id	sid	tid	token	lemma	pos
t722h705513	1	1	the	the	DET
t722h705513	1	2	bacterium	bacterium	NOUN
t722h705513	1	3	pseudomonas	pseudomonas	PROPN
t722h705513	1	4	aeruginosa	aeruginosa	PROPN
t722h705513	1	5	is	be	AUX
t722h705513	1	6	ubiquitous	ubiquitous	ADJ
t722h705513	1	7	in	in	ADP
t722h705513	1	8	the	the	DET
t722h705513	1	9	environment	environment	NOUN
t722h705513	1	10	and	and	CCONJ
t722h705513	1	11	is	be	AUX
t722h705513	1	12	also	also	ADV
t722h705513	1	13	an	an	DET
t722h705513	1	14	opportunistic	opportunistic	ADJ
t722h705513	1	15	human	human	ADJ
t722h705513	1	16	pathogen	pathogen	PROPN
t722h705513	1	17	.	.	PUNCT
t722h705513	2	1	a	a	DET
t722h705513	2	2	hallmark	hallmark	ADJ
t722h705513	2	3	characteristic	characteristic	NOUN
t722h705513	2	4	of	of	ADP
t722h705513	2	5	p.	p.	PROPN
t722h705513	2	6	aeruginosa	aeruginosa	PROPN
t722h705513	2	7	is	be	AUX
t722h705513	2	8	its	its	PRON
t722h705513	2	9	ability	ability	NOUN
t722h705513	2	10	to	to	PART
t722h705513	2	11	readily	readily	ADV
t722h705513	2	12	develop	develop	VERB
t722h705513	2	13	attached	attach	VERB
t722h705513	2	14	-	-	PUNCT
t722h705513	2	15	growth	growth	NOUN
t722h705513	2	16	biofilms	biofilm	NOUN
t722h705513	2	17	on	on	ADP
t722h705513	2	18	many	many	ADJ
t722h705513	2	19	surfaces	surface	NOUN
t722h705513	2	20	under	under	ADP
t722h705513	2	21	various	various	ADJ
t722h705513	2	22	growth	growth	NOUN
t722h705513	2	23	conditions	condition	NOUN
t722h705513	2	24	.	.	PUNCT
t722h705513	3	1	during	during	ADP
t722h705513	3	2	colonization	colonization	NOUN
t722h705513	3	3	of	of	ADP
t722h705513	3	4	a	a	DET
t722h705513	3	5	surface	surface	NOUN
t722h705513	3	6	,	,	PUNCT
t722h705513	3	7	initial	initial	ADJ
t722h705513	3	8	biofilm	biofilm	NOUN
t722h705513	3	9	development	development	NOUN
t722h705513	3	10	for	for	ADP
t722h705513	3	11	p.	p.	PROPN
t722h705513	3	12	aeruginosa	aeruginosa	PROPN
t722h705513	3	13	is	be	AUX
t722h705513	3	14	aided	aid	VERB
t722h705513	3	15	by	by	ADP
t722h705513	3	16	surface	surface	NOUN
t722h705513	3	17	motility	motility	NOUN
t722h705513	3	18	mediated	mediate	VERB
t722h705513	3	19	by	by	ADP
t722h705513	3	20	both	both	DET
t722h705513	3	21	type	type	NOUN
t722h705513	3	22	iv	iv	PROPN
t722h705513	3	23	pili	pili	NOUN
t722h705513	3	24	(	(	PUNCT
t722h705513	3	25	tfp	tfp	PROPN
t722h705513	3	26	)	)	PUNCT
t722h705513	3	27	and	and	CCONJ
t722h705513	3	28	flagella	flagella	NOUN
t722h705513	3	29	.	.	PUNCT
t722h705513	4	1	understanding	understand	VERB
t722h705513	4	2	the	the	DET
t722h705513	4	3	environmental	environmental	ADJ
t722h705513	4	4	cues	cue	NOUN
t722h705513	4	5	that	that	PRON
t722h705513	4	6	influence	influence	NOUN
t722h705513	4	7	motility	motility	NOUN
t722h705513	4	8	could	could	AUX
