id	sid	tid	token	lemma	pos
mk61rf5885n	1	1	the	the	DET
mk61rf5885n	1	2	overwintering	overwinter	VERB
mk61rf5885n	1	3	physiology	physiology	NOUN
mk61rf5885n	1	4	of	of	ADP
mk61rf5885n	1	5	aquatic	aquatic	ADJ
mk61rf5885n	1	6	insects	insect	NOUN
mk61rf5885n	1	7	has	have	AUX
mk61rf5885n	1	8	not	not	PART
mk61rf5885n	1	9	been	be	AUX
mk61rf5885n	1	10	well	well	ADV
mk61rf5885n	1	11	characterized	characterize	VERB
mk61rf5885n	1	12	.	.	PUNCT
mk61rf5885n	2	1	the	the	DET
mk61rf5885n	2	2	stonefly	stonefly	NOUN
mk61rf5885n	2	3	,	,	PUNCT
mk61rf5885n	2	4	nemoura	nemoura	PROPN
mk61rf5885n	2	5	arctica	arctica	PROPN
mk61rf5885n	2	6	,	,	PUNCT
mk61rf5885n	2	7	was	be	AUX
mk61rf5885n	2	8	collected	collect	VERB
mk61rf5885n	2	9	from	from	ADP
mk61rf5885n	2	10	the	the	DET
mk61rf5885n	2	11	headwaters	headwater	NOUN
mk61rf5885n	2	12	of	of	ADP
mk61rf5885n	2	13	the	the	DET
mk61rf5885n	2	14	chandalar	chandalar	PROPN
mk61rf5885n	2	15	river	river	NOUN
mk61rf5885n	2	16	,	,	PUNCT
mk61rf5885n	2	17	where	where	SCONJ
mk61rf5885n	2	18	streambed	streambe	VERB
mk61rf5885n	2	19	temperatures	temperature	NOUN
mk61rf5885n	2	20	measured	measure	VERB
mk61rf5885n	2	21	as	as	ADV
mk61rf5885n	2	22	low	low	ADJ
mk61rf5885n	2	23	as	as	ADP
mk61rf5885n	2	24	-12.7ìü	-12.7ìü	NOUN
mk61rf5885n	2	25	ác	ác	NOUN
mk61rf5885n	2	26	in	in	ADP
mk61rf5885n	2	27	midwinter	midwinter	NOUN
mk61rf5885n	2	28	.	.	PUNCT
mk61rf5885n	3	1	when	when	SCONJ
mk61rf5885n	3	2	in	in	ADP
mk61rf5885n	3	3	contact	contact	NOUN
mk61rf5885n	3	4	with	with	ADP
mk61rf5885n	3	5	ice	ice	NOUN
mk61rf5885n	3	6	,	,	PUNCT
mk61rf5885n	3	7	n.	n.	PROPN
mk61rf5885n	3	8	arctica	arctica	PROPN
mk61rf5885n	3	9	froze	freeze	VERB
mk61rf5885n	3	10	at	at	ADP
mk61rf5885n	3	11	-1.5ìâå±0.4ìü	-1.5ìâå±0.4ìü	PROPN
mk61rf5885n	3	12	ác	ác	PROPN
mk61rf5885n	3	13	,	,	PUNCT
mk61rf5885n	3	14	but	but	CCONJ
mk61rf5885n	3	15	individuals	individual	NOUN
mk61rf5885n	3	16	survived	survive	VERB
mk61rf5885n	3	17	temperatures	temperature	NOUN
mk61rf5885n	3	18	down	down	ADV
mk61rf5885n	3	19	to	to	ADP
mk61rf5885n	3	20	-15ìü	-15ìü	VERB
mk61rf5885n	3	21	ác	ác	NOUN
mk61rf5885n	3	22	,	,	PUNCT
mk61rf5885n	3	23	making	make	VERB
mk61rf5885n	3	24	this	this	PRON
mk61rf5885n	3	25	the	the	DET
mk61rf5885n	3	26	first	first	ADJ
mk61rf5885n	3	27	described	describe	VERB
mk61rf5885n	3	28	species	specie	NOUN
mk61rf5885n	3	29	of	of	ADP
mk61rf5885n	3	30	freeze	freeze	VERB
