id	sid	tid	token	lemma	pos
5999n298v5h	1	1	the	the	DET
5999n298v5h	1	2	role	role	NOUN
5999n298v5h	1	3	of	of	ADP
5999n298v5h	1	4	ecology	ecology	NOUN
5999n298v5h	1	5	in	in	ADP
5999n298v5h	1	6	speciation	speciation	NOUN
5999n298v5h	1	7	is	be	AUX
5999n298v5h	1	8	a	a	DET
5999n298v5h	1	9	central	central	ADJ
5999n298v5h	1	10	issue	issue	NOUN
5999n298v5h	1	11	in	in	ADP
5999n298v5h	1	12	evolutionary	evolutionary	ADJ
5999n298v5h	1	13	biology	biology	NOUN
5999n298v5h	1	14	.	.	PUNCT
5999n298v5h	2	1	here	here	ADV
5999n298v5h	2	2	,	,	PUNCT
5999n298v5h	2	3	five	five	NUM
5999n298v5h	2	4	fundamental	fundamental	ADJ
5999n298v5h	2	5	questions	question	NOUN
5999n298v5h	2	6	concerning	concern	VERB
5999n298v5h	2	7	ecological	ecological	ADJ
5999n298v5h	2	8	speciation	speciation	NOUN
5999n298v5h	2	9	-	-	PUNCT
5999n298v5h	2	10	with	with	ADP
5999n298v5h	2	11	-	-	PUNCT
5999n298v5h	2	12	gene	gene	NOUN
5999n298v5h	2	13	-	-	PUNCT
5999n298v5h	2	14	flow	flow	NOUN
5999n298v5h	2	15	are	be	AUX
5999n298v5h	2	16	investigated	investigate	VERB
5999n298v5h	2	17	in	in	ADP
5999n298v5h	2	18	the	the	DET
5999n298v5h	2	19	rhagoletis	rhagoletis	PROPN
5999n298v5h	2	20	pomonella	pomonella	NOUN
5999n298v5h	2	21	species	specie	NOUN
5999n298v5h	2	22	complex	complex	NOUN
5999n298v5h	2	23	,	,	PUNCT
5999n298v5h	2	24	a	a	DET
5999n298v5h	2	25	model	model	NOUN
5999n298v5h	2	26	for	for	ADP
5999n298v5h	2	27	sympatric	sympatric	ADJ
5999n298v5h	2	28	ecological	ecological	ADJ
5999n298v5h	2	29	speciation	speciation	NOUN
5999n298v5h	2	30	:	:	PUNCT
5999n298v5h	2	31	(	(	PUNCT
5999n298v5h	2	32	1	1	X
5999n298v5h	2	33	)	)	PUNCT
5999n298v5h	2	34	what	what	PRON
5999n298v5h	2	35	constitutes	constitute	VERB
5999n298v5h	2	36	a	a	DET
5999n298v5h	2	37	sympatrically	sympatrically	ADV
5999n298v5h	2	38	-	-	PUNCT
5999n298v5h	2	39	formed	form	VERB
5999n298v5h	2	40	species	specie	NOUN
5999n298v5h	2	41	?	?	PUNCT
5999n298v5h	3	1	(	(	PUNCT
5999n298v5h	3	2	2	2	X
5999n298v5h	3	3	)	)	PUNCT
5999n298v5h	3	4	is	be	AUX
5999n298v5h	3	5	the	the	DET
5999n298v5h	3	6	initiation	initiation	NOUN
5999n298v5h	3	7	of	of	ADP
5999n298v5h	3	8	ecological	ecological	ADJ
5999n298v5h	3	9	speciation	speciation	NOUN
5999n298v5h	3	10	in	in	ADP
5999n298v5h	3	11	sympatry	sympatry	NOUN
5999n298v5h	3	12	a	a	DET
5999n298v5h	3	13	common	common	ADJ
5999n298v5h	3	14	or	or	CCONJ
5999n298v5h	3	15	exceptional	exceptional	ADJ
5999n298v5h	3	16	phenomenon	phenomenon	NOUN
5999n298v5h	3	17	?	?	PUNCT
5999n298v5h	4	1	(	(	PUNCT
