id	sid	tid	token	lemma	pos
00000001r5t	1	1	almost	almost	ADV
00000001r5t	1	2	all	all	PRON
00000001r5t	1	3	living	living	NOUN
00000001r5t	1	4	organisms	organism	NOUN
00000001r5t	1	5	require	require	VERB
00000001r5t	1	6	iron	iron	NOUN
00000001r5t	1	7	(	(	PUNCT
00000001r5t	1	8	fe	fe	PROPN
00000001r5t	1	9	)	)	PUNCT
00000001r5t	1	10	to	to	PART
00000001r5t	1	11	live	live	VERB
00000001r5t	1	12	.	.	PUNCT
00000001r5t	2	1	although	although	SCONJ
00000001r5t	2	2	iron	iron	NOUN
00000001r5t	2	3	is	be	AUX
00000001r5t	2	4	abundant	abundant	ADJ
00000001r5t	2	5	on	on	ADP
00000001r5t	2	6	earth	earth	NOUN
00000001r5t	2	7	,	,	PUNCT
00000001r5t	2	8	aerobic	aerobic	ADJ
00000001r5t	2	9	conditions	condition	NOUN
00000001r5t	2	10	and	and	CCONJ
00000001r5t	2	11	circumneutral	circumneutral	ADJ
00000001r5t	2	12	ph	ph	NOUN
00000001r5t	2	13	restrict	restrict	VERB
00000001r5t	2	14	it	it	PRON
00000001r5t	2	15	to	to	ADP
00000001r5t	2	16	the	the	DET
00000001r5t	2	17	3	3	NUM
00000001r5t	2	18	+	+	NUM
00000001r5t	2	19	oxidation	oxidation	NOUN
00000001r5t	2	20	state	state	NOUN
00000001r5t	2	21	,	,	PUNCT
00000001r5t	2	22	in	in	ADP
00000001r5t	2	23	which	which	PRON
00000001r5t	2	24	it	it	PRON
00000001r5t	2	25	is	be	AUX
00000001r5t	2	26	bound	bind	VERB
00000001r5t	2	27	up	up	ADP
00000001r5t	2	28	as	as	ADP
00000001r5t	2	29	insoluble	insoluble	ADJ
00000001r5t	2	30	fe(iii	fe(iii	NOUN
00000001r5t	2	31	)	)	PUNCT
00000001r5t	2	32	(	(	PUNCT
00000001r5t	2	33	hydr)oxide	hydr)oxide	PROPN
00000001r5t	2	34	minerals	mineral	NOUN
00000001r5t	2	35	(	(	PUNCT
00000001r5t	2	36	i.e.	i.e.	X
00000001r5t	2	37	hematite	hematite	PROPN
00000001r5t	2	38	,	,	PUNCT
00000001r5t	2	39	goethite	goethite	NOUN
00000001r5t	2	40	)	)	PUNCT
00000001r5t	2	41	,	,	PUNCT
00000001r5t	2	42	or	or	CCONJ
00000001r5t	2	43	,	,	PUNCT
00000001r5t	2	44	in	in	ADP
00000001r5t	2	45	a	a	DET
00000001r5t	2	46	human	human	ADJ
00000001r5t	2	47	host	host	NOUN
00000001r5t	2	48	,	,	PUNCT
00000001r5t	2	49	tightly	tightly	ADV
00000001r5t	2	50	sequestered	sequester	VERB
00000001r5t	2	51	by	by	ADP
00000001r5t	2	52	proteins	protein	NOUN
00000001r5t	2	53	such	such	ADJ
00000001r5t	2	54	as	as	ADP
00000001r5t	2	55	transferrin	transferrin	NOUN
00000001r5t	2	56	and	and	CCONJ
00000001r5t	2	57	ferritin	ferritin	NOUN
00000001r5t	2	58	.	.	PUNCT
00000001r5t	3	1	hence	hence	ADV
00000001r5t	3	2	,	,	PUNCT
00000001r5t	3	3	although	although	SCONJ
00000001r5t	3	4	a	a	DET
00000001r5t	3	5	typical	typical	ADJ
00000001r5t	3	6	cell	cell	NOUN
00000001r5t	3	7	needs	need	VERB
00000001r5t	3	8	a	a	DET
00000001r5t	3	9	concentration	concentration	NOUN
00000001r5t	3	10	of	of	ADP
00000001r5t	3	11	~10	~10	PROPN
00000001r5t	3	12	-	-	SYM
00000001r5t	3	13	6	6	NUM