mk61rf5885n	3	31	tolerant	tolerant	ADJ
mk61rf5885n	3	32	stonefly	stonefly	NOUN
mk61rf5885n	3	33	.	.	PUNCT
mk61rf5885n	4	1	nemoura	nemoura	PROPN
mk61rf5885n	4	2	arctica	arctica	PROPN
mk61rf5885n	4	3	produce	produce	VERB
mk61rf5885n	4	4	both	both	CCONJ
mk61rf5885n	4	5	glycerol	glycerol	NOUN
mk61rf5885n	4	6	and	and	CCONJ
mk61rf5885n	4	7	ice	ice	NOUN
mk61rf5885n	4	8	thermal	thermal	ADJ
mk61rf5885n	4	9	hysteresis	hysteresis	NOUN
mk61rf5885n	4	10	factors	factor	NOUN
mk61rf5885n	4	11	in	in	ADP
mk61rf5885n	4	12	response	response	NOUN
mk61rf5885n	4	13	to	to	ADP
mk61rf5885n	4	14	low	low	ADJ
mk61rf5885n	4	15	temperature	temperature	NOUN
mk61rf5885n	4	16	.	.	PUNCT
mk61rf5885n	5	1	hemolymph	hemolymph	NOUN
mk61rf5885n	5	2	glycerol	glycerol	NOUN
mk61rf5885n	5	3	concentrations	concentration	NOUN
mk61rf5885n	5	4	increased	increase	VERB
mk61rf5885n	5	5	from	from	ADP
mk61rf5885n	5	6	3	3	NUM
mk61rf5885n	5	7	mm	mm	NOUN
mk61rf5885n	5	8	to	to	ADP
mk61rf5885n	5	9	930ìâå±114	930ìâå±114	NUM
mk61rf5885n	5	10	mm	mm	NOUN
mk61rf5885n	5	11	in	in	ADP
mk61rf5885n	5	12	response	response	NOUN
mk61rf5885n	5	13	to	to	ADP
mk61rf5885n	5	14	acclimation	acclimation	NOUN
mk61rf5885n	5	15	at	at	ADP
mk61rf5885n	5	16	-8ìü	-8ìü	NOUN
mk61rf5885n	5	17	ác	ác	NOUN
mk61rf5885n	5	18	and	and	CCONJ
mk61rf5885n	5	19	glycerol	glycerol	NOUN
mk61rf5885n	5	20	concentrations	concentration	NOUN
mk61rf5885n	5	21	increased	increase	VERB
mk61rf5885n	5	22	even	even	ADV
mk61rf5885n	5	23	after	after	SCONJ
mk61rf5885n	5	24	nymphs	nymph	NOUN
mk61rf5885n	5	25	were	be	AUX
mk61rf5885n	5	26	frozen	freeze	VERB
mk61rf5885n	5	27	.	.	PUNCT
mk61rf5885n	6	1	although	although	SCONJ
mk61rf5885n	6	2	,	,	PUNCT
mk61rf5885n	6	3	the	the	DET
mk61rf5885n	6	4	hemolymph	hemolymph	NOUN
mk61rf5885n	6	5	of	of	ADP
mk61rf5885n	6	6	individual	individual	ADJ
mk61rf5885n	6	7	cold	cold	ADJ
mk61rf5885n	6	8	acclimated	acclimate	VERB
mk61rf5885n	6	9	nymphs	nymph	NOUN
mk61rf5885n	6	10	occasionally	occasionally	ADV
mk61rf5885n	6	11	exhibited	exhibit	VERB
mk61rf5885n	6	12	more	more	ADJ
mk61rf5885n	6	13	than	than	ADP
mk61rf5885n	6	14	a	a	DET
mk61rf5885n	6	15	degree	degree	NOUN
mk61rf5885n	6	16	of	of	ADP
mk61rf5885n	6	17	thermal	thermal	ADJ
mk61rf5885n	6	18	hysteresis	hysteresis	NOUN
mk61rf5885n	6	19	,	,	PUNCT
mk61rf5885n	6	20	typically	typically	ADV
mk61rf5885n	6	21	the	the	DET
mk61rf5885n	6	22	hemolymph	hemolymph	NOUN
mk61rf5885n	6	23	exhibited	exhibit	VERB
