id	sid	tid	token	lemma	pos
ff365427r9h	1	1	dynactin	dynactin	PROPN
ff365427r9h	1	2	is	be	AUX
ff365427r9h	1	3	a	a	DET
ff365427r9h	1	4	multi	multi	ADJ
ff365427r9h	1	5	-	-	ADJ
ff365427r9h	1	6	subunit	subunit	NOUN
ff365427r9h	1	7	complex	complex	NOUN
ff365427r9h	1	8	implicated	implicate	VERB
ff365427r9h	1	9	in	in	ADP
ff365427r9h	1	10	cytoplasmic	cytoplasmic	ADJ
ff365427r9h	1	11	dynein	dynein	NOUN
ff365427r9h	1	12	-	-	PUNCT
ff365427r9h	1	13	driven	drive	VERB
ff365427r9h	1	14	transport	transport	NOUN
ff365427r9h	1	15	of	of	ADP
ff365427r9h	1	16	membranous	membranous	ADJ
ff365427r9h	1	17	organelles	organelle	NOUN
ff365427r9h	1	18	,	,	PUNCT
ff365427r9h	1	19	mitotic	mitotic	ADJ
ff365427r9h	1	20	chromosomes	chromosome	NOUN
ff365427r9h	1	21	and	and	CCONJ
ff365427r9h	1	22	other	other	ADJ
ff365427r9h	1	23	cargos	cargo	NOUN
ff365427r9h	1	24	.	.	PUNCT
ff365427r9h	2	1	the	the	DET
ff365427r9h	2	2	p150glued	p150glue	VERB
ff365427r9h	2	3	subunit	subunit	NOUN
ff365427r9h	2	4	of	of	ADP
ff365427r9h	2	5	dynactin	dynactin	NOUN
ff365427r9h	2	6	contains	contain	VERB
ff365427r9h	2	7	a	a	DET
ff365427r9h	2	8	microtubule	microtubule	NOUN
ff365427r9h	2	9	(	(	PUNCT
ff365427r9h	2	10	mt)-binding	mt)-binde	VERB
ff365427r9h	2	11	domain	domain	NOUN
ff365427r9h	2	12	composed	compose	VERB
ff365427r9h	2	13	of	of	ADP
ff365427r9h	2	14	a	a	DET
ff365427r9h	2	15	cap	cap	NOUN
ff365427r9h	2	16	-	-	PUNCT
ff365427r9h	2	17	gly	gly	NOUN
ff365427r9h	2	18	motif	motif	NOUN
ff365427r9h	2	19	and	and	CCONJ
ff365427r9h	2	20	an	an	DET
ff365427r9h	2	21	adjacent	adjacent	ADJ
ff365427r9h	2	22	basic	basic	ADJ
ff365427r9h	2	23	domain	domain	NOUN
ff365427r9h	2	24	(	(	PUNCT
ff365427r9h	2	25	bd	bd	PROPN
ff365427r9h	2	26	)	)	PUNCT
ff365427r9h	2	27	.	.	PUNCT
ff365427r9h	3	1	despite	despite	SCONJ
ff365427r9h	3	2	the	the	DET
ff365427r9h	3	3	widespread	widespread	ADJ
ff365427r9h	3	4	conservation	conservation	NOUN
ff365427r9h	3	5	of	of	ADP
ff365427r9h	3	6	this	this	DET
ff365427r9h	3	7	structure	structure	NOUN
ff365427r9h	3	8	,	,	PUNCT
ff365427r9h	3	9	the	the	DET
ff365427r9h	3	10	requirement	requirement	NOUN
ff365427r9h	3	11	for	for	ADP
ff365427r9h	3	12	mt	mt	PROPN
ff365427r9h	3	13	-	-	PUNCT
ff365427r9h	3	14	binding	bind	VERB
ff365427r9h	3	15	remains	remain	VERB
ff365427r9h	3	16	controversial	controversial	ADJ
ff365427r9h	3	17	.	.	PUNCT
ff365427r9h	4	1	to	to	PART
ff365427r9h	4	2	test	test	VERB
ff365427r9h	4	3	the	the	DET
ff365427r9h	4	4	hypothesis	hypothesis	NOUN