00000001r5t	3	14	m	m	NOUN
00000001r5t	3	15	fe	fe	ADJ
00000001r5t	3	16	to	to	PART
00000001r5t	3	17	survive	survive	VERB
00000001r5t	3	18	,	,	PUNCT
00000001r5t	3	19	readily	readily	ADV
00000001r5t	3	20	available	available	ADJ
00000001r5t	3	21	,	,	PUNCT
00000001r5t	3	22	soluble	soluble	ADJ
00000001r5t	3	23	fe	fe	PROPN
00000001r5t	3	24	is	be	AUX
00000001r5t	3	25	only	only	ADV
00000001r5t	3	26	~10	~10	NUM
00000001r5t	3	27	-	-	SYM
00000001r5t	3	28	18	18	NUM
00000001r5t	3	29	m.	m.	NOUN
00000001r5t	3	30	microbes	microbe	NOUN
00000001r5t	3	31	have	have	AUX
00000001r5t	3	32	responded	respond	VERB
00000001r5t	3	33	to	to	ADP
00000001r5t	3	34	this	this	DET
00000001r5t	3	35	enormous	enormous	ADJ
00000001r5t	3	36	discrepancy	discrepancy	NOUN
00000001r5t	3	37	by	by	ADP
00000001r5t	3	38	evolving	evolve	VERB
00000001r5t	3	39	very	very	ADV
00000001r5t	3	40	efficient	efficient	ADJ
00000001r5t	3	41	strategies	strategy	NOUN
00000001r5t	3	42	for	for	ADP
00000001r5t	3	43	iron	iron	NOUN
00000001r5t	3	44	acquisition	acquisition	NOUN
00000001r5t	3	45	,	,	PUNCT
00000001r5t	3	46	primarily	primarily	ADV
00000001r5t	3	47	the	the	DET
00000001r5t	3	48	production	production	NOUN
00000001r5t	3	49	and	and	CCONJ
00000001r5t	3	50	secretion	secretion	NOUN
00000001r5t	3	51	of	of	ADP
00000001r5t	3	52	fe	fe	PROPN
00000001r5t	3	53	-	-	PUNCT
00000001r5t	3	54	chelating	chelate	VERB
00000001r5t	3	55	siderophores	siderophore	NOUN
00000001r5t	3	56	.	.	PUNCT
00000001r5t	4	1	however	however	ADV
00000001r5t	4	2	,	,	PUNCT
00000001r5t	4	3	the	the	DET
00000001r5t	4	4	long	long	ADV
00000001r5t	4	5	-	-	PUNCT
00000001r5t	4	6	held	hold	VERB
00000001r5t	4	7	assumption	assumption	NOUN
00000001r5t	4	8	that	that	SCONJ
00000001r5t	4	9	siderophores	siderophore	VERB
00000001r5t	4	10	alone	alone	ADV
00000001r5t	4	11	deliver	deliver	VERB
00000001r5t	4	12	mineral	mineral	NOUN
00000001r5t	4	13	fe	fe	PROPN
00000001r5t	4	14	to	to	ADP
00000001r5t	4	15	microbes	microbe	NOUN
00000001r5t	4	16	at	at	ADP
00000001r5t	4	17	a	a	DET
00000001r5t	4	18	rate	rate	NOUN
00000001r5t	4	19	sufficient	sufficient	ADJ
00000001r5t	4	20	to	to	PART
00000001r5t	4	21	support	support	VERB
00000001r5t	4	22	microbial	microbial	ADJ
00000001r5t	4	23	growth	growth	NOUN
00000001r5t	4	24	has	have	AUX
00000001r5t	4	25	recently	recently	ADV
00000001r5t	4	26	been	be	AUX
00000001r5t	4	27	called	call	VERB
00000001r5t	4	28	into	into	ADP
00000001r5t	4	29	question	question	NOUN
00000001r5t	4	30	by	by	ADP
00000001r5t	4	31	abiotic	abiotic	ADJ
00000001r5t	4	32	measurements	measurement	NOUN
00000001r5t	4	33	of	of	ADP
00000001r5t	4	34	siderophore	siderophore	NOUN
00000001r5t	4	35	-	-	PUNCT
00000001r5t	4	36	promoted	promote	VERB