t722h705513	4	9	provide	provide	VERB
t722h705513	4	10	insight	insight	NOUN
t722h705513	4	11	into	into	ADP
t722h705513	4	12	biofilm	biofilm	NOUN
t722h705513	4	13	formation	formation	NOUN
t722h705513	4	14	.	.	PUNCT
t722h705513	5	1	this	this	DET
t722h705513	5	2	thesis	thesis	NOUN
t722h705513	5	3	examines	examine	VERB
t722h705513	5	4	how	how	SCONJ
t722h705513	5	5	cations	cation	NOUN
t722h705513	5	6	act	act	VERB
t722h705513	5	7	as	as	SCONJ
t722h705513	5	8	environmental	environmental	ADJ
t722h705513	5	9	signals	signal	NOUN
t722h705513	5	10	to	to	PART
t722h705513	5	11	affect	affect	VERB
t722h705513	5	12	p.	p.	NOUN
t722h705513	5	13	aeruginosa	aeruginosa	PROPN
t722h705513	5	14	bacterial	bacterial	PROPN
t722h705513	5	15	tfp	tfp	PROPN
t722h705513	5	16	motility	motility	NOUN
t722h705513	5	17	in	in	ADP
t722h705513	5	18	a	a	DET
t722h705513	5	19	concentration	concentration	NOUN
t722h705513	5	20	dependent	dependent	ADJ
t722h705513	5	21	manner	manner	NOUN
t722h705513	5	22	.	.	PUNCT
t722h705513	6	1	low	low	ADJ
t722h705513	6	2	levels	level	NOUN
t722h705513	6	3	of	of	ADP
t722h705513	6	4	calcium	calcium	NOUN
t722h705513	6	5	,	,	PUNCT
t722h705513	6	6	magnesium	magnesium	NOUN
t722h705513	6	7	,	,	PUNCT
t722h705513	6	8	iron	iron	NOUN
t722h705513	6	9	,	,	PUNCT
t722h705513	6	10	potassium	potassium	NOUN
t722h705513	6	11	,	,	PUNCT
t722h705513	6	12	or	or	CCONJ
t722h705513	6	13	sodium	sodium	NOUN
t722h705513	6	14	stimulated	stimulate	VERB
t722h705513	6	15	an	an	DET
t722h705513	6	16	increase	increase	NOUN
t722h705513	6	17	in	in	ADP
t722h705513	6	18	p.	p.	PROPN
t722h705513	6	19	aeruginosa	aeruginosa	PROPN
t722h705513	6	20	tfp	tfp	PROPN
t722h705513	6	21	surface	surface	PROPN
t722h705513	6	22	motility	motility	NOUN
t722h705513	6	23	at	at	ADP
t722h705513	6	24	the	the	DET
t722h705513	6	25	single	single	ADJ
t722h705513	6	26	cell	cell	NOUN
t722h705513	6	27	level	level	NOUN
t722h705513	6	28	.	.	PUNCT
t722h705513	7	1	more	more	ADV
t722h705513	7	2	specifically	specifically	ADV
t722h705513	7	3	,	,	PUNCT
t722h705513	7	4	there	there	PRON
t722h705513	7	5	was	be	VERB
t722h705513	7	6	an	an	DET
t722h705513	7	7	increase	increase	NOUN
t722h705513	7	8	in	in	ADP
t722h705513	7	9	the	the	DET
t722h705513	7	10	tfp	tfp	PROPN
t722h705513	7	11	motility	motility	NOUN
t722h705513	7	12	mode	mode	NOUN
t722h705513	7	13	called	call	VERB
t722h705513	7	14	crawling	crawling	NOUN
t722h705513	7	15	,	,	PUNCT
t722h705513	7	16	mean	mean	ADJ
t722h705513	7	17	square	square	ADJ
t722h705513	7	18	displacement	displacement	NOUN
t722h705513	7	19	,	,	PUNCT
t722h705513	7	20	and	and	CCONJ
t722h705513	7	21	mean	mean	VERB
t722h705513	7	22	velocity	velocity	NOUN
t722h705513	7	23	.	.	PUNCT
t722h705513	8	1	this	this	DET
t722h705513	8	2	tfp	tfp	ADJ
