IMAGE EVALUATION TEST TARGET (MT-3) // i 1/ ;<" c?^ ^^ & ^ z ^ 1.0 I.I ■^i^ lllliM <^ 1^ III 2.2 :: lis 12.0 1.8 11-25 11.4 11.6 m Va /: '/ /^ CIHM/ICMH Microfiche Series. CIHM/ICMH Collection de microfiches. Canadiar Uistitute for Historical Microreproductions Institut Canadian de microreproductions historiques 1980 Technical Notes / Notes techniques The Institute has attempted to obtain the best original copy available for filming. Physical features of this copy which may alter any of the images in the reproduction are checker^ below. L'Institut a microfilm^ le meilleur exemplaire qu'il lui a 6t6 possiLle de se procurer. Certains d6fauts susceptibles de nuire d la quality de la reproduction sont notds ci-dessous. Coloured covers/ Couvertures de couleur Coloured maps/ Cartes g6ographiques en couleur □ Coloured pages/ Pages de couleur Coloured plates/ Planches en couleur D Pages discoloured, stained or foxed/ Pages d6color6es, tachetdes ou piqudes Show through/ Transparence E Tight binding (may cause shadows or distortion along interior margin)/ Reliure serrd (peut causer de I'ombre ou de la distortion le long de la marge intdrieure) D Pages damaged/ Pages endommagdes D Additional comments/ Commentaires suppidmentaires Bibliographic Notes / Notes bibliographiques D Only edition available/ Seule Edition disponible n Pagination incorrect/ Erreurs de pagination D Bound with other material/ Reli6 avec d'autres documents D Pages missing/ Des pages manquent □ Cover title missing/ Le titre de couverture manque Plates missing/ Des planches manquent D Maps missing/ Des cartes gdographiques manquent n Additional comments/ Commentaires suppldmantaires The images appearinf; here are the best quality possible considering the condition and legibility of the original copy and in keeping with the filming contract specifications. Les imagas suivantes ont 6t6 reproduites avec le plus grand soin, compte tenu de la condition et de la netteti de I'exemplaire filmd, et en conformity avec les conditions du contrat de filmage. The last recorded frame on each microfiche shall contain the symbol — »► (meaning CONTINUED"), or the symbol V (meaning "END"), whichever applies. Un des symboles suivants apparaltra sur la der- nidre image de cheque microfiche, selon le cas: le symbole — ^ signifie "A SUiVRE", le symbole V signifie "FIN". The original cooy was borrowed from, and filmed with, the kind consent of the following institution: Library, Geological Survey of Canada Maps or plates too large to be entirely included in one exposure are filmed beginning in the upper left hand corner, left to right and top to bottom, as many frames as required. The following diagrams illustrate the method: L'exemplaire film6 fut reproduit grdce d la g6n6rosit6 de I'dtablissement pr§teur suivant : Bibliothdque. Commission Gtologlque du Canada Les cartes ou les planches trop grandes pour dtre reproduites en un seul cliche sont filmdes d partir de i'angle sup6rieure gauche, de gaurhe d droite et de haut en bas, en prenant le nombre d'images ndcessaire. Le diagramme suivant illustre la mdthode : 1 2 3 1 2 3 4 5 6 DEPAKTMKNT OF THE INTERIOR D. 8. GEOLOGICAL AND GKOOIIAPHICAL SURVEY OF THE TEBaiTOBIBS F. V. IIAYDEN, Oroi.oowt-in-Ohabo* MISCELLANEOUS FUBLICATIONB-No. 12 HISTORY OK ISrORTH AMERIOA-ISr PINNIPEDS A MONOfiKAIMI or THE WALRUSES, SEA-LIONS, SEA-BEARS SEALS OF NORTH AMERICA. Bv JOEL ASAPH ALLEN AisMantin lln Mimeiim of Comparative Zoology at Cambridgi Special Collaborator of the Survey WASlMNCiTON GOVERNMENT I'llINTINO OFFICE 18S(» <• ^ :• : PIIEFA'I'ORV NOTE. United States Geological and Geograpuioal Survey of the Teuritories, Washimiton, D. C, 'fitly 1, 1880. Tlii' picsciit si'iies ol' iDonofiiaplis of the North Anit'ricHii Piiiinji(nl>li- cation of the part already i)repared relatinji' to the Peiniipeds, As nearly all of the K])ecies belonjiinj^' to this {:ronp foinid in the nortlunii lienMsjdiere are nuMubers of the North American fauini, the ]n'esent treatise is virtually a mono}.';raph ol all the species occurring;' north of the ecpiator, anrinter, it i»roves to be, to only a small degree, a rei>etitioii of the account given by ]Mr. Elliott, also rejuoduced at length. The history Captain liryant gives of the changes in the numbers and relations of the different classes of these animals at the rookeries, under the present system of management of tl>e Fur Seal business, forms a valu- able l)asis for generalization in regard to the future regulation of this industry, and is aiso an important contxibution to the life-history of the species. The cuts, some thirty in munber, illustrating the cranial char- uctt'rs of the Walruses, were drawn for the present work by Mr. J. H. Blake, of Cambridge, and engraved by ^lessrs. Bussell and Eichardson, of Boston. The Survey is indebted to Professor Baird, Secretary of the Smithsonian Institution, for aseries of six- teen original tigui'es, engraved «y Mr. H. H. Nichols, of Washing- ton, from photographs on y «M,d. vllustrating the skulls of CaWoWti- nus ?( rsin MS, Peale's ^'■Haiivhwmaantarcticuii,''^ (Jystophora crLstata, ^-^.»f-^^f-:rf*f-,^ (»f London for electros of (7 ray's ^^HalUyon richardsi," and of a seri''H yf historic tignres of the walrus i>ublished in the Society's »' l»roceedinj;s," Ity tiie late Dr. Gray, and to the proprietors of "Science (Jossii),"for electios of the full-lenj;th figures of seals. These were reci'ived throuyh Dr. Cones, who also furnished the full-length \iews of EnmetopiaN .stelleri and CaUorhinus ursinns. Mr. Allen desires nie to express, in tiiis connection, acknowl- edgnients of liis indebtedness to IMof. Si)encer F. Bdrd, Secre- tary of the Smithsonian Institution, and to Prol". Alexander Agassiz, Director of the iNluseum of Comparative Zooh)gy, for the liberality with which they have placed at his service the rich material relating to this gTon]) of aninuds ( )lt.K.\ 1 1 >.K— Wiilnmcs 5-180 Sjuniiviiiy .I tii'Uriiil olMcrvatiiuis iuid cliiinicteM of the j^roiip 5-12 (Jfiicia 12-14 S.Miii|i»i.-( III till- iii'iina 14 til'IIUS (•IHIII.EM > 14 S.viniiiyiiiv anil lii.stmv 14-17 Species 17-23 ( M)Oiia;m s KosM Alii s— Atlantic Wi'lrus 23-147 Syuoiiviuy anil liililiii!;iai>liii'al ict'iTcnces 23-20 Kxtcrnal iliarartcis 2()-;)8 Srxnal ilill.Ti iiics 3S-43 Inii'.iilual viiiiat inns anil vaiiatiouH dcprmlL'Ut npou age 43-45 McaHiilrnirnts 111 .sliiills 40 Unutitii)n 47-37 l''o«sil ri'iiiains ."i7-fl5 (rfogiapliiral ilistrilmliim. pivsrnt ami pUHt 05-79 f '(last lit Ndi til Ainriira Oi5-71 ("oast lit' Kniopc 71-79 NoinciiclatuiT 80 EtyuKdoiiy bO-82 Litcratiiiv . , 82-107 ( icnt ral history 82-92 Fi)j,nivs 92-107 Habits and tlu« cliaBe 107-133 Products 133-134 Food 134-137 Fiiui'tiiins of the timks 137-1"" Knciiiics 13H- Diiincstiiation 140- Oiioha;m K oiiKsif—l'acitlc Walrus 147-.' Synonymy and liildiiijiiaphical references 1 Kxlenial iliaraeters and skeleton 147-1 Measurements of ' deton 14!»-1 Measinenielits ot . ..tills 15 Uill'ereiitial eliaracters 150-171' Nomenclature . .. 170-171 General history 171-172 Figures 172-174 < teou;rapliical distribution 174-178 llaliits, food, commercial products, and the chase 178-180 Family tVl'AKI ID.K— Fared Seals 187-411 Synonymy and characters of the group 187 Technical history 188-207 Higher gi-oups 188-190 (leuera 190-193 Species ia3-207 IX ■j.iii •j;i7-j:is J44 X TABLE OF CONTENTS. Page. t'li.aiirliT.H 111' I he I'lNMl'KIH \--('()ntuiiir(l. KmiiiUv OTxVIUIDJ:— KmitiI ScalH. S\ iiiipsiH of till, ^iiiiia unci .spiicics 2U8-ai3 M.vlhic.'il :iu(i uiiilitiiiiiiMiibli' spcciea 2U->l(i tliDm^liliical ilistiiliiilion 216 f'o»,sil ( Itiiiics ''l?.-!'"! Milk (Iciititidii 2''l-2"4 •Infj,'iiliiiilips of (Iciititimi ^24 I'lLsition of (h,. 1,181 upper piriiiiincnt mou.r ^iri (rfui rill oliaervationn "ja.'i-'J''" Habits ""' .„"- J J 1 Products .,.,g Pestrurlioii of Fur Si iils for tiirir peltries i;'Ji*-J31 Genus Ml MKTOIMA.S ._i;j1 Ei;Mi:roi'iA.s biklleui— StfUci s Sea-Lion 2;i'j-274 S.vuou.viiiy iiml l)il)lio-raphi(Ml n iVjicncos L':)2 Kxtt'ina! cliaraclir.s Extoriial niia.surt'uicnts Skull ' '" .McaauriTiiiiits of skulls .. TlM'l ll ^rji, Sk(li't(Mi 24(1-241 ili'a.siiiriUfUt.s of skrlt'ton 212-244 Sexiuil, ailolfsifut, ami iuilividual variation 244 Gpofir.ipliiial \ aiiatiou Comparison with allied .speeies 244-24 .McasiiiTiiii ills of skulks of OTAiilA JliDATA 247 Geouiapliical iKsiiiliiiliuu 248 General bistm y and nonienelature 248-254 Habits 254-274 Genus ZAUii'iirs 275 Zai.01'hi-,s ( Ai.itniiMANrs— Califcuiiiau Sea-Lion 27ii-;il2 Syuouyniv ami bibliosiiapbieal references 270 External eliaraelrrs ]'_' 27ii-27^ Younu .,7^ •'''l"!i'- 27b Si^«; 27>^-28;i External measurements 2711-280 Measuieiiients of skeleton (d' female 281-283 ^"^"11 283-285 ileasnieiuenls of skulls 285 Dentition .jjjg Sexual dirt'erenees 287 Variation with age 287-289 Coraparison with allied speiiies 289 Geographiealdistiibiition 280-291 General liistory and nomenclature 291-290 H"l"'« 296-312 Ge.nusOAi.LoimiNis 312-4l(i UALi.oHHiiM s imsLNUb— Northern Fur Seal 3i;j Synonymy and bibliographical references 313-314 External eharaetiTS ;j14 Color jjl^ Pelage ;j|,-, Size 31»-319 External measurementa ... 319 Ears ;j2u Fore limbs ;j2i) Hind lindis 320 TAIJLK OF CONTENTS. XI Page. Chiira«tci sol' till- I'i.vmikiha— ('(intinind. raniily OTAIillD.K— Kiiicd S.uls. Gi'nuH('ALi.ij|;iii.Niis. CAi.LOiiiil.Ni'8 UU8I.NLS — Northern Kur Seal. Skull 820-323 Moasuvcmonts of skulls 323 Tc-etli 3i;4 Skclctdii 324-32t) ^Mtasiin nil Ills of ski'letun 325 Si'XUiil (lill'.iiniiH 325-327 Difl'iicnccH iiHuhiii;; frtiiii age 327 Inilividiiiil variiitioii 'S'JS Comiiaiiwin with alliccl Hpocies 329-331 Mcasurciiu uts nf skulls of AliCTOCEPHALUB AUfiTIIALIS 331 Oi'Diiiniiliiral ilistriliution and niifiration 332-335 GeiH'i'al liistmy and ncninuclature 335-33'J Fijiuii's 330-3 U Ilalnts 341-371 Till' cliasi! 371-3Tr Miidr lit Caiiluri- STl'-:;"- llistiiiy aud pKisjiicts iif the Kur Seal business at the Prybilov Islands 37H-3«1 EufUiifS (if llic I'm- Seals Ilj^l History of tlie Kiir Seal Fishery at thi; I'rybllov Islauda, Alaska. iVimi 186!) 1(1 1«T7, liy Ciiaui.hs Huyani' :)i?J-tll I'reliiniuary aud acueral oliservatioiis ;ihj-:j:<.- Kecelit ehauuis in the lialiits and relative uiimbers (d' the ditlerellt classes (if Seals :il?>?-;i!(5 Cause (if tile eliani;es in I lie lialiits (if the Seals, &e 3ilh-4U2 Albinos and sexually almoriual individuals 4(13 Descriiition of the youni; ; vaiiatiou in eohn' with ago, &c. . . . 403 M.iltin.i.' 404 Sexual orj^aus, &e 405 I'ower (if siispendiug respiration 40G Natural enemies 400 Eflbet of elimatie iutluences 407 Number of Seals reiiulred for the subsistence of tlie natives. . 409 Winter resorts aud habits of the Seals 410 Family PUUCIDiE—Karless Seals 412-7.10 Characters of the tirmiii 412 Technical liistory 412-4«li Highet groups 412-414 Genera 414-421 Species 421^00 Classilication 400-407 Synopsis of sub-families and genera 401-463 Synon\ niatic list of the species 463-407 Geographical distribution 407-409 Fossil remains 409-481 North America 409-470 Europe 47(i-481 Milk dentition 481-484 General habits and instincts 484-486 Food 486 Enemies 487 Mitiratious 487-491 Locomo.^ion on land 481-490 Seal huutin); 490-540 XII TAMLK OF CONTliNTH. Chiini' HiH (il llic I'lNsirKiifA— Colli iiiiiiil. F .iiiil.v I'llOCID.K- KiiiliHH SciiIh. Si-aliiif; iliHii'iriN WimI I in I'll III mi I Nrw rdiiriilliiiiil illin MllMTl 111 "(Mrl'lllllllll ' SlIlM Niivii /I'liililii ami Kiini Sni Wllltr Sill ('aH|iiiiii Sisi Niirlli I'acilii Son III rarilir ami A iilanl ii' Siiim MiIIioiIh of i:i|iliiii , ,V 1 Slioif liiiiiliii^ l')Hii|iiniaii\ iiiiilioilH Hv tlliMlls ilT IlrlH. . 'Jill' HI -a I I io\ , ..... ... 'I'll!' .sfal lioiik 'I'llr ■ SKiiiikla" Ici' liiiiil ill'.: In llii' liiiHol' liolliiiia (Ill I 111' 1 oa.sl 111 Si'uli. Ilami In I III' .Ian Mavi II Slim |laii;:i'i K anil iiinri tajnlir.s ol' \ir liiiiiliiii; .SpcrirH liniilril Alililiilaiii 1 III' Si alH III |iai I Iriilai liM'iilllirH I'rodinlH . . I'li'iiiiralion 111' llii' |iioiliirlM WiiHli Till ili'Hli III linn 111' Siiils Di'i'i'i'iiHi' rriiiii in.jiiilii'i'iii.s liiinUiit; Si'iii.s anil Sral liiliilili); in llirnlilrii llinr in llir ( iiilt' ill' St. laiwri'iici' Sub-Ciiinily I'liiii in.i, fii'nim I'liiii ,\ I'jioi A VII I i.iNA— Ilailinr Si'iil S\iionwii> anil liililii>nia|iliiriil 11 Iii'iihth K\ 111 nil I I'liarai Ins HlHlini'livi' rllllliU'll TH Iliiliviiluiil iiml HI' X mil vaiialioii ... . Mt'iiMiii'1'tnrnln iif (lie HkiillH -r)4» .')41l-,'^^,0l :.::.{ .■i.'.:!-.",.-.7 .-,.''.7 .V)7-.'i,-.!) .'"i.''ili-.'"i(i7 .")i!.*>-ri7 1 liTI-.W! .-.74 r>7.''>-.'>H4 .'H4'-.'iHH ,MI7-»K» .SIt7-tlOI) i>iHi-(iu:i IID.'I-DU.'') U(I6 01)7-1114 Ulilt-lllll •11 -.Mil;! IIM-Iiltl uii) rdu (illl-tlL'il iYM-im tw()-(i;rj o:)'j-ii:i7 ii;i7 urn ii;iu TAIlLi: OK ('ONTKNTS. CIlllliK I' IH III lll> riSMI'I'.IllA -('(llltlllUl'll. I'.uuil.v I'llOClK.lv Siili liiiiiil\ I'iKii l\.i:. (Ic'lillH rilni A. I'lKlc A ( I'Ai.cil'llll.l I l.l|c|„NI.ANI)|l'A— Illllp HiMtl. tii> ;iiiil liililiii;;rapliiital icfi^iciKtoM Kxiriiinl rliiiiai'lciM SUiill and mLcIi lull MraHiirriiii'iitH 1)1' I hi- Hki'li'tnn MiaNiiii'iiii'iilH III' III! sKiill (ii'lii'ial IiIhIiii V and iihiih nrlaliirr • li'ii:;ia|ilil< al ili^l li lull inn II aliil-i, iHiiiliirlM. anil liiiiilln;: (JllllH lll^l ninriKir \ IIimHIIiiIIKh A I'Ahi I M A — Itiiiliiin Sial Syiiiiii.Nins anil liililliij,'ia|iliiral 1 1 I'liinrrM KxliTiial I liaiaili IN Si/i- tii'iii'ial liiHlni \ Iirii;:ra|iliiral iIInI i iliiillnii lialiiK (irllll^ II Mil IIH.Ill H - < ii-ni-ral IiInIiii \ anil iliHriisMJun nl' the ''(ii-nim IN'ha " iit' Sropoli . IlAI,|rl|i|;UI - i.in I'l ,■^— (ila\ Si al S\ 111 my in \ anil liilillnurapliiral irrrri'iirnH • lOsIri nal iliaiarli IH MiaNlllrllH lit H III' mU nils (li'iiV'iapliiral ilisli iliiil ion (inirial li ill I II V anil niniii inlatiiK^ Ilaliils (iriin« Mll.NArill n .. Mi.NAi iii.s iiail'irAl.lH— WihI Inillan Siiil ... (Miaiai Ills I>anipli'i\s arriiiiiil 1 1 ills anil (iiissc .s ai riainlH. iKi:), I HP)! (iiav H ai riinnlN. IXI'.I, |h7I (■ill nil (llr W'rnl Inillall Sial.s. I Hid Aiial\Mi« anil ili.siii.ssinii nl' I In Intiiiiniiii^ Alllnilli n nl' I III' .laniaiiaii oi I'liliii Siiil ( li'nL'l;i|illlr.ll ilisit ilillljiill Sail rainih CVSl'orillilMllN.K (il'llllrt ('Vhl'll'IMKA (!VKHiriiiii!A I iii>i Ai \— llonilcd Sial . S\ 111 iiiv Illy and lillilin;.'iaphi('al ii riTriir.tm Kxiri nal rliarai lii.s Skill Inn and sUiill .Mrasiiii nnnl.-i 111' HiiiillK MiiLsiininiiilti 111' till' Hliili'tiin liin^iapliii m1 iIimI I IIiiiI inn and nii mil nniiii'iirlatilKt llalillM Iliinliliu and jiiiidinls XIII Pa(jc<. (14(1 (m-(M7 (M7 llal (i.'il (I,'.'J flr.u-ii.'M ll.-lj (ir.,-i-(i7r. (irir>-(i.';7 ():.7 B,-.*<-(Itll (ItiO litii (Kl'J lilili lillll (170 (17n-(i7"i 117:1 -ii7ii (17(MWJ Ii7() (I7(M17'< tYjH (17H-(iM| (iKI-liK-.- ()«:; (W'J-(i«!t liH.'!- i;«:i (lH!l-7lMi (in'.l^li'.tll (iiHi li'.i:: (IIM (m.'i-(i!Hi lillli-li'Jn (i!lil-7il(i 707-7nx 7iiK.7'j:t 7llH 7(IH 710 7|ip 7l"i 17.". 7 M -\H 7IK-7'JI) (Un-TJl 7'J 1-723 7'j;i 7-j:i 7:'i-7»:: 72.1 72(i 7'J(i 721* 7:111 7:1:1 7:12 7:i:t 7:1:1-7:17 7;w-71o 7ln-7H 741-712 XIV TAIU.K OF CONTENTS. Chara4'U)i'M of tin- I'inmi'I hia— Cciiitiiiiiril. Family riKMID.K. Sub- family— ('vi-i(pi'iii iiiiii.N a;. .i: 761-704 I'liuiij viliiliiia 761 Plioca fotiila 702 Plioca fiiii'iilaliillcu 701! Eritfiiatlms liarlmlus .. 703 m.stiio|ilioia fasi'iata .. 703 Hulicliiiiiis ;;iy|iiiH 704 ryHli>|ilioia ri i.slata 764 .Mariorhiiiiisaiicim' 'mtris "04 B. Adtiitidiin iiiiU I'lncrtii. n 765-774 Family OlMiii.i,M!>.v, 705-7IW (Moha'iiiiK riixiniirun — .\tlaiitlo WaJruB 765-708 .Ailditioiial iiliiiiiciH 705 Sizi anil rstiiiial ainicanilKM! 705 (ii'oniaiiliiciil ilistiihuliou 700-707 Nova Zi'inbla Kraiiz-.Iomf Laud Aliiiiiilaii' (' ill Wolatenhobne Sound SpitzlMljirll, &c lirlaiid SiipiMiHi'il pi'cHi'Mi'i^ of Walraflos in the Autarotio Soaa . 'I'lw 'Wall iirt a foriiiiilalilt> nnta^oniHt Ciiiiosiiv ami liailcHsncsH of thr Walrus I/oi iiniotioii : iiHC of tin) UukH ill climbing KiHuriH of till' Walrus Odobcmiig obemin — I'aciQo Walrus , Disliiliiiiioii Family OtaiiiidjE ( )tai iiH at t he (iolapagos Tslnnda FoHHil Olarii'S ('apt nil- of Sea Lions for menafjerles Zalophun cal{f. 41. Fi};. 2, p. 4J. Fie. 'M'- 4;). Vig- 4.1>- u:t. Fi^:. r.,p ii:i. Fiji. O.p. 1)4. Fif.'. T,p. '.»4. Fi«. H,p. !t4. Fin. l»,p. !l."i. Fiti. 1", p '.Hi. Fi«. ll.p. Kill. Fi};- 1-. !'• 101. Fi-. r!,l>. l.-.:i. Fili. M, p. I-'Im I."i II i.-.(i. 1 ."i7 . 1 IL. !•'. P- Fi;;. Ii-i. p. i:.s. Fi;;. 17. p. i."i«. Fi-. 1.S, p l.-.!t. Fi^;. l!".p I.V.I. Fi;;. Jii. p ICO. Fi^'.'Jl.p Kil. FiL'. -JJ, P ll!J. Fi^'.-J:i,i) l(i^. Fi.-. •-•4, P i(i:i. Fi;.. -J.-., p IGli. Fi«. -jr,. p 104. Fii; -7, p Km. Vi'A--Jx.\< II 10. Fi-.21l, p 100. Imk- :• 107. Fit'.iil.p . 107. Fi-. .'l^, p lOK. Fi-. ;)3, 1 . IIW. Fi-. :;i, 1 . 101). Fi;;. ;t.->, 1 . lO'.t. Fi-. :ili. I . i7;i. Fi,;;. ;i7, | . ;;,■>». Fi-. -.iX, 1 .:ii7. Fi;;. :i!l, 1 . :i'.'i. Fi-. -li), 1 .:i'.'i. Fi-. 41,1 :rj^'. Fi;;. 4J. i . :cj;;. Fi-. lit, 1 . .')0:i. Fi;;.4J. 1 . .V(). Fis.4.'-..| . "iHO. Fiu. 4(i, I .581. Fi-. 47, 1 . .182. Fig.4H,|). '8:1 Fi(f. 4«, p. Udl . Odvbimiwi rimmarut. skull nl' ((oinilf in pnilili' anil lowiT.juw from above. OdAibniPuim rnnmariui, skull nl' fiiiiiilf fiDiii iilinvf. Odob-iiiuH riiKiiKinin. .•^KuU nl !• miilr tVniii liclcw. OlauH M;i-iiiis s " Udsuianis srii .Mnrsiis \iiivf;;icuH." (•laiis MaLiiiiis's I'linus alniistniMis Ocriiui (li'nniiuici." (H'Siii'i'.s '■ KiisMianis." tiisiici s •■ Varca iiiaiiiia " (.\ililii)(ia tii Icoiifs .\iiiiiial). (ii'sui r's •■ KoHiiianiM" (Innics Aniiiiill.. l.'iOli/. l)r Vri'is 'Sia IIkisc," 100!I. HisHil (Ji'iaiil's " \ValiU8s," lOia. Martin's '• Wall-niss," 17ti.">. l!llll'(ill's ■' I.r Mdl'Sl'.' 17*i."i. OiliihiriiiiK iiIk^hhii, llilri- \ ii WH i)f licail. (Mliihitiynim iilieiiit'<. sliiiU in imililc. Odiilxi'iiiis riiKiiiiiniK, skull in pmliJi-. Oil'ihifiniH iiJHiiinntn, skull liniii I'riiiit. Oiliilni'iDiH ubi'siiK, skull rriiiii liniit. (J(tiib(riiUK niHiiKirim. m ripilal viiw of xkuU. (JduhiviiiiK iibexiin. iMii|iii.il \ ii-w nl' skull. (Miihii'itiiH roHmaruH, skull IVom almvf. (Jdiil'd'UUX (ibf.Kiis, m1;u11 I'l nil almvi-. OdiihifiiiiH iibfKtiu. .Minn- skull fimii almvi'. Odiibdniw nmiiKinin, Mnni- skull I'lnni above. (Jddbunian obebitu, .miiiu- skull I'miu I'riiut. Odobwnnn rumiMnui, .viniii.i; skull f'lniu t'niut. Odobii'iiu.i riiKiiioruii, .skull I'lcmi liclovv. OdiibiviiHH ohemix. skull riimi lirl.iw. Odiihii'nnn rnKKinriiK. |ci«ir.j,,« ridiii aliiivi'. Oddbiiuun (ibentin, lowri jaw I'linu abuvo. Odobimius rimmaniH. Inwcijaw t'rnni side. (JdnhiiiniH tilnKim, liiwci- jau I'lnni Bitlc. nddbiiiitm riiHiitdiKu, liiwii .jaw nl' young from above* Odvbd'uiiH (ihcKiiK, Inwi-r jaw nl \ nuug from nbovo. Odulnriiiin riiniiiariDi, lower, j.iw nl' young from Hide. OilnhirnuH iibi'xu)!, lower jaw nl' ynuiig I'rom Biilc Odobd'iiiin ubt'sim. Cnnk'.s fl-ure of tlif aiiironl. KumeUijniiH stelUri, li-uriB of animal VaUi/rliiniin nrninun, li-urcM of animal. (JaUurlii'ii^KK iiitiintin. skull nf fiinalr iu protilc. ('allurliiniig urtiiiniii, skull of I'l luiiln from ubovi-. Cnlliirlnniin iimintiii, skull nf fiinali lower jaw. ('ulliiihuiiin iirniniiK. skull of finialr from below. J'liiicd litiiliitii. auiinal. . " llalichwriiM ant'irrliCHH," I'liiln, Hkull iu prollle. " llulii'lidriin <(iiliir<:t.icuK," I'rali', okall from above. " niilirliwrus iintdrctiKng," I'lule, Bkiill lioin below. " Ilalichiernii antnrclicut" I'ealc, lower juw. " tlalicyun richtirdgi" (iray. skull iu profile. Fhiica firtida, uuinuil. XVI IJST OK IIJ.l'riTltAI lONH. I'iu ■'", l> n:i:i Ki«.r.i,|. )ll)l Kili.V.M. 7'.' ( Klu.r.:i,|. 7-JH V\H. M, |. 7211 Ki«. r,r., p Tlsn KiK. .'>«, I, 7;ii KiK ■■•T, |. 711 Ki-. r.K, |, 71:. KiU'. . ■■,!), p 7 Hi Hlmio, p 717 t'Imrti ^fruliinflif-ii, ;ilMiii;il. llntirliirniH 'iri^jiiiK. ;iiijiiiiil ('ilKliiiilinrii frinlii>ii. :iiiiiti;il I'llHli'iiliiifii ••iiHldld mIuiII ill piiillli'. I'liKfiiiiliinii irinldlii ^^liiill hciiii iiliiiviv ('jltiliiithdrii rrinldlii. sKiill ficiiij ImIuvv. (hlKhi/iliiirii ftisliilii. lipHi r |ii» Mnrriifliiiiiiii(iiiiiiiiiliriiHlr\H, hIiiiII in piiilll«. MiirrnrliiniiH tiii;iiiiilini/ihin. hKiiII limii :iIm)V«. Miifriirhiiiiix iiiiiiiiKlirdHhix. hUiiII ('miii lii'low. Mdvriirhiddn dii'iUKtim/ilriii. lnwi r.jiiw. HISTORY OV Till'] NORTH AMKRICAN riNNIPKDS. 'i'lic l'iiiiii|>iMls, <)!• I'iiuilixtliti, niiliracin^ llic Seals iiiid Will- riis<'s, iuv (■oiiiiiioiily r<'<'(><^iii/«'d by r«M'«Mi(, sysleriiiitiit wiitcrs as coiislitiilinjf a siilxndcr ol" Mic order hWtv, or Oariiivoioiis Matiiiiials. Tliey are, in slioit, ti'iie Cortniutni, iiiodiiied lor an a<|iiatie existence, and have <-onse<|iieiitly l»eeti sometimes termed '■'•Amphihinus ('antirnra.''^ Theii- whole I'orm is modilied for life in the water, whieh element is their true liom«-. Ileie they display extreme activity, lint on lan transi'ormed into swimmin;;' orj;ans. As tiieir riiime impli(^s, they are Jin- footed, (lencially speakin;^, the body may be compared to two cones ioined basally. I'niikethe other niaiine mannnals, the I'imiipeds arc all well clothed with hair, while several of them ha\e, underneath the exterior coars<-r hair, a thick, soft, silky nndcr liM. In contrasting' thetn with the ordinary or terrestrial mammals, we note that the l)o(ly is only ex<'eptionally laiscd, and the limbs are contined within the common internment to beyond the knees ami elbows, and \\\v. hence to only a slight dejirce sciviceabh^ for terrestrial locomotion.^ The first di^it of the manns is generally lenj^thened and enlarj^cd, as arc both the outer dibits of the pes. As compared with other Fcm\ they present, in osteolo;^ical characters, many ob\ioiis poinisof dilVercnce, especially in relation to the strnctnrc of the sknii. limbs, and pelvis, and in (h-ntition. The sknil is ,s hct wcfii tli<- piilatiiic iiiid Iron .ntition, the canines are j;i'catly de\eIoi)ed, soinetiines enoiMKHisly so; the ](»\ver incisois aie never more than tour in nnndter, an-' sometimes only two; the iippt-r incisors usually nund>er six, hut sometimes only Ibui', or even two; the jrrindin^ teeth (premolars and molars) are j;enerally simjile in structure, and usually didei' from each other merely in resi)ect to size, or the number of roots by which they are inserted. The pelvis dilVei's from that of the teirestrial /'V/vr in the sliortness of the iliac portion and the eversion of its anterioi- border; the ischiac bones Itaiely meet tbi' a short distance in the male, and arc usually widely separated in the female, the i»elvic ai'ch thus remaining' in the latter permanentl,\ open \entrally. The existing rinnipeds constitute three xcry distinct minor iiroups or families, ditl'criii;^' ipiitc widely from each other in importaid characfeis: these arc the Walruses, or Otlolxniithr, the Eared Seals, or Ohirilthv, and the Marless Seals, or /'hociila: The tirsi two arc far more nearly allied than arc cither of these with the third, so that the (hlolxcnidic and (Htnilthr ma\ be together contrasted with the I'hochUv. Tiie last named is the lowest oi mo>l .i;cncralizcd yroup. while the others a[)pcar to stand on nearly the same plane, and about e([ually iemot< from the Pho- viiJd. The AValruscs arc really little more than thick, clumsy, obese Ibrms of the Otarian type, Avith the canines «'normously develo]>cd. and the whole skull i-orri'lat ively modilied. The lind)-structiire. tin' mode of life, and the whole economy are essentially the same in the two fiioups, and, aside iromthe cran- ial modifications ])rescnted by the (hlohdiiuhv, wldeh ar<^ obvi ously related t(» the development of thecauines as huj;(! tusks, the Walruses are merely elephantiueOtariids, the absence or presenee of an external ear beiuo- ju reality a feature of minor importance. The characters of the suborder and its three families amy be more foriually stated as follows : * "The (.■liiUiicf'.r.'s Iktc j;ivun uio in part tlioso collated !•>• 1 >r. I'licdddrc ». Gill ill 187:'. (• Ariim;;('iii(Mit of tho rainilifs .>t' llir Mai iiiials.' Sniillisniiiaii Mi<('('lliiiiciiiis ('(illcct ions. \o. ■.>:!(•. )i|i. ."iCi. (H, li",M. l'\ wlioiii I lir dist inctivc t'latnicN ol' llirx uiiiii|is well' lirst loriiiiilati'i!. 'I'licv Jiav; , liowcvcr. been ansncjaiiil. CIIAIfACTKHS (»l' IMNMITlilA. Limbs i>iniiil'iii'iii, cir iiinilHii'il intn >.