id	sid	tid	token	lemma	pos
ff365427r4s	1	1	pacific	pacific	PROPN
ff365427r4s	1	2	salmon	salmon	PROPN
ff365427r4s	1	3	(	(	PUNCT
ff365427r4s	1	4	oncorhynchus	oncorhynchus	PROPN
ff365427r4s	1	5	spp	spp	PROPN
ff365427r4s	1	6	.	.	PUNCT
ff365427r4s	1	7	)	)	PUNCT
ff365427r4s	1	8	provide	provide	VERB
ff365427r4s	1	9	resource	resource	NOUN
ff365427r4s	1	10	subsidies	subsidy	NOUN
ff365427r4s	1	11	to	to	PART
ff365427r4s	1	12	stream	stream	VERB
ff365427r4s	1	13	and	and	CCONJ
ff365427r4s	1	14	riparian	riparian	ADJ
ff365427r4s	1	15	ecosystems	ecosystem	NOUN
ff365427r4s	1	16	throughout	throughout	ADP
ff365427r4s	1	17	much	much	ADJ
ff365427r4s	1	18	of	of	ADP
ff365427r4s	1	19	the	the	DET
ff365427r4s	1	20	northern	northern	PROPN
ff365427r4s	1	21	pacific	pacific	PROPN
ff365427r4s	1	22	rim	rim	PROPN
ff365427r4s	1	23	.	.	PUNCT
ff365427r4s	2	1	the	the	DET
ff365427r4s	2	2	dynamics	dynamic	NOUN
ff365427r4s	2	3	of	of	ADP
ff365427r4s	2	4	salmon	salmon	NOUN
ff365427r4s	2	5	runs	run	NOUN
ff365427r4s	2	6	and	and	CCONJ
ff365427r4s	2	7	environmental	environmental	ADJ
ff365427r4s	2	8	conditions	condition	NOUN
ff365427r4s	2	9	of	of	ADP
ff365427r4s	2	10	streams	stream	NOUN
ff365427r4s	2	11	in	in	ADP
ff365427r4s	2	12	which	which	PRON
ff365427r4s	2	13	they	they	PRON
ff365427r4s	2	14	spawn	spawn	VERB
ff365427r4s	2	15	can	can	AUX
ff365427r4s	2	16	vary	vary	VERB
ff365427r4s	2	17	in	in	ADP
ff365427r4s	2	18	space	space	NOUN
ff365427r4s	2	19	and	and	CCONJ
ff365427r4s	2	20	time	time	NOUN
ff365427r4s	2	21	,	,	PUNCT
ff365427r4s	2	22	potentially	potentially	ADV
ff365427r4s	2	23	creating	create	VERB
ff365427r4s	2	24	or	or	CCONJ
ff365427r4s	2	25	reinforcing	reinforce	VERB
ff365427r4s	2	26	heterogeneity	heterogeneity	NOUN
ff365427r4s	2	27	in	in	ADP
ff365427r4s	2	28	these	these	DET
ff365427r4s	2	29	recipient	recipient	ADJ
ff365427r4s	2	30	ecosystems	ecosystem	NOUN
ff365427r4s	2	31	.	.	PUNCT
ff365427r4s	3	1	enrichment	enrichment	NOUN
ff365427r4s	3	2	of	of	ADP
ff365427r4s	3	3	stream	stream	NOUN
ff365427r4s	3	4	biota	biota	NOUN
ff365427r4s	3	5	by	by	ADP
ff365427r4s	3	6	salmon	salmon	NOUN
ff365427r4s	3	7	-	-	PUNCT
ff365427r4s	3	8	derived	derive	VERB
ff365427r4s	3	9	nutrients	nutrient	NOUN
ff365427r4s	3	10	and	and	CCONJ
ff365427r4s	3	11	energy	energy	NOUN
ff365427r4s	3	12	may	may	AUX
ff365427r4s	3	13	be	be	AUX
ff365427r4s	3	14	offset	offset	VERB
ff365427r4s	3	15	by	by	ADP
ff365427r4s	3	16	spawner	spawner	NOUN
ff365427r4s	3	17	-	-	PUNCT
ff365427r4s	3	18	mediated	mediate	VERB
ff365427r4s	3	19	physical	physical	ADJ
ff365427r4s	3	20	disturbance	disturbance	NOUN
ff365427r4s	3	21	of	of	ADP
ff365427r4s	3	22	sediments	sediment	NOUN
ff365427r4s	3	23	during	during	ADP
ff365427r4s	3	24	migration	migration	NOUN
ff365427r4s	3	25	and	and	CCONJ
ff365427r4s	3	26	nest	nest	NOUN