00000001r5t	4	37	dissolution	dissolution	NOUN
00000001r5t	4	38	.	.	PUNCT
00000001r5t	5	1	this	this	DET
00000001r5t	5	2	dissertation	dissertation	NOUN
00000001r5t	5	3	presents	present	VERB
00000001r5t	5	4	an	an	DET
00000001r5t	5	5	approach	approach	NOUN
00000001r5t	5	6	and	and	CCONJ
00000001r5t	5	7	bioinformatic	bioinformatic	ADJ
00000001r5t	5	8	analyses	analysis	NOUN
00000001r5t	5	9	based	base	VERB
00000001r5t	5	10	on	on	ADP
00000001r5t	5	11	tools	tool	NOUN
00000001r5t	5	12	from	from	ADP
00000001r5t	5	13	molecular	molecular	ADJ
00000001r5t	5	14	biology	biology	NOUN
00000001r5t	5	15	,	,	PUNCT
00000001r5t	5	16	such	such	ADJ
00000001r5t	5	17	as	as	ADP
00000001r5t	5	18	knockout	knockout	NOUN
00000001r5t	5	19	mutant	mutant	NOUN
00000001r5t	5	20	and	and	CCONJ
00000001r5t	5	21	biosensor	biosensor	NOUN
00000001r5t	5	22	strains	strain	NOUN
00000001r5t	5	23	,	,	PUNCT
00000001r5t	5	24	in	in	ADP
00000001r5t	5	25	conjunction	conjunction	NOUN
00000001r5t	5	26	with	with	ADP
00000001r5t	5	27	classic	classic	ADJ
00000001r5t	5	28	biochemical	biochemical	ADJ
00000001r5t	5	29	and	and	CCONJ
00000001r5t	5	30	physiological	physiological	ADJ
00000001r5t	5	31	assays	assay	NOUN
00000001r5t	5	32	,	,	PUNCT
00000001r5t	5	33	to	to	PART
00000001r5t	5	34	investigate	investigate	VERB
00000001r5t	5	35	this	this	DET
00000001r5t	5	36	process	process	NOUN
00000001r5t	5	37	in	in	ADP
00000001r5t	5	38	a	a	DET
00000001r5t	5	39	living	live	VERB
00000001r5t	5	40	system	system	NOUN
00000001r5t	5	41	.	.	PUNCT
00000001r5t	6	1	the	the	DET
00000001r5t	6	2	versatility	versatility	NOUN
00000001r5t	6	3	of	of	ADP
00000001r5t	6	4	this	this	DET
00000001r5t	6	5	approach	approach	NOUN
00000001r5t	6	6	allowed	allow	VERB
00000001r5t	6	7	questions	question	NOUN
00000001r5t	6	8	in	in	ADP
00000001r5t	6	9	both	both	CCONJ
00000001r5t	6	10	environmental	environmental	ADJ
00000001r5t	6	11	and	and	CCONJ
00000001r5t	6	12	biomedical	biomedical	ADJ
00000001r5t	6	13	contexts	context	NOUN
00000001r5t	6	14	to	to	PART
00000001r5t	6	15	be	be	AUX
00000001r5t	6	16	addressed	address	VERB
00000001r5t	6	17	.	.	PUNCT
00000001r5t	7	1	in	in	ADP
00000001r5t	7	2	studies	study	NOUN
00000001r5t	7	3	with	with	ADP
00000001r5t	7	4	the	the	DET
00000001r5t	7	5	common	common	ADJ
00000001r5t	7	6	environmental	environmental	ADJ
00000001r5t	7	7	aerobe	aerobe	NOUN
00000001r5t	7	8	p.	p.	PROPN
00000001r5t	7	9	mendocina	mendocina	NOUN
00000001r5t	7	10	,	,	PUNCT
00000001r5t	7	11	it	it	PRON
00000001r5t	7	12	was	be	AUX
00000001r5t	7	13	found	find	VERB
00000001r5t	7	14	that	that	SCONJ
00000001r5t	7	15	the	the	DET
00000001r5t	7	16	contribution	contribution	NOUN
00000001r5t	7	17	of	of	ADP
00000001r5t	7	18	siderophores	siderophore	NOUN