\viiiiiiiiu;;' nruiins, ;iiiil ciirlnsril li> ni Ix'yitiiil III!' I'lhiiws ,'tii(l ktii't's witliiii I lie I'lHiiiiiiiii hiti'^iiiiiriit. Dij^ilsiil' till' liiiillM.s tlcircii.siii;; ill Iclljilll illltl si/c iVmii t Ik' I'iinI to tilt' lil'lli; III' lliiisi' III' tilt' |M's, the lii'Nt, iiiiil lit'tli l;ir;;i'xt iiliil lolly•('^*l , llir llircc iiiiililli' mil's .slioitiT iiiiil Miliri|ii;il. I't'lvis with till' ilijir |iiiitiiiii MTV ^limt, ami tin' .iiili'iioi- Ipnriji'r iniiili cvri'lt'il; isilijii liairl.N iiii'i'l iiiji l(\ a sliiirl >viii|(li\.siN (ncvrr liucliylosrd) anil in tlir IViiiali' iisiiall.v wiilrly .sr|iaiali'il. Skull P'lit'rally ;;ii'atly cDiiipirssi'il iiili'iitrliitally : I'arial |Miitii(ii iiHiiiilly sliiirt anil ratliiT liri>ail, ami llir luain-iaM' aliiiiptly t'X- pandi'il. liarliryiiial liuiir iiii|M'i'tbi'iitr ami Jiiiiii'il In tlir maxil- lary, cncl'isi'il wliidly williin tlir nlliit. ralatiiirs usually si'pa- ratril liya variiil.v, kIIiii nrnuisiilcralili' si/c rriuii llir tViuiials. Tyiii|uinic lioni's si'parali'il alsoliya varuityrrom tlii'i'XDccipil- als. Di'iititinii sini|ilr. jii'iH'i'ally unsprriali/ril, till' imdais all .similar in striii'liirr. Itii'liliiuus dcnliiiiin riiiliim-nlary, iii'vcr truly luni'tiiHial, and ^riirrally not jit'isisti'iit ltr,\iiiid tin- ln'tal slaj,'r id' llir animal. I'rrniani'iit im'isius usually !; lu |. snmi- times .] {('fiKUti'liiint and Miifi'inliiiiiis). nr rviii > ( Diliilm iiiik); raniiics .; ; nndars- ;, ' , nr '; I 'INNiri",! )rA A. llimi li';^sr;i>)ialdr id' Idillii I llltird tiU'waid and llsiil in li'l irstri.'ll jdi'o- miiliiin. Nick lrii;;l liriH'd (isjii riallv in laiuilv ,1 1 i. .^kullwilli till' masliiid proii's>i's lar^i- and salient ^espeeially in I In- mult.'S), ami with disiinel .'liispliennid eaiials. Anteiinr I'eei nearly as larjie as the )iipsteriiu'. tlieir di,i;its rajiidly deeicasiii'^ in lenntli I'riim tile liisi Id the lillli, wiilniut ilistinet ela.\s, and with a hroad earl i la;; i noils liorder ex I end in;; heyoiid I he di;;il-. Hind Icct .sileepi ible of Mical expansion, I he I liiee iiiiddii' di^ils only willi e laws, and all I he di;;ifs terminat in;;' in lon^i, narrow, i ar- tilai;iiioiis tlaps. united li.'isally. I'eiiinr wiili the Irocliaiiter ininiir well developed < !lii;,«SK;nAUA. I. Witlioiit external ears. I'orm Ihiek and heavv. .\iiterior por- tion of the skull <>reatly swollen, ffivin;:" support to the enor- mously developed eaiiiiies, wliieh i'orm li>n>^, protrudiiiff tu.slroee.s.ses, ami tlie surface of the mastoid processes contiiiuons with the auditory Imlhc Odolnvnldcc. II. With small oxtonial ears. I'-orm slender and clon'^ated. Ante- rior portion of the skull not uiin.siially swollen, anc' tlio canines not highly sj)i'ci;ilized. Incisors of deciduous dentition -}, only the outer on either side cuttinj^ the j^um ; of permanent denti- tion 4, the two central pairs of tlus npi)cr with a transverse irroove. I'ostorhital ]ii'()ce.sses strongly developed. Siirfteo of the ma.^^i/id jiroeesses not eonfimious with the anditorj'' hnlhe Otarilda: B. Hind li';;s not ca|>;ildi' of heiuf; tiirneil forward, and not sorvicoablo for terrestrial locomotion. Neck short. Slcull with the mastoid •Jr view of the uncertainty respecting the proper notation of the grind- ing teeth, they will in the jiresent work be designated simply as molars, "with noiiltemiit at distingnishiiig "premolars" from "molars.'' CIIAI.'ACTKKS or I'INNII'KDIA. IH'iM'i'.s.si's swiillrii, lull IK 1 1 smIu'IiI, ,\\\i\ w it liiir' ilistiiicl ali,H|ilu'- iioiii 1','iiiiils. Anli'iior iimli.s siiiiiiji'i' ijiim t hi' iioHtriltii', Ihr first. ilit;it litllc, il'iiii\, liiiij;cr tliiiii tlir iirxl Niicccciiiii;; diics, all iU'int'il with Nti'<)ii<; rhiw.s, \\ hicli jiii' tri'iiiiiiiil. liiiiil iVi'i I'a- ]itllil(' of iiioiii I'jitr <'\|iMiiMi()ii, shiil'l : (limits (iiHiiaJl.v) all aiiiinl witii Niroii;;- claws, anil witJKMit icrmlnal laitilayiimiis llaps. I'riiiiii with nil tiarr i>t' the ti'iM'haiilt'r niiimr. . . Iii:i'i iiiijAPA.* III. Without cxtci'iial cat's, rostoi'liital ju'ori'SHcs watititi;;, m' very small. Itn'isiits vaiialiji' (';, :|. i>i'}). lirridnoiis ili'titil ion not |M'i'sislrnl licvotiil I'd'lal life I'limidii. The riiiniptulH prcs'iit ii lii;;li drorcc of cci'dtriil ilcNclop- ment, iiiul arc easily «loiiH'sti<'al('(l under liiv(»ial)le e<»iirs to certain favorite hreedino' oromids, and diirino' \]w .season of re ])rodnction !ea\<' the water, and pass a coiisideraltle period upon land. The ICarle.ss Seals, on the other hand, with the exception of the Sea I'Mcphants, do not so nniforiiily resort to |»articnlar lireedin;; oronnds on land, and leave the water (»idy for very short intervals. They nsiially hrinj;' forth their yonn;; on the ice, most (»f the sp"cies l»«'inj;' conlined t]>ears to be strictly tropical, and very few of them rauj^e into troi>ical waters. As a i^ronp, the ['innipeds ari^ distinctively character- istic- (tf the Arctic, Antarctic, and Temperate ])ortiortions of the Northern iremisidiere. The Otariidw and riiocida', on tlie other hand, are abumlantly represented on both sides of the equator, as will be noticed more in detail later. *l''or the .suKj;osti()ii to my friciid l>r. Kllio. ho terms Grcsmgrada and UcpUgrada I am indiOiteil ^011f.S. FAMILY ODOHyl^.NlDiK. Wl»!r:j;;j:;. '• 'J'rkhnidii, (iJt.w, l.niiiliMi Mfd. K'riius., If^-JI, ItiKl" (I'iirnil.v ). .lj(«(i Gray, Jri(7»((7/((/ri, (iit.w, Ann. ol' I'liilos.. l^•,>^), :tl(i; Ann. and .M.ij;. Niil. Hint., ^d H (laniily). Tiivhiciiia, (iu.VY, London's .Mai;. Nat. lli.st., i, 1K!7, r>:{H ; "Zotil. Krthiis and Tt'n<)i',;r' (.sulil'aniily). Jn pait only, oi('xt. ii, 1850, 'ill; Cat. Srals and WIialcK. H(i(i, :i:5 (snlifaniily). In jiart only _- Trichicina (jiny, 1837. " Triihtrliiild stn Cuiiipodoiiliti, IfiiooKKs, Cat. Anal, and Zoiil. Muh. 18a8, 37." Trirhnlioiilni, (iiKlii:!,. Fanna . '27. (ill, 7(1 ( -.= " TtiihivhUhv MrooUcs, (irrvais").— Ai.i.r.x, Hnll. ,Mns. Comp. Zoiil.. ii. 1^70, *,M. J!iiniiiiili33; Diet. ."^iLNal., lix, l^'JO, Iti.'i (family'). (|i;m;i;ai. ohsi'-uvations. .Viiioiij; llic disliiictiv*' li'adin'.s ol" (lie (hlolnvnUkv iiio the ('iu»iiii()ii.s(l('v»'l(»|)iiM'iit of I he iij)i)cr('imiii('s, aiul tlie <'on.s»'quont gic'iit rnliirjicinciit of tlu' aiilcrior ])(>rti<)n of tlic skull for their reception jiihI sii])j)oit, tiie early losis of all the iiu'isors excej)t the outer i)air of the upper jaw, the; caducous character of the posterior molars, auar vertebra' iciiiaiiiiny' of proj)ortioualely the same lenj;th. \n consecpu'uce of their obesity, the ribs and tile proximal s(>o|iients <»f lhelind)s are lonjicr in the Walni.ses '' than ill tile iOared Seals, wliiU' tlie distal sej^ineiits (d the li.Mbs are iclatively sliortei. The scapula is long and narrow, instead of short and broad, a.s in the Otariida', iwid it.s ciest is ithiced (i FAMILY tiDoll/KNIlM;. irioif iiiilnidilv. Acnii'diii^lN , ill rcspccl to L; iiiid liiiilis in the one r(»litrastC(I with ;^rr;il tliicKiicss ol' IxMly. iiiid distall.v ii dispi »])()rti()iiiito K'diitiidii (>r I lie cNti'diiitics in llic (»tlicr. TIic iiKid stiikin;;" tlirrticiiccs^ liowt'Vci', exist ill llic ciaiiial riniriiclcrs. roiiitinj;' Ikhii the unreal devt'lopincnlor tln' iippci- caiiiiics in ihr W ;il- I'liscs, ;iiid tlic ('oiis('(|ii('iil iiiodilit'iilions ol' (lie facial portion of tlic skull. Ill llic (}hir!!(l(i\ tin- j;<'iit'ial contoiir of tlic skull is stroii,L;l,\ I isint'; in llic (hlolKiiiida, \\ is iiiii(|iic. owiii;; toils ;iicat expansion antcri(»il,v. In respect toolliereianial features, the WalinsesdilVeilioin the Eaved Se:ils in liavinjiiio post-ovb- ital ]»i'oee.sHos, and in the niastoiil [troeesses lieinj;' not . separated from the auditory hiilhe. Tiie teeth are all sin^ile rooted, and iiave in the periiianent dentition no distinct crowns. On coiiipariiiji' the Odolxviiidiv with the I'lioriihv, the ditVer- enee.s in j^oneral .stnieture are Ibiuul to be far ••reater than ob- tain between the Wabnses and I'iared Seals, espe<'iall,v in rej^ard to the hind exti'eiiiities; these in the J'liocidd- beinji directed ba<'kward. and useless as organs of teirestiial Ioconioti(»n. Hence, in s(. .ar as tL<' (hlohwnUUv and ()t(irii(l(v n'^vvv in linib- aiid skiiU-strneture, they both similarly depart from the Phocine tyi»e. As already indicated in the synopsis of the sidxtrdei' Z'/«- nijx'ilid, the I'Ikh'kUv dill'er far more from either the OdolxvnUlw and OtariUUv than do these latter from each other. This ditier- onee is esi»ecially emphasized in the' skull; for whih; tlie Odo- h(vni(l(v and Otariiilw aji'ree in all important cranial characters, aside from the special features correlatf'd with tlu; immense enlar;;<'ment (»f the u])per canines in the former, they widely differ from the riiocida: This is es])ecially seen in the absence in the latter of an alis]dienoid canal, in the greatly swollen andi- toiy bulla', the position of the carotid foramen, and the non- salient character of the mastoid ]m>cesses. The few points in which the Walruses dilfer in myology from other Pimiipeds, Dr. 3Iurie statv's to be ''the presence of a co- I'aco-brachialis, a llexor brevi.s manus, a pronator quadratus, an op])onens jtoUieis, and a palmaris brevis," in the ])Ossession of Avhich it differs both from Otarid and J'Iiock, but tliat in other resj)ects they " muscularlyju'esent general a jireement."' ''('om- ]>ared with the Seals [I'liora .^] there are two extra peionei and a llexor Imca ishallucis." "•Though deficient in concha, the auri- cular muscles aic remarkably large."* I'roi-. Zoai. Soc. 1,(111(1.. 1*370, 1). .')4.'). OMM'.l.'AI. (tllSl-.KVATlONS. i ••( 'nlisi<|»'rill,n tin' Nt'fV (litTcifllt ;lttitll liltic (l(!viii(iii of tlii- iin sclcs of tlir liiiid Icj;', aixl in this rcsiioct himmIc iVdiii tlic Sell, vet Imt slijilitly."* Ill n's]icct to fill' iMisitjdii iuiil cliiiriictcr of till' visc«!ra, a yen- fill! a;ii'tM'iiifiit lias Itcccii iiutcd with lliosc of tlic other IMiiiii- pcds, and tlicv i»icsciit iiolliin;; that calls lor special notice ill tlic present connection. As Dr. ^Inrie lias stated, tlu-re is littlo api>recial»le dilferciice exiiihited tlironj^lioiit the IMnnipeds in the construction of the alinitMitary canal. " It is .siiiij)ly that of a Carnivore, with. lio\v«'ver, a moderate-sized ca'ciini. The great ylaiidnlar snperticics and correlated large lyini>hatie.s point to means of sjicedy and freipient digestion; and in the Walrns these apparatus are extraordinarily develoi)ed.''t In a<*cor(lance with the characters already given (p. .'i), if any snbdivision of the Pinnipeds into groups of higher rank than families is to be made, it seems evident that the OdohaunJcc and Otarlidn' are to be collectively contrasted with the PhocUlw; in other words, that to nnite the Otarildw and Phocidw as a group of co-ordinate rank with the Odohwnuhc is to lose sight of the Avide ditterences that sei)arate the two first-named fami- lies, as well as of the many important features shared in com- mon by the (hlolxviiidw and Otarndtv, by Avhieh both are trench- antly se])arated from the Phocidw. Although the Walruses are now \ery generally recogniz«'d as constituting a natural faunly of the Pinnipeds, ranking co-ordi- nately with the I'^ared Seals on the one hand and with tho Earless Seals on the other, the affinities of few groui)s have been more diversely intcrju-eted. As early as the thirteenth c«'ntnry. the author of the"kSi)eeidnm Regale'', — one of the earliest works re- lating to natural history, in which the Walrns is mentioned, — stated distinctly that the Walrus Avas an animal closely related to the .Seals: and wetin^7'i. p. 4()I. 8 FAMILY ODOH/EXlDyi:. llicm with forms with whicli they hiul no lohitionshii*. Tn the ii'fuiR'v of scioncc, nothhig was perhaps more natural tliau that an'inals sliouhl be chissitiecl in accordance Avith their mo(h' of ]ife, their liabitat, or their external form, and w<' are heiu-c not surprised to find ihat llondelet, (lesner, Ahh'ovaiuhis, Joustou, and other pre-Linna?an writers, arranged tlie Pinnipeds, as well as the Sirenians and Cetaceans, with the iishes. or tiiat other early Avriters should term all four-footed creatures "Qiiadin peds,*' and divide them into "Land Quadrupeds" and "Quadru- peds of tlu' Sea." While all marine animals were by some early writers classified as ''fishes,'"* the Pinnipeds were much sooner «lisassociated from the true fishes than wer«» the Cetaceans and Sirenians, the mannnalian aflinities of which Avere not at first lecognized by even the great Linne himself, who, as late as the tenth edition of his "Systema datura'" fl75S). still left them in the class •• J'isrcs." In view of the several exceUeut descri])tions and very credit- ,d)le figures ol' the Atlantic Walrus that ap])earcd as early as the sixteentli and seventeenth centuries (a detailed account of which will be given later), it is sur])rising iii;'t the early sys- t^ematic writcis sliould display such romi>lete ignorance of souu- of the most olnious external characters of this animal, as was Jiotably the case with Linne, Klein. Brissou, ICrxleben, and Gmelin, who strangely associated th<' AVidriis and the ]\binatee as niend)er> of the same (/eiiKs, ami groujied them with such diverse creatures as Sloths and Elci)hants. Linne, it is true, in the earlier editions of the '' Systema Xatura>,'' placed the Wal- rus with the Seals in the genus Phoro, in his order Fem\ — a near hit at their true affinities. Later, however, foil wing probably Klein and Brisson, he fell into the grave error of r.'nioviiig tluMii to nearly the most unnatural association possible. In this con- nection, it may prove not uninteresting to sketch, in brief out- line, the strange history of the classification of this singula)" grou]) of fin-footed Carnivores. .\.s already stated, l^inne's first allocation of the grou]» was the natural one. Brisson,t in IToO, led in the long role of error by forming his third "'order" of mammals of the lOIcphant, the *Mosf modfi'ii l!ii)<'iiii!L{c8 still ivtiiin nlics of (liis iincii'iii cnstoiii, as «'\iiict'ii, for ('\;iiiiiilf. ill siicli I'hi^'ii.sli words ;is s7((7/-//.s//, crdii-jisli. irlidh- Jlslitri/. >ic(il-ji>flu'rji, etc.. wliilc hrdljifili (Swcilisli), ividrixclt (Djiiiisli), irallJlKk (GiTiiiiiii), ftc.. iirc coimiioii Ncrnariiljir iiiiini's jiiipiii'd to Cclaccuns. tRi'^nr Aiiiiiial. I7.")(i, p. t-'. (!i;m:i{Ai. ousKiaATioNs. 1) NValius, and llic .Manatee, tlie two h.s\ iiaiiKMl eoiistitiitiiiu Iiis •• j;euus Odohnni.s." This was a iiiaiki'd i<'ti(K'es.si(»ii IVoin even the system of Ivleiii,* of a lew years' eailier date, w1h» l)ioiitilit to<;etlier as one family tlic Seals. ( )ttei's, Meaver, NN'alrns, and Manatee. Linne, in 17(»<1.+ not <»niy removroui)s than ;;('nera. plaeed the Waliuses and Seals tojiether im- uiediately after the ('avidvores. Schrel)ei',§ at about th«? same date (1777), ad»»j>t<'d a siudlar ehi.s.sitication, t!u' Walrus .staud- iuy next aftei' the Hlejthant fiid lU'.'cedinfi' the Seals. Scliro- l)Oi*s ;;enus 'I'ricliirliKs contained also the Dufionj;' and tlio Manatee. ris.son to (rinelin. standiu;;' next to the l']le[»hant, and as.sociated yeueric- all\ with the Siiciuans. r>lunu'nl)acli.'| from 17HStillas late even as ISi',"). still arranged the Walrus and the Sireniaus in the ucnus 'rriclii'clnis. in other res|)('cts. the Walius appears with new associal"s. the ji^cnus 1', IciiccIiks Iteinji' uinted with Oniitliiirln/iirlnis to form a "faniily"(I) of his "(»rder" I'abiintd. The order I'dliiiiiia. as the name implies, was composed of the web-footed manunals, and divided int(> three •• families," namely, "A. (Hires" (eonsistinjiof the yenus Castor); -']>. Fcrw"' {I'lioco and Liitra); and "C. Jlrnfii" {OrnHhorlii/iicJiii.s and Trichechm). Tills is essentially also the ariauyvmeid j»ro]«)S(;d by Klein in 1751. The lirst step toward disinendteriiij^- ;.! divided the gcuu-s Tricluchiis of former autluu-s into throe yenera, namely. ManutKN, for the Manatee; JIifdioiiKtlis. for Steller's Sea-Cow {= lihi/tiiiH Illij-er, ISll); an. lit. tSyst. \li"^. Aiiiiii.. 1T77. p. .".!»:!. vNSiiiip'di.. ii. [I77t'..'l. 1.. v'lio. II Sy.';!. \!i(.,i, .-.!). ^Hiindlt. <1. Xiiliiriifscli.. I7"<-, p. II-.', and lalcr iililimis. "Koiigl. NrinivK. Acad. ii,\,i llaiidliii;;,. \\. I7'.M, iip. •J-r)-;!0(l. 10 lA.Mii.v (ii»(»r..i;Mii.K. I [ \\\ii Iiotli llif \\';iliii> iiikI llic 1 )ii,niiii,u. NVliilc lliis \v;is in tilt' iiiiiiii a iiiosi iiiipdi taiil ami iiro.mc -^'x e iiiiio\ati»iii. Kc; zius sociiis t(i \i;\\{' laltdi'i'd, liUt- >('\('ral still carlici- wiitns. iiiidcr tlic iiiipicssioM lliaf tlic Walrus, like ilic iMi.^oiiu. /""/ /"< hind fat. ( )/.('i('tsK()\ sk_\ . * alioiit a year later. aiil\' iuiKiiaiit nf li'ct/ius's [>a|M'|-. also placfd. as cuiioiisly liapiiriird. ilic Walrus and the hii^diiu to.iActliff in tlic ufims Triilucliits, iMM-ausc lie siii)ji(>scd the Dw^owj: lii((l hliid Jrct.Wki' tlif Walrus I These euiious aiit it lift ieal mistakes iiitlieate how little was known Ity systematic wiiteis ai»(>ut tlic stiiictuic ol' these ani- mals as late as the close of the last .s'/M«r«.v"; '•!'. TricluTiis i and the Manatee, to which he gave the generic names lespeetively of Phiti/sfonnis { = ll(ili<-uri . niiger, 1811) and OxystoiiiKs { = M((nafii,s. Ifetziiis. 17!»4). lea\ in;.; only the Walrus in TriclurhKs. The genus TrirlKTlins. however, as first instituted by Artedi (IT.'W) and Liniic (ITriS). as will lie shown later, did not rehite in any way to the Walrus, being applied exelnsively to the ^Fanatee. It was not till 17«>(i that the term was lirst made to cover both the then known Sirenians and the "Walrus, althongh the iMubroilment of the two groups began with Urisson, ten years eailier. The l*innipeds and Sirenians, collectively considered, were lirst separated as disthict groups by illiger§ inl.Sll, who raised them to the rank <»f orders, they forming respectively his orders Piiiiiipcdut and \(it(iiitii».. xiii. KDti. jij). :{:i-:i7ri. t'l'illll. l^I.'IM.Mlt., \>. IT','. t Diifi Natioiiiii-.Ntiiscmii (Irr Xatiirj^i'scjiiciih', li. IS(i;!, [vp. iMl-lt.'jS. V^ riixhiiiiuisSvstciiiatis.Maiimialiuiint Avium. l?'lt,iip. Uri. V.VJ; Aliliamll tier Akacl. Wi'^scnscli. /ii Itcrliii. ISOl-l-'JI. (IKJ.'.V i>lt. :!H !:">!•. imsnim. CENEUAL OIJSEHVATIONS. 11 (•(iui\ iilfiil ill extent with the order Pinniimlla. The i»i'0[niety of the eliaiiyes introducetl by Illigcr was not speedily leeognizcd by con temporary writers; (Javier, and many subsequent syste- matists tor hall' a eentury, ])lacin,L: tlie Pinnipeds amonn' the (jtinirora and tiie ttirenians amon.n' the CtUicca, with the rank respectively ol" I'anulics, the lanuly P//oc/(/ft' einbrachiy all the Pinnipeds. Dr. .1. 1'. Gray, in ISL'l,* and af:;ain in 182"),! widely separated the Walruses from the Seals as a laniily, Trichcchida', which he most stranjicly placed (together with the Sirenians) in the order ('t lier I 'inniiieds. Wagler,** in 18.'{0, made the Walruses merely a genus of his order Crsi. Nilssou,tt in 1837, divided the Pin- nipeds into two sections, the second of which embracd not only TriclurliUN, but also JlalichaiKs^ ('!iNto2>hora, aiul OtarUt. Tur- ner, |f in 1848, from a study of tiie skulls, separated the Piiiiii- l)eds into three natural groups, considered by him to hold the rank of stdifamilies, namely: Arctoccphalina. embra<'ing Otdria and An'toccphuluN ; Trichcciua, consisting (»f the genus '• Trivht- c«6'"; and ritovina, embracing all the other Seals. Gill, §§ in 1800^ * " London Mi'd. JJcpos., 1821. p. :?0'2," upiul (Jniy. t Annals of riiilosopliy. 'Jd sit., vol. x. ia.'.'>, p. niO. t Eoiidoii's Miifj. Nat. Hist., vol. i. p. '>"':!. v> I'ani. Rt'g. Anini.. \). .M. !| Diet. Sci. Nat., t. lix, p. 307. *'. "t'al. of Ills Anatoni. and Zoiil. Mns.. ]i. ;{(!." aiiud (tint. **\aiiiil. S.\(st. Ainpli., p. '27. ttVrtcnsk. Akiid. llandl.. 18117, !i:{i'>; WiP};mann's Aiob. f. Natnrg., 1811, jt. :i(lO(transl.) ttl'mc. Zoiil. Soc. I.oik:,, 1848, pp. 85, 88. VH^ I'ldc. I".ssc\ Ins|if!ii('. \ol. V, 11. 7. rJ (iKMI.'A oi rill. lA.Mll.V. \\;is tin- IH\( :iiillior \\Im» r(r();^iii/('(l I Ik- \\ iilriisc.s iis loiiiiiii;; a disliiict lUiiiilN , w iiicli lie h riiicil lioxnmriihi . lit I liis step, lie \\;i,s iiiiiiir(li;itrl\ lulluwcd l>,\ ro ordiimlc in niiik with his '■ /'Af/r«/Wr«," roiisistin;; of tlic I'Iiiu'kUv :iiiiI i>l((t'iiilties of hotli tiie I'iiinipeds aiil., :',. ('(Plllp. /cic'il., \ip|. ji. |). '.'1. I I'.'IIIIIM . •>'.•. ||\:iii I'lrmijiii, Ami. Mm-, illli^l. NmI . ili iii'lt;ii|iic. i, I-T7, |p. Till. J. (iKNKIUI, OUSKUVATIONS. caiiiiH's were roiiiiil :it . \ii\ ns, l»iit \';iii Knu'dcii i.s .s|i(»ii;;l,\ (»!' the ii|iiiiioii tliiit IIh' It'clli (l(',s( rilx-d l»\ l{ii.\ liiiiikcslcr,* Irmn the Ii'rd < 'ni;; <»(" I'lii^iliiiid, ill IHJJ."*^ iiiid iiiiiiwd Triclircliothm li".r //■///, ;ir<' I hose of his ,\li((lliiit ramus. The other hones hi'lieved to represent it are nine \erlehra', |»art of a pel \ is, a hinnerus, a femur, several metatarsal, meta<-ai'pal, and phalangeal hones, etc., ainl part of a tusk. Says Van Meneden: '• I lie lnanchede max ilia ire est tout ce < pie in ills possedoiis de la tele. liCS dents maiiipieiit, liiais le h(»id esl asse/ coiiijdel, pour tpi'oii piiisse hieii iii;.;er de leiirs caracleies par les ah i-oles. Nous poiiMins, (111 reste, fori hieii aiissi apprt'cier la forme dt; cet OS, dislinj^iier sa symphyse et sa hrievcte. •' l/os est luise a son c\tn''iiiile anti'-iieiire, la syin|(liyse est, fori cniirteet Tos ii'a pas plus dV-paisseiir siir la, li;;ne mediaiie que siir le coh'-. Lesalveolessoiil compaiat i\ emeiit (hit ;;randes: h's trois dernieres sont. a pen ]>re semlilahles, ranti'-iieiire est, la plus petite. ( "est Till virse dans Ic Morse, liaca nine (leva if «'tre tort ;;raiide. II iTs a ipi'iiiie settle alvi'ole |iiiiir ane dent iiiei si\e. '* \a' i(ii]>s (111 maxillaire esl remai<|iiahle jioiir sa coiirhiire. Toui(!la ]iarlie jiost(''rieiire, (pii const Hue, la hranrhcdn maxillairi^ maiKpie. On xoil siir la face e\tern(; trois Irons mentonniers. -> Mil coiiiparanl ce masillaire a celiii dii M(M'se vivaiit, (Ui Noif ((Ue la SMiiphyse est loiite ditferelile, (jii'il e\ist(^ line ;;ran(le ahcole pour la dent caniiie el (les traces d'line petite al\(''oUi pour line iiicisi\c ipii restait prohahleineiit (•a(!li(''(! sous les j;(,'U- civcs. Dans le. iMors(^ vivaiit, il try a pas de place jioiir line canine |yraiide| an maxillaire, infc'iriciir." • Sec hiVDlid. p. arts of AUuthcfiuui. Van Hencden's descriptions and liyuri's of the lower Jaw Iraguu'ut indicate features widely ditferent from those of the coiTespondiny part in the Walrus, especially in the shortness ol the symphysis and in the curvature of the i)art represented, but above all in the nuMd)er, rcLitiv*' size, and foriu of the alveoli, and particularly in the larjjic size of that of the canine, which must have been almost as hiyhly specialized as in the Sea Lions. That the tusks referred to it by Van IJencden (those described by Lankester especially, as well as the frajiuient he himself fig- ures) beloiiji' here, there seems to be at leasi ro(»m for reason- able doubt.* The dilfcrences presented by the Jaw fragment of Trichechotlou as compannl with the <'orrcs])onding part ol Aladheriitm arc c\cn still Juorc marked. The more obvious characters distincti\c tif the three genera oi \\vii OiUthanidn . a^aX picscnt Iciiowii, may Ite luietly indicated as follows : Slliiiijiniy (if //(( ivi'ly to the Walrus in a generic sense). — IJr.issON, Kegnc Anim.. 17.">*i, 43 (used strictly in a generic sense, hut emhracing ■•!. t.a \'aelie marine — OiloheHHn" = \\;i\v\\s: "•,'. Le Lamaulinc — Mdinilii"." The characters given apply almost exclusively to the Walrus). *Van Beneden himself says: --M, U'ay I.ankaster avait vu en Angleterre diffdrcutes grandes dents, i)rovenaul duii Inu'hiii h'ay I.ankaster" as a syno- nym of Trkliiihodoii liOiiiitncliii. dc.scrilpcil l.y himself much later! In view of the iiueertaiuties of (he ease, it is to he regretted that Ik; did not ])ropo5e il new generic as well as specific name lor his 'iv'uhivhiuhtii hnnihirlii. ♦ With refereiKe only Jo the lower Jaw, tlw only ktiown )iart, in case of the extinct tyjies, readily susceptilile ol' eomiiaiison. SYNONYMY AND N< )Mi:NCI,ATri{K. 1; ;> Diluhiiiiiis. MAi.M(ii;i;N,<"»rv(rs. K. N'.t. AU;iil. Forli. IHti:i, (iHlUj. i;!ii. ]:,>s,iiitni.-<. Ki.i:i\,(^ii;nl. Dis-i. I{icv. Hist. Niil., IT.M, M\'M (;iii]>lir(l in a ;;cii- fiic sense exeiiisivcly to llie \Yiilnis), — "S ui'oi.i, IntiKil. llisl. Nat., 1777. —." — (iii.i, ("ex Stdpoli "), I'loe. Kssex Inst., v. If^titl. 7. I'lioni. I.inm';, ."^vst. Nat.,i. 17.> (in pail uniy). Trichi 'litis, I.iNNi';, .^yst. Nat., 170li, 4i>(iniiarl (ini\ ; ni(| of Linne, 17riH, nm Artedi, I7rw ; liaso of the )iresent eentiiry. (idiihiimlhiriinii. (in.vTiOLlCT, Ihill. Soe. (ieol. (h> I'ranee. •," ser.. xv, 18")?. (I'J4 ( = ■■ Tiirliirlnin ri>siiiiirnx'' anet. — fonnded on a supposed fossilV Oilmitobinins, Sfxi»i;\ ai.i„ Ofvers. K. Vet. Akad. Forh., 1-.V.», 441. .' I'll ell I'll oilim, l..v\Ki;.srKU, (Quarter. .loiuu. Cicol. .'^oe. Loud., xxi, Ifftio, 'J-iti, pi. X. xi (hased on I'ossil tusks from tlie K'ed Crajj-, i'.n^iland). Tlic iiiiiiH- 'rriclin-liiis, lur so Imiii' :i tiiiu- in m ficnil use loi tlif \V;iIiiiscs. ]»>ov<'.s iMd, iis loiij; ;|o(> sliuwii l>v \Vi»'o||i;uiii,* von liiicr. .Miillcr. Stiiiiiiiiis. ;m«l liitcf lt\ idlu r "viiti'is. to Itt'loiij^' at .ill to llicsc iiiiiiimls. lull to tilt' .M;iiiiit<'('. The iiiiiiic Trli'ltirlnix oi'i;.;iii;itiMl with .\rtt'tli in IT^IS. insi posilniiuoiis worl^ t clitcd ii\ li:siiiii iiniM'iinc. Fistiihi . . . . " 'I'lic cita lions nntlcr TriilurliKs fiiilnacf no allusion to the W'aliiis. luii i«'Iatt' wholly to Siicnians, of to tln' .Maiiatcf. a.s the latr«'i' was tlnii Uiiown.T Artt'. llC. tlehthyolo-ria, 1738, pars i, ]). 