* H3. IS ENCYCLOPAEDIA BRITANNIC A SEVENTH EDITION. ENCYCLOPAEDIA BRITANNICA OR DICTIONARY OF ARTS, SCIENCES, AND GENERAL LITERATURE. SEVENTH EDITION, WITH PRELIMINARY DISSERTATIONS ON THE HISTORY OF THE SCIENCES, AND OTHER EXTENSIVE IMPROVEMENTS AND ADDITIONS; INCLUDING THE LATE SUPPLEMENT, A GENERAL INDEX, AND NUMEROUS ENGRAVINGS. VOLUME III. ADAM AND CHARLES BLACK, EDINBURGH; M.DCCC.XLII. ENCYCLOPAEDIA BRITANNICA COMPARATIVE ANATOMY. PART I. ANATOMY OF THE ORGANS OF RELATION. CHAP. I.—COMPARATIVE OSTEOLOGY. Compara- X) ED-BLOODED ANIMALS only can be said to pos- tive A*' sess that assemblage of bones denominated skeleton; and as in these the most constant part is the vertebral ' column, it furnishes the general character of Vertebrated. The shells of the Mollusca and the Crustacea have been generally regarded as a species of internal skeleton ; and in the circumstance of affording mechanical support and external protection, they indeed resemble the skeleton of the Vertebrata. But neither in mode of developement nor in chemical constitution can they be regarded as of the same nature. Hence it is only in the vertebrated classes that it is requisite to study the peculiarities of the skeleton. Bones ge- In general characters the bones of the Mammalia re- nerally. semble those of the human subject. Like them, they are white, firm, elastic, and incompressible. They consist also of compact and reticular or cancellated tissue. In the lower animals the latter kind of structure is in gene¬ ral coarser and looser than in man; and in the Cetacea especially the cavities are large. In the carnivorous ani¬ mals the compact structure is exceedingly dense, and gives the bone much greater weight than in other animals. In the Cetacea also the acoustic or lithoid portion of the temporal bone is of a marble hardness. The bones of the Mammalia may, like those of man, be distinguished, according to their mechanical form, into long, flat, and short bones. Though the first class in ge¬ neral possess a medullary canal, this cavity is imperfect or wholly wanting in the bones of the Cetacea and Am¬ phibia. I he cavities denominated sinuses are much more com¬ pletely developed in several of the Mammalia than in the human skeleton. In the pig these cavities extend into the occipital bone ; in the elephant they not only give the frontal bone extraordinary protuberance, but they extend into the parietal, temporal, and even the occipital bones, and contribute much to augment the volume of the head. In the ox, deer, and sheep, they communicate with the cavity of the horns. The bones of Birds are in general whiter, firmer, and smoother than those of the Mammalia ; and they con- sist of a firm, compact substance, which is elastic and hard VOL. in. in the bones of the trunk, and extremely brittle in those Compara- of the extremities. With the exception also of some of ^ve the thin, flat bones, as the sternum, they consist of thin, ,^iatom.v- compact walls, inclosing large capacious cavities, which contain not marrow, but air, and which communicate by one or more minute holes with the windpipe and lungs. While these cavities, which may be regarded as the most perfect and advanced form of sinuses, diminish consider- ably the weight of the whole skeleton, by the density and completely cylindrical shape, they rather augment the strength. In the chick, and at birth, the bones are ho¬ mogeneous and without cavities; afterwards they contain marrow; and eventually this disappears and gives place to air. The bones of the Reptiles are not remarkable in any respect, unless in being void in general of medullary ca- vity. dhe absence of this canal was originally observed by Caldesi, and afterwards by Cuvier, in the tortoise; by d roja in the bones of the frog and toad; and by Carus in those of the turtle. In the crocodile, however, and in several of the lizard family, they are large and distinct. The bones of Reptiles also are less firm than those of Birds and Mammals. The bones of Fishes are remarkable for great softness, flexibility, and elasticity. Those of the lamprey, shark, ray or skate, and sturgeon family, are soft, flexible, sec- tile, of a bluish white colour, translucent, and so elastic that a cutting instrument forced into them is speedily re- truded by the resilient nature of the bony matter. From these characters, the bones of these families have been regarded as cartilaginous, and the fishes themselves have been distinguished by this character. (Pisces cartila- ginei, PISCES chondropterygii.) In the other fishes, the bones, though softer than those of Mammals, Birds, and Reptiles, present a greater degree of firmness and so¬ lidity, are whiter and more opaque, and are much less sectile, than those of the cartilaginous division. As in this respect, therefore, they approach the genuine bone of the Mammals, these are distinguished as fishes with osseous skeletons. (Pisces ossei.) The bones of both classes of fishes consist of a large quantity of gelatine, with a small proportion of phosphate A ANATOMY. In the osseous fishes, however, the latter sub- Compara- oflime. tive stance is most abundant. • Anatomy. colour of the bones of fishes, though in general whitish gray, is observed to vary in certain genera. In the crar-pike (esox belone), for instance, they aie green, , . • • i-i /A/vi/»*/»/c* 'ii'iw'nr/'r'i/s). tnP and in the viviparous Uiiau (Bradypus didactylus). The most common number Compara- is 12, which is that not only of man, but of the ourang- ^t.ve outang, silky monkey (sirma mankim), patas (a. patas), maimon (simia maimon), macaca (sirnia cynomolgus), baboon (s. sphynx), magot (s. inuus), mandrill (s. maimon). Plate XXXIV. fiff. 1. The spine. Cervical vertebrae. . blenny LtJoTel Zy assumeyi green ^ nu£ colour after boiling. The causes of these vanet.es m ^ white beqar, cive,, the cat tribe (felis), the dog, wolf and fox, the didelphis tribe, the cavy, guinea pig and paca, the mouse tribe, excluding the two exceptions already mentioned, the long¬ tailed manis, the stag, the antelope genus, the goat, sheep, and ox, and the dolphin and porpoise. The num¬ ber is 14 in the gibbon, coaita, and weeping monkey, in the colour are unknown. SECT. I.—OSTEOLOGY OF THE MAMMIFEROUS ANIMALS. The skeleton of the Mammalia bears a general resem¬ blance to that of the human subject, in the construction, Shed howling ape (simia UelzeM), the tarsius, the brown bear, recognise "ufe importance of the trunk, and especially of raccoon and coat,, the weazel gmus the porcup.ne, hog, the°spine, in the different classes of mammiferous ani- and giraffe. It ,s 15 m the Ion, hedgehog and tenrec m I ’ the badger, pangolin, and seal. The number is 16 in the m The spine consists of separate vertebrae, which are con- glutton, hyena, ant-eater, American lamantin, and rnega- veniently distinguished, as in man, into cervical, dorsal or thenum. In the horse, quagga, and dugong, iey are costal, lumbar, sacral, and coccygeal or caudal 18 ; in the rhinoceros 19 ; in the Indian elephant and tapir The number of cervical vertebrae is the same in animals 20 ; and m the Unau or two-toed sloth ,.3, which, as al- with the longest and shortest necks,—in the horse, camel, ready stated, is the greatest number yet know n, and giraffe, and in the mole and ant-eater. They are al- In the ape family the dorsal vertebrae resemble those ways seven. The only exception is observed in the A.i or of the human subject, but then spinous piocesses are three-toed sloth (bradypus tridactyla), which has 9 cervical long, and erect in the macaca and magot. . In the bats, vertebrae (Cuvier); and an apparent exception is presented instead of spinous processes, which are wanting, there are in the dolphin and porpoise, in which the first two are con¬ solidated into one; and in the cachalot or large-headed whale, in which the last six, sometimes the whole seven, are united or ankylosed. The last six are also united in the ant-eater and manis (Cuvier). Even in this state, however, the traces of the original separation are distinct. In the ape the cervical vertebrae are distinguished from those of man chiefly by the spinous processes being stronger and not bifid, and in their bodies being projected more over each other before, so as to support the head more minute tubercles. The want of these, however, in sun¬ dry species, leaves the column comparatively smooth be¬ hind. In the proper quadrupeds these processes are larger, straighter, and stronger, as the head is weighty or sup¬ ported on a long neck, in order to give attachment to the strong yellow cervical ligament. This peculiarity is very distinct in the giraffe, camel, ox, rhinoceros, and elephant. In the dolphin they are straight, and smaller than those of the loins. The lumbar vertebrae vary in number still more, per-Eumbar the cervical vertebrae are flattened from behind backwards, and those of the atlas are very large, both for supporting the head and giving attachment to the strong muscles employed in defence, attacking prey, or bearing it off. For the same purpose the spinous process of the axis is very prominent, while the others are short and directed towards the head. In the mole and shrew the cervical Dorsal vertebrae. perfectly. In the Zoophaga the transverse processes of haps, than the cervical and dorsal; and this variety may oc- vertebraa casionally be traced to the greater or less distinctness with which the sacral and coccygeal are distinguished. The smallest number is 2, which is that of the twro-toed ant- eater, ornithorhyncus, and American lamantin; and the greatest 9, which is that of the lori. The most frequent number is 7, which is that of the greater part of the monkeys, the macauco, the great bat (iioctula), the hedge- vertebrae, which are void of spinous processes, are simple hog, shrew, raccoon ; the tiger, panther, puma, and cat, in osseous rings, which move easily on each other, probably the feline genus; the wolf and fox in the dog; the hare and to facilitate the frequent motions requisite in these ani- rabbit; the whole murine genus except the hamster; and in mals in burrowing. In the hog the cervical transverse the camel and dromedary. The next number in frequency processes are compressed and broad before, so as to ap- is 6, which is that of the horse-shoe bat, the colugo (galeo- peai double. In the elephant the cervical vertebrae have pithecus), the white and brown bear, the coati, the weasel shoi t single spinous processes, and the bodies projecting genus, the civet, the lion, among the feline, and the dog over each other as in the ape. In the Ruminants the among the canine genus, the didelphis and cavy genera, length of the spinous processes diminishes as the neck is the hamster, the stag, antelope, goat, sheep, ox, horse, and elongated. _ Ihus they are almost wanting in the camel quagga. The gibbon, coaita, Ai, Echidna or Ornithorhyn- and giraffe, in which the arched neck is much retro-flected; cus hystrix of English zoologists, six-banded armadillo, and the same peculiarity is recognised m those of the and dugong, have only 3 lumbar vertebrae; the ourang- 10T?e‘ c , -s. ii I, outang, pongo, and howling ape, 4; the vampyre bat 4; the from these facts it results that the length of the neck hyena, armadillo, Unau, and tapir, 4; the jocko, tarsier, offiS but on ,lie lo"g,tud,nal ^~bot 5CeSer andglrn’ ,her cuT’ The doisal, thoracic, or cotta, vertebne are distinguish- chl^Sti/aKT nd ed by forming the central fixed basis of the ribs; and rel have 8 8 ' ° 4 their number depends on that of the latter class of bones, In theQuADKUMANA and ZooeHAqA generally the outer varies iomYl whidt tha'ofTnd ° COS“‘vert** side “f each posterior articular process'presel an apex vtiiiLS om ii, wmen is tliat oi trie L/hiiiese rooukoy* com- turned tiapkwnrd cr» tUof * 4.* i * o .non hat, armadillo, helmet-headed dolphin glo- the next vertebra s l£ked£^ bxeps), and Gangette dolplun, to 23, winch is that of the confine its movement much Though Ufis a^eTk fotd ANATOMY. 3 Compara- in the Rodentia, it is there shorter; and the arrangement live is wanting in the other tribes. The size of the transverse Anatomy. processes indicates the strength of the loins,—a fact which is evinced especially in the instance of the horse, porpoise, &c. Sacral ver- The number of sacral vertebrae is still more various, tebrae. even in the species of the same genus. Thus, while in several of the ape genus, in the lori, in the vampyre bat, the colugo (galeopithecus), the coati, and two of the didelphis, there is one sacral vertebra only, most of the apes have sacra consisting of 2, 3, 4, 5, or 6 pieces ; the majority of other animals have 3 sacral vertebrae; the hedgehog, porcupine, guinea pig, paca, hare, tiger, several of the murine genus, the ant-eater, rhinoceros, camel, dro¬ medary, chamois, goat, sheep, and ox, have 4; the ele¬ phant has 5; the Ai 6; the Unau 7; and in the mole, white bear, and quagga, they also amount to 7. The fre¬ quency of the three sacral vertebrae in the lower animals shows that Galen, who ascribes only 3 to the human sub¬ ject, must have derived this inference from the former. These vertebrae are in the mammalia narrower than in man, and their direction forms with the spine, instead of receding backwards, a straight line ; an arrangement evi¬ dently connected with the horizontal position of the for¬ mer. The shape of the sacrum in the lower mammals is that of an elongated triangle ; and it is further remarkable, that in those species which occasionally assume the erect attitude on the hind leg, as apes, bears, and sloths, the width of the sacrum is proportionally greater. The sa¬ cral spines, which are short in man and the ape, become longer in the ZooPHAGA,and form a continuous ridge in the rhinoceros, most ruminants, and especially in the mole. In the vampyre bat the sacrum forms a long compressed cone, the extremity of which is united to the ischial tu¬ berosities, without sustaining a coccyx. The seal has two sacral bones; but the Cetacea, e. g. the dolphin and por¬ poise, are void both of sacrum and coccyx. The coccygeal bones constitute the tail of the lower animals, and in many instances they are extremely nume¬ rous. The smallest number is 3, which is that of the magot (simia sylvanus, pithecus, et inuus) or Barbary ape ; and the greatest yet known is that of the ant-eater, in which they amount to 40, and the long-tailed manis, in which they amount to 45. Next to these may be placed that of the coa'ita 32, the baboon 31, the phalanger (di- delphis orientalis) 30, the marmoset (didelphis murind) 29, the pangolin 28, the silky monkey {simia rosalia) and black rat 26, the weeping monkey and howling ape 25 ; the panther, mouse, dormouse, and elephant, 24; the lion, beaver, water-rat, Norway rat, and field-rat, 23 ; the flying- cat, puma, cat, dog, marmot, and rhinoceros, 22; the otter, 21; the Chinese monkey, raccoon, civet, hare, and rabbit, 20; the tiger and wolf, 19; the macauco, glutton, marten, fat dormouse, dromedary, giraffe, and quagga, 18; the tarsier, shre^V, camel, and horse, 17 ; and other genera and species, without any determinate order, descending so low as to 9, 8, 7, 6, and 4. The quilled duckbill {echidna, ornithorhyncus hystrix) has only 12 caudal vertebrae, while the common one (ornithorhyncus paradoxus') has at least 20. The gibbon and vampyre bat are the only mammifer- ous animals, excepting the Cetacea, in which there are no coccygeal bones. It sometimes happens that a monkey or opossum loses a portion of its tail, when the truncated end is converted into a knotty excrescence, sometimes carious, always different from the taper point of the last coccygeal vertebra; and in this case it is difficult to de¬ termine the exact species. In the Cetacea, in which the absence of pelvis affords no mark to distinguish the lower vertebrae into lumbar, sacral, and coccygeal, those below the dorsal may be re- Coccygeal or caudal vertebrae. garded as lumbo-coccygeal; and their number is estimated Compara- by deducting that of the cervical and costal from the total tive number. The following table, which shows the number Anatomy, of the costal, the lumbo-coccygeal, and the total number of vertebrae, indicates that their number varies much in various genera of this family. d. Lamantin 16 Dugong...... 18 Dolphin 13 Tursio ...13 D. Globiceps 11 D. Griseus 12 D. Gangeticus 11 1-c. t. d. l.c. t. 24 46 Porpoise 13 40 60 28 46 Narwal 12 35 54 47 67 Hyperoodon.... 9 29 45 38 58 Cachalot or White 37 56 Whale 14-15 39 60 42 61 Greenland Whale... 15 37 59 28 46 llorquhal 14 31 52 In general, however, if we reckon the first 2, 3, or 5 ver¬ tebrae after the costal as lumbar, it may be said that the caudal vertebrae of the Cetacea vary from 22 or 25, which are the numbers respectively in the lamantin and dugong, to 34, 38, and 42, at which they may be estimated in the dolphin. We shall see that, in the dugong at least, we are guided in this estimate by the rudimental bones of the pelvis. The coccygeal or caudal vertebrae of the Mammalia may be distinguished into two kinds; those which con¬ tain a canal in continuity with that of the vertebral co¬ lumn and sacrum, and those in which the pieces are solid. The first, which are next the sacrum, have articular, transverse, and spinous processes, distinct in proportion as the animals move their tails. The latter are generally prismatic in shape, diminish in size towards the extre¬ mity, and have only slight tubercles for muscular attach¬ ments. Animals with prehensile tails, as the American ape (sapajous), have above, at the base of the body of each vertebra, two small tubercles, between which pass the tendons of the flexor muscles. By means of this mechanism these animals can twine the tail round the branch of a tree with sufficient force to support the weight of the body. The Mammalia with long mobile tails have often two or three small supernumerary bones placed on the lower surface of the junctions of several of the coccygeal ver¬ tebrae, from the 3d or 4th to the 7th or 8th. These sesamoid bones give attachment to muscles. In the beaver, which employs its tail as a trowel, the transverse processes are remarkable for size, while the lower spinous processes are larger than the upper ones,—an arrangement which enables it to depress the tail forcibly when it beats the ground. The shape of the chest in the Mammalia varies in The chest, general as the clavicles are present or wanting. In ani¬ mals provided with clavicles, as the Quadrumana, bats, the squirrel, beaver, mole, ant-eater, hedgehog, and sloth, the shape of the chest approaches to the human, or is conoidal, and flattened before and behind. In those void of clavicles it is compressed laterally, from the smaller in¬ curvation of the ribs; and the sternum makes a remark- The ster- able prominence, so that the transverse or intercostal dia-num- meter is less proportionally, and the sterno-vertebral is greater proportionally, than in man and the claviculated animals. In the long-legged animals, as the giraffe and those of the stag kind, this prominence of the sternum is sufficient to give it a keel-like appearance (thorax carina- tus). In the carnivorous animals the chest presents its greatest longitudinal extent. The number and shape of the ribs varies in the differ- The ribs, ent tribes. In number, indeed, the ribs always correspond with that of the costal vertebrae. Thus, in the Quadruma¬ na, Zoophaga, Rodentia, Edentata,and Ruminantia, they vary from 12 to 15 pair, with only three exceptions, the glutton, hyena, and ant-eater. In the Chinese monkey, common bat, and armadillo, they are a pair less than in Compara- man. While the quilled duckbill {echidna, ornithorhyncus tive hystrix) has only 15 ribs, the common duckbill {ornithor- AnatQni)r. ]Lyncus paradoxus) has 17; the horse and quagga have t]ie rhinoceros 19, the elephant and tapir 20, and in the Unau or two-toed sloth they amount to 23, which is the greatest known number. On the whole, the most prevalent number is 13. In the carnivorous animals they are narrow and dense in structure. In the herbivorous they are large, broad, and thick. In the armadillo the two first ribs are large compared with the others. In the two-toed ant-eater, which has 16 pairs, they are so broad that they are imbricated over each other like the plates of a corslet, and render the parietes of this animal’s chest exceedingly solid. In the two species of duckbill {orni¬ thorhyncus paradoxus and hystrix ; echidna of Cuvier), the true ribs, in number 6, consist of two portions—a long or vertebral joined to the spine, and a short or sternal at¬ tached to the sternum. These portions are united by cartilage, so as to resemble the ribs of birds. The 9 or 10 false ribs terminate before in broad, flat, oval plates of bone, which are mutually connected by elastic ligaments. The sternum, which is broad in the ourang and pongo, is narrow in the other species of ape, and consists of seven or eight pieces. In the vampyre and all the bat family it is narrow, but presents before or below rather a promi¬ nent azygous ridge or keel {carina), and an anterior ex¬ tremity, broad on the sides, like a T, for receiving the clavicles. In the mole the clavicular extremity of the sternum is continued before the ribs, and is flat on the sides for receiving the two short clavicles. In the hog the sternum is broad behind and narrow before. In the rhinoceros, horse, and elephant, it is prolonged before and flat on the sides. In the Cetacea it is broad and thin, especially before. Cranium. Though the Quadrumana have 8 cranial bones, the sphenoid often consists of two portions, one forming the orbitar wings and the anterior clinoid processes, the other the temporal or large wings, the posterior clinoid processes, and the basilar fossa. The two parietal bones are early united into one in the Chiroptera and the other Zoo- phaga, in which, however, the frontal remains biparted by a middle suture. The temporal tympanum is separat¬ ed from the rest of the bone by a suture, which is seldom obliterated in the feline, canine, and viverra genera. The temporal tympanum is also separate in the Rodentia, and the frontal ununited. The parietal is united in some, as the hare, the porcupine, cabia, marmot, rat, and squirrel; se¬ parate in the mouse, fat dormouse, and rabbit. The frontal and parietal bones of the elephant are early united with the other cranial bones, and form a vault without trace of suture. In the hog, tapir, and hippopotamus, the two parietal bones form one piece, while the frontal bone is biparted; and though in the rhinoceros both are biparted, the frontal is early united into one portion. The sphe¬ noid bone of the animals of this tribe long consists of two pieces, one forming the orbitar wing; the other the tem- poial wings, which, it is to be further observed, are the smallest, in opposition to their proportional dimensions in man. In the Ruminants and Solidungula the frontal remains long parted by its middle suture; but the two panetals are represented by a single bony vault. The tympanum is always distinct from the temporal bone. In the seal and walrus the parietal and the frontal con¬ sist of two pieces. The lamantin has only one bony arch representing the two parietal and the squamous part of the temporal bones, while the temporal tympanum is de¬ tached from the rest of the bone. In the other Cetacea the parietal bones are at an early period united to the oc¬ cipital and temporal bones, so that the five form one solid poition. I he auditory or pyramidal bone is always de- ANATOMY. tached from the temporal, and adheres to the cranium by Compara. soft parts only. The sphenoid is not only long separate, tive but consists of several portions. > v Though, among the Quadrumana, the cranium of the ourang-outang approaches that of man in shape, it differs nevertheless in the connections of the constituent bones. The temporal wing of the sphenoid bone is very narrow, does not reach the parietal, and touches the frontal only by its upper extremity, so that the temporal bone is part¬ ly articulated with the frontal. The temporal suture is not imbricated, but serrated. The same mode of connec¬ tion is observed in the mandrill, macaca {s. cynocephalus), magot, and guenon {Cercopithecus), or tailed monkey tribe. In the American monkey the temporal wing of the sphenoid touches neither the frontal nor the parietal bones ; but the temporal bone is articulated directly with the malar by its flat portion. In the American monkeys the frontal bone does not touch the temporal wing of the sphenoid, and the parietal is articulated to the malar. In the howling ape {simia beelzebul) the connections are as in man. The connections of the cranial bones are in the Zoo- Coimec- phaga the same as in man. In the Rodentia the sphenoid tions. is joined to the frontal and temporal, without touching the parietal; and the orbitar and temporalare very small. In the armadillo, pangolin, and sloth, the connections are as in the Rodentia; but in the ant-eater the parietal bone, continued below the cranium, is united to the sphe¬ noid at the posterior part of the orbito-temporal fossa. In the elephant, though the cranial bones are at an early period consolidated into one, the auditory is always distinct from the temporal bone. In the hog, tapir, rhi¬ noceros, and hippopotamus, the sphenoid is united to the parietal bone, and its temporal wings occupy a small space only of the orbitar and temporal fossae. The orbi¬ tar wings, though larger, appear small externally. The auditory bone, though distinct, is, however, united by its base to the margin of the auditory canal of the temporal bone. The sphenoid of the ruminants is articulated, as in man, with all the cranial bones; but its orbitar wing, which is extensive, is principally concealed within the cerebral cavity, and covered by the orbital part of the frontal bone. In the Cetacea generally, all the sutures which remain after early life are squamous or imbricated. I he outline of the frontal bone in the ourang-outang is more irregular than in man, and the orbitar arches are less surbased. In the American monkeys its outline is tiiangular, and terminates in a point towards the vertex. others of this family {Simia), this bone is almost elliptical, and the orbitar arches are nearly straight; and m the whole family these arches form, as in man, the an- tenor border of the frontal bone, in consequence of the narrowness of the root of the nose. In the makis it be¬ gins to widen, and the eyes become oblique,—a circum- stance which gives their frontal bone a rhomboidal shape. ie fronta. bone in the Zoophaga, and in all the subse¬ quent Mammalia, except the Cetacea, forms an irre¬ gular prismatic or cylindrical surface with three faces—a superior, bounded before by the muzzle, behind by the cranial convexity and two lateral, descending into the or¬ bitar and temporal fossa on each side. The hedgehog, mole, shrew ant-eater, some of the phoca, the morse or 3 a"d thf rhinoceros, have no proper orbitar arches; and np!rfir0n arbi01?e’it l0Ugh br°ad behind> is contracted and nearly cylmdrica! between the orbits. In the hippo- enl p!’ ™minants, and the one-hoofed animals, it enlaiges, and forms a vault over each orbit. Lastly, in blbi- a fitlpf Aflfc ;S,narow from before backward, resem- beneatl^ he m •nhed,aCr°SS the cranium> but descends beneath the maxillary bones to form the floor of the orbit. ANATOMY. Com para, live Anatomy The occi¬ pital bone. [ The tem- joral bone. ,1’he facial lones. intermax¬ illary lines. The parietal bones of the ourang-outang differ from those of man only in their temporal margin being nearly straight. Those of the ape are narrower, and become more oblique-angled as the cranium is flattened. In the Zoo- phaga and Edentata they are almost rectangular. The single parietal of the Rodentiais nearly quadrilateral; but it is sometimes flat, sometimes rounded, sometimes sur¬ mounted by a crest. Of the single parietal bone of the ruminants, that of the stag, most of the antelope genus, the sheep and the goat, is broad, and sends on each side a narrow process into the temporal fossa before the occi¬ pital arch; in the camel it is narrower, and bears a lon¬ gitudinal crest; and in the ox and antilope bubalus it is placed behind the occipital crest, and resembles a fillet surrounding the back of the head transversely. In the Solidungula the single parietal is nearly quadrilateral, and placed before the occipital crest. The occipital bone in the lower mammalia is remark- 1 able for five characters. 1. The proper occipital surface, instead of being oblique or horizontal, and inferior or ba¬ silar, becomes vertical and posterior. 2. The plane of the occipital hole forms with that of the orbits an angle constantly diminishing, becomes parallel to the orbitar plane, and at length crosses it above the head. 3. The plane of the occipital condyles, instead of being transverse and horizontal, becomes oblique, and at length vertical. 4. The basilar or cuneiform process is not only hori¬ zontal, but forms with the occipital a right angle. And, 5. The mastoid process, which in man and the ape forms part of the temporal, belongs in the other mammalia to the occipital. In the polar bear, however, the mastoid process constitutes part of the temporal bone. From the 1st, 2d, and 3d characters, it results that the head of quadrupeds is not balanced on the spine, but is suspended by muscles, tendons, and ligaments, especially the strong cervical, which connects the occipital spine to the spinous processes of the cervical and dorsal vertebra;. This ligament, therefore, though feeble and indistinct in man, is strong, particularly in quadrupeds with heavy head or long neck, in order to counteract the disadvantage of the long lever. It is strongest in the elephant, and is almost wholly ossified in the mole—a condition requisite for the burrowing faculty exercised by that animal. The temporal bone is naturally distinguished in the Mammalia into two parts ; a fiat or proper temporal, cor¬ responding to the squamous part of the human temporal bone, and the pyramidal, acoustic, or auditory, correspond¬ ing to the pyramidal or lithoid portion of the human sub¬ ject. The first only, which is proper to the scull, claims attention here. In the ourang-outang and most of the genus simia it forms a trapezium with the longest side above, and the height of which varies with that of the scull. In the American apes it is smallest in this direc¬ tion. In the Zoophaga the proper temporal bone is as in the ape. Being narrow in the Rodentia behind, it is a little rounded in the short-muzzled Edentata, the Rumi- nantia, and Fachydei{mata. -The ethmoid is, strictly speaking, the olfactory bone, and shall be mentioned under the organs of sense. The sphenoid, among other offices, may be regarded as the essential ophthalmic bone. I he facial bones ot the lower Mammalia differ from those of man; first, in the number of separate pieces; and, secondly, in the form and proportional horizontal ex¬ tent. The difference in number consists in each superior max¬ illary bone being divided into a maxillary bone proper, and an anterior or labial portion; which being interposed be¬ tween the proper maxillary, are commonly denominated the intermaxillary (ossa intermaxillariaf As it bears also the 5 superior incisors, it is named by Haller the incisive bone Compara¬ tor mcisivum); but since it is found not only in the rumi- dve nants, which, excepting the camel, are void of incisors, but Anatorny- in the Edentata and Cetacea, this denomination is less applicable than the former. It may be doubted whether these should be regarded as additional bones, as they are generally represented by zootomists ; for they are in truth merely the incisive or anterior portion of the superior maxillary bones. In other respects, the difference be¬ tween the human and the animal superior maxillary bone is, that in the former each bone is in one piece, in the latter it is in two. Even in the human foetus the trace of the separation may be recognised ; and we have seen it in the human scull some years after birth. Conversely, it is early obliterated in some quadrupeds. Thus, though distinct in the ourang-outang seen by Cuvier, it was not found by Tyson or Daubenton, and is wanting in one pre¬ served in the Hunterian museum. In a young specimen of the jocko also, noticed by Cuvier, no trace of the in¬ termaxillary suture was observed. It appears also to be wanting in the perforated bat, the horse-shoe bat, and the three-toed sloth. Mutually united on the mesial plane, the intermaxillary bones are united to the maxillary by sutures, which pass from the outer angles of the latter, near the incisive holes, towards the palate, where they intersect. In form and size it varies in the different orders and genera. Small in many of the Zoopkaga and the walrus, it is large in the Rodentia, in the hippopotamus, porpoise, and cachalot, and prominent in the wombat. In the duckbill it consists of two unciform portions, united by a broad intermediate cartilage. . Tlie peculiarity of the animal face consists in the ho-Elongation rizontal elongation of the two jaw-bones. In the monkey °f the face, tribe this elongation is trifling; and all that is remarked is, that the palate and maxillary bones are more elongated in proportion to their height, and that their anterior part, instead of being vertical, is more or less inclined forwards. The degree of this elongation, which differs in different genera, may be estimated by the acuteness of the facial angle. The narrowness of the interorbital space is another Interorbi- character of the animal countenance. In the guenon and tal region. American ape it is a mere septum; but in the ourang- outang, magot, and howler, it is larger, by reason of the nasal fossae ascending to this height. From these the face of the Zoophaga is distinguished by the following circumstances. 1. The breadth of the ascending maxillary processes throws the orbits to the sides; 2. these orbitar surfaces form the anterior wall instead of the floor of the orbit; 3. the malar bone is united neither to the frontal nor to the sphenoid bone, and forms only the zygomatic arch and the lower margin of the orbit; 4. the orbit, clos¬ ed neither behind nor below, communicates freely with the temporal fossa; and, 5. the palate bones are much elongated and form a considerable space of the internal wall, to which the ethmoid bone does not contribute. In the Rodentia the interorbital space is still larger, by rea¬ son of the size of the intermaxillary bones throwing the maxillary backwards and to the sides, where they form the inner orbitar wall, in which the palate bones occupy only a small space. The anterior wall is formed by a process of the maxillary, which contributes to form the zygomatic arch, while the malar is suspended in the middle between the process and that of the temporal bone. Very similar is the face of the elephant, except that the height of the alveoli from the tusks, thrusting the nose upwards, and shortening its bones, alters entirely the expression of the head of this animal. In the sloth, in which the face is short in proportion to 6 Compara¬ tive Anatomy. Orbits. The lower jaw. ANATOMY. the scull, the malar bone attached to the maxillary only, mvoRA.most f ^^^InS^ffSleS-idah pZfkZ'u. ht^et’eral of the Uodentia,for instance the paca, beaver, forms the inner wail of these fossa. The zygomatic arch, and porcupine, and the armadil o, ant-eater, and duck- which ^interrupted in the ant-eater and pa?i|olin, is com- bill, among the Edentata In the Zoophaoa, however, nMed in the Cape ant-eater and the armadillo. In the in which the prehensile and masticatory muscles are large Sp r and rhinoceros the maxillary bone passes beneath and powerful, the « becomes broad, and its coronoid the'orbit; and the nasal bones form a sort of vault, which process is extensive. The angle which the « forms supports in the first animal the trunk, and in the second with the body of the jaw, and which is almost right m the the horn adult human subJect’ becomes obtuse m the lower am- In the CETACEA the maxillary and intermaxillary bones mals, nearly at the same rate at which the ramus dis- form a sort of flattened beak, distinguished into four paral- appears ; and indeed the transition of the angle into a tit.- iiaoal fossa is a vertical opening , . , , r- ai i • ai surrounded before and laterally by the intermaxillary after these parts have been seen for the last time in the bones. The maxillary ascend in the same manner, and amphibious Mammalia. . . . _ cover that part of the frontal bone which forms the orbitar When the mammiferous cranium is considered gene- vault, hut do not themselves contribute to the formation rally, and the relative direction and proportion of the of this cavity. The nasal bones are two minute tubercles cranial and facial part of the head examined, we recog- implanted on the frontal bone above the narrow aperture, nise more distinctly the characters by which the lower The malar is in the shape of a style, suspended by cartilages orders of that class are distinguished from man. This beneath the orbit; and the latter cavity is completed be- character consists in the position of the occipital bone and hind by a process of the frontal, which joins the zygoma- hole, the position and direction of the facial bones in re- tic of the temporal bone, and below which the orbitar lation to the frontal, the elongation of the former, and and temporal fossce communicate. large size which they present in relation to the cranial. The direction of the orbits, the shape of their base or In the human subject, it has been already observed, the facial border, and their relation to the temporal/ossa, are important circumstances in the animal face and cranium. In the simice the angle of the orbitar axes is rather smaller ; and the shape of the margin, which is quadrilate¬ ral in the jocko, becomes oval in the ourang-outang and American monkeys. The angle of the axes enlarges in the other Mammalia; and the base or anterior margin becomes nearly circular in the Zoopiiaga, Rodentia, Edentata, and Pachydermata ; but the arch is incom¬ plete behind. In the Ruminants and Solidipeda, however, in which it is also circular, the border is complete. In the Cetacea the orbitar vault is semicircular, their axes are rectilineal, and there is no floor. In the human scull the junction of the malar bone with the frontal and.sphenoid completes the orbit exter¬ nally, and prevents it from communicating with the tem¬ poral fossa ; and the same arrangement is observed in the simice. In the Carnivora, Rodentia, Edentata, and Pachydermata, however, in which the malar bone is position of the occipital bone is oblique and horizontal, and the plane of the occipital hole is horizontal, while its position is anterior. In most quadrupeds, while the bone assumes a vertical position, the hole becomes posterior, and its plane vertical or oblique, in proportion as the face is elongated. The plane of the occipital hole forms with that of the horizontal a considerable angle, which Dauben- ton undertook to determine, by drawing one line through the plane of the aperture, and another from its posterior margin through the lower edge of the orbit. (Mem. de VAcad, des Sciences de Paris, 1764, p. 568.) In the horse this angle is about 90°, while in the ourang-outang it is only 37°, and in the lemur 47°. In other respects, however, it furnishes an imperfect result, since in most quadrupeds which differ very much it ranges between 80° and 90°. The direction of the face in relation to that of the Camperian cranium, determined according to the method of Camper, line and furnishes more accurate results. While in the human ansle* subject it varies, according to the races, from 70° to 80°, united neither to the frontal nor the sphenoid, the orbit in the ourang-outang it is only 65°; in the American is not only incomplete on the external posterior border, and long-tailed monkeys about 60° ; in the macaca and but communicates freely with the temporal/omt. In the baboon about 45° ; and, lastly, in the mandrillo, the most Ruminants is observed an arrangement intermediate be- vicious and ferocious of the monkey tribe, only 30°. In tween that of the Quadrumana and that of the Carni- some species in which the ear is elevated and the guttu- vora. The malar bone, united to the frontal, completes ral fossa deep, for instance in the pongo and alouate or ^ T?' i ^ 18 T? United t0 the sPhenoid’how\er, the small size of this angle does not indicate pro- mhii nf Hw tnl3 ^P0™1/^0communicate. The portional elongation of muzzle; and to rectify this incon- sahS° suPeificial> that 11 can scarcely be venience, it is requisite to draw the basilaf line of the The lower imv nf fLe ’c i i rir fac^ angle parallel to the base of the nostrils. With from th t Of i chRflvrr ?l7S •quad-ruped dlffers thlS modification, however, the Camperian line admits of TlTtWmla^ circumstances, correct application to the human race and Quadrumana w R °nly’ mTwhlch the frontal ^ses are small and not promi- r ? mostdistinct in the Caucasanrace, begins to be- nent. In quadrupeds, for instance the carnivora several cotne faint in the negro, and is altogether lost in the monkey of the ruminants, and in the elephant the frontal sinuses nfYho ]ln ourang-°utang’. indeed, the animal character are so large and prominent as to affect the results y a large facette. The collar-bone of the echidna and ornithorhyncus is very singular. It consists of a broad central bone, surmounted by two trans¬ verse branches spreading out on each side, so as to give the whole bone some resemblance to the letter T, but sinuated so as to make the diverging branches like the Greek v. In young animals this bone consists of three portions. The two diverging branches are genuine collar¬ bones, and may be regarded as a bifurcated bone; while the middle is supported on the sternum, and has articu¬ lated to each side a part of the scapula, corresponding to the coracoid process. The collar-bone, indeed, is a power¬ ful buttress, which prevents the arm-bone from being thrust too much forward. The shoul- Of the shoulder-blade or scapida, which is present in all der-blade. red-blooded animals with thoracic extremities, and hence in all the Mammalia, the principal point is to remark the varieties which its shape presents. Though in man, most of the Quadrumana, the Chiroptera, and the elephant, the vertebral margin or base of several authors on human anatomy is the longest, it becomes the shortest in most quadrupeds, especially those which, like the Ruminants and the Solidungula, have long legs and narrow chest. In most of them, also, this margin, instead of being straight, is rounded, as in the Carnivora and Rodentia. In the Car¬ nivora without collar-bone, the hedgehog and didelphis, the acromion is less prominent; there is another eminence directed backwards, almost perpendicular to the spine. The coracoid process, also, which is present in the Chi¬ roptera, the hedgehog, and didelphis genus, is wanting in most ot the zoophagous tribe. In the hare the acromion terminates in a long slender process, rising at right angles and bending backwards, which may be named the recur¬ rent. In the Ruminantsand SoLiDUNGULA,notonlyare this and the acromion, but even the coracoid, wanting. The scapula, again, of the hog and rhinoceros is remarkable for the disappearance ot the spine at the glenoid angle ; while from its middle proceeds a prominent process towards the costal or inferior margin. In the mole the scapula is long and narrow, like a cylindrical bone, placed parallel to the spine,—an arrangement which, together with the shoitness and thickness of the clavicle, already mentioned, is evidently connected with the burrowing habits of this animal. Lastly, in the echidna and ornithorhyncus, which in so many characters of organization approach the Am¬ phibia on the one side, and the Birds on the other, the scapula is a single sinuous bone, attached by one extre¬ mity to the sternum and middle part of the clavicular bone, with the other loose ; and in the middle an articular cayHy, m which the head of the humerus is placed, and winch evidently corresponds to the glenoid. In this in¬ stance, therefore, the clavicle and scapula may be regarded as united into a single bone. Thehume. The humerus, which exists in all the animals with tho¬ racic extremities, undergoes considerable variations. In the lower animals generally it is much shorter than in man • and it is invariably shorter in proportion as the metacar¬ pus is elongated. I hus, in animals with what is named a camwn bone, that is, one metacarpal, as in the horse and the ruminants, the humerus is so short that it is conceal¬ ed m the soft parts as far as the cubit. In the Cetacea it may be said to attain its maximum of brevity. In the bat and sloth it is long in proportion to the rest of the body dina v fST ;';,m0le is ^rl,aPs tlle extraor- i , ' , 16 mamn“ferous animals. Not only IS it articulated with the scapula by a small head hut It .S connected with a facette of the clavicle by another be longtng to the great tuberosity, and between which and ANATOMY. the head is a deep pit. The crest of the small tuberosity Compara. is so large, that it represents a square placed vertically, tive so that the linea aspena is above. The rest of the body^j^^ of the bone, which is very short, is arched above, so that the cubital extremity is directed upwards. From this arrangement it results that the cubit is elevated above the shoulder while the palm is turned downwards,—a disposi¬ tion necessary for the burrowing habits of the animal. In the simice the radius and ulna are arranged as in Radius and man, except that in the cynocephalus, mandrill, magot, and ulna, guenon, the coronoid process of the ulna is narrower, and the radial facette deeper. In the other Mammalia the idna very generally disappears or becomes rudimentary only. In the bat family and the colugo (galeopithecus) the ulna is wanting or is represented by a slender style placed below the radius. These animals are therefore des¬ titute of the power of pronation and supination. In the Zoophaga, the radius and ulna, though separate, are void of rotatory motion; and the olecranon is compressed, and continued farther back than in man. In the Pachy- dermata the radius is before and the ulna behind, and, though distinct, there is no rotation. In several of the Rodentia, for instance the marmot, porcupine, &c. the coronoid process is small, and in others, e. g. the cavy, hare, and mouse family, it is altogether effaced. In the Ruminants the ulna is united immovably to the radius ; and in the Solidungula it is represented by an olecranon adhering to the posterior surface of that bone. In the Cetacea, though both bones are present, they are much flattened. The carpus of the ape genus contains one bone more Carpus, than that of man, situate between the pyramidal and large bone, and which seems to result from the trapezium be¬ ing divided into two parts. Conversely, in the Zoophaga, but especially in the dog, cat, hedgehog, shrew, bear, and seal, the scaphoid and semilunar are united into one large bone. Those which have a vestige of thumb, as the hyena and glutton, have the trapezium very small. The mole has not only 9 carpal bones, as the ape, but a large sickje-shaped bone, which is attached to the radial mar¬ gin of the fore paw, and which gives it the shape proper to the habits of the animal, ffhe toes are further very short. Ot the Rodentia, the hare resembles the ape; but in the beaver, marmot, squirrel, and cat, the scaphoid and semilunar make one bone; while in the porcupine the su- pei numciary bone is between the pisiform and metacarpal oi the fifth toe. In the two-toed ant-eater there are only 6 caipal bones, 4 in the first row and two in the second; m the three-toed sloth there are only 5, 3 in the first row and 2 in the second; the pangolin has 7; the cachecame 8 and a rudimental small toe; the elephant 8, 7 wedge- shaped and one elongated, corresponding to the pisiform ; and the other Pachydermata 8. In the rhinoceros, which las only 3 toes, the trapezium only is wanting; but there is a supernumerary bone on the margin of the scaphoid, and on that of the unciform, as in the porcupine. The first range consists, in the Ruminants and Solidungula, of 4 hones; m the former, excepting the camel, the second consists of 2, and the latter of 3. Those of the Cetacea, which are much flattened, are 3 in the first row and 2 in the second. hnnil6 IV!AMM.tLI^ Senerally have as many metacarpal Metacar- bones as toes, that is never fewer than 3 or more than 5,pus. are in e C^tlon 01 ^ Ruminants, in which these bones hone I 7 •lfe coHsohdated into one named the cannon wldch ie SI * WhlGh Walk on the tjPs the toes, or Minch use them as organs of prehension, the metacarpal the^^mT r^/0 ^ ^ ? -d henceTn ^l! ^ mefacxarlms 18 erroneously named the oie leg, and therefore it has been imagined, that in several ANATOMY. 9 Compare- of our domestic animals the different parts of the lower ex- the single toe which constitutes the foot are distinguished Compara¬ tive tremity are articulated in opposite directions to those of as the pastern bone, which is the first phalanx ; the coro- tive - 1 r J_._ .1 net, wj1icj1 is t]le middle or second; and the coffinbom, Anatomy. Anatomy. Fore toes. man. Thus the fore leg of the horse, deer, sheep, and dog are in truth the metacarpus of these animals ; and what is vulgarly named the fore knee or cannon bone of the horse, is actually the carpus or wrist-joint. It is there¬ fore convex on the dorsal, and concave and inflected on the palmar aspect, exactly as the carpus of the human subject. In the three-toed sloth, the three bones of which the metacarpus consists are mutually consolidated at the base and with the rudiment of a fourth toe. In the Cetacea, the metacarpal bones, which are much flattened, are also mutually united. In the Mammalia generally, if we include imperfect or rudimental phalanges concealed in the skin, there are never fewer than 3, nor more than 5. The Unguiculated ani¬ mals generally have 5, perfect and imperfect- The cha¬ racter of the perfect fore toe or finger is to consist of 3 rows or phalanges, excepting the first of the radial side, which has only 2. In the Quad rum an a this is separate, and opposable to the other toes, constituting a thumb, and giving this tribe of animals a prehensible organ en¬ titled to the epithet of hand. It is, however, shorter and less perfect in other respects than the genuine thumb of the human hand. In the codita (simia paniscus) it is con¬ verted into a rudimental bone, concealed under the skin. In the Zoophaga, which have no power of grasping mi¬ nute objects, the thumb or first toe is parallel to the others, and, though equal in length to these in the ursine family, it is shorter in the mustela, viverra, canine and feline ge¬ nera. In the latter, which have the power of erecting the claws, to prevent them from being blunted in walking, the shape of the middle and unguinal phalanx is remark¬ able. The former is triangular prismatic, with two late¬ ral and a plantar or palmar inferior surface. The third which is the third or unguinal phalanx, which has the' shape of the hoof, rounded before, convex above, and flat below. To the back of the pastern joint are connected two sesamoid bones; and to the coffin bone is attached another, named the shuttle bone. In the Cetacea, all the phalanges, which are flattened, and often cartilaginous, are united in the fin or paddle. The thigh-bone, which is single in all the classes, fol-Thigh- lows the type of that of the human frame in general figure bone, and parts. In the Mammalia it is, however, proportion¬ ally shorter, and its length diminishes as that of the me¬ tatarsus augments. In the Ruminants and Solidungula, for instance, it is so short that it is concealed by muscles against the belly; and hence it is too often overlooked and confounded with the leg. In other respects the ge¬ neral characters are, that it is not arched ; that, excepting in the bear and some of the simia genus, e. g. the ourang- outang, it is shorter than the leg-bones; that its neck is very short, and more perpendicular to the axis of the diaphysis than in man; and that the great trochanter is raised above the head, which is directed inwards. In the simice it is quite cylindrical, and void of linea aspera. In the tapir the middle part is found flattened; and at the external margin there is a prominent crest, terminating in an unciform process. In the rhinoceros the great tro¬ chanter and the unciform process are so elongated as to unite almost, and form a hole between them and the dia¬ physis. The unciform process is observed also in the horse, beaver, and armadillo. The thigh-bone of the seal is so short, that the half of its length consists of the two articular extremities. Though the leg-bones of the Mammalia bear a general Leg-bones. similitude to those of man, the tibia alone is constant; and or unguinal phalanx is shaped like a hook, consisting of the fibula, after becoming unusually slender, and chang the fore nnon one. two parts. One, directed forwards, sharp and pointed, receives the nail or claw, in a long groove like a sheath. The second part of the hook, which is behind, rises verti¬ cally from the lower part by which it is articulated, and is produced into two processes, to which are attached the erecting muscles of the claw, which are flexors of the phalanx. Among the Rodentia there is a perfect but short thumb in the hare, beaver, and jerbois ; a two-phalanx but conceal¬ ed one in the squirrel, mouse, and rat family, porcupine, paca, agouti; and a one-phalanx concealed one in the cavy, guinea-pig, marmot, &c. In the Edentata the number of fore toes varies much; in the Tamanoir, and Tamandua or four-toed ant-eater, the thumb-toe is oblite¬ rated ; in the Ai or three-toed sloth, both that and the fifth toe are obliterated; and in the two-toed ant-eater, and Unau or two-toed sloth, these, with the second toe also, are obliterated. The elephant has 5 perfect toes, all concealed under the thick, callous hide of the foot. In hoofed animals with 4 toes, for instance the hog, tapir, and hippopotamus, the thumb-toe is in the shape of a Small rudimental bone. In the Ruminants the single metacarpal bone (Chesel- den’s figure of the Deer, Plate I.) is articulated with two digital phalanges, which constitute one of the distinguish¬ ing characters of this order—the cloven foot. In some genera, at the root of these two perfect toes are two small bones, often covered with horn, which represent two other toes. The last or unguinal phalanx is always trilateral in shape. In the horse and the Solidungula generally, the two lateral toes are represented only by two bony styles, named the splint bones, situate on the two sides of the metacarpal or cannon bone. The three phalanges of VOL. m. ing its position from the outside to the posterior part of the tibia, is converted into a mere appendage, and at length disappears entirely. Thus, though it is distinct, and occupies its usual position in the simice, in the Chirop- tera it is extremely slender; and since the femora are directed backward, the fibulae are turned towards each other. In several of the Edentata, for instance the pha- tagin, armadillo, and sloth, it is large, curved, and remote from the tibia. In the dog family and the Rodentia it is altogether behind the tibia. In the mole and murine genus it is consolidated to the lower third of the tibia, leaving an empty trilateral space above. In the rhinoceros, ele¬ phant, and hog, the fibula is flattened and united to the whole length of the tibia. In the ruminants it is repre¬ sented by a small bony appendage, placed on the outer margin of the astragalus, below the tibia, and forming the external or fibular ankle. Lastly, in the horse and Soli- dungul a, thefibula is reduced to a styloid rudimental pro¬ cess, which is firmly consolidated in the adult animal to the upper part of the tibia. Between the tarsus of man and that of the other Mam¬ malia the following are the principal differences. In the simice the fibular facette of the astragalus is The tar- nearly vertical, and the tibial is very oblique; and thesus* calcaneum wants the tuberosity, except in the pongo. In the ordinary bat family the calcaneum is elongated into a styloid process, concealed in the substance of the mem¬ branous ubiform expansions; but in the roussette (jptero- pus) the tuberosity projects beneath the foot. In the Rodentia the calcaneum is produced consider¬ ably backwards, while the scaphoid, which consists of two parts, forms a tubercle on the sole. Among the Eden¬ tata the three-toed sloth is peculiar in having a tarsus. 10 Compara¬ tive Anatomy. ANATOM Y. consisting of four bones only, the astragalus, calcaneum, and perfect toes; the bog 4 ytl|e^1|'an^[hin^^1°sSaf two cuneiform bones, the first of which is articulated not Ruminants have two perfect toes on une metataisa bone Anatomv only with the tibia, fibula, and calcaneum, but with the and two small ones attached behmdite^ large cuneiform bone, without any intermediate scaphoid lidungula have one perfect toe, ^ ’ni^ T bone. Its connection with the tibia is by means of a con- are reduced to a single styloid b,°f • vex articular surface, which rolls on the external part of the body is supported in walkl^ ^ th^f- not ^ the tarsal end of the tibia. From this mode of articular phalanx alone; and hence the term foot is ot c the tion it results that the foot of the sloth admits neither of same import as m the human subject and animals simi- being elevated nor depressed, but simply of performing larly constructed. While indeed man supports his person lateral motions of adduction and abduction, to which it in progression on the os ca/cis and the postenor or meta- owes the power of clasping the trunks of trees and climb- tarsal phalanges, m the other mammiferous animals the imr but which renders "progression difficult and laborious, former bone touches not the ground, but is always elevated The hog has a scaphoid with three ordinary cuneiform above it a considerable height. All the zoophagous or bones, and a rudimental great-toe bone beneath the first, unguiculated animals, excepting the plantigrade, support In the tapir and rhinoceros there are only two cuneiform themselves chiefly on the unguinal and middle phalanges bones. All the animals already enumerated have the both of the fore and hind foot; and neither the posterior same number of metatarsal bones as of toes. phalanges nor the calcaneum touch the ground, as is easi- In the Ruminants the cuboid and scaphoid bones are ly demonstrated on observing the gait of the hedgehog, united, unless in the camel, in which they are distinct, dog, fox, cat, or similar individuals of the same family. The At the outer margin of the pulley of the astragalus is a animals distinguished by the name of Plantigrade are bone which represents the lower head of the fibula, and believed to support themselves on the entire foot. But which is farther articulated to the upper surface of the though the foot is certainly spread on the ground more os calcis. In this side also there are only two cuneiform freely than in those already mentioned, by the bear, bones, which are united in the giraffe. The two meta- glutton, badger, and others, it appears that not the heel, tarsal bones are always united, as in the metacarpus, into but the metatarsus, is allowed to touch the ground in pro- one, which forms a posterior cannon bone. The Soli- gression. In the Ruminants and Solidungula, as already dungula resemble the camel in this, that the scaphoid is mentioned, the only part of the foot which is applied to distinct from the cuboid bone, and that there are two the ground is the unguinal phalanx; and it is well known that the horse supports himself on the plantar surface of the coffin bone only. Lastly, in the Amphibious Mammals, while the extreme brevity of the humerus and femur unfit them for progression on land without extreme awkwardness and difficulty, the expanded shape and oblique position of the metacarpal Third can¬ non bine. Hind toes. cuneiform bones, while the peroneal rudiment and the corresponding articular surface of the calcaneum are want¬ ing. The metacarpal are also consolidated into a single piece, named the hinder cannon bone, each side of which is provided with a minute bony style. The toes of the Quadrumana and the Marsupialia are longer than those of man ; but the great toe is shorter bones and phalanges, the length of the tibia and fibula, and than the others, and its metatarsal bone is susceptible of the greater length of the first and last than the middle separation and opposition, as the thumb or thumb-toe of metatarsal phalanges, all concur to give these animals the hand. Hence Cuvier, in his first classification, dis tinguished the latter by the name of Pedimana. The Aie-aie among the Rodentia appears to possess the same faculty. Among the Zoophaga the great toe remains always conjoined with and parallel to the others; and in the canine and feline genera it is obliterated. Among the Rodentia, that of the beaver is nearly equal to the other toes; those of the marmot, porcupine, and the murine genus, are shorter; in the paca it is almost obliterated; it is reduced to a single bone in the Cape gerboa; and the leporine genus have no trace of it. In the cavy, agouti, and guinea-pig, the great and small toes are each great facility in swimming. (Cuvier, Ossemens Fossiles, tome v. partie i. septieme partie.) In the Cetacea, again, while the total want of pelvic extremities renders motion on land quite impracticable, the fin-like disposition of the metacarpus and metacarpal phalanges, with the great strength of the lumbar, and the length of the coccygeal vertebrae, peculiarly qualify them for locomotion in the waters. SECT. II. OSTEOLOGY OF BIRDS. Plate The number of vertebrae of which the different regions of the spine consist, is not less variable in Birds than in ’ by Cuvier. in the following table Species. Vertebra? of Neck. V ertebra? Sacral of Back. Vertebrae- Vultur. V ulture Falco fulvus. Eagle — halia'ctus. Bald buzzard — biiteo. Buzzard — nisus. Sparrow hawk — milvus. Kite Strix bubo. Efigle owl Strix ulula. Brown owl Mutcicapa grisola. Fly-catcher Turdus merula. Blackbird.., Tanugra tatao. Tanagra Corvus corone. Crow pica. Magpie glandarius. Jay Stumus vulgaris. Starling... Loxia coccothraustes. Grosbeak — pyrrhula. Bullfinch... 13 13 14 11 11 12 13 11 10 11 10 13 13 12 10 10 10 Coccygeal Vertebrae. reduced to one bone. In the gerbon (nius jaculus) and m An, at, a c • i c o ■ , alactaga (mus sagitta) the three middle metatarsal bones from th T* ld^K.0/,these.va”atl0ns may> are united into a single one similar to the cannon bone of ^ fr0m the nUmber exhlblted the Ruminants and Solidungula ; and while the two la¬ teral toes are distinct, though short, in the former animal, they are obliterated altogether in the latter. Among the Edentata, the ant-eater, orycteropus, pan¬ golin, and armadillo, have five toes, of which the great is the shortest in all. In the sloth the great and small toe are reduced to one small bone. The other metatarsal bones are umted at their base. The toes have only two phalanges, of which the unguinal are the largest. In the subsequent families the metatarsal bones de¬ serve particular attention. In the elephant and Pachyder- mata, their tarsal extremity has a flat surface, and the phalangeal consists ot a convex tubercle, which presents below a prominent line in the middle of the bone. In the feoLiDUNGULA this line is above and below both. In the Ruminants, in which the cannon bone consists of the two metatarsal bones, the line of union is represented by a deep line like the tract of a saw. The elephant has 5 11 17 11 10 11 11 12 11 10 10 9 13 13 11 10 12 11 Spim ANATOMY. 11 Compara¬ tive ' Anatomy. Kervi.'al Wei-tebne, Jorsal ertebrae. lock. Fringilla domestica. Sparrow... cardaelis. Goldfinch Parus major. Titmouse Alaiida arvensis. Lark Motacilla nubecula. Ited-breast Hirundo urbica. Swallow... Species. . Caprimulgus Europceus. Goat¬ sucker Trochilus pella. Colibri Upupa epops. Hoopoe Alcedo ispida. King’s fisher... Picus viridis. Woodpecker. Ramphastos. Toucan Psittacus erithacus. Parrot.. Columba cenas. Stockdove.. Pavo cristatus. Peacock Phasianus colchicus. Pheasant Meleagris gallopavo. Turkey... Crax nigra. Curassow bird. ) Hocco j Struthio Camelus. Ostrich Casuarius. Cassowary... Phcenicopterus. Flamingo Ardea cinerea. Heron - alba. Stork - gnus. Crane Platalea A'iaia. Spoonbill Rccurvirostra. Avoset Charadrius pluvialis. Plover... Tringa vanellus. Lapwing Scolopax ruslicola. Woodcock... arquata. Curlew.... Hcematopus. Oyster-catcher... Rallus crex. Hail Fulica atra. Coot Parra. Jacana.... Vertebras of Neck Pelicanus onocrotalus. Pelican... carbo. Cormorant... Sterna hirundo. Sea swallow... Larus. Gull Procellaria. Petrel Anas cygnus. Swan - anser. Goose - bernicla. Bernacle - boschas. Duck - tadorna. Sheldrake - nigra. Black diver Mergus merganser. Merganser Colymbus cristatus. Grebe 9 11 11 11 10 11 11 12 12 12 12 12 12 13 14 13 15 15 18 15 18 18 19 19 17 14 15 14 18 13 12 13 15 14 1C» 16 14 12 14 23 15 18 14 16 15 15 14 Vertebrae of Back. 9 7 7 8 8 9 7 7 7 7 8 8 11 7 7 7 9 7 9 8 8 7 8 9 8 9 8 7 9 8 8 8 11 10 10 8 11 9 8 10 Sacral Vertebrae 10 11 11 10 10 11 11 9 10 8 10 12 11 13 12 15 10 10 20 19 12 10 11 12 14 10 10 10 13 10 15 13 7 12 14 14 10 11 14 14 14 15 11 14 13 13 Coccygeal Vertebra:. 8 7 7 9 7 8+ 7 8 In this table the most remarkable circumstance is the great number of dervical vertebra, which are much more numerous than in the Mammalia. They vary from 9, the number in the sparrow, to 23, which is that of the cervical vertebra of the swan. The most common num¬ ber is 11, which is that of 10 genera. The next most fre¬ quent is 12, 13, and 14, which are equally the numbers of 9 genera. The next is 15, which is that of 8; 10 oc¬ curs in 6, 18 in 4, and 16 in 3. In the stork and crane they are 19. The next remarkable circumstance is, that the dorsal or costal vertebra are greatly fewer than in the Mam¬ malia, never exceeding the number of 11, and being more frequently about 7 or 8. Thus, while they are 11 in the cassowary, swan, and sheldrake, 10 in the goose, bernacle, and grebe, and 9 in the sparrow, lark, humming-bird, parrot, crane, avoset, oyster-catcher, cormorant, and black- diver, they are 7 or 8 in all the other genera, and only 6 in the bullfinch. There are no lumbar vertebra strictly so named, for those which extend from the chest to the tail are con¬ solidated into one piece with the iliac bones. The tail, which is short, consists of from 7 to 9 vertebra. Ihe part most variable in proportional length is the neck. It is so much longer as the feet are elevated, ex- Anatmny, cept in some of the swimmers, in which it is greatly Compara- longer, because they require to seek their food below the tive surface of the waters on which they float. The bodies of the cervical vertebra are articulated not' by plane facettes, which would admit obscure motion only, but by portions of cylinders, which allow extensive mo¬ tion. The 3d, 4th, or 5th superior vertebra allow of an¬ terior inflection only, and the others of posterior inflection. This gives the necks of birds an alternate serpentine in¬ flection ; and it is by rendering the two arches, of which this curvature consists, straight or convex, that the ani¬ mal elongates or shortens his neck. The articular pro¬ cesses of the superior vertebra are directed upwards and downwards; those of the lower are turned anteriorly and posteriorly. Instead of transverse processes, the cervical vertebra of birds are provided with a tubercle above, and the anterior extremity of which terminates in a narrow style, descend¬ ing parallel to the body of the vertebra. Only the most superior and inferior vertebra have dis¬ tinct spinous processes, and these have anterior as well as posterior ones. The middle ones have before two crests, which form a half-canal, and.behind a tubercle, often bifid, or, when they are elongated, two rough lines. The atlas, which is articulated with the occipital bone by a single facette, has the shape of a minute ring. As the neck of birds is movable, the back is fixed. The Back, spinous processes of these vertebras are in mutual contact, and they are connected by strong ligaments. Most of these processes are generally consolidated into a single continuous crest, extending along the whole back. The extremities of the transverse processes terminate in two apices, one directed forwards, the other backwards; and occasionally they are consolidated into a continuous mass like the spinous. That this arrangement is requisite for the trunk to remain fixed during the violent motions which take place in flying, is rendered probable by the fact, that in birds which do not fly, as the ostrich and cassowary, the spinal column retains its mobility. The last dorsal vertebra are often placed on the crest of the iliac bones, and they are then united, as the lumbar, on the large piece of the iliac bones, from which it re¬ sults that the number of vertebra can often be estimated in no other mode than by that of the holes of the nerves which issue from the chord. The caudal vertebra are most numerous in the species Caudal which move the tail with most energy; for instance, the vertebrae magpie and swallow. They have spinous processes be¬ low as well as above, and very long transverse processes. The last of all, to which the pinions are attached, is long¬ est, and has the shape of a ploughshare or a compressed quoit. In the cassowary, which has no visible tail, the last bone is conical; in the peacock, on the contrary it has the shape of an oval plate, situate horizontally. It was early observed by the original zoologist and tra- Cranium, veller Pierre Belon, that the crania of birds were void of sutures; and that in a few only Avere these lines of dis¬ tinction into separate bones recognised. The explanation of this peculiarity is found in the history of the ossifica¬ tion of the head in young birds, which shows that the cranium consists at that period of separate bones, corre¬ sponding in number and situation to those of quadrupeds. Thus, there are two frontal bones, which are continued fonvards to form the vault of the orbits ; two small parie¬ tal bones behind the frontal; a temporal bone on each side of the scull; a sphenoid united to the occipital, even in subjects in which the other sutures are distinct; or a spheno-occipital bone, which is early united with the tem¬ poral. These sutures, however, are distinctly seen only in ANATOMY. 12 Compara- young birds and those recently hatched; for the bones tiye are very early united, and in the adult bird the cranial 'Anatomy.^ sutures are invariably obliterated. Thus, in the domestic fowl and turkey the scull is one piece ; and the only trace of suture that remains is a linear depression in the mid¬ dle of the frontal bone, indicating the original formation in two halves. In the recently hatched bird, also, the sphenoid is separated from the occipital bone by a trans¬ verse suture, extending from the one ear to the other. The occipital bone is at the same time a ring, consisting of four parts; a superior, two lateral, and an inferior which is small. The sphenoid, which forms the greater part of the base of the cranium, is nearly trilateral, with a small anterior process, to which the palatine arches are articulated. It has no pterygoid processes, and does not touch the posterior aperture of the nostrils. The tem¬ poral bone, though void of zygomatic process, has a pointed style, which contributes to form the posterior margin of the orbit. The frontal bone, after covering part of the cranium, is continued forwards in a broad, thin plate, which forms the vault of the orbits, while these ca¬ vities are separated by a thin vertical bony plate which descends at right angles from the frontal bone, and is con¬ nected behind with the sphenoid. The long eminences observed on the heads of the cassowary, curlew, pintado, and some species of hocco, are produced from this supra¬ orbital part of the frontal bone; and their interior, which consists of loose diploe, communicates with that of the same bone. Facial The face in birds is rarely so firmly consolidated as the bones. cranium. It is composed of two lacrymal bones, forming the anterior margins of the orbits, and united on the me¬ sial plane; two nasal bones anterior to the lacrymal; two bones corresponding to the superior maxillary, and form¬ ing the external lateral parts of the upper half of the bill; two inter-maxillary bones; two anterior palate bones, cor¬ responding to those of the Mammalia ; two posterior pa¬ late bones corresponding to the pterygoid processes of the sphenoid ; and the lower jaw a paraboloid bone, consisting of two rdmi united before, where they are covered by the horn of the lower half of the bill. Besides these, there is in The quad- the whole class an irregular-shaped bone, common to the rilateral cranium and lower jaw, and connecting these two together. This bone, which has been rather improperly named the square, quadrangular, or quadrilateral bone {os quadratum), consists of a body with curvilinear hollow margins, termi¬ nating in two elevated and rather pointed processes, one of which is connected with the cavity named tympanum, while the other, projecting into the orbit, affords attach¬ ment to several muscles. The anomalous character of this bone has perplexed several of the most distinguished zootomists; and while Geoffroy gives it the name of os Tympano-styloideum, Spix considers it analogous to the annular process of the temporal bone, which in the human foetus is separate; and Carus regards it as representing the incus, to which it bears a remote resemblance in shape and in one of its connections. Maxillary Both maxillae are void of teeth; but the hard, horny bones. matter of the bill covering the margins and extremities of each jaw, and constituting the mandibles {mandibula), is manifestly constructed to perform for Birds what teeth do for the Mammalia. But the most remarkable pe¬ culiarity of the, facial bones of this class is, that the upper jaw admits of more or less motion. In the majo¬ rity of instances this is effected by the jaw being united to the cranium by means of thin, flexible, elastic, bony plates; but in the parrot family the upper jaw is entirely distinct, and is connected by a proper articulation. The base of the palatine surface of the upper jaw is divided into 4 branches, which diverge backwards. The two external ones, which correspond to the zygomatic Compara- ( arches of the Mammalia, and which are very slender, tive are articulated to the quadrangular bone which moves on | the temporal before the ear. The two intermediate ones, which have been already stated to correspond to the pte¬ rygoid processes, and which are parallel, are placed beneath the septum of the orbits, and are articulated by their pos¬ terior extremities with a small bone, variable in shape, but named omoid by Herissant, which is also articulated ! with the quadrilateral. From this arrangement results a singular species of broken lever, not dissimilar to the parallel joint of the piston and lever of the steam-engine, and the effect of which is, that whenever the lower jaw is depressed by its proper muscles, it necessarily causes the quadrilateral bone to perform a slight rotatory mo¬ tion, in consequence of which, by means of the omoid bone, the upper jaw is at the same time elevated on the ] elastic plates; and as soon as the lower jaw is raised, the elasticity of these plates forces down the superior one. The upper jaw is immovable in a few instances only, and of these the calao or rhinoceros bird is one. The breast bone {sternum) is a trilateral, boat-shaped Sternum bone, concave internally, convex with a middle longi- °r breast- f tudinal crest externally, with the base of the triangle bone‘ above, and the apex, which is also incurvated backwards, below. The middle longitudinal crest, which is occa¬ sionally named the keel {carina), is shaped something like a spherical triangle, with the broadest side above, the base before, and the apex behind; and its prominence forms large spaces on each side for the attachment of the pectoral and other muscles used in flight. In the male wild swan {anas cygnus), in some species of curlew, in the crane, and in the guinea-fowl, this crest forms a cavi¬ ty for the reception of the windpipe. In the ostrich and cassowary, which do not fly, the sternum is void of crest, and is merely arched strongly. The ribs, which rarely exceed 10 pairs, may be distin-The ribs. ; guished into sterno-vertebral and vertebral. Though the latter are generally before, they are sometimes also be¬ hind. The vertebral end terminates in two diverging pro¬ cesses, one of which is articulated with the vertebral body, the other with the transverse process. The sternal extre¬ mity consists of a bony process, which performs the part of the sterno-costal cartilages of the Mammalia by uniting the rib to the sternum. The ribs of birds, however, are further distinguished by presenting near their middle a flat long process, projecting from the rib backwards at an acute angle, and resting on the rib immediately below, so that each rib is supported not only on the vertebrae and sternum, or the vertebrae alone, but on the next rib below. These processes are obliterated in the lower ribs. The coxal bones constitute one piece with the sacrum Basin or and lumbar vertebrae. The ischial portion is united with Pelvis- the sacrum, and the ischiatic notch is converted into a hole. The part which corresponds to the os pubis of the Mammalia is not consolidated before so as to form a symphysis, but proceeding directly backwards, terminates in a styloid process, variable in length and slenderness. The only exception to this mode of structure occurs in the ostrich, in which the pubal bones are united below. The infra-pubal or oval hole is present in the whole class not- 1 withstanding. It is worthy of remark, however, that in young birds this and the ischial aperture are still notches, in consequence of the deficient ossification of the parts. The direction of the pelvis in birds is nearly that of the spine, that is, obliquely backwards, and deviating but little from the horizontal line. The wings or thoracic extremities are connected to the trunk by three bones, the collar-bone or clavicle, the sea- Collar- pula, and the bifurcated bone. The collar-bones, which bone. ANATOMY. 13 s- ’ompara- are straight, strong, and cylindrical, are articulated by a tive large head with the anterior and lateral part of the ster- j Anatomy. numj jn jts motion is rather limited. It forms be- fore and laterally two short processes, one anterior-infe¬ rior and internal, articulated with the bifurcated bone; the other posterior-superior and external, uniting with the scapula, and forming a cavity, in which the head of the humerus is lodged. • ■boulder- The scapula is a long bone, flattened, but narrow, and lade. slightly incurvated, with the convex side turned towards the spine, to which it is nearly parallel in position. The head or anterior extremity is thick and extensive, oblique from before backwards, and is articulated behind with the clavicle, before with the humerus. The free extremity is thin, flattened, and sharp. The whole bone is not dissi- rtiilar in shape to a scimitar. lifurcated Besides these, which Birds possess in common with the one, or Mammalia, we find an azygous bone, situate on the mesial urcula. plane, denominated in ordinary language the Merry thought, and, from its shape, the fork-like or bifurcated bone. It consists of two long, rounded, converging branches, united at an acute angle, and forming a broad process, flat in the vertical direction, and by which it is articulated to (the anterior extremity of the crest or carinated part of the breast-bone. To the posterior or free extremities of the divergent branches are articulated the humeral ends of the collar-bones, which are thus enabled to sustain the violent motions of the humerus during flight. The branches of the bifurcated bone are separate in the os¬ trich, and each is united with the clavicle and scapula of the same side, so that the three bones form only one, much flattened, and with a hole towards the sternal ex- Itremity. In the cassowary the bifurcated bone is re¬ duced to a mere rudimental process at the inner margin of the head of the clavicle. From these facts it results, that the bifurcated bone is particularly useful in the ener¬ getic and continued efforts of the wings in flight, and not only serves to keep the clavicles apart, but, by lengthen¬ ing the distance between the collar-bones and sternum, enables the animal to use a longer lever. It is freest, strongest, and most elastic in the birds which fly best. In birds which do not fly, and which use the wings mere¬ ly to sustain the equilibrium, as the ostrich and casso¬ wary, it is reduced to almost nothing, or it is in such a rudimental and imperfect form, that it cannot keep the collar-bones apart. rhoracic The bones of the thoracic extremities, or those of the ixtremi- wings, correspond in general to those of the Mammalia. They consist of a single cylindrical humerus, articulated with the scapula and collar-bone above, two bones of the fore wing corresponding to the ulna and radius, two bones of the carpus, two of the metacarpus, consolidated by their extremities, one styloid bone as a thumb, a long finger con¬ sisting of two phalanges, and a short one consisting of one. The thumb supports the bastard pinions, the large finger and metacarpus the primaries, while the small one, which is covered by the skin, is destitute. In several of the web¬ footed divers, for instance the duck and penguin {alca im- pennis and spheniscus), these bones are flattened like thin plates. r elvic In the pelvic extremities the thigh-bone is provided with Ktremi- one trochanter only, is shorter than that of the leg, and is almost invariably straight; and is arched only in the cormorant, duck, and dobchick. In the ostrich its dia¬ meter is about four times that of the humerus. The tibia ! differs from that of the Mammalia chiefly at its lower extremity. M' bile the fibula adheres to it like a slender appendage as far as the middle, the tarsal extremity ter¬ minates in two trochlear condyles, with an intermediate pulley-like groove. The tarsus and metatarsus are repre¬ sented by a single bone of considerable length, and the Compara- head or tibial end of which consists of a middle promi- tive nence and two lateral depressions, and which, therefore, Anatomy, moves in cardinal opposition, but does not admit of exten- sion beyond the straight line. Though variable in pro¬ portion to the length in different orders, this bone is very long in the order Grall^: (Grallatores). It termi¬ nates below in 3 pulley-shaped processes, to which are attached the bones of the 3 anterior toes, with an inter¬ nal margin for that of the great toe. In the ostrich there are 2 processes only, corresponding to the two toes. In the penguin tribe, however, the tarsus and metatarsus con¬ sist of 3 bones, separate from each other in the middle, but united at the tibial and digital extremities. To the tarso-metatarsal bone of the cock, and others of the Galli¬ naceous tribe, is attached the spur, a conical pointed ex¬ crescence of hard horny matter. SECT. III.—OSTEOLOGY OF THE REPTILES. The number of vertebrae, and all the other attributes of the spinal column, vary more in this class than in all the others. In the Cheloniad or Tortoise family there are 7 cer-Vertebra?, vical, 8 dorsal, connected with the shell in an immovable piece, so as to have neither processes nor articular facettes; from 3 to 5 lumbar and sacral, consolidated in like manner ; and about 20 caudal or coccygeal. (Plate XXXIV. fig. 5.) In the Saurial or Lizard tribe, the number 7 predomi¬ nates in the cervical, being that of the crocodile and most lizards. In several, however, there are 8, as in two of the monitor genus, the American safeguard, the lizard of Fontainebleau, the dragon, the iguana, the anolis, and the gecko and seine; and in a few, as the Nilotic monitor, and an undetermined species of monitor, they amount to 9. In the chameleon there are only 5 cervical vertebras. Here, however, a singular peculiarity is observed. Instead of the cervical vertebras being, as in the Mammalia, distinguished by being unconnected with ribs, to those, from the third to the seventh inclusive, short ribs, un¬ connected with the sternum, are attached. The atlas and axis, therefore, alone are proper cervical vertebrae; but the general analogy is observed in the cervical ribs being exceedingly short and almost rudimental. The dorsal vary from 11, which is that of the crocodile and iguana, to 29 and 30, which are the numbers in the New Holland seine and Nilotic monitor. In the American safeguard, cordy- lus, stellio, crested basilisk, dragon, guana, and great ano¬ lis, they are 16 ; in the chameleon, black safeguard, and ameiva, 17 ; in the tupinambis, spotted gecko, and golden seine, 18; in the green lizard and spotted guana (poly- chrus) 19; in the Fontainebleau and gray lizard 20; 21 in the Levant seine and undetermined monitor; and 22 in the Java and New Holland monitor. TheBATRACHOiD or Ranine reptiles are void of ribs, and it is impossible therefore to distinguish the first three orders of vertebrae from each other. In general, however, there are from the nape to the pelvis 8 vertebrae, all provid¬ ed with long transverse processes, and which are longest in the last. The sacrum is represented by a long flattened but pointed bone, without coccyx. In the Surinam toad (rana pipa) the last vertebra is consolidated with this bone; and the transverse processes of the second and third vertebrae are so much larger than the others, that they resemble ru¬ dimental ribs. In the Salamander family there are from the head to the sacrum 14 vertebrae, all alike in shape ex¬ cept the first, which receives the occipital bone, and the last, which is articulated with the sacrum. These two are distinguished by wanting rudimental ribs, which are small elongated bones, movable, and articulated with the trans- ANATOMY. 14 Compara- verse processes, which are directed backwards. The arti- tive cular processes are large and imbricated, the posterior Anatomy. resting on the anterior, so as to resist the motion of the spine backwards. The sacrum consists of one vertebra only, but the coccyx or tail is composed of 27. In the Serpentine tribe the vertebrae may be said to attain the most extensive numerical developement. With the exception of the head and rudimental ribs, they con¬ stitute the whole skeleton. (Fig. 3.) From the head to the tail their shape is the same, and may be distinguished into body, articular and transverse spinous processes. In some species, for instance the boa, the spinous processes of the back are so much separated as to allow mutual motion to a considerable extent. In others, conversely, for instance the rattlesnake, these processes are so long and broad as to touch each other, while the oblique processes, which form their bases, are imbricated over each other. In con¬ sequence of this arrangement the motion of the spine is limited behind, but more extensive on the ventral sur¬ face. The vertebral bodies, which move easily on each other, are provided with a sharp spine directed towards the tail, which somewhat limits motion in this direction. The first vertebra differs from those of the rest of the body in supporting short or rudimental ribs ; there are therefore no cervical vertebrse and no proper neck in the serpent family. The caudal vertebrae are distinguished by not supporting ribs, and by their spines both dorsal and ventral being double, and forming two rows of tubercles. The articulation of the bodies of these vertebrae is pecu¬ liar. On the anterior part of the body is a round hemi¬ spherical tubercle, while the posterior presents a corre¬ sponding cavity, so that each vertebra forms a cup and ball joint with the following one. The number of costal vertebrae varies from 32, which is that of the blind worm (Angiiis fragilis), to 204 in the ringed snake ( Coluber natrix), 244 in the snake, and 252 in the Boa constrictor, and which is perhaps the greatest known number. Of intermediate numbers, the Amphisbama has 54, the viper {Coluber berus) has 139, the rattlesnake 175, and the cobra di capello 192. The caudal vertebrae vary in number from 7, which is that of the Amphisbcena, to 112, which is that of the Coluber natrix. Of interme¬ diate numbers, the blind worm has 17, the rattlesnake 26, the boa 52, the viper 55, and the cobra 63; from which it appears that the number of the caudal is not in pro¬ portion to that of the costal vertebrae. Cranium. Of the heads of the Cheloniads, the most remarkable characters are, that the facial line is horizontal, and quite continuous with the cranial line; that the orbits, though complete without, are continuous behind with the temporal fossae ; that the parietal and occipital bones are compressed laterally, while the latter terminates above in a sharp spine, projecting behind. The occipito-parietal and occi- pito-temporal sutures are distinct. The cranial cavity is small compared with the volume of the scull. These characters are not less remarkable in the Saurial or Lacertine Reptiles. The cranium of a crocodile mea¬ suring from 13 to 14 feet is scarcely capacious enough to admit the thumb ; and Cuvier estimates the area of the cranial section, which is oblong, at about of that of the whole head. In these animals, indeed, the bones of the superior and inferior jaws are so much prolonged, and oc¬ cupy so large a proportion of the head, that small space is left for the proper cranial cavity, which indeed is an immediate continuation of the vertebral. In these ani¬ mals, also, the anatomist can trace, much more distinctly than in the more perfect, the resemblance between the cranial bones and the vertebral. In the Cheloniads, and Saurial especially, the occipital bone is very distinctly a cephalic vertebra. Still more manifest is this arrangement in the Ranine Compara, or Batrachoid and Serpentine or Ophidial Reptiles. tlve In the former, as exemplified in the frog, the occipital bone, which forms the posterior cranial vertebra, consists of four pieces, and has two articular processes. The middle cra¬ nial vertebra is represented by the parietal bones above and the posterior part of the sphenoid below, while be¬ tween it and the occipital or posterior is contained the temporal as the organ of hearing. The third or anterior cranial vertebra is represented by the anterior part of the sphenoid bone below and the two narrow frontal bones above. The face, which may be regarded as the organ of the senses, is elongated anterior to the head, somewhat after the manner of the Cheloniad family; while an approximation to the Birds is indicated in the articulation of the lower jaw, which is connected to the head by the intervention of a quadrilateral bone. In the Serpentine family, the cranium of which is very similar in other respects, the most remarkable deviation is in the want of ethmoid bone. .The lower jaw is con¬ nected to the cranium by an intermediate bone, corre¬ sponding to the quadrilateral, but of an oblong shape, and something like a collar-bone. The chest of the Reptile class varies much in the Chest, mode of formation. While true ribs are recognised in the Saurial family only, the Batrachoid reptiles have a sternum without ribs, the Serpentine ribs without ster¬ num, and the Cheloniad ribs united into the dorsal shell, and a sternum expanded into the abdominal one. In the Saurial family the ribs correspond in number Ribs, to that of the costal vertebrae already mentioned, that is, 12 in the crocodile and iguana, two of which are not con¬ nected to the sternum, 17 in the chameleon, 18 in the tupinambis, and 27 in the monitor. The Saurial rep¬ tiles, however, are peculiar in having from 1 to 6 ribs at¬ tached to the cervical vertebrae, and the opposite ends of which are not connected to the sternum. These, which have been named cervical ribs, form a transition to the rudimental ribs of the Serpentine family, which are larger in the neck than elsewhere. The sternum of the crocodile consists of two parts,—an anterior or thoracic, which is osseous, supporting the two collar-bones,—and a posterior or abdominal, which is cartilaginous, and extends to the pubis, and furnishing to the abdominal parietes eight cylindrical cartilages. In the East India crocodile it appears that these lateral processes are converted into a single broad piece of cartilage on each side. (Fig. 4.) The ribs of the Cheloniad family are represented by Dorsal and the dorsal shell, which consists of eight broad incurvated sterno-ab- plates, identified behind with the dorsal vertebra;, and terminating before in the margin of the shell, and which are doubtless genuine ribs. In the ordinary land-tortoise ( Testudo Or (tea) these are seen in the shape of elevated bony ridges, proceeding from the head of each rib in a trans¬ verse concave bend to the margin of the dorsal shell. On each side of these ridges the bone is depressed, and is united at its lowest point by a genuine suture with the ad¬ joining ones. These sutures, however, are not continuous with those of the sterno-abdominal shell, but meet it in the intermediate points. (Fig. 5.) The sterno-abdominal shell consists, in like manner, of several transverse pieces consolidated into one. The ordinary number is eight on each side of the mesial plane, and a ninth azygous, gene¬ rally placed in the centre of the shell. In a specimen, however, of the tabular tortoise ( Testudo tabulata), in our possession, the number of the sterno-abdominal pieces is 11, of which 8 are in pairs, united on the mesial line from before backwards, and 3 azygous at the posterior tip of the shell. In young animals it is easy to recognise the unions of these constituent bones, which consist of sutures ex- ANATOM Y. 15 ‘ompara- actly similar to those of the cranium in the Mammalia. tive So feeble is the union, that it often happens that the ab- inatomy. s]ieu especially separates at the lines of junction, in the attempt to detach it from the dorsal. The Batrachoid Reptiles, though void of ribs, are provided with a sternum, which before is a cartilaginous process, terminating on a disc placed below the larynx, where it receives the collar-bones, and forms behind a broad plate placed below the abdomen, and giving attach¬ ment to the muscles. In the Salamander tribe, which are without sternum, the ribs consist of twelve pair of small rudimental processes, articulated with the vertebrae, but admitting of very limited motion. Lastly, in the Serpentine family, though there is no sternum, the upper vertebrae are provided with costal processes, quite rudimental. The great number of these costal rudiments, amounting in the rattlesnake to 175, in the cobra di capello to 192, in the coluber natrix to 204, and in the boa constrictor to 252, and the freedom of their anterior extremities, enable the animals of this tribe, wdiich are destitute of locomotive members, thoracic or abdominal, to employ the spinal column and the ribs as organs of progressive motion. On this point the reader will find some interesting observations by Sir Everard Home (Phil. Trans. 