ff365427r9h	4	5	that	that	SCONJ
ff365427r9h	4	6	mt	mt	PROPN
ff365427r9h	4	7	-	-	PUNCT
ff365427r9h	4	8	binding	binding	PROPN
ff365427r9h	4	9	is	be	AUX
ff365427r9h	4	10	important	important	ADJ
ff365427r9h	4	11	for	for	ADP
ff365427r9h	4	12	transport	transport	NOUN
ff365427r9h	4	13	,	,	PUNCT
ff365427r9h	4	14	we	we	PRON
ff365427r9h	4	15	engineered	engineer	VERB
ff365427r9h	4	16	mutants	mutant	NOUN
ff365427r9h	4	17	that	that	PRON
ff365427r9h	4	18	disrupt	disrupt	VERB
ff365427r9h	4	19	normal	normal	ADJ
ff365427r9h	4	20	mt	mt	PROPN
ff365427r9h	4	21	-	-	PUNCT
ff365427r9h	4	22	binding	bind	VERB
ff365427r9h	4	23	and	and	CCONJ
ff365427r9h	4	24	developed	develop	VERB
ff365427r9h	4	25	a	a	DET
ff365427r9h	4	26	shrna	shrna	PROPN
ff365427r9h	4	27	/	/	SYM
ff365427r9h	4	28	rescue	rescue	NOUN
ff365427r9h	4	29	assay	assay	NOUN
ff365427r9h	4	30	to	to	PART
ff365427r9h	4	31	replace	replace	VERB
ff365427r9h	4	32	endogenous	endogenous	ADJ
ff365427r9h	4	33	p150glued	p150glue	VERB
ff365427r9h	4	34	with	with	ADP
ff365427r9h	4	35	each	each	DET
ff365427r9h	4	36	mutant	mutant	NOUN
ff365427r9h	4	37	.	.	PUNCT
ff365427r9h	5	1	assaying	assay	VERB
ff365427r9h	5	2	er	er	INTJ
ff365427r9h	5	3	-	-	PUNCT
ff365427r9h	5	4	golgi	golgi	NOUN
ff365427r9h	5	5	transport	transport	NOUN
ff365427r9h	5	6	first	first	ADV
ff365427r9h	5	7	,	,	PUNCT
ff365427r9h	5	8	wild	wild	ADJ
ff365427r9h	5	9	-	-	PUNCT
ff365427r9h	5	10	type	type	NOUN
ff365427r9h	5	11	gfp	gfp	NOUN
ff365427r9h	5	12	-	-	PUNCT
ff365427r9h	5	13	p150glued	p150glue	VERB
ff365427r9h	5	14	was	be	AUX
ff365427r9h	5	15	successful	successful	ADJ
ff365427r9h	5	16	in	in	ADP
ff365427r9h	5	17	rescuing	rescue	VERB
ff365427r9h	5	18	normal	normal	ADJ
ff365427r9h	5	19	motility	motility	NOUN
ff365427r9h	5	20	.	.	PUNCT
ff365427r9h	6	1	in	in	ADP
ff365427r9h	6	2	contrast	contrast	NOUN
ff365427r9h	6	3	,	,	PUNCT
ff365427r9h	6	4	a	a	DET
ff365427r9h	6	5	gfp	gfp	NOUN
ff365427r9h	6	6	-	-	PUNCT
ff365427r9h	6	7	p150glued	p150glue	VERB
ff365427r9h	6	8	g59s	g59s	VERB
ff365427r9h	6	9	mutant	mutant	NOUN
ff365427r9h	6	10	that	that	PRON
ff365427r9h	6	11	displays	display	VERB
ff365427r9h	6	12	reduced	reduce	VERB
ff365427r9h	6	13	mt	mt	PROPN
ff365427r9h	6	14	-	-	PUNCT
ff365427r9h	6	15	binding	bind	VERB
ff365427r9h	6	16	incorporated	incorporate	VERB
ff365427r9h	6	17	into	into	ADP
ff365427r9h	6	18	the	the	DET
ff365427r9h	6	19	dynactin	dynactin	NOUN
ff365427r9h	6	20	complex	complex	NOUN