ff365427r4s	3	27	construction	construction	NOUN
ff365427r4s	3	28	.	.	PUNCT
ff365427r4s	4	1	these	these	DET
ff365427r4s	4	2	two	two	NUM
ff365427r4s	4	3	contrasting	contrast	VERB
ff365427r4s	4	4	roles	role	NOUN
ff365427r4s	4	5	of	of	ADP
ff365427r4s	4	6	salmon	salmon	NOUN
ff365427r4s	4	7	are	be	AUX
ff365427r4s	4	8	normally	normally	ADV
ff365427r4s	4	9	studied	study	VERB
ff365427r4s	4	10	at	at	ADP
ff365427r4s	4	11	reach	reach	NOUN
ff365427r4s	4	12	scales	scale	NOUN
ff365427r4s	4	13	but	but	CCONJ
ff365427r4s	4	14	may	may	AUX
ff365427r4s	4	15	vary	vary	VERB
ff365427r4s	4	16	at	at	ADP
ff365427r4s	4	17	larger	large	ADJ
ff365427r4s	4	18	scales	scale	NOUN
ff365427r4s	4	19	for	for	ADP
ff365427r4s	4	20	enrichment	enrichment	NOUN
ff365427r4s	4	21	and	and	CCONJ
ff365427r4s	4	22	at	at	ADP
ff365427r4s	4	23	smaller	small	ADJ
ff365427r4s	4	24	scales	scale	NOUN
ff365427r4s	4	25	for	for	ADP
ff365427r4s	4	26	salmon	salmon	NOUN
ff365427r4s	4	27	-	-	PUNCT
ff365427r4s	4	28	induced	induce	VERB
ff365427r4s	4	29	physical	physical	ADJ
ff365427r4s	4	30	disturbance	disturbance	NOUN
ff365427r4s	4	31	.	.	PUNCT
ff365427r4s	5	1	i	i	PRON
ff365427r4s	5	2	studied	study	VERB
ff365427r4s	5	3	the	the	DET
ff365427r4s	5	4	response	response	NOUN
ff365427r4s	5	5	of	of	ADP
ff365427r4s	5	6	benthic	benthic	ADJ
ff365427r4s	5	7	biofilm	biofilm	NOUN
ff365427r4s	5	8	(	(	PUNCT
ff365427r4s	5	9	i.e.	i.e.	X
ff365427r4s	5	10	,	,	PUNCT
ff365427r4s	5	11	the	the	DET
ff365427r4s	5	12	complex	complex	NOUN
ff365427r4s	5	13	of	of	ADP
ff365427r4s	5	14	autotrophic	autotrophic	ADJ
ff365427r4s	5	15	and	and	CCONJ
ff365427r4s	5	16	heterotrophic	heterotrophic	ADJ
ff365427r4s	5	17	organisms	organism	NOUN
ff365427r4s	5	18	growing	grow	VERB
ff365427r4s	5	19	on	on	ADP
ff365427r4s	5	20	submerged	submerged	ADJ
ff365427r4s	5	21	surfaces	surface	NOUN
ff365427r4s	5	22	)	)	PUNCT
ff365427r4s	5	23	to	to	ADP
ff365427r4s	5	24	the	the	DET
ff365427r4s	5	25	presence	presence	NOUN
ff365427r4s	5	26	of	of	ADP
ff365427r4s	5	27	salmon	salmon	NOUN
ff365427r4s	5	28	spawners	spawner	NOUN
ff365427r4s	5	29	in	in	ADP
ff365427r4s	5	30	seven	seven	NUM
ff365427r4s	5	31	southeast	southeast	PROPN
ff365427r4s	5	32	alaska	alaska	PROPN
ff365427r4s	5	33	streams	stream	NOUN
ff365427r4s	5	34	.	.	PUNCT
ff365427r4s	6	1	autotrophs	autotrophs	PROPN
ff365427r4s	6	2	and	and	CCONJ
ff365427r4s	6	3	heterotrophs	heterotroph	NOUN
ff365427r4s	6	4	were	be	AUX
ff365427r4s	6	5	nutrient	nutrient	NOUN
ff365427r4s	6	6	-	-	PUNCT
ff365427r4s	6	7	limited	limit	VERB
ff365427r4s	6	8	by	by	ADP
ff365427r4s	6	9	n	n	NOUN
ff365427r4s	6	10	or	or	CCONJ
ff365427r4s	6	11	p	p	NOUN
ff365427r4s	6	12	(	(	PUNCT
ff365427r4s	6	13	or	or	CCONJ
ff365427r4s	6	14	both	both	PRON
ff365427r4s	6	15	)	)	PUNCT
ff365427r4s	6	16	in	in	ADP
ff365427r4s	6	17	all	all	DET
ff365427r4s	6	18	study	study	NOUN