00000001r5t	7	19	to	to	ADP
00000001r5t	7	20	fe	fe	PROPN
00000001r5t	7	21	acquisition	acquisition	NOUN
00000001r5t	7	22	from	from	ADP
00000001r5t	7	23	hematite	hematite	PROPN
00000001r5t	7	24	depended	depend	VERB
00000001r5t	7	25	strongly	strongly	ADV
00000001r5t	7	26	on	on	ADP
00000001r5t	7	27	the	the	DET
00000001r5t	7	28	presence	presence	NOUN
00000001r5t	7	29	and	and	CCONJ
00000001r5t	7	30	concentration	concentration	NOUN
00000001r5t	7	31	of	of	ADP
00000001r5t	7	32	other	other	ADJ
00000001r5t	7	33	organic	organic	ADJ
00000001r5t	7	34	ligands	ligand	NOUN
00000001r5t	7	35	common	common	ADJ
00000001r5t	7	36	in	in	ADP
00000001r5t	7	37	the	the	DET
00000001r5t	7	38	soil	soil	NOUN
00000001r5t	7	39	.	.	PUNCT
00000001r5t	8	1	mineral	mineral	ADJ
00000001r5t	8	2	particle	particle	NOUN
00000001r5t	8	3	size	size	NOUN
00000001r5t	8	4	was	be	AUX
00000001r5t	8	5	also	also	ADV
00000001r5t	8	6	shown	show	VERB
00000001r5t	8	7	to	to	PART
00000001r5t	8	8	be	be	AUX
00000001r5t	8	9	an	an	DET
00000001r5t	8	10	important	important	ADJ
00000001r5t	8	11	parameter	parameter	NOUN
00000001r5t	8	12	,	,	PUNCT
00000001r5t	8	13	as	as	SCONJ
00000001r5t	8	14	nanohematite	nanohematite	ADJ
00000001r5t	8	15	proved	prove	VERB
00000001r5t	8	16	to	to	PART
00000001r5t	8	17	be	be	AUX
00000001r5t	8	18	far	far	ADV
00000001r5t	8	19	more	more	ADV
00000001r5t	8	20	bioavailable	bioavailable	ADJ
00000001r5t	8	21	to	to	ADP
00000001r5t	8	22	p.	p.	PROPN
00000001r5t	8	23	mendocina	mendocina	PROPN
00000001r5t	8	24	,	,	PUNCT
00000001r5t	8	25	even	even	ADV
00000001r5t	8	26	supporting	support	VERB
00000001r5t	8	27	robust	robust	ADJ
00000001r5t	8	28	growth	growth	NOUN
00000001r5t	8	29	in	in	ADP
00000001r5t	8	30	the	the	DET
00000001r5t	8	31	absence	absence	NOUN
00000001r5t	8	32	of	of	ADP
00000001r5t	8	33	siderophores	siderophore	NOUN
00000001r5t	8	34	.	.	PUNCT
00000001r5t	9	1	of	of	ADP
00000001r5t	9	2	biomedical	biomedical	ADJ
00000001r5t	9	3	relevance	relevance	NOUN
00000001r5t	9	4	,	,	PUNCT
00000001r5t	9	5	ferritin	ferritin	PROPN
00000001r5t	9	6	(	(	PUNCT
00000001r5t	9	7	a	a	DET
00000001r5t	9	8	protein	protein	NOUN
00000001r5t	9	9	"	"	PUNCT
00000001r5t	9	10	cage	cage	NOUN
00000001r5t	9	11	"	"	PUNCT
00000001r5t	9	12	with	with	ADP
00000001r5t	9	13	a	a	DET
00000001r5t	9	14	core	core	NOUN
00000001r5t	9	15	of	of	ADP
00000001r5t	9	16	nanomineral	nanomineral	ADJ
00000001r5t	9	17	ferrihydrite	ferrihydrite	PROPN
00000001r5t	9	18	)	)	PUNCT
00000001r5t	9	19	was	be	AUX
00000001r5t	9	20	shown	show	VERB
00000001r5t	9	21	to	to	PART
00000001r5t	9	22	be	be	AUX
00000001r5t	9	23	an	an	DET
00000001r5t	9	24	excellent	excellent	ADJ
00000001r5t	9	25	source	source	NOUN
00000001r5t	9	26	of	of	ADP