5999n298v5h	4	2	3	3	X
5999n298v5h	4	3	)	)	PUNCT
5999n298v5h	4	4	what	what	DET
5999n298v5h	4	5	traits	trait	NOUN
5999n298v5h	4	6	are	be	AUX
5999n298v5h	4	7	important	important	ADJ
5999n298v5h	4	8	to	to	PART
5999n298v5h	4	9	host	host	VERB
5999n298v5h	4	10	shifts	shift	NOUN
5999n298v5h	4	11	and	and	CCONJ
5999n298v5h	4	12	reproductive	reproductive	ADJ
5999n298v5h	4	13	isolation	isolation	NOUN
5999n298v5h	4	14	for	for	ADP
5999n298v5h	4	15	phytophagous	phytophagous	ADJ
5999n298v5h	4	16	insects	insect	NOUN
5999n298v5h	4	17	?	?	PUNCT
5999n298v5h	5	1	(	(	PUNCT
5999n298v5h	5	2	4	4	X
5999n298v5h	5	3	)	)	PUNCT
5999n298v5h	5	4	what	what	PRON
5999n298v5h	5	5	is	be	AUX
5999n298v5h	5	6	the	the	DET
5999n298v5h	5	7	source	source	NOUN
5999n298v5h	5	8	of	of	ADP
5999n298v5h	5	9	genetic	genetic	ADJ
5999n298v5h	5	10	variation	variation	NOUN
5999n298v5h	5	11	fueling	fuel	VERB
5999n298v5h	5	12	adaptive	adaptive	ADJ
5999n298v5h	5	13	divergence	divergence	NOUN
5999n298v5h	5	14	?	?	PUNCT
5999n298v5h	6	1	(	(	PUNCT
5999n298v5h	6	2	5	5	X
5999n298v5h	6	3	)	)	PUNCT
5999n298v5h	6	4	what	what	DET
5999n298v5h	6	5	role	role	NOUN
5999n298v5h	6	6	does	do	AUX
5999n298v5h	6	7	genome	genome	NOUN
5999n298v5h	6	8	structure	structure	NOUN
5999n298v5h	6	9	play	play	NOUN
5999n298v5h	6	10	in	in	ADP
5999n298v5h	6	11	speciation	speciation	NOUN
5999n298v5h	6	12	-	-	PUNCT
5999n298v5h	6	13	with	with	ADP
5999n298v5h	6	14	-	-	PUNCT
5999n298v5h	6	15	gene	gene	NOUN
5999n298v5h	6	16	-	-	PUNCT
5999n298v5h	6	17	flow	flow	NOUN
5999n298v5h	6	18	and	and	CCONJ
5999n298v5h	6	19	how	how	SCONJ
5999n298v5h	6	20	is	be	AUX
5999n298v5h	6	21	this	this	PRON
5999n298v5h	6	22	reflected	reflect	VERB
5999n298v5h	6	23	in	in	ADP
5999n298v5h	6	24	patterns	pattern	NOUN
5999n298v5h	6	25	of	of	ADP
5999n298v5h	6	26	genomic	genomic	ADJ
5999n298v5h	6	27	differentiation	differentiation	NOUN
5999n298v5h	6	28	as	as	ADP
5999n298v5h	6	29	speciation	speciation	NOUN
5999n298v5h	6	30	proceeds	proceed	NOUN
5999n298v5h	6	31	?	?	PUNCT
5999n298v5h	7	1	i	i	PRON
5999n298v5h	7	2	addressed	address	VERB
5999n298v5h	7	3	these	these	DET
5999n298v5h	7	4	questions	question	NOUN
5999n298v5h	7	5	by	by	ADP
5999n298v5h	7	6	investigating	investigate	VERB
5999n298v5h	7	7	two	two	NUM
5999n298v5h	7	8	components	component	NOUN
5999n298v5h	7	9	of	of	ADP
5999n298v5h	7	10	the	the	DET
5999n298v5h	7	11	r.	r.	PROPN
5999n298v5h	7	12	pomonella	pomonella	PROPN
5999n298v5h	7	13	species	species	PROPN
