind. Occ, 1633, p. 41.
^ See von Baer, 1. c, p. 128; Gray, Proc. Zool. Soc. Lond., 1853, p. 115.
86
ODOBiENUS ROSMARUS — ATLANTIC WALRUS.
Walnis at these islands, and in one place a quaint description
and some very curious figures of the animal.*
In J 07.5, the Walrus was again described and wretchedly
flgured by Martens, t who is said to have been the first " natu-
ralist" who ever saw the Walrus in its native haunts. Zorg-
drager, | in 1720, supplied by fiir the fuliost account of these
animals, as observed by him in Spitzbergen, that had appeared
up to that date. He gives not only a quite detailed and truth-
ful account of their habits, especially under persecution, but also
of their wholesale destruction at that early time in the Spitz-
bergen seas, and of their extermination at some of the points
at which they had formerly been accustomed to land in immense
herds. He also notes the increasing difl&culties of their capture
owing to the great shyness of man they had acquired in conse-
quence of persecution, and describes the manner in which they
were captured, and also their products. Copious extracts from
Zorgdrager's account of the Walrus are given by Buffon (trans-
lated into French from a German edition), and he has also been
extensively quoted by even much later writers.
The Greenland Walrus was described by Egede§ in 1741, by
Anderson II in 1747, by Ellis If in 1748, by Cranz** in 1765^
and by Fabricius tt in 1780, some of whom added much infor-
mation respecting its habits and distribution, its usefulness t'j
the natives and their ways of hunting it, as well as respecting
its external characters.
The above-cited accounts of the Walrus formed the basis of
numerous subsequent compilations, and most of those last given
are cited by the early systematic writers, few of whom, as pre-
^^ou8ly shown (see antea, pp. 8-11), had any just appreciation
of even its most obvious external characters. Linn6, as already
noted {anted,, p. 8), profited little by what had been written
by preceding authors, while Brisson, Erxleben, and Gmelin
manifest a scarcely better acquaintance with this badly misrep-
resented and poorly understood creature. No little confu-
sion has hence arisen in systematic works respecting its posi-
* See anted,, p. 74-78, and. posted.
t Spitzbergen, pp. 78-83, pi. P, fig. h.
t Bloeyende Opkonist der Alonde en Hedendaagscho Groenlandsclio Vis-
Bchery, etc., ed. 1720, pp. 165-172.
$ Det gamle Gr(^nland8 nye Perlustration, etc., 1741, p. 45.
II Nachrichten von Island, Gronland und der Straw e Davis, p. 258.
IT Voyage to Hudson's Bay, p. 134.
** Historie von Gronland, pp. 165, 167.
+t Fauna Groenl., p. 4.
■I (
GENERAL niSTORY.
87
tion and aainities (see antca, pp. 7-12). The accounts by Hoiit-
tuyn, Buftoii, Pennant, P. S. L. Miiller, and Sclireber are excel-
lent for their time. These authors all recofjnized the close
relationship of the Walrus to the Seals, and (piite con-ectly
indicated its external characters and habits. Some of these
accounts, however, include references to both species.
Daubenton, in Buffon's "Histoire Naturelle,"* gavo a de-
scription and figure of a Walrus's skull, and made the first
contribution to our knowledge of its internal anatomy, based
on the dissection of a foetal specimen.
Since the beginning of the present century, the Walrus liaa
been the subject of almost numberless notices, as well as of sev-
eral elaborate papers, devoted in most cases to special points
in its anatomy, very few of which need be here enumerated, t
The elder Cuvier, beginning with his " Legons d'Anatomie
compar^e " (1800-1805), and ending with the third edition of his
''Ossemens fossiles" (1825), contributed considerably to our
general knowledge of its structure and affinities, especially of
its osteology ; he in 1825 1 first figuring and describing its skel-
eton. A paper by Sir Everard Home, § in 1824, figured and de-
scribed the stomach and feet from specimens taken to England
from Hudson's Bay, preserved in salt. This paper is noteworthy
mainly on account of the singularly erroneous interpretation
there made of the structure and functions of the feet, Home
supposing that these organs were provided -with sucking discs,
by means of which th« creature was enabled to adhere firmly
to the ice in climbin ^. The skeleton of the Walrus was again
figured and described by Pander and d'Alton|| in 182G, and
still later by BlainviUe ^ about 1840. Von Baer, ** in 1835,
published some account of the axterial system of the Walrus,
based on a dissection of a young specimen. Its general anato-
my, especially its limb-structure, myology, vascular and respi-
ratory syste. , viscera and generative organs, and external cha-
* Tome xiii, 17*', pp. 415-424, pll. liv, Iv. The skull had been previously
figured by Houttuyu (in 1761), as will be noticed later.
t Those relating to its dentition have been already noticed in detail (see
anted, pp. 47-57) ; several others have also been specially referred to, and
nearly all are cited in the references given at pp. 23-26.
t Ossom. Foss., 3« dd., tome v, ii™" pt., pp. 521-523, pi. xxxiii.
$ Phil. Trans., 1824, pp. 233-241, pi. iv.
II Skeloti der Robben und Lamantine, pll. 1, ii.
IT Osteographie, Des Phoques, pll. i and iv.
** M6m. de I'Acad. St. Pdtersb., vi"" s4r., Sci. mpt?i., phys. et nat., tome
')■"«. 1835, pp. 199-212.
88
ODOB^.NUS ROSMARUS — ATLANTIC WALRUS.
racters, Avere quite fully and satisfactorily treated by Dr. J.
Murie* in 1872.
lUiger, in 1811, in a paper on the geographical distribution of
the mainmals of the Northern Hemisphere (see antea, p. 18), first
nominally r cognized the Pacific Wah'us as a species distinct
from tie Atlantic animal, while Fremery, in 1831, recognized
three species, and Stannius, in 1842, admitted two,t but, as
already noticed, only one species of Walrus has been commonly
recognized. The matter of variation dependent upon sex, age,
and individual peculiarities, has received, as already noticed
(see anteu,, pp. 38-43), special attention at the hanas of Wieg-
mann, Stannius, Jaeger, and other writers.
Unquestionably, the most important paper relating to the lit-
erature, geographical distribution, and habits of the Walruses
is the well-known an 3. Geziihmte Rob-
ben. § 4. Wahro Cetaceen. ^^ .'>. Gesellschaftliches Leben. ^ 6. Liebe dei'
Aeltem zu den JuugCii und dor Jungon gegen die Aelteru. ^ 7. Gatten-
liiebe. $ 8. Allgemeine Begriinduug dieser Verhaltnisse.
Cap. V. Verbreitung dor Wallrosso (pp. 172-204). ^1. Sio wohnen in zwci
getrcnnten Verbreituugs-Bezirken. 'J 2. Ocstlicher Verbreituugs-Bezirk.
GENERAL HISTORY,
89
published in 1837. This elaborate memoir, so often already
cited in the prescsut article, gives a general summary of nearly
all papers, referenees, and figures relating to the Walruses that
api)cared prior to 1835, the date of its presentation to the Impe-
rial Academy of Sciences of Saint Petersburg for publication.
It also contains iiumy original biological and anatomical ob-
servations, based on a young living specimen brought to Saint
Petersbmg in 1828, which, surviving for only a week after its
arrival, soon fell into his hands for dissection. *
Von Baer, after a few preliminary remarks respecting the
occasion and objects of his paper, and a few words on the
anatomy of the Walnis, devotes some thirty pages to a critical
and exhaustive historical resume of the literature relating to the
general subject. Then follow some eighteen pages detailing
his observations on the living animal, in which he gives some
account of the few young indiAiduals that had, up to that time,
been taken alive to Middle Europe ; also a detailed account of
the external appearance of the specimen he had examined in
life. lie notes especially its attitudes, movements, and limb-
structure, and compares it in these points with the Seals. After
describing the position and character of the limbs in the Seals,
and the restriction of their movements on land to a wriggling
movement, with the belly lying on the ground, he refers to the
freer use of the extremities possessed by the Walrus, Avhich he
found was able to truly stand upon its four feet, and says that,
i 'S. Westlicher Verbreituugs-Bezirk. $ 4. Periodische Wanderuugen der
Wallrosse. ^ 5. Pliysischo VerliuUnisse, welcho die Verbreitung der Wall-
roHse bedingou.
Cap. VI. Ehemalige Verbreitung der Wallrosse (pp. 20.'>-228). $ 1. Mei-
uuugen hieriiber. ^ 2. Veriinderungen ini Vorkommeu der Wallrosse in den
drei letzten Jahrhunderten. $ 3. Ob an den Orkadischen Inseln Wallrosse
bis ins 16te Jalirhundert sich aufgebalten haben 1^4. Beweiss, dass, so weit
liistorische Nachrichton zuriickgehen, kein Wallrossfang an der Kiiste von
Lappland getrieben worden ist. $ 5. Ob die Wallrosse im Mittelalter bei
Island liiiufig waren. $ 6. Verbreitung der Wallrosse zur Zeit der Romer
uud Griechen. $ 7. Ehemaliges Vorkommen an der Nordkiiste der Conti-
uente. . t , , .
Cap. VII. Paarung (pp. 228-230).
Cap. VIII. Nabrung der Walbosso (pp. 231-233).
Cap. IX. Stellung des Wallrosses im Systeme, oder Verwandtscbaft mit
andorn Thieren (pp. 234, 2.35).
* He seems, however, to have never published in full the results of his
observations upon it.s anatomy, he apparently reserving the anatomical
part of his memoir in the liope of perfecting it through the study of addi-
tional m.aterial.
90
ODOBiEXUS ROSMARUS — ATLANTIC WALRUS.
in respect to the use of its limbs, it occupies an intermediate
place between the Pinnipeds and the ordinary Ibnr-lboted IMam-
mals, amony which latter its less pliant feet jiive it the appear-
ance of a crijiple. If we should call, he says, the Seal a crawler
or slider, we should have to term the Walrus a waddler, since in
walking it throws its pluuij) body to the right and left. Here avc
have fairly described, lor the first time, the tlexibility of the ex-
tremities, — the bending of the hind feet sometimes forward,
sometimes backward, and the free turning of the fore feet, —
although an allusion was made to this by Vorstius* two centu-
ries before, yet the fact of flexibility remained generally unre-
cognized till 1853, when a young living specimen reachetl London,
Yon Baer points out the fallacy of Sir Everard Home's notion
that the feet of the Walrus are provided with suction -discs, ami
the "blowing-' of the Walrus mentioned by Martens, \.-ho de-
scribed it as throwing wattr from its nostrils like a whale.
Following this chapter on its external features, movements,
temperament, behavior, etc., is an interesting dissertation of
some twenty or more pages on the domesticability of the marine
mammals in general, which is devoted largely to a history of
the behavior of the Seals in captivity, with a short notice of
the different examples of the Walrus, the Sireniaus, and the
smaller Cetaceans that had been observed in confinement. The
next thirty pages are given to a discussion of the geogTaph-
ical distribution of the Walruses, the treatment of which subject
is marked by the same pains-taking research that characterizes
the other parts of this learned monograph. He shows that
Walruses are confined to two widely separated habitats, and
not, as previously supposed, found all along the Arctic coasts.
He describes them as limited to two regions, an eastern and a
western, the first including, the northwestern coast of North
America from the Peninsula of Aliaska northward, and the
corresponding parts of the neighboring Asiatic coast. To the
eastward he could trace them only to the \icinity of Point Bar-
row, and to the w^estward only to a few degrees beyond East
Cape.
The western region, he affirmed, embraces only the Arctic
coast of Europe eastward to the mouth of the Jenesei Biver,
* "Pedes nnterioreH antrorsum, posteriores retrorsum siiectabant cmii iii-
grederetur," says Vorstius as quoted by De Laet (see anted, p. 37). The hind
feet are also represeutcd as turned forward in Hessel Gerard's figure, pulj-
lished in 1C13 (see posted).
GENERAL HISTORY.
91
and, on the other side of the Atlantic, the shores of Greenland
and Arctic America westward to the western shores of Hudson's
Bay a 'd Fox Channel. There is thus left between these two
regions nearly the whole of the coast of Asia bordering on the
Polar Sea on the one hand, and almost the whole of the coast
of Xorth America formed by the Arctic Sea on the other.
In the later portion of his chapter an the distribution of the
Walruses he devotes a few pages to a consideration of their
migrations, and the physical causes which limit their distribu-
tion. Their migrations, he believes, are very imperfectly known,
but he inclines to the opinion that they only periodically visited
such points in their former range as Sable Island and other
southerly lying islands. The causes which limit their range he
considers to be mainly temperature, since he finds the southern
boundary of their distribution is deflected northward and south-
ward in accordance with the curves of isothermal lines.
The former range of the Walruses is also considered at length,
to which subject are devoted nearly twenty-flve pages. A short
account is given of their reproduction and food, the paper clos-
ing with an inquiry into their systematic relationship to other
animals. The map accompanying his memoir shows not only
the distribution of the Walruses as at that time known, but
indicates also the region over which they are known to have
formerly occurred, and also the habitat of the Bhytina, or Sea-
cow of Steller.
The reception in London, in 1853, of a young living Walrus
gave rise to a paper by Owen* on its anatomy and dentition,
and another by Gray,t " On the Attitudes and Figures of the
Morse." A short paper was contributed by Sundevall J in 1859
on its general history.
Leidy, in 1860, published an important paper on the fossil
remains of Walruses found on the eastern coast of the United
States, while Gratiolet, Defrance, Lankester, and Van Beneden
have also written about those that have been met with in
France, England, and Belgium.§
Malmgren, in 1864, in a paper on the Mammalian Fauna of
•Proc. Zoijl. Soc. Lond., 1853, pp. 103-106.
tibicl., pp. 112-116, figs. 1-10.
tOm Walrosseu, Ofversigt K. Vet. Akad. Forb. (Stockh.), xvi, Ib.j'J, pp.
441-447 ; also translated in Zeitschr. gesammt. Natnrw. Halle, xv, 1860, pp.
270-275.
ij See anted, pp. 61-65.
92
ODOB^NUS ROSMARUS ATLANTIC WALRUS.
Fiumark and Spitzbergen,* jiublisbed many interesting notes
relating to its babits and food, and biter a special paper on its
dentition (noticed anteu, p. Hi.) Malmgren's observations on
tbeii" bal)its, distribution, etc., also appear in tbe history of the
Svredisb Expedition to Spitzbergen and Bear Island in the year
1801,t together with a somewhat detailed and very interesting
general history of the animal, with several illustrations.
Brown, in 1808, in his " J^^otes on the History and Geogi'aph-
ical Kelations of thePinnipedia frequenting the Spitzbergen and
Greenland Seas,"^ devotes several pages to the Walruses (pi).
427-435), in which he considers especially their habits and food,
geographical distribution, and economic value.
In addition to the special papers cited in the foregoing pages,
their general history has been more or less fully presented in
several general works treating of the mammalia, and in several
faunal pubhcations, § Much information respecting their general
history may also be I'oiind in the narratives of various Arctic
€xi)lorer8, as Parry. Wrangell, Keilhau, Kane, Ilayes, Lamont,
and others, whose co"tributions will be more fully noticed in the
following pages relat i ; to the habits of the Walruses.
2. Figures. — As von Baer fiicetiously remarlcs, no animal
has had the honor of being depicted in such strange and widely
diverse representations as the Walrus. These, as has been
previously stated, began with Glaus Magnus, about the middle
of the sixteenth century, who opened the series with half a
dozen phantastic flgui'es, based apparently upon this animal,
only one of which, however, bore the name liosmarus {Bosnia-
* lakttagelser och antcckuiugar till Finmarkcns och Spetsbergens Diigg-
ecially the anterior.
Another, called Vacca marina, represents the head of an ox,
with a long beard on the chin. A fourth represents a dolphin-
like body, with four feet, a fish's tail, a pair of long, ascending,
*My remarks respecting Olaus Magnus's figures are based on Gesnei's
(Hist. Animal. Aquat., 1558, pp. 247-249), and Gray's copies of tbem (Proc.
Zool. Soc. Loud., 1853, p. 113), Olaus Magnus's work not being accessible to
iiM'. The figures herewith given (Figs. 4-12) are from electros of Gray's
lignrt's.
t Sec Fig. 4, copied by Gray from Ohu.nic literature of
the Walrus by furnishing a figure, respecting which he says :
"I shall not detain the Reader with an Account of a Creature
[" Sea Horse"] so often described, b*t refer him to the Cut, in
which he will find^it very truly repre8ented."§ The figure, how-
ever, is one of the worst imaginable, considering the oppor-
tunity Ellis evidently had for observation. In some respects
it bears some resemblance to that given by De Yeer. EUis's
figure combines a Lynx-Uke f ace withlldon-liko fore limbs, short,
* IJloeycndo Opkonift der Aloude cu Hedendaagsche Groenlandscho Vis-
schciy, 1720, plate facing p. 1()2, upper left-hand figiu-e.
tBfscLrcibung inid Natnrgeschichto von GriJnland, 1763, p. 106, pi. vi,
lower left-hand ligiirc, Knniitz's German translation. The wo .: appeared
in Danish as early as 1741.
{ Voyage to Hudson Bay, pi. facing p. 134, middle figure.
$Loc. cit., p. 236.
100
ODOBiENUS ROSMAUUS — ATLANTIC WALRUS.
roiiiid, pi'oiiiiiH'iit fill's, Miiiall, [jointcd, inwardcurviii^' tUMkH, no
liitHl t'(\ ',, jind a body tapeiiii}; to a doubly t'liiaiyiiiatc tish-liko
tail, poH^iblv intended to i'(^]ueHt'.it hind lind)s.
J'ontoppidon, in liis Natnral J li story of Norway, published
in IT.')!, ^ave a tljfuie of the WaLns in whieh the resenibhniee
consisted mainly in the presenei! of two huge tusks in the upper
jaw. Only tlu! head, neek, and upper portion of the body are
re]uesented ; but the {general outline, as far as seen, is sug-
gestive «)f the animal it was intended to represent.
Houttuyn,* in 17(Jl, gave a very fair figure of the skull and os
I)enis of a Walrus. As 1*. L. S. Miiller, in 1773, used Houttnyn's
plates in his " Natursystem," these llgures are there again called
into service, to which was added a noteworthy representation
of the animal.t This represents an apparently young Walrus
as lying partly on the side, with the diminutive hind feet
Fig. 11.—" Wall-Iioaa, Marten's Spitzbergen, &c. 1675, t. P, tig, ft. (Reduced
threo-tenths.)"
turned foricard. The general outline of the body indicates
the obese form of the Walrus; but the head, with its small,
short tusks, has scarcely the faintest resemblance to the head
of that animal.
* Natuurlyke Tlistorie of oitvoerige Beschryving der Dieren, Planten en
Mineraalon, volgens het Samenstel van den Heer Linnaeas. Met naaw-
keurige Af beoldingeu. Eerste Deels, tweede Stuk, 1761, pi. xi, figg. 1, 4.
t Des Hitters Carl von LinntS Koniglich Schwedischen Leibarztes, &c. &c.
vollstUnih^'is Natursystem nacb der z wolf ten lateiuisclieii Ausgabe und nacb
Anleitung des holliindisclien Houttuynischen Werks mit einer ausfUhrliohen
Erklarnng ausgefertiget von Philipp Ludwig Statins Miiller, etc. Erster
Theil. Niimberg, 1773. PI. xxix, fig. 2. This is one of the few original
plates added by Miiller to Houttnyn's series
FKirUKS.
101
111 170.'), ii most wrotc'lu'd iiiid IndicrouH cariciitiirc of tho
Wiilriis was coiitrihutcMl l).v ^Murtciis.* In tills lijiuro, tlio iiiucli-
alms«'(l Walrus is iepivs(Mito«l as liaviiif,' an oiHnnioiisly largo
and sliap«'less head, in which tho small tusks are sot widely
up "t : it has small Seal-like tore feet, and no hind lindis, or, if
prf M tit. lliey are directed backward, and look more like a fish's
tail tiian distinct limbs. The tusks alone give tho figure any
suggestion of what it wa,« intended to represent.
The next figure of which I have knowledge was published by
Buffon,t also in 170a, and soon after copied by Schreber.J This
Fig. 12.— "y.c Morse, Huffon, xili, t. 548, 1765. (Reauced two-fifths.)"
was evidently drawn from a stuft'ed specimen, to which the taxi-
dermist had given the attitude cand general form of a common
Seal. In 1827, a very fair figure of tho head (the animal being
sup])osed to be in the water, with only the head visible) was
published in Griffith's Animal ICiugdom (vol. ii, pi. v), which
was later repeated by lIamilton,§ and also elsewhere. In 1830,
a very fiiir, colored figure (evidently from a stuffed specimen),
barring the posterior direction of the hind limbs, appeared in the
" Disciples edition '' 1 1 of Cuvier's E6gne Animal, copied from Pal-
* Spitzbergische Eeisebesclireibiuig, pi. P, fig. i. This fig. is also repro-
duced by Gray (1. c, fig. 7), and is here copied as Fig. 11.
tHistoire Niturelle, t. xiii, pi. liv.
t Siiuget., pi, Ixxix.
$ Amphibious Caiaivora, p. 106, in Jard. Nat. Library, Mam., vol. viii.
II Le E^gne Animal, etc., par Georges Cuvier. "Edition accompagn^e des
planches gravies, .... par une reunion de disciples de Cuvier," etc. Paris,
1836 et seq.
The Walrus is figured in "Mammif feres," pi, xliv. The history of the
figure is given as follows: " Figure dessiii^e d'apriis cello qu'a donn^e Pal-
las dans la Zoographia Rosso -Asiatica, et r^formde, pour lo pose, d'aprfes un
croquis in<5dit de Choris; an vingtifeme environ de la. gi-andour uaturelle,"
The only copy of Pi^Uas's "Icones" accessible to me i.? imperfect, and has
not the figure here copied. There i,;, however, a quite dLforent one, which
will bo noticed later in another connection. Whether Fallas's figure hero
copied represents tho Atlantic or the Pacific species cai:not well be deter-
mined.
102
ODOBiENUS EOSMAEUS — ATLANTIC WALEUS.
las. Another much like it was publislied soon after in Macgil-
livray's British Quadrupeds, * and still another, also quite simi-
lar, in Hamilton's Marine Carnivora.t The vignette-titlepage of
the last-named Avork also rei)resents a "Walrus hunt," in which
a boat's ciew are depicted as attacking a gioup of live old Wal-
ruses. The plate in Hamilton's "Amphibious Carnivora " pui'-
ports to have been drawn from a specimen in the Edinburgh
Eoyal Museum, and seems to be essentially the some as that
in Macgillivray's British Quadrupeds, with a somewhat altered
position and different background. In each of these plates
are represented two other distant figures of the Walrus. In
each, the tusks are long, and seem to represent the Pacific
rather than the Atlantic species, as is also the case in the "Dis-
ciples edition" of the Eegne Animal. In all these last-named
figures, the hind limbs are directed posteriorly, but in other
respects they are fair representations.
Dr. Kane, | in 1856, gave several illustrations of the annual,
and also of its breathing-holes, and of the implements employed
by the Innuits in Walrus-hunting. In Sonntag's "Narrative
of the Grinnell Exploring Expedition," § published in 1857, a
* Jardiue's Nat. Library, Mam., vol. vii, 1838, pi. xx.
tibid., vol. viii, 1839, pi. i.
t Arctic Exploration, vol. i, jip. 141 ("Walrus Sporting"), 142 ("Wah-us-
hole), 419 (''portrait") ; vol. il, plate facing p. 214 ("Walrus Hunt off Pi-
kantlik" — a nearly full tigure.
$Thi8 curious and apparently little known brochure, by the eminent
astronomer of the Expedition, is well worthy of attention, notwithstanding
the ludicrously sensational character of the titlepage affixed by the en-
terprising publishers. The titlepage, transcribed in full, is as follows:
"Professor Sonntag's Thrilling Narrative of the Grinnell Exploring Expe-
dition to the Arctic Ocean, in the years 1853, 1854, and 1855, in search of
Sir John Franklin, under the command of Dr. E. K. Kane, U. S. N. Con-
taining the History of all previous explorations of the Arctic Ocean, from
the year 1618 down to the present time ; showing how far they advanced
northward, what discoveries they made and their scientiiic observations.
The present whereabouts of Sir John Franklin and his party, if they are
still alive. A statement of the only practicable method by which the North
Pole may be reached; the reasons why all exploring expeditions have
hitherto failed to penetrate the icy barriers of the Polar Eegions. Highly
important astronomical observations, jiroving that there is no such thing as
apparent time at the North Pole ; sulferings of Dr. Kane's exploring party;
how they Avere buried for two years in the ice, enduring a degree of cold
never experienced by any human being before; their miraculous escapes
and unprecedented hardships ; their abandonment of the ship ; and perilous
journey of four hundred miles over the ice. With nearly one hundred
Bplendid engravings. By Professor August Sonntag, Astronomer to the
FIGURES.
103
yioui) of four old AValruses is figured (full-page woodcut, p. 113).
The animals are disposed in various attitudes, and represent
admirably tlie grim visage, postures, and uncouth proportions
of the Atlantic Walrus. The figure in the foreground is pre-
sented in inoflle, with both fore and hind limbs in a natural
pimtion; behind this are two old veterans seen in half-profile,
and behind these a third lying on its back with the hind limbs
thrust upward. Tliis illustration, evidently a study from life, is
by far the best representation of the adult Atlantic "Walrus with
which I am acquainted. In 1857, ;!>i'. Gray reproduced, as pre-
viously detailed {antea, pp. 93-100), i series of the early figaires
from Olaus Magnus, Gesner, Jonston, Gerard, Martens, Buffon^
and Cook.
The next original figures of the Walrus with which I am
acquainted were drr.wn from the living specimen in the Gar-
dens of the Zoological Society of London by Mr. Wolf, and
appear in Wolf and Sclater's " Zoological Sketches,"* published
in 18G1. In plate xviii is represented a group of Walruses in
various attitudes. Those in the foreground are young and tusk-
less, with a heavy array of long mystacial bristles, and much
thinner necks and shoulders than the Walrus is commonly repre-
sented as having, doubtless owing to the very emaciated condi-
tion of the Uving original.
At about this date (1861), some very good pictures of groups
of Walruses were published by Mr. Lamont ir his entertaining
and instructive book entitled " Seasons with the Sea-horses."
In a spirited plate (called " Chase of the Walrus"), facing the
titlepage, is portrayed a group of Walruses in the sea, attacked
Expedition, formerly of the Royal Observatory at Vienna, and late of the
U. S. National Observatory, Washir, ^n City, D. C. Philadelphia, Penn. :
Jas. T. Lloyd & Co. Cincinnati, Ohio: Jas. T. Lloyd & Co." No date.
Large 8vo, pp. 176, paper. Copyright dated 1857.
The publishers state: "The undersigned having purchased Professor
Sonntag's Narrative of the Grinnell Expedition, some months sinco, have
used their best judgment and abilities in preparing this thrilling nairativo
for the press, to make it as acceptable to tlio . ^idiug public as possible," etc.
The name of the author is alone suflQcient guaranty of the trustAvorthy
and iustructive character of the work, which, despite the dime-novel aspect
of its exterior, is a valuable contribution to the history of the Arctic Re-
gions. Pages 80 to 85 are devoted to a general account of the Walrus At
page 83 is a sketch of a "Desperate attack of Walruses on the English
Boat," based apparently on Captain Becchey's account of an adventure with
these animals.
*Vol. i, pi. xviii.
104
ODOIJ^ENUS ROSMAKUS ATLANTIC WALRUS.
by a boat's crew, oue of the poor animals having been already
harpooned. Another plate, facing page 72, entitled " Walruses
on the Ice," represents it .-erd on the ice in various attitudes,
most, but not all, of which have the hind feet extended back-
wartV in the manner of Seals. In his later work, " Yachting in
the Arctic Seas," he has given (plate opposite p. 50) a very flue
side-view of the head, and on p. 221 a large vignette figure of
the head seen in front.
Mr. Brown also refers to " the excellent figures of the Wal-
rus taken by the artist of the Swedish Expedition," namely, a
"chromolithograph and head, both drawn by Herrvon Yhlen," —
" under the direction of such well-informed naturalists as Torell,
Malmgren, Smith, Goes, Blomstrand, &c.," in which "the fore
flippers are represented as rather doubled back, and the hind
flippers extended." This work ("Svenska Expeditioner til
Spetsbergen ar 1861, pp. 168-182, pi. facing p. 169, and head
p. 308 ") I have been unable to see, but presume the figures are
the same as those in the German translation of this work,
which appeared in 1869. * The frontispiece of this work repre-
sents a group of ioiu* old Walruses resting on the ice. with a
fifth in the water in the foreground. A woodcut of the head of
a young, or mote probably a female, is given on p. 132, and on
p. 130 a hunting-scene.
In 1867 appeared figures of the second living specimen
received at the Zoological Society's Gardens. According to Dr.
Murie t these were published in " The Field," " Laud and Water,"
" Illustrated London News," and elsewhere. The figure origi-
nally appearing in " The Field" (drawn by Mr. Wood) is repub-
lished by Dr. Murie in his " Memoir on the Anatomy of the
Wah'us " f from the original wood-block. This is a rather more
robust figure than those published by Wolf and Sclater, but is
likewise tuskless (being also that of a very young animal), and
shows similarly the long, descending, curved mystacial bris-
tles.
In 1870, Dr. Gilpin figured a male Walrus killed in March,
1869, in the Straits of Belle Isle, Labrador. In this figure, the
general form of the body is very well represented, but the hind
* Dio Hchwedischen Expeditiouen nach Spitzbergeu iind Btircn-Eiland,
ausgefUbrt iu deu Jalircu 1861, 1864 und 1868, otc. (for full trauscript of
the titlepago sec ante^, p. 92).
t Trans. Zool. Soc. Lend., vol. vii, 1872, p. 413.
t Loc. cit., p. 416.
riGl'RKS.
105
limbs are tniiied l)a('kwurd, as in the coiiunoii Seals. A view
of tlu! muzzle forms a second liguie, and the I'oim of one of the
fore limbs is given in outline.
Wells, in liis "Gateway to the Polynia,"' iniblislied hi 1S73,
gives a plite (faeing page 201) in illustration of the Wakus.
The ligure in the foreground represents an individual tlat on
its belly with all the limbs directed i)osteriorly. Other figures
represent other individuals reijosing in various attitudes.
The above-enumerated figures of the Walrus embrace all the
original figures of the Atlantic si)ecie8 thus far knoAvn to me, and
all to which I have seen references, so far as figure^ of the entire
animal are concerned. In recapitulation, it may be stated that,
Gesners figure, published in 1558, is the first that had an actual
foundation in nature, all t^e preceding (the mythical ones of
Olaus Magnus) being purely fictitious or based on erroneous
conceptions. Gesner's, as already noticed, was a curious combi
nation of reality with myth, the head only being drawn from
nature, and a fanciful body added ! The first really drawn from
nature ("ad vivum") was Hcssel Gerard's excellent figure pub-
lished in IGIS. Subsequently appeared numerous figures in the
works of travellers, drawn apparently either from memory or by
artists who had never seen the aninml they so confidently
attempted to depict.
The first representation based on a museum specimen api)ears
to have been Bufton's, in 1705, which has been aptly described
as being merely a common Seal with tusks. Other figures fol-
lowed later, as those in the so-called " Discip'es edition" of Cu-
vier's Regue Animal, and in the two already cited volumes of
Jardine's Naturalist's Library, drawn also from stuft'ed speci-
mens, in which the hind limbs were always placed in a whoUy
false attitude, though in other re8i)ects passably fair figures. I^^ot
until a living specimen reached London, in 1853, did the cor-
rect attitudes of the animal and the natural position of the hind
limbs become generally known to naturalists, and not until
then was the tnithfulness of Gerard's early figure duly recog-
nized and appreciated, notwithstanding that von Baer, nearly
twenty years earlier, testified to its excellence, and correctly
described the flexibility of the limbs. Now, through the two
living specimens seen and figured in London, and through
excellent recent figures of tlie Pacific Walrus, the attitudes and
oxternr.l bearing of few of our marine mammalia are better
uiown than thoi-,e of the Walruses.
.UG
( )])OI{^E\L'.S I{( )SMA1{U.S ATLANTIC WALIUIS.
In Jidditioii to the iibovtMlcsi'ribcd ti;;iiros of the general ani-
niiil, rcpresentiitions of viirious iinatoniiciil details, both of the
osteohtgy and the soft parts, have been from tinu^ to time ])ub-
lished. As early as 17(51, the skull, as previously stated, was
liyured by llouttuyn, and ajiain by Daubenton* in ITO.j, these
boinj;' the earliest liyin-es of the skull to vhieh I find referen(X'.
Goethe, in his "Morphoh),';ie" (see antca^ V''^^)i gave impcutant
figures illustrati\'e of tlie dentition and strueture of the ante-
rior portion ot the skull.
Home, t in 1824, published a series of exeellent figures of the
extremities and stomach. G. Cuvier, f in 1jSL»5, figured skulls
and the skeleton, his figures of the skull being also reprodueed
in the "Diseiples" edition of Cuvier's liegne Animal. § Pan-
der and d'Alton, in 1820, in their " Vergleichende Osteolo-
gie,"|| gave an excellent figure of the skeleton and detail illns-
trations of the skull and limbs. In the figure of the skeleton,
the hind feet are turned forward in a plantigrade position, and
the fore limbs are given their natural jiose. Von Baer,^] in
1835, figured the blood-vessels of the limbs, and, in ISIO.
De Blainville •* figured the skeleton and the skull. Gray, tt
in 1850, gave a view of the skull, the same figure being re
peated in some of his later works. || Owen, in 1845, figured
the dentition in his " Odontograi)hy " (pi. cxxxii, fig. 8), tlu'
skull and dentition in 1854, §§ and gave another figure in
1868. nil In 1857, Walrus skulls were figured by Blasius, HJI ami
Leidy*** the same year figured a fossil skull from Monmoutli
County, New Jersey. Later, as already noticed {atttea, p. .")4).
the milk dentition was figured by Malmgren, and also by
* Buft'on's Histoire Nat., tome xiii, pi. Iv. An artistically inucli improvtil
(but unaccredited) copy of Daubeuton's figures appears in Hamilton's "Am-
phibious Caniivora" (Jardine's Naturalist's Library, Mam., vol. viii, jip.
100,101).
t Phil. Trans., 1824, pp. 2:i5-241, pll. iv-viii.
t Ossem. Fossiles.
ij Mam., pi. xliv.
II Lieferuiij; xi. Die llobbeu und Lauiantiue, pll. 1, ii.
f Mdm. de I'Aead. St. Pdtersb., Sci. Nat., vi'n« s6r., 1835, t. ill, pi. — .
**Ostdoj^., De.H Phoqiies, pll. i (skeletou) and iv (skull); eight figures.
tt Cat. Mam. Brit. Mus., p. lU, fig. 11 (small woodcut).
tt Cat. Seals and Wliales. IHOli, p. 3,'), tig. 12.
ijij EueyeJ. Brit., artic^le Odoutogr.apliy, p. 4(i;J, fig. 112.
nil Comp. Anat. and Phys. Verteltr., vol. iii, p. 3;]8, fig. '2 ^5.
Hlf Fauna Wirbelth. Deut.schl., ]ti>. 2{)l, 2()2, figg. 14.5-150.
*** Trans. Amcr. Pliil. Sor. Phila., (2), vol. xi, pll. iv, v.
HABITS AND THE CHASE.
107
Peters. Dr. Murie, in 1874, gave immorous figures illustrative
of its external characters, myology, dentition, generative, di-
gestive, and vocal organs, based on a dissection of the young
AValrns that died in the Garden of the Zoological Society of Lon-
don in 1SG7, tliese being the only figures, so far as known to me,
dii voted to the general anatomy. Doubtless other figures of the
skull, and possibly of the dentition, have appeared that are not
here noted.
Habits and the Chase. — The Walruses are at all times more
or less gregarious, occurring generally in large or small compa-
nies, according to their abundance. Like the Seals, they are ^
restricted in their wanderings to the neighborhood of shores or |
large masses of floating ice, being rarely seen far out in the open I
sea. Although moving from one portion of their feeding-grounds-
to another; they are said to be in no true sense a migratory ani-
mal.* They delight in huddling together on the ice-floes or on
shore, to which places they resort to bask in the sun, pressing one
against another like so many swine. They are also said to repair
in large herds to favorable shores or islands,! usually in May
and June, to give birth to their young, at which times they some-
times remain constantly on land for two weeks together, with-
out ever taking food.| They are believed to be monogamous,
and to bring forth usually but a single young at a time, and
never more than two. The period of gestation is commonly be-
lieved to be about nine months. The young are bom from April
to June, the time probably varying with the latitude. Malm-
gren states that the pairing of the Walnises takes place about
the end of May or the beginning of June ; that the female gives
birth to a single young in May or June ; and that the period of
pregnancy lasts probably for a year. He states that Dr. A. von
(loes found a month-old foetus in the uterus of a female on the
Sth of July, in latitude 80° N., but adds that females with ma-
ture young in the uterus have been taken as late as the end of
* Sco Browu, Proc. Zotil. Soc. Loncl., 1868, p. 433.
t Says Zorgdrager (writiug in 1750), as quoted by Buft'ou, in referring to
tills habit : " Aucieunenient & avant d'avoii- 6t6 pers6cnt<^s, lea morses s'avau-
voltiit fort avant dans les terres, de sorto que dans les bautes mardes ils
1 idii'iit asst'z loin do I'eaii, & quo dans le touq)s dc la basse mer, la distance
jt'tupt fiicoro beaucoup plus grande, on le ubordoit aisdmeut. "—Ifisf. Xaf.,
Utinv xlii, p. IWG.
{ Soe SUuldham, Pliil. Trans., vol. Ivi, 1777, p. 249, quoted anteH, p. C7.
X
108
ODOBiENUS UOSMAUl S ATLANTIC WALRUS.
June or July. The females, he believes, suckle tlieir young for
two years, and that hence not less than three years elapse be-
tween each birth. The feuuiles with their newly-born 5'ouiig
are said to keep alocf from the society of other AValruses, and
that females are never found to be pregnant daring the year
following the birth of their young. Females in the second year
of suckling their young collect in large herds and live apart
from the full-grown males. Of thirty full-grown Walruses killed
by Malmgren's harpooner in Henlopen Straits, in the month of
July, not one was a male. Where the full-grown males were at
this time was unknown, but they were believed by the hunters
to be " on the banks," remote from the land, while the females
with their young sought the bays and open sea near the shores,
the two sexes thus living in separate herds.*
Notwithstanding the explicitness of Malmgren's account,
who no doubt correctly details his own experience in the
matter, there is much rebutting testimony, most observers
reporting that both sexes and the young occur in the same
herds, t The only detailed account of the pairing and repro
* Sco further Malmgren's paper, as translated in Arch, fiir Naturgescb.,
1804, pp. 70-72.
t Says Dr. Kane : " The early spring is the breeding season, ... at
Avliicli time the female with her calf is accompanied by the grim-visageil
father, surging in loving trios from crack tc crack, sporting around the berg-
water, or basking in the sun." — Ardic Exploration, vol. ii, jt. 13L
Dr. Hayes, referring to a herd upon which he made an attack, thus ob-
serves : " Besides the old bulls, the group contained several cows and a fev
calves of various sizes — some evidently yearlings, others but recently boru,
and others but half or three quarters grown. Some were without tusks.
while on others they were just sprouting ; and above this they were of vari-
ous sizes up to those of the big bulls, which had great curved cones of ivory
nearly three feet long." — Open Polar Sea, p. 406.
Lament also refers to the presence of young and old, males and females.
in the same herd, and to the custom of the Walrus-huutcrs of striking a j
young one in order to detain the herd, which, through sympathy, join cod-
certedly in its defense, thus aflfording the hunters opportunity for furthot |
slaughter. — Seasons with the Sea-horses, pp. — .
Through the kindness of Prof. Henry A. Ward, of Rochester, N. Y,, I am !
in receii)t, in answer to inquiries respecting +ho habits and reproduction of |
the Walrus, of the following information from the i)en of Captain Adams.
of the whaling-steamer "Aretiirus," from Dundee to Baffin's Bay. Captain j
Adams, writing from long experience in Walrus-hunting, says: "I am of
opinion that the female Walrus prefers low flats of land on which to biinf
forth her young. The time is in mid-spring. In early May I have seen vcrj
young Walruses on the ice with their mothers. I have also seen afterbirths
on the ice, but atill think that low flat land ispreferrcd when attainable. I d"
HABITS AND THE CHASE.
109
(lu(;tlon of the Walrus is that long since given by Shulrtliaui,
l>as('«l on ()l)scrvations made a century ago at the Magdalen
Islands in tlie Gulf of St. LavTcnce (• .e autcCt, p. 07), to which
he says they repair " early in spring " and immediately bring
fortli their young. Captain Parry states that he met with
females a<'Comi)anied by their young in Fox Channel, Jidy 13,
and Mr. Iianu)nt speaks of meeting with y in succession as carefully as pos-
sible, we did not succeed in getting within harpooning distance
of a single Walrus. Many of them were asleep ; but there were
always some moving about who gave the alarm to their sleep-
ing comrades by flapping them with their fore feet, and one troop
after another manage to shuffle into the sea always just a second
or so in time to avoid a deadly harpoon." t
''With reference to the Walrus," says Captain Hall, " Mr.
Kogers told me that one day, when out cruising for Whales, he
went, with two boats and crews, half way across Frobisher Bay,
and then came to an iceberg one hundred feet above the sea,
and, mounting it, with a spy-glass, took a look all around.
Whales there were none ; but Walrus — 'Why', to use his figu-
rative but expressive words, ' there were millions out on the
pieces of ice, drifting with the tide — ^Walrus in every direc-
tion — millions on millions '."J While these numbers are not,
doubtless, to be taken literally, they certainly imply an immense
number of Walruses. The context states that while the whalers
* Seasons with the Sea-horses, p. 73.
tibid., pp. 80, 81. ; ^ ■,:.!;
t Arctic Researches, etc., p. 234.
Misc. Pub. No. 12 8
114 ODOUiENUS R0.SMARU8 — ATLANTIC WALlvi;...
in Frobisher'H IJtiy liad met with no VVlniU's, "Waliais in any
numbers could l»e obtained, and many liad been secured for
their Hkins and tnsk.s."
The WahiiscM in the Spit/.bergcn waters, according to Mr.
Lamont, usually conpegate in August in great niunbers on
hmd, " sometinu's to the number of several thousands, and all
lie down in «ome secluded bay or some rocky island, and there
remain in a semi-toii)id sort of state, for weeks together, with-
out moving or feeding." They do not usually do this, he adds,
till near the end of August, or some nu)nths later than thoy
were found to do in the sixteenth and seventeenth centuries ou
the shores and islands of the Gulf of St. Lawrence. This is
possibly owing to the difference in tlie climate, although it
seems hardly i>rol)able that this can be the whoh; cause of the
difl'ereiu'e. ]Mr. Lamont, in this connection, makes no reference
to the time of bringing forth of the young, and does 7,iot gi>'e
this as one of the reasons for tlieir ^•isiting the land. lie alludes,
however, to their sudden disappearance at this time from the
ice-lloes. lie says the Wahus-hunters consider themselves
fortunate if they lind one of these resorts, as then they can kill
in a few hours a " small fortune'cj-worth of them." His account
of these " trysting-places," however, is mainly at second hand,
and possibly the date is not carefully given.*
3Ir. Lamont's account of ^A)e great havoc the hunters often
make with the then helpless beasts, destroying many hundreds
in a few hours, is quite similar, so far as the destruction of hfo
is concerned, to the account given by Lord Shuldham of theii-
destruction a century and a half ago at the Magdalen Islands.
Referring to one of the southwesternmost of the Thousand
Islands, Mr. Lamont says : " It seems that this island had long
been a very celebrated place for Walruses going ashore, and
great numbers had been killed upon it at different times in by-
gone years. In August, 1852, two small sloops sailing in com-
pany approached the island, and soon discovered a herd of Wal-
ruses, numbering, as they calcidated, trom three to four thousand,
reposing upon it. Four boats' crews, or sixteen men, proceeded
to the attack with spears. One great mass of Walruses lay in
a small sa '.dy bay, with rocks enclosing it on each side, and on
a little mossy flat above the bay, but to which the bay formed
the only convenient access for such unwieldy aninmls. A great
many hundreds lay on other parts of the island at a little dis-
* Seasons with tho Soa-horses, pp. 173, 174.
HABITS AND TUF. CHASK.
115
taiico. Tlio boiitH liiii(U'«l u little way oil", tso iia not to liis'iten
tlioni, and tlit; Nixtcon men, crccpinjLf alonj^ slioro, not lu'twoen
the sea and tlu^ bay lull of Walruses betbro mentioned, and ini-
iiu'diately ((Uiunenced stabbinj;' the animals next them. The
Waliiis, althou**!! so active ami fierce in the water, is vi-ry nn-
M ieldy and helpless on shore, and those in front soon sueeund)ed
to the lances of their assailants; the i)assa<;e to the shore soon
jjot s() Idocked uj* with the dead and dyiu^' that the unfortunate
wretches belli u«l«',ould not pass over, and were in a manner bar-
ricaded by a wall of carcasses. Considering"' that every thrust
of a lance \>'as worth twenty dollars, the scene must have been
one of terrific excitement to men who had very few or no dol-
lars at all; and my informant's eyes si)arkled as he related it.
lie said the Walruses were then at their mercy, ami they slew,
and stabbed, and slauj-htered, and butchered, and murdered
until most of their lances were renderetl useless, and them-
selves were drenched with blood and exhausted Avith iatigue.
They went on board their vessels, ground their lances, and had
their dinners, and then returned to their sangninary work;
nor cory.
The smell of the island was perceptible at several miles' dis-
tance, and on landing w^e found the carcasses lying as I have
described them, and in one place two and three feet deep. The
skin and flesh of many remained tolerably entire, notwithstand-
ing the ravages of Bears, Foxes and Gulls. So many Wal-
ruses have been killed on this island at different times that a
ship might easily load with hones there. . . ."* The worst
feature of this wholesale slaughter was the fact that their small
vessels, aheady parily loaded, could carry away only a small
portion of the spoil. A subsequent attempt to reach the island
later in the season for the purpose of securing the rest failed,
owing to its being surrounded by impenetrable ice.
Respecting the parental affection displayed by the Walruses,
Mr. Lament relates the following : " 1 never in my life witnessed
anything more interesting and more affecting than the wonder-
ful maternal affection displayed by this poor Walrus. After she
' Seasons -with tlio Sea-horses, pp. 175-177.
116
ODOBiENUS ROSMARUS — ATLANTIC WALRUS.
was fast to the barpoon ami was dragging the boat furiously
through the ict oergs, I was going to shoot her through the head
that we might have time to follow the others ; but Christian
called to me not to shoot, as she had a 'junger' ■ ith her. Al-
though I did not understand liis object, I reserved my fire, and
upon Jooking closely at the Walrus when she came up to bre.ithe,
I then i)erceived that she held a very young calf imder her right
arm, and I saw that he wanted to harijoon it ; but whenever he
poised the Aveapon to throw, the old cow seemed to watch the
direction of it, and interposed her own body, and she seemed
to receive with pleasure several harpoons which were intended
for the young one. At last a well-aimed dart struck the calf,
and we then shortened up the lines attached to the cow and
finished her with the lancet.. Christian now had time and breath
to explain to me why he was so anxious to secure the calf, and
he proceeded to give me a i)ractical illustration of his meaning
by gently ' stirring up ' the unfortunate junger with the butt
end of a harpoon shaft. This caused the poor little animal to
emit a peculiar, plaintive, grunting cry, eminently expressive of
alarm and of a desire for assistance, and Christian said it would
bring all the herd round about the boat immediately. Unfor-
tunately, however, we had been so long in getting hold of om*
poor decoy duck that the others had all gone out of hearing,
and they abandoned their young relative to his fate, which
quickly overtook him in the shape of a lance thrust from the
remorseless Christian.
" I don't think I shall ever forget the faces of the old Walrus
and her calf as they looked back at the boat ! The countenance
of the young one, so expressive of abject terror, and yet of con-
fidence in its mother's power of protecting it, as it swam along
under her wing; and the old cow's face showing such reckless
defiance for all that we could do t;0 herself, and yet such terrible
anxiety as to the safety of her calf!
" This plan of getting hold of a junger and making him grunt
to attract others is a well-known 'dodge' among hunters; and,
although it was not rewarded on this occasion, I have several
times seen it meet with the full mo-asure of success due to its
hiunanity and ingenuity."*
WTien in the water, to again quote from Mr, Lamont, " the herd
generally keep close together, and the simultaneousness with
which they dive and reappear again is remf»jrkable ; one moment
* Seasons with the Sea-horses, pp. 70, 71.
HABITS AND THE CHASE.
117
you see a hundred grisly heads and long gleaming white tnsks
ahove the waves; they give one spout* from their blow-holes,
take one breath of fresh air, and the next moment you see a
hundred bro^\ni hemispherical backs, the next a hundred i)air of
hind liippers flourishing, and then they are all down. On, on,
goes the boat as hard as ever we can pull the oars ; up come
the Seahorses again, pretty close this time, and before they can
draw breath the boat rushes into the midst of them : whish !
goes the harpoon: birr! goes the line over the gunwale: and a
luckless junger on whom Christian has kept his eye is 'fast':
his bereaved mother charges the boat instantly with flashing
eyes and snorting with rage ; she quickly receives a harpoon in
the back and a bullet in the brains., and she hangs lifeless on
the hne: now the junger begins to utter his plaintive grunting
bark, and flfty furious Walruses are close round the boat in a
few seconds, rearing up breast high in the water, and snorting
and blowing as if they would tear us all to pieces. Two of these
auxiharies are speedily harpooned in th*^ir turn, and the rest
hang back a little, when, as bad luck would have it, the junger
gives up the ghost, owing to the severity of his harpooning, and
the others no longer attracted by his cries, retire to a more pru-
dent distance. But for the ' untoward ' and premature decease
of the junger, ihe men tell me we should have had more Wal-
ruses on our hands than we could manage. We now devote our
attention to ' polishing oft"' the two live WaLruses — well-sized
young l)ulls — who are still towing the heavy boat, with theu'
two dead comrades attached, as if she were behind a steam-tug,
and straggling mad' ,' to drag us under the icebergs: a vigor-
ous appUcatiou of the lances soon settles the business, and we
now, with some difl&culty, tow our four dead victims to the near-
est flat iceberg and fix the ice-anchor, by which, ^vith the pow-
erful aid of block and tackle, we haul them one by one on the
ice and divest them of their spoils. . . .