ff365427r9h	6	21	but	but	CCONJ
ff365427r9h	6	22	failed	fail	VERB
ff365427r9h	6	23	to	to	PART
ff365427r9h	6	24	rescue	rescue	VERB
ff365427r9h	6	25	normal	normal	ADJ
ff365427r9h	6	26	er	er	INTJ
ff365427r9h	6	27	-	-	PUNCT
ff365427r9h	6	28	golgi	golgi	NOUN
ff365427r9h	6	29	transport	transport	NOUN
ff365427r9h	6	30	.	.	PUNCT
ff365427r9h	7	1	to	to	PART
ff365427r9h	7	2	test	test	VERB
ff365427r9h	7	3	if	if	SCONJ
ff365427r9h	7	4	dynamic	dynamic	ADJ
ff365427r9h	7	5	mt	mt	PROPN
ff365427r9h	7	6	-	-	PUNCT
ff365427r9h	7	7	binding	binding	PROPN
ff365427r9h	7	8	was	be	AUX
ff365427r9h	7	9	required	require	VERB
ff365427r9h	7	10	,	,	PUNCT
ff365427r9h	7	11	we	we	PRON
ff365427r9h	7	12	generated	generate	VERB
ff365427r9h	7	13	a	a	DET
ff365427r9h	7	14	gfp	gfp	NOUN
ff365427r9h	7	15	-	-	PUNCT
ff365427r9h	7	16	p150glued	p150glue	VERB
ff365427r9h	7	17	-	-	PUNCT
ff365427r9h	7	18	map4	map4	PROPN
ff365427r9h	7	19	chimera	chimera	NOUN
ff365427r9h	7	20	which	which	PRON
ff365427r9h	7	21	exhibited	exhibit	VERB
ff365427r9h	7	22	stable	stable	ADJ
ff365427r9h	7	23	mt	mt	PROPN
ff365427r9h	7	24	binding	binding	PROPN
ff365427r9h	7	25	.	.	PUNCT
ff365427r9h	8	1	this	this	DET
ff365427r9h	8	2	construct	construct	NOUN
ff365427r9h	8	3	also	also	ADV
ff365427r9h	8	4	failed	fail	VERB
ff365427r9h	8	5	to	to	PART
ff365427r9h	8	6	rescue	rescue	VERB
ff365427r9h	8	7	er	er	INTJ
ff365427r9h	8	8	-	-	PUNCT
ff365427r9h	8	9	golgi	golgi	NOUN
ff365427r9h	8	10	transport	transport	NOUN
ff365427r9h	8	11	but	but	CCONJ
ff365427r9h	8	12	resulted	result	VERB
ff365427r9h	8	13	in	in	ADP
ff365427r9h	8	14	immobilization	immobilization	NOUN
ff365427r9h	8	15	of	of	ADP
ff365427r9h	8	16	vesicles	vesicle	NOUN
ff365427r9h	8	17	along	along	ADP
ff365427r9h	8	18	mts	mts	NOUN
ff365427r9h	8	19	.	.	PUNCT
ff365427r9h	9	1	to	to	PART
ff365427r9h	9	2	test	test	VERB
ff365427r9h	9	3	the	the	DET
ff365427r9h	9	4	contribution	contribution	NOUN
ff365427r9h	9	5	of	of	ADP
ff365427r9h	9	6	the	the	DET
ff365427r9h	9	7	bd	bd	PROPN
ff365427r9h	9	8	,	,	PUNCT
ff365427r9h	9	9	a	a	DET
ff365427r9h	9	10	naturally	naturally	ADV
ff365427r9h	9	11	occurring	occur	VERB
ff365427r9h	9	12	p150glued	p150glue	VERB
ff365427r9h	9	13	isoform	isoform	NOUN
ff365427r9h	9	14	with	with	ADP
ff365427r9h	9	15	an	an	DET
ff365427r9h	9	16	internal	internal	ADJ
ff365427r9h	9	17	deletion	deletion	NOUN
ff365427r9h	9	18	in	in	ADP
ff365427r9h	9	19	the	the	DET
ff365427r9h	9	20	bd	bd	PROPN
ff365427r9h	9	21	was	be	AUX