74; pars iii, \>. 71>: pars i\, ]>. Itlil. In Arie- di's work tile name is twice written Trkhrcliiis ami twii'c Tliricliirlimt, On p. 71 iif pars i. wliere it lirst occurs, its deri\alioii is given, namely : •• Trhln- r'nin ;i \^iti^ rriiiii .J'' i \;fl<)c piicis ipda solus inter pisces fere hirsulus sit." : The references in a eenerid way ajipcarto include .'ill the ."sircnians then known. v^']. g.. "Deniiuin duo utrini|iie I'mimiil. louuit udine :>pii liani;e crassii u poUicis." m u; 'llll. (il.M S Mixili.lAI S. <<(iii\iil(iil lu I he T/Vr ul Linin- (S\.s|. N;il ., •il. \, I 7.">Sj. I.iiiiif, ill I7.")H. liisl iiitindiirt-il AltcdiVs ;;fiiiis Tritli)flnis, ;il wliicli lime Ik- plmc*! in il uiilvtiic Miiiuitcf, I Mi;;()iiu, iiikI Slflh-i's Sea ('(»\\, I<-:i\ in;; t III' Waliiiscs still in /'linen. Mi,, (lia^iimsis of till' ;;ciilis* ciiiliiactMl mtiic ul' the ilisi ilirt i\ c rliaiarti) > nj llic Waliiis. Ill I7«I«; (iLMli «■l\ aMcrilii'd to tlic Walrus in loiiiH-r fditifuis, when I In- Walinst ; wi'ii- plan-il iiniii'i' I'linm. Ilnicc, to uliatcvei tli<- ;;fii(iic naiiif I'lichvclms iiia\ he ic|'rral>l<', it r»itaiiil,\ is not prrtimnt to tlic Walrus. 'I'liis liciii;^ scltltMl, tin- ipn-stion arises. What jfi'iH'i'ic iiaiiic is oCiiiHpiestioiiaMc appliraiiilit.N to the Walruses .' Here the real iliMienlt.N in the tase he^^ins, tor aiilhors who ailiiiil the inapplii-ahilitN ot' 'I'rifli clnis to this ;,iioiip are not agreed as to what shall lie snitst it iite<| lor it. .Seandinav ian Avriters, as .Maliii,i;ren (Isiilj and lalljehorj,^ (|s7l;, ;iiid I'eitrs ( JSOI j aiiionj;(Ierman ant horil ies. Iia\ e lor some \ ears eiM|iIo,\ ed (hlolxniiis, a name apparent l\ ori;:inat in;^ with l-inne (as Oiluht iiiis) in 17."!.'», and adopl<'d in a j^eiieiie sense li\ Itiisson in I7.'i(i. A iiKdlilied I'orm ol" it ((hhnihihitniis) w as als(» emplox ed li\ Sun de\aliin I.S,"i't. (lill,in isiKi, and other recent Aineiiean wrileis. have l»roii;ih( into some pidiniiieiiee t he name 1,'iisiiiiirii.s. \\\si ii.sed in a ;4eneiie isense li\ Klein in I7."»|, li\ Seo|M»l: in 1777, h.N l*allas| in I.S.'JI, and In liamont i in Isilj ; while the ;:;reat mass of Mn^iiish and ('onlineiital w riti-rs still elin;.; to '/'riclicilnis. 'Ilie ;;eiiera (hl(tl)iiH)lli( rlinii and 'I'riclicrliixluH, based on fossil ri^iiiaiiis of file Walrus, iiavealso heeii reeeiillv introduced into 1 lie literal lire of the sill ijecl, the former li\ (Iratiolet in ls."»,s. and the. lafter li.\ l,aiilvesler in ISIm; Iml these (espe tlielirst) *"l)riillil:iiii. nmlaii-s r\ o-^c lii;ji;iiM> III riii(|iii' iiilriiiiN (Inn. l.iiliJM ^iiiiiiil.'i. I'cilis ikoIci inns ((lailiiniil i ill |iiiiii:iiii." \(/«/. .V((/., 1(1. \. i, 17.".'', p. lit. t 'I'lic. sec I III (I (liM^iinsis dl' I'rii llll liiiK is, in In II, :is IfilldU > : •• ilcnii s |ii lino rcH niilli nljiiii|ni . I.aniiiiii .sn|ii rinics snlil.irii. .Mnlaics i \ ns.sc iii^nsn lilliminc : iiili rins duo. I.iiliia ^cniiiiala. i'lilis |in.sl( rinics (:niii|i('ii( .s cn- iidiiiiali ill piiiiiaiii.'" — Ai/f/. .\V(^, ed. sii. ITiK'i. i, |i. If". ;y,(«il. l{:iNS()-.\hial., \nl. i. •Jfi'.l. ii\S('ilsn!i.s willi llic Scjl linrscs. |i|i. Id. IIm. ■«! 'nil; si'KciKs OF tiik gknijm. 17 ii]t|M-iii' to Im* ictnTiil)!*' lo the «-\istiii;; \V:ili'iis<-s, and of ciiiiisc, IXTOIIM' lllcr«>l,V HVIIOII.VIIIM of (Ml'licl- IIIIIIICS. ( 'oll.SC(|IM'llt l,V t Im) clKticc, rvidciilly lies Ih'Iwccii (hli)h(viiits:\\\\\ h'nsitianis. ihhha:- viis liiis sixlrcii \<'iirs' piiorilv over liosiinttiis, if we ^o Itjirk to till' riii'lifst, iiitrodiictioii of IIk-sc iiaiii<-s into svstfniiitic nonicn- flii.lMi'f.* It is tin*- tliiil liiisiiiiirii.s \\i\s{\u' *':ii\U's\ liiilin nanio a|i|)li<-i'(;lian;;cal)l,v witit Mots and Mnrnii.s l»y Olans Ma^^nns, (h'simt, Ih-ilMTstain, anil {){\\v\n^ hill onlji ill (I rcniitvnlar sciisr. Altlion;;li used l»\ Kh^in systrniatiraliy in ITr*!, , as did also i>riss(»n in I7ri(j. 'I'lie. wliolo «pU'stion tnrns on what siiall Im- considerrd as the proper start- in;4 |ioint lor ;;enerii- nonienejatnre, ahoni which opinion is still dixidrd. ir the early ;^i-iieric names of Artedi, Klein, llrissori, ami liinne (prior to J7r)(',cics Oiliiliiiiniit, Liiiiic, "l)i;;ili ;iiil., |Mist. .'>, )i;iliili|M'S. \iiis>i Mormin, Di'iitOS iiilciiiicilii Hii|)rriiiiiH Iminissiitii.''— .s'//«/. AVt/., \7'.i7) (cd. fir), f/.t. — Jiosmarua, Kli ill, (^iiii.l. I)i:.ii. r.ri\. Ilisl. Nat., 17r>l, 40, 1)'^'. till :icr(inl;iii( T willi (ii.sldiii ill Kiiiiiliir riisi'H, llin iiiiiiD'. i>l' I In-, riiiiiily 1)0- coiiii's thlohii iiiild', — Ml illur HiiHiiiariiUv, nor Trlchcchiilw lining tciiiiltio. Misc. Tub. No. IL' li 18 Tin; OKM'S (>IK)H/KMS. I ,.1 with those ol' th(! (iiilph ol' St. Lmiirmr. The tusks (if those of the I'l'o/eii iSea arc iiiuch loii^^ci'. uhmc sh'ii(h'r. iiikI liavc a. twist and inward curvatiiiv. '* iShaw, a few years later, tlioii^iht that tlie W'ahiis (h'seiihed ami liuiiied in the aeeonnt of Cap- tain Cooli's last \(»ya;;(', Ihonnh perhaps n(»l speeilieallv distinct. I'loni lh(»se of tlie Arctie shores of llnrojie, slmuld he regarded as l)eh)nj;in;;' to a dilferent \aiiet,\.t lie appears, li(»wever. to Lave based his opinion wholly on li^^uresof the animals, and par- tienlarly on those j^iv-en l»y ("ook and .lonston (the latter a copy of lierrard's, at second-hand fi'oia I)e l.ai't). Illiucr, in isll, formally reco;inized two s]»ecies in his '• rehcrltliek der Siin;;;- thiorc iiach ihrer \'ertheihiii,i;- iiher die \\'elttheih>,"| namely, Tnvhcchux >v>,s;/ho'».s', occurring' on the northein shoresof (West- '.ni ;') Asia, Europe, and North Ameiiea, and 7'. o/>r.s7/,v, occur- riufi' on the northwestern shores of North America anil the ad- joinin*;' iiortheasteru sbor<'s of .Vsia. While I do n()t lind that he bus anywhere j'iven the distinctive eliaraet<'rs of those two species, he, in the abov»'-cited ]»aper, also nauu'd the ainnial described and li<;iired by Cook, T. (livvtujiUN. V. Cuvier, in ISiT), ill describinji' the dent it ion of the ''Morses," says: "Ces dents out ete decrites d'apres plusieurs tetes qui senddent avoir ai)i)artenu 11 deux especes, a en Juj;er du nioius par les proportions de cpiel- ques uues de lours parties, et noii seulenient par lY'tendue de lours defenses, caraetore (pi: iiv.dt d(^ji\ fait soup^ouuor a SUaw * Arctic Zoology, vol. i, 1792, pp. 170, 171. nF<> says: "An excellent represeutalioii is also given in pi. 52 of the Inst voyage of our illnstrions navigator, Captain Cook. It is easy to sec, liowtjvcr, a reinarkalile ditlerence lietween the tusks of this last, and those of the former kind Hgured in Jonston, and it clearly appears, thiit though this (lilTorenco is not such as to justify our considering them as two distinct species, yet it ohliges us to remark them as v.arieties; and it should seem, that, in the regions tlien visited liy Captain Cool<, viz. tlie iey coasts of the American ctrntinent, in lat. 7(», the Walrus is found with tusks nnicli Icuigcr, thinner, and iar more sharp-jiointed, in proportion, tlian the eiiiii(!i'iiK, and 7'. cndh-i. Tile llrst (V. roKinarus) was juineipally eliMi'acteiized l>y lia\ inj,' di\ cijiini;' tnsks, idtont as !un,u' as the lenu'lli tit'tlie whole he;id, tiiiMll\ grooved on the outside, and with two distinct ^iroovcs on the inside: by the ])()ssession of live Itat'k teeth, the last iwo very small; l»y t lie lower ed^c of the uasal openin;;' lieinn but little produced ; lt,\ the occipital crest heinn' stionyiy developed ; and iiy the yicat specilic i^ravity of the Itoiiesof the skull. The second ( '/'. UtnijUhnx) was piiucii)ally characterized hy the tusks equallinii'or ex('eores, p. 23.">. tWicf^mann, in coiniuonting upon Fremcry'.s supposed spccifu' tlitlcvonces, observes as follows respecting probal>l(» sexual and individual ditl'erenceft in the tusks any Fn'iiiciy as distinctive of several species, iiiid alter nieiitioiiiiif;' at leiinth otlar features of varia- tion observed by biin in a eonsiderable series of skidls, describ- inn several of Ids sj>ecirnens in detail, and arriving;' at< the con- clusion that up to that time all th(^ supposed sjiecies of Walrus constituted really l)ut a sinjile sjiecies, a(hled another, under the ajtpropriate uaine TrichechuH lUibim. This with sid)se(|uent autintrs has shared the fate of Freniery's si)eeies,* beinj;- (fonsid- ered as based merely on individual variation. As will be more fully noticed later, two nonunal species have been foiuuled on the fossil remains of the NN'alrus, namely, Tri- chirhiiH viniiniduits, DeKay, ISlli, and Ihlobvnoihn'ium larte- tiaiiiii.. of Gratiolet, the former based on remains from Accomac County, Virginia, and the latter on n^mains from near Paris, France. LanUester, in 180"), added still another, based on tusivs I'rom the. lied Cray of Knyland, under the na/ne Trichc- chodon hnxhj/i. Dr. Leidy, in ISOO, in a pajx-r on fossil remains of the Wal- nis from the, eastern coast of the United tStates, a<;ain noticed the dill'erences in the s'v/.v, leuj^th, and curvature of the tusks in specimens fronx the northwest coast of Xorlh .Vuu'rica and the ((unmon Walrus of the ZS'orth Atlantic, lie says : ''In the course of the precedin;;' iuvestiji'atious |referiin,t>' to previous portions of his i)aper], I was led to examine ;• specimen, in the cabinet of tile Academy of Natural Sciences [of IMiiladelpliia], consisting;' of llie sluifed skin of a j)ortion of the head envelop- ing' the Jaws of a species of Walrus apparently ditferiu";' from tl'e true 'I'rIclHTKN ro.siiianis. of whii "i, as characteristic, 1 have viesved tin- ti;;'ures of the skull and skeletons as ^iven by Dau- benton, Cuvier, and De Blainville. The specimen was pre- sented by Sandwith Drinker, Esq., of Canton, China, and was I)robably derived from the Asiatic shore of the Arctic Ocean. From the worn condition of the upper incisor and molars, it appears to have l)elon!;'ed to an old individual ; and in the case of tlie lower jaw, the teeth appear to have been entirely worn out. The tusks are verv nuu-li larger and arc narrower tlKin in 11- *^Giubcl, ill ld55, refcirctl to I remoiy's and St.iimius'.s species as still need- ing cf)niirniati()u: "Die von ri/iiicrv nacli (lev licsclial't'ciilieit dcr Ziilme uuterscLiedenen Ai'teii, 'Ti: longidcim mid Tr. Cooli; siiid liingst als unlialtbar erkanut wovdeii luul audi die von Staiiuius uiil" SuliiideldiD'crenzen begriiu- detc Art, 2V. dubiiiH, entbelirt noch der wcitoru Bestiitigung." — Siiugcthicrc, p. ViS, footnote. t-.'l Tin: si'Kf'irs nv the oenus. 21 the 7'. ro.siiKirii.s, niid llicy ciu-vc dowiiwiird, outward, and in- wiinl, instead of contiiiiially divi'i'j;iii<;' as in this species. At tlieir eMiei'.ncnce tVoiii '.lie aheoli the tusks are Iwo and threo- «|U;utei' inelies ajtart, near (lie middle livc^ and ii (juarter ini'hos, and at their tips only one inch. Their len;Lith is twenty-two inelies and their diameter at the alveolar hord. r autero-posteri- orly \\\o and a (piarter inches, and transversel\ one and a half inches. Towards their lower part tliey are twisted from within, forwards and outwardly." After (piotiny Pennant's remark (already ju'iven, see [i. 17) about siuular ditlerencos noted by him, he adds that "the superior incisor and molar teeth are also very nuieh smaller than in the fossils of T. ronmarKs,''^ and he i^ives measurement sshowinjit his diU'erence. He then says: "The hairs of tlu? upper lip of the T. romiKintti are stated by Shaw, to be about three inches lonj^', and almost ecpud to a straw in diameter.* lu tlio specimen innler consideration, the hairs of the moustache are stiff-pointed spines, not more than one line long- at the upi)er l)art of the li[>, and they j;radually increase in size until at the lower and outer part of the lip they are about one inch in lengtli." lie further achls, in the same connection : " Since presenting tlie above comnnniication to the Society, the Academy has received from ]\Ir. Drhdcer, of Canton, an entire specimen of the AVah'us from Xorthern Asia. In this individual, Avliicli measures in a straight line eight feet from the noso to the tail, the tusks aro ten inches long, and diverge from their alveoli to the tips, where tliey are five and a half inches apart, but they are slender, as in the stuflfedhead above mentioned, and appear as if they would ultimately have obtained the same length and direction. Per- haps the pecuharities noticed may prove to be of a sexual ehar- acter."t As will be shown later, we have hero the more prominent ex- ternal differences characterizing the two species of AValrus for the first time expUcitly stated from direct observation of speci- mens. If Dr. Leidy had had at that time good skulls of the two species for comparison, the other important cranial ditfer- euces (noted beyond) could not have escaped him, and he per- haps would have been led to formally recognize the Pacific Wal- rus as a species distinct from the Atlantic Walrus. I have met with nothing further tonching this subje(;t prior to Mr. U. W. EUiott's report on the Seal Islands of Alaska, * "Shaw's Zoology, vol. i, pt. i, p. 2'M." tTraiis. Amer. Phil. Soc. Phila., vol. x.i; i)p. d"), 8G. 90 THE GKNUS ODOli.EXrS. piildislied in l.STo, in Miiicli, iiiidcr the li('ii(lin,n' "The Walrus of IJcriii*;' 8('a, (liosiniirns Jircticus)" lie says: — "I write Mlio Walrus of IkriiKj >Sva\ bewuisc; (liis auiiaal is (jaitc distinct from the AValrus oi'llie North Atlantic and (Irccnland, dilTciiny from it specilieall.v in a very .strilduj;- manner, l»y lis <4r('alcr size and semi-liairlcss skin."* Tliis is all lie says, lio\v(i\ cr, icsjx'ct- mtX tlieir dilTcroiiCL'S, no reference Ix'in;^' made to the I'eally dis- tinctive features. Tluis (he matter rested lill, in J.S7(), (lili Ibr- mally lecoji'uized two si)ecic.s in his "List of the I'rincipal Use- ful or Injurious .Mamnuds,"t in a eatalo,i;ue of a "CoHei-tion to Illustrate th(; Animal Resources o''lhe Uidted States" in tli«' ex- hibit of the National ^Museum at tiie International l']xhibi!ion of 187(5, hohl in Piiiladelphia. This is inerely a Jiominal list, in which ap])ears, under ^•Eofimarlda'," the foUowinj^, which I hero fully and literally transcribe : KasMAitcs (iiiKSis, (Illijrcr,) Gill. TIic [Atl;iii(ic] Wiilnis. Athnif ic (.'nasi. i ]i()S.MAia;.s CooKii, (I'lvnicry,) Cij!. Till' [Piicific] V'alrii.s. I'aciilr ( 'i:|isi . Here is simply a no.uinal recognition of two species without ex])ressed I'easons tlierefoi'. in an article on the UoHinnridiC, published in 1877, Dr. (lill a.^aiu says: "Two siK'cies ii]>])earto exist — one {It. ohesiin) inliabiliuin' the northern Atlantic, and the other (/i. t'ooldi) the n(n'tlieru l'acilie."| Van !>ene(h'n, on the other hand, in 1877, distinctly afUrms his disbelief in tin; existence of two spe(;ies. In referring to the subject he says : "Xous ne croyons jtas (|ue h.'S IMorses du d6- troit de Lehrinj;' diil'erent specih(|uement de eeux de la mer de Batiiii ou de la Nouvelle-Zendde, et c'est a tout, a notre avis, que Fremery a essaye de les repartir en especes distinctcs d'a])res les modifications de leurs dents." He further adds the festinuniy of von liaer as follows: "Von ]>aer s'est occnp6 en lS;ir> de cett(! qnestion s\ FAcademitj de St. r(5tersbourg ct I'il- lustre naturaliste m'eerivait, ])eu de temps avant sa uiort, an sujet de la difference legere des Morses, a I'Est et a I'Ouest de *lii']iortoii the Pry'iilov Groin) or Seal iMlandH').' Alaska, 187:5 (not paged). Also, Keport on Ihe Condition of Ad'airs in Alaska, 187.'), p. lO). tTliis "List" is aiioiiyuions, and is licnce, perhaps, not properly qnotablo in tins eomieclion, althonyh its anthorsliip Is known to tlio present Awiter. tJcdnison'sNew Universal Cyclopsedia, vol. iii, 1877, p. 1725. 1:^ SYNONYMY. 23 lamer (llMciiilc, (|ii"il ••c^iinliiit Ics (lil'liTcnccs coiimic (U's iiio- (liliciitioiis loc'iik's*. ('" ii'csl piis r;i\ is dc Henry \V. IClliot, qui coiisi(lei'(! ]<' ^lorsc dii iiniia]ili. (,)iiad. lie. 1777. :i:!0. jyjuiiK )iii(riinis (I nippiijtohimiiK J'lilso diihin, Mornc or S< a Jlornf. 1>av, Syii.. hvx,, iin. 7r(T//co.w, JMauti.xs, Sjiii/li,, 1(J7.'), 7'-^, jd. J*, tij;-. I). — V.c.r.in:, licsilir. mid Xatiir- (jlcs( li. (J Win la lid, 171^, ijl; 17(>;>, lOli; ]Jc.sorip. el: J list. Nat. dii Gio'iil., 17()."), 5; En.— Goi-niK, ".Moi-jdioL, 1, 1817, ^11"; Act. Acad. Cius. Lcoi>. (Jarol., xv, i, 1"^:'>1, H, pi. iv (dentition, etc.).— Vox Baku, M(;ni. Acad. «t. Petersb. Math, etc., vi'= Kcr., ii, lH:ir», 199 (blood- vessels of limbs).— Jakgkr, Miiller's Aieh. llir An.'it., 1844, 70 (den- til ion — Labrador Hi)ecinien; . Walross, Mauticx.s, Zocdogischc Garten, xi, 1870, 283 (etymology). Wallrun sen Mors, RUYSCU, Theatr. Animal., 1718, 159, pi. xliv (liguro same as .lonstou'i)). Walrus, WuUM, Mus. Worm., 1G55, 289 (fig. from Do Lact).— Wymax, Proc. IJost. Soc. Nat. Hist., iii, 1850, 242 (relation to l*acliyderma). — Lea, Proc. Acad. Nat. Sci. Pliila., 1854, 2C5 (nsc of the skin). — Wheat- LAXi>, Proc. Essex Institute, 1, 1854, G2 (remarks on a sknll). — SoxxTAG, Nar. GrinncU Exj)!. Exp. 1857, 113 (woodcnt — group of Walruses). — Murray, Geogr. Distr. Mam., 180(5, 128, map, xxviii* (distribution; in part).— Hayes, Open Polar Sea, 186t, 404 (hunt- ing).— Packard, Lull. Essex Institute, i, 18G9, 137 (fm-mer exist- ence in Gulf of St. Lawrence). — Atwood, Proc. Bost. Soc. Nat. * " Lcs Morses des cOtos d(^ Sibdrio ou do I'est do I'AhIo out les dents cani- nes phis lortes quo les Morses do Spitssbcrg ot do GroDuland, nics tlisait-il dans une, lettro " t Ann. du Mus. Roy. d'llist. Nat. Bclgique, pt. 1, 1877, 45. tibid., p. 17. I A 24 OD015ii:\i:S UOSMAIM'S — ATLANTIC WALRUS. H Hist., xiii, L~7(l, ii'JO (icnuaks on a wkiill IVuiii IIk; (iiill' of St. Law- rence). — TuiiNEK, .Toiini. Aiiat. iiihI I'liys., v, 1^70, lir)(rcliifions of luTicardiiim). — L'ink'. l»inii'.li (ircciiland, Ij-T", 1'.'() ((listiil)nliun), !J1>', -'.V,', -.;:•,' (fliiisc). Arctic Wtilnix, I'kxxant, .'^yimii. (^'ikkI., IT"!, :s;i,">; Aiftic /in".]., v,'(l cil., i, ITU--', 1(W (ill jiart). Fossil It'iilnix, I'.AiM'dN, Lfiiicloii I'hil. Mil;;., wxii, 1-(C>, '.)> (iii> locality). — MnciiKi.i,, .Smith, iV (.'(h)1m;i!, Ann. Now York L.vc Nat. Hist., ii, li^'i^, 'J71 (Ibssil, Aciiiac (-'o., \'a. — doiibf I'iiIIn' ict'rircil to tlu' cxist- iiij; species) ; IMinli. New I'liil. .Jiiiiiii., y, 1~'J~. :!-.") (alisiiact of tlio last). — ![a!;i,.\\, Kdiiil). New I'iiil. .loiirn., xvii, ISM, ;>ti(l; Tiaiis. (Jeol. Soe. reini., i, l-^li.") 7.">; Med. and I'livs. Ke.caiclics, Itfll.i, 277 (same si»eeinieli). — Lyki.F, iV OWI'.N, I'loe. Lond. (ieid. Soe., iv, I'ri;], ;W (fossil. Mavtiia'a ^'illeya^d, Mass.): Aiiur. .loiiiii. Sci. and Arts, xlvi. If};'., :U'.» (same). La Viichc nuiritic. IJiiis.sox, lliii. Anini.. 17.">(i, i^. "Morscli, (i.mi:m\. L'eisc dureli liusslaiid, iii, 17;')!. It!.'). Morse ml \ (tvlic iiiuniK. lU rrox. Hist. Nat., xiii, ]7t).'), :C)f', pi. liv (animal). — l>Ai:iii:xr(»x, Million's Hist. Nat., xiii, I7(i.">, tl.'i. i>l. Iv (sknll). — HoL- i.AXKiii;. Alire^ed'llist. Nat. des Quail. Vivip.. i, 17'.l(l, pi. xii, ti;;-. :!. — 1'. C'rviKi;, Diet, des Sci. Nat., xxxiii, Ir^K!, '..'7; Dents des .Mam.. lft>5, )i^:i, 1)1. xcv. Morse, HUET, Coll. de Mam. dii Mus. dllist. Nat., l^Ot*, .V.), pi. liii (ti^-. from IJullou). Sea Cow, Siiuldhasi, Phil. Trans., Ixv, 1775, 249. PAoca, IJoxXAXio, KenimNat. Hist., i, (no date), !.'>'.), ]d. xxxix, tii;. •i7 {a i)oor veiji'esentiition of J)e Laefs ii;;ine, witli the yonn;; one omitted). P/joca rosmaritx, LlNXl5, Sy.st. Nat., i, l?")!^, 38. 2Vich<:chHs rosmariis, LiXiXi':, Syst. Nat., i, 17110, 411. — M(r,t,i;i{, I'l'od. Zool. Dan., i. 177ti, 1.— Sciii!i:iJi:i!, Sihii;etli., ii, 177."). -'(W, j)!. Ixxix (from Ballon). — Zl.MMKUMAXX, Geogr. (Jesehielite, i, 1778, iiDD; ii. 17H), 4t>l.— FAitiMcns. Taiina (inenl., l7Kt, 4.— Li{Xi.i;iii;x, Syst. He;,'. Aniin., 17."'7, .">'.»:{.— (;.mi:i.ix, Syst. Nat., i, 17."'-*, :")'.•. — SllAW. Nat. Miscel,, 171)1, pi. e( Ixxvi ; (Jen. Zoill., i, lH(t(», '2:U (in part), lig. (W, (from Jonston). — nu:Mi;xitA<'i[, Handh. derNatnrjjeseli., 178s. ij-^; 18-Jl, l:S(); It^-J.'). 11-J; Abl.ild. natnr. (;e, U71.— I?AUT()X, I'liil. Ma^^, xxxii, 180.-., 98 (fos- sil; locality not stated),— (i. Cuvji:i!, Tableau el<'!menl., 171)8, 17'i; Lemons 1."), .''.0, 01, 04, 0"^, 7.") (dis- tribution). — l)i:.sMAi!i:sT, Noiiv. Diet, d'llist. Nat., xxi, l-^l>, ;!90; Mam., 1820, 2.'j:{,— ScouicsDY, Account Ant . Kcfxions, i, 1820, 002 (K«^n- oral history). — •' Ki:i;sri;i!\. CapitisTrielieelii IJosmari Descrip.Ost,, 1821,— ."■— ScAinii, Kdinb. I'liil, .Jonrn., ii, 182."), 28',! (Orkney): .»ar- diuc'.sNat. l.ilirary, .Mam., vii, 1838, 219, pi, .xx (orij,'inal li,u;uro ifa SYNONYMY. 25 of animal).— Harlan, I'auii. Anier., 1^'2'i, 111 ; Ediul). Now Pliil. Jouni., xvii, WM, ;?G0 (fossil); T.van.s. (ieol. Soc. renii., i, lf*\'>:>, 72 (eaiiic); M('*l. and Pliys. IJcpos., 1835, L'77 (same).— GoDJLVN, Anicr. Nat. Hist., i, 18-^0, a.Vt.— Sciiix/, Nairn-. il.-r.Siiii-.-tli., 1>'-J7, 1(;'.>, pi. Ixv (two liyiiics — " AbbilduM!^ nacli BliiiUL'ubaLli imd .SL'lniiid'"). — Lkssox, Man. do Mam., 1827, 'J08.— Ross, App. Puitv'.s Fourth Voj-., 1828, ID'2; Api>. Koss'.s Second Voy., I8l!5,xxi.— Fi.kminc, I'.iit.Anini., 1828, 18.— liAi I", Natuiw. Aldiandl. Wiirtonib., ii, l-^2-\ lii7 (denti- tion) ; "Bull. Sei.Nat., xvii, 1821), 280" (abstract).- Fisciiicit, Synop. Mam., 1821), 213. — GuicKiN-Mi'iNT.vn.i.K, Icon, du I'ei^ne Auim. do (i.Cuvi-r. ilani., 18211-18:58, It), pi. xix, li^'. .") (animal).— FuKMr.iiV, lji,jdra;;t()t de natuurk. Wctcnscii., vii, 18:?1, 384.— Dkloxgciiami'S, Mom. Soc. I>inn. do Norm.'iudio, v, 131. 101 (doutitiou). — WlL- SOX, Nat. llisl. Quad, and AVIiales, 1837, 14.", pi. cccxxxiv, lig.2 (iininuil); Kncycl. Brit., 7th ed., xiv, 12"). — BKr.r., Brit. Quad., 1837, 2r)8 (animal and skull; original ligures). — Vox Baku, Mem. Acad. .St. I'etersb. Math., etc., (5" s('V., iv, 1838, it7, pi. xlvii (distribn- tion). — WllXi.MAXX, Arch, llir Natingesch.. 1838, 11:5 (dentition). — IIamii.tun, Jaid. Nat. Libr., Mam., viii, 1831), 103, pi. i (animal, and woodcut of skull, — original ligure). — IJiCltAUD.SOX, Zool. Boeciiey's Voy., 1831), ().— Br.Aixvir.r.i;, Ostoojiraphie, I)esl'ho«|ues, 1840-.'>1, 19, pi. i (skeleton), pi. iv (skull).— Dkaay, Nat. Hist. Now York, ZoiJl., i, 1842, 50.- ZiMMKKMANX, Jahrb. fiir Mineral., 1845, 73. — WA(iNr,i{, Schrcber's S,'iuy.(tli., vii, l<|i;. , V'!^; Odoutog., 1855,(^2, pi. xxxvi, fig. 5 (dentition). — Nii.ssox, Skand. Faini., 1847, 318.— (iKKVAis, Zool. et I'al. Fraiicais, i, 1^4-'-.'>2, 14(1.— Uuay. Cat. Seals in Brit. Mus., l.-.'.O, :;2; I'roe. Zo.il. .^oc. Loud., 1S53, 112 (ou iittitiules.'ind ligures): fat. .S'alsand AVluiles. 1K!1!, 3(>, 307.— OWKX, Proc. Zool. Soe. Loud., 1>'.")3. I(i3(anat. and dentition) ; Ann. ami Mag. Nat. Hist., xv, 185.'>, 22ti (from th • foregoing); Cat. Osteol. Coll. Mus. Collego Surg., 1853, 031 (skeleton); Fucycl. Brit., xvi, 1854, 403, iig. 112 (.skull); Odontography, 1'^.>I. 510, pl.exxxii. tig. 8 (dentition); Orr's Circhuif the Sciences, Zoiil., i, li-54, 230, tig. 27 (skeleton); Comp. Anat. an, 4".t(». 498, 507 ; iii, 1808, 338, 524, 780.— Blasiu.S, Faun. Wirb. Deutschl.. i, l-i57, 202, ligs. 148-150 (skull).— Van dkij Hokvicx, Ilandb. Zo,.|. Kugl. Ed. ii, 18,">8, 007.— Vo\ SciiUKNCK, h'eiseu im Aiuur-Lande, i, 1-59, 179 (in part only).— Lkidy, Trans. Amer. Phil. Soe. Phila., xi, I'OO, pis. iv, v (in part); .lourn. Acad. Nat. Sci. Phila., 2d ser., vii, l-ti9, 410.— Woi.i' A- Sc[.ATi:i!. Zool. Sketches, i, 1801, No. 10.— Gkijuai'.d, Cut. Bones .M;iui. Brit. .Mus., 1^02, 14."). — Nkwtox, Proc. Zo.ll. .8oc. Loiul., IftU, 499.— .<< i.ATiMt & Bautu-.TT, Proc. Zool. Soe. Lond., 1HI')7, Hl-i, ^^p.t.— Vox .MiDDr.NDOitri', SiliirisehoReise, iv, 1S(;7, 934 (in l)art only).— Buown. I'roe. Zoiil, Soe-. Loud., L-*lH, ;!35, 427 (liaMts and disiribulion) ; Man. Nat. llisl. (Jreeuland, 1-T."), :!,'>.— .Mii;ii;, Proc. Zoid. Soe. I, Olid.. I-'(W, t;7 (rejiori on cause of death of sprci- meu in Zoiil. (laid., I.oiid.); H70, 5"'l ; Trans. ZoiU. Soe. I.nud., \ii, pt. vi, 1H71,411. pis, li-lv (anatomy), — (Jii.ri.v. I'roe. and Trans, Nova Scotia Inst. "at.. Sci.. ii. p(.3. l.-'70, 123(witli ,i plate).- I.'ki;;;' . /j id- 20 ODOILKNUS :.>lSMAUrS ATLANTIC WALlfUS. ay, XcwtiiuiiillMiid). — lli:c<;i.iv, Rei- MMi iiacli (IriiiXoiditdliiiiiiccr, iii, 1>74, ■11! (I ml "its anil ]., :!'• stT., ii, 1^74, 1G4 (IoksII. Fraiici.').— (Iri.i,i\i;i;, I'loc. '/.oiW. Sue. Loiiil.. 1-74, ">"^!l (size 1)1' liliMiil-coi'jiii.s- ck's). — 1'i:ii.1)i:n, Zuiild^iisi, ;!(! scr., i. l-^TT, ;!(!(> ((lisiriliiiiinii and t'lKHl).— ^'A^■ lirNKDiix. Ann. Mns. l.'oy. li ibnlion, ^rncral lialiits. and liissil remains). — IviXK, Hanisli (iifinland. its l'c(i|)l(' and its I'lodncts, 1~77, VHi. Trichicluin loiKjitli iin. l'i:i:Mi;i!Y. l>ijdra;j; lot dc Xalunil;. Wchnsidi., \ i, l^'M, :W4. Trirluclnix vlr, j)l. \ix, lijjs. I, a, h (I'ds.sil. Acconiai' C'i>., Va.). t Tricliivliiis (Inhius, 8i.\N'Nirs. MiiUcr's Aicli. iur Aunt., l.';4"J, 4(t7 (witliout locality). lioxiiittfUfi (ircliciiK, LiLLlKiic )!!