" While we were engaged in cutting up these ^/^alruses, there
were at least fifty more surrounding the iceberg, snorting and
bellowing, and rearing up in the water as if smelling the blood
*It is, perhaps, almost needless to say that the " spouting" hero relx^rred
to is merely the spray thrown upward by the forcibly expelled breath as they
rise to the surface, although a "spouting from their blow-holes" has ujca-
sionally been attributed to them since the time of Martens, who says they
" blow water from their lostrils like a whale." See on this point von Baer
(1. c, pp. ISO-l'i?), who has discussed the matter at length in his above-
cited memoir on the Walruses.
118 ODOBiENUS ROSMARUS — ATLANTIC WALRUS.
of their slaughtered liiend.s, and curious to sec what we were
doing to them now. They were so close that I might have shot
a dozen of them ; but, as they would have heeu siu'e to sink be-
fore the boat could get to them, I was not so cniel as wantonly
to take their lives. AVhen the AValruses were all skinned, we
followed the herd again with success ; and when W{? left oft", in
consequence of dense fog suddenly coming on, we had secured
nine altogether — a very fair morning's bag we thought. . . .
During this morning's proceedings I realized the immense
ad\'antage of striking a junger lirst, when inacticable. This
curious (tlanuish practice of coming to assist a calf in distress
arises from their being in the habit of combining to resist the
attacks of the Polar Bear, which is said ofteu to succeed in kill-
ing a Wahiis. If, however. Bruin, pressed by hunger and a
tempting opportunity, is so illadvised as to snap a calf, the
whole herd come upon him, drag him under water, and tear
him to pieces with their long sharp tusks. I am told this has
been seen to occur, and I quite believe it."*
Capt. William Edward Parry, in his narrative of his second
voyage for the discovery of a northwest passage, makes frequent
reference to the Walrus, and his report of encounters with them
shows that serious and even fatal consequences sometimes re-
sult to the boats' crews who venture to attack them.
" In the course of this day [July 15, 1822, in Fox Channel]
the Walruses," says Captain Parry, "became more and more
numerous every hour, lying in large herds ui)on loose pieces of
(kift-ice ; and it having fallen calm sit one r. M., we despatched
our boats to endeavor to kill some for the sake of the oil they
afford. On approaching the ice our people found them huddled
in droves of from twelve to tiiirty, the whole number near the
boats being i)erhaps about two hundred. Most of them waited
quietly to bo fired at, and even after one or two discharges did
not seem to be greatly disturbed but allowed the people to land
on the ice near them, and, when approached, shewed an evident
disposition to give battle. After they had got into the water,
three were struck with harpoons and kille«l from the boats.
When first wounded they became (juite furious, and one, which
had been struck from Captain Lyon's boat, made a resolute
attack upon her, iuuX injured several of the planks with its enor-
mous tusks. A nuiul)er of the others canu; round them, also
repeatedly striking the wounded animals with their tusks, with
* y«'ii8oii!s with tlio tsea-horses, pp. 81-8;}, 84.
m smm
IIAIJITS AND THP: CHASE.
119
the intention of either gettinj;- tliem iiway or else of joining in
the attaek npon them. Many of these animals had young- ones
wliich, Avhcn assaulted, they either took between their fore-tlip-
])('is to carry off, or bore away on their hacks. Both of those
Idllcd by the Fury's hoats were females, and the weight of the
larjii'st Avas lifteen hundred-Aveight and tAvo quarters nearly;
but it was by no means remarkable for the largeness of its
dimensions. The peculiar barking-noise nmde by the Walrus,
when irritated, may be heard, on a calm day, with great dis-
tuictness at the distance of two miles at least. We found raus-
(luet-balls the most certain and expeditious way of despatching
them after they had been once struck with the hai-poon, the
thickness of the skin being such, that whale-lances generally
bend without penetrating it. One of these creatures, being
accidentally touched by one of the oars of Lieutenant Nias's
boat, took hold of it between its flippers and forcibly twisting
it out of the man's hand, snapped it in two."*
Again, says the same writer, " The Heckla's two boats had
one day a very narrow escajjc in assaulting a herd of these ani-
mals [Walruses] ; for several of them, being wounded, made so
fierce an attack on the boats with their tusks, as to stave them
in a number of places, by which one was immediately swamped
and the other much damaged. The Fury's being fortunately
in sight prevented any further danger ; two of the Walruses
were killed and secured, and the damaged boats lightened and
towed to the shore, from which they had been several miles dis-
tant." t
In addition to the foregoing testimony respecting the power
and courage of these animals when in the water, I add the fol-
lowing : Mr. Lamont states that " a boat belonging to a sloop
from Tromsoe had been upset two or three days before, in our
imniQdiate vicinity, and one of the crew killed by a W.alrus. It
seemed that the Walrus, a large old bull, charged the boat, and
the harpooner, as usual, received him with his lance full in the
chest; but the shaft of the lance broke all to shivers, and the
Walrns, getting inside of it, threw himself on the gunwale of
the boat and overset it in an instant. While the men were floun-
dering in the water among their oars and tackle, the infuriated
animal rushed in among them, and, selecting the unlucky har-
pooner, who, I fancy, had fallen next him, he tore him nearly
*Nuirutivo of Parry's Second Voyage, p. 268. tibid., p. 4159.
120
ODOBiENUS ROSMARUS — ATLANTIC WALRUS.
into two halves with Iiis tusks. The rest of the men save«l them-
selves by clambering on to the Ice until the other boat came to
their assistance.
"Upon another occasion I made the acquaintance of the skyp-
par of a sloop who had been seized by a bereaved c<''\ Walrus,
and by lier dragged twice to the bottom of the sea, but without
receiving any injury beyond being nearly drowned, and having
a deep scar plowed in each side of his forehealetely, l.viny
huddled tofi'ether, h)uu,i>iiij'- in the sun or hizily roUinj;- and
twistiiiji' themselves about, as if to ex])ose some fresh part of
their uuwichly bodies to the warmth — }>r«'at, uj;iy, wallowing'
sea-hoys, they were evidently enjoyinj? themselves, and were
Avitliuut ai>i)rehension of a])i>ioachiny' danger. We lunired them
slowly, with nuiffled oars.
"As the distance between us and the game steadily narrowed,
we began to realize that we were likely to meet with rather
formidable antagonists. Their aspect was forbidding in the
extreme, and our sensations were perhai^s not unlike those
which the young soldier experiences who hears for the first time
the order to charge the enemy. We should all, very i)0S8ibly,
have been quite willing to retreat had we dared own it. Their
tough, nearly hairless hides, which are about an inch thick, had
a singularly iron-plated look about them, peculiarly suggestive
of defense ; while their huge tusks, which they brandished with
an appcaran(;e of strength that their awkwardness did not
diminish, looked like Aery formidable weajmns of offense if
apphed to a boat's idanking or to the human ribs, if one should
happen to find himself Uoundering in the sea among the thick-
skiiuied brutes. To <-oinplete the hideousiu^ss of a facial exi)res-
sion which the tusks rendered formidable enough in appearance,
Xature had endowed them with broad flat noses, which were
covered all over with stiff whiskers, looking much like porcu
pine rAKUs — att.antic walrus.
asleep. This was exliibitiiifja degree ol'eoolness rather alarming.
If they had slioAved tlie least timidity, we should have found some
exeitemeut in extra caution ; but tln'y seenu'd to nudce so light
of our ai)])roa<'h that it was not easy to keep up the bold front
with wiiich we had eonuneneed the adventure. But we liad
come (|uite too far to thiidc of baeking out; so we judled in and
made i-eady for tlu> fray.
'' Beside the old bulls, the group contained several cows and
a lew calves of various sizes, — some evidently yearlings, others
but recently born, and others half or three quarters grown.
Some were without tusks, while on others they were just sprout-
ing; and above this they were of .ill sizes u]) to those of big bulls,
which had great curved cones of ivory, nearly three feet long. At
length we were within a few boat's lengths of the ice raft, and
the game had not taken alarm. They had prol)ably never seen
a boat before. Our jireparations were made as we approached.
The Walrus will always sink when dead, unless held by a harpoon
line ; and there were therefore but two chances for us to secure
our game — either to shoot the beast dead on the raft, or lo get
a harpoon well into him after he was wounded, and hold on to
him until he was killed. As to killing the animal where he lay,
that was not likely to happen, for the thick skin destroys the
force of the ball before it can reach a vital part, and indeed, at
a distance, actually Hattens it; and the skull is so heavy that
it is hard to penetrate with an ordinary bullet, unless the ball
hapi>ens to strike through the eje.
" To Miller, a cool and spirited fellow, w ho had been after
whales on the 'nor- west coast', was given the hariwon, and he
took his station at the bows; while Kuorr, Jensen, and myself
kept our places in the stern-sheets, and held our rifles in readi-
ness. Each selected his" aiumal, and we fired in concert over
the heads of the oarsmen. As soon as the rifles were discharged,
I ordered my men to 'give way', and the boat ^hot right among
the 8tartlefain llall, in his "Arctic llesearches," also thus nuikes
reference to a Walrus-fight in Frobisher Bay: "On their way
back, ^Ir. Lamb, in charge of the second boat, had a fight with
some Walrus in the following manner. Approaching a piece of
ice on wiiich some of these creatures were basking, he attacked
one , as the de
scription went, after the fashion of his sylvan brother, the wild
boar. The st on the ice-Hoes, the ferocious beast
])hui}4cd in foaminji' cinles. si-ekinji' fresh victims in that part
of the sea which was discolored by his blood.
" Some idea may be formed of the lerocity of the AValrns,"
continues Dr. Kane, " from \\w fact that the battle which
3Iortou witnessed, not without sharinj;' some of its danger,
lasted four hours; duriufi; which the animal rushed continually
at the Esquunaux as they ap]noached, tearing off great tables
of ice with his tusks, and showing no indication of fear what-
ever. He received upward of seventy lance-wounds,— Morton
counted over sixty; and even then he remained hooked by
his tusks to the margin of the ice, unable or unwilling to retire.
His female fought in the same manner, but tied on receiving a
lance-wound. The Es(iuimaux seemed to be fully aware of the
danger of venturing too near; for at the lirst onset of the Wal-
rus they jumped back far enough to be clear of the l)roken ice.
Morton described the last three hours as wearing, on both
sides, the aspect of an unbroken and seendngly doubtful com-
bat."*
From the foregoing it appesws that the early accounts of the
courage of the Walrus and its attacking and ev<»n destroying
boats in defense of its young, or in retaliation for an assault,
finds ami)le corroboration. I conclude the abundant evidence
on this subject by the following from the ])en of ]VIr. llobert
* Arctic Kxploiation, vol. i, pp. 414-417.
HABITS AND THE CHASE.
127
Brown, who sa,v«: "When attacked, unliko the othcT Seals
(uiih'ss it be the Vyntophora [Iloodel Seal]), it [the WalniSj will
not retreat but boldly meet its enemies. I wa8 one of a party
ill a boat whieli harpooned a solitary Walrus asleep on a i)iece
oi" i('«'. It immediately divetl, but jnesently arose, and, not-
witlistandinj;' all our exertions with lance, axe, and rille, stove
ill the bows of the boat ; indeed wo were only too };lad to (ait
the line adrift and save ourselves on the tloe which the Walrus
had left, until assistance could rejich us. Luckily for us the
eiirafietl ]\lorse was nmj^nanimous enough not to attack its
oli()i)-falleu enemies, but made otf j>runting iiulignantly, with a
;> an harpoon and a new whaleliue danyliuf"- from its bleeding
llauk^."*
TIh^ foreyoiny i)ayes sufliciently indicate the methods and im-
lileiiients commonly employed in destroyinj,' the Walrus for com-
mercial or other ])uri)ctses. To com])lete the aecount of the
«liase it is only necessiuy to note the special equij)ment of a
Walrus-hunter, and to desciibe the iiiannev of disimsinj;" of the
aiiinial when captiuvd, with a 1»rief account of its products and
tlieii uses. This will be jiiven from Mr. Lamont's work, already
s(i often ([noted, who, in a (jhapter devot(»d to the subject, has
riiniished the only connected and detailed account known tome.
I'l'om this 1 ('onde'is(^ the following':
A well-api)ointe(l Walrus-boat for Ave men is twenty-one feet
lonji by five feet beam, haviu};' her main breadth about one-third
from the bow, and strongly built. She is how-shaped at both
ciuls, and should be light, swift, and strong, and easy to man-
ajic, and hence has the keel well depressed in the middle. She
is always " carvel-built," being thus much less liable to injury
from ice or the tusks of the Wa' uses than if "'clinker-built,''
and easier to repair when daumged. She is braced with thick
and strong stem- and stern-pieces, to resist con(5U8sions with the
ice. There is a deep notch in the centre of the stem-piee^, and ,
three others in a block of hard wood on each side of it, for the^
lines to run through, in addition to which there is also some- 1
times an upright post on the bow for making fast the lines, but
usually the foremost thw art is iiied for this purpose. Each nmn
rows with a i)air of oars hung in grumniets to single stout thole
pins. The steersman rows with his face to the bow, and steers
with his pair of oars instead of Avith a single oar or rudder ; and
each man rowing with a pair of oars enables the crew to turn
* Proc. Zooi. Soe. Loud., WiS, p. 4'^
■■■i
128
f)UOH/ENUS HOSMAUrS — ATLANTIC WAIiRUS.
tho boat much (flicker tlmii it could be done otlu'rwiw, while
the Mhortness of tlu; oars renders them easier to handler and less
in the way amonj;the ie«' than lon;;'erones would be. The har-
pooner rows the bow-oars and is the connnander of the boat, he
ahnie usin^" the weapons and tin' telescope. The strongest man
in the boat is placed next the harpocmer, to haul in the line wlien
a Walrus is struck and to bo the assistant of tlu^ harpooner.
The boats are always painted white outside to assimilate their
(jolor to that of the ice. Each boat ir provided with six har-
poons, placed in racks, three on each side of the bow (inside),
and protected by a painted canvas curtain. To each harpoon
is attached twelve or tifteen fathoms of line, each coiled sepa-
rately in flat boxes under the front thwart, the en'^ beiufr linnly
fastenejl to some strong part of the boat. The liiies should b(^
of the finest (luality of two-inch tarred rope, "very soft laid,"
of the best workmanship and materials. Four shafts for the
hari)oons are usually earned, made of white-i)ine poles about
twelve feet long, and about an inch and a half in thickness,
fitted at one end to enter the socket ol the harpoon. The har-
poons are iised for either thrusting or darting, and a skillful
harpooner is said to be able to secure a Walrus at a distance of
four or five fathoms. When possible, they are thrust into the
victim, and a precautionary twist given in order to disengage
the shaft and more securely entangle the barbs in the monster's
blubber or skin. In addition to the harpoons are usually car-
ried four or five very large lances, with heavy, white-pine shafts
r *■ line feet long, and increasing in thickness from an inch
iialf to two and a half where it enters the socket of the
.e. This is for the double purpose of giving the necessary
strength to the shaft, and to afford buoyancy enough to float
the lance-head in ease it becomes disengaged from the animal,
the lance-head being secured to the shaft by a double thong of
raw seal-skin. Each boat is also provided with five "haak-
picks," or boat-hooks, which may be used in dispatching Seals, as
well as for the ordinary uses of a boat-hook ; also with several
axes, a large one for decapitating the dead Walruses, and a small
^ one for cutting the line in case the Walrus proves too fierce and
mischievous, or in case of accidents ; five or six large, sharp
" flensing " knives ; an ice-anchor, with tackle for hauling the
dead Walruses on to flat icebergs ; lockers supplied with vari-
ous smaller implements and a small outfit of provisions, to guard
against the uncertainties arising from accidents and thick
msm
HABITS AND THE CHASE.
129
woiitbjT. In the way of additional weapons, heavy rifles with
]>h'uty of iuunuinition are considered desirable, and often prove
of iivant service when the Walruses are too wary to permit a
near ai)proach, as often happens. Generally a mast and sail
are, or should be, also carried, though by no means always
needed.*
Ai'cordinj;; to the same writer, the manner of " flensiny;,'' or
taking otf and securing the skin and blubber, is as follows:
The liufie beasts being drawn up on to an ice-floe, the skin, with
the blubber adhering, is then removed by dividing the skin into
iialvest by a slit along the ventral and dorsal lines of the body.
It is then loaded into the boats and taken to the ships and
thrown into the hold in bulk. Afterward, as leisure or oppor-
tunity oft'ers, the skins are drawn up, spread across an inclined
platform erected on deck for the purpose, and the blubber re-
moved. This is done by two men who act as " blubber-cutters,"
clad in oil-skin suits, and armed with large, sharp knives hav-
ing curved edges. The blubber is then dexterously removed
from the skin, c»it into slabs of twenty or thirty pounds' weight,
and thrown down the hatchway, where two men are stationed
to receive it and slip it into the square bung-holes of the casks.
From its oiliness it soon finds its own level in tlie casks, which,
when full, are tightly closed, f
Captain Hall describes the Esquimaux method of taking the
Walrus as follows : " The hunter has a peculia • spear, to whicli
is attached a long line made of Walrus hide ; t^is line is coiled,
and hung about the neck; thus prepared, he hides himself
1. ^oug the broken drifting ice, and awaits the moment for strik-
ing; bis game. The spear is then thrown, and the hunter at
once slips the coil of line off his head, fastens the end to the ice
by driving a spear thi'ough a loop in it, and waits till the Wal-
nis comes co the surface of the water, into which he has plunged
on feeling the stroke of the harpoon; then the animal is quickly
dispatched by the use of a long lance. The recklessness and
cool daring of the Innuit is forcibly shown in this operation, for
if he should fail to free his neck of the coil at just the right
moment, he would inevitably be drawn headlong beneath the
ice."§
* Compiled from Lament's Seasons with the Sea-horses, pp. 43-51.
tin the case of full-grown Walruses; but in the case of "calves," the skin
is left entire.
t Compiled from Lament's Seasons with the Sea-horses, pp. 76, 77.
5 Arctic Researches, etc., p. .500.
Misc. Pub. No. 12 9
130
ODOB^NUS ROSMARUS ATLANTIC WALRUS.
11
"In attacking the Walrus in the water they [the Esquimaux]
use the same gear [as in attacking Whales], but much more
caution than with the Whale, always throwing the katteelik from
some distance, lest the animal should attack the canoe and
demolish it with his tusks. The Walrus is iu fact the only
animal with which they use any caution of this kind."* This
"gear," oi Mtteelik, is said to be the largest-of their weapons,
and to be used only in attacking Whales and Wabuses. It
has a shaft of light wood, about four feet in length, like those
of their weapons used in killing Seals, but the shaft is much
thicker than in the others, especially near the middle, where is
lashed a small shoulder of i jry for the thumb to rest against,
in order to give additional force iu thr wing or thrusting the
spear. The spear-point is of ivory, litter- into the socket at the
end of the shaft, where it is secured by double thongs, in such
a way as to give it steadiness when a strain is put upon it in the
diiectiou of its axis, but provided with a spring that disengages
it when a lateral strain endangers its breaking. To the line
attached to the katteelik a Avholc Seal-skin, inflated like a bladder,
is fastened, for the purpose of impeding the progress of the
animal iu the water when struck, f
Dr. Kane gives a graphic account of a Wakus hunt by a party
of Innuits. They set off with three sledges drawn by dogs, for
the open wat'^r, ten miles distant. As they ueared the new ice,
they would from time to time remove their hoods and listen in-
tently for the animars voice. Myouk, one of the party, becom-
ing convinced, by signs or sounds, or both, that the Walruses
were waiting for him, moved gently on and soon heard the cha-
racteristic bellow of a bull. The party now forming in single
file followed in each other's steps, winding among hummocks
and approaching in a serpentine course the recently frozen ice
spots surrounded by firmer ice. " When within half a mile of
these, the line broke, and each man crawled toward a separate
pool ; Morton on his hands and knees following Myouk. In a
few minutes the Walrus were in sight. They were five in num
ber, rising at '• ^^ervals through the ice in a body, and breaking
it up \ (th an explosive puff that might have been heard for
miles. Two large grim-looking males were conspicuous as the
leaders of the group.
" Now for the marvel of the craft. When the Walrus is above
* Narrative cf Parry's Second Voyage, p. 510.
tSeo Parry's Second Voyage, pp. o07,T)08, and pi. facing j). 550, flgs. 20,21
HABITS AND THE CHASE.
131
water, the hunter is flat and motionless ; as he begins to sink,
alert ami ready for a spring. The animal's head is hardly be-
lov\- the water-line before every man is in a rapid run; and
again, as if by instinct, before the beast returns, all are motion-
less behind protecting knolls of ice. They seem to know before-
hand not only the time he vnM be absent, but the very spot at
which he will reappear. In this way, hiding and advancing by
turns, Myouk, with Morton at his heels, has reached a plate
of thin ice, hardly strong enough to bear them, at the very brink
of the water-pool the Walrus are curvetting in.
'' Myouk, till now phlegmatic, seems to waken with excite-
ment. His coil of Walrus-hide, a well-trimmed line of many
lathoms' length, is lying at his side. He fixes one end of it in
an iron barb, and fastens this loosely by a socket upon a shaft
of Unicorn's [Narwhal's] horn : the other end is already looped,
or, as sailors would say, ' doubled in a bight'. It is the work of
a moment. He has grasped the harpoon : the water is in mo-
rion. Pufling with pent-up respiration, the Walnis is within a
couple of fathoms, close before him. Mj-ouk rises slowly ; his
right arm thrown back, the left flat at his side. The Walrus
looks about him, shaking the water from his crest : Myouk throws
up his left arm ; and the animal, rising breast-high, fixes one
look before he plunges. It has cost him all that curiosity can
cost : the harpoon is buried under his left flipper.
'' Though the Awiik [Innuit name of the Walrus] is down in
a moment, Myouk is running at desperate speed from the scene
of his victory, paying off" his coil freely, but clutching the end
by its loop. He seizes as he runs a small stick of bone, rudely
pointed with iron, and by a sudden movement drives it into the
ice : to this he secures his line, pressing it close down to the
ice surface with his feet.
" Now comes the struggle. The hole is dashed in mad com-
motion with tb ; struggles of the wounded beast ; the line is
drawn tight at one moment, the next relaxed : the hunter has
not left his station. There is a crash of the ice ; and rearing up
through it are two Walruses, not many yards from where he
stands. One of them, the male, is excited and seemingly terri-
fied : the other, the female, collected and vengeful. Down they
so again, after one grim survey of the field ; and on the instant
MyoiLc has changed his position, carrj ing his coil with him and
fixing it anew.
" He has hardly fixed it before the pair have again risen,
132
ODOBJENUS ROSMARUS — ATLANTIC WALRUS.
breaking up an area of ten feet diameter about tlie very spot
lie left. As they sink once more be again changes his place.
And so the conflict goes on between address and force, till the
victim, half exhausted, receives a second Avound, and is played
like a trout by the angler's reel."
The method of landing the beast upon the ice is thus de-
scribed : " They made two pair of incisions in the neck, where
the hide is very thick, about six inches apart and parallel to each
other, 80 as to form a couple of bands. A line of cut hide, about
a quarter of an inch in diameter, was passed under one of these
bands and carried up on the ice to a firm stick well secured in
the floe, where it went through a loop, and was then taken back
to the animal, made to pass under the second band, and led off
to the Esquimaux. This formed a sort of ' double purchase ',
the blubber so lubricating the cord as to adm'i of a free move-
ment. By this contrivance the beast, weighing some seven
hundred pounds, was hauled up and butchered at leisure." *
Referring again to the chase of the Walrus, Dr. Kane says
the manner of hunting varies considerably with the season of
the year. In the fall, when the pack is but i)artly closed, they
are found in numbers about the neutral region of mixed ice and
water, when the Esquimaux assail them in cracks and holes
with nalegeit and line. This fishery, as the season grows colder,
darker, and more tempestuous, is attended with great hazard,
and scarcely a year passes without a catastrophe. The spring
fishery begins in March. Tlxe Walrus is now taken in two ways.
Sometimes when he has come up by the side of an iceberg or
through a tide-crack to enjoy the sunshine, he lingers so long
that he finds his retreat cut off by the freezing-up of the open-
ing through which he ascended. The Esquimaux, scouring the
ice-floes with keen hunter-craft, then scent him out by the aid
of their dogs and despatch him with spears. Again thoy are
found " surging in loving trios from crack to crack, sijorting
around the berg-water or basking in the sun," when they are
^tacked by their vigilant enemies Avith the spear and harpoon.
This mode of attack " often becomes a regular battle, the male
gallantly fronting the assault and charging the hunters with
furious bravery. Not unfrequently the entire family, mother,
calf, and bull, are killed in one of these combats." t
* Arctic Exploration, vol. i, pp, 407-414, 417. t Ibid., vol. ii, pp. 131-133.
PRODUCTS.
133
ruoDUCTS. — The couiiuercial products of the Walrus are its
oil, liido, and tusks. The oil is said to bo much inferior in quality
to that of Seals, but is used for nearly the same purposes.* The
yield is also much less in proportion to the size of the animal, in
the larj;est specimens seldom exceeding five hundred pounds.t
The hide is said to be a valuable commodity, and '' sells for from
two to foiu' dollars per half skin, calves only counting for a half;
it is principally exported to Russia and Sweden, where it is used
to mauufacturo harness and sole leather ; it is also twisted into
tiller-ropes, and is used for protecting the rigging of ships from
chafing. In former times nearly all the rigging of vessels on
the north coasts of Norway and Bussla used to be composed of
Wahnis-skin. [J] When there is a superfluity of the article in the
market I believe it is boiled into glue. It is from an inch to an
inch and a half thick, very pliable in its green state, but slightly
spongy, so that I shoidd doubt the quality of the leather made
from it."§
As noted in the earlier j)ortions of this paper, the tusks were
in very early times a valuable article of traffic among the bar-
barous tribes of Eastern Europe and Northern Asia. Brown
states that "there is said to be a letter in th(; library of the
Vatican proving that the old Norre and Icelandic colonists in
Greenland i)aid theu- ''Peter's Pence' in the shape of Walrus
tusks and hides." || The ivory afforded by the tusks, though
*Lainont says it is usual to mix tlio Seal and Walrus oil indiscriniiuatoly
together, and that "the compound is always exported into Southern Europe
under the name of Seal oil." — Yachting in the Arctic Scaa, p. 89.
t Scoresby states that ho " -lover met with any that afforded above twenty
or thirty gallons of oil." — Account of the Arctic Regions, vol. i, p. 503.
t [In the instructions given to Jonas Poolo by the Muscovie Company in
March, 1610, occurs the following : "And in as much as we have agreed here
with a Tanner for all the Morses hides which wee kill and bring into England,
and have sent men of purpose for the slaying, salting, and ordering of the
same, whereof we have appointed one to goe in your ship : We would have
you reserve the hides, and stooro yonr ship therewith in stead of ballast.
And if you obtayne a greater quantitie then you can bring away with you,
having alwayes regard to commodities of more value, which are Oyle, Teeth,
and Whales flnnes [whalebone], that none of them be left behind ; We would
have you leave the said overplus of hides in some convenient place, till the
next yeere, that we send more store of shipping." — Purchaa his Pilgrimes, vol.
iii, p. 709.]
i Seasons with the Sea-horses, p. 77.
llProc. Zoiil. Soc. Lond., 1868, p. 434.
134 ODOHiENUS U0SMA1UI8 ATLANTIC WALitUS.
Si
inferior in (jUiili*^" to Elephant ivory, is used for nearly the saui*^
])iiri)()ses. It is ^..id, howevc^r, to sooner become yellow by ex-
posure, to be of coarser texture, and lieuce to have less commer-
cial value. I have met with Jio statistics relating; to the amount
annually obtained, or the price it brings in market.*
The llesh of the Walrus is sometimes used as food by Arctic
voyayers, and foi-ms an important article of diet \vith the Esqui-
maux and Tsehuchchis. Captain Hall states that while his
party remained at Cape Tru(5 they were never in want of food.
''Walrus," he says, "was abundant, and was indeed almost
exclusively our diet. We had Walrus bruins for supper; stewed
Walrus, or Walrus boiled, for dinner ; but always Walrus, and
uo bread." f Kichardson states that ''their llesh is preferred
by the Escpumaux before that of the Small Seal {Fhoca JuHjyida),
their feet or tins are considered delicacies, and the heart and
liver were pronounced by our navigators to be excellent. The
tongue is said to be good when fresh, but becomes oily by kv*ep-
iug." I In the narrative of Cook's last voyage it is stated that
the fat of the Pacitic Walrus " is as sweet as marrow," but that
it soon grows rancid unless salted, when it will " keep much
longer." The lean flesh is described as being coarse and black,
and as having a rather strong taste, but the heart is said to be
"nearly as well tasted as that of a bullock." § Captaiii Parry,
in a passage already quoted {antea, p. 119), states that the meat
was not only eaten by his men, but was " eagerly sought after
on this and e>ery other occasion throughout the \oyage, b;\ all
those among us who could overcome the prejudice arisuig chiefly
from he dark color of the llesh. In uo other respect that I
couM <'ver discover, is the meat of the Walrus when fresh-killed
iii the slightest degree oflensive or unpalatable. The heart and
liver are indeed excellent," 11
Food. — The food of the Walruses has long been a subject of
dispute, not less from the varied character of the substances
*Mr. Lainont sajN, rospccting products of flic Walni.s juid their value;
"Cnriosify Icil inc once to wcijfli inid value tlio marketable jjaits of a larj;e
lUill Walrus, au. X\8.
^S Cook's l^ast Voyiige, vol. ii, p. 457,
II Narrative of Pairy's Second Voyage, p. 268.
FOOD.
135
luiiiid ill tlioir .stomachs l»,v (liU'cicnt ob.scrvcrs than from tho
IMciiIiarrontbriuiitiou of (hoirtciUh. i\Iartens,.jiKlg'injj^ from tho
apiK'araiico of their cxcroincnt, thought it must subsist mostly
upon si'a-jiTass. Auclcrson, liowcver, correctly stated that they
siiltsistcd uiion ^lolhisca, Avhich tliey (»l)laiiied from tho bottom
of the sea by dijifiiuf;- Avith their t usks. Crauz also says its food
scciiis to (;onsist \vholly of ''muscles and such kind of shell-
lish" and "sea-grass." F. Cuvier, Bell, and others, thought tho
dentition indicated tliat their diet must be mainly, if not wholly,
\ cgetable. Most modern observers who have given attention
to tlie matter state that they have often found vegetable mat-
ter mixed with other food in their stomachs, some claiming the
food to bo in small i)art vegetable, but mainly animal, while
otliers think the fragments of sea-weed so frequently met with
in their stomachs are only accidentally i)resent. Mr. Browji, who
appears to have had excellent opportunity of obtaining infor-
mation on this point, observes: "I have generally found in its
stomach various species of shelled Mollusca, chiefly Mya trun-
cata, a bivalve very common in the Arctic regions on banks and
shoals, and a quantity of green slimy matter which I took to
be decomposed Algai which had accidentally found their way
into its stomach through being attached to tho shells of the
^loUusoa of which the food of the Walrus chiefly consists. I
cannot say that I ever saw any vegetable matter in its stomach
which could be decided to have been taken in as food, or which
could be distinguished as such. As for its not [sic] being car-
nivorous, if further proof were necessary, I have only to add
that whenever it was killed neu. where a "Whale's carcass had
been let adrift, its stomach Avas invariably found crammed full
of the Iranf) or flesh of that Cetacean. As for its not being
able to hold the slippery cuirass of a fish, I fear the distin-
guished author of ' The British Mammalia' [Bell ] is in error. Tho
Narwhal, which is even less fitted in its want of dentition for
an ichthy()])hagous existence, lives almost entirely upon i)la-
ticlithyoid fishes and Cephalopoda. Finally the ex})erim the moUusks, that
almost exclusively constitute their food.* Other writers, how-
ever, who appear to have had equally as good oi>portunity for
observation, refer to the tusks as being of considerable service to
the animals in climbing. Cranz says: "The use the Sea-cow
Tiakes of these tusks seems to be in part to scrape muscles and
such kind of shellfish out of the sand and from the rocks, for these
and sea-grass seem to be its only food ; and also to grai)i>le and
j;et along by, for he fastens them in the ice or rocks, and thus
«hin\ s up his unwieldj' helpless trunk ; and fiimlly 'tis a weapon
of defence both against the white bear on the land and ice, and
the sword-fish in the sea."t
Most of the other early accounts of the "Walrus contain simi-
lar statements respecting the use of the tusks as locomotive
organs, and many later writers also refer to this use of them.
Mr. Brown says: "I have seen it also use them [the tusks] to
' Say« Maluigieu ; "In lii'trett' der eigeutlicheu Beatimmuug dcr ZUlme
bin ich iia Stuiulc die iiotliigo AufklUniug zu gelicii. Es liisst sicli uiclit
bcstifiten, dass dieselben als Wafleu angewendet werdou uud als solchc audi
furchtbar sind ; dass sic ivbei- anch als Lokoraotionsorgane dienen sollten,
ist cine Fabel, und daher der Nam** Odoutobwmts Steenstr. nicht pasaend.
Gleich dcu Robbeu bewegen sioli die Walrosse iiur init HUlfe iliier Fiisse,
sowohl auf dera Eise als au den saiidigeu Meeiesgestadeii, an deneu sio bis-
weilcu biuaiifsteigeu, luu zu schlafeu, oft zu Hundorten neben eiiiander.
Die Bestininiung der Ziiline ist eiue ganz andorc uud fiir die Existeuz des
AValrossea bei weiteni wiclitigere, deun niu* niit HUlfe derselben kann ea zu
Ht'inrr Nabrung koninieu. Ich fand, dass das Walroas sicli auaaclilieslich
vein zwei Mus',:belu, Mya truncata und Saxkava rugosa, niibrt, welcbo in oiner
^^'llssertiet■e von lO-fjO Faden 3-7 Zoll in deni Bodenlebm eingegraben lebeu.
I'm au diese zu kouuneu, muss das Walrosa sie aus dcni Lelim aufgraben." —
Ofarmijt Vvknuk. Akad. Forhaiidl. Sioclholm, 18(5.1, p. VM, as translated in
Ardiiv fiir Naturgesch., 18G4, p. UP.
*Tbe History of Greenland, etc., Brethren's Society's English translation,
LkikIou, 17()7, p. 127.
138 ODOBiENUS ROSMARUS — ATLANTIC WALRUS.
tlnig its huge body on to the ice. In progressii'-g' on shore it
aids its chnnsy progression by their means."*
Dr. Kane observes : "l"]veu Avhen not excited, lie manages liis
tusks bravely. They are so strong that he uses them to grapple
the rocks with, and climbs steejis of ice and land Avhich Avould be
inaccessible to him without their aid. He ascends in this Wiiy
rocky islands that an; sixty and a hundred feet above the level
the of sea; and I have myself seen him in these elevated posi-
tions basking with his young in the cool sunshine of August and
September.'' t
Enemies In respect to the enemies of the Walruses, nuui is,
of course, their chief foe ; but, after man, all writers rank the Polar
Bears as their i)rincipal adversaries. In their contlicts with this
formidable antagonist, the Walrus is usually the reputed victor.
Says Mr. Brown : " The Eskimo used to tell many tales of theii
battles ; and though I have uever beeu fortunate enough to see
any of these scenes, yet I have heard the whalers give most
circumstantial accounts of the Walrus drowning the Bear, etc.
These accounts may be taken merely for what they are worth ;
but still this shows that they are not wholly confined to Eskimo
fable, and ought therefore not to be hastily thrown aside. There
is no doubt, however, that the Bear and Walrus are (like all
the Piunipedia) but indiflferent friends." |
Captain Hall, however, relates the following story, rife amoiij;
the Inuuits, of a very ingenious w ay the Polar Bear has of kill-
ing the Walrus. The bear is said to take up his position on a
cliflf to which Walruses are accustomed to resort in tine weather
to bask in the sun on the rocks at its base. The Bear, mounted
on the cliif, watches his opportunity, and " throws down upon
the animal's head a large rock, calculating the distance and
the curve with astonishing accuracy, and thus crushing the thick,
bullet-proof skull. If the Walrus is not instantly killed — simply
stunned — the Bear rushes down to the Walrus, seizes the rook,
and hammers away at the head till the skull is broken. A fat
feast follows. Unless the Bear is very hungry, it eats only the
blubber of the Wjilrus, Seal and Whale." Captain Hall accoiii
panics his account with a i)icture of a Bear in the act of hurlinj; ii
stone upon the head of a Walrus ! § The story, doubtless without
» Proc. Zool. Soc. LoncL, 1868, p. 430. t Proc. ZotiL Soc. Loud., 18G8, p. 4:W.
t Arctic Exploration, vol, i, p. 415. ^S Arctic Researches, etc., p. 581.
OBM
ENEMIES.
139
basis in fact, is of interest in i*^^ bearingupon tLo mythical history
of tlio Walrus. In fact, Dr. Kane, on the other hand, says : " The
goneiiilly-received idea of the Polar Bear battling with the Wal-
rus ]ii(H>ts little favor among- the Esquimaux of Smith's Straits.
My own experience is directly adverse to the truth of the story.
The Walrus is never out of reach of water, and, in his peculiar
(,'leiuent. is without a rival. I have seen the Bear follow the
Ubsnk [Bearded Seal, Erignathns harbatiis] by diving ; but the
toui^ii hide and great power of the Walrus forbid such an at-
tack." *
The Walrus is also greatly persecuted, with parasites. These,
according to Brown, are two species of Hccmatopimis, one of
which invariably infests the base of the mystacial bristles, and
the other its body. " I have seen," says this writer, "the Wal-
rus iiu'itking loudly on the ice, tumbling about and rushing back
from the water to the ice, and from the ice to the water, and then
swimming off to another piece, and repeating the same ope-
ration, as if in pain. A few hours afterwards I saw a flock
of Saxicola cenanthe (it was on a land floe, close to the Fni
Islands) alight on the spot. On going over, I found the ice
speckled with one of these species of Hcematc^inus, on which the
birds had been feeding ; and the unfortunate Walrus seems to
have been in the throes of clearing itself of these troublesome
friends, after the approved fashion. Subsequently I have seen
these and other small birds alight on the back of the Walrus to
peck at these insects, just as crows may be seen sitting on the
backs of cattle in our fields." t It seems also to be infested with
intestinal parasites. Dr. Murie, J in his report iii>on the causes
of the death of the specimen in the Zoological Society's Gai'dens,
found it infested by a species of Ascaris {A. blcolor, Baird) to
sncli an extent that it was probably the cause of its death. He
states that he removed from its stomach about " half a pailful"
of small round worms, two and. a half to three inches in length.
Their presence had evidently induced chronic gastritis, death
resulting from ulceration. Circumstances seemed to indicate
that they had not been introduced with its food since its cap-
ture, but that it was infested with them before its capture and
confinement.
•Arctic Exploration, vol. i, j). 203.
tProc. Zool. Soo. Loiul., 1868, p. 430.
t Ibid., pp. (57-71.
140
ODOBiTlNUS ROSMARIIS — ATLANTIC WALl J8.
Domestication. — The AV'alruH i)osse.s.se.s ii high «h^yrcc of
c't'n;brul d('VC'h)i)UK'nt, und seems to be easily 8UH('ei)til)le of
domestication, ft appears, liowevor, to be dillicult to ]nn)\)
aliv*' in conflnenient, especially when taken far south of its
natural home. Doubtless the long period occupied in its trans-
portation from the Arctic; regions to the /ooloyical gardens of
European cities, during which time it is necessarily subject to
very unnatural conditions and unsuitable food, does nuicli
toward reducing it to a greatly enfeebled state before it reaches
European i)ort8. It appears, however, that three specimens
have at difterent times reached England, while two at least
have been taken to Ilollaud and one to St. Petersburg. In each
case they were quite young animals, probably less than a year
old. The first specimen seen alive in England reached London
August 20, 1G08. The account of the capture of this si)ecimen
and of its arrival in Loudon is thus detailed by Purchas. It was
brought in the ship "God-speed," commanded by Thomas Wei
den, on its return from a voyage to Cherie (now Beai) Islanci.
The account says : " On the twelfth [of July, 1008,] we took into
our ship two young Morses, male and female, alive : the female
died before we came into England : the male lived about ten
Aveeks. When wee had watered, we set sayle for England about
foure of the (ilock in the morning. . . . The twentieth of
August, wee arrived at London, and having disi>atched some pri-
vate business, we brought our living Morse to the Court, where
the king and many honoiu^able personages beheld it with ad-
miration for the strangenesse of the same, the like whereof had
never before been scene alive in England. Not long after it
fell sicke and died. As the beast in shape is very strange,
so is it of strange docilitie and apt to be taught, as by good
experience we often proved."* It hence appears that this spe-
cimen lived for only about three weeks after its arrival in Lon-
don.
Another is reported to have been exhibited alive in Hol-
land in 1612. This specimen was secured with its mother, which
died on the voyage to Holland, but its skin was preserved and
stuflfed, the two forming the originals of Gerard's famous draw-
ing already noticed. Von Baer,t however, raises the question
whether the London and Holland specimens were not really the
*Furchas his Pilgriines, etc., 1()24, vol. iii, p.
tLoc. cit., p. 1!U.
,60.
domestication'.
141
same indiviiliial exhibited at ditfen^nt times in the two eoun-
tri«>s, and devises an ingenious explanation lor the origin of the
supposed discrepaney of dates. He seonis to be led into tlieso
doubts by the similarity of some of the eirennistanees attend-
in;; tlie ( iiptuni and exhibition of these animals, and the elose
jigreenient of the dates. Master Welden's aecount of the eap-
tir.'e and transportation of his speeimen to London, and of its
early death there, seems, however, too oxplieit to be overthrown
by mere (jonjecture. There is apparently no reason for suppos-
ing that the Lon«lon specimen was ever seen alive in Holla icd.
From a statement in Camper's writings, it would appear that
a living s]>(!eimen reached Amsterdam sibont or before 1 780, as
he refers to having seen the living Walrus in that city.* But
of this specimen there appears to be no further record. The
specimen taken to St. Petersburg from Archangel, and described
by von lUier, lived only a week after its anlval in St. Peters-
burg.
In isr>3, a second living specimen reached London, and was
placed in the Gardens of the Zoological Society, where, how-
ever, it survived only a few days, dying apparentlj' of improper
and insufficient food. A third specimen, captured in Davis's
Straits, August 28, 1807, reached the Zoological Society's Gar-
dens in London about October 28 of the same year, where it
lived till December 19, or for nearly five weeks, when it died of
chronic gastritis induced by the immense number of intestinal
worms {Ascaris), by which it was unfortunately infested.t The
first London and Holland specimens were quite young animals,
as were also probably all the others. The second London speci-
men (1853) was a " very yoimg" female, but I have seen no fur-
ther statement respecting its probable age or its size. The
third London specimen (1807), a male, was judged to be less
than a year old, but measured 8 feet in length and weighed
about 250 pounds. No other specimen has thus far, so far as I
can learn, been taken alive to any point south of the Scandi-
navian ports, to which, according to Brown, they have of late
been frequently carried. J
That the Walrus, when young, possesses, like the common
Seals, a high degree of docility and intelligence, is amply evi-
* Camper says: "... . et que j'en avois vii plusieitrs mfime un
vivaut ii Amsterdam."— fflwvj-es, tome ii, 1803, p. 481.
tbeo Proc. Zool. Soc. Lond., 1867, p. 818, and 1868, p. 67.
tProc. Zool. Soc, 1868, p. 431.
■■
142
ODOBiENUS R0SMARU8 — ATLANTIC WALRUS.
1
,:i
i
dent from observations made upon it in captivity. In fur-
ther illustration of tliis point I quote the following from Mr.
Brown's important paper on the Seals of Greenland and Spitz-
bergen, from wliicli T have already quoted so largely. Mr.
Brown says, in referring to the subject of its naturalization in
zoological gardens : " I cannot better conclude these notes on
the habits of the Walrus than b.\ lescribing a young one I saw
on board a ship in Davis's Straits, in 1861, and which, had it
survived, was intended for the Zoological Society. It was
caught near the Duck-Islands off the coast of North Green-
land, and at the same time its mother was killed ; it was then
sucking, and too young to take the water, so that it fell an easy
prey to its captors. It could only have been jiupped a very few
hours. It was then three feet in length, but filready the canine
tusks were beginning to cut the gums. When I first saw it, it
was grunting about the deck, sucking a piece of its mother's
blubber, or suckitig the fskin which lay on deck, at the place
where the teats were. It was subsequently fed on oatmeal and
water and i)ea-soup, and seemed to thrive upon this outre nour-
ishment. No fish could be got for it ; and the only animal food
which it obtained was a little freshened beef or pork, or Bear's
flesh, which it readily ate. It had its likes and dislikes, and its
favorites on board, whom it instantly recognized. It became
exceedingly irritated if a newspaper was shaken in its face,
when it would run open-mouthed all over the deck after the
perpetrator of this literarj^ outrage. When a 'fall'* was
called it would immediately run at a clumsy rate (about one
and a half or two miles an hour), first into the surgeon's cabin,
then into the captain's (being on a level with the quarterdeck),
apparently to see if they were up, and then out again, grunting
all about the deck in a most excited manner ^awuk! moult V
When the men were 'rallying'! it would imitate the operation,
though clumsily, rarely managing to get more than its own
length before it required to turn again. It lay during the day
basking in the sun, lazily tossing its flippers in the air, and ap-
peared perfectly at home and not at all inclined to change its
condition. One day the captain tried it in the water for the
* " When a boat gets ' last' to a whale, .all tlio rest of the crew run shout-
ing about the decks, as they got the other boats out, 'A fall ! a fall! ' It is
apparently derived from the Dutch word ' Val', a whale."
t "When a ship gets impeded by loose ice gathering around it, the crew rush
in a body from side to side so as to loosen it, by swaying the ship from
beam to beam. This is called 'rallying the ship'."
MMMia
■HM
DOMKSTICATIOX,
i4n
first tiino; Imt it wiLs arently heartily sick
ut its mother element. After surviving for more than three
months, it «lied,.just before the vessel left for England. As T
wits not near at the time, I was unable to nmke a dissection in
ordei' to learn the cause of its death.''*
3Ir. Lamont thus describes a young Walrus he saw on board
the Norwegian brig "Nordby,"' in the possession of Captain Eric-
sou : " IJetbre parting layi'ul as a kitten. It was, of course, a great pet with all
on board, and scented much nn)re intelligent 'ived a voyage from Buenos Ayres,
* Yachting iu the Arctic Seas, p. H2.
tTliis specimen was captured iu Davis Strait, August 25, "by a noose
swung over his head and one foro limb from the ship and hauled on board.
For some days the captive was kept tied to a ring-bolt on deck, and refused
food altogether. Subsequently ho was induced to swallow thin strips of
boiled pork, and was thus fed until the vessel reached the Shetlands, when
a supply of fresh mussels was provided for its use. A large box with openings
at the sides was fabricated ; and the animal, secured therein, was brought
safely to Dundee on the 26th ult. [October]. From that port to London
the Walrus had been conveyed in the steamer 'j^glia' under the care of the
society's superintendent."— Proc. Zoiil. Soc. Land., 1867, p. 819. Mr. Bartlett
furtbei' says, in referring to the specimen : " As regards the present animal,
I may state that on my arrival at Dundee, on the 29th of October, I found
tlio young Walrus in a very restless state, and, as I thought, hungry ; it was
bcinij; fed upon largo mussels ; about twenty of these were opened at a meal,
and the poor beast Avas fed about three times a day. [1] I immediately told
\\\o. owners that I thought the animal was being starved. Stevens at once
iii;r(('(l and a codfish Avas procured ' the neighborhood, and by lue cut
into long tliin strips. On offering thc;c (lieccs of cod to the animal, he greed-
ily devoured Ihoni. Since that time I have fed the Walrus upon ^is/t, mussels,
whvlhs, clams, and tlie stomachs and inteatincs and other soft parts of fishes cut
suiall; for I find that it ortiou 525
Length of eleventh rib, cartilaginous portiou 380
Length of eleventh rib, total 905
150
ODOB^NUS OIJESUS — PACIFIC WALRUS.
mm.
Length of twelfth rib, osseous portion GOO
Length of twelfth ril>, cartilaginous portion '^'i{)
Length of twelfth rill, total d'JU
Length of thirteenth rib, osseous portion 4r)0
Length of tliirteeuth rib, cartilaginous portion 210
Lengtli of thirteenth rib, total OtiU
Length of fourteenth rib, osseous portion 3G5
Li'ugth of fourteenth rib, eavtilagiuous portion 120
Length of fourteenth rib, total 48.')
Length of tifteenth rib, osseous poi'tion 70
Length of lifteentli rib, cartilaginous portion 00
Length of tifteenth rib, total 70
Length of sternum, osseous portion 540
Length of steniuui, total 650
Length of scapula 420
Breadth of scapula * 24")
Oreatest height of its spine (at base of acromion) ^t,\
Length of the humerus 390
Tr.'insverse diameter of its head 110
Antero-posterior diameter of its head 132
Transverse diameter of distal end 138
Length of radius 273
Length of ulna :Ui2
Longest diameter of proximal end of ulna 130
Length of carpus 48
Length of first digit 124
Length of metacarpal of second digit 87
Length of thii'd digit 68
Length of foiirth digit 68
Length of tifth digit 75
Length of femur 250
Circumference of neck of femur *.. 135
Least transverse diameter of shaft 55
Transverse diametttr of shaft at end 116
Length of tibia 380
Length of tibula 375
Length of tarsus 172
Length of metatarsal of first digit 142
Length of second digit 126
Length of third digit 1 123
Length of fourth digit 132
Length of fifth digit 158
Length of innominate bone 430
Greatest width of pelvis anteriorly 320
Length of ilium 475
Ijcngth of ischio-pubic bones 245
Length of thyroid foranuai 153
Length of os penis 710
Width of manus at base of metacarpus 140
Width of pes at base of metatarsus 130
rnggmM
MKASUREMENTS AND EXTKKXAL CHARACTERS. IT)!