5999n298v5h	7	14	complex	complex	PROPN
5999n298v5h	7	15	:	:	PUNCT
5999n298v5h	7	16	1	1	X
5999n298v5h	7	17	)	)	PUNCT
5999n298v5h	7	18	populations	population	NOUN
5999n298v5h	7	19	of	of	ADP
5999n298v5h	7	20	r.	r.	PROPN
5999n298v5h	7	21	pomonella	pomonella	PROPN
5999n298v5h	7	22	infesting	infest	VERB
5999n298v5h	7	23	an	an	DET
5999n298v5h	7	24	array	array	NOUN
5999n298v5h	7	25	of	of	ADP
5999n298v5h	7	26	hawthorn	hawthorn	NOUN
5999n298v5h	7	27	species	specie	NOUN
5999n298v5h	7	28	in	in	ADP
5999n298v5h	7	29	the	the	DET
5999n298v5h	7	30	southeastern	southeastern	ADJ
5999n298v5h	7	31	u.s	u.s	PROPN
5999n298v5h	7	32	.	.	PROPN
5999n298v5h	7	33	and	and	CCONJ
5999n298v5h	7	34	(	(	PUNCT
5999n298v5h	7	35	2	2	X
5999n298v5h	7	36	)	)	PUNCT
5999n298v5h	7	37	the	the	DET
5999n298v5h	7	38	flowering	flower	VERB
5999n298v5h	7	39	dogwood	dogwood	NOUN
5999n298v5h	7	40	fly	fly	NOUN
5999n298v5h	7	41	,	,	PUNCT
5999n298v5h	7	42	the	the	DET
5999n298v5h	7	43	sister	sister	NOUN
5999n298v5h	7	44	species	specie	NOUN
5999n298v5h	7	45	to	to	ADP
5999n298v5h	7	46	r.	r.	PROPN
5999n298v5h	7	47	pomonella	pomonella	PROPN
5999n298v5h	7	48	.	.	PUNCT
5999n298v5h	8	1	i	i	PRON
5999n298v5h	8	2	found	find	VERB
5999n298v5h	8	3	that	that	SCONJ
5999n298v5h	8	4	southern	southern	ADJ
5999n298v5h	8	5	populations	population	NOUN
5999n298v5h	8	6	of	of	ADP
5999n298v5h	8	7	r.	r.	PROPN
5999n298v5h	8	8	pomonella	pomonella	PROPN
5999n298v5h	8	9	are	be	AUX
5999n298v5h	8	10	differentiated	differentiate	VERB
5999n298v5h	8	11	both	both	CCONJ
5999n298v5h	8	12	phenotypically	phenotypically	ADV
5999n298v5h	8	13	and	and	CCONJ
5999n298v5h	8	14	genetically	genetically	ADV
5999n298v5h	8	15	and	and	CCONJ
5999n298v5h	8	16	represent	represent	VERB
5999n298v5h	8	17	partially	partially	ADV
5999n298v5h	8	18	-	-	PUNCT
5999n298v5h	8	19	isolated	isolate	VERB
5999n298v5h	8	20	host	host	NOUN
5999n298v5h	8	21	races	race	NOUN
5999n298v5h	8	22	.	.	PUNCT
5999n298v5h	9	1	furthermore	furthermore	ADV
5999n298v5h	9	2	,	,	PUNCT
5999n298v5h	9	3	these	these	DET
5999n298v5h	9	4	populations	population	NOUN
5999n298v5h	9	5	may	may	AUX
5999n298v5h	9	6	have	have	AUX
5999n298v5h	9	7	acted	act	VERB
5999n298v5h	9	8	as	as	ADP
5999n298v5h	9	9	a	a	DET
5999n298v5h	9	10	reservoir	reservoir	NOUN
5999n298v5h	9	11	for	for	ADP
5999n298v5h	9	12	standing	stand	VERB
5999n298v5h	9	13	genetic	genetic	ADJ
5999n298v5h	9	14	variation	variation	NOUN
5999n298v5h	9	15	,	,	PUNCT
5999n298v5h	9	16	enabling	enable	VERB