(;. Fanna rd'vcr.s Svcrij^rs ocli Xorgos Uyj^gr., 1874, »i74. Eobiiitiniii iriclicdnix, T..\.M()NT, Seasons w if litho Sea-borHes, 1801, 141, 1(57 (two ]ilatcs). — (in. I., Johnson's Xcw Univ. Cycio])., iii. 1^77. (i:!li. liOgniurus ohcmis, GiLi., I'roc. llsscx Inst., v, ISGd, 11! (in i)ait only); Interna- tional Exliil). 1871), Anini. Resources U. S., No. x!, l'^7(), 4 (Aflantic Waliiis; no deseiii)tion) ; JoImisou'.h X^(^w Univ. C'ycl.. iii, 1877, 17-,'.">.— r.\cK,\i:i), I'loc. Host. Soe. Xal. Hist., \. \>i\i'<. v!71 : .Mem. Dost. Soe. Xal. llisl.. i, 18()7,'J4(i (Ib.ssil).— Lkidv. .louni. Aead. X'at. .Sei. Pliila., viii, 1-77, "JU, ]il. xxx, fig. (i (I'o.ssil. .Sontli ('ar(din;i). Odohiiiotlivriiiiii hirtiliiuiinn, (.ii;ATiiH.i;T, Hull. Sue. (li'ol. de I' ranee, •."' si'r.. XV, l.-3."^. I'l'.' 1, 111. V (fossil, near I'aris, Trance). OdoHldbiniiix ro.s/, -f//(s, SrXDKVALl., (il'ver. K. Vet. Akad. I'orli.. 1>.V.». 441 ; Zcit.x li. (Ji'sammt. Xatnrw. Hallo, xv, 18()0, '2711. Odobainix ronmanix, Mai,mi;iii:x, (JlVer. K. Yet. Akad. I'orli, 18il;!. (18(14), 1"0 (food and liabi(s), 5(1."), ]il. vii (dentition) ; AViegniann's .Arcliiv f. Xatnrgcschichte, ]8i)4. Cw (tran.slated from (UVers. K. Vet. Akad. Forli., l-r,;!. i:iOet sell.). — I'KiT.Hs, Monatsli. Akad. Wiss. Jicrlin, 18(;4, ti8,"), jd. (tlcntition) ; Ann. and JIag. Xat. llisl., (;J), xv, lS()o, 3u5 (abstract).— Rink, Danish lireeulaud, 1877, 4;>0. t Tr'nhicodon huxli\j\, Laxkicstkh, Quarter. Journ. Gcol. ,Soc. Loud., xxi, 18(ir), yjC), ids. X, xi (fossil; Rc8. trhixii iiraitia, Fahricius, Fauna Grusnl., not Callorhinus uraintta ; seeBKOWN, Proc. ZoiJl. Soc. Lend., 18G8, 357, 348. Morse; Vuvhc marine ; Chvral marine ; B6ic d, la grande dent (French). Bon hiitriniis, RuY.scil, 1. e. Hvalroxs (Swedish and Danish). Harhcst; Ilrahuns (Norwegian). Morsk (Lapp). Wallroiis; MtrrjiJ'rrd (Gennan). Walrus; Sea Cow; Sea Home (Fnglisli). External Char acters.— As regards general form, the head, ill conipiirison witli the size of tlie body, is rather small, squar- EXTKUNAF- CIIAHAfTKlIS. '21 isli ill (Hitliiif, Itiil iiiiicli l()U,u('r lliiin lii'(i;i(l. witii llic iiiii/.zle iihiiipllv iniiiciilcd :iiul •soiiicwliiii l)i!oltc(l i>y llic llfs. Tin- nnsiiils arc soaicwliat crcsccntic ill --h.ipi', placi'd \ I'iiicaliv. \\ilii tlif uppiT i)ai'l iiimc ('\piUirly. The tail is scarcely, if at all, \ isible, bein^i;' enclosed within tlie te.iiUineiitsof the body. The fore limbs are free only from tlieelltow; as in the J*iniiii>eds generally, (hey are jireatl\ ex])aiided, llai, and somewhai fin- like, !)ut wilii miK'li more t'reedom of motion than is the case in tJK' PltDvidiv. They are armed with live small llat nails, i>laeed at considerable distance from the end of the cartilaginous toe- tlap. The first or inner diyit is sli;^litly the htni^'est, the others beiii;;- each stu-eessively a little sliorter till the fifth, w hich nearly equals the first. The hind liinl) is enclosed within the te;;u- nieuts of the body nearly to the heel; the free portion when expanded is lan-sha])e(l, but w hen (!losed the sides are nearly parallel. The first and fifth digits are eousiderably longer and larger than the middle ones, the lifth being also rather larger than the lir.st. They are all ])r(n'ided with snndl nails, placed at some tlistanee from the Knu\ of the toe-Hap. The soh's of both lore and hind extremities are l)are, rough, and '' warty,"' and the dor.sal surf aee of the digits as far as tin; jn'oximal jdialanges is also devoid of liair. In the young and middle-aged, the body is rather thiekly eovere \\ lii're more or less wriiiUled and thrown into Idhls, es|ireially o\erlIie slionldei's, where t lie folds are dee|i ami liea\y. 'IMie axerau'e Icii^i'lli of lour adiill males e\amined is aliont 10.', Icei. \ar\in,u' from!).] lo 11 i'eet. .\iltliors, however, eoiiiiiionl.\ .ui\e rather lar.u'er dimensions, and a length ot'lwehc feet is said to lie not inrrei|iieiitly attained. The lar,!;vst Itristles \ary in length iVoin 'J.L'."» lo LI.Tl inches. Vnn>> Dr. Miirie's paper on the ;4('iieral anatomy ol' a yoiinji individual I add a few riirtiicr details. Dr. Miiiie descrihes the muzzle as capahli' of ureal niohilily, and the mxstaeial hristlesas eiuviii;;' in dilfereiil diicctioiis aei-ordiiiu to the imiseiilar tension of the ]»arts to which they are alt ached. •'When the nostrils are rt'lax<'d they drop fmwards and the hristles inwards. At such tiiiu'.s tiie n arcs are apart liilly ].\ inch; Itnt when they are <'oii- trii. Il'.t. KXTKk.NAL CIIARAOTKHS, 2\) III respect lo llie iiiystacial l)ii.stles, Dr. Millie's li^^iii* s of tiie lieail iiiid iiiii/zle of tln^ .Voiiiijf s|)e<-iiiieii desciilted \ty liini (drawn lioiii |)lioti<;;i'apii,s, sniiie tVoiii the liviii;; animal) rejt- resent tlieiii as (|iiite Ion;;', tlie loii;L;<'st lieiii^' sailles lu'come slioilcr in adult life, heiiii;- |)erliai)s worn oil hy coiistaiil friclioii. Tlie itrislles in the specimens I have sfteii JMiic no rcsciiildaacc lo l!ie lon.i;' curving' luistles i'i;;iired and dcsciilied hy Dr. Miiiir as existing' in the yoiiiiu animal. They were coiisider.ihly (oiiethirdj longer, however, in tin; ycainu'est of four specimriis in I'rofessor Ward's collection than in the oldcsi. uiviiiu siipjioit to llic oiiiiiion already slated that llie\ itccome shorter as the animal advances in a^'ct .\s already noted, the fore feel are formed much as in other IMniiipeds, more nearly aurecin;^', however, with those of the ()liirli(l(f {h:i\\ with those of the t'kocitliv, esjtecially with res])ect to freedom of movement, having' the power of pronation and supination to a considerable decree. "In the Walrus," says Dr. Millie, '•ihe hiimeriis, radius, and ulna can he so placed that liiey meel at ail acute aii^le, the lower limb of which is in a uivat measure fre<'. '{"lie dibits, on the other hand, can to.i'ellier he t iiriied backwards at a sliari» alible with the radius and ulna, >o that the bones of the limbs altouellier forai an S ■shaped liUiire. In the Heal tlii^ anlibraciiiiiiii and dibits bend on each i'' f: Sfc l';isMir;i(''.H GcniiMii tiimsliilidii, p. i:i'J. t hi J',ill;is's li;; I 111 ■fill lliH "Icoili's'") of ;i \ nini;; i\;nii|il<' oC llir I'lnilic Wnl- nis, llic iiiysliU'iiil luisllcH arc irin'csculcd as very luiii;-, as in llic vnmi^ of 11k: Allaiilic species. 30 ODOILENTS }{()SMAl{rs — ATLANTIC WAI.KTS. I'. V, ■ r'\ other more iiiiniiliiily, tliiis < In tlic net of swiniiniiiii the Wiilnis cv itlciilly ciiii use its lore limb iis \\\v jis tlic cUiow. with ;i kind of rotiiry movi'iiiciit (>; the maims and iiiitiliracliimii; Init in the Seal liic lotaiy a<'tii)n takes phice only at the wrist, and al»o\(' that a sort ol'^inylynioid of hack and I'oiwaid mo\c- nient." ••The pahnar snrfaee or sole of {lie nianns is not r.iiiiKe a par- hji' sliovel in li;iiire. 'I'heic is a .!;reat eallons. ron,i;hei;»'d and warty pad at the ]>roximal eml or l»all of tlie hand: an, while. La- nioiit says a lull ;; row ii old male will wei;,di at least. .'5,000 ])oiiiids.f Aside, t'loiii Dr. .Miirie's iiieasiireiiieiits ol'a .voiiiit^spe- ciiiieii, I have iii(;t, witli iiodetaihid liieasiii-eiiieids «)!' the, Atlan- tic Walrus, exeepi thosi! j,'iv(^n by I)i'. iii,|: whieli are, as fol- lows : I'-t. III. JlxiniiK! length 12 3 j,iii-iii <.r ii.aii I r, 111 cm I 111 of iiiii/,/lc 1 J li--l,iiir<' I'lniii ii(>s<^ tr> vyi' H l)i^I;ilii(j hcl Wren ryi'M lii J", \lrliMion (if t ll-.k lir\(ili(l 1 lie llionlll I II ] li,->liilicc 111' IiisUn ;i|i:i|'I ;it Iiusc J ] lisl.inii' of lilsUs :i|i;irl :it (i|is II Ij(Ii;;||i (if fciiv ||i|i|i(f U U ISlvjiillli ,,f foil llipiirr I I Li'iiy;lli of hi Ml I lli|i|pir 1 In Urcadlli of liiiiil lli|i|ii T, ilislimliil 2 I'eet, with a ciieiiiiit'erenee at. the shoulders of Id feet ; and the leiiii'th of the tiisUs as L'tl iiH-hes. Ilaniiltoii I savs an iiiilix idiial Uilled inOrlc- lie\ . ill ISLM, whieli he saw, '• was alioiil ten feet in leiio(li," with the head \'.'>S inches in lenj^lli. From the si/e of the lusk.s (ex- serted S.j inches) it ajipears to lia\c been far from fidly ;(rowii. Daiilieiitoii j^i\es the leiiulli of the specimen he described as lU feel, with a eirci I inference at t In-shoiilders of S I'eet. Ijuniont 'Acromil of the .\irl. I.'c;^., vol. i, |i. .'ifi-,'. >.Mr. I.aiiioiil, ill his •'Sca.soiis willi the Sca-hor.scs" (p. — i, ;;ivc.s llic vii;;iil of an old male as il.noii |ioiiinN. Imf in his " Yachfiii^; in I Ik; Arctic Seas" (p. "'.I), he says, "A fiill-si/eil ohl hull Walrus imist weiyli iit least. GjCiMi Ijis.. anil ,-.iich a Walrus, if \eiy fat, will inoiluee Ci.'iO Ihs. of Iilnbber, bul sehloiii niore I hail 7MI) Ihs., whieli is I think the iinnuic :inioiint yiclileil li\ till 1I10-.I ohese of our \ ill iiiis." lie speaks, luiwever, ill aiioflicu place (p. !-;(), of one that ''yiehleil hetwccn 70it ami -0(1 ]ii)un .IS II I'tTl ."» iiiclics.* Mvnuri iiM-a^iiic- iiK'iils III' lliii'i- ihIiiII iii:ilrs rruiii iilisl iin'iil (Milltilj sKiii-i :il'i- :ih I'liHows: (I) li'lljilli (I'lnni liosi' ti» lilil), 10 I'ril ■"» imliivs; (J)!» I'l'i-I, i'l iiirlii's; (;;) 10 li'it III iiiclics; (l)S fi-cl o inilics. 'i'lii- liist, lliirr Will" I'lilly inliill, w liili- uiii- ■if I Ill-Ill. Id jilili^r I'imiii il^ hroKiii, wmii liisks;iii«l |>iiilly ii;ik«Ml, sciincd skin. Wiis \<'r\ nlil ; tlic oIIkt \v;is iiiiI iiiori' tli;iii Iwu lliiids ;;r"\\"- 'llii'si' iii;i\ :ill li;i\r lii'cii s|i('riiiH'li.s ol' less tliiili I he ;i\ riilj^i' sizi-. AiMilij;, liowiMT, l."» lo IH iiu'lu's lor I he l<'n;;lli ol" llic liiiHlliml* (in>' iii'ii- ilirlllilfil), wiiiilil ;^i\i';i irii;4tli ol' ;il>i»iit IL' I'lrt lor lln- l;ii;;»'r iliili\ iilliills.^ Most ol' t lie old w lilcis wcir coiilctil wit li stilt in;;' it to lie iis hiip'iis iiii o\ :iiiil ;is thick iis ;i Iio^^sIkmiI. 'I'Iic iiccoiiiits of lli<^ color iirr iilso ilisci'c|Kiiit ; I'liiiriciiis's stiitciiiciit Hint llic color \;iiics V ''li ii;^!', Ilic voilli;,'^ liciii;; liliick, tiicji (|iisk.\, liitcr |»;ilcr, iiiid liii.ill.\ ill old ;i,iic white, luiviiij;' Itccii (|iiot(d it.v most suit- sc(|iiciit coiii|)ilcis. Writers who liiivc ;;i\cii the color I'loiii iictiiiil oliscr\!itioii li;i\c never, however, coidiriiied l''id»riciiis's ;iccoiiiit, they iisiiiilly descrilMii;;' tln' c'olor of IIm^ hair iis "yel- lowish hrowii," "yellowish jiijiy," " tauiiy," ''very Ii<;ht yellow- ish-;;ray," etc., some of whom explicitly state that after (;xteiid<'(l oliservatioiis tliey have never met with the chaii;^e,s of color with a;;(' noted hy l-'aliriciiis. TlmH, Mr. Ivohert, l»rown say.s that althonjili he has seen Walruses of all staj^os, from Itiitli until nearly mature aj;e, he never saw any of a l)la<;k color, all hf'inj;' of" the ordinary hrown color, though, like mo.st animals, they ;,'et lijilifer as tln^y {i;v()\v old."| Scorcsby says that the skin of the Walrus is coverod "with a short 'llowish-brown coloied liair."§ Dr. (lilpin stat<'H that Ids Jiahrador siHtcinuMi was thinly cov- ered with " a
  • resse,d li};ht yellowish-f^reen hair," about an inch in len^'th. lie jwlds that the, siirfju!»! of the. whole skin was mmm 34 ()Doba:m:s uosmaruh — atlanthj walrus. 1 I i covcivd by "sciU's iiiul bald warty patches," and tbat tbo skin itsi'b" was tbrowii hito " welts and folds'' <»ii tlu' neck and slioul- deis. Mv. Ibown Ihrther says that " the, very circumstantial account of tlie number of mystacial bristles given in some accounts is ni(»st erroneous; they vary in the lunnber of rows and in the numlM'i' in each row in almost every specimen. They are ele- vatt'd on a minute tubercle, and tiie spaces between these bris- tles are covered with downy whitish hairs."* Many other writers also note the scars and warty i)atches and partial absence of hair n'ferred to above by l>r. (rili)in. Mr. Brown, in speakinj; of those he met with in Davis .Straits, says: ^' I iiavc seen an old Walrus <|uite spotted with leprous-lookinji' marks consisting of irregular tubercular-looking whiter carti- laginous hau'less blotches; they api»eart d to be the cicatrices of woumls inliicted at dilfercnt times by ice, tlu' claws of the Polar Uear, or nu't witli in the wear and tear »>f tiie rough-and- tnndde life a Sea-horse must lead in X. hit. 7 P."* ^Ir. Lamont further adds that in the Spitsbergen seas the "old bulls are always very light-colored, from being nearly devoid of liair: their skins ai'e rough and rugose, like tliat of a Hidnoceros. and they are gt-nerally quite covered with scars and wounds, inlbcted l)y harpoons, hinces, aiul bullets wiiich they have escaped from, as well as by tlie tusks of one another in tights .init.ug themsehes.''t From these reports, especially tiiat of Mr. IJrowM, Dr. 'riei ]ias iid'erred that the Walrus is subject to skin diseas hat the; ''glandular spots*' thus produced are mista' liealed cutaneous \\ounds.'' However this may be. etty well established that many of these marks are reall^> ars of wounds. Respecting other external characters, especially the tusks, and their variations with age, sex, and accidental causes, I transcribe the following from Mr. Lamont's entertaining book, which will be found s(» freely ([noted in subsequent ])ages: "Old bulls," he obser\-es, •<■ very fre(iuently have oue or both of their tusks broken, whi(!h may arise from using them to assist in chunbering up the i(,'e and rocks The calf has n(» tirsks the lirst year, but the second year, when he has attained to about the size of a large Seal, he has a i)air al)out as large as *l*ioc. ZoiJl. .Soc. Loud., IciiJd, p. l-'cr. t Seasons with tlio .Smi-Uoi'ses, p. i;57. ITraiiy. Zool. Soc. Loud., vol. vii, 167'.i, p. 122. 44 KXTF.HNAL CIIAIIACTKRS. 35 I tlu' fiiiiiiieteethofa lion ; tlu' tliinl year tln'y lire about .six inches lon^. "Tnsks vary very ""K'h i" >^^'/'^' i"'^■ ^1^ iK m !^-'' 3G ODOBvENUS KOSMAKUS — ATLANTIC WALRUS. suspected by AVieji'inaiiu luitl Staniiius (see (intca, p. 11>), who believed that the Jeiiiale liad longer, sk'udeier, and more eoji- veryiny tusks thau the male. Tlu.'re is also a speeimeu in the collection of the Museum of Comparative Zoiilogy, Cambridge, ill which the tusks are very long and slender, and eoii^^-ge to such a degree that their points actually o\erlai). Ill concluding this rat!ierrand)ling notice of the external cliar- acteis and aspect of the Atlantic Walrus, I append the (juaint and very correct description of tins animal, Avritti'U by the mis- siouary Egede as early as 1740. I give it from Kriinitz's Ger- man translation from the original Danish : "Der Walh'oss, oder das Meerpferd, ist eine Art \on Fisch, desseu Gestalt einem Seehunde gleicliLjmmt: jed .-h ist es Aveit grosser uiul stiirker. Seine Pfoten sind mit fii f Klaueu verseheu, wie die I'foteu des Seehuudes ; doch kiir/er von Xil- geln ; uud der Kopf is dicker, ruuder und stiirker. Die Uaut diescis Thieres ist, vornehmlich am liaise, einen Daumen dick, imd aller Orten faltig, uud runzlig. Es luit eiu dickes und bratmes Haar. In dem obern Kiuubacken sitzeu zwey ki-umme Ziibne, welche aus dem Muiide liber der Unterlipi)e hervorrageu ; und eiueu oder zwey Fuss lang, uud bisweileu aueh wohl noch liinger siud. Die WaUi'osszalme sind in eben solcliem Werth, als die Elephanteuziihue. luwendig siud sie dicht iiud fest, au dor Wurzel aber hold. Seiu Maul ist wie eiu Ochsenmaul; uuton iind oben mit staclilichten Borsten, in der Dicke eines Strohhalms, besetzt, uud dieso dieuen ihm anstatt eines Bartes. Oberhalb des Mundes sind zwey Naselocher, wie bey dem See- hunde. Seine rothe xUigen seheu ganz feurig aus ; und weil sein Hals ganz ausserordentlich dick ist, kanu er nicht leicht um sicli herum selien ; und dieserhalb dreht er die Augeu im Kopfe horum, waun er etwas ansehen will. Er hat, gleich dem Seehunde, einen sehi* kurzen Schwauz. Seiu Fleisch hat eine Aehnlichkeit mit dem Schweinenlleische. Es pflegt sich tlieses Thier mehrentheils auf dem Else aufzuhalten. Indesseu kann es so lange auf dem Lande bleiben, bis es der Hunger niithigt, in die See zu gehen ; iudem es sich von deuen Fischen und Meer- Insokteu uiiter} ilt. Wdun es im Zorne ist, briillt es wie ein Ochs. Die Meerpferde smd beherzt, und stehen sich eiuander bis in dea Tod bey. Sic leben in bestiindigem Kriege mit denen Biiren, denen sie mit ihreii grossen und starken Ziihnen genug zu schaffen machen. Oefters tragen sie den Sieg davon; und .il KXTKHX/ T> (FrARACTKUS. '■> 7 ■woniji'stciis kiiiiii>l<'ii sic so hiiiuc, I»is si(> todt ziir lOnlc iiicdcr- faiU'ii."* AiHitlicr iiftiMiiit of the Wiilnis, lioiii its hciiii;' one of tiic Oiirlicst extant, is also of espcciiil iiilcrest in tlie present cou- iieetioii. 'riionj»li repeatedly eo})ie(l, in part or wlioll.v, l»y tlio earliei' authors, and also l»y von J':u'r, I tliink it deseivin.n' of reprodnetion here. It was written by Irof. A. E. Yorst, and was based on the yonnj; specimen taken to llolhuitt in lOl.'J. It is liere c()pi<'d from I)e Laet (l)escrip. [ndiic Occident.), by wlioni it was pnbli.slied in HJ.'>;>: "BeUuain lian<; niaiinani vidi, inagnitndlne vituli, aut eanis Britannici majoris, Plioc.T non diasimilem; capite rotnndo, ocu- lis bovillis, naribns depcessis ac patulis, (luos modo contralie- bat, modo dicbicebat, auriiim loco ntrinquc foramina; rictns oris rotnndo nee ita \asto, snperiori parte aut labro mystaca gestabat setis cartilaj>ineis, erassis ac rilane carebat. I'ostica parte repebat magis quam incedebat. Cute (^rassa, coreacea, pilisque brevibus ac temiisibus obsita vestiebatur, colore cinereo. Grun- nitum apri instar edebat, sen crocitabat voce gravi et valida. Kepebat per aream extia aiutiendo, herumque gestantem cibuni ac offerentem magno nisu ac gruuuitu accedebat, sequebaturque, nidore ejus allectus. Lardum ejus gustantibus baud insuave visum est. * Hfvni tIauM Egi'rtc, Missioniirs mid liischolV's in Groulaml, Ijescliioibung uutl Xatuv-Gcsfhu'lito von Gninland, iibcrsctzet von D. Joh. Co. Kriinitz. Jlir Ivii]>ri 111. ]{( rlin, vorlogrs August Mylius, 17G3. i)}). l()(i-10f^.— Since transrvil'iiig the above I have met with an early (I7(i8) Englitsli transhitiou of this Avork, in whieh nn English rondoring o ' the above description may be found at p. 12.'>. 11 ns ()1)( )B.'E\US I?OS.MAHUS — ATLANTIC WALllUS. ■1 '; (.:' Coiispii-icbiiiilur il»i(Uiii duo iiuijoiimi ciipitM, ilciitilms duobus exertis ICl('j)liaiit<>iiiiii iiist;ii'. loii^is ;if cmssis ct iilbiciiiitilni.s iinmitii. fjtii (Icoi'siiiii \<'rsiis |»fctiis s]»('('t;il>;iiit. iMinuii.coria cere mil IC ixnulo iicudissc I'ci'cliaiit .\ii,i:li (jiii attidciiiiit. Ilisct' dt'iitiltii.s rui)('s asccndci'c siMino snstiiicrc ;ij('l»;iiit. rt pro- dfilii! ill (•(iiitiliclitclii sell tfiTMlii \\\ soimiiiiiii il;i r;i|ii;ilil .^li'j^a- tiiii. i'iil>iiliii!i ajcliam illis esse lolia nltl()i::^a ac iiiauiia, lu'rhtc ciijusdaiii t' I'liiidoiiiaii.s nascent is. NVc piscihiis \ iNcrcaut ra.i'ni- vormii <'ss('. \idi iltidcm pcncin i-Jusdcni aiiiiiialis osst'iini, ro- tunduiii, i'ultitiiiii ct aiiiplins loii^iini. ciassinii, |)ondoi-(i,sinn ac solidniii, in tine ]n'(>i)e j^landcMi lon.uc cras.sioieni ac jotundioreni. llujns pulvcre ad calcidiini jtcllcndinn .Moscovita' rctnntur."* A still earlier desciij)ti()ii ol'tlie Walrus is juivcn hy Purcliast in liis account of the first voyage ''into llie North Seas," by ^^'illialll Uari'nts, a Dutch iiavij;atoi', \vli(» met with AN'abnses oil Oraii.iic Island, in l.?,tt, Iraiislaled from the Dutch b; \y. Philip. The account says they "'^ went to one «if those Islands [of OraiigeJ, where th<^y I'ouiid about two hundred Walriisheu, or Sea-horses, lyinjj,- upon tlie shore to bast tlieiuselves in the Suune. This Sea-horse is a Avoiulerful strou<;' ]Mouster of the Sea, uuich biji<>er than an Oxe, which keeps continually in tlu^ Seas, haviiiji' a skin like a Sea-ealfe or Seale, with very short hayre, mouthed like a Lion, and many times they lye u]»on the Ice; they are hardly killed unlesse you strike them , just upon the forehead, it hath foure Feet, but no Eares, and commonly it hath one or two youiif>- ones at a time. And when the Fisher- men chance to find them upon a Hake of Ice with their young ones, sliee easteth her younj;' ones before her into the water, and then takes them in her Amies and so i)luiiji('th up and downo ■with them, and when slice will revenge her-selfe upon the IJoates, or make resistance against them, then shee casts her young ones from her agaiiie, anil with all her force goeth towards \\u' Doate thinking to overthrow if They have two teeth sticking out of tluiir mouthes, on each side one, each being about half an Ell long, and are esteemed to bee good as any Ivory or EleiHiants teeth, especially in Mil scon via, Tiirtiiria, and thcreal i •; where they are knowne, for they are as white, hard, and c\eii as Ivorie." Skxial Dii'KK1m:>;ci;s. — The subject of sexual dilfciences ni the Walruses has received very little attention at the hands of ' Nov lis Oil lis sell l)cs( riplid Jmliii' Occident:! lis, |)]i. lis, '.V,), 1(i;i;), tlli< I'll'^riiiics, \(il, iii, p. ITii. SKxr.M. ini'i r.i;i:\rKS. ao sysl('iiiiiti<' wiilt'i's, wlio li;i\c, imlft'd, no jjositivc iiilni'iiiatidii to olVci', ;mm1 \<'r\ little can he yh-aiH'd IVoiii other .soiiircs. All that 1 have met \vith,atfeii»retlye\leiisive researt'h.1iasalrea(l\ Iteeii iiieiileiilally ix'ivvu in the loreyoiii.u' aeeoiiiit of t'le extenial char Meters. All that (an hi' .^atheredis that in the lemale the tusks are smaller and thinner, and th<^ ji'enenil size oi' i!ie animal may lie inleni'd ti> he somewhat smaller than in the mal<-. In tact, the external eharaeters in the adult animal of t lie speeies under eonsideiation have never as yet heen j;iven with much detail, the few naturalists who have met with it in life seemin-; to take it lor .granted that an aniiaal so long- known, and so familiar to tliom, nuist he well known, thereby ronderinj;- a careful and d*;- tailed descript ion unnecessary. The very good descrijttion j;ivon by Ur. (iilpin (see (intri), ]»]>. '"U, 32, ;?.">) of an adult is about all that I have met with in the way of detailed descriptions of the adults of either sex. The lijiures and descri|)tions given of the young, especially those rce<'ntlyi»ublished by Dr. J. 3[urie,* leave lit tie to be desired as regards the external eharaeters in early life. The absence of references to any strongly marked sexual dift'erenees in the adult might ])erhaits be taken as negative evidence that none exist; but on the basis of analogy Avitli the other IMnnipeds, especially with the OtnriUl(V, we shonI;1 hardly exi»ect their absence. Even in the case of the skulls, leT sexed specimens appear to have come under the observation of specialists. Wo " re and there, however, meet with references to supposed sex- ual difl'erences in the size and character of the tusks, and also in respect to the size of the skull and the density and weight of the bones in those of supposed females as comi)ared with tlK)So of supposed males. Thus, Wiegmann, in 1832, in referring to the species described by Fremery, in 1831, says, in remarking ujwn Fremery's " Trlchcchm Cooliii,''^ that he remeud)ers having heard from a Greenland traveller that the female Walrus has longer and slenderer tusks than the male, and states, on the au- thority of Fremery, that a young specinuMi in the IJoyal Museum of JloUand, having long, slender tusks, was reganU'd by Tem- minckas a fenude. IJe als(» considers, (ui the ground of analogy, that the greater or less develo]>ment of the occipital and other crests of the skull, as well as the relative weight of the bones, * " Kfst:nxli(s iqioii tlic Anatomy of tlio riuniitedia.— I'fiit I. < »ii I lie Wal- rus ('/Vi./Kc/iH.f ivmiittniH, liiim.)." — Trans. Zi)t')l. Soc. Loud., vol. \ii, 1>7'.*. pp. 411-1(11, witji wdoilciits, and iilatus li-lv. ■M i iiE M If* )■ 40 (•DOIS.KNIS IJOSMAKUS ATLANTIC WAMJIS. 'i ■ ( i. 1(» be only (litlrrciiccs of a scvual cliaractcr.* Staimiiis.f icii years later, citcil the views >.. reiiiiniiick ami \Vi(L;iiiaiiii (as above .uiveii) resi»ee(iiij;' sexual «lil]'ereiu'es iu Walruses, but adds nothing' u«'\v to the subject. Laniont (see (inlci'i. \>. ."!.">) states that the ''tusks vary very nuu-h in si/e and slia|H' aeeoidiiiii to the a^'e and sex of the animal." "(Jows' tusks,'' he says, ••will areraije fully as lonji' as bulls', Irom beiuj^' less liable to l»e broken, but they are seldom more than twenty inches lonu and three pounds each in weight. They are j^i'nerally set nuich closer together than the bull's tusks, sometime o\('rhii)i»inji- at tlio points, us in the case with the stuftcd specimen at the I>ritish Museum." lie gives the UMigth of tusks in the male as 2t inches, aud the weight as i jjounds each. A skekton, marked as that of a female, in the Museum of Comparative Zoology, coHeeted in the Greenland seas by ])r. Kane, has the bones very light, soft, and i)orous, as coni]»ared with those of male s]>eelmens. The skull (see tigg. l-;») is nuich smaller, with the (;rests and ri'('lnod(')ilicit sciii hoIIIc. mil Aiialofricii beU'gt werden." — Arch, fur Xafiirficsfh., 18:]2, pii. l'^-', I'J!*. tMiillcr'sArcli. fiir. Aiiat., 1^11, j.. :«>•,'. Sr.XIAI. DIFl'KRKNCKS. 