id sid tid token lemma pos 4x51hh65r1b 1 1 swarming swarm VERB 4x51hh65r1b 1 2 is be AUX 4x51hh65r1b 1 3 a a DET 4x51hh65r1b 1 4 surface surface NOUN 4x51hh65r1b 1 5 - - PUNCT 4x51hh65r1b 1 6 associated associate VERB 4x51hh65r1b 1 7 motility motility NOUN 4x51hh65r1b 1 8 where where SCONJ 4x51hh65r1b 1 9 bacteria bacteria NOUN 4x51hh65r1b 1 10 spread spread VERB 4x51hh65r1b 1 11 in in ADP 4x51hh65r1b 1 12 groups group NOUN 4x51hh65r1b 1 13 , , PUNCT 4x51hh65r1b 1 14 often often ADV 4x51hh65r1b 1 15 as as ADP 4x51hh65r1b 1 16 a a DET 4x51hh65r1b 1 17 precursor precursor NOUN 4x51hh65r1b 1 18 to to ADP 4x51hh65r1b 1 19 biofilm biofilm NOUN 4x51hh65r1b 1 20 formation formation NOUN 4x51hh65r1b 1 21 . . PUNCT 4x51hh65r1b 2 1 the the DET 4x51hh65r1b 2 2 bacterium bacterium NOUN 4x51hh65r1b 2 3 pseudomonas pseudomonas PROPN 4x51hh65r1b 2 4 aeruginosa aeruginosa PROPN 4x51hh65r1b 2 5 is be AUX 4x51hh65r1b 2 6 known know VERB 4x51hh65r1b 2 7 to to PART 4x51hh65r1b 2 8 swarm swarm VERB 4x51hh65r1b 2 9 using use VERB 4x51hh65r1b 2 10 its its PRON 4x51hh65r1b 2 11 polar polar ADJ 4x51hh65r1b 2 12 flagellum flagellum NOUN 4x51hh65r1b 2 13 and and CCONJ 4x51hh65r1b 2 14 a a DET 4x51hh65r1b 2 15 self self NOUN 4x51hh65r1b 2 16 - - PUNCT 4x51hh65r1b 2 17 produced produce VERB 4x51hh65r1b 2 18 surfactant surfactant NOUN 4x51hh65r1b 2 19 called call VERB 4x51hh65r1b 2 20 rhamnolipid rhamnolipid NOUN 4x51hh65r1b 2 21 . . PUNCT 4x51hh65r1b 3 1 this this DET 4x51hh65r1b 3 2 thesis thesis NOUN 4x51hh65r1b 3 3 details detail NOUN 4x51hh65r1b 3 4 swarming swarm VERB 4x51hh65r1b 3 5 in in ADP 4x51hh65r1b 3 6 response response NOUN 4x51hh65r1b 3 7 to to ADP 4x51hh65r1b 3 8 two two NUM 4x51hh65r1b 3 9 types type NOUN 4x51hh65r1b 3 10 of of ADP 4x51hh65r1b 3 11 environmental environmental ADJ 4x51hh65r1b 3 12 conditions condition NOUN 4x51hh65r1b 3 13 : : PUNCT 4x51hh65r1b 3 14 surface surface VERB 4x51hh65r1b 3 15 hardness hardness NOUN 4x51hh65r1b 3 16 and and CCONJ 4x51hh65r1b 3 17 carbon carbon NOUN 4x51hh65r1b 3 18 source source NOUN 4x51hh65r1b 3 19 . . PUNCT 4x51hh65r1b 4 1 it it PRON 4x51hh65r1b 4 2 was be AUX 4x51hh65r1b 4 3 found find VERB 4x51hh65r1b 4 4 that that SCONJ 4x51hh65r1b 4 5 swarm swarm NOUN 4x51hh65r1b 4 6 zone zone NOUN 4x51hh65r1b 4 7 expansion expansion NOUN 4x51hh65r1b 4 8 and and CCONJ 4x51hh65r1b 4 9 bacterial bacterial ADJ 4x51hh65r1b 4 10 population population NOUN 4x51hh65r1b 4 11 are be AUX 4x51hh65r1b 4 12 not not PART 4x51hh65r1b 4 13 linearly linearly ADV 4x51hh65r1b 4 14 correlated correlate VERB 4x51hh65r1b 4 15 during during ADP 4x51hh65r1b 4 16 swarming swarm VERB 4x51hh65r1b 4 17 and and CCONJ 4x51hh65r1b 4 18 further far ADV 4x51hh65r1b 4 19 that that PRON 4x51hh65r1b 4 20 increased increase VERB 4x51hh65r1b 4 21 surface surface NOUN 4x51hh65r1b 4 22 hardness hardness NOUN 4x51hh65r1b 4 23 significantly significantly ADV 4x51hh65r1b 4 24 decreases decrease VERB 4x51hh65r1b 4 25 rhamnolipid rhamnolipid NOUN 