5999n298v5h	9	17	the	the	DET
5999n298v5h	9	18	rapid	rapid	ADJ
5999n298v5h	9	19	evolution	evolution	NOUN
5999n298v5h	9	20	of	of	ADP
5999n298v5h	9	21	the	the	DET
5999n298v5h	9	22	apple	apple	NOUN
5999n298v5h	9	23	race	race	NOUN
5999n298v5h	9	24	of	of	ADP
5999n298v5h	9	25	r.	r.	PROPN
5999n298v5h	9	26	pomonella	pomonella	PROPN
5999n298v5h	9	27	.	.	PUNCT
5999n298v5h	10	1	comparisons	comparison	NOUN
5999n298v5h	10	2	of	of	ADP
5999n298v5h	10	3	the	the	DET
5999n298v5h	10	4	flowering	flower	VERB
5999n298v5h	10	5	dogwood	dogwood	NOUN
5999n298v5h	10	6	fly	fly	VERB
5999n298v5h	10	7	to	to	ADP
5999n298v5h	10	8	the	the	DET
5999n298v5h	10	9	hawthorn	hawthorn	NOUN
5999n298v5h	10	10	and	and	CCONJ
5999n298v5h	10	11	apple	apple	NOUN
5999n298v5h	10	12	races	race	NOUN
5999n298v5h	10	13	of	of	ADP
5999n298v5h	10	14	r.	r.	PROPN
5999n298v5h	10	15	pomonella	pomonella	PROPN
5999n298v5h	10	16	revealed	reveal	VERB
5999n298v5h	10	17	that	that	SCONJ
5999n298v5h	10	18	species	specie	NOUN
5999n298v5h	10	19	-	-	PUNCT
5999n298v5h	10	20	level	level	NOUN
5999n298v5h	10	21	differentiation	differentiation	NOUN
5999n298v5h	10	22	was	be	AUX
5999n298v5h	10	23	accompanied	accompany	VERB
5999n298v5h	10	24	by	by	ADP
5999n298v5h	10	25	the	the	DET
5999n298v5h	10	26	formation	formation	NOUN
5999n298v5h	10	27	of	of	ADP
5999n298v5h	10	28	range	range	NOUN
5999n298v5h	10	29	-	-	PUNCT
5999n298v5h	10	30	wide	wide	ADJ
5999n298v5h	10	31	genetic	genetic	ADJ
5999n298v5h	10	32	clustering	clustering	NOUN
5999n298v5h	10	33	based	base	VERB
5999n298v5h	10	34	on	on	ADP
5999n298v5h	10	35	allele	allele	NOUN
5999n298v5h	10	36	frequency	frequency	NOUN
5999n298v5h	10	37	differences	difference	NOUN
5999n298v5h	10	38	.	.	PUNCT
5999n298v5h	11	1	i	i	PRON
5999n298v5h	11	2	observed	observe	VERB
5999n298v5h	11	3	a	a	DET
5999n298v5h	11	4	striking	striking	ADJ
5999n298v5h	11	5	correlation	correlation	NOUN
5999n298v5h	11	6	in	in	ADP
5999n298v5h	11	7	the	the	DET
5999n298v5h	11	8	topology	topology	NOUN
5999n298v5h	11	9	of	of	ADP
5999n298v5h	11	10	differentiation	differentiation	NOUN
5999n298v5h	11	11	between	between	ADP
5999n298v5h	11	12	the	the	DET
5999n298v5h	11	13	host	host	NOUN
5999n298v5h	11	14	race	race	NOUN
5999n298v5h	11	15	and	and	CCONJ
5999n298v5h	11	16	species	specie	NOUN
5999n298v5h	11	17	,	,	PUNCT
5999n298v5h	11	18	consistent	consistent	ADJ
5999n298v5h	11	19	with	with	ADP
5999n298v5h	11	20	a	a	DET
5999n298v5h	11	21	genome	genome	NOUN
5999n298v5h	11	22	-	-	PUNCT