mk61rf5885n	6	24	only	only	ADJ
mk61rf5885n	6	25	hexagonal	hexagonal	ADJ
mk61rf5885n	6	26	crystal	crystal	NOUN
mk61rf5885n	6	27	growth	growth	NOUN
mk61rf5885n	6	28	and	and	CCONJ
mk61rf5885n	6	29	recrystallization	recrystallization	NOUN
mk61rf5885n	6	30	inhibition	inhibition	NOUN
mk61rf5885n	6	31	.	.	PUNCT
mk61rf5885n	7	1	the	the	DET
mk61rf5885n	7	2	freeze	freeze	ADJ
mk61rf5885n	7	3	tolerant	tolerant	ADJ
mk61rf5885n	7	4	beetle	beetle	NOUN
mk61rf5885n	7	5	upis	upis	PROPN
mk61rf5885n	7	6	ceramboides	ceramboide	NOUN
mk61rf5885n	7	7	is	be	AUX
mk61rf5885n	7	8	the	the	DET
mk61rf5885n	7	9	only	only	ADJ
mk61rf5885n	7	10	animal	animal	NOUN
mk61rf5885n	7	11	species	specie	NOUN
mk61rf5885n	7	12	known	know	VERB
mk61rf5885n	7	13	to	to	PART
mk61rf5885n	7	14	accumulate	accumulate	VERB
mk61rf5885n	7	15	high	high	ADJ
mk61rf5885n	7	16	concentrations	concentration	NOUN
mk61rf5885n	7	17	of	of	ADP
mk61rf5885n	7	18	threitol	threitol	PROPN
mk61rf5885n	7	19	(	(	PUNCT
mk61rf5885n	7	20	250	250	NUM
mk61rf5885n	7	21	mm	mm	NOUN
mk61rf5885n	7	22	)	)	PUNCT
mk61rf5885n	7	23	as	as	ADP
mk61rf5885n	7	24	a	a	DET
mk61rf5885n	7	25	cryoprotectant	cryoprotectant	NOUN
mk61rf5885n	7	26	;	;	PUNCT
mk61rf5885n	7	27	however	however	ADV
mk61rf5885n	7	28	the	the	DET
mk61rf5885n	7	29	biosynthetic	biosynthetic	ADJ
mk61rf5885n	7	30	pathway	pathway	NOUN
mk61rf5885n	7	31	has	have	AUX
mk61rf5885n	7	32	not	not	PART
mk61rf5885n	7	33	been	be	AUX
mk61rf5885n	7	34	investigated	investigate	VERB
mk61rf5885n	7	35	.	.	PUNCT
mk61rf5885n	8	1	in	in	ADP
mk61rf5885n	8	2	vivo	vivo	ADJ
mk61rf5885n	8	3	metabolism	metabolism	NOUN
mk61rf5885n	8	4	of	of	ADP
mk61rf5885n	8	5	13c	13c	NUM
mk61rf5885n	8	6	-	-	PUNCT
mk61rf5885n	8	7	labeled	label	VERB
mk61rf5885n	8	8	glucose	glucose	NOUN
mk61rf5885n	8	9	isotopomers	isotopomer	NOUN
mk61rf5885n	8	10	was	be	AUX
mk61rf5885n	8	11	monitored	monitor	VERB
mk61rf5885n	8	12	by	by	ADP
mk61rf5885n	8	13	13c{1h	13c{1h	NUM
mk61rf5885n	8	14	}	}	PUNCT
mk61rf5885n	8	15	nmr	nmr	NOUN
mk61rf5885n	8	16	spectroscopy	spectroscopy	NOUN
mk61rf5885n	8	17	,	,	PUNCT
mk61rf5885n	8	18	demonstrating	demonstrate	VERB
mk61rf5885n	8	19	that	that	SCONJ
mk61rf5885n	8	20	c3	c3	PROPN
mk61rf5885n	8	21	-	-	PUNCT
mk61rf5885n	8	22	c6	c6	PROPN
mk61rf5885n	8	23	of	of	ADP
mk61rf5885n	8	24	glucose	glucose	NOUN
mk61rf5885n	8	25	become	become	VERB
mk61rf5885n	8	26	c1	c1	PROPN
mk61rf5885n	8	27	-	-	PUNCT