t722h705513	8	3	-	-	PUNCT
t722h705513	8	4	specific	specific	ADJ
t722h705513	8	5	behavior	behavior	NOUN
t722h705513	8	6	is	be	AUX
t722h705513	8	7	further	far	ADV
t722h705513	8	8	exaggerated	exaggerated	ADJ
t722h705513	8	9	in	in	ADP
t722h705513	8	10	the	the	DET
t722h705513	8	11	&	&	CCONJ
t722h705513	8	12	#	#	SYM
t722h705513	8	13	916;flim	916;flim	NUM
t722h705513	8	14	mutant	mutant	NOUN
t722h705513	8	15	,	,	PUNCT
t722h705513	8	16	as	as	SCONJ
t722h705513	8	17	this	this	DET
t722h705513	8	18	flagellar	flagellar	ADJ
t722h705513	8	19	mutant	mutant	NOUN
t722h705513	8	20	is	be	AUX
t722h705513	8	21	strictly	strictly	ADV
t722h705513	8	22	tfp	tfp	ADJ
t722h705513	8	23	-	-	PUNCT
t722h705513	8	24	dependent	dependent	ADJ
t722h705513	8	25	.	.	PUNCT
t722h705513	9	1	calcium	calcium	NOUN
t722h705513	9	2	starvation	starvation	NOUN
t722h705513	9	3	was	be	AUX
t722h705513	9	4	found	find	VERB
t722h705513	9	5	to	to	PART
t722h705513	9	6	affect	affect	VERB
t722h705513	9	7	the	the	DET
t722h705513	9	8	resultant	resultant	ADJ
t722h705513	9	9	biofilm	biofilm	NOUN
t722h705513	9	10	phenotype	phenotype	NOUN
t722h705513	9	11	as	as	ADP
t722h705513	9	12	homogeneous	homogeneous	ADJ
t722h705513	9	13	flat	flat	ADJ
t722h705513	9	14	biofilms	biofilm	NOUN
t722h705513	9	15	formed	form	VERB
t722h705513	9	16	under	under	ADP
t722h705513	9	17	calcium	calcium	NOUN
t722h705513	9	18	starvation	starvation	NOUN
t722h705513	9	19	as	as	SCONJ
t722h705513	9	20	compared	compare	VERB
t722h705513	9	21	to	to	ADP
t722h705513	9	22	the	the	DET
t722h705513	9	23	structured	structure	VERB
t722h705513	9	24	architecture	architecture	NOUN
t722h705513	9	25	of	of	ADP
t722h705513	9	26	biofilms	biofilm	NOUN
t722h705513	9	27	that	that	PRON
t722h705513	9	28	formed	form	VERB
t722h705513	9	29	under	under	ADP
t722h705513	9	30	ample	ample	ADJ
t722h705513	9	31	calcium	calcium	NOUN
t722h705513	9	32	conditions	condition	NOUN
t722h705513	9	33	.	.	PUNCT
t722h705513	10	1	the	the	DET
t722h705513	10	2	influence	influence	NOUN
t722h705513	10	3	of	of	ADP
t722h705513	10	4	these	these	DET
t722h705513	10	5	cations	cation	NOUN
t722h705513	10	6	upon	upon	SCONJ
t722h705513	10	7	tfp	tfp	PROPN
t722h705513	10	8	motility	motility	NOUN
t722h705513	10	9	at	at	ADP
t722h705513	10	10	the	the	DET
t722h705513	10	11	community	community	NOUN
t722h705513	10	12	level	level	NOUN
t722h705513	10	13	as	as	SCONJ
t722h705513	10	14	determined	determine	VERB
t722h705513	10	15	using	use	VERB
t722h705513	10	16	standard	standard	ADJ
t722h705513	10	17	twitch	twitch	NOUN
t722h705513	10	18	and	and	CCONJ
t722h705513	10	19	chemotactic	chemotactic	ADJ
t722h705513	10	20	twitch	twitch	NOUN
t722h705513	10	21	assays	assay	NOUN
t722h705513	10	22	was	be	AUX