1812, p. 163). In the region of the neck, where the ribs acquire peculiar length, they are employed in erecting that region, and producing the ex¬ pansive swelling peculiar to this tribe of animals. It is an important link in the same series of facts, that in the animal absurdly named the flying lizard (draco volans), the five posterior ribs are recurvated and elongated to form the bony skeleton of the membranous sails by which the ani¬ mal supports itself in its desultory flight from tree to tree, rhoracic It is in the Saurial family that the locomotive extre- Ixtremi- mities of Reptiles ought first to be studied. In these we find an elongated scapula without spine, and one short flat bone, constituting the clavicle, united to the sternum. In the iguana and chameleon this bone is broad and nearly quadrilateral, while in the tupinambis it is large and oval¬ shaped, with its greatest length from before backwards, ■and with two unossified points. In the Ranine tribe, while the scapida consists of two articulated pieces, the upper towards the spine, each shoulder is provided with two collar-bones attached to the two extremities of the sternum, and the two anterior of which correspond to the bifurcated bone of birds. The sternum, collar-bone, and first part of the scapula, form one piece. In the salamander, in which the same conso¬ lidation is observed, the scapular portion is most distinct and directed to the spine, while of the clavicular portion the part connected to the sternum stretches below the chest, but, without uniting with that of the opposite side, the right glides over the left,—an arrangement which fa¬ cilitates the dilatation of the chest during inspiration. A nearer approach still to the bifurcated bone than is seen in the Ranine may be recognised in the Cheloniad family. In these animals three bones are united to form the humeral cavity. The first is a flat, trilateral bone, situate below the abdominal and thoracic viscera, close to the abdominal shell, and which, notwithstanding its situa¬ tion, is evidently the scapula. The second is a bone about the same length, flat, and like the feather of an oar at one extremity, which is free, round in the middle, and flatten¬ ed in the opposite direction at the other end, which is firmly united at a right angle to a long slender cylindrical bone. At the angle of union of these two bones is part of the glenoid cavity, which is complete in the small end of the scapula. The first of the two bones is the collar¬ bone proper; the second is the lateral branch of the bone, which forms the bifurcated, and which is occasionally united with its fellow. (Plate XXXIV. fig. 5.) The abdo- Compara- minal shell we have already stated to represent the ster- tive num or breast-bone. Anatomy. The humerus in the Saurial and Cheloniad family is arched and incurvated in a serpentine direction. It is articulated with a radius and uhia, which are succeeded by three rows of carpal bones, one row of four metacarpal bones and digital phalanges, varying in number in differ¬ ent genera. In the skeleton of a fossil animal belonging to the Saurial tribe, originally delineated by Collini, and Fossil ske- afterwards by Cuvier, and named by him the Pierodac- }e.ton tyle or Wingtoe (Pterodactylus, Ossemens Fossiles, tome^11^ v.), the metacarpal bone and phalanges of the index are '" J “ prolonged to about twenty times the ordinary length, for the purpose, apparently, of giving attachment to the mem¬ branous web by which the animal occasionally elevated it¬ self into the atmosphere. This animal, which, like the dra¬ gon (draco volans) of modern times, must have combined the contradictory characters of a flying reptile, may be re¬ garded as forming the link between the Reptiles and Birds, as the Ichthyosaurus does between Reptiles and Fishes. In the pelvis of the Cheloniad family it is remarkable Pelvis, that the pubal and iliac bones appear to change places. Thus the ilium on each side is a narrow bone proceeding backwards to the sacral part of the spine, which is re¬ ceived between its posterior aperture; while theap¬ pears in the shape of a broad, trilateral, flat bone, uniting before With its fellow on the mesial plane, behind with the ilium, and below with a flat, thin, quadrilateral bone, cor- • responding to the ischium, with which it forms the oval aperture. The inner of these three bones presents, as usual, the cotyloid cavity. It is further to be observed, that the two iliac bones, and consequently the whole pel¬ vis, are movable on the vertebral column. (Plate XXXIV. fig. 5.) In the Saurial Reptiles the pelvic bones are arranged Pelvic ex- and shaped nearly as in the Cheloniad. In the Ranine tremiues. the iliac bones are much elongated, and the pubal and ischial are consolidated into one piece, the symphysis of which forms a rounded crest. The femur is short, thick, and incurvated sinuously, with the convexity before towards the tibial end, and the con¬ cavity towards the pelvic. Trochanters, though present in the Cheloniad, are wanting in the Saurial and Ra¬ nine Reptiles. In the leg we find both tibia and fibula distinct, and of nearly equal size, in the Cheloniad and Saurial family, but conjoined in the Ranine family. The tarsus consists of five bones, and sustains four or five metatarsal ones, on which are supported three rows of phalanges. The metatarsal bones, which vary in length, are longest in the crocodile and others of the Lacertine tribe. In the Ranine, again, the astragalus and calca- neum are the bones of greatest proportional length. The anatomical characters now enumerated are proper to the skeletons of Reptiles at present existing on the surface of the earth or in its waters ; and in these we find a gradual transition from the Saurial and Cheloniad, by means of the Serpentine, to the finny inhabitants of the ocean. Even the Batrachoid Reptiles, in the early period of their existence while tadpoles, we shall have occasion to see, approach to the Fishes ; and in one sin¬ gular genus, if not two, the Proteus anguinus and Siren Lacertina, the characters of the Reptile are combined with those of the Fish, in having at once lungs or internal respiratory cells, and gills or external ciliated branchiae. The transition thus indicated is still more strongly de¬ monstrated in the osteological characters of two Genera of animals now extinct, so far as is yet known,—the Ich¬ thyosaurus and the Plesiosaurus. 16 ANATOMY. Compara* From the specimens of the Ichthyosaurus hitherto dis- tive covered, it appears that the number of vertebrae varies from . on^i 80 to 90 or more; in one entire specimen they amounted to 104 (Conybeare and De la Beche); that they are flat- cal pecifli- tened, with the transverse diameter greater than the longi- arities of tudinal, and the two articulating surfaces of the bodies the Ich- calycoid or cup-shaped as in Fishes. Though the annular thyosaurus part is distinct from the body, it is united to its sides. The spinous processes, which are long and prominent, form a continuous ridge above the spine, and are connected to each other by a process from the front of the one spine, which is inserted into a pit in the back of the other. In¬ stead of proper transverse process, a certain number of the vertebrae are provided with two tubercles on each side of the body, of which the superior, convex, is articu¬ lated to the tubercle of the rib, while the other, which is concave, receives the head. In the inferior part of the vertebral column, these two tubercles, after approximat¬ ing, are eventually identified into one. The ribs, which are numerous, and extend from the occiput to the pelvis, are slender and trilateral in shape, bifurcated above, and attached to the vertebrae by a head and tubercle. In the perfect specimen of Mr de laBeche they amount to 31, and of these 17 appear to be cervical or anterior false ribs, with single tubercles; thus affording another mark of resemblance to the Saurial family in osteological characters. The bones of the head, distinguished by the extraordi¬ nary size of the orbit, are similar to those of the Saurial Reptiles. The sternum, collar-bone, and scapula, though also similar to those of this family, bear a much closer re¬ semblance to the figure of these parts in the Echidna and Ornithorhyncus. The humerus is short, thick, and sinuat- ed; the bones of the fore arm flat, and probably constitut¬ ing part of the fore or thoracic fin. The Carpus consists of three rows, the first containing three bones, the other two, four each. These are followed by five or six rows of flattened, irregularly cuboidal bones, gradually diminishing in size and number to the tips, and which represent at once the metacarpus and phalanges of the fore paw, used appa¬ rently chiefly as a fin or paddle. The pelvic extremities appear to have been less strong and perfectly construct¬ ed than the thoracic. The femur is smaller and shorter than the humerus ; the tibia and fibula are flattened like the ulna and radius ; the tarsus consists of two rows only, the first containing three, and the second five bones ; and this in like manner terminates in five ranges of flattened bones, gradually diminishing in size, and which represent the metatarsus and metatarsal phalanges of the hind naw or paddle. and Pie- From the specimens hitherto discovered of the Plesiosau- siosaurus. ^ jt appearS that the total number of vertebra? amounts to 90, of which 35 appear to be cervical, while the other 55 are dorsal and caudal, the regions of which are propor¬ tionally short. The head of this animal also is small and compressed, nor has it the large orbit of the Ichthyosau¬ rus. Each rib consists of a vertebral and sternal portion, united at an obtuse angle, the former articulated by a single head to the transverse process, and the latter con¬ nected with its fellow by a transverse slip, so that the lower or abdominal ribs appear to have surrounded the abdomen with a complete cincture. The anterior part of the chest is occupied by two trilateral bones uniting in the middle, which, from their connection with the scapula, are believed to be the coracoid bones; and above these is a transverse piece, with a middle notch and lateral sinuated elevations, which is regarded as the sternum; while the scapula extends on each side like a buttress be¬ tween the two. It is. not improbable, nevertheless, that the middle portions named coracoid bones are the ster¬ num, and the transverse bone the clavicles; and it is Companj " worthy of remark, that not only this bone, but the middle tive ^ piece, closely resembles in figure and disposition those of Anatom1f the Echidna and Ornithorhyncus. The pelvis consists of' three bones, a vertebral or superior, corresponding to the ilium, narrow and slightly incurvated ; an anterior, ascend¬ ing forwards, and broad, separating the pubis ; and a pos¬ terior, short, forming the ischium. The humerus and femur are longer than in the Ichthyosaurus. There is a very short radius and ulna, and tibia and fibula, articulated with five carpal and tarsal bones; and the rest of both paddles consists of successive rows of flattened but long bones, contracted in the middle, and expanded at the ex¬ tremities, representing the metacarpal and metatarsal di¬ gital phalanges. (Home, Phil. Trans. 1816, 1818, 1819, 1820; De la Beche and Conybeare, Geological Transac¬ tions, vol. v. p. 559 ; and Cuvier, Ossemens Fossiles, vol. v. I part ii.) I r; SECT. IV. OSTEOLOGY OF THE FISHES. ' The Serpentine or Ophiad Reptiles present in their osteological characters an approximative transition to those of Fishes. While in the former order the skeleton is reduced to the spinal column, ribs, and head, in the latter class the spine and head only are left; and in some tribes the transition is still more distinctly marked by the presence of ribs. The vertebra of a fish is distinguished from that of any Vertebra other animal by the shape of its body. The cephalic and caudal, or anterior and posterior surfaces, are hollow cup¬ like cones, so that the union of each two vertebrae forms a double conical cavity, united by the base, containing a substance composed of concentric fibro-cartilaginous layers, with intermediate albuminous or gelatinous matter. By this cartilage the vertebral bodies are united; and on this the motions of the spine are effected. This motion, how¬ ever, is chiefly lateral; for the spinous processes are so long, and the articulation so complex, that antero-poste- rior inflection or extension is nearly impracticable. In the cartilaginous fishes, for instance the shark, stur¬ geon, and lamprey, the vertebral bodies form simple tubes, which, from the extreme elasticity of the constituent car¬ tilage, propel the contained fluid to a considerable dis¬ tance. Thus Sir E. Home saw the fluid projected to the height of four feet from the intervertebral cavities of the shark. {Phil. Trans. 1809.) In this order, also, the spine is infinitely more flexible, and its resilient power, when bent by the muscles, is almost incredible. On each T'01 side, also, the vertebrae are excavated, to form a canal for lodging the large blood-vessels. Ihe vertebrae of fishes are numerous, and not easily distinguished into classes. They may, however, be dis¬ tinguished into two, according as the spinous process is above only, or above and below at once. Those with the dorsal spine only are denominated dorsal or abdominal verte¬ bra, and have commonly at the sides transverse processes |>e for the attachment of the ribs. Those with the dorsal and ventral spines are distinguished as the caudal verte¬ bra. The last caudal vertebra is generally trilateral, flat¬ tened in the vertical direction; and its tip is marked with articular pits, which indicate the attachment of the small elongated bones which sustain the caudal fins. Ihe number of vertebrae varies. In the uranoscopus or star-gazer there are only 25, in the balista 17, and in the four-spined trunk-fish (ostracion) only 13; while in the sturgeon the number is 84, in the eel 115, and in the shark 207. Though Fishes have no chest, and require none, since their respiratory organs are gills, all of them are not void of ribs. The ray, shark, syngnathus, tetraodon, diodon, cy- ive Otti V\ ebrs ■arj ANATOMY. 17 ompara- tive natomy. clopterus, jistularia, «Src. have indeed no vestige of rib. But in the sturgeon, balista, eel, uranoscopus, pleuronectes, sea-wolf, and dory, they are in the shape of short rudi- mental processes; in the trigla and loricaria their sides are |ie head, anium. posterior margin of their aperture. This belt consists of Compara- a single bone on each side, articulated to the posterior- tive superior angle of the cranium, and uniting below the Anatomy, breast with that of the opposite side. This bone, which horizontal; in the perch, carp, pike, and chetodon, they may be regarded as a scapula, varies in shape and the f^ctoral encompass nearly the whole upper region of the abdomi- angle which it forms with its fellow in different species. ' nal cavity ; and, lastly, in the silver-fish {zeus vomer), the In fishes flattened vertically, the angle of union is acute; herring, rhomboidal salmon, &c. they are united to a in those which are depressed, the angle is so obtuse as to sternum. In the little animal named sea-horse (syngna- form nearly a straight line. In many fishes, especially thus hippocampus), several series of osseous tubercles of those of the order Thoracici, e.g. pleuronectes, coitus, zeus, the skin, surrounding the body like belts, are supposed to chetodon, perch, &c., in the small unicorn (balista), and others, the superior part forms a large spine, which de¬ scends immediately behind the fin, and to which the ad¬ ductor muscles are attached. This spine, which is mov¬ able, has been improperly named a clavicle. The rays by which the membrane is supported are not represent false ribs. The sternum is limited to a small number of Fishes. Besides those already mentioned, in the dory there is a series of minute flat bones disseminat¬ ed along the lower edge of the belly, which is supposed to represent a rudimental sternum In size and number the ribs vary, though in the silurus, directly articulated to this belt, but are connected by a carp, and chetodon they are of largest proportional size ; row of minute flat bones, which may be compared to the in the herring they are as fine as hairs. carpus in the other three classes. When the first ray of The head in the finny tribes is more an object of zoolo- the pectoral fin, however, is thorny, as in the harness-fish gical than anatomical description. The chief points to be (loricaria), and some species of silurus, it is articulated di- remarked are, that the cranium forms but a small part of rectly with an osseous belt; and it is remarkable that some the head; that the orbits are separated by a some- fishes, as the silurus and stickle-back, have the power of times membranous, occasionally, as in the wolf-fish, bony; retaining this spinous ray erected against the body as a and that there is on each side a large movable bone, cor¬ responding to the quadrilateral of Birds, not square, however, but oblong, which supports not only the lower jaw and palatine arches, but the gill-cover. In the car¬ tilaginous fishes the sutures are early obliterated, and the cranium consists of an inseparable mass of cartilage. In the bony fishes the cranium is separable into numerous pieces, and in the perch they amount to 80. In the cra¬ nium of fishes the anatomist recognises more distinctly than in the superior orders the formation according to the vertebral type. Small in proportion to the whole head, the cranium appears like a direct continuation of the ver¬ tebral column. In the osseous division of the class es¬ pecially, the cranium may be distinguished into the occi¬ pital or posterior vertebra, the spheno-parietal or middle, means of defence. This is effected by a cylindrical tu¬ bercle, on which the spinous ray is articulated by a hol¬ low, bounded before and behind by an elevated process. When the spine is erected, the anterior process, entering a hole in the cylindrical tubercle, is locked in it by the spine revolving slightly on its axis, so that it cannot be inflected unless by the spine revolving in the opposite di¬ rection. The pectoral fins are so long that they answer the pur- Pterygoid pose of wings in several species of trigla, the trigla hi- pectoral rundo, the flying gurnard (trigla volitans), the springing ^ns- gurnard (trigla evolans,) in the scorpcena volitans, the tropical flying fish (exoccetus volitans), and some others. Their situation also is liable Icomo- jRe mem- l(j-s or fins. to vary. In the exoccetus _ they are near the gills, but in the blennius and others they and the frontal or facial vertebra. I he cavity thus formed are remote. Lastly, they are totally wanting in a small is very small; yet small as it is, it is not exactly filled by number only, as the lamprey (petromyzon), the hag-fish the brain, between which and the bones there is inter- (myxine, Lin.; gastrobranchus), the murcenu, the eel genus, posed a pellucid fluid, contained in fine cellular tissue, the sphagobranchus, &c. I he cranium of the osseous fishes also is widest between The abdominal or ventral fins, which correspond to the Abdominal the ears, because the organ of hearing is contained within pelvic extremities of the other classes, are so denominated or ventral its cavity with the brain. In the cartilaginous it is quite because in the majority of fishes they are situate below different. the belly, and nearer the anal outlet than the pectoral. By 1 hough Fishes are destitute of extremities similar to this circumstance a numerous order are distinguished by the name of Abdominal Fishes (Abdominales). In a small number of fishes, comprehending the gadus, blennius, kurtus, callionymus, trachinus, and uranosco- those possessed by the other three classes of the Verte- brata, they are not, however, without locomotive mem¬ bers. The thoracic extremities are represented by the Ihtoral f pectoral fins, and the pelvic by the ventral. In short, pus, the ventral fins are placed under the throat, below it may be said that the bones of the thoracic and abdomi- the aperture of the gills, and before the pectoral fins, nal extremities are converted into osseous rays in the This order is therefore distinguished by the name of Ju- finny tribes. gulares. In the Ray genus, in which the wing-like disposition of In the most numerous order of all, the ventral fins are the pectoral fins gives the body a rhomboidal shape, they situate behind and below the pectoral fins. These have consist of numerous radiating cartilaginous lines, all at- therefore been denominated Thoracic Fishes (Thora- taehed to a cartilage parallel to the spine, divisible into two or three others, and articulated above to another ad¬ herent to the spine. Below there is a strong transverse bar common to the cartilages of both fins, and separat¬ ing at once the sternum and clavicle. This transverse bar is also seen in the shark tribe ; but their pectoral fins, which are much smaller, are not articulated with the spine cici). The ventral fins consist of two parts—one formed of rays covered by a double membrane, apparent externally, and constituting the proper ventral fin ; the other internal, representing the coxal bones of the pelvis, always support¬ ing the pinnal rays, and often articulated with the bones of the trunk. It is never articulated^ however, with the spine, nor does it form an osseous belt round the abdo- Iq the osseous fishes, and in many others usually re- men. The bones of which it consists are generally flat ferred to the cartilaginous division, e. g. the balista, the - tened, varying in shape, and in mutual contact by the in¬ pectoral fins are fixed to an osseous belt, which sur- ternal margin. In the shark and ray genera only is there rounds the body behind the gills, and which supports the a single transverse bone, nearly cylindrical, to the extre- vol. in. c 18 ANATOM Y. Compara- mities of which the fins are attached. The direction of tiye the pelvic plane to the walls of the abdomen varies ac- ^Anatomy. corc|jng shape of the body of the fish. In the fiat Ventral ^iey are directed obliquely, and their inner margin fins. forms the keel of the belly. In fishes with a broad or cy¬ lindrical belly they form a plane more or less horizontal. In the Jugular and Thoracic Fishes, the pelvic bones are always articulated with the base of the belt which sustains the pectoral fins; and they vary much in shape and situation. In the trachinus, uranoscopus, cottm, scicena, chetodon, and perch, these two bones are united by their inner mar¬ gin. In the cuckoo-gurnard, in which they are united by the posterior tip only of their internal margin, they are broad, flat, and oval. In the sole and flounder genus (pleu- ronectes), in which the fins are attached to their anterior tip, they are united in a quadrangular pyramid, the apex of which is directed backwards and upwards, and the base forwards. In some of the stickle-backs these bones are altogether separate, and being long, receive in their middle a movable spine, which supplies the place of the ventral fin. In the dory (zeus faber, L.) they are flat and triangular, in mutual contact by their whole surface. In the silver-fish (zeus vomer) they are small and cylindrical. In the whole of the Abdominal order, on the contrary, the pelvic bones are equally unconnected with the bones of the shoulder and with the osseous belt of the pectoral fins, and are confined to the middle-inferior part of the belly, not far from the anus. In general these two bones are separate from each other, and are retained in their si¬ tuation by ligaments. In the carp, in which they are elongated, they touch only by their posterior third. In the herring, in which they are small and approximated, they are continuous with the minute bones of the sternum. In the pike they are broad and trilateral, approximated by their anterior tips, separate behind where the fin is at¬ tached. In the silurus, in which they are united, they form a round and often spinous shield before, while the fins are attached to the exterior-posterior margin. Last¬ ly, in the cuirassier or harness-fish (loricaria), the pelvic bones are united in one piece, the posterior notch of which forms the anal aperture, while the fins are attached to its external margin. The proper fin consists of a certain number of osseous rays, simple or bifid, supported by one or two rows of mi¬ nute bones placed between them and the pelvic bones. On these small bones the constituent rays move, diverging or converging like the rods of a fan, while the whole fin may be inflected or extended by the minute bones moving on the pelvic, so as to adduct or abduct the fin. In the cartilaginous fishes the structure is different. To the tip of each pelvic bone are articulated two principal cartilages, one external, forming a kind of toe with seven or eight joints; the other internal, supporting all the other rays of the fin, which often exceed thirty in number. Analogy or If we suppose these bones, like the minute ones of the unity of pectoral fin, to represent the tarsus of the other three Principle in c]asseS) must follow that, in the locomotive extremi ties, 3msa- the humerus, with the ulna and radius, and the femur, with the tibia and fibula, are obliterated. It is not un¬ important to observe, that the general structure of the Vertebrata tends through various transitions to this ter¬ mination. In the Amphibia the long bones of the extre¬ mities are shortened by removing the diaphysis, and leaving their articulating ends only. In the Cetacea the pelvic extremities are removed altogether. In the Cheloniad and Saurial Reptiles the same long bones of the extre¬ mities are much abridged; and in the Ichthyoid Rep¬ tiles, now extinct, but sharing by their structure a form of animal existence partaking of the reptile and fish at once, and perhaps intermediate between the two, this ab- Compara. t breviation is carried perhaps to its greatest possible degree, bve in leaving the articular ends only of the four locomotive '^nat0U1^ A extremities. Lastly, this reduction is merely prepara-P tory to that exhibited in the whole class of Fishes, in which the three longitudinal bones so conspicuous in the higher classes of animals are completely obliterated, and those representing the hand or forepaw and foot are arti- v culated directly to the shoulder and pelvic bones. L Besides the bones already mentioned as constituting ti the skeleton, there are observed in the osseous fishes n minute bones, generally fork-like in shape, disseminated through all the muscular parts of the body. The purpose of these bones, which, as being totally insulated from the other parts of the skeleton, are denominated ossicula mus¬ culorum, is chiefly to afford points of support; and they are probably to be regarded as rudimental representatives of osseous parts, more completely developed in the higher animals. It is further a curious circumstance, that the skeleton, Violation which is so symmetrical in all the other classes and orders, of the law begins to exhibit a deviation from this first in the skele- ton of the finny tribes. In the Sole genus (Pleuronectes) this deviation is very conspicuous. Both eyes are placed on the same side of the mesial plane; and the side on which the eyes are placed is broader than the opposite one. The former is bounded by a convex margin, the lat¬ ter by a concave one. The orbit towards the former is large, the other small and imperfect. Conversely, it is to be observed, that in the latter the maxillary and intermax¬ illary bones are larger than in the former. The sides of the inferior jaw are less discordant; and though in the Sole and Plaice those of the eyeless side are more straight and elongated than those of the other, in the Turbot (Pku- ronectes maximus) they are nearly symmetrical. CHAP. II.—COMPARATIVE MYOLOGY. Though this is the proper place to consider the pecu¬ liarities of the muscular system of animals, the limits as¬ signed to this sketch will not allow us to enter into de¬ tails. We shall merely, therefore, take a cursory view of those points in which the myology of the lower animals differs from that of the human subject. In general, in the lower animals, especially the Mam¬ malia, Birds, and Reptiles, the muscles correspond in situation to those of the human subject; and whatever modifications they undergo consist in changes of figure, and in some few instances in changes in attachment. The former kind of changes may be in all cases pretty accu¬ rately estimated by the osteological characters of the class, order, or genus; for when the position, shape, or direc¬ tion of a bone is altered, in the same proportion nearly are the attached muscles altered in their attributes. Though in the lower animals, however, the zootomist Deficiencr traces muscles in general quite analogous to those of the in number, human subject, in several instances this analogy ceases to be observed. In general the muscles of the lower animals are less numerous than those of the human subject; and this deficiency in number, though not much observed in the Quadrumana, is very remarkable in all the inferior orders of the Mammalia, and still more in the Birds and Reptiles. In general, also, these variations are most conspicuous in the locomotive extremities. Thus the small pectoral muscle, which is present in the Quadru¬ mana, is wanting in the Carnivora and the whole of the Ungulated Animals and the Reptiles. The short supinator is present in the Canine and Feline genera, but the long is wanting; and both are absent in the Chirop- tera, Rodentia, Pachydermata, Ruminantia, and M IF iir II ANATOMY. 19 )]«n v\ )1 il 0 ion la« m. « ii n Jompara- Solidungula, and in the whole class of Birds. Both tivc pronators (teres and qimdratus) are present in the Qua- drum ana and Carnivora, but wanting in the Chirop- tera, Ruminants, and Solidungula. The Rabbit, and perhaps the Rodentia generally, have the pronator teres ; but as the radius is not very movable, its influence is trifling. fyological In the mole the rhomboideus is inserted into the cervi- pculiari- cal ligament, which is ossified; and it therefore elevates esofthe head and neck on the scapula with singular force. ,0 e‘ This is effected still more remarkably by the occipital part of the rhomboideus, the fibres of which being parallel to the spine, pass below the proper rhomboideus to be attached to the transverse ligament and the middle of the cranium. The strong, thick, quadrangular collar-bone has two muscles, a supradavius and a subclavius. The large pectoral is very thick, and nearly as large as in birds. The common extensor of the fingers or fore toes is the only muscle which is common to man and all the quadru¬ peds. Of the proper extensors the horse has two on the side of the common extensor, but acting as an extensor of the fore pastern ; another between the common exten¬ sor and the extensor of the pastern, and which seems merely an appendage to the former. The proper extensor of the index is wanting in the Rodentia, Ruminants, and Solidungula ; and while the two latter orders are destitute of the long and short extensors of the thumb, and the feline, canine, ursine, and leporine have the former, they are destitute of the latter. Lastly, the lower animals are wholly destitute of the short muscles of the hand, which in man produce flection, abduction, adduc¬ tion, and opposition. In the Ghiroptera only is there one extensor, and flexors of the fore toes. Among the muscles of the pelvic extremities the glu- tmus maximus, or large muscle of the buttock in man, di¬ minishes much in the Quadrumana ; and in the other orders is reduced to a very small size. The buttock in the Mammalia generally consists chiefly of the glutceus mediiis and minimus ; and while the glutceus maximus is in the horse in a great part aponeurotic, the g. medius is so large as to produce those forcible and sudden extensions of the hind leg which constitute the kick. In the leg the sartorius of the horse, the animal in which the muscles have been most studied, is large, and is distinguished by the name of the long adductor, in op¬ position to the gracilis, which constitutes the short ad¬ ductor. The muscle representing the biceps of man is in all quadrupeds a uniceps, and the single head is attached to the ischium only. In the horse and dog it has been denominated the vastus longus. The gastrocnemius externus etinternus (gemellus), which constitute the calf of the human subject, diminish consi¬ derably in the lower animals; and the solceus, which is placed belotv them, also becomes small, and is particu¬ larly slender in the Ruminants and Solidungula. luscles The following muscles are wanting in the whole class ■ birds! ®IRDS• The diaphragm, the recti abdominis, and the T ‘ pyramidales; the muscles of the dorsal part of the spine, the splenitis, the brachialis externus, or third head of the triceps ; the supinators of the fore arm or wing, as already mentioned, all those corresponding to the short muscles of the hand and fingers; the quadratus lumborum, the psoas parvus, the psoas magnus, iliacus internus, obturator externus, and the extensor longus pollieis pedis. Two muscles, which occupy the situation of the prona¬ tors, act as flexors, showing the connection between the actions of inflection and pronation, and the occasional substitution of the latter for the former. In tins class, also, the glutccus maximus is of a pyra¬ midal shape, while the true pyriforinis is wanting. The glutceus minimus, which is attached to the anterior edge Cotnpara- of the iliac bones, is the iliacus. In place of the pectineus tive there is a slender muscle, which extends to the knee, Anatomy. over which its tendon passes, and gliding behind the leg, its tendon is bifurcated, one slip going back to be inserted into the posterior part of the metatarsus, the other to be united to the perforated flexor of the first and last toe. This muscle, which is named the accessory femoral flexor, is the one by which birds are enabled to clasp a perch during sleep. In Birds the great pectoral is a remarkable muscle in Muscles point of size. It consists indeed of three muscles, thelis^hi large pectoral, the middle, and the small, which occupy 'in£’ the sides of the vertical crest of the sternum, and consti¬ tute what is named the breast of the animal; and which are chiefly employed in the energetic motion of the wings in flying. These muscles are sometimes so large that they weigh more than all the other muscles of the animal together. In birds which fly much they are dark coloured and firm; in those which fly little, as the domestic poul¬ try, they are white coloured, and in general soft. The same distinction is observed in the muscles of the two ex¬ tremities. In birds much on the wing these muscles are dark coloured and firm, while those of the legs are com¬ paratively lighter and more tender; and, conversely, in birds little on the wing and mostly on the legs, as the domestic poultry and many of the Grallce, the waders, swimmers, &c. the muscles of the wings are light coloured and tender, while those of the legs are dark coloured, firm, and strong. The flexor muscles of the leg and toes of Birds merit Mecha- notice. They consist of muscles corresponding to thenism f long flexors, which are divided into three masses. Thefiercain«' first consists of five portions, three of which may be re¬ garded as constituting a single common perforated flexor. It rises by two bellies, one attached to the external con¬ dyle of the femur, forming a perforated tendon, which receives one of those of the muscle corresponding to the peronceus; the other to the posterior surface of the fe¬ mur, forming the tendons of the index and small toe. This muscle is further connected by intermediate fibres with the accessory femoral flexor,—a muscle placed on the internal surface of the thigh, and sending its tendon over the knee; and as the tendons are inserted into the unguinal phalanges, when the accessory femoral bends the thigh the flexors of the toes inflect them also, and retain them in the inflected position. By means of this arrangement birds are enabled to clasp a perch or other small body when roosting, without continued muscular effort, and thereby to sleep on the perch. This mode of explanation, which was originally given by Borelli, has been controverted by Vicq d’Azyr; but apparently not On good grounds. Among the class of Reptiles, while the muscles of the Ophidial family are confined to those of the vertebra: and rudimental ribs, in the Cheloniad these are oblite¬ rated, and the muscles of the neck, head, and tail, and those of the locomotive extremities alone, are left. In the other two classes of reptiles the muscles are in gene¬ ral analogous to those of the Mammalia. There are nut many instances of muscles which, though Cutaneous unknown in man, are found in the lower animals. Ofmuscle. these the most remarkable are the cutaneous muscle (pan- niculus carnosus), and the suspensory of the eye. The former was absurdly maintained to exist in the human subject, especially by Nicolaus Massa ; but it is manifest that the assertion was derived from the dissection of the lower animals only. It was not long after demonstrated by Charles Etienne, that no fleshy pannicle or cutaneous muscle exists, sUch as is found in the lower animals; ami ANATOMY. 20 Compara- tlmt the only cutaneous muscles in man are the latis- Uve simus colli, the epicranius or scalp-muscle, and those Anatomy, are attached to the face, and which by their mo- tion give expression to the countenance. The cutaneous muscle even is not found in the Quadrumana, nor does it exist in the pig. In various other animals, however, it is found in different degrees of distinctness. It is very well marked in the hedgehog and porcupine;—by its means they have the power of erecting their spines, and rolling themselves up;—and in the armadillo and the ant- eater tribe. In the mole, also, we have seen it pretty well marked. Suspen- It is an interesting fact, that Galen originally observed sory mus- that the lower animals possess a seventh muscle of the cle of the eyej or one more than man. The suspensory or infundi- to certain1 hular muscle (musculus choanoides), as it has been named, animals. fr°m its shape, especially in the Ruminants and Soli- dungula, has the apex fixed to the margin of the optic hole, and its base inserted a little behind the four straight muscles. In the Zoophaga and Cetacea it consists of four parts, so that these orders appear to be provided with 8 straight muscles. In the rhinoceros it consists of two portions. Tail, There is yet another part, the muscles of which can scarcely be said to exist in the human subject, but which attain a very great degree of developement in the lower animals. The coccyx of the human subject is expanded in the lower animals into a highly flexible prolongation denominated the tail (cauda), variable in length, but al¬ ways consisting of separate vertebrae, articulated and movable on each other. While the coccyx of the human subject possesses two muscles only, the ischio-coccygeus and sacro-coccygeas, which are so insignificant in size that they scarcely serve to move the part, the caudal verte¬ brae of the lower animals are moved by muscles greatly larger, more numerous, and more powerful. The tail is to animals a much more useful and power¬ ful organ than the coccyx to man. It is a member which peculiarly belongs to them; and though in ordinary cir¬ cumstances pendulous, it is made to assume a variety of motions of which no other organ is susceptible, and to perform duties which would be otherwise impracticable. With many, as the long-tailed monkeys, the sloths, the ant-eater, and the squirrel tribes, it is indispensable as an organ of prehension. The majority of animals, as the Ruminants, Solidungula, &c. use it as a whip or lash to drive away insects. The lion, tiger, and others of the feline tribe, lash their sides with it when enraged. The Cetaceous swimmers employ it as a rudder and oar in the waters. The beaver uses it as a trowel, to enable him to construct his clay-built dwelling. An organ employed so variously must consist of a muscular apparatus rather complex. and caudal The different motions of which the tail of the Mamma- muscles. lia is susceptible may be referred to three heads,—one by which it is extended or elevated, another by which it is inflected or depressed, and a third by which it is made to beat the sides. The combination or succession of these motions gives rise to secondary ones more complex in character. It may be twisted on its axis, or turned in a spiral direction. These motions are effected by three classes of muscles. ls£, The muscles which raise the tail are situate above ; they are muscidi sacro-coccygcei superiores. Commenc¬ ing at the base of the articular processes of the 3 or 4 last lumbar vertebrae, or those of the sacrum and the cau¬ dal vertebrae, by fleshy slips, they are connected to ten¬ dons, which are inserted into the base of the first of the caudal vertebrae, which are void of articular processes. The second tendon goes to the next following vertebra, and so on to the 13th, each contained in a ligamentous Conipara- groove, which forms an investment. The muscles of both tive sides acting together, elevate or incurvate the tail up- wards. The interspinalis and spinalis obliquus or lumbo-sacro- coccygeal are the continuations of the interspinales dorsi et lumborum. The spinous processes, however, becoming indistinct, or being represented by two tubercles, the at¬ tachments vary. 2d, The muscles which depress or inflect the tail down¬ wards take their origin within the pelvis, and are pro¬ longed to various extents along the inferior surface of the tail. Of these there are four pairs, the ileo-coccygeal of Vicq d’Azyr, the inferior sacro-coccygeal, the inter-coc¬ cygeal muscles, and the pubo-coccygeal of the same author. The insertions of these muscles vary in different genera, according to the number of vertebrae of which the tail consists. The pubo-coccygeal is wanting in the raccoon, but it is distinct in the dog and opossum. The effect of the ileo-coccygeal and it, is to depress the tail and apply it forcibly to the anus. 3d, There are only two muscles which carry the tail to the sides of the animal—the ischio-coccygceus externus, and the intertransversalis of Vicq d’Azyr; the former proceed¬ ing from the pelvic surface of the ischium below and be¬ hind the acetabidum, to the transverse processes of the cau¬ dal vertebrae, the second extending in a continued band between all the transverse processes. The tail, therefore, in the Mammalia, consists of a series of successively decreasing vertebrae, moved by eight pairs of muscles. In Fishes it is not easy to trace any analogy between the muscles and those of the other classes. Though the spine, head and fins, have appropriate muscular bundles, the natural or fascial distinctions are less evident than in the other three classes. It is important, however, to re¬ mark, that while the muscles which move the spinal co¬ lumn are placed in these classes, partly before, and chief¬ ly behind the vertebrae, those of Fishes are placed on each side. Hence the lateral motion of the spine, which is inconsiderable in Mammiferous animals, Birds, and Reptiles, becomes very conspicuous in the finny tribes, especially in the motion of swimming, while the antero¬ posterior inflection or extension is altogether insignificant. It is almost superfluous to remark, that, in the greater part of the finny tribes, the muscles are white or pale coloured. In a few only, for instance the salmon, trout, gwiniad (coregonus'), herring (ciupea harengus), carp (cy- prinus), and some others, are the muscular fibres of a pale flesh red. The circumstances on which these differences depend are not known; but it is supposed that in the latter sorts the proportion of oleo-albuminous matter is more abundant than in the former. The proportion of albumen, however, in the muscles of fish, seems in general to be small. They abound in gelatine and isinglass; and in some of the cartilaginous fishes especially, the greater part of the muscles seem to consist chiefly of gelatine in various degrees of consistence. This is particularly the case with the lamprey, the hag-fish (myxine glutinosd), and even with the sturgeon. The sterlet especially (aci- penser Ruthenus), a small species of sturgeon found in the rivers of Russia, both European and Asiatic, abounds in gelatine ; and the presence of this principle enables the inhabitants to use it in the preparation of a species of soup, the sterlet, which is esteemed a great delicacy. In some of the genus Pleuronectes this principle is also very abundant. Thus the Plaice {Pleuronectes Platessd), sole (P. iSb/m), and especially the turbot (P. Maximus), con¬ tain a considerable proportion of gelatine. On the pro¬ portion of this principle depends the quality of fish used CL. ^ ANATOMY. 21 Cmpara- as an article of food in nourishing without exciting. All tive fishes which abound in gelatine uncombined with oleo-al- atomy, buminous matter may be safely used as articles of food; while those in which the latter ingredients predominate are invariably eaten with the risk of disordering the sto¬ mach and producing indigestion. CHAP. III.—COMPARATIVE iESTHIOLOGY, OR THE COM¬ PARATIVE ANATOMY OF THE ORGANS OF SENSATION. SECT. I. THE ORGAN OF SMELL. The organ of smell consists of the nasal cavities, those of the ethmoidal and turbinated bones, and the frontal, sphenoidal, and superior maxillary sinuses, all of which communicate with the nasal. The whole of these parts are invested by fine periosteum, lined by mucous mem¬ brane. The ethmoid bone is the essential organ of smell; and the others appear simply to multiply the extent of the membrane. The ethmoid bone consists of a perforated plate, with a middle vertical one attached at right angles to it, and lateral portions composed of thin bony plates convoluted with various degrees of complexity and minuteness in Ipmoid different orders and genera of animals. These convoluted c4s. plates form what are denominated the ethmoidal cells. They may be represented as numerous tortuous canals, proceeding from the perforated plate forwards and out¬ wards, approximating mutually, and forming numerous communicating cavities. Such nearly is the structure of these plates in the Edentata, Ruminantia, Solidun- gula, Pachydermata, and Carnivora, the last of which have more complicated cells than the first. In the dog they are numerous and extensive. In the Rodentia, for instance the porcupine, they are few—not above 3 or 4 on each side. In Birds the internal side of each nostril is occupied by three orders of plates; the inferior turbinated or spongy bone; the middle, consisting of one plate convo¬ luted on itself two turns and a half; and the upper, shaped like a bell, adhering to the frontal and lacrymal bones. These form three tortuous passages, varying in size and tortuosity in different genera. Though generally carti¬ laginous, these turbinated bones are membranous in the cassowary and albatross, and bony in the calao and toucan. In the nostrils of Reptiles there are convoluted pro¬ minent plates, which, however, are merely membranous productions, unsupported by any bone. In the Fishes, in like manner, there are membranous folds, the disposition of which is tortuous. They are, however, more regularly arranged than in the other classes. In the cartilaginous fishes they consist of semi¬ lunar folds placed in parallel tracts on each side of a broad plate, which divides the one side of the nasal cavity from the other. In the sturgeon, however, they are ar¬ ranged in diverging plates, which are subdivided into more minute ones, like the branches of a tree. In the osseous fishes generally they consist of radiating plates disposed round a prominent central tubercle. In these three classes the olfactory nerve is distributed to the membrane much in the same manner. This nerve, however, does not proceed farther than the superior tur¬ binated bones; and the middle and inferior appear to be supplied with filaments of the fifth pair, the naso-oph- thalmic branch of which is distributed to the nose in all the vertebrated animals. In the Mammalia, further, the spheno-palatine ganglion sends several filaments to the posterior part of the narine membrane. By most zootomical authors the trunk of the elephant has been described as an organ of smell peculiar to that animal. We are satisfied, however, from observing the mo- Compara- tions of this body in the living animal, that it is an organ, tive not of smell, but of prehension. Cuvier, after adopting ^natomy- the ordinary view, has relinquished it, and, on the ground of personal inspection, admits that the sense of smell in the elephant is confined, as in other animals, to that por¬ tion of the nasal cavities which is contained within the bones of the head. The trunk of the elephant, therefore, will with greater propriety be noticed under a subsequent head. The nasal cavities of the Cetaceous animals are not so much organs of smell as channels of respiration, and must also be noticed afterwards. It is sufficient here to remark, that in these animals the part of the cranium cor¬ responding to the ethmoid bone is penetrated by no aper¬ ture, or, in other words, is not an ethmoid bone. It has therefore been asserted that the Cetacea have no olfac¬ tory nerve, and no sense of smell. This, however, is by no means established. Blainville and Jacobsen have ob¬ served in the dolphin nerves which they regard as olfacient; and Treviranus delineates nerves of the same character. By Otto and Rudolph!, on the contrary, who have had frequent opportunities of dissecting the dolphin and whale, the existence of these nerves is denied. Though almost all the invertebrated animals give proofs Smell in of the existence of the sense of smell, in none of them do inverte- we find any organ in which this sensation appears with brated uni- certainty to be exercised. That these animals possessmals’ the faculty of smell, is inferred from the fact, that insects recognise their food at a distance; that male butterflies scent the female even when inclosed in cages ; and that the ordinary flesh-fly deposits her eggs on tainted meat, and occasionally on fetid plants, in the belief that they are the proper nidus, though in the latter case the larvce perish for want of the necessary sustenance. Since odorous particles are evidently applied to the ol¬ factory membrane of all aeropnoic animals by the me¬ dium of the atmosphere, and since the organ of smell is therefore situate in connection with the wind-pipe, it was conjectured by Raster, that, in insects at least, the organ of smell is situate at the entrance of the tracheae or air- tubes. This conjecture derives some probability from the fact, that the inner tracheal membrane in these animals is soft and moist, and that those in which it is expanded into convoluted lacunae and tortuous vesicles, for instance beetles, flies, and bees, are remarkable for the nicety of their sense of smell. The antennae, in which this sense has been placed by some anatomists, appear to be rather organs of touch than of smell. In the Mollusca the whole cutaneous covering seems to combine the character of an organ of touch or tact, and of smell. Like an extensive pituitary membrane, it is soft, villous, moist, and liberally supplied with nerves. The Articulata and Zoophytes seem much in the same state. But on all these points information is rather con¬ jectural than positive. SECT. II. THE EYES ; THE ORGANS OF VISION. All red-blooded animals, without exception, are pro¬ vided with two movable eyes, consisting of the same es¬ sential parts as those of man, forming globular organs, and placed in the cranio-facial cavities named orbits. In none are there more or fewer; and the exceptions to the general rule, either in relation to the presence of these organs, or number, are only apparent. Among the Mammalia, Blind indeed, there are two instances of blindness,—in the though not or blind rat (Mus typhlus, Lin.; Spalax typhlus, Pall.), and eyeless the golden mole (Talpa Asiatica, Lin.; Chrysochloris,aniITials- Lacep. and Cuvier). But in neither of these animals are 22 ANATOMY. Anatomy. Compara- the eyes absolutely wanting; they are merely very mi- tire nute, and covered by a thin fold of hairy skin, in which there is said to be no aperture. Much in the same man¬ ner the murmna coecilia, and the hag-fish (inyxine, Lin., gastrobranchus emeus), though provided with eyes, are de¬ prived of the use of these organs by the opacity of the con¬ junctiva. In the Anableps (Cobitis Anableps, Lin.), the cornea and iris are biparted by transverse bands, so as to give the animal the appearance of having two pupils in each eye, though the crystalline lens, vitreous humour, and retina, are single. This animal affords the only ex¬ ample of this structure among the vertebrated animals; but a similar arrangement is observed in the Cephalopo- dous Mollusca and cuttle-fish family. The general shape of the eye depends on the medium in which the animal lives. It is nearly spherical, or ap¬ proaching the spherical shape, in man and the quadrupeds moving along the surface of the earth; that is, in the lowest and most dense region of the atmosphere. The cornea merely forms a slight convexity, in consequence of being the segment of a smaller sphere than the rest of the eyeball; yet in the porcupine, opossum, &c., this difference is inconsiderable. To show the degree of this convexity, it is merely requisite to compare the axis or antero-posterior diameter with the transverse diameter of the ball, as exhibited in the following table:— Figure of the eye- hall. Axis and diameter. Axis. Tr. Diam. Whale C 11 Porpoise 2.... 3 Owl 13 12 Vulture 13 16 Man and ape 137 136 Dog 24 25 Horse 24 25 Ox 20 21 According to the measurements of the younger Soem¬ mering, the axis of the human eye, taken in a beautiful Tyrolese girl of 20, is to the transverse diameter as 100 to 95; that of the eye of the magot (simia inuus) as 85 to 84 ; and that of the bat (vespertilio auritus) as 12 to 11. In the raccoon (ursns lotor) and lynx (fells lynx') alone the axis is exactly equal to the diameter. In all the other ver¬ tebrated animals, it is, as in the measurements of Cuvier, less than the transverse diameter at the rate of from 9 to 33 or 45 per cent. In the horse it is as 186 to 212, in the seal as 130 to 142, in the Indian elephant as 135 to 180, and in the black whale (balocna mysticetus) as 20 to 29. In the owl it is as 17 to 18, in the golden falcon as 14^ to 16, in the ostrich as 18 to 19^, and in the swan as 7 to 10. In the Reptiles and Fishes it is always less at the rate of from 3 to nearly 10 per cent. In the cuttle-fish, which may be taken as a general example of the inverte- brated classes, it is much greater, the axis being to the diameter as 80 to 57. (D. W. Soemmering de Oculorum Hominis Animaliumque Sectione Horizontali Commentatio. Goetting. 1818, fob) In Fishes and the Cetacea which inhabit the sea, the anterior part of the eyeball is much more flattened, and in many fishes it resembles a hemisphere with the plane surface before and the convex behind. In the ray genus the superior part is also flat, so as to give the eye the ap¬ pearance of the quadrant of a sphere, cut through two large circles perpendicular to each other. In some fishes, especially the burbot ( Gadus Lotaj, the cornea is convex. In Birds which occupy the elevated regions of the at¬ mosphere, the deviation from the spherical shape is in the direction opposite to that of fishes. On the anterior part, which is sometimes flat, sometimes shaped like a truncat¬ ed cone, is chased a short cylinder, closed by a very con¬ vex, and occasionally hemispherical cornea, always be- longing to a much smaller sphere than the posterior con¬ vexity. bonoiJaom vkuoiymo ad) ni fiedt !■ H • Aqueous These differences in shape depend on the proportion be- humour. tween the density of the medium in which the animals live and that of the aqueous humour. In the higher regions of Compara. the atmosphere, in which the air is very much rarefied, the tive refracting power of the aqueous humour is much more con- ^ato^7; siderable than at the surface, occupied by quadrupeds; ^ and hence it is more abundant in the former than in the latter class. Its refracting power, however, would be almost extinguished in a watery medium, from which it could differ but little in density ; and hence it is either trifling or absolutely wanting in the inhabitants of the deep. In the cuttle-fish family it is entirely wanting. The crystalline lens in Fishes, which is nearly spheri¬ cal, projects through the pupil, and leaves little room for the aqueous humour. The lens is also very convex in the Cetacea, the Amphibious Mammalia, the diving birds, as the cormorant, and the marine and aquatic Reptiles. Affecting the oblate spheroidal shape in the Mamma¬ lia, it becomes extremely so in man, and still more in Birds. Its consistence is greatest in animals in which it is most convex ; and hence it is matter of common obser¬ vation, that the crystalline of fishes is particularly firm. It also contains rather more albumen than the lens of the Mammalia. The crystalline lens occupies least propor¬ tional space of the eyeball in man, and most in fishes. The comparative spaces occupied by each of the hu¬ mours may be understood from the following table, in which the axis of the eye, or the space occupied by the whole three humours, is represented by unity. Aqueous Humour. Crystalline Humour. Vitreous Humour. •37 T? •if Man Log ..sV— 58t,......... OX Ml Sbeep t? T7-- Horse y......... Owl 8 II 8 VJVV1 57 57 57 Herring £ f * On the proportion of the total volume occupied by each of the three transparent parts there are few accurate facts. It may be remarked, however, that the human eye among the Mammalia is that in which the vitreous humour is proportionally most abundant. It is estimated to be twenty times more copious than the aqueous. In the ox it is only ten times, and in the sheep nine times the quantity of the aqueous. In the Mammalia generally, the sclerotic is compa-Sclerotic, ratively elastic, soft, and yielding ; but in all animals in which the eye deviates from the spherical shape, as the Cetacea, Fishes, and Birds, this membrane is strength¬ ened by greater solidity and thickness of tissue, or sup¬ ported by accessory parts of a hard unyielding structure. In the eye of the whale these two parts, the hard and soft, are naturally distinguished in a very striking mariner. The lateral parts of the sclerotic are nearly an inch thick, and very hard. The posterior part is about one and a half inch thick, and softer, because the spaces between the firm fibres are filled with oily substance. The posterior part presents for the optic nerve a canal one and a half inch long, the walls of whiph are formed chiefly by fibres in direct continuity with the dura mater,—-the only fact, it may be observed, which favours the statement of the ancient anatomists, that the sclerotic is derived from the dura mater. The sclerotic of the porpoise, though only two or three lines thick, has the same structure as that of the whale. In the seal it is thick before and thicker behind, but the middle zone is thin and flexible. ^ The sclerotic m Birds is thin, flexible, and elastic be-Osseous hind, with a bluish glistening aspect, and without distinct zone of fibres. The optic nerve enters, riot by a hole, but an ob-bhris? lique cqnal. The anterior part consists of two plates, be¬ tween which is enchased a zone of thin, hard, oblong, OSse- ’'it' ANATOMY. 23 Ompara- tive Jiatomy. ail rep- KS. fl rnea. (.meal ii) usijunc- t n. Cioroid ci'-nt and liyschian Wembrane. ous scales, varying in number from 11 or 12 to 14 or 15, imbricated over each other so as to give the anterior part of the eyeball a great degree of hardness, and a figure unsusceptible of change. Though these plates are nearly flat in most birds, and form an annular zone slightly con¬ vex, they are broad, arched, and concave internally in the owl gmus, and form a bell-shaped tube, with the posterior aperture oval and the anterior round. This may be de¬ nominated the osseous ring {annulus osseus, zona ossea). In the ostrich it is narrow and flat. Among the Reptiles, the Cheloniads possess an osse¬ ous zone, consisting of plates inclosed in the membrane without being continuous with its substance. They are also found at the lateral part of the sclerotic in the cha¬ meleon and some of the Saurial Reptiles, as the Croco- dilus Sclerops and Lucius, the monitor, and the iguana (D. W. Soemmering). It is also an important character in the structure of the eye of the Ichthyosaurus, which indicates the connection of that animal with the Saurial tribe, that its sclerotic was provided with an osseous zone, con¬ sisting, as in these, of 13 separate pieces. In Fishes the sclerotic is cartilaginous, homogeneous, semi-translucent, elastic, and, though thin, firm enough not to collapse. In the ray it is expanded into a tubercle, by which the eye is attached to a peduncle or stalk. The sclerotic of the sturgeon is so thick that it resembles a cartilaginous sphere, with the external part hollowed for the humours and membranes. In the Cephalopodous Mollusca it forms behind a truncated cone, with the apex at the bottom of the orbit containing the gangliform swelling of the optic nerve, and several glandular parts, with the eye before. The cornea has often been represented to be merely a continuation of the sclerotic; and though this is easily disproved by accurate dissection of the human eye and that of our ordinary domestic animals, its inaccuracy is much more manifestly demonstrated in the animal world at large. In the whale and rhinoceros the margins of the two membranes penetrate reciprocally. In man and the ox the corneal margin is enchased within the sharp im¬ bricated edge of the sclerotic. In the tope-fish {squalus milandra, Lin.; galeus, Cuv.) the cornea is observed dis¬ tinctly passing within the sclerotic in the manner of im¬ brication. The cuttle-fish is destitute of cornea ; and as there is no aqueous humour, the crystalline lens is cover¬ ed by a fine thin membrane, extended beneath the con¬ junctiva. In all animals provided with eyelids, the mucous mem¬ brane, after being folded behind the eyelids, is reflected forwards over the sclerotic and cornea, in the form of a thin, transparent membrane. In those void of eyelids, as most fishes are, the skin, passing into mucous membrane, is continued directly over the cornea, without forming any angular fold, and adheres strongly. This is very dis¬ tinct in the eel, which may be flayed without leaving any trace at the site of the eyes, except a round, translucent spot. The same peculiarity is remarked in Serpents and in the cuttle-fish family. In the zemni, golden mole, blind eel, and hag-fish, it has been already stated that the cornea is covered by opaque mucous membrane. The choroid coat exists in all animals yet examined. It is always very vascular. The inner layer, which has been distinguished by the name of tunica Ringschiana, can scarce¬ ly be said to exist in man, small quadrupeds, and birds. In the large quadrupeds, however, especially the Cetace¬ ous animals, it appears in the form of a distinct simple membrane like epidermis. The lateral and anterior parts of the membrane are always invested by a semifluid, viscid substance, of different shades of black or brown black {pig- mentum nigrum); chocolate brown in the hare, rabbit, and pig; deep red brown in some birds; and purple red in Coropara- the calmar. The absence of this dark-coloured pigment, dve which is not unfrequent, is observed in albinos, both hu- ^riatomy. man and animal, for instance white rabbits and white mice. The transparency of the Ruyschian membrane then shows the choroid of its natural red colour ; and the pupil is red and contracted, and the eye intolerant of light. In the Zoophaga, Ruminantia, Pachydermata, So- The ta- lidungula, and Cetacea, the concave or inner surface PetuI?. or of the Ruyschian membrane is diversified with colours metallic lustre, more or less brilliant and something irides¬ cent. In the ox it is of bright metallic green, changing to sky-blue; in the horse, buck, buffalo, and stag, it is a silvery blue passing to violet; in the sheep of a pale golden green, sometimes bluish ; in the lion, cat, bear, and dol¬ phin, of a pale gold yellow; and in the dog, wolf, and badger, of a pure white, surrounded by blue. This coloured part of the inner choroid surface, which occupies chiefly the side opposite to that on which the optic nerve enters, is named the tapetum. The use of it is by no means obvious. The explanation of Monro in reference to the tapetum of the ox, that it represents more distinct¬ ly to that animal the colour of his natural food, is not only frivolous, but inapplicable to the other genera. The tapetum is wanting in all Birds and Fishes, ex¬ cepting the ray, in which there is at the bottom of the eye a beautiful silvery-coloured space, produced by the trans¬ parency of the Ruyschian tunic, through which the tint of the choroid is seen. In Fishes generally the choroid consists of two dis-Thecho- tinct separable membranes; the external, the proper cho- glarul roid, white, silvery, or golden, very thin and not vascular ;° s 1Ch' and the inner or Ruyschian, black, and consisting of a net¬ work of vessels. Between these two membranes is a body of a bright red colour, consisting of numerous tortuous vessels, convoluted and inclosed in pulpy filamentous tissue. Its general shape is that of a thin cylinder, en¬ compassing the optic nerve like a ring, which, however, is incomplete at one side. This is the choroid gland,—a body about Avhich there has been some difference of opi¬ nion, but which appears to be glandular rather than any thing else. Its vascular structure is well seen in the globe-fish, perca labrax, and cod, in which they are very large, and form numerous anastomotic communications. They are genei’ally covered by a white, opaque, viscid fluid. The choroid gland is wanting in the Cartilaginous Fishes, the eye of which approaches more nearly to that of the Mammalia in this as in other circumstances. The choroid of the ray and shark genera is a threefold tissue of vessels, thick and consistent; the tunica Ruyschiana is very thin and semi-transparent; and between these is a layer of silvery matter with metallic lustre. In the cuttle-fish genus, though between the sclerotic and choroid there are several glandular bodies, there are none between the choroid and Ruyschian tunic. The choroid is thick, soft, and vascular; the Ruyschian thin, firm, and dry; and though there is no tapetum, the whole interior surface of the eye is covered by deep purple, semi¬ fluid, viscid varnish. Ciliary processes are found in all the Mammalia, Birds, Ciliary several Reptiles, and even in the cuttle-fish among theProce?ses Mollusca ; but they are wanting in almost all fishes. an< ciic The indented border of these processes is more dis¬ tinct, and is converted into a genuine fringe in the large animals, as the ox, horse, rhinoceros, and whale, in which the angle applied to the capsule is more acuminated than in other animals. In the Carnivora, particularly the lion, the base of the plates is shorter in proportion to the other sides than in the previously mentioned animals, so that the opposite angle is more prominent; nor is the 24 ANATOMY. Compara¬ tive Anatomy. The iris and uvea. border indented. In all the species every third or fourth plate is shorter than the others, but without determinate order. The ciliary plates of Birds are mere serrated striai, without sufficient prominence to make them undulate in fluid. In the owl they are fine, closely set, and nume¬ rous ; in the ostrich they are larger and more numerous; and ill all, their extremities adhere firmly to the capsule of the lens. In the tortoise the ciliary processes are so short that they are recognised only by the impression left on the vi¬ treous humour; in the crocodile, however, they are dis¬ tinct, and terminate each by an angle nearly right. They are indistinct in the toad, and imperceptible in the ordi¬ nary lizards and serpents. The ciliary body and processes are large and distinct in the tope-fish; but if they are seen in any other of the car¬ tilaginous fishes, they are wholly wanting in the osseous, in which the Ruyschian tunic is directly continuous with the uvea. The utility of these processes in retaining the lens in its position is nowhere so distinctly seen as in the eye of the cuttle-fish family, and especially the many-feet of the ancients {polypus octopodd). In these the ciliary pro¬ cesses form a large diaphragm or zone, in the aperture of which the crystalline lens is truly chased. They pene¬ trate a deep annular furrow which surrounds the lens, dividing it in two unequal hemispheres, and cannot be de¬ tached without laceration. The iris, of the same intimate structure as in man, is of a deep tawny or brown in the Mammalia, and marked with fewer coloured strice than the human iris. In Birds it is of a uniform lustreless colour, varying according to the species, bright yellow, red, or clear blue. In the microscope it appears like a net-work formed by the in¬ tersection of numerous very minute fibres. The uvea is so fine that when the viscid varnish is removed it be¬ comes transparent, and the iris appears of the same colour on both sides. In Fishes, conversely, the iris is so thin and transparent that the uvea is seen through it, of a golden or silvery brilliance, showing its direct connection with the choroid. Intermediate in metallic splendour is the iris of the Reptiles. The vessels, however, are greatly more conspicuous, especially in the crocodile. The central aperture or pupil, though round in man, the Quadrumana, many of the Carnivora, and Birds, is not of that shape when contracted in all animals. In the feline family it consists of two elliptical ^ogments, which form angles above and below, and which approach mutu¬ ally so as to form a slit nearly vertical. In the Ruminants it is oblong transversely, and forms at its greatest contrac¬ tion an oblong or transverse slit. In the horse, in which it is also transverse, its upper margin is distinguished by a five-pointed festoon. In the whale it is oblong transverse¬ ly, and in the dolphin it is heart-shaped. The pupil of the crocodile resembles that of the cat; in the frog and gecko it is rhomboidal; and round in the tortoise, chameleon, and common lizards. In the ray among Fishes the upper margin forms several radiating slips like the branches of the palm-tree, gold-coloured without, dark within. In the dilated state these slips are folded backwards between the upper margin of the pupil and the vitreous humour; but when the eye is pressed they are erected, and close the pupil like a blind. The motion of the pupil is voluntary in the parroquet, and is indistinct in most of the Fishes. The pupillary membrane is well known to exist in the foetuses of all the Mammalia ; but it is not determined whether it is found in the chick of birds. On the subject of the retina in the lower animals the most important point is the structure of the melanoplectic Compara- or pectiniform membrane {pecten, marsupium nigrurn) of five Birds. In this class the optic nerve forms not a round disk Anatoinv. as in the Mammalia, but a narrow white line, the margins and extremity of which are in continuity with the n A Along this line is suspended a plicated or convoluted membrane membrane, very fine and vascular in structure, like the or mamu. ’ choroid, from which, however, it is quite distinct, and en-fuim, ni. tering a depression of the vitreous humour almost like afr"’?’ an(l wedge. Its vessels, which proceed from a proper branchlts *0 ( s' of the ophthalmic artery, are distributed in a minute arbo¬ rescent form, among the folds of which the marsupium con¬ sists; and from these vessels the black viscid pigment with which its folds are covered appears to be secreted. The plicae or membranous folds vary in number. In the casso¬ wary they are only 4; in the brown owl ( S. aluco) 5; in the common owl, ostrich, Guiana macaw, and merganser, they are 7 ; in the flamingo 9 ; in the falcon and swan 11; in the vulture and goose 12 ; in the duck, large heron, woodcock, and coot, 13; in the stork and partridge 15; in the crane 17 ; in the pheasant 20; in the turkey 22 ; in the jackdaw 25 ; and in the thrush 28. According to the observations of the elder Soemmering, to whom we are indebted for these numbers, the number of folds, though variable in different species, is the same in the same. In most birds the folds are arranged in a pectiniform order. On the use of this organ different opinions have been entertained by Petit, Haller, and Home ; but all of them are conjectural. In the Reptiles and many Fishes, between the optic nerve and the retina is a small tubercle, from the margins of which the latter membrane appears to rise ; and radiating fibres are perceived more distinctly than in most quadru¬ peds. In many other Fishes the connection of the reti¬ na with the optic nerves resembles that of Birds. Thus, in the salmon, trout, herring, mackarel, cod, dory, and moon-fish, the optic nerve, after passing through the Ruyschian tunic, appears to be parted into two long white processes, which, following the outline of this membrane parallel but not contiguous to each other, are connected with the retina by their opposite margins. In all animals provided with ciliary processes the retina terminates at and is connected with the gray pulpy zone denominated ciliary ligament. In those without ciliary processes, as the Fishes, it terminates suddenly at the attached or large margin of the uvea. In several of the reptiles the retina presents the yelloAv spot of Soemmering. The principal peculiarities of the humours have been already mentioned. Of the appendages the most important are the lacrymal gland and nictitating membrane. In the Ruminants the lacrymal gland consists of two or Lacrymal three bodies, each composed of granules, each provided glanii- with a separate short excretory duct. In the hare and rabbit, in which the gland is large, there appears to be only one excretory duct, which perforates the upper eye¬ lid near its posterior angle. A gland peculiar to certain species, and wanting in man, that of Harder, is situate at the external or nasal angle, and presents an aperture under the third or nicti¬ tating eyelid, from which issues a thick viscid fluid. It is found in the Ruminants, the Rodentia, the Pachy- dermata, and in the sloth genus. The caruncle exists in the Ruminants as in man, and appears to consist of numerous aggregated follicles. It is wanting in the Rodentia. In the Cetacea, as in most animals which live under water, there is neither gland nor lacrymal passages; and they are represented apparently by lacunae below the up¬ per eyelid, which discharge a thick mucilaginous fluid. Birds, though destitute of caruncle, have both lacry- 1 ' ANA' 4>mpara- mal gland and that of Harder, the latter large, oblong, tive and flesh-coloured, placed betwixt the levator and adductor, inatomy. an(j discharging by a single canal, opening at the inner surface of the third eyelid, a thick yellow fluid. The la- 2*ucipa- crymal, which is small, round, and very red, is provided Wis Mil- in general with two or three canals, which, though small, are distinct. In most of the Orally and Palmiped Birds there is, in place of the lacrymal gland, a hard granular body, occupying the upper part of the orbit, and following in situation the curvature of the eye. It has, nevertheless, no visible excretory duct. In the turtle there is a reddish granular lobulated body, of considerable size, extending beneath the temporal vault. In the tortoise, frog, and toad, there are two small blackish glands without apparent excretory ducts. Neither in Serpents nor Fishes has any glandular apparatus in the eye been recognised. He third Though in man and the monkey tribe the eyelids ei lid or consist of two semilunar cutaneo-muscular folds, with a ndilating minute mucous duplicature at the nasal angle, the latter nnnbrane.acqUjres sucfl a developement in the lower animals as to constitute a genuine third eyelid, often distinguished by the name of nictitating membrane. This duplicature is semi¬ lunar in shape in the Ruminants, Edentata, and Pachy- dermata. In the rhinoceros it is thick and fleshy; but of this the Cetacea present no trace. In the Birds, on the contrary, in which the eye is covered by the eleva¬ tion of the lower eyelid, which is also the largest, the third eyelid is large, and covers the eye like a blind drawn before it; yet it is in some degree translucent, for it is evident that birds see objects through the membrane. In the owl and goatsucker the eye is closed by the depres¬ sion of the upper as well as the elevation of the lower eyelid. Though the Serpentine reptiles are void of eyelids en¬ tirely, in the crocodile, tortoise, and Batrachoid, there are three, as in birds, the third being vertical in the two former orders, and horizontal in the latter. In the Saurial and Cheloniad,also, the third, which is translucent,moves from before backwards by means of a single muscle, and may cover the whole eye. In the lizard genus the eyelids consist of a circular veil drawn before the orbit, and per¬ forated by a horizontal fissure, which is shut by a sphinc¬ ter, and opened by a levator and depressor. The gecko has no movable eyelid. Cenpound In insects, the eye consists of innumerable hexagonal of the surfaces, slightly convex, and mutually separated by mi- Anricu- nute furrows, containing fine hairs variable in length. Each of these hexagonal surfaces, which constitute a hard, elastic, very transparent membrane, may be regarded as a cornea or crystalline lens, convex externally, concave within, and thicker in the centre than on the margins. Immediately behind is an opaque, viscid coating, varying !in colour in the different species, analogous to the choroid pigment of the vertebrated animals, and completely ob¬ structing the transmission of light. Beneath this varnish are short, whitish filaments, corresponding in number to the corneal surfaces, and mutually joined like mosaic or tessellated pavement, separated only by the dark-coloured pigment, and which appear to correspond to the retina of the Vertebrata. Behind these again is a delicate, dark- coloured membrane, which appears to correspond to the choroid; and exterior to this is a membrane continuous with the optic nerve, and which seems to be a general re¬ tina, forming, by subdivision of its parts on the anterior part of the choroid, the divided or multiplied retina. This is the structure of what are named compound eyes. That of the simple eyes of insects is too minute to be accurate- ly demonstrated; but analogy gives probability to the in¬ ference that they are not dissimilar. VOL. in. O M Y. 25 sect. hi.—the EAR. Compara¬ tive In warm-blooded animals generally, that is, in the Mam- malia and Birds, the labyrinth or essential part of the organ consists of three semicircular canals, with a globular enlargement to each {ampulla), a cavity common to these canals named vestibule {vestibulum), and a conical taper¬ ing canal, divided into two compartments by a longitudi¬ nal septum. This may be named the bilocular cone (conus bilocular is). These parts consist of membranous substance inclosed in the bony walls of the pyramidal or auditory bone. In all the Mammalia the bilocular conical canal is convo¬ luted in a spiral form, and hence is denominated, as in man, the cochlea—a name, however, which is applicable to it in this class only. The organ of hearing in the Mammalia consists of the same parts nearly as in man. In some; indeed, for in¬ stance the guinea-pig (cabiai), and porcupine, the cochlea Cochlea makes three turns and a half; and, conversely, in the Cfi-and canals. tacea only one and a half. In most of the Zoophaga, and in the hog, elephant, and horse, the cochlea is much larger in proportion than the semicircular canals ; but in the hare it is small, and in the mole very small. In the Cetacea, while the cochlea is very large and fully deve¬ loped, its spiral is on the same plane throughout; and the semicircular canals are so small, that their existence was long denied by Camper, till they were demonstrated by Cuvier in a foetal whale. In general the labyrinth of the Mammalia is greatly smaller in proportion to the head than in Birds. This part, which is membranous, is inclosed in the solid compact substance of the temporal pyramid, so closely that its existence appears to be identified with the latter. Re¬ searches, however, on the labyrinth in the foetus of the Mammalia, and especially in those of whales, demonstrate the fact that it is in a completely membranous form, dis¬ tinct from the bony inclosure; that in shape and consti¬ tuent parts it exists previous to the bony inclosure ; and that the latter is afterwards moulded round the different parts as they acquire their full developement. It is also to be observed, that in the mole the semicircular canals are seen within the cranium without preparation, and the cochlea is merely inclosed in fine cellular tissue. In the bat family, also, both parts are seen without bony inclo¬ sure. The tympanum forms a cylindrical or spheroidal cavity in Tympa- most of the Mammalia. In most of the Digitata the num. mastoid process consists of a slight prominence of the tym¬ panum only as it is identified with the latter; but in the cabiai, guinea-pig, hog, the Ruminants, and Solidungula, it is represented by a long process of the occipital bone. In most of the Zoophaga and Rodentia the parietes of this protuberance, which are thin and hard, form by their se¬ paration a large cavity. In the hog only it is occupied by a firm cancellated structure. All the Mammalia, except the ornithorhyncus, have the Tympanal tympanal bones as in man; the hammer (malleus), anvil bones. (incus), orbicular bone and stirrup (stapes). The lenti¬ cular bone, which is rarely found in the adult, is probably only an epiphysis of the anvil. They are articulated with each other so as to admit of motion, and are moved by the same muscles as in the human subject—the internus mallei, externus mallei, laxator tytnpani, and stapedius. In the ornithorhyncus, however, there are only two tympanal bones. In all the Mammalia, except the Cetaceous, the ear External is provided with a bony external canal (meatus); and aperture, most of the Mammalia, except the Cetaceous, have a cartilaginous funnel-shaped opening (concha) attached to the outer margin of the bony meatus, and which serves to x> 26 ANATOMY. Compara- collect the sonorous vibrations, and direct them to the tive meatus. The other exceptions are among the Insecti- Anatomy. V0RA? the mole, and some of the shrew genus; among the Rodentia, the zemni or blind rat, and some of the rat- mole genus; among the Edentata, the pangolin or scaly ant-eater; and among the Amphibia, the morse and se¬ veral species of seal; and the ornithorhyncus paradoxus. The tympanum of the Cetacea is peculiar. It con¬ sists of a bony plate, convoluted on itself like a buccinum, unless that the thick side, instead of containing a spiral cavity, is entirely solid. The opposite side is thin, with an irregular margin. The anterior extremity of the tym¬ panum is open, and there commences the Eustachian tube, which ascends along the pterygoid process, and, pe¬ netrating the maxillary bone, terminates at the upper part of the nose. This direction of the tube and position of its orifice is so much more necessary, because, since these animals have no external bony meatus, and the ear-hole scarcely admits a pin, the vibrations of the air reach their organ of hearing entirely by the Eustachian tube, and because the Eustachian tube also in these animals conveys odorous impressions to the part in which the sense of smell appears to reside. The aperture by which it communicates with the nose is provided with a mem¬ branous valve, which prevents the water from entering when the animal expels it by his nostrils. In Birds, of the three semicircular canals the vertical is largest, and obliquely directed forwards and outwards; the second is horizontal and turned outwards; and the third, which, like the first, is vertical, crosses the second, and is turned in the direction opposite to that of the first. Bilocular The vestibule is small and nearly spherical. The bilocu- cone no iar cone, which is obtuse at the apex, is situate obliquely longer spi- backwards and outwards below the inferior part of the cranium. The longitudinal septum consists of two nar¬ row cartilaginous plates connected by a thin membrane. The posterior canal is short, and, as in the Mammalia, is separated from the tympanal cavity by the membrane of the fenestra rotunda; while the anterior, which is larger, communicates directly with the vestibule. The whole of these parts are inclosed, as in the Mammalia, in the compact bone of the pyramid. The posterior and inferior ^walls of the tympanal cavity are formed by part of the occipital bone ; the lateral aper¬ ture is large, and the cavity superficial; and its anterior part is closed by the posterior superior cornu of the quad¬ rilateral bone and a membrane. The inner wall presents the two apertures—the oval or vestibular, and the round or cochlear. In this class, however, while the upper is round or triangular, the lower is distinctly elliptical,—a disposition the reverse of what is observed in man. The Tympanal Eustachian tube or tympano-guttural canal is entirely os- bones dis- seous. The tympanal cavity contains only one ossiculum, appearing consisting of two branches ; the first attached to the tym- fied10 '*' panum, corresponding to the malleus ; the second closing by an oval or triangular plate the vestibular aperture, and therefore corresponding to the stapes of the Mammalia. By Carus the incus is supposed to be represented by the quadrilateral bone. The external meatus is short, and opened by a simple aperture, while the absence of external ear is compensat¬ ed by a ring or zone of fine elastic feathers with thin barbs, between which the air passes very easily. In the owl tribe it terminates in a large cavity, the margins of which are covered by a smooth valvular fold of skin. Bilocular The ear of the Reptiles is remarkable for the last ap- cone disap-pearance of the bilocular cone, and the first of the sac- pears. cular apparatus which is found in the Fishes. In the cro¬ codile and lizard this part appears, as in Birds, in the shape of a conical tube, divided by a cartilaginous partition into a double canal, one separated by the membrane of the round Compara. , hole from the tympanal cavity, the other communicating tive with a membranous sac containing three very small friable . stones, not harder than starch. There are also three semi- ' circular canals of considerable size, each forming a large circumference. In the frog and toad, while the three canals Lithopho. ' form almost a complete circle, the sac contains an amyla-rous sacs ceous friable stone; but the bilocular cone is no longeramy- observed. In the salamander, also, in which the three ^nesS canals form together a sort of equilateral triangle, the sac appear. which is below contains a single amylaceous stone. The same arrangement is observed in the Cartilaginous Fishes, unless that the sac contains two amylaceous stones, near¬ ly oval in shape, suspended in a gelatinous semifluid pulp. In the Osseous Fishes it is a little different. The three semicircular canals terminate in a membranous sac, which is divided by septa into compartments v/hich contain one, two, or three small stones suspended in gelatinous fluid. These minute stones, however, instead of being soft, fri¬ able, and amylaceous, as in Reptiles and cartilaginous Fishes, are as hard as rock, and white as porcelain. These parts are situate on the sides of the cranial cavity, and are fixed to it by cellular tissue, vessels, and osseous or cartilaginous processes. This sac, in the fluid of which the extremities of the auditory nerve are distributed, is believed to correspond to the bilocular cone of the higher classes. These membranous cavities are contained, in the bony fishes, in the general cavity of the cranium; and while only the middle of the canals is inclosed in the bone of the cranium, the extremities and the sac are quite free. The sturgeon, which belongs to the cartilaginous order, is the first in which the canals are entirely inclosed in the cra¬ nial cartilage; but even in this a membrane is interposed between the cranium and sac, which is free. In the ray and shark genera, again, these organs are entirely in¬ closed in the cartilage of the head. The tympanal cavity, in like manner, is modified, and eventually disappears as we descend in the scale. Though present in the tortoise, crocodile, and lizard tribe, it is superficial and open; it becomes membranous behind in the Ranine tribe, and communicates directly with the back of the mouth; and in the Serpentine reptiles it entirely disappears, so that the handle of the osseous plate by which the oval aperture is closed is suspended in the soft parts with its free extremity below the skin, near the articulation of the lower jaw. In the lizard tribe, also, Cochlear , the round or cochlear aperture is seen for the last time, aperture j In the Cheloniad, for instance, the Batrachoid, and the c^saPPeaB Serpentine, this aperture disappears, and the oval or ves¬ tibular alone is left; and in the salamander both disap-Vestibula: pear, and there is no communication between the external aperture part of the cranium and the labyrinth. This arrangement (hsaPl)eaB is continued in the fishes. In the molluscous animals the labyrinthine membrane is a simple sac, globular or ovoidal, containing pulpy matter, in which is suspended a small body, which is osseous in the sepia and amylaceous in the many-feet (polypus'), in which the filaments of the auditory nerve are distributed. Our limits do not allow us to enter into the detailed description of the organ in the other Invertebrated animals. SECT. IV. THE ORGAN OF TASTE. Though the sense of taste is seated chiefly in the tongue in animals, yet that organ performs, in all the classes, so important a part as an instrument of prehen¬ sion, that it cannot with much justice be distinguished by the former title only. In the present section, therefore, we must regard it as one of prehension as well as of taste. ANATOMY. 