00000001r5t	9	27	fe	fe	PROPN
00000001r5t	9	28	to	to	ADP
00000001r5t	9	29	the	the	DET
00000001r5t	9	30	pathogen	pathogen	PROPN
00000001r5t	10	1	p.	p.	PROPN
00000001r5t	10	2	aeruginosa	aeruginosa	PROPN
00000001r5t	10	3	regardless	regardless	ADV
00000001r5t	10	4	of	of	ADP
00000001r5t	10	5	the	the	DET
00000001r5t	10	6	presence	presence	NOUN
00000001r5t	10	7	of	of	ADP
00000001r5t	10	8	siderophores	siderophore	NOUN
00000001r5t	10	9	.	.	PUNCT
00000001r5t	11	1	although	although	SCONJ
00000001r5t	11	2	close	close	ADJ
00000001r5t	11	3	cell	cell	NOUN
00000001r5t	11	4	-	-	PUNCT
00000001r5t	11	5	mineral	mineral	NOUN
00000001r5t	11	6	interaction	interaction	NOUN
00000001r5t	11	7	was	be	AUX
00000001r5t	11	8	a	a	DET
00000001r5t	11	9	requirement	requirement	NOUN
00000001r5t	11	10	for	for	ADP
00000001r5t	11	11	siderophore	siderophore	NOUN
00000001r5t	11	12	-	-	PUNCT
00000001r5t	11	13	independent	independent	ADJ
00000001r5t	11	14	fe	fe	PROPN
00000001r5t	11	15	acquisition	acquisition	NOUN
00000001r5t	11	16	from	from	ADP
00000001r5t	11	17	nanohematite	nanohematite	NOUN
00000001r5t	11	18	by	by	ADP
00000001r5t	11	19	p.	p.	PROPN
00000001r5t	11	20	mendocina	mendocina	PROPN
00000001r5t	11	21	,	,	PUNCT
00000001r5t	11	22	this	this	PRON
00000001r5t	11	23	was	be	AUX
00000001r5t	11	24	not	not	PART
00000001r5t	11	25	observed	observe	VERB
00000001r5t	11	26	for	for	ADP
00000001r5t	11	27	ferritin	ferritin	NOUN
00000001r5t	11	28	and	and	CCONJ
00000001r5t	11	29	p.	p.	PROPN
00000001r5t	11	30	aeruginosa	aeruginosa	PROPN
00000001r5t	11	31	.	.	PUNCT
00000001r5t	12	1	other	other	ADJ
00000001r5t	12	2	possible	possible	ADJ
00000001r5t	12	3	mechanisms	mechanism	NOUN
00000001r5t	12	4	include	include	VERB
00000001r5t	12	5	secreted	secrete	VERB
00000001r5t	12	6	proteases	protease	NOUN
00000001r5t	12	7	and	and	CCONJ
00000001r5t	12	8	reductants	reductant	NOUN
00000001r5t	12	9	.	.	PUNCT
00000001r5t	13	1	overall	overall	ADV
00000001r5t	13	2	,	,	PUNCT
00000001r5t	13	3	this	this	DET
00000001r5t	13	4	work	work	NOUN
00000001r5t	13	5	shows	show	VERB
00000001r5t	13	6	that	that	SCONJ
00000001r5t	13	7	microbial	microbial	ADJ
00000001r5t	13	8	acquisition	acquisition	NOUN
00000001r5t	13	9	of	of	ADP
00000001r5t	13	10	fe	fe	PROPN
00000001r5t	13	11	from	from	ADP
00000001r5t	13	12	mineral	mineral	NOUN
00000001r5t	13	13	sources	source	NOUN
00000001r5t	13	14	is	be	AUX
00000001r5t	13	15	complex	complex	ADJ
00000001r5t	13	16	,	,	PUNCT
00000001r5t	13	17	and	and	CCONJ
00000001r5t	13	18	reveals	reveal	VERB
00000001r5t	13	19	a	a	DET
00000001r5t	13	20	variable	variable	ADJ
00000001r5t	13	21	role	role	NOUN
00000001r5t	13	22	for	for	ADP
00000001r5t	13	23	siderophores	siderophore	NOUN
00000001r5t	13	24	.	.	PUNCT