Ivospcctiiij;' tlir si/c iiii over the inequalities of the
siuracel and lias 10 or 12 feet of jfirth, and an old bull, shot by
the natives on Walrus Island, July 5, 1S72, Avas nearly l.'J feet
lonu. with the enormous <>irth of 14 feet. The immense mass
of l»I.il>ber on the shcmlders and around the neck makes the
1m ad and i)osteriors look small in proportion and attenuated."*
He estimates the ^iross weij^ht of a well-eonditioned old bull at
>'t\vo tliousan
J fore 2 4
Length of the fina / , . , „ ^
fore 1 2i
I
Rrcadtli of the fins n , . , -. n.
hind 2
(breadth - 5J
•'"""M depth 1 3
Circumference of the neck close to the ears 2 7
Circumference of the body at ftie shoulder 7 10
Circumference near the hind tins ") 6
From the snout to the eyes '• 7t
This was evidently either a female or not fully grown. The
circumference, as here given, is somewhat less than the length.
Respecting the external appearance of the old males as ob-
served in life by Mr. Elliott on Walrus Island, Mr. Elliott says :
" I was surprised to observe the raw, naked appearance of the
hide, a skin covered with a multitude of pustular-looking warts
and pimples, without hair or fur, deeply wrinkled, with dark
red venous lines, showing out in bold contrast through the thick
yoUowishbrown cuticle, which seemed to be scaling off in
places as if with leprosy. They struck my eye at first in a
'This is well shown in Mr. Elliott's tigures.
t Condition of Affairs in Alaska, pp. 161, 162.
+ Voyage to the Pacific Ocean, etc., vol. ii, p. 459. ' ''
ir)2
ODOn^ENUS OIJESIIS rAf'IKK" WAKHIS.
Ill
■
I?,
most unploasant iiianiu'r, lor tlit-y looked, like bloated, iiiortity-
iny, shapeless masses of fleshy the elusters of swollen, warty
pimples, of a yellow, i)arboiled flesh-eolor, over the shoulders
and around tlu^ neek, sujjgested lunvholesonieness foreihly." *
The old male, in tlie Mu.-- eum of Comparative Zoiiloj^y, of whieli
measurements are given, above, is almost wholly naked, except
about the numerous old healed pishes and sears, which arc
generally bordered with very short, stitf, brownish hair, ('aj)-
tain Scammon, however, who has also observed them in their
native waters, states that the hair that covers " most individn
als is short and of a dark brown ; yet there is no lack of exam-
ples where it is of a much lighter shade, or of a light dingy
gray. . . . The young, however, before its cumbrous canines
protrude . . . is of a black color." t
The mystaizc iind U'lif^tli, next to the long tusks, oiieol' tlu' most striking
Iciinuvs of tlio pliysioguouiy. Jn Cook's and Elliott's figures of
tlic I'iicilic siH'cies, liow-
ivt'i', tliey arc I>y no means
ii prominent feature, and
tlit'ican' nosueli allusions
to the formidable asi>eet
tliey give to the facial ex-
picssion as are (commonly
iiu't with in the a(!Couuts
of tlm Atlantic! species. A
duect comparison of speci-
HK^ns of (•orrespouding ages
shows them to be much
shorter than in the Atlan-
tic Walrus.
The eyes of the Atlantic
\N'idrus are described as
tiery retl, one writer (jom-
l»iiring them to glowiiig
(•(Jills. Mv. Elliott refers to
th(>s(M»f the Pacific species
iis having the sclerotic coat
••of a dirty, mottled coii'ee-
ycllow and brown, with
an occasional admixture of
white; the iris light-brown,
with dark-brown rays and
spots" ; and in no case have
I seen any referen(!e to
their being "red." While
most writers who have de-
scribed e Atlantic AVal-
riis from life refer to the
redness of the eyes as a
remarkable and striking-
feature. Cook, Scammon,
and others (Mr. Elliott ex-
oei»ted) make no reference to the color of the eyes, which would
hardly have escaped them had they possessed the redness char-
acteristic of the Atlantic species.
3Ir. Elliott further describes the eyes as small, but pronnneut,
"protruding from their sockets like those of a lobster," and
11). — Odohiiinii ohenH».
154
ODOli.KM S OMKsrS— PACIFIC WAr^HUS.
stnti's tliiit the iiiiiiiiiils liiivc the powiTot' rolliiifj; tlM'in about in
every (lin'ctioii, so that wlicn arousotl tlu'.v schloui move tlie
head more tliaii to <'levate it, the position of the eyes near tlic
top of the head Ki^'i"!^ them the needed raiiye of vision.
The nostrils, as in the Athinti«' species, are at the toj) (►f the
muzzle; they are ''oval, and about an inch in their <;Teat('st
diameter." The auricular openinj;' is placed nearly in a. line with
the nostails and eye, and henc«' near the top of the head in a
fold of the skin. Tin* animal is said to have a keen sense of
smell and an a(!ute ])ercei)tion of sound, but a limited power of
vision. *
An idea of the lun'onth and peculiar facial aspect of tbe Pa-
cific Walrus may be derived from the above-fjfiven figures (Fig.
13) drawn by Mr. Elliott, to Avhose kindness I am indebted for
tlielr presentation in tin- present cimnection.
I append herewith nn ., iirements of a considerable series of
skulls, of different ages, one oidy of which is marked as that of a
female, they being mostly skulls of middle-aged or very old
uiales.
* See Elliott, 1. c, pp. 161, 162.
t; I
M
MEASUREMENTS OF SKULLS.
155
•8S,),)0Jll
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-^ ra B fl O
O S S S 3
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rs -3 TJ a B >■! >■! >•,
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s s s;
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s s s
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e5fiSi«NciriiH«S)Si
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■q)Saai 'Boaoq Ibsbx
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15G
ODOUiENUS OBESUS — PACIFIC WALRUS.
Differential Characters.— As already stated, the I'a-
cilic Walrus differs I'roiu the Atlautic Walrus very little in
external characters, except iii facial outline and in the size aud
Fig. 14. — Odohauus obeaus.
*' set " or curvature of the tusks. The skulls, however, afford
many important differential characters, and on these differ-
mttmm^
DIFFERENTIAL CHARACTERS.
167
enccs I veuturo to predicate tbo existence of two 8i)ecies, using
the term " species " in its commonly accepted sense. To show
inoro readily what these differences are, I present herewith a
.series of figures of skulls of both old and young of the two
Ibrnis. Tlio skulls selected for this purpose are average exam-
](les of ii considerable series, the adiUt skulls being those of
males of strictly comparable ages. ^s
Fig. 15.^ — Odobcenus roamarus.
The skulls of the two species seen in profile (Figs. 14 and 15)
exhibit the following differences: The first and most obvious
158
ODOH^NUS OBESUS — PACIFIC WALKUS.
is pel uaps that preseiitetl by the tusks, which, iu the Pacific
species (Fig. 14), are mnvAi longer and thicker than in the other
(Fig. 15), less incurved aud
more convergent; their alve
oli are deeper and thickev.
with heavier walls, theryby
giving much gxeater fullness
to the front wall of the skull,
j even modifying the form of
the nasal bones. The front
outline, as seen in profile, is
very oblique in the Atlantic
species, while iu the Pacific
species it is nearly vertical,
the front edge of the nasal
bones being verj- little liostc-
rior to the/Vow* border of the
base of the tusk, while iu tbc
1 Atlantic species the., scarcely
Fig. 16.- Odob'niKHnxiiiumt^. pass beyond a vertical liuc
drawn from the himler border of the tusk. Tlie orbits in the
Pacific species arc
placed more ante-
riorly than iu tlie
other.
In the trout view
of the skulls, the
muzzle is seen to be
much smaller in the
Atlantic spades
(Fig. IG) than in the
Pacific (Fig. 17),
with, however, not
very marked differ-
ences in outlines and
proportions. The
receding ujiper bor-
der in tlie latter is a
marked feature.
The difiereuce iu
size hero shown is
an important one,
since the '■tino skulls compared dift'er very little in geuei'iil size,
Fig. 17.— ^loho'iiuH ohcsun.
DIFFERENTIAL CHARACTERS.
159
tliey giviug ve v nearly tlie same measurements in respect to
extreme dimensions oi length and breadth. The difference is
Fig. \fi. — OdoUrnm romnaruH.
lieuce one of proportion, resulting from the far greater develop-
ment in both breadth and depth of the anterior portion of
the skull in the Pacitic species. IJut while the skull of the
Fig. l!). — ()tl
(Figs. 22 and 24) of an extension posteriorly of the interniaxilla
ries for two-thirds of the length of the nasals. In the Atlantic
skull (Figs. 23 and 25), the intermaxillaries do not enter into tlie
dorsal outline of the skull, but terminate at the anterior l»i
der of the nasals. This difference is open to exceptions, and is
not offered as a character of importance, since the same modifi
cation or backward prolongation of the intermaxillaries occurs
occasionally in the i^ tlantic species, and is sometimes absent in |
the Pacific species, while in some exami)les the intermaxillaries
reach the dorsal surface only as isolated ossicles between tlie
nasals and maxillaries. As a rule, however, the conditions in |
this respect shown in the young skulls here figured appear to
be diagnostic of the two species.
DIFFERENTIAL CHARACTERS.
163
A comparison of tue skulls as seen from below (Figs. 26 and
27) sliows not only che considerably greater contraction of the
skull anteriorly, and the greater massiveness and different form
of the mastoid processes in the Atlantic Walrus, but other
weighty differences. These are especially seen in the size and
fonn of the auditory buUte, and, to a less extent, in the form of
the occipital condyles, the form of the glenoid cavity, the orb-
ital fossa), etc. In the Atlantic Walrus (Fig. 26), the auditory
Fig. 24. — Odobcenus obesus.
Fig. 25. — Odohwnus ro»maru«.
bullic are relatively larger than in the other (Fig. 27), more
])areiit in this aspect of the skull is the
relative posterior extension of the condylar portion, which, in the
Pacific species, extends nuu;h further beyond the posterior bor-
der of the mastoids than in the other. This is obviouslj- due to
greater length of the basioccipital segment of the skull in the
Pacific species, which is clearly shown in the annexed figures
FiG-^G.— Odolcrnusrosmarus.
(Figs. 20 and 27). The position of the foramina of the basal
portion of the skiUl is ilso quite different in the two, as is
especially seen in respect to the condylar foramina, which are
situated more posteriorly in the Atlantic species than in tbo
other, due, ]»erhaps, to the shortness in this form of the basi-
occipital region.
■to
DIFFERENTIAL {'IIAHACTERS.
165
Aiiotlu'v (liftbreiicc not yet noted consists in the greater
[('ii;itli iind massiveness ol' tiie /yjioiiiatsi in the Tacific species,
II which tliey are fully oue-thinl heavier than in the Atlantic
species; they being in the former both deeper and thicker.
This is well shown in the above given figures of the skulls as
5CCI1 in prolile and from above and below, but especially as
IS seen from below.) The orbital fossa' are also quite dift'erent,
:licy being relatively long and narrow in the Paciflo, and shorter
uul broader in the Atlantic Wahois.
Fui. 27. — Dilohivinis obvHux.
To sum ui) in a word the above-detailed cranial tlifterences
between the two species of AValruses, the skuU of the Pacific
animal is heavily developed anteriorly and relatively much less
so posteriorly, while in tlie Atlantic Walrus just the reverse of
this obtains, the skull in the latter being heavily developed
posteriorly ixnd relatively less so anteriorly. The axis of vari-
ation being- at the posterior border of the orbital fossae, the
166
ODOB^NUS 0BE8U.S — PACIFIC WALRUS.
zygomata share the general character of the anterior half of the
skull.
Fig. 28. — Vdohanua rosmarua. Adult.
But equally striking differences are seen in a comparison of
the lower jaws of the two species. These differences correlate
Fni. '«i'.). — OiLbfenus ohesus. Adult,
in a most interesting manner with those that characterize the
cranium. Tims, in the Atla-itic species (Figs. 28 and 30), the
DIFrKKENTIAL CHARACTERS.
167
iiiinidiblc is far less iiiiis.sive aiitciiorly than iii the Atlantic
"Wiilrns (Fi^s. IM) ami .'U), while it is much more massive pos-
tcriuily. There is also eoiisiderahle l ri-KUKX TI A L ( 1 1 A MM TEU'S.
I(i9
sililc serins ovidcnl iVoiii tlic tact cil' the «'\islc'iu'(', (luiiii;;' \un-
lions of the year at least, of areas of open water alon;;' those
|i(>rtioiis (»f the Aretie eoast supposed to separate the haltitats
111' ihe two species. Further fiiau tliis, I luive seen a skull (now
ill the 3Inseum of the Boston Society of Natural History) which
L'apt. Charles Hryaiit (certainly a trustworthy authority)
issiu'cs me was taken by his assistant, on Wali'us Islaud, in
the suminer of 1871 or 1872, that agrees in every particular
rtitli the skulls of the Atlan-
tic species. This skull hav-
iiif;' been somewhat fantas-
tically painted (the lower
surface deep red and tlie
upper yellowish-white), led
QIC at iirst to doubt the cor-
rectuess of the alleged local- FiO- ^i.—Odohamus ronmarun. Youug.
ity. su])posing that if really obtained at the Prybilov Islands
it lai^iht have been brought there from some distant point.
This quaint ornamentation proves, however, an aid in fixing
the locality of its capture as Walrus Island. It differing so
widely from the form usually occurring in those waters, it at
once attracted attention, and was mounted on a bracket and
preserved as a curiosity, the paint being applied, as Captain
Bryant inforius me, to facilitate its being kept free of dust!
Cai)tain Bryant states (in a letter to the Avriter) that he has
hinist'lf "seen two specimens like it," but adds that he "did not
succeed in killing them." Hence, of course, their resemblance
to the one now in question is
ouly presumed, the animals
being only seen alive. He
Aviites, further, that this
"head" was recognized as
"different from any before
seen there." I will merely add
that this skull is indistin- ^^*^- '^^^•—Odohamua obesus. Young,
giiisliable in any essential detail from skidls of corresponding
age from the Atlantic waters, and points to the occasional oc-
currence of Odobwnus rosmarus within the habitat of Odohcvnm
ok'niis. As von Middendorft" has showai (see iintea, p. 78), the
Ti alrus (presumably" the Atlantic species) has occurred much
further to the eastward than the limits assigned it by von Baer,
lie having traced it, satisfactorily to himself, api)arentl.y, to
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170
ODOB^NUS OIJESrS — PACIFIC WALRUS.
witliin thiity (lf,'noos of the western limit of the range of tlie
Pacific animal. Ii vicAv of these facts, the question arises ns
to "whether the Atlantic species may not occasionally pass aloiiji'
the northern coast of Asia so far as to sometimes reach tlit'
habitat of the Pacific species.
Nomenclature. — The first specific mime applied to the Ta-
ciflc Walrus is ohisufi, given by Illiger in 1815, in his " Uebei-
blick tier Saugethiere nach ihrer Vertheilung iiber die AVelt-
theile."* In this paper this name is three times used as a dis-
tinctive appellation for the Pacific Walrus, namely, (1) in his
list of the species of Northern Asia, in which " Triclteehus ros-
inarus^^ and " Trichcchus ohesus^' are both given; (2) in his list
of the species of North America ; and (3) in his remarks respect-
ing the first-named list. In these remarks (1. c, p. 75) he says,
" Die beiden Arten des Wallrosses, Trichcchm ohesus uud [T.]
Eosmarus, sind schon bei Nord-Asien vorgekommen." For Eu-
rope he gives only T. rosmariis (I. c, p. 5G), respecting the dis-
tribution of which he says, " Der Trichechus Eosmarus, das Wiill-
ross, lebt an den eisigen Kiisten von Nord-Europa, Nord-Asieu.
und des ostlichen Nord- America " (1. c, p. 61). It is thus not
quite clear whether he considered his T. rosmarus to have u
complete circumpolar range, with T. ohesus as a second species
occurring only on the northeastern shores of Asia and the north
western shores of North America, or whether, as is more probable,
hemerely meant that T. rosmarus ranged eastward along the Arc-
tic coast of the Old World to the northern shore of Western Asia
(as is the fa<;t), and was replaced on the Pacific shors^s of Asia and
America by T. ohestis. In cither case he I'ccogiiized as a distinct
species, under the name T. obesus, the Walrus of the North Pa-
cific and adjacent portions of the Arctic Ocean. In the same
paper is also a reference to a Trichechus " lUvergens,'" respctiiig
which he thus observes : "Auser dem schon bei Europa erwiilin-
ten Wallross, Trichechus Eosmarus, flndec sich an der westUclien
Nord-Amerkanischen und nahen Oat-Asiatischen Kiiste, uikI
dem Else dieser Meere, vielleicht aber auch an der ganzen Iviiste
des Eismeers das von Cook beschriebene und abgebildete Wall-
ross, das ich wegen mehrerer Verschiedenheiten, besonders . (>.S). lie thus, in the same paper, appears to
recognize two species of Pacific Walruses. The name din'rueim.
' * Al)liaii. 178.
176
ODOIJ^NUS OBESUS — PACIFIC WALRUS.
in 1823. Cook, in 1799, found tliem nuinerons in the neighbor-
hood of Icy Cape. They were also met with by lieeehey on
Diomede and Saint Lawrence Ishinds* and on other i.shmds
more to the soutliward.* Liitke found great herds at Saint
Mathew's Island, in latitude (JOo,t where their teeth were seen
later by BMlings-f They formerly resorted in summer in large
numbers to Saint Paul's and Saint George's Islands, Avhere, ac-
cording to Sarytschew, 28,000 pounds of their teeth were ob-
tained in a single year. They still resort, in small numbers, to a
neighboring islet (Walrus Island), and even to the easternmost of
the Aleutian chain, as will be presently more fully noted. For-
merly they were also abundant on Nunivak Island, situated to
the eastward of Mathew's Island, and not far from the Alaskan
coast.
On the coast of the mainland they have been met with in
great herds at different times in Kotzebue and Norton Sounds
and in Bristol Bay. Captain Cook appears not to have ob-
served them south of latitude 58° 42', at which point he found
them in Bristol Bay, as well as more to the northward.§ There
appears to be no certain proof that they were in early times ever
met with on the outermost of the Aleutian Islands, || and no
early reference to their occurrence anywhere south of Bristol
Bay and the Prybilov Islands. Brown, however, as late as
1868, says: "On the northwest coast of America I have known
it to come as far south as 50° north latitude.''^] Of this I can
find only a partial confirmation, and think that possibly there is
a mistake in respect to the latitude here given.** Elliott says,
* Narrative of a Voyage to the Pacific ami BcLringV f*traits, vol. ii, p. 'J71,
t Voyage autour du Monde, torn, ii, p. 176.
t Sauer's Account of Billings' Expcd. to the North Parts of Russia, p. 2'i'>,
$ Voyage to the Pacific Ocean, etc., vol. i, i)p. 43n, 455, 457; vol. ii,»l»p.
245, 248, 249, 259.
II On this point, see vou Baer, 1. c, i». 182.
U Proc. ZoiJl. Soc. Loud., 18(W, p. 4'.52.
**Mr. Brown further states in'the same connection that "It [the Walrus]
is found all along the circumpolar shores of Asia, America and Europe," and that
"Jt is not unlikely that it may cren be found in the Antarctic regions" ! L. c, p.
432. This idea I liave not seen elsewhere revived since tlie early part of the
present century. (On this point see vou Baer, 1. c, p. 173, and footnoti'.)
Dr. Gray refers to the reported occurrence by Bouelli of "Sea Horses" on
the Island of Saint Lorenzo, Callao. As this author describes "the two
great white tusks projecting from the mouth on either side," and further
says that "the tusks are of great value and form an important article of
commerce," Dr. Gra.V concludes these remarks "cannot apply to the tusks
of the Sea Bear"; but ho adds tliat he had "never heard of the genus Trichc-
ill
ill
.jaiuiL iJLJU.il
OEOORAPHICAL DISTRIBUTION.
177
AVI it ill}? in 1874, tluit "not more tban tliirty or thirty-fivo years
iif^o small numbers of these animals were killed now and then
on islands between Kodiak and Oonemak Pass" (hit. 55° to 57°).
lb' adds none "arc now found south of the Aleutian Islands."*
Kespectinfj their present €listribution, Captain Scammon, writ-
iiif; in 1S74, from personal observation, says: "Great numbers
of Walruses are found where the waters of the Arctie Sea unite
with those of Behring Straits, and also in Behriug Sea, and
that innumerable herds still resort in the summer months to dif-
ferent points on the southern or central coasts of Alaska, par-
ticularly at Amak Island and Point MoUer, on the northern
shore of the Alaskan peninsula. Within the last ten years many
of these animals have been destroyed by the whalers, both in
the Arctic and Behinng Seas."t
According to Mr. Elliott, the Walruses are now to be seen
in the Prybilov Islands only on Walrus Island,^ they being
so shy and timid that thej' deserted the other islands as they
became poi)ulated by man. In early days, or when the Rus-
sians first took possession, a great many Walruses were found
at Northeast Point, and along the south shore of Saint Paul's
Island, but with the landing of the traders and seal-hunters
the Walruses abruptly took their departure, and Walrus Island
alone is now frequented by them, being isolated and seldom
visited during the year by tne natives. He adds that they
are now most numerous, outside of the Arctic circle, in Bristol
Bay, where "great numbers congregate on the sandy bars and
flats, and Avhere they are hunted to a considerable extent for
their ivory ."§
They are now far less numerous than formerly, having greatly
decreased in numbers within the last fifty years. So numerous
were they in Behring's Straits about 1821, that a Eussian writer
elms living out of tho Arctic Ocean, and should have believed that he [Bo-
nolli] had uiistakcn the Sea Bear {Otaria leonina) for the Sea Horse," if he
biid not so particularly described the tusks. — Cat. Seals and Wlialea, p. 37.
Tile reference by Bonclli to the great white tusks of the " Sea Horses" relates,
in all probability, to tho large canines of the Sea Elephant, which were for-
merly eniiiloyed for a variety of uses.
'Condition if Affairs in Alaska, p. 1G4, footnote.
♦ Marine Mammalia, p. 180.
t A low rocky island, about half a mile long by one-eighth of a mile in
lireadtli, situated a few miles to tho southeastward of tho eastern end of
Saint Paul's Island.
H'ondilion of Affairs in Alaska, pp. 161, 164.
>Iisc. Pub. No. 12 12
178
ODOBJENUS OBESUS — PACIFIC WALRUS.
mi
i^f!
reports meeting with herds there embraciug thousands, and
even hundreds of thousands, of individuals.*
During recent years, in addition to the number killed by the
natives, the whalers are said to have destroyed as many as 12,000
annually, so reducing their numbers that the natives have be-
come anxious lest they shall soon lose this source of subsist-
ence, upon which they are so dependent.
Habits, Food, Commercial Products, and the Chase.—
The Pacific Walrus appears to agree quite nearly in habits ^v itli
its closely allied congener of the Atlantic waters. It has tlie
same gregarious propensity, the same intense affection for its
young, the same strong sympathy for a distressed comrade,
lives upon similar food, and is limited in its distribution by
about the same isotherms. Its leading characteristics were
concisely stated nearly a century since by Captain Cook in the
following words :
" They lie, in herds of many hundreds, upon the ice ; hud-
dling one over the other like swine ; and roar or bray very loud ; so
that, in the night, or in foggy weather, they gave us notice of
the vicinity of the ice, before we could see it! We never fouud
the whole herd asleep ; some being always on the watch. These,
on the approach of the boat, would wake those next to them ;
and the alarm being thus gradually communicated, the whole
herd would be awake presently. But they were seldom iii a
hurry to get away, till after they had been once fired at. Tlieu
they would tumble one over the other, into tlie sea, in the utmost
confusion. And, if we did not, az the first discharge, kill those
we fired at, we generally lost them, though mortally wounded.
They did not appear to be that dangerous animal some autliors
havo described ; not even when attacked. They are rather more
so, to appearance, than in reality. Vast numbers of them would
follow, and come close up to the boats. But the flash of a iiius-
quet in the pan, or even the bare pointing of one at them, would
send them down in an instant. The female will defend the
young one to the very last, and at the expense of her own life.
whether in the water or upon the ice. Nor will the youny one
* Von MiddendorfF says, " Taasende ja Hunderttansende im lebensfiisclie-
reu Berings-Eismeere," and cites as authority a Russian writer named
HUlsen. Von Middendorff continues, " Jm Jahre 1821 Uber Bah er [Hiilsen]
dort im Dezember Tausende, zu Ende des Juni Hunderttansende von Wal-
Tossen zugleich,welche die Luft mit ihrem Stuhnen erfiillten und von dcneu
einige, fnichtlos kratzond, sinTi bemiihten an den Schiffswandungeu ompor-
zuklimmen." — Sibiriacke Beiae, Bii. iv, p. 913, and footnote.
iiii:
HABITS.
179
quit the dam, though she be dead ; so ;'iat, if you kill one, you
arc sure of tlie other. The dnni, whou iu the water, liolds the
young one between her fore fins."*
In Captain King's continuation of the narrative of Cook's
last voyage, reference is made to a "Sea Horse" hunt. "Our
])e(»i)k'," says the account, " were more successful than they
had been before, returning with three large ones, and a joung
one, besides killing and Avounding several others. The gentle-
inou who went on this party were witnesses of several remark-
able instances of parental affection in those animals. On the
approach of our boats toward the ice, they all took their cubs
under their fins and endeavored to escape with them into the
sea. Several, whose young were killed or wounded and left
floating on the surface, rose again and carried them down. Some-
times just as our people were going to take them up into the
boat, and might be traced bearing them to a great distance
tluough the water, which was colored with their blood. We
afterward observed them bringing them, at times, above the
surface, as if for air, and again diving under it with a dreadful
behowing. The female, in particular, whose young had been
ileslioyed and taken into the boat, became so enraged that
she attacked the cutter and struck her two tusks through the
bottom of it."t
Tlie accounts given by subsequent observers confirm the
general truthfulnt ss of this brief but comprehensive sketch,
and supply some further details respecting its interesting his-
tory. Mr. II. W. Elliott, recently an agent in the employ of
the Treasury Department of the United States Government,
stationed at the Prybilov Islands, has made these animals a
special study, under opportunities unusually favorable for
observation. On Walrus Island, well known as being still a
favorite resort for a large herd of old males, he was able to ap-
proach within a few yards of a herd of several hundred old
bulls, which lay closely packed upon a series of low basaltic
tables, elevated but little above the wash of the surf. Here he
studied and painted them from life,f seated upon a rocky ledge
a few feet distant from and above them. He describes these
scarred, wrinkled, and almost naked old veterans as of by no
means prepossessing appearance. He says they are sluggish
* Cook's Voyage to the Pacific Ocean, etc., vol. ii, p. 458.
tibid., vol. iii, p. 248.
t See anteh, p. 174.
180
ODOU^NUy 0UE.SU8 — PACIFIC WALRUS.
and chiinsy in the wutor siiul almost helpless on land, their im-
mense bulk and weif^ht, in comparison with the size and strength
of their lin)bs, ri'ndering them qnite impotent for terrestrial
movement. " Like the seal, it swims entirely under water when
traveling, not rising, however, quite so frequently to breathe;
then it 'blows' not nnlike a whale. On a cool, quiet May
morning, I watched a herd off the east coast o*" the island,
tracing its progress by the tiny jets of vapor thrown off as tli«j
animals rose to respire.
" In lauding and climbing over the low rocky shelves," he
continues, " this animal is almost as clumsy and indolent as
the sloth ; they crowd up from the water, one after the otht^r,
in the most ungainly manner, accompanying their movements
with low grunts and bellowings ; the first one up from the sea
no sorner gets composed upon the rocks for sleep than the sec-
ond one comes i)rodding and poking with its blunt tusks, de-
manding room also, and causing the first to change its position
to another still tiirther off from the water ; and the second is
in turn treated in the same way by t) third, and so on, until
hundreds will be i)aeked together on che shore as thickly as
they can lie, frecpiently i)illowing their heads or posteriors upon
the bodies of one another, and not at all quarrelsome ; as they
pass all the time when on land in sluggish basking or deep
sleep, they seem to resort to a very irregular method of keep-
ing guard, if I may so term it, for in this herd of three or fonr
hundred bulls under my eye, though all were sleeping, yet the
movement of one would disturb the other, which would raise
its head in a stupid manner, grunt once or twice, and before
lying down to sleep again, in a few moments, it would strike
the slumbering form of its nearest companion with its tusks,
causing that animal to rouse up for a few minutes also, grunt
and pass the blow on to the next in the same manner, and so
on, through the whole herd; this disturbance among then Ives
always kept some one or two aroused, and consequently more
alert than the rest.
" In moving on land they have no power in the hind limbs,
which are dragged and twitched up behind; progression is
slowly and tediously matle by a succession of short steps for-
ward on thfe fore feet. How long they remain out from the
water at any one time I am unable to say. Unlike the seals,
they breathe heavily and snore.
" The natives told me the walrus of Bering Sea is monoga-
wammmm
IIAUriS — FOOD.
181
iiious, iind that the ai»er:s treatinji' of this j?rou]> in 1871;* but in 1872 1
he i»i()|>()se«l a new arrangement of the "Sea Bears." The sub-
division of this group into "tribes" is not here eleaily hidi-
cated, although he arranges the genera in four unnamed sec-
tions. In 187o| he proposed another arrangement of the "Sea
Hears," in which they were ^naced iu two i^rimary divisions, in
.C— 6
aeeordaneewith whether the number of molars is^ or '~.
His
later moditicationw were more formally jnesented iu his last gen-
eral account of the group published in 1874,§ in Avhicli theclas-
silication then presented diflered very much from that adopted
l>y him in 1808 and 1871. Although a new "tribe" ("Tribe 2,
(Ijllisophociua^^) was instituted, his former "tribes," CuUorhi-
hUuu Arctoc€2)hulina, and Eumeto]}iina, were united into one,
under the luime Arctoccphalina, thus reducing the whole num-
ber of "tribes" to four, as follows: 1. Otariina; 2. Gypsopho-
cina; 3. Arctoccphalina; 4. Zalophina. As before, he recognized
two primary "sections," by means of which Otaria is opposed
to all the other genera as a group co-ordinate in rank with all
the rest. Also the " sections," or primary divisions, are still
based on the posterior prolongation of the bony palate, and the
"tribes," or secondary divisions, on the number of the molars
aiul the position of the hinder pair relative to the "front edge
of the zygomatic arch." It is needless to add that a more purely
artificial and valueless basis could scarcely be devised. In his
later schemes, Eumctopias is placed under the division charac-
terized as having the molars ^, on the wholly theoretical
ground that "the fifth upper molar on each side [is] wanting,"
leaving "the sixth separated from the fourth by a wide space."
Ou similar grounds his Phocarctos elongatus, — based, as I shall
later give reasons for believing, in part on an adult female
Eumetopias stelleri and iu part on the young of the Japan species
of Zaloplms, — is considered as lacking the " fifth grinder" when
adult, though i)osses8ing it when young. As late as 1873, Eu-
mvtftpiafi is placed in a group explicitly characterized as having
*Supi>l. Cat. Soiils ami Wluiles, 1871, p. 11.
tProc. Zool. Soc. Lond., 187^, p. G55.
tProc. Zool. Soe. Loutl., 1873, p. 779.
^Hand-List of Seals.
190
FAMILY OTARIID^.
'thick under-fur" ! In his latest notice of the species (in 1874) his
synonymy of the species shows that he still believed the skin
of a young Callorhinus ursinus, referred in 18CG to his Arctoce-
phalus montcriensis, belonged to this species, although in 1871*
he properly assigned it to Callorhimis ursinus, which I had
shown in 1870 was its proper allocation.
Dr. Gill, in 1871,t made two primarj" divisions of the family, the
genus Zalophus alone constituting one division, which was thus
contrasted with all the others. The characters cited as the basis
of this division are the rostra^ profile (whether " more or less
decurved," or "straight or incurved") and the sagittal crest.
The last distinction was based wholly on a misapprehension
of the facts in the case,t and the first proves to be open to very
obvious exceptions. Although Dr. Gill, in his later papers on
this group, retains these divisions as originally proposed by
him, he has adduced no additional characters in support of
them.
ill
Genera. — The first generic division of the Eared Seals was
made by F. Cuvier in 1824,§ who separated them as "Arctoc^-
phales" (Arctocephahis) and "Platyrhinques" {Platyrhinclius),
with '■''Phoca ursina^^ (= Arctocephahis (lelalandi,V. Cuvier; A.
aw/flrcfjcMS, Gray) as the type of the former and ^^ Phoca leonimP
(= Otariajvhata of recent authors) as the type of the latter. Suc-
ceeding writers very generally adopted the name Arctoceplialus
for the greater part of the species, while PlatyrMnchus was con-
sidered as equivalent to Otaria of Peron, of prior date. Otaria
has, by some writers, even down to the present time, been used
in a generic sense for all the species of the family, sometimes
with and sometimes without subgeneric divisions. In 1859, Gray
separated generically the .'N'orthern Fur Seal from ArctocepMlm
under the name GallorJdnus, and the group has been since very
generally recognized a3 of generic or subgeneric value. Prior
to this date the only commonly recognized genera were Otark
and Arctocephalus. The next generic subdivisions of the Ota-
ries were institiited by GUI in 1866,1] namely, Sumetopias and
Zahphu ^ the former having for its type and only species the
Uorthem Sea Lion, or Leo marinus of Steller, while the latter
* Suppl. Cat. Seals and Whales, p. 15.
' ^"^-^ t Amer. Nat., vol. iv, Jan., 1871, p. 681.
tSee Am. Nat., vol. v, Marcli, 1871, p. 41.
$M^m. duMus. d'Hlst. Nat., vol. xi, xd24, 205.
II Proc. Essex Institute, vol. v, pp. 1-13, March, 1866.
il ■ ti
-m
- .^'^'-*t
OENEKA.
191
AViis f()iiii(lt'. 564.
UMonatsb. d. k. P. Akad. d. Wissenseb. zn Bnlin, 1877, pp. 505-507.
SPECIES.
193
IJiiir Sciils ( = tlic sul>Henerii EumctojHitfi, Zalophun, and Phoc-
(tirtos (if IN'tt'i's's «'avli('r papers).
SPKriKS. — I'l'ior to about the begiuninj^' of the present cen-
tui'\ , the Eared Seals then known were eoninionly referred to
two species, cue of which was termed, in eoinnion parlance, the
Sea Hear, Ours niarin, Meerbiir, etc., and the other Sea Lion,
Lion iiiarin, Meerlowe, etc. They were hardly mon; definitely
known in technical terminology, the "Sea Bear" being Phoca
uysina, and the " Sea Lion" the Phoca jubata. The first of these
names originated witii Linn<5 in 1758,* and the other with
Forstor in 1775.t Phoca urnina was based originally on Stel-
ler's Ur.su8 marinus, and Phoca jubata on the Southern Sea Lion,
or "Lion marin," of Penietty, to which species these specific
names have of late been properly restricted. Zimmermann, in
[1782,1 named the Southern Sea Bear Phoca «M«^rflZ/»(=" Falk-
land Seal, Pennant II, p. 521," the Sea Bear of Forster), which
Shaw, in i800, renamed Phoca falklanilica. Both names were
based on the "Falkland Isle Seal" of Pennant, but Zimmer-
mann's seems to have been entirely overlooked by subsequent
rritors. As it has eighteen years' priority, it must be adopted
in place of falklandica.
During the last half of the last century and the early part of tlie
present, the early voyagers to the southern seas (as Anson, Per-
netty, Forster, Weddel, Pdron and Lesueur, Quoy and Gaimard,
Lesson and Garnot, and Byron, among others) met with different
species of Sea Lions and Sea Bears. They described these ani-
mals very imperfectly, their accounts relating mainly to tlieir
liabits and localities of occurrence, and they brought with them
to Europe very few si3ecimens.§ Desmarest in 1817, and Lesson
in 1828, gave names to the species thus obscurely indicated, the
latter renaming several that had already received names. To
these authors, and to the often-quoted remark of P^ron that he
believed there were not less than twenty species of Otaries, we
are indebted for much of the confusion and obscurity that must
ever be inseparable from the early history of this group. Des-
marest alone, in his article on the Otaries in the " Dictionnaire
•I'Histoire naturelle" (vol. xxv, 1817, pp. 590-603), lecognized
'S.vst. Nat. i. 17r)8, 37. '
tDpsciip. \iiiin., pp. 6(5, 317.
tGeograiili. GoHcliicute, Thell iii, 1V82, p. 276.
^ G. Cuvicr, accorrtiug to Gray (Catalogue of Seals, 1850, p. 2), had skulls
of only two Hpccies of Eared Seals when ho wrote the "Ossemens Fossiles."
Misc. Pub. No. 12 13
194
FAMILY OTAimi)iE.
niiio spocios, only two of which hiiv«i any tan^^ibh) basis, or cau
bo dt'torminod except conjecturally, and mainly on the basis of
their habitat. In fact, it is abnost impossible to say whotlier
they are "hair" Seals or "far" Seals; the dt'scriptions sluiw
merely that they were sona^ kind of Ear* il 8e'>l. Desmari'st's
81)ecies are the following;: 1. Otnr'm konimi (= ^farin juhnUij-
EHmctopinH sh'lhri)', 2. Otaria ursimi {=(Muo/,t nun ursiintii):
3. Otaria pcroni (n. sj)., based on a vaj^n^ account by M. IJiiillv'
of an Eared Seal seen in great numbers on llottnest Island, west
coast of Aust ralia. Desmarest doubtfully refers to it two mounted
skins in the Paris Mtiseum, both of very young animals, the
larger only about two feet and a half long, brought from " Ter
res Australes ") ; 4. Oiaria vinvrea ( l*6ron et Lesutnir, Voy. an
Terr. Austr., ii, 77; habitat, "lie Decres," coast of Australia;
an Eared Seal, with rough hair, described only in general terms,
ami undeterminable ; probably = Zalophm lohatus) ; 5. Otam
alhkollis (Perou et Lesueur, 1. c, 118 ; habitat, " lie Engtiie,"
coast of Austraba ; an Eared Seal, eight or nine feet long, char
acterized by a white spot on the middle and upper part of tlie
neck ; perhaps the same as the last, but not certainly detti
minable) ; 0. Otaria flarescens (ShaAV, Mus. Lev. ; Gen Zoiil., i,
200, pi. ixxiii; habitat. Straits of Magellan; a "Yellowish Seai,
with pointed ears " ; not determinable, but probably = O. juhata]
7. Otaria falklandiaa {— Plioca faJldandim Sh aw = iVtoca «««
iralis, Zimm. ; " Cinereous Seal, witli small pointed ears, aiultli.,
cutting-teeth marked with furrows " ; presumably the common
Fur Seal of the Falkland Islands) ; 8. Otaria porcina (= P/iotd
poreina, Molina ; habitat, coast of Chili ; wholly uudetermiuable ;
9. Otaria punilla { = '■'■ I*hoca pusilla, Linn."; a wholly mythical
" Otary " as described by Desmarest, supposed to inhabit theMdi-
terranean S^^a ! t Of these nine siiecies, only one ( Otaria iirsina).
* P6ron ct Lcsueur's Voy. Terr. Auatr., vol. i, p. 189.
t In view of recent attempts to revive tlie name imsilla as a touablo desig-
natioji for some species of Eared Seal, it seems desirable to slate fully tk
original badis and early history of th^s name. It was given originally')?
Schreber, in 1776, to "Le Tetit Phoqne" of Buffon, Schreber oven copyinj
Buflbn's figure (Hist. Nat., xiii, 1765, pi. liii). Buflfon introduces his uotiff
of this species as follows : " Le second [ osp^co] (planche LIII ) qui est le phoqne j
de la Mdditerrande & des mcrs du Midi, & que nous pr^sumons 6tre \cpho»
des Ancienq, paroit 6tro d'une autre espfece, car il difffere des autres par )°
quality & la couleur du poll qui est ondoyant & presque noir, tandis qnek
poll des premiers est gris & rude, il en difffere encore par la forme des dents
& par celle des oreilles; car 11 a une espfece d'oreille exteme tres-petiteili
v€rit^ ..." Then follows a good description of a young Fur Seal; lint in
SPKCIKS.
105
or possiltly a socoiul {O./alklamlien), is positively referablo to
any ])ai"ti(!nlar specicH as now known.
Tlii'C'c years later (in 18LM)) J)esniarcst again, in his "Manuna-
loj-ic'' {EiK'yclop(i(lie Metliodicpie, vol. clxxxii, pp. 248-252), re-
(li'siiilKMl the Otaries, reducing the number of species to eight
by uniting his Otaria pimUa to his Otaria peroni under the
latter name, which now relates not only to the Fur Seals of the
western coast of Australia, but also to those of the Cape of Good
Hope.
Lesson, eight years later, in his article on the Otaries (Dic-
tionnaire classique d'Histoire Naturelle, vol. xiii, 1828, pp. 419-
426), raised the number to fifteen. One is purely mythical ; live
or six can be determined as equivalent to species now commonly
recognized, but the greater part are not satisfactorily identifia-
ble. His species are the following : 1. Otaria fabricii ( = " Plioca
?/r«/««Fabricius"; habitat, Greenland; wholly undeterminable;
certainly not an Eared Seal, and probably wholly mythical) ; 2.
0tariastelleri{=Leomarinu8, iitQ\ler,=Eum€topias stelleri, which
here receives its first distinctive name) ; 3. Otaria californiana
(="jeune Lion mariu de la Califomie," of Choris, and hence =
Zabj)hus (jillespii of recent authors, which here received its first
specific name*) ; 4. Otaria Jcraschenninikoicii {= Ursus marimtSy
n long footnote to tbis description he gives quotations from Glaus Magnus,
Zorgdrager, Charlevoix, and from collections of voyages, which relate to the
Seals of both the Arctic and Antarctic regions, none of which probably re-
fer to any species of Eared Seal. On the following page ho says: "C'est
par line convenauce qui d'abord parolt assez Idgiire, & par (inelques rapports
fiigitifs que nous avons jugd que ce second phoque (pL LIU) <5toit le phoca
de» anoieus ; on nous a assurd que I'individu que nous avons vu venoit des
Imles, & il est an moins tr6s-probable qu'il venoit desmers du Lev.ant; . . ."
—Hist. Nat., xiii, 1765, pp. 340, 341. Though assumed to be a Medttci-^aueau
species, the origin of the specimen here described and figured as " Le Petit
Phoque " is .avowedly unknown, and a certainly ereoneous habit.at is as-
signed to it. This is the sole basis, however, for the Phoca pmilla of .all the
earlier systciuatists, and of some modem ones. As already stated, Desma-
rests Otaria piisilla is purely mythical; for while he describes an Eared Seal,
lie elainis for it a Mediterranean habit.at, and deems; it to be the species
ileserihed by Aristotle, Pliny, and ^Elian, and figured by Belon, .and even
goes so far as to say, "Buffon et Erxleben parolssent avoir confondu, avec
ee plwKiue, de jennes individus d'autres espfeces particuliJ^res aux Torres Aus-
trales, et particulierement i\ I'ours-marin de I'Ue de Juan-Femandez. Quant
a lui, il seiiible p -pre b, la Mdditflrrande." The Phoca pmilla of Erxleben and
Gineliu is a heterogenous compound of Eared and Earless Seals from both
lieiuisphereM.
* See further remarks, poetea, unJer Eumetopiaa atelleri and Zalophm califor-
mnus.
196
FA:HILY OTAKIID/E.
SteWer =C((lJorhiin(s urshius); 5. Otariu pernetUji { = Otaria juh-
ta) ; 0. Otariaforateri (cnibraces all the Fur Seals of the Soutlievn
Hemisi)here) j 7. Otaria mollissina ("Lesson et Ganiot, Zoolope
dehiCoqiiille,pl. iii,p. 140"; habitat, " lies Malouiues " ; the long
description contains nothing in itself distinctive of any species,
but it has Ijeen determined, by Nilsson and Gray, from the skull
and skin in the Paris Museum, to be a young Otaria juhata)',S.
Otaria peroni {='"'■ Otaria peroni, Desm., sp. 382"; embraces
^'' Plioca piisilla Linn.", "Petit Phoque, Buffon," "Otarie de La
lande, F. Cuvier," and "Loup marin, Pag^s"; habitat, Cape
of Good Hope ; formerly referred by Gray to his Arctocephalm
delalandi, to which it is mainly referable); 9. Otaria coronak
("Desm., spec. 383; Phoca coronata, Blainv."; undetermhiaUe,
and habitat unknown); 10. Otaria cinerea ("P<5ron et Lesueiir/'
as above); 11. Otaria alhicollis ("P^ron et Lesueur," as above);
12. Otaria flaveseens (= ^^ Phoca Jlavescens, Shaw," as above; uot
determinable); 13. Otaria sliawi {= Phoca falJdandicus, Sha.\!,
therefore = Arctocephalus falklandicus, auct.) ; 14. Otaria hmi-
villii ("G. Cuvier, Oss.Foss.jt.v, p. 220"; = Arctocephalus fall
landicus, auct.; habitat, "Hes Malouines"): 15. ^'•Otaria moll-
naiV {=^^ Phoca porcina, Molina"; no tangible description, and
wholly undeterminable).
Fischer, in 1829-30,* appears to have recognized fifteen (only
twelve have numerals prefixed) species of Eared Seals, whicli
are the same as those describ'^d by Lesson in 1828, with the ex-
ception that Lesson's Otaria fabricii is not admitted, and Gray's
Arctocephalus lohatus is added.
Hamilton, in 1839,t recognized twelve species, as follows: 1.
" Sea-Lion of Steller" (= Uui etopias stelleri) ; 2. " Sea-Lior, of
'FoTnter^ {= Otaria jubata)', 3. " Sea-Lion of Pernetty" (=Of. OUtria (jodcfrotji m
sp.=:(>. Jnhaid); -l. ^*^ ? OUiria hi/roiiid"' { — I'liocd hiiroiiia, ]>lniii
\'\\\c. — , ci)i<'rvd, I'eroii and LesiK'ur.
Qnoy and Claimard"; " 0. stcllcri, Sehlef;el,'' in part,ete.); ?('.
Ota rid falklditdicd (= An'tocephalus falhldndicus, anet.); 10.
Otdria uvsiud { = Cnllorhinus iirshris); 11. Otdrid stcUcri {Eimt
topias ,stelle)-i); 12. Otdria gillcspl {= ■ <■ phius californianus); 13.
Otdrid lobdta (= Arctocephalus lob. , Gray, 1.S2S, Otarid am-
tralis, Quoy and Gainiard, 1830, 0. uteUeri, " Schlegel," in
part, = Zalophxts hbatua)-, 14. Otaria philippii (n. sp. = Arda-
eephalusfalklandicuH, anet.). Six months later, on again review
ing the group,! the same writer redneed the number of speciis
to ten. In this paper he referred the 0. byronia, 0. leonina, aud
0. (jodeffroyi of his former paper to O.jubata, and his O.philipph
to 0. fallxlandica. 0. idlow is still retained as a valid specitvs
and " Otaria steUeri, Sehlegel," is determined to be the 0.
ffillcspi, M'Bain.
In 1808 f Dr. Gray deseribed as a new species Arctoccphalm
uirosus {= A. autarcticus, H. 7>«s///«s) from the Cape of Good
Hope, and Professor Turner added,§ as a new species, Anio-
ccphaluH schisthyperol's (later corrected to nchisitKperus by Giiu-
ther), from Desolation Island, considered later by Gray,|| aftoi
an examination of the type, to be referable to his A. dcJaUmli
(therefore = A. antarcticm). !M'Bain, tlie same year,^ described
an imperfect skull of what he called "0. uUowf'' {=Otark
jubata, fem.), adding that in case it proved to be a new species it
might be called " 0. graiiP
In 1870 ** I was able to recognize only six species as well
established, but gave two more as probably valid, the latter
*Mouat8b. d. k. I'. Akucl. Wissonsch. zu Berlin, 18GG, pp. 261-'281.
tibiil., 1860, pp. r)(]!>-G7'2.
t Ann. and Mag. Nat. Hist., 4th ser., vol. i, p. 21'J.
^ Journ. Anat. and Pliys., vol. iii, pp. 113-117.
II Ann. and Mag. Nat. Hist., 4th scr., vol. iv, p. 264.
• H Ibid., pp. 109-112.
"'Bnll. MnH. Comp. Zoiil., vol. ii, pp. 44, 45.
SPEC IKS.
199!
licinj;' A rctoiTphalm cinercvs, ( Jray (Australia and New Zcalaiul),
;iii(l A.((iit((r('ticiis,(\vi\y (Capo of (Jood Hope). The othev.s are:
1. OitifUi jubatn (iiudcr \vlii<'li was wionjily included 0. haolcen,
(;i;iy);2. Eumctopinx stiKcrl ; 'A. Zdloplius (iHlc.spi ; I. Zalophus
IdIiiiIiis; 5. ('allorliiniot urniitHN ; (». Afcto(Tpii(iliis/. idUnv of
Peters). L*. Callorhhius nrninus. 'A. riiocarctoxhool-eri. i. Arcto-
ccph(th(s auta)'cticus (Cape of Good Hope = Fhoca nnturcfica,
Thunherg", 1811, and Fhoca [s. Otaria] jno^ida and dchilandi^
iUiot.). r). Arctoccpludus nif/rcscohs {=A. OKstrtdis). (». Arcto-
vvpludus ci»crc»s. 7. ArctoccphiduH footer i ("IN^ew Zealand, "'=
''•riwca nrxina, Forster," = 0^flr(Vt forsteri, Lesson, formerly re-
ferred hy him to his A. /(dJdaudicits!). 8. ArctoccpMho^ fidlc-
hmlicus. i). ^^Arctoeejyhainsf nivosus^^ {= A. antarcticiiN). 10.
Zalophiis (/dlcspi. 11. Kcophoca loibata. VI. Eumetopias KtcUcri
(embracing his Arctoccphalm montcrkiisis and A. califoniiduns).
13. Aritophoca philipjiii (= Arctoccphalm aus'traJis). A.forsteri
is tbeonly species added, while no less than six species, recognized
by either himself or Peters in 18GG, are reduced to synonyms.
Gray, in ]87L*,{ added Gypsophoca trojiictdis, based on a young
skull from Auckland Island, to which siK^cimens from X()rtk
Australia are also referred. This Clarlv§ believes to be in part
based on the young of Otaria hooker i, and in part referable to
A rctoccph (diis cin ere us.
Scott, in 1873, in his account of the Otariid(V,\\ described (p.