27 «3 r;« ;]IIf 0 IB lompara- In the Mammalia and Birds the tongue is a muscular tive organ invested by mucous papillated membrane, supported Jan atomy. ^ a proper bone, the hyoid, which serves as a point of support in its various motions. In the Ranine Reptiles t mue. it is aiso muscular, attached to the margin of the lower n jaw. In the salamander, however, it is attached as far as the tip, and is movable on the sides only. In the croco¬ dile it is attached so generally, both by tip and margins, that it was long asserted that the animal was tongueless. In the stellio and iguana it is as movable as in the Mam¬ malia ; and in the seine and gecko to this property is added that of being bifid, or divided by a longitudinal notch into two pointed tips. In the ordinary lizard, tu- pinambis, monitor, &c. the tongue is remarkable for its great extensibility, and terminates in two long, flexible though semi-cartilaginous extremities. That of the cha¬ meleon is still more extensible, and forms, by a peculiar arrangement of vessels, a cup-like extremity. The tongue of the blind worm (anguis fragilis) and amphisbeena is also bifid at the tip. The cartilaginous fishes are void of tongue, while in the bony division of the class this organ is represented by a hard protuberance, attached to the middle branchial bone. In some of the Mammalia, however, the tongue is not exclusively muscular. In the singularly long, extensi¬ ble, and tortuous tongue of the giraffe, Sir Everard Home describes a peculiar arrangement of vessels, which he re- I-ectile presents as a substitute for muscular motion. Though Sir arange- Everard does not appear to understand the exact nature of *?nt in this arrangement of vessels, all the circumstances tend to He tongue sjlow tbat denominated erectile. These vessels, (].je from the account given, are large, numerous, and com¬ municate freely; and it would be impossible to discover the reason of such a vascular system, unless for some purpose of this description. (Phil. Trans. Comp. Anat.) When the tongue is protruded it becomes perfectly black or bluish-black, evidently from the injection and detention of the blood in its elongated and anastomosing veins. By means of this mechanism the giraffe not only elongates the tongue to the distance of about twenty inches or two feet beyond the mouth, but twists it round the soft leafy twigs of the trees on which he feeds. It is not improba¬ ble that a similar vascular arrangement exists, though in less degree, in the tongue of the deer, and in the long projectile tongue of the animals of the ant-eater tribe, as the Tamanoir, Tamandua, &c. Id of the The erectile arrangement is still more distinctly pre- oiameleon. sented in the tongue of the chameleon. The researches of Mr Houston of Dublin show that the tongue of this animal consists of two parts,—a prehensile, which is ante¬ rior, and provided with a glandular apparatus for secret¬ ing the viscid fluid by which its tip is covered, and in¬ sects are entangled; and an erectile, which is posterior between the prehensile and the hyoid bone, in the form of a trellis-work of innumerable minute anastomosing blood¬ vessels, not very dissimilar to those of the cavernous body in animals generally, and inclosing a central tube connect¬ ing the prehensile portion to the hyoid bone. The effect of this arrangement is, that when the vascular network is injected with blood, the anterior part of the tongue is ra¬ pidly darted out at the insects on which the animal lives. The injection of these vessels, and the consequent pro¬ jection of the tongue, is not independent altogether of the will of the animal; for the veins by which the blood is returned pass through a slit in the tendon of the internal cerato-maxillary muscles, which are always contracted in order to protrude the hyoid style, and thereby tend, by compressing the veins, to inject the erectile part, and pro¬ ject the tongue. ( Trans. R. I. Acad. 1828, and Dublin Hospital Reports, vol. v. p. 487.) The same arrangement is in all probability found in several of the lizards, the Compara- tongues of which, like that of the chameleon, are darted tive out suddenly, and become of a dark-blue colour at the Anatomy, moment of projection. In all the Mammalia the tongue is invested by a pa-Papilke. pillated muco-villous membrane, in which the papillae are of the same general characters as in man—granular, mush¬ room-like, or fungiform, tubercular or calycoid, and conical or acuminated. The only differences consist in the size and abundance of the fungiform papillce, in the number of the calycoid and the mode of their arrangement, and in the shape of the conical papillae and the mode of their termination. In the Ruminants especially, the conical papillae are numerous, long, slightly incurvated, and each terminating in a horny but flexible style slightly incur¬ vated backwards. The tongue of the dolphin and por¬ poise, examined even by the microscope, presents no dis¬ tinct conical papillae, but is covered by minute eminences, each penetrated by a small aperture. In the tongue of the dog genus there is a ligamentous Worm of substance extended longitudinally from the hyoid bone to the dog; its the tip of the member. This, which has been vulgarly distinguished by the name of worm of the tongue, and has been absurdly supposed to be the seat of hydrophobic rabies, is merely a central pillar of support for the mus¬ cular fibres to act with greater steadiness and effect, and which enables the animal to protrude and expand the tongue in lapping water or other fluids better than he could have otherwise done. A similar central ligament is found in the opossum. The tongue of Birds is generally more or less horny, and almost cartilaginous. That of the woodpecker and wryneck is peculiar in consisting of two parts,—a basilar or posterior, loose and fleshy; and an anterior projec¬ tile, long, smooth, acuminated, and covered laterally with four or five stiff spines directed backwards, which make the organ a sort of barbed arrow. The soft, loose, or basilar part of the tongue contains the aperture of the glottis; and the surface is covered with minute spines pointed backwards, and each of which is placed in the centre of a fleshy papilla. As a prehensile organ of very singular construction, Trunk of the trunk of the elephant deserves particular notice ; and A16 ele- it cannot be more conveniently introduced than under the phant" present section, since it is used not only to convey food, but drink, into the mouth. The trunk may be described as a cylindrical tubular organ, consisting of integument, a sort of fibro-cartilage, muscles, fat, and a membrane of villous character internally. This tube contains two long canals continued from the nostrils, parallel to the axis of the trunk, and separated throughout by a partition of adi¬ pose substance about two fifths of an inch thick. From the extremity to the middle part of the intermaxillary bone, in which the tusks are fixed, these canals are nearer the anterior-superior than the posterior-inferior part of the tube, the latter wall being thickest; and their diameter is the same throughout. At this part they undergo a sudden incurvation, approaching the anterior surface of the inter¬ maxillary bone, and form a semicircular bend with the convexity turned forwards. Here also they are so narrow that, without a muscular effort on the part of the animal to dilate them, fluids could not ascend beyond this point; and hence this forms the only valvular contrivance, either to impede the progress of fluids upwards, or to propel them downwards, at the will of the animal. Above this curvature each canal is dilated before the upper part of the intermaxillary bone, and again is con¬ tracted where it bends back to enter the bony nostril; and the curvature is protected before by the nasal cartilage, which is oval, convex in the male, and flat in the female. 28 ANATOMY. Compara- Both canals are lined by a dry, greenish-yellow coloured tlve membrane, marked with superficial intersections {ruga:), ^closing rhomboidal spaces, and some venous branches. Muscular Though the muscular fasciculi of the trunk are nume- apparatus rous, they may be referred to two orders,—those forming of the the substance or inner part of the organ, and those by trunk which it is invested. The former, which are transverse, highly an(j cu). |.jle axjs jn different directions, consist of nume- Ca * rous smaii muscular packets proceeding in various direc¬ tions, some running from the inner membrane to the cir¬ cumference of the tube, others directly from right to left, and others crossing the two former obliquely. All these little muscles are inclosed in cellular tissue, containing white homogeneous fat; and all of them terminate in slen¬ der tendons, some of which cross the layers of the longi¬ tudinal muscles in their course to the external covering, while others are attached to the internal membrane. Cuvier calculates the number of these minute transverse muscles in the trunk of the elephant to be not fewer than 30,000 or 40,000. (Plate XXXVII. fig. 13.) The longitudinal muscles, which are external, may be distinguished into anterior, posterior, and lateral bundles. The first extend from the anterior surface of the frontal bone, above the nasal bones and cartilages, in parallel bundles, connected by tendinous intersections downwards on the trunk. The posterior extend from the posterior surface and inferior margin of the intermaxillary bones, and form two layers which meet on the median line along the lower surface of the trunk. The lateral muscles form two pairs, one of which, descending between the anterior and posterior muscles to the middle of the trunk, may be regarded as a continuation of the orbicular muscle of the lips, or the representative of the nasalis labii superioris ; while the other, which is attached to the anterior margin of the orbit, and is expanded over the root of the former, may be supposed to correspond to the levator of the upper lip. The whole of these muscles are supplied by a very large branch of the infraorbital or second branch of the tri¬ facial nerve, which, entering on each side between the lateral and superior muscle, is distributed to the whole of the trunk. With such a construction, it is not difficult to under¬ stand the numerous motions of the elephant’s trunk. While the longitudinal muscles are employed either to shorten the tube, to bend it upwards or downwards or to the side, or by means of the tendinous intersections to give it peculiar inflections, it is manifest that the trans¬ verse ones, which act as antagonists to the longitudinal, may also either dilate or close the canals, or incurvate or alter the direction of particular parts. Vacuefy- The foot of the Lacerta Gecko and the house-fly pre- ing appa- sents a prehensile apparatus of peculiar construction for the1 feet 0fw.alkinS along surfaces, in opposition to the action of gra- certain vity‘ 1° the former animal the plantar surface of each animals, toe presents sixteen transverse slits, leading into an equal number of pouches, which by means of appropriate mus¬ cles are capable of forming an equal number of vacua, so that the atmospheric pressure is employed with muscular effort to support the animal in his unnatural position. A similar apparatus is found in the upper surface of the head of the sucking fish {echeneis remora); and something ap¬ proaching to it, though less distinctly, in the foot of the walrus. (Home, Phil. Trans. 1816.) CHAP. IV.—COMPAKATIVE ANATOMY OF THE ORGANS OF VOICE. _ Under this head our limits allow us to mention very few circumstances. In the American long-tailed monkeys {sapajous) the cuneiform cartilages form, by means of adipose cellular Compara. 31 tissue, before the upper extremity of the ventricle of the glottis, a large cushion like a spherical segment, which, ^ touching that of the opposite side, causes the air to whistle through the canal in its course to the mouth, and occa¬ sions the flute-like voice of some of these animals, as the weeper {s. apella) and the capuchin {s. capucina). In the Voice of j howler {s. seniculus), so remarkable for its morning and the howler evening yelling, though the larynx is similar in general aPe> characters to that of the common sapajou, in having the two rounded cushions before the ventricles, the hyoid bone is arched in the form of a spherical chamber, with a large quadrilateral aperture, and each ventricle opens into a membranous sac, lying between the epiglottis and the adjoining wing of the thyroid cartilage. The air, which passes between the vocal chords, is therefore partly im¬ pelled into this osseous and elastic cavity of the hyoid bone, and probably by its resonance in this situation gives the voice of these animals the deep-toned howl by which they are known in the American forests. Among the Zoophaga, in the dog the cuneiform carti¬ lages are large, the arytenoid small, the vocal chords well marked, and the ventricles deep. In the feline tribe the anterior ligaments, though destitute of cuneiform carti¬ lages, are thick, and separated from the back of the epi¬ glottis by a broad, deep furrow. The posterior ligaments, though neither free nor sharp-edged, are distinguished from the anterior by an appearance of greater firmness, more regular fibres, and by an intermediate furrow. The approximation of the anterior ligaments towards the gloU tis forms a sonorous vault, in which the air may be forci- r bly vibrated by the posterior. In the bear the cuneiform cartilages assume the shape of styles, and their posterior extremity forms a distinct eminence, not above, but with¬ out the arytenoid cartilages, while the ventricles are merely deep fissures. The kangaroo has neither cuneiform cartilage, anterior ligament, nor ventricle; and it may even be said to be void of vocal chord, while the margins of the glottis are much separated in the middle. This arrangement appears to indicate that the animal is almost destitute of voice. In the opossum, in which there is merely a small infe¬ rior ligament susceptible of tension, voice is limited to a whistling sound. In the Solidungula, in which the cuneiform cartilages are completely concealed by the mucous membrane, there is neither superior ligament nor proper ventricle; but an aperture in the lateral wall of the laryngeal membrane, above the vocal chord, leads into a large, oblong, sinuous cavity, situate between this membrane and the thyroid cartilage, and covered chiefly by the thyro-arytenoid muscles, by which it may be compressed; and above the anterior commissure of the vocal chords, or below the base of the epiglottis, is an aperture on the mesial plane, leading into a cavity below the vault formed by the ante¬ rior margin of the thyroid cartilage. This cavity, which may be named the infrathyroid, is superficial in the horse, of the ass, and its aperture is large; while in the ass, with a small, round aperture, the cavity is large, capacious, and globular in every direction, and allows the latter animal to make his voice re-echo in the singularly harsh sound denominated the bray. Conversely, though the lateral cavities are equally large in both animals, the apertures in those of the ass are small, round, and situate nearer the epiglottis than the vocal chord, while those of the horse are large, oblong, and situate immediately above the vocal chord on and horse, each side. On the latter peculiarity appears to depend the neigh of the horse. In the Cetacea we recognise neither vocal chords nor glottis, that is to say, an aperture variable in size ao ANATOMY. 29 Ciipara- cording to the will of the animal; but the superior part j,1 ‘ve of the trachea, which represents the larynx, forms a hollow 'n pyramid or funnel, rising into the posterior part of the ' nostrils, in which only it opens, while on the sides is left a passage for the food. This pyramidal funnel is formed by an elongated triangular cartilage, corresponding to the of epiglottis, attached by membrane to the arytenoid carti¬ lages, which also take the shape of scalene triangles, with the small side connected to the cricoid cartilage. Strictly speaking, therefore, the Cetacea have no larynx, and probably no vocal organ; and the superior part of the trachea, with the nostrils, serves merely to admit the at¬ mospheric air for the purposes of respiration. Birds are distinguished by possessing not only a glot¬ tis or laryngeal aperture similar to that of the Mamma¬ lia at the upper end of the trachea, but a second, deno¬ minated the inferior glottis or larynx, at the lower end, near its bifurcation. The former, which is composed of four cartilages, or six, according as the cricoid consists of one or three pieces, on the middle of the posterior part of which is a small round bone, articulated with two oblong longitudinal bones, parallel, and forming between them in the posterior wall of the windpipe a longitudinal slit, sus¬ ceptible of approximation by means of muscles, is intend¬ ed merely to regulate the admission of air into the wind¬ pipe, or its expulsion from that tube, and to close more or less accurately its superior orifice. : Inirior The inferior larynx consists of a membrane projecting l 0<1|U" from each s‘de ^ie inferior aperture of the trachea. [ ' This aperture is divided into two, occasionally by an os- seous anterior-posterior middle bar, occasionally by the angle at which the two bronchial tubes unite. Since the first bronchial arc has the same curvature as the last tra¬ cheal ring, the second and third, which are arcs of larger circles, are less convex without, but more prominent with¬ in, than the former. Over this prominence the tracheo¬ bronchial membrane forms a fold, which, half closing on each side the inferior tracheal aperture, forms a plate sus¬ ceptible of vibrating by the motion of the air, and produc¬ ing sound. This apparatus, which constituteswhat is named the inferior larynx, or rather glottis, is of two kinds, one void of proper muscles, the other provided with muscles. In the former kind of larynx the state of the glottis is altered only by those muscles which depress and elevate the trachea. The depressors are two pairs, the sterno¬ tracheal and the glosso-tracheal, the latter attached to the bifurcated bone and trachea. There are no proper elevators; but the windpipe is raised by the mylo-hyoid muscle through the ligaments which connect the hyoid bone to the superior larynx. In the quiescent or relaxed state, and while the trachea is depressed, the bronchial rings approximate, and the second and third even gliding below the first, the glottis may be elongated. When the trachea is elevated by these pairs of muscles, the bronchi are at the same time dragged upwards, and the second and third arcs are separated from the first; and while the ex¬ ternal prominence of the glottidal membrane diminishes in length, its tension is augmented. These forms of larynx without proper muscles may yet be subdivided into two sorts, as they have or have not lateral pouches, membra¬ nous or osseous. These are observed in the male duck {anas) and the merganser (mergus), but never in the fe¬ male ; and to this perhaps the harsh and deep tone of the voice of the male bird is to be ascribed. The larynx without muscles and without pouch is observed in all the gallinaceous order without exception. The forms of larynx provided with proper muscles may be distinguished into three subdivisions. The first, which has only one proper muscle on each side, is observed in the whole of the falcon genus, e. g. the eagle, hawk, falcon, buzzard, sparrow-hawk, and goss- Compara- hawk; in the owl genus and the majority of the waders tiv6 and swimmers, as the heron, bustard, woodcock, lapwing, Anatomy, rail, coot, gull, cormorant, and some of the passerine birds. In these birds, in which the motions of the lower larynx are necessarily limited, the voice is not variable or exten¬ sive in its notes. The second form of larynx has three pair of proper muscles, a constrictor of the glottis, an auxiliary constric¬ tor, and a laxator or opener of the glottis. This kind of larynx is observed in the whole of the parrot genus and psittacoid birds generally, as the toucan, macaw, calao, &c. In the third kind of musculo-membranous larynx there are no fewer than five pair of muscles, the longitudinal levator of the demiannular cartilages, the posterior le¬ vator of the same cartilages, the small levator, the ob¬ lique levator, and the transverse levator. This quinque- muscular larynx is found not only in all the birds proper¬ ly named whistlers or warblers, as the nightingale, hedge- sparrow, blackbird, thrush, goldfinch, lark, linnet, canary, chaffinch, &c., but in others whose tones are more mono¬ tonous, as the swallow, sparrow, stork, crossbill, &c., and even in some the tones of which are harsh and positive¬ ly disagreeable, e.g. the jay, magpie, crow, raven, &e. The differences remarked in the notes of these three divisions of birds with the quinque-muscular larynx de¬ pend not so much on anatomical peculiarities as on the timbre of their larynx, and on the mobility of the trachea in relation to the larynx, and on the tracheal membrane having dilatations and contractions. On the whole, the inferior larynx of birds is to be re¬ garded in three lights: Is#, As the reed of a wind-instru¬ ment, like a hautboy or clarionet, in which the notes vary as the lower glottis varies in its position to the wind¬ pipe ; 2d, as an instrument susceptible of uttering differ¬ ent tones, according to the distance between the mouth¬ piece and vent, or as the windpipe is elongated or short¬ ened ; and, 3d, as an instrument capable of uttering dif¬ ferent notes by varying the diameter of the mouth-piece, or as the superior glottis is widened or contracted. The only bird in which the inferior larynx is wanting is the vulture. The vocal organ of the Reptiles consists of the supe¬ rior larynx only, analogous to that of the Mammalia. This is a cartilaginous apparatus, composed in general of five distinct pieces at least in the large individuals of the Saurial genera, as the crocodile and alligator, and form¬ ing a broad cavity behind, before a narrow slit, bounded by two vertical pillars. The glottis, however, is entirely membranous; and there are neither vocal chords nor ven¬ tricles. There are nevertheless two muscles, one for opening and another for shutting the glottis. When these act, and the air is made to vibrate against the anterior pillars, it gives a slight whistling sound only. In the iguana, tupinambis, lizard, tortoise, and serpent, the ar¬ rangement of parts is nearly the same ; and these animals, therefore, can utter only slight hissing sounds. In the chameleon the pillars are furnished with a tense, vibrating membrane, a fleshy tubercle which contracts the glottis, and a membranous pouch opening below, between the lower laryngeal cartilage and the first tracheal ring. In the frog tribe, so remarkable for their croaking noise, the vocal chords are large and prominent. The males have also two membranous pouches, opening by a small aper¬ ture, not in the larynx, but deep in the lateral part of the mouth. When the frog croaks these pouches are inflated, and swell the skin on each side below the ear. Though these sacs are wanting in the female frog, and toad both male and female, as well as the tree-frog, there is beneath the throat a single pouch on the median plane. ANATOMY. 30 Compara- CHAP. V.—COMPARATIVE NEUROLOGY, txve Anatomy. ^ general view of the nervous systems in different classes of animals shows that the only common part is an azygous tubercle, situate at the anterior extremity of the spinal chord, connected by means of two lateral bundles or peduncles to the rest of the system. This mass, which corresponds to what is denominated the cerebellum in man, is connected in the vertebrated red-blooded animals with several pairs of tubercles, forming generally a larger mass than itself, and connected to the rest of the system by two longitudinal bundles or limbs, which mingle with and intersect those of the cerebellum. These anterior-supe¬ rior tubercles, which constitute the brain proper, present numerous modifications in figure, disposition, and magni¬ tude, and even in presence, in the different orders of the animal world. In the vertebrated animals the brain or central part of the nervous system consists of the vertebral or funicular portion, named the spinal chord, and the cranial or cepha¬ lic portion, properly denominated the brain. Spinal The funicular portion has the shape of a cylinder flat- chord. tened on its superior or dorsal and inferior or sternal surfaces. It consists of two similar chordiform produc¬ tions, united on the mesial plane, and marked at the line of junction by linear longitudinal furrows, of which the superior is most deep and distinct. This, however, which was observed by Blasius in several animals, is denied by Bellingeri, who asserts, from various observations, that though in some parts the superior furrow penetrates more deeply than the inferior, in the general course of the chord the latter is the deepest. ( De Medulla Spmali Annota- tiones Anatomico-Physiologicce. Auctore C. F. Bellingeri, Aug. Taurinorum, 1823.) This discordance appears to depend on the circumstance that the sternal or inferior furrow is in truth deeper and more distinct in man, the monkey tribe, and a few Mammalia, than the dorsal; whereas in most others* as the coati, raccoon, horse, &c. the dorsal furrow is of the same depth as the sternal; and . in the mole, murine, and leporine genera, it is deeper and more distinct than the sternal. In Birds and the carti¬ laginous Fishes the sternal furrow is deepest; in Rep¬ tiles the dorsal and sternal are nearly equally deep ; and in the osseous Fishes the dorsal is the deepest. On each side of the chord, also, there is a slight longitudinal depression commonly called the lateral (sulcus lateralis). The breadth and thickness of the spinal chord vary in dif¬ ferent regions. The lower cervical portion is in general the broadest and thickest; at the upper dorsal region it is some¬ what more slender, becoming thicker again at the lower dor¬ sal region; and after again becoming more slender in the lumbar region, it is expanded into a filamentous brush-like termination of different lengths in different classes, and even in different orders. These enlargements at the superior dorsal and lumbar regions are believed by Serres to cor¬ respond with the origins of the thoracic and pelvic extre¬ mities ; and each swelling he represents to predominate over the other, as the animal habitually employs the one or the other kind of members. It is remarkable in illus¬ tration of this principle, that in the Cetacea, which are void of pelvic extremities, the lumbar enlargement is wanting; and in the amphibious Mammalia, the pelvic extremities of which are feeble, this enlargement is also inconsiderable. The spinal or funicular brain consists of white cerebral matter on the surface and gray matter in the centre, the former being most abundant, unless in the sacral region, where both are nearly equal in quantity, or the gray rather predominates. In the Mammalia the gray matter is in greater proportion to the white than in the other three classes, in which the white matter progressively aug- Compara.! ments. t've The sternal and dorsal longitudinal lines indicate the J original formation of the chord in two lateral portions, ^ * with an intermediate cavity denominated the spinal canal. This exists only during the formation of the chord in the human foetus, in some of the Mammalia when adult, and in the other three classes. Its disappearance in the hu¬ man foetus, and in that of various Mammalia, is repre¬ sented by Serres to depend on the progressive deposition of gray cerebral matter on the inner or central surfaces of the component pillars of the chord. Though obliterated in man, in whom the gray matter is abundant, it is not hermetically sealed in the monkey tribe, in which some traces of it are left. In the Amphibia and Cetacea it is larger than in the monkey tribe; its diameter augments in the Carnivora, feline, canine, and ursine, in which the gray matter is thinner than in the former; and in the Rodentia it is said to be largest of all among the Mam¬ malia. Lastly, in the birds, reptiles, and fishes general¬ ly, in which the gray matter is scanty and the white pre¬ dominates, the spinal canal is large and distinct. The lateral regions of the spinal chord are connected with a double series of nerves by means of two rows of ner¬ vous filaments, an anterior and posterior, separated by a longitudinal membrane of fine white tissue, with a serrat¬ ed or festooned border. This membrane, which is named the denticulate, is the same in the Mammalia and Birds. That these nerves do not issue from the spinal chord, must be inferred not only from the phenomena attending the original developement of the nervous system, but especial¬ ly from what is observed of their comparative size, and that of the chord in the inferior classes. In that of fishes especially, while the chord is small and slender, and by no means fills the vertebral canal, it is remarkable that the nerves which supply the voluntary muscles are ex¬ ceedingly large. Thus, in many both of the cartilagi¬ nous and osseous divisions, as the sturgeon, dog-fish, ray, wolf-fish, cod, &c. the nerves which supply the pectoral fins are large, broad chords, two or three of which seem to contain more substance than the whole spinal chord itself. The chord is expanded at its cephalic end into a thick Bulb oftlf eminence denominated the spinal bulb, the surface ofch°rd. j , which presents three pairs of eminences. These eminences are developed in various degrees in the different classes and orders. In some of the Mammalia the anterior pyramids or pyramidal bodies are more distinct than in man, for instance the ape tribe, the Cetacea, Carnivora, Ruminants, and Rodentia. They are small in most Birds, all the Reptiles, and the Cartilaginous Fishes. In the osseous division they assume the appearance of two parallel chords at the base of the brain. The olivary bodies, less prominent in the ape than in man, are still less so in the Cetacea, and progressively through the Amphi¬ bia, Carnivora, Ruminantia, Rodentia, and Insecti- vora, and the other three great classes of vertebrated ani¬ mals. Conversely, the restiform bodies or posterior pyra¬ midal eminences, from man through the ape tribe, the Cetacea, Amphibia, to the Ruminants, Carnivora, and Rodentia, increase in size. Though the brain of the Mammalia presents the same parts, and is arranged nearly in the same order, as that of man, it varies in its proportions to the rest of the body; in its proportions to the cerebellum and spinal bulb ; in ge¬ neral figure; in the presence, absence, and number of con¬ volutions ; in the configuration of its central surface; in the communication of its central with its external surface ; and in the manner of its connections with the cerebral nerves. As it is impossible in this sketch to examine all 1 ANATOMY. 31 live jiiatomy. On para- these circumstances fully, we shall confine our attention to the notice of a few only. It is not easy to ascertain the proportion of the mass of the brain to that of the rest of the body. Excluding as much as possible the ordinary sources of fallacy, in small animals the brain is proportionally larger; yet in this respect man is surpassed only by a small number of ani¬ mals, habitually lean, and with little muscle, as bats, small birds, &c. While the proportion of brain in man to that of the whole person varies from a 22d to a 35th part, that of the monkey tribe varies from a 22d to a 42d part; and in the baboon it is only the 104th part of the body. Among the Mammalia, the Rodentia have in general the largest proportion of brain, and the Pachydermata the smallest; and while the hare has a brain about the 300th part of the size of the body, that of the elephant, the most sagacious of animals, is about the 500th part the size of his body. It is also remarkable, that while the brain of the horse is only a 400th part of the size of his body, that of the ass amounts to a 254th part. The Reptile brain becomes excessively small, that of the turtle being rather more than the 5000th part the size of his body; and in some of the Fishes, not all, it appears to attain the maximum of decreasing proportion, that of the tunny be¬ ing so small as the 37,000th part of his body, while the brain of the carp is so large as to approach the proportion ’ of the elephant. It may be doubted whether, under such circumstances, any precise conclusions can be drawn from results so variable and so little to be expected. The proportional weight of the brain to that of the cere¬ bellum is, excepting in the case of one species of ape, the saimiri, greater in man than in any other animal. The ox is equal to man in this respect, and the dog approaches him. The animals most remote are the Rodentia, as the beaver, rat, and mouse, &c. The convolutions, which are so numerous and so deep in man, diminish both in number and size in the Quad- rumana and Carnivora, and are nearly obliterated in the Rodentia. In the Ungulated animals, however, and especially in the Ruminants and the horse, the con¬ volutions are numerous; and even in the dolphin among the Cetacea, they are numerous and deep. In all the Mammalia the cerebellum is foliated. On the whole, the peculiar character of the brain of man and the ape family consists in the existence of the posterior lobe and digital cavity. The brain of the Zoo- phaga is remarkable for the small size of the nates or anterior pair of the bigeminous eminences in proportion to the testes or posterior pair. In the Rodentia the or¬ gan is distinguished for the large size of the nates, and the want or superficial nature of the convolutions. In the Ungulated division ,of animals, i. e. Pachydermata, Ruminantia, and Solidungula, the brain is remark¬ able for the large size of the nates combined with the number and depth of the convolutions; while that of the Cetacea is remarkable for its height and breadth, and the want of olfactory nerves. It is further to be observed as a general distinction between herbivorous and carni¬ vorous or zoophagous animals, that in the former the nates are larger than the testes, whereas in the latter the testes are largest. Lastly, Man and the Quadrumana are the only animals which possess genuine olfactory nerves. In the other quadrupeds they are represented by the mam¬ millary processes of the ancients; and in the Cetacea they have not yet been unequivocally demonstrated. The brain of Birds is at once recognised by consisting of six distinct tubercles, two representing the cerebral hemispheres, two representing the optic eminences, one the cerebellum, and one the bulb of the chord. The hemispheres are void of convolutions, but the cerebellum is marked by transverse parallel striae corresponding to Compara- the lamince of the mammiferous brain. There is neither tive middle band (corpus callosum), vault, nor septum. The ^natorny- ceiling or vault of the aqueduct or passage from the third to the fourth ventricle is not, as in the Mammalia, sur¬ mounted by the bigeminous eminences, but is merely a thin plate corresponding to the valve. Each optic emi¬ nence contains a cavity communicating with the others by the Sylvian aqueduct. The anterior eminences (cor- pora striata) are not striated with alternate white and gray matter, as in the Mammalia. Between the anterior and the optic eminences are four rounded tubercles, best seen in the ostrich, which are to be regarded as entirely hete¬ rologous to the structure of the mammiferous brain, and connecting the cerebral structure of Birds with that of Reptiles and Fishes, in which also these tubercles are ob¬ served. The Reptile brain is smooth and unconvoluted. The optic eminences, which are situate behind the hemispheres, are uncovered, and contain a ventricle communicating with the third. At the extremities of the latter are the anterior and posterior commissures, but there is neither soft commissure nor bigeminous eminences. The hemi¬ sphere presents an anterior eminence, which, however, in the brain of Birds is unstriated. The cerebral valve is, like that of Birds, unsurmounted by bigeminous emi¬ nences. In the class of Fishes the structure becomes still more simple. The tubercles of which the brain consists are placed in a row; and their increase in number only de¬ monstrates the decomposition of the organ, and its reso¬ lution into simple integrant parts. The two representing the hemispheres are ovoidal, unconvoluted, and contain a ventricle, in which is seen the eminence analogous to the striated bodies. The optic eminences, situate beneath the hemispheres, though small, contain each a cavity, as in the two oviparous classes already noticed. Lastly, there are in several genera, under the common vault of the hemispheres, occasionally two, occasionally four tu¬ bercles, variable in shape and proportions, but which would be analogous to the bigeminous eminences, were they not, like those already mentioned in Birds, situate before and above the optic chambers. In the cartilagi¬ nous fishes, in which these tubercles are not observed, the anterior or striated eminences are obliterated. The cerebellum does not cover the fourth ventricle. Behind the cerebellum are two tubercles, which in the ray give origin to the fifth pair, and are very distinct in the pike, trout, salmon, and perch. These tubercles are peculiar to this class. The cavities in the interior of the optic eminences in Birds, Reptiles, and Fishes, are observed in the foetal brain of the Mammalia during its early growth. It is almost superfluous to mention, that, in the two warm-blooded classes, Mammalia and Birds, the brain, with its investments, fills completely the cranial and verte¬ bral cavities. In the Reptiles, however, in which the brain does not approach the cranial walls, we remark the first departure from this arrangement; and in the Fishes it is so completely violated that the brain and chord occupy but a small proportion of the cranio-vertebral cavity; and be¬ tween the former and the osseous walls there is a quantity of fine but very loose filamentous tissue, containing in its cells a large quantity of pellucid fluid. Though this ar¬ rangement gives this the appearance of a white jelly-like substance, it is not gelatinous, as is generally represented, but merely a pellucid fluid, sometimes pale straw-coloured, occasionally with a reddish tint, contained in numerous communicating cavities of a tissue which appears to repre¬ sent the arachnoid of the warm-blooded animals. ANATOMY. 32 Compara- The pia mater in the reptiles and fishes is reduced to a t^ve filamento-vascular web, accompanying the blood-vessels. The dura mater undergoes some peculiar modifications jn difFerent orders. In the duckbill a bony plate is con¬ tained between the laminae of the falx; and the same structure is found in the porpoise, perhaps in the other Cetacea. An osseous tentorium with a quadrangular aperture is found in the coa'ita and marten, and the feline and ursine families; and an osseous partition consisting of three parts is found in the dog, horse, Cape ant-eater (pryc- teropus), the wombat, and the seal. The bony tentorium is also found in the woodcock and others of the feathered class. In the red but cold-blooded animals the dura mater forms neither falciform nor tentorial process. On the nerves or ramified chords of the nervous system a few words must suffice. In fishes the tenth or pneu- mogastric consists not of a common trunk, but of three orders of filaments, the first and largest of which are dis¬ tributed to the gills, and correspond to the pulmonary nerves of the Mammalia ; the second, slender, are dis- Com pan ' tributed to the muscles of the tongue and the surface of tive the oesophagus; and the third terminate in a large nerve which traverses the body longitudinally immediately be¬ neath the lateral line. The phrenic nerve is wanting in birds, reptiles, and fishes. In the Mollusca the nervous system consists of a number of whitish cerebral masses distributed in different parts of the body, with one or two more conspicuous than the rest, and supposed therefore to represent the brain, placed transversely over the oesophagus, which it encom¬ passes with a nervous collar. In the Articulata the nervous system consists of two long chords extending along the belly, and expanded at various intervals into gangliform knots or enlargements. The first of these, which is situate on the oesophagus, rarely exceeds the others in size. Among the Zoophytes hitherto examined the nervous system assumes either a radiated or an arborescent form. PART II. comparative anatomy of the entrophic organs. CHAP. I. THE LIMITROPHIC ORGANS. SECT. I. Though genuine teeth are found in three classes of ani¬ mals only, viz. the Mammalia, the Reptiles, and the Fishes, yet all the orders of these classes are not provid¬ ed with teeth. Thus, among the first class, the ant-eater tribe, the pangolin, the echidna and ornithorhyncus, and the whales—among the second the cheloniads—and among the third the sturgeon—are altogether destitute of these organs. In all the invertebrated classes, the jaws, when present, are provided with notches varying in number. The echinodermata alone have genuine teeth, inserted in a mechanical apparatus very different from ordinary jaws. Though in general structure the teeth of the lower ani¬ mals resemble those of man, in some respects they differ considerably. These varieties consist either in some change or modification of the constituent parts of the teeth, or in the addition of some other substance to those parts. The first variety to be noticed is of the former descrip¬ tion. Bone of Though in the Quadrumana and Zoophaga the bony tooth. matter of the teeth is quite similar to that of man, in other orders this substance appears in the form of a very hard, compact, and more regularly fibrous substance than bone, and to which the name of ivory (ebur) is applied. It is chiefly in the canine or tearing teeth that this sub¬ stance is found to represent the bony pillar of the teeth ; and it is principally among the Pachydermata, and some of the Amphibia and Cetacea, that this change is ob¬ served. Ivory of The ivory of the elephant is the most tender, and that different which most rapidly becomes yellow on exposure to air. anxma s. jt jg reacjj]y distinguished from the ivory of other animals by the curve lines which radiate from the centre to the circumference of the tooth in various directions, and which form by intersection regular curvilinear lozenges. The ivory of the hippopotamus is greatly harder and whiter, and is on that account preferably employed for the preparation of artificial teeth. A transverse section of this substance shows since extremely delicate and re¬ gular. In this animal, also, not only the canine but the incisor teeth consist of this substance. The tusks of the Ethiopian boar (sus JEthiopicus) consist of ivory similar to that of the hippopotamus. In those of the ordinary boar, though no stria are recognised, there is sometimes a mix- the organs of bigestion.—§ 1. the teeth. ture of brown substance disposed in layers. The ivory of the teeth of the morse, though void of since, is compact and susceptible of polish nearly as brilliant as that of the hippopotamus; and its character is, that the central pillar of the tooth consists of minute round grains, indiscrimi¬ nately aggregated, like pebbles in puddingstone. The axis or pillar of the molar teeth of this animal, which are without internal cavity, consists of similar minute grains. The ivory of the dugong is homogeneous and without striae. That of the teeth of the white whale or cachalot resembles the bone of human teeth in its satin-like ap¬ pearance. The ivory of the tusk of the narwal is very compact and homogeneous in appearance. The most singular structure of teeth among quadrupeds is observed in those of the Cape ant-eater {prycteropus'). The teeth of this animal, which have the appearance of two cylinders conjoined, consist of an infinite number of minute straight parallel tubes, so that their transverse section resembles that of a rush. As these tubes are closed only at the triturating surface, it is there only that the tissue of the tooth is compact; and when the enamel is worn, the upper orifices of these tubes begin to be exposed. There is, therefore, no general cavity in the in¬ terior of the tooth. These teeth are also void of root. A similar tubular structure is observed in the two molar teeth of the ornithorhyncus, and in the teeth of some fishes. The enamel {lamella vitrea, cortex striatus) presents Enamel peculiarities in the lower animals, as well as the bone of the tooth. While the enamel of the human tooth is confined to the crown, in several of the lower animals, as the morse, it envelopes the tooth all round; and in the molar teeth of this animal, which, indeed, are void of cavity, it is thicker under the root than at the crown. A similar ar¬ rangement is observed in the old or adult teeth of the cachalot, which, when their cavity is obliterated by the full deposition of osseous substance, are also covered with enamel below. The texture or constitution of the enamel is best seen in the grinders of the elephant. The section of a tooth in the germ exhibits fibres similar to those of asbestos or fine velvet. The fracture of the enamel is more dis¬ tinctly fibrous than that of the bone, and the fibres are everywhere perpendicular, or nearly so, to the surface of ANATOMY. 33 ‘H 0it>para- the tooth. The hardness of this substance may be in- ive ferred from the fact that it strikes fire with steel. These tomy. component fibres, however, are not always rectilineal. Most frequently they describe curves with the convexity of incurvation towards the crown and the concavity to¬ wards the root. This arrangement at least is observed in the ruminants. The distinction between the enamel and bony matter is recognised by a gray line, and another whiter which belongs to the latter substance. The enamel varies chiefly in thickness in different ani¬ mals. The tusks which project from the mouth are ge¬ nerally observed to be less white, less hard, and more similar to bone or ivory than the other teeth; and on this account, probably, the existence of enamel has been de¬ nied in the tusks of the elephant. It is nevertheless cer¬ tain that the external layer of these tusks presents radiat¬ ing fibres, though it is by no means so hard, or possesses the same grain, as the enamel of the other teeth. Enamel is more apparent, though thinner, in the tusks of the morse, dugong, and boar; and it is quite as distinct in those of the hippopotamus as in the other teeth of that animal. Lastly, the enamel of the teeth of the cachalot, which is very thick, shows in its section only stria parallel to the surface of the osseous substance. Teeth may be distinguished according to the mode in which their component tissues are arranged into three sorts. ls£, When the enamel invests the axis all round, and does not penetrate the latter, the tooth is said to be simple (dens simplex). Such is the character of the human teeth, and those of the Quadrumana and Zoo- phaga, and several other animals, and all the reptiles, pound 2d, When the enamel is folded as it were round the bony part, but without inclosing it, so that the latter forms a continuous band several times folded on itself, and sec¬ tions of the tooth in every direction divide repeatedly the component substances, the tooth is said to be compound or complex (dens multiplex vel compositus). A good exam¬ ple of this structure is seen in the grinders of the ele¬ phant. 3c?, When the base or root of the tooth is simple, and the folds of the enamel and bone penetrate only to a certain depth, they are said to be semicompound. Ex¬ amples of this modification of arrangement are seen in the grinders of the ruminating animals. In the compound, and part of the semicompound teeth, the enamel is covered by a third substance; and as the latter is arranged, especially in the former sort, so as to leave intervals between it and the next layer of enamel, this substance serves to fill all these intervals, and conso¬ lidates the component lobes of the tooth even before their osseous parts are united below. This substance, which is denominated by Cuvier cement, by Tenon cortex osseus, and by Blake crusta petrosa, though less firm than either bone or enamel, is dissolved by acids more slowly than the former, and sooner becomes black in the fire. In the teeth of the elephant and cabiai it forms half their mass at least. In most genera it presents no apparent organiza¬ tion, and resembles a sort of crystalline tartar incrusted on the tooth. In the cabiai, however, it presents nume¬ rous pores very regularly arranged. Tenon was of opi¬ nion that it arose from ossification of the membrane which enveloped the tooth ; blake ascribed it to deposition from the opposite surface of the enamel membrane; but Cuvier ascertained that it is deposited by the same membrane and the same surface as the enamel. This accurate observer found, on inspecting the germs of the teeth of the elephant, that when the internal membrane of the dental capsule has deposited the enamel, it undergoes a change of structure, and becomes thick, spongy, opaque, and reddish, to furnish the cement, which is then deposit¬ ed, not in regular crystalline fibres, but in random drops. VOL. m. t ce¬ lt or The teeth of the Reptiles consist of hard, compact, os- Compara- seous matter, invested by a thin covering of enamel, and tive without cement. Anatomy. The teeth of Fishes vary much in structure. They are oj.. either simple or compound. The simple teeth are those fishes. which consist of bone invested by enamel. They may be distinguished into two species, according to their mode of attachment. The first are the simple teeth, which are not implanted in alveoli, but merely attached to the gum, or fixed by articulation to the jaw, as those of the shark tribe ; the second are simple teeth growing in alveolar cavities, as is observed in the majority of fishes, the pike, dory, &c. The simple teeth attached to the gum are chiefly distinguished by their fibres intersecting in the manner of the cancellated tissue of bones, and being therefore at first light, porous, and spongy, and becoming afterwards uniformly hard and compact like ivory. The compound teeth, which consist of an infinite num¬ ber of minute tubes mutually aggregated and invested by a common covering of enamel, form plates of different sizes, adhering to the bones of the jaws or palate by an intermediate membrane only. In some they affect the disposition of the quincunx; in others they occupy the whole breadth of the upper jaw at least, as in the ray as seen on the small scale, and in the same manner in larger fishes ; others are in straight transverse bars ; others as¬ sume the shape of a circular segment, or the figure in heraldry denominated the chevron. In the wolf-fish the jaws are provided with eminences composed of fibres or tubes proceeding from the base to the circumference, and which are connected to the jaw by a substance more spongy than the rest of the bone. After their first formation the teeth retain nearly their Mechani- original shape in the Zoophaga, man, and the Quad-cal detri- rumana. In the two latter only their crowns begin to be the worn, rendering the incisor and canine less pointed by thetee use of food partly vegetable; but in the zoophagous tribes they undergo no detrition whatever. In the herbivorous animals, however, the crown begins to undergo detrition more or less rapidly; and in no long time the superior layer of enamel is entirely worn off, and the surface of the tooth exposes the succession of bone or ivory, enamel, and cement. These substances are well seen in the teeth of the Rodentia, for instance the hare; those of the Pachydermata, as the elephant; the Ruminants, as the stag, sheep, and ox ; and the Solidungula, as the horse. In all these animals the enamel, which is hardest, forms prominent lines or ridges; while the bone and cement are indicated by depressions. This detrition, which is purely mechanical, might pro¬ ceed to such an extent in the herbivorous quadrupeds as to destroy the whole of the crown of the tooth,'and leave the process of mastication to be performed by the jaws only. It appears to be chiefly to obviate this incon¬ venience that the dentition of the elephant, the Ethiopian Successive boar, and perhaps all the Pachydermata, is conducted dentition in a successive manner through a series of six or eight0: !e sets of teeth at least. In the former animal, in which 1 this process has been best observed, and was ably explain¬ ed many years ago by Mr John Corse Scott (Phil. Trans. 1799), each half-jaw, whatever it contains, exhibits at one time only one complete grinder and part of another behind it, the prominent parts of which are placed ob¬ liquely to the horizon, forming an inclined plane, so that the anterior parts are worn before the posterior. The anterior complete one, which is employed in mastication, undergoes progressive detrition till its anterior portion is worn down to the level of the jaw. In this state the fangs of the anterior part of the tooth begin to diminish, render¬ ing the tooth narrow before; while the crown of the poste- E ANATOMY. 34 Compara- rior begins to be worn, and undergoing the same detri- tive tion, the posterior fangs also begin to give way. While Anatomy, procesSj however, is advancing, the posterior tooth, 'i of which only the anterior part was appearing above the gum, gradually rises, with its crown forming a plane in¬ clined from before backwards, similar to that of the ante¬ rior grinder. When this posterior tooth has been raised sufficiently to allow its anterior margin to be used in mas¬ tication, the residue of the anterior tooth drops out al¬ together, and the posterior one continues to rise and ad¬ vance rapidly, until it is completed, when it is found to be much larger than the previous tooth, and to consist of a greater number of plates of ivory and enamel (denticuli). In no long time this new tooth, which undergoes the same process of detrition, is succeeded by another one, the anterior margin of which rises first behind the poste¬ rior one of its predecessor, and which passes through the same stages of growth, detrition, and shedding. This pro¬ cess is repeated at least seven or eight times, and each succeeding tooth is larger, and contains a greater number of ivory and enamel plates than its predecessor. The elephant has thus 7 or 8 grinders in each half-jaw, or 28 or 32 grinders respectively; yet there are never more than one tooth and part of another, or at most two, that is, eight teeth in the upper and lower jaws, at the same time. Though the disappearance of the fangs of the an¬ terior tooth is ascribed to absorption—which indeed is a good general name—yet the true reason is the fact that the maxillary or dental vessels of the elephant are unable to sustain more than one tooth in each half-jaw at once; and that since these vessels gradually transfer their blood to the new tooth, while those of the old one shrink and are obliterated, the new tooth grows as the old one is actually dehematised or atrophied. The order in which the teeth of the elephant succeed each other is nearly the following. The first or milk grinder, composed of 4 eburneo-vitreous plates {denticuli), cuts the gum eight or ten days after birth, is well formed in six weeks, and completely out in three months. The second, which con¬ sists of 8 or 9 plates, is completely exposed at the age of two years; the third, consisting of 12 or 13 plates, at six years. The fourth to the eighth grinder consist of plates varying in number from 15 to 23; but the period at which these teeth appear has not yet been determined. This process has been shown to have taken place also in the gigantic fossil animal named the mastodon. A similar process of displacement and renovation takes place in the poison-teeth of serpents, and in the teeth of the shark, ctiodon, and tetraodon tribes. In the wolf¬ fish {anarrhicas lupus) the teeth are shed along with the spongy membrane in which they are contained, exactly as the horns of the stag. Dentition In the horse, in which the process of dentition has been of the carefully observed, it is usefully employed to determine horse. ^}ie age 0f (;]ie animal. The milk incisors appear at the end of 15 days; the four middle ones, or the nippers, are shed at 30 months; the four following ones at 42 months; and the four external, or the corner teeth, at 54 months. The permanent corner teeth do not grow so quickly as the other incisors ; and by these especially the age of the horse is determined. At first they scarcely rise above the jaw. Their middle then presents a hollow filled with blackish tartar, the margins of which are worn down as the tooth rises from the gum, and is rubbed against the corresponding one; and it diminishes pro¬ gressively from 54 months to 8 years, when it is alto¬ gether obliterated. The hollow of the other incisors is obliterated at a later period than that of the corner ones; and the age of the animal is then estimated from the length of the incisors, which continue to increase. The first two molar teeth appear in each jaw and on each side about the 8th day, the next at the 20th, and Compara. the complementary or small anterior grinder about the five 5th or 6th month. The first posterior molar appears Anatomy, about the 11th month, and the second in the 20th. At the 30th or 32d month the first two milk grinders are shed, the third in the 3d year; and about the 5th or 6th year the last posterior grinder appears. The milk grinders are longer from before backwards than the permanent ones, which are themselves contracted in this direction, as they are pressed by the posterior grinders; from which it results that the dental crowns of young horses are ob¬ long, while those of the old are quadrangular. In the Mammalia the teeth are always implanted in the jaw-bones, and never, as in other animals, in the tongue, palate, &c. The only exception to this rule is the echidna. The three kinds of teeth, incisor, tearing, and grinder, Arrange, are found together only in Man, the Quadrumana, thementof Zoophaga, the Pachydermata except the elephant^ and two-horned rhinoceros, in the hornless Ruminants, and in the Solidungula ; but, of all these animals, in man only are the three forms of teeth arranged in an unin¬ terrupted series, and in such a manner that those of the lower jaw are applied to those of the upper. In one other animal only, now extinct, the anoplotherium, is this conti¬ nuity of arrangement observed. In the Quadrumana and Zoophaga, and all those in which the canine are larger than the other teeth, there is a gap on each side of the jaw to receive the canine of the opposite one. In the ursine genus there is a large empty space behind each canine tooth. In the hedgehog, shrew, phalanger, and tarsier, in which the canine are shorter than the other teeth, a space is left between them and those opposite. In the maki tribe, proper bat, colugo, and camel, there is a large interval between the upper incisors. Last¬ ly, the Ruminants want the incisors of the upper jaw, and the morse those of the lower. Some animals provided with the three classes of teeth lose the incisors at a certain age; for instance several of the bat tribe, and the Ethiopian hog. Other Mammalia have only two sorts of teeth, for instance incisors and grinders, separated by an interval without canine, as the wombat and all the Rodentia, in which there are only 2 incisors in each jaw; the kangaroo, which has two be¬ low and 6 or 8 above; and the cavy or hyrax genus, which have 2 above and 4 below. The elephant has grind¬ ers and two tusks planted in the superior intermaxillary bone, but no inferior incisors or canine teeth. Animals may possess grinders and canine teeth with¬ out incisors, as the sloth tribe and the dugong. The grinders, which are most essential, are most rarely want¬ ing ; and when others are deficient these are present, as in the armadillo tribe, the orycteropus, the ornithorhyncus, two-horned rhinoceros, and lamantin. The jaws of the dolphin are provided with uniform conical teeth all round, while the cachalot or white whale has them in the lower jaw only. In the narwal there are only two long spiral tusks implanted in the intermaxillary bone, and of these one is often wanting. Lastly, teeth are entirely wanting in the ant-eater tribe, Absence of pangolin, and echidna, which are therefore arranged among the teeth, the Edentata. In the whale the teeth are represented by plates of the laminated, fibrous, bluish substance dis¬ tinguished by the name of whale-hone. § 2. ORGANS OF INSALIVATION. Under this head ought to be noticed the modification which the salivary glands undergo in the lower animals. Our limited space, however, obliges us to proceed imme¬ diately to notice the peculiarities of the other divisions of the alimentary canal. I ^ St:nach. J S'i ple c stcnach. i Blcular i. stonach. ANATOMY. 3o § 3. CESOPHAGUS, STOMACH, AND INTESTINAL TUBE. The muscular tissue of the oesophagus consists, in most of the Mammalia, of spiral fibres twisted in two opposite directions, the external from before backwards, the inter¬ nal from behind forwards. This arrangement, which was first observed in the Ruminantia, was supposed to ex¬ plain the process of rumination. This opinion, however, is refuted by the fact that the arrangement is not con¬ fined to this order, but is very general among the zoopha- fous and other animals which do not ruminate. In the angaroo the direction of these fibres is, as in man, trans¬ verse in the internal layer, and longitudinal in the external. The cesophageal mucous membrane is covered by epi¬ dermis, which extends to the cardiac opening of the stomach in man, the Quadrumana, and all the Zoopha- ga. This membrane, as well as the mucous, is thrown, by the action of the muscular tunic, into longitudinal folds, which are effaced only when the oesophagus is distended. In the tiger, lion, and lynx, there are large transverse val¬ vular folds, and smaller ones in the civet and couguar— an arrangement connected probably with the carnivorous habit. The stomachs of the lower animals vary considerably in shape, in the insertion of the oesophagus, in the disposi¬ tion of their muscular tunics, and in the simplicity or complication of their cavities. These characters it is im¬ possible in such a sketch as the present to consider in de¬ tail ; and we shall confine our attention to those peculiari¬ ties which are most striking in the digestive organs of the animal world. The stomachs of the Mammalia may be distinguished into the simple and compound. Those of man, the Quad¬ rumana, zoophagous and most of the herbivorous tribes, belong to the former order. This simple form of stomach, however, may be generally distinguished into two parts, a cardiac and a pyloric, more or less separated from its other by a central transverse contraction of its annular muscular fibres. This is particularly seen in the horse, man, murine family, and many other animals which occa¬ sionally feed both on animal and vegetable matter. In the human stomach this contraction is represented in Plate XXXVI. fig. 4. In the porcupine, however, there are three pouches. This contraction depends on a strong annular band of muscular fibres at this part of the organ. In the pure carnivorous animals, however, as the feline family, the annular fibres, which are very thick, are nearly equally so from the cardiac to the pyloric end. The compound stomachs, or those which contain more cavities than one, are found in the sloths, and the ruminant and cetaceous animals chiefly. In the first tribe the stomach of the Unau, or two-toed sloth, is two-fold. The first cavity is large and globular, but tapering behind into a conical appendage, separated by a semilunar fold; while a large cul de sac on the left of the cardia opens into a canal which proceeds at first back¬ wards, and then turning to the right, enters the second cavity by a narrow aperture. The second, which is small, tubular, and folded under the former from left to right, is distinguished by a semilunar fold into two halves, the first of which opens into a small cul de sac on the right side of the first cavity. The inner membrane of both cavities is smooth, and without villi. A similar arrangement is found in the Ai, or three-toed sloth, with this exception, that the appendage of the second gastric cavity is divided into three compartments by two longitudinal bands. This ca¬ nal seems analogous to the arrangement of the ruminating stomachs, in so far as it may allow the alimentary matters to pass occasionally from the oesophagus directly into the second stomach. The stomach of the hyrax, ashkoko, or Cape cavy, also consists of two pouches, separated by a middle partition, Compa,ra- in which there is an aperture for mutual communication. tive In the hippopotamus the cardia communicates with three Anatomy, pouches, two of which only are cognizable without, and ''“■’''V'’-'' with a long tubular bowel, the interior of which is divided across by several valvular folds. In the kangaroo the stomach receives the oesophagus near its left extremity, which is small and bifid (Plate XXXV. fig. 8); and forming a larger cavity on the right, passes upwards, making a turn, and crosses to the left be¬ fore the oesophagus, makes another turn, and again crosses the mesial plane to the right, where it terminates in a ta¬ pering cavity at the pylorus. In this course it presents internally a longitudinal band (l, l, l), extending all round to near the pyloric end, and crossed by valvular membra¬ nous folds, which divide the cavity into cells not unlike those of the colon, especially in the horse. The mucous epidermis is continued from the oesophagus over the space marked c, c. The stomach of the Ruminants consists of four distinct Quadrilo- but communicating cavities. The first, denominated the cular sto- Paunch (xo/X/a, rumen, penida, ingluvies; la pause, Vherbier, ™ach of la double), is a large bag occupying the left side of the ^a®^gumi* abdomen chiefly, marked externally by two saccular appen¬ dages, and separated within into four parts. (Plate XXXV. fig. 1, a, a, A, a.) Its inner surface, upon which the epider¬ mis is continued, is occupied by flat papillce. By a pretty wide aperture (b, b), with rounded margins, this communicates with the second cavity named the Kings- hood (xsxpu are Provided with fimbriated or pole!6 aa" cibated processes attached to the neck, and which are in all respects sinv'lar to the gills of fishes. These gills dis¬ appear as the animal grows; and wdien it assumes the true ranine or reptile character, vesicular lungs like those of other reptiles, and which had continued in a hitherto latent and rudimental state, are developed, and the ani¬ mal breathes as others of the same tribe. A peculiar form of respiratory organ is found in the lamprey or seven eyes, and the two species of hag-fish, (Myxine, Lin.; Gastrobranchus, Bl.; and Gastrobranchus Dombey). The former has on each side seven apertures leading into cylindrical tubes, in which the branchiae are contained. (Plate XXXVII. fig. 3.) In the two species of hag these tubes are dilated into ovoidal cavities, in which the water is received, and on the membrane of which the branchial vessels are distributed. In this re¬ spect, therefore, the hag-fish approaches to the mode of respiration among the cephalopodous Mollusca, in which the bratichice are inclosed in a cavity. Lastly, in the Aphrodite aculeata, which may be taken as an example of the respiration of worms, there is a series of tubes like tracheae and bronchi, proceeding from the surface to the interior, and in which the water containing the air requi¬ site for respiration is received. (Fig. 6.) the erectile movable tubular fangs. Fig. 15 is the head of Compaq the innocuous, and 16 of the poisonous serpent. five' Anatom PART III.—REPRODUCTIVE ORGANS. Under this head we mention only the nipple-bag (marsu- pium mammillare), or secondary uterus of the Marsupial animals (Plate XXXVII. fig. 7); the nipples (fig. 8); and the manner in which the foetal animal, in a very imperfect and embryal form, becomes attached by the mouth to the nipples (fig. 10). The Marsupium, therefore, ought to be re¬ garded, not as a mere pouch in which the young may take refuge after they are grown, but as a subsidiary uterus, com¬ bining the character of the Mammae of the other orders.* The Mammalia are peculiar in possessing a uterus. In the other classes this organ is withdrawn, and the ovary (fig. 11) and oviduct alone are left. In the ovo-viviparous animals, as the ovo-viviparous shark, the oviduct (fig. 12) resembles that of the common fowl. In the lower classes the ova are hatched out of the body entirely. Urinary organs of the ovi¬ parous classes. Poison gland and fangs of serpents. SECT. III. SECRETORY ORGANS. Under this head ought to be described the urinary or¬ gans of the four vertebrated classes. Those of the Mam¬ malia agree in consisting of kidneys more or less lobu- lated, ureters or excretory tubes, a reservoir or urinary bladder, and a urethra opening on the same mucous sur¬ face with the organs of generation. In the three oviparous classes considerable changes are made. Though in Birds and Reptiles the glandular organs denominated kidneys are left in the shape of aggregated glands with the two excietoiy tubes, the bladder is withdrawn, and the ureters open in the cloaca. The only apparent exceptions are the ostrich and cassowary, in which the cloaca is so orga¬ nized that it may serve as a bladder or temporary recep¬ tacle of the urinary secretion. In the Reptiles the pre¬ sence of this organ is variable, being found in the Che- loniad and Batrachoid ; and the iguana, tupinambis, chameleon, stellio, and dragon, among the Saurial tribe ; but wanting in the crocodile, lizard, agami, gecko, and the whole Ophidial tribe. In Fishes it is not less variable. While the ray and shark tribe are destitute of bladder, and the ureters terminate in a cloaca, this receptacle exists in the sea-devil, lump-fish, globe-fish, and others of the cartilaginous division. A peculiar secreting organ, deserving notice, is the poi¬ son gland of the poisonous serpents. It is a glandular body situate on each side above the upper jaw, behind and be¬ low the eyes, with a considerable cavity, which opens into a long excretory tube, lying along the outer surface of the upper jaw, and opening in the tubular tooth, represented at fig. 17 and 18; and which is movable in an articulation, and may be erected, as in fig. 18, or depressed, as in 16, at the will of the animal. The poisonous serpents are there¬ fore distinguished from the innocuous by the presence of In the space assigned to this article, it was impos¬ sible to treat fully of a subject so extensive as the struc¬ ture of the animal world; and while the author has ar¬ ranged its divisions in such a manner as to show in what order it may be most easily and advantageously studied, he has introduced only those topics which are most indis¬ pensable, and most require illustration. For more com¬ plete details, therefore, he rfefers the reader to the follow¬ ing works. 1. Lemons d'Anatornie Comparee de G. Cuvier, Membre de 1 Institut National, &c.; recueillies et publiees sous ses yeux par C. Dumeril, chef des Travaux Anatomiques, &c. Cinq tomes. Paris, tome i. 1799;—tome v. 1805. 2. Blumenbach’s Manual of Comparative Anatomy ; with additional Notes by William Lawrence, Esq. F\ R. S. Se¬ cond edition, revised and augmented by William Coulson. Lond. 1827, 8vo. The notes are derived chiefly from the work of Cuvier and the papers of Sir E. Home in the Philosophical Transactions. 3. Gore’s Translation of Ca- rus’s Introduction to the Comparative Anatomy of Animals. Lond. 1827, 2 vols. 8vo. The arrangement of this work, in which the author examines the forms of organs as they ascend, from the lowest to the highest classes, diminishes its general interest. 4. Lectures on Comparative Anato¬ my, in which are explained the Preparations in the Hunte¬ rian Museum. By Sir Everard Home, Bart. Lond. 1823, 6 vols. 4to. This work consists of the papers read by the author at the Royal Society, and published in their Iransactions. Though entitled, therefore, Lectures on Comparative Anatomy, it embraces a much more exten¬ sive field, and contains a great number of physiological and pathological papers. This renders it at once rather desultory and prolix. It contains, nevertheless, a-great number of facts illustrative of peculiarities of structure in the animal world; and it is particularly valuable by the number of engravings with which it is embellished. It can scarcely be said to possess any arrangement whatever. 5. Recherches sur les Ossemens Possiles, oil Von retablit les characleres de plusieurs animaux dont les Revolutions da Globe ont detruit les especes. Par M.le Bar. G. Cuvier, &c. Nouvelle edition. Tome i. 1822, Osteology of the Elephant, Hippopotamus; tome ii. partie i. 1822, Osteology of the Rhinoceros, Horse, Hog, Daman, and Tapir; tome iv. 1823, Osteology of the Deer and Ox, the Bear, Hyena, Lion, Glutton, Wolf, and Dog; tome v. partie i. 1823, Osteology of the Reptiles, the Ichthyosaurus and the Plesiosaurus; partie ii. 1823, Rodentia, Edentata, Monotrema, Amphibia, and Cetacea. Paris, 1824. I hese papers contain much accurate osteological descrip- tion. (v.) 41 VEGETABLE ANATOMY. Int 'iluc- All the plants which collectively form the vegetable fry kingdom have been arranged under two great divisions. Kearks. 'i'|lose which possess visible organs of reproduction, as stamens and pistils, have been termed phcenogamous, and constitute the first 23 classes in the sexual system of Linnaeus; while those in which the reproductive organs are either obscure or have not yet been discovered, have been termed cryptogamous, and form the 24th class of that system. Humboldt estimates the total number of known species at 44,000; of which 38,000 belong to the former division, and 6000 to the latter. M. Brogniart, jun. sup¬ poses the present known Flora to embrace 50,350 species, of which 10,200 may be considered as cryptogamous, and 40,150 species as belonging to the phaenogamous division. On the present occasion we propose to describe the struc¬ ture of those plants only whici. are included in the division of phaenogamous plants. Of the vast number of plants which cover the surface of our globe, it belongs to the botanist to describe the exter¬ nal forms in such manner as may serve to discriminate species, and assign to each its place in a methodical sys¬ tem of arrangement: it is the province of the anatomist to demonstrate, by dissection, their internal structure, so as to prepare the way for a rational explanation of their functions. In pursuing this object we may examine suc¬ cessively each of the parts as they severally present them¬ selves to view; or we may, in the first place, study the elementary organs common to all plants, and then consi¬ der how their combinations form the different parts of vegetables. The former method is the analytic, and was necessarily that of the first observers ; but, now that all the different parts are recognised as being formed out of the same elementary organs, we gain, both in conciseness and clearness, by adopting the latter or synthetic mode. We shall accordingly describe, first, the elementary organs, and the primary textures which they form; and proceed afterwards to the consideration of the individual members and more complex organs of the plant. These elementary organs have been denominated vessels and cells; and they form, either singly or conjointly, what are called the vascular system and cellular tissue of plants. Their combination gives rise to certain textures, which appear in the well-known forms of skin, of bark, and of wood. On these we shall bestow the appellation of common textures, and exhibit a general view of their structure and disposition in the several parts of the ve¬ getable body. A brief description of some minuter parts, as of hairs and glands, will terminate this division of the subject. In the second part we shall begin with a description of the general structure of seeds, and afterwards treat more particularly of those bodies under the two great divisions of monocotyledonous and dicotyledonous seeds, tracing also the changes of form and of structure which they exhibit in their evolution and progress to the state of the mature plant. The structure of the mature plant itself will next claim our attention ; and we shall accordingly exhibit the anatomy of its several members, as of the trunk, the branch, and the root, in their more remarkable varieties and forms. After this, the structure of the organs that spring from these several members, as buds and bulbs, leaves, flowers, and fruits, will be separately and distinctly examined; and having thus followed the progressive changes of form and of structure exhibited in the several stages of vege- VOL. in. table existence, we shall terminate our descriptions by Introduc- anatomical representations of the organs in which the tory seed was produced, and the series of appearances succes- Bemarks. sively displayed in its formation. Through the whole of the descriptive detail we shall adhere as closely as we can to the language of demon¬ stration ; supporting and illustrating our representations of structure by continual reference to figures, selected in great part from authors of repute, and in some instances from dissections made by ourselves. We are aware, that of many reputed anatomical facts very different represen¬ tations have been given, all equally professing to rest on microscopical observation. In such circumstances, we can do no more than report concisely the statements of different observers; but shall dwell chiefly on those de¬ scriptions and representations which seem best entitled to credit, and appear most conformable to the analogies of other organized structures. When we consider the immense number of species that compose the vegetable kingdom, and call to mind that in form, in size, and in structure, each species differs from every other through every period of its existence, it must appear altogether impracticable to describe and delineate any considerable number. Fortunately, however, these diversities arise not from differences in the elementary organs, but chiefly from their varied proportion, disposi¬ tion, and texture. In numerous species the disposition of the internal organs is very similar, where the external form and texture widely differ. In other instances the arrangement and composition of the internal parts vary not less than that of the external figure. Of these varie¬ ties we shall exhibit different examples. In describing individual parts or organs, we might have brought many concurring examples, and exhibited many similar representations, to confirm the views of structure under consideration; but, in general, we have dwelt only on one or two examples, and these we have selected from plants which are either important in themselves, or whose structure has been most satisfactorily displayed, or which seemed to afford the best illustration of the peculiarities we were engaged in describing. From one example clear¬ ly given, the reader will readily apprehend the nature of analogous structures, and escape the perplexity and fatigue which unnecessary repetitions might occasion. Instead, also, of describing the vegetable at one or two stationary points of its existence, in some of which its size is so minute as to be scarcely capable of demonstration, we have followed it through the several stages of its growth. In this way we really study it as a living body, continually exerting its vegetative powers, and daily ex¬ hibiting the most striking variations in external form, and frequently in internal structure. We hope thus to have conferred an interest on the descriptive part, which may in some measure relieve the unavoidable dryness of ana¬ tomical details; to have exhibited, in some instances, clearer views of vegetable organization; and to have given a continuity to the subject which isolated dissections, at a few stated periods, could not alone have bestowed. It remains only to add a few remarks on the nomencla¬ ture employed in the present article. In the description of external parts we have adhered chiefly to the Linnaean nomenclature; but some of the terms employed by Linnaeus, in relation to anatomical structure, are exceedingly vague and inappropriate; others ANATOMY, VEGETABLE. 42 Introduc- are manifestly erroneous, and, however well suited to the tory purposes of botany, are not at all to be tolerated in any Remarks. thjng that aspires to correct anatomical description. In the anatomy of seeds we have adopted many of the terms employed by Gsertner, in his excellent work De Ffuctibus et Seminibus Plantarum, most of which had pre¬ viously been used by Malpighi and Grew. Thus, in every instance, we have exercised our own judgment in the selection of terms, and, where it seemed necessary, have subjoined the synonyms of different wri¬ ters. Though we presume not to say that we have uni¬ formly chosen the best, we trust they will always clearly express the idea we designed to convey; and that, in ge- Introduc. j neral, they have been used in one and the same sense, and tory ] in no other. Except in one or two trivial instances, we RemarH , have not ventured to introduce new terms, but have stu-V'^vv diously sought to avoid it, retaining even an inappropriate expression, sanctioned by use, if it did not, at the same time, lead to ambiguity, or convey an idea evidently false ; and we have in general resisted that torrent of new and barbarous terms, founded often on fancied refinements and pretended discoveries, with which several continental writers have of late attempted to deluge this branch of science. PART I. OF THE ANATOMY IN GENERAL OF THE ELEMENTARY ORGANS AND COMMON TEXTURES OF VEGETABLES. CHAP. I. OF THE ELEMENTARY ORGANS. Prelimi- Before we proceed to describe the structure of the nary ob- individual parts of vegetables, it may be useful to exhibit beivations. a genera| view 0f the elementary organs of which they seem to be composed. Such a view will prepare the reader for understanding more clearly the descriptive lan¬ guage hereafter to be employed, and will even much abridge the extent to which that description must other¬ wise be carried. Every one is familiar with the natural division of plants into herbs and trees, and is aware that, how differ¬ ent soever they may appear in form and texture, they all possess, in common, certain parts or members which we name the root, the trunk, and the branches, from which proceed the leaves, the flowers, the fruits, and seeds. Infinitely varied as these several parts are, in figure, size, and texture, they all originate from a few constituent or elementary organs, whose situation, proportions, and com¬ bination, give rise to all the diversity that we see. “ Up¬ on the anatomical analysis of all the parts of a plant,” says Grew, “ I have certainly found, that in all plants there are two, and only two, organical parts essentially distinct, viz. the pithy part and the ligneous part.” (Ana¬ tomy of Plants, p. 19.) “ And as every part hath two, so the whole vegetable, taken together,” he adds, “ is a composition of two only, and no more. All properly woody parts, strings, and fibres, are one body; all simple barks, piths, parenchymas, and pulps, and, as to their sub¬ stantial nature, peels and skins, are all likewise but one body; the several parts of a vegetable differing from each other only by the various proportions and mixtures, and variated pores and structure, of these two bodies.” (Ibid. p. 47.) In the anatomical descriptions of Malpighi, the compound structure is resolved, in like manner, into two constituent parts, called by him the ligneous and utricu¬ lar portions. To these parts may be assigned the gene¬ ral appellations of the Vascular System and Cellular Tissue of plants, the description of which shall form the first subjects of consideration. Section I. Of the Vascular System. Art. I.— General Characters of the Vessels. Definition. By the vascular system may be understood, in a ge¬ neral sense, all those parts of a plant which do not exhi¬ bit the form either of membrane or of cells. It consti¬ tutes almost the entire bulk of the more solid parts of trees, and by Grew and Malpighi was denominated the ligneous body, in contradistinction to the cellular tissue which accompanies it, and which forms by far the largest portion of many herbaceous plants. To common observa¬ tion, a piece of dry wood appears to be a mass of solid fibres, that is, a series of particles arranged in a filiform figure, and destitute of any continuous canal. Thus Tournefort and others considered the ligneous parts of a plant to be a mass of minute solid filaments, placed paral¬ lel to each other, like the threads in a skein of silk, be¬ tween the interstices of which the sap ascended; but the anatomical researches of Grew conducted him to a differ¬ ent conclusion. “ If it he asked,” says he, “ what all that part of a plant, whether herb or tree, which is pro¬ perly called the woody part—what all that is ? I suppose that it is nothing else but a cluster of innumerable and most extraordinary small vessels or concave fibres.” (Anat. of Plants, p. 20.) Malpighi held similar opinions con¬ cerning the vascularity of plants, which was farther at¬ tested by the microscopical observations of Hooke and Leeuwenhoeck. Du Hamel, though he admits that, un¬ der maceration, the parts of plants seem capable of inde¬ finite subdivision, yet, from many circumstances, avows his conviction of their vascularity; and Hedwig maintains that the oldest and most compact plant is but a congeries of vessels and cells, which have nothing of the character of a fibrous solid, except in the thin membranous coats by which they are formed. (De Fibrce Veyetab. Ortu, p.17.) Few circumstances have contributed more to per-Names of plex and retard our knowledge of the structure of plants, than the vague and erroneous nomenclature that has been employed to designate their constituent organs, more par¬ ticularly in relation to the vascular system. Thus the several terms filaments, fibres, strings, threads, and nerves, which, in their ordinary acceptation, are understood to express a solid substance, have been constantly made use of in describing the tubes or vessels of plants. The same organs, however, even by the same writers, are fre¬ quently called tubular bodies, ligneous tubes, concave fibres, ducts, canals, arteries, veins, and vessels. In our future descriptions we shall employ the term vessel in a generic sense, to express all the diversity of names just enumerated ; and the different kinds or species of vessels we shall hereafter attempt to discriminate by appropriate appellations. Vessels, as -we have said, exist in almost every part of a plant. In the higher orders of animals the fluids contained in the vessels are conveyed to a central reser¬ voir called the heart, from which they are again sent out to all parts of the body. Near to this reservoir the ves- ANATOMY, VEGETABLE, 43 n;; ikientarysels are few in number and large in size ; and they gra- ([gans. dually lessen in size and increase in number as they re- r cede from it. In plants there is no such reservoir, but the fluids which enter by innumerable mouths at the root (are at once distributed equally through all parts of the vegetable that are fitted to receive them. Hence in plants there is little variation in the diameter of the ves¬ sels ; and their general figure is therefore cylindrical, ir size. From the extreme minuteness of the vessels, it is scarcely possible to compute their number with accuracy. By driving off their fluids without destroying their figure, as is done in the preparation of charcoal, Hooke number¬ ed in a line, ygth part of an inch long, not fewer than 150 vessels; therefore, in a line an inch long, there must be 2700, and in a surface of a square inch, 7,290,000 vessels ; “ which would seem incredible,” says he, “ were not every one left to believe his own eyes.” These facts he ve¬ rified by other observations on decayed wood, in which the vessels were empty; and also on petrifactions of ligne¬ ous bodies, in which the places of the vessels were very conspicuous. In very close and dense wood, as that of guaiacum, the vessels were still more minute than in the examples just quoted. (Micrographia, p. 101, 108.) In a piece of oak of the size of about Ygth part of a square inch, Leeuwenhoeck reckoned 20,000 vessels; so that in an oak-tree of no more than one foot in circum¬ ference, or about four inches in diameter, there will be found, according to his computation, 200,000,000 of such I vessels. (Select Works, translated by Hoole, vol. i. p. 3.) The largest vessel observed by Hedwig (De Vegetab. Ortu, p. 26) in the stem of the gourd appeared through his microscope about j^th of an inch in diameter; and as his instrument magnified 290 times, the true diameter must be reckoned the 3480th part of an inch, which would give for the square inch 12,110,400 vessels. In certain plants, however, the vessels are large enough to be discerned by the naked eye, and in some cases they acquire a large size. ■“I ji d(n fas- The vessels of plants do not, like those of animais, cicri. exist single, but are collected into fasciculi or bundles, which, however, have often the appearance of single ves¬ sels. In the stems of herbs, and in roots, Grew discover¬ ed each small fasciculus to be composed of from 30 to 100, or sometimes many hundred vessels. (Anatomy of Plants, p. 20.) The direction of these fasciculi in the trunk is generally perpendicular, but in other parts their course is often oblique, and in their smaller ramifications they pro¬ duce all sorts of figures. In herbs the fasciculi are more or less numerous, and placed often at considerable dis¬ tances from each other, exhibiting the appearance of small columns dispersed through the cellular tissue: in other instances they are much more numerous, but destitute of any symmetrical arrangement; while, in trees, they are disposed regularly around the axis, presenting in their transverse section the well-known appearance of concen¬ tric circles in the wood. fy, llaiify. In some parts, where the fasciculi stand at a distance from each other, some vessels often quit one parcel to unite with another, and return afterwards to that which they had previously left. In this manner they are said to ramify, and frequently, by their ramification, a reticu¬ lated appearance is produced, as occurs especially in the bark and leaves. In the wood of the trunk, where they stand collaterally in a perpendicular direction, they very seldom, if ever, run into one another, but keep, says Grew, like so many several vessels, all along distinct; as, by cutting, and so following any one fasciculus, may be ob¬ served. (Anatomy of Plants, p. 20.) In branches and roots, though the direction of the fasciculi be changed, they seem only to break into smaller parcels, and run Elementary side by side, never inosculating with each other, nor being Organs, ramified, so as to be successively propagated one fromv^v^w/ another, as in the vessels of animals. Neither, adds he, are they wound together like threads in a rope, but are only contiguous or simply tangent, like the several cords in the braces of a drum. (Anatomy of Plants, p. 66.) Even in the leaf, where the vessels seem to ramify out of greater into less, as in the arteries of animals, yet, if the skin and pulp of the leaf be removed, and the vessels laid bare, it will appear that they are all of the same size everywhere in the leaf, and all continued through it, as distinct tubes, like the several threads in a skein of silk. The distribution of the vessels is not effected, therefore, by their ramifying out of greater into less, but by the di¬ vision of a greater fasciculus into several smaller fasciculi, till at last they come to be single. (Anatomy of Plants, p. 155.) It may be doubted whether, when the vessels of dif- Do not ferent fasciculi come into contact, they ever actuallyanasto- unite and are lost in each other, forming that kind of con-111080, nection which is called inosculation or anastomosis. Grew strenuously contends against any such connection of the vessels in any part of the plant. “ On a superficial view, indeed, the vessels of the leaf,” says he, “ seem to be inos¬ culated, not only side to side, but the ends of some into the sides of others: but neither is this ever really done, the lesser fasciculi being only so far diducted as to stand at right angles with the greater; but they are never in¬ osculated, except end to end, or mouth to mouth, after they are come at last to their final distribution.” (Ana¬ tomy of Plants, p. 155.) Malpighi, however, from the fact of the alternate separation and conjunction of the vessels, and from analogy with what occurs in the animal system, speaks always of the anastomosis of the vessels, but he nowhere gives us any thing like proof of the fact; and Du Hamel, from actual dissection of several fasciculi, regards them in their union as resembling more the nerves than the blood-vessels of animals. When, indeed, we consider the extraordinary minuteness of the vessels, and the cir¬ cumstance of their possessing nearly the same size in every part, there is no room for that continual ramification out of one into another, and consequent diminution of diameter, that occur in the vessels of animals; and the im¬ mensity of their number, together with the endless sepa¬ rations and re-unions which their fasciculi make, seems cal¬ culated to fulfil the purposes of less general distribution in the plant, which successive division and perpetual anas¬ tomosis effect in the animal system. Another general circumstance in the vessels of plants Have no is, that we do not discover in them any structure which valves, has the true nature and use of valves, similar to what is met with in the veins and absorbent vessels of animals. Dr Hooke could never observe in their canal any thing that had the appearance of valves. (Micrographia, p. 116.) Did such a structure exist, the absorption of nutrient matter from the lobes of the seed, and its con¬ veyance, in a backward course, to the embryo, could not, says Grew, have place; neither could the root, as it often does, grow upward and downward both at once. (Anatomy of Plants.) If the piece of a root of elm-tree be cut in autumn, the juices, says Du Hamel, are found to escape indifferently at either end, as the one or the other is al¬ ternately held downward; a circumstance, he observes, inconsistent with the opinion of Mariotte, who maintained the existence of valves. (Phys. des Arbres, tom. i. p. 56.) It is well known, also, that many plants may be made to grow in an inverted position, so as to put forth leaves and flowers from their roots; and large trees have been 44 ANATOMY, VEGETABLE. Elementary nourished by juices received through the extremities of Organs, their ingrafted branches, after all connection between the earth and the roots had been cut off. In general, in¬ deed, the extreme minuteness of the vessels seems almost to preclude the possible existence of valves in their ca¬ nal ; but in some instances, where the vessels in aged trees have become enlarged, membranous productions have been found to occupy their cavities, which some have alleged to perform the office of valves. They occur, how¬ ever, only at an advanced period of growth, and form no necessary part of the structure of the vessel, and, instead of promoting, contribute only to obstruct the course of the fluids. Thus far with regard to the general nature of the ves¬ sels of plants: let us next .discriminate their several spe¬ cies or kinds. Art. II.