5999n298v5h	11	23	wide	wide	ADJ
5999n298v5h	11	24	reduction	reduction	NOUN
5999n298v5h	11	25	in	in	ADP
5999n298v5h	11	26	effective	effective	ADJ
5999n298v5h	11	27	migration	migration	NOUN
5999n298v5h	11	28	.	.	PUNCT
5999n298v5h	12	1	these	these	DET
5999n298v5h	12	2	findings	finding	NOUN
5999n298v5h	12	3	support	support	VERB
5999n298v5h	12	4	the	the	DET
5999n298v5h	12	5	following	follow	VERB
5999n298v5h	12	6	conclusions	conclusion	NOUN
5999n298v5h	12	7	regarding	regard	VERB
5999n298v5h	12	8	the	the	DET
5999n298v5h	12	9	five	five	NUM
5999n298v5h	12	10	questions	question	NOUN
5999n298v5h	12	11	:	:	PUNCT
5999n298v5h	12	12	(	(	PUNCT
5999n298v5h	12	13	1	1	X
5999n298v5h	12	14	)	)	PUNCT
5999n298v5h	12	15	sympatrically	sympatrically	ADV
5999n298v5h	12	16	-	-	PUNCT
5999n298v5h	12	17	formed	form	VERB
5999n298v5h	12	18	species	specie	NOUN
5999n298v5h	12	19	can	can	AUX
5999n298v5h	12	20	represent	represent	VERB
5999n298v5h	12	21	host	host	NOUN
5999n298v5h	12	22	races	race	NOUN
5999n298v5h	12	23	writ	writ	NOUN
5999n298v5h	12	24	-	-	PUNCT
5999n298v5h	12	25	large	large	ADJ
5999n298v5h	12	26	;	;	PUNCT
5999n298v5h	12	27	(	(	PUNCT
5999n298v5h	12	28	2	2	X
5999n298v5h	12	29	)	)	PUNCT
5999n298v5h	12	30	the	the	DET
5999n298v5h	12	31	initiation	initiation	NOUN
5999n298v5h	12	32	of	of	ADP
5999n298v5h	12	33	ecological	ecological	ADJ
5999n298v5h	12	34	speciation	speciation	NOUN
5999n298v5h	12	35	may	may	AUX
5999n298v5h	12	36	proceed	proceed	VERB
5999n298v5h	12	37	readily	readily	ADV
5999n298v5h	12	38	in	in	ADP
5999n298v5h	12	39	rhagoletis	rhagoletis	NOUN
5999n298v5h	12	40	;	;	PUNCT
5999n298v5h	12	41	(	(	PUNCT
5999n298v5h	12	42	3	3	X
5999n298v5h	12	43	)	)	PUNCT
5999n298v5h	12	44	both	both	PRON
5999n298v5h	12	45	olfactory	olfactory	NOUN
5999n298v5h	12	46	and	and	CCONJ
5999n298v5h	12	47	diapause	diapause	NOUN
5999n298v5h	12	48	traits	trait	NOUN
5999n298v5h	12	49	are	be	AUX
5999n298v5h	12	50	important	important	ADJ
5999n298v5h	12	51	to	to	ADP
5999n298v5h	12	52	ecological	ecological	ADJ
5999n298v5h	12	53	adaptation	adaptation	NOUN
5999n298v5h	12	54	and	and	CCONJ
5999n298v5h	12	55	reproductive	reproductive	ADJ
5999n298v5h	12	56	isolation	isolation	NOUN
5999n298v5h	12	57	in	in	ADP
5999n298v5h	12	58	rhagoletis	rhagoletis	NOUN
5999n298v5h	12	59	,	,	PUNCT
5999n298v5h	12	60	but	but	CCONJ
5999n298v5h	12	61	there	there	PRON
5999n298v5h	12	62	are	be	VERB
5999n298v5h	12	63	limits	limit	NOUN
5999n298v5h	12	64	to	to	ADP
5999n298v5h	12	65	their	their	PRON