10) what he regarded as two new species of ArctocephahiSf
namely, '■'■ Arctoccphalm GrayW'' and '■'■ Arctoccphalm eulophus.^^
The first is equivalent to Gray's A. faVdandicm of his Cata-
' Mdiiiitsh. a. k. r. Akiul.Wissoiiscli. zn Berlin, 1871, pp. 5.'>d-.'>()6.
tiSupltl. (Jilt. Sfiils aiul WiUik\s.
tPn.c. Zool. Soc. I<()iul., 187i*, pp. CW, 743.
^^ I'roc. Zd.il. Soi'. Lond., l'-'7:{,p. 759.
ilMiiiiiiimlia, Kocciii and Extinct, nn elemcntnry treatise for tlio iiso of the
imlili, s(li,)<)lsotN<'\vSonthWak'H. By A. W. Scott, M. A. Sidney, 1873. Sec-
tion H, rinnata, Seals, DnjfongN, Whales, &c. &c. &c. — Otariidse, pp, 7-25.
200
FAMILY OTAllIlDi^i:.
m
mi-
I: i
I"-) ''1-
logiu' of Seals and Whales (1S0(5, j). r»r)),aiKl the "Supplein«'iit"
to the same (1871, p. 2~)), whieli ]\rr. Seott gives as a syiionym.
After quoting Gray's deseriptioii oi A. /((nda)uliciis, he says:
"This is <;learly a s])eeies distiiiet from the eonimon Southern
Fur Seal. . . . The speeitie name Fidkbtmlieus haviiijj;
been ai)pro])riate'wj//i." The Aretocephalm
eulopliUH is based on verbal information from Mr. Morris, an
experienced sealer, Avho infornied him " that during his sealing
voyages he occasionally met with a fur-seal, which he aud
those connected with him in the trade readily recognized as a
distinct kind — by the diminutive size of the adult animal ; by
a top-knot of hair on the crown of the head; aud by the soft,
beautiful under-fur, unlike in colour to, and mucli more valna
ble for articles of ladies' wear than that of any other fur-seal
they were in the habit of captiu'ing." " This seal," continues
Mr. Scott, '* appears to be rare, only a few specimens having
been taken ; some were seen on the south-east coast of New
Zealand, evidently stragglers driven far away from home. i\Ir.
Morris has been told that they were formerly common on tho
shores of Patagonia and the Island of Juan Feriumdez." AVith
all due deference to the opinions of ]\Ir. Morris and Mr. Scott,
this information hardly forms a satisfactory basis for the erec-
tion of a new species in this obscure group, where external
characters, when well known, are of slight distinctive value.
The Arctocephalus eulophiis can only be assigned to the category
of vaguely described and indeterminable species, of which the
writings of P6ron, Desmarest, and Lesson were so prolific half
a century ago. Only six other species were recognized by Mr.
Scott, namely: 1. Arctocephalus nmnun {= Callorhluus ursi-
mis). 2. Arcfnceplialus falklamlieus (embracing all the Southern
Fur Seals, with the exception of his two " nes? species," already
noticed). 3. Zalophus gillespi. 4. Z.lobatus. 5. Otaria stelleri.
6. 0. jubata {= 0. jiibata and Phocarctos hoolccri Grdy).
In 1873, Dr. Gray described* a Eumetopias elongatus, ])ased
in part on a skull from Japan he had the previous yeart referred
to U. stelleri, and in part on a joung skull, also from Japan,
which, doubtless, is the same as the Otaria stelleri of Temminck
(Fauna Japonica).
*Proc. Zool. Soc, 1873, p. 776.
tibid., 1872, p. 738.
-;i'S*=ii
SPECIKS.
201
In 1874, the sauie author* addrd two more •' now species" of
Otaria, this time whoUy from old material, from unknown loj-ali-
ties,which he had had before him in tlu' 1 >ritish ^I useum for uearly
twenty years, and which he had hitherto uniformly referred to
Otaria juhata ! Ilaving', however, found that the lower Jaws
ilitl'ored from those of the other sp«;cimens in beinji " straight,
not bowed on the side, and elong.ate," and that " the scar of the
temporal muscle is elongate, nanow in front," instead of being
" broad, rounded in front." One of the species, basef this grouj*. In
this work two other "new sj)ecies" are added, making in all
eighteen species of Otariidw now recognized by Dr. Gray!
These are: 1. Otaria jnltata. 2. 0/arm »M/«or (see above, hist
paragraph). ,3. Otaria tdloa' {=0. iiHoa, you Tschudi and
Peters, and O. pygmcca, Gray, both formerly, and, I believe, cor-
rectly, referred by him to 0. Jubata). 4. Gypsonhnca tropicalis
{=Arctocephalus cinereus). /). Phocarctoa hoolioi >. H. Phocarc-
tns clongatm { = Eumctopia8 stelleri, in part, and Otaria stelleri,
Tciiiniinck, in part). 7. CaUorhimis ursinus. 8. Arcfocephalun
oittarcticuH. 9. Euotaria cinerea (includes Aretocephalus fomtcri
of Gray's Suppl. Cat. Seals and Whales [see above, p. 199] ).
10. Euotaria nigrcscenn {= Arctocc2}haluii austral is). 11. Euoia-
via hitirontri.'i (n. sj)., based on a skull supposed to have come
from the Falkland Islands, formerly releried to his A. nigrcs-
ccns. lie now says, " The skull may belong to the Arctocepha-
hiH faUdandicu.'i, of which [/. c, his A./allcIandicus] the skull is
i
*Ann. nudMag. Nat. Hist., 4th ser., vol. xiii, p. ;i;'4. t Hiuul-List of Seals, etc.
202
FAMILY OTAKIIDiE.
not known, or it may he a (listiuct upccics'^). 12. Euotarin com-
prcssus (n. sp. ; lial>. '^ South Africa? ^^'arwi(•k^^•, formeily iv-
foiTcd by liini to Arvtocc.phalaH hoolerl a.s "9 .skull, South
Sea, 3Ir. Warwick's colh'ction"*). !.">. Eitotaria .svh isf In/ jk nil's
(=.Arclocq>li<(li"< ■sclilstlii/jxrors/Vuvuvv, formerly referred, witii-
out reservation, by (f ray liiniself to his. l>T^o(r/>/<«/w.s««/«r(//V».s).
14. Einnctopids Ntclltri. 1~k ZolopliKs jfifJeKpi. 10. Xeopliacn
lobiita. Two other species are also <>iven, as follows : 17. ^'Arrto-
(■(pliahis .^ iiivoNiis"' ( = .l. (i)if((rcticus)', IS. ''Arctorcphahis.'/dJh
h(H(licuH " ( = ,1 . ((Kstralis), These are Fur Seals, referred (h»ubt-
fully to Arctoceplialus from lack of knowledji'e of the skulls.
The first, he says, "nniy be the skin of Euotaria coi)ipri'N,su or
schisthyporoi'ti^'', to the latter he refers the '-'■Arvtocipluilux iirdjiii"
and "('h/o/>/<».s" of Scott (see abov<', p. 200), the latter, however,
doid)tfully.
In 1875 Dr. Peters described t still another si)ecies, based uu
two .specimens, an old male and a youus" female, brought home
by the German Transit-of- Venus Expedition (supposed by liiiii
to have both come from Kergfuelen Island), to which he ^^avc
the name Arctophoca yazeUa. Externally A. gazella appears to
differ little from the other Southern Sea Bears, the distinctive
characters resting in the form of the hinder border of the bony
palate, which has ji triangular projection at the middle, in the
very small size of the tympanic bones, and in other «letails of
the skuU-structure.f Later he found that only one of the spcei
mens on Avhich A. yazella was based came from Kerguck'ii
Island, the other having been brought either from "der Insel
St. I'aul o«Ier Amsterdam." In 187(»,§ therefore, in referriu;i
again to these specimens, after the discovery of the error in
locality respecting one of the specimens, he renamed the Saint
Paul or ^Vmsterdam Island skin Otaria {Arctophoca) elegaiis.
In 1877, Dr. Peters again reviewed! | the Avhole group of
Eared Seal, of which he at this time recognized three genera
and thirteen species. He refers to having had access to uuicU
new material, and it is greatly to be regretted that he has not
* Cat. Seals, Brit. Mils., 1850, p. 16; Cat. Seals and Wlialos, 18(56. p. .'.4.
tMonatsb. einy here delinitely allocated,
lie liavinji' here made ri!di» Australea, t. ii, p. 54," The Otary of the Cape here referred
t(» is the one biought by M. Delalaude, whicli is the Fur Seal of the Cape
of Good Hope.
^ '11
9
;]
204
FAMIL^' OTAPilDiE.
Phoque," on which the name pusiUa rests,* must have come
from tlie Cape of Good Ilope.t Tlie Fur Seals of South
Ameriea are reeoyiiized as behuiyiu}^ to two si)eeies, those of
the east coast, tlie l-'alklaml Islands, the southern extremity of
the continent, antl the west coast northward to Ciiili i>ein};' ro-
fened to ArctoccphaliiH falkhuHlicm, while those from Juan Fer-
nandez and Masafuera Islands are assigned to A liliiUppi. We
are therefore left to suppose that his and Gray's A. nigrescens,
bis A. argentata,X Gray's Euotaria latirostris, and Scott's ^1.
grayi and A. euhphm, are regarded by him as synonyms of
these species. The Fur Seal of Australia he calls Arctocepluilm
brevipes, citing " Otaria einerea Quoy et Gaimard, Voy. Astro
labe, Zoolog. i, p. 89 (non P6ron)." He also recognized A. ek-
gam from Saint Paul and Amsterdam Islands (to which he
doubtfully referred A. tropicalis, Gray); A. gnzella, from Ker-
gueleu Island; and the A./orsteri, Lessoji, from New Zealand
and the Antarctic Seas to the southward of New Zealand. Four
of his species, namely, Arctocephalus ckgan,s, A.forfiteri, A. (fa-
zella, and A.pMUppii, ai)pear to me to be invalitl, while under
his Eunietoinati giUespi, I l>elieve he has c«mf()unded two quite
distinct species, namely, ZaIo2)hus aiUcspi and Z. lohatus, Pe-
ti'rs's thirteen species i. .» the foUowinji :
8. Arctocppliiiliis Itrcvipcs, Peters.
9. Arctocepliiiliis elcjfans, Peters.
10. Arctoeeitlialiia foiHteii, Lesson.
11. Arctoceph.alus gazellii, Peters.
12. Arctocephalus pliilippi, Peters.
! 13. Arctocoplialus ursinus (Liuiid).
1. Otaria jubata (Forster).
2. Eumetopias stelleri (Lesson).
3. Enmetopias gillespi (M'Baiu).
4. Eumetopias einerea (Pdron).
5. Eumetopias liookeri (Gray).
6. Arctocephalus pu8illus(Schreber).
7. Arctocephalusfalklandicua, Shaw. |
Five are Hair Seals and eight are Fnr Seals. Three only are
given as found in the northern seas, while ten are recognized
as occurring in the southern.
From the foregoing it will be seen how widely opinions have
differed respecting the number of species and their generic
afBnities among recent writers on this group, and how unstable
have been the Aiews of the two leading authorities in this field
*See antm, p. 194, second footnote.
tG. Cuvier supposed it to have como from the Cape, because! Pagiia (see
Bufion's Hist. N.at., Suppl., vi, 357) had reported the young Otaries ol' the
Cape as of a l)lack color (Oss. Foss., 3d od., v, 220) ; but it is now Avell known
that all Fur Seals ar<3 black when young. On the other haml, Daubcutnii
insisted that Buffon's "Petit Phoiiuo" (sec Desmarest, Mam., p. 251) came
from "I'Indc."
{ J (i7ehfUppi i fvom Juan Fernandez and Masafuera, says:
" These three skulls have nearly the same teeth, and appear to
nie to belong to one group; btit whether they are three distinct
species (two from the west coast of South America and one from
North Australia) 1 will not attempt to determine, as 1 have only
seen the skins and skulls of the one from the latter region; but
they are all Fur-Seals and may be distinct.''t Dr. Gray says
his genus Qypsophoca "is most like Arctophoca in the position
of the teeth ; hut the pala*<) h much narroiccr, the face shorter, and
the hinder part of the ulcuU much larger and more ventricose^^',1^ but,
as Clark has shown,§ and as is evident from Gray's figures, Gyp-
sophoca was based on a young skull, and young skulls of Otaries
differ from adult ones of the same si)ecies iu just these characters.
It nuiy here be noted that in several instances the so-called "spe-
cies" of Fur Seals differ from others recognized by the same
authors only through differences that can be demonstrated to
be, ill other Avell-known allied species, simply sexual. Hence,
until writers ou this group have learned to discriminate the
sexes, and to make due allowance for the great changes iii
contour and details of structure that result from age in the
skidls of Otaries, we can hardly hope to have the subject of
species i)laced on a jiroper basis.
* Hand-List, p. 36. ~
t Hand-List, p. 28; first printed in Proc. ZooLSoc. Lond., 1872, p. 661.
tProo. Zool. Soc. Lond., 1872, p. 659.
$ Ibid., 1873, p. 759.
ii
.SI'KCIHS.
207
TIh' (listrilmtiiMi of Iho Kiir Srals in tlic Southoni Sea.s pix'
.st'iits no obstacle to the siijuiosition of tliolr t'Oiispeiillc rola-
tionsliij). TlH'y occur not only on both the Atlantic and Pacific
coasts of the South Aniciican contiiu'ut, about its southern ex-
tremity, and on all the outlying islands, including' not only the
Falklands, the South Shetland, and South Georjiian, but at
otiicr small islands more to the eastward, ut Prince Edward's,
the t'rozets, Kerguelen, Saint l*aul, and Ajusterdam, the south-
ern ami western shores of Australia, Tasmania, New Zealiind,
and at the numerous smaller islands south of the two last
named. They have been found, in fact, at all the islands nuik-
iuji up the chain of pelagic islets stretching somewhat inter-
rnptedly from Cape Horn and the Falkland Islands eastward
to Australia and New Zealand, including among others those
south of the Cape of Good Hope, so famous in the annals of the
soal-tishery. It has been stated by Gray and others that the
Cape of Good Hope Fur. Seals (really those of the Crozets and
neighboring islands) are far inferior in commercial value to
tliose of other regions ; but in tracing the history of the sealing
business I have failed to notice any reference to the inferioi-
(luality of those from the last-named locality, or that there has
been any difference in the commercial value of the Fur Seal
skins obtained at dift'erent localities in the Southern Seas. The
(jnality differs at the same locality, wherever the Fur Seals are
found, with the season of the year and age of the animals, so
that skins may come not only from the Cape of Good Hope,
but from any other of the sealing-i)laces, that one " might feel
convinced could not be dressed as furs," being " without very
thick under fur."
In this connection I may add that Gray's figure (Hand-List
of Seals, pi. xxiii) of an old male skidl of Aretocephalm antarc-
ticus so closely resembles an aged male skull (No. 1125, M. C.
Z. Coll.) of Arctocephalus amtralis { — falMandums, auct.), that
the latter might have served as the original of the figures ! while
other skulls of the last-named species bear a striking resem-
blance to Gray's figures of his Euotaria cinerea (Hand- List, pi.
xxvi) and Ids E. latirostris (ib., pi. xxvii). In fact, the series of
skulls of J.rctocepAa?M« australis in the Museum of Comparative
Zoology, from the Straits of Magellan and the west coast of
South America, presents variations that seem to cover all of
Gray's species of Arctocephalus and Evotaria as figured by him
iu the Hand-List of Seals.
208
FAMILY OTAKIIDA:.
Syiiiipsin of the (linvnt itiul SpccicH,
A. I'i'liimiharNli. witlidiit iiiiilcr-fiir. KiiiMNliort. MolaiH
(1— (I.
fi — 5'
Yo' SiH'cics j;t'iu'iiilly of liu'nc nI/.c. Color yt'llowish-ltrowii ; rcil-
(liHh-hrown y\hm youiin Tuicnoi-irocAiA:.
I. dcHUH Otauia, dill ex I'cron.
Otaria, P^:j{on, Voy. aux 'ft-rr. AuHtr., ii, 181(i, :J7, lootiiotti (in part).
[Platiirhhirhm] J'latyrhiiDiue, V. CUVIEK, Mi^in. dii Muh., xi, 1824,208, pi. iv,
li«. 2.
riatyrhhuua, Y. CuviER, Diet, des 8c. Nat., xxxix, 1827, T").— LK8.son,
Man. de Maui., 1827, 20:i (in part).
Otaria, Git.l, Proc. Essex IriHt., v, 1860, 7.
Char. Gen.— I'alatine liones extending nearly to the pterygoid i»roce88e8,
deeply eoneuvo, trunente behind. Molars ^^ = ^^.
1. Otaria jubata ("Forster") Blaiuvillo.
Phocajiibata, "Forster, 177.'j"; Sciirebeu, Erxleben, Gmelin, and other
early writers.
Phoca jubata, Forster, DeHerij). Anini. ndLicht., 1844, 317 (" Terra Statuum ;
Insula Novi-anni").
Otaria jubata, Desmarest. Mam., 1820, 248 (in part), and of most recent
writers.
t Phoca flavcscena, Suaw, Gen. Zool., i, 1800, 2C0 (young).
Otaria leonina, P£rox, Voy. aux Terr. Austr., ii, 1816, 40. Also of Desmarest,
Gray, Peters, and some others. ,
Platyrhincua honinun, F. CuviER, Lesson.
Phoca byroni, Blainville, Jonm. de Phys., xci, 1820, 287.— Desmarest,
Mam., 1820, 240 (Jide Gray, Suppl. Cat. Seals and Whales, 1871, 13),
Otaria moUosmna, Lesson vl Garnot, Voy. Co(|., Zool., i, 1820, 140, pi. iii
("IlosMalouines").
riatyrhyneus motlosainua et urauiw, Lesson, Man. de Mam., 1827, 204.
Otaria pernettyi, Lesson, Diet. Class. d'Hist. Nat., xiii, 1828, 420.
Otaria platyrhinchu8 et chilrnniH, MtUxER, Wiegmann's Archiv fWrNaturgesch.,
1841, :m.
Otaria honina, goihffroyi, hyronia, et uUow, Peter.'^, Monatsb. Akad. Berlin,
18(i6, 2(J4. 2»)(), 2()y, 270, (i/O, (>71.
Otaria uUoa; von Tsciiudi, Fauna Peruana, 1842-44, 13.'). 136, pi. vi.
Arctocephahta falklaiidiciia, Burmei.ster, Ann. and Mag. Nat. Hist., 3d ser.,
xviii, 186(), pi. ix, ligs. 1-4 (at least in part).
Otaria minor, et pyymmi. Gray, Ann. and Mag. Nat. Hist., 3d ser., viii,
326.
Otaria hookeri, Sclater, Proc. Zool. Soc. Lond., 1806, 80.
Habitat. — Galapagos I>*land8 (Coll. Mus. Comp. Zool., from Hassler
Expedition), and coasts of South America from Peru and Chili on the Pacific
side, and Rio de la Plata on the Atlantic side southward.
SPECIES.
209
II. Genun riiocARCTOS, Peters.
Aniocvpnnliis, ni part, of GHAY, i»rior to IHtJG.
I'liocanioH (milij;i'iuiM), I'l/iKiis, ALoiiatHb. Akatl. Berlin, IrffiO, 269.
CiiAii. (iKN. — I'lilatiiu' lioiu'H eniliii}; conHnltTably in front of tho pterygoid
iiKKcssci*, (li'ci»ly t'oucavo in front, nurrowotl and fniarginutc behind. Mo-
tl-t! 12
'i. PhooarctOB hookerl (Gray) PetcrH.
.iirhctphahiH hookcri, GnAY, "ZoiU. Voy. Kn-buH and Terror, pll. xiv, xv";
Cat. ScalH and WbaleH, Ir^fiO, .'»:», tig. 15.
VhiidrvloH lioohrri, Ghay, iSnppl. Cat. Seals and Whales, 1871, l.'i; Hand-
List Seals, 1H74, "29, pi. xx.
(iliiria jiiliata, Ai.LKN, Unll. Mns. Coinp. Zool., ii, 1870, 45 (in jtart).
iiliirid hookni, Ci-AHK, Proe. Zool. See. Lond.. 187:{, 754, ilgs.
llAiiiTAT. — Auckland Islands. (Orif^inally (h'serihed from specimens »«j>-
liimil to have come from tho " Talkhmd Islands and Cape Horn." See Clark,
US aliove cited, and Gray, Ann. and Maj;. Nat. Hist., 4th ser., vol. xiv, 1874,
\i\>. -jf;-;!!'.)
III. Getni8 El'METOPlAS, Gill.
Oliirid, in part, of earlier aiithors.
Kiimi1(>ii)(ts, Gll.L, I'roc. Essex In8t.,v, 18()(), 7, 11.
( iiAU. Gkx. — PaIatin(^ hones ending very far in front of pterygoid pro-
(csscs, Hat, or nearly so; hinder border hollowed or cniarginate. Molars
lzl~ fO' *'^*'' 1"*"' separated by a considerable space from the fourth pair.
3. Eumetopias stelleri Peters. *
IIaihtat. — Pacific coast of North America from California to Alaska;
I'acitic consf of Asia from Ja])an ncrthward.
IV. Genus Zalopiius, Gill.
Anloccphaliix, in part, of GiiAY, prior to 1866.
/.(dopliiis, Gii.l, Proc. Essex Inst., v, 1866, 7, 11.
XivpliDra, GuAY, Ann. and Mag. Nat. Hist., 3d ser., xviii, 1866, 231; Suppl.
Cat. feo-xls and Wh.ales, 1871, 28.
EumetopiuH, Peters, Monatsb. Akad. Berlin, 1877, 506 (in part).
Char. Gex. — Pala+ino much as in Eumetopias. Sagittal crest very high.
Interorbital region greatly constricted. Molars ^^^ = ^, in a continuous
.■'Ones.
4. Zalophus californlanus (Lesson) Allen.t
Habitat.— Coast of California.
5. Zalophus lobatus (Gray) Gill.
'HOtaria uWicollis, P^ron, Voy. Terr. Austr., ii, 1816, 118.
(^iaria cincrea, Gray, King's Narr. Austral., ii, 413.
ArdocepMhis lohatus, Gray, "Spic. Zoolog., i, 1828, pi. — "; Cat. Seals, 1850,
_ 44 ; Cat. Seals and Whales, 1866, 50.
* For synonymy, see infra, under the general history of Eumetopias stelleri.
*lor synonymy, see infra, under tho general history of the species.
Misc. Pub. ISo. 12 14
210
FAMILY OTARIID^.
m
■j
Ncophoca lobata, Gray, Ann. and Mag. Nat. Hist., 3d scr., xviii, 1866, 231;
Suppl. Cat. Seals and Whales, 1871, 28 ; Hand-List Seals, 1874,
43, pi. ..XX.
Otaria auatralia, Quoy &, Gaimakd, Zool. Voy. Astrolabe, i, 1830, 95; 1833,
pi. xiv (animal), xv, figg. 3,4 (skull), "Nouvelle Hollande."
Arctocephalus auatralia, Gray, Cat. Seals and Whales, 1866, i")7 (not Pima
auatralia of Zimmemianii and Kerr).
Otaria atelleri, Temminck, Faura Japon. (at least in part).
Phocarctoa clongatua, Gray, Hand-List of Seals, 1874, 30, pU. xxi, xxii.
Eumeiopiaa clnerca, Peters (ex l^dron), Monatsb. Akad. Berlin, 1877, 506.
Habitat. — Australian Seas. Japan??
B.— Pelage soft, with abundautnnder-fiir. Earelonger. Molars 5E5=Jq- Size
smaller. Color gray ; black when young OuLiPHOCAca;,
V. Genua Callorhinus, Gray.
Callorhinua, Gray, Proc. Zool. Soc. Loud., 1859, 357.
Arctocephalua, Gill, Proc. Essex Inst., v, 1866, 7, 11 (not of F. Cuvier).
Char. Gen. — Facial portion of the skull short, convex,
Molars 5^:1 = io^
G. Callorhinus ursinus Gray.
Habitat. — Shores of the North Pacific, from California and Japan (Pe/m)
northward.
VI. Genua Arctocephalus, F. Cuvier.
lArctocephalua'l Arctocephalea, F. Cuvier, Mdm. du Mus., xi, 1824, 205, pi.
iv, fig. 1.
Arctocephalua, F. Cuvier, Diet, dcs Sci. Nat., xxxix, 1827, 554.
Hdlarctua, Gill, Proc. Essex Inst., v, 1866, 7, 11.
Arctophoca, Peters, lionatsb. Akad. Berlin, 1866, 276.— Gray, Suppl. Cat.
Seals and Whales, 1871, 31.
Euotaria, Gray, Ann. and Mag. Nat. Hist., 4th ser., iv, 1869, 269; nanJ-
List Seals, 1874, 34.
GypaopUoca, Gray, Ann, and Mag. Nat. Hist., 4th ser., iv, 1869,269; Hand-
List Seals, 1874, 27.
Char. Gen. — Facial portion of skull slender, elongated, pointed, gently
declined. Molars ^^^^ = ^, much larger than in CallorMnua.
7. Arctocephalus australis (Zimincrmann) Allen.
Phoca uraina, in part, of various early writers.
Phoca auatralia, Zimmermann, Geograph. Geschichte, iii, 1782, 276 (= "Falk-
land Seal, Pennant, ii, 521")-— Kerr, Anim. King., 1792, 12^
(= "Falkland Seal, Penu., Hist. Quad. N., 378").
. PlMcafalklandica, Shaw, Gen. Zool., i, 1800, 256 (= "Falkland Isle Seal" of
Pennant — the Fur Seal of the Falkland Islands).
Otaria falklandica, Desmarest, Diet. d'Hist. Nat., xxv, 1817, 601, and of I
many subsequent writers.
Otoria s. Arctocephalua falklandicua. Gray, Peters, and others.
•For synonjrmy see infra, under the general history of the species.
SPECIES.
211
Olurla nhawi et hanviUei, Lksson, Diet. CIush. tVIIist. Nat., xiii, 1828, 425.
ArdoccphnluK nigresccns, Gray, Pktkrs.
Otaria (Arvtophoca) philippii, I'KTKns, Mouatsb. Akad. Berlin, 1866, 276,
pi. ii.
Utitria (Ardophoca) argcntata, I'lULim & I'etkijs, Moiiatab. Akad. Berliu,
1871, 560, pll. i, ii.
AntocqihaluH grayi, Scott, Mam. lieeeut and Extiuet, 1873, 19.
Kiiotarid latirosfris, Gray, Hand-List Seals, 1874, 37, pi. xxvii.
Habitat. — Galapagos Islands (sitecimens in Mns. Comp. Zoiil., Hassler
Exp. *) and shores and iglands of South America, from Chili and the Rio de la
Plata soatlnvard.
•Specimens of both Otaria juhata and Arctocephahis misiralis were col-
lected by members of the Hassler Expedit ion at the Galapagos Islands, show-
ing that they both rangi> much farther northward than has hitherto been gen-
erally supposed. For the following observations respecting their numbers
iinrl habits I am indebted to my friend Mr. J. H. Blake, artist of the Expe-
dition, who h.as kindly transcribed them from his note-book:
'^Charles Island, Galapagos Group, June 10, [1872]. — On an island at the
easteru side of Post-Offlce Bay is a Sea Lion rookery, where at almost any
time can be seen hundreds of Sea Lions lying at a little distance from the
water. Two of our company, in a little boat about ten feet long called the
'Dingy', went near the shore Avhere they were, when the Seals innnediately
ran into tlu^ water and surrounded the boat. The Seals came close to and
iindcr the boat, so that there was danger of their capsizing it, some of them
being as largo as the boat, and some were even hirger; hence it was deemed
Vi'iuk'ut to leave them. Toward evening the Captain, with others, took a
largfr boat and landed on the shore below the Seals, and while they were
nuniiug toward the water one measuring six o^ seven feet in length was shot.
Many of them were of enoiinous size, and great numbers could easily have
boon killed. They made a noise when rushing to the water louder than 'ho
waves on the shore. We saved one skeleton, and next day two half-grown
Seals were brought on shipboard and also saved.
'••fun'w hliiml, June 16, 1872. — At this island we saw many Seals, and some
were killed, one small one being preserved in alcohol. I went on shore in
the second boat, and as our boat landed we were surrounded with Seals of
different sizes, which came near the boat. Near where we landed was a
mother Seal und her two young ones lying together in a shallow excavation
tliey had made in the sand. They lay very quietly and appeared to be not
much disturbed by our presence as we gathered about them, excei)t when
we offerod to touch the young The mother was about six feet long,
and of ii light grayish color, with the head small and shaped like that of a
dog. The young resembled their mother but had shorter noses and were
about three or four feet long.
"In walking along the beach I came to another small rookery where
tlierc were family groups similar to that above described, lying about in all
kinds of positions, and so comfortably situated I did not disturb them.
One Seal, about six or seven feet long, which I met with at some distance
from the water, I drove some distance to study its movements in walking
and running. It would nearly raise itself from the ground and walk like
212
FAMILY OTARIIDiE.
i
ell
Hi' ''''
1
-1
i
H. Arctocephalus antarcticus (Tluiultirg) Allen.
Phom iirsina, roitsTKit, and in part of many early writirs.
t Phoca pmiUa, SriiKicniMi, Sihifj^et., iii, [177(>f], 1511 (=Le Petit Plioque.
Bnllon, based on .a yonnf^ I'lir Seal, from an nnknowu locality,
but stipposed to have come from India or t\w Levant,* but as no
Seals exist tbere, and as many ;Miinials Avbieb, in fonner ycarsi.
jtnr]torted to liave been brouj^ht from India were found to lie
really African, some late writers have assumed that Butl'iiu's
"Petit Pboque" must liave been also Afrean, but the pertineiicc
of the name 7)«s(7/« to tlie African I'ur .Seal is not beyond reason-
able doubt). t Also of EuxLEBKX, Gmklix, and others.
f OtariapiiKilla, Dksmakkst, ^ouv. Diet. d'Hist. Nat., xxv, 1827, <>0*i (liiiswl
on (he sauuO.
Otaria puailla, Pktkus, Monatsb. Akad. Herlin, 18()(), 271, f)71 (nauio adopted
from Sehrebtu').
Arctovt'plinhix piixUliiK, Pktkks. Jlonatsb. Akad. IJerlin, 1H77, 506.
/ ^* I'lioca parra, Hoddakkt, Elenehus Anim., pi. Ixxvii" (= liutt'on's IVtit
Phoque, as above).
Phova antarrtiva, TiifNHKm;, Mem. Acad. St.-P<pl. Cat. Seals an9, 107, IlfiO, pi. Ixix;
Cat. Seals and Whales, IHWJ, .Vi; Anu. and Mag. Nat. Hist., 3d
M'l"., xviii, IdlU), 235.
Anloic]iliahtaf(ilklati(HviiH {hi iKirt), Gkay, Cat. Seal.s, 1850, 42.
Jrcloeephalun nrhinthyprrovx, Ti'itXKit, .lonrn. Anat. and Pliys., iii, 1868,
113, fig.
Antuccjilitiluii tiiroHiiH, Gkav, 8ui>i)1. Cat. Seals and Wluvles, 1871, 27.
/ Euotar'iH ivmjtreami, Gray, Hand-last, 1874, 38, pi. xxiv ("South Africa T").
Habitat. — Cai)c of Good Hope.
TO. ArctocephaluB foreterl (Lesson) Gray.
Phoca ursina, FORSTEB, Descrip. Auiin. (ad Lichtenstein), 1844, 64 (New
Zealand) = Sea Bear, Forster, Cook's Second Voyage, 1777,
= Ours Marin, Biiffon, Hist. Nat., Suppl., vi, 1782, 336, pi. xlvli,
so far as it relates to Forster's figure and notes).
Otaria fomtiri, Lesson, Diet. Class. d'Hist. Nat., xiii, 1828, 421 (=:Sea Bear
of Forster, whieh became, later, Phoca Hr»ina, Forster, exclusive
of references to Stollei-'s Ursus mnrinun).
AntocephaIii»J'orntiri, GRAY, Snppl. Cat. Seals and Whales, 1871, 25.
Otaria ciiima, QuoY &. Gaimard, Zool. Voy. Astrolabe, i, 1830, 89; Atlas,
1833, pU. xii (animal), xiii (animal), xv, tigg. 1,2, skull ("Nouvello
Hollaude"; probably not Oiaria vhnrid, Perron, Voy. Terr. Austr.,
ii, 18(5(5, 54, 77, which, however, is indeterminable). — Peters,
Monatsb. Akad. Berlin, 180(5, 272, (571 (exclnsive of some syno-
nyms).
Anton phaliix diifreim, (Jray, Cat. Seals, 1850,43; Cat. Seals and Whales,
18G(), 5(5; Snppl. Cat. Seals an\l Whales, 1871, 24, etc.
Liiotaria nncrva, GRAY, Hand-List Seals, 1874, 34, pi. xxvi.
f Otarui lamarii, MCller, Wiegmann's Arch. f. Naturgcs., 1841, 334 (in part
Hi \onHt,—fide Peters, Monatsb. Akad. Berlin, 1866, 271, 272).
(Ijipsophoca tropicalis, GRAY, Proc. Zoiil. Soc. Lond., 1872, (5.59, figg. 5, 6;
Hand-List Seals, 1874, 28, pi. wiii.
fAntoplioca gazelln, Peters, Monatsb. Akad. Berlin, 1875, 396 (Kergueleu
Island).
J Otaria (Arctophocn) cleyaitH, Peters, Monatsb. Akad. Berlin, 1876, 316 (St.
Paul Odd Amsterdam Islands).
Arvtoceplialm Vrcripes, forateri, fclegans, ot f gazeUa, Peters, Monatsb. Akad.
Bovlin, 1877, .507.
llAniTAT. — Australia, New Zealand, Auckland Island ; f Kergueleu Island ;
f Saint T'uul and Amsterdam Islands.
*" Otaria ihlalamlii, F. CuviER. Diet. Sci. Nat., xxxix, 423," cited by
lisflicr (Syu. Mam., 232), and repeatedly by Gray and by Peters, is evidently
on'oiu'ous, lis the article " Phoqmi " begins on p. 540, and no spocics of Seal or
Otaiy is nienticmed on p. 423. The correct citation is not " Otaria dela-
kndii" Init " Olarie de Delalandc," as given above. G. Cuvier refers (Oss.
Fos.. ;til (Ml., v, 1825, 220, pi. xv5" , fig. 5. skull) to it as "Otarie du Cap" " recu
Piir M. Delalande."
214
FAMILY OTARIIDiE.
MYTIIICAIi AND UNDiyi'lOKMINAllLE SPECIES.
In tlic. |)n!(;(i«liiif;' piij^c's r<'i<'i'('.n<'('- has bocii iiiiido to v.'irious
siMMrics (IcsciiluMl loo iiiipcilecMy to iidinit of rcoo/^iiHioii.
Sonic ol" lln'sc, 1 liiivo (loiilttl'iilly ;illo(!iito) (lalls it ''Ic
IMitit phoipie noil' 5, described iiiid
lij;iii'cd under that name by Parsons in the Phil Trans, xlvii, t.
(i, and iH)ticed inrennant's '(J|iia«lrupeds', ii,p. ii74. Dr. Slinw.
ill his 'Zoolojiy', i, ]». 250, Iranslatcd the iiaiiic. into I'lutni Ion-
{ficollis, ami copied Parsons's lijiiir<'s. The name; and the loriii
of the fi'ont feet are enough to show that it is an Fiare. 240, overlookinji' this character aiul the (h's<'rii»tion of tlic
front feet, (considers it as the saiiK*. as tin* Sea-Leopard of AVcd-
dell {I'lioea Weddcllii) from the Antarctic (hreaii, an Earless
Seal. Thou{;h the habitat is not ;;iven, there can be no donhf,
when we consider the, ••eojiraphieal distribution of the lOiircd
Seal, that it must have been receiv«'d either from the soiithcni
part of South America or from the (Jape of (rood Flope, as tlic
animals of the North Paeille and of Australia were not known
or brou^jht to l<^ny,iand in 1(JS(». As no a<'count of the i'olorot
the fur is j;iven, it is impossible to (h^termine to which species
inhabitiufjf these c(mntries it should be referred. It is most
I)robably the Sea Lion {Otaria leonina), as that is the aniiiiiO
which is most {generally distributed and commonly brou{,ditt(!
England. The sailors sometimes call it the 'Long necked
•Ann. iinil Mii^;. Xuf. HiMl., 4tli Hcr., i, 18(W, j)]), 217, 218,
mtgmmm
SPECIES.
215
Soul'." ^Jniy, liow(;v('r, liinl Ibniicily iclornMl if (l()ul)tliilly
(C)it. Seals, lS.j(>, i;{; O.it. S«':i1h aiul WliiilcH, ISfX;, r.(5) to
An-lorcplitiliiN foll.hniiliciis.
.!. I'lioni JhirrNcnis, SlIAW, < Icii. Zoiil., i, LSOO, li(J(), ii .siiiiill,
••vfllowislr' l')iir<'(l Seal, (l«'S<'ril)('(l ('loiii a spcciiiH'ii in llio
liCvcriaii .Miisciirii Itioiijilil from llic Straits of Ma;;«'l!aii. It is
tlic "l-laicd Seal" of I'ciinaiil ((^>iia(l., ii, L'TS), antted with yellow, the Ibro
feet are said to hsiv. Otariafabricii, LESSON, Uict. Class. d'Hist. Nat., xiii,1828,
419,= Phoca ursina, Fabricius, Faun. Grcenl., 6. Based on a
216
FAMILY OTAKIID^.
■ii
supposed species of Eared Seal erroneously believed by Fabri-
eius to exist in the Greenland seas, but wlio never saw tbe
animal, and described it mainly from what were tcfl?i»."
218
FAMILY OTARIID/E.
indications of tlioirf(»ni!('ri)i('scncT in Knioiicofsiu'cinl intorcsi,
and calls l"oi- a <'ntical examination of the sn]ti)oscd evidence ot
tlieir loiiner existence tliere.
(iervais, many years sin<-e,* described and liyni'ed a tooth
which he releired, with (hnibt, to Otaria {'■^Otnria i priscu"),
Itnt Van Ueneden has since determined it to be referable to
Sqtidlndon. ]M. K. l)elfortrie,+ in 1871*. described two fossil teeth
fi'om the bone l)reecia of Saint-Medard-en-.Ialle, near Uordeaiix.
which he considei'cd as represent injj;" two s]»ecies of Otary, wiiich
he named, iv ])ectively, 0/tO(l as satisfactory ])r()of of tlie presence of Otaries iu the
Tertiary fauna of lOiiropo.
N'mii lU'iK'dcii also n'fcrs t(» a Immorns oi'un Otary iu the Mu-
s(Miiii of tli(^ CJeoloyieal Institute of Vienna, supposed to have
hccii taken from the bed of the Danube, and adds that it bears
ii close resenibhince to the same part in Otariajnhata, if indeed
it is not referable to that species, but adds: "Get os, en
tout cas, n'est pas fossile," He also refers to a skull found by
Valenciennes on the shore iu the department of Laude, meu-
tioned by Gervais,* and says it is still unknown how it came to
bo fount on the coast.
Van r>enedcu, however, believes that he has proof of the ex-
istence of fossil Otaries in Europe iu his Mesotaria ambigua,i a
species i)reseuting many remarkable charactei's, which ally it,
he believes, in some points, to the Otaries. This species is rep-
resented by the greater jiart of the bones of the skeleton aud
iiuinerous teeth, but the skull is not known.| The teeth, he
says, arc unlike those of any other genus, while the bones indi-
cate a s])ccial mode of life, and a size about equal to or rather
larger than that of Phoca (jnenlundka.^ The ilium is described
as resembling more the san)e part in the Otaries than the Seals,
and as indicating a mode of life more terrestrial than aquatic.
Tlie humerus, on the other hand, is stated to more resemble
tiiat of the Seals than that of the Otaries.
Of tlie fenuu' he says: "Nous avons trois fenuirs assez complets
qui indiquent que cet os s'^loigne par sa conformation des autres
Aui]>hit(''riens. La tete, ainsi que le col, tieuLcnt de I'Otarie,
coiiune les condyles, et le (jrarnl trochanter, pen large, ue s'eleve
I>as au-dessus de la tetc de I'os. La tete est comparativement
petite. La cavit6 trochant<5rique est profonde et etroite vers le
milieu taries etaient moins aqua-
ti([ues que les Phoques actuels."
Upon careful comparison of his excellent flgures (pi. ix) of
the fenuir, humerus, scapula, and fragment of pelvis, with the
*Zo()logie ct la Pal^-ontologio fran5ai.es, p. 27<>.
tAim. (In Mns. Roy. d'Hist. Nat. do Belgiqne, i, 1877, p. 5C, pi. i.
{Van IJenedcn reports having two canines, three molars, seven cervical
vcrtcline and an axis, eix dorsal and seven Ininb.ar vertebrie, aright ilinni and
11 left iscliinui, the distal end of a scapnla, four right and five left humeri, a
loft and a right femur, six tibiic, and four metatarsal bones.
^ The parts of the skeleton figured by Van Beneden correspond very nearly
iu size with the corresponding parts of Cj/ntophora cristata.
220
FAMILY OTARIID^:.
rorrespondiiifj; i)arts of tin: skelt'ton in Iiv«' siu'cies of Otarics,
vcpirst'iitiii;'' all tlio j-ciu'ia of that yronp, aiul with tlio primi-
l»al types of tlu' IMiocids, 1 fail to api)r.t'<'iat«> any iiiij)ortaiit up
l)roa('h toward thf fonnor, or any niarketl departure from tiic
latt«'r, especially tiie snl)fauiil> Ci/ntophorina'. In tii«' femur, f(»r
e.\anJi)le, there is in Memtaria no trace of a trochanter mi)tree fnnetional,
1111(1 the dental formnla of the temporary teeth is snbstan-
tially the same in all. In the Walrnses, however, two of the
posterior npi)er milk nudars and the last lower one often remain
till a comparatively late jjeriod of life, but all traces of the
otliers disappear soon alter birth. The two nuddlo pairs ot*
incisois probably never pieree the <;ums, and the others scarcely
]i('isist beyond the fcetal period. The feruuda for the temporary
dentition of this <;roni) is usually reeofjfnized as I. i|~;J, C. \~^,
M. ,|~..| iov M. ]~l)- I" tl'^ Seals, however, the nund)er of
molars appears to never exceed ij^jj • l'» the P^arless Seals ''the
milk-teeth are extremely rudimentary in size and form, and per-
tl'c'tly lunetionless. The majority of them never ent the gums
and are absorbed actually before birth, and certainly within a
week after birth scarcely a trace of any of them remains.''*
The milk molars are three in inind)er on each side, both above
aud below, and are replaced respectively by the second, third,
and fourth molars of the permanent set. The canines are
all represented in the temporary set. The number of tem-
porary incisors varies in the ditterent genera, it corresponding
up arently with the juimber in the permanent set. In Phoca
I- ilina, 1\ j/r th above and below, the middle one was much
smaller than the others. These alveoli are exterior to the per-
uianent teeth, which do not vertically reiduce them, that of the
Hist upper milk molar being opposite the spaee between the
tirst and second permanent molars; the second opposite the
space between the second and third permanent molars, while
the third is nearly opposite the fourth tooth of the permanent
set. In the lower jaw the alveoli of the milk molars are re-
spectively just behind and exterior respectively to the second,
third, and fourth permanent teeth.
Ill three foetal skulls of ZaUphiu (No. (•15G, Mus. Comp. Zool..
Xos. 15G60, (J , 15661, 9 , Nat. Mus., all from the Santa Barbara
Jslands), the milk teeth are all still in situ, except the middle
incisors, which are replaced by permanent incisors that were
apparently about ready to pierce the gum. As in EumetopiuH
and Arctoeephalns, the middle molar, both above and below,
is much the smallest, and is placed very close to the third, leav-
224
FAMILY OTABIIDiE.
IM
m
k--
inj; ii vory brand ijitcrval Ixitwcon tli
ree«)rd the followiiiff instanees: In (UillorhiniiH ursinus I have
noted the following' irref,adarities: Skull No. L'DL'l* (M. ('. Z.
(!«;ll.) has M. J^~J, the normal niiiiiber cent, of the skidls of these two spexamined not less than thirty of ea«!h) lU'esent one <»'
more supernumerary molars. I have also found su|)i)ressi(»ii <>t
molars in ArvtoatjihabiH amUutliH (M. i\ Z. Coll., N<». ll.'U,—
M. :■-•;).
The supernumerary molars are placed (in all the iiistance.s 1
OKNERAL OHSKRVATIONS.
225
liavt' seen) hcliiiid tin- liist, molar ol' flic, noniiiil ,s«>ri<'H. They
iii'c iisiimII.v smaller than the iiocrna! inolai'H, sonictiiiieH alinoHt
iiidiiiifiitai'.v, usually wilhont accessory cusps, and with ii
siiiiiotli or nearly Ninooth ciii^'iiliiiii. They are lieiKre <>;eiierally
M-ce^^iii/altle by their size and I'orni. In (^ases of snppressiou
:', is usually th<'. antepennltiniate nu)hir rhaf is missing'. This
iiiohir also I'rccpHMitly tails late in lite, Init traces of an alveolus
ill siicli cases usually atti'st its t'onner presence.
I'OSlTKtN ()!•' I'lli; LAST IJIM'KK I'lOUMAM'-N'r MOLAK.
Ill species with the snp<'rioi' molars a — 5, the last (<'.\cept in
Eiiiiirtojti(iH) is j»la<*eoHte«ior to that of th«
last molar in tlu', six-molare,di Hair tS(!als, arc* hitermediate in size betwetuithe jtlier
Hair Seals and the l''iir Seals. All th<', Ifair S(^als hav, «;osu'.se,
iianl, stilf hail', varyinn" in length with at;e siml seiison, and are
wliolly wit hout soft underfnr. All tlie Fur Hcnils hav
^"^^O" {=75-100.
76-100 5
«1-100 ). 4.100.
66-100 i
88-100 )^g5.1,X).
83-100 >
77-100)^73.100.
75-100 >
HABITS.
227
times tliat of the adult I'euiales of tlio sanu' spocios. There
aro also veiy great ditt'ereiKH'ts in the form of the skull, esi)e-
cially in iv'spect to the (leveh>]uneut of crests and i»rotul)erauees
for inuscuhir attaehnieut, these being- only slightly (h'veloped
in females and enonuously so in the males. Witli such reunirk-
iible variations in eolor and eianial characters, di'pendent upon
age and sex, it is not a matter of suri)rise that many nominal
species have arisen througli a uiisai)i)reeiation of the real signiti-
cauce of these differences.*
HABITS.
Tlie Eared Seals show also a remarkable resemblaiu'e in tlieir
gregarious and polygamous habits. All the species, wherever
occurring, like the Walruses and Sea Elei)hants, resort iu
great numbers to i)articular breeding stations, which, in seal-
ers' parlance, have acquired the strangely inappropriate name
of " rookeries." The older males arrive tirst at the breeding
grounds, where they immediately select their stations and await
tbe arrival of the females. They keep up a perpetual warfare
for their favorite sites, and afterward iu defense of their harems.
The number of females acquired by the successful males varies
l'roh> a dozen to tifteen or more, which they guard with the utmost
joalous;;, — ujight being with them the law of right. The strong-
est males are miturally the most successful iu gathering about
tlu'ui large harems. The nuiles, during the breeding seasouT^
remain wholly on laud, .and they will sufl'er i»ed by short, very
soft, tine, j^ray overhair. Later in tlu' season, and esi)eciiill.\
iu the old aninuils, the overhair is coarser aiul longer, and even
somewhat harsh, beneath which, however, is still the heavy
soft underfur. Dealers sort the skins into grades, in accord
auce with the size of the skins and the quality of the fur, these
features depending upon the age and sex of the animal, ratlior
than ui>ou the species. i)r. Gray refers to what he <'alls Ardo-
ceplialux faMandkm as being "easily known from all other
Fur Seals in the British Museum by the evenness, shortness.
i)ears not to difler ai)]>re('ia-
l)lv from llie otlu'rs.*
DKSIinCTION »»1' TlIK Fll{ SKALS I'OJI TlIKlll I'KLTlllES.
Tilt' value of tlie jx'lti^es of the l-'nr Seal has led to whole-
sale destruction, amounting at some localities ahnost to I'xter-
iiiiiiatioii. TIh^ tratlic in their skins appears to have begun
toward tlie end of the last century. Captain Fanning, of the
sliii)"r)etsey,"ofXew York, obtained a full cargo of choieo Fur
Seal skius at the island of ^Nfasafuera, on the coast of Chili, in
179S, Avliich lie took to the Canton market. Captain Fanning
states that on h'aving the island, after procuring his cargo, he
ostiinated there were still left on the island between 500,000
and 700,000 Fur Seals, and adds that subsecjuently little less
than a million of Fur Seal skins wei'e taken at the island of
Masafuera alone.t a small islet of not over twenty-five miles
ill circiuiiferencc, iind shijiijed to Caiiton.| Captain Scammon
states that the sealing fleet otV the coast of Chili, in ISOl,
aiii(»iiiited to thirty vessels, many of which were ships of the
h\Y\n'v class, and nearly all carried the American tlag. Xot-
witlistaiidiug this great slaughter, it ai)pears that Fur Seals
ooiitiiiued to exist there as late as 181."), when Captain Fanning
apiiii obtained them at this island.§
In the year 1800, the Fur Seal business appears to have been
at its height at the Georgian Islands, where, in the single season,
112.000 Fur Seals are reported to have been taken, of which
57,000 were secured by a single American vessel (the "Aspasia,"
iiiulor Captain Fanning). Vancouver, at about this date, re-
ported the existence of large numbers of Fur Seals on the south-
west coast of New Holland. Attention was at once turned to
this new field, and in 1804 the brig " Union," of New Yoik, Capt.
Isaac Pendleton, visited this part of the Australian coast, but not
finding these animals there in satisfactory numbers, repaired to
Border's Island, where he secured only part of a cargo (14,000
skins), owing to the lateness of the season. Later G0,000 were
(•btained at Antipodes Island. About 1806, the American ship
'S. :i64.
ni'.,p.299.
li
2:30
FAMILY OTARIIDiE.
-f
"Catliiuiue," of New York (Capt. IT. raiiniiifi), visited flio Cio-
zetto Islaiids, whore tliey landed, and found vast nunibeis ot
Fur Seals, but obtained tlieir earj^o from Prince Kdwnid's
Islands, situated a few liundred miles southeast of the Cape of
Good Hope, where other vessels the same year obtained lull
carfi'oes.