—Of the Common Sap-Vessels. Sap.ves- To ascertain the kinds and situation of the vessels sels. 0f plants, various means have been employed. The plant has been dissected both in its dry and recent state; the natural qualities and movements of its fluids have been observed; and its vessels have been filled with coloured liquors, by causing it to vegetate in them. By the com¬ bined use of these several means, many important parti¬ culars have been ascertained; but it must be acknow¬ ledged that the question is still beset with doubts and dif¬ ficulties, and that, with relation to it, great diversity of opinion continues to prevail. A concise statement of the facts ascertained with respect to the movements of the vegetable fluids may perhaps contribute to define the situation and kinds of the vessels that convey them. Course of It has been proved that, early in spring, before the the sap. leaves appear, a watery fluid rises abundantly in the woody part of the trunk of trees, and continues visibly to ascend to the very extremity of the branches, until the leaves are developed, when, to appearance, it ceases to flow, and can no longer be collected by perforating or tap¬ ping the tree. This fluid has been shown to ascend through the wood, and to rise, in general, most abundant¬ ly through its youngest or outmost circle; but in trees whose vessels have not been obstructed from age or other causes, it rises through every circle to the very pith, and, as far as can be judged, in all the vessels that compose those circles. At this early period of vegetation no fluid is found in the bark, nor between it and the wood, nor in the pith; but the vessels of the bark are perfectly dry. These facts are deducible from observations on the natu¬ ral flow of the fluids by Grew, Du Hamel, Walker, and others; and are supported by various experiments of M. de la Baisse, Bonnet, Reichel, and others, made by caus¬ ing plants and parts of plants to grow in coloured liquors, in which the vessels of the wood alone became filled; but no tinge of colour was communicated to those of the bark. To these vessels the several names of lymph-ducts, sap- vessels, ligneous tubes, ascending and adducent vessels, have been applied: we shall in future denominate them sap-vessels. Kinds of The vessels which thus form the mass of wood have sap-ves- by some writers been distributed into different kinds, sds. an{] supposed to exercise very different functions. At certain periods of vegetation they appear empty; and hence Malpighi supposed two species of vessels to exist in the wood, one destined to carry sap, and the other to convey air: to these latter, from their supposed office, he gave the name of tracheae, and from their structure called them spiral vessels. {Anatom. Plantar. Idea, p. 3.) Grew also believed these empty tubes to be air-vessels, but ad¬ mitted that at certain seasons they carried sap. At an Element^U early period, however, Ray maintained that the vessels Organ» thus supposed to convey air were truly sap-vessels; andv^v> Du Hamel, in common with Grew, admitted that they carried sap in spring. Hill considered them altogether as sap-vessels; and Reichel, Hedwig, and others, by filling them with coloured fluids, proved that such was their true office. On the other hand, no experiments, says Ludwig, have yet shown that there exist in vegetables vessels destined to convey only air; and in this opinion subsequent writers, with few exceptions, have acquiesced. It will, however, be convenient to treat of their general nature and form under the distinct appellations of sap- vessels and spiral vessels, by which they are commonly known. By Grew and Malpighi the common sap-vessels were Opinion regarded as entire tubes, having no apertures but in of Grew the direction of their length. Grew represents a single vessel as having the appearance exhibited in fig. 1, Plate XXXVIII., the aperture or canal of which is not visible un¬ less highly magnified, as in fig. 2. According to Malpighi of Mai. these vessels send off numerous capillary filaments to the pighi, cellular tissue, so that the cells are surrounded by a plexus of vessels, as is particularly seen in the pith of elder and some others; and these ramifications, he adds, spring probably from the perpendicular vessels both in the bark and wood. {Anatom. Plantar, p. 29. Lugd. Bat. 1688.) These lateral ramifications were observed also by Leeuwenhoeck in a piece of fir-wood newly felled. Of0fLeeu. this wood he procured a longitudinal section, so extreme- wenhoscl ly thin that he could see distinctly the particles of fluid moving in the vessels, as represented in the upper portion of fig. 3, Plate XXXVIII.; while lower down, on many parts of these vessels, small points or dots were visible, which he at length discovered to be round apertures; and as he did not see these apertures in any other parts than those in which he had separated the horizontal cellular tissue from the perpendicular vessels, he concluded that, at these points, these two organs were connected. He farther separated two of the vessels from the remainder, and through the microscope they appeared as in fig. 4; but the “ engraver,” he adds, “ said that he could not possibly draw all the jagged parts that he saw; and we both of us perceived, in the broken membrane or coat of the tube, many excessively minute vessels, which, by rea¬ son of their smallness, he was unable to express in the drawing.” {Select Works of Leeuwenhoeck, by Hoole, vol. i. p. 12.) # - It may however be said, that this communication be¬ tween the vessels and cells is maintained not by rami¬ fications from the vessels, but through apertures or pores in their sides; and, accordingly, many appearances have been remarked as existing on the sides of the vessels, of which different authors have given very different repre¬ sentations, and which some have regarded as pores. Thus, on the vessels of the fir Malpighi observed certain dotted appearances, which he describes as roundish tubercles, and which were so numerous that they appeared to cover the vessels. On the vessels of the elm, the beech, and the willow, Leeuwenhoeck saw similar particles, which re¬ sembled small globules. {SelectWorks,\o\. 'n.Tp. \f Dr Hill of Hill, describes the vessels of the alburnum, or newly formed wood of the willow, as connected with each other by a flocculent interstitial matter. When, by long maceration, this matter is detached, the vessels then exhibit a dotted appearance; and, if examined by a highly magnifying power, these dots, according to him, are so many oval swellings, and each has, as it were, a mouth. Through these mouths, which he represents as innumerable, and ANATOMY, VEGETABLE. 45 Cletentary existing on all parts of the vessels, he conceives the fluids < tons, to he discharged into the cellular tissue. (On the Con- struction of Timber, p. 18.) In fig. 5, Plate XXXVIII., is a representation of these vessels connected by flocculent mat¬ ter, with their extremities collapsed from the escape of their juices, and their sides sprinkled with the little mouths which he mentions. These mouths, If they exist at all, are probably not pores in the sides of the vessels, but the little apertures seen by Leeuwenhoeck, and produced by the separation of the cellular tissue while the parts are still young and tender. The same author, speaking of the vessels of the mature wood of the pear, states them to be close canals, as in fig. 6, with no lateral apertures in them. >p i m A still later writer, M. Mirbel, declares, that not I ^I'bel, fewcr than fjve species of vessels are to be found in the woody part of plants. These he denominates porous tubes, cleft tubes, tracheae, mixed tubes, and vessels en chapelet, from their supposed resemblance to a string of beads. Of these several species we have given representations in figures 7, 8, 9, 10, 11, and 12, Plate XXXVIII. The first species, or porous tubes, according to this writer, exist in every part of the plant where the sap moves with free¬ dom. Their sides are covered with small eminences or projections, in the centre of which is to be found a small pore. (Exposition de la Theorie de V Organisation Vegetale, p. 107.) Improving a little on Hill, he represents these pores not as promiscuously placed, but as ranged in trans¬ verse lines (fig. 7); and through them he conceives the fluids of the plant to percolate, not, however, into the cells only, but out of one layer of tubes into another, in a lateral direction. In this manner he conducts the fluids from the centre to the circumference of the wood, and at length, by a route not so easily followed, contrives to get them into the vessels of the bark, the sides of which he declares to be perfectly entire, and alike destitute of pores and clefts. (Ibid. p. 297.) Several writers have sought to discover these al- nd nhers. leged pores in the sides of the vessels. With Malpighi, some regard the appearances observed not as pores, but as elevations on the surface of the vessels; others, as par¬ ticles contained within them. Under a very highly mag¬ nifying power, Kieser thinks he has, in some vessels, de¬ tected the existence of pores; while M. Dutrochet pro¬ nounces the same objects to be corpuscles, filled with nervous matter; and De Candolle suggests the probabi¬ lity of their being small glands, destined in some way to contribute to the nutrition of the plant. These different opinions, formed on viewing the same objects, sufficiently manifest the difficulty and uncertainty of microscopical observations made with highly magnifying powers, as was long since pointed out by Hooke, who first applied the microscope to the examination of the structure of plants. “ Of such minute objects,” says he, “ there is much more difficulty to discover the true shape by an instrument than by the naked eye; the same object quite differing in one position to the light, from what it really is and may be shown to be in another. In some objects,” he adds, “ it is exceedingly difficult to distinguish between a pro¬ minency and a depression, between a shadow and a black stain, or a reflection and a whiteness in the colour; and the transparency of most objects renders them yet more difficult than if they were opaque.” (Micrographia, Preface.) >e.sc ion. Art. III.—Of the Spiral Vessels. Various as have been the opinions of writers respect¬ ing the common sap-vessels of plants, they have dif¬ fered yet more in their views concerning the position, number, size, structure, and uses of those which have Elementary been denominated spiral The common fathers of Vege- Organs, table Anatomy, Grew and Malpighi, who at the same'^,^v^*^ time, but in different countries, prosecuted their inquiries without any knowledge of or communication with each other, are nearly of one opinion on all the more impor¬ tant points in relation to these vessels. Later writers have differed alike from them, and from each other, on almost every point. As the subject is of fundamental im¬ portance in the economy of vegetables, we shall endeavour to set before the reader the leading facts and opinions concerning it; to canvass their relative merits; and de¬ duce from the whole such conclusions as seem most nearly to approach the truth. Malpighi describes spiral vessels as existing in the Opinion of ligneous parts of all plants. He called them spiral Malpighi, because, when extended, they were resolved, not into se¬ parate rings, but into a single zone, which might be drawn out to a great length. In general they form continuous tubes, but are sometimes contracted at regular distances, so as to resemble somewhat a series of oblong cells. One of these contracted spiral vessels, as delineated by Malpighi, is represented in fig. 17, Plate XXXVIII., at the extremity of which the spiral filament is in part drawn out; and si¬ milar appearances of the spiral structure are exhibited in figures 9 and 10, by Mirbel. In herbs, according to Mal¬ pighi, these spiral vessels constantly accompany the com¬ mon sap-vessels, and are ensheathed by them; in shrubs, they occur in every part of the wood, single or in clusters; and in trees, an intermixture of spiral vessels with the common sap-vessels is observed in every part of the wood. In the fir, they are found immediately beneath the bark, and are so numerous as to constitute the chief bulk of the wood. They exist with the sap-vessels in the petioles and ribs of the leaf, and likewise in the petals of the flower. In the roots they are also met with, and in some roots are so abundant, as to exceed in bulk all the other parts. They exist, he adds, in every part of the plant ex¬ cept the bark; and are annually formed with the alburn- ous vessels of trees. (Anatom. Plantar, passim.) From the writings of Grew we collect also that of Grew, spiral vessels exist in every part of a plant except the bark. In the root they are numerous, of very various size, and their bore is generally larger than that of the common sap-vessels. In the trunks both of herbs and trees they are equally visible, and in position, size, and number, subject to great variation. Sometimes they are collected into fasciculi, at other times they are disposed in rays, and in other instances they are arranged in a cir¬ cular form: they stand sometimes next to the pith, in other instances next to the bark, and in other cases again they alternate with the common sap-vessels in every part of the wood. They have a more ample bore than the common vessels, and vary in size to at least twenty differ¬ ent degrees. In the leaf they always accompany the sap-vessels, and both in its petiole and ribs are constant¬ ly surrounded or ensheathed by them. They have a similar position in the petals of the flower, and in the vas¬ cular parts of the fruit. (Anatomy of Plants, passim.) Hence, then, it appears, from the dissections of Malpighi and Grew, that, in every plant in which vessels are to be seen, and almost in every part of it, spiral vessels abound; they exist, however, only in the ligneous portion, or that part in which the sap ascends, and are never to be found in the bark. By Du Hamel, the spiral vessels, supposed to con-of Du Ha- vey air, are described as existing in the leaves and the “el- flowers, the petals of which are almost wholly composed of them. In the herbaceous portion of young branches 46 ANATOMY, VEGETABLE. Elementary they are also well seen; which portion afterwards be- Organs. comes ligneous; so that it cannot be doubted that they exist in the wood, though he could never discover them but in young branches. (Phys. des Arbres, tom. i. p. 42.) If, however, all the empty vessels seen in a transverse section of the wood be deemed air-vessels, and all air- vessels have a spiral structure, then, says this author, they would, in many plants, form a great part of the lig¬ neous body. From these large vessels, however, he has seen, in autumn, fluids to escape; so that they are not properly air-vessels, or, as Grew observed, they sometimes carry sap. They are not to be found in the bark. Opinion Under the common name of sap-vessels, Hill deli- of Hill, neates all the varieties of vessels that constitute the wood; they are largest in the outer circles, smaller in the others ; they contain, says he, in spring and at midsummer, a limpid liquor; but at all other seasons they appear emp¬ ty, and have therefore been erroneously deemed air- vessels. He says nothing of their spiral construction; but describes the vessels which form the chief mass of the wood as possessing solid and firm coats, forming an arrangement of plain and simple tubes, as in fig. 6, Plate XXXVIII., resembling those of the alburnum, except that they have no mouths in their sides. (On the Con¬ struction of Timber, p. 8 and 19.) ofReichel, M. Reichel maintains the existence of spiral ves¬ sels in almost all parts of the plant. By causing plants to vegetate in coloured fluids, he traced them from the roots, through the trunk and branches, to the extremity of the leaves, and into all parts of the flower,—as the calyx, the petals, the style, the stamens, and the anthers. In fruits and in seeds, and in the radicle and plume of the latter, they were equally apparent. The coloured liquors, as they rose, communicated a tinge to the cellu¬ lar tissue of the wood, as was previously observed by De la Baisse ; but no trace of colour was ever observed either in the bark or pith, which therefore contain no spiral ves¬ sels. He considered the spiral vessels as the organs everywhere conveying nutrient matter to the plant, and as having no title to the appellation of air-vessels. (En- cyclop. Methodique, article Physiol. Veget. p. 288.) Si¬ milar experiments of Hedwig and others confirm these facts as to the general distribution of the spiral vessels, and their bearing to every part the fluids absorbed by the roots. of Mirbel, We have already enumerated the five different spe¬ cies of vessels which Mirbel regards as constituting the woody part of plants. According to him, true spiral vessels (which he continues to denominate tracheae) are not to be found in the root, but only in the trunk and the parts which are produced from it. Even in the trunk they are to be found only around the pith, and never in the exterior ligneous layers. He admits that they exist in all soft and succulent parts, and that coloured fluids rise in them as well as in the other varieties of vessels; but they never are to be found either in the bark or pith. {Exposition de la Theorie de l Organisation Vegetale, p. 74, 78.) of Kieser. According to M. Kieser, spiral vessels are found in all the more perfect plants, from the earliest period of their existence; and in all parts of them, except the bark and pith. They vary much in size, and in aged plants are frequently obstructed by a species of vesicle, which ori¬ ginates from the interior sides of the vessels. According to him, nothing but air is found in their cavity, though, in the wood of guaiacum, he has seen all the spiral vessels, in common with the cellular tissue, completely filled with resinous matter. He does not know the anatomical rela¬ tion subsisting between the spiral vessels and the other organs; but thinks it proved that no direct commumca-Element,•'I tion exists between them and the cells. {Mem. sur F Or- Organ! ganisation des Plantes, 1814, p. 116, 117.) Lastly, M. de Candolle admits the existence of spiral vessels in the wood of the trunk, but not of the roots; and, with Kieser, re¬ gards them as conveying only air. {Organographie Vegetale, tome i. p. 39.) From the combined observations of all the preced¬ ing writers, with the exception of MM. Mirbel and de Candolle, we learn that spiral vessels exist not only in every part of the trunk, but in the root, and in every other part of the plant except the bark and the pith. From the actual presence of sap in these vessels early in spring, and again in autumn, and from the entrance of coloured fluids at all seasons, we also learn, that their true office is not to convey air, as Malpighi and others formerly, and Kieser and others more recently, maintain; but to carry sap. Are we therefore to regard them as a species distinct from the common sap-vessels, or are both to be held merely as varieties of one common kind ? This question may receive some illustration from a little farther inquiry into the formation and structure of the sap and spiral vessels. Art. IV.— Of the Structure of the Sap and Spiral Vessels. Malpighi seems to have regarded the common sap-Structu: vessels as being formed of a series of small vesicles or of sap.\ cells, mutually opening into each other. {Anat. Plantar.^5, p. 28.) In this instance he appears either to have mis¬ taken a series of elongated cells for vessels, or to have conceived that the contractions, occasionally formed in the vessels themselves, afforded evidence of their having been constructed originally by cells. Grew, on the other hand, considered these vessels to be composed of straight fibres, placed parallel to each other, so as to form a cylin¬ drical tube. {Anat. of Plants, p. 112.) Leeuwenhoeck held them to be composed of two fine transparent coats, formed of other vessels excessively minute. ( Select Worhs, vol. i. p. 11.) And Du Hamel, by submitting thin layers of wood to long maceration, obtained bundles of longitu¬ dinal fibres of extreme minuteness, by which he consi¬ dered the lymphatic vessels of the wood to be formed. {Phys. des Arbres, tome i. p. 32.) Hill describes the al- burnous vessels as being formed of the same material as the cells, and to collapse when emptied of their fluids. In one instance, where a strong light was made to penetrate the vessel, it appeared as if composed of numerous cells; but on farther examination, these seeming divisions al¬ tered their places, and were found to proceed from por¬ tions of watery sap still retained in the vessel. This ap¬ pearance, as he properly observes, may be a very neces¬ sary lesson against hasty judgments. {Construction of Timber, p. 33.) According to M. Mirbel, “ the entire mass of the plant is nothing but cellular tissue, the cells of which differ only in form and dimensions.” The cells and ves¬ sels are thus considered to be formed out of one and the same membranous tissue. In forming cells, this mem¬ brane is supposed to dilate in every direction: in pro¬ ducing vessels, it increases only in length. {Exposition de la Theorie de F Organisation Vegetale.) The manner in which he supposes the vessels to be formed out of the pri¬ mitive membrane is very fanciful; and he is probably right in thinking that “ no one before himself had formed a similar conception of vegetable organization.” Concerning the structure of the spiral vessels, opin-Structu ions have varied still more than in relation to those justofspira considered. Malpighi describes them as composed of'resse,s’ met | sap, .s,h| ANATOMY, VEGETABLE. 47 fffiiilaentarya thin pellucid plate, of a silvery colour, somewhat broad, JjLans. and which, being placed spirally, and united at its edges, v»LX~y-constructs a tube. When this tube is drawn out, it does not )p !oa °f separate into distinct rings, but is resolved into a continu- ous gpjrai zone. At particular places these vessels are sometimes contracted, so as to exhibit the appearance of oblong cells opening into each other, as in fig. 17, Plate XXXVIII. In size they greatly surpass the ordinary sap- vessels ; and their canal is then frequently occupied by membranous vesicles, which nearly fill their cavity. (Anat. Plantar, p. 3-26.) These vesicles were also observed by Leeuwenhoeck, and their appearance, in the transverse section of an enlarged vessel, is represented in fig. 25. f rew, According to Grew, the thin plate that forms the spiral vessel is not flat, and, instead of being single, consists of two or more round filaments or threads, placed collate¬ rally, but perfectly distinct. These component filaments he regards as united by other smaller transverse filaments, and the thin plates which they form by their connection constitute the spiral vessel,—“ as if we should imagine,” he adds, “ a piece of fine narrow riband to be wound spirally, and edge to edge, round a stick, and the stick being drawn out, the riband to be left in the figure of a tube, answerable to a spiral vessel.” As, however, the riband is composed of numerous threads, placed parallel to each other, so is the plate that forms the spiral vessel; and it is according to the greater or less delicacy of the vessel examined, and the manner of its dissection, that it appears to be constituted either of a flat plate or a round filament. The spiration of the filaments he considered to be made in the root from right to left, and in the trunk from left to right. (Anat. of Plants, p. 73 and 117.) u Ha- The opinion of Du Hamel with respect to the con¬ struction of these vessels was very similar to that of Grew, and he employed the same analogy of a riband edwig, twisted round a stick to illustrate it. Hedwig gave a very different account of them. Reconsidered the spiral vessel to be composed of two distinct parts ; one a mem¬ branous canal conveying air, and the other a spiral tube rolled round it, by which the fluids were conveyed. The spires of the tube, in some instances, are represented as close; in others they are separated, and the intervening portions of the membranous canal exhibit a dotted appear¬ ance. He considered all the sap-vessels, from their first formation, to possess this compound structure, and, by a series of changes, which he professes to describe, to be ultimately transformed into the solid fibre of the wood. (De Fibrce Vegetal). Ortu, p. 25.) Others, also, have believed the spiral vessel to be formed of a membranous tube, but have denied that it conveyed air; and the spiral tube of Hedwig they have regarded as a solid filament. From the different appearances which they exhibit, MM. Treviranus and Bernhardi distinguish several varieties of spiral vessels; as does also M. Kieser, whose account of these vessels is not only the most elaborate, but that probably which approaches nearest to truth. We shall therefore subjoin a brief abstract of his observations on the structure of these vessels, and more particularly on the series of transformations which they seem to undergo. The construction of these vessels M. Kieser professes to have studied with the greatest care, and to have esta¬ blished incontestably the following points. Sometimes, says he, only one fibre, sometimes many, go to form the spire ot a vessel. These fibres are commonly round, some¬ times a little flattened, and are twisted spirally about an empty space, so as to form a tube. The spiral fibres in young plants, and sometimes in mature ones, are in close contact; in other instances they are separated, and the interstices are then occupied by a dotted or punctuated membrane, as is very frequent in trees; or sometimes Elementary they are connected by ramifications which proceed from Organs, the spiral fibres themselves. From the minuteness of the' spiral fibre, it is difficult to decide whether it is a solid or tubular body, and often, from the same cause, to pronounce whether it is round or flattened.. It is transparent, has considerable consistence and tenacity, and in some plants appears to possess elasticity. The number of fibres that form the spire of a vessel is very various ; sometimes, as before remarked, there is only one, but more often several, which are twisted in the same plane and the same direction. He has seen nine fibres thus united; and when unrolled, the spires of the vessels then seem to form a kind of riband. When many fibres are employed to form a spire, they always run parallel, and do not cross ; so that the side of the vessel has never more than the thickness of a single fibre. At their first formation the fibres are not united; it is only in more advanced age that they become united, either by ramifica¬ tions from their edges, or by a peculiar membrane. The direction of the fibre in twisting is sometimes from right to left, and sometimes the contrary. The size also is very various. In young plants it is very small, so that a milli¬ metre comprises the diameter of 2600 of these fibres ; and in some instances they have little more than half that size. In older vessels they have a larger size. Between the knots of the trunk they are simple, and do not ramify; but in the knots they undergo great changes of form, and are variously ramified and combined. Art. V.— Of the Transformations of the Spiral Vessels. It was the opinion of Grew, that the “ common sap- Opinion of vessels begin to be formed in spring, but the spiral ves- Grew, sels not till the end of summer, at least not till about that time do they appear.” (Anat. of Plants, p. 131.) Malpighi also describes these vessels as gradually appearing in the alburnum, and augmenting in size in every successive ring of wood ; an observation made also by Grew, who describes the “ spiral vessels as being amplified in each annual ring, so as to form a vessel of a wider bore.” These facts in¬ dicate that the spiral vessels, after their formation, under¬ go considerable changes in the progress of vegetation, or, to use the language of Grew, that “ they are postnate, and seem produced by some alteration in the quality, position, and texture of their fibres.” In treating of the transformations or metamorphoses of of Kieser. the spiral vessels, M. Kieser illustrates his ideas by a series of dissections of the stem and root of the gourd (cucurbita pepo), made apparently with great care. These designs occupy several plates of his work ; and being taken from different parts of the same plant, at different stages of its growth, and viewed with the same magnify¬ ing power, they show the progressive changes in form and appearance of these vessels as the plant increases in age. M. Kieser exhibits corresponding vertical and ho¬ rizontal sections of the same parts; but our object will be answered by copying only certain portions of the vertical sections. The ligneous portion of the stem of the gourd possesses ten bundles of vessels, symmetrically disposed around the pith, and standing at unequal distances between it and the bark. In fig. 13, Plate XXXVIIL, we have copied the representation of a vertical section of the vessels in one of these bundles, taken from a portion of the plant near the summit of its stem, and made with the aid of a micro¬ scope magnifying 130 times. The number of vessels in each bundle at this period is said to vary from three to five. They are constructed of one or more fibres, placed ANATOMY, VEGETABLE. 48 Elementary contiguous to each other, and twisted into a spiral form, Organs, producing a round cavity within. These are called simple spirals. They are found in every young plant, and in the spiral6 newly formed parts of old ones. Their size is smaller P ' than that of the other varieties to which they give origin. The number of fibres in each spire varies from one to nine ; and they are the only variety met with in some of the inferior vegetables. The next figure (fig. 14, Plate XXXVIII.) exhibits a simi¬ lar section of the vessels of the same plant at the internodial space below. Of the four vessels contained in this bundle, the outer one, «, is larger than the rest, but its spires are contiguous, as in those of the former figure: the inner one, is formed of a series of rings, disposed in a per¬ pendicular line. These rings are very like the spires of the preceding variety, and often combined with them in the same vessel. They are sometimes separated from each other only by a space equal to their own diameter; in other instances to eight or ten times that distance, forming then the substratum of the next variety. In their Annular present form they constitute the annular spiral; and in spiral. herbaceous plants occupy frequently the same position as the simple spiral, that is, next to the pith. From the two simple varieties of form just described, other more complex forms are produced, as the plant ad¬ vances in age. In the next section, made still lower down on the same plant, the vessels exhibit the forms represented in fig. 15 of the same plate, which brings the third varie¬ ty of spiral vessel into view. The exterior vessel, b, of this fasciculus, or that nearest to the bark, is of much larger size: its spires, represented by the white lines in the figure, are separated to nearly equal distances from each other, and the intervening spaces are filled up with a membrane, sprinkled over with small obscure points or Punctuat- dots, constituting the punctuated spiral vessel. In this ed spiral. variety the rings are never contiguous. Such vessels oc¬ cur only in the more advanced age of herbaceous plants, but are said to be original formations in the alburnum of trees: they acquire a much greater size than the two for¬ mer varieties, especially in the stem ; but do not become so large in the root. In herbs they occupy a position ex¬ terior to that of the two preceding varieties ; but in trees the largest vessels of the annual layers are nearer to the pith. The three other vessels in this figure are simple spirals, in part unrolled. The spiral fibre in these vessels is of various size, be¬ ing largest in the greatest vessels: it is sometimes so small as to be scarcely visible. It is often difficult to de¬ termine whether its form is spiral or annular: for the most part it is spiral in herbs, and annular in trees. The membrane which connects the spires or rings in this va¬ riety is not visible in young vessels, but appears in more mature age. It is at first transparent, but becomes opaque from age. In this punctuated variety of vessel the spires cannot be unrolled without tearing the connecting mem¬ brane. As to the points or dots observed on this mem¬ brane, they have been taken by some for pores on its surface; by others for clefts produced by the joining of the spires. They are clearly not clefts, says M. Kieser, and it is against the supposition of their being pores that their position has a relation always to the spires, and not to the adjoining cells. In the vessels of some plants, as in those of the French bean, when magnified 400 times, these points, fig. 21, a, seem as if pierced by a hole ex¬ ceedingly small; but in other instances they are quite dark, and similar to those found on the membranous productions that fill the cavity of aged vessels, where there is not the smallest reason for considering them as pores. There seems a great analogy between these con¬ necting membranes and those which thus fill the cavity of Element , aged vessels. In some instances the size of the dots is OfgMi [ large, and they appear transparent at the centre; in . others the dots are extremely small. Their form is ob¬ long, and they are ranged generally in determined lines, parallel to the direction of the spires. In the next vertical section of a fasciculus of vessels, taken from about the middle of the stem of the same plant, all the peculiarities of size and form are still more dis¬ tinctly seen. Of the twenty-three vessels which appeared in the horizontal section of a fasciculus at this period of growth, six only were apparent in the vertical section, and are exhibited together in fig. 16, Plate XXXVIII. Of these, the first two, or outer ones, are by much the largest, and belong to the punctuated variety. A portion of the an¬ terior wall of these two vessels has been removed by the sections at /, l, so that the inner surface of the posterior wall is brought into view; and in the former vessel, d, a portion of the posterior wall itself has also been carried away. In both vessels the spiral fibre, indicated by the white line, is prominent in the section ; and in the second, e, it is unrolled at m, n,—where, at m, it is seen single, transparent, and round, while at n it is united with the punctuated membrane that connects the spires, and has a flat or riband-like form. The third vessel, f, of this fasciculus is also of the punctuated class, and the three last, g, h, i, are simple spirals ; the last of all, i, being in the progress of formation, and its spires not yet brought into a state of contiguity. There is a fourth variety of vessel that is said to have the same origin as the last, being formed in part by spires or rings, but the separations between which are filled up, not by membrane, but by small productions or ramifica¬ tions from the spires or rings themselves; and these ra¬ mifications are often so implicated as to form a net-work; whence the names of ramified and reticulated spiral vessels. Ramifielr Like the former variety, they do not exist in the young spiral' \ plant, but are formed gradually in a more mature age, and by the same series of actions ; that is, the spires or rings, which were at first contiguous, separate at a later period, and the intervals are filled up by ramifications from the spires themselves. At first these ramifications are few, as in the vessel, a, fig. 21, when they are term¬ ed ramified spirals; and as these ramifications increase, the vessel becomes reticulated, as exhibited at the letter Reticuk ^ c, fig. 20. They are found but in few plants, and ap- e4 pear to differ from the punctuated variety chiefly in the position of the rings, which are more or less obliquely placed, and at diflerent points send off one or more branches. In his ninth plate, M. Kieser has given similar repre¬ sentations of the vessels of the same plant, as they appear from sections made near the root. The fasciculus at this part of the stem contained twenty-nine vessels, as seen in the horizontal section, of which only eight were visible in the vertical section. These retained nearly the characters of those given in the preceding section, fig. 16; but ex¬ ceeded them in number and size. The large ones, situ¬ ated next the bark, were of the punctuated class, while the smaller ones, next the pith, were simple spirals. In fig. 18 we have copied the three smaller vessels. In one of these, o, which belongs to the punctuated variety, the interior of the vessel is nearly filled up with small portions of punctuated membrane, produced from the inner wall of the vessel itself, as is still more clearly seen in horizon¬ tal sections of similar vessels in fig. 25. The two vessels p, q, of fig. 18, are the youngest, and consist only of the simple spiral fibre ; the last, it is said, being composed of two such fibres, which, in consequence of the transpa- ANATOMY, VEGETABLE. 49 Spills of theoot. Ehsentary rency of the posterior wall of the vessel, appear to cross Cj^ans. each other. But M. Kieser has not confined his examination to the stem of this plant. In his tenth plate he has given simi¬ lar sections of a portion of the root. The piece of root he examined presented, in the horizontal section, four fas¬ ciculi, each of which contained about thirty-seven vessels, varying in size; the larger ones being placed exteriorly, and the smaller ones towards the centre of the root. The cells of this part were small, and nearly of the same size and form throughout, differing much from the varying size and form of the cells in the stem. Of the thirty-seven vessels before mentioned, only ten were retained in the • vertical section, all of which belonged to the punctuated variety. Of these, two had united so as to form but one cavity ; and two others were filled up with membrane. In fig. 19 we have copied the appearance of three of these vessels. One of these, indicated by the letter r, is from the middle of the figure, and its size is intermediate be¬ tween those on each side of it. A portion of its anterior wall has been carried away in three places, which brings the inner side of the posterior wall into view. The vessels a t of the same figure are the smallest, and situated at the centre of the root: all the three belong to the punctuat¬ ed variety, no simple or annular spirals being found at this period in the root. In the knots of plants the vessels undergo great changes of form, the larger vessels being contracted in different parts, and giving off productions or ramifications which form new vessels, and give to the whole a very irregular appearance, as may be seen in the upper part of fig. 20, taken from the appearance of the vessels in the knot of the balsam (Impatiens Balsamina L.). In this figure M. Kieser has also given an ideal representation of the various metamorphoses already described, viz. of the annular vessel into the simple spiral, and of that into the other varieties: thus the letter a represents the rings of the annular vessel, which at b begin to form the spiral: at c the simple spiral passes into the ramified variety, and this again, at d, into the reticulated vessel: at e the vessel becomes contracted at different parts, as previously observed by Malpighi, fig. 17, and by Mir- bel, fig. 11 and 12. This latter author called this the repls en vessel en chapelet, from its resemblance to a string of beads. From these vessels go off the branches /y f, to form the ramified spirals of the knot. In fig. 12 of the same plate M. Mirbel has given a similar ideal represen¬ tation of the five species delineated by him in figs. 7, 8, 9, 10, and 11; but they do not carry with them the same evidence of accurate observation as the several varieties indicated in the alleged transformations of Kieser. Such is the account given by M. Kieser, of the deve¬ lopment and appearances of the vessels in herbaceous plants, and of the successive changes in character which, in the progress of growth, they undergo. All the figures except the last are taken from dissections of the same plant, beginning with that part of the stem which was of latest growth, and proceeding in succession to the parts of earlier growth, or older formation. At first the vessels are comparatively few in number, of nearly uniform size, and belong to the varieties of annular and simple spiral: at a later period those first formed become augmented in size, and assume the characters of punctuated and ramir fied vessels ; they are now also removed farther from the centre, and their place is supplied by new vessels of simpler forms. With the age of the plant, they all con¬ tinue to increase in size and number, and to acquire the new characters which distinguish the several varieties in the manner already described. VOL. m. In the early growth of trees, the formation and develop-Elementary ment of vessels go on at the centre, adjacent to the pith, in Organs, the same manner as in herbs ; but at a later period, whenv^"v^s^y/ the additional growth is made at the circumference of the tree, the first formation of vessels must occur in the albur¬ num, and the subsequent changes of character be effected in the several layers of wood; which accords with the fact that spiral vessels are rarely to be detected in the alburnum, but are found with all their different characters, and of all sizes, in the several layers of wood, as may be seen in all the trans¬ verse sections of wood delineated by Grew, Malpighi, and others. In truth, the descriptions of Kieser with respect to the structure and transformations of the spiral vessels accord, in most respects, with the opinions of Grew, as briefly stated, page 47 of this article, and as may be far¬ ther seen by referring to pages 73 and 177 of Grew’s Anatomy of Plants. It is further to be remarked that, in these representa¬ tions of Kieser, which exhibit all the vessels visible in all parts of the stem and root of this plant, only one kind of vessel is described, which at different periods of growth assumes different forms ; that no vessel answering to the common or lymphatic vessel of authors is mentioned; and consequently, that all the vessels described, however differing in size and external character, are in truth but varieties of one common kind. Calling to mind also that, in the earliest production of leaves and of flowers, and even of stamens and pistils, vessels either possessing the spiral character, or capable of assuming it, are everywhere met with,—that they occur also in the stem adjacent to the pith in succulent plants, and in the alburnum of trees, and every successive ring of wood produced from it—in short, in every part and portion of the plant, except the bark and pith,—and believing, as we do, in those changes of form suggested by Grew, and subsequently demonstrat¬ ed by Kieser,—we are led to the general conclusion that only one kind of vessel exists in the ligneous portion ofOnly one other plants, as well as in that of the gourd; and that °fsaP- different plants, and in different circumstances and condi-vesse^' tions of vegetation, this vessel is capable of assuming the different forms, sizes, and characters, which have led many writers to describe these organs as constituting dif¬ ferent species, and serving different uses. M. de Candolle, in his late work on the Organography Opinion of of Plants, seems also to reject the separate existence of^e ^an* the ordinary sap-vessels or lymph-ducts of authors, and todolle‘ consider, with Kieser, all the vessels collectively to pos¬ sess a spiral character, which appears under five different forms in the vessels severally named tracheae or simple spirals, annular spirals, punctuated and reticidated spirals, and the vessels en chapelet. (Oryanographie Vegetale, tome i. p. 32.) Without deciding positively in favour of Kieser’s views, he admits that the different orders of ves¬ sels have among them very intimate relations; and in fa¬ vour of the opinion that they are really modifications of each other, he alleges that almost all these orders exist together in certain classes of plants, and are wanting alto¬ gether in others. Still, though he has no theoretical ob¬ jection to make to these alleged transitions of form, he has never been able to verify them by actual observation, and thinks the facts require further investigation. (fLbid. p. 48, 51.) With respect to the existence of pores in the spiral vessels, as alleged generally by Mirbel, and even occasionally by Kieser, but denied by a great number of observers, M. de Candolle has been led by his own ob¬ servation to entertain doubts, and to believe that what has been regarded as a pore is a luminous point, such as is exhibited by bubbles of air in water, when placed under the microscope. (Jibid. p. 55.) G 50 ANATOMY, VEGETABLE. Elementary The late opinions of M. Dutrochet on the structure of Organs. plants seem to us to have drawn attention more from their novelty than their value. He criticises the opinions of ofDutro Mirbel in relation to the structure of the vessels, and says diet. that those denominated by that author jootws tubes, are altogether destitute of pores, but have corpuscles adhering to, or rather implanted in their sides, on which account he calls them corpusculiferous tubes. (JElecherches Anatom, sur la Structure des Vegetaux, &c. p. 23.) In like manner the tracheae are said to have similar corpuscles attached to their sides, but which adhere only weakly, and form no necessary part of the organization. (Ibid.) With respect to the uses of these two kinds of vessels in the wood, he con¬ siders the corpusculiferous tubes to be the canals by which the sap ascends from the roots; while the spiral vessels, or tracheae, convey not air, but a diaphanous liquid, pre¬ pared in the leaves, and which descends, by the spiral vessels, to impart its vivifying influence to all parts of the plant, (ifo'e?. p.29-31.) No direct communication, it is said, exists between the cells and vessels which compose the vegetable texture; they are only contiguous. Fluids, nevertheless, are transmitted from one to the other; not however through holes or passages expressly designed for that purpose, as Mirbel contends, but through intermole- cular spaces, left between the integrant globular mole¬ cules, of which the organic solids are composed. (Ibid. p. 47, 48.) Such is an outline of the different opinions that have been held regarding the structure of the sap-vessels. Dif¬ ferences not less great still continue to exist with regard to their uses. Our reasons have been already given for coinciding in opinion with those who hold the vessels to be the proper channels in which the sap is conveyed. M. Kieser, however, contends that the spiral vessels of the wood are always empty, or contain only air, and considers them, with Malpighi, to be organs by which air is convey¬ ed. To this opinion M. de Candolle also inclines, though he admits that in some cases they may convey lymph. (Organog. Veget. tome i. p. 61.) Those physiologists who thus oppose the opinion that the vessels carry fluids, suppose the sap to be conveyed by certain minute channels, said to exist between the cells. The place of these alleged channels is marked out in a sec¬ tion of cellular tissue (fig. 29, Plate XXXVIII.) taken from M. Kieser, in which the black points at the angular junc¬ tions of the cells are said to denote them : from their po- Intercellu- sition they have obtained the name of intercellular canals. lar canals, gy many, however, the existence of such spaces at the angles of the cells is denied; and, indeed, when we con¬ template the various size and form of the cells, the dif¬ ferent degrees in which they are filled with fluid or so¬ lid matters, and consequently the greater or less degree of compression which these alleged intercellular spaces must undergo, we can hardly admit the sufficiency of such minute, tortuous, and precarious channels to convey the ascending sap with that force and velocity which the ex¬ periments of Hales and others have proved it to possess. On the other hand, no direct communication between the spiral vessels of plants and the external atmosphere, si¬ milar to that which occurs between the tracheae of insects and the air, has been shown to exist; and the total ab¬ sence of spiral vessels in the bark, where they would be near to the atmosphere, and their presence in the root, where they are almost beyond its influence, are alike re¬ pugnant to the notion that they exercise a respiratory function. Lastly, the absence of sap in these vessels at certain seasons, on which alone the notion of their being tracheae or air-vessels was founded, depends, as we have seen, on the period at which the examination is made; for at certain periods they have been found filled with Elementarj sap; and at all periods, when in active vegetation, they Organs. ^ will absorb and convey coloured fluids. It should also beV-''~v“^ J borne in mind that the respiratory function in vegetables is 1 performed, not by the trunk, but by the leaves.of the plant. d Dismissing then these notions concerning the uses of vl the spiral vessels, and adhering to the opinion, that in d all their variety they serve the office of conveying sap I through the plant, we shall conclude this branch of the subject with a brief notice of their modes of termination. If we begin at the root, we may consider these vessels Termina. as terminating at that part, by continuation of canal, intionsoftlK the capillary absorbents of the rootlet. As, further, thevesse^s' vessels of plants, like those of animals, are the only organs which convey the materials out of which not only the fluid, but the solid parts of the plant are formed, it may, in a general sense, be said that their modes of termina¬ tion are as numerous as the kinds of distinct parts or or¬ gans which the vegetable system contains. Hence, there¬ fore, as the cellular tissue of plants contains various mat¬ ters often precisely similar to those which exist in the vessels, it may be inferred that the vessels have a termi¬ nation in those organs; and this inference may perhaps carry more weight with many than the attempts before related of Malpighi and Leeuwenhoeck to demonstrate it anatomically. Another termination of these vessels must be in certain minute and ill-defined organs called glands, which separate from the mass of fluids peculiar secre¬ tions ; and a fourth mode may, in the leaves, be in other vessels which carry back the juices from those organs. The last and fifth mode of termination is into transpiring or exhalent organs, by which a certain portion of the con¬ tents of the vessels is discharged: so that in plants the sap-vessels terminate externally at one extremity in ab¬ sorbents, by which fluids are received; and at the other in exhalents, by which these fluids are in part discharged. Art. VI.— Of the Proper Vessels. By observation of the natural flow of the sap, com¬ bined with the results of experiments made with coloured liquors, we have endeavoured to determine the situation and kinds of vessels by which it is conveyed in the wood. The same method will best assist us in ascertaining the nature and place of the vessels met with in the bark. . t'l It was before stated, that, early in spring, the sap Course of t« of plants rises through the wood alone, and that no fluid the sap in whatever is then to be found in the bark. At a later^16"00^ period, however, the case is completely reversed; for the vessels of the wood no longer appear to carry sap, and those of the bark then become abundantly supplied with it. This difference is very clearly and concisely stated by Grew. “ The sap,” says he, “ in many plants, as the vine, ascends visibly through the wood for a month, in March and April, and rises through every ring of wood to the very centre, yet at the same time there ariseth no sap at all out of the bark, nor between it and the wood.’ “ But late in spring,” he continues, “ and in summer, the sap is no longer visible in the wood, but is abundant in the bark, in the inner margin adjacent to the wood.” Du Flame!, too, remarks, that when the lymph rises abundantly through the wood in spring, the bark is in ^ dry, and adheres to the wood, and no sap then issues from’ it, nor from between it and the wood; but, later in the season, he adds, the bark yields abundance of sap. These statements have been verified by the multiplied observa¬ tions of various subsequent authors. But why is the sap thus present only in the wood at its first rising in spring ? Why at a later period does ANATOMY, VEGETABLE. 51 ir!:;ie|jnrary it cease to be visible in that part? And how and why ' >r|ms. does it afterwards find a passage into the bark? Some -TT^ observations of Du Hamel, Hales, and Walker, point, we ClfL think, to the true cause. In spring, says Du Hamel, vlwncnt" when tlie saP rises vigorously, the buds have not appeared ; ,f A when they begin to open, the sap then flows less freely ; ea*; and when the leaves are fully developed, then the flow of sap entirely ceases. Dr Hales also remarks, that, to¬ wards the end of April, when the young shoots come forth, and the surface of the vine is greatly increased by the expansion of the leaves, the sap then ceases to flow in a visible manner till the return of the next spring. All bleeding trees, he adds, cease to bleed as soon as the young leaves begin to expand enough to perspire plenti¬ fully, and draw off the redundant sap. The bark of oak, too, separates easily when lubricated with sap; but be¬ fore the leaves appear and perspire, the bark will no longer run (as they term it), but adheres most firmly to the wood. ( Vegetable Statics, 3d edit. p. 126.) In like manner, in an experiment of Dr Walker, a birch-tree bled from every perforation in its trunk, and from every cut extremity of its branches, until vernation or budding began ; then the bleeding was almost immediately checked; and when the young leaves had pushed beyond the //?/- bernaculum, the bleeding entirely ceased. (Edin. Phil. Trans, vol. i. p. 31.) It is however certain, that though the sap was no longer visible in the wood after the leaves were developed, it continued nevertheless to rise through it; for in no other way could the leaves obtain the large portion of fluid which it is known that they constantly discharge by transpiration ; and coloured fluids manifest their presence in the vessels of the wood, as well after as before the de¬ velopment of the leaves. The leaves, therefore, must be regarded as the organs which, by their perspiration, draw off, as Dr Hales observes, the redundant sap; and hence, in an experiment where a notch was cut two or three feet above the lower end of a stem, though a great quan¬ tity of sap passed by the notch, yet was it perfectly dry; because, stfys he, “ the attraction of the perspiring leaves was greater than the force of trusion from the column of water.” {Vegetable Statics, p. 111.) as*t Not only, however, does the development of the leaves render the sap no longer visible in the wood, but they htyre" also appear to be the organs by or through which it 0 finds its way to the bark. In all the experiments just re¬ cited, the bark continued dry until the sap disappeared from the wood; in other words, until it was drawn off by the leaves; and then, and not till then, the bark became moist, and continued laden with sap through the rest of the summer. Now, as it has been before shown that no sap enters the bark by the roots, nor gets into it directly from the wood, there is no other known channel by which it can be conveyed, except through the leaves; and these, therefore, necessarily appear to be the organs by which it is apparently carried off from the wood, and by or through which it at the same time finds its way to the bark. This inference appears to follow, not only from the fact of the bark continuing dry until the leaves are developed, but from the circumstance that it is again rendered dry af¬ ter having become moist, if these same leaves be removed. If, says Du Hamel, we remove the leaves of a young tree when in full sap, and whose bark is easily detached, in a few days after the same bark will adhere as closely to the wood as it commonly does during winter. This direct connection between the leaves and bark is also well illustrated in an experiment of Hales, employed by him for a very different purpose. From two thriving shoots of a pear-tree he cut, in several places, half an inch of the bark off all round. All the ringlets of bark between these incisions had a Elementary leaf-bud upon them except one, and all but this one ring- Organs, let grew and swelled at their bottoms till August; andv-^v^“*>' the larger and more thriving the leaf-bud was, so much the more did the adjoining bark swell. ( Veg. Statics, p. 149.) Mr Knight also found the bark of the vine to become shrivelled and dry when the leaves were stripped off; but in those parts in which it communicated directly with the leaves, it continued moist and flourishing. {Phil. Trans. 1801, p. 335.) By connecting, therefore, the circumstances attending the flow of sap with the development of the leaves, we gain satisfactory reasons for all the apparent anomalies observed to attend its course. Early in spring, before the appearance of the leaves, no natural outlet for the escape of the rising sap exists, and therefore, when the vessels are cut or perforated, they readily pour out their sap, or bleed; but late in spring, and in summer, when the leaves are developed, the more watery parts of the sap are thrown off by transpiration; and while this process proceeds, the fluids do not accumulate in such quan¬ tity in the minute vessels of the trunk as to be effused or bleed through their cut or perforated sides. A cold day, however, or a moist and still atmosphere, by check¬ ing transpiration from the leaves, restores more or less the propensity to bleeding from the trunk ; and in au¬ tumn, when fructification begins, and vegetation makes a pause, the same disposition to bleeding recurs in the trunk, from the check imposed on the more active powers of growth. It is further evident, that, when the vessels of the Qualities bark become supplied with fluid, they cannot have de-of sap in rived it immediately from those of the wood, since, in die bark; these different parts, the fluids have frequently no sort of agreement in properties. Thus, Grew remarks, that al¬ most all plants, late in spring, and in summer, bleed from their bark; and the sap has either a sour, sweet, hot, bitter, or other taste. At this period the bark of the vine yields a sour sap; but, “ what is vulgarly called bleeding in the vine is,” he adds, “ quite another thing, both as to the liquor which issueth, and the place whence it issues; that is, it is neither a sweet nor a sour, but a tasteless sap, issuing, not from any vessels in the bark, but from the air-vessels of the wood.” {Anatomy of Plants, p. 125.) Malpighi, also, was well aware of the difference in the qualities of the sap, and thought every plant possessed its peculiar sap; but he has not so accurately defined the situation of the vessels that contain it. Du Hamel, how¬ ever, points out distinctly the difference of quality in the sap of the bark and wood. In the bark of some plants it is white, in others red, and in others yellow. It is in some instances milky, in others resinous, and in others gummy. In many plants it has a sweet taste, in some it is caustic, and in others insipid. It has sometimes much odour as well as flavour, and frequently it is destitute of both. {Phys. des Arbres, tome i. p. 68.) But if the qualities of the sap in the bark be thus acquired in different from those of the sap in the wood; if these pe- the leaves, culiar qualities are detected in it only after the develop¬ ment of the leaves, and the leaves be the organs by which alone the sap can be conveyed from the one part to the other; then it seems to follow that the sap must acquire these new properties in the leaves during its transmission through them. Malpighi remarked the existence of this altered sap in the leaves, and held them to be the organs which prepared nutrient matter for the plant; and Dr Darwin, by immersing plants of spurge in coloured li¬ quors, not only saw, as others had previously done, the red fluid ascend through the leaf, but another fluid, of a 52 ANATOMY, VEGETABLE. Descrip- Elementary white colour, returning at the same time from the ex- Organs. tremities of the leaf, and descending into the petiole. (Botanic Garden, vol. i. notes, p. 450-53.) This same returning fluid Mr Knight observed in similar experi¬ ments on branches of the apple and horse-chesnut trees, and even traced it through the petiole into the inner bark, by the vessels of which it seemed to be conveyed to the extremities of the roots. (Phil. Trans. 1801.) The motion, therefore, of the sap in the bark is not that of ascent, as Grew and Malpighi and many others have be¬ lieved, but of descent, as the observations and experiments of M. de Sarrabat, Du Hamel, Knight, and others, abun¬ dantly prove. To these vessels of the bark Grew assigned particular names, according to the apparent quality of the fluid they conveyed. Malpighi gave them the general appellation of vasapeculiaria, from their containing a fluid different from the common sap. By others they have been called cortical vessels, a term, however, not applica¬ ble to many tribes of vegetables which are entirely desti¬ tute of bark. Lastly, Du Hamel and others denominated them proper vessels, which term differs but slightly from the appellation of Malpighi, and, though not very precise, is that we shall continue to employ. From regarding the newly formed vessels of the wood tion ot the as 0f jjle bark, Grew uniformly represents the bark sels ^ VeS"as Possess*ng tw0 distinct species of vessels, in which he has been since followed by several other writers; but the vessels which he calls lymphceducts belong rather to the wood, so that we may regard him as describing only one species oi'proper vessels. In herbs, these vessels stand sometimes in distinct parcels or columns ; sometimes they are disposed in a ring; sometimes they have a radiated position; and sometimes they are more intermixed with the sap-vessels, and seem to alternate with them. In trees, the vessels of the bark are more distinct, and have a much more regular appearance. They are commonly postured near the inner margin of the bark, and, when viewed in a longitudinal direction, seem collected into fasciculi, which are more or less numerous, and the compo¬ nent vessels of which continually diverge and join with others, so as to form a reticulated appearance, as in fig. 24, Plate XXXVIII. Of these reticulated fasciculi, many layers exist in an old tree; and to these layers the thickness of the bark is chiefly owing. As they proceed inward, the direction of the fasciculi is less oblique, so that near to the wood they are almost straight. Hence the spaces formed by the reticulations are very unequal, often large in the exterior part of the bark, and diminishing in size towards the wood: they are. everywhere filled with cel¬ lular tissue. Such is a general description of the situa¬ tion of the proper vessels of the bark, as given by Grew, Malpighi, and Du Hamel. The vessels which thus form the vascular portion of the bark appear to differ but little in structure from the more simple vessels of the wood. Grew considered them to possess a similar structure, from believing them to be formed by the inner bark at the same time with the ves¬ sels of the wood; and therefore, he adds, “ they may be reasonably thought similar in the bark and wood.” (Anat. of Plants, p. 112.) Malpighi regarded them as simple tubes, containing sometimes peculiar juices; but advances nothing particular respecting their structure. (Anat. Plantar, p. 3.) M. Mirbel’s opinion deserves some no¬ tice, inasmuch as he declares the structure of these ves¬ sels to differ entirely from all those of the wood. Their sides, says he, are perfectly entire; they have neither pores nor clefts, and may therefore be deemed simple tubes. (Exposit. de la Theorie de V Organised. Veget. p. 109.) A single vessel of this kind is represented in Plate XXXVIII. their structure fig. 22; and in fig. 23 a fasciculus of the same vessels Element! ill magnified is given as delineated by M. Mirbel. A re- Ofgani, mark also of Hill, if it be deemed to rest on correct ob-^' servation, is entitled to great attention. “ The vessels of the bark that form the fasciculi are not,” says he, “ united to each other, but are connected with the cellular tissue at numerous places; and, when separated from it, there appear on the sides of the vessels small oval depressions, dotted as it were with pin-holes.” ( On the Construction of Timber, p. 28.) These appearances he regards as of a glandular nature, but their description corresponds better with that of Leeuwenhoeck respecting the lateral ramifica¬ tions from the vessels of the wood; and they may proba¬ bly be the points at which communication is effected be¬ tween the vessels and cells. The cause of the reticulated appearance which the their ret vessels of the bark exhibit in trees, is doubtless to be at-culafiro tributed, as we think Du Hamel somewhere remarks, to their peculiar mode of growth. A new layer of cortical vessels is every year added to. the inner surface of the bark, as well as a new layer of ligneous vessels to the out¬ er surface of the wood; so that, as Grew observes, the new matter of the tree is every year distributed two con¬ trary ways,—one part falling outward towards the bark, and the other part retaining its situation inward to consti¬ tute the wood. At first the newly formed cortical ves¬ sels are straight, and stand parallel, like those of the wood ; but, by the continual growth of the new parts formed between the bark and the wood, the older vessels of the bark are gradually forced outward; and being thus every year disposed around a large cylinder, are necessa¬ rily more and more separated from each other, and pro¬ duce at length that net-like form which we observe them to possess. The newly formed vessels of- the wood, on the other hand, retain their original position, and there¬ fore preserve their parallelism, seldom or never exhibit¬ ing those flexures and reticulations so common to the ves¬ sels of the bark. In the bark, as well as in the wood, the vessels are their sia found to possess different sizes. In the pine, vessels con¬ taining turpentine are represented by Grew, which are very much larger than the comjnon sap-vessels, and are surrounded by smaller ones, exactly as the large spiral tubes of the wood are said to be ensheathed by the com¬ mon sap-vessels. The milky juice of a species of sumach is contained in very large vessels, disposed so as to form a ring, and each large vessel is surrounded by many small¬ er ones. (Anat. of Plants, tab. 20.) The appearance of these large vessels in one species of pine is well repre¬ sented by Hill, and the account he gives of their forma¬ tion is probably correct. In this tree (pinus orientalis), some of these vessels form oval openings, large enough to admit a straw; these openings occupy the centre of the bark, and are surrounded by a ring of smaller vessels. As their contents are soluble in alcohol, it is easy to obtain them empty. In fig. 26, Plate XXXVIII., the vessels of this tree, as they appear in the bark, are displayed; the woody portion of the tree having been scooped away, so that the longitudinal aspect, as well as the transverse sections of them, is exhibited. From a strict inquiry into their na¬ ture, Dr Hill concluded that these larger vessels were originally the same as those of the smaller fasciculi in the bark of the same tree ; “ so that if we conceive one of these smaller fasciculi opened in its centre, and the vessels driven every way outward, till they are stopped by the substance of the bark, we shall have an idea of the structure of this large vessel, which is nothing more than a great cylindrical hollow formed in the centre of such a fasciculus.” (Hill on the Construction of Timber^ ■y lemitary p. 29.) It is in trees that have copious and viscid juices j Oi^is. that these enlarged vessels are formed; and where the juices do not concrete, it is probable that, as the vessels annually recede from the centre, they suffer a reduction in size from the continued effects of desiccation and com¬ pression to which they are exposed. ‘‘ einsi- The foregoing varieties appear alone entitled to the )ny appellation of the proper vessels of the bark. In many tribes of vegetables, however, as will afterwards be shown, no distinction of bark and wood exists ; but one uniform distribution of vessels extends from the centre to the cir¬ cumference of the plant. Such plants have also their - proper vessels, but the place and disposition of these ves¬ sels are not so precisely ascertained. From the mode in which their growth is accomplished, as well as from ob¬ servation of their structure, it may be inferred that their J proper vessels are distributed through the whole substance of the plant, accompanying in every part the sap-vessels. J 111 some plants possessing this arrangement of parts, such as different species of, wheat,‘Malpighi {Anal. Plant. tab. 4, fig. 15) describes and delineates & vas proprium forming a part of each fasciculus of vessels; and a simi- * lar intermixture, of the two kinds must, we conceive, 1 exist in ah similar structures. Even in many plants possessing a distinct bark, ves- ! sels containing proper juices are found in the wood. In 1 every circle of wood, from the inmost that surrounds the pith, to the outmost in contact with the bark, vessels containing a gum, turpentine, or some other concrete or coloured juice, Vnay frequently be found. Malpighi con¬ ceived ihem to exist in all plants, though, from the nature of their fluids, they could not always be distinguished ; and he believed them to afford a highly perfect juice for the nutrition of.the plant. According to Grew, “ the tur- pentine vessels that are scattered up and down the wood of the pine and fir are the self-same which did once ap- j pertain to the bark; but being pinched up by the wood, 1 they are become much smaller pipes.” (Anat. of Plants, 1 p. 115.) Du Hamel also regarded them as similar to t those of the bark, but rendered much smaller by com- 1 pression. In the ,pine and fir they are disposed circularly a around the axis, much like, the sap-vessels, and alternate ' with them. (Phys. des Arbres, tf»me i. p. 41.) In Piscidia erythrina, the proper juices are of a scar- 1 let colour, and the vessels that contain them are there- f fore readily discerned wherever they exist. This plant has been selected by Hill to demonstrate the position of t these vessels. In the bark they are collected into fasci- c culi, all the vessels of which contain coloured juices, and a are disposed in a ring on the inner margin of the bark. 1 Within this ring stands the alburnum, through the sub- s stance of which many smaller red vessels are distributed ; a and similar red vessels are more sparingly seen in every 1 layer of wood, particularly in that which envelopes the j pith. (Construction of Timber.} Now the red vessels thus observed in the wood of the ; above-mentioned plant must either have been formed in the situations they occupy, or transported from some other ] place. The latter supposition is inadmissible, inasmuch : as the wood of trees is formed by layers of new vessels s superimposed on one another ; and no removal of the old i vessels, nor reproduction of new vessels within the old 1 layers, ever takes place ; consequently no actual transpo- s sition of vessels could occur, nor could new vessels be c developed in the wood after it had been completely form- e ed. If, however, an alternate deposition and absorption o of matter go on into and from the cells, it is possible that t the vessels might in this way become filled with a mat- t ter different from that which they originally possessed; 53 but in the case before us, a readier explanation presents Elementary itself. The new matter of the wood is formed at the same Organs, time, and in the same place, as that of the bark; andv>^~v^,,-/ through this new woody matter the red vessels were dis¬ persed as well as in the bark. Consequently every addi¬ tion made to the ligneous layers would furnish some vessels that contained these proper juices; and this, being annu¬ ally repeated, would exhibit that intermixture of proper vessels with sap-vessels which is observed in all the lig¬ neous layers of this and many other trees. Hence these proper vessels of the wood must be held to retain the po¬ sition in which they were originally produced, and cannot be said to have approached nearer to the centre, but only, by the addition of new layers exterior to them, to be placed farther from the circumference of the tree. As Grew, therefore, held the alburnum to be a part of the bark, he might correctly say that these vessels “ did once appertain to it.” Art. VII.— Of Collections of the Proper Juices in the Cellular Tissue. Besides this accumulation of the proper juices in cer-In single tain vessels of the wood, it frequently happens that de-ceUs- positions of similar matter occur in all parts of the cel¬ lular tissue of plants. In the bark of the oak and poplar, and of other trees, resinous concretions are often found in the cells; they are situated irregularly, and, according to Malpighi, are observed even in very young bark. Some¬ times the cells containing milky and resinous juices are so postured in the bark, says Grew, as to form cylindrical channels, which are neither parallel nor anywhere inos¬ culated, but run, with some little obliquities, distinct from one another. They appear to be formed out of the cells, and are not bounded by any walls or sides proper to them¬ selves, but only by those of the cells. (Anat. of Plants, p. 112.) They are often short and tortuous, always iso¬ lated ; and are sometimes placed irregularly, at others disposed in a circular form. They occur sometimes in the pith, and possess very different sizes and figures. They have frequently been deemed a species of proper vessels, from the mere circumstance of their containing similar juices, and from possessing sometimes an elongat¬ ed form; but they are organs which, neither in form nor in function, bear any resemblance to vessels. Mirbel pro¬ poses to name them secretory canals, and M. Link cellular reservoirs—the term certainly most generally applicable, and involving no hypothesis respecting either their forma¬ tion or functions. The manner in which these cellular reservoirs may In cellular be produced in the bark or pith is readily explained, on reserv°irs- the supposition that a communication everywhere subsists between the vessels and the cells. One set of vessels has been shown to receive and carry sap to the leaves, and another to bring it back from them to the bark; and these two sets of vessels are everywhere in their course surround¬ ed by cellular tissue. Hence the cells in every part may receive a portion of the fluid which the vessels are em¬ ployed to convey. Thus in herbs, the cells both of the bark and pith are filled with fluid, which led Grew to believe that the sap was actually transmitted through those or¬ gans ; but at the same time he delivers facts which, even in his own opinion, prove that it is derived directly from the vessels. Not only in herbs, but “ in every annual growth, whether of a sprout from a seed, of a sucker from a root, or of a scion from a branch, the pith is always found the first year full of sap; but in the second year the same individual pith always becomes dry, and so it continues ever after. One cause whereof is, that the ANATOMY, VEGETABLE. 54 ANATOMY, VEGETABLE. Elementary lymphasducts of the bark being the first year adjacent to Organs, the pith, they do all that time transfuse part of their sap into it, and so keep it always succulent. But the same lymphseducts the following year are turned into wood, and the vessels which are then generated and carry the sap stand beyond them in the bark; so that the sap, being now more remote from the pith, and intercepted by the new wood, cannot be transfused with that sufficient force and plenty as before into the pith, which, therefore, from the first year always continues dry.” (Anat. of Plants, p. 124.) All that is here said respecting the transfusion of the common sap from the vessels into the cells as it ascends through the wood, is equally applicable to the proper juices as they descend through the bark. During the first year of growth, both the sap and proper vessels are adja¬ cent to the pith as well as to the bark, and each order may' therefore transfuse its fluids into the cells of either. The common sap, from retaining its fluidity, is frequent¬ ly removed by absorption, and the cells that contained it appear empty and dry; but where the proper juices are transfused, and become viscid or concrete, they are re¬ tained, and appear in different quantities and forms both in the bark and pith, according to the nature and proper¬ ties of the fluids from which they are derived, and the texture and situation of the tissue into which they are poured. In inter- To these collections of the proper juices in the cells cellular of the cellular tissue may be referred the opinions of canals. those who describe them as existing sometimes in vacui¬ ties between the cells. Grew had remarked the occa¬ sional presence of these juices in cylindrical channels formed by these vacuities, to which M. Treviranus has given the name of intercellular canals. According to him, these canals not only contain the proper juices, but con¬ vey them to all the cellular parts of plants, and are the true proper vessels of the bark. We before saw that M. Kieser, after making the wood to consist almost entire¬ ly of spiral vessels, destined only to convey air, supposed the sap to be carried by these same intercellular canals. (Kieser, Mem. sur V Organisat. des Plantes, p. 36.) With M. Treviranus, he also considers them as the real proper vessels of the bark, by which alone the juices are convey¬ ed. (Ibid. p. 86.) Opinion of M. de Candolle considers the proper juices to possess DeCan- no special organ for their conveyance, but to nestle in dolle. tiie cavities that surround them, and form sacs or reser¬ voirs of various size and form. He distinguishes several kinds of these reservoirs, which vary with the figure of the containing sac. Thus, what others call “ proper vessels,” he designates by the names of tubular and fascicular reser¬ voirs. In this, however, he appears to confound the “ proper juices,” destined immediately to the functions of nutrition and growth, with various fluid or solid matters, either simply deposited in the cellular tissue, or that have undergone subsequent alteration by the agency of a secret¬ ing organ. The reservoirs of the proper juices, and the juices themselves, are, he says, readily seen in many fami¬ lies of dicotyledons ; but have not been observed with cer¬ tainty, either in monocotyledonous or acotyledonous tribes. ( Organog. Veg. tome i. chap. 11.) The reason of this seems to be, that in dicotyledons the bark, in which the pro¬ per juices are seen, is distinct from the wood, in which the common sap flows; and the colour, consistence, and quali¬ ties of the two' fluids, therefore, at once point out their respective places; but in plants belonging to the two lat¬ ter tribes there is no proper distinction of bark and wood; and the two orders of vessels, as shown by Malpighi in the instance of wheat, are associated together through the whole substance of the plant, which renders the discrimi- Element;, nation of their common and proper juices more difficult. Orgai, ^ Besides two orders of vessels existing in the ligneous part of trees, M. Dutrochet describes a particular form of^™0” cell, which, in the wood, serves only as a receptacle forchel , fluids; but to which, in the bark, he assigns a vascular function. This is the elongated cell observed by Mal¬ pighi, and represented by him in fig. 18, Plate XXXIX. M. Dutrochet, from the resemblance which its figure is said to bear to a spindle, has given it the name of clostre; and his representation of it is given in fig. 17, Plate XXXIX. These elongated cells form what Kieser and others have called the ligneous fibres. By M. Du- trochet they are regarded as reservoirs of the nutrient juices, and as containing secretions which impart soli¬ dity to the wood. In the bark, however, these same clostres are declared to be the organs in which the ela¬ borated sap is contained, and by which jt is transmitted to the roots to furnish the materials of their growth. The ^ ^ clostres, which thus convey the nutrient matter, ought not, he adds, to be confounded with the “ proper vessels,” which are the true secretory organs. These vessels are tubes of greater diameter than the clostres, and contain various excrementitious substances, as resin, &c. in differ¬ ent plants. (Recherches Anatoyniques, 8zc. SI.') Ac¬ cording to M. Dutrochet, therefore, the elongated ceils and vessels which exist in the bark and wood exercise functions precisely opposite; for whilst in the wood the corpusculiferous vessels convey the sap, and the cells are reservoirs for containing it, in the bark the cells are the organs which transmit the elaborated sap, and the vessels those in which the excrementitious matters are contained. In our view, these notions are the reverse of fact, since we consider the vessels, both in the wood and the bark, as the organs by which the fluids are conveyed, and the various forms of the cellular tissue as the general recep¬ tacles into which they are in part deposited: they may, however, in some instances be retained in the vessels themselves, as well as in the cells. Section II. Of the Absorbent and Exhalent Systems. Art. I.— Of the Absorbent System. tlw Connected with the vessels that distribute the fluids through organized bodies, is another system of vessels, by which extraneous matter is taken -up to support the growth of parts, and supply the waste occasioned by the exercise of the various functions. To these vessels anatomists have given the name of Absorbents. The function which they perform is carried on either from the external surface, or from some internal part of the body; and its exercise, in animal bodies, may be distin¬ guished into three kinds or stages. The first is that in which new or extraneous matter is taken up and added to the system, as in the absorption of substances from the Sp skin, or of chyle from the intestines; the second is that in which substances previously separated from the fluids by secretion, but without becoming organized, are again taken up by the absorbents, and reconveyed into the blood-vessels, as in the absorption of milk, bile, and fat; the third kind is that in which the secreting organs them¬ selves, and successively all the solid parts of the body, are removed by the action of the absorbent vessels. By plants this function is exercised to less extent, and seems to comprise only the first two kinds or degrees of it, viz. the primary absorption of extraneous matter, and the re¬ absorption of certain secreted matters; but in them there ANATOMY, VEGETABLE. 55 n ■/np'vlL' b > tioin nU* ^erj, eipif r t n ISOft- nuts | n- r* tl P tl s; v fi ti w a e oi’ffliles, does not appear any power of re-absorbing what has once formed an organized part of the system. Hence the for¬ mation of organized parts in plants is not accompanied, as in animals, by the unceasing removal of old particles; but the particles which have once become organized con¬ tinue permanent until removed by some cause or process foreign to the living powers of the plant. When once, therefore, the organs of plants have become mature, they are exposed to decay only from the operation of foreign causes, and cannot be mined from within by that gradual loss of balance between the secreting and absorbing func¬ tions which the advances of age bring on the animal sys¬ tem. Neither, in the vegetable system, can the removal of organized parts under disease, any more than in health, have place; and, consequently, the several modes or stages of ulcerative absorption, so finely illustrated by the researches of Mr Hunter, belong not to the economy of plants. As the function of absorption is thus of more limited extent in plants than in animals, so may we expect to find the arrangement of organs destined to its exercise. In the more perfect animals, the absorbent vessels are quite distinct from those which carry blood, are pro¬ vided everywhere with glands, and, like the veins, are furnished with valves. From their beginnings, in all parts of the body, to their termination, they continually unite with one another, until, after a long course, they form at length two trunks, which deliver their contents into the large veins near the heart. As no common reservoir exists in plants, there is no such single point to which their ab¬ sorbed fluids require to be carried. Hence their absor¬ bents seem everywhere to have a very short course, to form no union with one another, but to deliver their con¬ tents at once into the sap-vessels adjacent to them. They appear to be destitute alike of glands and valves, and in¬ deed, in an anatomical view, they can scarcely be con¬ sidered distinct from the sap-vessels, but may rather be deemed ramifications from them; so that although we grant, with Grew, that the ordinary sap-vessels do not ramify one out of another, yet they certainly send olf those fine ramifications which, from their office, we de¬ nominate the absorbents of plants. In the root, the absorbents are capable of being de¬ monstrated. When a plant is immersed in coloured fluids, many of its capillary absorbents become tinged through their whole course to their termination in the sap-vessels, proving them to be simple ramifications from these vessels themselves. A further proof of the identity of these two systems is derived from the fact, that any part of a sap- vessel is not only capable of emitting capillary absorbents from its sides, but of exercising itself an absorbent func¬ tion, whenever its cut extremity is brought into contact with a fluid. These absorbents are formed very speedily and in great multitudes on the roots of annual plants; and even in perennial plants they appear, like the leaves, to suffer an annual decay, and to be reproduced with the re¬ turn of vegetation. ■ M. de Candolle describes a structure as existing at the extremities of the roots and of the pistil, and also on the outer coat of seeds, which possesses a peculiar aptitude for absorption. To this structure he gives the name of spon- giole: it is composed of small round cells, and in roots is situated at all their fibrous extremities. The entire body of each radical fibril is indeed made up of an analogous cellular tissue; but it is the extremity alone which actual¬ ly absorbs, and which therefore possesses a peculiar ab¬ sorbent or hygroscopic power. The pistillary spongioles are in like manner situated at the extremity of that organ, and absorb the fecundating liquor. The third variety, or seminal spongioles (spongiolce seminales), are Elementary placed on the surface of the grain, apparently with great Organs, regularity; and by them the absorption of moisture is ef- fected. Not only water, but coloured liquors, are ab¬ sorbed by these spongioles, though the molecules of such liquids never enter the ordinary pores of plants, which pores must be infinitely greater than the apertures with which the spongioles may be furnished. This is the more remarkable, says M. de Candolle, when we call to mind that these colouring molecules traverse the compact and almost stony coats of seeds, and yet do not enter leaves whose tissue is so lax, and which are visibly furnished with pores that have the power of absorbing water when in contact with it. ( Organog. Vegetale, tome i. chap. 7.) But plants absorb fluids by other parts of their sur-Absorption face as well as by the roots. This absorbing power ex- by leaves, tends, in some of the lower tribes, over the entire surface of the vegetable, which is destitute of any organ analogous to a root; in other instances, where the roots are small and the soil arid, the plant derives almost all its moisture by the absorption of dews through the leaves. The ex¬ periments of Bonnet show that all leaves, both those of herbs and of trees, when brought into contact with water, are capable of absorbing it, and that the moisture thus ab¬ sorbed is communicated through the vascular system of the leaf. The leaves of herbs he found to absorb nearly alike from either surface ; but those of trees absorbed best by the lower surface. The petiole and larger riblets ap¬ peared to absorb much less than the other parts of the leaf. So great and general is this absorbing power, that vegetables, says he, may be said to be planted in the air, nearly as they are in the earth, the leaves being to the branches what the capillary rootlets are to the roots. {Recherches sur l' Usage des Feuilles, p. 22, 47.) What then are the organs by which this function is External carried on in the leaf? M. Bonnet imagined the vessels al)SOrPtioa of the leaves to receive their fluids through the pores ^ P01’68, adjacent to them; and that the leaves, which had only few pores, possessed but little absorbent power. {Ibid. p. 20, 22.) He thought, also, that the hairs frequently dis¬ tributed over the leaf attracted moisture, and might even act as absorbents; though he admits that many leaves which have only slight inequalities on their surface, with¬ out hairs, exercise an absorbent function. {Ibid. p. 47.) This subject has been since investigated by M. de Can¬ dolle, whose researches appear to confirm the account of Bonnet as to the absorbing power of the pores. Accord¬ ing to him, these pores are found on all parts of the leaf except the riblets, which have none, but are covered with hairs. At the mouth of the pore a vascular net-work is always to be found, which he regards as a production from the larger vessels of the leaf. He asserts that pores are found only in those parts where vessels go to terminate, and not in others; and that in trees this structure occurs chiefly on the lower surface of the leaf, while in herbs it is equally seen on both surfaces. The stem in general has few or no pores, except where it is soft and her¬ baceous ; and even then the pores occur only in the deeper green furrows, not on the prominent lines which bound them, and are usually covered with hairs. No pores are to be found on roots or bulbs or fleshy fruits, but most of the organs above ground are more or less furnished with them. Exposure to the air seems necessary to their formation; for plants, or parts of plants, that live beneath water or earth, are destitute of pores, but acquire them if brought into the free air. Exclusion of light prevents also the for¬ mation of pores; and hence etiolated plants are not fur¬ nished with them. {Mem. de VInstit. Nat. tome i. p. 351.) In general, his anatomical researches respecting the ex- 56 ANATOMY, VEGETABLE. Internal absorp¬ tion : Elementary istence of pores in leaves agreed with the results of Bon- Organs, net’s experiments on the absorbing power of their surfaces ; and when we consider that all plants and parts of plants secluded from the air are at the same time destitute of pores, and of the power of absorbing by their surfaces, it may be inferred that the organs of absorption on the ex¬ ternal surfaces of plants are the minute vascular produc¬ tions which, in tender and succulent parts exposed to the air and light, everywhere perforate the cuticle, and form in it those innumerable orifices which we denominate pores. But the function of absorption in plants is not con¬ fined to the taking up of extraneous matters. Many facts prove that, in every part where active vegetation exists, internal absorption is continually going on. The organs by which this absorption is immediately performed can¬ not, perhaps, from their extreme delicacy and minuteness, be rendered capable of anatomical demonstration; but certain facts which occur in plants, coupled with certain analogies derived from other organized textures, must, we think, carry complete conviction of their existence. In almost every part, and on every surface of animal bodies, the vessels which exercise absorption may be traced ; but the mouths or orifices by which they actually absorb are scarcely ever to be seen. In one instance only, viz. in the intestines, have they been followed to their begin¬ nings, and discovered in the act of exercising their appro¬ priate function. (Gordon’s System of Human Anat. vol. i. p. 70.) But though their orifices remain in other parts undiscovered, no anatomist hesitates to admit their exist¬ ence when he sees the canals of the vessels themselves laden with blood, or milk, or bile; and seeing them thus to convey fluids whose colour manifests their presence, he equally believes them capable of absorbing and convey¬ ing other fluids, though they may not be visible to the eye. Believing further, that no solid part of the body, nor even fluid part that has been deposited in close cavi¬ ties, can be removed in a natural manner from its place, but by the agency of these vessels, he comes to regard the simple fact of the disappearance of such solid or fluid part as sufficient evidence of its absorption. Now the facts and analogies on which internal ab¬ sorption rests in the vegetable system are precisely of the same nature, and the evidence of its existence is scarcely less complete. In every part of the cellular tissue of plants various substances have been found which must have been primarily derived from the vessels, the only organs which furnish new materials to the plant. These substances, however, often disappear from the cells, and are again to be detected in the vascular system. Thus, in the seed, as will afterwards be shown, the cells of the cotyledon contain a solid unorganized matter, which could have been originally deposited in them only by means of the vessels. During germination, this solid matter is ren¬ dered fluid, disappears from the cells, and is again to be traced in the vessels on its way to afford nutriment to the radicle and plume. We say, then, that this unorganized matter must have been taken up from the cells of the co¬ tyledon, and conveyed into the vascular system, by the agency of absorbent vessels, which, it is probable, are dis¬ tributed everywhere on the inner surface of these cells ; just as, in the animal system, absorbent vessels are con¬ sidered to take up the fat from the inner surface of the cells in which it is deposited, and convey it into the vessels of the animal. This alternate absorption and deposition of the nu¬ trient matter of seeds is sometimes strikingly display¬ ed in the growth of potatoes. It frequently happens that potatoes lying in a damp cellar put forth shoots which grow to a considerable size, without the access of any in seeds; foreign agents except heat, water, and air. On these Elen*, shoots young bulbs, as large as the eggs of pigeons, are °fgii sometimes to be found, and the substance of the oldS^ bulb has in great part disappeared. In such cases, the111 matter from the cells of the old bulb must be consider¬ ed as removed by absorption, and conveyed into the vessels of the shoot, where it was in part employed in forming the new organs of the young bulb, and in part deposited, to experience, perhaps, in some future growth, similar successions of removal and deposition. In the living parts of perennial plants, also, nutrient matter ap¬ pears to be alternately deposited and absorbed from the cells during the active periods of vegetation ; and in the cellular tissue of herbaceous plants a similar deposition and absorption of fluids seems to be frequently taking place; so that, in all the vegetating parts of plants, these alternate functions of secretion and absorption are more or less constantly exercised. A good illustration of the manner in which these func-in tmi tions are alternately exercised is afforded by an ex¬ periment of M. de Candolle. The parasitic plant called misletoe draws its nourishment, as is well known, from the tree on which it grows. M. de Candolle placed a branch of an apple-tree, bearing a stalk of misletoe, in an infusion of cochineal for five days. He then dissected it, and observed the coloured liquor to have risen through the wood and alburnum of the apple-branch, and reached the place of junction between it and the misletoe, which it strongly reddened; and from thence it penetrated into the woody part of the misletoe. There did not, however, appear to be a true anastomosis between the vessels of these different plants; but, at the base of the misletoe, where the parts were so deeply reddened, a minute cel¬ lular structure was observed. Into these cells the vascu¬ lar system of the apple appeared to deposit its sap, and from them the capillary absorbents of the misletoe, distri¬ buted upon the cells, seemed, like the ordinary absorbents of roots, to take it up. {Mem. de Vlnstit. Nat. tome i. p. 370.) From these and many similar facts it may be in¬ ferred, that absorbents, communicating with the vessels of the plant, exist in every part, and that the removal of all secreted matters from the cells and other close cavities of the vegetable, when effected by the living powers of the plant, is accomplished, as in animal bodies, by the exer¬ cise of an absorbent function. Art. II.