41 ttisks (see li:r. 1) is iloiibllcss iihiioriiiiil. nml is doiilttl<'ss owiiii;- to their iiiisyiiiiiictviciil ared imnnrous specimens of lioth sexes, whicii were lost v.iili the ill- fated vessel. till theXalioual Miiseinu there is also a female skull of (he I'aciii'' \\ alms that ])rese)its corresiiondin}; diHerences as compared with male skulls of the same Hpceies. I VARIATIONS DEPENDENT UPON AGE, ETC. 43 coiijecturo.s of Wk'giiuuiu ami Tcumiinck. What othor dilier- cucc'S obtain, especially in external (^Isaiactois, can as yet bo only conjectured. It i.s to be hoped, lio\v<'ver, that wc shall not have lony to wait for detailed accounts of the external characters of the adults of both sexes. Fio, 'X — Odoha'uiifi ronmarus, 9 . Individual Vaeiatio>>!, and ^'Al{IA^IONs dependent UPON Age. — That a wide rauye of individual variation obtains in this si)ecios is sufflcicntly evident from an examination of even a limited series of skulls. These dilierences have been noted in (.'onsiderable dcitail by Fremcry, Wicgniann, Stannius, and Jaeger, as will be presently noticed more in detail iu ju'e- senting the j^enend histoiy of the species. Still greater difler- ences, of course, result from dilierences of age. These collect- ively, as A\ill be noted later, have formed the basis of several nominal species. All the Pinnipeds appear to be subject to a wide range of variations of thischaraitter, and none more so than the AValruses. Tliese atVect to a considerable extent tlie general l)roportions of the skull, and especially tlie form and relative development of different I)ones. Tliese latter dilierences are best seen in eom})arativfly young skulls, since most of the sutures close at a ratliei" early age. Among tliese variations are especially noteworthy those of the nasal bones, the inter- maxillaries, and the frontals, and to a les.-. degree those of the base of the skull. Tlu' crests and ridges for nuis. 20. \.VliIATK>N.S DKl'ENDKNT UPON ACJE, KTC. 45 TIm! iVoiitiils Viiry * Z 0^ <>* 1 '< 1- o il « : S f2 •qjSnoi '.ttiifao.ttoi •)JV(Ib 09aC|8Il> 'BJ0ilI3U| J,)dd£], •j«ioni yiv\ IB o)BiBd JO mPIAV O rt g -T • O M ?1 ?i CI H ■ Si cI s s s s s •8ojao8 iiiaojt.iBium .loddn jo i(^noi I*; f5 C5 XI 1^ « ci .- S » •sdj} IB ?,iB(Iu ojun^Bip 'KjuinBO •oBtiq )B K.iSpa jBn aojxa uooAvj"! ojubjbii) 'souiub,') S 3 fi S fS ?? S HI MSBq IB ^i.nMi.i|iMii.).ii.) 'K.mniiQ | J,' -(SJCIOOI JO onc[)Bll!lI JO pu) .loi.wjsod O) Q!ii!XBnu.>(nt jd.i.ip.ioq .wi.wjuv' Tt h. ja e = X t- S ?1 H fi ? S ^5 s fS X 1^ M I- I- !• is {I ri I- » *H -H r-< fj ffl 1^ M •.\I.iou.itm! ipi'lAv 's niiH| n:ni:.>v •.\'t.xoiaojS()4l uipiAV 'Kouofj iiisBX •i[iSaoi '8.>no(i iiisBX I vussoj I iBiodmo} uoD.ttjoq CO fS •XOg Of O O \3 \3 O "to r', a O O O o O ^ .3 -3 -a '53 -a "TS •jaqninn ouSoxbjbo '?5 -I « « « to© 5 ^ 5 t-< •« rt » O lias DKNTITION. 47 I)i",.NriTif»N. — The (It'iititioii of Wiilniscs, for \iirioii.s rcu-sons, lias hccii :i |M'i|ilt'.\iii;; siilijcci, and lias cii^a^ii'd tlif attoiitioii of iiiaii\ niiiiifiit wiitcrs. In tlic adult stajnc it pivsi'iits many altiioniialitics, and, ln'sidcs, issuhject to nmcli individual varia- tion. Iiotli in tin- tnnporarv and adult series. For a Ion;;' time its deviations from tli«' normal type were not well untler.stood, and even now leadin;;' authorities do not appear to be roi)er notation. A.s previously staled, iIm' incisors of both Jaws, except the outer i)air in the upper, disa]>pear soon after birth, and before middle life is reached the last tttotli of the molar series on each sule iu both jaws also usually disapjiears. A brief history of the princii)al iiiv<'sti^'ations. and opinions held at different times resi)ectiug thetlentition of llie Atlantic \Valrus (for the investigations resjicctin;;' the dentition of this ^roup appear to have been based almost wholly uiton this species), is herewith appended as form- ing a lii;.thly interesting chajjter in the technical history of the species. Ill this historical sketch will lie found noted many facts relating to the general subject, given by the authors whose papers ai'<' here biiclly summarized. The dentition of tlie Atlantic Walru.; has bt'cii discussed in greater or less detail by iJai)p, von l.aer, Wiegmaun, Frcmery, k>tannius, .lacgcr, ( )weii. Mahiigreii, Peters, and various other writers. NViegmanii, in IS.'JS, pretty fully presented the early history of the subject, noting the almost total lack of informa- tion respecting the matter shown by Linne, who evidently paid little attention t(> the references to the subject made by previ- ous writers. The credit of first giving any definite statement respecting the nnnd)er of the teeth and their character is due to Anderson, who, in 1734, gave the niunber of molars as four above and three below. Brisson, in IToG, gave the number as four both above and below; while Crantz, in 170,'), again gives foiu' above and three below, and ipiite fairly describes the nor- mal dentition of the adult.* In the same year, Daubenton gave also again four below on each side as well as four above. ' I iii>pt'n(l iu lull Ci'uutz's description: '• It liail no sliiiip («ci\s<)/T« in its mouth, anil noiio at all liifoiv, Init only four tcotli on I'adi side ; on tlierii^lit siilcol" tlii" nntltT-jaw three pretty hroad eoncave jrrindors The two lonj^ t usks or horns grn\viii more an imiiodinient than a liel]) to It Tlierijht tusk is aliont an inch lon<;er than the left, and its wliole lengtli ' -HI m\\ • 'I > C. f I 1' :, ''.. :M 4H (Mum.KMS I.'OSMAIMS— Ari.AM'lC WALias. No ;intli<>r ]niof to Sclirchci- (177"i) iippcars to li;i\(' iiicr •vith (Iccidiioiis iiici.voi's. \\Im» I'omul l\V(» such ii|»|»i'i' iiicistus on ciu'li si.s'.S7( iiifcrnKt.rilhirid (W^ Wiiliiosses mid (h'r Sehlieide/.iihiie i || ' dessellu'ii liiit keiiiieii ieiiieii, iiideiii er iiiir cine vortiefllielie Abhiindhiiiji' iiiit sehruieii Zeiehiimi.u'eii dieser Kiioeheii \fr- sc'hiediMier Thieve zii^escliickt li;itte.'' Caiiipev, in eritieisin.i;' Limio's errors rejiiirdiiijn' the Wiilnis,t K'ves four imisois (,^^,)' aud four iiiohirs altove and livt^ lielow (-,Z-,) ^*'^' soiueliiiies only four below). Tlu^ observations of Sehreber, (loethe. and Taiii- j , per ap])ear to luive been ucneiaily oxcilooked by siibsiMineiit writers, so that it was h'ft for (r. Ciivier to iliseover anew the liresenee of deeiibious ineisors in the yoniifi" Wahiis. lietweeu the canines he recoj;nized two incisors similar t.> the molars, which he says the majority of observers had overlooked, because they are not lixed in the interiiiaxillaiy, and between these again two i)ointen alleni Anselien naeh auserhalb dem Zahnlieische nicht zn bemerkeu seyn, /umal da sie nicht am Rande der obern Kinnlade, sondern mehr hineinwiirts stehen. Ich linthi sie an einem zur Natnraliensanindnng hiesiger Universitilt gehorigen Wall- rossschiidel ; xmd da derselbe, besage seiner Grosse, vcm einem jungen Thierc ist ; so glaube ich beynalie gar, dass si(^ bey zunehiiiendem Alter des Thieres ausfallen nnd nicht wieder wachsen. Sie konimen also hier in keine weitero Betrachtung, als d.iss sie dem Systeniatiker (;inen Wink geben, dis Thier nicht zu weit von dem Robbengeschleehte zn entfenien." — Saugcthiere, Th. ii, p. 2C0. tl qtiote the French edition of C.'miper's works ((Eu^Tes, torn, ii, p. 480, Paris, 1S0I>), the only one accessible to me. 4M KHNTITION. 40 Inwcrjiiw.* I''. (Juvicr jiiivc latt'i'tilsu the siiiiic dciitiil loninila. IIcdcciiM'd tliiit tlic peculiar or imoiiiiilons deiititioii of the Wal- ruses indicated that tiiey were ait isohited ^inaip, lutviii;^ atliui- ties. on tlu' (»ne iiaud, with the Carnivura, and, on the other, witli the Ikiuninantslt Accordin.i; to \Vie;uniann, I'lidolpliil (in ISOL*) reco;;ni/ed liie lirst ofliie series ui' loircr ^rindin^' teetii as a canine. 5 Thus, as \Viej;niann lon;^' sin<'e oliserved, liu' suhject reniairu-d till Ifapp was so I'ortuuate, iu ISiiS, as to Inive opportunity to exanuue a tuMal specinu'ii. in this example, he Ibund six inci- sors in the upper Jaw and live in the lower (ij^^)- He also expressed it as his heliet' that the lirst low«'r molar should he rej;arded as a canine, hecause (1) it was somewhat further removed from the rest than tlu; others wore frcuu each <»t her; because (li) of its j;reater length and thickness in the adult ani- mal: hccause (.'{) it stands close to the temporary ov nnlk inci- sors, and shuts aj>ainst the outernu)st of the ui)per incis(»rs; and hecause ( t) it lacks the transverse de])ressiou seen on the inu'-r side of the crown of the back-teeth. The dental '')rmula recojiuized hv him for the Walrus may be considered as 1. I-'renu'ry. in 1S,")1, also made reference to the dentition of the Walruses; hut his paper bears mainly upon the question of whether there are one or more species of these animals, and will be further noticed in another connection. He notes partic- ularly the presence, in some of his skulls, of two small molars abo\ e, behind the largo ones. Wieginann,|| in 1838, contributed facts additional to those already recorded, but his memoir is largely devoted to a discus- sion of the obsen-ations of preceding Avriters. He assents to Rudolphi's and Kapp's interpretation of the honiologieal rela- tion of the lirst large tooth of the lower jaw; refers to finding ' Ki'jfiio Aniiual, toiii. i. 1817. p. 1G8. t Dents (li'sMiiiii., p. -23^. tAiiiitoini;>cli-itliysiolojfis(li(> Abliaiulluugi'ii, p. 145. vNVVifjfiiiaimsiiy.s: ". . . . UelJcnVu's ist liiiddlplii dcrorstr, dordic iintorca 1"< lli of six incisors l)<)(ii iihovf^ iiiid below, iuul to tlic (-.irly (Iccidiioiis ciiiiractcr of tin; last (liltli) upper molar, and I lie rreipicnt disappearancui t/f Mic loiirMi. lie ('(/ncludes thill I lie norniiil number of the back-teeth is ^~-' and lliat in 4 — 'I' earl.N lile the dent it ion of the Wall lis is not \vi(h'ly ditl'eicnt from that of other Pinnipeds. 'I'he same year (l.S.'{.S), Mae;;illivray* eonsi(h'red the normal dentition of the Walrus to l)e \.'^.', <'• ',1^; TJii. + M. ^-^' = j^ = ;;:;. ills conclusion was based on th(^ examination of a ])er Jaw there are <>n each side tliiee imisors, the first oi- inner extremely small, the seciond a little lari^cr, the third or oiiltr disj>ro|>or(i()nately iar;;e, beiii;; e((iial to the laij^cr .yiind- eis. The socket of this tooth is placed in tiie iiiterniaxillar\ bone, but towaids its mouth it is|»ai'lly i'oniied Ity the maxillaiy. The small incisors ha\e deej) conical sock<'ts. The canine" tooth is disi)Iaccd, beiiij;' thrust outwards licyoiid the line of the other teeth, and caiisin.in' the peculiar l)ulj;in;;' of tlic head. The lateral iiuMsor is on llic IcncMiI' its aiitcrioi' liiai.niii. and the first urindcr is o|»posite to its middle, 'i'liere arc ti\e j^rindcrs. ]ia\iii,u ccoiid, and third nearly c(pial. the latter beinu' a little larger, the fourth much smaller, and the IllVli vciy small, flic tusks, or enormoiisls de\cl(»ped canine teeth <»f the upper jaw, arc comiu'csscd, <"oiiicai. a little curved backward, directed dow nwards and a little forwards, and somewhat ili\cr;4in,i;-, but in some iiidi\ idiials, when \ery loiri;. they ajiain coii\(r;.:e towards the points. In adults, lhcincisor> are obliteiatetl, exceptiii!.; the lateral pair of the up|)er jaw; the fifth jj,rlndcr in liotli Jaws has al-o disappeared, and sometimes the fourth in one or both Jaws." ■ iiriti.'dniiuidniiicds, JsiJH, j.p. -^Jl i, -J-J 1 . if ;(;;! DKNTITION. 51 StiUiiiiii.s. ill ISl'J, riiiUicr coiitrilmti'd to llic siilijrct hy suldiii;; <»i).s(a'v;ili<)ii.s icspccliii^ Viirinlioii i'l the iiiuiiIh'I' of t\H\ l(M>tli icsiiltiii^' IVoiii ;i;;(', ' in di'tail llic incisive driitilioii ol" ii sciics of four skulls of (liU'erciit, )i|?«'S. In two youii^' skulls, tln! outer tcin|)onii',v incisor of the iipiHT Jaw on either si(l(5 icmainod; tlu5 alveoli of IIk; s;'cond pair were still distinct, while the alveoli of tiie iiddd' : paii' weit^ nearly oblit- erated. In another, the alveoli of the, inin'r pair of ineisorH were wholly obliterated; those of the second pair wen; baiely re(!Og- nizalih', while those; of the outer j>air wer»} distinct, the teeth havin^i' fallen later. He was also able lo i'eeoj:;nize the alveoli of six incisors in the lowei- Jaws of tin; skulls Just mentioned, and states that Ik; lhouj;ht Kapp's view of the homology of the first lower bac k-tooth (considered as a canine) was jtrobably correct. He liirthei- takes exceptions lo the value of tin; char- acters assumed by I'^remery as the l»asis of several species of \Valriis(\s.* lie adds, in respect to the lusks, that in old a.nc they become \vlioll,\ solid to the base. In regard to the ii|)pcr molars, lie notes thepicseiice of live in sev<'ral instaiicciS, and liiids tiiat, as a rule, the fourth disappears before the fifth, or, at least, that, its alveolus becomes s(»oner obiilerate(l. llo also ctudirms tiie siMtemeiit jucviously made by Wiegmann, that th(! alveoli becoiin! lilled by depositions of bony iimtler in concentric layers at the bottom and on tin; sid( s. ''I'lirsc ;illc)j;c(l sprcilic, cliiO'dclrrs ln>, not ices in dotail, jinil coiKsititTH thorn ii.s (l('|icii(li'iii, iijiim ji;;c. lie, says: " iJiiH Mis.siiclMi (liener Clianiktcro <,'ilu'lll KclioM iiiiK(l('iiilJiiis)aii(lt',tiawH(ii('.s('lln'ii iiiir riirv(illi;r aiiHyrwachseiic'J'liicre, iiiciit abcr I'iir Jniij^c! uiiwciidliai' Hind, dcnn das Htiiik.sto Waclistliiim der Eckziilnio fall), erst, (>ti'cid)ar in cino Hpiitero Lcbciispdriodi) in widclier naniciitlicli dii', Ix'idcn iiincrstcn Sclincidcziilinti nnddio bcidcn lolztcn iJack- /.iibno j(!diT HciU; drx oljcrcn ICinnlado schon j^escliwundcii Hind. Uiorzu kouuiit noch dor Unintand, dass aiuh din ICok/illino Ixii sehr alton 'i'liioreu an del- Spitzo ItodcnfcnJ al);;('nnizt; .sii.il, dcnniacli in siiiittTon Lclx-nsHta- tlicn an Jiiin;^o waLrschcinlich wicdor abnclinicn. Endlidi Hclicint cs ja scllist, alsol) dio Liiiifjo dioscr Ziihno jo iiach don Gcsolilrclitorn vorscliioden wiiri'. " i^lioii so wcni;;- (i('\vi(lii nuiclito ich util" di(i l"urclinn;,;on dicscr Ziihno Ic^iin. llinr Zaid, wic iiinr fStiirko uach Hind Hio ))oi vorschiodonon iibri- Hiiis niijif \()ii cinaMiloi' aliU(:icliii<;, lie notes lliiit tlie l:ist m|i|mt iiioliii' Itiid already fallen tVoin the ri^lit side, hut still remained on tlio left, belli lid wliieli, as well as heliind the ah et tins of the fourth iiiohir on the ri<;ht side, was a little shallow pit, in wliieli, diir- irij; fd'tal life, a tooth hiid perhaps stood. In front :A' these heforeiiientioiied teeth were three molars on eaeli side, and in front of these a eonieal iiieisoi, and the alxcoli of the others were traceable, altlioii;4h alieiid.v filled with a spongy substance. f the lower jaw, there were li\(' teeth on ea<'li side, with traces of three already lalleii ludal incisors on one side and of two on tli(! other, '{'he second skull (!>•/ inches by Trj) was somewhat older, the canines bein^' about live inches lony. There were present in this skull three ii|>per molars cui e:icli side, and a lilled-wp alveolus behind them. Of tlu'se te<'tli fas also in tin; other skull), tlu^ middle one wu.s the lar^^est and most worn.t In front of these, and somewhat distant from them (')'"), was an incisoron (;ach side, and in addition to these another jiair of small c — .'{; .lae^icr thus recoj;ni/in^, as had Ifudolphi, Uapp, \Viej;iiiann, Fj'emery, and Stannius, the lirst of the; low«'r-Jaw series as a canine. Iteliind these were 1 races of lli«s alveoli of the fourth jiair of molars. In the third skidl (leii«rth 12.\ inches, breadth lOA), still older, with tusks a foot lonj;', wen^ three u]»[»er "baek-liMith " on each side, close to;;('tlier, the middh; one beinj; the lar;;est, and in frontof these a eultinj;-tooiii. The low<'r Jaw had also lour teeth on each side, homiil\ in IJH' iipprr liiiiii I lie lowci-Jiiw : ;iii(l lliiil. iiiiHc i;ii<'j\ , ;i siii;ill InoIJi \\;is siiiM'imlflcdbcliiiKl I lie nor mill loiii-, ill llic upper jiiw, iiiid still inure rniely in Hie lower j;i\\ ; llie roriniilii -A' tlie ileiitilion of siieli \iirielies, in exeess, iH.in-.— i.r,-:, <■• 'ni. ;;, M, I —I 1—1 = LM»." < hveli liereinnkes. no relereiiee totlie lileinl lire of llic, sii'oject, aiid evldcntl.y ^avc- a very erroneous inlerprelnlion ol' tiic; denial loriiiida. In Ids ■atei' relereiiees lo llie siihjecl lie, ^ives an enliiely diHereiit in- terpretation, and one more, nearly af^reeiiij;' with lliat now <;oin- monly a(M;ei>led. In his lalesi i«!leicn(U', lo llie, suhjecl,* iitj writes: " In Hie W a!iiis(7V/(7/a7i».s' rfmnariin) \hv normal incisive roriiiiila is I ransitorily represented in Hie veiy yoiin;;' animal, wliieli lias three teeth in each i)rema\illary and I \v(» on each side of the lore part of the lower jaw; Iheysoon disappeai' except the outer pair above, which rcmaiiM'htse, to the maxil- lary suture, on the inner side of Hie sockets of Hie eiiormoii.-, canines, and commence the series of small and simple, niolus which they resemhie in size and form. In the adult. Hien^ iMe, usually three such molars on c k h side, hehiiid the permaiienf incisor, and four similar teeth on each side, of the lowei' jaw; the anterior one passing' into the intcrsiiace. between the npi»er incisor and the lirst molar Tin; canii'es iii'e of enormous si/e Their hom()lyp(i below I'clain.s Hicsizoan.l sliajte of tile siiceeedin;;' molars," Tlit^ formulii of Hie normal dentition a])parciitly hove re('o;;nize Ihe lower Jaw, the lirst permanent looHi is I'c^iardeil as a canine, because it is thicker and rounder than tluj jtostciior leelli, and lacks tiie eross-furrow that marks the oth- .\ii,ilinii\ 1(1" Vi'i'ti^hriitcs, vol. iii, y. ',V.W. tOilniil,.-., 1,. M-J, |,1. :!(;, lijr. 7; Siill}irl ll., p. !v!;>. ^riirs iiic, iiowcvi!!', US mIiiiwii liy Miiliiigrcii (kim^ licyoiid), incccdcd in tho riiii)iy() liy (cinporary tccili. ;l li- hn ■ V 54 ( MM tU.KXrs K( )S.MAUrS ATI-ANTIC WALUUS. ; ! ors. The \(miij;aiiiiiia! \^i^ live iipiici iii()liiiil\»i'iii teeth (•' llaclc- zslline''), tlie lii.st t\v'll dentition: 1. J^J, C. \-\, M. :-;. While (iieliel ae('e]»ts the first ])ernianent tooth of the in<>lariforiM s<'rie.s (if the Mi»i>er jaw as an ineisor, and the lirst in the lower Jav as a canine, he recoj-nizes only two iiersisteid nioLirs on each side, above and below. Malm.uren,* in ISG4, ti;.>ured the dentition from a Iti'tal s]K»ci- men, and i»nblished an elaborate i»a]»er on the deidition of the Walrns, in which he leviewed at some lenjith the history of the snbject, notieinjj;- (^nite fully the writings of the early authors, from ( 'rautz to the Ciiviors, and the papers of liapii, Owen, Wiegmaun, Xilsson, and other later writers. The fonnula he I)resonts as that of the peraumcnt dentition is : I. ^-^J, C. ~\, M. !|5J|=^=18; and for the deciduous dentition: 1. 1~^, 0. ]^\, M, 4—4 Hi J" 'JO + 4 — 4~Iii The specimen figured shows both the permanent and deeidu- ons dentition. The deciduous teeth are most of them separately lignred, of natural size, as minnte, slender, spindle-i'ooted teeth, with short, thi(;kened crowns. The permanent teeth are already in place, altlumgh even the n]»per <'anines liad inobably not pierced the gum. The middle i»air of incisors of both jaws had ah'eady disappeared, leading only their distinctly recogiuzablo alveoli. His specimen api)ears to have liad bnt a single cadn- cous molarbehind the permanent series, Iron Avhich he assumes the nnniber of upper molars to be 4 — 4 insteid of 5 — ."). The following year, Peters f referred to Malmgren's paper, jiublishing a jdate illustrating the dentition as existing in a some- what older skull (received from Labrador) than that tignred by Malmgren. I'eters here takes e.\ce])tion to Malmgren's assumed numberofba'.'k-teeth, which, in accordance with the views of Kapp and Wiegmaun, Veters believed shoidd be I'—.', instead of j~i >^ . i) — u' 4 — 4 •M»rvcrsiy;t ill' Kiiii.ul. \'(t.-Ak;iil. I'orlmmll., It^O.i, i)]). r)(lo-r)-J:.', jil. vii. ■ tTln' ii;i|M T liiMim |>iililislifit ill Swcdisli. I am iiiuihli' to I'ollnw liiiii in his llisCllSsioll (if lIlC SlllljlMl . t M(iiial>l). I\. P. Akail., l-^'i."), \)]>. (!t!r)-(i87, id. faciny p. Cc^."). u ■■■i k DENTITION. ;JD t'ii According to lA-U-rs, .M;il!ii,i,'i\'ii, froui not liuditi.^' niorc than I'oiu' iii>[t('r hiick Ici'tli ill any of I lit' many skulls of \arious aj;os lie liiid cxaniiiicd, concluded lliat wjicn a lifth is present it is aliiioriaal. TIk' youn}^' skull li,uui'ed and desciihed l)y P»;ters, JHtwcvcr, lias in the ui)]»ei' Jaw tlie I'onrth and liltli hack-teeth still in place on tiie lij^lit sidi', and the t'ourtli on the left side, with an aheolusot'a liltli. This Peters considered as allording new proof of the correctness of Wiogniainfs formula. As already noticed, live molars have been recognized l»y I'remery, l^app, (iiehel, and Owen, and, though perhaps not always pres- ent, are frequently to be met with. The dental formuhi of the Wabus, as determhied by liapp and AVieginauu, hivs l)eeu adopted by Van der Iloeveu * and Blasius, f as well as by Peters, and essentially by (liebel. Gie- bel, however, gives oidy four deciduous lower incisors, instead of six. Owen, in his later works, agrees in this point with Gie- bcl, but takes apparently no cognizance of the deciduous fourth and fifth molars, to which he refers, however, in Ids earlier papers. Gray, | in 18G(>, althouuh quothig the formida given by Eapp, 'vlo ^s the folk-vlug: "Cutting teeth ^ in young, I in adult; grniders —^ in adult, truncated, all single-rooted; ca- nines, upper very large, exserted." He, however, quotes Iiapi)'s fonnula, and also that given by Owen in his " Catalogue of the Osteological Series of the IMuseum of the Eoyal College of Sur- geons" (1853, p. 030). Professor Flower, § in 18G0, gave a diagraui of the dentition of the Walrus based on many observations made l>y himself and on "those of others, especially' Professor Malmgren,'' in which both the temporary and permanent dentition is indi, p. 738, Eutjlisli cd. \ Siiiiy;i'tlu(i'e Di'iUst'lilaiidw, l>i7u, iip. 2G1, :i0'2, I Cat. Seals and Wliaks, p. :35. vN .IiiMin. Aiiat. and Pliys., iii, ]). 27'2. .1 ■mm m ' .")<) oltoli.KNl S HOSMAHIS AI'LANTIC WAI.IM'S. ,,' , ; UIifi'cqiK'iitly pfl'sistciil to cxd'ciiic old iiyf, iilllioii.uli ('(Hiiliiolily ,, ' losi ill inacciMtcd skulls. TIk'si- iii(liiiiciiliii',v tcctli iirc iisiuilly ' ii dosciilicd as ' iiiilk-tcctli '; (ivcn llic postciior ones arc sonic- times so called, luit it apjsoars to iiic an open (lucstion wlicllicr tlio\ rcscnt pennaiiciit teeth in a nidiinciitary I or aborted condition." , J Huxley, ill liis "AnatoHiN of \'ertcl>rated Animals" (pp. I'A'A), j . i 301), piiltlislied in l.sTl', ado])ts tlie f'ollowinji' as the dental for- ;; ' ; mulaofthe Walruses: "I. ,'-^, ('. J-[, i>.ni. in. :;-;!+ ;::^V' lie I C ! says: >• Tlu^ dentiTioii of the Walrus is extremely peculiar. In \\.< the adult, there is one simple conical tooth in the out«'r part of I : the i)remaxilla, followed by a huge tusk-like canine, and three, short, simple-fanji'od teeth. Sometimes, t\V(» other teeth, which ' ' i soon fall out, lie behind these, on each sid«' of the upper Jaw. In , the mandible theie are no incisors, but a sin;;le slntrt canine is ' 1 ; followed by three similar, simple teeth, and l)y one other, which I is caducous." * Txdh here and in the formula no reference is made to the deciduous incisors, althouj;li the caducous molars are recognized. In the fore^oiiiii risitmr, we have seen how va^tui Avas the information bcaiiii.u' on this .viibj;'t possessed l)y all writers I, l)i-ior to altout tile Ix'^iiniiiiiu' of the present ceiitiii\ : how the jji ' ' earlier notices of the existence of iiicisois in i lie noiiiii:' were overlooked and rediscoxcrcd l»y hiter writers, as well as how ' slowly the tirst periiianeii', tooth of the molariform series of the ui>per Jaw came to be ^('neially reco^iiixed as a true incisor and not a molar; how, later, the numi)er of incisors in theyonn^was fouiul to be six in tlii' upper Jaw and six in the lower Jaw, with, as a rule, two small caducous molars on each side in the upper jaw, and one on each side in the lower behind the i>ermanent ' . grindinj;' teeth ; that the tirst permanent molariform tooth of the lower Jaw was a canine and not a molar; and that l>y dif- ferent writers the inimber of incisors i-eco^nized in the lower jaw has been sometimes four and sometimes six, and the cadu- cous upper mohirs re<;arded sonK'tiii:"s a.s one and sometimes as two. Finally, that the true formula of the full dentition of the Walrus is I. i!^!*; C. \-\; I'm. M. ]-;;=l;?=;U. It lience appears that the dentition of the Walruses is peculiar and some- what abnormal in four featui'cs, namely, (1) the early disap- pearance of all the incisors except the outer pair of the upper * AiiMt. N'crti'ln-. Aiiiiii., \)\k ;?(jO, ;?G1. Hfc^ffl FOSSIL IIKMAINS. i)l jaw, (L') llic ('iionnoiis (l('V<'lo]iiin'iit of fluMiiipcr caiiincs. (.'») tlic sli.uiil spcriali/.alinii of l lie liiwt'i' caiiiiics, aiitl (I) t he <'a(!iic()!is ciiaiiictci' III' till' two postciiur |iaiis nl' molars ol' 1 hi' iipiu r jaw anil till' itostcrioi' |)air in tlic litwi r jaw. 