In ISoO, the supply of Fur Seals in the southern seas luul
so ^ireatly decreased that the vessels enjiaged in this enterpiist'
"{generally made losinj;' voyages, from the fact that those
places which were the resort of Seals," says Captain r>enjaiiiiii
Pendleton, "had been abandoned by them, or cut oif from
them," so that the discovery of new sealing grounds vas
needed. Undiscovered resorts were believed to exist, from tlie
fact that large nund)ers of Fur Seals Avere seen while eruisinj;
far out at sea, which must repair once a year to some favorite
breeding station.*
Captain Weddell states that during the years 1820 an,(HHi
in bulk, which, through hack of care in curing, si)oiled on the
way, and on the arrival of the ship in London the skins were
dug out of the hold and sold as manure! At about the same
time there was a similar wasteful and indiscriminate slaughter
of Fur Seals at the Aleutian Islands, Avhere for some years
they were killed at the rate of 200,000 a year, glutting the luiii
ket to such an extent that the skins did not bring enou},'h to
defray the expenses of transportation. Later the destruction
of Fur Seals at these islands was placed under rigid restrio
tions (see infra the general history of the Northern Fur Seal).
ill consequence of which undue decrease has been wisely pw-
* Faniiiiig's VoyngeH, p. 487.
m
m
GENUS I.UMEIOPIAS.
231
\ ciitod. But ii()\\ii('r<> ('l.s»> liiis there been systematic protection
of tlie Fur Seals, or any nn>iisnres taken to prevent wasteful or
iiiidiic destruction.
(iENu:>« EUIMETOPIAS, (fill
(Hiiria, ill imit of various smllKirs.
AfilKirphuhi^ (ill |>;nl), OitAY. Cat. Seals and Wliali's, 18()(), 51.
IjiiiiiUpiux. (ill, I,. I'nic. Kssi'x IiiHfitnto, v. 7, 11, July, li^6(i. Type '• Ofaria
t)ilii'iinii(iiiiin, I/i'ssou, = Arclorrphalua montt'rienmn, Gray."
Molars ,|~ J =|[;; the ujjper hinder pair separated from the
others by a considerable interval; the last only double -rooted.
Postorbital ]>roccsses (juadrate. Palatine surface of the inter-
iiiaxillaries tiat, only slightly de])re.ssed* and greatly contracted
posteiiorly ; tlu^ i)alatals moderately jtroduced, extending about
three-fourths of the distance from the anterior end of the zygo-
matic arch to the pterygoid ]>rocess; their posterior margiu
straight, ()!• slightly or deeply emarginate; rarely deeply so in
old age.
JjKinetuinas differs from Oturia, as restricted by Gill, in hav-
ing one pair less of upi>er molars, a inU(;h less posterior exten-
sion (»f the palatine bones, and in having the posterior i>ortiou
of the palatal surface less than one-third, instead of more than
one half, the width of the anterior jjortion, and but slightly in-
stead of deei)ly dei)ressedj also in the greater depth of the
skull anteriorly, and in the less de^■elopmellt of the occipital
and sagittal crests. In Euimtopins the depth of the .skull at
the anterior border t)f the orbits is nearly as great as in the
plane of the occiput, while in Otaria these i»roportions are as
13 to 18, there being in the latter a marked declination anteri-
orly in the superior outline of th(? skull. The breadth of the
skull at the temiioral fossai is also much greater than iu Otaria ;
that is, the skull is nuich less constricted behind the orbits.
The postorbital processes also differ considerably iii form iu
the two genera, while another noteworthy difference is the uu-
usnally great development in Otaria of the i)terygoid hamuli.*
*A coniparisou of mlult lualo skulla oi Eumeiopias and Otaria, of strictly
t'oricsiiondiug ages, shows the following differences :
Eumvioima stcUeri (No. 176.")): height of skull iu occipital plane 155 mm.;
lieiglit of skull at anterior edge of orbits 152 mm.
Otaria jubata {No. 1095): height of sknll in occipital plane 180 mm.; height
of skull at anterior edge of orbits 130 mm.
Comparing the same skulls in respect to the development of the pterygoid
hamuli it is found that when placed on a plane surface the skull of E. aiel-
'2:\-2
EiMKToriAs .sti:lm;ri — s tellers sea lion.
Kuiiutopias (lilVcis IVoiii Zdlnitlin-s tliioii^ii the jnosciM'o ol' a
wi(U' space iK'twecii the lotutli iiiid lit'fli i)airs of ii]»ju'r molars,
the less t'lnaijiiiialioii of llic jiostcrior lioidcr of the palatiiu'
l)oiu's, tlio (luadiate instead oi' the tiiaiij^ulav and posterioily
pointed form of tin- postorbital proeesses, li'c less relative
breadth of the i)osterior nares, and the larger size of tlu' faciiil
angle; also throngh its nuieh broader mnzzle. the less degree
of the postorbital eonstrietion of the sknll, and its nineh less
developed sagittal crest.
Enmcf(>pias difl'ers too widely fr»)ni ('(lUorhinUN and Ardoce-
2)hal)is, in dentition and cranial characters as well as in size and'
pelage, to render comparison necessary. The genns is at once
distinguishable from all the others of the family by the wide
space between the fourth and fifth upi)er molars. In distrihu
tion it is restricted to the shores and islands of the North Pa-
cific Ocean, ranging from Southern California northward to
Behring's Straits. Its geographical rt'presentative is the Otarin
jiihata of the Southern Seas, which ranges from tlu^ equatorial
regions (Galapagos Islands) southward.
EUM1':T0PIAS STF.LT.EIU, {/.mo») PHern.
StcUer's Sra Lion.
Leo mariniis, Stki.lku, Nov. Coniiii. rctvop., xi, 1751, :5(iO.
Phoca juhaia, Scin?EBKn, Sangetli., iii, 1778, ;?0(), i»l. Ixxxiii 11 (iu part only;
uot P. Jnhala, Forstpr, with wliicli, however, it is iu part con-
foundi'c)). — Gmkmx, Syst. Nat., i, 1788, 63 (iu part ; = P. _/H&a<(i,
Schrol»cr). — Pandeu &■ D'Altox, Skolcte dor Eobbou und Lamau-
tine, 182(i, ])!. iii, figs, d, e, /.— Hamiltox, Marino Aniphib., 1839,232
(iu part — not the fignrc of the skull).
Phoca (Otaria) jubata, Richardson, Zool. Beechey's Voy., 1839, 6.
Otaria jubata, P£uox, Voyage Terr. Austr., ii, 1816, 40.— NiLSSON, Arch. i.
Naturgeseh., 1841, 329 (iu part only ; includes also the true Otaria ju-
bata). — ? Veatcii, J. E. Browne's Resources of the Pacific Slope,
[app.], 150 (probably only iu part, if at all).
Otaria steUeri, Lesson, Diet. Class. Hist. Nat., xiii, 1828, 420.— J. MCller, Ar-
chivf. Naturgeseh., 1841, 330, 333.— ScHiNZ, Synop. Mam., i, 1844,
473.— Gray, Cat. Seals iu Brit. Mus., 1850, 47 ; Cat. Seals and Whales
iu Brit. Mus., 1866, liO.— Sclater, Proc. Zool. Soc. Lend., 1868,
190.— Scott, Mam. Recent and Extinct, 1873, 22.
Phoca atelleri, Fischer, Synop. Maai., 1829, 231.
leri rests anteriorly on the mastoid processes and the points of the canines,
the points of the pterygoid hannili being several millimetres above the plane
of rest, while in O. jubata the skull in the same position, rests posteriorly on
the pterygoid hamuli, which project f)™"* below a plane connecting the mas-
toid processes and the points of the canines.
^i »ii
EXTEKN.M. CHARACTERS.
233
Olai ill [,/■*: iiK'lop'nix) f^tillcii. ri".Ti;i!s, Moimtsl). Akml. liirliii, l-iliJ, ■•i7l,(')71.
Einnrliijiinn stclhri, ('>U\\. Aiiii. mihI Maj;, Xal. llisl., :!il scr., xviii, 18:i;'), 'i:'.:'. ;
Sni)i>l. Cat. Seals ami ^V hairs in lit it. Mils.. 1-^71. :'(l ; I'nir. Zoiil. Sor.
Loud., IHT'J. 7;!7 (in jiart), liK«. ••, ■'> ('l'" yoiiim skull nii wliii li
.iriloci'ithtilii-^ itioiilcniiixin,livn\. was in [larl liascih; I'luc Zixil. ."^or,
Loud., 1-'7;i, 77<) (its (•(•curiciici' in ,lai>an siarcd lolic ^74, -in.— .\u.l'.x, Bull. .Miis. Coui]). /ool., ii.
1870. ir). jdl. i, iiijiyy;. it-lT), and li.u^'. 1 ■.'>in text.— ScammoN, Maiin--
Maui., l~7-4. i'J4. (our woodcuis of iniinial \<\>. l-'li, l\;7. — Li.i.ioiT,
Kf'l)oit oil the I'lyldlov or S (iioimlar aciounl, with liffiircs).
'Stemmiito^nan Htdlcri, \\s Ukni-.dk.n, Ann. dn Mus. Roy. d'Hist. Nat. dn IJcl-
yiciui', ]it. i, ]H77, l.">(in text), — la]iHus pcniia' foi' L'liiiiclopiun nlvlhri .*
Pltoca Iconiini, Pai.la.'^, Zoog. Rosso-Asiat., i, IKU, 104 (^— /'.,/«/«(/«, (iincliu).
Arctocrphahiit monU'ru'tigin, GliAY, Proc. Zool, Soc. Lond., Ifihi), :{5H, ;U>0, pi.
Ixxii, .sknli (in part only; the .skin rcffvaMt' to ('uUorUiuHn urxiinis):
Cat. Scal.s and Whahs, IHdO, 4t).
ArctiHq)halusmUj'oniiaiuiii,GnA\', Cat. Seals andAVhalcs, IHtili, .'>! (:^A. nioiili-
/•(('M.SI.S, Gray, lf')\), in part ; not r^- Oluria cnliforninnn, Loason).
IJimitiii>iiin caliJ'oniianii>i, (iiLL, I'roc. Es.scx lust., v, IHtitl, 115 {^^ .Inlocepliuluii
iiioiitn-icimH, Gray, 1859," and i)ossil)ly al.so [identical] with O/ana
xUlleri, Miiller": lienee not =::- (>. lalifoniiaiKi, Lesson).
? Eiiiiiiliiiiidti rloiigulim, Glt.vv, I'roc. Zool. Soc Lond., l<-'7;{, 77t), tigs. 1. 'J
(— A', stelleri, Gray, ib., 187-J, 7;?7, tigs. 1-:?, .Japan ? ).
n'liiiinrclnx cldiiiiafiifi. GlJAY, Hand-List Seals, etc., 1H74, :?n, pi. xxi, xxii.
]ll., vi, 178*2, X\7 (in ]>art only).
LcniiiiK Seal, ricxxAxr, Arctic Zool., i, , '200 (in ])art only).
Lion Moriii, CllOUls, ^'oyag•c PittorcsiiiU', lies Alt5outieuue, 18'iiJ, V2 (not
— Lion mariii dr In Califoniic, pi. xi, ''I'ort Sau-Fiancisco et scs
Habitants").
Leo muriiiiiM, the Sra Kuit/, Em.iott, Scribner's Monthly, xvi, 87!), Oct., 1878.
"Sec-Vitchic," Russian; Lion maiin, French: iSccldice, German; Sea Lion,
Hair Seal, English.
Habitat. — Shores of the North Paciti(% from Behriii<>'s
Strait.s .southward to California and Japan.
External Charactersi. — Length of full-grown male eleven
to twelve and a half or thirteen feet, of which the tail forms
three or four inches ; girth about eight to ten feet ; weight vari-
ously estimated at from one thousand to twelve hundred or
thirteen hundred pounds.* The weight of the full-grown female
* A .skull of this species in the National Mnsonm (No. 4702), collected at
Fort Poiut, Bay of San Fraucisco, .luly, 1854, bears a l.-ihel with the fidlow-
ing l('j;i'iid: "Length 13 ft. 8 in. ; weight, by estini.ate, one ton."
Captain Bryant, in some MSS. notes on this species recently received,
states that the full-grown male measures 13^ to 14 feet from the tip of the
nose to the end of the ouMretchcd hind-feet, and from 7i to 9 feet in girth
231 Kl'MllTOIMAM STFJ.LKRI — STELLKH's SEA LION.
is said to rnnjjc from lour ImiMlicd tollNc liiiiidrcd poiiiHls, with
ii lon^rtli of ci^ilil to nine tWi. Tlic color Viirics uitli ;i^M' iiiul
scnsoji. Tlic yoiiii;;' sue ''of :i ricli diirk ('lit'stiiut-l)r(»\vii."
TIk' iiduUs, on their first iirrivid iit tlit'lr breed iti^-;,n'oiiiHls in
spriiij;-, present no sexiinl dissiniilarity of color, wliieli is tlicii
lij-iit l)ro\vnisli-rnfoiis, «'arker lteliinatches of dark brown on a yellowish-rufous ^-round [KUiott).
\\\ two adult males in the .Museum of Comparative Zoiiloj^y.
and another adult male in tin* Xational Museum, the jjcnt ral
color of the upi)er side of the body ^'arios from pale yellow isli
brown to reddish-brown, beeominy; much darker toward the
tail. Tlie sides below the mediairiine are re\vn.
ax»proaching black, especially «'xtenially. Th«' hairs are indi-
vidually variable in color, some beiny entirely pale yellowish.
others yellowish only at the tips and dark below, while othcis
are wholly dark reddish-brown or nearly black throujihoiit.
The relative proportion of the light and dark hairs d«'teriniiics
the general color of the body. The pelage consists of two kinds
of hair, the one abundant, straight, stiff", coarse and Hattemd.
and constituting the outer coat ; the other very short, exceed
ingly sparse .and finer, and in such small cjuantity as t(» he
detected only on close inspection. The hair is longest on the
anterior upper portion of the body, where on tlu; neck and
shoulders it attains a length of 40 mm.; it decreases in length pos
teriorly, and toward the tail has a length of only 15 mm. It is
still shorter on the abdomen, becomes still more reduced on the
limbs, and disappears entirely toward the ends of the digits.
The end of the nose, the soles and palms, the anal region, and
the extra-digital cartilaginous flaps are naked and blacdi (in
around the clicst, and that tlu* average weight is over one thousand ])oiimls.
He gives the length of the full-grown fen Ue as 8^ to 9^ feet, and the circiiui-
ference at the shoulders as 4, the I'emal s being relatively much BJciiden'r
than th( males. The weight of the fem 'o he states to bo one-third tbiit
of the male.
EXTERNAL CnARACTERS.
235
lil'r ••(lull MiU'-MiK'lv "). TIh' whiskers are louy, shMider, ami
ryliiulrical, wliilc <»!• hrownisli-wliite, and set in four or five
nitlior iiKlislliifl rows. Some ol" tlie I(m: est liave sometimes ii
l(ii;;tliof")OOmm.,orab(nit twt'iityiuehes, witlia iiiaxiinimi thiek-
iicss o\''2 mm. Tliey are set in several rows, and number between
thirty and forty, incrcasinji' in lenf;tli from tlio inner ones totlio
outer, wliieli are longest. The ears are short an, whieh is slightly and somewh-»t irregularly in-
dented on the inner side. The terminations of the digits are
indicated by snudl (drcular horny disks or rudimentary nails.
'Die liind feet are broad, ami gradmilly widen from the tarsus,
reaching their greatest breadth at the eml of the toes. Their
length is short as compared to their breadth, tlui distance be-
tween the ends of the outer toes when spread exceeding half
of the length of the foot, nuiasured from the tarsal joint. The
toes terminate in strong cartilaginous tlaps, covered Avitli a,
thick leathei\v naked membrane, which is deeply indented oppo-
site the intervals between the toes, and serves to connect the
(li\ crging digits. The three middle toes are provided with long,
welldeveloped nails; the outer toes a^e without true nails, but
ill place of them are thickened, horny disks. The outer toes
are slightly longer than the three middle ones, which are sub-
e(|ual. The nails on all the feet are bluish horn-color.
The following table of external measurements of two males,
one very aged and the other adult, both from St. Paul's Island,
\la8ka, indicates the general proportions of the body. A part
were taken from a moist flat skin before stuflfing, and the others
from mounted skins.
1 ■■•K .
••• H
Mi
-» S o ^ * •
9 r» t ^5
23(J i;UMKT0PIA8 8TELLKHI — STELLEU's 8EA LION.
Mcanurcnicnls from iwo Skhm of Kdmktoi'Iah stki-M'.im.
'No. SO-'l, C..11.
|Ko. 2020,('(>1I. MiiH. Cdiiip. Zim''1.,| Mus. Ciuiiip.
I (/.nljoiit 10 ji'iirsolil, I / from end of noso to ear
Distuneo between the eyes
Distance between the ears
Lengtli of the ear j
Lengtli of longest harbulo
Dist. between points of longest barbules
Circumference of the body at fore limbs.
Cireumfereuce of tho body uearthotall.
Circumference of tho head at the ears. .
Length of body to end of hind limbs . .
13 years old.
Unmounted.
.Miiuuted.
Motiuted.
2, 750
2, 700
3,010
100
100
110
2, 302
575
500
flfiO
337
335
•JOO
559
S40
010
210
210
230
483
445
440
200
200
220
17&
ISO
210
152
147
190
104
150
100
104
150
165 1
215
100
170
308
303
380
100
105
210
372
370
420 1
37
35
35 1
342
342
1
800
800
2,250
2,600
1,000
1,020 '
....
1,000
980
3, 450
8,700 !
il#<
i ,1
CaptJiiii Scanimon gives the following external measurenieiits
of a full-grown male taken at the Farallone Islands, July 17,
1872.
ft. in. mm.
Length from tip of iiose to end of hind flippers 12 :^3, 060
Length of hind flii)pers 2 2= OGO
Breadth of hind flippers (expanded) 9 = 220
Circumference of hody behind fore limbs 7 =2, 150
From nose to fore limb.s 5 =1,526
Length of fore flipper 2 tJ = 753
Breadth of foro flipper 1 4= 470
Distance between extremities of fore limbs 10 0=3,035
Length of ear li= 4*
\ Length of tail 7= 17"
■ Length (.flon'f. eat vhiskers 1 6= 457
. ' Lengtli
•.««(' ao^ui JO njSaai
O O Cl to O h-
-H 00 oc » o t--
PS M IM CI M M
1(5 O O O
gj
•B9i^pnoo xBitdjooo JO q^puaig; I ^feSSSSSS
•^n«»qTO^»»^! linns JO nipno-iq^suoi 3K3Si3"SK
P
^
,2
■saniOBt) %v iin^iB jo q^po^-iS
-*M0530CeiBS6njo i]}pB9j(i; ; ^ ^
i» (M l.'S O O
M CO CO ^ TO
•oSpa i>UO.IJ IB BIBSBO JO njpBa.ifi
CO OO t* l-O w ■.*<
L'S '^t -^ ^ Irt -}'
•89Uoq IBSBH JO qijSnai S § S g S S
CO CO C5
00 >* l» o
^ ^ ^ CO
W -^ © ^H »-(
in to » o ^
•asnrxflni JO pua jopa:)
■Bod ^B oanjjns oa;;BiBd jo q^peojg
CO ta O M T* ID IfS 00
».0 ^ O ^ ^ Ift -^ CO
•A'aEJ
"liXBUi JO lapjoq JoioaAiB jo qjSnoi
•sseiDoid jBxnmeq jo pno oj nani | 3 g >«, w
-TIB >faBnraBni-o!)BiBd u MJ oonBjsjQ I '^ '^ r^ ^
ti
•BBSoojd pionai8}sod 0; bsubhixbui
•aajnt jo japioq jnojj tuojj ooub^biq
SoQto»ooo»;5 0ot»
O^OOCaOOCt^SoOrH
CI CO M ?1 f> CO M « (N e-l M
• jBioni }8Bi JO japjoq japmq o; BajJBni
-xBtniajni jo aSpa :juo.ij rao jj aouBjsjd
pto3^JO!jdjos8aDOadaBinHiBqo}sauBiii
sBUU»nn JO o3pa ^nojj nio.ij a.iUBjBid
i*: O »?5 o ts 10 h- Q O Tl ^H
CO o tD 00 ^ uo e» S o i--^ ci
BaiiaiB oi!>Buio9.Jz ;« q;pBa.iq ;bo jBa-iQ
ociooinot-otooco
O'^^CO'.fCOCOCi'tCIt^
CI CJ CI CI CI CI CI CI fj r^
5?
■Baasaaoid pio^sBin ;b q^psa.ia;
O O IS O O CO I"} 10 O O CO
COCOCICICO^CIOCOOl-O
C5 CI CI CI CI CI CI CI CI CI 1-1
s
•q^na'!
CO IB
CO CO
>* O O 00 O 15 M< 00
cO"*cocoS5cococJ
■sag
^^•b"D'D•D^^•o'D"D*DO^■ o
:^
4
a
O
I
I
IS
I
o o
I
I
o o
a
c«
o
.1
i
'laqmnu enSoiB^BO
CI r-l
iH t- r- o> lio
O iH to >b t-
t- M t- CO fe
^ M *H Irt fH
?
:i'i
i^^^i^l:
TEETH.
239
Tkk'J'II. — TLc lust upper Jiioi.ir is double rooted, sind its erown
directed backward. All the other luolais are single-rooted, with
a sliylit median longitudinal groove on the outside. Their
<;io\viis are irregidarly conical, pointed, aiul jut out over their
contracted necks; inner side of the crowns hollowed. Surface
id the crowns roughened with minute longitudinal grooves and
ridges. The upper molars have no trace of the supplemental
points to the crowns seen in many species of this family. The
lower molars, particularly the third and fourth, ha\e very
slight accessory cusps. Necks of the molars uniform in size
with tho upper part of the fangs. Fangs of the molars gradu-
ally tapering, those of the first and second upper much curved
inward; that of the third less so; that of the fourth straight;
the two fangs of the fifth are directed abruptly forward, the
posterior one much the smaller. Canines of both jaws very
large; the upper, however, much the larger; the lower more
curved. Of the six incisors of the upper jaw, those of the
outer pair are mucli larger than the middle ones, two-thirds
as long as the canines, and much like them in form. The mid-
dle ones have their antero-posterior diameter nearly twice their
lateral diameter, and their crowns are divided transversely.
The fangs of the inner pair are slightly bifid. Of the four
lower incisors, the outer are much the longer. *
MeaaurcmcHtu of the TeethA
A. — Teeth of the UrPEn Jaw.
Holara.
Total length
Length of the crown
" neckt
" ioot§.
1 Aiiteio-postcrior diameter || j 11. 5
j Lateriil fliameter || ^ 6. 5
•
.a
Incisors.
t
i
1,
1
1
1
37
13
6
18
40
11
6
23
11.5
8.5
84
34
6
03
23
7
29
5
7
25
4
7
13
10
24
20
15
12
7
5
«
<
* For figures of the teeth, see Bull. Mas. Comp. Zool., vol. ii, j)!. i, ligg.
"i-.V (one-half natural size).
tThps"' measurements are takt^n from a middle-aged specimen, in which
the dentition is perfect and normal. Tn old age many of the teeth are usually
1iink(>n, and a portion of them often entirely wanting, through loss from
accident. As the lower canines could not be removed without removing a
portion of the jaw, they have not been fully measured,
{The distaice from the crown to the alveolus.
^ The portion of the tooth inserted in the jaw.
II At tlie base of the crown.
fi
240 EUMETOPIAS STELLERI — STELLER's SEA LION.
B. — Tketii of the Lowi;k Jaw.
Molurs.
Iii('is
10..". 20
K..') IV
31
H
4
HI
Skeleton. — Vertebral forimila : Cervical vertebra', 7: dorsal.
ir»; lumbar, 5; caudal (iucludiufj the four sacral), variable;
probable average, IG.
Ten of the fifteen ribs articulate with the .st<'rniini; their
sternal portions are entirely cartilaginous. Their os.seous ])(»r-
tions evidently increase niucli in length alter middle age. Tlio
apophyses of the vertebne are well devt;loi>ed. Of the neiinil
spines of the dorsal vertebne, the first, .second, and third are
sub-equal, 130 mm. long; they gradually shoi-ten jwsteriorly, tlie
last having a length of oidy 75 mm.
The sternum is normally composed of nine thick ami broad
osseous segments, the first and last very long, the eighth short-
est. Between the eighth and ninth a shorter cartilaginous
one is sometimes intercalated (as in specimen No. 2020).
The pelvis is well developed. The ilia are very long and nar-
row antero-posteriorly. The i)ubic bones are unanchylosed,
they being merely approximate at their posterior exti'cmitiois.
Probably in the femjiles (as in Callorhimm ursinus), they are
widely separated, and the Avhole pelvis is much smaller than in
the males and differently shaped.
The humerus, as in the other Pinnipeds, is short and thiek.
with the greater tuberosity enormously developed. The bones of
the forearm are also very large and strong, with all their i)io
cesses greatly developed ; in length they but slightly exceed the
humerus. The length of neither of the segments of the arm
quite equals the length of the bones of the first digit (inch'dinjr
its metacarpal Ixme) of the hand. The first digit of the hand is
the longest, twice as long as the fifth, and very thick and strong.
*Tho distance from the crown to the alveolus.
tTho portion inserted in tho jaw.
t At the base oi' tho crown.
SKELETON.
241
The bones of the hinder limbs are also sliort and thick, espe-
cially llie leinur, whieli is scarcely more than one-third as long
as The tibia. The latter in length about e(iuals the foot. The
relative lenj^th of the digits is as follows, tln^ lon{j;est being
niciitioiied lirst: 5th, 1st, 2d, 3d, and 4t[». The third and
lourtli are of equal length, and but little shorter than the sec-
ond. In respect to size, the metatarsal and phalangeal bones
(if the lifth digit are nearly twice as large as those of the lirst,
while those of the lirst are about twice the size of those of
cither of the other three. As previously noticed, the three
iiiitldle digits of the foot are supplied with long narrow nails;
the tirst and fifth with rudimentary ones.
McamremcnUi of the Bones of the Hand (Metacarpal and I'ludamjial).
I
• length of metncarpal and plip-
langcs -
lA'Dgtli of iiKitacaipal bone
Length of Ist phalanx
Length of 2(1 phalanx
L -ngtli of 3(1 phaloux
Middle-aged Hpcclmen.
Very old specimen.
. I .
I
•a
4. 1 •■?. :a
352
152
140
60
310
110
05
80
25
a
S
I -^
240
85
70
60
25
200
SO
55
45
20
•a
.4
•a
« . M
I
177
80
65
20
12
357 320
160 I 110
140 05
57 { 80
, 35
*5
•a
'3
250
00
70
65
25
■a
-3
205
80
60
45
20
185
85
65
18
17
Measuioncnta of the Bones of tlie Foot (Metatarsal and Phalangeal).
Length of metatarsal and phalan-
ges
Length of metatarsal bono . .
Hugth of Ist phalanx
Length of 2d phalanx
Length of 3d phalanx
Length of nail
Middle-aged specimen.
310
120
140
50
«i
•«i
•a
•a
•a
-d
s
S
290
05
00
75
30
40
290
95
90
75
30
40
•a
305
110
90
80
25
37
:a
328
130
93
70
35
Very old specimen.
•a
■a
320
145
130
45
•a
■■5
317
110
100
80
27
50
•a
327
110
105
85
27
55
•a
■■3
350
120
105
05
30
50
-a
■s
350
130
110
75
35
Tli(3 liyoid bono is greatly developed. Each ramus consists
i)f five segments, its two rami being connected together by a
transverse segment articulating wi*^^h the juncture of the fourth
iiud llfth segments. All the parts Oi" the hyoid bone are very
tliick, esixicially the transverse and anterior segments; rela-
tively much more so than in GallorhintyS, In the common Phoca^
mac. Pub. No, 12 IG
242 EUMETOPIAS 8TELLERI — STELLER's SEA LION.
the byoid bone is reduced almost to a bony filament. The length
of the hyoid bone in the present species is 270 mm. ; of the trans-
verse segment, 05 mm. ; circumference of the transverse segment,
45 mm.; of this segment at the thickest i^art, 95 mm.
The OS penis is 170 mm. long, slightly arched, somewhat Hat-
tened above, especially posteriorly, sharply convex below, and
abruptly expanded and squarely truncate at the end. Its cir
cumference at the base is 72 mm. ; just behind the terminal ex
pansion, 32 mm. ; of the terminal expansion itself, 65""".
Measurements of the Skeleton.
Whole length of skeleton (including skuU) . .
Length of sknll
Length of cervical vertebras
Length of dorsal vertebras
Length of lumbar verl /bras
Length of caudal vertobras
Length of first rib
Length of first rib, osseous portion
Length of first rib, cartilaginous portion
Length of second rib
Length of second rib, osseous portion
Length of second rib, cartilaginous portion . .
Length of third rib
Length of third rib, osseous portion
Length of third rib, cartilaginous portion
Length of fourth rib
Length of fourth rib, osseous portion
Length of fourth rib. cartilaginous portion...
Length of fifth rib
Length of fifth rib, osseous portion
Length of fifth rib, cartilnginous portion
Length of sixth rib
Length of sixth rib, osseous portion
Length of sixth rib, cartilaginous portion
Length of seventh rib
Length of seventh rib, osseoua portion
Length of seventh rib, cartilaginous portion..
Length of eighth rib
Length of eighth rib, osseous portion
Length of eighth rib, cartilaginous portion. . .
Length of ninth rib
Length of ninth rib, osseous portion
Length of ninth rib, cartilaginous portion
Length of tenth rib ,..
Length of tenth rib, osseous portion
Length of tenth rib, cartilaginous portion
Length of eleventh rib, osseous portion only.
No. 2920,
rf.lOy'rs
old.
2,750
374
600
1,050
340
440
260
130
130
345
175
170
410
230
180
470
280
190
535
320
215
580
360
220
640
400
240
670
420
250
710
420
290
750
420
330
430
No. 2921,
rf.lSy'n!
old.
2,935
3S5
540
1,090
400
520
240
140
100
305
185
120
410
270
140
470
330
140
5J0
370
160
300
4M
170
eo
440
160
670
4S0
190
6M
4«
200
745
4»i
IHi:;
SKELETON.
243
Meaaurvnients of the Skeleton — Coiitinui-d.
No, 2920, I No. 2921,
,^, 10 y'rs 1(^,15 y'rs
old. old.
i Lengtli of twelfth rib, osseous portion only
Length (if thirteenth lib, osseous portion only
I.c'ii);th of fourteenth rili, osseous portion only ,
Len^'l h of flft<'cnth ril), osseous portion only
Lt'ii>;tli of sternum (ossified portion)
L(uj,'th of sternum, Ist scfpiient
Li'iij;th of sternum, 2d segment
Length of st^-rnum, M segment ,
Length of stenimu, 4th segment
Length of sternum, 5tli secment
Length of sternum, 0th segment
Lengtli of gtemum, 7tli segment
Length of sternum, 8tli segment
Lengtli of st^-'inum, 9th segment
Length of supemumery eartiLige (between 8th and 9th)
Lengtli of scupulii
Breadth of scapula ,
Greatest height of its spino ,
Length of humerus ,
( 'ircunifevence of its head
Least eireuraference of the humerus
Length of radius
Length of ulna
Longest diameter of upper end of ulna
Length of earpus
Length of metaearjius and 1st digit
Lengtli of iiiftaeaipus aud 2d digit
Length of metacarpus and 3d digit
Length of metacarpus .and 4th digit
Length of metacarpus and 5th digit
Length of femur
Cheumference of neck •
Length uf tibia •.
Length of fibula
Length of tarsus
Length of metatarsus and Ist digit
Length of metatarsus and 2d digit
Length of metatarsus and 3d digit
Length of metatarsoe and 4tli digit
Length of metatarsus and 5th digit
Length of innominate bono
Greatest width of the pelvis anteriorly
Length of ilium
Length of ischio-pubic bones
Length of thyroid foramen
Lengtli of OS penis
■^iiUhof hand at base of digits
Width of foot at base of digits
490
450
410
340
700
130
70
70
65
63
60
60
55
70
30
830
350 I
45 i
300 j
300 I
170 I
260 I
310 I
100 I
80 I
350 j
310 \
240 i
200
170
170
125
320
310
140
310
290
290
305
227
320
140
140
140
500
470
460
350
840
180
90
85
80
85
75
73
65
77
370
380
52
285
290
180
260
310
130
80
360
320
250
205
185
220
120
340
330
160
270
290
270
285
310
360
160
160
200
200
170
140
p'iM
244 EUMETOPIAS STELLKRI — STKLIKU's SEA LION.
The above, table j^ivcs the priiK-ipal iiieasiiroiiu'iitsofthebonofj
of tbt' skcU'toii. .McasiirciMcnls of two spcciiiiciis are j^iveii, as
in l>revioiis tables, foi'tlie purpose of illiistratiiij;' the variations
thai oeeur in the relative si/e of (litVerent parts after maturity
is attain«'(l, and also for the [>nrpose of illnstratin<>- individual
variation, whieh in some i)artieulars these specimens exhibit
in a marked dej-Tee. The ribs, it will be observed, diifer hut
isliji'litly in total lenf;th in the two; riot nearly so uuieh as
would be expeeted from the nuieh jL>reater bulk of the body of
the older specimen. It will be noticed that the principal diller
ences in the ribs consist in the relative length of the bony to
the cartilaj-inons portions, in the older the ossified portion be
inj;' nnu'h lonjier and tli«' cartilagin material, consisting merely of
three adult males, does not furnish many facts toiichii g these
l)oints. These si)ecimeus, however, dift'er considerably from
each other, not only in color, but in size and i>roportioiis. Some
of these dift'erences are clearly due to age (one of the specimens
being nuich younger than the others), but others equally great
cannot be thus explained. The body increases greatly in bulk,
and the bones in size and density, after the animal has reached
its adult length. The crests of the skull are almost wholly
developed after this period, and in great measure also the spines
or ridges of the scapula. The tuberosities for the attachment of
muscles also increase in size, as do the vertebral or osseous por
tious of the ribs, as shown by the measurements already given.
The teeth also change nuich in size and form after maturity is
attained, and in old age often become much worn and broken
by long use. The general form of the skull in the males differs
considerably in dilBfereut individuals of the same age, and also
undergoes great modification with age.* As already stated, tliis
consists mainly in the development of the crests and processes
for the attachment of muscles, and in the size and form of the
teeth.
Mr. Elliott states that the young, when first born, have ;i
■weight of about twenty to twenty-five pounds, and a l^ugtli of
*Seo Bull. Mu8. Comp. Zool., vol. ii, pp. 56-60.
^TBjf
COMPARISON WITH ALLIED SPECIES.
245
about two i't'oi, and dcst'iiLvf^ their color at tliis ago as being
••(laik clioeolate-brown." When they are a year okl ho says
tlicy liave tlie sain<> (•oh)ras the achilts. On their arrival at the
I'nbilov Islands in s]>rinji', Mi". l''^lliott slates that he was nu-
\[h\v to (liseern any marked dissimilarity of e()!orin;n' between
the males and females, and adds that the "young males and
yearlings" have the same color as tlu' adults, with here and
tluTc an animal marked with irregularly disjjosed patclies of
(lark brown. After their arrival, the general color gradually
becomes .somewhat lighter or more golden, and darker agaiu
after the moult.
As already noted, the .sexual differences in the skull are
strongly marked. They are, however, only parallel Avith those
seen in the other .s])ecics of Otaries. The skeleton of the female
is still unknown to me, but nuiy beiuxvsumed to differ from that
of tlie male very much, as is found to be the case in the Fur Seal,
as described further on.
(lixxHJAPniCAL Vauiation. — The material at hand seems to
indicate that there is no marked variation in size with U)cality.
A considerable series of skulls from the California coast indi-
cates that the species attains fully as large a size there as at the
I'rybilov Islands. One of the largest skulls I have seen came
trom the Farallone Islands, the extreme southern limit repre-
sented by the s})ecimens before ine.
Comparison with Allied Species. — Eumetopias stelleri is
tlie largest of the Eared Seals, very much exceeding in size any
of the other species of the family except Otaria juhata^ which
alone it sufBciently resembles in external features to render com-
parison necessary. While widely distinct from the latter in
cranial characters, it seems to quite closely resemble it in exter-
nal features, so far as may be judged from descriptions. The
character of the pelage, the color, and the conformation of the
linib.s are nuicli the same in both. In neither is there a distinct
"mane," so often attributed to them, and especially to the
Soaiiicrn Sea Lion, although the hair on the neck and shoul-
ders is longer than elseAvhere, the resemblance to the mane of
the Lion being duo to the heavy folds of skin over the shoul-
ders when tlie head is raised, more than to the existence of an
abundance of lengthened hair that can in any true sense be
cousiddied as forming a mane such as is seen in Leo.* The skins
* AtLordiug to Captain Bryant, "At tho fourth year of ago the uceli and
sliouldcrs tlijcken, from having a thick layer of fat nndc-r the skin, tlu' skin
n
246 EUMETOPIAS STELLERI — STELLER's SEA LION.
of tbese two species at my coinniand arc in the one casd those
of very young animals, and in the other of very old males. A
fine series of the skulls of each enables me, however, to s])eiik
with confidence in respect to the matter of comparative size.
The largest old male skull of Eumctopias steJlcri has a length of
400 mm., Avhile none fall below 375 una., the average being about
390 nun. In Otaria jiihata, the largest old male skull in a series
of a dozen barely reaches 372 mm., and several fall below 340 nini.;
the average being about 355 mm., or about 50 mm. shorter than
tlie average of a similar series of Eumctopias steUeri. Adult
female skulls of the last-named species reach 290 to 300 mm,,
while old female skulls of 0*ariajubata about 265 nun. Accord-
ingly it seems fair to conclude that the linear measurements of
Otaria jitbata are about one-eighth less than those of Eumetopm
stelleri, with a corresponding difference in the bulk and weight of
the entire animal in the two species. As very few measure
ments of the skulls of Otaria jubata have been as yet published.
I append the following for comparison with those of Eitmeto-
plan stcUcri already given {antea^ p. 238). The wide differences in
dentition and cranial structure liave already been suflBciently
indicated.
itself beiug loose and flabbj'. When the animal is at rest ou a rock with its
hind flippers folded under its body, its head erect and the shoulders tliiown
back, the loose skin and fat lies in folds, looking like the mane of a Lion;
hence its name Sea Lion. This thickening of the neck is peculiar to the
adult male." — MSS, notes.
mmmmmmmmmasm^sam
COMPARISON WITH ALLIED SPECIES.
247
<
9
1^
o
"5"
I
g) to 6(1
oooS?!S?^3SoSS
•jeojD iBj^iSna jo ^mSjou
•saiimiij %v iniMAV
Or-lf-HF-(0*-tC5CTOlrtlrtlO
■Xisoouajiij JO aopjoti !)nojj
•snitiraoq pioSA'aojd o) sauv\
C-l t» ® O -* »5 Q »0 t- O 00 O
» C^ C^ C-l ^ ft CJ n IH IH CI iH
CI C) CI CI CI CI -M CI i-t »H iH iH
•^iaou.t4nii inp'vv 's.moq pim :
^[jo]aa)8ud q4Pt:ti 'Bduoq \vb\s^
•S'jiqio uooM'faq 93uw,8i(i
L-j f5 ifl ClrH€0H
ClrHMCIrHCJrti-li-Ji-lt-lH
&ICOClrtrHCIrH»-i^'««J»^^
CI CI d CI d CI C4 C4 »-4 i-t ri iH
^inincit*co^coiot*io
coweocococ^wcocidCiw
•X9g
"o^^^3^3"D^D"D"nOl■ooo»■
{
"5 -(t~t-{0r-lr^
II r
JM ,
24 S EUMETOPIAS STELLKRl— STELLEU's SEA LION.
CiEOGKAl'IIKAL J^I^TKim TIOX. — TllC klloMli riin;;(' of tills
sporh's cxtciids iiloii;;' tlu' west coiist of XorUi AiiH-rica fioiii
the FiniilloiK' Lslaiids, in liititiidc ;57o 10' N., to tlir Prvbilov
Lsliiiids. Its iiortliciii limit ol' distribution is not (h'linitcly
known, bnt it docs not iipia-iir to Imvc been nu't with north nl'
about the latitude of St. Matthew's Island (about latitude OP).
Keither m-. ^^ . II. ])all nor 3Ir. II. \V. Elliott nt one is led
to infer that he had not then seen six-cimens of it, and tliat he
rested his belief in the existence of sueli a species mainly on
^teller's account of it, as he himself expressly states in his
later papers. Tlie skull received sul>se(pu?ntly at the British
^[nsenm from ^lonterey, California, and ti};nred and described by
Gray, in 1859, as a new sjjecies, under the name Arctocephalus
moiitvrlcHffiK, proved, however, to be of this species, as lirst
atliriiied by Dr. Gill, and latei- by Professor Peters and by Gray
himself. AVith the exce])tioii of the (ij;ures of an imi>erfect
skull of Steller's Sea Lion from Kamtchatka, fiiven by Pander
ami D'Alton in 182G, Dr. Gray's excellent ti<4ure* (a view in
laolile) is the only one of its skull published i)rior to 187(5. The
(iiilv specimens of the animal extant, up to about ten years
since, in tU<' Euro])ean museums, seem to have consisted of the
two skulls and a stull'ed skin in the JJerlin Museum mentioned
h,v Peters, and the skull in the British Museum tiyured and
4k'scribed by Gray.
With the Monterey skull abov(^njentioned, Dr. Gray received
another very youny skull, and the skin of a Fur Seal, both of
which were said to have belonged to one animal, and which he
liesitatinyly leferred to his Aretocephalus inonteriemuA Later,
however, he regarded them as representing a new species,!
which he called Arctocephalus caiifornimitis. Still later lie refer
red his A. caUfornianm to Eametopias isteUeri § {=Arctocephalii.s
moHteriemifi, Gray, of earlier date), and in 1872 1| published tigures
of this young California skull. Concerning the skin above re-
ferred to he remarked at one time as follows: "If the skin sent
last year by Mr. Taylor to Mr. Guruey, and by that gentleman
presented to the Museum, is the young of this species [A. mon-
' I'roc. ZoiJl. Soc. Load., 1859, pi. Ixii.
'i'roc. Zool. Soc. Loud., Id59, i». ;358.
;C;it. Seals ami Whales, 18iJ6, p. 49.
v^Aun, and Mag. Nat. Hist., M sories, 186C, voL xviii, p. '2'S'S; Hand-
list of Seals, etc., 1874, 40.
ilPioc. Zool. Soc. Loud., 1872, pp. 740,741. . ; .; '
250 EUMETOPIAS STELLERI — STELLER's SEA LION.
teriensifi], tlie yoimg: aiiiinal is blackish, silvcrtMl by the slioit
white tips to the short black hairs; those on tho nape and
hinder parts of the body with lonj,a'r Whitii tips, iniikin}^' thoM'
parts whiter and more silvery. The under-fur is very abuinl
ant, reachiny nearly to the eiul ol" the hair. The eiid ot the
nose and sides of the face are whitish. The whiskers are i-loii
;,'uted, rigid, smooth, and white. The hind feet are cloiijAiitc.
with rather long- tlaps to tlM' toes. The skull is small for the
size of the skin, and I should have doubted its belotigiiiy to
the skin if it were not accompjMned by the following labtl:
'Sknll of the ]<\tr Seal I sent last year. It is very imperfect,
from my forgetting where I had put it; but it must «lo until airi
dent throws anotlu'r in the way; the other bones were lost.—
A. S. T.'"* Dr. Gray, in his "Hand-List." published in 1S74.
refers the skidls of both A. mov.tcriennis and A. caJifornionn*
to Eumetopim (ttellvri, but makes no reference to the skin. As
he seems, however, to have become settled in his opinion tbat
this skin is identical with his A. monteriensis, this nisiy nc
count for the statement made by him in 180(5, t and sultse
quently reiterated, | that the Eumetopiax stelleri is a species in
which '' the fur is very dense, staiuling nearly erect from the
skin, forming a very soft, elastic coat, as in O. falldandha and
0. steUeri, which,'' he erroneously says, "are the only Seals
that have a close, soft, elastic fur." §
Lesso; ; gave the name Otaria californiana toasupposed species
of Eared Seal based solely on a figure entitled "Jeune lion nun in
de la Califbrnie," published by Choris.|| The following is tlie
only allusion Choris makes to this animal, in this eoiuieetioii,
in his text: "Les rochers dans le voisinage de la bale Sau
Francisco sont ordinalrement converts de lions marins. PI.
XI." In his chapter on the "lies A16outiennes," in describing
the "Lions marins," he says: "Cesanimaux sontaussi tr^s-com
muns au port de San-Francisco, sur la c6te de Californie, oil
*Proc. Zool. Soc. Lond., 1850, p. :?58.
t Ann. and Mag. Nat. Hist., 4tli series, 1866, vol. i, p. 101.
tlbid., p. 'ilij.
$ Dr. Gray's mistake seems to have misled others in respect to the real
characters of Eumetopiaa stclleri, which Dr. Veatch, on the authority of Gray,
refers to as tlie "fur-coated Eumatopias," which he supposed to be the
proper name of the Fur Seal of the North. (See "Report of Dr. John A.
Veatch on Cerros or Cedros Island," in J. Ross Browne's "Resources of the
Pacific Slope," [appendix], p. 150, 1869.)
II Voyage Pittoresque, pi. xi, of the chapter entitled "Port San-Francisco
et ses habitants." Tho date of this work is 1822.
iBiai£S£t III tc'tc ])lu.s line: (piaut a la couleur, •■Ho passe lortoiiiont an
lirmi, tandis que ceux des iles Aleouiicunes sont d'une coulcur
plus yiise, out le corps plus lond, les mouvemcnts plus diOiciles,
la tote plus grossc ot plus (5paisse; la coulcur du poll des
iiioustaohcs plus uoiratro que celui des iles Aleoutieunes." *
It thus ai)pears that Choris clearly recognized the larger aud
the smaller Sea Lious of the west coast of North America, aud
coirootly pointed out their more obvious points of external
difleronce. Hence Lesson's name Otaria californiana, founded
on Choris's "Lion marin do la Califoraie," must be considered as
api>lying exclusively to what has till now been commonly known
as ZolopliHK (fiUespii.
Dr. Gill, however, in his "Prodrome," adopted provisionally
Lesson's name [californiana) for the present species, but at the
same tinu' asserted its identity with the ArctocepliahiH - . ,ac-
rlnisin of Gray (1859), and also suggested its probab' identity
with the so-called Otaria stelleri of MUller. Peters, a few months
later, came to the conclusion that Gill's suggestion was correct,
since which time the name stelleri has been universally accepted
lor the laiger northern Hair Seal. The Otaria stelleri of Tern-
uiinck,t formerly supposed by Grayf and also by Peters §
to include both the Australian Eared Seals (viz, Arctoceplialm
cincreits and Zalophvs lobattis), has finally been referred by
the latter, after an examination of the original specimens in
the Leyden Museum, to the so-called Zalophus gillespii. \\ I be-
lieve, however, that the skull of the J'oung female figured
in Fauna Japonica (pi. xxii, flgg. 5 and 6) belongs to some
other species. It certainly differs greatly in proportions, as
well as in dentition, from the other skulls figured in that work
(same plate), and called 0. stelleri.
The northern Sea Lion having become generally recognized
as specifically distinct from the Sea Lion of the southern seas,
Dr. Gill, in 1866, separated the two generically. This had
indeed already been done practically by Dr. Gray, inasmuch as
* Voy. Pittor. aut. tlu Moudo, Iles Al^outieuues, p. 13.
tFauuii Japouica, Mam. inarius, p. 10.
{Ann. and Mag. Nat. Hist., 3d series, 1866, vol. xviii, p. 229.
Hlouatsberichte Akad. Berlin, 1866, pp. 272,276.
II Ibid., p. 669. See further on this point posted, under Zalophus cal\for-
nianuH.
tlv
252 KUMETOPIAS STEIXERI— STKI-LKU'S 6KA LION
111' ])lacod liis A. montericmiH ( = 0. .sfc//er/, auct.) in the geims
Arc.occpJiahts, and llie .southern Sea Lion in Otaria, with which
he associated the 0. stcUcri. He faiUnl, however, to rccoguizo
the identity of hi« A.montericnxifi with his O.Hfelleri, and hence
the entire jieneric diversity of the northern and southern Sea
Lions seems lo have escape! his observation. Tlie hitter fact
was lirst pointed out by Dr. Gill in his "Prodrome,'' as above
stated.
Dr. Gray has recently described and figured the skull of what
he at first regarded as a second species of Eiwietopian from
Japan, and which he called Eiimetopian elonyafus,* but he sub-
sequently transferred it to his " genus*' Phocarctos.] In his first
mention of it, however, he referred it to Euinetopias stelleri.l
The ^^Phocarctos clongatus'^ was first described from a "nearly
adult " skull (pl. xxi, Hand-List), eleven inches long and seven
and a half broad at the condyles, and placed "in the genus
Eumetop'uiH, because it had a space in the place of the fifth
upper grinder." Judging from the figures § and Dr. Gray's de-
scription, it seems to differ in no important point from the skull
of an adult female, E. sfcllcri. Later !•<: received from Japan
a younger sku?l (pi. xxii, Hand-List), " seven and a Iialf
inches long and lour and a half inches broad," which agrees in
general form with the other, but has a " shortei pa'atej" six
uXiper uiolars (instead of five), and differs "in the form of the
internal nostrils." He considered the two as both belonging
to the same species, and, from the presence of six upper molars
in the young skull, transferred the species to " Phocarctos."
Judging from Dr. Gray's figure of this skull (Hand-List of
Seals, pl. xxii), it seems to be referable to Zalophits (the Japan
species, probably Z. lobatus), the last pair of upper molars
be ng in all probability supernumerary, as they are smalk:
thaJi the others and differ from those jireceding them just as do
the jL'ipcrunmerary molars in skulls of Zaiophiis californixmis.