—Of the Exhalent System. But from their external surface, and from the same Organs parts as we have seen to exercise an absorbent function, exhalat plants, in certain circumstances, give off a large quantity of fluid by transpiration; and the organs by which this function is performed seem, from many considerations, to be the same as those by which, under other circumstances, absorption is accomplished. This function of transpiration is common to all terrestrial plants, and, with few excep¬ tions, all are more or less furnished with pores; hut it does not occur in aquatic plants, according to De Candolle, which are destitute of pores. Fleshy leaves and fruits, and the petals of flowers, which have but. few pores, transpire little; and etiolated plants, which.are destitute of pores, do not transpire at all. On the contrary, herba¬ ceous stems and plants, which have numerous pores, throw off most fluid by transpiration. This general agree¬ ment between the existence of pores and the exercise of the transpiratory function leads to the presumption that they are the orifices through which the fluids are dis¬ charged ; and if it be admitted that these pores are situ¬ ated at the extremity of the fine ramifications that come ANATOMY, VEGETABLE. 57 ] emetary off from the vessels, their fitness for such an office cannot be denied. Comparing these facts regarding transpiration with those previously stated concerning absorption, M. de Candolle is led to conclude that the pores on the sur¬ faces of plants are the organs by which these functions are alternately carried on, according to the existing con¬ dition of dryness or humidity in the surrounding atmo¬ sphere. ; -ae Repulsive as this conclusion may at first seem to tlio»‘ of our ordinary conceptions of organized bodies, yet there sopon are many circumstances in the structure and habits of plants that give to it great probability; so much so, that we ourselves had long since reached the same point by a route different from that pursued by M. de Candolle. It is highly probable that the exhalents of the leaves are simple ramifications from the larger vessels, like the ca¬ pillary rootlets ; and as they have no valves in their canal, there is no mechanical impediment to their exercising an inverted action. The sap-vessels themselves readily ab¬ sorb even coloured fluids when inverted; and though their exhalent terminations are too fine to receive such fluids, yet why may they not, like the trunks from which they spring, be capable of taking up ordinary fluids in that manner? The fluids absorbed through the leaves must at once enter the sap-vessels, for there is no com¬ mon reservoir to which they can first be carried; and it is extremely improbable that, from the same parts of the same vessels, exhalents and absorbents, capable of exer¬ cising only opposite functions, should at the same time arise. In the animal system the exhalents spring from arteries, and the absorbents terminate in veins; but in the less complex structure of plants it seems demonstrable that both orders of vessels must at once communicate with the same sap-vessels. It is therefore more proba¬ ble to suppose that, instead of two distinct orders of vessels, as in animals,, one only should be provided, capa¬ ble, under different circumstances, of exercising different functions. This vicarious office of the organs under con¬ sideration leads to no confusion in its exercise; for the condition of the atmosphere, which favours transpiration, is that which removes from the leaves the power of ab¬ sorption ; and, on the contrary, absorption occurs only in a humid atmosphere, when, as Hales has shown, little or no transpiration takes place. Besides exhalation carried on through visible pores, plants, like other bodies, lose their humidity by simple evaporation. This evaporation seems to go on in a mo¬ derate degree under favourable circumstances by day and by night, and is quite distinct from the copious transpira¬ tion that is carried on by the agency of solar light through the pores. That the pores are exhaling organs, was an opinion first advanced by Hedwig, and subsequently illustrated by De Candolle, Sprengel, Link, and Rudol- phi. (Organog. Vegetale, tome i. p. 86.) In like manner, though absorption may be carried on in certain circum¬ stances through the pores of leaves, yet there are plants which largely imbibe water, but in which few or no pores have been detected. Thus, fleshy leaves which have lost their moisture quickly regain it either when plunged in water or exposed to rain, though they possess but few pores; and';The aquatic algae, says M. de Candolle, evi¬ dently imbibe water over their whole surface, though they are wholly destitute of pores. He supposes that, beside the ordinary pores which may be readily seen, plants may be furnished with others that are invisible from their extreme minuteness. By such pores we might account for the loss of weight which plants destitute of visible pores gradually undergo when exposed to the free air, and also for the increase of weight which mosses, and vol. in. apciv n. other plants equally destitute of pores, rapidly acquire Elementary when plunged in water. (Organog. Veget. tome i. p. 88.) Organs. The view thus presented of the external absorbent and exhalent vessels may probably be extended to the Inte™al minute vascular productions which seem everywhere toancj ak_ spring from the vessels internally. If secreting and ab-sorbing sorbing vessels be held to exist in every part of the plant, organs, they must everywhere communicate with the vascular system ; for it is from the vessels of this system that the matter of their secretions is primarily derived, and it is into the same vessels that, in many cases, these secretions are subsequently returned. Nor does the exercise of the two functions of secretion and absorption in plants pre¬ sent any apparent obstacle to the supposition of their being performed at different times by the same organs. Thus, when nutrient matter is deposited in the cellular tissue of the seed, it is destined only for a future use; and the purpose of nature would be defeated were an ab¬ sorbent function to be at the same time employed for its removal. On the other hand, when this matter is again taken up during the germination of the seed, no secreting function seems then to be exercised in that part; for the organ itself, in many seeds, gradually wastes, and no fresh matter is deposited in it. Even when the cotyledon to a certain extent augments in size, its nutrient matter is continually drawn off for the support of the radicle and plume, and no fresh matter of the same kind seems to be then deposited; so that the same vessels which formerly exercised the function of secretion may, without disturb¬ ing the economy of the plant, be now employed in the ex¬ ercise of absorption. As thus the two functions do not require to be performed in the same part at the same time, they may, if nothing else forbid, be exercised at dif¬ ferent times by the same organs. In the animal system, where the organs themselves are removed, secreting and absorbing vessels must necessarily co-exist; and to main¬ tain the integrity of parts, their functions must proceed at the same time, and to a certain extent balance each other; but as no similar operations appear to be carried on in the vegetable system, no such complex organization is required to sustain them. Section III. Of the Cellular Tissue. Art. I.—Description and Structure of the Cells. The elementary organ denominated cellular tissue may Definition, be said to consist of a membranous substance, disposed into a great number of small circumscribed cavities, con¬ nected with each other, and arranged in rows or suites, generally in a direction opposed to that of the tubes, which represent the vascular system. From Grew it re¬ ceived the appellation of parenchyma, a term still often used in describing different parts of this tissue ; by Mal¬ pighi, it was called the utricular substance; and it owes, we believe, its present name to M. du Hamel. The cavities which distinguish its construction were Descrip- called indifferently bags or bladders, pores and cells, by Aon. Grew; by Malpighi, utricles; by others, vesicles; and more commonly cells. The form of these cells varies so much, not only in different plants, but in different parts of the same plant, as to have authorized, in some degree, these different appellations. The tissue which they con¬ stitute enters into the composition of every organ in the more perfect plants. Of many herbaceous plants it forms the chief portion, and some of the lower tribes of vege¬ tables, as the algae, lichens, mushrooms, &c. are said to be wholly composed of it; in other words, no vessels can be H ANATOMY, VEGETABLE. 58 Elementary actually demonstrated in them. In most cases it contri- Organs. butes greatly to modify the form of organs, and adds al- ways to their bulk and strength. Nothing can exceed the diversity of appearance in figure, bulk, and texture, which it exhibits in the several parts, circumstances, and conditions in which it is placed. It represents sometimes a lax cellular substance, all the parts of which are succu¬ lent and transparent; in other instances it is compressed into a solid, opaque body, retaining but faint traces of its former cellularity; and in others, again, it is spread out into a most thin and delicate membrane, in which the cellular character is wholly lost. It everywhere envelopes and holds together the vascular system, and seems to be the general receptacle of almost all the vegetable secre¬ tions. Figure of The figure of the component cells of this tissue is the cells; exceedingly various. Sometimes they have nearly a globular or spheroidal shape; in other instances they are angular, and exhibit in their section a greater or lesser number of sides and angles, being in a few examples tri¬ angular ; in others square, but more commonly hexago¬ nal, the figure which collections of soft cells, mutually impressing each other, seem naturally disposed to assume. This form is represented in the transverse section of cells, fig. 28, Plate XXXVIII.; they seem in this figure, and in most of those given in different works, to possess double sides; but, as M. Kieser has remarked (Mtm. sur F Or- ganisat. des Plantes, p. 91), this appearance is produced by the borders of the subjacent cells being seen through the transparent sides of the superior layer. In Plate XXXIX. fig. 16, is a representation of a transverse slice of the cellular part of sugar-cane, drawn from nature, and so thin as to exhibit only one layer of cells, in which the sides appear distinctly single ; but, in a thicker slice of the same plant, fig. 21, comprehending more than one layer, the double appearance becomes very evident. their size; The size of the cells varies not less than their figure in different plants and in different parts of the same plant. In one of the plates of Grew they are represented as possessing twenty different sizes, from that of a minute pore, to the size of a common pea. Hooke examined them in cork, and in the pith of many plants. In cork he reckoned several lines of these cells or pores, as he calls them, and found there were about sixty placed endwise in one eighteenth part of an inch, or somewhat more than a thousand in the length of an inch; and, therefore, in a square inch above a million, and in a cubic inch above 1200 millions. (Mierograpkia, p. 114.) In this substance the cells are not visible by the naked eye, but become very distinct when highly magnified. In most plants, however, they are readily visible, and their appearance is familiar to every one. When viewed in a longitudinal section, their hexagonal form is much less distinct, and is sometimes wholly lost. In fig. 22, Plate XXXIX., is a series of single columns of the cells of sugar-cane, in which each cell is, to appear¬ ance, bounded only by four sides. Similar representations are given by Hooke of the cells in cork, and by Kieser in most of the figures which exhibit longitudinal sections of the cellular tissue; but in some instances the hexagonal form is visible even in these sections. In fig. 23 of the •came plate we have given, in outline, the appearance of two series of columns of these transparent cells, in which one series is seen behind the other, and gives somewhat of the confused double appearance exhibited in the trans¬ verse section, fig. 21. contents; The nature of the matter contained in the cells of this tissue varies according to the part in which it exists, and the peculiar powers of the plant. Both Hooke and Grew remarked, that in the pith and bark of succulent Elemen, plants the cells were often filled with aqueous juices, and Orgat, in the same plant, at other periods, they appeared empty, or filled only with air. In the seed, the cells of the coty¬ ledons contain minute unorganized particles, which, at a 1 future period, serve as nutriment for the young plantule. Other particles of still smaller size, of a resinous nature and a green colour, exist in other parts of this tissue, and bestow on the plant its verdure. In every part of the plant these cells are also the occasional receptacles of the peculiar fluids which both the sap-vessels and the proper vessels convey; and hence various gummy and resinous substances, corresponding in quality to the fluids previous¬ ly existing in the vascular system, are frequently detected in them. In the pulp of fruits, the various acid, saccha¬ rine, and austere juices that we meet with, are contained in different modifications of this tissue; and it is into its cells that the osseous secretions, which constitute their shells and stones, are made. These facts prove not only the great importance of this tissue in the construction of the vegetable organs, but the active share it bears in the economy of their functions; and demonstrate likewise an universal communication between the vessels and the cells. The sides of these cells, when emptied of their con-struck tents, and viewed through the microscope, appear to be formed by a very fine transparent membrane, which some maintain to be everywhere entire, and others to be perfo¬ rated with pores. The same sources of error exist here as before noticed in similar microscopical observations on the vascular system; and, accordingly, the respective dis¬ putants maintain, with equal confidence, the same opinions with regard to the porosity or non-porosity of the cells, as they had previously held concerning the vessels. We must therefore call in the aid of other means besides those of the microscope, for determining the important fact whether the cells have or have not any direct com¬ munication with each other. Dr Hooke examined the films or sides of the cells ofare ck cork, of the pith of elder, and of many other plants, forcavities the very purpose of discovering whether any direct com¬ munication existed between them; but “ each cavern or cell,” says he, “ is distinctly separate from the rest, with¬ out any kind of hole in the encompassing filmsnor could he, with his microscope, nor by his breath, nor by any other way that he tried, “ discover a passage out of one of those cavities into another.” (Micrographia, p. 116.) Dr Grew describes the little cells or bladders that com¬ pose the bark of roots as possessing a spheroidal shape in most plants. When viewed with the microscope, their sides are as transparent as water; and “ none of them,” he adds, “ are visibly pervious from one into another, but each is bounded within itself.” (Anat. of Plants, p. 64.) Both Hooke and Grew, however, believed a communica¬ tion to exist between the cells, from the fact of their con¬ taining liquor; and Malpighi held the same opinion from similar considerations ; but they nowhere describe the mode or structure by which they conceived it to be ac¬ complished. Later writers have not only adopted this opinion, but professed to demonstrate the structure by which the communication is maintained. M. Mirbel describes the sides of the cells as composed of an extremely thin, colourless, and transparent membrane, which is common¬ ly perforated with pores, the diameter of whose aperture is not perhaps the 300th part of a millimetre. These pores are ranged generally in transverse series, and through them, it is said, the cellular tissue both receives fluids from the vessels, and transmits them very slowly tie C ANATOMY, VEGETABLE. 59 lerolu caials. through its cells. {Exposition de la Theorie, &c. p. 105.) M. Sprengel and some others adopt this view of the po¬ rosity of the cells; but it is denied by Link, Treviranus, and Kieser. The latter author declares, that notwith¬ standing all that has been said concerning the pores in the sides of the cells, his observations, made with the greatest care and exactness, have not enabled him to discover the slightest trace of them. The sides of the cells, he adds, are always formed by a membrane extremely thin, but al¬ together smooth and uniform; and the cells themselves have never an open communication with each other. (Kieser sur V Organisat. des Plantes, p. 94.) If, then, no pores exist in the sides of the cells for the reception and transmission of the fluids they contain, some other means must be provided for the accomplish¬ ment of these objects. M. Link, accordingly, supposes the juices to pass from one cell to another by transudation. M. Rudolphi thinks that a decomposition of the fluid is effected by the cells themselves, during which it is trans¬ mitted through their sides; and M. Dutrochet calls in the aid of electrical agency. To us there occur no pro¬ bable means of accomplishing these operations, consistent¬ ly with the integrity of the cellular structure, but the exercise of those alternate functions of secretion and ab¬ sorption which, from so many other considerations, we have supposed to be carried on in every living part of the vegetable system. Another question of importance in relation to the sides of the cells is, whether they are single or double; that is, whether each cell has a side of its own, or whether one side is in every position common to two cells. Mirbel asserts the former, and Kieser maintains the latter opi¬ nion. In fig. 29, Plate XXXVIII., is an outline representa¬ tion of these double sides as given by Kieser. From the extreme thinness of the membrane, it is very difficult, he says, to distinguish this double structure; but where the cells are large, and a glass that magnifies highly is em¬ ployed, each partition that separates two cells is distinctly seen to be composed of two membranes, which are some¬ times separated about the middle of the partition, and united towards the angular points. {Mem. sur l Organi¬ sat. des Plantes, p. 91.) MM. Amici, Dutrochet, De Candolle, and others, support this opinion. (See Organog. Veg. tome i. p. 21.) The existence of this double struc¬ ture receives some countenance from the fact observed in the construction of the honeycomb by the late Dr Bar¬ clay, who says that each side of every cell in the comb is composed of two plates, or is double. ( Wernerian Trans¬ actions, vol. ii.) It may still, however, be more properly said, that each side of every cell is truly single, and is rendered double only by coming into contact with the corresponding side of an adjacent cell. ■ When the cells have a regular hexagonal figure, and are equally distended with their appropriate juices, there is no reason to suppose that any vacuities are left be¬ tween their sides or angles. Mathematicians have long since demonstrated a regular hexagon to be one of those figures that completely fill up a given space; and that no vacuities can exist either about its sides or its angles. Where, however, the cells deviate from this regular figure, and more or less approach to a spherical form, va¬ cuities or interstices may readily be conceived to occur. These vacuities are said to have been first noticed by Grew and Leeuwenhoeck, and afterwards by M. Trevira¬ nus, who describes them as interstices left between the cells in their mutual approximations towards each other. He gave them the name of intercellular canals, as already stated. Gn the other hand, MM. Mirbel and Rudolphi alto¬ gether deny their existence ; but M. Kieser contends stre-Elementary nuously for it. He describes them as small interstices Organs, situated at the angles of the hexagonal cells, and formed not by any sides of their own, but by the mutual approach of three contiguous cells, and possessing, therefore, a pris¬ matic form. These interstices he conceives to exist at every angle, and thus every cell to be surrounded by them. In fig. 29, Plate XXXVIII., the black angular points denote their place. By their conjunction with each other they form a canal, which, when the hexagonal figure is perfect, and the cells are ranged horizontally, extends both in a longitudinal and transverse direction; and when the cells are placed obliquely, the canals have a similar direction. Their size varies according to that of the cells, by the sides of which they are constructed; they contain and convey the proper juices in the bark, but in the pith are often dry; and their course is said to terminate only with that of the cells themselves, at the surface of the plant. {Mem. sur V Organisat. des Plantes, p. 104.) Such are the organs which, as we have seen, M. Kieser considers to convey both the sap and the pro¬ per juices in plants. That in some circumstances they may exist, and become reservoirs of the sap or other juices, seems highly probable; but of the impropriety of ascribing to such casual productions the performance of the primary functions of the vegetable system, we have already spoken. The cellular tissue, as described above, is that form Changes of of it which must be regarded as the most perfect. From f°rm in the various causes, however, it is subject to great alterations.ce^s‘ In herbs, and in the pith and succulent parts of trees, the cells preserve their original form and appearance for a considerable time; but by the growth of the other parts, and consequent extension and compression they expe¬ rience, they acquire in the bark and wood an elongated figure, and this both in a transverse and longitudinal di¬ rection. In the latter case they surround and connect the layers of vessels with each other, constituting what has been named the parenchyma of the bark and wood. In this form their size is often greatly reduced, their ca¬ vities sometimes obliterated, and their cellular character altogether effaced. In other instances traces of a cellular structure are occasionally visible, appearing in detached portions among the perpendicular vessels. In some plants the cells become elongated in a longitudinal direction, and yet preserve their capacity nearly unchanged. It was probably this form of cell that led Malpighi to regard sometimes as a vessel, what, in reality, appears to be only a series of elongated cells. To his representation of these cells, as given fig. 18, Plate XXXIX., we have before refer¬ red. M. Kieser considers these elongated cells as forming the ligneous fibres of the wood, and that the bark is almost entirely formed of them. {Mem. sur V Organisat. des Plantes, p. 99, 101.) These cells, he says, were originally round; but in trees they become so much lengthened as to exhi¬ bit the form of a tube. It is easy, however, he adds, to detect transverse partitions in these seeming tubes, which, having often a diagonal direction, give to these cells the , form of a double pyramid with sharp points. The mem¬ brane which forms these cells is, in all trees except the ConifercB and some others, smooth, without pores, and transparent. {Ibid. p. 299.) To these organs Treviranus gave the name of fibrous utricles; by De Candolle they are called tubulated cells; and Dutrochet, from their spindle shape, denominates them, as we have seen, clostres. His representation of them, fig. 17, Plate XXXIX., cor¬ responds with the foregoing description of Kieser. From suffering compression in a transverse direction, the cells have frequently their longer diameter thrown 60 ANATOMY, VEGETABLE. Elementary into that position, and thus extend from the centre to Organs, the circumference of the plant. This position, as will af- terwards be shown, was fully noticed by Grew and Mal¬ pighi. Leeuwenhoeck also observed it, but mistook the cells thus elongated for vessels, and considered their par¬ titions as valves,—errors which M. Kieser, as well as others, duly points out. In this transverse direction the tissue forms partitions more or less large between the ves¬ sels, as will afterwards be shown; and by the obliteration of its cells it is frequently reduced to the condition of a solid membrane. Ituptures Besides these more constant and necessary changes of the cells. in the figure and character of the cellular tissue, it often suffers others of a more casual and accidental nature. In the pith, as the plant grows up, divers ruptures, says Grew, occur, oftentimes very regularly, and observed constantly in the same species of plant. These ruptures are some¬ times prolonged, so as to form a tube of considerable length. {Anatomy of Plants, p. 120.) Others have ob¬ served similar canals in the pith, formed not by sides of their own, but by those of the adjacent cells, and very various in size and form. They have been called lacunce, or reservoirs, contain a variety of substances, and Cells con- sometimes, especially in aquatic plants, only air. As we tain vesi- have seen the cavities of the larger spiral vessels to be filled with vesicles, so the larger cells of the pith, accord¬ ing to Grew, frequently contain smaller ones, or are di¬ vided by cross membranes. A similar observation is made by Kieser, who likewise remarks that, in the empty cells of Calla, JEthiopica, he has sometimes seen small round- headed bodies, supported on little peduncles, which spring from the sides, and point towards the centre of the cells. Small crystallized bodies are also occasionally found in the cavities of the cells, and within the intercellular ca¬ nals. {Mem. sur T Organisat. des Plantes, p. 94.) Of those changes in the character of the cellular tissue, by which its cells are converted into receptacles and reser¬ voirs of the proper juices of the plant, we before dis¬ coursed when treating of the proper vessels. To such an extent does this change sometimes proceed, that, in aged oaks, and, according to Kieser, in guaiacum, and probably in many other plants, the whole cellular tissue becomes filled with these secreted matters, and the distinctive cha¬ racters of the cells, and almost of the vessels themselves, are obliterated and lost. cles. CHAP. II. THE ANATOMY IN GENERAL OF THE COMMON TEXTURES OF VEGETABLES. Preliminary Observations. Nature of The elementary organs, whose description has so long the com- occupied our attention, form, either singly or by their tures eX" combination, all the other parts of plants. Some of the lower tribes of vegetables consist entirely of cellular tis¬ sue, in which no vessels are at any period to be seen ; and, even in the higher orders, many parts exhibit no ap¬ pearance of a vascular structure. There can be little doubt, however, of the existence of such a structure, since, physiologically speaking, we can form no just con¬ ception of the growth of an organized body, without asso¬ ciating with it the existence of a vascular system. In all plants the pith consists of a cellular tissue alone. In her¬ baceous plants this tissue forms their greater portion; but in trees the number of vessels is so great as to constitute the chief bulk of the plant. To certain forms of these elementary organs, whether existing singly or in combina¬ tion, we have given the name of common textures, because Comma they are very generally to be found in all plants, and in Text**, almost all parts of them, howsoever varied in quantity,v proportion, and arrangement. These textures are fami¬ liarly known under the names of cuticle or skin, of bark, of wood, and of pith; to which may be assigned the ge¬ neral appellations of the cuticular, the cortical, the lig¬ neous, and medullary textures. All the several textures just enumerated are readilyTheirp, distinguished by their different places and characters don var;, in the section of most arborescent plants, in which theyous* commonly appear well defined, and perfectly distinct from each other. In many plants, however, both herbs and trees, this distinction of parts is not preserved; but, with the exception of the cuticle, all the other textures are blended together through the entire substance of the plant, as was long since noticed both by Malpighi and Grew. “ In the stalk of maize or Indian wheat,” says Grew, “ the work of nature appears less diversified; in which, although there are the same parenchymous and lig- r neons parts as in all other plants, yet is there neither bark nor pith, the vessels being dispersed and mixed with the parenchyma, from the circumference to the centre of the stalk.” “ The like structure,” he adds, “ may also be seen in the sugar-cane and some other plants.” {Anat. of Plants, p. 104.) Similar observations were made by Mal¬ pighi, not only on different species of wheat and sugar¬ cane, but on ferns and palms. “ In ferns,” says he, “ the vascular fasciculi are numerous, but placed without order, and are everywhere sustained by the intervening cellular tissue, the cells of which are sometimes much smaller than the orifices of the vessels themselves.” Anat. Plantar. p. 24, 25.) This structure is represented in the transverse section of the sugar-cane, Plate XXXIX. fig. 25; and in a similar section of the palm, fig. 26 of the same plate. This variety of structure, thus clearly described, and Theory I distinctly delineated in the works of Malpighi and Grew, Desfon. has likewise been noticed by M. Desfontaines. ye_ tames; getables, according to him, may be distinguished into two divisions: Is#, Those which have no distinct concen¬ tric layers, whose solidity decreases from the circumfe¬ rence towards the centre, and whose pith is interposed among the vessels, and does not extend in divergent rays: 2d, Vegetables which have distinct concentric layers, whose solidity decreases from the centre towards the cir¬ cumference, and whose pith is contained in a longitudinal canal, and extends in divergent rays. The former struc¬ ture he considers as peculiar to plants whose seeds are monocotyledonous, and the latter as belonging to those which have dicotyledonous seeds. {Mem. de Vlnstitut. Nat. tome i. p. 478.) This opinion, though partly correct, is not universallyimperfei- applicable. That many plants which spring from mo¬ nocotyledonous seeds are destitute of concentric layers, and have no distinct bark or pith, is certain; but it is not less certain that many herbaceous plants, which are pro¬ duced from dicotyledonous seeds, are pretty much in the same condition, being equally destitute of concentric ' layers and of divergent rays, and in which the bark and the pith must be regarded as one continuous structure. On the other hand, some monocotyledonous plants, as M. Desfontaines admits, may deviate a little from the pre¬ scribed conditions. In a paper on the organization of such plants, M. Mirbel, who regards this doctrine as the most important step made of late years in Vegetable Anatomy, says, nevertheless, it would be erroneous to assert that they have never a bark. In several species of plants he produces examples to the contrary; and adds, that in some instances their diametral growth goes on at the cir- ANATOMY, VEGETABLE. 61 yornron cumference, which would seem to approximate them to I exta:9. dicotyledons. As, however, there is no appearance of di- /v«-Vw/vergent rayS? or 0f concentric layers, these examples are c considered by him rather to confirm than overturn the 1 theory of M. Desfontaines. ( Annales du Mus. d’Hist. Nat. I tome xiii. p. 67.) But if in this theory its second divi- s sion embrace only those plants in which the concentric ' layers are perfect, and divergent rays exist, then it ex- ! eludes a great number of herbaceous plants, whose seeds have two cotyledons; and if the absence of these regular layers, and of divergent rays, serve as a passport to the [ first division, then many of these same plants must be a admitted among those whose seeds have but one cotyle- ^ don. The theory of M. Desfontaines, therefore, rests on II too partial an observation of the structure of plants. ie P51- P r a P n e f t ■scr|. in. t s. St SI ai ei C( C( fli C( rr 0 a li w tl U| I p Cl d sselfin 0“ p4- l ! r ! 1 n h i w oi in st w se Section I. Of the Pith or Medullary Texture. The pith (medulla) of plants, when present, occu¬ pies the centre of the stem, where it is commonly sur¬ rounded by a circle of vessels which construct for it an appropriate canal. In the succulent shoots of trees, its proportion to the other parts is generally large ; but it di¬ minishes as the tree advances in age, and is frequently entirely obliterated. Where the vessels of the wood are few in number, as in herbs, only a few fasciculi are seen to surround the pith, and the intervening spaces are oc¬ cupied by a boundary of thickened cellular tissue. In some plants, again, no pith whatever exists, but the stem is hollow or tubular. In other instances, and especially in roots, the centre of the stem is occupied by vessels; and in others, both cells and vessels, promiscuously blend¬ ed together, constitute the centre of the stem. In those plants where the pith is present, and pos¬ sesses its most perfect form, it is seen to be composed entirely of cellular tissue, possessing often very different shades of colour, but, in its anatomical characters, re¬ sembling exactly the description already given of that tis¬ sue. Its bulk in different plants is exceedingly different, as are also the form and size of its cells. It is frequently entirely insulated by the surrounding vessels, but is often continuous with the cellular tissue of the bark. Its cells contain, especially in the early age of the plant, aqueous fluids, which afterwards disappear, and then the cells be¬ come filled only with air. The “ proper juices” of the plant may also be sometimes detected in the cells of the pith. Of the ruptures produced in it by desiccation and other causes, we have already spoken in discoursing of the cel¬ lular tissue: they occur particularly in succulent plants, where the cells are large, and their sides thin; so that as the plant advances to maturity, the pith breaks and shrinks up, making the trunk a pipe. (Grew’s Anatomy of Plants. p. 129.) We have also noticed the fact, that, within the cavities of the larger cells of the pith, new vesicular pro¬ ductions are sometimes found. Grew speaks of the existence of vessels in the pith of certain plants, as in that of the fig and the pine; but he adds, that they are usually so postured as to form a ring about its margin. (Anatomy of Plants, p. 119.) They are doubtless to be considered as enlarged proper vessels, which made a part of the first ligneous circle, and have retained therefore nearly the situation in which they were originally formed. Hence, as he observes, they are of divers kinds, answerable to those of the bark, contain¬ ing in the fig a milky juice, and in the pine a resinous substance. Similar vessels, containing a “ proper juice,” were observed also in the pith of elder by Malpighi, who seems to regard such appearances as common where the contained juice concretes or possesses a dark colour. (Anat. Plantar, p. 4.) It is probable, however, that the organs here considered to be vessels may in some cases be cells, into which these juices have been poured; but where real vessels of this kind are found, they are not to be considered as a part of the original structure of the pith, but occurring only in consequence of the changes which the vegetable body undergoes in the progress of its growth. Common Textures. The general nature of the pith is thus clearly an- Its nature, nounced by Grew. “ Although,” says he, 11 it have a dif¬ ferent name from the parenchyma in the bark and the in¬ sertions in the wood, yet, as to its substance, it is the very same with them both ; whereof there is a double evidence, viz. their continuity and the sameness of their texture so that all these parts are “ one entire piece of work, be¬ ing only filled up in divers manners with the vessels.” (Anatomy of Plants, p. 119.) This continuity of the pith with the cellular tissue, of the bark, by means of the inser¬ tions or transverse ranges of utriculi, as he calls them, is also adduced by Malpighi as evidence of the similarity of their nature, and of the pith being, as it were, an inter¬ cepted portion of the bark (Anat. Plantar, p. 4, 30); an opinion which seems abundantly confirmed by the inter¬ mixture of the medullary and cortical textures in many plants, in which, as already remarked, the distinctive characters of bark and pith are alike lost, and the entire stem exhibits only one uniform appearance of structure. The term medulla, employed by the ancients to note Errors this texture, derived its origin, no doubt, from the re-concerning semblance which the pith in the centre of trees bore1*'’ to the marrow in the bones of animals; and as the same term, in animal anatomy, was incorrectly employed to ex¬ press alike the marrow in the bones and the nervous substance in the vertebral column, so the same latitude of signification has been extended to the vegetable system. Hence, as Malpighi remarks, the medulla in vegetables was regarded as analogous in its nature to the brain of animals,—a doctrine which even later writers have conti¬ nued to espouse. It is not our present intention to de¬ scribe the uses of the pith, but only to remove erroneous opinions concerning its nature, and restore to it that just anatomical character long since assigned it by Malpighi and Grew, and which some writers have of late put forth as a considerable novelty. Section II. Of the Wood or Ligneous Texture. Immediately surrounding and enveloping the pith is The wood, the part called the wood (lignum vel lignea portio of Mal¬ pighi). It is essentially composed of vessels and of cel¬ lular tissue, but combined in such an infinite variety of proportions, and exhibiting such a boundless diversity of forms, that it is difficult to seize even its more general features, without the risk of extending our description beyond the limits which our plan necessarily prescribes. Except in those vegetables in which no vessels have j)escrip. been hitherto demonstrated, but in which they must ne-tion. vertheless be presumed to exist, this texture may be con¬ sidered to form a part not only of every plant, but of all its organs; for into whatever part fluids are conveyed, vessels must be supposed to extend; and wherever ves¬ sels are present, cellular tissue is to be found: hence, in its distribution, it may be considered the most universal of all the textures. In trees, the vessels, as we have fre-jn 0rdi- quently remarked, are very numerous, and, when viewed nary trees, in a transverse section, are seen to be disposed in layers or concentric circles around the axis, and to stand also iu 62 ANATOMY, VEGETABLE. Common lines or radii, diverging from the centre of the tree. (See preserve always the same position, the fasciculi being Com, < Textures, fig. 4 Plate XL.) Between each line or ray of vessels a placed at the same relative distance fiom each other and exturq "■^~'tlhn partition of cellular tissue is interposed, which ex- from the common centre of the pith. Sometimes, insteadH tends in the direction of the ray, through the entire sub- of a few solitary fasciculi, they consist or several ranges, stance of the wood. At certain distances, varying in dif- forming an imperfect sort of concentric layers ; and in such ferent trees, thicker transverse portions of the same sub- examples the ligneous texture is commonly separated by stance are placed, and are readily distinguishable in almost distinct but irregular marks from the two other textures, every species of wood. Between each layer that is an- In these plants the cellular tissue preserves its characters, nually added to the wood, and each of the smaller layers and exhibits no appearance of divergent rays, that cro to the formation of the larger one, cellular tissue The three modes of arrangement above described Causey seems also in some trees to be longitudinally interposed ; appear to constitute the chief varieties of structure in the^s varie, so that it is probable that in both directions each fasci- ligneous texture ; but in each variety, and through every cuius of vessels is intercepted by cellular tissue, and that stage by which they graduate into one another, the great- in such trees no two fasciculi are on any side in immediate est diversity of forms prevails. Each species of plant has contact with each other. It is even probable that the its peculiar internal structure, as well as its external form; individual vessels which contribute to form the fasciculi and this seems to be in a great measure determined by are themselves connected by intervening cellular tissue, the number of vessels that enter into its composition, and which acts like the neurilema that holds together the fila- their peculiar mode of arrangement. If the vessels are ments of the fasciculi in the nerves, or the cellular sub- few, the cellular tissue is large in proportion, and its stance that connects the primary filaments in the muscular characters are distinct and well preserved; if they are fibres of animals. In this manner, the whole vascular numerous, and disposed in rays, the tissue is compressed system of the plant is everywhere connected and held in various directions, loses more or less completely its cel- to^ether by cellular tissue. Of this tissue, and the dif- lular character, and forms alike those divergent rays or ferent figures its cells acquire, from the different modes transverse partitions already so often noticed, and those and degrees of compression to which they are exposed, thin membranous expansions or fascia; which, both in the we have already spoken. bark and wood, are seen sometimes to cover the vessels In palms. In many trees, however, as palms, the vascular fas- in a longitudinal direction. Though the whole of the ciculi, though numerous, are much less abundant than ligneous texture is thus made up of vessels and cells, van- in the examples just referred to. They are consequently ously formed and blended together, yet many wiiters placed at a greater distance from each other, and, not speak of ligneous fibres, which some describe as consist- being disposed in regular lines, do not constitute that ing of vessels, others of cells, and others of an assem- jd radiant appearance so common in ordinary trees, but are blage both of cells and vessels. _ U promiscuously dispersed through the cellular tissue. (See The manner, too, or rather the place, m which the fio-. 26, Plate XXXIX.) As this tissue itself is not, from the vessels are developed, in perennial plants, will greatly same causes, compressed either in the direction or to the contribute to vary the appearance of the ligneous texture, extent before described, the smaller membranous parti- In those trees whose diameter is annually increased by tions that divide the vascular radii from each other are the formation of new vessels around the cylinder of older not produced; neither, for similar reasons, is there any wood, the new parts must necessarily present in then- distinct appearance of the larger partitions that, at certain longitudinal section the appearance of annual layers su- distances, intersect the diameter of other trees. The cel- perimposed on one another, and, in their transverse sec- lular tissue, therefore, in such plants, retains more of its tion, that of concentric circles; but in palms and similar primitive character, and appears everywhere to surround trees, where the development of new parts seems to be the vascular fasciculi, but nowhere to be so compressed accomplished in a different manner, their appearance, as to form solid partitions between them. In some plants under similar sections, may be expected to be different, which possess this structure, as the sugar-cane (fig. 25, In a longitudinal section of the palm, says M. Desfon- Strut Plate XXXIX.), the cells indeed retain their perfect forms ; taines, we discover an assemblage of large ligneous fibres ot pal and even the fasciculi of vessels, though standing at consi- (that is, vascular fasciculi), solid, smooth, and flexible; derable distances from each other, have towards the centre and these are composed of others still smaller, which are of the plant a symmetrical arrangement. This latter cir- firmly united together: they mostly run parallel to the axis cumstance is observable in many other plants, which have of the trunk, from the base to the summit, without inter¬ even fewer vessels than the sugar-cane; so that it is pro- ruption ; but some proceed obliquely, and cross the others fiable that, in the first instance, it takes place in all; and at angles more or less sharp. (See fig. 2|, Plate XXXIX.) that the irregular position of the vessels in the palm and In a transverse section of the same stem, continues the similar trees, particularly towards their circumference, author, we remark neither concentric circles nor trans¬ proceeds from the peculiar modes of their growth, and are verse partitions; but the fasciculi of vessels, placed without not a primary condition of their structure. This inter- order by the side of each other, are enveloped by the cel- mixture of the vessels and cells in the plants now under lular tissue, which fills up all the intervals : they sensibly consideration extends from the circumference to the approach each other, harden and diminish in size in pro¬ centre, so as to constitute their entire bulk, to the exclu- ceeding from the centre to the circumference (see fig. 26, sion of bark and pith ; unless we choose rather to say Plate XXXIX.) ; so that the stem has much more strength that in such plants the medullary, ligneous, and cortical and solidity near the surface than in the interior,—an or- textures, are all blended together. ganization quite distinct from that of ordinary trees. In herbs. In other examples the vessels form a still smaller por- The cause of this diversity of structure seems to Thei tion of the ligneous texture, consisting only of a few be amply accounted for by the different modes in whichtfro'v fasciculi, which stand at considerable distances from each the growth of these trees is accomplished. When the other, the intervening spaces being occupied by cellular seed of a palm is sown, the leaves, says M. Desfontaines, tissue which forms the chief bulk of the plant. (Fig. 15, successively develope and augment in number for four or Plate XXXIX.) Though few in number, the vessels, how- five years ; the neck of the root augments in the same ever, are symmetrically disposed, and in the same species proportion ; the bulbous part, formed by the re-union of r ANATOMY, VEGETABLE. -onaon the petioles of the leaves, increases insensibly; its soli- Fevires. dity augments, and at length the stem rises above the ^ earth with all the size it ever acquires. Its figure is cy¬ lindrical from the base to the summit; and if the diame¬ ter be measured at different epochs, it will be seen, as Ksempfer had already remarked, not to increase. The palm, therefore, is a regular column, whose summit is crowned with leaves, disposed above each other circular- 1 ]y: those which grow in spring shoot always from the top; the older ones, placed below, dry, and when they fall leave circular impressions, which furrow the surface of the stem, and mark its years until it has ceased to grow. If next we examine the interior, we discover, as M. Desfontaines thinks, the true reason why the stem rises in a column, and does not, like other trees, yearly aug¬ ment in size. This was done by M. Daubenton, who states that every leaf of the date-palm, in proceeding from the bud, is formed by a prolongation of the vascu¬ lar fasciculi and cellular tissue which exist in the trunk of the tree, as is apparent in the petiole of the recent leaves, and of the dried ones that adhere to the trunk. The elongation of the trunk is produced, therefore, by the leaves which annually proceed from it; and as the parts which form these leaves spring from the centre, they always, as they shoot, force the older leaves outwards. Hence, therefore, as the augmentation of these trees ori¬ ginates at the centre, all the parts capable of displace¬ ment must be pushed outwards, just as the new layers of bark and wood, formed annually in ordinary trees, force outward the older layers of bark exterior to them, ffe ii of In these latter trees, continues M. Daubenton, the re- ■oHti on cession of the bark has no limits so long as new parts rutare* continue to be formed beneath it, because the new cor¬ tical layers are flexible, and the older ones readily break and give way; but in the palm the substance of the trunk has more compactness as we approach the circum¬ ference, and, at a certain point of density, it no longer yields to the central force of the interior parts; so that when this point is reached, no further enlargement takes place; and hence the date-palm scarcely exceeds ten inches diameter. It is for similar reasons that the trunk of the date-palm is of the same size through its entire length; for, in proportion as the tree rises, the exterior parts of the trunk lose successively their flexibility; and when they have acquired a certain degree of density they no longer yield to the force from within; and as this is equally the case in all parts, the trunk is necessarily of the same size throughout. (Mem. de VInstit. Nat. tome i. p. 482, &c.) It is further evident that, in this mode of growth, no appearance of concentric circles, similar to those of ordi¬ nary trees, can have place; for, by the growth at the centre, the exterior vessels are continually displaced from their original positions, are more and more compressed as they are forced towards the circumference, and pre- Isent in their transverse section that irregular distribution which they have been described to possess. Hence the cylindrical figure and the absence of concentric layers are as necessary consequences of the mode of growth in these trees, as the presence of those layers and the conical figure are of the mode of growth in ordinary trees. The greater solidity of the parts at the circumference is clearly to be ascribed to the same cause; and even the want of regular transverse partitions must in part also have a similar origin, and be ascribed, perhaps, in part to the smaller number of vessels which these plants possess, as well as to their irregular distribution, from this difference in their mode of growth, these two great divisions have received different names: those which, like our ordinary trees, augment in size by the de- Common position of new matter near the surface, are named Ex- Textures, ogenous ; and those which, like palms, grow' from within, Endogenous. Section III. Of the Bark or Cortical Texture. This texture, in its component parts, resembles that of The bark, the wood, being made up, like it, of vessels and cellular tissue intimately connected with each other. Its structure, as a distinct texture, is best characterized in the bark of in trees, ordinary trees, as it is there separated, in a great measure, from the ligneous texture. As in these plants a new layer of vessels is annually made to the wood, so a simi¬ lar but much thinner layer is yearly added to the bark, to which the name of liber has commonly been applied. These vessels are at first straight, and run parallel to the axis of the trunk ; but, by the successive formation of new layers beneath them, they are gradually forced outward, become separated more and more from each other, and, touching in a few points only, exhibit at length a reticu¬ lated figure (see fig. 24, Plate XXXVIIL), the meshes of which yearly augment in size, from the greater space over which they are continually spread. Between the vessels thus annually formed, a consider-is united able portion of cellular tissue is interposed, which, in the to the young and succulent state of parts, contributes chiefly towo°d- the thickness of the bark. This tissue is variously com¬ pressed by the vessels, so as to form transverse partitions between them, which, in the vine, the oak, and many other trees, as both Grew and Malpighi remarked, are seen to be continuous with those of the wood ; and in this way the two textures are united together. In the ex¬ pressive language of Grew, the bark, therefore, does not “ merely surround the wood as a scabbard does a sword, or a glove the hand, but is truly continuous with it, as the skin of the body is with the flesh.” In the willow and other trees, when full of sap, the bark is so easily separat¬ ed that it seems to have no connection with the wood; but this is supposed by Grew to arise merely from the ex¬ treme fineness and tenderness of the vessels that are an¬ nually formed in that part, and which on that account oppose no obstacle to the separation. (Anat. of Plants, p. 129.) It is probable, however, that the cellular tissue forms the only direct connection between the cortical and ligneous textures; and that, if a vascular communication exist, it is only, as in all other cases, through the medium of that tissue. Besides the transverse compression which the cellular Altered by tissue experiences from the vessels, it is compressed in pressure, the opposite direction by the formation of the new layers of bark and wood beneath the older bark. These in the progress of their growth exert an expansive force out¬ wards, so that the cells of the tissue are made to assume an oblong or flattened form in the direction of the vessels of the trunk, or sometimes to form a thin fascia upon the vessels, in which the cellular character is nearly or entire¬ ly obliterated. It is by the continued exertion of this force acting on the exterior and desiccated layers that these latter ultimately crack, producing figures of differ¬ ent sizes, which have frequently the shape of rhombs, the fissures of which represent, according to Grew, the posi¬ tion and track of the vessels in their reticulations. (Anat. of Plants, p. 129.) The spaces or meshes formed by these reticulations are always filled up by cellular tissue, which, in the opinion of Malpighi, originates from the vessels themselves. Both in the vessels and cells of this texture, collections 64 ANATOMY, VEGETABLE Common Textures. Contains proper juices. of the “ proper juices” frequently occur; especially in plants t]^jer^ Xe - U in which these juices are of a viscid nature, and disposec these several textures, as they exist in leaves^k^'\ e! The bark in herbs; of wood, and even in the pith of certain trees, it is only in plants in which the “ proper juices” are coloured, or dis¬ posed to concrete, that this intermixture of the cortical and ligneous textures has been traced through the whole substance of the tree; but it is probable, from the con¬ junction of the vessels of the bark and wood at the period of their formation, that it is common to other plants, the nature and properties of whose juices afford no clew to its detection. , . . ^ In most herbaceous plants, the cortical texture is not so clearly distinguished from the ligneous, since in them the greatest variety obtains both in regard to the number and relative position of the sap and “ proper ves¬ sels and very frequently the cellular tissue is quite con¬ tinuous and of uniform appearance through the entire substance of the plant. In general, however, the sap-ves¬ sels occupy the inner place, and are surrounded by the “ proper vessels,” disposed either in rings or distinct fasci¬ culi, more or fewer in number. Sometimes the sap-vessels seem to be placed exterior to the others ; and hence it is difficult to discover the true place of the “ proper vessels Section IV. Of the Connection subsisting between the Vessels and Cells J in the several Textures. in palms; in roots and branches; In our discussion of these several textures, we have Mode d noticed only in a general way the direct means by connect which the vessels and cells that construct them are con-otvj nected with each other ; but, when treating of the sap- vessels, of the absorbent vessels, and of the cellular tissue, we endeavoured to show that an universal communica¬ tion obtains between these elementary organs, and conse- quently inferred that some mode of connection, by which it can be accomplished, must have place. Grew consider-Opinioc ed the vascular and cellular parts to be connected with of Gm each other, not only by the transverse partitions of cellu¬ lar substance that intercept the vessels, but “ per mini- mas partes organicas; that is to say, the paienchymous fibres are wrapped round about the vessels, or at least in¬ terwoven with them, and with every fibre of every vessel, as in very white ash or fir wood may be observed. {^Anat. difficult boundaries of the cortical and^igne- Ct1^ph^:StrS=: of “ proper vessels” cells of the pith of elder (samhucus), ^ of some other with those that carry sap seems to be general through plants, the vessels are very abundant, and in every case thfwhole plant and consequently no distinction can be are probably derived from the straight ligneous fibres made be^veeiTithe^gneous and corfical textures. In such (vessels) both of the bark and wood. P* 29*> n- nlants the “ proper juices” must be considered to exist in deed he held it probable that the nutrient fluu s m g every part • and accordingly Malpighi, as we before re- through the vessels were in all parts poured into the cells, marked points out a vas proprium, or “ proper vessel,” as and there undergoing a certain preparation, were after- accompanying every fascicukis of vessels in different spe- wards mixed with more recent juices, and with them take •p. J wheaX (Anat Plantar, p. 24.) From the simi- up and applied to the support of the young buds a Sv of slue Jrf to pita and especially from their leaves, p. 30.) This doctrine has smee been hold y •. . . i-..i i ^ ]-jy Darwin and Knight, and it necessarily supposes a vas cular connection between the vessels and the cells, by which the functions both of secretion and absorption can be performed. The microscopical observations of Leeu- wenhoeck, already noticed, supply further evidence in sup¬ port of this opinion. In the hypothesis of Mirbel both cells and vessels are of considered as formed out of one and the same membrane. racter and appearance which uiey a, ... He rejects, therefore, the aid of all intermediate orga™* as , simple as indleir more complex forms, and as they exist necessary to connect them together, and supposes a c0m. either separately or variously intermingled together. Our mumcation to be everywhere maintained betwee descriptbns have been confined entirely to the trunk or vessels and cells, by the medium of pores in their sides, stem-but, with slight variation, they are applicable equal- As, however, these pores are nowhere proved to have iv to the root and branch, in which a similar combination existence but in the imagination of the author, we J of the elementary organs obtains. In the root, however, altogether reject their agency in maintaining a commun- they commonly exisUn a more compressed and compact- cation between the vessels and cells of plants. In the of I ed form so that the ligneous texture is seen chiefly to opinion of M. Kieser, the conjunction of the cells with t predominate, frequently to the entire exclusion of the vessels is extremely simple the sides of the cells, says he, pith and often in great part also to that of the cellular being contiguous to the sides of the vessels. {Mem. sur portion of the bark. This, however, is not universal, es- V Organisat. des Plantes, p. 94.) But mere con^guity of neciallv in annual plants, some of which, as the carrot and parts does not amount to connection, much less does it al- others,particularly in a cultivated state, are distinguished ford any information concerning the actual commun mode of growth, there can be little doubt that a similar intermixture of the two kinds of vessels prevails every¬ where in them; and with respect to these plants, what has already been said of the construction of the ligneous texture, is equally applicable to that of the bark. We have thus given a very brief and general view of the principal textures that enter into the construction of plants, and pointed out the more prominent diversities of cha¬ racter and appearance which they exhibit, as well in their ANATOMY, VEGETABLE. 65 etii ice eta e nil 1 ,ur . |l n tion that exists between these organs. In addition, there- xtu:S. fore, to connection by cellular substance, it seems abso- ■v -'lutely necessary to suppose also the existence of a vascu¬ lar structure, which shall at once serve as a medium both of connection and communication. Before concluding this branch of the subject, we may observe that the structure of the cellular tissue, and its relation to the vascular system in plants, appear, in many points, to resemble that of the adipose cells, and their relation to the vascular system of animals. These cells are described as minute close cavities, possessing no apparent communication with each other; and within them adipose matter is alternately deposited and removed. Now the deposition of this matter could only be accom¬ plished by secreting vessels which terminated in the cells, and its removal be effected by absorbents which origi¬ nated from them ; and accordingly both blood-vessels and absorbents are found to be present in this texture; but neither the secreting nor absorbing orifices have ever been actually observed. Within the cells of the cellular tissue of plants the alternate deposition and removal of various matters are not less certain ; and in the germinating seed the matter that actually existed in the cells is found af¬ terwards in the vessels. We are led, therefore, or rather we are driven, not only by the direct exclusion of all other alleged means of communication, but by a close analogy in the exercise of these animal and vegetable functions, to conclude, that secreting and absorbing vessels must be employed to deposit and remove the secreted matter from the cells of plants, in the same way as they are considered to effect similar depositions and removals of adipose mat¬ ter from the cells of animals: and as this alternate func¬ tion seems to go on in every part of the plant capable of active vegetation, it may further be inferred that a vascu¬ lar communication exists between the vessels and cells in all parts of the vegetable system. By means of this general communication between the vessels and the cells, we are enabled to assign satisfactory reasons for some puzzling phenomena, which have occur¬ red in relation to the movements of the sap. It is by this alternate action of secretion and absorption that in young plants we must suppose the cells of the pith, during the first year, to be filled with fluid, and to be rendered dry for the most part ever after. In like manner, the surface of the bark in contact with the wood appears in some trees, as the birch, to be rendered moist during the rise of the sap in spring; which led Dr Walker and others to suppose that the sap rose in part between the bark and the wood,—an opinion not at all probable in itself, and cer¬ tainly not supported by what is observed in most other trees. The fact, however, is easily explicable, on the supposition that the sap was transfused from the alburnous vessels of the wood, in the same manner as, at a later pe¬ riod, it is secreted in the same part, but in a different form, by the vessels of the bark, to form the new matter that is annually added to the tree. That the sap of plants was capable of moving in a la¬ teral direction, was inferred by Malpighi, from the fact that parts lived and grew when the perpendicular vessels that supplied them with nutriment had been destroyed. (Anat. Plantar, p. 13.) The experiments of Hales afford more decisive evidence regarding this lateral movement of the sap. He cut two large gaps in the opposite sides of an oak-branch, at four inches distance from each other, carrying the incisions down to the pith: the branch nevertheless absorbed and perspired water, but only in half the quantity that another similar but uncut branch did. In a branch of cherry-tree he made four similar cuts down to the pith, at four inches distance from each VOL. m. eg.:ir ve-i ntilf Silt of other, and opposed to the four points of the compass : the Common branch notwithstanding absorbed, in forty-eight hours, Textures, twenty-four ounces of water. ( Veg. Statics, p. 128, edit.) And when similar incisions were made on branch¬ es while still attached to the tree, their leaves continued green nearly as long as those of other branches in a na¬ tural state ; whence he justly inferred that, at these gaps made in the branch, a lateral movement of the sap must have taken place. Experiments of a similar nature have been made, and like results obtained, by Mr Knight (Phil. Trans. 1808); so that it seems clear that in certain cir¬ cumstances a lateral movement of the sap must have place. In what manner, then, must we suppose this movement How ef« to be accomplished ? Grew supposed the cellular tissue, fected. that stretches from the circumference to the centre of the plant, to be the organ by which such a communica¬ tion could be maintained; but the impermeability of this tissue to fluids opposes such an opinion. Malpighi thought the lateral communication to be made by an anastomosis of vessels; but in the vessels of plants no such mode of communication appears to exist. From the ascent of the sap in branches in which the vessels had been thus pre¬ viously cut through, Mr Knight infers that this fluid does not rise in the vessels at all, but is conveyed through the cellular tissue. This opinion necessarily implies the per¬ meability of this tissue by fluids, which, as we have shown, is contradicted by direct experiment, as well as by micro¬ scopical observation. Since, therefore, this lateral move¬ ment of the sap cannot be accomplished, either by simple percolation through the cells or vessels, or by direct anas¬ tomosis of the vessels with one another, no other known means of effecting it remain, but those of alternate depo¬ sition and absorption by the vessels into and from the cells. And if, as we have seen, the sap-vessels of plants deposit coloured fluids in the cells, which the capillary absorbents of parasitic plants are able to take up, there seems no reason for denying to the vascular productions, which have been supposed everywhere to spring from the perpendicular vessels, a like capacity of absorbing fluids from the adjacent cells. These fluids must, however, in all cases, have been deposited before they could be ab¬ sorbed ; and, by the alternate exercise of these functions, there is no difficulty in conceiving how a lateral move¬ ment of the sap might be accomplished in parts where, by the incision of the vessels, a stop was necessarily put to its perpendicular ascent. Section Y. Of the Skin or Cuticular Texture, and its Appendages. Art. I.—Description and Structure of the Skin. The skin, rind, cuticle, or epidermis, as it has been va-The skin, riously named, is the last of the common textui’es that remains to be described. It is the general envelope which invests all parts of the plant and all its productions, being equally common to the trunk and branches, the root, the leaves, the flowers, and the fruit; but in these different parts, and even in similar parts of different plants, it ex¬ hibits the greatest diversity of appearance and form. In herbaceous plants, and in the young shoots of those Descrip- which are arborescent, it resembles a thin membrane, but tion. is generally thicker on the stem than on the roots or leaves. In some leaves, however, it is thick and dense, in young as is the case also in several fruits, and is thereby fitted trees, to resist the effects of too rapid desiccation. On the upper surface of some leaves, on many fruits, and on roots, it is an entire membrane, destitute of any apertures I 66 In old trees. Common or pores; but on many stems on the under surfaces of Textures, leaves, and sometimes on the upper, it is frequently fur- -^v***^ nished with numerous pores, often visible to the naked eye, and with other luminous points of smaller dimensions, which Du Hamel also regards as apertures. It is readily separable from the bark in recent and succulent paits, 01 after maceration in water; and in certain leaves it is very completely separated by a species of caterpillar, named by Reaumur the miner. It appears then to be a thin transparent membrane, often destitute of colour, and deriving, therefore, its appearance from the colour of the parts beneath; but both in leaves and flowers it is often itself coloured. It is frequently seen to extend in all its dimensions, in common with the parts it covers. \eiy often, too, as will be noticed hereafter, its surface is cover¬ ed with hairs; and sometimes small follicles or utricles are met with, which exercise a glandular function. ^ The characters above enumerated belong chiefly to the cuticle in its young and succulent state. In perennial plants it commonly possesses others that are quite dissimilar. It is of a different colour not only on different trees, but on different parts of the same tree. It is white and shining on the trunk of the birch, and browner on the branches; greyish on the plum-tree ; red and silvery on the cherry ; green on the young branches of the peach ; and ash-co¬ loured on the larger branches. In these and many other instances it does not, says Du Hamel, merely, participate in the colour of the body it covers, but contributes itself to give colour to the exterior bark; for when it is strip¬ ped off, the substance below has frequently a different colour. (Phys. des Arbres, tome i. p. 10.) By the gradual enlargement of the trunk it is stretched and dried, and at length loses its vitality, and, as well as the bark beneath, is variously cracked and broken. Before this happens, however, it often undergoes considerable extension in all its dimensions, enlarging in breadth, and stretching lon¬ gitudinally over the young shoots. This expanded state is particularly remarkable in certain fruits, in which, when they enlarge slowly, the cuticle is extended without rup¬ ture to a very large size ; but if the expansion be very rapid, as after considerable rains, the cuticle then gives way. In certain trees the cuticle is more susceptible of expansion than in others; and in very vigorous trees it breaks more slowly than in those whose growth is lan¬ guishing, although these latter push forward more slowly than the former. (Ibid. p. 11.) In some vigorous trees of this description it altogether resists rupture ; and in this state the tree is often said to be hide-bound or bark- bound. In most instances the cuticle, when taken from young branches, appears to consist of a single layer; but on the branches of many species, says Du Hamel, after one plate or layer has been removed, another may be seen beneath, which resembles the former in its texture, but is much thinner and more green and succulent. From the birch- tree he has removed more than six layers, very thin and very distinct from each other, and is of opinion that more might still be separated. Sometimes the original cuticle seems to be entirely thrown off, and the exterior covering is formed by a portion of the cellular tissue of the bark. Grew thinks that this substitution takes place annually, the older skin being cast off, like the skin of an adder, by the generation of a new one beneath. (Anat. of Plants, p. 114.) Du Hamel describes also the existence of small leaflets or scales, which are continually detached from the cuticle of some trees ; and these he considers to be as con¬ stantly replaced, by the formation of new ones beneath Is regene- them. rated. ■ Concerning the regeneration of the cuticle on parts from ANATOMY, VEGETABLE. which it has been removed, Du Hamel observes, that Con, when the wound is covered with waxed cloth, a new cuticle Tex| is promptly formed without any separation of a portion^, of the bark beneath. When the exterior portion of the bark is removed with the skin, the inner part of. it.is equally capable of regenerating Composed of lavers. a cuticle ; but if the wound be noTp'rotected from the" air, a certain degree of exfoliation first occurs, and, under the decayed parts, a new skin forms. Even where the bark of a cherry-tree was entirely removed from the trunk, he found that, the wood was capable of regenerating a new bark and cuticle, if the parts were properly protected from the air. This cuticle did not originate from that which remained on the roots and branches, but was reproduced in isolated por¬ tions on different parts of the trunk; it continued, how¬ ever, after the lapse of fifteen years, always different from that of the natural growth. In other instances, he adds, the cuticle does not seem to be regenerated at all. He remarks certain analogies to exist between the cuticle in some plants and in animals. In both, he adds, it seems in certain circumstances capable of great extension; in both it is easily regenerated, and that too in isolated portions, and not by continuity of organs, as is common in other instances ; and in both, lastly, it is perpetually obliterated, and continually and imperceptibly renewed. (Phys. des Arbres, tome i. p. 11.) With respect to the nature of the cuticle, very differ-Its ent opinions have been advanced, and still continue to prevail. Are we to regard it as a peculiar organ, formed immediately by the proper exercise of the vegetative functions, or is it produced in a sort of secondary manner, by some changes induced on some previously constructed organ ? Grew asserted it to be sometimes original, and in some instances produced out of the exterior layer of the cortical texture beneath it; and this first view of its ori¬ gin seems to be generally supported by the descriptive character which has been assigned to it. Its co-existence with the first traces of vegetable organization, its con¬ tinued growth and expansion, and its subsequent regene¬ ration after removal, all seem to favour its primary and in¬ dependent origin, which is also supported by investiga¬ tions into its minute structure. Mirbel, however, and some other writers, after Hill, have regarded it, in all cases, not as an original membrane, but formed by the exterior sides of the common tissue of the plant; and where there is no separation of these sides in the form of a membrane, such plants are held to be destitute of a cu¬ ticle. (Exposit. de V Organisat. Veget. p. 103.) Another question relating to this organ is, whether it It: must be considered a simple membrane of uniform struc-tu ture, or a compound of two distinct parts, like the true skin and the cuticle in animals. Grew seems to have re-0 garded it as a simple body, but constructed both of vesselsol and cells, the cells being continuous with those of the bark. (Anat. of Plants, p. 62.) Such, too, seems to have been nearly the opinion of Malpighi, who describes it as of constructed of horizontal ranges of cells, but often deli-pi neates reticulations of vessels as constituting a part of its structure. (Anat. Plantar, p. 2, 19.) In the birch, the plum, the cherry-tree, and others, Du Hamel declares thed component fibres of the cuticle to possess a directionm transverse to that of the trunk; but this is not general. In the birch-tree the fibres seemed to be placed parallel to each other, and to be connected together by lateral fibres; but he could see nothing of the vesicular structure of Malpighi and Grew, and therefore regards the struc¬ ture of this texture to lie altogether fibrous. (Phys. des Arbres, tome i. p. 8 and 9.) M. Desfontaines, on theo other hand, describes it as a membrane resembling in ap^t: 4 ANATOMY, VEGETABLE. 67 >pi on jijQion pearance a thin plate of parchment, and perforated by im- t iures. perceptible pores, which give issue to the insensible trans- ’T’^piration. Its structure he regards as unknown, but con¬ siders it capable of regeneration, (d/ewz. da l Instil, Nett, tome i. p. 4$1.) M. Kieser, who professes to have studied this texture falser, w|th great attention, adopts nearly the opinion of Grew, pronouncing the cuticle to be constructed of a very fine cellular tissue, and of extremely minute vessels which run through its whole extent. These vessels form an exceed¬ ingly delicate and subtile net-work, the meshes of which possess very different forms, and their vessels terminate at the orifice of a pore. His observations were made on the cuticle of leaves. On the inferior surface of the leaf of Amaryllis formosissima (fig. 14, A, Plate XLI.), magnified 260 times, these vascular meshes of the cuticle have an elongated hexagonal form; and four of their vessels pro¬ ceed always to terminate at the orifice of the little oblong aperture or pore situated at their junction. In Canna Indica, the vessels of the meshes on the lower part of the leaf, which thus terminate in the pores, are said to origi¬ nate from a fasciculus of the spiral vessels that ramify through the leaf, as is represented at a' in fig. 14, B, Plate XLI.; and within the areas of the larger meshes a still finer net-work of vessels is seen. On the inferior sur¬ face of the leaf of a species of fern, the vessels of the cuticle, instead of forming meshes of different figures, ex¬ hibit the appearance of sinuous lines, which run in every i direction through the cuticle. See fig. 15, Plate XLI. which represents the central part of the little adjoining , leaf, magnified 130 times. These sinuous vessels often I j°in> and, after making a half-circle, terminate by one ex- i tremity in the minute pores everywhere spread over the e leaf, and by the other in the larger vascular fasciculi that ramify through it. At the letter b', in this figure, the e hexagonal cells that construct the parenchyma of the e leaf are distinctly visible through the vascular sinuosities i- of the cuticle. It was by the examination of this leaf :■ that M. Kieser was first enabled to discover the origin and i- termination of the vessels that construct the cuticle, I- having in all his previous investigations examined the cu¬ ll tide in its separated state, after it was detached from its II connection with the other organs; but the researches e made on this leaf rendered every thing clear. (Mtm. sur d f Organisat. des Plantes, p. 141-2.) jf: (rocker The vascular net-work of the cuticle, thus described ]. withers, by M. Kieser and others, had been regarded as a decep¬ tion by M. Krocker, who considered these reticulated figures as no part of the real structure of the cuticle, but merely as the sides of the subjacent cells; in which opi¬ nion Sprengel, Link, Jurine, and Mirbel, concurred; but M. Kieser, in opposition to this opinion, maintains that, in the fern and other leaves, the real cellular structure of the parenchyma is seen entire through the vascular re¬ ticulations of the cuticle, with the meshes of which the sides of the subjacent cells do not anywhere coincide. He observes that these cells are commonly much smaller than the vascular meshes which cover them; and that the ves¬ sels of these meshes may be traced, as before remarked, to the larger fasciculi that construct the leaf. In the fern the vascular structure of the cuticle is the same on both sides of the leaf, but the superior side does not pos¬ sess pores. Saus- A very different view of the structure of the cuticle was taken by the late celebrated M. de Saussure. He regarded it not as a simple, but a compound texture, con¬ sisting of a very delicate external pellicle or membrane, beneath which was placed a net-work of very fine vessels. The external membrane he describes as perforated by ■H pores of unequal figure, between which he observed some Cominon opaque and tortuous filaments, disposed in a reticulated Textures, manner, each mesh being formed by six filaments, four ofVw^v-v^ which terminated at each pore. To this arrangement of filaments he gave the name of cortical net-work, and re¬ garded it as quite distinct from the cuticle that covered it. The meshes of this net-work differ much in size and figure in different leaves; and, when minutely examined, they are often seen to form junctions, but never to cross each other; whence he was led to regard them as vessels derived from those of the expanded petiole, and thus con¬ stituting a very fine vascular net-work. A similar struc¬ ture was observed in the petals of the flower. (Encyclop. Method, tome i. p. 67.) M. de Candolle considers the cuticle in certain parts Opinion of to be a simple and proper membrane; in other parts to be Le Can- formed by layers of cellular tissue. The cuticle of leaves, dolle. and probably of all annual shoots, appears to be formed of a layer of flattened cells, differing in form and in other conditions from those of the parenchyma of the leaf. It may be regarded in this state as a proper membrane. When stripped off and examined by the microscope, it exhibits small spaces, bounded by lines or rays, or a sort of net-work, the meshes of which have different figures in different plants. The lines, says he, appear in the form of single or double filaments, and may be considered hollow, and as forming a system of cuticular vessels which terminate in the pores. In more advanced age, and espe¬ cially in the trunks of old trees, this primitive cuticle gradually changes its character, and finally disappears. It is then succeeded by a thicker membrane, which no longer exhibits the net-like figure of the former, and ap¬ pears evidently to be formed by the exterior cells of the cellular tissue, which, by the combined effects of disten¬ tion and desiccation, assume a membranous appearance, to which the term epidermis is properly applicable. This epidermis may be considered single when composed of one layer of cells, and double, triple, or multiple, when formed of two or more successive layers. It is thus that Ulloa, in describing a tree in Peru, speaks of having de¬ tached more than 150 epidermoidal layers, when he lost all patience in counting them, seeing that he had not then reached to half the thickness of the bark. A similar ap¬ pearance is exhibited in the numerous layers that form the epidermis of the white birch, amounting sometimes to 15 or 18 in advanced age, and which ends by breaking into that cracked condition of the bark that presents only discontinuous portions of white epidermis on the remnants of its cellular envelope. At an early period the cuticle is most easily torn in a longitudinal direction, which is that of its growth; but at a later period, when the diametral growth has increased, the cells of the tissue are drawn out more in that direction, and therefore more readily break transversely than in length. (Organog. Veget. tome i. chap. 5.) Art. II.—Of the Pores of the Skin. The pores of the cuticle, called sometimes cortical or Descrip- miliary glands, sometimes exhaling pores, sometimes cor- tion of tical pores, and more lately, by Link and De Candolle, ^ stomata, were first noticed by Grew, who describes many orifices as existing on the leaves of different plants, which vary in size, number, shape, and position. In the white lily they are of an oval shape, of a white colour; and each is surrounded by a slender border. When viewed through a good glass they appear as if standing about one sixth or one eighth of an inch apart all over the leaf, but not arranged in any regular order. In the pine, also, 68 oval shape, but have no rising border, and Descrip¬ tion of pores by 3)e Can¬ dolle. Common they have an oval shape, But nave no rising - Texiures. are arranged in lines from one end of the leal to tne (Amt. ofPhnts.r-1S3.) Hedwig considered the borders mentioned by Grew as produced by a ring of one or more vessels, which terminated in the pore. I he n ber of pores he represents as exceedingly great. In the square of a line of the cuticle of a bulbous lily he reck¬ oned 577 pores. „ , „ _ofo The characters and position of these pores or stomata have been further examined by M. de Candolle on more than 600 plants. They occur most frequently on the leaves, occupying both surfaces in some herbs, and in trees chiefly the inferior surface. Stems in general have no pores, except, as in the Graminece, where they are succulent, and have the character of leaves ; or where the plant is altogether destitute of leaves, as the Cactus. Un the prominent lines or vessels of the leaves and stems no pores are to be seen, but only in the grooves or depressed surfaces of the parenchyma. They are never observed on the root, not even on bulbous roots, where the scales of the bulb are true leaves. The small leaflets called stipula} and bractece. sometimes have and sometimes have not pores. The calices of the flower in general have pores, but the petals have not. Pericarps of a foliaceous consistence have pores: when fleshy, they are destitute o them. The envelopes of the seed are destitute of pores, but they are found on all seminal leaves that rise above the ground. The lower tribes of vegetables, as the fad, musd, hepatica, fungi, &c. are destitute of pores. The occurrence of pores in those plants where they are found seems to be much influenced by external circum- the pores; stances. They are never met with but on vegetables, and those parts of vegetables that are exposed to the air; and therefore the internal surfaces of many leaves that embrace the stem are without pores, though on the ex¬ ternal surfaces of the same leaves they are abundant. No plant that is completely aquatic, nor any part of it that is habitually under water, is provided with these or¬ gans ; but the parts which rise above the water are furnish¬ ed with them. In ranunculus aquations the leaves that are constantly under water are destitute of pores, while those that float on the surface are provided with them, but only on their superior face. Even leaves which do not naturally possess pores when under water, acquire them if they are made to grow in air; and land plants, on the contrary, when made to grow under water, may, by such treatment, be deprived of their pores. Thus, the leaf of green mint, when growing in air, possesses not fewer than 1800 pores on its lower surface; but if kept for a month under water, its leaves fall, and the new ones that succeed are destitute of pores. Light seems to be necessary also to the production of res, for etiolated plants do not possess them. When anatomy, vegetable. or rhomboidal. In size they very much varied in differ- Co*J 0g ent plants, but in the same plant the size was uniform. ^ The largest pores were seen on the leaf ot the white lily,v^Y\ the smallest on that of the French bean. (Kieser s Mem. sur l' Orqanisat. des Plantes, p. 144.) Although the pores are generally dispersed over the parenchyma of leaves at nearly equal distances, yet where the vessels run parallel, they are disposed in one or more lines between them. Sometimes, again, they are collected into little clusters, as in the leaves of crassula cordata, where the roundish dots or points, visible by the naked eve are clusters of pores or stomata. This appearance Pore,, suggested to M. de Candolle the idea that the stomata orifices might be the orifices of vessels, as each of the little points768^ mentioned above is the termination of a fibre, which is itself a bundle of vessels. This idea is further strengthen¬ ed by the fact that these pores are not found in plants Influence of air on of light on pores. Research¬ es of Itu- dolphi. grown by the light of lamps, the leaves possess a few pores ; and, in all cases, the parts secluded from light and air are destitute of these organs, but acquire them if they are duly exposed. {Mem. de flnstit. Nat. tome i. p. 351.) This general account of the pores of plants is confirmed by the researches of M. Rudolphi. In most herbaceous plants he found the pores to occupy both sides of the leaf, but in trees only the inferior surface. They were not often met with on the parts of the flowers, or on fruits ; they were never seen on roots, nor on the trunks of trees; nor ever on aquatic plants, except on such parts as were raised above the water. The lower tribes of vegetables seemed to be universally destitute of them; the leaves also of those plants that were covered thickly with hairs on both sides had no pores. The form of the pores was commonly oval or elliptical, but in a few instances square that are destitute of vessels; and though he could never trace the continuity of a vessel with a pore, yet M. Com- paretti assures us he has seen the vessels terminate in them. Mirbel, on the other hand, considers the pores to be the orifices of cells; and Kieser regards them as con¬ nected with the intercellular canals. (Organog. Veget. tome i. chap. 6.) Some have considered the pores as organs, by which the resinous or waxy matter found on certain leaves is exhaled; others, as organs for the absorption and trans¬ piration of air or gas; and others, with more probability, as those by which moisture is thrown off by the leaf. In Trans; support of this last opinion, it may be stated that porestlonli: exist in all leafy parts that transpire, and are more mi-Pore!' merous in membranous leaves that transpire most, than in fleshy leaves which transpire little. 4 hey are wanting altogether in aquatic leaves and in etiolated leaves; also in fleshy fruits, in roots, and in the petals of flowers, which do not transpire in any degree analogous to leaves. In darkness too, when transpiration ceases, the pores are closed ; and they again open and transpire when light re¬ appears, and especially in bright sunshine. It is always ne¬ cessary to distinguish the evaporation, more or less great, that goes on through the tissue in all organs by day and by night, from the active transpiration which occurs in sunshine, but only in organs furnished with pores, and which seems to be executed by them. Besides transpiration, M. de Candolle believes that, inAbsoi certain cases, the pores may also serve for absorption ; but '’J P this he considers to be rarely the case, and out of the natural course of vegetation ; and that the experiments which fa¬ vour this opinion may be explained by the hygroscopic powers of the vegetable tissue. He thinks the results ob¬ tained by Bonnet, who kept leaves a long time in life by lay¬ ing their porous surfaces on water, may be explained by supposing this treatment to prevent decay by checking transpiration, rather than by promoting absorption. ( Or¬ ganog. Veget. tome i. p. 87.) This suggestion will not, however, explain the phenomena of an experiment con¬ tinued for many years in the conservatories of the Royal Botanic Garden here, under the care of Mr Macnab. He has kept a species of fig {ficus elastica) for several years fastened against the wall, with its roots, stem, and branches entirely exposed to the air; and in that situation it has not only lived, but actually grown in every part, so as to have increased greatly in size, only by being thus kept in a warm and moist atmosphere, and occasionally sprinkled with water. In addition to these visible pores, it is probable, says In'* M. de Candolle, that the surface of vegetables is furnished Pore with others that are invisible, but so small that with the strongest microscopes we are unable to recognise them: i ;i* ANATOMY, VEGETABLE. 