ff365427r4s	6	19	streams	stream	NOUN
ff365427r4s	6	20	before	before	SCONJ
ff365427r4s	6	21	the	the	DET
ff365427r4s	6	22	salmon	salmon	NOUN
ff365427r4s	6	23	run	run	VERB
ff365427r4s	6	24	.	.	PUNCT
ff365427r4s	7	1	during	during	ADP
ff365427r4s	7	2	the	the	DET
ff365427r4s	7	3	salmon	salmon	NOUN
ff365427r4s	7	4	run	run	NOUN
ff365427r4s	7	5	,	,	PUNCT
ff365427r4s	7	6	biofilm	biofilm	NOUN
ff365427r4s	7	7	nutrient	nutrient	NOUN
ff365427r4s	7	8	limitation	limitation	NOUN
ff365427r4s	7	9	was	be	AUX
ff365427r4s	7	10	partially	partially	ADV
ff365427r4s	7	11	alleviated	alleviate	VERB
ff365427r4s	7	12	,	,	PUNCT
ff365427r4s	7	13	but	but	CCONJ
ff365427r4s	7	14	biofilm	biofilm	NOUN
ff365427r4s	7	15	responses	response	NOUN
ff365427r4s	7	16	to	to	ADP
ff365427r4s	7	17	salmon	salmon	NOUN
ff365427r4s	7	18	varied	varied	ADJ
ff365427r4s	7	19	from	from	ADP
ff365427r4s	7	20	net	net	ADJ
ff365427r4s	7	21	biomass	biomass	NOUN
ff365427r4s	7	22	accrual	accrual	ADJ
ff365427r4s	7	23	to	to	ADP
ff365427r4s	7	24	net	net	ADJ
ff365427r4s	7	25	loss	loss	NOUN
ff365427r4s	7	26	among	among	ADP
ff365427r4s	7	27	streams	stream	NOUN
ff365427r4s	7	28	and	and	CCONJ
ff365427r4s	7	29	years	year	NOUN
ff365427r4s	7	30	.	.	PUNCT
ff365427r4s	8	1	when	when	SCONJ
ff365427r4s	8	2	data	datum	NOUN
ff365427r4s	8	3	from	from	ADP
ff365427r4s	8	4	multiple	multiple	ADJ
ff365427r4s	8	5	streams	stream	NOUN
ff365427r4s	8	6	and	and	CCONJ
ff365427r4s	8	7	years	year	NOUN
ff365427r4s	8	8	were	be	AUX
ff365427r4s	8	9	considered	consider	VERB
ff365427r4s	8	10	,	,	PUNCT
ff365427r4s	8	11	the	the	DET
ff365427r4s	8	12	95	95	NUM
ff365427r4s	8	13	%	%	NOUN
ff365427r4s	8	14	confidence	confidence	NOUN
ff365427r4s	8	15	interval	interval	NOUN
ff365427r4s	8	16	of	of	ADP
ff365427r4s	8	17	relative	relative	ADJ
ff365427r4s	8	18	salmon	salmon	NOUN
ff365427r4s	8	19	enrichment	enrichment	NOUN
ff365427r4s	8	20	was	be	AUX
ff365427r4s	8	21	positive	positive	ADJ
ff365427r4s	8	22	only	only	ADV
ff365427r4s	8	23	for	for	ADP
ff365427r4s	8	24	chlorophyll	chlorophyll	NOUN
ff365427r4s	8	25	a	a	DET
ff365427r4s	8	26	(	(	PUNCT
ff365427r4s	8	27	16	16	NUM
ff365427r4s	8	28	-	-	SYM
ff365427r4s	8	29	411	411	NUM
ff365427r4s	8	30	%	%	NOUN
ff365427r4s	8	31	increase	increase	NOUN
ff365427r4s	8	32	)	)	PUNCT
ff365427r4s	8	33	.	.	PUNCT
ff365427r4s	9	1	the	the	DET
ff365427r4s	9	2	wide	wide	ADJ
ff365427r4s	9	3	range	range	NOUN
ff365427r4s	9	4	of	of	ADP
ff365427r4s	9	5	biofilm	biofilm	NOUN
ff365427r4s	9	6	responses	response	NOUN
ff365427r4s	9	7	reflected	reflect	VERB
ff365427r4s	9	8	,	,	PUNCT
ff365427r4s	9	9	in	in	ADP
ff365427r4s	9	10	part	part	NOUN
ff365427r4s	9	11	,	,	PUNCT
ff365427r4s	9	12	variability	variability	NOUN
ff365427r4s	9	13	among	among	ADP
ff365427r4s	9	14	streams	stream	NOUN
ff365427r4s	9	15	and	and	CCONJ
ff365427r4s	9	16	years	year	NOUN