mk61rf5885n	8	28	c4	c4	NOUN
mk61rf5885n	8	29	of	of	ADP
mk61rf5885n	8	30	threitol	threitol	PROPN
mk61rf5885n	8	31	,	,	PUNCT
mk61rf5885n	8	32	which	which	PRON
mk61rf5885n	8	33	is	be	AUX
mk61rf5885n	8	34	expected	expect	VERB
mk61rf5885n	8	35	for	for	ADP
mk61rf5885n	8	36	four	four	NUM
mk61rf5885n	8	37	-	-	PUNCT
mk61rf5885n	8	38	carbon	carbon	NOUN
mk61rf5885n	8	39	saccharides	saccharide	NOUN
mk61rf5885n	8	40	arising	arise	VERB
mk61rf5885n	8	41	from	from	ADP
mk61rf5885n	8	42	the	the	DET
mk61rf5885n	8	43	pentose	pentose	NOUN
mk61rf5885n	8	44	phosphate	phosphate	NOUN
mk61rf5885n	8	45	pathway	pathway	NOUN
mk61rf5885n	8	46	(	(	PUNCT
mk61rf5885n	8	47	ppp	ppp	NOUN
mk61rf5885n	8	48	)	)	PUNCT
mk61rf5885n	8	49	.	.	PUNCT
mk61rf5885n	9	1	in	in	ADP
mk61rf5885n	9	2	vitro	vitro	X
mk61rf5885n	9	3	experiments	experiment	NOUN
mk61rf5885n	9	4	show	show	VERB
mk61rf5885n	9	5	that	that	SCONJ
mk61rf5885n	9	6	threitol	threitol	PROPN
mk61rf5885n	9	7	is	be	AUX
mk61rf5885n	9	8	synthesized	synthesize	VERB
mk61rf5885n	9	9	from	from	ADP
mk61rf5885n	9	10	erythrose	erythrose	NOUN
mk61rf5885n	9	11	4	4	NUM
mk61rf5885n	9	12	-	-	PUNCT
mk61rf5885n	9	13	phosphate	phosphate	NOUN
mk61rf5885n	9	14	(	(	PUNCT
mk61rf5885n	9	15	e4p	e4p	PROPN
mk61rf5885n	9	16	)	)	PUNCT
mk61rf5885n	9	17	,	,	PUNCT
mk61rf5885n	9	18	a	a	DET
mk61rf5885n	9	19	c4	c4	NOUN
mk61rf5885n	9	20	intermediate	intermediate	ADJ
mk61rf5885n	9	21	in	in	ADP
mk61rf5885n	9	22	the	the	DET
mk61rf5885n	9	23	ppp	ppp	NOUN
mk61rf5885n	9	24	.	.	PUNCT
mk61rf5885n	10	1	threitol	threitol	PROPN
mk61rf5885n	10	2	4	4	NUM
mk61rf5885n	10	3	-	-	PUNCT
mk61rf5885n	10	4	phosphate	phosphate	NOUN
mk61rf5885n	10	5	appears	appear	VERB
mk61rf5885n	10	6	to	to	PART
mk61rf5885n	10	7	be	be	AUX
mk61rf5885n	10	8	the	the	DET
mk61rf5885n	10	9	preferred	preferred	ADJ
mk61rf5885n	10	10	substrate	substrate	NOUN
mk61rf5885n	10	11	of	of	ADP
mk61rf5885n	10	12	the	the	DET
mk61rf5885n	10	13	sugar	sugar	NOUN
mk61rf5885n	10	14	phosphatase(s	phosphatase(s	NOUN
mk61rf5885n	10	15	)	)	PUNCT
mk61rf5885n	10	16	,	,	PUNCT
mk61rf5885n	10	17	promoting	promote	VERB
mk61rf5885n	10	18	threitol	threitol	ADJ
mk61rf5885n	10	19	synthesis	synthesis	NOUN
mk61rf5885n	10	20	over	over	ADP
mk61rf5885n	10	21	that	that	PRON
mk61rf5885n	10	22	of	of	ADP
mk61rf5885n	10	23	erythritol	erythritol	NOUN
mk61rf5885n	10	24	.	.	PUNCT
mk61rf5885n	11	1	in	in	ADP
mk61rf5885n	11	2	addition	addition	NOUN