Dr. Gray seems to have believed that Eumetopias hac in early hfe
six upi)er molars on each side, and that the fifth, or last but one, is
deciduous, thus leaving a vacuity between the last '^"io molars
on either side. Of this T have seen no evidence ; on the con-
trary, I have found in a very young skull the same number of
molars as in the adult. Thus skull Xo. 4703 ( National Museuir'
"i'roc. Zoiil. Soc. Loud., 187;i,7T(i, (igg. 1, '2,
tilaud-liist S'•
s; ;
256 EUMET0PIA3 STELLERI — STELLER's SEA LION
to brinf;^ tlioir skins to the dryiiij^-frames. It soiiiotiiiM's
roqiiircs live days to make the journe}', as at n'oiiiiont intervals
they have to be allowed to rest. It is a somewhat daujicroiis
animal, and the men frequently j;et seriously hurt by it in driv-
ing and killing' it. They are driven together in the same man-
ner as the Fur Seals are; and v.hile eoniining eaeh other hy
treading upon each otliers' Hippers the small ones are killcil
with lances, but the larger ones have to be shot.
" This animal is the most com[)letely consumed of any on the
island. Their Hesh is preferred to that of the Seal for dryiiijr
for winter use. After the skins are taken off (two thousand of
which are required annually to snpply tin; trading-posts of the
Territory), they are spread in ])iles of twenty-flve each, with
the flesh side down, and left to heat until the haii- is loosened;
it is then scraped off, and the skins are stretched on frames to
dry. The blubber is removed from the carcass for fuel or oil,
and the flesh is cut in strips and dried for winter use. The lin-
ings of their throaty are saved and tanned for making the kj^s
of boots and shoes, and the skin of the flippers is used for tho
soles. Their stomachs are turned, cleaned, and dried, and are
used to put the oil in when boiled out. The intestines are
dressed and sewed together into water-i)roof frocks, which are
worn Avhile hunting and Ashing in the boats. The sinews of tlie
back are dried and stripped to make the thread with whi<^li to
sew together the intestines, and to fasten the skins to the canoe-
frames. The natives receive thirty-five cents apiece for the
skins when ready for shipment. But these skins are not so nnich
valued by the trader for the i)rofit he makes on their sale, as
for the advantage it gives him in bargaining with the luuiters,
since by buying these they are able to secure a right to the
purchase of the hunter's furs on his return, the natives always
considering such contracts binding."*
The following careful description of their movements on land
was also communicated to me by Mr. Theodore Lyman in ISTD.
who had recmtly observed the Sea Lions on the '' Seal Kocks"
near San Francisco. His remarks may, howcer, relate in part
to the smaller species.
" These rocks," he says, "are beset with hundreds of these
animals, — some still, some moving, some on the land, and sonic
in the water. As they ai»proach to ell'cct a landing, the head
only appears decidedly above water. This is their familiar
*Bull. MuH. Coinp. Zool., ii, pp. 64,65.
um
HABITS
257
ileincnt, and they swim with great speed and ease, qnitc im-
iiiiuU'iil of the heavy surf and of the breakers on the ledges.
[ii liiiuling, they arc apt to take advantage of a heavy wave,
ivhich helps them to get the forward flipj^ers on terra firma.
\stlie wave retreats, they begin to struggle up the steep rocks,
:wisting the body from side to side, with a clumsy worm-like
notion, and thus alternately work their flippers into positions
svbero they imiy force the body a little onward. xVt such times
they have a general appearance of sprawling over the ground.
It is quite astonishing to see how they will go up surfaces having
3ven a greater inclination than 45°, and where a man would have
to creep with much exertion. When the surface is nearly
horizontal, they go faster, and often proceed by gathering their
liin(I-(iuarters under them, raising themselves on the edges
[)f their fore-limbs and then giving a push, whereby they make
a sort of tumble forwards. In their onward path they are
accompanied by the loud barking of all the Seals they pass }
and tliese cries may be heard a great distance. Having
arrived at a good basking-place, they stretch themselves out in
various attitudes, — often on the side, sometimes nearly on the
back, but commonly on the belly, with the flipi)ers somewhat
extended. They seem much oppressed with their oicn weight
(which is usually supported by the water), and it seemed an
exertion for them even to raise the head, though it is often kept
up for a long time. They play among themselves continually
by rolling on each other and feigning to bite. Often, too, they
\rill amuse themselves by pushing off those that are ti-ying to
land. All this is done in a very cumbrous manner, ami is
accompanied by incessant barking. As they issue from the
water, their fur is dark and shining; but, as it dries, it becomes
of a yellowish brown. Then they appear to feel either too dry
or too hot, for they move to the nearest point from which they
may tumble into the sea. I saw many roll off a ledge at least
twenty feet high, and fall, like so many huge brown sacks, into
the water, dashing up showers of spray."*
From the accounts given by various observers, the Sea Lions
evidently move with much less facility on land than do the P r
Seals, doubtless mainly from their much greater size. The young
and the females of several of the different species of these
auiniiils are described as walking with much greater ease and
rapidity than the half-grown and the more unwieldy old males.
*Bull. Mns. Conjp.
xMisc. Pub. No. 12 17
Zool., ii, pp. ()(), 67.
l t .,
li-'i
ffi
258 EUMETOPIAS STELLERI STELLER's SEA LION.
Captaiii Bryant states that the Fur Seal may be driven at the
rate of a mile and a half per hour, while, according to the Kuiue
authority, the Sea Lions can be driven with .safety but about
two miles a day.
Captain Scammou, in 1874, published a very interesting
account of the Sea Lions of the Aleutian Lslands, particularly
as respects the methods employed in their capture, portions of
which will be quoted later. His ac the latter. At about the same time
appeared Mr. H. W. Elliott's more detailed history of the north-
ern species, which is so full and explicit that I transcribe it
almost entire.
The Sea Lion, he says, "has a really leonine ai>pearance and
bearing, greatly enhanced by the rich, golden-rufous of its coat,
ferocity of expression, and bull-dog-like muzzle and cast of eye,
not round and full, but showing the white, or sclerotic coat,
with a light, bright-brown iris.
" Although provided with flippers to all external view as the
fur-seal, he cannot, however, make use of them in the same free
manner. While the fur-seal can be driven five or six miles in
twenty-four hours, the sea-lion can barely go two, the conditions
of weather and roadway being the same. The sea-lions balance
and saving their long, heavy necks to and fro, with every liitcli
up behind of theii' posteriors, which they seldom raise from the
ground, tlrawing them uj) after the fore feet with a slide over
the grass or sand, rocks, &c., as the case may be, and pausing
frequently to take a sullen and ferocious survey of the tield and
the drivers."
"The sea-lion is polygamous, but does not maintain any snch
regular system and method in preparing for and attention to
its harem like that so finely illustrated on the breeding-grounds
of the fur-seal. It is not numerous, comparatively speaking,
and does not ' haul ' more than a few rods back from the sea.
It cannot be visited and inspected by man, being so .shy and
wary that or the slightest approach a stam})ede into the water
is the certain result. The males come out and locate on the
narrow belts of rookery-ground, preferred and selected by
them; the cows nmke tlieir ai)pearance three or four weeks
after them, (1st to (Jth June,) and are not subjected to that
intense jealous supervision so characteristic of the fur-seal
HAUITS,
259
,- - s«<%r jaw-j^c' '
i- ■ :!
Vm. 'M.—i:umtUq)>.m »Ulhri. Adult uuilc, fer.tiiU's, mul youug.
I
260 EUMETOPIAS STELLERI STELLEU's SEA LION.
liaiTiii. The bulls lijilit
ly anioiifi' tliomsolves, ami tiim
■ I
ott'lVoni the lueediiiji-jiiouiid all the youiijicr and weak iiialos,
"The cow sea-lion is not (^uite half the size ol" the male, and
will measure from 8 to 1) feet in length, with a weij^ht of I'oiu'
and five hundred pounds. She has the same general ciist of
countenance and build of the bull, but as she does not siistiiiu
any lasting period of over a week or ten days, she never comes
out so grossly fat as the male or ' see-catch.'
" The sea-lion rookery will be found to consist of about teu
to fifteen cows to the bull. The cow seems at all times to have
th(^ utmost freedom in moving from place to place, and to .start
with its young, i)icked up sometimes by the nape, into the
water, and ])lay together for spells in the surf-wash, a move-
ment on the part of the mother never made by the fur-seal,
and showing, in this respect, much more attention to its off-
spring.
" They are divided up into classes, which sustain, in a gen-
eral manner, but very imperfectly, nearly the same relation one
to the other as do those of the fur-seal, of which I have already
spoken at length and in detail ; but they cannot be approaclied,
inspected, and managed like the other, by reason of their wild
and timid nature. They visit the islands in numbers compara-
tively small, (I can only estim.ate,) not over twenty or twenty-
five thousand on Saint Paul's and contiguous islets, and not
more than seven or eight thousand at Saint George. On Saint
Paul's Island they occupy a small portion of the breeding-
ground at Northeast Point, in common with the CaUorhiiim,
always close to the water, and taking to it at the slightest dis-
turbance or alarm.
" The sea-lion rookery on Saint George's Island is the best
place upon the Seal Islands for close observation of these ani-
mals, and the following note was made upon the occasion of
one of my visits, (June 15, 1873 :)
" * At the base of clift's, over 400 feet in height, on the east
shore of the island, on a beach 50 or 60 feet in width at low
water, and not over 30 or 40 at flood-tide, lies the only sea-lion
rookery on Saint George's Island — some three cr four thousand
cows and bulls. The entire circuit of this rookery-l»elt was
passed over by us, the big, timorous bulls rushihg off into the
water as quickly as the cows, all leaving their young, ^^any
of the females, perhaps half of them, had only just given birth
to their young. These pups will weigh at least twenty to
^ii
HABITS.
261
t\v<'iit,\ fivo poniids on an avorage wlicn born, arc of a dark,
cli(unliU('-l»i<)\vii, Avitli tlir vyv as large as tlio adult, only being
a siilViiscd, wateiy, gray-blue, where the S(;lerotie eoat is well
iiiid sharply delined in its maturity. Tlioy are about 2 feet
ill length, sohk' longer and some smaller. As all the imi)S seen
today were veiy young, som«^ at this instant only born, they
ucic (lull ami apathetic, not seeming to notice ns unudi. There
arc. I sliould say, about one-sixth of the sea-lions in number on
tills island, when compared with Saint Paul's. As these iini-
luiils lie here under the (jliffs, they cannot be approached and
(liivoii : but should they haul a few hundred rods up to the
sdiith. then they can be easily captnred. *They luive hauled
ill this manner always until disturbed in 18G8, and will undoubt-
edly do so again if not molested.
'"These sea-lions, when they took to the water, swam out to
a distance of lifty yards or so, and huddled all up together in
two or three packs or squads of about five hundred each, hold-
ing their heads and necks up high out of water, all roaring
in concert and incessantly, making such a deafening noise that
we could scarcely hear ourselves in conversation at a distance
I'loin them of over a hundred yards. This roaring of sea-lions,
thus disturbed, can only be compared to the hoarse sound of a
tempest as it howls through the rigging of a ship, or the play-
ing of a living gale upon the bare branches, limbs, and trunks
of a forest grove.' They commenced to return as soon as we
left the ground.
"The voice of the sea-lion is a deep, grand roar, and does not
have the flexibility of the Callorhinns, being confined to a low,
mntteriug growl or this bass roar. The pups are very playful,
but are almost always silent. When they do utter sound, it is
a sharp, short, querulous growling.
" The natives have a very high api)reciation of the sea-lion,
or see-vitchic, as they call it, and base this regard upon the supe-
rior quality of the flesh, fat, and hide, (for making covers for
tbeir skin boats, MdarJcies and Udarrahs,) sinews, intestines, &c.
"As I have before said, the sea-lion seldom hauls back far
from the water, generally very close to the surf-margin, aind in
this position it becomes quite a difficult task for the natives to
approach and get in between it and the sea unobserved, for,
unless this silent approach is made, the beast will at once take
the alarm and bolt into the water.
" By reference to my map of Saint Paul's [not here repro-
262 EUMETOPIAS STELLERI — STELLER's SEA LION.
duced] a small poiiit, near the head of the northeast neck of
the island, will be seen, upon which quite a lar^e number of
sea-lions are always to be found, as it is never disturbed exwpt
on the occasion of tliis annual driving. The natives step down
on to the beach, in the little bight Just above it, and begin to
crawl on all fours tiat on the sand down to tlie end of the iit'ck
and in between the dozing sea-lion herd and the water, always
selecting a semi-bright moonlight night. If the wind is i'avor
able, cand none of the men meet with an accident, the natives
will almost always succeed in reaching the point unobserved.
when, at a given signal, they all jump up on their feet at once,
yell, brandish their arms, and give a sudden start, or alarm, to
the herd above them, for, just as the sea-lions move, upon the
first impidse of surprise, so they keep on. For instance, if the
animals on starting up are sleeping with their heads j^ointed in
the direction of the water, they keep straight on toward it ; but
if they jump up looking over the land, they follow that course
just as desperately, and nothing turns them, at first, either one
way or the other. Those that go for the water are, of course,
lost, but the natives follow the land-leaders and keep nvflnn
them on, and soon have them in their control, driviug them
back into a small pen, which they extemporize by means of
little stakes, with flags, set around a circuit of a few hundred
square feet, and where they keep them until three or four liuii
dred, at least, are captured, before they commence their drive
of ten miles overland down south to the village.
"The natives, latterly, in getting this annual herd of sea-
lions, have postponed it until late in the fall, and when the ani
mals are scant in number and the old bulls poor. This they
were obliged to do, on account of the pressure of their sealing-
business in the spring, and the warmth of the season in August
and September, which makes the driving very tedious, lu this
way I have not been permitted to behold the best-conditioned
drives, i. e., those in which a majority of the herd is made up
of fine, enormously fat, and heavy bulls, some fonr or five liun
dred in number.
" The natives are compelled to go to the northeast point of
the island for these animals, inasmuch as it is the only place
with natural advantages where they can be approached for the
purpose of capturing alive. Here they congregate in greatest
number, although they can be found, two or three thousand of
them, on the southwest point, and as many more on 'See
vitchie Cammin ' and Otter Island.
HABITS.
263
" Cai)turing tlie sea-lion drive is really the only serious busi-
ness these people on the islands have, and when they set out
for llu' task tlie picked men only leave the village. At North-
east Point they have a barrabkie, in which they sleep and eat
while {gathering the drove, the time of getting which depends
upon tlie weather, wind, &c. As the squads are captured, night
utter night, they are driven up close by the barrabkie, where
tlic natives mount constant guard over them, until several hun-
dred animals shall have been secured, and all is ready for the
drive down overland to the village.
"The drove is started and conducted in the same general
manner as that which 1 have detailed in speaking of the fur-
seal, only the sea-lion soon becomes very sullen and unwilling
to move, requiring spells of frequent rest. It cannot x>ick itself
up ii'om the ground and shamble off on a loping gallop for a
few hundred yards, like the Callorhimts, and is not near so free
and agile in its movements on land, or in the water for that mat-
ter, for I have never seen the Eumetopias leap from the water
like a dolphin, or indulge in the thousand and one submarine
acrobatic displays made constantly by the fur-seal.
'•This ground, over which the sea-lions are driven, is mostly
a rolling level, thickly grassed and mossed over, with here and
there a fresh-water pond into which the animals plunge with
gxeat apparent satisfaction, seeming to cool themselves, and
out of which the natives have no trouble in driving them. The
distance between the sea-lion pen at Northeast Point and the
Tillage is about ten miles, as the sea-lions are driven, and occu-
pies over five or six days under the most favorable circum-
stances, such as wet, cold weather ; and when a little warmer,
or as in July or August, a few seasons ago, they were some
three weeks coming down with a drove, and even then left a
hundred or so along on the road.
"After the drove has been brought into the village on the
killing-grounds, the natives shoot down the bulls and then sur-
roinid and huddle up the cows, spearing them just behind the
fore-dippers. The killing of the sea-lions is quite an exciting
spectacle, a strange and unparalleled exhibition of its kind.
• . . . The bodies are at once stripped of their hides and
much of the flesh, sinews, intestines, (with which the native
water-proof coats, &c., are made,) in conjunction with the throat-
linings, {oesophagus,) and the skin of the flippers, which is ex-
ceedingly tough and elastic, and used for soles to their boots or
Harhosars.''
''iiK
IMAGE EVALUATION
TEST TARGET (MT-3)
^
1.0
I.I
l^|28 |2.5
J Kli 12.2
« iM 12.0
III 1.8
IL25 ii.4
km
%
^
v:
/A
4>^
V
:\
;v
\
V
"^'^^^-
-. -rj."-
u
1
i
264 EUMETOPIAS STELLERI — STELLER's SEA LION.
"As the sea-lion is without fur, the skin has little or no om-
mercial value ; the hair is short, and longest over the nape of
the neck, straight, and somewhat coarse, varying in color greatly
as the seasons come and go. For instance, when the Eumetopias
makes his first appearance in the spring, and dries out upon
the land, he has a light-brownish, rufous tint, darker shades
back and under the fore flipiiers and on the abdomen ; by the
expiration of a month or six weeks, 15th June, he will be a bright
golden-rufous or ocher, and this is just before shedding, which
sets in by the middle of August, or a little earlier. After the
new coat has fairly grown, and just before he leaves the island
for the season, in November, it will be alight sepia, or vaudyke-
brown, with deeper shades, almost dark upon the belly ; the cows,
after shedding, do not color up so dark as the bulls, but when
they come back to the land next year they are identically the
same in color, so that the eye in glancing over a sea-Uon
rookery in June and July cannot discern any noted dissimilar-
ity of coloring between the bulls and the cows ; and also the
young males and yearlings appear in the same golden-brown
and ocher, with here and there an animal spotted somewhat
like a leopard, the yellow, rufous ground predominating, with
patches of dark-brown irregularly interspersed. I have never
seen any of the old bulls or cows thus mottled, and think very
likely it is due to some irregularity in the younger animals
during the season of shedding, for I have not noticed it early
in the season, and failed to observe it at the close. Many of
the old bulls have a grizzled or slightly brindled look during
the shedding-period, or, that is, from the 10th August up to the
10th or 20th of November ; the pups, when born, are of a rich,
dark chestnut-brown; this coat they shed in October, and
take one much lighter, but stUl darker than their parents', but
not a great deal.
"Although, as I have already indicated, the sea-lion, in its
habit and disposition, approximates the fur-seal, yet in no
respect does it maintain and enforce the system and regularity
found on the breeding- grounds of the Callorhinus. The time
of arrival at, stay on, and departure from the island is about
uie same ; but if the winter is an open, mild one, the sea-hon
will be seen frequently all through it, and the natives occasion-
ally shoot them around the island long after the fur-seals have
entirely disappeared for the year. It also does not confine its
landing to these Prybilov Islands alone, as the fur-seal uuques-
HABITS.
265
tionably does, with reference to our continent ; for it lias been
aud is often shot ni)on the Aleutian Islands and many rocky
islets of the northwest coast.
" The sea-lion in no respect whatever manifests the intelligence
and sagacity exhibited by the fur-seal, and must be rated far
below, although next, in natural order. I have no hesitation
in putting this Eumctopias of the Prybilov Islands, apart fiom
the sea-lion common at San Francisco and Santa Barbara, as a
distinct animal ; and I call attention to the excellent descrip-
tion of the Calilornia sea-lion, made public in the Ai)ril num-
ber for 1872 of the Overland Monthly, by Capt. C. M. Scammon,
in which the distinguishing characters, externally, of this animal
are well defined, and by Avhich the difference between the
Eumetopias of Beiing Sea and that of the coast of California
can at once be seen ; and also I notice one more point in which
the dissimilarity is marked — the northern sea-lion never barks
or howls like the animal at the Farralones [sic] or Santa Bar-
bara. Young and old, both sexes, from one year and upward,
have only a deep hass growl, and prolonged, steady roar ; while
at San Francisco sea-lions break out incessantly with a 'honk-
ing' bark or howl, and never roar.
" I am not to be understood as saying that all the sea-lions met
^vith on the Californian coast are different from E. stelleri of
Bering Sea. 1 am well satisfied that stragglers from the north
are down on the Farralones, but they are not migrating back
and forth every season ; and I am furthermore certain that not
a single animal of the species most common at San Francisco
^'38 present among those breeding on the Prybilov Islands in
1872-'73.
"According to the natives of Saint George, some fifty or sixty
years ago the Eumctopias held xlmost exclusive possession of
the island, being there in great luimbers, some two or three
hundred thousand; and that, as the fur-seals were barely per-
mitted to land by these animals, and in no great number, the
Russians directed them (the natives) to hunt and woriy the
sea-lions oft' from the island, and the result was that as the sea-
lions left, the fur-seals came, so that to-day they occupy nearly
the same ground covered by the Eumctopias alone sixty years
ago. This statement is, or seems to be, corroborated by Ghoris,
ui his description of the lies S.-George's et S.-Paul's [sic], visited
by him fifty years ago; * but the account given by Bishop Ven-
"* Voyage Pittorusque autoiir du Monde."
"il
T? i 1
266 EUMET0PIA8 8TELLEEI — STELLER'S SEA LION.
Mk
1
i
i
i
f
i
ii
laminov,* differs entirely from the above, for by it
almost as many fur-seals were taken on Saint George, during
the first years of occupation, as on Saint Paul, and never have
been less than one-sixth of the number on the larger island.
.... I am strongly inclined to believe that the island of Saint
George never was resorted to in any great numbers by the fur-
seal, and that the sea-lion was the dominant animal there until
disturbed and driven from ii;s breeding-grounds by the people,
who sought to encourage the coming of its more valuable rela-
tive by so doing, and making room in this way for it.
"The sea-lion has but little value save to the natives, and is
more prized on account of its flesh and skin, by the people liv-
ing upon the islands and similar positions, than it would be
elsewhere. The matter of its preservation and peipetuation
should be left entirely to them, and it will be well looked after.
It is singular that the fat of the sea-lion should be so different
in characters of ta ;te and smell from that of the fur-seal, being
free t''om any taint of disagreeable flavor or odor, while the
blubber of the latter, although so closely related, is most repug-
nant. The flesh of the sea-lion cub is tender, juicy, light-col-
ored, and sUghtly like veal; in my opinion, quite good. As the
animal grows older, the meat is dry, tough, and without flavor."!
Captain Scammon gives a few particulars respecting the
"drive," not especially refeiTed tr by Mr. Elliott. "This
'drive,'" he says, "to the good-natured Aleuts, is what the
buffalo hunt has been to the red-skins on the plains of the
Platte, or matama-time with the old Californians, for the party
starts out as on a sporting foray, and at night they stealthily
get between the herd of Sea Lions and the water ; then, with
professional strategy, they manage to 'cut out' six or eight of
the largest at a time, and drive them a short distance inland,
where they are guarded until a band of two or three hundred
aie assembled. Formerly the implement used in driving was a
pole with a small flag at the end ; but, since our adopted coun-
try-folk have become Americanized, that Yankee production, a
cotton umbrella, has been substituted, and it is said that any
refractory siutchin the 'drive' is instantly subdued by the sud-
den expansion and contraction of an umbrella in the hands of
a pursuing native.
" To collect the desired number for the yearly, supply involves
"•Zapeeskie ob Ostrovah Oonahlashkenskaho Otdayla, St, Petersburg,
1840."
t Report upon Condition of Affairs in Alaska, pp. 152-159.
HABITS.
267
Beveral days ; therefore a throng of villagers, it is said, sets out
prepared with everything needful for the campai - j. As tlie
work of driving goes on only at night, the day is passed in
sleeping and cooking their food by smoldering lires of drift-
wood and seal-fat, sheltered by their umbrellas, or a sort of
tent contrived by spreading blankets and garments over whales'
ribs in lieu of tent-poles — never forgetting in their repast the
fragrant c/w, which is quaffed in numberless cups from the
steaming sam-o-var. At length, the whole troop of animals
being assembled, a flash of umbrellas here and there, with the
call of the herdsmen, brings all into a moving phalanx. But
the tir > for driving must be either at night, aftei the dew is
fallen, or i vn a dark, misty, or rainy day ; as the thick mat of
grass that covers the land must be wet, in order that the ani-
mals may easily slip along in their vaulting gait over the green
road to their place of execution. Under the most favorable
circumstances, the march does not exceed six miles in the
twenty-four hours; and it being a distance of four leagues or
more to the village, three days and nights, or more, are spent
before they arrive at the slaughtering-place. There they are
allowed to remain quiet for a day or two, to cool their blood,
which becomes much heated by the tedious journey; after
which they are killed by shooting. The dead animals are then
skinned, and their hides packed in tiers until fermented suf-
ficiently to start the hair, when they are stretched on frames to
dry, and eventually become the covering or planking for the
Aleutian haidarlcm and baidarras. The fat is taken off and
used for fuel, or the oil is rendered to burn in their lamps. The
flesh is cut in thin pieces from the carcass, laid in the open air
to dry, and becomes a choice article of food. The sinews are
extracted, and afterward twisted into thread. The lining of the
animal's throat is put through a course of tanning, and then
made into boots, the soles of which are the under covering of
the Sea Lion's fln-like feet. The intestines are carefully taken
01)' cleaned, blown up, stretched to dry, then tanned, and
worked into water-proof clothing. The stomach is emptied of
its contents, turned inside out, then inflated and dried for oil-
bottles, or is used as a receptacle for the preserved meat ; and
what remains of the once formidable and curious animal is
only a mutilated .skeleton."
Captain Scammon adds the following respecting their cap-
ture on the Asiatic coast : "Crossing Behring and the Okhotsk
!: ' P-
268 EUMETOPIAS STELLERI — STELLER's SEA LION.
Seas, to the coast of Siberia, including the iieninsula ot Kaiu-
schatka and the ishind of Saghalien, the mode of cai)ture by tlio
natives chanj^es from that of the eastern continental siiores.*
The inlets iind rivers ot these Asiatic regions swarm with sal-
mon from .June to September, and at this season the Seals fol-
low and prey n])on them as they ascend the streams. Tlie
natives tlien select such i)laces as will be h^ft nearly bare at
low tide, and then set their nets — which arc made of seal-
thonys — to strong stakes, so placed as to form a curve oj)en to
the continence of the stream. These nets are similar to gill-
uets, the meshes being of a size to admit a Seal's head, — which
gives free passage to the shoals of flsli — and tin* ])ursuing ani-
mal, as soon as entangled in the net, struggles forward in his
efl'orts to escape, but is held tirndy in the meshes, where it re-
mains till low water, when the natives, in their Hat -bottomed
skin-boats, approach and dispatch the victim with their rude
bone implements. As the season becomes warm, the animals
of both sexes congrcgjite in their favorite rookeries, and the
females climb to the most inaccessible places among the rocks
and crags to bring forth and nurture their ofispring. But here
they are Inuited by the natives accustomed to the use of fire-
arms, who shoot them for the skins of the young ones, which
are used for clothing.
"In this region also, during the spring and fall, after the
'net-sealing' is over, great numbers of Sea Lions are captured
upon the floating ice, with gun or s])ear ; and during the rigor-
ous months, the seal-hunters cut through the congealed mass
what they term 'breathing-holes'. Through these the Seals
emerge, to the frosted surface, and, if the sun peers through the
wintry clouds, the creatures, warmed into new life, may stroll
hundreds of yards away; the watchful hunter, secreted beliiud
a cake of ice or a bank of snow, rushes out from his covert, and
places a covering over the hide, eft'ectually preventing the an-
imal's escape, and then disiiatches it with knife and si)ear. Its
jkin is stri[)ped off, scraped clean', closely rolled, ami laid away
'Although Caitiiiu Scimmon purports to i)e speaking of "Sea Lions," I
have recently beiome conviuced (since the copy of this article was sent to
the printer) that very little, if anything, in this paragraph and the next
relates to any species of Eared Seal. In the first place, the localitv is one
not known to be frequented, except casually, by Otaries, while the acconnt
of the capture in rets and in the ice, and especially the reference to
"breathing holes," renders it almost certain that the animals refened to
are Phocids.
^"^i
\l^
HABITS.
269
until the hair starts-^tliis process is called 'scouring'; then the
Lair is scoured off and tlie l)arc hide is stretched to season — a
process usuallj' requiring about ten days — when it is taken
down and rubbed between the hands to make it pliable; this
coiii,>letes the whole course of dressing it. The i)repared skins
are then converted into harness for the sledge-dogs and rein-
deer, and water-proof bags ; if Avanted for the soles of mocca-
sins, or to cov.er their skin-boats, they are dried with the hair
on, and become nearly as stiff as plates of iron. The blubber
of the auhnals, if killed in the fall or winter, is preserved by
freezing, and is vzed for food, fuel, and lights, as desired ; while
the same part of those taken in the spring and summer is put
in the skins of young Seals, and placed in earthen \aults, where
it keeps fresh until required for consumption. The residue of
the onimal is tumbled into a reservoir, sunk below the surface
of the ground, where it is kept for the mnter's supply of food
for the dogs, which live upon the frozen flesh and entrails of
the Seals, whose skin furnishes the tackle by which they trans-
port the primitive sledge over the snow-clad wastes of Siberia
and Kamschatka."*
Since the foregoing was transmitted for publication I have
received from Captain Charles Bryant a very full account of
this species, based on his many years' observations as United
States Treasury Agent at the Fur Seal Islands, and kindly pre-
pared by my request for use in the present connection. Although
so much space has already been devoted to the history of this
species, it seems desirable to give Captain Bryant's report
nearly in full, although repeating in substance some of the
details which have already been presented, since it contains
some new points, and is at least based on long experience.
Some portions, relating especially to the products of the Sea
Lion and their uses, are omitted, since they are fully anticipated
hy Avluit has already been given.
"Froni fifteen to twenty thousand Sea Lions," says Captain
Bryant, "breed annually on the Prybilov or Fur Seal Islands.
They do not leave the islands in winter, as do the Fur Seals,
to return in spring, but remain during the whole year. 1 aey
bring forth their young a month earUer than the Fur Seals,
landing during the months of May and June. They advance
but little above high tide-mark, and those of all ages land
together. Th0 strongest males drive out the weaker and mono-
* Marino Mammalia, pp. 136-138.
/
/
J
m
ill i .1 i.
270 EUMETOPIAS STELLERI — STELLER'S SEA LION.
polize the females and continue with them till September. They
yo with them into the water whenever they are disturbed, and
also watch over the young. When in the water they swim
about the young and keep them tr^^^e'lier until they have au
opportunity to land again. The females also keep near, rushiug
hither and thither, appearing first on one side and then on the
other of the groups of young, constr gives the
girth behind the axilhe as 1220 mm. (" 4 ft.").
A fully-adult female (M. C. Z. Coll., No. 5787) gives a length
(from tip of nose to end of tail) of 1800 nun.; to end <>f out-
stretched hind-rtipi)ers, 2054 mm. (collector's measurement from
fresli specimen, "0 ft. in."); girth behind axilla*, 1247 mm. (col-
lector's measurement, "3 ft. 9 in."); hind-flipper, 270 mm.; fore-
flipper, 310 mm.; tail, 70 mm.; ear, 30 mm.; longest whisker,
110 uuu.
Another adult female (M. C. Z. Coll., No. 5788) gives the fol-
lowing : Nose to end of tail, 1570 mm. ; nose to end of outstretched
* This measurement, is l)y estimate baaed on the collector's measurement
of the total length to «m>(1 of outstretched hind-Hipper, taken from the fresh
spctinien, the calculation being based on a study of the skeleton. The
total length of head and body, as taken from the mounted specimen, is
obviously much too short.
:1k
-im
280 ZALOPHUS CALIFORNIANUS — CALIFORNIAN SEA LION'.
liiiil»('rs, iOOGnuu. (collector's measurement ironi fresh s])('(i-
meii, 'MJ J't. 7 iu.")> girth behind axilltii, IOCS ikim. (coHectoi's
measurement, "3 ft. G iu."); tail 80 mm.; car, 34 mm.; lonj^cst
whiskei", 100 mm. The skeleton of an adult female (M. C. Z.
Coll., Xo. 0) has a total length to end of tail of 1700 mm. ; to end
of phalau; n of hind flipper, 1908 mm.
The collector's measurements of the young (two or tlirec
months' old) specimens are : Male, nose to end of outstretcliod
hind-flippers, "4 ft." (1220 mm.); glrtb, "2 ft." (GIO mm.).
Female, nose to end of outstretched bind- flippers, "3 ft. 8 in."
(1120 mm.) ; girth, ^' 1 ft. 11 in." (583 mm.). The fcetal specinioii
(M. C. Z. Coll., Ko. 5839, stuffed), already described, measures
from nose to end of tail 850 mm.; from noso to end of out-
stretched hind-flippers, 970 mm.; hind-flippers (from heel),
115 mm.; fore-flipper (from ax'Ua), 200 mm.; tail, 45 mm,;
ear, 25 mm.; longest whisker, 55 mm.
Captain Scammon gives thefoUowing measurements of "Sea
Lions," taken at Santa Barbar;i Island in April and May,
1871-73. They inchule an "adult male" (column 1); a male
(column 2), "about ten months old," taken April 4, 1872; a
fennile (column .i), "supposed to be a yearling"; and "a new-
born feui' .e Sea Lion pup " (column 4), taken May 3, 1873.
Length from t ip of nose to end of hind-flippcra —
Length from tip of nose to base of tail
Length of hind-flippers
Length of fore-flippers
Girth behind nxillic
Girth at base of hind-flippers
From tip of nose to eye
From tip of nose to car
Length of ear
Length of tail
Length of longest whiskers
From base of tail to posterior teats
From base of tail to anterior teals ] j
Distance between posterior tejits I o
Distauce between anterior teats i o
Thickness of blubber o
ft. in.
6 4
1
4
3
6
3i
8
u
2
6
2
10
5
8
Oi
2t
ft. in.
4 10
3 lOi
Hi
1 3
2 8
Hi
I
1
24
5i
Oi
3:
ft. in.
4 10
3 lOi
11
1
2
1
i^
21
7
1
3
6
li
24
0}
* Adult female.
t Male, ten mouths old.
t B^emale, .iliout one year old.
J Female, newly bom.
ft. in.
2 4
1 11
54 j
7 j
3
Oil
"i
4
14!
oil
SIZE.
281
Mcamrcmenix of Ihc Skeleton of (in Adult Female.''
mm.
\Vliolt' li'iiytli of skeleton (inclndiuf; skull) 1706
U'listlK'f sliiill 236
I.cii<;f li of cervical vertebr.T 320
Lciiiitli of dorsal vertebra- 640
L(iij;ili of lumbar vertebne 230
U'ligtli of caudal vertebra? (+ sacral) 280 '
Length of lirst rib, total 140
Leugt h of iirst rib. osseous X'ortiou 7.5
Length of lirst rib, cartilaginous portion 6.">
Li'iiKth of second rib, total 17:5
Length of second rib, osseous portion 100
Length of second rib, cartilaginous portion 7;{
Length of third rib, total 240
Length of third rib. osseous portion 158
Length of third rib, cartilaginous portion 82
Length of fourth rib, total 280
Length of fourth rib, osseous ))ortion 185
Length of fourth rib, cartilaginous portion 95
Liiigthof lifthrib, total 335
Length of fifth rib, osseous portion 220
Length of fifth rib, cartilaginous portion 115
Length of sixth rib, total 370
Length of sixth rib, osseous portion 250
Length of sixth rib, cartilaginous portion 120
Length of seventh rib, total 395
Lengt h of scventli rib, osseous portion 270
Length of seventh rib, cartilaginous portion 125
Length of eighth rib, total 445
Length of eighth rib, osseous portion ,.... 295
Length of eighth rib, cartilaginous portion 150
Length of ninth rib, total 445
Length of ninth rib, osseous portion 290
Length of ninth rib, cartilaginc^'s portion 155
Length of tenth rib, total 430
Length of tenth rib, osseous portion 280
Length of tenth rib, cartilaginous portion liJO
Length of eleventh rib, total 413
Length of eleven til rib, osseous portion 280
Length of eleventh rib, cartilaginous portion 133
Length of twelfth rib, total 395
Length of twelfth rib, osseous portion 260
Length of twelfth rib, cartilaginous portion 135
Length of thirteenth rib, total.. 362
Length of thirteentli rib, osseous portion , 247
Length of thirteenth rib, cartilaginous i ortion 115
Lengtli of founeentb rib, total 310
Length of fourteenth rib, osseous portion. 215
*No. 6159, Collection of Museum «)f Comparative Zoology.
H :;«
282 ZALOPHUS CALIFORNIANUS — CALIFOKNIAN SEA LION,
mm.
Length of fourteenth rib, cartilaginous portion %
Length of fifteenth rib, total -wii
Length of fifteenth rib, osseons portion 1«0
Length of fifteenth rib, cartilaginous portion 4u
Length of sternum, ossified portion :,:,{)
Length of sternum, 1 st segment llii
Length of sternum, 2(1 segment ;'ii
Length of sternum, 'M segment .")3
Length of sternum, 4th segment ii)
Length of sternum, oth segment 4^
Length of sternum, (5th segment 4T
Length of sternum, 7th segment 4il
Length of sternum^ 8tli segment ;!,«
Length of sternum, 9th segment 5")
Length of scapula 18(t
Greatest breadth of scapula 250
Greatest height of its spine It:
Length of humerus 15.J
Antero-posterior diameter of proximal end of humerus G3
Transverse diameter of proximal end of humerus lu
Transverse diameter of distal end of humerus ."
Length of radius l.j.')
Length of ulna. im
Longest (liiiuieter of upper end of ulna , tit
Length of carpus P
Length of 1st metacarpus and its digit ".'b
Length of 2d metacarpus and its digit 1"
Length of I5d metacarpus and its digit 1'"
Length of 4th metacarpus and its digit 1-"
Length of oth nu'tacarpus and its digit ' '."'
Width of niauus at base of metacarpals '"
Total length of fore limb (excluding scapula) •'>'!"
Length of femur '•"
Longest diameter of proximal end of femur 4ij
Longest diameter of distal end of femur ^'
Least antero-posterior diameter of shaft of femur !•'
Length of tibia 1^'
Length of tarsus J"
Length of Lst metatarsus and its digit '"-
Length of 2d met atarsus and its digit 1- '
liCngth of :{d metatarsus and its digit '""
Length of 4th metatarsus and its digit 1"^'
Length of 5th metatarsus and its digit l"'
Width of pes at base of metatarsals •"
Total length of hind limb •''■'
Length of innominate bone 1'''
Greatest width of pelvis anteriorly '""
Length of ilium '"
Length of ischio-pubic bones '^
SKULL. 283
][ii(nn>rmc)it8 of the Metacarpal and Phalanriral Hones of an Adult Female*
L('nj,'tli of mauna to end of .
Lfiiglli iii' mctiicarpal of
Liii^lh 111' 1st plinlanx of . . .
LiiiStli of 2(1 plmlanx of —
Length of 3d phalanx of . . . .
1st digit.
2d digit.
ad digit.
4th digit.
5th digit.
258
226
187
147
112
90
58
55
48
42
78
CO
55
35
31
50
47
3C
23
15
20
15
12
10
Measurements of the Metatarsal and Phalangeal Hones of an Adult Female.*
Length of pes (posterior end of os
call' is) to end of
Leiiglli of inetncari)al of
Length of Ist ph.Manx of
Length of 2d phalanx of
Length of ;id phalanx of
Length of nail of
Ist digit.
280
79
75
40
2d digit.
2ti0
Gl
60
40
16
10
3d digit.
60
57
48
16
18
4th digit.
250
CO
54
48
10
15
5th digit
250
65
58
37
Skull. — The skull in Zalophus californianits, as cojnpared
with the .skull in allied genera, is remarkable for the narrow-
ues.s and great elougation of the facial portion, which is even
iiuich more elongated and slenderer than in Arctoccphalus. In
its general configuration (excepting, of course, the great devel-
oimicnt of the sagittal and occipital crests in the very old males)
it more resembles the Arctocephaliue type than any other.
The maximum breadth (i. e., at the zygomata) in the females
barely equals or falls a little short of half the length, while hi
the old males it rather exceeds this proportion. In very old males
the crests of the skuU are enormously developed, and, con-
trary to what usually obtains in the other genera of this family,
are considerably developed in very old females. The superior
outline (in old males) slopes rapidly from the high sagittal
crest to the end of the nasals. The postorbital processes are
long and rather narrow, and are directed backward in old age;
the nasals are long and narrow, decreasing in width posteriorly.
The superior edge of the intermaxillai is very narrow, and is
prolonged backward nearly to the middle of the nasals. The
postorbital cylinder is long and narrow, and often abruptly
contracted posteriorly. The bony palate is nearly flat, but little
depressed, and is rather deeply emarginate posteriorly. The
'Specimen No. G159, Coll. Mns. Comp. Zoology.
I
1
284 ZALOPHUS CALIFORNIANUS — C\LIFORNIAN SEA LION.
paliito-iiiiixillaiy .sutuio is about oppos'tc the liiiulor odge of
the last molar. Tlie ptcryj^oid hamuli are small. Tlu- postciior
uarial ojx'uiuj;' is wider tlian deep; the autcrior lias those Iwo
dimeusious about equal.
Ill ZaiophiiH the superior aspect of the skull, before the devol
opiiu'ut of the erest, is strikingly like that of Arctoc('itli((liiN,;\s
indeed is also the inferior aspect, aside from the dental I'or-
mula. In Zaloplnis the auditory Indian are rather less swollen
than in Arcton^ihahis, but in all other resi)eets there is a strik-
ing resemblance. The anteorbital portion of the skull, how-
ever, is more attenuated, and relatively much longer. With
this exception there is little difterence in the general confor
mation of the skull in middle-aged females of thes<^ two genera,
while both differ widely from Otaria, Eiimctopias, and Callorhi-
mis. The great development of the crests of the skull late in
life in Zalophtis gives it at that time a highly peculiar confor-
matiou.
SKULL,
285
•A
%
M
<
■J
n
S
<
•Si
a «
O V
11
to . to
"S a "5 c3
. "o . i . "o . i
fi c- a ? fi c a ^
t.
s
■)80JD HI)I(Il03(> JO ■UlSl'lH
•BOj.Cpno,) 0} Aii!('aoA\o[ JO njSaii
C CO t-1 -^ CC M CI O »3 'f
CI -H CI CI i-H 'H r-i i-H tH ft
•Hojaos ai![oui .idM3n.rj
'SJi.it's .iu[oia jjdcin jo qjSnoi
•soi^pnoD ii))|dpao jo q'jpBajg;
IBJOdlUJ) ■)» 1111518 JO TIlpBajq !}8tl31
O 00 M« 00 t^ t-
lo ^ ^ ^ ^ ^
s?
s
^ Irt iH CI « t-
e% ro c^ C4 CI CI
•soujuii;) ^u tiniiB JO iijpuojfi
-xutu-ojitoaj ^11 sonoq xii8nn jo inpuoag;
'^[i0U3')nn 8aaoq {"^^'^ JO qtpv^'iQ
'eodoq \WBVL JO q^Snai
•asnjxBin JO paa io\i
■0)80(1 !)B oaejine oni^uied jo qjpu.UH
'jap
-joq jt)xoaA|n Suop DnjxBra jo q'jSnsi
■B8a30id JtipniBq jo paa o;
oin^iiB iSiBHl^^ni-o^BiBd mojj aotiB^Bid
•BPOooad pionaiS^Bod o; bjubhixbui
•jo;ut JO o3pa jotjinuc lucij aDUB^siQ;
-JBIOm ')8BI O) BapBIIIXBUI
■ao)ni JO oSpa jot.io;nB raojj oauB^ejd
•pioS.vjojd JO es.i.i
-Old aBiniuoq jo pno 04 soiaBnixBui
■10)01 JO oSpo jouojUB mo.ij ooob^biq;
■B38Ba30jd pio)SBiu )B qipBajQ
iHrH ISSqoooSS
''
: S3
rH
T^^
i ^
g i
; S8
• 1—1
t '
11 ^ : 2 S 3
Moicr-O'^iooi^
•i^oiB OT)QaioSiz ^B q^puajq ;83;b9JO
CO t' W C>l i-<
•qi^Suoi
s
w
•X3g
•D*D^^"D"oOfOCH.OO♦■
I
,a I
2
SI
c o o o o
.0 c3 "^ 'IS 'd TS 'd
'jaqmim onSoiB)B3
rH ■* in
.
loll
.g s esj
a .58 g
iai*
« s s
^ o
o ee «
'
28G ZALOPIIUS CALIFORNIANUS — CALIFORNIAN SEA LION,
Dentition.— Tlu> teeth in ZahpJim are uU stronj^ly (level-
oi)e(l and very ttriiily implanted. All are single-rooted except
the last inolai', wliieh is imperfectly double-rooted. The molars
all have a distinely beaded citijjulum on the inner side. The
lower molars and the fifth and sometimes the fourth ui)j)or
molars have a small but distinct anterior cusp. The canines
and the incisors present the usual form seen among the Otaries.
The teeth of a roiddleaged m.ale skull i)re8eut the following
measurements :
McasurcmentH of the Teeth.
A.— Teeth of tuk Upper Jaw.
Molars.
CaninrB.
Incisors.
5th.
4th. i 3d.
10
7 9
8. 5 j 10
2d.
10
8
9.5
Ist.
Outer.
Middle.
1
Imicr.
Antero-posterior diameter.
Xransvcrse diameter
9
7
8
7
r
9
17
13
20
4
15
5
3
5
1
5
2.5
4
B.— Teeth of the Lower Jaw.
t ■
1 '
1
i
■ i ■ 1 ■-'■
i
■
Molars.
Canines.
Incisors.
5th.
4th. ' 3d. ' 2d. Ist.'
1
Outer.
Inner,
Antero-postenor diameter . . .
Transverse diameter
Height of crown
9
6.5
8
12 11 10 ' 8
8 9 7 1 7
9 9 7.5 G
18
11
25
5
C
G
4
4
The molars arc usually closely approximated, but sometimes
there is a small space between the two hinderraost of the series,
and occasionally they are all slightly and evenly spaced. The
hinder edge of the last upper molar is generally anterior to or
about <}ven with the posterior border of the zygomatic process
of the maxillary. When more than Ave upper molars are pres-
ent, the sixth or supernumerary is postt^rior to the fifth, and is
usually smaller than the fifth (sometimes almost rudimentary)
and lack' le accessory cusps seen in the fifth.
The milK dentition (fully represented in three skulls before
me and partly so in five othei^s) does not differ from that of the
other species of the family, and has been already fully described
(see antea, p. 223).
VARIATION WITH AGE.
287
Skxual Diffkre>'cks. — From the testimony of Captain
Swiiimion, ami from the matetial I have been able to examine,
the Iciiiah' ditfers from th«i male in eohn- in being rather lighter,
or(»f a more yellow isli-brown. The most notable ilitferenco is
ill size, the female being very nmch the smaller, but not quite
so j;reat a sexual disparity in size obtains in this species an
in Kunwiopias stelh-ri and CoUorhinnn ursinus.* Unfortunately
die material at my (command will not enable me to give full
statistics on this point. iMost of tin; nude si)ecimens in a large
series sent to the jVIuseum of Couii)arative Zoology, by Mr.
Schumacher, from Santa Barbara Islands, are young or middle-
iXiiod, only one having the teeth perceptibly worn or the crests
of the skull very highly developed. ^V comparison of very old
skulls of both sexes shows tluit nearly the usual amount of sex-
ual variation in size common to the Otaries obtains in the pres-
ent species. The table of measurements (on page 28i5) of ten
skulls — five nude and live fenmle — all fully adult and most of
them very old, gives all tlui information I am able to ofter re-
specting sexual wiriation in size.
As usual in this group, the dental armatiue (especially the
canines and caniniiVu'm incisors) is much weaker in the female
than in the nude, by means of which the skulls of females can
lie readily ilistinguished from those oi males of about the same
size. The whole skull is slighter and weaker, and all the pro-
eossos and ridges for the attachment of muscles much less devel-
oped. There are, however, in very old female skulls, , to 35 or 40 mm. The limbs are also nnich weaker
and slenderer, as of course are all the bones of the skeleton.
Variation with Age. — As already noticed, the color of the
young at birth is dark gray or slaty, and the pelage has at this
inu) a delicate softness, due to the silky texture of the hair.
Tlie pelage is wholly devoid of a second coat of true under-fur,
like that of the Fur Seals, but from its softness might readily
lie mistaken on casual observation for true fur. This is very
soon replaced by a coarser and harsher, but still quite soft
pelage, in comparison with that of adults, of nearly uniiorm
chestnut or dark reddish-brown color. This is succeeded by the
harsh, stiff pelage of the middle-aged and adult animals.
•V .flU
' See flHr. Tlic
rupiU'ity of tlu' brain-ciiso tlocs not j,'i"Oiitl\ incrciwciiftci" Itiitli,
it enliiifiiiij;' mainly by the tliiclvcninj;' of its walls. Tim ante-
rior half of the sknll dovt'lops rapidly and Utt'rs very nuicli in
form. At birth the inter- and anteorbital jtortions of tln^ skull
are very short, they to^etuer fonninjjf rather less than half the
lenjjfth of the sknll, whihun fnlI-}»rown sknlls theyeomprisc alxnit
tw r>-thirds of its length. In a young skull (Nat. ^lus., Xo. irtddo,
4 taken from an animal killed a few days after birth), which lias
J a total length of 140 mm., the brain-case alone lias a length of
78 mm., the interorbital region* a length of .'U mm., and the ante
'orbital a lengtli of 37 mm., giving a total length from the an-
, terior wall of the brain-case to the front border of the interinax-
illiu of 08 mm. In this skull (as in several others before iiw of
about the same age) the occipital condyles are wholly anterior
to the plane of the occiput. In a very old fenuUe skull (M. C. Z.
Col., No. 0150), with a total length of 233 mm., the o(!cipital col^
dyh's project 15 mm. behind the occipital plane. Of the remain
lug -18 mm. of the length, the brain-case occupies 8.'} nnii., tLc
interorbital region 05 mm., and the anteorbital 71 mm., and the
two regions together 130 mm. In the first the ratio of the
length of the brain-case to that of tln^ rest of the skull is as 78
to 08; in the last as 83 to 130, In a middle-aged male skull,
the total length is 282 mm., of which the condylar extension is
22 mm. Of the remaining 200 mm., the brain-case occMipies 0.)
mm., the interorbital region 78 mm., and the anteorbital 87 mm,,
making the proportionate length of the brain-case to the rest of
the skull as 95 to 105. The ratios between the dift'erent regions
in these three skulls are as follows :
Batio of brain-case to whole skuU
Katio of interorbital rcRion to whole ekiill • . -
Ilatio of anteorbital region to whole skull
Condylar extension to whole skull
Yoting.