ff365427r4s	9	17	in	in	ADP
ff365427r4s	9	18	environmental	environmental	ADJ
ff365427r4s	9	19	conditions	condition	NOUN
ff365427r4s	9	20	,	,	PUNCT
ff365427r4s	9	21	such	such	ADJ
ff365427r4s	9	22	as	as	ADP
ff365427r4s	9	23	in	in	ADP
ff365427r4s	9	24	temperature	temperature	NOUN
ff365427r4s	9	25	and	and	CCONJ
ff365427r4s	9	26	nutrient	nutrient	NOUN
ff365427r4s	9	27	concentrations	concentration	NOUN
ff365427r4s	9	28	.	.	PUNCT
ff365427r4s	10	1	however	however	ADV
ff365427r4s	10	2	,	,	PUNCT
ff365427r4s	10	3	those	those	DET
ff365427r4s	10	4	environmental	environmental	ADJ
ff365427r4s	10	5	variables	variable	NOUN
ff365427r4s	10	6	explained	explain	VERB
ff365427r4s	10	7	only	only	ADV
ff365427r4s	10	8	a	a	DET
ff365427r4s	10	9	small	small	ADJ
ff365427r4s	10	10	amount	amount	NOUN
ff365427r4s	10	11	of	of	ADP
ff365427r4s	10	12	the	the	DET
ff365427r4s	10	13	overall	overall	ADJ
ff365427r4s	10	14	biofilm	biofilm	NOUN
ff365427r4s	10	15	variability	variability	NOUN
ff365427r4s	10	16	,	,	PUNCT
ff365427r4s	10	17	even	even	ADV
ff365427r4s	10	18	if	if	SCONJ
ff365427r4s	10	19	partitioned	partition	VERB
ff365427r4s	10	20	among	among	ADP
ff365427r4s	10	21	spatial	spatial	ADJ
ff365427r4s	10	22	scales	scale	NOUN
ff365427r4s	10	23	(	(	PUNCT
ff365427r4s	10	24	e.g.	e.g.	ADJ
ff365427r4s	10	25	,	,	PUNCT
ff365427r4s	10	26	stream	stream	NOUN
ff365427r4s	10	27	,	,	PUNCT
ff365427r4s	10	28	reach	reach	VERB
ff365427r4s	10	29	,	,	PUNCT
ff365427r4s	10	30	riffle	riffle	ADV
ff365427r4s	10	31	)	)	PUNCT
ff365427r4s	10	32	.	.	PUNCT
ff365427r4s	11	1	the	the	DET
ff365427r4s	11	2	only	only	ADJ
ff365427r4s	11	3	consistent	consistent	ADJ
ff365427r4s	11	4	predictor	predictor	NOUN
ff365427r4s	11	5	across	across	ADP
ff365427r4s	11	6	scales	scale	NOUN
ff365427r4s	11	7	was	be	AUX
ff365427r4s	11	8	sediment	sediment	NOUN
ff365427r4s	11	9	size	size	NOUN
ff365427r4s	11	10	,	,	PUNCT
ff365427r4s	11	11	especially	especially	ADV
ff365427r4s	11	12	during	during	ADP
ff365427r4s	11	13	the	the	DET
ff365427r4s	11	14	run	run	NOUN
ff365427r4s	11	15	,	,	PUNCT
ff365427r4s	11	16	demonstrating	demonstrate	VERB
ff365427r4s	11	17	the	the	DET
ff365427r4s	11	18	importance	importance	NOUN
ff365427r4s	11	19	of	of	ADP
ff365427r4s	11	20	salmon	salmon	NOUN
ff365427r4s	11	21	disturbance	disturbance	NOUN
ff365427r4s	11	22	.	.	PUNCT
ff365427r4s	12	1	high	high	ADJ
ff365427r4s	12	2	-	-	PUNCT
ff365427r4s	12	3	frequency	frequency	NOUN
ff365427r4s	12	4	sampling	sampling	NOUN
ff365427r4s	12	5	revealed	reveal	VERB
ff365427r4s	12	6	that	that	SCONJ
ff365427r4s	12	7	effects	effect	NOUN
ff365427r4s	12	8	attributed	attribute	VERB
ff365427r4s	12	9	to	to	ADP
ff365427r4s	12	10	salmon	salmon	NOUN
ff365427r4s	12	11	spawners	spawner	NOUN
ff365427r4s	12	12	may	may	AUX
ff365427r4s	12	13	reflect	reflect	VERB
ff365427r4s	12	14	other	other	ADJ
