id sid tid token lemma pos 8336h130n2q 1 1 prior prior ADV 8336h130n2q 1 2 to to ADP 8336h130n2q 1 3 follicle follicle NOUN 8336h130n2q 1 4 selection selection NOUN 8336h130n2q 1 5 , , PUNCT 8336h130n2q 1 6 granulosa granulosa ADJ 8336h130n2q 1 7 cells cell NOUN 8336h130n2q 1 8 are be AUX 8336h130n2q 1 9 actively actively ADV 8336h130n2q 1 10 maintained maintain VERB 8336h130n2q 1 11 in in ADP 8336h130n2q 1 12 an an DET 8336h130n2q 1 13 undifferentiated undifferentiated ADJ 8336h130n2q 1 14 state state NOUN 8336h130n2q 1 15 by by ADP 8336h130n2q 1 16 epidermal epidermal ADJ 8336h130n2q 1 17 growth growth NOUN 8336h130n2q 1 18 factor factor NOUN 8336h130n2q 1 19 ligands ligand NOUN 8336h130n2q 1 20 ( ( PUNCT 8336h130n2q 1 21 egfl egfl PROPN 8336h130n2q 1 22 ) ) PUNCT 8336h130n2q 1 23 that that PRON 8336h130n2q 1 24 signal signal NOUN 8336h130n2q 1 25 via via ADP 8336h130n2q 1 26 the the DET 8336h130n2q 1 27 mitogen mitogen NOUN 8336h130n2q 1 28 activated activate VERB 8336h130n2q 1 29 protein protein NOUN 8336h130n2q 1 30 kinase kinase NOUN 8336h130n2q 1 31 ( ( PUNCT 8336h130n2q 1 32 mapk)/extracellular mapk)/extracellular ADJ 8336h130n2q 1 33 regulated regulate VERB 8336h130n2q 1 34 kinase kinase NOUN 8336h130n2q 1 35 ( ( PUNCT 8336h130n2q 1 36 erk erk PROPN 8336h130n2q 1 37 ) ) PUNCT 8336h130n2q 1 38 pathway pathway NOUN 8336h130n2q 1 39 inhibiting inhibit VERB 8336h130n2q 1 40 follicle follicle NOUN 8336h130n2q 1 41 stimulating stimulate VERB 8336h130n2q 1 42 hormone hormone NOUN 8336h130n2q 1 43 receptor receptor NOUN 8336h130n2q 1 44 ( ( PUNCT 8336h130n2q 1 45 fshr fshr NOUN 8336h130n2q 1 46 ) ) PUNCT 8336h130n2q 1 47 . . PUNCT 8336h130n2q 2 1 at at ADP 8336h130n2q 2 2 the the DET 8336h130n2q 2 3 time time NOUN 8336h130n2q 2 4 of of ADP 8336h130n2q 2 5 follicle follicle NOUN 8336h130n2q 2 6 selection selection NOUN 8336h130n2q 2 7 , , PUNCT 8336h130n2q 2 8 there there PRON 8336h130n2q 2 9 is be VERB 8336h130n2q 2 10 a a DET 8336h130n2q 2 11 release release NOUN 8336h130n2q 2 12 from from ADP 8336h130n2q 2 13 inhibitory inhibitory ADJ 8336h130n2q 2 14 mapk mapk PROPN 8336h130n2q 2 15 / / SYM 8336h130n2q 2 16 erk erk PROPN 8336h130n2q 2 17 signaling signaling NOUN 8336h130n2q 2 18 initiating initiate VERB 8336h130n2q 2 19 granulosa granulosa ADJ 8336h130n2q 2 20 cell cell NOUN 8336h130n2q 2 21 differentiation differentiation NOUN 8336h130n2q 2 22 , , PUNCT 8336h130n2q 2 23 causing cause VERB 8336h130n2q 2 24 an an DET 8336h130n2q 2 25 increase increase NOUN 8336h130n2q 2 26 in in ADP 8336h130n2q 2 27 fshr fshr NOUN 8336h130n2q 2 28 in in ADP 8336h130n2q 2 29 one one NUM 8336h130n2q 2 30 selected select VERB 8336h130n2q 2 31 follicle follicle NOUN 8336h130n2q 2 32 . . PUNCT 8336h130n2q 3 1 the the DET 8336h130n2q 3 2 present present ADJ 8336h130n2q 3 3 study study NOUN 8336h130n2q 3 4 was be AUX 8336h130n2q 3 5 conducted conduct VERB 8336h130n2q 3 6 in in ADP 8336h130n2q 3 7 cultured cultured ADJ 8336h130n2q 3 