t722h705513	10	23	unclear	unclear	ADJ
t722h705513	10	24	,	,	PUNCT
t722h705513	10	25	suggesting	suggest	VERB
t722h705513	10	26	that	that	SCONJ
t722h705513	10	27	the	the	DET
t722h705513	10	28	effect	effect	NOUN
t722h705513	10	29	of	of	ADP
t722h705513	10	30	cations	cation	NOUN
t722h705513	10	31	on	on	ADP
t722h705513	10	32	tfp	tfp	PROPN
t722h705513	10	33	motility	motility	NOUN
t722h705513	10	34	are	be	AUX
t722h705513	10	35	complicated	complicated	ADJ
t722h705513	10	36	and	and	CCONJ
t722h705513	10	37	dependent	dependent	ADJ
t722h705513	10	38	upon	upon	SCONJ
t722h705513	10	39	multiple	multiple	ADJ
t722h705513	10	40	factors	factor	NOUN
t722h705513	10	41	.	.	PUNCT
t722h705513	11	1	the	the	DET
t722h705513	11	2	affect	affect	NOUN
t722h705513	11	3	of	of	ADP
t722h705513	11	4	calcium	calcium	NOUN
t722h705513	11	5	starvation	starvation	NOUN
t722h705513	11	6	on	on	ADP
t722h705513	11	7	tfp	tfp	PROPN
t722h705513	11	8	was	be	AUX
t722h705513	11	9	further	far	ADV
t722h705513	11	10	characterized	characterize	VERB
t722h705513	11	11	since	since	SCONJ
t722h705513	11	12	calcium	calcium	NOUN
t722h705513	11	13	has	have	VERB
t722h705513	11	14	direct	direct	ADJ
t722h705513	11	15	links	link	NOUN
t722h705513	11	16	to	to	ADP
t722h705513	11	17	virulence	virulence	NOUN
t722h705513	11	18	and	and	CCONJ
t722h705513	11	19	tfp	tfp	PROPN
t722h705513	11	20	.	.	PUNCT
t722h705513	12	1	several	several	ADJ
t722h705513	12	2	types	type	NOUN
t722h705513	12	3	of	of	ADP
t722h705513	12	4	experimental	experimental	ADJ
t722h705513	12	5	approaches	approach	NOUN
t722h705513	12	6	were	be	AUX
t722h705513	12	7	utilized	utilize	VERB
t722h705513	12	8	in	in	ADP
t722h705513	12	9	an	an	DET
t722h705513	12	10	effort	effort	NOUN
t722h705513	12	11	to	to	PART
t722h705513	12	12	pinpoint	pinpoint	VERB
t722h705513	12	13	a	a	DET
t722h705513	12	14	specific	specific	ADJ
t722h705513	12	15	gene	gene	NOUN
t722h705513	12	16	and/or	and/or	CCONJ
t722h705513	12	17	regulatory	regulatory	ADJ
t722h705513	12	18	mechanism	mechanism	NOUN
t722h705513	12	19	that	that	PRON
t722h705513	12	20	would	would	AUX
t722h705513	12	21	link	link	VERB
t722h705513	12	22	calcium	calcium	NOUN
t722h705513	12	23	response	response	NOUN
t722h705513	12	24	with	with	ADP
t722h705513	12	25	tfp	tfp	PROPN
t722h705513	12	26	action	action	NOUN
t722h705513	12	27	.	.	PUNCT
t722h705513	13	1	there	there	PRON
t722h705513	13	2	is	be	VERB
t722h705513	13	3	a	a	DET
t722h705513	13	4	clear	clear	ADJ
t722h705513	13	5	population	population	NOUN
t722h705513	13	6	effect	effect	NOUN
t722h705513	13	7	under	under	ADP
t722h705513	13	8	calcium	calcium	NOUN
t722h705513	13	9	-	-	PUNCT
t722h705513	13	10	starved	starve	VERB
t722h705513	13	11	conditions	condition	NOUN
t722h705513	13	12	as	as	SCONJ