ff365427r4s	12	15	environmental	environmental	ADJ
ff365427r4s	12	16	variables	variable	NOUN
ff365427r4s	12	17	that	that	PRON
ff365427r4s	12	18	change	change	VERB
ff365427r4s	12	19	during	during	ADP
ff365427r4s	12	20	the	the	DET
ff365427r4s	12	21	salmon	salmon	NOUN
ff365427r4s	12	22	run	run	NOUN
ff365427r4s	12	23	,	,	PUNCT
ff365427r4s	12	24	such	such	ADJ
ff365427r4s	12	25	as	as	ADP
ff365427r4s	12	26	increased	increase	VERB
ff365427r4s	12	27	discharge	discharge	NOUN
ff365427r4s	12	28	later	later	ADV
ff365427r4s	12	29	in	in	ADP
ff365427r4s	12	30	the	the	DET
ff365427r4s	12	31	run	run	NOUN
ff365427r4s	12	32	.	.	PUNCT
ff365427r4s	13	1	salmon	salmon	NOUN
ff365427r4s	13	2	tissue	tissue	NOUN
ff365427r4s	13	3	decomposition	decomposition	NOUN
ff365427r4s	13	4	and	and	CCONJ
ff365427r4s	13	5	macroinvertebrate	macroinvertebrate	NOUN
ff365427r4s	13	6	assemblages	assemblage	NOUN
ff365427r4s	13	7	differed	differ	VERB
ff365427r4s	13	8	between	between	ADP
ff365427r4s	13	9	terrestrial	terrestrial	ADJ
ff365427r4s	13	10	and	and	CCONJ
ff365427r4s	13	11	aquatic	aquatic	ADJ
ff365427r4s	13	12	habitats	habitat	NOUN
ff365427r4s	13	13	likely	likely	ADV
ff365427r4s	13	14	due	due	ADJ
ff365427r4s	13	15	to	to	ADP
ff365427r4s	13	16	necrophagous	necrophagous	ADJ
ff365427r4s	13	17	guilds	guild	NOUN
ff365427r4s	13	18	in	in	ADP
ff365427r4s	13	19	terrestrial	terrestrial	ADJ
ff365427r4s	13	20	ecosystems	ecosystem	NOUN
ff365427r4s	13	21	.	.	PUNCT
ff365427r4s	14	1	integration	integration	NOUN
ff365427r4s	14	2	of	of	ADP
ff365427r4s	14	3	different	different	ADJ
ff365427r4s	14	4	spatial	spatial	ADJ
ff365427r4s	14	5	and	and	CCONJ
ff365427r4s	14	6	temporal	temporal	ADJ
ff365427r4s	14	7	scales	scale	NOUN
ff365427r4s	14	8	indicates	indicate	VERB
ff365427r4s	14	9	that	that	SCONJ
ff365427r4s	14	10	the	the	DET
ff365427r4s	14	11	ecological	ecological	ADJ
ff365427r4s	14	12	effects	effect	NOUN
ff365427r4s	14	13	of	of	ADP
ff365427r4s	14	14	salmon	salmon	NOUN
ff365427r4s	14	15	on	on	ADP
ff365427r4s	14	16	recipient	recipient	ADJ
ff365427r4s	14	17	ecosystems	ecosystem	NOUN
ff365427r4s	14	18	are	be	AUX
ff365427r4s	14	19	complex	complex	ADJ
ff365427r4s	14	20	,	,	PUNCT
ff365427r4s	14	21	and	and	CCONJ
ff365427r4s	14	22	constrained	constrain	VERB
ff365427r4s	14	23	by	by	ADP
ff365427r4s	14	24	multiple	multiple	ADJ
ff365427r4s	14	25	environmental	environmental	ADJ
ff365427r4s	14	26	factors	factor	NOUN
ff365427r4s	14	27	potentially	potentially	ADV
ff365427r4s	14	28	operating	operate	VERB
ff365427r4s	14	29	at	at	ADP
ff365427r4s	14	30	scales	scale	NOUN
ff365427r4s	14	31	different	different	ADJ
ff365427r4s	14	32	from	from	ADP
ff365427r4s	14	33	those	those	PRON
ff365427r4s	14	34	of	of	ADP
ff365427r4s	14	35	biotic	biotic	ADJ
ff365427r4s	14	36	responses	response	NOUN
ff365427r4s	14	37	.	.	PUNCT