mk61rf5885n	11	3	,	,	PUNCT
mk61rf5885n	11	4	erythritol	erythritol	PROPN
mk61rf5885n	11	5	is	be	AUX
mk61rf5885n	11	6	catabolized	catabolize	VERB
mk61rf5885n	11	7	under	under	ADP
mk61rf5885n	11	8	conditions	condition	NOUN
mk61rf5885n	11	9	that	that	PRON
mk61rf5885n	11	10	promote	promote	VERB
mk61rf5885n	11	11	threitol	threitol	ADJ
mk61rf5885n	11	12	synthesis	synthesis	NOUN
mk61rf5885n	11	13	,	,	PUNCT
mk61rf5885n	11	14	leading	lead	VERB
mk61rf5885n	11	15	to	to	ADP
mk61rf5885n	11	16	the	the	DET
mk61rf5885n	11	17	accumulation	accumulation	NOUN
mk61rf5885n	11	18	of	of	ADP
mk61rf5885n	11	19	high	high	ADJ
mk61rf5885n	11	20	concentrations	concentration	NOUN
mk61rf5885n	11	21	of	of	ADP
mk61rf5885n	11	22	threitol	threitol	PROPN
mk61rf5885n	11	23	without	without	ADP
mk61rf5885n	11	24	concomitant	concomitant	ADJ
mk61rf5885n	11	25	accumulation	accumulation	NOUN
mk61rf5885n	11	26	of	of	ADP
mk61rf5885n	11	27	erythritol	erythritol	PROPN
mk61rf5885n	11	28	.	.	PUNCT
mk61rf5885n	12	1	upis	upis	PROPN
mk61rf5885n	12	2	ceramboides	ceramboides	PROPN
mk61rf5885n	12	3	also	also	ADV
mk61rf5885n	12	4	produces	produce	VERB
mk61rf5885n	12	5	a	a	DET
mk61rf5885n	12	6	novel	novel	NOUN
mk61rf5885n	12	7	,	,	PUNCT
mk61rf5885n	12	8	highly	highly	ADV
mk61rf5885n	12	9	active	active	ADJ
mk61rf5885n	12	10	antifreeze	antifreeze	PROPN
mk61rf5885n	12	11	glycolipid	glycolipid	PROPN
mk61rf5885n	12	12	.	.	PUNCT
mk61rf5885n	13	1	this	this	DET
mk61rf5885n	13	2	glycolipid	glycolipid	NOUN
mk61rf5885n	13	3	exhibits	exhibit	VERB
mk61rf5885n	13	4	3.7ìâå±0.3ìü	3.7ìâå±0.3ìü	NUM
mk61rf5885n	13	5	ác	ác	NOUN
mk61rf5885n	13	6	of	of	ADP
mk61rf5885n	13	7	thermal	thermal	ADJ
mk61rf5885n	13	8	hysteresis	hysteresis	NOUN
mk61rf5885n	13	9	at	at	ADP
mk61rf5885n	13	10	5	5	NUM
mk61rf5885n	13	11	mg	mg	NOUN
mk61rf5885n	13	12	/	/	SYM
mk61rf5885n	13	13	ml	ml	NOUN
mk61rf5885n	13	14	,	,	PUNCT
mk61rf5885n	13	15	whereas	whereas	SCONJ
mk61rf5885n	13	16	fish	fish	VERB
mk61rf5885n	13	17	af(g)ps	af(g)ps	NOUN
mk61rf5885n	13	18	exhibit	exhibit	NOUN
mk61rf5885n	13	19	only	only	ADV
mk61rf5885n	13	20	1.5ìü	1.5ìü	NUM
mk61rf5885n	13	21	ác	ác	NOUN
mk61rf5885n	13	22	of	of	ADP
mk61rf5885n	13	23	thermal	thermal	ADJ
mk61rf5885n	13	24	hysteresis	hysteresis	NOUN
mk61rf5885n	13	25	at	at	ADP
mk61rf5885n	13	26	20	20	NUM
mk61rf5885n	13	27	-	-	SYM
mk61rf5885n	13	28	40	40	NUM
mk61rf5885n	13	29	mg	mg	NOUN
mk61rf5885n	13	30	/	/	SYM