'I'lir ciirly (Iciititidii of tln' Walrus (lillcrs mainly iVom tli.il ol' iiiost ullici' riiiniprils in liaviH,u six lower incisors instead of lonr. I lit' incisi\»' Ibi'Miiila ol' otJMT Pinnipuls, as^ciicially I't'coj^uizcd, Itcinj;' usually v^ !. IVc- qucnlly i;"!;, and sonu'tiMics (as in Mocrorli i mis iuu\ t'l/stopliura) •-^-, — never, at least in the i>einiaiient dentition, ^-~;|, but I am far Iroiii sure this number may not sometimes appear in the deciduous dentition, in the Sea Otter {Knlii/dris), there are said to be six lower incisors in the yonn^i', wliile only four are present in adult lile. The middle i)air of h)wer incisors so early disai>]tear that e\cn in very yonny specimens they are some- f'lnes wanting;. Ifapj) found in a f(etal s])e('imen tliree on one side and only two on the other, and ([uite a inimber of ]>romi- nenl writers on the subject have reco^iiii/ed two j>aii's of lower incisois as the normal number. In many specimens, the alveoli of three pairs have been found, and, in addition to tin? instances already jiiven, 1 may ad '. that theic is a younu' skull in the Museum of ('om])arative /o(lloi;y lliat shows decided traces of thri'c paiis. the outer incisor on one side bein^■ still in j>lace. In view of all that is at jtresent known respectinj^' the sid)- ject, 1 a(h)pt the followinji formula; .is beini;- well-established, — premisinji', however, that they are substantially in accord with the view of the ease presented by Professor Flower in 18(I!» : — Temporary dentition: 1. i|i;|; <'.JeI; M. i5^ = ||; = ;w. Permanent dentition : I. i-;^-; t\}5-|; M. •^^^=JJ = 2G; the last two upper molars and the last lower one on each side being rudimentary and often absent. Fossil Re3IAIx\S.— llemahis of the Atlantic Walrus, in a fossil state, have been fonnd at various points along the Atlantic coast li'om Elaine to South Carolina, and in Europe as far south as l-lngland and France. The first noticed front American locali- ties was thus mentioned by Barton in 1S05, but the locality is not given, lie says: "The bones of one of these large animals have been foun»l. Tines*? ap])ear to have belonged to as]»ecies <»f trichcchns; j)erhaps to the triclicchun rosinarus or ?Ho/-.sc."* Messrs. Mitchill, Smith, and Cooper described, hi *Liiii(l(.ii Phil. Mill;., vol. .\xxii, !->()'>, p. Dd Mm »aJU,>J*- I i:' oS olxiB.EMS IJOSMAIiCs — ATLANTIC WALKIS. 1tfMllv fcftTicil it: hut hitcr it wiis rc^nrdcd 1>\ l)fi\ii.\ Jl-S r('i»i'('S('ntiii;n' ii distinct species, to wliicli lie ii;i\e liie n;i:iie Trlcli cell IIS ririiiiiiiiiiiis.\ In iSlI, l,,veil described a tusk ul» tainod from the Tertiary Clays of (iardiner. Maine, wliicli Owon reii'ardod as pi'ol»al»l\ belonijiHiU' to an extinct s|»ecies.§ Lyellll also refers to a skidl lie ohtained at .Martha's X'ineyard, Massaeliiisetts. Ilcdcseiihes tliis skull as >• lilferin.L;fr(Mn skulls of the cxistiiiij;' species {Tricliivliii.s rosmiini.s, IJnn.). with wliich it was eoiiipaicd hy Professor Owen, in lia\ in,u' only six molars and tw(» tusks, whereas those of the recent lia\ ti four molars on each side, besides occasionally a rudimentary one. The front ' I tusk is rounder than that of the recent walrns."<| In 1S."»7, Dr. Leidy** described and li^iired a skidl found on the sea-beach at Lonj;' l>ranch, 3Iouuiouth Couidy, New .Jersey, where it was (»btain»'d by Prof. .1. V. l''razer in IS."),'), and refers to another specimen (consistin;j;' of the facial portion of a skull) J discovered at the same locality by Prof. (Jeo. II. Cook. The first-nanK'd specimen, says Dr. Leidy, ''has lost a portion of the craniuui i>roi)er, and the exserted ixution of one tusk, but other- Avise, except beiny a little water-worn, is in a ^ood state of preservation. It is luichan^^ed in texture, and nearly so in colour; and it helonyed to an old individual, as all the sutures are com])letely obliterated. The form of the facial ]>ortion of this specimen corresponds with that of the specimen from Vir- ginia, [described by JJeKay and precediuji' Avriters,J abox'e men- tioned ; and the entire skull closely resembles that of the recent Walrus, TrichccliiiN rosmai'iis, as represented in Ihi^ li<;ures of Danbenton, (.'iivier, and De lilainviUe ; and its measurements are also sufllciently near those j^iven by the lirst-named auth(»r to recoj^nize it as the same s]iecies. * Anil. bye. \:it. Hist. New Ycuk, vol. ii, Is^S. j.. -JTl. tEdiuli. New I'iiil. .loiirii., vol. xvii, l-^IM. p. :!C.(l. tNat. Hist. Xfw Yoik, Zoiilof^ty, \>t. i, t>'l'J, [p. aCi, |il. \ix, (ins. t, I'lH-kiiitl. .A[. 'ilC. IIAincr. .(.lurii. Sci. and Arts. vol. xlvi, IsM, )>. oil*. ^1 As is well known, t lie cxistiiij;' Wah'is lias occasionally only I lie imiiilni oftoctli foiiiid in tlic .Marllia"s Vini'yard siu'cinicn. **Traiis. .Via. I'liil. Soc, \(d. xl. \. -:>. ids. i\, \, li^-. 1. m4il "I, FOSSIL REMAINS. 50 "Tlif tusks in tliu fos.sil ciirvod (lowmvai'dly in a e "^he remains of the sanu' species which prol>ably during the glacial period extended its habitation very far south of the latitude in which it has been found in the historic iieriod."! In view of the now well-known former extension of the habitat of the Moose, (Jaribou, lieindeer, Musk Ox, and other northern manuiials, southward to Kentucky, the latter hypothesis seems the more probable one, and that the species in glacial times inhabited the eastern coast of the United States southward to Virginia, if not even l)eyond this point. 3[ore recently. Dr. Leidy has announced the occurrence of AValrus remains in the phosphate beds of Ashley Jtiver, South Carolina, and has described aud figured a tusk from that locality. * Trans Aiiii'f. IMiil. Soi'., vol. xi, jiji. f^l!, 84. tlliid.. p. .-^l. ^ii, 'i; \^ GO < »1 >( )nA:NIJS 1{0SMAHL.S — ATLANTIC WAMJl'S. ( ■':!' ''Til is spcciiiicii," lie siiys, " is its Idacli as cboiiy, dense, licstvy, and Itiittlc, and is iicaily coiiiplctt', cxccpl a( llic tliin border oi" the |»idp eavil.v. 'rii(MMu\atiiri' is ,«;li>;lit, and it indii-ates the tootli li> he oi' the h'l't si(U'." He <;iveH its . FOSSIL li'K.MAlNS. 01 '• ]| \v;i,s])iirli;ill,v imbedded in :i liiyt-r t»l" blue el;iy ii tu(»t in fliiek- iiess, (tveiliiid by i) lii.verofli-iliterela.v l\\<> I'eel two iiielies (liick, ('()iil;iiiiiii,n (lists iiiid f^\n'\ls iA' Mi/d (iriiKiria, Maroniii iinhiilBull. Soe. Gdol. de Frauce, 2-= «6r., xv, 1858, p- C24. i i ; • \'S' *S J.« ,:i*^aj^ fi2 (tltlill.r.MS K'OS.MAKMS — ATLANTIC WALKI'S. }|. 1 H'" t sont (I'liillcins »|ii(' ties piiiticiiliMitf's relatives ])oiir lii plnpart i\ IVi;;*' ct ill! scxr, iiiiisi <|ni> Til <'tiil)li M. (icivais."* Vail r.('ii»' oceiiju'et ipie (Iratiolct u (leeiit sons le nom iVOdohnmtlurv. Lai'tet I'avait reinis s\ (Jratio- let. Xous avons examine cette tete avee tout le soin mlcessairc et lions partapjeons conipletenient I'avis que M. I'anl (lervais a exj>i'iin6 t\ son snjet dans la Zoolof/ie et la PnU'ontoUxjic frani;ai8es (p. SS), c'est-a-(liie, (pie ee erane IVaittnre et .pit a subi I'actiou du fen, n'est antie (!hose (pi'iui erane de >rorse vivant qui 6ii(t rapporte da Xord. '• Nons avons etndie eett snr nil t'rajiinent de crane (hmt la eavite een'-hiale a (''t('' ntiUstJe pour iiii iisa;;'e (piehMtiapie et (pii anra ('id apportc dans eet etat ])ar (piehpie i»("('lieiir dii Nord. C'est h; e('»t«!i droit et iioii le e(»t('' jianehe (pii est eoiiservtJ. "(-ehii qui se Ironve devaiit ee rra<;iiieiit de crane et (ini a devaiit Ini nii choix de seel ions dcs diverses le^ioiis s. He ennmerates no less than twelve or tifteen spechneiis of these remains, mostly I'ragments, collected from varions hicalities, all from the so-called "IJed ("rag" forniati(ni of England, or its eqni\'alent. The principal hxtalities are Sntton, Felixstow, and Bawdsey, hi England, hnt he refers also ty tlnMr o(!cnrrence at diilerent points in Jjelginin. The iiiaj(nity of the specimens of the tusk obtain(!(l, Avrites .Mr. Lankester, "are its ])ointcd 'Dull. Soc. G<;ol. de rranec, I^' st^r., ii. 1H7-I, pp. ICO, 170. tDcsL'iip. (tcsOsHcineutsFossilcsdcslJivironsd'Aiivcvs, Aim. Mtis. d'llist. Nat. (le I]clKi,i,'iu\ i, 18T7, pj). 10. 11. t'vt.i; lid's, I.'iiiniiiiK fds^ih, l';ti7, p. 1)1)4. — Id., Jlouitininl dca mora . , . , , p. 48, 1^70." FO.SSIL KKMAINS. <;;; toniiiiiiitioiis; hill otiifi- s|icciiiii'iis, of tlu^ bust' aiiil iiitcnncili- iitc |»orti()iis, liu\«' collie lo li;;lil. 'riirtdif-lioiil its U'iij;tli," Mr. I.;iiilv«'»ti'i- cuiiliiiiu's, •• w iiirli in soiih' cxamplrs must Iiaxc Ih'uii I'lillv liiivc I'l't't, llic liisU is sli;^lilly curved; but in those wliicli apptar In 1»<' I'lilly j;ro\vii tlic curve, is (loiisiilerably ;;i'e;ilei' lowanls the leriiiiiial point, the (lirccttioii of the ciii\c, probabl^N <>i\iii^ ihe tiislv, it' its l'iiini;;raih^ atllnities Ixt established, a j'elrollccted position, as in tlic Diiiotluiriiun. Tiie Cia;;' tusk is \eiy iiiiuih compressed hiterally, so liiat. its transverse sec- tion lias an elUptical outline, whilst that of th»i Dinothrriuin- tusk is nearly cireidar. 'I'he amount of lateral compression is, however, extremely variable, as it is also in tin; iixiii^^- VV'al- nises; the amount also of the lat«!ral as well as the antcro- j)osieii(»r llection <»f the tusk appears to vary, as in the recent Tfivlnriis, the xariability of which in the si/e and form of its ni>ks is well known. A single lar^e furrow on the outer siir- I'acc. two on the inner, and one on tin; inner curved margin, ex- tend aloiij; the wiiole length of the tusk ill many specimens, exactly similar to those oil some liisks of Walrus; but in both the recent and fossil specimens llie_\ are subject to much varia- tion, i.i I heir major or minor dcM-lopmci it. No appearance of aii\ weaiili;;' of Ihe point of the tusks b_\ use diirill.y life is obsiTVa- l)!r: and indeed the j;reater backward cui'vatnre of that part sei'iiis to result from its freedom from iisan'c, since in the Walrus the jioint of the tusk is rapidly worn away, which of course checks any tendency to curvature which mi,i;lit become. a|)par- cut if the tusk were not used against such hard substances us rocks and blocks of ice. '• I'loiii ail cxaniinatiun of the yeuoral contour and form of the tusks, without reyard to their substance or structure, one would unquestionably be led to reyard tlieui us belonj^in;;- to an animal siiiiihu' to the existiu,';' Walrus, inasmuch as it is in this animal alone that this form of tusk, \\*ith its longitudinal furrows, j;reat h.'iijitli, and j;entle curvature, is found."' .Vfter describin}^' ill detail the structure of ihese fossil tusks, as shown in seittiyns and as revealed by (he microscope, ^Mr. Laiikester further observes: -'In its microscopical striutlnre, the dentine of the fossil tusks presents a complete resemhlauee lo that of the Walrus,* .... The dentinal tubes are verv *Tli('ir inicii)s('(>])it'!il siructiu'i', iis wot! ;ih cxttTiKiI \'tn\\\, tar illustrated liy uuiiu'i'iius liL;iii'.'s, loniiin;^' jilatcs x and \i, afcoiniiaux iiii; Mr. t.aidvf.s- ii'r's jiaprr. rM 04 (»I)()IL1>NLJ.S K'OMMAIMS ATI.ANI'IC WAMJl'-S. I 'I- '■'1 ^i(^^-:1!^l nearly of the Siiiiic : izc, and ('(inally closclv paclccd, and ate connected with stellate lacnna' in some nnnilteis near the jjcri- pliery of the tooth. This Vtinclnic, which is not pc ^n liar to the Walrns, is, nevei'lheh'ss, a l«'st of alllnitv, inasnnicli as the form of the lacuna- vaiies in dillercnt animals, 'i'liev are not .net with in the tnsks of the I'roboscidea or tin' liippopotannis, init occnr in the cnrions incisors of the l)n,n(»n,ii. i'he ■dentinal cells' of the < 'ra,n' tnsks also reseiiihle those <»f the W'ali'iis In Htrucfiin^ the cement exactly resembles thatof the Walrus, displayin,:;' vascnlar canals, honc-lacnna', and canalicnli. of the .siune form and disj)osition ; hnt the |>rop(»rtion which it hears to tln! tlii(;kiiess » I" the othei' tooth tissues appears t(t he lar;>«T in the Walrus than in the; lossil. " I'^'rom the foicn'oini;' remarks it will he apparent that w;- have in thesivfossil tusks characteis which ally them most closely to the lai'ii'c canines of the ;j,'einis lVo.S(; to establish the ^enus Trichccodoa to r<'ceiv(( the animal tinis iiidicated. The Jusu(i(;ation of a e,en- erio .sepiirutioii mu.st bes(m<.dit iuthefaetof the yreat anticpiity of the 14od Crag, and the consequent probability of the associ- ation of other and more distinctive attributes with those of the tusks." As re^'urds its ji'eolo^^ical posithni ami associations. Mi". Lau ke.ster adds: " It a])i)ears that the Trichaothm IlKxlciji, like the Cetice."n remains of the Cray and hw^v. Sharks' teeth, is a ;, 1>.\ the ;^(.'(>l(»;4i(.'al «'\ idciM'c ol' llirir ;iiiti(i- iiit\ i!s Itv tli<' ;ictii!ill,\ ■»l).s('i'V('(l and inliriittcdlN s!ij;lit dilVci- ciicrs of roiMi and striifl inc. Mr. liaiiUcslcr docs not iiiloiiii iis rcsi»;'(i iii.u I Ik- locality wIicik-c <'aiiic liis spcciiiicns oi' t lie t iisks ol' ilir li\iii.';' Walrus with wiiicii lie coinitais'il tin- fossil liisks. Ill I his coll licet ion it iiia.v l»<' added (see further on this point the Mccoiint nl' 0(l()l)(r'tiis<)h(Niis'j;]\<'n\t{'y{m(\){hn{ (he disks of the i'a- cilic species (^/(/o/yrr/*»f.s- o/>r.s».s) are not only !oii;;ciand slenderer than I hose of liie Atlantic species {(). rosiiHiriis), hut are shaiper- j)oiiilcd and more inciiiNcd, and do not present the worn and hntkeii appearance so often (indeed, iisiially) seen in the tasks of lid individuals of the latter. Whet her oi- not tlic\ present dilfercnces of striK'tiirc has not, so far as known ,0 me. hceii iiiicroscopieally determined. The tusks of the l*acilic species, flirt herm(»re, sometiiiK's atfain the si/(^ indicated for the tusks of" TriclKcodon hnxhuiW For the jircscnt I niiist consider Laii- k(!Stci"s Tricltcwdon hiudcjil as certainly not j;«'iierically separ- able from the exislin^' Walruses, although it may have; differed from the exisliny Atlantic species in larger si/*; and jiossihly in other cl'.aracters, as so often ha[>pen,s among' the iiniiiediate [»ro- genitorsof existing speeios in other gToui>s of inainmals. \'an IJeiieden has recently reviewed iit considerable length the hi loryof the supposed and actual fossil remains of tin; Walrus,* sliowing that most of those reported as found in dill'er- eiit ]»arts of l^'ranc • aiuKjcrmany were really those of d.iHereut >speci(!s of extinct tSireniaus or other animals than the Walrus. Van Ucneden, liowever, describes and iigures a dorsal vertebra Ik; considers as that of the Walrus, found near Deurne, and a sea- ]»hoid hone from Anvers. (iKiKiKAl'IIlCAI. DlSTUnU'TlON, PKKSJINT AND PAST. — 1. (Uxtst oJ'Nortk America. — As already shown (^/H/m,pp.'~>7-<»l ), the Walrus, like the Musk Ox, the (Jaribou, and the Moose, langed (hiiiiig the great lee Period nn.ch beyond the southern limit of its Itoiindary at the time the eastern coast of North Ainericii was first visited by lOiirop 'aiis. AVhile its icmains have been found as far south :;s Xew .Jersey, Vii'ginia, and even South < -arolina, thert; is no (;vi. .Misc. I'ul>. No. IL' r» fid nl>0|',,i;M S IJOSMARIS All-AN'I H' WAMMIS. I No\ ;i S(i»ti;i, ;i.^ ucll ;i - t lit- sliorc-i mid isliiiids to I In- iioiHiWiird; hill llii^ ;i|i|M-:irs lo liiivi- \u-i\i :il lliiil liiiic llicji' soiif licrii liinit Ol l|i>ll illlllidll. Ill M;i\, I."».'I1, II1C3 well' iiicl uilli l»\ .hiiiics ( iniicr, nhoiit llic i>l;iiiil ol' '" K':ilii<-;i"' (piol^iMv Siihic I.s|;iii(lj, w li(» t liiis irlcis 1(» IImiii: "Ahoiil Hh- siiid Isliiiid {li:iiii(;i| jiic v»'r\ j^icmU; Ix'iislcs ;is ;ii(';it iis oxen, wliicli li;i\c luo ^^ifiit tcctli in tlicir fiioiitlis Iil I'reiyhted with these lislies." Another writer sa_\s tiiat he had seen ;i "drs Hal lidl al oiiee" ol' llieir teeth, ••uliieh ale a i'oole and -iiiiielinies more in len'^IJi."" 'riie\ al^o. al ahoiil I li< ,-ame time, rieipieiited the >o called •• l;ir "' oil' < 'ape Kiel on. Sa.\ s < hai'les lau ;iieate store of Morsses or Sea Oxen, wlii<'h were a sleepc upon liic rock<'S: Imt when we apinnachcd nere unto them uilli our lutale tlie,\ east liieiir-el\e> into tiie >ea and j)iirsiied ii.> with such fiirie as that ue were ;^lad to lice from lliein." It is later said that the nnmlMr of these "Sea Oxen'" was "ahoiil thirt\ or fortv.":j l''rom Ihe accounts of e -'Sea < )xen " u ere a<'custoiiied to resort to these \arioiis islands during Apiil, .Miiv and .liine. loi' Ihe. purpose of luin;;ili;; forth t heir \oiiii;:. 'I'liiis, ■■'rhoinas .lames ol' iJristoll,"' in sjieakin^ of t he " Isle of l.'amea.'" sa.\ s it was situated "in 17 de;irees, some lift ie lea;4ues from thedrand IJas. lie. re New foniidlaiid : and i-^alioiil I w eiit ie lea.uiu s aiioiit . and soiiH' pail of the Island is Hal Sands and shoiilds: and IIk list I commeih on liaiike do do ilieir kiiidcj in Ajiril, Mav i\: ■liiiie. by iiiiiiihers of I liou>aiids. w hicli lish is verv hi;;: ami halh tun ^I'eat tcelh; and I he skiniie of I lu'iii is lik< Iliiffi'S l« al her : ami l apjiiaiaiice ol' rain, tlie,\ immediately iclicil Id the water with ;4real preeipilal ion. The\ are, when oh! of I he w aler. \ er\ iiii w ield \ , aiioiin;.; about nine, moiitli<. Tl.>e> never ha ve more than two at a time, and seldom more t hail one. '•'I'lie eehoiiri<'s are t'ornieil principally by nat me. bein;4a j^rad- iial slope ol'sol'l rock, with w hich I he .Ma;;(lalen Islands abound; aiioiil so to 100 yards wide at the ualer side, and spreadiii;^ so ;i> lo contain, near Hie siimmil, a very considerable number. I b II- t lie\ are siiirered lo coiae and amuse t liemseh cs lor a eon •idi lable time, till t lie_\ acipiire a boldness, bein;^at their lirsl, laiidin;^ so e.\ceediii;;|\ timid as lo make, il impossible lor any peiMiii lo ajipioach theiii. In a few weeks t hey asscmlle in •^leal numbers; roriiierl,\, wlich iindisi iirbed by tiie AmericanM, !'• Ill'' a moil III of .e\ en or <'i;.;hl ( I ion sand ; and the foi iii ol' the ecjioin ie not allow inj; 1 hem !oremainconlij4iioiis lo the water, llie. ion most ones ine, insiMisibly piisiicd abov(! Ilie slojie. vVlicii ! ■'.3 iij n J fl 'i"r IImI;IiivI, Vi)y(i;4cs, Mil. iii,|i.y;!7. M'li.'iilivoix, viil. \, 11. 'iMi. y-A'l SJti&kiM 68 ODOIi/KM S l.'(tS.M.\i;rs ATI.AN IK UAMM'S. :|i:-'^ '>' '■. ■' f llicy lire Miiivcd to a cuiivciiiciit (lisliiiicc the lislicriiicii, liii\ iiij;' '|ir»»\i(Ic(l tlic iicccssiny apparaliis, kiUc tlic adsaiila.nc <»!' a sea wIihI, tlie slope of the cclioiivlc. the darkness of the nij^lit de- prives tlicm of e\cry direction to the water, so that they stray al»oiil and are killed at leisnre, those that arc nearest the shore bein;; tiie lirst \iclims. In this manner there has Iteen killed (iftecn or sixteen hundred af one cut. They then skin them, and take oil' a coat of fat that always surrounds them, which they dissolve by heal into oil. 1'he skin is cut into slices of two or tlax'c inches wide, and e\]>ortcd to Ameiica for <'arriaye traces, and lo iaiuland for ^^liu'. The teeth is an iid'erior sort of ivory, and is manufactured ibr the same jturposcs, but soon luins yellow."* Accoidin.!.;' to Dj. A. ►S. I'ackard, Jr., its bones are still ibund at th(^ localities mentioned by Hhuldham. "According' to tra- dition," he further says, "it also iidiabited sonu; of the harbors of Cape Breton ; audi have l)een informed by a lisherman in Maine, whose word 1 d(» not doubt, that on an islet near Cape .Sal)le, Nova Scotia [probably tin- '' isle of liamea" of the early voyagers abcady (|uotedj, its bones are found abundantly on the sandy shore, litteen to twenty feet abov<; the sea. In the St. Lawrence (lulf they were extciininated during;' the jniddle of the last ceidury. The last one seen (tr hca.rin, wilting a few years later (in bSOD), in referring to the former oocurrence of the Walrus on the shores TJiil. Trims., yoI. Ixv, p. '^41). tProc. Bust. .Soc, Nat. Ili.sl., v(.l. x, ]«)(>, p. >J71. (UKMIKAI'IIK AL DISIIMIll 'IfoX. no ;iii(l i'-l;iii\iil ('itiii|i;iiiv of .Mis( on", roiiiided (iiiiinj;' tlie e;iilier piiil oT tlie .■.e\eiitee:illi eeiililiA. 1>\ lll<' l\ill,u iil' I'^nilice, ;ind \vli(i>e epIieiiM I'lil eily of New l.'oclielle. iiiliiiiierilii^' ;it one time some llioiisiiiids, li;is jiiissed i!\v;iv le;i\iiiji iio siyn. 'I'lie miir (lered Se;i liorses iinxc lel'l :i more eiidiirini;' monument timii tlie miirdeiers." lie liirtlier iidds: '■'l"lioii,i;li we li;i\e no iiecoiints Inter timn the sevenleentli and ei.uliteentli eeiitiiries of tlieir inliahitin;:' Sable Island,-^*! it is very proltable tliat tliey continued to icsort tliere until tliey entirely left, tliese, latitudes. Its dillieiilty of access, its heiiif^' nninliahitcd, and its sandy liars frinj^cd Avilli a ceaseless snrf, ])oint it out as tlieir last hold."* Di'. (iilpin also r(M'(mls the ('a])tiire of a Walrus in tin; Stiaits (if lielle Isle, Lalirador, in IMarch, ISO!*, which was diaimcd on liie ice lor li\(' miles, and then taken by ship to S!. .lohii's. New I'onndlaiid. aiid thence to Ilaliiax. No\ii Scotia, where it was desciilied and li'^iired Ity Dr. (''lusion."§ *!' . <'»J» ■ I'ror. .'iiiil 'I'raiis. Xova Si^oria JumI. Nat. Si'i., vol. ii, pt. I!, ])p. lv''M27. t lliid.. 11)1. l-.':!-1"^7, witli a plate. i/.o.'iinMi-i, 1871, -rjoO. ^Vv'H'. /,oul. Si)c. liond., 1808, p. 4',V.\. '() !iii(l:iiil ;il)(iiit Tori iMJiillic, on llic wcslciii coiist ol' (licchl.'iinl. ill liililiidr Til', but tlicyscciii Id Innr, >iiic(f I li;il ihilc. nical ly tlrcrcwsrd in niiiiilicis iiloiiu I lie w liolr nf tlic ( 1 l((ill;l!|(| en; I si. ClI pill ill i'ciN Icll. Ill ili- | i:i piT oil t lie •• .Mlllli II 111 11; I (li Noilli ( Irccnliiml niid ( IiiiiimH I-iiiid," oIiscia cd in is;,') h,\ llic r.iilisli Arctic I'Apcdil ion, iil'lcr iilliidiii.L; to tlicir roiMici' ;iliiiiid;iiicc .ihoiil I'ort I'onllvc. ns ol»scr\ fd hy l\;iiic;iiid lliiycs, Miys: •• < 'iiiioiisly ciioii;;ii. wcdid not sec one ol I licsc iiniiiiiils ill llic \ iciiiily of Toil i'oiilkc nor in Sinitli Soi !«•, until wt; rciiclicd l''i;iiil3 and (tS'5 >;. lat., where it oc<'iirs pretty numerously ;it times, '^riie darin,n' task of enteiin;^' into c(»iitest with this animal from the kayak on the open sea forms a re,L,uIar sport to the natives (d' Kati.i;amiut in (>i'P X. lat. Tiie num'ier yi'urly killed has not been separatel\ calciilaled, . . . but they can hardl.\ ex(i<."|: 'I'lie westeininost point at which it has been oliserxcd is .--aid to be t he western shore of lliidsop's liay. Mr. -I. ( '. itoss stales it to bean iiihabitanl of tlie wi'Si coast of I'atlin's May and "'I'lir /(ii'i|(i;;h t, :',(! ^c|■., \n\. i, |i. ol)!!, Sc|)l .•llllicr, l'"^/". ( III MSS. IKllrs lie JKIS liilldJS' pJMCld ill IMV ilis|ios;il. I Diiliisli < ;i((iil.niil, ils i'((p]p|i' :iii(i its I'inilncls. pp. I'JIi-JiT, tH77. n fJI'.ocIv'AI'lIK ,M, DISTKIItTITION. 71 : ri' hi m i;r|(lll>i' I!;IV- mill lo lie ((r!';isi(tli;ill\ liH'l willl ill I ln' IMilllicni |;iil (,r riiiicf IJci^cnrs Inlci, lull siiys it is iiiikiiow ii lo llic ii;ili\ (--ol Itoiilliiii.* Dr. Kicliaidsoii s;i\s: ''Tiic \\ iiliiiscs wcrii \ci\ iiiiiiicroiis ill I.L;I(»(»lik iiiid uit Ilic oilier |t;iits uT I lie <'(iii,sl Id iIm' cMstwiird of lilt' l''iiry siimI llccia's Sirait. Tlirs arc not loiiiKi. Iiowcxcr. al IIh' iiioiiIIi of llic Copper Aline Ijiver, ;il!li(i!i,'^li liie hiaek w hail' liati lieeii soiiKjIiiiies (Irilled I liillier."t lie also rel'ers lo lis heiiin iiiikiiown lo tlie Ivskiiiio.' ol' the ('opperiiiiiie and Mat-ken/ie ifivers. ;j; No species of Walrus ;ippeais lo lia\t' excr Itceii seen on tin' Arctic c(tasl of America helwceii llic !l7lli and l."),Slli iiierialnied luiiong th('ir Jolk-lore, may he th(^ Waliiis."H I''!eming- states that one was killed in the Sound of Stockness, on the east (!oast of Iluiris, in December, 1817, 1[ while another, accoidin;;- to Maegillivray and others, was killed in Oikney in .Fiine, 1S2."».** Mr. \i. Brown adds that onci was seen in Orkney in IS57, and another in IjTor' Isles abont tlio siune time. ft It appeals tohave !iever ocenrred in Iceland, ex- cept as a li' (; straggler. Many years ago tliey are said to have lived on the shonss of I'^inmark, and at a innch lat< <• date to haveabonndedonsonujof the i.slands oil" this coast. Mi'. Lamout says: " We learn from the voyag(! of OhthcMv, wiiich was per- formed aboid a tlnaisand yeais ago, that tlie AN'alriis then abounded on the coast of I'inmarken itself; tiieyhave. however, abandoned 'hat coast for some eentnries, although indi\idnal stragglers lia\e been occasionally cajttnred there ii|» to within ■' Ivd.ss's ■.'(! \()y., All]!., is;!."), p. wi. tSiiii|)l. I';iri-.v".s ','(1 Voy,, |i. ;!:if. t Ziii'ilo^y of IJccclii'y's V^ty;i;j;c, Mam., !-•:'>'.', p. <>. VI lli'ctor lidccc's History of ScotliiiHl. .■is(|iiot('cl liy I'rili.sli /(iiilni;i>its. 1! I'loc Zoill. S(ir. I.ohil.. HiW, |i. \:\\\. • isiiiisii \iiiiiKii>, p. i:i. ■'lidiiili. New I'liil. .(oiini., \ (il. ii, p. !!'■'!