69 mom their existence, therefore, can only be presumed from xtun« physiological phenomena. Ihus, a pair of a vegetable destitute of pores loses weight when exposed to the air, by the escape of the fluid it contained; or a portion of moss equally destitute of pores, if placed in watei, speedily acquires additional weight. Whether these results are obtained through the medium of minute pores, or of un¬ organized apertures, such as are admitted to exist between the molecules of all matters,—and whether through the same pores or apertures a passage is afforded to gases or other fluids, or to the oily or waxy secretions that cover certain surfaces,—are questions, adds M. de Candolle, to which no answer can at present be given. (Organog. Veg. tome i. p. 88.) Art. III.—Of Hairs. ;cri]i From the surface of the cuticle, in many parts of her- \ ai baceous plants, and in the succulent parts of arborescent ie.t1, ones, hairs (pili) are seen to spring. They possess very iaU" different forms, and vary likewise greatly in texture. In a strict sense, they may be defined small filaments pos¬ sessing considerable stiffness, which project from the sur¬ face, and stand out pretty erect. When they are very numerous, a little soft, and less erect, they take the name of villi; when still softer and less numerous they are termed down (pubes). Sometimes this down is composed of long hairs nearly resembling wool, at other times it approaches more to the character of cotton. When the hairs are stiff and ranged along the edge of a surface, like the lashes of the eye, they are named cilia ; and if, with these characters, they are produced to a greater length, as in the beard or awn of wheat, they acquire the name of barba or arista. Sometimes they resemble the bristles of the hog, and are then called seta:. Many other varieties are enumerated by botanists, who further distinguish them by various names, according as they terminate in a single point, or are hooked, or forked, or branched, or feathered, &c. In some instances, instead of appearing like one con¬ tinuous substance, they are composed of many joints, or are said to be articulated. In fig. 16 and 17, Plate XLL, we have copied from Du Hamel a few of the varieties, both of single and jointed hairs ; but the forms they exhibit are so numerous and diversified, that we must refer to the writers on botany for minuter information. In some ex- ® amples the point of the hair is terminated by a small rounded globulet, and sometimes by a fine filament, that seems to proceed out of the hair, ir} si- Some writers distinguish hairs into two general classes, 11 > —the glandular and lymphatic. In the former class, the hair or filament is sometimes the excretory duct of the small gland situated at its base; and in other instances, as in the chick-pea, the glandular or secreting organ is seated at the extremity of the hair. The lymphatic class embraces a much greater variety of hairs, which differ widely from each other in consistence, direction, and form. They spring only from the parts of plants which are ex¬ posed to the air, and are not found, therefore, on the parts concealed below the earth or in water: they are also rare on plants that grow in the shade, are want¬ ing altogether in etiolated plants, and are most abundant, in general, on plants that grow in warm places, and are well exposed to solar light. With respect to their seat or origin, as compared with that of the pores, although, says M. de Candolle, these two kinds of organs appear some¬ times mingled, yet each has a determined place; for the pores are seated on the cellular or parenchymatous parts, while the lymphatic hairs constantly spring from the vas¬ cular or ligneous parts. Yet the hairs, he adds, are al¬ most always placed on the same surface as the pores ; and Common hence the superior surface of the leaf, which is common- Textures, ly destitute of pores, has in general but few or no hairs. The pores, as before stated, seem to be the true organs of transpiration ; and the various kinds of hairs serve to pro¬ tect the plant against the excess of solar light, against variations of temperature, against humidity, or sometimes against insects. (Organog. Veg. tome i. chap. 10.) Hairs, also, of various kinds are met with on the different parts of the flower. These often have the colour of the pe¬ tals or other parts on which they grow, and are distin¬ guishable from the true lymphatic hairs, though some¬ times associated with them on the same organ. With regard to the structure of these minute bodies, structure, little that is satisfactory can be said. They seem to ori¬ ginate either directly from the cuticle or from the corti¬ cal texture beneath it; but not often from the ligne¬ ous texture, except in those instances where they are very long and rigid, as in the awns of wheat. Du Hamel observes that almost all of them are implanted on small bodies, similar to the bulbs which give origin to the hairs of animals. (Phys. des Arbres, tome i. p. 183.) They commonly resemble simple filaments, but often appear like elongated cells threaded on one another, and, instead of terminating in a sharp point, end in a small papilla or utricle, which yields in many instances a viscid or oily matter, or sometimes a coloured liquor, which has led many to regard them as exercising a glandular function. One species of these supposed glandular organs has been more particularly examined, and their fluid analyzed, by M. Deyeux, who gives the following account of it. Soon after the seeds of the chick-pea (cicer arietinuni) Hairs of are sown, its first leaves are seen to be covered with hairs, chick-pea. at the extremity of each of which is a transparent glo¬ bule, about the size of a small pin-head, consisting of a fluid matter. It abounds most in mid-day, when the air is warm and dry, and is scarcely perceptible at night, or when the air is cold and moist; after rain, indeed, it does not again appear for two or three days. When these fluid globules were removed in a dry day by blotting-paper, they soon re-appeared; they were acid to the taste, reddened litmus paper, and caused an effervescence in carbonate of potash when brought in contact with it. He regarded them as composed of oxalic acid, the properties of which they precisely resembled. (Mem. de Vlnstit. Nat. tome i. p. 157.) Art. IY.—Of Prickles and Thorns. It is not easy to discriminate between some of the harder Definition; species of hairs, described in the former article, and those to which the appellation of prickles (aculei) has been as¬ signed. They are defined by Du Hamel to be excres¬ cences, often hard, and always terminated by a sharp point, which are developed with the other productions of plants, but are not inclosed in particular buds; so that they may for the most part be regarded as hard and so¬ lid hairs. They spring equally from the stem, the branch-P0Sltl0n> es, the petioles of the leaves, and also from the leaves themselves in various plants; and in the chesnut and some others they are seen to cover the fruit. They are frequently straight, but in the rose and many others are curved at the point, as in fig. 18, Plate XLI.; and, accord¬ ing to Malpighi, possess sometimes in this plant a little head, which yields a viscid fluid. Regarding their structure, Grew remarked that they sf were connected only with the skin or the bark, and he therefore named them cortical, to distinguish them from thorns properly so called, such as those of the hawthorn, 70 anatomy, vegetable. (Lectures on Inflammation, p. 318.) Besides these more Co® simple structures, it is well known that most of the inter- lestjl nal viscera are likewise denominated glands, though fering in all their characters from those just mentioned. The ambiguity which thus prevails in animal anatomy,in in relation to the use of the term gland, has been increasedtable; tenfold in the applications that have been made af it toH' the organs of vegetables. It is justly observed by M.de CandoUe, in reference to this subject, that the numerous which spring from the wood, and which he denominates^ neous. These latter, he adds, always ascend whde the co - tical thorns commonly point downwards. 5 p. 33.) In proof of their origin from the bark, DuHame remarks that if, after maceration in boiling water, the ta* of such Plante be stripped off nil the prickles come away with it, and leave not the smallest impression on the wood, nor even on the more interior layers of the baik it¬ self. When a section also is made of the branch and pricklei as~in figiYs, Plate XLL, the wood y and the pith approximations r. 18,1 late AljA*» . f 1 , • A.]e, mais have often promoted our researches into the former, z are both seen to have no connec ,, sometimes led physiologists astray, and intro- f are ootn seen J ‘'V'W •“ internosed between the but have sometimes led physiologists astray, and intro, but the inner layer a- of *eba k P does not> duced int„ the language of botany many inexact expres Prickle of In animal anatomy the term gland is understood to express some organ that exercises a secretory function; but in vegetable anatomy this term has often been applied to bodies that are not known to be real secretory organs. Thus the cells of the cellular tissue, which frequently con¬ tain resinous or oily matter, have been sometimes named cellular glands ; the little globulets or utricles at the ex- tremities of the hairs on the edges of leaves, utricular I 01* i ■ v base of the prickle and the wood. The prickle does not however, spring from the skin, for it is formed of many layers like the bark. As the parts become more solid, it is less freely supplied with juice, and therefore hardens and turns brown. (Pln/s. des Arbres, tome i. p. ■ In the nettle (urtica dioied), Malpighi states, that be- the nettle. gJdg the common prickles on the leaves, there are among them others of a different description. They possess more of a lio-neous character, are hollow internally, and contain - r j -u o i p a iuicfvvh eh" when i gains admission beneath the skin, glands; the small organs formed by the pores on the leaf, excites itching and tumour. (Anat. Plantar, p. 137.) Dr cort/mZ or miliary f/W, ; certam fleshy tubercles on the Hooke had previously given a much more minute account leaves, urceolar glands ; and the little sea es that cover of the sting of this plant. Almost every part of it, says he, the fructification in ferns, scahform glands. The mtar h covered with prickles like sharp needles. Each prickle rium of the lower commonly contains a sweet juice, an consists of two parts, very different in shape and quali- is therefore deemed a gland; but Lmnaeus, with his usual ty from one another; one is shaped much like a round disregard both of the structure and function of organs, bodkin, is very hard and stiff, exceedingly transparent and considers as a nectary, not only the body which may se- clear, and hollow from top to bottom. When this bodkin crete, but any other that may serve as a receptacle of the is thrust into the skin, it does not at all bend ; but a cer- secretion ; and indeed is said to comprehend under this tain liquor is then seen to move up and down in it, rising term all those bodies which have no resemblance to the towards the top, when the point is pressed down on the other parts of the flower, in whatever variety of form they base. This base is formed by a little bag, is more pliable may appear, or whatever purpose they may serve. (Will- than the bodkin part, and within it is a cellular structure, denow’s Principles of Botany, p. 87.) in some other m- which contains a thin transparent liquor (see fig. 19, stances the term gland has been used, not to express the Plate XLI.): it is this liquor that rises in the tube, and, be- secreting organ itself, nor even the receptacle of the se- ing deposited beneath the skin after it is punctured, ex- cretion, but the solid excreted matter on the surface of cites the irritation that succeeds. (Micrographia, p. 142.) certain leaves; and others consider hairs, and every other The true thorn (spina) derives its origin, as Grew re- protuberance that projects from the surface, and contains marked, from the ligneous part of the plant, and in some a fluid different from the common sap, as entitled to the plants is produced from the degeneration of some other distinctive appellation of gland. organ, as of the leaf, or even the branch itself: we shall Amid such diversity of opinion concerning the struc-iat therefore defer the further consideration of its structure, ture, position, and function of these minute organs, and ini* until we come to treat of that of the branch. such vagueness in the methods employed to characterizeg» them, it is extremely difficult to define their true nature, Section V. or declare the principle on which this definition should „ „ . „ Tr , ,7 proceed. The mere existence of a fluid, distinct from the Of the Glands of \ egetables. common snn in anv or nan. cannot be considered as be- A common sap in any organ, Ambiguity Perhaps in the whole science of anatomy there is no stowing on it the title of gland, otherwise the greater por- of the term word that has been employed with such latitude of signi- tion of some plants would come to be regarded as glancm* ';land, lar: those varieties of structure which exercise no secre¬ tory function may also be excluded from the list of glands; and so likewise the hairs of plants, though containing pe¬ culiar fluids, may be excluded, since these peculiarities appear to arise frequently from circumstances foreign to fication, and is therefore exposed to so much ambiguity, as the term gland. In animal anatomy it was doubtless used at first to denominate certain organs, from the exter¬ nal resemblance which they bore to certain fruits or seeds; in Animal and in that sense it is still employed on several occasions. „rr ^ v _ ^ ^ Anatomy, Afterwards it was understood to signify not so much the the action of the organ itself; and even if they do not, external form as the internal organization, and was con- some specific variation of the general name they bear is sidered to express a certain structure, by which alone the preferable to the employment of so ambiguous a word as function of secretion could be exercised; but it is well gland. But where any organ is distinct from the com-^ observed by Dr Thomson, in his valuable work on Inflam- mon textures of the vegetable, and by the peculiarity oi mation, that “ the definition of a secreting glandular part its structure is fitted to produce those changes on the ve- must be taken from its function, and not from its struc- getable fluids which we name secretion, it may be deem- ture; for nothing can be more various than the internal ed a secreting organ. This secretory function, however, structure of those organs that are denominated glandular may sometimes be exercised, as in animal bodies, by roem secreting organs: they consist sometimes of convoluted branous surfaces, and sometimes by small isolated bodies, vessels, sometimes of follicles or small hollow bags, and to which, perhaps, may properly belong the denomination sometimes of transparent membranes, in which neither of glands. convoluted vessels nor mucous follicles can be perceived.” But even though this method of defining glands were Uli 1 ANATOMY, VEGETABLE. 71 i* ^aB lopted, it still is a matter of no small difficulty to distin- *2 |s. jish their species by appropriate appellations. In ani- al anatomy no settled rule obtains; but the name of the ‘ and is assigned from some accidental circumstance of tuation, figure, use, &c. In vegetable anatomy the bo- .nist, regarding glands only as aiding the discrimination ' species, refers commonly to their situation, and speaks ' foliaceous, stipular, or petiolar glands, according as ey happen to be seated on the leaves, the stipules, or the itioles. The anatomist imposes names according to their rms, as they chance most to resemble a globule, an ricle, or some other figure; and the physiologist is fiefly directed by ideas which indicate their functions, stinguishing them into mucous, oily, resinous, or necta- ferous glands, according to the nature of the fluid they rnish. Of these different modes, that which proceeds i the apparent form, where it can be discovered, seems e most precise; but as this cannot always be accom- ished, the situation of the organ, or the nature of the creted fluid, must occasionally be had recourse to. ced ! Of these bodies it is to be remarked that they differ in hy ie respect from most of the corresponding organs in ani- als, almost all of them being seated on the external parts i the plant, like several of the more simple glandular bo- seeds in which it occurs have been term¬ ed exalbuminous ; and in a few instances it seems to be entirely wanting. Its quantity, situation, and figure, in different seeds, are subject to very great variation. In the seeds of the Graminece, where the embryo acquires only a very small size, the albumen constitutes almost the en¬ tire bulk of the seed, and is placed wholly exterior to the embryo. In the Leguminosce, on the other hand, the em¬ bryo is more completely developed, and the whole of the albuminous matter is contained within the cotyledons. In beet (beta), and many others, the albumen is partly re¬ ceived into the cotyledons, and lies in part exterior to them; and where this occurs the embryo sometimes en¬ circles the albumen, and is sometimes encircled by it. In Rheum the embrjm is placed in the centre of the albu¬ men, in Rumex and some others it is applied on the side of it, in Atriplex the long cylindrical embryo sur¬ rounds the albumen, in Bberhaavia the embryo and its ISlti s 7o cotyledons cover entirely the granulated substance of the Of the Seed, albumen. (M. Jussieu, An. du Mus. d'Hist. Nat. tome p. 224.) In the onion (allium cepa) the embiyo makes several curves within the substance of the albumen, and in dodder (cuscida) it is twisted around it in a spiral form ; so that the relative positions of the embryo and al¬ bumen, as well as their quantity, proportion, and figure, are subject to endless variation. But however much in these respects the albumen may Albumen vary, it is always contained within an organized structure, of wheat. Sometimes this structure is that of the cotyledon, as al¬ ready exhibited in the bean (fig. 32, Plate XXXVIII.), the cells of which contain this albuminous matter. . Where the albumen is placed exterior to the embryo, as in the seeds of wheat, it is nevertheless contained in a cellular tissue. This is exhibited in fig. 27, Plate XXXVIII., copied from Leeuwenhoeck, in which cells of an hexagonal form are seen to be filled with the albuminous particles that con¬ stitute the white matter or flour of that seed. This mealy part of wheat he describes as consisting of minute globules, inclosed in a kind of membrane so exquisitely thin as scarcely to be observed, within which the globules are contained, as it were, in cells. The globules appeared to be of different sizes, not perfect spheres, but having an indentation on one part, which led him to suppose that they were not formed by simple accretion, but by some mode of growth, and that “ the membranes which inclose them in cells must be provided with so many veins or vessels, that every particle of meal may have its separate vessel.” He even conceived the globules themselves to be inclosed individually in a thin skin or shell; but this opinion he never brought to ocular demonstration. (Select Works by Hoole, vol. i. p. 169.) Similar observations on the albuminous part of wheat have since been given by Mirbel; and Kieser and others have delineated the glo¬ bular particles contained in the cotyledonous cells of the bean and other seeds; so that whether the albumen be situated in the cotyledons, or be placed exterior to them, it is in every case contained in a similar and distinctly organized structure. In consistence the albumen is said to be either fari-Varieties naceous, fleshy, or cartilaginous; and it may exist in°falbu- various intermediate states. The farinaceous kind ismen- readily reduced to powder, and is dissolved by water into a viscous mass. The embryo is generally placed exterior to this species of albumen, as in the Graminece. The fleshy albumen is more frequent. It is softer than the former, and dissolves by water into a gelatinous mass. It is often entirely contained within the embryo and its cotyledons, and yields the thick oil that is express¬ ed from many seeds. Lastly, the cartilaginous species has a horny consistence, is difficultly soluble in water, and not easily reduced to powder. The embryo is never placed exterior to it, and when it contains oil, this is usu¬ ally very thin. (Gaertner de Fructib. &c. vol. i. cap. 10.) In many seeds the albumen serves as a support and de- Uses of al- fence to the embryo, as well as for nutriment. If it be re- humen- moved previous to germination, as was done by Mirbel (An. du Mus. d'Hist. Nat. tome xiii. p. 157) in the seed of the onion, and by Dr Yule ( Wern. Trans, vol. i. p. 591) in different species of Graminece, the embryo, though plant¬ ed in a rich soil, and carefully tended, grows but feebly, and for the most part dies. Besides the albumen above described, Gaertner has re- Vitellus. vived the use of the term vitellus, but employed it to de¬ signate a very different part from that to which it was originally applied by Grew. The latter made use of this term to designate the inorganic matter of the mature seed, which in the early stage of its production he called ANATOMY, VEGETABLE. albumen (Anat of Plants book iv. chap. 3); but it is tunics be raised and thrown back, as is done in fig. 36, Oft L albumen \Anat. Plants, book P , nnus the little ob]on. body h, and its semilunar appendage^ employed by Gaertner to indicate a^placed behind it, are brought into view; and, together, body, which in many seeds is placed between the em- u ^ brvo and albumen, and is closely connected with the for- they constitute the embryo. - oac;i,r from the latter. The fisure of Let next a vertical section ot anotner seea ne maaein j^T/heilt'^erv virious^ndif^ the direction oT'theVrrow that runs along its flatter side, this small body is described as being very various m dil- tne Qirecuon 01 u.e - ‘embrvo as is rente! ferent seeds. It is said not to rise out of the earth during and let this the seed to be com germination; but, like the albumen, seems destined to sented in fig. 37 . ^ ^hen observe the seed to be com afford nutriment to the embryo. In the Graminece it re- posed almost entirely of albumen «, with winch the em- presented thfn scale interposed between the albumen and bryo consisting f “Tf “Sied'^aC 'embryo, to which, from its shield-like form, he g.ves the “ E’lSt namtfof Tmtdium. "(De Fructib. Plantar, vol. i.”cap. 11.) the albumen is the cetytede^ whiA Mt that surface is There can be no doubt that this scutellum of Gmrtner is convex, and on the opposite one concave, ti e little “ conglobate leaf first observed in wheat by In fig. 38, the entire embryo has been removed from its Malpighi, and which later writers have denominated the connection with the albumen, and a front view of con. PS ’ - ■ I„ the next section its form and siderably magnified, is there given, m which the letter m denotes the cotyledon, in the concavity of which the cotyledon of that seed, situation will be clearly displayed, Section II. Of the Structure of Monocotyledonous Seeds, as displayed in their Evolution. Classifica¬ tion of seeds: their evo¬ lution ob¬ served. Evolution of wheat. Descrip¬ tion of wheat. plume n is lodged, and o indicates the protuberances from which the radicles afterwards spring. If now this same embryo be reversed, as is done in fig. 39, then the convex back of the cotyledon only is seen, with the extre¬ mity of the principal radicle at the base. It is this side of the cotyledon that was applied against the albumen ; All seeds have by some botanists been distinguished and its polished surface, says M. Poiteau, proves that it into such as possessed one or more cotyledons, and such nowhere adhered by any organic structure. Gaertner also as were entirely destitute of them. In treating of seeds remarks that the connection between these parts is not under the two divisions of mono and di-cotyledons, we organic, but merely superficial,—an observation that is would not be understood to deny the existence of seeds true as far as relates to the embryo itself and the albu- that possess more than two. minous matter, but not as applied to the tunics which Some seeds are so extremely minute, that, until lately, envelope them; for, at the base of the seed, the inner their existence was not clearly ascertained; and it is membrane, which contains the albumen, appeals to be only during their germination that their general form continuous, as Leeuwenhoeck remarked, with that which and character can be detected. In many others, the covers the cotyledon, being reflected from the albumen organized parts are so small as to be scarcely capable over the cotyledon, much in the same way as the pleura of demonstration, except by following the progressive and peritonaeum, that line the sides of the great cavities changes of form they exhibit in their evolution. We pro- in animal bodies, are reflected over the viscera they con- pose, therefore, to select, from each of the two divisions tain. Such are the appearances presented by this seed of mono and di-cotyledonous seeds, an example or two of antecedent to germination: let us next follow it through the successive appearances displayed in their evolution, the several stages of that process. ! which will, besides, form the best introduction to a know- After a seed of this species has been in contact for 24ci ledge of the structure of the mature plant. or 30 hours with the humidity necessary to its germina-ti In most of the monocotyledonous seeds the cotyledon tion, its embryo becomes swollen, and, when removed from" does not appear above the soil during germination, but is the other parts, and moderately magnified, presents thest retained within the coats of the seed, and consequently appearance exhibited in fig. 40. In this figure the radicle undergoes but little alteration in size. As an example of is rendered more protuberant, and the fine tunic that the evolution of a monocotyledonous seed, we shall select invests it has undergone an alteration, being changed from that of wheat (triticum hibernum), because its develop- a smooth, opaque, and solid texture, to one that is villose, ment has been studied with great care, and, in common with transparent, and cellular. A vertical section of the same^ some others of the same natural family, it exhibits some embryo, as exhibited in the next figure (41), shows the striking peculiarities, which add greatly to its productive elongation of the principal radicle p, which caused the powers. The successive appearances exhibited in its evo- protuberance belowr, and the sprouting of the two lateral lution have been given with great accuracy by Malpighi, radicles/> which push forth more slowly on the sides, who has carried its anatomy farther, in some points, than These three radicles soon force their way through the sac most of his successors. In the earlier stages of growth, that envelopes them, which then forms sheaths around some very accurate representations of it have also been their origins. In the same figure, the letter q denotes the given by M. Poiteau; and Dr Yule has likewise obliged plume, consisting of several convoluted leaves, and resting us with some valuable observations. From these different on the cotyledon. In fig. 42 the appearance of the seed, authorities, confirmed generally by our own observations^ in a stage a little more advanced, is exhibited. The we shall endeavour to present a concise view of the struc- plume r is now seen to have risen above the cotyledon s, ture and evolution of this very important seed. and the three radicles, surrounded at their origins by their If we take a grain of wheat, and examine its convex proper sheaths, have greatly increased in length, and in-* side, we observe, at its base, a small oblong body (fig. 35, numerable capillary rootlets are emitted from their sides. Plate XXX\ III.) lying in a semicircular depression, which (An. du Mus. d'Hist. Nat. tome xiii. p. 383.) is well defined through the tunics that cover it. These tunics are two in number; an outer one, to which the chaffy filaments at the vertex of the seed are attached, and which readily separates when moistened; and an inner one, which everywhere adheres closely to the cel¬ lular tissue that contains the albumen. If these two The daily appearances exhibited in the evolution of\ this seed, as previously given by Malpighi (Anat. Plantar^ p. 103), accord well with the above representations of M. Poiteau; and he has noticed some additional particulars of considerable importance. On the first day of germination, cj ie he represents the plume of the embryo as beginning toe ANATOMY, VEGETABLE. 0f Ifth'ieed. open, and the protuberances, which indicate the eruption 3 of the three radicles, as beginning to form. The radicles at this period are completely enveloped in a membranous sac or involucrum; and the body of the embryo is closely connected with a “ conglobate farinaceous leaf, by which nutriment is administered.” This conglobate leaf is the cotyledon before mentioned, and its connection with the radicle and plume is well shown by Malpighi. In fig. 43, t, he exhibits a front view of the radicle and plume, as they appear when separated from the cotyledon ; and at the letter v of the same figure a back view of the same body is displayed, in which the letter x points to the mark or scar that denotes the place of separation. Malpighi believed these parts to be united with each other by a little node, hereafter to be described; but it is by the medium of vessels that this connection between the cotyle¬ don and the other parts of the embryo is maintained; and by this route alone can the nutrient matter or albumen be conveyed through the cotyledon to the radicle and plume. To these vessels M. Bonnet gave the distinctive appellation of mammary: the union they form between the different parts of the embryo is so close, that at this part, says Gaertner, the cotyledon, and radicle, and plume, form one undivided body. (Gaertner de Fructib. Plantar. vol. i. p. 149.) If ic 2-lay, On the second day of germination the exterior tunic of the seed, according to Malpighi, gives way ; the plume [rises upward; the radicles do not as yet pierce their in¬ vesting sac, but this sac is turgid with juice, and is cover¬ ed exteriorly by a fine white down : the cotyledon also, at this period, is rendered moist, jieli lay, During the third day the cotyledon is quite turgid with juice; the plume is much enlarged, and begins to look i green; the three radicles have pierced the enveloping s sac, and are everywhere thickly covered with hairs ; and above the first two lateral radicles, two small protuber¬ ances, y, z, fig. 44, the origins of two more radicles, are now seen to emerge, while the sac that envelopes them is observed sensibly to waste. :jc4li|(lay, When the third day has elapsed, the plume, inclosed in a fine transparent membrane, is still more elevated, ,> and acquires a greenish colour : the protuberances of the f two new radicles are more prominent, and the three for¬ mer radicles have greatly augmented; the cotyledon is much softer, and, as if milky, yielding, when compressed, j a white and sweetish liquor. eo lilay, After the completion of the fourth day, the plume, con- . turning to ascend, pierces the membranous covering a' , (fig. 45), and pushes into day a permanent leaf, green and convoluted, around which the membrane forms a sheath. Inferiorly, the first three radicles have greatly extended, and the two others, b' b', are much increased : the outer coat of the seed now begins to lessen, but still contains a sweetish liquor. M. Poiteau gives a section of the entire plantule about this period of its growth, which agrees very exactly with the figure of Malpighi. In -this sec¬ tion (fig. 46) the plume d is seen to have pierced the membrane d! that formerly inclosed it; the albumen d is diminished; the cotyledon/' retains its situation and form ; and the five radicles / / are nearly of a length, and cover- i, r with hairs. L 1 a' ‘ About the sixth day the plantule, still invested by its sheath, begins to open and expand; the seminal tunics , shrink, and the surface of the outer coat is corrugated. , ^ these tunics are cut open, the cotyledon within is ob- I served in some parts to be firmer than before, and has the s appearance of a concave leaf; but in other parts it is more vascular and filled with juice, especially in that part near to the mammary vessels. After the eleventh day these tunics still adhere to the Of the Seed, plantule, but appear much wasted, and the juice they con- tain is mixed with bubbles of air; while the stem form-llth clay, ing many knots, and the radicles emitting innumerable rootlets, continually augment in size. Where the vegeta¬ tion has been very active, the whole original contents of the seminal tunics are by this time exhausted, and, when compressed, they yield only a v/atery fluid. After the lapse of a month, when the parts already de- 28th day. veloped are still farther advanced, new buds break out from the primary seat of growth and rise upward, and new radicles push forth and descend. So readily are these radicles produced, that sometimes, if the primary ones be removed, others in crowds spring forth ; at the same time new buds or shoots, protected in their proper sheaths, arise from the same part, and surrounding the primary plantule, are borne upward with it. Of these ap¬ pearances accurate delineations are given, and they may be observed in every field of growing wheat. The foregoing descriptions of Malpighi are in general Error of very correct, and his figures, though somewhat rude, ex-Malpighi, hibit faithful delineations of the objects they are destined to represent. In one or two points, however, he has fallen into error, which, in the above statement of his opinions, to avoid confusion, we corrected as we went along. Thus, though he distinctly points out the “ conglobate farinace¬ ous leaf” as the organ by which nutriment is administer¬ ed to the radicle and plume, he assigns to the sac that, in an early state, envelopes the radicles, the function of placentula, and even gives to the exterior tunic of the seed the title of seminal leaf. The true cotyledon, how¬ ever, which never in this seed is produced into a seminal leaf, is the little conglobate body above mentioned; and the common tunics of the seed have no title to the ap¬ pellation of seminal leaves. To this cotyledon Gaert-of Gaert¬ ner, from its shield-like form, gave, as before observed, ner5 the name of scutellum. He held it to be characteristic of the Graminea;, and analogous to the organ to which, in some other seeds, he gave the name of vitellus. (De Fructib. Plantar, vol. i. p. 139.) But later writers, as Jussieu, Smith, Brown, and Poiteau, have all restored to it its proper office of cotyledon. M. Poiteau has gone even farther, and asserted the ex- of Poiteau, istence of a second cotyledon in this seed, and in the oat, which he describes as situated directly opposite to the former. (An. du Mus. dHist. Nat. tome xiii. p. 388.) In this instance, however, he has mistaken the rudiment of the second bud for a second cotyledon, as Dr Yule ascer¬ tained by “ tracing the growth of this supposed cotyledon from its first becoming visible to its final development as a plant.” ( Werner. Transac. vol. i. p. 594.) M. Mirbelof Mirbel. considers the sac that invests the plantule to be the co¬ tyledon of this seed, and this cotyledon to form the first ensheathing leaf. (An. du Mus. dHist. Nat. tome xiii. p. 148.) But, as already remarked, the cotyledon never in this seed rises out of the tunics; and, as Dr Yule ob¬ serves, differs totally in situation, structure, and consist¬ ence, from the ensheathing leaf of the plantule. A very remarkable peculiarity in these plants is their great reproductive power, as displayed in the indefinite number of new plants which we have seen to be evolved from one primary seed. Malpighi not only observed this peculiarity, but has described the structure from which it originates. He considered the radicle and plume of the embryo to be connected with the cotyledon, not by the mammary vessels, as we have stated, but by a little body which he called the umbilical node. In a section of the Peculiarity lower part of the stem of the plantule, made after the of strut-* third day of germination, he delineates this "node as situ- 78 ANATOMY, VEGETABLE. Of the Seed, ated at the junction of the radicle and plume, as repre- sented by the letter h! (fig. 48, Plate XXXVIII.) ; and describes it as solid exteriorly, and softer and more me- Opinion of dullary within. If a section of the same part be made on Malpighi, the fourth day, as in fig. 47, the stem i of the plantule will be seen, says he, to spring from this node, from which also the radicles equally take their origin, of Leeu- This peculiar property was also observed by Leeuwen- wenhoeck, hoeck, though he seems not clearly to have apprehended the nature of the organs from which it proceeded. In the embryo of wheat he describes three points, from which not only three distinct radicles spring, but they are also, he adds, “ the beginnings of three several spires or stalks of wheat; so that from every grain of wheat (which is well worthy of observation) there will arise not merely a single stalk, but three distinct ones, which are formed in the seed itself.” Select Works by Hoole, vol. i. p. 169: and in vol. ii. p. 289, are to be found similar ob¬ servations on the seeds of oats, barley, and rye. c? Mirbel, By M. Mirbel, the umbilical node of Malpighi is con¬ sidered as a fleshy knot (un nceud charnu), by the me¬ dium of which the plume and radicle are united. The lateral radicles which issue from it he regards as distinct in their nature from the primary one, and as resembling those which spring from knots in the stem ; he therefore names them articular roots, les racines articulaires. (An. of Yule, clu Mus. d'Hist. Nat. tome xiii. p. 149.) According to Dr Yule, however, this fleshy knot is to be considered as a tuber, analogous to the tuberous substance interposed be¬ tween the bulbs and roots of the Liliacece and other mono- cotyledonous tribes ; and which is destined to produce an indefinite number of young plants, a greater or less num¬ ber of which are subsequently evolved by the joint agen¬ cy of the roots and leaves. The “ articular roots” of M. Mirbel he regards as in reality young plants, the roots of the Graminece being invariably fibrous. It is by means of these lateral shoots and their tubera that bushes, consist¬ ing of from sixty to several hundred stems, are sometimes • seen to originate from one seed. The above important peculiarities in the germination of the seeds of the Graminece are very perspicuously dis¬ played by Dr Yule in the three figures which we have co¬ pied from his Memoir. In fig. 49, Plate XXXVIII., Dr Yule represents the embryo of wheat as it appears when detached from the albumen, a short time after germina¬ tion has commenced; the ascent of the plume covered with its membrane, and descent of the three primary radicles, which have pierced their containing sac, are clearly exhibited; and the letter A' points to the little co¬ tyledon placed at the junction of the two parts just men¬ tioned. In fig. 50 the germination of the same seed is shown in a more advanced stage; the plume l! has now risen to a considerable height, and pierced the investing membrane ; and at m! a second bud or plume (which M. Poiteau mistook for a second cotyledon) is seen to shoot from the tuber like the first. The letter v! denotes the seminal tunics. At a still more advanced period, four young plants, o'o'o' o', fig. 51, of the second month, with their sheaths in part withered, are seen to have sprung from the same part; but the two seminal tunics of the seed, exhausted of their contents, still remain attached, as indicated by the letter p'. ( Wernerian Trans, vol. i. p. 589.) The description given above of the evolution of wheat is applicable, with little variation, to the seeds of all the cerealia. The seed of the oat emits from four to six ra¬ dicles, all of which break through their enveloping sac at the same place, and thus appear to be contained in one sheath. Such too is the case with barley, the plume of 0 which extends beneath the seminal tunics, and pushes out Oft at the vertex of the seed. 'Jfcf, This peculiar constitution of the seeds above mentioned ^ is attended with important advantages in their culture, and explains the source of their great productive power. A single grain of barley was observed by Du Hamel to have produced 200 ears, each of which yielded 24 grains; so that one single seed planted in a good soil has produced 4800 grains. Curtis and others, by transplantation of thePrC: several plantules of wheat, obtained still higher returns from tin single seeds. For the same reason, these plants are bet-wll( ter enabled than others to resist the injurious effects of accident or disease. If a seed, says Dr Yule, be buried under a stone or lump of indurated clay, the seminal plan¬ tules cannot shoot upward; but stems are then sent off in a horizontal direction, until they can effect their escape upward. Sometimes it happens that a small insect (Mus- ca pumilionis) deposits its egg in wheat, and the grub is lodged in the very centre of the stem, just above the root, by which the stem is invariably destroyed, and the root so materially injured as to prevent its throwing out fresh shoots on each side, or stocking itself, as the farmers term it. Nevertheless, the plants thus attacked are not per¬ manently injured; for, in the instance where these de¬ predations occurred, the crop of wheat was good, and the ears large and fine through the whole field; so that these injured plants, by the production of lateral shoots, yielded an abundant crop. (Lin. Trans, vol. ii. p. 76.) In the germination of other monocotyledonous seeds aEv similar succession of phenomena present themselves, with the exception of those which relate to the multiplicationsee of so many individuals from a single seed. In all, the ra¬ dicle first shoots forth, and the plume soon follows; the cotyledon is commonly of small size, and is retained with¬ in the tunics. As the embryo grows, the albumen is taken up and conveyed through the cotyledon to the young plantule; and before the albumen is exhausted the em¬ bryo is enabled to draw its nutriment from the soil in which it grows. In many instances it appears that the primary radicle ^ of seeds, after a short time, becomes dry and falls off,*™ and is replaced by a great number of secondary rootlets. na| M. Poiteau regards this last circumstance as common and peculiar to monocotyledonous seeds. He has re¬ marked it in many hundred palms, not one of which had a descending or tap-rbot. No plant in the numerous family of the Liliacece is said to possess such a root. The radicle of the Cyperacece does not, perhaps, perish so soon ; but it does not continue long. This premature and constant de¬ struction of the radicle he considers as the cause of the bulbs and truncations which occur, particularly in the Li- liaceee ; for the lateral roots not being capable of receiving all the descending sap, it collects at the lower part of the stem, and occasions these different enlargements. (An. du Mus. d'Hist. Nat. tome xiii. p. 392.) The effects which thus succeed to the spontaneous de¬ struction of the radicle occur partly in other plants, in which the first radicle is naturally permanent, if it be ar¬ tificially removed. Du Hamel found that if the extre-ty mity of permanent radicles were cut off, lateral rootlets were produced; that even mechanical obstruction to the descent of the radicle frequently gave rise to divisions in it, and the production of these lateral rootlets. He ascer¬ tained also, by experiment, that roots extend invariably, not by an elongation of parts already formed, but by new matter added to their extremities; and hence it is that roots, whether ligneous or herbaceous, do not elongate if even the smallest portion of their extremity be cut oft. (Phys. des Arbres, tome i. p. 83.) The results of observa- 1 ANATOMY, VEGETABLE. 79 ;'jjSrl. tions on the growth of carrots in different soils, by Mr * ^ Knight, correspond with those of M. du Hamel. rC Section III. Of the Structure of Dicotyledonous Seeds, as displayed in their Evolution. [* les i From the greater number of seeds which have two coty- * utic. ledons, the phenomena of their evolution may be expect¬ ed to exhibit at least as great variety as those of the di¬ vision last described. In different species they differ in this respect as much from one another as they do from monocotyledonous seeds. Some seeds of this class raise their cotyledons above ground during germination; in others these organs are wholly retained within the tunics. Of each of these modes of evolution we propose to give an example, selecting, as before, those seeds which have been most accurately observed; or which, by the forms they exhibit, seem best calculated to illustrate the gene¬ ral laws by which their evolution is accomplished. In the seeds of this division the embryo is commonly much more completely developed than in those of the class last described, so that the several organs of the plantule become distinctly visible. The radicle and plume are readily distinguished, and the cotyledons are frequently so large as to form nearly the entire mass of the seed. Within the cotyledons the albuminous matter provided for the nutrition of the embryo during its evo¬ lution is often entirely contained; and these organs, as before remarked, rise sometimes out of the earth, increase greatly in size, and after a certain period decay. In other instances no increase in size nor elevation above the surface occurs, but, like the greater number of mo¬ nocotyledons, they remain beneath the soil, and yield gradually their nutrient matter for the support of the embryo. Even in plants of the same natural order, the Papilionacecc for example, some, as lupinus, says Dr Smith, raise their cotyledons into the air and light; while others, as lathyrus, retain them under ground, concealed within the tunics of the seed. As an example of the lat¬ ter, we shall give from Malpighi an abridged account of the successive appearances exhibited by the common pea (pisum), which, in its evolution, approaches in some re¬ spects to that of the seeds last described. Nwilut i The figure and size of this seed are familiar to iepL , u every one. After being placed for a day in circumstances i lk day ’ favourable to its germination, it is much increased in size ; its outer coat is rendered softer, and becomes more white and thin; the umbilical aperture continues closed, but near to it an irregular opening or laceration is visible. If the outer coat be now stripped off, the nucleus comes into view. It is seen to consist of two distinct parts or lobes, which are the proper cotyledons of the seed. These cotyledons are closely invested by the inner tunic; ex¬ ternally they have a convex surface, but internally, where they are in contact, their surfaces are nearly plain. Be¬ tween them, in a small depression formed in their sub¬ stance, lies the plume; it is composed of a number of yellowish leaves folded on each other, and bent inward, and is united by a little curved stem to a small white conical body, the radicle. These appearances are exhi¬ bited in fig. 1, Plate XXXIX., in which one of the coty¬ ledons has been removed, so that the inner surface of the other, together with the plume and radicle, is fully brought into view : the letter a denotes the cotyledon, b the plume, and c the radicle. This radicle at its neck, or point of junction with the stem, sends oft' on each side a little stalk or pedicle to each cotyledon. In the above figure one of these pedicles has been cut through, and the other, that remains attached to the cotyledon, is con-Of the Seed, cealed behind the plume. It is by these pedicles alone that the two cotyledons are connected with each other. When the second day of germination is completed, the 2d day. cotyledons are rendered more tumid, the tunics give way, and the radicle begins to protrude. Soon after, the coty¬ ledons separate a little, and become somewhat concave internally. After the third day the radicle has pushed out through the tunics; it is white, except at its point, which is more deeply coloured, and it emits on all sides fine capillary rootlets; the cotyledons are now farther separated, and by degrees the stem of the embryo, with its curved plume, is disclosed. About the fifth day the stem d, fig. 2, mounts up-5th day. wards: it is white, and bears on its summit the plume e, still curved, and now becoming green : the stem now also begins to exhibit the marks of knots at particular parts: the radicle/is farther advanced, and small protuberances, the origins of future rootlets, appear on it: the cotyle¬ dons g retain their place, are turgid and solid, and still surrounded by the lacerated tunics. At the close of the seventh day the plantule is much 7th day. more advanced: the knots on the stem h h, fig. 3, are quite distinct, and its apex is furnished with broad green leaves, but which are not as yet unfolded. The substance of the cotyledons is still solid, and when compressed yields a bitterish juice: the radicle is much elongated, and has emitted numerous rootlets. After the ninth day the plantule is completely formed :9th day. its stem i, fig. 4, is now erect, and the leaves of the plume k are expanded: the cotyledons l are reduced in size: and the radicle m, or root as it may now be termed, with its numerous rootlets, is greatly augmented. Every part of the plantule except the cotyledons continues daily to increase: at the end of a month these organs are still found to adhere; but are wrinkled, thin, and ex¬ hausted of their nutrient matter, with which the stem and other organs are filled. The progress of evolution in those seeds that raise Evolution their cotyledons above the earth is exhibited by Malpighi of the in that of the gourd (Cucurbita). This seed is of an ob-Sourth long figure, and has a flattened form. It possesses three distinct coats or tunics: the outer one is thick, tough, and coriaceous; the middle one thin, membranous, and of a greenish colour; and the inmost is that transparent colourless pellicle that closely invests, and is inseparably connected with, the cotyledons of the seed. After this seed has been made to imbibe moisture, the outer and middle tunics readily separate, and expose the nucleus, which is seen to consist of two leaf-like cotyle¬ dons, which have no connection with each other, except by the medium of the little conical body, or radicle, at their base. The size and figure of these cotyledons, and the situation of the radicle that connects them, are repre¬ sented in fig. 5, Plate XXXIX. Their external surface ex¬ hibits to the naked eye prominent lines, which indicate a vascular structure, the vessels of which proceed from the radicle at their base. They are commonly five in number, and from their main fascicular trunks ramifications are produced, which, in their distribution, form a finely reticu¬ lated appeai’ance over the wrhole organ. On their inner side the cotyledons are quite plain, and closely applied against each other, but, as already remarked, are nowhere connected, except at the base. This surface is displayed in fig. 6, in which the great vascularity of the organ is rendered more apparent. It is between the two coty¬ ledons that the plume, consisting of minute convoluted leaves, is lodged and cherished. In fig. 7 a part of the nucleus of this seed is represented a little enlarged, and so Of the Seed 1st day. 4th day. ANATOMY, VEGETABLE. , , of rliflfprpnt places vounsr plant continues to increasej and acquires at lengthOf ^ .the two cotyledons have been removed at . P j ft full magnitude, in the progress towards which the co-^ v 'by transverse sections, to show more clearly the situation ^dually, and finally fall, of the radicle and plume. The letter » ^no e p ^ } The geeds of the radish (Raphanus), of the lettuce EvjL from which one of the cotyledons has bee , Lactuca), and of the kidney-bean (Phaseolus), are repre-of as to bring the plume o into view; and p points to the (^“^’^pighi as exhibiting, in their evolution, a si-seeP conical radicle below. _ ., .^irrri cnrcession'of appearances. In all these seeds, and When this seed has been twenty-four ^ m ,c rcum- of thisdivision, the radicle first pierces the stances favourable to its germination, i is 1(- , ’ spmjna] tunics • next the cotyledons come into view, and and the umbilical aperture at its base .a enhrged by he X form JleaveS) betwee„ which the swelling of the parts within: t e co y et on“ e tender plume is for a time concealed, and at a later period gid, and the plume is augmented in size. After the se ender^m In different scedSj however! ,he forras of cond day the outer coat >s N*e ™'ds ,‘lllin these organs, and the periods of their successive evolu- appears as if torn and decaying, “"^ aU the parts within “ese »rM ; the greatest variation, not only as re- ScSViro^^Iirb^Lker, Tcotl £ mthe Ipecies of sled but to the soil, the cUte, ledons are more swollen, their vessels more conspicuous, and season m which it is destined to grow, and the radicle pushes out through the umbilical aper- Having thus surveyed the changes in external formic ana tne raaicie publics & which the germinating seed exhibits, we shall concludestl we tUWhen the fourth day has elapsed, the plantule is still our description by a display of the peculiarities of its in« !« retained within the tunics; and if these te now removed ternal structure. Th.s we shall hud to consist entirely lid examTned the middle one is found to be dry and thin: of vessels and cellular tissue, variously proportioned and and examined, u e , combined. In a longitudinal section of the radicle of the pea in an early stage, Malpighi represents it, as in fig. 11, to be composed entirely of cellular tissue exteriorly, in the centre of which the vascular system, separating at the the cotyledons (q, fig. 8) are whitish, soft, and flexible, but the vessels on their surface are much more distinct: the radicle r is elongated and covered with down, as likewise is the stem s. At the top of the radicle a protuberance is seen, which is white and soft; and far- top into three divisions to supply the plume and cotyle- fith day. ther down appear several smaller tumours, indicating the dons, is placed. A simi ar section of the radicle (fig. 14) nlaces of rootlets about to break out. on the seventh day exhibits corresponding sections of 1 About the sixth day the cotyledons t, fig. 9, emerge the rootlets it puts forth ; which are seen also to consist of from beneath the tunics, representing the u dissimilar cellular tissue, and of vessels that come ofr from the cen- leaves” of Grew, and the “ seminal leaves” of Malpighi, tral fasciculus of the radicle. but which we shall in future denominate cotyiedonous The stem, when about a month old, is composed of a leaves. They are thick, soft, and a little separated from thick bark formed of cellular tissue, within which are each other; their position is pendent; the colour yellow- several fasciculi of vessels. In some very thin slices of ish ; they are very vascular, and between them the plume the stem of the pea, viewed through a microscope of con- is still concealed. The radicle u, at this period, is much siderable power, the arrangement and distribution of the elongated, and rootlets everywhere spring from its sides; vessels and cellular tissue appeared to us as represented the stem v is also lengthened and curved, and, in com- in fig. 15. The centre of the stem was occupied by cel- 9th day. mon with the radicle, is everywhere covered with a white curling down. Towards the ninth day the cotyiedonous leaves (x, fig. 10) assume an erect position. At their points they are still yellowish, but elsewhere green, and their cellular tis¬ sue is filled with a greenish-yellow juice; they begin to separate a little, but still entirely conceal the plume. The stem y, at this period, is greatly elongated, and its lower extremity has become green ; the protuberance that existed at this part is greatly lessened; and below it the radicle z is continued, from which numerous rootlets, covered with capillary productions, break out. lular tissue, round which was a zone of vessels, c,' forming four principal fasciculi. Exterior to this zone was a small ring of thickened cellular tissue, and beyond this the pro¬ per cellular substance of the bark. Near the circumference of the stem were four larger fasciculi, which may proba¬ bly be considered as the “ proper vessels,” while those near the centre may be deemed the sap-vessels. In the plantule of the gourd a similar structure is ob-',, servable. In fig. 19 is represented a longitudinal sec-11 ’ tion of its stem and radicle on the ninth day, as given by Malpighi. In this stem the vessels are disposed in a cir¬ cle that surrounds the pith ; but as they descend towards 21st day. During the following days the cotyiedonous leaves con- the root, they approach each other, and give off ramifica- tinue to enlarge, the stem to elongate, and the plume to augment in size; but it is not yet unfolded. About the twenty-first day the development of the plantule appears to be completed. A representation of its foliage at this period is given in fig. 13; the cotyiedonous leaves, a!, have now reached a great size, are of a deep green colour, and very vascular: they rise by a short pedicle from the summit of the stem. On each leaf seven fasciculi of vessels are visible, which, beyond the middle, terminate in a net-work, from which is produced the cellular struc- tions to form the rootlets. In a transverse section of the stem, on the 21st day, Malpighi describes it as hollow in the centre, around which six fasciculi of vessels are dis¬ posed, and the intermediate portion is occupied by cellu¬ lar tissue. Hedwig and Kieser, however, enumerate not fewer than ten fasciculi of vessels in the stem of this plant, some of which are placed next the pith, and others near to the bark, as in fig. 28. . To these representations of the structure of the pea and hei gourd by Malpighi, we shall add that of the bean in its ture that contributes to form the breadth of the leaf. In early stages, as "delineated by Grew, who has traced the the axil, formed by the cotyiedonous leaves, the plume distribution of the vessels in the radicle and cotyledons U lay concealed; it is now disclosed by the removal of with great minuteness. In fig. 12, Plate XXNlX., is exhi- one of those leaves. At first the leaves of the plume are bited a vertical section of a young bean, which is made to curled and convoluted, but afterwards they expand, and pass through the cotyledons, the plume, and radicle. From their figure is then seen to differ entirely from that of the the extremity of the radicle the vessels ascend in fasciculi cotyiedonous leaves; they have notched margins, and to the neck of the plantule, where they are seen to diverge their suiface is coveted with down. In this manner the towards each cotyledon, and ramify through it, while a !t ANATOMY, VEGETABLE. 81 G the T nk. Btai- Nat's of Btet f tan central vascular portion is continued to the plume. In fig. 20 the vascular and cellular structures of the germinating /bean are shown in conjunction, in a highly magnified re¬ presentation, presented here in a reduced size from Grew. In this figure d denotes the cotyledon, e' the enveloping tunics,/' the cellular tissue, cf the vascular system con¬ tinued from the fasciculus /f in the radicle, and ramified through the substance of the cotyledon; the letter i-points to the plume, which also receives vessels from those of the radicle; and U indicates a depression in the cotyledon, in which, antecedent to germination, the plume was partly lodged; a similar depression existed in the other corre¬ sponding cotyledon. CHAP. II. THE ANATOMY OF THE MEMBERS OF VEGETABLES. Section I. Of the Structure of the Stem or Trunk. Art. I.— Of the Stem in Herbs. In the foregoing chapter we have traced the successive changes of form which the seed exhibits in its progress to constitute the perfect plant; we have next to display the structure of the plant itself in its more remarkable varieties and forms. The leading features of this struc¬ ture have already been laid before the reader when dis¬ coursing on the common textures of plants : it remains now to exhibit individual examples of it, as they occur in the several members of the trunk, the branch, and the root. Botanists employ different terms to distinguish the dif¬ ferent kinds of stems or stalks that support tlie leaves and the organs of fructification. These are the stem (caulis), which is considered peculiar to herbaceous plants; the trunk (truncus), which is proper to herbs and trees; the straw (culmus), which is the appropriate stem of the grasses; and the stalk (scapus), which differs from the other varieties in bearing flowers only, and not leaves, lor the peculiarities in external form and character which distinguish these several kinds of stems, as they oc¬ cur in different species of plants, we must refer to the writers on botany ; and shall proceed to exhibit a general outline of their internal structure. Perhaps there is no plant in which the simplicity of • vegetable organization is more clearly displayed than in the sugar-cane, which belongs to the family of Graminece. In its stem or culm the cells and vessels are comparatively large, retain much of their more perfect forms, and are quite distinct from each other. When treating of the cel¬ lular tissue we referred to the cells of this plant exhibited m the very thin transverse slice, fig. 16, Plate XXXIX., as illustrative of their hexagonal figure, and of their be¬ ing bounded on every side apparently by a single mem¬ brane. In some parts, when the observer is viewing these cells through the microscope, some of them appear quite transparent, from the upper and lower bounding membranes being entirely removed, and the light, in con¬ sequence, being freely transmitted; but in others one or both of these membranes remain, and though they are exquisitely thin, yet, when viewed by a strongly reflected hght, a degree of refraction seems to be produced, which communicates to the surface of the membrane an irregu- ar appearance, such as it has been attempted to express m the darker cells of the same figure. The deception t lat arises from viewing two layers of these cells in con¬ junction, which imparts the appearance of double sides, Of the Trunk. as in fig. 21, was before noticed; and the longitudinal ap¬ pearance of the same organs, as seen both in a single and double series of columns (fig. 22 and 23) was at the same' time described. It is through this cellular structure that the vessels which constitute the other component part of the culm of this plant are distributed. They occur in fasciculi, which, towards the centre, are placed at considerable distances from each other, and preserve a symmetrical arrange¬ ment ; but nearer to the circumference they are more nu¬ merous, and their distribution is much less regular. In fig. 25, Plate XXXIX., a very thin transverse slice of this plant is exhibited, in which this regular disposition of the ves¬ sels at and near the centre s' s', and their crowded state near the circumference t, are well shown; the cells, too, at the centre are larger, and have a more perfect form than those near the circumference of the figure. In fig. 27 a very thin longitudinal slice of this same plant is de¬ lineated ; it is considerably magnified; the letter u! de¬ notes the cellular tissue, and v' v' two fasciculi of vessels which ascend through it. In the palm, which, though belonging to the division Stem cf of trees, we shall notice in this place, a similar disposition die palm; of the elementary organs is observed; but both the ves¬ sels and cells are smaller than in the sugar-cane, and the fasciculi of vessels are also much more numerous. This structure is exhibited in the transverse section of the trunk of the palm, fig. 26, in which the dark spots indicate the vascular fasciculi, and the whiter portion the cellular tissue. As before observed in the sugar-cane, the vessels are seen to be less numerous at the centre than at the circumference, where they are much crowded together, and very irregularly distributed, in consequence of the peculiar mode in which the growth of these trees is ac¬ complished. In fig. 24 of the same plate we have also copied from M. Desfontaines a portion of the longitudinal section of the trunk of another species of palm {dracaena draco), which displays more clearly the irregular direction which the vessels take in the lax cellular tissue through which they are distributed. It was before remarked that these plants do not naturally produce branches, but that their vascular system is expended wholly in the production of leaves at their summit, and their trunk is perfectly cylin¬ drical. If, however, the top of the plant has been cut off, or broken by accident, a division into branches is said to take place. {Mem. de VInstit. Nat. tome i. p. 486.) As in these plants the ligneous and cortical textures are uniformly blended together through the entire stem, it must be presumed that the s«/?-vessels and proper vessels are everywhere associated. This fact is accordingly pointed out by Malpighi as occurring in several species of the Graminece, who, as we before remarked, deli¬ neates a proper vessel as existing in each fasciculus of sap-vessels. The next variety of structure we shall notice is that of of the certain herbaceous plants, in which the proportion of cel-gour .V Malpighi and Linneeus, employs the term gem- and in not possessing the principle of multiplication w i ^ ^ jndjcative^0f that species now under consideration. resides in the tuber. Ti ; tilprPfnrp as the most generally received appellation, The third species of bud is a compound germen, and Th^, theretore, as tim y named bulbus. Its figure is^ somewhat globular , i w summer? says Du Hamel, buds are graduallyD formed in the axils of the leaves, viz. in the angle which tk the petiole forms with the branch. They are at first ex- ceedino-lv minute ; are seen in winter chiefly on the young nameu uucuu*. — - o , nearly leafless, and formed of a very short keel- (canna) and thick succulent scales, and at length separates spon¬ taneously from its parent. Of this species there are two varieties, the one solid, constructed ot a solid fleshy body, varieties, the ope solid, constructed 0 a 80 1 ^ i i* sometimes on the larger ones, and more rarely and having generally the rudiments o t ie nfw I ^ tjie tr’unb> They exhibit different forms, according to outwardly upon it; the other coated, ^ing the k}nd 0f tvee that bears them, and are attached to it veral concentric scales, m the cen xe y & b very short pedicle. Their position on the branch 1>1TheShstC"nec?e'S of bud, and that which is strictly so was considered by M. Bonnet to be reducible to five xhe last species or uuu, unu a j riqsc;pS. sometimes they are situated on opposite sides of leaflets: it «h‘e but phced resembles a branch in miniature, and never separates spon- are placed exactly opposite to each other. taneously from its parent. It is named an eye (oculus) when it puts forth flowers alone, or flowers and leaves to¬ gether; and simply a gem {gemma) when it is unfolded into leaves alone. Of these several species, the last, gemma, is generally considered as alone entitled to the appellation of a true bud by those who consider scales and leaves as essential to its constitution; but this definition, says Gaertner, would exclude even bulbs, and it is better to derive our idea of a bud from a general agreement in properties, and parti¬ cularly from similarity of origin in formation and evolu¬ tion, than from the ever-varying condition .of external form. The two former species of buds occur in the lower tribes of vegetables; the two latter are observed on the stems and branches, and roots of various plants. On the present occasion we shall notice the latter only. That variety of bud which is called a bulb, and which springs from, or is variously attached to, the roots of many plants, has already been stated to exhibit considerable difference in structure. It is sometimes constructed of stances they form a kind of ring round the branch. Some¬ times they have a spiral disposition, and at otners consti¬ tute a sort of double spiral around the branch. In those cases where the buds stand opposite on the branch, the extremity of the branch is frequently terminated by three buds; but where the buds are only alternate, the young branch is commonly terminated by a single bud. In the pine the true buds are placed, not in the axils ot the leaves, but at .the extremity of the branch alone. Some buds stand out a considerable distance from the branch; others are placed in close contact with it; and these va¬ rieties occur sometimes on the same branch in regard to the buds that issue from its sides and extremity. The shape of buds is also very various—some being long and pointed, others short and round; some again are hairy, others smooth ; some very small, and others large. {Phys. des Arbres, tome i. p. 99.) _ * _ Beside these varieties in position and form, which serve l| ijJ to distinguish the buds of different genera and species^ there are also many sorts of buds to be observed on the' difference in structure, it is sometimes consirucieu ui men. cue tu&u wauj uj. ™ ^ several thick scales or leaves enveloping each other, and same tree, whose characters are discoverable by their is sometimes formed of a more continuous and solid sub- form. Those which are pointed usually produce leaves stance. To the first description belongs the bulbous sub- and branches, and from those which are large and round- stance of the lily and tulip. Grew has given a section of er commonly proceed flowers. The former are named the latter, made in the month of September. It displays leaf or wood buds {gemmae foliiferce), the latter flower or the tunicated structure of the bulb, at the base and in the fruit buds {gemmae floriferce) : others which possess both centre of which the young flower, destined to appear in leaves and flowers have been called mixed buds {gemma the following spring, is observed. (See his Anatoyny of mixtce). In some trees, as those of the apple and pear, Plants, table 63.) Of the tunicated variety, the common two varieties of wood-buds occur, one of which is small, onion affords also a good example. The coats that com- Names of branch- buds. onion attords also a goou example, me coais mat com- produces only a small bunch of leaves, and in the end pose its bulb are to be regarded as fleshy leaves, and the becomes a fruit-bud; the other is larger, and gives origin true root, according to Du Hamel, is the fleshy plate that to a branch. supports the bulb, from which the rootlets spring. The As the rudiments of the flower appear in the bulb of; common potato affords an example of the solid bulb, on roots the season before they are destined to bloom, soi whose surface numerous buds, all capable of producing those of the leaf and the flower are distinguishable at the entire plants, are seen. Botanists enumerate a great many same period in the bud of the branch. They are to be other varieties of bulbs, for which we must refer to their perceived, says Du Hamel, in autumn, and continue to writings; and to the works of Malpighi and Du Hamel grow even during v, inter, appearing to be clandestinely for the anatomy of many of them. formed in that season, when the movements of the sap The true bud of the stem and branch was distinguish- seem to be suspended; and are thus prepared to shoot ed by the ancients into two kinds, according as it produced forth on the return of spring. It is, however, only in ea ny tne ancienxs mio two kiiius, accorumg as u prouucea tortn on me return ot spring. It is, either a leaf or a flower, and to it they assigned different perennial plants that these phenomena are observed. An names. “ Germen autem est id quod ex ipsisarborum sur- nual plants do not produce buds, and even those * culis primo vere exit, ex quo deinde folium producitur,” roots survive the fall of the stem produce buds or says Pliny; and this he distinguishes from the flower-bud, their roots. {Phys. des Arbres, tome i. p. 103.) , “ nam gemma propriZ fioris esC By Grew the term ger- Climate appears to exert the greatest influence on the ; ’ men, and by Malpighi the word gemma, is employed to formation and evolution of buds. In cold regions, as we " ANATOMY, VEGETABLE, 91 Of i cK have just observed, the bud is formed many months before ir\ Oft is destined to shoot into a leaf or branch ; but in warm¬ er regions scarcely any interval occurs between the periods of formation and evolution. The buds, in such climates, are said to unfold themselves immediately from the bark into branches, without having remained in the form of buds for any length of time. Sometimes, in the milder seasons of our own climate, the evolution of buds rapidly succeeds to their formation, and the vegetative process with us emulates the productive powers of more favoured climes. In some examples, however, the specific charac¬ ters of particular plants overcome these general tendencies of climate; and thus hot climates are said to possess some bud-bearing plants, and in colder climates there are a few shrubs which are said never to bud. (Willdenow’s Principles of Botany, p. 273.) It seems, however, more correct to consider all plants as bearing buds, from which the branches, the leaves, and the flowers, successively t proceed; and to say that in warm climates in general no suspension of the vegetative process occurs, as in cold ones, and no marked interval is observed, therefore, be- . tween the formation and evolution of buds. The few ex¬ ceptions that occur respectively in warm and cold climates i must be considered in reference to the specific characters of the individual plants. The process by which buds are f actually formed has been called Gemmation or Gemmiji- ji cation. iris:' Having made these few general remarks on the nature I, ds. and formation of buds, we have next to exhibit a few ex¬ amples of their structure and evolution, confining our¬ selves at present to a description of those which produce i either branches or leaves. The mature bud consists of i. two parts ; one that forms the new branch or leaf, and in t the language of botanists may be termed persistent; the other serving only a temporary purpose, and falling when that purpose is accomplished. To the former may pro¬ perly be applied the term germ or gem, and the latter, from its office, may be called hybernaculum. In its lead¬ ing characters the bud bears a near analogy to the more perfect seed, for the germ very exactly resembles the plume; and the hybernaculum, as we shall see, in struc- I ture, office, and duration, approaches near to certain co- a tyledons. ^rbem- The leaves or scales which constitute the hybernaculum, (, u“ and which in future we shall denominate the hyberna- , cular leaves, vary much in number, size, and figure, in dif- r ferent buds. Even in the same bud the inner ones are thinner, and much more tender and succulent, than the outer, and are besmeared with, a viscid humour which in¬ timately unites them; while the outer ones are common¬ ly hard, hairy, and of a scalyr texture. Like the cotyle- donous leaves of seeds, those of the hybernacle sometimes grow for a certain time with the germ, and fall succes¬ sively at periods more or less distant: they are also not less ( I distinct in figure from those of the germ, than the leaves of the cotyledon are from those of the plume. This aris¬ es from the peculiarity in the distribution of their vessels, i which do not spring from one common central trunk, as in ordinary leaves, but are derived from several distinct fasciculi at the base, like those of the cotyledonous leaf, as seen in that of the gourd, a', fig. 13, Plate XXXIX. According to the manner in which they are disposed or folded up in the bud, botanists have assigned them differ¬ ent names, for an account of which we must refer to their writings. In the opinion of Du Hamel, they all derive their origin from the inner layer of the bark, of which they seem to be only a prolongation. (Phys. des Arb. tome i. p. 103.) ern . ^le gem of the bud, which is contained within and protected by these enveloping leaves, is composed of one Of Buds, or more leaves generally folded and curled, but in some instances open and expanded. At first they are very small, and their form is indistinct, so that the pedicle alone is distinctly visible ; from which branch oft' the vessels that form the middle rib, and are afterwards expanded to construct the lobes of the leaf. From the figure which the germ possesses before its expansion being like that of a keel, its vascular portion at this period has been nam¬ ed carina, and its softer part medulla or pith. With respect to the particular portion of the branch Origin of from which the germ internally derives its origin, opinions the germ, have much varied. Some have held that it proceeded from the pith alone ; others from the first circle of ves¬ sels that immediately surrounds the pith; others from the tender wood alone ; and others from the pith, the wood, and the bark conjointly. Grew held this latter opinion. (Anat. of Plants, p. 28.) Malpighi describes the germ as a tender, ligneous substance, formed of vessels and cellu¬ lar tissue, and surrounded by its proper cortical texture. (Anat. Plantar, p. 45.) According to Du Hamel, it ori¬ ginates from the ligneous texture and the pith (Phys. des Arbres, tome i. p. 103) ; and Hill considered it to spring from the first circle of vessels alone, but not to carry with it any portion of the pith. As every germ is composed of the cortical and ligne¬ ous textures, it may be said to originate in part from both, as all these writers seem to admit. Sometimes also the pith of the germ is continuous with that of the branch ; but in other instances no such connection subsists, and there is nothing in the character of the pith of the trunk that renders it at all essential to the constitution of the germ. That in many instances, also, the germ springs from the first circle of vessels, is most certain; but it is not less certain that buds spring from trees long after this first circle of vessels has lost its vegetative power, or has been entirely destroyed. In the oak Malpighi describes the entire bud as con- Bud of the sisting of many scales enveloping each other, and forming oak. an oval body. When evolution commences, these open and expand, and in part fall; but two generally remain and protect the springing germ for a long time. In fig. 15, Plate XL., is represented one of the hybernacular leaves d of the oak-bud, at the base of which the germ e is placed. The leaf d is described as possessing an ob¬ long form, and is very evidently vascular; the germ is exceedingly small, and is represented as possessing at this period only one fasciculus of vessels. By degrees three fasciculi become apparent, which are continued through the germ, and form three pointed extremities, as in fig. 16. These parts augment, and the vascular fasci¬ culi separate, so as to produce successively the appear¬ ances in fig. 17 and 18. At length the small curled leaf/^ fig. 19, rising between the two hybernacular leaves g g, is seen to resemble the plume of the seed, and to possess a form altogether different from the enveloping leaves, which in appearance resemble more the cotyledonous leaves of certain seeds. In fig. 20 is given a vertical sec¬ tion of the germ, considerably magnified, in which h de¬ notes the pith that occupies the centre : it is inclosed by vascular fasciculi, i, that send off through the bark ramifi¬ cations to the several little processes that compose the serrated border of the leaf. (Anat. Plantar, tab. 10 and 14.) But the method of nature in the evolution of buds, con- Other tinues Malpighi, is not always the same; for the hyberna-buds, cular leaves do not always waste and fall as those of the germ increase. On the contrary, in many trees, these leaves, especially those about the base of the bud, losing 92 in horse, chesnut Of Buds, their primary figure, assume new forms, and are convert- ed into the permanent leaves with which the branch is adorned. Examples of this sort occur in laurel, in the apple-tree, the almond-tree, and many others. In other instances, as in the rose, the permanent leaves seem to be generated out of those of the hybernacle, from the apex of which they emerge ; and gradually the latter is chang¬ ed into a sort of petiole, to the sides of which two slendei appendages, the remains of the former hybernacular leaf, adhere. Similar transformations are said to occur in many other plants. Buds on The buds, such as they have been described, pullulate the ends of variously from the sides of stems and branches, and al- branches, ways above the insertion of the fallen leaves; but they spring also from their extremities, and produce the annual elongation of the branch or stem. The manner in which this takes place, and the appearance which the parts ex¬ hibit, have been well illustrated by the observations and dissections of M. du Hamel. In the evolution of the seed, the plume, as we have seen, rises above the earth, and produces the stem, which puts forth leaves, \vhen these leaves fall in autumn, the stem continues, and is terminat¬ ed by one or more buds. The roots, as already remarked, do not increase in length but at their extremity, and therefore never elongate after the smallest portion of their extremity has been cut off. It is not the same with branches ; for the newly formed part of the young shoot actually elongates, especially at its extremity, where it is most succulent and tender,—less in the parts lower down, where it is harder,—and in its more ligneous parts not at all; as Du Hamel, by very simple and decisive experi¬ ments, ascertained. (Phys. des Arbrcs, tome ii. p. 14.) The appearance, structure, and evolution of a bud, at the extremity of a branch of the horse-chesnut tree, are given by the same ingenious author, whose candour, abi¬ lity, and success in the prosecution of these curious re¬ searches render him worthy to rank by the side of Mal¬ pighi and Grew. In fig 1, Plate XLI., is represented part of the annual shoot of this tree, terminated by its appro¬ priate bud, formed in autumn, and which is the com¬ mencement of the next year’s shoot. In fig. 2 a vertical section of the same bud is exhibited, in which the num¬ ber and disposition of the hybernacular leaves that enve¬ lope and protect the germ in the centre are displayed. In the stem of the shoot the letter a denotes the pith, which is surrounded by the wood b b, and this again is covered by the bark cc. In fig. 3 is represented a section of part of the bud detached from the woody part of the shoot, and a little magnified, to show that the leaves of the hy- bernaculum take their origin from the inner portion of the bark. their Proceeding next to the interior part of the bud, Du structure. piamel represents it as composed of numerous small leaves (fig. 4), which are more and more minute as we proceed inward, and are covered with fine hairs. In fig. 5 is a branch-bud of the peach, as seen in February, after all the enveloping scales have been removed. It is com¬ posed of greenish filaments, ranged nearly as they appear in the figure. When some of these filaments were detach¬ ed and viewed with the microscope, they appeared tooth¬ ed at the edges, as in fig. 6, and were covered with hairs. All these filaments were afterwards detached in order to disclose a small body lodged within them, and which ap¬ peared to consist of two small leaflets, folded and toothed at their edges, but not covered with hairs. It is repre¬ sented in fig. 7. It occupied the centre of the shoot, and seemed to be connected with the pith. In the next figure, 8, is represented the bud of the horse- chesnut in the state of evolution. The letters d d indi- ANATOMY, VEGETABLE. cate the scales of the hybernaculum thrust aside by the cL shooting of the germ e, accompanied by two permanentSfv ' ‘ The letter g denotes the place of a second bud. . • l* a. 11 / i id o r»ti rr» Q leaves A vertical section of the same bud is shown in fig. 9, from which all the scaly envelopes have been removed It ex¬ hibits an entire shoot of one year’s growth attached to part of one of two years. From h to i denotes the growth of two years, and from i to k that of one year, with the germ k in the centre, and the lateral leaves as in the pre¬ ceding figure. The letter l marks the pith, m m the wood, and nn the bark of the young branch. From l to n the pith is white ; from n to o greenish ; and towards i it is of a brownish-red colour. From i to k, which marks the ex¬ tent of the annual shoot, the pith is green and succulent, and at p p it is seen to be prolonged into the lateral branches. The wood of two years’ growth, from m to i, is white, and forms a continuous tube round the pith, except where the branches go off. It is covered by another layer so thin as to be scarcely visible, but which will in the end become wood; and this layer is covered by the bark. The ligneous layer of the annual shoot appears to be a prolongation of the new layer of the older one, and, like it, possesses at first an herbaceous character. The cortical layer also seems to be a prolongation of that of the older shoot. As to the pith, though in both shoots it is conti¬ nuous, it is to be observed that that of the older branch is white and dry, and that of the young shoot green and succulent. (P/iys. des Arbres, tome i. p. 117.) It is thus by the formation of a bud in autumn at theE extremity of a branch, and the shooting and growth ofthat1' bud in the succeeding spring and summer, that the trunk*11 of the tree and its branches are annually elongated. Dur¬ ing the first season the shoot retains in great part its herbaceous characters, but in the second it becomes per¬ fectly ligneous. In the axils of the permanent leaves of the young shoot, the rudiments of new buds become ap¬ parent, even in the first season. It was before remarked that trees receive an additionalas circular layer every year, and that from these new layerse1 buds successively spring, so that the earliest branches may contain as many ligneous layers as the trunk, and those of later formation a smaller number, according to the year in which they shot forth, and the circle of wood from which they sprang. Combining this growth in breadth with that in length, it will appear, says Du Hamel, that at the base and centre of a tree 100 years old there is wood of 100 years of age; whilst at the exterior part of the same tree, and at the extremities of its branches, there is wood of one year’s age only. As the latitudinal growth was before illustrated by a diagram, a similar mode may be adopted to explain the longitudinal increase. Let fig. 10, Plate XLI., represent in q r the ligneous portion of a tree that has proceeded from a seed in spring, and is observed in autumn. The following spring a second shoot proceeds from the bud r, which reaches as far as s; but at the same time there is a second ligneous layer formed on the first shoot q r, by which its thickness is proportionally augmented ; and at the end of the second year the tree has the form and extent of the unshaded portion of the figure q s. The next spring the bud s opens and sends out another shoot to t; and ligneous layers are added as before to the two preceding shoots; and thus the tree is extended from q to t. The fourth year the same processes are repeated, and the tree extends from q to w ; and each annual shoot, from the base to the sum¬ mit of the figure, is seen to be composed successively of four, three, two, and lastly of one layer in thickness. This figure, therefore, illustrates the mode in which trees in¬ crease at the same time in height and breadth. The lig- Ej el ANATOMY, VEGETABLE. 93 I 0f ,lds> neous layers may be compared to a series of cones which envelope each other, and which annually augment the diameter of the tree by the two thicknesses of the layers. It shows also that trees grow much more in height than in breadth, and that this growth is effected by the suc¬ cessive formation and evolution of buds at the extremity of the stem, precisely as the first shoot issues from the seed. (Phys. des Arbres, tome ii. p. 50.) It is well known that if a bud be taken from one plant and inserted beneath the bark of a kindred species, it will grow, and retain in its new situation its original qua¬ lities and habits; or some buds, if directly planted in the earth and protected from too rapid exhalation, will f ndiiu. produce a tree similar to the parent. Hence some wri- t Jitv f ters, as Dr Darwin, have considered each bud as an indi- k- mils vidual plant, and that a tree, properly speaking, is a family or swarm of buds, which annually produce as many new plants. The bulbs produced beneath the soil by the tulip I i and many other plants resemble, in this respect, the buds produced in the atmosphere. Vegetables, therefore, are propagated by two methods—the oviparous, as by seeds, and the viviparous, as by buds and bulbs ; and the indi- | i vidual plants, whether springing from seeds or from buds and bulbs, are all annual productions. The reproduction of plants, therefore, appears to be of two kinds, solitary and sexual; the former being accomplished by buds and bulbs, the latter by seeds, or through the agency of sexual in¬ tercourse. | f Section II. Of the Structure of Leaves. fthrtof »;eav ; 4' These are organs of great importance in the vegetable economy: they are not however universal; for the Cac¬ tus, some species of Schcenus, and a few other plants, are considered to be destitute of leaves. Like other parts of the plant, they may be regarded with reference either to their external form or their internal structure. The for¬ mer view belongs more especially to the botanist, who has very happily applied his descriptive language to portray the almost infinite diversity of figure, size, and character, which the leaves of different species of vegetables exhi¬ bit. On this branch of the subject we shall but lightly touch, recalling simply to notice the leading distinctions of the botanist, and such only of them as may appear to be more immediately connected with the structure of these organs. The leaves {folia) are distinguished and denominated according as they are simple or compound. Simple leaves are such as have only a single leaf on the stalk or petiole that supports them, and where all the parts of the leaf are continuous with one another. Compound leaves are those which are mqde up of more than one piece, or where the leaf is formed of parts or leaflets articulated together. In regard to their place, situation, and insertion, leaves are said to be determinate. By the place of a leaf is meant the part of the plant to which it is attached. By situation is understood the disposition of the leaves on the stem or branch, which corresponds to that of the buds. By insertion is expressed the mode of connection between the leaf and the stem or branch; and the direc¬ tion of leaves is considered to bear reference to the posi¬ tion in which they stand to the stem. The foregoing observations apply to leaves considered in connection with other members: when we regard them sing!y> we remark several parts which are common to al¬ most all leaves, and to which particular names have been assigned. The part at which the leaf springs from the branch or stem, whether directly or by means of a pe¬ tiole, is called the base, and the point opposed to this, the Of Leaves. apex, of the leaf. The expanded portion is termed limbus, which has its two sides, faces, or surfaces, as they are in¬ differently called. The prominent lines that appear on these surfaces were named riblets (costulce) by Malpighi, and have very improperly been called nerves by most wri¬ ters ; for the term nerve denotes an organ of a totally different nature, of which not even the existence has yet been demonstrated in any part of the vegetable system. To the line that circumscribes and forms the boundary of the leaf the term margin or border is commonly ap¬ plied. The figure of leaves exhibits the greatest diversity, to their cha- express which various terms are employed. Their marginx™Xzrs. also is either entire or variously fissured, notched, or toothed. Their surfaces axe naked and smooth, or clothed with hairs and studded with excrescences; sometimes they are plain and flat, at others furrowed or plaited; and in all these particulars the opposite surfaces present also the greatest diversity. In size, leaves exhibit the most remarkable differences; and, with respect to substance, some are exceedingly thin, and have a membranous tex¬ ture, others very thick, and of a fleshy nature. As to colour, they exhibit every shade of green, from the most gay and lively to the deepest and most obscure. In some the tint of colour approaches to blue, in others to red; and previous to their fall they all undergo those changes of colour which produce the diversified beauties of an au¬ tumnal scene. The period, however, at which this occurs is very different. Some leaves fall earty, before the sum¬ mer has passed, and are termed caducous; others retain their place till autumn, and then fall, and are named de¬ ciduous; others continue beyond the summer, and are styled persistent; and others, which have a still longer duration, are denominated perennial. In the preceding section on the structure of buds we Origin exhibited the form and appearance of leaves in the earlier from buds, periods of their existence. They were shown to possess a very minute size, and to be curiously folded up and concealed within their various scales or coverings, which effectually protected them from the rigour of the winter season. The germ of the bud, from which the leaves ori¬ ginate, was stated to be composed of the same elementary parts as the stem and bi'anch from which it sprang. From the ligneous ring in the annual shoot it derived its vessels, which, after having traversed obliquely the cortical layers, were prolonged into the pedicle by which it remained attached to the branch. This pedicle was continued to form the keel of the germ in its folded state; and, in its more expanded forms, to constitute the vascular riblets of the leaf. By subsequent growth it gradually extends and elongates, becomes more ligneous, and is formed at length into the slender stalk or petiole by which the leaf remains attached to the branch. {Phys. des Arbres, tome i. p. 123.) The vessels which are given off from the branch to Structure form the petiole are not collected into one bundle, but of the pe- constitute several fasciculi, which are disposed in differ- ent ways about the centre of the stalk. In some stalks there are three, in others five or six, and in others seven or more fasciculi, all placed, says Grew, either in an an¬ gular or circular posture, and at a greater or less distance from the centre. This centre is generally occupied by a pith, but sometimes it is hollow or tubular. In table 49 of the Anatomy of Plants, several transverse sections of the petioles of the leaves of different plants are given, ex¬ hibiting the different modes in which these fasciculi are dispersed through the cellular tissue that forms the greater portion of the stalk. In every instance the pith and bark ANATOMY, VEGETABLE. Structure of the leaf. Vessels of the leaf. Its “ pro¬ per ves¬ sels.” of these petioles are represented as forrmng.one conti¬ nuous substance. In some plants the fasciculi of vessels that come off from the stem are not collected into a cy¬ lindrical figure, but, after being variously implicated with each other, expand at once into a leaf. . Such leaves rise by a broad base, and, from being destitute of a petioie, are termed sessile, forming often a sort of sheath about the stem, of which several varieties are enumerated by botanists. The petiole, as it springs from the branch, may be com¬ pared to a small stem, which at the basis of the leaf ex¬ pands into numerous branches. Frequently it is conti¬ nued through the centre of the leaf, forming its middle rib, and giving off in its course numerous branches. At other times, on reaching the base of the leaf, it sepaiates at once into three or more equal portions, which form as many distinct leaves or parts of a leaf, supported by their respective petioles. The structure of the petiole is the same as that of the branch, being composed of the lig¬ neous and cortical textures, in which sap and “ proper ves¬ sels” are discoverable; and the whole is invested by the cuticle. Where the petiole terminates, the proper or expanded portion of the leaf (limbus) commences, the figure of which is determined by the number and distribution of the vessels. These vessels divide and ramify in various modes, till at their termination they form a finely reticu¬ lated structure. In many recent leaves this structure is very visible; but when the softer parts are removed by spontaneous decomposition, the vascular system of the leaf remains nearly entire, and its extent and form are then more completely exposed. In fig. 11, Plate XLL, this distribution of the vessels is exhibited in a leaf of holly. From the central fasciculus branches are every¬ where given off, which further subdivide, and form small¬ er ramifications, that terminate at length in a minutely reticulated structure. Around the margin of the leaf the vessels are continued, and at certain parts are prolonged into the thorns that bound the circumference of this leaf. In this and similar instances the vessels appear to be ramified out of greater into less; but, as already observed, this does not appear to be really the fact, the vessels being all of the same size everywhere in the leaf, and all continued through it, like so many distinct tubes. This structure Grew represents in a highly magnified view of the leaf of borage, in table 50 of his work; part of which is represented, on a reduced scale, in fig. 13, Plate XLI., designed to show that the ramified vessels o are all clus¬ ters of tubes of the same size, which, though they sepa¬ rate continually, and come into contact, are never pro¬ duced or ramified one out of another. Neither do they ever inosculate or anastomose with each other, until, ac¬ cording to Grew, they come to their final distribution. The vessels thus distributed through the leaf belong chiefly to the order of spiral vessels, as was shown by Grew in the leaf of the vine and many others ; and, from the obser¬ vations of Malpighi, Darwin, and Knight, already related, it appears that “ proper vessels” are everywhere asso¬ ciated with them. It would seem, from the observations of Darwin, that these two orders of vessels communicate in the leaf by continuation of canal, as the arteries and veins do in animal bodies. In the stalk he considered them to be disposed in two concentric rings, the inner one of which carried out the sap to the leaves, and by the outer one it was returned to the bark. (Phytologia, p. 43 and 58.) In branches of the apple and horse-chesnut, Mr Knight also observed coloured fluids to rise through the vascular fasciculi in the petiole of the leaf. These fasciculi were surrounded by others free from colour. which conveyed a different fluid, and, on being traced Of , j down the stalk, were found to enter the inner bark, and*^ g to have no communication with those of the wood. The returning vessels he describes as being situated parallel to and surrounding those which carry up the sap. (Phil. Trans. 