ff365427r9h	9	22	prepared	prepare	VERB
ff365427r9h	9	23	.	.	PUNCT
ff365427r9h	10	1	this	this	DET
ff365427r9h	10	2	construct	construct	NOUN
ff365427r9h	10	3	was	be	AUX
ff365427r9h	10	4	successful	successful	ADJ
ff365427r9h	10	5	in	in	ADP
ff365427r9h	10	6	rescuing	rescue	VERB
ff365427r9h	10	7	er	er	INTJ
ff365427r9h	10	8	-	-	PUNCT
ff365427r9h	10	9	golgi	golgi	NOUN
ff365427r9h	10	10	transport	transport	NOUN
ff365427r9h	10	11	but	but	CCONJ
ff365427r9h	10	12	displayed	display	VERB
ff365427r9h	10	13	reduced	reduce	VERB
ff365427r9h	10	14	run	run	NOUN
ff365427r9h	10	15	-	-	PUNCT
ff365427r9h	10	16	lengths	length	NOUN
ff365427r9h	10	17	for	for	ADP
ff365427r9h	10	18	motile	motile	NOUN
ff365427r9h	10	19	vesicles	vesicle	NOUN
ff365427r9h	10	20	.	.	PUNCT
ff365427r9h	11	1	because	because	SCONJ
ff365427r9h	11	2	dynactin	dynactin	NOUN
ff365427r9h	11	3	is	be	AUX
ff365427r9h	11	4	also	also	ADV
ff365427r9h	11	5	thought	think	VERB
ff365427r9h	11	6	to	to	PART
ff365427r9h	11	7	augment	augment	VERB
ff365427r9h	11	8	dynein	dynein	NOUN
ff365427r9h	11	9	-	-	PUNCT
ff365427r9h	11	10	driven	drive	VERB
ff365427r9h	11	11	chromosome	chromosome	NOUN
ff365427r9h	11	12	movement	movement	NOUN
ff365427r9h	11	13	during	during	ADP
ff365427r9h	11	14	mitosis	mitosis	NOUN
ff365427r9h	11	15	,	,	PUNCT
ff365427r9h	11	16	we	we	PRON
ff365427r9h	11	17	applied	apply	VERB
ff365427r9h	11	18	the	the	DET
ff365427r9h	11	19	same	same	ADJ
ff365427r9h	11	20	shrna	shrna	PROPN
ff365427r9h	11	21	/	/	SYM
ff365427r9h	11	22	rescue	rescue	NOUN
ff365427r9h	11	23	approach	approach	NOUN
ff365427r9h	11	24	to	to	ADP
ff365427r9h	11	25	assay	assay	PROPN
ff365427r9h	11	26	kinetochore	kinetochore	PROPN
ff365427r9h	11	27	-	-	PUNCT
ff365427r9h	11	28	mt	mt	PROPN
ff365427r9h	11	29	capture	capture	NOUN
ff365427r9h	11	30	,	,	PUNCT
ff365427r9h	11	31	chromosome	chromosome	NOUN
ff365427r9h	11	32	movement	movement	NOUN
ff365427r9h	11	33	,	,	PUNCT
ff365427r9h	11	34	and	and	CCONJ
ff365427r9h	11	35	the	the	DET
ff365427r9h	11	36	timing	timing	NOUN
ff365427r9h	11	37	of	of	ADP
ff365427r9h	11	38	anaphase	anaphase	NOUN
ff365427r9h	11	39	onset	onset	VERB
ff365427r9h	11	40	.	.	PUNCT
ff365427r9h	12	1	whereas	whereas	SCONJ
ff365427r9h	12	2	wild	wild	ADJ
ff365427r9h	12	3	-	-	PUNCT
ff365427r9h	12	4	type	type	NOUN
ff365427r9h	12	5	gfp	gfp	NOUN
ff365427r9h	12	6	-	-	PUNCT
ff365427r9h	12	7	p150glued	p150glue	VERB
ff365427r9h	12	8	rescued	rescue	VERB