•; I5ri)isli (,!ii;iut still in the southeast <'orii<>i'|, and Jiyk Yse Island, which in their turn aic now veiy inleiior lniiitin;^-j;round to the Itanks and sUeiiies lyinj;' to the north of Sititzherficn. " J'oilninit«'ly Cor the persecuted Walruses, liowever,tliese lat- tei' disti'icts are only accessilile, in open seasons, or ])erliaps once in three or loui' summers, so that, tliey }ivt a little breathinji' time there to i)reed and re])lenisli their inimhcrs, oi' undoubt- edly the next twenty or Ihiity years would witness the total extinction of Hosmtirns Irirlicrns on the coasts of the islands of Northern I'^urope. "Th(^ Walrus is also loundail round the (!oasls (»f No\a Zeni- l>la, but not in such ninubejs as at Spitzberj^en; and he inider- g'oes, if |)ossible, more jx'rsecution in those islands frctin some colonies of Itussians or Samo'icdc's, who, I am told, reuularly winter in Xova Zembia ibr tlu' purpose of hunt inj^' and lish- in,yv'f ''The war of extermination,'' sa\s Mr. Kamoiil, in his later Avork. •• which has been cariied on Ibr many years in Spitzi)er- j;en and N(»vaya Zendya has driven all the Arctic faiuia jmain- iiialsj from their old haunts, and. in seekinj;' I'ctreats more inac- cessible to man, it is piobable that they have had in some defiTce to alter their habits. I'or exaMi]>le, up to about twenty years aj^o it was eustomai'y for all ^\'alrus hunteis to entertain a reasonable lio|»e that by waitinj^' till lat(> in the season all Ibr- mer ill-luck miyht be com])ensated in a few fortunate hours by Ifillinj;' some hundreds on shore; in fact, favorite haunts were AV( II known to the lishers, and were visited successively before linally lea\ in;; the liuntinj;j;i'ounds. >iow, althouj>h the Arctic seas ai'c ex])l(U'e(l by steamers and visited anraially by as l)o]d and enlerprisinj'' hunteis as foimerly, siu-h a windfall as a herd of Walruses ashore is seldom heard of, " Each year better found \ essels and mor(^ elaborate weai)ons * Jb\ LiiiiKiJil liassiiMT rcpoilcd the iiipl iiic of :i Ijuj^o IhiII " in Mm^^cii") Soiuid near Ihc, NortJi ('.i|ic aliunl iHCiri." — Vdrhl'nuj in ihc Arctic licua, \\. 58, foodioic. t Hi'ii.sdii.s with the (Sra-lior.scs, pp. 1(17, Kib. (;i;(»(ii;Ai'iii(\i. distimisi ikin. I ■( jUf sent "iiil In li;iri\ tin- Wiilriis; ;is ii n»iisc(|iifii<'(' cn civ mm- .■soii tlicrc is niiMicr «lir(i('iill v in olttiiiiiiii;;- ;i (miuh — lor two rc;i- sniis. I iKisriiliiliKiis wliirli li;i\r \i ■mi I led iliti) w li;il w iis siifc Irr*! iiiy ;;!()iiml liisl ,vc;ir iiict'l liiciiciH-iuv, ;iii«l liiiil' iiic killed, u liilc the (itlicr liiiir csciiiiiii,'^ will he IoiiimI iirxl ,\ riir ;i sIc)) r;nl Ikt ii\\;i\. Tliis iiilclliuciil rcli'ciitiii^' ul' the Wiiliiis hciorc ;i siipc lior nicDiN will, 1 hclicxc, pi'csrrNr I lie species iii'iei' its sciii'eily ill aecessihle w;ilers reinlers it no loiter ;iii oliject of sport niiiiiiiierce. Tliiit tli( \\ ;ilriis, ... is heiiij^ (Iriveii IVoiii every (iistrict where llie li:iii(l of iiiiiii is lelt, is eerlaiii."* Mr. Alfreil Newton, wriliiij;' in lS(>t, iMjsjx'ctiii^' llicir loniier ])reseiiee on the eoiists of J<''iiMii:irk, iiiid theii'distriliiitioii at tliiit .serves : '• I se«' no icason to doiild (he a.sseition, oi jier- liaps it would he safer to say the inference, that in former days Walruses hahitiially frecpieiited the coasts of i'lniiiark'. fn tlie sixteenth and seAcnteeiith ceiitiirios t hey were certainly ahniid- aiit altoiit l!ear I.-laiid; llie,\ are spoken of there as ' iyin;^' Iik(^ houses upon heaps, '|t| . . .: ye) for the last thirty years proli- alily not one has lieeii seen there. Now they are lieninied in l»y (he packed ice of the i'olarSea on the one side and (heir nicrciless eiieiiiies on the other, 'i'he result cannot admit of any r);;-l."».i|, we r<'ad: '' 'I'o the North part of that Conn trey are the places wliere they have their i''uircs. as .Sahles, Martenis, <;re('se iievci's, Foxt's white, hiacke, and icd, Minkiis, ]']riiiiiies, Minivers, ami Harts, Therc^ are als(» a lisiios te(3th, wliicli fish is called a ."Morsse. The, takers fiicreof dwell in a place called i'ostesoia, which hrin;; them njxni Harts to Liim- pas to sell, and from LitiujxiN carrie tiieiii to a place calh.'d Col- inoyro, wiien^ the hij^li Market is holden on Saint Nicolas day,"§ On Ilondins's iiia|> of I'nssia a(;(;onipanyin|). 2i:i, 'Z\\. ' 'm ><■! :f Ife t K;y Miit^-*'' 'I ,1' 'V , 74 oi.oit.r.Ms i;nsM,\i;i s — \i i.amic \v am; rs. ' ■{' ' 'Wf ;.:' 'I'lir Wiilrii.scs ii|i|i(;;!r lo lia\ (' Itccii 111^1 iiici wiili on ( 'liciii Island in H>n.">, inid lolinxc iH-cnmc n,'arl\ r\|ci minalcil iIhtc w il liin a \ civ I'l'W ,\cai' •. 'I'lic liislois nl' I hric dc-l i iicl ion 1 lnic and a! S|iit/.lHi!^ pari oT ilic ■.(•\cnli lul li cent ni.\ is ;^i\rn in tin- lollow in.i: cxcci pis : •• In I lie v( iiic hio."). Sirjdicn llcnni't was iinplo\cd lt\ I lie roiapanic." in a ."-Wiip called llic ^//vfrf. |o iIkisc parls X((il liuards ol' ijic <'aiic|--or Ni!r\\a\ "|, ainl \\a^ a I ( 'lici ii' I hind :\\\i\ Ixillcd ;.ohic S( aiiorsc.s. and ltroii;^lil home head Oaic I'loin tlicn<'c . . . •■ 1 1 cere It is to lice mid erst ood, that t he ( 'on i panic !ia\ iii.^ I>\ (d'teii I'csorl and iiiiplo\ iiient to lliose parts. oiisei\c," says tliat as they approa'lie. wc went alioard." 'I'lie next da,\ they went on shore ayain and * liii( irpiii.it til ,.(iiiir I iiiic pridi' to 1 i.i' yciir iririti, uih1r('ak(! it, if it lice not nil the heller phited. Tliey will also strike with their Tectli ill him tliiit is next llieni: lint l)eeaiis(^ their Teeth <;row downward, I heir str(>]ces are of small force and danj^er." They took in ••ele\('n t mines of Oyle, and the teeth of all the beasts ;i(on's;ii(l." '{"he I'ollowin;; year (KJOfI) they aj^ain set oid for Cherie Island, arriving theic Jidy .".. They found Die ice still about tin; island, and the Walruses not yet on slioic; '' I'or their nature is sucli, thill tlie.\ will not come on land iis lon^j; as any lee is about the liiiid." ( )n the Ilth they ])erceived on shore "of the beiists sidlicieni to iiiiike our xoviijic, \ve(' ])repared to ;•■()(' killing. Miister Wrhlcii and Miister /iV'?//(f7 appointed nice totidv'e eleven Mien with mee, and to }^o(! l)eyond the beasts wlieic they lay; tiiiit they iuid w(M' mi;4ht meet at the middest of them, and so enclose them, thiit none of them should ;i('t into the Sea, . . . . iind before six lionres were ended, we li;id slayne ;d)oiit sexcii or eijiiit hundred Ueasts. . . . l''or ten diiyes spae(^ we jilved our businesse veiy hard, and brought it almost to nn end." They took in "two and twentie tuns of the Oyle of the IMorses, and three hogsheads of their Teeth." in KiOS they :ig;iin reaehed Cherie island towaid the (Mid of -bine, and on the 22d "came into a CoveAvherc the .Morses were, and slew abont OOO. or ]()()(». of them in less than seven hoiires: and then we plied our business untill the sceond of .Inly : at wliiit time Ave Imd tidcon into our shi]) L'L'. tuniies and three hogs- heads of Oyle." On their icliirn they look with theia two live young Walruses, on(! of which lived till they reached London.* The voyage in 100(1 w;is less successful. They slew at one tiiiK! eighty, at iinotlier one hundred and liity, iind id siill iin- "I'mcliii.s liis I'ilniiincs, vol. iii, |i|i. ri.">7-r)(j(). -. -wn .r.^ IMAGE EVALUATION TEST TARGET (MT-3) ^m /. ^ i ^ 1.0 I.I 1^ 1^ 1^ US ^ y£ III 2.0 1.8 11.25 1111.4 ill 1.6 % 7 f W > ^^ ^^^.-^^ >!^ O? 'ni 4^ I 76 ODOB^NUS ROSMARTJS — ATLANTIC WALRUS. other time forty-five; but they lost most of tliem in coiiseqiieuce of bad weiitber. " In the yeore IGIO. the Coini>anie set out iwo Ships, viz. the Lionesse for CJicrie Hand, Thonun IJdpe Com- mander ; and the Amitie, for a Kortherne Discoverie, tlie Mas- ter of which ship was Jonas Foole : who in the moneth of May fell with a Land, and called it Greenland, this is the Land that was discovered by Sir Hugh WiUonghby long before [Spctsberg of the Hollanders], which Shij) Anitie continued upon the coast of Greenland, discovering the Harbours and killing of Morces [the first killed by the English on Spit>.bergen], untill the moneth of August, and so returned for England, having gotten about some twelve Tunnes of goods, and an Unicornes home. "In the yeere 1611. the Companie set foorth two Ships, the Mark Margaret Admirall, burthen one hundred and sixtie tunnes, Thomas Edge Commander ; and the Elizabeth, burthen sixtie tunnes, Jonas Poole Master, well manned and furnished •with all necessarie Provisions, they departed from Blackwall the twentieth of Aprill, and arrived at the Foreland in Green- land in the Latitude of 79. degrees, the twentieth of May fol- lowing, the Admirall had in her six Biskayners expert men for the killing of the Whale : this was the first yeere the Compa- nie set out for the killing of Whales in Greenland, and about the twelfth of June the Biskayners killed a small Whale, which jeelded twelve Tunnes of Oyle, being the first Oyle that ever was made in Greenland. The Companies two Shalops looking about the Harbour for Whales, about the five and twentieth of June rowing into Sir Thomas Smith his Bay, on the East side of the Sound saw on the shoare great store of Sea-horses: after they had found the Morses they presently rowed unto the ship, being in crosse Eoad seven leagues off, and acquainted the Cap- tayne what they had found. The Captaync i nderstanding of it, gave order to the Master, Stephen Bennet, that he should take into his Ship flftie tunnes of emptie Caske, and set sayje with the Ship to goe into Foule Sound. The Captayne went pres- ently aw{iy in one Shallop with sixe men unto the Seamorse, and tooke with him L&nces, and comming to them they set on them and killed five hundred Morses, and kept one thousand Morses living on shoare, because it was not profitable to kill them all at one time. The next day the Ship being gone unto the place & well mored where the Morse were killed, all the men belonging to the Ship went on shoare, to worke and make Oyle of the Morses ; and when they had wrought two or three dayes, GEOGRAPHICAL DISTRIBUTION. 77 it fortuued that a small quantitie of Ice come out of Foule Sound, and put the Ship from her Moriiig, . . . The Ship being cast away without hope of recoverie, the Commander Thomas Edge gave order, that all the Morse li^aug on shoare shoold be let goe into the Sea, and so gave over making of Oylc. . . ." Fitting np their boats as Avell as they could they soon after abandoned the coast of Spitzbergen (" Green- land "), and set sail for Cherie Island, ^\ here they found the "Elizabeth" and returned to Spitzbergen "to take in such Goods as the sayd Edge had left in Foule Sound, woorth flfteene hundred pounds.'*" As early as the year 1611, the previous persecutions of the Walruses at Cherie Island had made them very wary. Thomas Finch, in his account of a visit to this island by William Gom'- don in August of that year, says: "At our comming to the Hand, wee had three or foure dayes together very fine weather: in which time came in reasonable store of Morses, . . . yet by no meanes would they go on those beaches and places, that formerly they have been killed on. But fortie or flftie of them together, went into little holes within the Eocke, which were so little, steepe and slipperie, that as soone as wee did approach towards them, they would tumble all into the sea. The like whereof by the Masters and William Oourdons leyort, was never done."t During the years 1612, 1613, 1614, and 1615, numerous vessels were sent out from England to Spitzbergen for the products of the Walruses and Whales, but generally met with indifferent success, being much troubled with Spanish, Dutch, and Dan- ish " interlopers." ' ' •' ' ■' " In the yeere 1616, the Companie set out for Greenland eight Sayle of great ships, and two Pinnasses under the command of Thomas Edge, v, ho following his course, arrived in Greenland about the fourtii of June, having formerly appointed all his ships for their severall Harbours, for their making of their Voy- age upon the Whale, and having in every Harbour a sufficient number of expert men, and all provisions fitting for such a Voy- age. This yeere it pleased God to blesse them by their labours, that they full laded all their ships with Oyle, and left an over- plus in the Countrey^ which their ships could not take in. They imployed this yeere a small Pinnasse unto the East-ward part of Greenland, Namely, the Hand called now Edges Hand, * PurcluiH his Pilgrimes, vol. iii, pp. 4(54, 465 tlbid., p. 536. 78 ODOBiENUS ROSMARUS — ATLANTIC WALRUS. and other Hands lying to the Korth-wards as farre as seventie eight degrees, this Pinnasse was some twentie tunnes, and had t\.elve men in her, who killed one thousand Sea-horses on ^flges Hand, and brought all their Teeth home for London^ In 1017, they " employed a ship of sixtie tunnes, with twenty men in her, who discovered to the Eastward of Greenland, as farre J^ or th- wards as seventy-nine degrees, and an Hand which he named Witches Hand, and divers other Hands as by the Map appearoth, and killed store of Sea-horses there . . ."* The Dutch, Danes, and Spaniards began, in 1612, also to visit Spitzbergen in pursuit of Whales and Sea-horses, but axe reported by the EngUsli to have made indifferent voyages. The company soon also had rivals in the " Hull-men," who, as well as the Dutch, did them much "ill service."! About the years 1611 and 1612, the Whale-fishery was found to be more profitable than Walrus-hunting, and subsequently became the main pursuit, not only by the English, but by the Dutch and Danes. Yet the Walruses were by no means left wh jily unmolested, having been constantly hunted, with more or less persistency, down to the present day, and, as already shown, were long since exterminated from Oherie Island and other smaller islands more to the northward, and greatly re- duced in numbers on the shores of Spitzbergen. Walruses have been recently reported as occurring on the outer or northwestern coast of Nova. Zembla, but as not exist- ing on the inner or southeastern coast. Von Baer, on the au- thority of S. G. Gmeliu and others, gave the eastern lunit of the distribution of the Atlantic Walruses as the mouth of the Jene- sei River, though very rarely single individuals wandered as far eastward as the Piasina Kiver. He even regarded the Gulf of Obi as almost beyond tlieir true home.| Von Middendorff, however, considers von Baer's eastern limit as incorrect, and cites old Eussian manuscript log-books ("handschriftliche Schiftsbiicher") in proof of their occurrence in numbers in Au- gust, 1736, as far east as the eastern Taimyr Peninsula, and of their being met with in August, 1739, as far east as Chatauja Bay. Still further eastward, in the vicinity of the mouth oi the Lena Elver, he gives similar authority for their occurrence in August, 1735, and says that Dr. Figurin attests tlieir presence *Purclia8 his Pilgrimes, vol. iii, p. 467. tibid., pp. 472, 473. tMdni. do I'Acad. des Sci. de St. P(5tersb.. vi° 8<5r., Sci. math., phys. etnat., tome iv, 2''" pars, pp. 174, 184. GEOGRAPHICAL DISTRIBUTION. n -on the shores of the delta-islands of the Lena. Eespecting the move easterly coast of the Siberian Ice Sea, he says it is cer- tainly known that the Walruses of B iring's Sea extend west- ward in great numbers to Koljutschiu Island. Only the males, however, reach this limit, th'j females not extending beyond the Aicinity of the mouth of the Kolyma Eiver.* It lience appears that about 1735 to 1739 Walruses were met with as far eastward as the mouth of the Lena Eiver; but Wrangell, nearly a century later, explored quite thoroughly this whole region without meeting with them, and I have found only one reference to their existence on the Siberian coast be- tween the Kolyma and Jenesei Elvers later than those cited by von Middendorff. According to a recent letter t from Professor iNordenskjold, of the Swedish Northeast Passage Expedition, "two Walruses" T\ere seen in August, 1878, a little to the eastward of the Jenesei River, and that open water was found as far as the mouth of the Lena. From this it would seem that there is nothing to prevent, at least in favorable years, the W^alruses from passing eastward to the mouth of the Lena. There still remains, how- ever, a breadth of some thirty degrees of longitude (between 13(P and 3 60°) where as yet no Walruses have been seen. They appear to have been only verj^ rarely met with to the eastward of the Jenesei (longitude 82° E.), and to be uncommon east of tlie Gulf of Obi. At present the Atlantic Wiilrus ranges along the northeast- eru coast of North America from Labrador northward to Ee- pulse Bay and Prince Eegent's Inlet, and along the shores of Greenland; in the Old World only about the islands and in the icy seas to the northward of Eastern Europe and the neigh- boring portions of Western Asia, where it rarely, if ever, now visits the shores of the continent. On the easttmi coast of North America, Walruses have been met with as fiir north as explorers have penetrated, and as far as the Esquimaux live. They winter as far north as they can find open water, retiring southward in autumn before the ad- vance of the unbroken ice-sheet. Kane speaks of their remain- ing in Eenssellaer Harbor (latitude 78° 37') in 1853, till the sec- ond week of September, when the temperature reached zero of Pahreuheit. | * Von MiddeiKloiiPs Sibiiischo Reise, Bd. iv, 1867, pp. 935, 936. t Soo Nature, vol. xix, p. 102, December 5, 1878. t Arctic Exploration, vol. i, p. 140. 80 ODOB^NUS ROSMARUS ATLANTIC WALRUS. Nomenclature. — Several specific ntunes have been iii moro or less current use for the Atlantic Walrus, or rather for the Atlantic and Pacific species collectively. Accepting Odoba'nus as the proper generic name of the group, there is nothing to prevent the adoption of rosmarm for the specific name of the Atlantic species. It Avas used for this species exclusively by Linn<;, Erxleben, and other early systematic writers, the Pacific Walrus being at that time unknown to the systematists. If lios- marus be used as the geneiic name of the group, as it lias been by a few late writers, as a substitute for the wholly untenable one of Trichechtis, it will be, of course, necessary to adopt some other name for the si)ecies. Dr. Gill has used obesus of Illiger ; but as this was applied by Illiger exclusively to the Pacific Wal- rus, it cannot properly be used for the Atlantic species. It would be difl&cidt to select a subsequent name that would not be open to objection, if one should stop short of trichechus, used (inadvertently?) in a specific sense {^^ Rosmarus trichechus^') by Lament in 1861. The name lonaidem of Fremery, 1831, was based on what subsequent writers have considered as probably the female, but the name is highly inai)propriate, inasmuch as it is the Pacific species, and not the Atlantic, that has the longer tusks. There are left virginianus of DeKay and dtMm of Stannius : the first is objectionable on account of its geograph- ical significance ; the other is only doubtfully referable to the Atlantic species. Adopting Odohcenus for the genus, leaves rosmartts available for the species, thus settling the whole diffi- culty. As aheady noticed [antea, p. 20), two species besides virgi- nianus have been based on fossil remains, and have been made the basis of new genera. The first of these is the Odobenothe- rium lartetianum of Gratiolet, since referred by Defrance to the existi":'ig species ; the other is the Tricheeodon huxleyi of Lan- kester, which there is perhaps reason for regarding as the large extinct progenitor of the existing Walruses. Etymology. — The term rosmarm was originally used by Olaus Magnus, about the middle of the sixteenth century, in a vernacular sense, interchangeably with morstis, the Latinized form of the Eussian word niorsz (or morss). It was used in tho same way by Geaner a few years later, as well as by numerous other pre-Linnaean authors. Eespecting the etymology of the word, von Baer gives the following : " In dem historisch-topo- ETYMOLOGY. m grapliischen Werke : Be gentium septentrionalium conditionibus cet. Komae 1555 lieisst es : Normgium littns maximoa ac grandes places elephantis hahet, qui morsi seu rosmari vocantur, forsitan oh asperitate mordendi sic appellati, (Eine recht witzige Etymo- logic !) quia, si quem hominem in maris littore viderint apprehen- dereque poterint, in eum celerime insiliunt, ac dente lacerant et in momento interimunty * The same author also gives the following from Herbeistain (1567): "Under andern ist auch ein thier, so grosse wie ein ochs, und von den einwonem Mors oder der Tod geheissen wird." t Hence, either from superstitious notions of the terri- ble character of this animal, or from the resemblance of the iiussian word morss to the Latin word mors, these terms be- came early confounded, and rendered by the German word Tod, or death, f In the account of the exploits of the Norman Othere, where the Walrus first finds its place in literature, it is termed Horse- wael. As noted by Martens § and other writers, equivalent words in other languages have become current for this animal, as Walross or Wallross of the Germans, Wallrus of the Dutch, *In an early (1658) English version of Olaus Magnus's work ("A Com- pendious > ''story of the Goths, Swedes, &, Vandals and other Northern Nations. .itten by Olaus Magnus, Arch-Bishop of Upsal, and Metropoli- tan of Sweden", i». 231), this passage is rendered as follows: "The Xorway Coast, toward the more Northern parts, hath huge great Fish as big as Ele- phants, which are called Morn, or Rosmari, may be they aiie so from their sharp biting ; for if they see any man on the Sea-shore, and can catch him, they come suddenly upon him, and rend him with their Teeth, that they will kill him in a trice." From this it would appear that Morsua, as used by Olaus Magnus, might bo simply the Latin word morsus, from mordere, to bite, tSee von Baer, M^m, de Acad, des Sci. do St. Pdtersb., vi« 8<5r., So. math., phys. et nat., tome iv, 2^" pars, pp. 112, 113. ^ Von Baer quotes a passage from the "Rerum Moscoviticarum auctores varii," originally published early in the sixteenth century, in which occurs the phrase " scandut ex mari pisces morss nuncupati," which he regards as the first introduction into Latin of the Sclavic name MopiKb. In Western Europe it a little ^ater became current in the form oi Morse, whicii was soon written Moras or Mors, from which Buflfon later forme*' the name Morse, which has since been the common appellation of this animal among French ^vrite^9. Von Baer further observes that the accidental resemblance in sound of this word to that of the Latin word for death (mors) appears to have contributed not a little to the strange conception of the torribleness of this animal whicli was early entertained and even still prevails in Western Europe, although the Russian accounts do not speak of it. «Zoolog, Garten, Jahrg. xi, 1870, p. 283, where the etymology of tho numes of tho Walvus is briefly discussed. Misc. Pub. No. 12 6 82 ODOBiENUS R08MARUS — ATLANTIC WALRUS. and Walrm of the English. By the early Scandinavian writer* it was termed Bosmhvzlr, which later became resolved into Boa- mul, from which, perhaps, originated the Latin term Boamarm, which has the same significance, introduced by Olaus Magnus and Gesner, and the Norwegian word Boatungr. Gesner and several subsequent writers also used the word Meerroaa, and we have in English the equivalent term Sea-horae^ as one of the ap- pellations of the Walrus, and also, but more rarely, Meerpferd in German, and Clwval marin in French. The current French term Morae appears, as already stated, to have been introduced by Buffon as a modification of the Eus- sian word moraa, used by Michow (1517) and Herberstain (1649). Among other old vernacular names we find in English Sea Coic, in French Vache marine, in Latin Boa marinm^ etc., while by the early French settlers in America it was commonly termed Bete f> la grande dent. ^ Literature. — 1. General Hiatory. — Passing over the by some supposed allusions to the Walrus by Pliny as too vague and uncertain for positive identification, * we meet, according to von Baer, with ^^he flist positive reference to the present species in the account of the exploits of the famous Norman ex- plorer Othere, or Octher, who, about the year 871 (890 accord- ing to some authorities), made a voyage to some point beyond the North Cape, where he met with large herds of Walruses, some of the tusks of which he is said to have taken to England as a present to King Alfred, t Walruses appear to have been * See K. E. von Baer, M6in, de I'Acad. Imp. des Sci. de St. Pdtersb., vi'"" s^r., Sci. math., pliys. et nat., tome iv, 3™« livr., 1836, (1837), pp. 101, 102. T .. this admirable monograph I am greatly indebted for information respecting the earlier publications bearing upon the history of the Walruses. To this exhaustive memoir the reader is referred for a full exposition of this part of the subject. The following short summary is based, so far as the early his- tory of the subject is concerned, mainly upon von Baer's monograph, an analysis of which will be presented at a subsequent page. (See posted,, p. 88, footnote.) t Hakluyt's rendering of this account is as follows: "The principall purpose of his [Othere's] traveile this way, was to encrcase the knowledge and discoverio of those coasts and countreyes, for the more comoditie of fish- ing of horsowhalcH, which have in their teeth bones of great price and ex- cellcncio : whereof lie brought some at his retume unto the king. Their skinnes are also very {\;ood to make cables for shippcs, and so used. This kind of whale is much lesse in quantitie then other kindes, having not in length above seven elles." — Hakluyt's Voyages, vol. i, p. 5. i GENERAL HISTORY. 