8 granulosa granulosa ADJ 8336h130n2q 3 9 cells cell NOUN 8336h130n2q 3 10 from from ADP 8336h130n2q 3 11 prehierarchal prehierarchal ADJ 8336h130n2q 3 12 follicles follicle NOUN 8336h130n2q 3 13 , , PUNCT 8336h130n2q 3 14 and and CCONJ 8336h130n2q 3 15 it it PRON 8336h130n2q 3 16 was be AUX 8336h130n2q 3 17 found find VERB 8336h130n2q 3 18 that that SCONJ 8336h130n2q 3 19 the the DET 8336h130n2q 3 20 effects effect NOUN 8336h130n2q 3 21 of of ADP 8336h130n2q 3 22 bone bone NOUN 8336h130n2q 3 23 morphogenetic morphogenetic ADJ 8336h130n2q 3 24 protein protein NOUN 8336h130n2q 3 25 2 2 NUM 8336h130n2q 3 26 ( ( PUNCT 8336h130n2q 3 27 bmp2 bmp2 PROPN 8336h130n2q 3 28 ) ) PUNCT 8336h130n2q 3 29 on on ADP 8336h130n2q 3 30 granulosa granulosa ADJ 8336h130n2q 3 31 cells cell NOUN 8336h130n2q 3 32 promote promote VERB 8336h130n2q 3 33 egfl egfl ADJ 8336h130n2q 3 34 expression expression NOUN 8336h130n2q 3 35 while while SCONJ 8336h130n2q 3 36 inhibiting inhibit VERB 8336h130n2q 3 37 fshr fshr ADJ 8336h130n2q 3 38 expression expression NOUN 8336h130n2q 3 39 , , PUNCT 8336h130n2q 3 40 by by ADP 8336h130n2q 3 41 either either CCONJ 8336h130n2q 3 42 promoting promote VERB 8336h130n2q 3 43 or or CCONJ 8336h130n2q 3 44 inhibiting inhibit VERB 8336h130n2q 3 45 indvidual indvidual ADJ 8336h130n2q 3 46 i i NOUN 8336h130n2q 3 47 d d PROPN 8336h130n2q 3 48 isoform isoform NOUN 8336h130n2q 3 49 expression expression NOUN 8336h130n2q 3 50 . . PUNCT 8336h130n2q 4 1 the the DET 8336h130n2q 4 2 inhibitory inhibitory ADJ 8336h130n2q 4 3 actions action NOUN 8336h130n2q 4 4 of of ADP 8336h130n2q 4 5 bmp2 bmp2 PROPN 8336h130n2q 4 6 on on ADP 8336h130n2q 4 7 fshr fshr ADJ 8336h130n2q 4 8 expression expression NOUN 8336h130n2q 4 9 were be AUX 8336h130n2q 4 10 neutralized neutralize VERB 8336h130n2q 4 11 by by ADP 8336h130n2q 4 12 treatment treatment NOUN 8336h130n2q 4 13 of of ADP 8336h130n2q 4 14 granulosa granulosa ADJ 8336h130n2q 4 15 cells cell NOUN 8336h130n2q 4 16 with with ADP 8336h130n2q 4 17 the the DET 8336h130n2q 4 18 bmp bmp NOUN 8336h130n2q 4 19 antagonist antagonist NOUN 8336h130n2q 4 20 , , PUNCT 8336h130n2q 4 21 noggin noggin PROPN 8336h130n2q 4 22 . . PUNCT 8336h130n2q 5 1 in in ADP 8336h130n2q 5 2 conclusion conclusion NOUN 8336h130n2q 5 3 , , PUNCT 8336h130n2q 5 4 prior prior ADV 8336h130n2q 5 5 to to ADP 8336h130n2q 5 6 follicle follicle NOUN 8336h130n2q 5 7 selection selection NOUN 8336h130n2q 5 8 , , PUNCT 8336h130n2q 5 9 bmp2 bmp2 PROPN 8336h130n2q 5 10 and and CCONJ 8336h130n2q 5 11 egfl egfl PROPN 8336h130n2q 5 12 act act PROPN 8336h130n2q 5 13 cooperatively cooperatively ADV 8336h130n2q 5 14 to to PART 8336h130n2q 5 15 prevent prevent VERB 8336h130n2q 5 16 the the DET 8336h130n2q 5 17 premature premature ADJ 8336h130n2q 5 18 differentiation differentiation NOUN 8336h130n2q 5 19 of of ADP 8336h130n2q 5 20 hen hen ADJ 8336h130n2q 5 21 granulosa granulosa PROPN 8336h130n2q 5 22 cells cell NOUN 8336h130n2q 5 23 . . PUNCT