ff365427r9h	12	9	each	each	DET
ff365427r9h	12	10	aspect	aspect	NOUN
ff365427r9h	12	11	of	of	ADP
ff365427r9h	12	12	mitotic	mitotic	ADJ
ff365427r9h	12	13	dynactin	dynactin	NOUN
ff365427r9h	12	14	function	function	NOUN
ff365427r9h	12	15	,	,	PUNCT
ff365427r9h	12	16	the	the	DET
ff365427r9h	12	17	g59s	g59s	NOUN
ff365427r9h	12	18	mutant	mutant	NOUN
ff365427r9h	12	19	disrupted	disrupt	VERB
ff365427r9h	12	20	kinetochore	kinetochore	NOUN
ff365427r9h	12	21	-	-	PUNCT
ff365427r9h	12	22	mt	mt	PROPN
ff365427r9h	12	23	capture	capture	NOUN
ff365427r9h	12	24	similar	similar	ADJ
ff365427r9h	12	25	to	to	ADP
ff365427r9h	12	26	cells	cell	NOUN
ff365427r9h	12	27	treated	treat	VERB
ff365427r9h	12	28	with	with	ADP
ff365427r9h	12	29	nocodazole	nocodazole	NOUN
ff365427r9h	12	30	.	.	PUNCT
ff365427r9h	13	1	a	a	DET
ff365427r9h	13	2	subset	subset	NOUN
ff365427r9h	13	3	of	of	ADP
ff365427r9h	13	4	g59s	g59s	SPACE
ff365427r9h	13	5	-	-	PUNCT
ff365427r9h	13	6	rescued	rescue	VERB
ff365427r9h	13	7	cells	cell	NOUN
ff365427r9h	13	8	entered	enter	VERB
ff365427r9h	13	9	anaphase	anaphase	NOUN
ff365427r9h	13	10	despite	despite	SCONJ
ff365427r9h	13	11	a	a	DET
ff365427r9h	13	12	loss	loss	NOUN
ff365427r9h	13	13	of	of	ADP
ff365427r9h	13	14	persistent	persistent	ADJ
ff365427r9h	13	15	kinetochore	kinetochore	NOUN
ff365427r9h	13	16	-	-	PUNCT
ff365427r9h	13	17	mt	mt	PROPN
ff365427r9h	13	18	attachments	attachment	NOUN
ff365427r9h	13	19	,	,	PUNCT
ff365427r9h	13	20	reflecting	reflect	VERB
ff365427r9h	13	21	a	a	DET
ff365427r9h	13	22	defect	defect	NOUN
ff365427r9h	13	23	in	in	ADP
ff365427r9h	13	24	the	the	DET
ff365427r9h	13	25	spindle	spindle	NOUN
ff365427r9h	13	26	assembly	assembly	NOUN
ff365427r9h	13	27	checkpoint	checkpoint	NOUN
ff365427r9h	13	28	.	.	PUNCT
ff365427r9h	14	1	together	together	ADV
ff365427r9h	14	2	these	these	DET
ff365427r9h	14	3	studies	study	NOUN
ff365427r9h	14	4	suggest	suggest	VERB
ff365427r9h	14	5	that	that	SCONJ
ff365427r9h	14	6	the	the	DET
ff365427r9h	14	7	mt	mt	PROPN
ff365427r9h	14	8	-	-	PUNCT
ff365427r9h	14	9	binding	bind	VERB
ff365427r9h	14	10	activity	activity	NOUN
ff365427r9h	14	11	of	of	ADP
ff365427r9h	14	12	p150glued	p150glue	VERB
ff365427r9h	14	13	is	be	AUX
ff365427r9h	14	14	essential	essential	ADJ
ff365427r9h	14	15	during	during	ADP
ff365427r9h	14	16	both	both	DET
ff365427r9h	14	17	interphase	interphase	NOUN
ff365427r9h	14	18	and	and	CCONJ
ff365427r9h	14	19	mitosis	mitosis	NOUN
ff365427r9h	14	20	.	.	PUNCT