88 an object of chase on the coast of Finmark as early as 980, and must have been met with by the Norsemen when they visited Greenland about the end of the tenth century. Their tusks were an article of commercial value among the Mongolian and Tartar tribes as early as the twelfth to the fifteenth centuries. Aside from the various notices by Scandinavian writers, the earliest unmistakable reference to the Walrus, other than that connected with Othere, as above mentioned, was, according to von Baer (1. c, p. 108), by Albertuf^ Magnus, in the first half of the thirteenth century. Says this writer (as quoted by von Baer), whose account is here paraphrased : The hairy Cetaceans have very long tusks, by which they suspend themselves to the rocks in order to sleep. Then comes the fisherman and separates near the tail as much skin as he can from the underlying fat, and then attaches a cord, which has at the other end a large ring, which he makes fast to a post or tree. Then when the fish awakens (by all of these operations he was not yet awakened), they cast a huge sliug-stone upon his head. Being aroused, he attempts to get away, and is held by the tail near to the place and captured, either swimming in the water or half alive on the shore. This ludicrous description von Baer believes had for its foundation misunderstood reports of the Walruses' habit of reposing upon the shore or upon ice-bergs, the use of their tusks in climbing up to these places of rest, and their deep sleep, and that the account of the mode of capture was based on an incorrect knowledge of the use of the harpoon; and that the account shows that as early as the thirteenth century the Walrus was harpooned on the coast extending from the White Sea north- wards. * * This curious legend is quoted by Gesner in his Historia Animalia Aqua- tilia, 1558, p. 254. The following rendering appears also in the above-cited English version of Olaus Magnus : ' ' Therefore, these Fish called Boamari, or MorA, have heads fashioned like to an Oxes, and a hairy Skin, and hair grow- ing as thick as straw or corn-reeds, that lye loose very largely. They will raise themselves with their Teeth as by Ladders to the very tops of Rocks, that they may feed on the Dewie Grasse, or fresh 'Vater, and role themselves in it, and then go to the Sea again, unless in the mean while they fall very fiat asleep, and rest upon the Rocks, for then Fisher-men make all the has.te they can, and begin at the Tail, and part the Skin from the Fat ; and into this that is parted, they put most strong cords, and fasten them on the rug- ged Rocks, or Trees, that are near; then they throw stones at his head, out of a Sling, to raise him, and they compel him to descend, spoiled of the greatest part of his Skin which is'fastened to the Ropes : ho being thereby 84 ODOB^NUS R08MARUS— ATLANTIC WALRUS. The Walrus is also referred to by Hector Boethius in 1526, in bis History of Scotland ; * by Herberstain (or Herberstein, as also written) in 1549; by Par6 about the year 1600; and by Al- drovandus in 1642. Herberstain also very correctly indicates the habits of these animals, which, he says, repair to the shore in large herds to repose, and that while the herd sleeps one of their number keeps watch. He compares their feet to those of the Beaver, and refers to the value of their tusks tc the Russians, Turks, and Tartars, and observes that they called them fish-teeth, t Even before the middle of the sixteenth century. Walruses had been met with on the eastern shore of North America. In May, 1534, they were seen by Cartier, and later in the same century by Fischer, Drake, and others, on the coast of Nova Scotia and adjacent islands, and later still by other explorers on the islands in the Gulf of Saint Lawrence (see antecl, p. 06), in the accounts of whose voyages | occur interesting notices of these animals. In the year 1553, Edward VI of England sent an expedi- tion under Willoughby and Chancellor to the White Sea, which resulted in still further increasing our knowledge of the Wal- ruses, especially of their distribution eastward along the Arctic coast of Europe and Asia. Chancellor's short account § refers especially to the uses made of the skins and tusks. The earliest delineations of the Walrus appear to have been made by Olaus Magnus in his " Tabula Terrarum Septentrio- nalium" (1555), where he has portrayed many strange and fabu- lous animal forms which there is reason to believe were based upon this animal. 1 1 Gesner a few years later (1558), in his " His- debilitated, fearful, and half dead, he is made a rich prey, especially for his Teeth, that are very pretious amongst the Scythians, the Moscovites, Eua- aiana, and Tartars, (as Ivory amongst the Indians) by reason of its hardness, whiteness, and ponderousnesse. For which cause, by excellent industry of Artificers, they are made fit for handles for Javelins : And this is also testi- f'ed by Mechovita, an Historian of Poland, in his double Sarmatia, and FuuIm Jovius after him, relates it by the Relation of one Demetrius, that was sent from the great Duke of Moscovy, to Pope Clement the 7th." — Loc. cit., pp. 231,232. " "Scotorum Regni Descriptio, p. 90," as cit«d by various writers. t Herberstain, as cited by von Baer, 1. c, p. 111. tSee Hakluyt's Voyages, vol. iii, ed. 1810, pp. 237,238,242,249,254, etc. \S See Hakluyt's Voyages, vol. i, ed. 1599, p. 237. II Olaus Magnus's figures will be noticed later under the section devoted to the fi};ure8 of the Walrus (posted, p. 92 et seq.). genp:ral history. 85 toriuiii Aiiiinalium" (in the volnino devoted to the " Animalia AquatiliJi "), tiiitlifully copied all ofOlaiis Magnus's figures under the heading "De Cotis," and then presents, under the name Ensmonis, the figure of the Walrus from Olaus Magnus. This figure, however, he judiciously criticises, stating that the tusks should be in the upper Jaw, and not in the lower, as they were represented by Olaus Magnus. This last-named author, in the later editions of his work " De Gentium Septentrionalium Con- ditionibus," etc. (as in that of 15G3), rightly places, iiecording to von Baer, the tusks in the upper jaw. Gesner (continues von Baer) knew only the first edition of this work, and took his figure from the above-mentioned " Tabula Terrarum Septen- trionalium." Also were unknown to him the aecounts of the Wal- rus given by "Herberstain, Chancellor, and Othere," so that he made extracts from only Michovius and Albertus Magnus. He also knew no better than to offer, as a figure of the Walrus, a drawing he had received from Strassburg, representing, pretty fairly, the head and tusks, while the rest was purely a fabrica- tion. Some rhymes, which he further inflicts upou his readers, show clearly how " awful" the conceptions of the Walrus then were (or, as von Baer puts it, " Wie schauerlich nocli die Yor- stellungen vom Wallrossewaren").* |, >, v ; . In 1008, a young living Walrus was taken to England, having been captured on Bear or Cherie Island off the coast of Nor- way, t while four years later (1012) another yf ung Walrus, with the stuffed skin of its mother, was taken to Holland. The first appears to have been very intelligently described by ^lius Ever- hard Vorstius, whose description is quoted by De Laet. J The specimen taken to Holland was well figured by Hessel Gerard, the young one doubtless from life, the figures being pu^^lished by him in 1613, § and subsequently repeatedly copied (as will be more fully noticed later). In 1625, Purchas, in his history of the voyages of the English to Cherie Island and Spitzbergen (then called "Greenland"), gives much interesting information respecting the chase of the * To ebow what these conceptions were, von Baer cites the passages ahtady quoted (anted,, p. 81), in reference to the singular misinterpretations given in Western Europe to the Russian name Moras. See von Baer, 1. c, p. li:{. . t Recuoil do Voy. au Nord, 2^ 6A., tome ii, p. 368. t Nov. Orb. 8. Doscrip. Ind. Occ, 1633, p. 41. ^ See von Baer, 1. c, p. 128; Gray, Proc. Zool. Soc. Lond., 1853, p. 115. 86 ODOBiENUS ROSMARUS — ATLANTIC WALRUS. Walnis at these islands, and in one place a quaint description and some very curious figures of the animal.* In J 07.5, the Walrus was again described and wretchedly flgured by Martens, t who is said to have been the first " natu- ralist" who ever saw the Walrus in its native haunts. Zorg- drager, | in 1720, supplied by fiir the fuliost account of these animals, as observed by him in Spitzbergen, that had appeared up to that date. He gives not only a quite detailed and truth- ful account of their habits, especially under persecution, but also of their wholesale destruction at that early time in the Spitz- bergen seas, and of their extermination at some of the points at which they had formerly been accustomed to land in immense herds. He also notes the increasing difl&culties of their capture owing to the great shyness of man they had acquired in conse- quence of persecution, and describes the manner in which they were captured, and also their products. Copious extracts from Zorgdrager's account of the Walrus are given by Buffon (trans- lated into French from a German edition), and he has also been extensively quoted by even much later writers. The Greenland Walrus was described by Egede§ in 1741, by Anderson II in 1747, by Ellis If in 1748, by Cranz** in 1765^ and by Fabricius tt in 1780, some of whom added much infor- mation respecting its habits and distribution, its usefulness t'j the natives and their ways of hunting it, as well as respecting its external characters. The above-cited accounts of the Walrus formed the basis of numerous subsequent compilations, and most of those last given are cited by the early systematic writers, few of whom, as pre- ^^ou8ly shown (see antea, pp. 8-11), had any just appreciation of even its most obvious external characters. Linn6, as already noted {anted,, p. 8), profited little by what had been written by preceding authors, while Brisson, Erxleben, and Gmelin manifest a scarcely better acquaintance with this badly misrep- resented and poorly understood creature. No little confu- sion has hence arisen in systematic works respecting its posi- * See anted,, p. 74-78, and. posted. t Spitzbergen, pp. 78-83, pi. P, fig. h. t Bloeyende Opkonist der Alonde en Hedendaagscho Groenlandsclio Vis- Bchery, etc., ed. 1720, pp. 165-172. $ Det gamle Gr(^nland8 nye Perlustration, etc., 1741, p. 45. II Nachrichten von Island, Gronland und der Straw e Davis, p. 258. IT Voyage to Hudson's Bay, p. 134. ** Historie von Gronland, pp. 165, 167. +t Fauna Groenl., p. 4. ■I ( GENERAL niSTORY. 87 tion and aainities (see antca, pp. 7-12). The accounts by Hoiit- tuyn, Buftoii, Pennant, P. S. L. Miiller, and Sclireber are excel- lent for their time. These authors all recofjnized the close relationship of the Walrus to the Seals, and (piite con-ectly indicated its external characters and habits. Some of these accounts, however, include references to both species. Daubenton, in Buffon's "Histoire Naturelle,"* gavo a de- scription and figure of a Walrus's skull, and made the first contribution to our knowledge of its internal anatomy, based on the dissection of a foetal specimen. Since the beginning of the present century, the Walrus liaa been the subject of almost numberless notices, as well as of sev- eral elaborate papers, devoted in most cases to special points in its anatomy, very few of which need be here enumerated, t The elder Cuvier, beginning with his " Legons d'Anatomie compar^e " (1800-1805), and ending with the third edition of his ''Ossemens fossiles" (1825), contributed considerably to our general knowledge of its structure and affinities, especially of its osteology ; he in 1825 1 first figuring and describing its skel- eton. A paper by Sir Everard Home, § in 1824, figured and de- scribed the stomach and feet from specimens taken to England from Hudson's Bay, preserved in salt. This paper is noteworthy mainly on account of the singularly erroneous interpretation there made of the structure and functions of the feet, Home supposing that these organs were provided -with sucking discs, by means of which th« creature was enabled to adhere firmly to the ice in climbin ^. The skeleton of the Walrus was again figured and described by Pander and d'Alton|| in 182G, and still later by BlainviUe ^ about 1840. Von Baer, ** in 1835, published some account of the axterial system of the Walrus, based on a dissection of a young specimen. Its general anato- my, especially its limb-structure, myology, vascular and respi- ratory syste. , viscera and generative organs, and external cha- * Tome xiii, 17*', pp. 415-424, pll. liv, Iv. The skull had been previously figured by Houttuyu (in 1761), as will be noticed later. t Those relating to its dentition have been already noticed in detail (see anted, pp. 47-57) ; several others have also been specially referred to, and nearly all are cited in the references given at pp. 23-26. t Ossom. Foss., 3« dd., tome v, ii™" pt., pp. 521-523, pi. xxxiii. $ Phil. Trans., 1824, pp. 233-241, pi. iv. II Skeloti der Robben und Lamantine, pll. 1, ii. IT Osteographie, Des Phoques, pll. i and iv. ** M6m. de I'Acad. St. Pdtersb., vi"" s4r., Sci. mpt?i., phys. et nat., tome ')■"«. 1835, pp. 199-212. 88 ODOB^.NUS ROSMARUS — ATLANTIC WALRUS. racters, Avere quite fully and satisfactorily treated by Dr. J. Murie* in 1872. lUiger, in 1811, in a paper on the geographical distribution of the mainmals of the Northern Hemisphere (see antea, p. 18), first nominally r cognized the Pacific Wah'us as a species distinct from tie Atlantic animal, while Fremery, in 1831, recognized three species, and Stannius, in 1842, admitted two,t but, as already noticed, only one species of Walrus has been commonly recognized. The matter of variation dependent upon sex, age, and individual peculiarities, has received, as already noticed (see anteu,, pp. 38-43), special attention at the hanas of Wieg- mann, Stannius, Jaeger, and other writers. Unquestionably, the most important paper relating to the lit- erature, geographical distribution, and habits of the Walruses is the well-known an 3. Geziihmte Rob- ben. § 4. Wahro Cetaceen. ^^ .'>. Gesellschaftliches Leben. ^ 6. Liebe dei' Aeltem zu den JuugCii und dor Jungon gegen die Aelteru. ^ 7. Gatten- liiebe. $ 8. Allgemeine Begriinduug dieser Verhaltnisse. Cap. V. Verbreitung dor Wallrosso (pp. 172-204). ^1. Sio wohnen in zwci getrcnnten Verbreituugs-Bezirken. 'J 2. Ocstlicher Verbreituugs-Bezirk. GENERAL HISTORY, 89 published in 1837. This elaborate memoir, so often already cited in the prescsut article, gives a general summary of nearly all papers, referenees, and figures relating to the Walruses that api)cared prior to 1835, the date of its presentation to the Impe- rial Academy of Sciences of Saint Petersburg for publication. It also contains iiumy original biological and anatomical ob- servations, based on a young living specimen brought to Saint Petersbmg in 1828, which, surviving for only a week after its arrival, soon fell into his hands for dissection. * Von Baer, after a few preliminary remarks respecting the occasion and objects of his paper, and a few words on the anatomy of the Walnis, devotes some thirty pages to a critical and exhaustive historical resume of the literature relating to the general subject. Then follow some eighteen pages detailing his observations on the living animal, in which he gives some account of the few young indiAiduals that had, up to that time, been taken alive to Middle Europe ; also a detailed account of the external appearance of the specimen he had examined in life. lie notes especially its attitudes, movements, and limb- structure, and compares it in these points with the Seals. After describing the position and character of the limbs in the Seals, and the restriction of their movements on land to a wriggling movement, with the belly lying on the ground, he refers to the freer use of the extremities possessed by the Walrus, Avhich he found was able to truly stand upon its four feet, and says that, i 'S. Westlicher Verbreituugs-Bezirk. $ 4. Periodische Wanderuugen der Wallrosse. ^ 5. Pliysischo VerliuUnisse, welcho die Verbreitung der Wall- roHse bedingou. Cap. VI. Ehemalige Verbreitung der Wallrosse (pp. 20.'>-228). $ 1. Mei- uuugen hieriiber. ^ 2. Veriinderungen ini Vorkommeu der Wallrosse in den drei letzten Jahrhunderten. $ 3. Ob an den Orkadischen Inseln Wallrosse bis ins 16te Jalirhundert sich aufgebalten haben 1^4. Beweiss, dass, so weit liistorische Nachrichton zuriickgehen, kein Wallrossfang an der Kiiste von Lappland getrieben worden ist. $ 5. Ob die Wallrosse im Mittelalter bei Island liiiufig waren. $ 6. Verbreitung der Wallrosse zur Zeit der Romer uud Griechen. $ 7. Ehemaliges Vorkommen an der Nordkiiste der Conti- uente. . t , , . Cap. VII. Paarung (pp. 228-230). Cap. VIII. Nabrung der Walbosso (pp. 231-233). Cap. IX. Stellung des Wallrosses im Systeme, oder Verwandtscbaft mit andorn Thieren (pp. 234, 2.35). * He seems, however, to have never published in full the results of his observations upon it.s anatomy, he apparently reserving the anatomical part of his memoir in the liope of perfecting it through the study of addi- tional m.aterial. 90 ODOBiEXUS ROSMARUS — ATLANTIC WALRUS. in respect to the use of its limbs, it occupies an intermediate place between the Pinnipeds and the ordinary Ibnr-lboted IMam- mals, amony which latter its less pliant feet jiive it the appear- ance of a crijiple. If we should call, he says, the Seal a crawler or slider, we should have to term the Walrus a waddler, since in walking it throws its pluuij) body to the right and left. Here avc have fairly described, lor the first time, the tlexibility of the ex- tremities, — the bending of the hind feet sometimes forward, sometimes backward, and the free turning of the fore feet, — although an allusion was made to this by Vorstius* two centu- ries before, yet the fact of flexibility remained generally unre- cognized till 1853, when a young living specimen reachetl London, Yon Baer points out the fallacy of Sir Everard Home's notion that the feet of the Walrus are provided with suction -discs, ami the "blowing-' of the Walrus mentioned by Martens, \.-ho de- scribed it as throwing wattr from its nostrils like a whale. Following this chapter on its external features, movements, temperament, behavior, etc., is an interesting dissertation of some twenty or more pages on the domesticability of the marine mammals in general, which is devoted largely to a history of the behavior of the Seals in captivity, with a short notice of the different examples of the Walrus, the Sireniaus, and the smaller Cetaceans that had been observed in confinement. The next thirty pages are given to a discussion of the geogTaph- ical distribution of the Walruses, the treatment of which subject is marked by the same pains-taking research that characterizes the other parts of this learned monograph. He shows that Walruses are confined to two widely separated habitats, and not, as previously supposed, found all along the Arctic coasts. He describes them as limited to two regions, an eastern and a western, the first including, the northwestern coast of North America from the Peninsula of Aliaska northward, and the corresponding parts of the neighboring Asiatic coast. To the eastward he could trace them only to the \icinity of Point Bar- row, and to the w^estward only to a few degrees beyond East Cape. The western region, he affirmed, embraces only the Arctic coast of Europe eastward to the mouth of the Jenesei Biver, * "Pedes nnterioreH antrorsum, posteriores retrorsum siiectabant cmii iii- grederetur," says Vorstius as quoted by De Laet (see anted, p. 37). The hind feet are also represeutcd as turned forward in Hessel Gerard's figure, pulj- lished in 1C13 (see posted). GENERAL HISTORY. 91 and, on the other side of the Atlantic, the shores of Greenland and Arctic America westward to the western shores of Hudson's Bay a 'd Fox Channel. There is thus left between these two regions nearly the whole of the coast of Asia bordering on the Polar Sea on the one hand, and almost the whole of the coast of Xorth America formed by the Arctic Sea on the other. In the later portion of his chapter an the distribution of the Walruses he devotes a few pages to a consideration of their migrations, and the physical causes which limit their distribu- tion. Their migrations, he believes, are very imperfectly known, but he inclines to the opinion that they only periodically visited such points in their former range as Sable Island and other southerly lying islands. The causes which limit their range he considers to be mainly temperature, since he finds the southern boundary of their distribution is deflected northward and south- ward in accordance with the curves of isothermal lines. The former range of the Walruses is also considered at length, to which subject are devoted nearly twenty-flve pages. A short account is given of their reproduction and food, the paper clos- ing with an inquiry into their systematic relationship to other animals. The map accompanying his memoir shows not only the distribution of the Walruses as at that time known, but indicates also the region over which they are known to have formerly occurred, and also the habitat of the Bhytina, or Sea- cow of Steller. The reception in London, in 1853, of a young living Walrus gave rise to a paper by Owen* on its anatomy and dentition, and another by Gray,t " On the Attitudes and Figures of the Morse." A short paper was contributed by Sundevall J in 1859 on its general history. Leidy, in 1860, published an important paper on the fossil remains of Walruses found on the eastern coast of the United States, while Gratiolet, Defrance, Lankester, and Van Beneden have also written about those that have been met with in France, England, and Belgium.§ Malmgren, in 1864, in a paper on the Mammalian Fauna of •Proc. Zoijl. Soc. Lond., 1853, pp. 103-106. tibicl., pp. 112-116, figs. 1-10. tOm Walrosseu, Ofversigt K. Vet. Akad. Forb. (Stockh.), xvi, Ib.j'J, pp. 441-447 ; also translated in Zeitschr. gesammt. Natnrw. Halle, xv, 1860, pp. 270-275. ij See anted, pp. 61-65. 92 ODOB^NUS ROSMARUS ATLANTIC WALRUS. Fiumark and Spitzbergen,* jiublisbed many interesting notes relating to its babits and food, and biter a special paper on its dentition (noticed anteu, p. Hi.) Malmgren's observations on tbeii" bal)its, distribution, etc., also appear in tbe history of the Svredisb Expedition to Spitzbergen and Bear Island in the year 1801,t together with a somewhat detailed and very interesting general history of the animal, with several illustrations. Brown, in 1808, in his " J^^otes on the History and Geogi'aph- ical Kelations of thePinnipedia frequenting the Spitzbergen and Greenland Seas,"^ devotes several pages to the Walruses (pi). 427-435), in which he considers especially their habits and food, geographical distribution, and economic value. In addition to the special papers cited in the foregoing pages, their general history has been more or less fully presented in several general works treating of the mammalia, and in several faunal pubhcations, § Much information respecting their general history may also be I'oiind in the narratives of various Arctic €xi)lorer8, as Parry. Wrangell, Keilhau, Kane, Ilayes, Lamont, and others, whose co"tributions will be more fully noticed in the following pages relat i ; to the habits of the Walruses. 2. Figures. — As von Baer fiicetiously remarlcs, no animal has had the honor of being depicted in such strange and widely diverse representations as the Walrus. These, as has been previously stated, began with Glaus Magnus, about the middle of the sixteenth century, who opened the series with half a dozen phantastic flgui'es, based apparently upon this animal, only one of which, however, bore the name liosmarus {Bosnia- * lakttagelser och antcckuiugar till Finmarkcns och Spetsbergens Diigg- ecially the anterior. Another, called Vacca marina, represents the head of an ox, with a long beard on the chin. A fourth represents a dolphin- like body, with four feet, a fish's tail, a pair of long, ascending, *My remarks respecting Olaus Magnus's figures are based on Gesnei's (Hist. Animal. Aquat., 1558, pp. 247-249), and Gray's copies of tbem (Proc. Zool. Soc. Loud., 1853, p. 113), Olaus Magnus's work not being accessible to iiM'. The figures herewith given (Figs. 4-12) are from electros of Gray's lignrt's. t Sec Fig. 4, copied by Gray from Ohu.