4x51hh65r1b 4 26 - - PUNCT 4x51hh65r1b 4 27 dependent dependent ADJ 4x51hh65r1b 4 28 swarming swarming NOUN 4x51hh65r1b 4 29 . . PUNCT 4x51hh65r1b 5 1 secondly secondly ADV 4x51hh65r1b 5 2 , , PUNCT 4x51hh65r1b 5 3 it it PRON 4x51hh65r1b 5 4 was be AUX 4x51hh65r1b 5 5 found find VERB 4x51hh65r1b 5 6 that that SCONJ 4x51hh65r1b 5 7 a a DET 4x51hh65r1b 5 8 rhamnolipid rhamnolipid NOUN 4x51hh65r1b 5 9 - - PUNCT 4x51hh65r1b 5 10 deficient deficient ADJ 4x51hh65r1b 5 11 p. p. NOUN 4x51hh65r1b 5 12 aeruginosa aeruginosa PROPN 4x51hh65r1b 5 13 strain strain NOUN 4x51hh65r1b 5 14 swarmed swarm VERB 4x51hh65r1b 5 15 upon upon SCONJ 4x51hh65r1b 5 16 the the DET 4x51hh65r1b 5 17 addition addition NOUN 4x51hh65r1b 5 18 of of ADP 4x51hh65r1b 5 19 the the DET 4x51hh65r1b 5 20 carbon carbon NOUN 4x51hh65r1b 5 21 source source NOUN 4x51hh65r1b 5 22 glutamate glutamate NOUN 4x51hh65r1b 5 23 . . PUNCT 4x51hh65r1b 6 1 this this PRON 4x51hh65r1b 6 2 is be AUX 4x51hh65r1b 6 3 unexpected unexpected ADJ 4x51hh65r1b 6 4 , , PUNCT 4x51hh65r1b 6 5 as as SCONJ 4x51hh65r1b 6 6 no no DET 4x51hh65r1b 6 7 previous previous ADJ 4x51hh65r1b 6 8 rhamnolipid rhamnolipid NOUN 4x51hh65r1b 6 9 - - PUNCT 4x51hh65r1b 6 10 deficient deficient ADJ 4x51hh65r1b 6 11 swarming swarming NOUN 4x51hh65r1b 6 12 has have AUX 4x51hh65r1b 6 13 been be AUX 4x51hh65r1b 6 14 documented document VERB 4x51hh65r1b 6 15 for for ADP 4x51hh65r1b 6 16 p. p. PROPN 4x51hh65r1b 6 17 aerguinosa aerguinosa PROPN 4x51hh65r1b 6 18 . . PUNCT 4x51hh65r1b 7 1 a a DET 4x51hh65r1b 7 2 mutagenesis mutagenesis NOUN 4x51hh65r1b 7 3 approach approach NOUN 4x51hh65r1b 7 4 was be AUX 4x51hh65r1b 7 5 used use VERB 4x51hh65r1b 7 6 to to PART 4x51hh65r1b 7 7 find find VERB 4x51hh65r1b 7 8 gene(s gene(s PROPN 4x51hh65r1b 7 9 ) ) PUNCT 4x51hh65r1b 7 10 responsible responsible ADJ 4x51hh65r1b 7 11 for for ADP 4x51hh65r1b 7 12 this this DET 4x51hh65r1b 7 13 rhamnolipid rhamnolipid ADJ 4x51hh65r1b 7 14 - - PUNCT 4x51hh65r1b 7 15 independent independent ADJ 4x51hh65r1b 7 16 swarm swarm NOUN 4x51hh65r1b 7 17 phenotype phenotype NOUN 4x51hh65r1b 7 18 by by ADP 4x51hh65r1b 7 19 probing probe VERB 4x51hh65r1b 7 20 for for ADP 4x51hh65r1b 7 21 mutants mutant NOUN 4x51hh65r1b 7 22 that that PRON 4x51hh65r1b 7 23 negate negate VERB 4x51hh65r1b 7 24 the the DET 4x51hh65r1b 7 25 glutamate glutamate NOUN 4x51hh65r1b 7 26 swarming swarm VERB 4x51hh65r1b 7 27 effect effect NOUN 4x51hh65r1b 7 28 . . PUNCT 4x51hh65r1b 8 1 two two NUM 4x51hh65r1b 8 2 thousand thousand NUM 4x51hh65r1b 8 3 mutants mutant NOUN 4x51hh65r1b 8 4 were be AUX 4x51hh65r1b 8 5 screened screen VERB 4x51hh65r1b 8 6 , , PUNCT 4x51hh65r1b 8 7 twelve twelve NUM 4x51hh65r1b 8 8 of of ADP 4x51hh65r1b 8 9 which which PRON 4x51hh65r1b 8 10 were be AUX 4x51hh65r1b 8 11 found find VERB 4x51hh65r1b 8 12 to to PART 4x51hh65r1b 8 13 not not PART 4x51hh65r1b 8 14 swarm swarm VERB 4x51hh65r1b 8 15 with with ADP 4x51hh65r1b 8 16 glutamate glutamate NOUN 4x51hh65r1b 8 17 . . PUNCT