53./V-100 I
21. -J— 100 I
2"). 3— 100 I
Female.
."s:.. 0—100
28 —100
30. 5—100
« —100
Male.
33. 7 -UK)
30. 8,')-100
27.CO-100
7. 7 -IIX)
The width of the brain-case in these skulls is respectively
90 mm., 97 mm., and 107 mm.
In adult skulls the breadth of the interorbital region is relu
tively, and generally absolutely, much less in adult skulls tbau
at birth, and the point of greatest constriction is placed much
* That is, the narrow portion of tlie sknll honntled laterally hy 'he tempo-
ral fossae and orbits.
jilHr
m
ili^i
GEOGUAI'IIICAL DISTRIBUTION.
289
iiiort' postcriorl.v, bciiij;' in the atliilt sit the posterior i'lul of tho
tciniMiiiil fossa', iiitd in lin' yomij'' at the orbits. T\n\ bicadtli
of tilt- siuill Just ill front of tln' luaiiiliox in very yoiinj'' skulls
(tiiosf taken a few " lys after birth) is 10 to 12 nun., in tiioso
three or Ibnr nu)Utli.« iy\ .'W inin.; in adult females, usually L'li to
;;(» iMiii. ; in adiili males, about .'50 to .'So mm. The amount of eon-
strict ion varies somewhat in atliilt skulls of the same sex, tho
(•(Histrietion increasiiif^ with the advaiuie of aj;e. There is a
correspondiuj,' contraction posteriorly of the palatal region. In
very youii}; skulls, the. palate is widest at the pterygoid ha-
iimli ; in those a few months old it is nearly straight, but later
in lite becomes narrowed posteriorly, the contraction being
},m'atest in aged sjiecamens, in wliicV. the width at the pterygoid
liimiuli is a third less than it is at the last molar.
The crests of the skull do not begin to develop until the ani-
mal roaches adult size, and attain their highest development in
V(rv old specimens. In a series of thirty skulls, only two have
tlio crests remarkably developed, these being the two old male
skulls described by nie in 1870.* In only one of the skulls of
tli(* series, aside from tho two above mentioned, are the teetli
much Avorn. The two very old skulls show, by tlieir large size
and lugose eliaracter, that the deposition of bony matter is
(ontiiiued to a very late period in life.
("05IPA11ISON WITH ALLIED SPECIES. — Zalophus mUfomianns
is too distinct in cranial characters and dentition to require com-
parison with any of the Hair Seals of other genera, while its
polaf-c and color aftbrd obvious points of difference from the
Fur Seals. As rcispects the conformation of the skull, it finds
its nearest allies in Arctocephalus, from which, however, it is
readily distinguished by its more elongated muzzle ami dental
t'onnulii. It appears to closely resemble its congener, Z. lobattis,
both ill size and color. Hm ing no specimens of that species
!it coiiniiand, I am unable to state the points of difference be-
twoou the two. The descriptions and figures of Z. lohatxis indi-
cate their close alliance.
Gkogkapiiical Distriiuttion. — Tho exact boundaries of
the habitat of Zaloplins californianus cannot at present be given.
Tlie only specimens I have seen are from the coast of California
1111(1 its islands, from San Diego and San Nicolas Island north-
ward to the Bay of S'ln Francisco. Captain Scammon (see
*i5ull. Mus. Coniii. Zool., vol. ii, p. 69; seo measurements at p. 70.
^lisc. Pub. No. 12 19
290 ZALOPHUS CALIFORNIANUS — CALIFORNIA^ SEA LION.
infra, pp. 301, 302) twice alludes Incideutally to its preseuce
'along the Mexican and Califoruiau coasts," and Dr. Yeatcli
states that "Sea Lions" (which he calls '■' Otaria juhata, but
which are, almost beyond doubt, the present species) laid
populous breeding stations twenty years ago, and doubtless
have still, on Cerros or Cedros Island, in about the latitude of
28i°, off the Lower California coast. Whether they occur south-
ward of this point at the present time I am unable to state, but
should infer that such was the case from Scammon's allusion
to their capture along the "Mexican" coast. In any case, it
appears probable that in Dampier's time they ianf>ed as for
south as thg Chametly aiul Tres Marias Islands, respectively
in latitudes about 23° and 21°, at which points he saw "$eals''
in the year 1G8G. In describing the Chametly Islands (the most
northerly of the two groujjs mentioned by him under this name),
situated off the "VVest coast of Mexico in latitude 23° 11', he
says: "The Bays about the Islands are sometimes visited with
Seals; and this was the first place where I had seen any of
these Animals, on the Xorth side of the Equator, in these Seas,
For the Fish on this sandy Coast lye most in the Laguues oi
Salt-Lakes, and Mouths of Rivers; For this being no rocky
Coast, where Fish resort most, there seems to be but little Food
for the Seals, unless they will venture upon Cat-Fish."*
He also met with Seals at the Tres Marias Islands (in liiti
tude "21° 5'"), and consequently two degrees south of the Cha-
metly Islands, in describing one of which islands, named by
him St. George's Island, he says: "The Sea is also pretty well
stored with Fish, and Turtle or Tortoise, and Seal. This is the
second place on this Coast where I did see any Seal: and this
place helps to confirm what I have observed, that they arc sel-
dom seen but where there is plenty of Fish."1
It is of course not certain that the Seals here alluded to arc
Zalophus calif or niamis, since the Sea Elephant of the Califor
nia coast also occurs at Cedros Island, and probably still further
south, the two species having apparently about the same rauge.
If they had been the latter, Dampier would probably have made
some allusion to their large size.
The species of Zalophus occurring in Japan has been by some
writers considered to be the same as the Californian one; but,
though doubtless closely allied, its affinities, as will be noticed
* A New Voyage round the World, 5th ed., vol. i, 1703, pp. 263, 264.
tibid., p. 276.
r
GENERAl, HISTORY.
291
later (see iufra, p. 29,3), appear to bo not as j-ot satisfactorily
{letcrniiiied. As XaJophits calij'ornianus has not yet been detec-
ted oil the Aiuericau coast north of California, its occurrence
on the Asiati(5 coast seems hardly to be expected.
(lE^E^^AL IIistouv. — This s]>ecics has hitherto bten believed
to be free fro:.>i any serious coujidications of synonymy, and to
liave be«'n tirst brought to the notice of the scientiiic world by
M'lJiiiu in 1858. The only synonyin hitherto (pioted has been
Otanu .steUeri, "iSchlej^el" (/. t',, Temminck), which Dr. Peters*
.stated, alter an examination of the original specimens i>reserved
ill the Leyden museum, to be identical with tlie O. (jillcspH of
M'Baiii. A re-exaininatit)n of the subject, in the light of much
new information and material, shows <^hat the lirst notice of the
sjiecies was published by Choris in 1822, under the name of
"Lion niarin de la Californie,'" who gave a rather poor figure
of it in plate XE of his chapter entitled "Port San-Francisco et
SOS Habitants." As already stated luider the head of Eume-
topias .steUeri, his only reference to it in the text of this chapter
is as follows: "Les rochers, dans le voisinage de la bale San-
Fraucisco sont ordinairement converts d^ lions marins, pi. XI."
Ill bis account of the Aleutian Islands, however, he again refers to
it. and clearly indicates its characteristic external features. lie
says: ''Ces animaux [Lions marins] sont aussi tres-comnmns au
port de San-Francisco, sur la cote de Californie, oil on les voit
ou noiii,»re prodigieux .^ur les rochers de la bale. Cette espece
in'a paru se distinguer de ceux (pii fre»sse Ibrtemeut au run, tandis
quo ecu .. des lies Aleoutiennes sont d'une couleur plus grise,
out le corps plus rond, les niouvements plus ditliciles, la tete
plus grosse et plus epaisse; la couleur du poll des moustaches
plus iioiratre que celui des iles Aleoutiennes."t
The importance of this reference turns upon itsbeingan « "olicit
iudication of the character of his "Lion marin de la Californie,"
the subject of "Tl. XI"; this being, as is well known, the basis
of Lesson's Otaria californiana, which has hitherto been re-
ferred to Eumctopias stclleri, but which is really the same as
the so-called Zalophns gillespii. Lesson says: "Cette espfece,
tVapres la figure de Choris, a le pelage ras, unilbrm^ment fauve-
brunatre,les moustaches peu fournies; le museau assez pointu;
*Monat8b. Akad. Berlin, 1866, p. 669.
t Voy. pittoresque, lies AliSoutiennes, p. 15.
I f^
m
292 ZALOpiirs ("amkormamts — californian sea licx
les iiKMiiUrcs imtc'iit'iiis sont iv;;iili«'is. nlus f;ran (){' Zaiophus, ill tlu'. same connection, ^^ Otarla Gillespii
>[;i(lKiiii," and subsequently, in tlie same paper, so character-
i/cs Ills li'enera EiimctopUix iu\([ Zohphm as to leave no doubt
tliiil Ennu'topUtH relates - only do the skulls
lijiured by ; • niinck show that the species is not Eumctopias
stcllrri, but . coin])arative remarks respecting its relationship
t(i <>. jiiltnta indicate unmistakably tlu^ same thing. Although
I at one time acee[)ted Teters's deterinination of Tenuninck's
Otnria stelleri, a subsequent examination, in the light of nuu-h
new material and information, has led nu' t(» doubt its correct-
iioss. Tlie range of Zalaphiis caliJ'oriiiainiN {=fiillespii) has not
liccn reported as extending northwanl on the .Vuu'rican coast
licvniid California, and no si)eeimensof this sjiecies (except one
citiMl by Cray, the identitication of which seems open to cpies-
ti(in) liave been thus far recognized frjm Japan or any i)ortion
of the Asiatic coast. Temminck, with good series of the Japan
species and of the ZaJophnx ?o/>«/«s before him (he seems not to
liiive had the true E. stelleri), was unable to recognize any ap-
piwiable differences between them. In comparing his Otaria
utelleri with the Otaria australis of Quoy and Gainiard, he says:
"Uii crane absolument send)lable A celui figur6 par les voyageurs
(lout nous venous de i)arler [Quoy et (iainuird] a et<5 ddcrit sous
le uoui d'Arctocephalus iobatus, par Gray, Spic. ZooL, 1, p. 1,
pi. 4, fig. 2 et 2 a; ce crane provenant de la collection de fen
Biookes fait maintenant partie du Musee des Pays-Bas ; 11 ne
«' distingue en effct par aucini caracterc essentiel de celui do
' I'roc. Essix Inst., v, 1860, pp. 7, 11, and IH (footnote).
tlVtiTs says: " UobrigeiiS zweillc ioh jetzt utu'li j^ar uieht luehr dur.an,
ilass o, friZ/icsjiH Machain und (>. ./"iw im Schlogt-l [Ms. — o. stcllrri, Fauna
JaiKiiiicaJzudiTsellxMi Alt goliihon, da die Schiidol bcidcv nichtallcin indcr
Form, soiidi'in aucL in dcr Griisso iuitciuandcr ill)ori'iii8tinnn«Mi. Denn dor
iilti' Scliiidol von (>. GiUk82>ii ist G."' 295 lang, wuhrend altc Schiidol dos
Lciiliiicr Musuunis von 0,japonka 0."> 270 bis 0."' :{10 lang sivV—Monatab.
Jhd. ItciUn, loGO, p. 601).
|i
m
ijj
1
294 ZALOPHUS CALIFORNIANUS — CALIFORNIAN SEA LION.
I'Otai'iu iiiistrali.s vt and [of Temniinck's x>late xxii] are evidently
GjIpHdphora,'' but thinks they may belong to an undescribed species.
{Snppl. Cat. Seals ami Whales, 1871, p. 24.
sU land-List of Seals, 187(5, p. 42.
GKNERAL HISTORY.
205
I'liUy a('('(']»t(Hl Pfttors's reference of it to the Zaiophus (/iUespii.
Peters liiinself (irst (in 18(»7) referred the species (except the
\\enerically from the other Eared Seals. This skull,
and another (beh)iij;injn' to the mus(Mim of the Chicago Acad-
emy of Sciences), also from California, I was able to describe in
(li'taii in 1S7().^| These Californian skulls sire the only ones thus
far described,** but Scammon,inhis 'IMarine Mammalia," under
the name " Eiinietnpias HtcUcri,^'' has given detailed measure-
*>[oii!itsl>. (lor Aknil. dcr Wiasensch. zu Boiiiii, ISW), (18()7), pp. 27-<». 276,
1)68.
tSupid. (.'lit. Seals and \Vhalf\s. y. ys.
t Hand-List of Seals, p. 41.
v\"l.")89/). Skull, l'2i iiu'lios 1<»U}>, with canimvsvoiy large ; no other toetli ;
110 lower jaw ; frontal crest very high. .Japan, 7;}. W. 12. 1."
II Dr. Peters cites Sohlogel as the author of that p.art of tho "Fauna Ja-
poniiir' relating to tho Mammals, although puhlished as "par C. J. Tem-
ininck."' Mii.led by Peters, I made the same enor in my paper on the
Eared Seals, puhlished in 1870.
1 They were not tigured in the regular edition of my paper ou tho Eared
Seals (Bull. Mus. Comp. Zool., vol. ii, No. 1), hut two photographic plates,
ropreseutir.g both specimens, were added to a few of the author's copies (ahoat
twcnty-iive), which were scut to some of the more prominent workers in this
fitltl. These interpolated plates have been referred to by Dr. Gray (Hand-
List of Seals, p. 42) as though they formed a part of tho original work.
"* During the last year, I may here add, as an indication of tho amount of
material relating to this species now accessible, that I have examined not
less than a dozen skins, representing adults of both sexes, and young of
various ages from a foetal specimen upward, and more than twenty skulls,
likoA\ se embracing young, even with tho milk dentition, and both sexes of
various ages, and two complete skeletons.
I .^ I
31
(liTl
i
2^H] ZALOPIIUS CALIFOKXIANUS CALIKORNFAN SKA Ut)X.
incuts <»r what I tak*' to he ('xainph's of tliis simtjcs IVoiii tlio
Faralloiic and Santa Uarbara Islands.*
IlAtUTS. — Several more or less fall aeeonnts ol" (lie iiabilsof
liie Calilbrnian Sea I^ions liavi- been .iniNcn by (iin'erent wiitcis,
who have, however, failed to distin;^nisii tlie two species occiu-
rin,ij' alon;^ the ( 'alifoniian coast, and conseiinentl.v their descrip-
tions are not wholly satisl'actory. The larj^e Xortiiern species
certainly (»ccnrs, an evidence of its prescnc*' at Santa liarbara Island.
Kven Captain Scannnon, in his account of the Sea Lions of
California, has not distintly r«>co}inizelaiii t^oaminoii publislu'd his liist ai-coinit ol' I lie Si-a l^ioiis in tlic
"Ovi'vlaiul .Mcintlily " iiiajia/iiio (vol. viii, i)it. *J()ii-"i7-', Man'li, lt*7'J), in an m-
ticlc ciitiflt'il ''Aliont Sea I^ions." whicli is siilisfantially tlic same as Unit in
the ••Marine ^fanimalia," with Ihc oinissio'i of li;;iu'fs and aliont two lui^cs
of taliiilatt'tl nicasurcnit'iits and other details, liascd on siiecinicns siilisr-
qucntly obtained at llie I'aralione and .Santa ISarliara Islands. In a loot-
note in the ••Marine Mammalia" (|>. l-"»), he refers to his loimer article , 'is
follows: '•Since the )in1dii'ation (d' the article 'Almnt Sea J. ions," in tln'
'Overland Monthly' of .*»e)deinlier. 1-71 [/c'/c ^larch. 1-T^Jl], we have had
opportunity (d" makiny; additional ohserva t ions npon these ainmals at tlio
Faralioue Islands, where we saw the largest females we Inive ever met with
oi\ the Califoriua coast, lleiici-, what we have formerly taken to hv the
Eumvtoputu StelUri may prove to lie the /.(ilophiin (Ullixpii .*; hnt if snchhe the
fact, Ixdli s[)ecies iidiahit the coast of California, at" least as far south as
the Farallones. Moreover, both species, if we may be allowed the expres-
sion, herd tojietlier in the .same rookeries. On niakin<;' ii series of olKserva-
tions npon the ontwavd forms of Sea Lions, it will he foiuul tlnit a confusing
variety exists in the lif^nres of these very interesting animals, especially in
the shape of the head — sonft^ having u slinrt muzzle with a full forehead
lEitmctopias gteUeril;; others with forehead and nose somewhat elongated
[_Zalophun (aHfornianus=gillciipi, auct.] ; and still others of a moditied shape,
between the two extremes [£. stellcri, female?].'' In this eonuection it may
he noted that four of the five specimens of wliicli Captain Scaranion gives
measurements in the "Marino jVIammalia," were taken a//crthe imhlicatiou
of the article in the "Overland Monthly," namely, No. 1, "lull-grown male,"
Farallones. July 17, 187*2; No. 3, male "about ten mouths old," Santa Bar-
bara Island, April 4, 187'2; No. 1 his, female, supposed to he a yearling, ami
No. 2 bis, female, new-lxu'ii pup, same locality, May 3, 1873. The other, No.',*
(referred to in the " Overland Monthly" paper), adult female, Santa Barhaia
Island, April I'i, 1871. The first (No. 1, full-grown male) I refer with little
hesitation to L\ stellcri, and the second (No. 2, adult female), to Z. culij'or-
niamis, especially as I lind skulls in the National Museum, receive J from
Captain Scammon, agreeing respectively with these in locality, sex. and
age.
i: II
HABITS.
207
iiii;; tlu'ic, uiul hits (U'S('iii)ti()ii doubtless ioIVts in ])art to botli
s]M'(its. hut nnquostiouahly n'latos nuiinly to t' .'. prcscul one*
Ill's ••Sketch of !i .sciiiliuH' srasou u[>oii Santa llarbara Island,"
ill isr>i,'. ])i(>siMnabVv relates exclusively to ZnlopliiiN nilifoi-
nidiiiis, ]ni{ in addition to this I <]uote a few paiaj^rajdis IVoni
his jicueral account of "the. Sea Lion," since it is the testimony
of a trust\v(U'tiiy ey«'-\vitiu\ss. "On approachinji' an ishind, <»r
point, occupied by a luunerous herd," he observes, '-one tirst
lioais their ion<>-, ]>laintive howHniisin|4' velocity, frequently
(liviii;; dill side the rollers, the next Tiioment cmerginjj from
tilt' crest ol' the foiiiniii;; breakers, and waddlin*;' up the
hcacli with head ereet, or, with seeiiiiiiy elfort, cliiabinj^- some
ki'lp rriii.i;<'d rock, to doze in the scorchin<;' sunbeams, while
(itlicrs would lie sleeping or playing among the beds of sea-
weed, with tlit'ir heads and outstretched limbs above the sur-
liice. l>ut a few days elapsed before a general contention with
the iidiilt males began for the mastery of the difi'erent rooker-
ies, and tlie victims of the bh)ody encounter were ♦^o be seen on
all sides of the island, witJi torn lips or mutilated limbs and
j,'ashed sides, while now and then an unfortunate creature
would l>e met with minus an eye or with the orb forced from its
socket, and, together with other wounds, presenting si ghastly
appeanmce. As the time for 'hauling-up' drew near, the island
hecaine one mass of animation ; every beach, rock, and cliff,
wlieie a seal could find foothold, became its resting-place, while
a countless herd of old males capped the summit, and the
united elamorings of the vast assemblage could he heard, on a
calm day, for miles at sea. The south side of the island is high
ami precipitous, with a projecting ledge hardly perceptible
from the beach below, upon which one immense Sea Lion man-
aged to climb, and there remained for several weeks — until the
.season was ovei*. How he ascended, or in what nmnner he re-
tired to the water, Avas a mystery to our numerous ship's crew,
as he came and went in the night; for 'Old Gray,' as named
by till' sailors, was closely watched in his elevated position dur-
ing the time the men were engaged at their work on shore.*
••None but the adult males Avere captured, AA'hich was usually
(lone by sliooting them in the ear or near it; for a ball in any
other i)art of the body had no more effect than it would in a
Grizzly Bear. Occs^pionally, however, they are taken with the
flub and lance, ordy shooting a few of the masters of the herd.
" * Ifcltitive to the Sea Lions leaping from giddy heights, an incident oe-
ciutimI at Santa Barbara Island, the last of the season of l&ij2, which we
will here mention. A rookery of abont twenty individuals was colle<;ted
nil iho brink of a precipitons clift", at a height at least of sixty feet above
the Kicks which shelved from the beach below ; and our party were sure
ill their own minds, that, by surprising the animals, wo could diivo them
over tlu' clift'. This was easily accomplished ; but, to our chagrin, when wo
arrived iit the point below, where we expected to find the huge beasts heli)-
lessly iinitilatcd, or killed outright, the last animal of the whole rookery
was seen plunging into the sea." ,
aOO ZALOPHUS CALIKORNIANUS — CALIFJKNIAN SKA LION.
.1;:
This is easily accoinplisiicd witii an cxpniciiccd crew, if (licro
is siilliciciit ^^roiind liaok IVoiii the beach for tiie animals (ore-
treat. Durinu,' our stay, an instanei' oeeurred, which not only
displayed the saj;aeity of 1 he animals, hut also their yieldinj;
disposition, when hard i)ressed in certain situations, as it' nat-
nrally desij;ned to he slain in nund)ei's e(|ual to the demands of
their human pursuers. On the south of Santa Uarhaia Islmid
was a i)lateau, elevated less than u hu'ulred feet above the sea,
«tretchin;;' to the brink (d" a <'lill' that overhunj;' the slum", and
a narrow ^orp! leadinjn' up from the beach, throuj;h which the
animals crowded to their favorite restinj^-jdace. As the sun
4lil)ped behiiul the hills, lifty to a hundred males would conjLiU'-
gate upon the spot, ami there remain until the boats were low-
ered in the morning, when inunediately the whole herd would
(puetly slip oil" into the sea and gand)ol about (bu'ing the day,
returiung as they saw the boats again h'ave the island for the
ship. Several unsuccessful attempts had been made to take
them ; but at last ti fresh breeze connneiKH'd blowing directly
from the shore, and inevented their scenting the hunters, who
landed some distance from the rookery, then cautiously ad-
vaui'ed, and suddenly yelling, and nourishing muskets, clubs,
and lances, rushed up within a few yards of them, while tlic
pleading creatures, with lolling tongues antl glaring eyes, were
quite overcome with dismay, aiul remained nearly motionless.
At last, two overgrown males broke through the line fornicd
by the men, but they paid the i)eualty with their lives befoir
reaching the water. A few moments passed, when all hands
moved slowly toward the rookery, which as slowly retreated.
This maneuvro is termed ' turning them,' ami, when once aecoin
plished, the di.sheartened creatures appear to abandon all hope
of escape, and resign themselves to their fate. The herd at
this time numbered seventy-five, which vffire soon dispatched,
by shooting the largest ones, and clubbing and lancing tlie
others, save one young Sea Lion, which was spared to see
Avhether he would make any resistance by being driven ov^^r
the hills beyond. The poor creature only moved along throngh
the prickly pears that covered the ground, when compelled by
his cruel pursuers ; and, at last, with an imploring look and
writhing in pain, it held out its fin-like arms, which Avere pierced
with thorns, in such a manner as to touch the symx)athy of tbc
barbarous sealers, who instantly pnt the sufferer out of its mis-
ery by a stroke of a heavy club. As soon as the animal is
irAHITS.
301
killed, tlu' l<)ii},'t'st spiiL's of its whiskers ur*; i)iilkMl out, tlu'ii it
is skinned, sinU its coatinj;' of fat cut in sections from its body
and lr!ins)»orte(l to the vessel, wliere, after Iteinji' 'minced,' the
oil is cxlriicted hy boilinj;. TIh' testes are taken out, and, with
Ihc selccitsl spires of whiskers, lind a market in China — the
former bciuj;' used luedieiually, and the latter for personal (una-
iiicnts.
'•At the <'lose of the season — which hists about three nuinths,
(111 the Californiu coast — a hirge majority of the ^jreat herds,
both males and fcuudes, return to the sea, and roam in all
directions in quest of food, us but few of them could find sus-
tenance about the waters contijjuous to the islands, or points
on tlic mainland, which are their annual resortiiig-places. They
live upon fish, mollusks, crustaceans, and sea-fowls; jilways
with the atldition of a few pebbles or smooth stones, some of
which are a pound in weight.* Their principal feathery food,
jiowever, is the penguin in the southern hemisphere, and the
ii\\\h in the northern ; while the manner in which they i)er and Lower Califwnia, and thousniuls
of barrel."^'. «»f oil obtaine»l. The luimber of Seals slain exclu-
sively for their oil would appear fabulous, when we realize the
fact that it requires on an average, throu^'hout th(^ season, the
blubber of three or four Sea Li(uis to i>roduce a barrel of oil.
Tlieir thick, coarse-grained skins were not considered worth
preparing foi- market, in a country where manual labor was so
highly valued. At tlu' present tiuu', however, they are valued
for glue-stot-lik<> Iiciid, aiui (HiiiipIiU'ciitly looking
iiltoiil liim. Tlu'.V lire n r(ni;;li s«'t of brutes, — liink Imllics, I
>limil(l sii.v ; lor I liiivc wiitcln'tl lliciii rcpciitcilly, as a hi;;- Cci
low slioul(U'i«', with tciith and
iii:iiii luivc, cxpt'lli'd the weakiT IVoiii his hid;;iiu'iit. The sinalh'r
Svii liioMs, at h'ast those whi<'h liave h'I't their mothers, api)ear
tit liiive no ri;ihts whii ' any one is honnd to respect. Th»'y get
out of tlie way witli abject promptness, whicli proves that thoy
live in terror of the stron^^er mend)ers of the community; but
tlicy d(> not ^-ive up their idaees without liarsli comphiint and
pitcoas ;;roans."*
Dr. John A. Veatch, in his account of the (!erros or Cedros
Island, situated otVtlu^ coast of Lower California (between the
paiidlels of -ISO anti\ ity these animals appear to become stronjjfly attached
to each otlier, so much so that in case one of a pair dies, the
other is very apt to die soon after, of {j^rief. They have also
been known to i)voi)a}iate in continement, an instance of which is
related by ]Mr. F.J. Thomi)Son in his interestiuj^ paper recently
jmblishcd in "Forest and Stream," on "Tlu* Habits and Breed-
iiij; of the Sea Lions in (Japtivity,''t based on observations
made at the Zooloji^ical Garden in Cincinnati, Ohio. As his
pa]»er contains, besides a general account of the habits of these
iiiterestin}>- animals, several novel points, I give it place iu
full :
"In the early part of June, 1877, I went, sent by the Zoolog-
ical Society of Cincinnati, to Chicago to receive some black
sealions {ZaJophm yiUespiei) which had arrived there from the
southern coast of California. On my arrival I found that the
female had calved on the jjrevious night, therefore thought it
Inist to lie over for a day in order that the yoiing might acquire
a little extra strength to bear the fatigue of the railway journey
to Cincinnati. They all arrived iu the garden in fine condition,
but had to be kept in their shipping crates for the first few days,
until an old beaver pond could be arranged as temporary quar-
ters for them, while the large basin intended for their perma-
nent home could be built. During this time, on account of
ii liea\y freshet in the Ohio River, the water iu the pond became
(luite muddy, whi(!h attected them so nmch that they were un-
alilc to retain their food, invariably vomiting up their fish some
one or two hours after feeding. By giving small doses of Ro-
ehelle salts for a few days, all recovered, but the calf «lied from
a violent attack of cholera infantum, caused no doubt by its
mother's milk being affected by the nviddy water.
"A sliort time before th(^ calf was taken sick my attention
was attra(^ted to the pe(;uliar appearance of the mother on
t'meri.ii|o from the water after taking her customary bath.
She was <'on'pletely covered with a whitish oleaginous sub-
stance, about the consistency of semi-fluid lard, which seemed
} ' « I
" Land and Water, Feb. 5, 1876, p. 104.
t Forest and Stream (uowspaper), vol. xii, p. 66, Fob. 23, 1879.
m^
308 ZALOPIIUS CALIFORNIANUS CALIFORNIAN SEA LION.
to ooze out all over her. As soon as she got into the crate
with the young one, she eonnnenced rolling-, so that in a sliort
time the young one and the inside of the crate were eoniph'tcly
covered with it. The (.'alf seemed to enjoy it hugely, and rolled
about until his coat glistened as if he had just left the hands ot
a first-class tonsorial artist. It instantly struck me that hi.s
mother had been i)re])aring him for the water, and I inninMli-
ately tested the matter by taking him out ard i)lacing him on
the edge of the pond, when in a few monu-nts he began to i)a(l-
die about in the water, something he had never before attemi)te(l
altl tough he had been almost daily placed in the same posi-
tion.
"As soon as the large basin was completed, and they were
transferred to it, I had a fine opportunity of observing the
tyrannical attentions of the male toward the female r('veiit
her egress. Tie never 8e<'nietlu> him by swimmiiijj; up and jiivinj? him a goo«l old-fiwh-
ioiM'd conjuf'al kiss. Finally they quieted down to the hum-
(liniii of regular wedded life, and early in October I noticed
tliiit the female was suffering from a violent catarrh, which
(jiiidually disappeared, followed by a dry cough, particularly at
nifilit. It was in INIarch when I first thought she showed signs
of i>regnancy, and in May, from her appearance when out of
the water, I became convinced of it. On Juiu^ 25 the young
OHO was born, nuiking the period of gestation as nearly as I
could judge about ten months, and it was some days before the
nutther would allow me to handle it, and when I did succeed in
so doing it was always at the risk of getting a nip, as he was
certainly the most ill-tempered, snarling little brute with which
11 dry nurse could be vexed. I soon found out that there was
hut one way of handling him with impunity, and that was by
suddenly catching him just back of the tlippers and liU'o«l it in his way, and lie iiiinieofthe lij) suv-
ronndin;;' the house, I had the carpenter c«)nstnu't a sniall.
shallow, woodi'n tank inside tlu^ lar«;er one, with a slopinji' plat-
forni leadinji' into it. So soon as the door was opened connect-
ing with it he followed his mother, and in a short time was
liaving high Jinks swimming anTieved so over the loss of his mate that for
8onit' time T was afraid we would lose him also, and at the end
ofabour six weeks he became so thin that 1 thouj'ht it best to
remove him to a small tank in-doors. Since, he has been improv-
iaS' slowly \\\) to within ten days, since when he shows a marked
iiiiinovement."
The Otaries, wherever occurring, ai^pear to closely aj^ree in
their habits, especially durin}^ the breeding season. As au
interesting supi)lement to the history of the two Northern spe-
cies of Sea Lions already given J transcribe the following
concise account of the great Southern Sea Lion {Otaria ju-
bnta), based on recent observations made at the I'^dkland
Islands, without, however, endorsing the author's "ballasting"
theory :
"The Sea Lion attains its full growth at nine years, and
annually <'omes back to the place it was born to breed and shed
its hair. The former o])eration occurs between the 25th of Decem-
l)er and the 15th of January, the latter in April aiiviMay. The
Lions connnence to arrive at their 'rookery' in November to
wait for the females^ who do not haul up until within two or
three days of pui)ping. They are fatter at this time than at any
other, and have to take in a quantity of ballast to keep them
down, without which they could not dive to catch tish. 1 have
opened them at this time, and found, in a pouch they have in-
side, ui)wards of twenty-five pounds of stones, some as large as a
goose-egg. As they get thin they have the power of throwing
these stones up, retaining only a sufiicient quantity to keep
them from coming up too freely to the surface.
"They are very savage in the breeding-season, and are con-
tinually lighting, biting large pieces out of esieh other's hide,
and sometimes killing the females. At this time they become
an easy prey to man, as they will stand and be killed without
trying to get away.
"The Lioness has her first pup at ttiree years of age, never
more than one at a time, and comes up to have intercourse with
iiilll
I't 1
li I *i
ti !,
i
312
GENUS CALLDUIIINUS.
the Lion at two, and as soimi as tho pup is born Tlioy
suckle their yoniij^ five niontlis before they are taken to tlic
sea, by which time the pu]* lias shed its first liair. Ilef'on- tlic
mother takes her pup to lish she has to ballast it, and I have
seen a Lioness tryinj;' for hours to make her pup swallow small
stones at the water's edye.
"The female keeps her pu]) with her until two or three weeks
before the next breedinji-season, when she drives it from licr.
About this time the yearlin{j[8 will be found some few miles
from the old rookery. . . .
" The Lions stay as lon^' as two months on shore, dr.riiiff the
breeding-season, without going into the water. During that
time their fat gives them suttieient nourishment. After the
season is over some of them are so thin and weak that they
are but Just able to crawl into the water. I have killed them iu
this state, and not one particle of stone have I found in them."*
Genus CALLOKHINUS, Gray.
CallorhinuM, Guay, Proc. Zoiil. Soc. Loud., 1859, 359. Type ''Arctocephahs
iirainiix, Gniy," —I'hoca umiia, Linii*^.
Arctocephalm, Gill, Proc. Essex Institute, v, 18(56, 7, 11. Typo " Vhoca
umina, Liimu'UH." Not Arciocephalm, F. CuviiT, IHiM.
Molars ~, small. Facial portion of skull short, broad, con-
vex, and but slightly depressed ; nasals short, rapidly narrow-
ing posteriorly. Palatal surface short, narrowed behind, with
the hinder border rather deeply concave. Toe-flaps very loiij;,—
nearly as long as the rest of the foot.
CaUorhinuSy in coloration, character of the pelage, size, ;;eu-
eral form, and dental formula, is rather closely allied to Arcto-
cephalus, from which, however, it is readily distinguished by the
form of the facial portion of the skull, which in Arctocephalus is
narrower, longer, and much less convex, with much longer na-
sals. From the other genera of the Otaries it is distinguish-
able not only by coloration and the character of the pelago, but
by its weaker dentition, and the strongly marked cranial ., 1839, 253, pi. xxi. — Nilsson,
Arch. f. Naturg., 1841, 331 (in part only).— MOller, Arch. f. Naturg.,
1841, 333.— Wagner, Schreber'sSiingt., vii, 1846, 65 (in part only) ;
Arch. f. Naturg., 1849, 39. — Von Sciirenck, Amur-Lanrte, i, 1859,
189.
Phoca {Otaria) ursina, RicnARDSON, Zoiil. Beechoy's Voy., 1839, 6.
Oiaria {VuUorhinus) ursinua, Peters, Mouatsb. Akad. Berlin, 1866, 373, 672.
Arctoccphahia ur8inu8, Lesson, Man. de Mam., 1827, 203. — Gray, Cat. Seals,
1850, 41 (not of F. Cuvier, or only in part) ; Proc. Zool. Soc. Lond.,
1859, 103, 107, pi. Ixxiii (skull). — Gill, Proc. Essex Institute, v,
18(i6, 13.— Scott, Mam. Recent and Extinct, 1873, 8.— Clark, Proc.
Zoiil. Soc. Loud., 1878, 271, pi. xx (colored figures of male, female,
and young).
Callorhiniw iirdnus, Gkay, Proc. Zoiil. Soc. Lond., 1859, ;}59, pi. Iviii (skull) ;
Cat. Seals and Whales, 1866, 44, fig. 16 (skull) ; Ann. and Mug. Nat.
Hist., 3d ser., xviii, 1866, 234 ; Suppl. Cat. Seals and Whales, 1871, 15 ;
Hand-List Seals, etc., 1874, 32, pi. xix (skull). — Allen, Bull. Mus.
Couip. Zoiil., ii, 73, pU. ii, iii (skull, etc.).— Scammon, Marino
Mamni., 1874, 141, pi. xxi, figg. 1, 2, and ligg. 1-6 in text (animal). —
Elliott, Cond. Aftairs in Alaska, 1874, 123. •»
i r: l
314 CALLORIIINUH IIRSINUS NORTHERN FUR SEAL.
"1
Phma niijrii , Pai.i.as, Z(>o>f. R7.
Le Chut marhi, Kkaschennimkow, Hist. Kanitsch., i, I7t)4, :{l(i,
Oiirn marin, Huffox. Hist. Nat., Siippl., vi, 178'2, 'A'M), pi. xlvii (in part).
Ursine Sml, 1'ennant, Synop. Quad., 1771, :{44 (liasod mainly on 8tila(!k.
Tilt' It >;ion iii'ouml tlu' mouth is ycllowisli l)rown. The neck in
Irtiiit is yiayish-l»hu'k. Tluj axilla" arc pale yellowish-brown; a
8(»iin'\viiiit darker sliadt' of the same color extends posteriorly
aiitl inward toward the median line of the belly, uniting; on the
aiitfi'itir portion of the abdomen. The {jfreat-er part of the lower
sinriu't", liowevcr, is dusky brownish-jjray, the rest being bhu'k,
lint less intensely so than the back. SpecinuMis of equal Jige
vary much in coh)r, some specimens corresponding nearly with
tilt' aliove descriptuMi, while others are much darker. On the
head and sides of the neck a portion of the hairs are foinnl on
ilitsc inspection to be obscurely tipped with gray. After the
lirst moult the pelage beccmies gra«lually lighter, through the
cxtcnsio' jf the gray at the tips of the hairs, especially in the
lliiialcs, the twt) s«>xes being at first alike. Contrary to what
lias been asserttul, the young are provided from birth with a
li)n;i coat of silky under-fur, of a lighter color and sparser than
tlic lunler-fur of the adults.
PklaCtE. — The pelage in this species consists of an outer cov-
ering of long, tiattened, moderately cojirae hair, l>eueath which
is a dense coat of long tine silky fur, which reaches on most
parts of the body nearly to the ends of the hairs. The hairs are
tlii('k«ir toward the end than at the base, but their clavate form
is most distinctly seen in the first pelage of the young. In
Ii'ngtli tlie hair varies greatly on the ditterent parts of the bcwly.
It is longest on the top of the head, especial' in the males,
which have a well-marked crest. The hair is nuich longer on
thti anterior half of the body tlmn on the posterior half, it being
longest on the hinder part of the neck, where in the males it is
very coarse. On the crown the hair has a length of 41i mm.; on
the hinder i)art of the neck it resiches a length of oO to 00 mm.
From this point posteriorly it gra^lually shortens, and in^ir the
tail has a length of only 20 mm. It is still shorter on the limbs,
the ui)per side of the digits of the hind limbs being but slightly
covered, while the anterior limbs are quite naked as far as the
carjiiis. The males have much longer hair than the females,
iu which it is much longer than in Eumetopias stelkrL*
' From tlic accountH j^iven by most writt-rs it wouhl seoin that (Hariajubata
is provided with a conspicuoiiB mane, but in the few accurate descriptions
in which tlie length of the longest hairs is stated, the so-called "flowing
mane"— which refers only to the greater length of the hair on the neck and
shoulders as compared with the other regions of the body — does not appear
31 n CALLORHIXrS rRSINL'S — XORTHKRN I'lTU SEAL.
The whiskers arc cyliiHlricul, I(Hl;,^ slender, and tapcrinn, and
vary with a^'c in h-nj^th and i-olor. In the .voun;;' (hey are l)lack;
hiter they ai'e h^ht colored at (he base, and -T ." n
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320 CALLOKIIINUS URSINUS — NORTHEKN FUR SEAL.
Ears. — The ears are long, narrow, and pointed, being abso-
lutely longer than those of the E. Hlelleri, though the latter
animal is two or three times the larger.
l^'OBE Limbs. — The handvS are very long and narrow, with a
broad cartilaginous flap extending beyond the digits, which
has a nearly even border. Both surfaces are naked the whole
length ; not covered above with short hair, as in EumetopinH and
Otaria. The nails are rudimentary, their position being indi-
cated by small circular horny disks, as in all the other Eared
Seals.
Hind Limbs. — The feet are very long, nearly half their length
being formed by the cartilaginous flai>s that project beyond the
ends of the toes. They widen much less from tlie tarsus to the
ends of the toes than these i)art8 do in iJ. steUeri, and the 1 ligth
of the toe-flaps is relatively many times greater than in the
latter species. The toes of the posterior extremities are of
nearly equal length. The outer are slightly shorter than the
three middle ones. The nails of the outer toes are rudiment-
ary and scarcely visible ; those of the middle toes are strong
and well developed.
Skull. — In adult specimens the breadth of the skull is a little
more than half its length, the point of greatest breadth Ix'in},' at
the posterior end of the zygomatic arch. The muzzle or facial por-
tion is broad and high, or greatly prmluced, much more so even
than In EumetopidH,* Tlie postorbital procasses vary from mh-
quadrate to sub-triangular, sometimes produced posteriorly into
a latero-posteriorly diverging point, as in ZaJophvH. Tlu^ post-
orbital cylinder is broad and moderately elongate [i!)i({
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324 CALLORHINUS URSINU8 NORTHERN FUR SEAL.
Teeth. — The dentition is relatively much weaker than in
either Eumetopias or Zalophus, or even in Aretocephalu-s. As
usual in the Otaries the outer pair of upper incisors is imioh
larger than the others and caniniform ; the two central pairs
are flattened antero-posteriorly, and in youth and middle a<;e
their crowus are deeply divided by a transverse groove. The
lower incisors are smaller than the upper and are hollowed ou
their inner face but are not grooved. The canines are larj;e
and shar])ly pointed, the lower somewhat curved. The molars
are small and closely approximated, with sharply conical crowns
and all siugle-rooted. They have no accessory cusp, or only
very minute ones in early life. The roots are usually grooved
both externally and internally, sometimes slightly so, but some-
times so deeply that the fang seems to consist of two connate
roots. The distinctness of these grooves varies not only in dif-
ferent individuals, but in the corresponding teeth of the two
sides of the mouth in the same skull,* so that it is not improb-
able that teeth may be found in which the grooves of the fangs
may be entirely obsolete, or so deep as to nearly or quite divide
the fang into two distinct roots. The roots of the molars are
very short, and but partly fill their alveoli ; hence when the
periosteum is removed they fit so loosely that they require to be
cemented in to prevent their constantly falling out whenever
the skull is handled. The canines and the incisors have much
longer roots, which more nearly fill their sockets.
Skeleton. — Vertebral formula: Cervical vertebrae, 7 ; dorsal,
15 ; lumbar, 5 ; sacral, 3 ; caudal, 8 to 10.
The skeleton in its general features resembles that of Eime'
topias Htelleri, already described. The bones of C. ursinns are,
however, all slenderer, or smaller in proportion to their lengta,
than in that species, the general form of the body being more
elongated. The scapulai are shorter and broader than in E.
stelleri, the i)roportion of breadth to length being in the one as
11 to 10 and in the other as 13 to 10. The pelvis is more con-
tracted! opposite the acetabula in C. ursinus than in E. delleri,
and the last segment of the sternum is also longer and nar-
rower. The diiferen(;es in the skull of the two forms have
already been pointed out in the generic comparisons. In pro-
portions, the principal difference, aside from that already men-
tioned as existing in the form of the scapula, consists in the
* See Bull. Mub. Comp. Zool., vol. ii, pi. ii, figs. 6 b and 7 e.
SKELETON.
325
loiiyor neck and longer hind feet in C ursinm; the ratio of
the length of the cervical vertehiw to the whole length of the
skeleton being as 15 to 100 in J-J. xtelleri, and as 2i5 to 100 in G.
ur.slniis; and the ratio of the length of the foot to the tibia
being in the former as l.'J to 10, and in the other as 10 to 10.
Tlie following nieasnrenients of two adnlt males and two adult
females indicate the length of the principal bones, and of the
different vertebral regions.
MeasurementH of the Skeleton.
Adult d
No. 2922.
■Vriiolt> longth of skeleton (including skuU) . .
Lcnjith of skull
1-criirtli of cervical vcrtebrsa
Lciict h of (loraal v('rtc#86
Lontstli of lumbar vertebral
I.cngtli of siicral vertebra>
Lcujitli of caudal vertebra)
Lenglli of first rib
Li'ii(;tli of first rib, osseous portion
I.ciifjtli of first ril), cartilaginous portion
Lin^tli of tbivrt ril>
Linjitli of third rib, osseous portion
[ Lengtli of third rib, cartilaginous portion
Li'n;;th 'ro\vii females, aecordiny' to Captain Bryant, being
If.ss tlioii ( (NK-siXTii that of the fiill-j,'ro\vu males. This estimate
Mr. Mlliott has siiutc found to be correct by actual weight of
liiiji'e series of specimens.
DirPERKNCES liESULTiNO KTIOM AGE.— The dift'erenccs in
color between the young and the adult consist, as already stated,
in tlic young of both sexes during the lirst three or four months
of their li\es being glossy black, and gradually afterwards
a('i|iiiriiig the coh)r characteristic respectively of the adult
iiiaU's and females. In res^iect to the ditierences in the skeleton
tliiit (listinguish the young, I can speak only of the skull, in
wliicli the relative de^■elol)ment of its different regions differs
\vi(h'ly fiom what is acau in the adult of either sex. The two
young skulls before me, said to be from si)ecimens thu'tyflve
(lays old, are both females, but at this age the sexes probably
do not differ in osteological features, especially in those of the
skull. In these young specimens the anterior or facial portion
of tlie skull is but little developed in com])arison with the size
of tlie Itrain-case. The nnizzle is not only excessively short,
but the orbital space is snudl, and the postorbital cylinder
is covrelatively I'educed almost to zero, the postorbital processes
bt'iiig close to the brain-case. The zygomatic arch is hence
very short ; the zygomatic foramen is as broad as long, instead
of being nearly twice as long as broad, as in the adult. On the
other hand, the brain-case is exceedingly large, the greatest
breadth of the skull being at the middle of the brain-case in-
stead of at the posterior end of the zygomatic arch. As will
be seen by the table of measurements of the skull already
given, the brain-case in nearly as large as in the adults, and the
bones being thinner, it must have a capacity as great as that
of the skulls of the adult males and females, there being, in
respect to this point, but slight difference in the sexes. As the
young advance in age, the anterior i)ortion of the skull, or that
part in advance of the brain-case, greatly elongates, especially
tlic ])ostorbital cylinder, and increases also in breadth, the
f^kaU in a great rn^asure losing the triangular form and the
narrow pointed, muzzle characteristic of the young. The post-
orbital processes also greatly change their form as they further
develop.
i:|«T:
*1 M
328 CALLORHINUS URSINUS — NORTHERN FUR SEAL.
The limbs Jire also rdatively much larger thitii in the adult, as
meutioutMl by Quoy and Gaimard in respeet to the Arctocephalus
cincrcHH of Australia,* which enables them to move on land with
greater facility than the adult, as the above-mentioned authors
ha\ft stated to be th«' ease in the Australian species.
It is not tnu', hoAvever, that tlie young of C ursinus are devoid
of under-fur, as some writers have alHrmed.t
Individual Variation. — The two males were both not only
full-grown, but (piite advanced in age, though in all i)robability
the crests of even the older skull (No. 2\)'22) would have been still
further develoi)ed. The other male (No. 21)23) was somewhat
younger, but alreat!v had the sagittal crest considerably pro-
duced; the teeth, however, were but moderately woi'u, the mid-
dle upper incisors still re+ainiug the groove dividing the surface
of the crowns. In the younger male skull the posterior outline
of the palatines is but slightly concave, whereas in the other it
is deeply and abruptly emarginato of these specimens of Fur Seals, Captsiin Bryant lias
I'l'spoiidt'd to my iiuiuiries as follows: "The grown females (the mothers of
till' jtiiiis) were average specimens. The only means I had t>f determining
tht'ii age was by tho evidences atforded by disscictiou. These were that
tlio older t'emalo had given birth to seven young and the other five, which
^vould nial. ■ their ages respectively ten and eight years. The two grown
iiialis wen Jilso selected as average specimens in size and color. Judging
U'om their general appearance and color, I estimated them to be ten years
old. The V o pups were thirty-tivo days old, and in that time had doubled
their size fr m birth. They were both females."
..M
■MMm
i'i
If' 1^
m\
(rs
330 ('AI.LOlMII.sUS rWSINUS -NOUTIIKHN FUR SEAL.
tln', toe-Haps of the jtosterior limbs, which in (!. urninm are
{jivatl.v developed, their extension beyoml the dij^its being
neiiily eijual to th(^ h'n^tii of the ro8t of the foot — in adult
• females \\{) mm. beyond th«! toes. In Arvloccitlndns aiinfrolis
( = /((Ihlaniliciis^ anet.), on the other band, they extend, as in
most of the other Otaries, only a sh()rt lA «4 «4
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II
332 CALLORIIINUS URSINUS — NORTHERN FUR SEAL.
Geographical Distribution and Migration.— The Fur
Seal is well known to luivo been formerly abundant on the
western coast of North America, as far south as California, but
the exact southern limit of their range I have been unable to
determine. Captain Scammon speaks of having seen them " ou
one of the San Benito Islands, on the coast of Lower Califor-
nia," and again says, " On the coast of California many beaches
were found fronting gullies, where [Fur] Seals in large numbers
formerly gathered ; and, as they there had plenty of ground
to retreat upon, the sealers sometimes drove tliem far enough
back to make sure of the whole herd, or that portion of them
the skins of which were desirable."* He also states that the
"Fur Seal and Sea El«;phant once made the shores [of Guadalupe
Island] a favorite resorting-place," and refers to their former
occurrence on Cedros Island, in latitude 28°.t Although at
one time abundant ou the California coast, they are by no means
numerous there now, having been nearly exterminated by un
restricted destruction by the sealers. The writer above cited
refers also to their capture by the Indians at the mouth of the
Strait of Juan de Fuca. The Seals appear here and ou the
neighboring coast, he adds, ^' some years as early as the lirstof
March, and more or less remain till July or August ; but tuey
are most plentiful in April and ]May. During these two months
tlie Indians devote nearly all their time to sealing wheu the
weather will permit." He reports their increase there in later
years, and that while only a few dozens were annually takeu
there from 1843 to 180-1, fully five thousand were taken in 1809,;
Captain Bryant has given a similar report, referring especially
to their abundance along the coasts of Oregon, Washington
Territory^ and British Columbia in 1869, as comi>ared with for
mer years. He says those taken "were mostly very yonug
Seals, none appearing to be over a year old. Formerly in
March and Ai)ril the nati vea of Puget Sound took large mini-
bers of pregnant females,§ but no places where they have
resorted to breed seem to be known oti' this coast." He thiiiks
it probable, however, that they may occupy rocky ledges off
* Marine Mamin., pp. loS, 154.
+ J. Ross Browne's Resources ol the Pacific Slope, second part, p. l^-
t Marine Mamm., p. 154.