t722h705513	13	13	less	less	ADJ
t722h705513	13	14	tfp	tfp	ADJ
t722h705513	13	15	-	-	PUNCT
t722h705513	13	16	dependent	dependent	ADJ
t722h705513	13	17	motility	motility	NOUN
t722h705513	13	18	is	be	AUX
t722h705513	13	19	observed	observe	VERB
t722h705513	13	20	at	at	ADP
t722h705513	13	21	high	high	ADJ
t722h705513	13	22	cell	cell	NOUN
t722h705513	13	23	density	density	NOUN
t722h705513	13	24	.	.	PUNCT
t722h705513	14	1	swarming	swarm	VERB
t722h705513	14	2	motility	motility	NOUN
t722h705513	14	3	,	,	PUNCT
t722h705513	14	4	which	which	PRON
t722h705513	14	5	is	be	AUX
t722h705513	14	6	a	a	DET
t722h705513	14	7	flagellar	flagellar	ADJ
t722h705513	14	8	motility	motility	NOUN
t722h705513	14	9	influenced	influence	VERB
t722h705513	14	10	by	by	ADP
t722h705513	14	11	tfp	tfp	PROPN
t722h705513	14	12	,	,	PUNCT
t722h705513	14	13	is	be	AUX
t722h705513	14	14	reduced	reduce	VERB
t722h705513	14	15	under	under	ADP
t722h705513	14	16	calcium	calcium	NOUN
t722h705513	14	17	-	-	PUNCT
t722h705513	14	18	starved	starve	VERB
t722h705513	14	19	conditions	condition	NOUN
t722h705513	14	20	.	.	PUNCT
t722h705513	15	1	this	this	DET
t722h705513	15	2	calcium	calcium	NOUN
t722h705513	15	3	effect	effect	NOUN
t722h705513	15	4	was	be	AUX
t722h705513	15	5	determined	determine	VERB
t722h705513	15	6	not	not	PART
t722h705513	15	7	to	to	PART
t722h705513	15	8	be	be	AUX
t722h705513	15	9	related	relate	VERB
t722h705513	15	10	to	to	ADP
t722h705513	15	11	acyl	acyl	NOUN
t722h705513	15	12	homoserine	homoserine	ADJ
t722h705513	15	13	lactone	lactone	NOUN
t722h705513	15	14	quorum	quorum	PROPN
t722h705513	15	15	sensing	sense	VERB
t722h705513	15	16	,	,	PUNCT
t722h705513	15	17	rhamnolipid	rhamnolipid	NOUN
t722h705513	15	18	production	production	NOUN
t722h705513	15	19	,	,	PUNCT
t722h705513	15	20	or	or	CCONJ
t722h705513	15	21	the	the	DET
t722h705513	15	22	tfp	tfp	PROPN
t722h705513	15	23	retraction	retraction	NOUN
t722h705513	15	24	gene	gene	NOUN
t722h705513	15	25	pilu	pilu	PROPN
t722h705513	15	26	.	.	PUNCT
t722h705513	16	1	in	in	ADP
t722h705513	16	2	separate	separate	ADJ
t722h705513	16	3	experiments	experiment	NOUN
t722h705513	16	4	,	,	PUNCT
t722h705513	16	5	methods	method	NOUN
t722h705513	16	6	were	be	AUX
t722h705513	16	7	developed	develop	VERB
t722h705513	16	8	to	to	PART
t722h705513	16	9	allow	allow	VERB
t722h705513	16	10	chemical	chemical	ADJ
t722h705513	16	11	analysis	analysis	NOUN
t722h705513	16	12	of	of	ADP
t722h705513	16	13	p.	p.	PROPN
t722h705513	16	14	aeruginosa	aeruginosa	PROPN
t722h705513	16	15	biofilms	biofilms	PROPN
t722h705513	16	16	.	.	PUNCT
t722h705513	17	1	these	these	DET
t722h705513	17	2	methods	method	NOUN
t722h705513	17	3	allowed	allow	VERB
t722h705513	17	4	for	for	ADP
t722h705513	17	5	a	a	DET