5999n298v5h	12	66	role	role	NOUN
5999n298v5h	12	67	in	in	ADP
5999n298v5h	12	68	diversification	diversification	NOUN
5999n298v5h	12	69	;	;	PUNCT
5999n298v5h	12	70	(	(	PUNCT
5999n298v5h	12	71	4	4	X
5999n298v5h	12	72	)	)	PUNCT
5999n298v5h	12	73	standing	stand	VERB
5999n298v5h	12	74	genetic	genetic	ADJ
5999n298v5h	12	75	variation	variation	NOUN
5999n298v5h	12	76	,	,	PUNCT
5999n298v5h	12	77	in	in	ADP
5999n298v5h	12	78	the	the	DET
5999n298v5h	12	79	form	form	NOUN
5999n298v5h	12	80	of	of	ADP
5999n298v5h	12	81	heterogeneous	heterogeneous	ADJ
5999n298v5h	12	82	resource	resource	NOUN
5999n298v5h	12	83	use	use	NOUN
5999n298v5h	12	84	in	in	ADP
5999n298v5h	12	85	allopatric	allopatric	ADJ
5999n298v5h	12	86	populations	population	NOUN
5999n298v5h	12	87	and	and	CCONJ
5999n298v5h	12	88	latitudinal	latitudinal	ADJ
5999n298v5h	12	89	clines	cline	NOUN
5999n298v5h	12	90	,	,	PUNCT
5999n298v5h	12	91	has	have	AUX
5999n298v5h	12	92	been	be	AUX
5999n298v5h	12	93	a	a	DET
5999n298v5h	12	94	major	major	ADJ
5999n298v5h	12	95	factor	factor	NOUN
5999n298v5h	12	96	in	in	ADP
5999n298v5h	12	97	the	the	DET
5999n298v5h	12	98	radiation	radiation	NOUN
5999n298v5h	12	99	of	of	ADP
5999n298v5h	12	100	the	the	DET
5999n298v5h	12	101	r.	r.	PROPN
5999n298v5h	12	102	pomonella	pomonella	PROPN
5999n298v5h	12	103	species	species	PROPN
5999n298v5h	12	104	complex	complex	NOUN
5999n298v5h	12	105	;	;	PUNCT
5999n298v5h	12	106	and	and	CCONJ
5999n298v5h	12	107	(	(	PUNCT
5999n298v5h	12	108	5	5	X
5999n298v5h	12	109	)	)	PUNCT
5999n298v5h	12	110	the	the	DET
5999n298v5h	12	111	transition	transition	NOUN
5999n298v5h	12	112	from	from	ADP
5999n298v5h	12	113	host	host	NOUN
5999n298v5h	12	114	race	race	NOUN
5999n298v5h	12	115	to	to	ADP
5999n298v5h	12	116	species	specie	NOUN
5999n298v5h	12	117	appears	appear	VERB
5999n298v5h	12	118	to	to	PART
5999n298v5h	12	119	involve	involve	VERB
5999n298v5h	12	120	the	the	DET
5999n298v5h	12	121	uplift	uplift	NOUN
5999n298v5h	12	122	of	of	ADP
5999n298v5h	12	123	genomic	genomic	ADJ
5999n298v5h	12	124	'	'	PUNCT
5999n298v5h	12	125	continents	continent	NOUN
5999n298v5h	12	126	of	of	ADP
5999n298v5h	12	127	speciation	speciation	NOUN
5999n298v5h	12	128	'	'	PUNCT
5999n298v5h	12	129	driven	drive	VERB
5999n298v5h	12	130	global	global	ADJ
5999n298v5h	12	131	reductions	reduction	NOUN
5999n298v5h	12	132	in	in	ADP
5999n298v5h	12	133	effective	effective	ADJ
5999n298v5h	12	134	migration	migration	NOUN
5999n298v5h	12	135	.	.	PUNCT