$ There arc six skulls in the National Museum from Pi;get Sound and the
neighboring coast (collected at several different points by Messrs. Scammon
an Swan), all of which anJ'emaUs.
GEOGRAPHICAL DISTRIBUTION.
33a
shore which are rarely visited by boats.* In his MS. report just
receiveier respecting the abundance of the Southern Fur Seal at the Isluul nf
Juan Fernandez, two hundred years ago, or ahotit a century heloif tlw
beginning of the Seal slaughter there, which in less thi'u a generation iicaily
exterminated the si)ecies at that locality. Danipier and liis party siicm
fifteen days on this ialainl in the year U]8',i. He sa., s: '^ Sealn swnriii as
thick about this Island ["of .John Fernando," as he terms it, J as if tlity had
no other place m lUt World to livi' in ; for there is not a Bay nor Rock tliat
one can get ashotir on, but is full of them. . . . These at John Fenm-
do's have line thick short Furr ; the like I have not taken notice of any where
but in these Se.as. Here are always thousands, I miglu say possibly iiiillious
of them, either sitting on the Bays, or going and coming in the Sea round tbo
Island, which is covered with them (as they lie at the top of the Water phiy-
ing and sunning themselves) for a mile or two from the shore. When tlii'v
come out of the Sea thoy bleat like Sheep for their young; and though tlie.v
pa«8 through hundn'ds of others yoting ones before they come to their own,
yet they will not suffer any of them to suck. The* young ones are Hkf
Pni)pie,^, and lie much ashore, but when beaten by any of n.s, they, as well
as the old ones, will make towards the Sea, and swim very swift and
nimble ; tho' on shore they lie very sluggishly, and will not go out of om
way unless we beat them, but snaji at us. A blow on tlie Nose soon kills
them. Large Ships might here load themselves with Seals Skins and Traue-
oyl; for they are extraordinary fat."— J Sew Voyage Hound the iVorld, "MA
edition, corrected," 1703, vol. i, pi>. 88,90.
t Cond. of Aff. in Alaska, pj). '217, 22i.
{ Alaska and its Resources, p. 41>;{.
GENEK'AL HISTORY.
335
ai)iK'iirL'd there, however, only in si)rinj^ and in Sei>teniber, none
being seen therefrom the bej;inning of .June till the end of
August, at wliieh time he .says they reiurti from the south with
tlieir young. Von Schrenek speaks of their oecurrenee in tlie
Ocliotsk Sea and the Tartarian (Inlf as far south as the 40th
degree of hititude, or to the soutlicrn ])ointof Sughalien Island.
The natives reported to him the ojjcurrenee of great numbers
of the animals on tlie eastern <;oast of that island. Captain
Scaiinnon also refers to their abundance twenty years since ou
the eastern side of Saghalieu.*
Excei)t during the season of reproduction, these animals ap-
pear to lead a wandering life, but the extent and direction of
their migrations are not yet well known. Steller spoke of their
migrations as being as regular as those of the various kinds of
sen-fowl, and they are recorded as arriving with great regularity
at the Prybilov Islands, but where they pass the season of
winter is still a nuitter of conjecture.
(iRNERAL History. — The Northern Fur Seal was first nuide
known to science by Steller in 17.51, under the name Ursus
mirinKs. During his visit to KamtschaJ^ka and its neighboring
isliuids, in 1742, he met with these animals ir. great numbers
*C;i]it;iiii Scainiiioii rcl.atoM in jiii oU'-liand way, iiicn'ly us tin ii'.rori'stiiig
iutuliiit (it sealing lilt', the lollowin;? : "In thi! midst of the crimtian War,
an I'litci'iirising iirni in New London, Connecticnt, fitted out iiflip])er bark,
wliich \va8 otticcrt'd and manned cxjtrcHsly for a sealing voyage in the
Okhotsk Sea. 'J'lie cai»tain was a veteran in the business, and many thought
him too old to eomniaud, but tlie result of the voyage proved him equal to
the tu«k. The vessel proeeedtul to Robin Island — a mere volcanic rock, situ-
ated on the eastern side of the; large island of Saghalien. Many outlying
rocks and reefs are about it, making it dangerous t»» approach, and affording
l)iit slijrht shelter for an anchorage. Here the vessel (of about tlireo hun-
ilml tons) lay, with ground-tackle of the weight for a craft of twice her
size. Much of the time fresh winds jirevailed, accompanied by the usual
ugly giniind-swell ; auil, iu eonsiMpu'nce of her b«ung long, low, and sharp,
tliPddk was at such times fre upon the former distribution and abundance of this and other
iieciesof Seal, and that he has not given mor<^ explicit references to the
loealitics at which the "ur Seals were formerly hunted on the southern por-
"on of the North American coast and elsewhere.
"Mill*
irivi ''i
i i\
fihip
I.*
:! 5' '.
336 CALLOKHINrs URSINUS NORTHERN PUR SEAL.
at Belli ing's Island, where he spent HOine time among them, and
carefully Htudied their habits and anatomy, a detailed account of
which appeared in his celebrated inemoir entitled " ])e Bcstiis
Mariuis," in the Transactions of the St. Petersburg Aj-adciuy
for the year Hi*.).* This important essay was the source of
nearly all of the accounts of this animal that appeared prior td
the bej^inning of the present decade. The tw^enty-eight <|narto
pages of Steller's nu'moir devoted to this s])ecies, gave not only
a detailed account of its anatomy, with an extensive table ot
measurements, but also of its remarkable habits, and tigurcs of
the animals themselves. A little later Krascheninikow, in Lis
History of Kanitschatka,t uiuler the nan»e " Sea Cat," ga\e also
a long account of its habits, apparently based mainly on Stel
ler's notes.f but it embraces a few particulars not given in "De
Bestiis Marinis." Steller's description of the habits of this
animal has been largely quoted by Button, Pennant, Schrel)ei,
Hamilton, and other general writers.
Button, Pennant, Schreber, Gmelin, and nearly all writers .11
the Pinnipeds, down to about 1820, confounded the Nortbeiu
Fur Seal with the Fur Seals of the Southern hemisphere, blend
ing their history as that of a single species. P^ron, in 181(5, first
recognized it as distinct from its southern allies, and it was so
treated somewhat later by Demarest, Lesson, Fischer, Gray, and
other systematic writers, § but its distinctive characters were not
*^Nov. Conim. Acad. Petrop., ii, pp. 331-359, pi. xv, 17.')1. Thiw, as is will
known, is a posthiimoiis pa])er, published six years sitter Steller's dfatli.
Steller dying of fever November 12, 1745, wbilii on his way from Siberia to
St. Petersburg. The description of the Sea Bear was written at Hcliniiu;'
Island in May, 1742.
tH'^t. Kamtchatka (English edition), trnnshited from the Russian by
James Grieve, pp. 12:M30, 1764.
t Krascheninikow, it is stated, "received all of Mr. Steller's pai)ers,"lo
aid him in the preparation of his " History of Kanitschatka."
^Nilssoii and Miiller in 1841, and Wagner in 184() and IHIS), on the otber
hand, still considered all the Sea Bears as belonging to a single spetii's.
Wagner, in 184'.) (Arch, fiir Natnrg., 1849, pp. 37-49), described the ostw-
logical characters of the Northern species from three skeletons in tlif
Munich Museum received from Hehring's Sea. One of these was ait)>arcutl,v
that of a full-grown female ; a second was believed to be that of a liall'-jti'owu
male, while the third belonged to a very young animal, in which the jht-
manent teeth were still not wholly developed. Wagner compares tiic speeif^
with Stelha's Sea Liy this author, in the paper in whicli the name Callorhinus was
liro])()se(l, as that of his Arctocephalus monteriemw, which is a
tirkii^f
'f 'IS
t \ !
^< \
m
i i :
n i
,! %i
*? -j,
* Proc. Zool. Soc. Lond., 1859, p. 102.
+ iiti■>
c>>^^r-
Hair Seal. This nkiii was accoinpaiiied by a young- skull, pur
poitiuft-, by the label it bore, to belong; to it, but Dr. Gray
observes that otherwise he should have thoujjht it too small to
have beloii}>:e(l to the same animal. Seven years later,* lie do
scribed the skull as that of ii new species {ArctocephaliLs mli
J'onn'ainiN), still associating with it, however, the skin of tlif
CtiUorhiuus ursinus. The skull ho subse(piently considcicd as
that of a young A . monierunsis ( = Eumctop'uu nicUvH) ; and refer-
ring his ^l. imliforuianm to that s])ecies, he was conseqiionth
led into tlu' double error of regarding the Unmctopias sleUeri as
a Fur Seal (as already explained under that species and olso
where in the ])resent paju'r), an
biology, f he «lescribiug at considerable length, from personal
observation, its habits, distribution, and ])roducts, as well a>
the various methods employed for its capture. The followini;
year Mr. W. IT. Dall devoted a few pages § to its history, in
which he made many important suggestions relative to tin.
sealing business. During the same year I was able to add nut
only something to its technical history, || but also to make puli
lie an important communication on its habits kindly placed at
my disposal by Captain Charles Bryant,^} Government agent in
charge of the Fur Seal Islands of Alaska. In 1874, Ca]ttaiii
Scammon republished his above-mentioned paper,** addiiis
thereto a transcript of Captain Bryant's observations already
noted. Almost simultaneously with this appeared Mr. H. AV,
Elliott's exhaustive Ileport on the, Seal Islands of Alaska,ttin
"Cat. Seals and Whales, 18()(i, j). ol.
t Suppl. Cat, Seals aii«l Whales, p. 15 ; Hand-List of Seals, 32.
t Overland Monthly, vol. iii, Nov. 1869, pp. 393-399.
ij Alaska and its Resources, 1870, pp. 492-498.
II Bull. Mus. Comp. ZoiJl., ii, pp. 73-89.
IT Ibid., pp. 89-108.
** Marine Mammalia, 1874, pp. 141-103.
- tt Report on the Pry bilov Group, or Seal Islands of Alaska, 4to, uupag'Hl.
1873 [1874].
FIOIIKKS.
339
wliicli the present species juoperly comes in lor si huge sliarc
of tlic iuitlioi's attention. Tlie work is richly illustrated with
|)luitoj;rapliic plates, taken from Mr. Elliott's skcti(!lies, about
twi'iity-live of which are chnoted to the; Fui' Seal. The text of
this viir*' and ])rivately distributed work has been sincte re-
printed,* with sonic changes and adhasc of the subject is treated at length. A few
iiiiiHtr notices ol" lhiss])ecies have since ajjpeared (mostly j)opu-
liir articles in illustrates, chiefly from the pen of Mr.
Klli(ttt), but nothing relating to its general history requiring
spi'c- 1 notice in the ])resent connection.
i''i(iri{ES. — The lirst figures of the Northern Sea Bear were
uivcii by Stcllcr, in liis i)aper already cited. They represent an
luhilt male, in a <|uite natural attitude, and a female reclining
nil licr back. In respect t< details, these early ligures were
naturally more or less rude and inaccurate. They were copied,
liowover, by iJutfon, Schreber, Pennant, and othei- early writers,
:iii(l are tlu^ only rejursentations of this species known to me
that wen; made ])i'ior to about the year 1831), except C/horis's
phitiMtf a group of these animals entitled "Ours marins dans
\"\\v (le St. Paul",t published in 1822. This represents three
ohl males, surrounded by their har«ms, and indicates very faitli-
tully tlie moch^ of grouping aiul the variety of attitudes as-
sniiied l»y these animals when assembled on the rookeries.
Ilaiiiiltoii, in 1839, gave a figure of the "Sea Bear of Steller
(OUir'td »>-,s/«rt)''| which he tells us is "from the engraving of
thi' distinguished Naturalist of the Euriek*,§ the original of
which I ha\e not seen. This represents a nmle and female, the
latter reclining on its side, with a pup resting on its right
Hipper.
The lirst llgure of the skull is that published by Gray in
i8,*,->,ll— a vioAv in profde of the skull of an adult male. A
wood-cut of the same was given in 1806,^ and a fine lithographic
"Condition of Afl'airs in Alaska, 1875, pp. 107-151.
t Yoy. pittor. uiitour dii Monde, lies Aldoutienues, pi. xv.
t Murine Ainphibiie, pi. xxi.
\Ubid.,p,2G6.
II Proc. Zoiil. See. Lond., 1859, pi. Ixviii.
1i Cat. Seals aud Whales, p. »5, fig. 16.
f;i
. Ji
340 CALLORIIIXUS ntSlXUS XOUTJIEHN FUR SEAL.
plate ill 1.S74,* ropn'seiitiiig tlit* .skull in piofile, liom above
iiiid from iH'low.t
In 1S7() I fiiive liju'iircs of t\vi» adult niak' skulls (two views
of eacli), of an adult female skull (three views), of a very youii^f
skull (tliree views), and of the scapula, dentition, ete. These,
so far as known to me, are the only tiijures of the skull orotiier
details of structure thus far published.
In 1874 Captain Seamnion j;ave fij>ures of the animal, f a /inc
ograph of an old male,§ from a sketch by Mr. Elliott, a woodcut
of the head of a female seen from beh)W (drawn by Elliott), || two
outline tigures representinjf the female as seen from below and
in proflle, and two others in oiitUne illustrating "attitudes of
the Fur Seals." 1| Mr. Elliott, in his lirst Report on the Seal
Islands,** in a series of over two dozen large photograpliic
plates (from India ink sketches from nature) has given an ex
haustive presentation of the phases of Fur Seal life so faithfully
studied by him at St. Paul's Isl.and. Among these may he
mentioned especially those entitled "The East Landing and
Black Buttes — The beach covered with young Fur Seals";
"The North Shore of St. Paul's Island" (giving an extensive
view of the rookeries) ; " Lukannon Bear'i " (Fur Seals playin;;
in the surf, and rookeries in the distance); "Old male Fur Seal,
or ' Seecatch ' " (as he appears at the end of the season after
three months of fasting) ; "Fur Seal Harem " (showing the rela
five size of nmles, females, and young, various atatudes, posi-
tions, etc.); 'Fur Seal males, waiting for their 'harems'" (the
females beginning to arrive); Fur Seal "llookery" (breeding-
grounds at Polavina Point) ; " Fur Seal Harem" (Reef Rookeiy,
foreground showing relative size of males and females); "Fur
Seal Pups at Sleep and Play"; "Hauling Grounds" (several
views at different points); "Capturing Fur Seals"; "Driving
Fur Seals"; "Killing Fur Seals — Sealing gang at work," etc.
The only other pictorial coutributions to the history of the
* Iluml-List of Seals, i)l. xix.
' 1 1 infer tins to bo tlio same specimen in each case, not only from there-
semblance tlio figures bear to ciich other, but from Ur. Gray, so lav as 1
can discover, referring to only tlic single skull from Beliring's Strait, re-
ceived in 1859.
t Marine Mammalia, \>\. xxi, two figures.
$ Ibid., p. 14:5.
||Ibid.,p. 145.
iribid., p. 14«J.
** Report on the Prybilov Group, or Fur Seal Itiiands, of Alaska, lUipageJ.
and jdates not numbered.
HAIUTS.
a4i
Fiir Seal of uotewoitiiy iuiitortancc is .Mr. Claik's coloicd ])lato,*
(III wliicli ai'c iei»i('S('iite(l a nearly fiill-^irowii male, a t'einule
and a |>u]>, i)r('i)ar('(l lioiii ^ikiiis .sent t(> the Jhitisli ]Mu.seum by
tlif Alaska Comiiu'rcial (J()ni]iaiiy. In these the attitudes are
cxct'llciit and the eolorin;-' lair.
llAiU'i'S. — The habits of the I-'iir Seal of the north .seem to
jiiive heen well known to Steller and hi.s companions a century
and a (piarter ajio, and their seemin;4ly marvellous accounts of
tlieni jirove to liave been only to sislifiht degree erroneous. As
a matter of historic interest, and for comi)arison with our pres-
ent kiiowii'dge of the subjely on tho.se of his fellow-trav-
('Ik'i', Stfciler. ''The Sea ('at ", says Krascheninikow, t '' is about
half the size of the Sea Lion; in form resembling the Seal, but
tldckor about the breast, and thinner towards the tail. They
have the snout longer than the Sea Lions, and hu'ger teeth ; with
eyes like cow's eyes, short ears, naked and black paws, and black
liair iiiix(;d with gray, which is short and brittle. Their young
are of a bluish black color.
" The Sea Cats are caught in the spring and in the month of
September, about the river Shcepanora ; at which time they go
from tlie Kxrllsloy Lsland to the American coast; but the most
are catched about the cape of Kro)iotzkoi/, as between this and
the cape Shi(2)lnfil:oy the sea is generally calm and affords them
pioperer places to retire to. Almost all the females that are
caught in the spring are pregnant; and such as are near their
time of bringing forth their young are innnediately opened and
the young taken out and skinned. Xone of them are to be seen
from the beginning of June to the end of August, when they
return from the .south with their young. The natives Avere
formerly at a loss to concei\'e where such great herds of preg-
uaut I'at animals retired in spring, and why they returned so
woalc and lean was owing to their fatigue.
.... ''The male and fenmle differ so much in the forin and
fu tlu' wiitcr. \\y \\m iiieiuis tlu! Ixmt is iVt'ijiUMitly over-
tni'iMMl iiinl the |»(!()|>1«' (h'owiu'd, unlt'.s.s liowho stcri's it bovciy
.skillliil, iiiid ()l*s«'rv»'s tlu' <'()urs(M)t't]u^ animal Tlicy
lastcn tln'ii' loiv paws in tlio rocks, und thus draw up tlicir body.
wldeli tlu>y can move but slowly in sucli places, but upon a
l»lain, one is in diinf^cr of beiny ov«iitakou by them, rjwu
Bifinifx Island there are such nund)ersot' them that they cover
.v^'-''^ . , the whole shore: so that truvellers are t'renuentiv oblij^ed tor
|v- r sal'etv to leave the sands jind level countrv and yo over tlu- hills
r
and rocky places. It is remarkable that in this island the Sen
Cats are found only upon the south coast which looks towards
Kamtsi'hntka. The reason for this may be, that this is the lirst
land they meet with goinji; east from tlu^ Krouotzhyy i)ass."*
Steller anntests with each other uncoii-
firmed by modern observers. In several respects the ac<'ounts
of these authors — in the main virtually identical — border uiwu
the mythical, but, generally speaking, are remarkably free from
exaggeration, considering the times at which they Avere writteu.
xVs already stated, they formed the source of all our knowl-
edge of these strange beasts prior to the beginning of the pres-
ent decade. Choris makes only very brief mention of them
and says very little about their habits, t Veniaminov, iu hi>
"Zapieska" published at Saint Petersburg, iuKussian, in 1840,
ami known to me only as quoted and translated by Mr. II. W,
Elliott,! has given valuable statistical information respcctiu;;
the sealing business as prosecuted by the Kussiaus at the Prv
bilo V Islands, but seems to have given no detailed account of their
habits. Our tirst important recent information respecting the
economy of these animals is that given by Captain Charles
* Krascheninikow'a Hist. Kamtschatka, Grieve's English translation, pi'.
12:5, 131.
t His .account iu full is as follows: " L'ours marius, on russt^ nivoutch, con-
vre par niillidrs les rivages des lies Kotoviya [Islands of 8aiut Paul ami
Saint Gcorgo], oil sont jetdes abondanimeut dcs plantcs marines (/«(•»«)•
On entend do tres-loin Ic cri do ces aniniaiix, lorsqu'ou est en nicr. h'-
fcnielles sont beauconp plus potites quo Ics milles ; elles ont Ic corps pln>
tluet et decouleur januiltre. Les mAles ont jusqu'st six pied do baut lois-
qu'ils Invent la tetc ; les jeunes sont ordinairenieut d'lm Ijrun noir ; il paraii
quo les fcni.clles no font jamais plus d'uu petit." — From the descriptiou ol
"Hcs S.Georgesct S.Paul," iu " Voy. luttorcsq. autour du Monde.''
* Condition of Aftairs in Alaska, pp. 241-24'i.
HAIUTS.
345
Ijryaiit, in 1870.* Mr. IClliott's iUM-oimt, puldisluMl tlii'0(! or
Ibiir vcais later, is tar mort' (Ictailrd, and ri\s|K'('tin<;- in(»st])oiuts
jiiiiy lie considered as fairly exhaustive of the sul>Jelaced
;ii iii\ disposal a coninninication embodying' the results of his
('i<:lit years" observations on these animals, i)rei)ared by my
r('(|iicst «'xi)ressly tor the present work. While replete with
new iiilbrmation, it (h)es not, to any };reat extent, dui)licate
t.;e account of the habits of the species ])ublished by Mr. Elliott,
l)einn ecies. I>elieving it desira-
ble to piesent in the present connectu)n a full and connected
history of the species, I offer no apology for the <;opious extracts I
t'loni Mr. Elliott's graphic account of the habits of the Fur Seal r
which here follow :
•'The fur seal {Callorhinus uriihiufi), which repairs to these
islaiuis to l)reed in nund)ers that seem almost fabulous, is by
far the highest organized of all the Pinnipedia, and, indeed, tin-
that matter, when land and water are fully taken into account,
there is no other aninml superior to it from a purely physical
point of view ; and few creatures that can be said to exhibit a
higher order of instinct, ai)proaching even intelligence, belong-
ing to the animal kingdom
"Observe it as it comes leisurely swimming on toward the
land ; how high above the water it carries its head, and how
deliberately it surveys the beach, after having stepped upon it ;
it may be truly said to step with its fore fli])pers, for they reg-
iilaily alternate as it moves up, carrying the head well above
them, at least three feet from the grouiul, with a perfectly erect
neck.
• ■ . . '"We observe as the seal moves along that, though it
hamlles its tore limbs in a most creditable manner, it brings up
its rear in (luite a different style ; for after every second step
ahead witli the fore feet it arches its spine, and with it drags
'* Bull. Mus. Coinp. Zool. il, pp. BO-lOt-.
346 CALL0RHINU3 URSINUS NORTHERN FUR SEAL.
and lift.s togetlier the hinder limbs to a fit position nnder its
body for another movement forward, by which the spine is
ajjain straij;htened (Mit so as to take a fresh hiteli up on the
posteriors. This is the U'isnrely and natural movement osition on the breeding-grounds
occupied by them during the preceding season. From my
knowledge of their action and habit, and from what I have
k'iU'iuMl of the ual^ives, I should say that very few, if any of
them, make such a selection and keep these places year after
year. One old bull was pointed out to me on the Eeef Gar-
hutch Ivookery as being known to the natives as a regular visitor
at, ch)so by, or on the same rock every season during the past
thi'oe years, but he tailed to re-appear on the fomth; but if
those, animals came each to a certain i^lace and occupied it reg-
iihiily, season after season, I think the natives here Avould know
it definitely; as it is, they do not. I think very likely, how-
ever, that the older bulls come back to the same rookery-ground
wlieie they spent the previous season, but take up their posi-
tions on it just as the circumstances attending their arrival will
permit, sucli as lighting other seals which have arrived before
tlici.., tS:c.
"With the object of testing this matter, the Russians, during
the early ^":trt of their possession, cut off the ears from a given
uuinbei" (»f young male seals driven up for that purpose from
one of the rookeries, and the result was that cropped seals were
found on nearly all the different rookeries or 'hauling-grouuds '
on the islands after. The same experiment was made by agems
two years ago, who had the left ears taken off from a hundred
young males which were found on Lukannon Eookery, Saint
Paul's Island ; of these the natives last year found two on !N"o-
vastosh-nah Eookery, ten miles north of Lukannon, and two
or tlnee from English Bay and Tolstoi Eookery, six miles west
l>y water; one or two were tak' n on Saint George's Island,
thirty-six miles to the southeast, and not one from Lukannon
was found among those that wore driven from there; and, prol>-
uhly, had all the young m.Jes on the two islands been driven
up and examined, the rest would have been found distributed
quite ecpially all around, although the natives say that they
tiiiuk the cutting off of iiio animal's ear gives the water such
access to its head as to cause its death ; this, however, I think
requires confirmation. These exp^riiu^nts would tend to prove
that when the seals approach tu islands in the spring they
ha\'e nothing but a geneial instinctive appreciation of the tituess
of the land as a whole, and no especial fondness for any partic
nlar q}ot.
'1(1
■'Si
348 CALLORIIINUS FRSINUS — NORTITEKN FUR SEAL,
'' Tlie landing of tbe soals upon tlio respective rookeries is
inliuen(,'e(l greatly by tiie direction of the Avind at the time of
ap]>roach to the islands. The prevailing winds, coming liom
the northeast, north, and northwest, carry far out to sea tlie
odor or scent (jf the pioneer bulls, which have located them-
selves on different breeding-grounds three or four weeks usu-
ally in advance of the masses; and hence it will be seen that
the rookeries on the south and soi;theastern shores of Saint
Paul's Island receive nearly all the seal-life, although there art'
miles of eligible ground on the north shore.
•'To settle this question, however, is an exceedingly difficult
matter ; for the identification of individuals, from one season
to another, among the hundreds of thousands, and even millious.
that come under the eye on a single one of these great rook-
eries, is really impossible.
"From the time of the first arrivals in May up to the 1st of
June, or as late as the middle of this month, if the weather be
clear, is *trength ; when on land, with the fore feet he does all climbing
*"'Si'c-i'atch,' native name for tho bulls ou the rookerios,c8pccially those
whith aie n'lle to maintain their position."
1^' t
IIH
k
1 ii
V
350 CALLOHHINUS UKSINUS — NORTHERN FUR SEAL.
over rocks, over the grassy hummocks back of the rookery, tlie
hiiul flippers being gathered up after every second step for-
ward, as described in the manner of walking; these fore feet
are the propelling power when in water, almost exclusively, the
hinder ones being used as rudders chiefly.
'' The covering to the body is composed of two coats, one be
ing of short, crisp, glistening over-hair, and the other a close,
soft, clastic pelage, or fur, whi(!h gives distinctive value to the
pelt.
"At this season of lirst 'hauling up' in the spring, the pre-
vailing color of the bulls, after they dry off" and have been ex
posed to the weather, is a dark, dull brown, with a spriiiklinjj
of lighter brown-black, and a number oi" hoary or frosted gray
coats ; on the shoulders the over-hair is either a gray or rulbus
ocher, called the 'wig;' these colors are most intense upon
the back of the head, neck, and spine, being lighter underiioatb,
The skin of the muzzle and flippers, a dark bluish black, fadin"
to a reddish and purplish tint in some. The ears and tail are
also similar in tint to fhii body, being in the case of the foriuei
a trifle lighter ; the ears on a bull fur-seal are from an incli to
an inch and a half in length ; the jyavil ions tightly rolled up on
themselves so that they are similar in shape and size to tlie lit-
tle linger on the human hand, cut off' at the second (phalaiiiical)
joint, a shade more cone-shaped, for they arc greater in diauie-
ter at the base than at the tip.
" I think it probable that the animal has and exerts the power
of compressing or dilating this scroll-like pavilion to its ear,
accordingly as it dives deep or rises in the water; and also, 1
am quite sure that the nair-seal has this control over the meatim
externus, from what I have seen of it; but I have not been able
to verily it in cither case by observation ; but such opj)ortniiity
as I have had, gives me untloubted proof of the greatest kceu
ness in hearing ; for it is iujpossiblc to approach one, even wheu
sound asleep ; if you make any noise, tmiuently no nifitter how
slight, t;;B alarm will be given instantly by the insignifioant-
looking auditors, and tlic aninud, rising uj) with a single luotiou
erect, gives you a stare of astonisliiuent, and at this season of
defiance, together with incessant surly roaring, grttwliug, and
' spitting.'
" This spitthig, as I call it, is by no means a fair or fi'.ll cxpres
sion of the most characteristic sound and action, peculiar, so
far as I have observed, to tlie fur-seals, the bulls in particuhU'
HABITS.
351
if
t
It is the usual prelude to their combats, aiul follows somewhat
in this way: when the two disputants are nearly within reach-
iii>;- or strikin*r distance, they make a number of feints or false
jjiisscs at one another, with the mouth wide open and liftinj;
the lii)s or snarling, so as to exhibit the glistening teeth, and
with each pass they expel the air. so violently through the
liiiyiix as to make a rapid choo-choo-choo sound, like the steam-
liutl's in the smoke-stack of a locomotive when it starts a heavy
train, and especially when the driving-wheels slip on tlic rail.
■•All the bulls now have the jmwer and fre(pient inclination
to titter four entirely distinct calls or notes — a hoarse, resonant
Kiai, loud and long; a low gurgling growl; a chuckling, sibi-
lant. pi]>ing whistle, of which it is imi)ossible to convey an ade-
(|tiatc idcii, for it must b(^ hetird to be understood; and this
spitting, jnst described. The cows* have but one note — a hol-
low. ])r()l<)nged, hia-a-ting call, addressed only to their pups ; on
all other occasions they are usually silent. It is something like
tlic cry of a calf or sheej). They also make a spitting sound,
and snort, when suddenly disturbed. The pups '/;/«- -
liickic," or bachelors. Tlie nami^ applied collectively to the fur-seal by
lliem is 'mor.skie-kot,' or sea-cat."
j.
n i
352 CAM.OKHIXUJS ritSIXrS XORTlIKlfX VVVv SEAL.
''Tlu' sound arising' from these {•reat breediiiy-fironiulsot'tlK
fur-seal, wIumc thousands ui>on thousaiuls of an}>iy, vi;iilaiit
hulls are roariny, ehuekliiij;', ])i]>iriy\ and multitudes of seal-
mothers are calling" in hollow, bla-atiu};' tones to their ycHiii;;,
which in turn resjmnd iiu-essantly, is simply inih'scribablc. It
is, at a sli'jht distance, softened into a deej) booming', as of a
cataract, and can be lieard a long distance off at sea, uiulei
favorable circumstances as far as live or six miles, and fre-
quently warns vessels that may be a])i»roaching the islands in
thick, foggy weather, of tlie positive, though unseen, proximity
of land. Night and day, throughout the season, the dhi of the
rookeries is steady and constant.
" The seals seem to suffer great inconvenience from a coni-
ptiratively low degree of heat ; for, with a temperature of 46=
and 48° on land, during the summer, they show signs of dis
tress from heat whenever they make any exertion, pant, raise
their hind flippers, and use them incessantly as fans. AVith tbe
thermometer at ooO-GOo, they seem to suffer even Avhen at rest.
and at such times the eye is struck by the kaleidoscopic appear
ance of a rookery, on ^vhich a million seals are spread out in
everj' imaginable position their bodies can assume, all indus-
triously fanning themselves, using sometimes the fore flippers
as ventilators, as it were, by holding them aloft motionk'ss, at
the same moment fanning briskly with the hind flipper, or tiip
pers, according as they sit or lie. This wavy motion of flapping
and fanning gives a peculiar shade of hazy indistinctness tu
the whole scene, which is difficult to express in language ; but
one of the most ])rominent characteristics of the fur-seal is this
fanning manner in which they use their flippers, when seen ou
the breeding-grounds in season. They also, when idling, as it
were, oft" shore at sea, lie on their sides, with only a partial ex
posure of the body, the head submerged, and hoist up a lore or
liind flipper clear of the Avater, while scratching themselves or
enjoying a nap; but in this position there is no limning. 1 say
' scratching,' because the seal, in common with all animals, is
preyed upon by vermin, a si)ecieK of louse and a tick, peculiiu
to itself.
''AH the bulls, from the vei;, first, that have been al)li' to
hold their positions, have not left them for an instant, iiiylit •'!
day, nor do they do so until the end of the rut ting-season, wlii^li
subsides entirely between the 1st and lOth of August, begin
ning shortly sifter the <;om ing of the cows in June. Of necessity.
HAIUTS.
353
tlKiclon', this causes tlieni to fast, to abstain eutirdy from food
of any kind, or water, for three months at least, and a few of
iliciii stay four months before goinjjj into the water for the first
time after haulinj; up in May.
"This alone is remarkable euouj^h, but it is simply v.uudtrful
wlicn we conie to associate* the condition .with the unceasing;'
activity, restlessness, and duty devolved upon the bulls as
hoads and fathers of large families. They do not stagnate, like
bears in caves ; it is evidently accomplished or due to the ab-
sorption of their own fat, with which they are so liberally sup-
jiliod when they take their positions on the breeding-ground,
and which gradually diminishes while they remain on it. But
still some most remarkable provision must be made for the en-
tire t()ri)idity of the stomach and bowels, consequent upon their
being empty and unsupplied during this long period, which,
however, in spite of the violation of a supposed physiological ^ —
law, does not seem to aflfect them, for they come back just as \^ Qjuuv^
sleek, tat, and ambitious as ever in the following season.
"I have examined the stomachs of a number which were
driven up and killed immediately after their arrival in spring,
and natives here have seen hundreds, even thousands, of them
tlnring the killing-season in June and July, but in no case has
anything' been fonnd other than the bile and ordinary secretions
of healthy organs of this class, with the exception only of find-
ing in erery one a snarl or cluster of worms {Nematoda), Irom
the size of a walnut to that of one's fist, the fast apparently
bavhig no etfect on them, for when three or four hundred old
bulls were slaughtered late in the fall, to supply the natives
with 'bidarkee' or canoe skins, I found these worms in a lively
• ouditiou in every jjaunch cut open, .and their presence, I
think, gives some reason for the habit which tliese old bulls
liave (if swallowing small bowlders, the stones in some of the
^toinaehs weighing half a pound or so, aiul in one paunch I
found about five pounds in the aggregate of larger pebbles,
^hich in grinding against one another must destroy, in a great
iiieasiirc, tliese intestinal i)ests. The sea-lion is also troubled
ill the saiin' way by a similar species of worm, and 1 have pre-
~^'i V('(l a stomach of one of these animals in which are more
ilian ten pounds of bowlders, some of them alone quite large.
li'' greater size of this animal enables it to swallow stones
^vliicii wcao], t^vj, .^,1,1 tiiive pounds. I can ascribe no other
|'''"i^e lor this habit anuuig these animals than that given, as
Misc. Pill). No. 11.' 1';5
354 CALLORIIINUS UBSINUS — NOUTIIERN FUR SEAL.
tLey are of the highest typo of the carnivora, eating fish as a
regular means of subsistence; [*] varying the monotony of this
diet with occasional juicy fronds of sea- weed, or kelp, and per-
haps a crab, or such, once in a while, provided it is small iiml
tender, or soft-shelled.
" Between the 12th and 14th of June the lirst of the cow-seals
come up from the sea, and the bulls signalize it by a universal.
spasmodic, desperate fighting among themselves.
'' The strong contrast between the males and females in size
and shape is heightened by the air of exceeding peace aud
iimiability which the latter class exhibit.
'' The cows are from -4 to 4i feet in length from head to tail.
,and uuich more shapely in their ])roportions than the bulls, th-
neck and shoulder being iiot near so fat and heavy in propor
tion to the posteriors.
"When they come up, wet and dripping, they are of a dull,
dirty-gray color, darker on the back and upper parts, but iu ;i
few hours the transformation made by drying is wonderful;
you would hardly believe they could be the same animals, foi
they uow fairly glisten with a rich steel and maltese-gray lustti
on the back of the head, neck, and spine, which blends into an
almost pure white cm the chest and abdomen. But this beauti
ful coloring in turn is altered by exi^osure to the weather, lor
in two or three days it will gradually change to a dull, rufoiu>
ocher below, and a cinereous-brown and gray-mixed above ; this
color they retain throughout the breeding-season up to the timi'
of shedding the coat in August.
" The head and eye of the female are really attractive ; tlio
■expression is exceedingly gentle and intelligent ; the large, luij'
trous eyes, in the small, Avell-formed head, apparently glcM
with benignity and satisfaction when she is perched up on some
•convenient rock and has an opportunity to quietly fan herself.
" The cows appear to be driven on to the rookeries by au ac
curate instinctive appreciation of the time in which their period
of gestation ends; for in all cases marked by myself, the pups
are bom soon after landing, some in a few hours after, but most
usually a day or two elapses before delivery.
[*Tho habit of swallowiug stones i.s one apparently foiiunon to nil oil"'
Pinnipeds. Tlie cominon belief among seab^rs and others is that llioy t.iKf
in these stones aw ballast. Compare on tliis point a ([notation alrcidy ^'ivtl'
respecting the Southern 8ea Lion ( OtariajiiVata), diitca, p. 31L Mr. KlHott'
oxplauatiou apiwar.s to ))e more reasonable tJian most that have been pro-
posed.]
HABITS.
355
"They are noticed and received by the bulls on the water-
line station with niu<;h attention ; they are alternately coaxed
and urged up on to the rocks, an«l are inmietliately under the
most Jealous supervision ; but owing to the covetous and ambi-
tious nature of the bulls which occupy the stations reaching
way buck from the water-line, the little cows have a rough-and-
tunihle time of it when they begin to arrive in small numbers
iit fust ; for no sooner is the pretty animal fairly established on
tilt' station of bull number one, who has installed her there, he
pciliajts sees another one of her style down in the water from
which she has just come, ami in obedience to his polygamous
feeling, he devotes himself anew to coaxing the later arrival
in the same winning manner so successful in her case, when
bull number two, seeing bull number one oft" his guard, reaches
out with his long strong neck and picks the unhappy but pas-
sive creature up by the scruft" of hers, .just as a cat does a kit-
ten, and deposits her on his seraglio-ground ; then bidls immber
tliiee, four, and so on, in the vicinity, seeing this high-handed
oiieiation, all assail one another, and especially bull number
two. and have a tremendous light, perhaps for half a minut or
so. and during this connnotion the cow generally is moved or
moves farther back from the water, two or three stations more,
where, when all gets quiet, she usually remains in ])eace. Her
last lord and master, not having the exi)0sure to such diverting
temptation as had her first, he gives her such care that she not
only is unable to leave did she wish, but no other bull can seize
upon Iter. This is only one instance of the many difterent trials
and tribulations which both parties on the rookery subject
tliemselves to before the harems are filled. Far back, fifteen or
twenty stations deep from the water-line sometimes, but gen-
wally not more on an average than ten or fifteen, the cows
crowd in at the close of the season for arriving, July 10 to 14,
and then they are able to go about pretty much as they pleast?,
''»r the bulls have become greatly enfeebled by this constant
I'Sliting and excitement during the past two months, and are
'liiite content with even only one or two partners.
'• The cows seem to haul in coinpact bodies from the water up
fo tlic rear of the rookeries, never scattering about over the
Ji'ound; ami they will not lie (piiet in any position oUiSide of
tlie great mass of their kind. This is due to their intensely
giogarious nature, and for the sake of protection. They also
select land witli special reference to the drainage, having ii
I
n^
h
350 CALLORIIINUS IR.SINUS NOUTIIKK'X I'Ull 8I-:AL.
This Is wi'U shown
oil
Kiciil dislike to watcr-piuldlcd fiiourul
Saint Piitil.
•• I hiivi^ Ibiiiid it (lilUciilt to as«'ertiiin the averaj{e nniuhci of
cows to one hull on the rookei-y, but 1 think it will be nciuiy
correct to assij^u to each male IVoiii twelve to lifteen leinales,
oc(Mi]>.vinj;' the stations nearest the Mater, and those back in the
rear from tive to nine. 1 have counted forty-tive cows all under
the charj;«' of one bull, which had them penned up on a tiat
table-rock, near Keetavie l*oint; the bull was enabled to do
this (|uite easily, as there was but one way to ^o to or come from
this ser.aglio, and on this i)ath the old Turk took his stand and
guarded it well.
"At the rear of all thes^ rookeries there is always a large
number of able-bodied bidls, who wait patiently, but in vain,
for families, most of them having had to fight as desperately
for the privilege of being there as any of their more fortunately
located neighbors, avIio are nearer the water than themselves;
but the cows do not like to be in any outside position, where
they are not in close company, lying most quiet and content in
the largest harems, and these la vge families i)ack the suiface of
the ground so thickly that there is hardly moving or turninj,'
room until the females cease to come up from the sea ; but the
inaction on the part of the bulls in the rear during the ruttinj,'
season only serves to q aalify them to move into the places va-
cated by those males who are obliged to leave from exhaustion,
and to take the positions of Jealous and fearless protectors for
the young pups in the fall.
"The courage wuth which the fur-seal holds his position, as
the head and guardian of a family, is of the very highest order,
compared with that of other animals. I have repeatedly tried
to drive them when they haxe fairly established themselves,
and have almost always failed, using every stone at my com
mand, making all the noise I could, and, finally, to put their
courage to the full test, I walked up to within 20 feet of a bull
at the rcfir and extreme end of Tolstoi Kookery, Avho had four
cows in charge, and commenced with my double-barreled
breech-loading ,shot-gun to pepper him all over with nnistiird
seed or dust shot. His bearing, in spite of the uoise, smell of
powder, and pain, did not change in the least from the usual
attitude of determined defense wiiich nearly all the bulls as-
sume when attacked with showers of stones and noise; lit'
would dart out right and left and catch the cows, which tini
lIAniTS.
357
idly atti'inptt'd to run after oiieli report, and tliuff and drag
tliciii hack to tlioir jdacos ; tlioii, strctcliing uj* to his lull height,
look MIC directly and defiantly in the lace, roaring and si>itthig
most vclieniently. The cows, however, soon got away from
liiiii: hut he still stood his ground, making little (charges on me
of 10 or I-"* fwt in a saceession of gallops or lunges, spitting
finiously, and then retreating to the ohl position, back of which
lie would not go, fully resolved to hold his own or die in the
attempt.
"This courage is all the more noteworthy from the fact that,
in re;,fard to man, it is invariably of a defensive character. The
seal, if it makes you turn when you attack it, never follows
you much farther than the boundary of its station, and no ag-
•fiavation will comi)el it to become otteusive, as far as I have
been able to observe.
"The cows, during the whole season, do great credit to their
amiable expression by their manner an^ ^> \. w^
358 CALLORHINUS URSINUS— NORTHERN FUR SEAL.
" The sleep of the fur-SQ^l, from the old bull to the young pup^
is always accompanied by a nervous, muscular twitching aud
slight shifting of the flippers ; quivering and uneasy rolliuup.s get together like a great swarm of bees, spread out
fl
360 CALLORIIINUS URSINUS NORTHERN FIR SEAL.
upon tlic ground in 'pods' or groups, while they are youiif;,
and not very hirge, but by the middle and end of Seiteinber,
until they leave in Xoveniber, they cluster t')gether, sleeping
and frolicking by tens of thousany connuon consent between the harems. Ou these lines
of i)ti><«age they are unmolested by the old and jealous bulls,
who guard the seraglios ou either side as they go and come ;
generally there is a continual file of them on the way, travel-
ing' up or down.
"As the two and three year old holluschukie come up in small
squads with the first bulls in the spring, or a few days later,
these common highways between the rear of the rookery-ground
and the sea get well defined and traveled over before the arrival
of the cows ; for just as the bulls crowd up for their stations, so
do the bachelors, young and old, increase. These roadways
may be termed the lines of least resistance in a big rookery ;
they are not constant ; they are splendidly shown on the large
rookeries of Saint Paul's, one of them (Tolstoi) exhibiting this
feature finely, for the hauling-ground lies up back of the rook-
ery, on a flat and rolling summit, 100 to 120 feet .above the sea-
level. The young males and yearlings of both sexes come
through the rookery on these narrow pathways, and, before
reaching the resting-ground above, are obliged to climb up an
almost abrupt bluff, by following and struggling in the little
water-runs and washes which are worn in its face. An this
is a large hauling-ground, on which fifteen or twenty thousand
commonly lie every day during the season, the sight always, at
all times, to be seen, in the way of seal climbing and crawling,
was exceedingly novel and interesting. They climb over and
np to places here where a clumsy man might at first sight
say he woidd be unable to ascend.
"The other method by which the 'holluschukie' enjoy them-
selves on land is the one most followed and favored. They, in
this case, repair to the beaches unoccupied between the rook-
cries, uud there extend themselves out all the way back from
the water a<5 far, in some cases, as a (piarter of .a mile, and even
farther. 1 have had under my eye, in one straightforward
sweep, from Zapad-nie to Tolstoi, (three miles,) a million and a
366 CALLORHINUS UR8INU8 — NORTHERN FUR SEAL.
half of seals, at least, (about the middle of July.) Of these I
estimated fully one-half were pups, yearlings, and * hoUuschu-
kiw.' The great majority of the two latter classes were hauled
out and pack'^d thickly over the two miles of sand-beach and
flat which lay between the rookeries ; many large herds were
back as far from the water as a quarter of a mile.
•'A sumll flock of the younger ones, from one to three years
ohl, will frequently stray away back from the hauling-ground
lines, 'out and up onto the fresh moss and grass, and tliere
sport and play, one with another, just as puiipy-d'ogs do; and
when weary of this gamboling, a general disposition to sleep is
suddenly manifested, and they stretch themselves out and curl
ui) in all the positions and all the postures that their flexible
spines and ball-and-socket joints will permit. One will lie upon
his back, holding up his hind flippers, lazily waving them in
the air, while he scratches or rather rubs his ribs with the fore
hands alternately, the eyes being tightly closed ; and the breath,
indicated by the heaving of his flanks, drawn quickly but regu-
larly, as though in heavy sleep ; another will be flat upon his
stomach, his hind flippers drawn under and concealed, while he
tightly folds his fore feet back against his sides, just as a fish will
sometimes hold its pectoral fins ; and so on, without end of va-
riety, according to the ground .and disposition of the animals.
"While the young seals undoubtedly have the power of going
without food, they certainly do not sustain any long fasting
periods on land, for their coming and going is frequent and
irregular ; for instance, three or four thick, foggy days Avill
sometimes call them out by hundreds of thousands, a million
or two, on the different hauling-grounds, where, in some cases.
they lie so closely together that scarcely a foot of ground, over
acres in extent, is bare ; then a clearer and warmer day will
ensue, and the ground, before so thickly packed with animal-
life, will be almost deserted, comparatively, to be filled again
immediately on the recurrence of favorable weather. They are
in just as good condition of flesh at the end of the season as at
the first of it.
"These bachelor seals are, I am sure, Avithout exception, the
most restless animals in the whole brute creation ; they frolic
and lope about over the grounds for hours, without a moment's
cessation, and their sleep after this is short, and is accompanied
with nervous twitchings and uneasy movements; they seem to
be fairly brimful and overrunning with warm life. I have never
HABITS.
367
observed anything like ill-humor grow out of their playing to-
{jetlicr; invtiriably well pleased one with another in all their
frolicsoiue struggles.
"The pups and j'earlings haA'e an especial fondness lor sport-
inj: on the rocks which are just at the watei-'s level, so as to be
alteniiitely covered and uncovered bj- the sea-rollers. On the
bare summit of these water- worn spots they struggle and clam-
ber, a dozen or two at a time, occasionally, for a single lock ;
the strongest or luckiest one pushing the others all ott", which,
liowever, simply redouble their efforts and try to dislodge him,
who thus has, for a few moments only, the advantage ; for with
the next roller and the other ])ressure, he generally is ousted,
and the game is repeated. Sometimes, as well as I could see,
iho same squad of 'holluschukie' pl.ayed around a rock thus
situated, off" ' Nail Sped' rookery, during the whole of one day;
but, of course, they cannot be told apart.
"The Miolluschukie,' too, are the champion swimmers; at
least they do about all tlie fancy tumbling aut"red by bemg clubbed.
he olo • ^ '''' ' *^'" *^' P"'" i« instantly withdrawn, and
sdZtH 1 ""I'^l'''^""*^ *^ ^'^^ floundering creatm-e, which
1 f^^^^^^^ :f ^f-e described. Indians from the Vancouver
^liorelieciuently Start uUhe night, so as to be on the bestseai-
* Condition of Affairs in Alaska, pp. 123-150.
t Hist. Kamtsch. (Englisli ed.), p. 130. , '
.(«
.'172 cAi.T.oiiiiixrs imsiNra — noktiikrn fuk seal.
>V;'
r\
\
W
iiij,' fjfiound in the inorninjf. This localitj' is .said to bo soutli-
Avcst of Cape Classet, five to fifteen miles distant."*
In liuntinj^ Seals for their eommercial prodncts the common
method of killing them appears to have generally been by club-
bing them, as is at present practiced on the Seal Islands of
Alaska, one or two lieavy blows npon the head being sulficieut
to dispatch them. The method of attack is very much like that
practiced in destroying herds of Walruses, already described.
A large party cautiously laud, when possible, to the leeward of
a rookery, and then, at a given signal, rush upon the Seals, with
loud shouting, and with their clubs soon destroy large number.s.
It has generally been practiced without system or restraint,
resulting in the speedy destruction of large rookeries. As is
well known, the Southern Sea Bears or Fur Seals {Arctocephalm
^^falklandicus," A. forsteri, A. ^^ cinereiiSj'^ etc.) were long since
practically exterminated at many localities where they were
formerly very abundant, as has been the case Avith the Northern
Fur Seal on our own Califomian coast. At one time thje same
destructive and ruinous policy was pursued by the Kussiaus at
the Prybilov Islands, but th^ folly of such a practice was soon
peiceived, and through government interference theii* extermi-
nation there has been happily prevented. Their destruction is
at present regulated by the United States Government, the
j whole matter being judiciously and systematically managed.
' The manner of taking and killing the Seals, and the method
adopted to prevent their decrease, has been described in detail
by Mr. Elliott, and is here appended.
"Taking t'ie Seals. — By reference to the habits of the fur
seal, it is plain that two-thirds of all the males that are born (and
; thoy are equal in number to the females born) are never per
mitted by the remaining third, strongest by natural selection,
" to land upon the same ground with the females, which always
■• herd together en masse. Therefore, this great band of bachelor
seals, or ' hoUuschuckie,' is compelled, wiien it visits land, to
live apart entirely, miles away frequently, from the brooding
grounds, and in this admirably perfect manner of nat.ne arc
those seals which can be properly killed without injury to the
'rookeries selected and held aside, so that the natives can visit
and take them as they would so many hogs, without disturbing
in the .slightest degree the peace and quiet of the breeding-
1 grounds where the stock is perpetuated.
* Marine Mammalia, pp. 1.54, L^S.
MODE OF (ArrrHi:.
373
-The niiimier in which the nativi's Ciiptuic iuid drive the
lioiliisclmckie up from the htiuliiig-;4roiiiuls to the sh»U{>hteriug-
liclds near the vilUige.s and elsewhere, cannot he improved upon,
;iii(l is most sutisl'actory.
"in the early part of the season large bodies of the young-
baclielor seals do not haul uj) on land very far from the water,
a liw '<>