t722h705513	17	6	combination	combination	NOUN
t722h705513	17	7	of	of	ADP
t722h705513	17	8	confocal	confocal	ADJ
t722h705513	17	9	raman	raman	NOUN
t722h705513	17	10	microspectroscopy	microspectroscopy	NOUN
t722h705513	17	11	(	(	PUNCT
t722h705513	17	12	crm	crm	PROPN
t722h705513	17	13	)	)	PUNCT
t722h705513	17	14	and	and	CCONJ
t722h705513	17	15	matrix	matrix	NOUN
t722h705513	17	16	-	-	PUNCT
t722h705513	17	17	assisted	assist	VERB
t722h705513	17	18	laser	laser	NOUN
t722h705513	17	19	desorption	desorption	NOUN
t722h705513	17	20	ionization	ionization	NOUN
t722h705513	17	21	mass	mass	NOUN
t722h705513	17	22	spectrometry	spectrometry	NOUN
t722h705513	17	23	(	(	PUNCT
t722h705513	17	24	maldi	maldi	PROPN
t722h705513	17	25	ms	ms	PROPN
t722h705513	17	26	)	)	PUNCT
t722h705513	17	27	to	to	PART
t722h705513	17	28	analyze	analyze	VERB
t722h705513	17	29	biofilms	biofilm	NOUN
t722h705513	17	30	.	.	PUNCT
t722h705513	18	1	a	a	DET
t722h705513	18	2	method	method	NOUN
t722h705513	18	3	was	be	AUX
t722h705513	18	4	developed	develop	VERB
t722h705513	18	5	to	to	PART
t722h705513	18	6	enable	enable	VERB
t722h705513	18	7	visualization	visualization	NOUN
t722h705513	18	8	of	of	ADP
t722h705513	18	9	individual	individual	ADJ
t722h705513	18	10	twitching	twitch	VERB
t722h705513	18	11	cells	cell	NOUN
t722h705513	18	12	with	with	ADP
t722h705513	18	13	confocal	confocal	ADJ
t722h705513	18	14	fluorescence	fluorescence	ADJ
t722h705513	18	15	microscopy	microscopy	NOUN
t722h705513	18	16	using	use	VERB
t722h705513	18	17	a	a	DET
t722h705513	18	18	plate	plate	NOUN
t722h705513	18	19	complex	complex	NOUN
t722h705513	18	20	that	that	PRON
t722h705513	18	21	has	have	AUX
t722h705513	18	22	been	be	AUX
t722h705513	18	23	used	use	VERB
t722h705513	18	24	to	to	PART
t722h705513	18	25	observe	observe	VERB
t722h705513	18	26	tfp	tfp	ADJ
t722h705513	18	27	-	-	PUNCT
t722h705513	18	28	dependent	dependent	ADJ
t722h705513	18	29	social	social	ADJ
t722h705513	18	30	motility	motility	NOUN
t722h705513	18	31	patterns	pattern	NOUN
t722h705513	18	32	of	of	ADP
t722h705513	18	33	myxococcus	myxococcus	PROPN
t722h705513	18	34	xanthus	xanthus	PROPN
t722h705513	18	35	.	.	PUNCT
t722h705513	19	1	this	this	DET
t722h705513	19	2	new	new	ADJ
t722h705513	19	3	approach	approach	NOUN
t722h705513	19	4	enables	enable	VERB
t722h705513	19	5	researchers	researcher	NOUN
t722h705513	19	6	to	to	PART
t722h705513	19	7	better	well	ADV
t722h705513	19	8	characterize	characterize	VERB
t722h705513	19	9	twitching	twitching	NOUN
t722h705513	19	10	and	and	CCONJ
t722h705513	19	11	its	its	PRON
t722h705513	19	12	role	role	NOUN
t722h705513	19	13	in	in	ADP
t722h705513	19	14	biofilm	biofilm	NOUN
t722h705513	19	15	formation	formation	NOUN
t722h705513	19	16	.	.	PUNCT