mk61rf5885n	13	31	ml	ml	PROPN
mk61rf5885n	13	32	.	.	PUNCT
mk61rf5885n	13	33	consistent	consistent	ADJ
mk61rf5885n	13	34	with	with	ADP
mk61rf5885n	13	35	our	our	PRON
mk61rf5885n	13	36	structural	structural	ADJ
mk61rf5885n	13	37	determinations	determination	NOUN
mk61rf5885n	13	38	,	,	PUNCT
mk61rf5885n	13	39	thermal	thermal	ADJ
mk61rf5885n	13	40	hysteresis	hysteresis	NOUN
mk61rf5885n	13	41	activity	activity	NOUN
mk61rf5885n	13	42	is	be	AUX
mk61rf5885n	13	43	unaffected	unaffected	ADJ
mk61rf5885n	13	44	by	by	ADP
mk61rf5885n	13	45	protease	protease	NOUN
mk61rf5885n	13	46	,	,	PUNCT
mk61rf5885n	13	47	but	but	CCONJ
mk61rf5885n	13	48	is	be	AUX
mk61rf5885n	13	49	rapidly	rapidly	ADV
mk61rf5885n	13	50	lost	lose	VERB
mk61rf5885n	13	51	upon	upon	SCONJ
mk61rf5885n	13	52	treatment	treatment	NOUN
mk61rf5885n	13	53	with	with	ADP
mk61rf5885n	13	54	endo	endo	PROPN
mk61rf5885n	13	55	ì_å_-(1?4	ì_å_-(1?4	NOUN
mk61rf5885n	13	56	)	)	PUNCT
mk61rf5885n	13	57	xylanase	xylanase	NOUN
mk61rf5885n	13	58	.	.	PUNCT
mk61rf5885n	14	1	this	this	PRON
mk61rf5885n	14	2	is	be	AUX
mk61rf5885n	14	3	the	the	DET
mk61rf5885n	14	4	first	first	ADJ
mk61rf5885n	14	5	high	high	ADJ
mk61rf5885n	14	6	molecular	molecular	ADJ
mk61rf5885n	14	7	weight	weight	NOUN
mk61rf5885n	14	8	antifreeze	antifreeze	VERB
mk61rf5885n	14	9	to	to	PART
mk61rf5885n	14	10	be	be	AUX
mk61rf5885n	14	11	isolated	isolate	VERB
mk61rf5885n	14	12	from	from	ADP
mk61rf5885n	14	13	a	a	DET
mk61rf5885n	14	14	freeze	freeze	NOUN
mk61rf5885n	14	15	tolerant	tolerant	ADJ
mk61rf5885n	14	16	animal	animal	NOUN
mk61rf5885n	14	17	.	.	PUNCT
mk61rf5885n	15	1	we	we	PRON
mk61rf5885n	15	2	have	have	AUX
mk61rf5885n	15	3	subsequently	subsequently	ADV
mk61rf5885n	15	4	isolated	isolate	VERB
mk61rf5885n	15	5	similar	similar	ADJ
mk61rf5885n	15	6	th	th	ADJ
mk61rf5885n	15	7	-	-	PUNCT
mk61rf5885n	15	8	active	active	ADJ
mk61rf5885n	15	9	glycolipids	glycolipid	NOUN
mk61rf5885n	15	10	from	from	ADP
mk61rf5885n	15	11	numerous	numerous	ADJ
mk61rf5885n	15	12	organisms	organism	NOUN
mk61rf5885n	15	13	,	,	PUNCT
mk61rf5885n	15	14	including	include	VERB
mk61rf5885n	15	15	other	other	ADJ
mk61rf5885n	15	16	insects	insect	NOUN
mk61rf5885n	15	17	,	,	PUNCT
mk61rf5885n	15	18	plants	plant	NOUN
mk61rf5885n	15	19	,	,	PUNCT
mk61rf5885n	15	20	fish	fish	NOUN
mk61rf5885n	15	21	and	and	CCONJ
mk61rf5885n	15	22	amphibians	amphibian	NOUN
mk61rf5885n	15	23	.	.	PUNCT