1801, p. 336.) Beside the vascular system which, by its ramifica-Cel] tions, forms the skeleton of the leaf, there is anothertissi structure that requires to be noticed. We have seen that11* the vessels, in their ultimate distribution, form a net-work more or less fine and minute in different leaves; so that a great number of intervascular spaces are produced. These spaces or areac, says Malpighi, are occupied by cellu¬ lar tissue, which springs from the vessels themselves, and seems to depend from them ; and by its means the thick¬ ness of the leaf is formed. In the recent part of the bo¬ rage leaf, represented in fig. 13, Plate XLI., this cellular structure, occupying the reticulated spaces formed by the vessels, is observed; but it is most clearly seen in thick leaves, when the cuticle has been removed, and the vessels in part dissected away. The cells or utricles which form this parenchyma, as it has been called, are of differ¬ ent figures, and are mutually contiguous; they are com¬ posed of a membrane formed into the shape of a little vesicle or bladder, from the middle of each of which a little vascular production issues; they are all connected with each other, and with the vascular system of the leaf. In fig. 12, Plate XLI., this cellular structure is represent¬ ed in the leaf of Cactus by Malpighi, in which the oblong cells are described as proceeding from the central vessel, and mutually communicating with each other. (Anat. Plantar, p. 52.) A similar idea of the formation of the cells seems to have been entertained by De Saussure. In different leaves the cells possess very different sizes and forms. In the experiments of M. de la Baisse, they are said to have become tinged by coloured fluids conveyed from the vessels. Whether they communicate with each other is not known; but analogy, derived from other si¬ milar structures, would lead us to suppose that they com¬ municate only with vessels, as seems to be the case in the cotyledonous leaf of the seed. M. de Candolle describes the leaf as consisting of aStn superior and inferior face, between which is an interme-0^ diate part, of very various thickness in different leaves, tojj\ which he gives the name of mesophylle, answering to the parenchyma of preceding writers. The leaf is formed, he says, by ramifications of the vessels, and by cellular tissue, which fills up the intervals of these ramifications. It probably comprehends twro systems; one which, re¬ ceiving the ascending sap, brings it into contact with the air, and permits the exhalation of its superfluous parts ; the other a system which receives the elaborated sap, and reconducts it to the stem, whqre it serves for nu¬ trition. Physiological phenomena, he adds, prove the existence of these two functions in leaves; but anatomy has not yet distinguished the organs by which they are accomplished. (Organog. Veget. tome i. p. 271, 273.) It has been before observed that M. de Candolle follows Kieser in regarding the alleged spaces between the cells, called intercellular canals, as the organs in which the vegetable fluids are conveyed; but the facts and argu¬ ments already stated establish, we think, with sufficient certainty, that, in the wood, the bark, and leaves, the sap is conveyed by appropriate vessels. The consistence of the leaf, continues M. de Candolle, depends much on the relative proportions of the cells and vessels that compose it. W hen the vessels are numerous, the cellular tissue is small in quantity, and the leaf is fibrous, firm, and thin; when, again, the vessels are few and distant, then ANATOMY, VEGETABLE. 95 save.' lives, the cells are more abundant, and the leaf is softer and more fleshy. r j)ifFences The two surfaces of this organ often differ much in : b tli >ur-appearance and structure. On the upper surface the vessels are commonly but little prominent; it is there¬ fore smoother, has but few hairs or pores, and is some¬ times destitute of them. The lower surface, on the other hand, presents the vessels more prominent, and is furnished with hairs and pores much more abundantly. These differences are more particularly observed in the leaves of trees: in those of herbs the two surfaces more nearly resemble each other. In aquatic plants, though the upper surface is smooth and green, and the under one pale and rough, yet the former only, as being exposed to the atmosphere, is furnished with pores. It is to be re¬ marked, however, that whether the two surfaces exhibit an aspect either very different or very similar, each seems destined to enjoy a special function. Hence, if the upper surface be turned downward, and the lower one upward, the leaf tends to resume its natural position ; and if it be retained by force in a position not natural to it, it dies. (Organog. Veget. tome i. p. 274.) In the cellular tissue of herbaceous stems, and of all lolou.ig latupl ]eaves, are contained small globules, which by the action sen-- ■■ mid of light are rendered green, and bestow on the plant its verdure. In the elongated cells of Conferva reticulata they are represented by Kieser as small green globules. (See Plate II. of his Memoir.') They are of a resinous nature, and have been named chlorophylle by chemists ; but as a matter similar in composition is said to reside in the flower, M. de Candolle deems the foregoing name im¬ proper, and proposes to call this matter chromule. {Ibid. tome i. p. 19.) In the substance of the leaf Malpighi describes small bags or follicles as existing between the cells and vessels, and which, in most leaves, yield a peculiar humour. They spring from the vessels, have a roundish or oblong form, and the lip that bounds their orifice is often furnished with hairs. This orifice points towards the upper surface of the leaf, or that which regards the heavens. In different leaves the follicles have different forms; and they afford a secretion which, adhering to the contiguous parts of the leaf, renders them shining. They are numerous in the leaf of the fig, lemon, orange, &c.; and are rendered more vi¬ sible as the cellular tissue of the leaf wastes and dries. {Anat. Plantar, p. 52.) There are other small leaves called stipidee and bractece, 5tl!' which are attached to leaves and flowers, and are of some importance to the botanist. In the- general features of their structure they resemble those already described. I he whole structure of vessels and cells that compose the leaf is covered by a cuticle variously furnished with hairs and pores, and other appendages already described. Section III. Art. I.—Of the Structure of Floicers. The flower is that assemblage of organs which, by the mutual action they exert on each other, give origin to hubs and seeds. All the parts hitherto described con- tybute to the growth and perfection of the individual plant, and have been named the Nutritive or Conservative System; but the organs composing the flower are des¬ tined to continue the race, and from thence have been denominated the Reproductive Organs. In a limited sense t ie bud, produced by the conservative system, may be deemed a reproductive organ, since new individuals may successively be propagated from it; but after a certain period, plants thus produced seem to degenerate, and be¬ come incapable of producing fruit. Reproduction with-Of Flowers, out fecundation presents no organic apparatus which proper to it; but sexual reproduction, or that accomplish¬ ed by fecundation, exhibits numerous and varied organs, which merit all our attention, not only because they exer¬ cise the principal functions of vegetation, but because, from the constancy and symmetry of their forms in the same species, though varied infinitely in different species, they afford the base on which Classification rests. In relation to the organs of reproduction, vegetables, as Sexual before remarked, have been divided into two great classes: organs in Is?, such as have flowers visible to the naked eye, and in dowers, which the sexual organs are distinct; 2dly, such as haveyisl.bJ®1or flowers (it there be any) which can be rendered visible11 V1S1^ e' only by the microscope, and in which the sexual organs are not distinct. The former class, named phaenogamous or phanerogamous, comprises all exogenous, and the great¬ er part of endogenous plants ; the latter, or cryptogamous division, comprehends some endogenous vascular plants, and all those which possess a simple cellular structure. At present we shall treat only of the first or flowering class. The flower is essentially constituted by the presence of Varieties one of the two sexual organs, or of two re-united by a in sexual common support, with or without the exterior envelope orSai!H- destined to protect them. In some instances the flower bears only a male or female organ, and is then named unisexual; but more commonly these organs are present in the same flower, and it is then called hermaphrodite. This unisexual flower, again, is simply male or female, Bescrip. according as it possesses either stamens only, or pistil only, lion of In some instances, though the same plant bears both male sexual and female organs, it is not hermaphrodite, since these or-orSans* gans occur in different flowers ; in others, again, the male and female flowers exist only in different plants. Lastly, we sometimes find both male, female, and hermaphro¬ dite flowers mingled together, either on the same or on different footstalks. The distinctions founded on this se¬ paration, re-union, and mixture of the sexual organs, have served as the basis of certain classes in theLinnsean system. •Sometimes the male or female organs alone, protect¬ ed in a small scale, constitute the flower; but in general they are surrounded and protected by others, named the corolla and the calix. All these parts are commonly borne on a stalk called the peduncle, which, expanding at its ex¬ tremity, forms the receptacle, or torus as it has been call¬ ed, upon which all the parts above mentioned are support¬ ed. In fig. 1, Plate XLIL, the sexual parts which consti¬ tute a perfect flower are exhibited. The letter a denotes the corolla, formed, in this flower, of six leaves called petals; b the stamens or male organs, which surround the pistil or female organ c ; and d indicates the peduncle of the flower. In the next figure (fig. 2) the corolla has been removed, and the parts within are then more clearly seen. They now consist only of the sexual organs supported on the peduncle, and at their base are embraced by the calix e. The pistil, in the centre, is composed of three parts, viz. the ovariumthe style g, and the stigma h ; while each stamen consists only of two parts, the filament i, and the anther k, which latter contains the prolific matter named pollen. Such an assemblage of organs constitutes a per¬ fect or complete flower; but it often happens, that in flowers where the sexual organs are present, one or other of the accessory organs is wanting; thus, in the tulip only one of the protecting envelopes is present, and botanists differ as to the name it should bear. Linnaeus considers it as a corolla, because it is coloured like ordinary petals ; but M. Jussieu regards it as a calix, from its analogy in structure and use to that organ. 96 ANATOMY, VEGETABLE. The flower is sometimes attached immediately to the of the flower are clandestinely formed, though still extreme-Of. Of Flowers. The iiower is someumes —Iv minute. As they enlarge, their position within the bud^ branch by its base, when it is termed sessile; butin g J vari0us in different species: they gradually expand Peduncle ral it is borne on a footstalk or peduncle. T his Ped , f/ tl perfect flower, whose several parts we have ^ of the flower, like the petiole of the leaf, is sometimes fl^etoptdescribe. single and sometimes ramified : in the latter case its i- llc^he Auricle of the flower, by which all the other partsStrl visions take the name of pedicels. c ,i nrp connected with the stem or branch, is frequentlyo/t] Around the base of the flower we often meet wi f, divides into several parts or pedicels, asdu'l leaves which, in size, colour, form af existence differ ^Sle’arbeUtn^ed; each of which supports one or more from ordinary leaves. They are called bractea, am i om e 7 Tbe modes in which the flowers are disposed on axils they form with the stem or branch the ' ' , pedicels have received different names from botanists, ten spring. From thus being the birth-place of the flovver, the pedicels have^ the ^ which draws from them a great part of the nutnen » florescence & The peduncle in structure resembles the the bractete are -aUer and lelftbelng formed, like it of the severa. com- E1*. . , t Tr> ci'zp. in lenp-th. and of the flower. Floral leaves or bractese. the flower. They must, however, be regarded as leaves, mo- mon textures dified by the circumstances in which they grow, as is demon¬ strated by the fact, that examples frequentlyoccur in which bractem are changed into true leaves. (De Candolle, Or- ganog. Veget. tome i. p. 438-9.) When these bracteae approximate so as to form a sort of envelope around the flower, they take the name of involucrum,—when this en In size, in length, and other external cha¬ racters, it exhibits all the varieties already noticed in the petiole of the leaf. , , At the extremity of the peduncle is placed the fiower-of cup or calix, which has very different forms, and has re-lk ceived in consequence different names. Even in its most common forms it exhibits great variety. It is sometimes and has a tubular form ; in vplnne becomes a sheath, it is named spatha,—in the composed of a single piece, G^mimTkk termed glum*, &c.; of all which forms others, of many p.eces called sepals (s^a), wh.ch vary m there are numerous modifications, to which botanists at¬ tach different names. The bracteae approximate more or number, position, and size, and for an account of which we must refer to the writers on botany. In some instances less in character to the calix, in proportion as they are this organ falls when the fruit has set; in others it con- coloured or exhibit the form of whorls ■ and the transition tinues till the fruit ,s mature; and in others the fruit is of the organs of vegetation into those of florescence is so formed within it, to which it becomes a permanent cover- gradual, says M. de Candolle, that the more we study it, ing, and exercises the office of pericarp. The colour of the better do we comprehend that unity of composition the calix is commonly green, but sometimes partly red, or which constitutes the base of philosophic organography, white, or yellow, ™ "owe., „ ,, eo.irelv want,on. (Organog. Veget. tome i. p. 447.) The flower, as well as the leaf, originates from a bud, and in many respects the buds of flowers resemble those from buds. 0f leaves. They are covered and protected in the same manner, but they are generally larger, and have a more rounded form. They consist of two parts, the gem or Flowers originate In some flowers it is entirely wanting. From the dissections of Grew and Malpighi it appears, like the leaves, to be constructed of a cuticle, a pretty thick cellular tissue, and of vessels, all of which exist in the peduncle, and are derived from the common textures of the plant. Above the calix, the corolla, the chief ornament ofof eye (oculus), as it is sometimes called, and the hyberna- flowers, is borne. It is inclosed by the calix, but sur-rt " -m . • 1 T a _ A F. ^ •»* /-v»-i ■»-» rl n 4-V»/-v i /“VT* Y'vO vFo /A'P FllO fWAT Q T4 fl IS Ol f'T* V C.O" rounds the interior parts of the flower, and is of every co¬ lour but green. It consists of one piece, or of several: these pieces are called petals, and according to the num¬ ber of these pieces the corolla is variously denominated. The monopetalous corolla, or that composed of one piece, is also distinguished into several parts, expressive of form, position, or quality; and in the polypetalous variety, culum, or protecting envelope. In some trees they spring from the extremity of particular branches; in others from the branches in common with the leaf-buds ; and in others from the axils of the leaves. In fig. 3, Plate XLII., is a portion of the branch of the peach-tree, bearing two flower-buds 11, between which the smaller and more pointed leaf-bud m is placed. x , * „ . . . .... . - Flower- Grew discovered and has exhibited many examples of which consists of several pieces, each petal has its claw buds form- the complete formation of the flower many months before (unguis) situated at the base, and by which it is attached ed in au- jt destined to bloom. (Anat. of Plants, tables 63, 64.) to the calix or receptacle, and its expanded part (lamina), tumn. of j-idg fact p)u Hamel has also given several examples, which is of very various figure, size, and colour. At the In fig. 4 is a longitudinal section of the flower-bud of the base of the petal a tuft of hairs is often observed, and its peach-tree, made in the month of February, to show the surface is frequently covered with a fine down, or some- complete formation of the stamens and pistils within the times with jointed hairs bearing globulets, as occurs on surrounding envelope of the bud. In fig. 5 is another leaves. With regard to structure, Malpighi describes it representation of a similar bud, from which the enveloping as composed of the same textures as the common leaf, as scales have been removed: it displays the calix, which at is distinctly visible in the thicker varieties of it. Grew this period completely envelopes the other parts of the showed it to possess spiral vessels,—a proof, as Du Hamel flower. When the leaflets of the calix are separated, as observes, that it is partly derived from the ligneous texture, is done in fig. 6, then the stamens and pistils are fully dis- since such vessels are not found in the bark. The odour closed: the petals also of this flower were visible, but at this period were very small. Even the anthers were found to contain a fine dust, but no rudiment of the future embryo could be detected in the ovary. In the flower- bud of the pear, examined in February, the parts of fruc¬ tification were also visible, but indistinct; a month later, all the parts were more advanced, and the stamens, petals, and pistils were distinct, and towards the end of March, even the rudiments of future seeds were visible in the base of the pistil. (Phys. des Arbres, tome i. p. 200.) Thus these organs possess leads to the belief that they contain also a peculiar juice. (Phys. des Arbres, tome i. p. 215.) The organ next to be noticed is the nectary (nectarium)^ In a strict sense this term designates any excretory gland which is situated on any one of the floral organs: and the juice it yields bears the name of nectar. The excretory glands observed on flowers deserve a common designation, because, whatever be their position on any part of the flower, whatever the nature of the juices of the plant, or whatever the size, form, or consistence of it appears that through the winter season the several parts the gland itself, the secretion it yields is always saccha- ANATOMY, VEGETABLE. 97 ne si in:! t tail. I 1 bef ent i he nj *r. , rs.rine, and very similar in all known plants. In regular A'flowers the nectaries may occur on all the organs, symme¬ trically disposed. Their most common seat is the recep¬ tacle or torus. In some flowers, however, they are seated on the ovary, in others on the corolla, calix, stamens, &c. They have the form sometimes of distinct tubercles; but in different plants they exhibit various other forms. In irregular flowers the nectaries are not thus symmetrically placed, but are found on one part and not on the corre¬ sponding part of the same flower. Sometimes they supply the place of abortive stamens or pistils. The juice they secrete is greedily sought by insects. (Organog. Veget. tome i. p. 535.) Art. II.—Of the Sexual Organs of Flowers. Having exhibited this general view of the flower, we must now describe somewhat more particularly the forms, characters, and structure of its most important parts—the sexual organs. The male organ or stamen consists, as before remark¬ ed, of two parts,—the anther, with the pollen it contains, and the filament on which the anther is borne. The fila¬ ment, though a common, is not an essential part, and is sometimes wanting, when the stamen is said to be sessile : the perfection of the stamen resides, therefore, in the an¬ ther, and especially in the pollen it contains. In regard to number, length, position, direction, &c. the stamens exhibit great diversity in different plants. The number of stamens in each flower varies from one to twenty or more; and on this number, progressively in¬ creasing in flowers truly hermaphrodite, the first ten classes in the Linnsean system are founded. In length, the stamens are equal or unequal; and this disproportion is sometimes symmetrical and sometimes not,—peculiari¬ ties which give rise to the formation of other classes in the Linnaean arrangement. In position, the stamens are some¬ times opposed to the divisions of the petals, and some¬ times they alternate with them. Sometimes, from being shorter, the stamens are wholly included within the co¬ rolla : in other instances they protrude beyond it. Their direction is erect, pendent, or horizontal; and their sum¬ mit is variously inclined to or reflected from the centre of the flower. In some flowers the stamens are united by their filaments; in others by their anthers; and in others they are joined to and almost confounded with the pistil, giving origin, in their various mbdes and degrees of union, to the formation of other classes in the Linnsean system. The filament which bears the anther is most commonly straight and filiform : sometimes, however, it is flattened or wedge-shaped, or awl-shaped. It is sometimes as small as a hair, sometimes large and flat like a petal, and its summit is either pointed or obtuse. To the summit the anther is commonly attached; but in some flowers this part is prolonged beyond the attachment of the anther. In structure, the filament resembles the corolla; and it often happens that these organs are transformed into one another. Thus the rose, in its wild state, has only five petals and many stamens; but by cultivation the stamens gradually degenerate into petals, and the flower becomes sterile. Malpighi describes the filaments as originating horn the ligneous texture, being formed of spiral vessels and elongated cells; and as they are sometimes produced h'om elongated floral leaves (petals), they must necessarily, he adds, be formed of the same parts. (Anat. Plantar. p. 64.) 1 v I he anther is that essential part of the stamen that contains the fecundating matter. It is borne on the sum¬ mit of the filament, and is generally formed of two small vol. m. membranous sacs, attached immediately to each other, orOfFlowers united by an intermediate connecting body. Anthers, therefore, are generally bilocular; but in some plants there is only one sac or loculament, and in others as many as four. The sacs are either round, oval, or elongated: each sac commonly exhibits on one of its sides a longitu¬ dinal furrow: this side is properly called the face, and the part opposed to it the back, of the anther. The an¬ ther is attached to the summit of the filament, sometimes by its face, sometimes by its back, &c. In form, the an¬ ther is subject to great variety; and the two sacs that compose a bilocular anther are joined together in very dif¬ ferent modes. In discharging the pollen the sacs open in different ways in different genera of plants, most common¬ ly by the longitudinal furrow that runs along the face of the anther; but in some plants the opening, or dehiscence as it is called, is through two small holes or pores situated at the summit of the two sacs; and sometimes it is by a sort of little valves in some one or more parts of the an¬ ther. As the filaments of the stamens sometimes cohere, so likewise do the anthers, in a manner to form a sort of tube. In other plants the stamens, instead of being free or simply united together, coalesce so as to make one body with the pistil. The colour of the anther is often yellow, orange, violet, white, &c., but never green or truly blue. The 'pollen, contained in the anthers, consists of nume- The pol- rous regularly figured small particles, which possess a very len. different figure, size, and colour in different plants. Many of their forms and sizes are delineated by Grew in table 58 of his Anatomy of Plants. In fig. 9, A, Plate XLIL, copied from Grew, one of the filaments, bearing its anther, as it appears when detached from the pistil of the mallow, fig. 8, is exhibited. The particles of pollen are consider¬ ably magnified, and still more highly in fig. 9, B. In fig. 10 of the same plate, several representations of the forms of the pollen, as given by Du Hamel, are also exhibited. The number of these particles in each anther extends, it is said, from a few hundreds to many thousands. In the cell of the anther they are said to float in a viscous liquid, and to be nowhere attached to its sides, though it is probable they once were; and M. Turpin has even designated a salient point in each cell as the part which produced the pollen. (De Candolle’s Organog. Veget. tome i. p. 465.) In some flowers the pollen consists of transparent grains ; in others they are of a white, purple, blue or brown, and more frequently of a yelloAV colour. Their surface is either smooth or rough, or covered sometimes with a viscous matter. When examined under the microscope at the pe¬ riod of maturity, they may be seen to burst, and yield a fluid, in which, according to Du Hamel, small granules are seen to float. This fluid is described as being thin, or viscid, or oily. If a grain of pollen be thrown on the surface of water, it may be seen to swell insensibly, and finally burst. At this moment a minute quantity of fluid matter escapes, which spreads on the surface of the water, and forms a sort of slight cloudiness. It is to this liquid matter that the fecundating property of the pollen has been attributed. (Elemehs de Botanique, par Achille Richard, p. 199.) This fluid is said by Hedwig to be discharged at once on the bursting of its containing cap¬ sule ; but Koelreuter considers it to be slowly transmit¬ ted through pores in the side, or hairs on the surface of the capsule. Instead of being formed of distinct granules, the pollen is sometimes met with in a solid mass. In many genera of the families Apocynece and Orchidece, fA the pollen con¬ tained in one sac is united into one body, having the form of the containing sac; in other instances it forms smaller N 98 ANATOMY, VEGETABLE. OfFlowers. masses, and these are united by a sort of elastic net-work; while in others the masses are granular. The pollen is very inflammable, and in many plants its odour is said, to bear a striking resemblance to the corresponding secretion in animals. (Elemens de Botanique, par Achille Richard, p. 200.) Sponta- It has lately been said by some, that the particles con- neous mo- tained in the grains of pollen exhibit spontaneous motions, lion in the]jke the spermatic animalcules of animals; others ascribe pollen. these motions to causes distinct from life; and others, though they allow similar motions to be exhibited by the molecules of unorganized matter, maintain, nevertheless, that in the granules of pollen they are independent of physical causes, and resemble the less rapid motions of some of the simplest animalcules of infusions. The pistil. The parts of the flower hitherto described are con¬ structed, says Malpighi, with reference to the female or¬ gans, in which the seed, the last result sought by nature, is curiously formed and matured. This organ, called the pistil (pistillum), consists of three parts. These are re¬ presented in the pistil of the almond, fig. II, Plate XLIL, where the letter jo indicates the stigma, q the style, and r the ovary. To this latter organ Malpighi and Grew gave the name of uterus, Linnaeus that of germen, and Gaertner that of ovarium. In different plants the pistil exhibits great variety in form, size, number, and mode of attach¬ ment ; upon which many of the orders in the Linnaean system are founded. The ova- The ovarium occupies almost always the inferior part rium. 0f the pistil. It is the organ in which the seed is pro¬ duced. When cut open it exhibits one or more cavities or cells, in which are contained the rudiments of the seeds or ovula; and it is in it that the change of the ovula into perfect seeds is accomplished. Its form is various, but most commonly ovoidal. It is seated commonly on the receptacle, together with the stamens; but frequently it is placed below the flower. Its cavity consists sometimes only of one cell or loculament, in which one or more ovula are found. More frequently there are two or more cells containing ovula: these are sometimes disposed in regular series, sometimes they are scattered and without order, and sometimes they are united to one another in a globular form. When these ovula are fecundated they become seeds; but in some plants a certain number are constantly abortive. The style. The style is a prolongation from the summit of the ova¬ rium, and supports the stigma. It is commonly so situate in the flower as to be surrounded by the stamens ; but sometimes it is entirely wanting, and the stigma is then said to be sessile. The ovary in different plants may be surmounted by one or more styles ; while in other plants there is but one style to many ovaries. Most frequently the style occupies the summit of the ovary, but sometimes it springs from its side, and very rarely from its base. The forms of the style are numerous :—the most common is the filiform; but frequently it is thick, angulated, or club-shaped. It is commonly a hollow tube which com¬ municates with the ovary: there is usually but one style to one ovary, but sometimes more. In some instances the pistils correspond in number with the loculaments into which the pvary is divided; in other instances every seed that is formed has its distinct pistil, while in other examples only one pistil is allotted to a great number of seeds. In some flowers the style is so connected with the ovary that it falls after fecundation ; in others it con¬ tinues after that event, and forms a part of the future fruit. The stig- The stigma, which forms the summit of the pistil, some- ma- times terminates the style by an open mouth. Sometimes it appears like a small bud; in other instances it is va- (jj riously divided or forked. Sometimes it is smooth; antN is sometimes covered with hairs. The number of stigma¬ ta is determined by that of the styles, or divisions of the styles, and therefore varies from one to six or more in dif¬ ferent plants. Sometimes the stigma is attached to the summit of the ovary, without the intervention of a style; but most frequently it is seated on the summit of the style, or occasionally attached to its side, or to that of the ovary. In consistence, the stigma is thick and fleshy, or thin and membranous, or formed of small glandular bodies. In form, it varies exceedingly; and its surface is smooth, pubescent, or plumous, &c. In structure, it is ordinarily glandular; and at the period of fecundation its surface is rendered moist by a peculiar viscous matter. When the pollen falls on this matter, its grains burst, and the gra¬ nules, called by some favillce, are supposed to be absorbed by the spongioles seated in this part, and conveyed to the ovula by the vessels which form the pistillary cord, and thus accomplish their fecundation. M. Bulliard is said to have traced coloured liquors, previously absorbed by the stigma, along the vessels in the interior of the style, to the ovula contained in the cells of the ovary. (De Can¬ dolle, tome i. p. 480.) In many plants, however, either from the structure of the pistil, the nature of the pollen, or the circumstances in which fecundation is effected, the pollen seems never to come into contact with the ovula. Section IV. Of the Structure of Fruits and Formation of Seeds. In the preceding section we described the structure of the flower antecedent to fructification: we have now to exhibit as concisely as possible the changes of form it undergoes after that event, and particularly as it regards the production of the seed. After fecundation has been effected, the calix, corolla, d stamens, and even style of the pistil, commonly fade andt! fall; the ovary alone remains, and undergoes very differ-*1 ent changes of form in different plants. In the latter pe-c riods of its enlargement it is usually called pericarp (peri- carpium), a term which is understood by botanists to ap¬ ply also in certain cases to the calix, the corolla, or any other apparatus of organs that serves as a support and defence to the seed. In its early state the ovary is described by Gaertner asij possessing at first a simple cellular structure, which at aa later period takes the form of distinct cavities or locula-jlj ments. Within these cavities minute globules are after-J wards seen, which are the rudiments of future seeds. Ac¬ cording to Mr It. Brown, the ovulum, in the unimpreg¬ nated ovarium, is attached only to a part of the mem¬ brane that lines its cell or internal cavity; but soon after fecundation in some cases, and still oftener during the growth of the seed, this membrane coheres so closely with the proper coat of the seed as to be no longer distinguish¬ able or separable from it. In the ovarium of plants of the family Compositcc he observed two slender filiform cords, which, originating from the base of the ovulum or its short footstalk, ran up, and were more or less connected with the parietes of the ovarium, until they united at the top ot its cavity immediately under the style, between which and the ovulum a connection was then formed. In some species of this family, as in tussilago odorata, these cords were easily separable from the ovarium, and could be removed entire along with the ovulum. These cords he regards rather as vessels conveying nutrient mat¬ ter, than as organs by which impregnation was accom¬ plished. A similar appearance of vascular cords con- 1 ANATOMY, VEGETABLE. 99 F ts. necting the ovulum with the style was observed in the ^ familyBrunonia; and in certain liliaceous plants the bulb¬ like seeds, which separate from the plant before the em¬ bryo becomes visible, are supplied with distinct spiral vessels, which, entering at the umbilicus, ramify regularly through the fleshy mass, and appear to have a relation to the central cavity, where the embryo is afterwards formed. (Linncean Trans, vol. xii.) Thus, then, the ovulum in the plant, like the egg in the animal, is, to a certain extent, pro¬ duced without fecundation; but if that function be not performed, it soon degenerates and wastes. The progres¬ sive changes that occur in the ovary itself, and in its con¬ tained ovula, subsequent to impregnation, have been ob¬ served by Malpighi in several plants, whose observations, as they relate to the almond, we shall briefly detail. In fig. 11, Plate XL1I., is represented the pistil of the almond, as it appears soon after fecundation, in which the la Jterovary r is seen to be somewhat enlarged, and has an oval Inc- form. In the next figure (fig. 12) a longitudinal section of the same pistil is exhibited, exposing the cavity of the ovary s, within which a small vesicle, t, is placed. As the ovary (fig. 13) enlarges, it becomes rounder, and its style is contorted and diminished in size. If in this stage it be laid open, as in fig. 14, its vessels, which in the peduncle w are disposed cylindrically, are observed to be dilated at the place of the calix x, and to give off branches to the ovary itself, and to the shell y that now begins to be formed within it. In the centre, the ovulum z is now seen to be much increased in size. All the parts conti¬ nue to augment; the ovary (fig. 15) becomes rounder, and the appearance of the style is obliterated. On exposing its cavity in this stage, as in fig. 16, the ovulum a' in the centre is observed to be much increased, and the outer layer b‘ of the shelly covering that invests it now begins to harden. Such are the changes of form and structure ex¬ hibited by the ovary: we have next to trace more mi¬ nutely those of the ovulum that is produced within it. The earlier appearances of this body have been exhi- hejyu-bited in fig. 12 and 14. When removed from its seat, a few days after fecundation has been accomplished, and viewed by a moderately magnifying power, it exhibits the form and appearance represented in fig. 17. Externally it is covered by a vascular tunic, d, derived from the inner coat of the ovarium, a part of which coat adheres to it at d'. If in this stage the ovulum be laid open by a vertical section, as in fig. 18, it is seen to be composed of two tunics or sacs, one within the other; the inner one is filled with a cellular tissue, that contains a transparent hijaa;juice. To this inner tunic the term chorion may be pro¬ perly applied. At a period a little later, when the ovulum is examined, a tubular body (e', fig. 19) is observed to extend through the chorion or tunic last mentioned. Shortly after, this tube expands at its apex, and is found to contain a small vesicle. In fig. 20, which represents a section of the en¬ tire ovulum, the outer tunic, the chorion, and the tube f expanded at its apex, are exhibited. To it the appella- ; tion of amnios may be given; for it is the organ in which the embryo, or corculum, as at this early period it has been called, is first seen to emerge. Through several successive days the expanded portion of this tube en¬ larges and forms a sort of sac, which is filled with cellular tissue, and the summit of which, saj^s Malpighi, the em¬ bryo is seen to occupy. In fig. 21 this amnios is separat- ■n yo.ed from the other tunics, and at its summit the embryo y' is observed. If removed from its seat, the embryo pre¬ sents the appearance Ji (fig. 22), and when expanded, as hi fig. 22, f, is seen to consist of a body and two little wings. T Having thus viewed the several parts of which the ovu- Of Fruits, lum is composed in their separate state, let us next ob-^~v^-^ serve them in connection, and trace the series of appear-The °vu- ances they exhibit, and the effects they produce on each t|'e other. In figure 23 is given a vertical section of the en-. tire ovulum in a more advanced state. The outer coat h! still envelopes the others; the embryo l occupies the summit of the amnios m', wdiose lower part, still tubular, is continued through the chorion rL In the next figure (fig. 24), from which the outer coat has been removed, the embryo o' and the amnios p are represented as en¬ larged ; but the chorion 9' is partly exhausted of its juice, and has fallen down in a collapsed state. At this period the embryo, when separated from the amnios, has the form r\ fig. 25, and in its expanded state is represented by the letter d of the same figure. The bulk of the em¬ bryo t, fig. 26, continually augments, and encroaches on the capacity of the two tunics v’, a/, whose forms are con¬ stantly changing; and from being successively emptied of their juices, with which the embryo becomes filled, they are gradually pressed downward. At last the em¬ bryo p, fig. 27, is so much augmented as to fill the cavity of the outer tunic; and by this time the amnios and cho¬ rion, exhausted of their fluids, exhibit the shrunk and corrugated forms in which they appear at the bottom of the figure. According to this representation, the outer tunic h!, fig. 23, derived from the ovary itself, and the fine membrane that immediately invests the embryo, form the only permanent coverings of the mature ovum or seed; for during the progress of formation the chorion and am¬ nios (which are successively produced subsequent to fecundation) are again obliterated by the growth of the contained embryo. Malpighi describes the process of formation in many other seeds to be nearly similar; for his descriptions of which we must refer the reader to his work (A?iat. Plantar, p. 71). In the above descriptions of Malpighi, the several parts seem to be clearly exhibited, except in one important particular, namely, the situation and course of the umbili¬ cal cord. In almost every instance he designates the tube, which we have represented as the first form of the amnios, as the umbilical vessel (vasculum umbilicale), which the subsequent appearances it exhibits show to be erroneous. In the descriptions of Grew this deficiency in the representations of Malpighi is supplied. He has particularly observed the formation of the seed in the apricot, which in many respects resembles that of the al¬ mond ; and we shall subjoin an abridged account of his observations. In this fruit the pericarp that envelopes the seed is in the seen, in its mature state, to be composed of the pulpy partaPrlcot, by a, fig. 28, within which is the osseous envelope b, and at^iew ’ the centre the kernel or true seed c. At an early period both the pulp and stone are observed to consist of cellular tissue; and through the stone the vessels passing from the peduncle are continued. At the base of the figure the letter d denotes one fasciculus of vessels continued through the stone, and turning inward, where it reaches the apex of the seed. These vessels form the umbilical cord or seed-branch of Grew, while the fasciculus e that runs on the opposite side is continued to the flower. In fig. 29 a vertical section of the ovulum, as well as peri¬ carp, is exhibited as it appears at a very early period; in which f denotes the pulp, g the stone through which the umbilical vessels pass and enter the outer tunic h of the ovulum, around which they make a ring. Within this tunic is another, i, filled with cellular tissue; and through its axis a small tube extends, at the apex of which the embryo k is first seen to emerge. 100 Of Fruits. in the pear, by Du Hamel. ANATOMY, VEGETABLE. In fig. 30 these several parts of the ovulum are ex- other instances, to be continued beneath the tunic to the Of 1 hibited on a larger scale; the letter l denotes the outer apex of the seed. (Phys.des Arbres,U)me i. p. 242.) tunic that immediately lines the stone; m the inner one, The last variety we shall notice in theformation of the0v„; corresponding to the chorion of Malpighi; and n the tube seed is that of wheat (triticum),^ given by Malpighuvfe answering to the amnios of the same author. Through In fig. 42 is represented the pistil of the flower of thisMalj the outer tunic Grew represents the umbilical vessels to plant, consisting of the ovarium q, the two styles r, and pass and be continued to the middle tunic, the cavity of the feathered parts that form the stigmata. I revious to which is occupied by large cells that contain a pure lymph, fecundation, the ovarium is found to contain a little At first this tunic is entire, but soon there appears in it vesicle, fig. 43, which is the rudiment of the future ovulum. the small duct n. This duct is not at first wider than a After fecundation the styles soon fall, and the ovarium hair and is dilated at each extremity into an oval cavity acquires a more pointed figure, as in fig. 44. If now it be that contains a pure lymph. A few days after a soft node opened, the little vesicle has changed its appearance, and is seen to emerge in the upper cavity of this tube. This contains within it another smaller vesicle, u, fig. 45. Ihe node (o fig. 31)1s described to possess a conical figure, and ovarium continues to alter its shape, and assumes a more to be another tunic filled with very minute cells. It is at oblong form, fig. 46, and the appearance of styles is now first entire, but when about the size of a carraway-seed quite obliterated. Gradually the little vesicle is formed it becomes a little hollowed near its apex, at which part into a small plantule, convex anteriorly, fig. 47, but more the vessels enter and terminate in another very small node hollowed within, fig. 48, and which is situated at the base (fio\ 32), which is the first appearance of the embryo of the of the ovarium. The two portions thus described in seed. This embryo, when about one fifth part as big as figures 47 and 48 are the minute germ and cotyledon of a cheese-mite, begins to be distinguished by a little fissure, this seed, which are represented in their appropriate place which marks the division of the lobes, as in fig. 33. When in fig. 49, in which the letter x denotes the germ, resting in the lobes have increased, and are more fully formed, the the concavity of the cotyledon ; the albumen that forms node contracts at its base (fig. 34), indicating the place of the chief bulk of the seed ; and z the ovarium, which in the umbilical cord, which subsequently becomes the ra- this seed continues permanent, and forms its outer tunic, dicle of the seed. (Anat. of Plants, p. 209.) (Anat. Plantar, p. 73.) This description corresponds nearly with that of Mai- In his description of the progressive changes exhibit-Oph pighi, as far as regards the situation and general form of edby the impregnated ovarium, Gsertner follows Malpighi6® the tunics, and the place in which the embryo is seen first and Grew, and refers to the figures given by those authors to emerge. It also displays the course of the vessels to as illustrative of them. The liquor, however, in which the form the umbilical cord; but the growth of the embryo embryo is first seen, he confounds with the sac that con- in this seed does not seem to produce the obliteration of tains it, calling it the amnios; but the term amnios desig- some of the tunics in the manner delineated by Malpighi, nates the membrane or sac only, and not the fluid con- An example of a different kind is observed in the pear, tained within it. Sometimes, he says, there is no proper Its structure has been described by Grew, and more mi nutely by Du Hamel. In its mature state it consists of a pulpy matter, in the centre of which are five locula- ments that contain each two seeds. These appearances are exhibited in the transverse section, fig. 35 ; and in the longitudinal section, fig. 36, the seeds are further shown sac for this fluid, but it is contained in the chorion, or in an appropriate cellular tissue. (fDe Fructibas, &c. tom. i. p. 60.) The changes in the ovary during the formation of the seed have likewise been observed by Mr Keith, of B whose account of them agrees with that of Malpighi and Grew. ( System of Physiological Botany, vol. ii. chap. 8.) to be attached by a small umbilical cord. The pulp of Very recently, also, the same subject has occupied the the fruit is made up of a very fine cellular tissue, filled with the proper substance of the fruit, and is everywhere furnished with vessels. Through this pulpy matter a number of solid particles are met with, which are more particularly accumulated at the top and about the core. They are formed of an assemblage of small particles, of a stony consistence, with which a little knot of vessels (fig. 37) is everywhere connected. In fig. 38 is a thin trans¬ verse slice, showing the relative position of these stony particles, as indicated by the knots of vessels with which they are associated. The stony matter is not observed at an early period, but seems to be deposited from the juices in a more mature state. By long maceration in water the pulpy matter is dis¬ solved, and the vascular system is obtained separate. In attention of several foreign writers, and particularly of M. Mirbel, whose observations we shall briefly detail. According to M. Mirbel, little has been done to advance and this part of physiology since the days of Grew and Mal-^“ pighi, till the late observations of M. Schmitz and Mr R. Brown. The latter remarked that we ought not to judge of the structure of the ovulum from that of the developed seed. Acting on this idea, M. Mirbel applied himself to examine the ovarium, from the moment the ovulum begins to appear, and followed its progressive changes of develop¬ ment in plants of the same species. The ovulum at first he describes as a small pulpy excrescence of a conical shape, attached to the containing cell, and which does not appear to have any proper envelope or aperture of any kind. In plants whose ovaria produce many seeds, the the peduncle of the fruit fifteen principal fasciculi of ves- ovula exhibit different degrees of development, according se s are contained. len of these are distributed to the as they are nearer to or more distant from the vascular seeds and flower, and the five others are dispersed through cords that supply nutriment. M. Schmitz, it is said, first the pulp. Ihis vascular structure is represented in fig. discovered in the apex of each ovulum a small orifice or 39 after the removal of the pulpy part; the larger vessels hole, which penetrates the two membranes or sacs, of embrace the core, and after variously ramifying, termi- which, at this early period, the ovulum is composed. To nate in the little vascular processes before described as these sacs, which are placed one within the other, M. Mir- connec e wi i ic sony matter o t ic pulp. In fig. 40 bel has given the names of primine and seconding. Thefni is represented one of the loculaments of the capsule, with orifices at their apex, which at first form but one aperture^ tie seed in it, receiving vessels from fasciculi continued externally, soon become more distinct from the enlarge-dim 1 K Tr G ’ aif' m f an en^ire ®e.e^ 18 rePre_ ment of the parts within, and gradually a third membrane Saf’veslk that ftp11 ° t- If m W \lch the um‘ is seen to protrude through them. No sooner has it be- b vessels that enter at the base are shown, as in come visible in the form of a small pulp, than it begins, ANA fn s. in some species, to dilate interiorly, so as to form a sac v-'V'with very thin sides, which afterwards becomes blended with the secondine: in other species it protrudes to a considerable extent, and in a conical form, through the primine and secondine, which appear to invest it as the cup of the acorn embraces the kernel. In this state it is free at its apex, but is attached at its base to the secon- •n (png. To this sac the name of tercine has been given. These three tunics, it is said, appear together as soon as the development of the ovulum begins; the primine is never absent, and the same is probably the case with the tercine, nor has he ever missed the secondine but in a very few instances. rti,. In addition to the three tunics or sacs just described, iju - two others, named the quartine and quintine, successively appear. The former, says M. Mirbel, is not rare, but has escaped observation by being confounded with the tercine, from which, however, it differs both in its origin and mode of growth. On its first appearance it forms a cellular lining over the inner surface of the cavity of the ovulum. By degrees it separates, and continues attached only to the summit of that cavity. In this state it forms a close sac, and so continues in the ovula of some plants; while in those of others its cavity becomes filled with cellular tissue, and forms a pulpy mass. The fifth envelope, or quintine, is always developed ei¬ ther within the tercine or quartine when they remain filled with cellular tissue. It is first seen in the centre of the sac like a fine tube, and is attached at one end to its summit, and at the other to its base. It begins to en¬ large at the summit, and the embryo is almost simultane¬ ously visible in it. As the enlargement proceeds down¬ ward, the quintine presses at all points on the sac that surrounds it, so as often to push down and occupy the place of the tercine or quartine. From the summit of the ovulum a very fine thread (le suspenseur) descends in the quintine, bearing at its extremity a globule, which is the nascent embryo. This quintine, adds M. Mirbel, answers to the vesicle called amnios by Malpighi, eitnt The foregoing observations were made on the ovula of M several plants, and in their more important points will be !’ seen to correspond nearly with the descriptions of Mal- |,a pighi and Grew. These latter authors have not so minute¬ ly examined the ovulum and its envelopes antecedent to fecundation ; but after that event the chorion of Malpighi may be considered as answering to the lercine or quartine of Mirbel; for in the centre of this chorion the tube e! (fig. 19, Plate XLII.) is seen to extend from the summit to the base, as in the tercine of Mirbel. The subsequent ANA 101 expansion of this tube at its summit (fig. 21), its gradual Of Fruits, enlargement downwards, and the appearance of the em- bryo at its apex as described to occur in the quintine of Mirbel, are all exhibited in the amnios of Malpighi; as is also, in fig. 23, the pressure which this amnios makes on the surrounding sac. In the representations of these appearances given by Grew in figures 28 to 34 inclusive, Plate XLII., a similar series of changes is exhibited; and in what relates to the vascular connection of the ovum with the parent plant, he has anticipated most of the ob¬ servations of those who have followed him. Thus, in fig. 28, 29, Plate XLII., he exhibits the course the vessels take after quitting the peduncle, and shows that they pe¬ netrate the several tunics to reach the point where the embryo, in the form of a small globule (fig. 32), is first seen to emerge; and in fig. 33 and 34 of the same plate he delineates the thread by which it is suspended. According to M. Mirbel, the vessels which form the funiculus or cord enter the ovulum at the point afterwards called hilum in the mature seed, and terminate at a point in the inner tunic or membrane, to which the term chalaza, or internal umbilicus, has been applied. This point or chalaza in the inner tunic corresponds, he says, in the ovula of some plants with that of the hilum in the outer ; but sometimes the vessels after their entrance are prolonged in the manner described by Grew, so that the place of the chalaza is opposite to that of the hilum. Wherever the chalaza may be situated, that part, Mirbel adds, is to be considered as the base of the seed. (i?e- ckerches sur la Structure de V Ovule Vegetal, par M. de Mirbel: Annales des Sciences Naturelles, Juillet 1829.) We have thus described in a general way the elemen- Conclu- tary organs that form the several textures of which the &i°n* vegetable system, in phaenogamous plants, is composed, and given examples of the various modes and proportions in which they combine for that purpose: we have next examined the structure of the mature seed, traced the changes of form observed in its evolution, and described the construction of the several members and organs that are successively produced : and, lastly, we have displayed the structure of the flower, and the formation and de¬ velopment of the ovum, from which a new seed origi¬ nates, fitted to undqrgo and exhibit the same series of changes. The means and conditions required for the accomplishment of these changes, the circumstances in which they act, and the modes of their operation, belong to the department of Physiology, and will form the sub¬ jects of future consideration. (w.) ANAXAGORAS, an eminent philosopher of antiquity, was born in the first year of the 70th Olympiad, or 500 years before Christ. Leaving his lands to be cultivated and enjoyed by his friends, Anaxagoras placed himself un¬ der the care of Anaximenes the Milesian. About the age of twenty he went to Athens and entered upon the study of philosophy, where he continued thirty years. Some suppose that he was the first disciple of the Ionian school, founded by Thales, a teacher of philosophy in Athens. When Anaxagoras assumed the character of a public teacher of philosophy, he quickly rose to high emi¬ nence, and produced many famous scholars, among whom were Euripides, Pericles, and Socrates. This philosopher contented himself with serving the republic in his own station, without interfering in any of the public affairs of hie state. Both by the principles of wisdom which he in¬ spired into the minds of the Athenian youth, and also by his daily advice in the most important affairs, particularly in the case of Pericles, he was of singular service to his country. But neither the friendship of Pericles, nor his own general disinterestedness of character, nor his im¬ mense stores of learning, could ward off the shafts of per¬ secution. Cleo accused him of impiety, and the intro¬ duction of new opinions concerning the gods, because he taught that the sun was a burning mass of stone, or an inanimate fiery substance. By this opinion he was said to rob the sun of his divinity, because in the popular opinion he was deemed Apollo, one of the greatest deities. But although Cleo made religion the avowed cause of the ac¬ cusation of Anaxagoras, it is highly probable that civil causes chiefly operated towards his condemnation. It is, however, abundantly evident that he did not hesitate to expose the vulgar superstitions on several occasions; but the evidence is not sufficient which pretends to prove 102 ANA Anaxago- that he was condemned for teaching the doctrine of a ras supreme intelligence, the creator of this world. His judges . 1- . condemned him to death; but Pericles appearing in his eh us!" defence, the sentence was changed from that of death to banishment and a pecuniary fine. Expelled from Athens, Anaxagoras passed the remain¬ der of his days at Lampsacus, teaching philosophy in the school of his deceased master Anaximenes, until the in¬ firmities of nature terminated his life in the year 428 be¬ fore Christ. Diogenes Laertius has collected, with little care and judgment, the details concerning this philo¬ sopher, which were scattered through various writings. It appears, that in the midst of some extravagant con¬ ceptions, Anaxagoras held opinions which indicate a con¬ siderable acquaintance with the laws of nature. His idea of the heavens seems to have been, that they were a solid vault, originally composed of stones, elevated from the earth by the violent motion of the ambient ether, in¬ flamed by its heat, and by the circular motion of the hea¬ vens fixed in their respective places. The testimonies of several writers, among which is that of Xenophon, unite in proving that he considered the sun to be a large fiery stone; and Xenophon introduces Socrates as refuting that doctrine, and delivering an unfavourable opinion concern¬ ing his other writings. From his perceiving that the rainbow is the effect of the reflection of the solar rays from a dark cloud, and that wind is produced by the rare¬ faction, and sound by the percussion, of the air, he seems to have paid considerable attention to the phenomena of nature. Our information is more correct concerning his opinions of the principles of nature and the origin of things. Fie imagined that in nature there are as many kinds of principles as there are species of compound bodies; and that the peculiar form of the primary particles of which any body is composed is the same with the quality of the compound body itself. For instance, he supposed that a piece of gold is composed of small particles which are themselves gold, and a bone of a great number of small bones: thus, according to Anaxagoras, bodies of every kind are generated from similar particles. That part of his system is more agreeable to reason which explains the active principle in nature. According to Diogenes Laer¬ tius, Anaxagoras taught that “ thg universe consists of small bodies composed of similar parts, and that mind is the beginning of motion.” “ He was the first,” says the same writer, “ who superadded mind to matter, opening his work in this pleasing and sublime language: ‘ All things were confused ; then came mind and disposed them in order.’ ” Plato informs us that this philosopher taught the existence of a disposing mind, the cause of all things. Anaxagoras, according to Aristotle, taught that mind was “ the cause of the world, and of all order; and that while all things else are compounded, this alone is pure and un¬ mixed and that “ he ascribes to this principle two pow- ers—to know and to move—saying that mind put the universe in motion.” Cicero expressly asserts that Anax- agoias was the first who taught that the arrangement and order of all things was contrived and accomplished by the understanding and power of an infinite mind.” ANAXARCHUS, a Grecian philosopher, who lived under Philip of Macedon and Alexander, was born in Ab- dera, and belonged to the sect generally known by the name of the Eleatic. He is said to have been conducted in the progress of his early studies by the skilful hands of Diomenes of Smyrna and Metrodorus of Chios. He had the honour to be a companion of Alexander; and a few anecdotes transmitted to posterity concerning him render it evident that he treated him with the usual freedom of ANA a friend. This philosopher candidly checked the vain-Am glory of Alexander, when, elated with pride, he aspired to the honours of divinity, by pointing to his finger when it bled, saying, “ See the blood of a mortal, not of a god.” , It is likewise reported, that, on another occasion, while in-^, dulging immoderately at a banquet, he repeated a verse from Euripides, reminding Alexander of his mortality. It is, however, to be regretted, that the fidelity of the philo¬ sopher was wanting at the time when the mind of Alexan¬ der was tortured with remorse at having slain his friend Clitus; for it is reported that he on that occasion endea¬ voured to soothe the agitated mind of Alexander, by saying, that “ kings, like the gods, could do no wrong.” It is reported that Nicocreon, king of Cyprus, exposed him to the torture of being pounded in a mortar, and that he endured this torture with incredible patience; but as the same fact is reported of Zeno the Eleatic, there is reason to suppose that it is fabulous; and it may be added, that this narrative is inconsistent with the general character of Anaxarchus, who, on account of his easy and peaceable life, received the appellation of “ The For¬ tunate.” ANAXIMANDER, a famous Greek philosopher, born at Miletus in the 42d Olympiad, in the time of Polycrates, tyrant of Samos. Fie was the first who publicly taught philosophy, and wrote upon philosophical subjects. He carried his researches into nature very far for the time in which he lived. It is said that he discovered the obliquity of the zodiac, wras the first who published a geographical table, invented the gnomon, and set up the first sun-dial in an open place at Lacedemon. He taught that infinity of things was the principal and universal element; that this infinity always preserved its unity, but that its parts underwent changes; that all things came from it; and that all were about to return into it. According to all appearance, he meant by this obscure and indeterminate principle the chaos of the other philosophers. He asserted that there is an infinity of worlds ; that the stars are com¬ posed of air and fire, which are carried in their spheres, and that these spheres are gods; and that the earth is placed in the midst of the universe, as in a common cen¬ tre. He added, that infinite worlds were the product of infinity, and that corruption proceeded from separation. ANAXIMANDRIANS, in the history of philosophy, the followers of Anaximander, the most ancient of the philosophical atheists, who admitted of no other substance in nature but matter. ANAXIMENES, an eminent Greek philosopher, born at Miletus; the friend, scholar, and successor of Anaximan¬ der. He diffused some degree oflight upon the obscurity of his master’s system. He made the first principle of things to consist in the air, which he considered as im¬ mense or infinite, and to which he ascribed a perpetual motion. He asserted that all things which proceeded from it were definite and circumscribed; and that this air, therefore, was God, since the divine power resided in it and agitated it. Coldness and moisture, heat and mo¬ tion, rendered it visible, and dressed it in different forms, according to the different degrees of its condensation. All the elements thus proceed from heat and cold. The earth was, in his opinion, one continued flat surface. Anaximenes, a Greek historian and rhetorician, was born at Lampsacus about 580 years before Christ. Some writers ascribe to him the Treatise on the Principles of Rhetoric, which bears the name of Aristotle; and it is reported that Philip of Macedon invited him to his court to instruct his son Alexander in that science. He at¬ tended Alexander in his expedition against Persia. The history of Philip, of Alexander, and likewise twelve books A N C nt on the early history of Greece, were the productions of |j his pen, but none of them has been preserved, iclf- ANBAR, a town of Asia, in Arabian Irak, situated v’“An the Euphrates. It was taken by Caled, lieutenant of the caliph Omar, in the year 632, and it was rebuilt by the first caliph of the Abapides. It is 35 miles west of Bagdad. Long. 43. 2. E. Lat. 33. 15. N. It is also the name of a town of Great Bukharia, in the province of Balk, 70 miles S. S. W. of Balk. Long. 64. 18. E. Lat. 36. N. ANBERTKEND, in the eastern language, a celebrat¬ ed book of the Bramins, wherein the Indian philosophy and religion are contained. The word in its literal sense denotes the cistern wherein is the water of life. The An- bertkend is divided into 50 beths or discourses, each of which consists of 10 chapters. It has been translated from the original Indian into Arabic, under the title of Moral al Maani, q. d. the marrow of intelligence. ANCARANO, a town of Italy, in the marquisate of Ancona. Long. 14. 54. E. Lat. 42. 48. N. ANCASTER, a town of Lincolnshire, situated in long. 30. W. lat. 52. 30. N. It gives title of duke to the noble family of Bertie. ANCENIS, an arrondissement in the department of Morbihan, in France. The extent is 310 square miles, or 197,120 acres. It is divided into five cantons, and these into 28 communes, having a population of 40,992. The chief place, of the same name, contains a population of 3295 inhabitants. ANCESTORS, those from whom a person is descended in a straight line. The word is derived from the Latin ancessor, contracted from antecessor, goer before. Many nations have paid honours to their ancestors. It was properly the departed souls of their forefathers that the Romans worshipped under the denominations of lares, lernures, and household gods. Hence the ancient tombs were a kind of temples, or rather altars, whereon oblations were made by the kindred of the deceased. ANCHISES, in fabulous history, a Trojan prince, de¬ scended from Dardanus and the son of Capys. Venus made love to him in the form of a beautiful nymph, and bore him Avneas, the hero of Virgil’s iEneid. ANCHOR, in Navigation, from the Greek ayxvoa, which Vossius thinks is from oyxri, a crook or hook, an instrument of iron or other heavy material used for holding ships in any situation in which they may be required to lie, and prevent¬ ing them from drifting by the winds or tides, b}r the currents of rivers, or any other cause. This is done by the anchor, after it is let down from the ship by means of the cable, fixing itself into the ground, and there holding the vessel fast. The anchor is thus obviously an implement of the first importance in navigation, and one on which too much attention cannot be bestowed in its manufacture and pro¬ per construction, seeing that on it depends entirely the lq ;y safety of the vessel in storms. The invention of so necessary no rs. an instrument remounts, as may be supposed, to the remot¬ est antiquity. The most ancient anchors consisted merely of large stones, or baskets full of stones, or sacks filled with sand, or logs of wood loaded with lead. Of this kind were the anchors of the ancient Greeks, which, according to Apollonius Rhodius and Stephen of Byzantium, were form¬ ed of stone; and Athenaeus states that they were some¬ times made of wood. These sorts of anchors retained the vessel merely by their inertia, and by the friction along the bottom. Iron was afterwards introduced for the con¬ struction of anchors, and also the grand improvement of forming them with teeth or flukes to fasten themselves into the bottom; whence the words odovrig and dentes are frequently taken for anchors in the Greek and Latin poets. The invention of the teeth is ascribed by Pliny to A N C 103 the Tuscans; but Pausanias gives the merit to Midas, king Anchor, of Phrygia. Originally there was only one fluke or tooth, whence anchors were called Irz^oGTogot; but shortly after¬ wards the second was added, according to Pliny, by Eupa- lamus, or, according to Strabo, by Anacharsis the Scythian philosopher. The anchors with two teeth were called afjjtpiZo'hoi or agpiGTogoi, and from ancient monuments appear to have been much the same with those used in our days, only the stock is wanting in them all. Every ship had se¬ veral anchors, the largest of which, corresponding to our bower or sheet anchor, was never used but in extreme danger, and was hence peculiarly termed Uga or sacra; whence the proverb sacram anchoram solvere, as flying to the last refuge. Anchors are now universally made of wrought iron, ex- Weight of cepting in Spain and some parts of the South Sea, where anchors of they are made of copper. One essential quality in every different anchor is a sufficient weight to fix itself in the bottom ; and s^zes- this has been determined by practice for different an¬ chors, and for vessels of different sizes. Large vessels have several anchors of different sizes, and stowed in different parts of the vessel. These are distinguished by different names, viz. the sheet-anchor, the largest of all, and which is only let down in cases of dan¬ ger, or when the vessel is riding in heavy gales of wind. The bower-anchors, of which there are sometimes two or three, are nearly of the same size with the sheet, and are distinguished into best bower, small bower, and spare an¬ chor. The stream-anchor is of a much smaller size than the above, used only for riding in rivers or moderate streams. It is not generally above one fourth or one fifth of the weight of the others. Lastly, the hedge-anchor is still smaller, being only about one half of the stream- anchor, and is only used when hedging in a river. In ships of war the sheet-anchor is stowed upon the after-part of the fore-channel, on the larboard side, with the stock vertical, and one of the flukes resting on the gang-way. The bower-anchor hangs to the cat¬ head, with the other extremity fixed up to the anchor- boards ; and the spare anchor is stowed away on the star¬ board fore-channel. Ships of the largest class carry seven or eight anchors, and the smallest class, as brigs, cutters, and schooners, three or four. The weight of anchors for different vessels is allowed by the tonnage. A pretty near rule for the principal an¬ chor of ordinary-sized vessels is to allow for the cwts. in the anchor one twentieth of the tonnage. Thus a vessel of 400 tons would require her principal anchor to be 20 cwts., or according to the following tables:— Merchant Vessels. Tonnage. Weight of Anchor. 20 tons., 35 50 65 70 80 100 130 160 190 220 250 280 320 380 430 480 520 720 1 cwt. If n 3i u 5f 7 8 9i Vessels of War. Weight of Anchor. 20 guns 25 cwt. .12 .14 • 151 .171 .20 .21 .25 .33 24. 28. 32. 36. 38. 40. 44. 50. 64. 74. 80. 90. 100. .30 .31 .35 .40 .44 .46 .48 .50 .57 .68 .76 .84 .90 104 ANCHOR. Anchor. Next to the weight, the form of the anchor, and the proportions of the different parts, are of great importance. Form and The most general form, and that which has indeed been dimensions almost uniVersally adopted all over the world, is that re¬ ef anchors. nted at plate XLIII. fig. 1, and in section at Plate XLIV. fig. 1 and 3, consisting of the two hooked arms tor penetrating and fixing themselves into the soil; the long bar or shank for attaching the cable ; and the stock, which is attached to the extremity of the shank, and serves to direct one of the points downwards into the soil. The weight of the anchor then causes the point to penetrate more or less according to the softness or hardness of the bottom ; and the action of the vessel on the cable, instead of loosening the anchor, tends rather, by the hooked shape of the arms, to fix these deeper and firmer into the soil; so that the vessel is held quite fast, unless either the cable itself gives way, or any part of the anchor, or the. anchor is dragged along owing to the looseness of the soil. The shank of the anchor is made long, in order that the stock may have the greater power in directing either the one or the other of the arms downwards. It also serves, when the anchor is to be weighed, as a lever for starting it out of its place ; the cable drawing upwards by the extremity, and turning the whole round the point of the fluke. The one end of the shank is made square, to receive and hold the stock steadily in its place without turning. To keep the stock also from shifting along the shank, there are raised on it from the solid iron, or welded on it, two square tenon-like projections, called nuts. The length of the square of the shank is about one sixth of the whole length of the shaft, and the thickness about one twentieth. From the end of this square the shank increases in thickness, tapering towards the extremity, where the arms are at¬ tached : in all this part it is either made wholly round, or with a flat on opposite sides, or polygonal. The end next the stock is called the small round. The other extremity, where the arms and the shank unite, is called the crown, and the point of the angle between the arms and the shank the throat. Here the thickness of the shank is from l^ inch in small anchors, to 3 inches in large ones, greater than at the small round. A distance equal to that between the throat of one arm and its bill is marked on the shank from the place where it joins the arms, and is called the trend. Near the extremity of the square part of the shank is the hole for receiving the ring for the cable, which is about half the thickness of the small round, and the dia¬ meter nearly equal to the length of the square. The ring used to be invariably lapped with cordage, to prevent the cable from chafing against the iron; but since the in¬ troduction of iron cables, this has become no longer ne¬ cessary. The arms make an angle of about 56° with the shank. They are made either round or polygonal like the shank, about half their length. The remainder of the arm consists of three parts, the blade, the palm, and the bill, Plate XT .TV. fig. 1, and in another view at fig. 2. The blade is merely the continuation of the arm in a square form. The palm or fluke is a broad, flat, triangular plate, fixed on the inside of the blade, the use of which is, by exposing a broad sur¬ face, to take a firmer hold of the ground. The bill is the extremity of the arm, where it is tapered nearly to a point, for the purpose of penetrating more readily into the soil. In some cases the arm is made quite straight from the crown to the bill; in others, and particularly in small anchors, the interior half is made with the arch of a circle, as at fig. 3. The whole length of the arm is nearly half the length of the round part of the shank. It tapers slightly from the throat to the blade, where it is about the same thickness with the small round of the shank. The palm is about one third of this in thickness, and the At| l# breadth of its base is nearly equal to its length. ^ /v* The stock of the anchor, represented at fig. 4, is made of oak, consisting of two beams embracing the square, and firmly united by iron bolts and hoops. The length of the stock is rather greater than that of the shank, the thick¬ ness in the middle about one twelfth of its length, and tapering to about the half of this at the extremities, the taper being all on the under surface next the arms, and the other quite straight. The taper is not quite regular. It commences at about half the breadth of the stock from the shank, and continues in one straight line to the ex¬ tremity. The beams of the stock are hooped close toge¬ ther at the extremities, but gradually open towards the centre, that, in case of the wood shrinking, the hoops may be driven farther in, fig. 5. Of late years the stock has frequently been made of wrought iron, the same as the anchor; and this plah is now very generally followed, particularly with smaller anchors. It has this advantage, that the stock can be at any time taken out and laid paral¬ lel with the anchor, which is very convenient for stowage. The iron stock, fig. 6, consists merely of a long round bar, about half the diameter of the anchor at the square. Instead of embracing the anchor, like the wood, it goes through a hole in the square, which is swelled out to re¬ ceive it. It has a shoulder in the middle, which rests against the square, and a key driven through a hole in the stock on the other side keeps it fast. When the stock is to be taken out of its place, the key is driven out: the stock then slides through the hole in the shank, and by means of a bend at its extremity, it is laid parallel with the shank. The operation of the anchor is easily understood. Being let down by means of the cable, the weight of the arms throws them downwards, and keeps the whole in a vertical position until it reaches the ground, where it lights upon the crown; and then falling over, the position of the stock at right angles to the arms, and its length and height, together with the weight of the cable, are sure to throw it with one of the arms pointed into the ground, if it does not take this situation of itself. This effect is aided by the anchor descending quickly— and hence it must be allowed to descend freely ; for which purpose, in throwing or casting the anchor, the cable is arranged along the deck in long coils, called by the sea¬ men a French flake, one end being attached to the an¬ chor and the other secured on deck. Every thing being prepared, the lashing of the anchor is cast off, and the men stand ready to let go; and when this word is given by the person in command, the fastenings are all cast off, and the anchor, falling into the sea, descends with rapidity, and draws the cable after it so violently, that it is often neces¬ sary to throw water on the hause holes to prevent their taking fire. When the anchor, again, is to be removed from its situation and drawn up into the vessel, the operation is termed weighing; which requires often a very heavy purchase, particularly at starting. This is obtained by means of the windlass, round which the cable is wound, and a number of hands applied to work it. With cables which are too large to be wound round a windlass, a small¬ er rope is used, termed a messenger, which being attached to the cable at different points, and wound round the windlass, serves to bring the cable forward. But since the introduction of chain-cables this contrivance is not so much required. V hen the anchor is brought above water, a tackle is got upon the shank, just within the flukes, and the arms are hove up so as to lie upon the gunnel and anchor-boards; the stock is then made vertical by heaving upon it with a non. 105 A N C \rt(y tackle, in which position it is secured by the stock-lashing. yV The ring is fastened to the cathead by the stopper, one end of which is fastened round the cathead, and the other is brought through the ring, then over the stopper-cleat, and is belayed round a timber head. To secure the shank Besides anchors of the common construction, there are various others of different forms occasionally in use. Small vessels often employ what are termed grapnels, which are merely common anchors with four or more arms instead of two, as shown in fig. 7. Following out the same principle, we have the mushroom anchor, fig. 8, much employed in the East Indies, to secure the vessels which they term grabs. In this the arms are continued in one segment of a sphere all round; it hence requires no stock, as it takes the ground in any direction. Attempts have frequently been made to introduce anchors with only one arm, but hitherto without any decisive result. A patent for an anchor of this kind, as represented at Plate XLIII. fig. 2, was taken out by Mr Stuard, which has attracted some notice. “ In order,” says he, in the Specification of his patent, “ that this anchor may be sure to fall the right way with the fluke downwards, I would have the shank very short, whereby, when suspended by the cable, it will cant the most, and when it has hold in the ground, the ship will ride safer; as a long shank has more power to loosen and break the ground, and is more likely to be bent or broken from its hold. Let the form of the shank and arm of the anchor be as AA, fig. 2; and, that the parts may be stronger than if made separately and shut together, I would have the bars which compose them in one length, so that there be no weld or joining in the whole length of the shank and arm. The hole B is to receive the ring for the cable, and the hole Cis for the stock, which is composed of a wrought-iron bolt, as A, fig. 3, covered with cast-iron at its ends, BB. The palm to be in shape as D, fig. 2, and shown detached in fig. 4, made either entirely of cast- iron, or a cast-iron shell filled with lead, which is of much more specific gravity than iron. The back of the palm to he formed either with concave surfaces or flat surfaces, making angles at the centre. The anchor is also to have a small shackle, fixed on the bend of the shank and arm, as at E, fig. 2, for the buoy-rope to be made fast to. The shank may be made without the hole C, and the hole B made octagonal; or if round, it should have a small fillet projecting from the stock, and a small cavity on one side vol. m. at the arms, a chain called the shank painter-chain is pass- Anchor, ed round it, and fastened to a timber head. The following table contains the dimensions of the parts of anchors of different weights. The letters g. l. denote the greatest and least diameters or breadths. of the hole B to receive it, thus to prevent the stock from turning round ; and instead of a ring for the cable, to have a shackle fitted on the stock, on each side of the shank; and, that the shackle may not turn on the stock and fall too low, a stop is to be fixed on each side at the upper end of the shank.” (See Repertory of Arts, &c. vol. v.) Mooring Anchors axe those which are fixed in certain New moor- situations in harbours or roadsteads, and to which any ofing anchor, the vessels frequenting the place may be secured. As these are no way limited as to weight like portable anchors, they often consist merely of a large block of stone, such as at fig. 9, with an inner ring fixed in the middle of the upper side ; or several such stones may be fastened together so as to act as one mass. Mooring anchors are also often made by choosing one of the largest anchors used for first-rate ships, weighing 80 cwt., and by bending one of the arms close down upon the shank, to prevent it catching the cable or mooring chains whilst the ships are riding. These anchors are lowered down into the water with a very strong iron mooring chain fastened to the ring, to which the ships are fastened: they are usually made from such as are damaged in one of the flukes or arms. A new kind of mooring anchor of cast-iron was described by Mr Hemman of Chatham, to .the Society for the Encourage¬ ment of Arts, 8$C', in 1809, for which he obtained a silver medal from the society. AA, fig. 5, Plate XLIII., repre¬ sents the palm or heavy part of the anchor, made very massive of cast-iron, and of considerable breadth, so that the edge B, or part which enters the ground, may have a great hold; the shank C is made also of cast-iron, and fixed firmly to the head by passing through it, and has a small ring at a, where the buoy-rope is fixed; the other end of the shank goes through the stock DD, which is formed of two large wooden beams hooped together in the same manner as the stocks for common anchors; the end of the shank projects through the stock, and has a strong wrought-iron shackle E fixed to it by a bolt passing through both, and with this the mooring chain is connect¬ ed. The great advantage of this over the common moor¬ ing anchors arises from its great weight and breadth of o 106 A N C Anchor, edge to act against the ground; and, being made of cast- iron, it is also more durable. A pair of these anchors, weighing 150 cwt. each, will, with the mooring chains, cost about L.874 less than a pair of the common anchors, which, with their chains, cost L.2472. (See Transactions of the Society for the Encouragement of Arts, Sfc. vol. xxvm.) Another form of this mooring anchor is shown at fig. 10. Floating This is the name given to a sort of anchor which has anchor. often been proposed, but never reduced to practice, for preventing a vessel from drifting, in cases where the great depth of the sea precludes the use of the cable and ordinary anchor. The plan suggested by Dr Franklin seems the most rational. This anchor consisted of two cross bars, secured together in the middle, and having sailcloth fastened to them in the shape of a parallelo¬ gram. To the centre of these bars the cable was at¬ tached, and being thrown overboard, it was thought the resistance of so large a surface would at least check the rapidity of the ship’s motion. Fabrica- In fabricating anchors the different parts are first forged tion of an- separately, as the square, the ring, the shank, the arms, and chocs. t^e paims The square is then welded on to the extre¬ mity of the shank, and the ring let through its hole, and then the joining welded. The arms are then welded on to the shank, and the palms to the arms. The forging of such heavy masses of iron as the larger anchors are com¬ posed of, and the welding of the parts together, is one of the most difficult operations in smithery, and one which it has required all the ingenuity of our most skilful artists, and a long course of experience, to bring to its present perfection. The shank forms the largest mass. It is formed of an assemblage of smaller bars, united together by welding them into solid masses. This mode is prefer¬ able to making a single bar of sufficient size by the forge- hammer in the original preparation of the iron, because the compounded bar is not liable to have any internal flaws or cracks, or at least they will not be in a transverse direction; for the bars are all examined separately before uniting them, and any which are imperfect are rejected; if, therefore, after the welding, any cracks are left between the bars, they must be in the length of the anchor, and will not deduct so materially from the strength of the whole. Formerly the small bars were laid together, so as to form a square, as in fig. 11; and round these, smaller bars were piled to complete the circle ; and then this circle was sur¬ rounded by bars to the number of 30 or 40, all arranged round the circle, and packed together like the staves of a cask; iron hoops were then driven over all to keep them to¬ gether, and the whole faggot thus united was welded by re¬ peated heats and elaborate hammering, applied successive¬ ly from the one end to the other of the bar. Great difficulty was always found in heating and welding together the inside or central bars along with the external ring, the latter becoming quite overheated before the former could be brought up to the welding point. In these circum¬ stances it became very difficult to weld the bar equally, so as to give it all the strength of which the material was susceptible; and it was often thought sufficient to weld merely the outside ring, so as to form a perfect covering or case for the inside bars, which were left in a raw or un¬ welded state. A more perfect method of welding the heavy bars of the anchor together was introduced some years ago at Plymouth, by Mr Perring, clerk of the cheque. This consists in using bars the whole breadth of the shank laid on one another, as at fig. 12, and hooped. These are then welded together in two heats, until the whole is one compact body, which is readily done in this manner, without overworking any part of the iron. If the bars do not extend the whole breadth, they are laid so as to break joint. H O R. The arms are formed and welded together into a mass An in the same manner as the shank. They are then united^! to the extremity of the shank, by shutting or welding them all together at the crown, for which purpose the end of the shank is formed with a flatted surface or scarf, having a shoulder on each side, and the ends of the arms with similar scarfs, as at fig. 13. A more perfect method of uniting the arms and the shank, by Mr Perring, is, to form the crown se¬ parately of bars similarly to the shank, uniting them into one for half the length only, and the other half remaining split in two, as at fig. 14 ; then opening this by means of two rods, welded on the ends on purpose, bending them round two steady pins, as at fig. 15, so as to form part of the arms, as at fig. 16. The angular opening being then filled up by a piece formed of bars properly welded, another piece half the thickness of the arms is bent over and unit¬ ed with the others, so as together to form the crown and part of the arms on each side. In this manner a much firmer joint is formed than by mere scarfing. The crown piece being thus formed, the shank is welded to one ex¬ tremity, and the two arms and the blades to the two others. The blades of the arms are formed in a manner similar to the shank. The palms are made by laying a number of plates one on another, and welding them all properly to¬ gether, after which the palms are shut on upon the blades. After the different parts of the anchor are thus forged and welded together, they are all smoothed and finished off by working with hammers till the metal becomes quite cold. This process, besides improving greatly the appear¬ ance of the anchor, is of real use in hardening the surface, and causing it to withstand better the corrosion of the sea¬ water. The following is a more particular account of the opera¬ tions of the anchor-smith on a large scale. The hearth AA of the anchor-smith’s forge, see fig. 6, Plate XLIIL, is built of brick-work raised about 6 or 9 inches above the ground, and 6 or 7 feet square ; in the centre is a large cavity, to contain the fire ; at the back of the hearth a vertical brick wall B is erected, supporting and forming one side of the chimney, which is little more than a dome placed over the hearth, and opening at the top with a low' chimney to carry off the smoke. Behind the wall the bellows CD are placed; the noses of the pipes being about the level of the hearth, and coming through the wall, which at that part is defended from the action of the fire by a facing of fire-stone. In this fire-stone the tue-iron is fixed ; it is a tube made of wrought iron, and very thick in the sub¬ stance, that it may not burn awray in the fire: the pipes of the bellows are inserted in the tue-iron, and thus con¬ vey the stream of air into the centre of the fire. The bellows are not like those which ordinary smiths make use of; but two large pairs of single bellows CD are placed horizontally by the side of each other, the pipes of both being inserted into the same tue-iron, and directed to blow to the same focus in the centre of the fire: these bellows are exactly like those in use for domestic pur¬ poses, which only throw out air when the upper board is pressed down. The two are worked alternately by means of chains c d attached to the ends of the upper boards, and united to the end of the working level's HI, placed over each pair of bellows. From the opposite extremities of these levers other chains ef are extended to the op¬ posite side of a long lever GG, which moves upon the pivots of a vertical axis E, and is loaded at the ends by heavy weights, to give it momentum. Now, two or more men pushing in opposite directions can give it a motion backwards and forwards, and by the communication of the chains and upper levers HI, they will alternately lift up the upper boards CD of the bellows, which being sufficiently loaded, will subside themselves, and force their contents $ T A N C hr of air into the fire. The men who work the lever G are >vVaided by six or eight more, who place themselves upon the board of one pair of bellows, and as soon as it subsides, they step upon the other pair, which also sinks, and then they return: they have ropes suspended from the roof to enable them to lift themselves, and mount from one bel¬ lows upon the other with more ease. The common tue- iron, which is simply a cone of wrought-iron, set with clay into fire-stone, composing the back of the hearth, is very soon burnt by the great heat. The most improved forges, therefore, are now furnished with what is called the water tue-iron, which is made hollow, and water introduced into it to keep it cool. For this purpose two cones are form¬ ed of thick iron plate, each with a small aperture at the vertex; these, when put one into the other, are welded together at their bases and their points, so as to form one cone, which is hollow, with a small space all round; two pipes communicate with the hollow, one bringing a con¬ tinual supply of cold water, and the other conveying away that which is heated by the fire. By this means the tue- iron is kept cool, and can never acquire such a degree of heat as to be burned away: this tue-iron is set with fire¬ clay into a frame of cast-iron, built up in the brick-work of the wall B. The anvil K is only a cubic block of cast-iron, placed on the ground much lower than the ordinary smith’s anvil; because, as the anchor-smiths always strike by swinging their hammers over their heads, at arms’ length, they have more force when the work lies low on the ground than if raised up. At a distance of eight or nine feet from the hearth AA a strong crane-gib LM is erected, so as to turn freely upon the vertical post M. It has no tackle, but the upper beam L, which must be horizontal, has a large iron loop n hung upon it, with a roller o, which admits it to run freely backwards and forwards upon the beam: the lower end of the loop suspends the anchor; therefore, by mov¬ ing the rollers along the beam of the gib, and by turning the gib round on its pivots, the anchor can be placed in any position in the fire or upon the anvil. To give mo¬ tion to the roller o, a rack p is connected with it; and this is moved by a pinion upon the axis of the wheel t, which has an endless rope hanging down, so that a la¬ bourer can reach it, and thus remove the anchor nearer or farther from the centre, however great its weight may be. The workmen employ scarcely any other tools than their sledge-hammers, and a few large punches, cutting chisels, and sets or prints, which, when urged by the hammers, will give any particular figure to the work: the hammers are of the largest kind, and weigh from 14 to 18 pounds, according to the strength of the workmen. In the Royal dock-yard great use is made of a stamping machine, which the workmen call Hercules, and which is very similar to the machine for driving piles. A heavy iron weight N, guided like the ram of the pile engine, is drawn up by the strength of several men, and let fall upon the anchor, to weld the bars, in the same manner as by a forge-hammer. The machine is erected on a large block of stone, which supports the anvil O: two square iron bars PP are fixed on each side of the anvil, in a vertical position, the angles of the bars being placed towards each other. These vertical bars are eight or nine feet high, and are fixed at the top to a beam in the roof of the building in which the machine is placed. The ram N, which weighs 47} cwt., is fitted to slide up and down be¬ tween the bars P, having notches in its sides, which re¬ ceive the angles of the bars: it is drawn up by a rope passing over an iron pulley Q, mounted upon pivots above the top of the vertical bars ; and the rope has eight or ten small ones 11 spliced into it, for as many men to act to¬ gether (which they do by a motion similar to that of 107 ringing), to elevate the ram, and let it fall upon the iron Anchor, placed upon the anvil O. The Hercules is placed in the'^-^v~^->' same sweep of the crane as the anvil K, so that the iron can be conveyed to either with equal ease. The first step in making the different parts of the an¬ chor is to assemble or faggot the bars. For the centre of the mass which is to make the shank, four large bars are first laid together; then upon the flat sides of the square so formed smaller bars are arranged, to make it up to a circle. The number is various, but in large anchors six or eight bars are laid on every side. This circle is sur¬ rounded by a number of bars arranged like the staves of a cask: as many as 36 are often used, and they form a complete case for the others. The ends are made up by short bars to a square figure. The faggot is finished by driving iron hoops upon it at sufficient distances; see W in the figure; and it is suspended from the crane in such a manner that it can be moved and turned in any direc¬ tion, by only one or two men, even when it weighs three tons. For this purpose an iron pulley k is hooked to the iron loop n of the crane ; and a short endless chain l passed over the pulley suspends the faggot in its loop. In this manner the weight of the iron is in reality borne by the pivot of the pulley k, and the mass can be easily turned round upon its centre to bring any side upwards. To give a power to the man who guides it, one of the four central bars is double the length of the faggot, and projects, see g} to form a long lever, by which it is steered; and two holes are made through the end of this bar to insert a cross lever A, by which the faggot is turned or rolled round upon its centre. As the faggot hangs very nearly on a balance in the loop of the chain l, the man, by weighing on the end of the long bar g, can easily raise up its end from the anvil K, and, swinging the crane on its pivots, move it into the fire, which is made up hollow like an oven. To effect this form, the fireman first spreads the coals evenly upon the hearth, and with his shovel or slice makes a flat surface about the level of the tue-hole: he then arranges some large cinders or cakes round in a circle upon this surface, and by other cinders builds it up like an oven or dome, leaving a mouth to introduce the iron. The oven is adapted in size to the magnitude of the mass of iron, and must be brought forwards upon the hearth, to leave a space between its interior cavity and the orifice of the tue-iron; in which space a passage is made from the tue-hole to the fire, and filled up with large lighted coals, and then covered up by small coals. The blast from the bellows passes through these hot coals, in order that the cold air may not enter the fire at once and blow on the iron, but be first converted into flame, which is urged forcibly into the oven, and reverberated from the roof and sides upon the iron placed in the centre. As the floor of the oven is nearly upon a level with the tue-hole, the flame from the coals between it and the fire also plays upon the bottom, and thus heats the iron on all sides. The outside of the dome is covered over with a considerable thickness of small coals, which cake toge¬ ther, and, as the inside of the oven consumes, settle down into a dome again, which the smith aids by striking the outside with the flat of his slice. If the Are breaks out at any place in the roof, the smith immediately repairs the breach with fresh coals, and damps them with water, that they may not burn too fast; for if the inside of the oven burns very fiercely, the flames will not be reverberated so forcibly as when it is in the state of burning cake. Care must likewise be taken to prevent the fire from burning back to the tue-iron. The mouth of the oven should be made no larger than to admit the work; and, that as little heat as possible may escape by the iron, the mouth is filled round it with coals. F is an iron screen hung on H O R. 108 Anchor. ANCHOR. hinges, to swing before the mouth of the fire when the iron is withdrawn, that the workmen may not be scorched by the heat. All the men unite to assist in blowing the bellows, which they work in the manner already described, from half an hour to an hour, according to the size of the an¬ chor, until they have raised the iron to a good welding heat. The mouth of the fire is opened occasionally to in¬ spect the process, and the faggot is turned in the fire if it is not found to be heating equally in every part. Eight men, and sometimes more, are employed to forge an an¬ chor: six of them strike with the hammers, one is sta¬ tioned at the guide-bar, and the eighth, who is master or foreman, directs the others, and occasionally assists to euide the anchor. When the whole of that part which is in the fire comes to a good welding heat, the workmen leave the bellows and take up their hammers; the coals are removed from the iron, w hich is swung out of the fire by the man who guides it, assisted by others, and the hot end placed on the anvil; during which time one or two labourers with birch brooms sweep off the coals which ad¬ here to it. The smiths now begin hammering, one half the number standing on one side, and the other half on the other: they use large sledges weighing from sixteen to eighteen pounds, and faced with steel, striking in regular order, one after the other, swinging the hammers at arms’ length, and all striking nearly at the same place. The foreman places himself near the man who guides, and with a long wrand points out the part he wishes them to strike, and at the same time directs and sometimes assists the guide to turn the faggot round, so as to bring that side uppermost which requires to be hammered. This is continued as long as the metal retains sufficient heat for welding. This process is exceedingly laborious for the workmen, and is much more effectually performed by means of the Her¬ cules, which strikes such powerful blows upon the iron as to consolidate the bars much more than the strokes of small hammers can do, however long they may be conti¬ nued. When the iron has lost so much of the heat that it will no longer weld, the foreman takes a number of pins, made like very thick nails without heads: one of these he holds in the end of a cleft stick, places its point upon the iron, and two smiths, with their sledges, strike on it with all their force, to drive it through the bars ; but this they must do quickly, or the pins will become hot and soft, so as not to penetrate the bar. These pins are in¬ tended to hold the whole together more firmly, and, by swelling out the sides, to fill up any small spaces there may be between the bars. The iron is now returned to the fire, another mouth being opened on the opposite side of the oven, to admit the end or part which has been welded to come through, that a part farther up the faggot may be heated; and when this is done the welding is performed in the same manner as before. Thus, by repeated heat¬ ings, the faggot is made into one solid bar of the size and length intended. It is then hammered over again at weld¬ ing heats to finish it, and make an even surface; and in this second operation the workmen do not leave off ham¬ mering as soon as the iron loses its full welding heat, but continue till it turns almost black. This renders the sur¬ face solid and hard, and closes all small pores at which the sea-water might enter, and by corroding the bars, expand them, and in time split open the mass of iron. The shank for an anchor is made larger at the lower end, where the arms are to be welded to it, and is of a square figure. A sort of rebate or scarf s is here formed on each side of the square, in order that the arms may apply more properly for welding. This scarf is made in the ori¬ ginal shape of the faggot, and finished by cutting away some of the metal with chisels whilst it is hot, and using k sets or punches, properly formed, to make a square angled to the shoulder of the scarf. The upper end of the shank is likewise square; and the length between these square parts is worked either to an octagon or round, tapering re¬ gularly from the lower to the upper end. The hole to re¬ ceive the ring of the anchor is pierced through the square part at the upper end, first by a small punch, and then larger ones are used till it is sufficiently enlarged. The punch is made of steel; and when it is observed to change colour by the heat, it is struck on the opposite end to drive it out, and is instantly dipped in water to cool it, and another driven in. The projecting pieces or nuts, which are to keep the stock or wooden beam of the anchor, and its place on the shank, are next welded on. To do this the shank is heated, and at the same time a thick bar is heated in another forge : the end of this is laid across the shank, and the men hammer it down to weld it to the shank; then the piece is cut off by the chisel, and another piece welded on the opposite side. Whilst this process of forging the shank is going on, the smiths of another forge, placed as near as convenient to the former, are employed in making the arms, which are made from faggots in the same manner as the shank, but of less size and shorter: they are made taper (seeX), one end of each being smaller than the other: the larger ends are made square, and cut down with scarfs, r, to correspond with those, s, at the lower end of the shank. The middle parts of the arms are rounded, and the outer extremities are cut away as much as the thickness of the flukes or palms m, that the palms may be flush with the upper sides when they are welded on. The flukes are gene¬ rally made at the iron-forges in the country, by the forge- hammer ; but in some yards they are made by faggoting small bars, leaving a long one for a handle. When finish¬ ed, they are welded to the arms, which have then the ap¬ pearance of X. The next business is to unite the arms to the end of the shank ; and in doing this particular care is necessary, as the goodness of the anchor is entirely de¬ pendent upon its being effectually performed. In so large a weld, the outside is very liable to be welded, and make a good appearance, while the middle part is not united. To guard against this, both surfaces of the scarfs should be rather convex, that they may be certain to touch in the middle first. When the other arm is welded, the anchor is complete, except the ring, which is made from several small bars welded together, and drawn out into a round rod, then bent to a circle, put through the hole in the shank, and its ends welded together. If the shank or other part is crooked, it is set straight by heating it in the crooked part, and striking it over the anvil, or by the Her¬ cules. After all this the whole is heated, but not to a white heat, and the anchor hammered in every part, to finish and make its surface even. This is done by lighter hammers, worked by both hands, but not swung over the head. This operation renders the surface of the metal hard and smooth; and if very effectually performed, the anchor will not rust materially by the action of the sea¬ water. The hammering is continued till the iron is quite black and almost cold. It is common with some manufac¬ turers, after they have made up the shank, to heat it again, and apply the end of a thin flat bar properly heated upon it; then by turning the large shank round, the har is wound spirally upon it, so as to form a complete covering to. the whole. I his method admits of emploving a kind of iron which is less liable to corrosion, but we fear it is sometimes resorted to to conceal the bad qualities of the iron of which the anchor is composed. I he iron from which anchors are made ought to be ol the best quality: that kind of it which is called red short * v A N C An, or will not bear sufficient hammering to weld the bars; and ' cold short, from its brittleness, is not to be depended upon Amfia. when the anchor is in use. A good anchor should be forrned of the toughest iron that can be procured. 'atei A new and improved method of fabricating anchors, for olio which a patent has been taken out by Lieutenant Rodgers, banM j^g i)een lately introduced. This consists in making the nflK ‘ shank hollow, and uniting it to a solid square at one end, and to a solid piece at the other, which forms the crown and part of the arms. The hollow part is bound at intervals by hoops, and the whole forms a much stronger anchor, of the same weight, than those of the common construction. The stock also is let on over the end of the square, and keyed up against a collar, by which means the anchor can be un¬ stocked with great facility. See Plate XLIII. fig. 7. The most extensive establishment for fabricating an¬ chors, &c. is that at Woolwich dock-yard. There the blow¬ ing apparatus, the working of the lift and tilt hammers, &c. is all done by a steam-engine of from 14 to 16 horse power. The improvements of Perring and Rodgers have been there introduced. (c.) To steer the ship to her Anchor, is to steer the ship’s head towards the place where the anchor lies when they are heaving the cable into the ship, that the cable may thereby enter the hause with less resistance, and the ship advance towards the anchor with greater facility. Anchor-Ground is a bottom which is neither too deep, too shallow, nor rocky; as in the first the cable bears too nearly perpendicular, and is thereby apt to jerk the anchor out of the ground; in the second, the ship’s bottom is apt to strike at low water, or when the sea runs high, by which she is exposed to the danger of sinking; and in the third, the anchor is liable to hood the broken and pointed ends of rocks, and tear away its flukes, whilst the cable, from the same cause, is constantly in danger of being cut through as it rubs on their edges. Anchor, in Architecture, is a sort of carving somewhat resembling an anchor. It is commonly placed as part of the enrichment of the boultins of capitals of the Tuscan, Doric, and Ionic orders, and also of the boultins of bed mouldings of the Doric, Ionic, and Corinthian cornices, anchors and eggs being carved alternately through the whole building. Anchors, in Heraldry, are emblems of hope, and are taken for such in a spiritual as well as a temporal sense. ANCHOVY. See Ichthyology. ANCIENT, or Antient, a term applied to things which existed long ago. Thus we say, ancient nations, ancient customs, &c. See Antiquities. ANCIENT Demesne, in English Law, is a tenure whereby all manors belonging to the crown in the times of William the Conqueror and St Edward were held. The numbers, names, &c. were entered by the Conqueror in the record called Domesday Book; so that such lands as by that book appeared to have belonged to the crown at that time are called ancient demesne. ANCIENTY, in some ancient statutes, is used for el¬ dership or seniority. The elder sister can demand no more than her other sisters, beside the chief mesne, by reason ot her ancienty. This word is used in the statute of Ireland, 14 Henry III. ANCONA, a delegation or province of the Papal States, in Italy, a part of the ancient mark of Ancona. It is bounded on the north by Urbino, on the east by the Adriatic Sea, on the south by Macerata, and on the west by Urbino. Its extent is 646 square miles, or 413,440 acres. It is a mountainous district, with scarcely any plains; but between the projections of the Appenines there are some valleys, watered by the Musone, the Esino, the Aspino, and several smaller mountain rivers, which A N c 109 yield good crops of corn, maize, garlic, fruit, almonds, Ancona tobacco, and some silk. Agriculture is much neglected, II but more attention is paid to cattle. The delegation con- Ancylo- tains 3 cities, 17 market-towns, and 34 villages; and in 1816 the number of inhabitants was 147,355, who, besides agriculture, find employment in manufactures of linen, silk, stockings, paper, and some other articles. The coun¬ try is remarkably healthful. Ancona, the capital of the delegation of the same name. The city is in lat. 43° 43' 36" N. and long. 12° 42' 27" E. It is situated on the declivity of a tongue of land project¬ ing into the Adriatic Sea, on which, after the destruction of the fortifications in 1815, a strong fort was built as a defence towards the sea. It has three gates towards the land. The most striking objects are the government palace, the town-house, the exchange, and the cathedral built at the extremity of the cape, on the site of an ancient temple of Venus. There is a college, an hospital, and several monasteries and nunneries. The inhabitants in 1816 were 29,792, among whom were 5000 Jews, who live in a separate quarter. The streets are narrow, and far from clean. There is a fine mole 2000 feet in length, on which a lazaretto and quarantine-house is built. The har¬ bour is capacious, but complaints are made of the mud gradually increasing and injuring it. The trade is carried on by means of about 1100 vessels, which arrive and depart annually. This place endured a long siege in the year 1799, but at length the French garrison wbo defended it surrendered to a combined army and navy of Russians, Austrians, and Turks. ANCONES, in Architecture, the corners or quoins of walls, cross beams, or rafters. Vitruvius calls the consoles by the same name. ANCOURT, Florent Carton d’, an eminent French comic writer and actor, was born at Fontainbleau, on the 1st November 1661. He died on the 6th of December 1726, being 65 years of age. The plays which he wrote were all, with one exception, of the comic cast. They have been frequently reprinted, and form, in the best edition, namely, that of 1760, a collection of 12 vols. 12mo. ANCRE, a small town of France, in Picardy, with the title of a marquisate, seated on a little river of the same name. Long. 2. 45. E. Lat. 49. 59. N. ANGUS Martius, the fourth king of the Romans, suc¬ ceeded Tullus Hostilius 639 years before Christ. He de¬ feated the Latins, subdued the Fidenates, conquered the Sabines, Volsci, and Veientines, enlarged Rome by joining to it Mount Janiculum, and made the harbour of Ostia. He died about 615 years before the Christian era. ANCYLE, in Antiquity, a kind of shield that fell, as was pretended, from heaven, in the reign of Numa Pompilius ; at which time, likewise, a voice was heard declaring that Rome should be mistress of the world as long as she should preserve this holy buckler. It was kept with great care in the temple of Mars, under the direction of 12 priests ; and, lest any should attempt to steal it, 11 others were made so like as not to be distinguished from the sacred one. These ancylia were carried in procession every year round the city of Rome. ANCYLOBLEPHARON (from ayxv'kog, bent, and /3Xs- cpagov, an eyelid), a disease of the eye, which closes the eyelids. ANCYLOGLOSSUM (from ayy^Xog, crooked, and yXuxJm, the tongue), a contraction of the ligaments of the tongue. Some have this imperfection from their birth, others from some disease. In the first case, the membrane which supports the tongue is too short or too hard; in the latter, an ulcer under the tongue, healing and forming a cicatrix, is sometimes the case. These speak with some difficulty. The ancyloglossi by nature are late before 110 AND Ancylosis they speak ; but when they begin, they soon speak proper¬ 'll ly. These we call tongue-tied. Andaman. ANCYLOSIS, in Surgery, implies immobility or stiff- ness of the joints, and is used to express such stiffness, whether proceeding from internal or external causes. ANDABATiL, in Antiquity, a sort of gladiators, who, mounted on horseback or in chariots, fought hoodwinked, having a helmet that covered their eyes. ANDALUSIA, an extensive province in the south of Spain, on the Mediterranean Sea. Though its surface is very unequal, and its soil and climate vary with the eleva¬ tions of the land, it must be considered the most rich and delightful of all the divisions of the peninsula. It is divid¬ ed into four districts, which, in conformity with ancient usage, are denominated kingdoms, viz. Jaen, Cordova, Se¬ ville, and Granada, the description of each of which will be found in their alphabetical place. Andalusia, New, a division of the province of Terra Firma in South America, whose boundaries cannot be well ascertained, as the Spaniards pretend a right to countries in which they have never established any settlements. According to the most reasonable limits, it extends in length 500 miles from north to south, and about 270 in breadth from east to west. The interior country is woody and mountainous, variegated with fine valleys that yield corn and pasturage. The produce of the country consists chiefly in dyeing-drugs, gums, medicinal roots, brazil wood, sugar, tobacco, and some valuable timber. To this pro¬ vince also belonged five valuable pearl fisheries. The capital of New Andalusia is Comana, Cumana, or New Corduba, situated in lat. 10° 5' N. about nine miles from the North Sea. Here the Spaniards laid the foundation of a town in the year 1520. The place is strong by nature, and fortified by a castle capable of making a vigorous de¬ fence. The country is better known under the name of Guiana. The revolution which began there in 1810 is not yet terminated. ANDAMAN Islands. These islands, which are situ¬ ated on the eastern side of the Bay of Bengal, are a con¬ tinuation of the archipelago which extends from Cape Ne- grais to Atchein Head, stretching from lat. 10° 32' to 13° 40' N., and from long. 90° 6' to 92° 59' E. They are called the Great and the Little Andaman. The Great Andaman, which is the northern island, is 140 miles in length, and only 20 broad. It was formerly supposed to be one island; but two straitshave been discovered, which open a clear pass¬ age into the Bay of Bengal, and divide the Great Andaman into three islands. The Little Andaman, which lies 30 miles south of the Great Andaman, is 28 miles long and 17 broad. It does not afford any good harbour, though tolerably safe anchorage maybe found near its shores. These islands have an extremely moist temperature. They are situated in the direct current of the south-west monsoon; and the central mountains, some of the lofty peaks of which, as Saddle Peak in the large Andaman, rise to the height of 2400 feet, intercept the clouds, which, for about eight months in the year, descend in incessant torrents of rain on the plains below. According to a meteorological table kept by an officer resident on the island, 98 inches of water appear to have fallen in the course of seven months. On the whole, however, the temperature is milder than in Bengal, and the heats not so intolerable. . The island is totally uncultivated, and the savage inha¬ bitants glean a miserable subsistence from the spontane¬ ous produce of the woods, in which the researches of the Europeans have hitherto found little that is either palat¬ able or nutritious. The principal trees are the banyan- tree, the almond-tree, the oil-tree, which grows to a crreat height and yields a very useful oil; the poon, the dam- mer, the red wood, which for furniture is little inferior to AND fine mahogany; the ebony, the cotton-tree, the soondry, AndjB, chingry, and beady; the Alexandrian laurel, the poplar, a Oc¬ tree resembling satin-wood, bamboos, cutch, the melon, aloes ; the iron-tree, of stupendous size, whose timber al¬ most bids defiance to the axe of the wood-cutter. There are many other trees well adapted for the construction of ships ; and, as in all the equatorial forests, there are num¬ berless creepers and ratans, which surround the stems of the trees, and are so firmly interlaced together, that the forests are impervious, except a road be previously cut through them. The only quadrupeds seen on the island are hogs, rats, and the ichneumon ; also the guana, of the lizard tribe; all which are very destructive to poultry. There are several species of snakes and scorpions, by which the labourers employed by the British in clearing away the underwood were frequently bitten; but in no instance did the bite prove mortal. The patient was frequently affected with violent convulsions, which gradually yielded to the ope¬ ration of opium and eau-de-luce. Fish abound on the shores, and are caught in great numbers during the prevalence of the north-east mon¬ soon, when the weather is mild: grey mullet, rock cod, skate, and soles, are among the best. There are, besides, various other species, such as guanas, sardinas, roe-balls, sable, shad, prawns, shrimps, cray-fish, a species of whale, and sharks of an enormous size. Shell-fish are in great plenty, and oysters of an excellent quality. The shores abound in a variety of beautiful shells, such as gorgonias, madreporas, murex, and cowries, with many other sorts equally beautiful. Birds are not numerous, and they are extremely shy. Doves, parroquets, and the Indian crow, are the most common. Hawks from the neighbouring continent are sometimes seen hovering over the tops of trees; and a few aquatic birds, such as the king-fisher, a sort of cur¬ lew, and the small sea-gull, frequent the shores. Within the caverns and recesses of the rocks are found the edi¬ ble birds’ nests so highly prized among the Chinese for their supposed medicinal and restorative qualities. The whole population of the islands does not exceed 2000 or 2500, and they are probably the most savage people on the face of the whole earth. They are far be¬ low the ordinary scale of barbarism; and in their modes of subsistence, and in their dwellings, they rise very little above the brute creation. They wear no clothes, and seem insensible to any feeling of shame from the exposure of their persons. The woods supply them with little in the way of food. They are provided with no pot or vessel that can bear the action of fire, and they cannot there¬ fore derive much advantage from such esculent herbs as the forests may contain. The cocoa nut, which thrives so well in the neighbouring islands, is not found in the Andamans, though the natives are extremely fond of it. The fruit of the mangrove is principally used by them. Their principal food consists of fish, in quest of a preca¬ rious meal of which they climb over the rocks, or rove along the margin of the sea, often without success during the tempestuous season; but they eagerly' seize on what¬ ever else presents itself, such as lizards, guanas, rats, and snakes, from their diseased and extenuated figures, it is plain that they have no abundant or wholesome nourish¬ ment. In stature the inhabitants of the Great Andaman seldom exceed five feet; their limbs are disproportionably slender, their bellies protuberant, with high shoulders and large heads ; and, what is singular and unaccountable, they have all the characteristic marks of a degenerate race of negroes, with woolly hair, flat noses, and thick lips; their eyes are small and red, their skin of a deep sooty black, while their countenances exhibit a mixed expres- AND ’ Ibi-. sion of famine and ferocity. Lieutenant Alexander de- ' my-vj gcribes the inhabitants of Little Andaman as far from be¬ ing a puny race. When he landed in a boat he counted six¬ teen strong and able-bodied men, many of them very lusty. The ingenuity of these savages is principally seen in the fabrication of a few simple weapons on which they de¬ pend for their subsistence. These are a bow from four to five feet long, with arrows of reed, headed with fish-bone or wood hardened in the fire, a spear of heavy wood sharp¬ ly pointed, and a shield made of bark. With these im¬ plements they shoot and spear the fish, which abound in their bays and creeks, with surprising dexterity. Their ' habitations display little more art than the dens of wild I beasts. Their canoes are hollowed out of the trunks of 1 trees by fire and instruments of stone, as they have no > , iron among them. Being much incommoded by insects, their first occupation in the morning is to plaster their 1 bodies all over with mud, which, hardening in the sun, forms an effectual defence. They paint their woolly heads with i a mixture of ochre and water; and, when completely j dressed, it is observed by Mr Symes, who visited the island > in his voyage to Ava, that a more hideous appearance is not i to be found in human form. Their language does not pos- 8 sess the slightest affinity to any spoken either on the conti- 111 nent of India or on the islands. It is not harsh to the ear. 1 Their songs are wildly melodious, and their gesticulations ^ when singing extremely impassioned. The men have all the c cunning and vindictive dispositions of savages; and they 8 showa marked hatred to all strangers. They have little idea o of a future state, but they adore the sun and the moon, and 1 the genii of the woods, waters, and mountains as inferior 8 agents. During the south-west monsoon, when tempests f prevail with unusual violence, they deprecate the wrath o of the malignant being who, according to their notions, 1' has occasioned the storm, by chanting wild choruses in s small congregations assembled on the beach or on some f1 *. rock that overhangs the ocean. These islanders have oc- c casionally been persuaded to come on board British ships; b but though they were well fed and kindly treated, no per- s e suasion could induce them to remain. The settlement o ; of these islands, with their negro inhabitants, so widely f different in their appearance not only from all those of the l § Asiatic continent, in which the Andamans are embayed, 5 but also from the natives of the Nicobar islands, presents a a curious problem, which has never been satisfactorily c explained. It is supposed, however, by Symes, that the of original stock must have been settled on the island by t « the accidental ship week of some Arab slave-ship. The 1 English made a settlement on the larger Andaman in the J year 1791. Their object was to procure a commodious ii harbour on the east side of the Bay of Bengal, to receive a and shelter ships of war during the continuance of the o I north-east monsoon; also to provide a place of reception i for convicts sentenced to transportation from Bengal. 1 But the settlement, proving unhealthy, has been since a abandoned. These islands have been frequently visited by ! British ships, but the barbarous natives reject all friendly i | intercourse. Lieutenant Alexander states that they pre- s sented a hostile aspect to the party who landed in search of fresh water, assailed them with volleys of arrows, and testi- t fied, by the most ferocious signs, their determined aversion t to them. In April 1824 the British armament destined ( |‘ against the Burmese touched at the great island, when t the natives persevered in their hostility, discharging flights ( of arrows at their boats, and then flying into the woods, tyiese islands, together with the Nicobar and other smaller islands, were included by Ptolemy in the general appella- 1 tlon °f Insulae, Bonce Fortunes, and were supposed by him to be inhabited bj^ a race of anthropophagi, though there ure no proofs of the modern inhabitants being addicted and in to this practice. (Symes’ Embassy to Ava ; Alexander’s Andante Travels from India to England, comprehending a Visit to II the Burman Empire, &c.; Hamilton’s Gazetteer.) (f.) Anderson. ANDANTE, in Music, signifies a movement moderate- ly slow, between largo and allegro. ANDEGAVI, or Andegavus, a town of Gallia Celtica (Pliny, Ptolemy); now Angiers: called Andecavi (Taci¬ tus). Long. 30. W. Lat. 47. 30. N. ANDELYS, an arrondissement in the department of the Lower Seine, in France. It extends over 390 square miles, or 256,440 acres; is divided into six cantons, which are subdivided into 147 communes; and contains 63,211 in¬ habitants. The chief place, of the same name, has a popu¬ lation of 5256 souls. ANDENA, in old writings, denotes the swath made in the mowing of hay, or as much ground as a man could stride over at once. ANDEOL, Saint, a town of France, in the department of Lozere, five miles south of St Viviers, whose bishop formerly resided there. Long. 2. 50. E. Lat. 44.24. N. ANDERAB, the most southern city of the province of Balk, possessed by the Usbec Tartars. The neighbouring mountains yield excellent quarries of lapis lazuli, in which the Bukhars drive a great trade with Persia and India. ANDERAVIA, or Inderabia, a low, level, and nar¬ row island on the Arabian shore of the Persian Gulf, about three miles in length. It is separated from the mainland by a strait three miles in length, and free from danger ; but ships running for shelter under this island must not come within a mile of its south end, until a remarkable tree, which will be distinguished standing by itself, bear west- north-west. Lat. 26. 40. N. ANDERNACHT, a city in the duchy of the Lower Rhine, belonging to Prussia. It is situated in a plain on the river Rhine, and is fortified with a wall, castle, and bulwarks. It has a trade in stone jugs and pitchers, which are sent to the mineral waters at Dunchstein. There are three monasteries here, and several churches. Long. 7. 4. E. Lat. 50. 27. N. ANDERO, Saint, a seaport town in the Bay of Biscay, in Old Castile, seated on a small peninsula. It is a trad¬ ing town, and contains about 700 houses, two parish churches, and four monasteries. Here the Spaniards build and lay up some of their men of war. Long. 3. 45. W. Lat. 43. 20. N. ANDERSON, Alexander, a very eminent mathema¬ tician, who flourished in the early part of the 17th century. He was born at Aberdeen, but passed over to the Conti¬ nent, and settled as a private teacher or professor of mathe¬ matics at Paris, where he published or edited, between the years 1612 and 1619, various geometrical and algebraical tracts, which are conspicuous for their ingenuity and ele¬ gance. It is doubtful whether he was ever acquainted with the famous Vieta, master of requests at Paris, who died in 1603; but his pure taste and skill in mathematical investigation had pointed him out to the executors of that illustrious man, who had found leisure, in the intervals of a laborious profession, to cultivate and extend the ancient geometry, and, by adopting a system of general symbols, to lay the foundation and begin the superstructure of algebraical science, as the person most proper for revising and publishing his valuable manuscripts. Anderson did not come forward, however, as a mere editor; he enriched the texts with learned comments, and gave neat demon¬ strations of those propositions which had been left imper¬ fect. He afterwards produced a specimen of the applica¬ tion of geometrical analysis, which is distinguished by its clearness and classic elegance. Of this able geometer we are ignorant both of the time of his birth and of Ins death. His brother David Ander- 112 AND Anderson, son, a small proprietor in Aberdeenshire, but engaged in business, had likewise a strong turn for mathematics and mechanics, which, joined to great versatility of talent, made him be regarded by his neighbours at that period as a sort of oracle. The daughter of this cleyei and active burgess was married to John Gregory, minister of Drum- oak, in that county, father to the celebrated James Gre¬ gory, inventor of the reflecting telescope; and is suppos¬ ed to have communicated to her children that taste for mathematical learning which afterwards shone forth so remarkably in the family of the Gregorys. The works of Anderson amount to six thin quarto vo¬ lumes, which are now very scarce. Ihese are, 1. Supplementum Apollonii lledivivi; sive, Analysis pro- blematis hactenus desiderati ad Apollonii Pergsei doc- trinam i/tvtfsojv, a Murine Ghetaldo Patritio liagu- sino hucusque non ita pridem restitutam. In qua exhi- betur Mechanice sequalitatum tertii gradus sive solida- rum, in quibus magnitude omnino data aequatur homoge- neae sub altero tantum coefficiente ignoto. Huic sub- nexa est variorum problematum practice, eodem Auc- tore. Parisiis, 1612, 4to. This tract refers to the pro¬ blem of inclinations, by which, in certain cases, the ap¬ plication of the curve called the conchoid is superseded. 2. AmoXoyia: Pro Zetetico Apollonian! problematis a sp jam pridem edito in supplemento Apollonii Redivivi. Ad clarissimum et ornatissimum virum Marinum Ghe- taldum Patritium Ragusinum. In qua ad ea quae obiter inibi perstrinxit Ghetaldus respondetur, et analytices usus clarius detegitur. Parisiis, 1615, 4to. I his is an addition to the author’s former work. 3. Francisci Vietae Fontenacensis de iEquationum Recog- nitione et Emendatione Tractatus Duo. Parisiis, 1615, 4to. To this work Anderson has supplied the dedica¬ tion, preface, and appendix. 4. Ad Angularium Sectionum Analyticen Theoremata KadoXauT^a, a Francisco Vieta Fontenacensi excogi- tata, at absque ulla demonstratione ad nos transmissa, jam tandem demonstrationibus confirmata. Parisiis, 1615, 4to. 5. Vindicise Archimedis; sive, Elenchus Cyclometriae Novae a Philippo Lansbergio nuper editae. Parisiis, 1616, 4to. 6. Alexandri Andersoni Scoti Exercitationum Mathema- ticarum Decas prima; continens Quaestionum aliquot, quae nobilissimorum turn hujus turn veteris aevi Mathe- maticorum ingenia exercuere, Enodationem. Parisiis, 1619, 4to. C (b.) Anderson, Sir Edmund, a younger son of an ancient Scotish family settled in Lincolnshire. He was some time a student of Lincoln College, Oxford, and removed from thence to the Inner Temple, where he applied himself diligently to the study of the law, and became a barrister. In 1582 he was made lord chief justice of the common pleas, and in the year following was knighted. He held his office to the end of his life in 1605. His works are, 1. Reports of many principal Cases argued and adjudged in the time of Queen Elizabeth in the Common Bench, Lond. 1644, fol.; 2. Resolutions and Judgments on the cases and matter agitated in all courts of Westminster in the latter end of the reign of Queen Elizabeth, Lond. 1655, 4to. » Anderson, James, LL. D. The subject of this article, who has been brought into notice principally from the more recent encouragement given to agriculture, and the versatility of his own genius, was born at the village of Hermiston, in the county of Edinburgh, in the year 1739. His parents were in humble life, and had possess¬ ed a farm for some generations, which he was destined to inherit and to cultivate. At that period improvement AND was in infancy, and the husbandman had to contend withAnde a climate whose uncertainty seemed to keep pace withW ; the progress of his skill, and which too often disappointed J him of the fruits of his industry. Anderson, while yet at an early age, lost his parents: however, his education was uninterrupted; and conceiving that an acquaintance with chemistry would promote his profession, he attended a course of lectures on that science, then delivered by Dr Cullen. None of the other pupils besides himself took notes of his lectures, which being afterwards surrepti¬ tiously obtained from him, with the design of publication, he defeated the intended purpose, apprehensive that his preceptor’s fame might be diminished by these imperfect transcripts. His own active occupations had already com¬ menced, and, along with the practice of husbandry, he prosecuted his original taste for literature. Enlarging the sphere of his employments, Anderson for¬ sook his first possession, for a farm of 1300 acres, which he rented in Aberdeenshire, though nearly in a state of nature, and where agriculture is still behind the southern districts. But previous to this he became known to men of letters, by some essays on planting, which, under the signature Agricola, he ventured to commit to the world through the medium of the Edinburgh Weekly Magazine, in 1771. Soon embarking in a higher sphere of literature, he composed the article Monsoon for the first edition of this Encyclopaedia, in 1773, wherein he threw out some luminous ideas, and, among other observations, predicted from physical facts, and the state of geographical know¬ ledge, that no polar continent would be found by the cir¬ cumnavigators then employed by government. In the year 1777 he published a considerable quarto volume, said to have been composed two years preceding, on the means of exerting a spirit of national industry with regard to agriculture, commerce, manufactures, and fisheries; and in this he enters into detailed views of many subjects of political economy. The interest of the Highlands and Islands of Scotland is in a particular manner considered; and the author maintains, that the only effectual means of increasing agriculture is by promoting manufactures; as also, that the neglect-which the agriculture of these parts of the kingdom experienced, resulted from the neglect of manufactures. Scotland, he affirms, is better adapted than England for the production of wool; and this, as well as other products, he thought, would be best encouraged by premiums. The advantages which might result from at¬ tending to the fisheries, he judged, would be very great, and the shoals of herrings frequenting the coast could be converted into a source of national wealth under suit¬ able establishments. Anderson, who soon after had the degree of doctor of laws conferred on him by the Univer¬ sity of Aberdeen, did not abandon these inquiries. He printed a tract regarding the fisheries, which was cir¬ culated among his friends ; and, in consequence of being more widely diffused, he was appointed by the Lords of the Treasury to survey the western coast of Scot¬ land, for the purpose of obtaining satisfactory informa¬ tion on the subject. This he did in 1784, and received the full approbation of his employers ; and he published a brief account of the Hebrides, a chain of islands then as little within the general acquaintance of the inhabitants of Great Britain as if they had been under the dominion of another country. The principal obstacles to the fishery, Dr Anderson considered, were to be found in a duty on salt and coals, and he recommended the repeal of both. It is certain that, from thenceforward, this great branch of national industry has received infinitely more patronage than before his report, and, while we only shared the labours of the Dutch for centuries, the fisheries on our own coast have since been monopolized by ourselves. No- AND non'thing can be more impolitic than to fetter the exertions of ^Mhe industrious by exorbitant duties, or equally oppres¬ sive with denying fuel to the poorer classes, where it is scarce, by duties on its importation. Dr Anderson had now withdrawn from his northern farm, where he resided above 20 years, and settled in the vicinity of Edinburgh. His agricultural speculations were still continued ; and when a parliamentary grant was about to be proposed to Mr Elkington for a particular mode of draining land, he reclaimed the practice as having been observed by himself many years anterior. Repeated ex- unples, indeed, prove that the rewards granted by Parlia¬ ment for improvement are attended with such slight in¬ vestigations, that the merits of real invention are over¬ looked. Dr Anderson now projected a periodical publica¬ tion called The Bee, consisting of miscellaneous original matter, which attained the extent of 18 volumes in octavo. It was published weekly, and a large proportion of it came from his own pen, which is seldom a prudent course in an editor. The relation of Great Britain and her colo¬ nies, and the political rights of mankind—subjects which had excited strong interests throughout Europe—also re¬ ceived some commentaries from Dr Anderson. He wrote a tract called The Interest of Great Britain with respect to her Colonies, and commenced a correspondence with Ge¬ neral Washington, which was afterwards published. Towards the year 1797 he again removed to Islesworth, in the neighbourhood of London, where he undertook another periodical publication, appearing at more distant intervals than the former, entitled Recreations in Agricul¬ ture, Natural History, Arts, and Miscellaneous Literature. This work was prefaced by two copious dissertations, the one on agriculture, the other on natural history; and opened with a discussion regarding a very curious and im¬ portant subject, namely, an inquiry into what are deno¬ minated varieties in plants and animals. Many useful and interesting remarks appear in the course of this pub¬ lication, a portion of which was supplied by other con¬ tributors; and it is embellished by beautiful vignettes from engravings on wood. Owing to some difficulties attending the publication, it ceasecH in 1802. Though natural history is rather predominant, the rest of his lead¬ ing subjects are not overlooked. Dr Anderson hence¬ forward lived in a great measure in retirement, though occasionally reminding the world of his wonted inqui¬ ries, by the publication of tracts on unconnected sub¬ jects. He obtained a patent for an improved hot-house, wherein no fuel was used; and employed himself in ex¬ periments regarding the degree of temperature and hu¬ midity most beneficial to plants. Likewise, having ob¬ served the uncommon depredations of wasps, he is said, after satisfying himself of their manner of increase, to have devisad a plan for their absolute extermination. This was chiefly by the destruction of the females before founding their respective colonies by the depositation of innumer¬ able eggs; and hand-bills were circulated under the aus¬ pices of an association formed by him, offering a reward for every female brought in dead within a specified time. It does not appear, however, that the breed was at all di¬ minished by the proposed expedient. Dr Anderson still remained in his retreat, enjoying the cultivation of his garden; and nothing of importance is known to have proceeded from his pen. After a gradual decline, partly occasioned by the over-exertion of the mental energies, he died in the year 1808, aged 69. He was twice married; first, to Miss Seton of Mounie; second¬ ly, to an English lady. By his first marriage he had thir¬ teen children, six of whom survived him. One of his sons made distinguished progress in the art of engraving on wood; and, if the vignettes of the Recreations in Agri- wliii. AND 113 culture were executed by him, they afford ample testimony Anderson, of his abilities. Dr Anderson was endowed with a vigorous understand¬ ing, which he chiefly displayed in treating of agricultural matters, and those connected with rural economy; but he was at the same time of a versatile talent, which could readily be occupied on transient facts and occurrences. Many of his works were of a fugitive nature, consisting of small impressions, which were not renewed, and hence are difficult to be obtained at present, if they have not totally disappeared. None of them soars to the more lof¬ ty regions of science ; they are directed to practical views in useful projects, and for the most part relate to subjects of ordinary detail. Of this the reader will be enabled to judge by the subjoined list, which we believe is the most copious that has yet appeared. The industry of Dr An¬ derson was indefatigable, whether in personal exertion or mental energy; and he possessed elevated sentiments of independence. During a period of overstrained political fervour, certain papers formed part of the periodical works already referred to, which were thought libellous on the government. Although Dr Anderson’s principles were noted for attachment to the existing administration, he was called upon to give up the author of the obnoxious compositions, which he steadily refused, and, even in the face of the civil magistrates, charged his printers not to violate their fidelity to him and the author in betraying his name. The business terminated here, until a factious individual insinuated to the same magistrates that the compositions had proceeded from one of the supreme judges, whose party politics were avowedly hostile to those of government. Dr Anderson having learned the reproach, hastened to relieve the object of it by divulging the name of the real author, who, to the universal surprise of the public, proved to be none other than the traducer himself. 1776. A Practical Treatise on Chimneys, containing full instructions for constructing them in all cases, so as to draw well, and for removing smoke, 12mo. 1776. Free Thoughts on the American Contest, 8vo. 1777. Observations on the Means of exciting a Spirit of National Industry, 4to. 1777. Miscellaneous Observations on Planting and Train¬ ing Timber Trees, by Agricola, in 8vo. 1777. An Inquiry into the Nature of the Corn Laws, in 8vo. 1777. Essays Relating to Agriculture and Rural Affairs, 8vo. A fifth edition, in 3 volumes, was published in 1800. 1779. An Inquiry into the Causes that have hitherto re¬ tarded the Advancement of Agriculture in Europe, 4to. 1782. The Interest of Great Britain with regard to her American Colonies considered, 8vo. 1783. The True Interest of Great Britain considered, or a Proposal for Establishing the Northern British Fisheries, 12mo. 1785. An Account of the Present State of the Hebrides and Western Coast of Scotland, being the Sub¬ stance of a Report to the Lords of the Treasury, 8vo. 1789. Observations on Slavery, particularly with a View to its Effects on the British Colonies in the West Indies, 4to. 1790. Papers by Dr Anderson and Sir John Sinclair on Shetland Wool, 8vo. 1791. The Bee, 18 vols. 8vo. 1792. Observations on the Effects of the Coal Duty, 8vo. 1793. Thoughts on the Privileges and Powers of Juries, with observations on the State of the Country with regard to Credit, 8vo. 114 AND Anderson. 1793. Remarks on the Poor’s Laws of Scotland, 4to. 1794. A Practical Treatise on Peat Moss, 8vo. 1794. A General View of the Agriculture and Rural Economy of the County of Aberdeen, and the Means of its Improvement, 8vo. 1794. An Account of the Different Kinds of Sheep found in the Russian Dominions, by Dr Pallas. With five Appendixes, by Dr Anderson. 8vo. 1795. Two Letters, to Dr Edward Home, on an Universal Character, in 8vo. 1797. A Practical Treatise on Draining Bogs and Swampy Grounds, with Cursory Remarks on the Originality of Elkington’s Mode of Draining Lands, 8vo. 1799. Recreations in Agriculture, 6 vols. 8vo. 1800. Selections from Correspondence with General Washington, 8vo. 1801. A Calm Investigation of the Circumstances that have led to the Present Scarcity of Grain in Great Britain; suggesting the means of alleviating that evil, and of preventing the occurrence of such a ca¬ lamity in future; 8vo. 1803. Description of a Patent Hot-house, which operates chiefly by the Heat of the Sun, and other Subjects, 8vo. Dr Anderson, besides, wrote many papers in periodi¬ cal publications, and an account of Ancient Fortifications in the Highlands, read to the Society of Scotish Anti¬ quaries. Anderson, Robert, the fourth son of William Ander¬ son, and of Margaret Melrose his wife, was born at Carn- wath in Lanarkshire on the 7th of January 1750. His fa¬ ther was a feuar, that is, a person who possessed some small parcels of real property by the tenure of a per¬ petual lease. Having received the earlier part of his edu¬ cation in his native town and in the adjacent village of Libberton, he was afterwards placed under the tuition of Robert Thomson, master of Lanark school, who had married a sister of Thomson the poet, and who has been commended for his uncommon proficiency in classical learning. Excellence however is always a relative term; nor must it be forgotten that the grammar schools of Scotland had about that period descended to their very lowest level. James Graeme, a native of the same town, was his companion and friend at all these seminaries: he appears to have been an amiable young man of promising talents, and to have been imbued with the love of lite¬ rature. One of their school-fellows at Lanark was John Pinkerton, who afterwards became a conspicuous mem¬ ber of the republic of letters. Anderson’s father had died in 1759, in the 40th year of his age, leaving his widow with a very slender provi¬ sion. Uniting considerable energy of character with a large share of piety, she made a vigorous effort to educate her four sons; and as she survived till the year 1796, she had the satisfaction of seeing one of them arrive at inde¬ pendence and distinction. Having discovered an early taste for reading, he soon made choice of one of the libe¬ ral professions. His first destination was for the church: in the year 1767 he was sent to the university of Edin¬ burgh, and in due time was enrolled among the students of divinity. Graeme, who had entered the university at the same time and with the same views, died of con¬ sumption in 1772, in the 23d year of his age; and, after a short interval, his faithful friend published a collection of Poems on several occasions, by James Grceme. Edinb. 1773, 12mo. These poems he inserted in the 11th vo¬ lume of the British Poets, together with an account of the author, in which his literary merits are estimated, not with the discrimination of sober criticism, but with, all the partiality of friendly zeal. About this period he relin- A N D quished the study of divinity, and betook himself to the Ai, study of medicine. He was for a short time employed 0 as surgeon to the dispensary at Bamborough Castle in Northumberland; and in a neighbouring town he then formed connections which had no small influence on his future destiny. On the 25th of September 1777 he mar¬ ried Anne, the daughter of John Grey, Esq. of Alnwick, who was related to the noble family of that name. Re. turning to Scotland, he took the degree of M. D. at St Andrews on the 20th of May 1778, after having been duly examined by the professor of physic. He now began to practise as a physician at Alnwick; but his general habits were rather those of speculation than exertion, and a moderate provision, acquired by his marriage, had emanci¬ pated him from the necessity of professional labour. In 1784 he finally returned to Edinburgh, where he continued to reside for the period of 46 years, in a condition of life removed from affluence, but perfectly compatible with genuine independence and comfort. He possessed the valuable secret of living within his income ; and his house was long distinguished by a hearty and unostentatious hos¬ pitality. His amiable and affectionate wife died of con¬ sumption on the 25th of December 1785, in the 39th year of her age, leaving three daughters, the youngest of whom speedily followed her mother to the grave. He did not re¬ sume the practice of physic, but being satisfied with his moderate acquisitions, he devoted much of his time to the education of his two children, and to miscellaneous and desultory reading, rather than to any systematical course of study. In 1793, after having remained a widower for eight years, he married Margaret, the daughter of Mr David Dali, master of Tester school in the county of Haddington. For several years his attention was occupied with his edition of The Works of the British Poets, with Prefaces biographical and critical, which was published at Edin¬ burgh, and extends to 14 large octavo volumes. The earliest volume, which is now the second in the series, was printed in 1792-3; the 13th was printed in 1795, and another volume was added in 1807. He was fre¬ quently solicited to revise his Lives of the Poets, and publish them in a separate form ; but after having collected some materials for such a work, he finally abandoned the design. In the mean time he had published The Miscel¬ laneous Works of Tobias Smollett, M. D. with Memoirs of his Life and Writings. Edinb. 1796, 6 vols. 8vo. The sixth edition of this collection was printed in the year 1820. Of his account of the ingenious author, the eighth separate edition appeared under the title of The Life of Tobias Smollett, 31. D. with critical Observations on his Works. Edinb. 1818, 8vo. But the most able and elaborate of his productions is the third edition of his Life of Samuel Johnson, LL. D. with critical Observations on his Works. Edinb. 1815, 8vo. The same service which he rendered to Dr Smollett he afterwards extended to Dr Moore, hav¬ ing published The Works of John Moore, M.D. with Me¬ moirs of his Life and Writings. Edinb. 1820, 7 vols. 8vo. At an earlier period he had published The Poetical Works of Robert Blair ; containing the Grave, and a Poem to tbs Memory of Mr Law; to which is prefixed the Life of the Author. Lond. 1794, 8vo. And his latest publication was The Grave, and other Poems, by Robert Blair: to which are prefixed, some Account of his Life, and Obser¬ vations on his Writings. Edinb. 1826, 12mo. Dr Anderson contributed his ready aid to many differ¬ ent publications, and was always influenced, not by the love of money^but by the love of literature. With many eminent men in England, Ireland, and America, he main¬ tained a literary correspondence; and having survived most of his lettered contemporaries, he enjoyed the AND AND 115 ,. esteem and consideration of a second and even of a third ^generation. In 1802, 1809, and 1810, he paid three long visits to Dr Percy, the late excellent Bishop of Dromore; and he likewise enjoyed the friendship of Dr Ledwich, Dr Ryan, Mr Boyd, Mr Cooper Walker, and many other li¬ terary men of the sister island. No part of his character was more conspicuous than his uniform and unabating zeal to promote the success of young men who discovered any promise, however moderate, of literary talent; and some of the more distinguished writers of our own age and nation were not without their obligations to his dis¬ interested friendship. Mr Campbell dedicated to him his earliest and most popular publication, The Pleasures of Hope. Miss Bannerman, a very ingenious and accom¬ plished lady, who died at Portobello on the 29th of Sep¬ tember 1829, dedicated to him her first collection of Poems. Edinb. 1800, 8vo. Thomas Brown, John Leyden, and Alexander Murray, who all died at too early an age, were among the most eminent of his young friends. For Dr Brown, who became professor of moral philosophy in the university of Edinburgh, he entertained a very cor¬ dial esteem, which suffered no abatement or interruption. Another prominent feature of his mind, was his ardent regard for the civil and religious liberties of mankind. This characteristic he displayed from the first years of manhood till the last day of his earthly existence; and, on the very evening before his death, he gave a signal proof of his unquenchable zeal, by sending for a map of Greece, in order, as he expressed himself, to form some notion of the general elements of this new state. His principles were those of a very decided Whig; and, du¬ ring the extraordinary excitement which attended the close of the last century, he was sometimes misrepre¬ sented as little better than a republican. The honest alarm of one class of individuals, and the hollow zeal of another, had a strong tendency to confound all nice dis¬ tinctions. His bodily frame had never been robust; but the uni¬ form temperance and regularity of his habits contributed to prolong a life which was marked by cheerfulness and benevolence. His faculties, mental and corporeal, be¬ trayed few or no symptoms of old age. During the greater part of his last winter he was confined to his own house by what was considered as a common cold, and was attended by his friend and contemporary Dr Hamilton. His lungs however seemed to be affected in an unusual manner; and, five days before his dissolution, his physician discovered that dropsy in the chest had supervened. As he had now reached a very advanced age, he indulged not even the faintest hope of recovery. Though his chief or only suffering proceeded from diffi¬ culty of respiration, the progress of his malady was swift and certain: but the native alacrit)r of his mind seemed inca¬ pable of diminution ; and maintaining much and varied con¬ versation during the last days, and indeed during the last hours ofhis life,heevincedmore than his usual continuity of thought and accuracy of expression. As the vital tide was rapidly receding, his heart seemed to expand with new kindness towards all those who approached his couch. 01 the immediate prospect of death he spoke, not merely with resignation, but even with cheerfulness; with the subdued but confident hope of one who had long and ha¬ bitually reposed on the assurances of the Christian faith. He died on Saturday the 20th of February 1830, after having completed the 80th year of his age; and, accord¬ ing to his own directions, his remains were interred in Carnwath churchyard. His eldest daughter, Anne Mar¬ garet, was married in 1810 to David Irving, LL. D. and died in 1812, leaving an only son. His second daughter, Margaret Susannah, lived to deplore the loss of a parent, whose declining years she had soothed by the most ex- Andes, emplary attention to all his wants and wishes. (x.) ANDES. The Andes are distinguished above all the known mountain chains by their immense extent and their prodigious altitude. They run almost parallel to the west coast of the southern continent of America, at a mean dis¬ tance of between 100 and 200 miles, rising in some places to the enormous height of 25,000 feet; and stretch from the mouth of the river Atrato, on the isthmus of Darien, in the latitude of 8 degrees north, as far as Cape Pilares, at the outlet of the Straits of Magellan, in the 53d de¬ gree of south latitude, a range of at least 4200 miles. They send out, nearly at right angles from their colos¬ sal ridge, between the latitudes of 14° and 20° south, three dependent branches, called by the Spaniards Cor¬ dilleras. Of these secondary chains, the first and most northern is that of the coast of Venezuela, which is likewise the highest and narrowest. With an irregular altitude, it bends eastwards from the Atrato, forming the Sierra of Abibe, the mountains of Cauca, and the high Savannahs of Folu, till it reaches the stream of Magdalena, in the province of St Martha. It contracts as it approaches the Gulf of Mexico, at Cape Vela; and thence extends to the mountain of Paria, or rather the Galley Point, in the island of Trinidad, where it terminates. This secondary chain attains its greatest known elevation where it rears the snowy summit, or Sierra Nevada, of St Martha and of Merida, the former being nearly 14,000 and the latter above 15,000 feet in altitude. These insulated mountains, covered so near the equator with eternal snow, yet dis¬ charging boiling sulphurous water from their sides, are higher than the Peak of Tenerifte, and can be compared only with Mont Blanc. In their descent they leave the Paramo or lofty desert of Rosa and of Mucachi; and on the west side of the lake Maracaibo they form long and very narrow vales, running from south to north, and covered with forests. At Cape Vela the mountain chain divides into two parallel ridges, which form three confined valleys ranging from east to west, and having all the ap¬ pearance of being the beds of ancient lakes. These ridges, of which the northern is the continuation of the Sierra Nevada of St Martha, and the southern the extension of the snowy summits of Merida, are united again by two arms which seem to have been placed by the hand of nature as dikes to confine the primeval collections of wa¬ ter. The three valleys thus inclosed are remarkable for their elevation above the sea, rising like steps one above another, the eastmost, or that of the Caraccas, being the highest. This plain was found by Humboldt to be elevated 2660 feet, while the basin of Aragua has only 1350 feet in height, and the Llanos, or reedy plains of Monai, spread within 500 or 600 feet above the level of the shore. The lake of the Caraccas appears to have forced a passage for itself through the quehrada or cleft of Tipe, while that of Aragua has been gradually dissipated by a slow process of evaporation, leaving some vestiges of its former existence in pools charged with muriate of lime, and in the low islets called Aparecidas. The medium height of the Cordillera of the coast is about 4000 or 5000 feet; but its loftiest summit, next to the Sierra Nevada of Merida, is the Silla (or saddle) of the Caraccas, which was visited by Hum¬ boldt, and ascertained from barometrical measurement to have an elevation of 8420 feet. Farther to the eastward the mountain chain becomes suddenly depressed, espe¬ cially its primitive rocks; the beds of gneiss and mica slate meeting as they advance with accumulations of se¬ condary calcareous substances, which envelope them com¬ pletely, and rise to a great elevation. The incumbent mass of sandstone, with a calcareous base, extending from ANDES. 116 Andes. Capelluarl, forms a detached range of mountains, in which no trace of primitive rock is found. The second branch, which stretches from the Andes across the American continent, and exhibits a chain of primitive mountains, is named by Humboldt the Cordillera of the Cataracts of Orinoco. This very enterprising tra¬ veller surveyed it over an extent of upwards of 600 miles, from the Black River to the frontiers of the Great Bara; but the rest of the chain is very little known, running through unexplored wilds and regions almost inaccessible, occupied by fierce and independent tribes of savages. It leaves the great trunk between the 3d and 6th degree of southern latitude, and runs eastward from the Paramo or high desert of Tuquillo and St Martin, and the sources of the Guaviari, rearing the lofty summits of Umamaand Ca- navami, and pouring forth the large rivers Meta, Zama, and Ymerida, which form the roudals or tremendous rapids of Ature and Maypure, the only openings existing at present between the interior of the continent and the plain of the Amazons. Beyond these cataracts the chain of mountains again acquires greater elevation and breadth, occupying the vast tract inclosed between the rivers Caura, Cavony, and Padamo, and stretching southward to the boundless forests where the Portuguese settlers gather that potent drug the sarsaparilla. Farther eastward this chain is not traced, no European or civilized Indian having ever explored the sources of the Orinoco; all access in that quarter being prevented by the ferocity of the Guaicas, a dwarfish but very fair and warlike race, and by the valour of the Guajaribos, a most desperate tribe of cannibals. Beyond these recesses, however, we are made acquainted with the continuation of the chain of the cataracts, by the astonishing journey performed by Don Antonio Santos, who, disguised like a savage, his body naked, and his skin stained of a copper colour, and speaking fluently the seve¬ ral Indian dialects, penetrated from the mouth of the Rio Caronis to the' Lake of Parime and the Amazons. The range of mountains sinks lower, and contracts its breadth to 200 miles, where it assumes the name of Serrania de Quimeropaca and Pacaraimo. A few degrees farther east¬ ward it spreads out again, and bends south to the Canno Pirara along the Mao, near whose banks appears the Cerro or hill of Ucucuamo, consisting entirely of a very shining and yellow mica slate, which has therefore procured from the credulity of early travellers the magnificent appella¬ tion of Dorado or Golden Mountain. East from the river Essequibo this Cordillera stretches to meet the granitic or gneiss mountains of Dutch and French Guiana, inhabited by confederated bands of negroes and Caribs, but giving birth to the commercial streams of Berbice, Surinam, and Maroni. The chain of the cataracts of Orinoco has only a mean height of about 4000 feet above the level of the sea. The greatest elevation occurs where the mountain of Duida rears its enormous mass from the midst of a luxuriant plain, clothed with the tropical productions of palms and ananas, and discharges from its steep sides, about the close of the rainy season, volumes of incessant flames. Jso one has yet had the resolution or perseverance to climb through the tangling and rampant bushes to its peak, which, measured trigonometrically, gives an altitude of 8465 feet above the sea. The whole mountain group which forms this Cordillera is distinguished by the abrupt descent of its south flank; nor is it less remark¬ able for containing no rock of secondary formation, or exhibiting any vestige of petrifactions and organic re¬ mains. It contains only granite, gneiss, mica slate, and hornblende, without any casing or admixture of sand¬ stone or calcareous matter. The third great branch sent out from the trunk of the Andes is that of the Chiquitos, which province it traverses, a making a sort of semicircular sweep between the parallels V of 15 and 20 degrees south latitude, and appearing to con¬ nect the colossal heights of Peru and Chili with the moun- tains of Brazil and Paraguay. It supplies the rivers that feed the Maranon on the one side, and the Plata on the other. The structure and disposition, however, of the Cordillera of the Chiquitos still remain almost unknown. These grand chains of mountains divide the southern continent of America, from the latitude of 19 to that of 52 degrees, into three immense plains, which on the west side are shut up by the enormous ridge of the Andes, but are all open on the east, and towards the Atlantic Ocean. The most northern is the valley of the Orinoco, consist¬ ing of savannahs or level tracts covered with reedy herb¬ age and scattered palms. The next is the plain of the Maranon, which is entirely covered with dense, impene¬ trable forests. The third and southmost valley is the Pam¬ pas, a dead flat of most prodigious expanse, clothed, like that of the Orinoco, with a coarse, rank herbage, and aban¬ doned to the occupation of countless herds of wild cattle. Of these immense plains, the subsoil resembles the com¬ position of the neighbouring mountains. In the valley of the Orinoco, the primitive rock is generally wrapt in a coat of sandstone, with calcareous cement, or covered with calcareous concretions, which betray the vestiges of re¬ cent organic remains, but show none of the older impres¬ sions, such as the belemnites and ammonites, so common in Europe. The woody plain of Maranon is distinguished by the thinness of its soil, and the total absence of any calcareous ingredients, the granite approaching close to the surface, which is in some places left quite bare over an extent of many furlongs. But the Pampas of Buenos Ayres are covered to a great depth with beds of alluvial deposites, in which the powers of vegetation, fomented by the rays of a burning sun, luxuriate in wanton profusion. The lake of Titicaca covers a surface of 16,000 square miles, being in some places 70 or 80"fathoms deep. It has an elevation of 12,800 feet above the sea, and terminates at the mountain Potosi, which rises to 16,000 feet, and is yet covered with the ruins of the ancient Peruvian civili¬ zation. Near this centre the volcano of Antiquipa stands, at the height of 17,800 feet, while the double peak of the Nevados, or snowy Illimani, tow'er to the enormous elevations of 24,200 and 24,450 feet, or about 3000 feet above the summit of Chimborazo, which was long regarded as the loftiest pinnacle of our globe. But, in the northern extension of the Cordillera, Mount Sorata rears its snowy head, at the stupendous elevation of 25,000 feet. In those tropical regions cultivation ascends to very near the limits of perpetual snow. Various prolific crops, and particularly wheat and potatoes, are grown at the height of 14,000 feet above the level of the sea. A con¬ siderable population, dispersed in towns or villages, occupy tracts about 1000 feet higher, and enjoy health and vi¬ gour in a keen atmosphere, twice as rare as at the level of the sea. I he ancient Peruvians had worked some gold mines at the vast altitude of 17,000 feet. I he disciple of Werner traces with delight, in the ma¬ jestic features of the American continent, the same or¬ der and succession of rocks which the sagacity of that il¬ lustrious geologist had discovered in the mountains ol Saxony. Granite appears still the oldest material of our globe: to it succeeds the laminated species, or gneiss; then mica slate, containing garnets; next primitive slate, with beds of native alum; now slate mixed with horn¬ blende ; above this greenstone or primitive trap, followed by amygdaloid; and last of the series, porphyry slate. Resting or flanked against those primary rocks, beds of ANDES. 117 mief,. the older limestone begin to appear, followed by a suite ‘■‘yX'''of’minerals bearing indications of organic remains—mica slate, hornblende, gypsum, and calcareous sandstone. The only formations which Humboldt did not meet with.in his extensive travels, were those of chalk, roestone, gray- wacke, topaz rock, and the compound of serpentine with granular limestone which occurs in Asia Minor. The grand ridge of the Andes is everywhere covered with por¬ phyry, basalt, phonolite, and greenstone; which being often broken into columns, appear at a distance like ruin¬ ed castles, and produce a very striking effect. Near the bottom of that enormous chain, two different sorts of limestone occur ; one with a silicious base, inclosing some¬ times cinnabar and coal; and another mostly calcareous, and cementing the secondary rocks. These formations are of enormous thickness and elevation. “ Beds of coal are found in the neighbourhood of Santa Fe, 8650 feet above the level of the sea; and even at the height of 14,700, near Huanco in Peru. The plains of Bogota, al¬ though elevated 9000 feet, are covered with gypsum, sandstone, shell-limestone, and even in some parts with rock salt. Fossil shells, which in the old continent have not been discovered higher than the summits of the Py¬ renees, or 11,700 feet above the sea, were observed in Peru, near Micuipampa, at the height of 12,800 feet; and again at that of 14,120, beside Huancavelica, where sand¬ stone also appears. The basalt of Pichincha, near the city of Quito, has an elevation of 15,500 feet, while the top of the Schneekoppe in Silesia is only 4225 feet above the sea—the highest point in Germany where that spe¬ cies of rock occurs. On the other hand, granite, which in Europe crowns the loftiest mountains, is not found in the American continent above the height of 11,500 feet. It is scarcely known at all in the provinces of Quito and Peru. The frozen summits of Chimborazo, Cayambe, and Antisana, consist entirely of porphyry, which, on the flanks of the Andes, forms a mass of 10,000 or 12,000 feet in depth. The sandstone near Cuenca has a thickness of 5000 feet, and the stupendous mass of pure quartz on the west of Caxamarca measures perpendicularly 9600 feet. It is likewise a remarkable fact, that the porphyry of those mountains very frequently contains hornblende, but never quartz, and seldom mica. “ The central Andes are rich beyond conception in all the metals, lead only excepted. One of the most curious ones in the bowels of those mountains is the pacos, a com¬ pound of clay, oxyde of iron, and the muriate of silver with native silver. The mines of Mexico and Peru, so long the objects of->envy and admiration, far from being yet exhausted, promise, under a liberal and improved system, to become more productive than ever. But na¬ ture has blended with those hidden treasures the active elements of destruction. The whole chain of the Andes is subject to the most terrible earthquakes. From Coto¬ paxi to the South Sea no fewer than forty volcanoes are constantly burning; some of them, especially the lower ones, ejecting lava, and others discharging the muriate of ammonia, scorified basalt, and porphyry, enormous quan¬ tities of water, and especially moya, or clay mixed with sulphur and carbonaceous matter. Eternal snow invests their sides, and forms a barrier to the animal and vege¬ table kingdoms. Near that confine the torpor of vegeta¬ tion is marked by dreary wastes.” (Edinb. Review, vol. xv. p. 233.) \\e may subjoin that, near Quito, the liquid mud eject¬ ed by the volcanoes often involves myriads of small dead hsh. ^ In some parts, the mountains, like the fabled cave of /Eolus, seem at times to let out their imprisoned air, and produce such furious gusts of wind as to sweep every dung before them to a vast distance. In other districts, the efforts of the contending elements are betrayed, es- Andes, pecially during the rainy season, by a doleful moaning noise, or hollow and portentous groans, enough to cast a darker shade on the gloom of superstition, and to fill the ima¬ gination of the remotest settler with secret awe and dread. A person who for the first time climbs the mountains of Switzerland is astonished to witness, in the space per¬ haps of a few hours, so rapid a change of climate, and such a wide range of vegetable productions. He may be¬ gin his ascent from the midst of warm vineyards, and pass through a succession of chesnuts, oaks, and beeches, till he gains the elevation of the hardy pines and stunted birches, or treads on Alpine pastures, extending to the border of perpetual snow. But within the tropics every thing is formed on a grander scale. The boundary of permanent congelation is 7500 feet higher at the equator than at the mean latitude of 45 degrees. Under a burn¬ ing sun ananas and plantains grow profusely near the shore ; oranges and limes occur a little higher ; then suc¬ ceed fields of maize and luxuriant wheat; and the tra¬ veller has actually reached the high plain of Mexico, or the still loftier vale of Quito, before he finds a climate analogous to that of Bordeaux or of Geneva. Now only commences the series of plants which inhabit the central parts of Europe. But the very magnitude of the Andes appears to have the effect of diminishing the impressions of awe and won¬ der which the sight of them so powerfully excites. The country on which they rest is heaved to such a vast alti¬ tude above the sea, that the relative elevation of their summits becomes diminished in comparison with that of the surrounding amphitheatre. The majestic forms of Chimborazo, Cotopaxi, and Antisana, though 6000 feet higher than Mont Blanc, and clothed, like it, with eternal snow, seem to a traveller scarcely more sublime from the plains of Riobomba and Quito, than the celebrated moun¬ tain when viewed from the vale of Chamouni. It requires some time for his imagination to expand itself to the new scale of grandeur. The central Andes, with all their magnificence, want a feature which, in the higher latitudes, contributes so much to the beauty and sublimity of the Alpine scenery. They have no vestige whatever of glaciers, those icy belts dropping from the limits of congelation, and spreading in concrete sheets, or hanging in disjointed columns fantas¬ tically thrown, which occur alike in the heart of Switzer¬ land and on the northern shores of Norway and Lapland. This defect is evidently owing to the almost uniform tem¬ perature which prevails near the equator. In those tor¬ rid regions the days are constantly of the same length, and the sun shines through the whole year with very nearly equal force. The limit of perpetual congelation is hence marked on the sides of the mountains of Quito with singular precision. The temperature decreases regularly in proportion as one ascends them, till at a certain altitude it comes to the point of freezing, where the permanent field of snow begins to appear, defined with an almost un¬ varying border. But in the higher latitudes the sun re¬ mains during the summer so long above the horizon, and shines with such augmented force, that the heat of the atmosphere, and consequently of the surface of the ground, suffers a wide alteration in the different seasons. To the general investure of snow is therefore annexed every winter a zone of considerable breadth, which again softens and partly melts away during the continuance of the sum¬ mer months. This alternate thawing and freezing occa¬ sions the production of glaciers, by converting succes¬ sively the lower detached masses of snow in the precipit¬ ous flanks of the mountains, into a collection of broken and intermingled pillars of translucid ice. ANDES. For the same reason, the Andes, though torn by flaming volcanoes, and convulsed by frequent and terrible eai t - quakes, are exempt from those avalanches and eboulements which in Switzerland and other mountainous parts of Eu¬ rope often bury the helpless traveller in a torrent of snow, and batter down whole villages by the sudden discharge of a shower of rocks. Under the equator the variation of temperature throughout the year is so small as not to disturb the solidity of the vast collections of snow; but on the flanks of the Alps or Pyrenees, as the heat of the summer increases, portions of the upper field of snow be¬ come loosened, and, sliding down, put other masses like¬ wise in motion, till spreading wider, and gaining accele¬ rated force, the whole tide precipitates itself to the plain, sweeping all before it. Such is the accident of an ava¬ lanche ; "but the occurrence of an eboulement, though less frequent, is more tremendous. When the alternation of frost and thaw detaches a mass of rock, it rolls down the side of the mountain with resistless fury, shivering into frag¬ ments and tearing every thing opposed to it, overwhelm¬ ing men, cattle, and houses, in one common heap of ruins. But the Andes are distinguished from the chains of the European mountains by frightful quebradas or perpendi¬ cular rents, which form very narrow vales of immense depth, whose terrific walls, fringed below with luxuriant trees and shrubs, seem to lift their naked and barren heads to the distant skies. The noted crevices of Chota and Cutaco are nearly a mile deep, the former measuring 4950, and the latter 4300 feet, in a vertical descent. The task of crossing such tremendous gullies is often a work of infinite toil and extreme danger. In those mountain¬ ous countries travellers are accustomed to perform their journeys sitting in chairs fastened to the backs of men called cargueros or carriers. These porters are mulattoes, and sometimes whites, of great bodily strength and ac¬ tion, who will climb along the face of precipices, bearing loads of twelve and fourteen, or even eighteen stone. The cargueros lead a vagabond life, exposed to incredible fa¬ tigue, but recommended to them by its irregular course. Often those wretched men toil over mountains for the space of eight or nine hours every day, till, like beasts of burthen, they have their backs chafed and made quite raw with the load. In this deplorable condition they are not unfrequently abandoned by unfeeling travellers, and left alone to sicken, pine, or die in the forests. Yet their earnings would appear inadequate to such violent and overpowering exertions, since they receive scarcely three guineas for performing the journey from Ibague to Car¬ tage, which requires fifteen, and perhaps twenty-five or thirty days. The Icononzo, remarkable for its natural bridges, is a small quebrada or cleft of the mountains, through which flows the river of the Summa Paz, descending from the highest upland desert. The rocks consist of two differ¬ ent kinds of sandstone, the one extremely compact, and the other of a slaty texture, divided into their horizontal strata. The rent was probably caused by an earthquake, which the harder portion of the stony mass had resisted, and now connects the upper part of the chasm. This natural arch is 50 feet long, 40 broad, and 8 feet thick at the middle. Its height is about 300 feet above the sur¬ face of the torrent, which has a medium depth of twenty feet. About 60 feet below the natural bridge another smaller arch occurs, composed of three slanting blocks of stone wedged together, which had probably fallen from the roof at the same instant of time, and struck against the sides of the crevice in their descent. The natural bridge of Icononzo has perhaps no coun¬ terpart in the old world ; but the writer of this article had the pleasure of seeing, in early life, a similar pheno¬ menon scarcely inferior to it in the United States of Aire- ^ rica. We allude to the famous arch described by MrU Jefferson, which crosses the Cedar creek in Rockbridge county, about a hundred miles beyond the Blue Ridge, in the higher district of Virginia. The divided rock is a pure limestone, leaving a chasm about 90 feet wide, of which the walls are 230 feet high, sprinkled with verdant bushes, and enamelled with gay flowers, among which the aquilegia is conspicuous. This bridge, viewed from a little distance below, has all the appearance of a Gothic arch ; and is of such solidity, that loaded waggons used formerly to pass along it, till a more convenient line of road was formed. In some places the natives of Peru connect the clefts of their mountains by pendulous bridges thrown fearlessly across, and suspended from both sides of a gap. They are composed of ropes made of the tough fibres of the agave, hanging in a gently sloping curve, and covered with reeds or canes, with occasionally a narrow border of bas¬ ket-work. The intrepid Indian, regardless of the horrors I of the unfathomed abyss which yawns from below, com¬ mits himself to his frail and floating arch, and swiftly glides along its bending curvature, till he gains the oppo¬ site bank. The Andes likewise give rise to waterfalls of immense height and amazing force. The cataract of Tequendama, considered in all its circumstances, exceeds any other in the known world. The basin which feeds its streams is the vast plain of Bogota, 7465 feet above the level of the sea, encircled completely with lofty mountains, except where the water, aided probably by the concussion of an earthquake, has cut for itself a narrow passage. The river Funcha, swelled by numerous feeders, gradually contracts its channel to the breadth of about 40 feet, and then ga¬ thering augmented force, dashes at two bounds from a perpendicular height of near 600 feet into a dark gulf. Owing to the excessive rapidity and depth of its current, it must discharge a prodigious volume of water, which quite stuns the ear by the roar of its crash; while it raises enormous clouds of thick spray and vapour, that continu¬ ally bedew, and perhaps quicken, the vegetation of the adjacent grounds. Every thing conspire's to exalt the beauty and grandeur of , the scenery. “ Independent of the height and mass of the column of water,” says Hum¬ boldt, “ the figure of the landscape, and the aspect of the rocks, it is the luxuriant form of the trees and herbace¬ ous plants, their distribution into thickets, the contrast ol those craggy precipices, and the freshness of vegetation, which stamp a peculiar character on these great scenes of nature.” The transition from a temperate to a warm cli¬ mate is rapid and surprising. The plain of Bogota bears rich crops of wheat, then succeed oaks and elms, inter¬ mingled with aralias, bigonias, and the yellow-bark trees; but immediately below the cataract a few palms appear, as if it were to mark the advance to a sultry soil. A lively idea of the character and grand features of the Andes may be conceived from the account which the celebrated Humboldt has given of his journey across that majestic chain. Our readers will be glad to peruse it in the author’s own words. “ The mountain of Quindiu, in the latitude of 4° 36', is considered as the most difficult passage in the Cordilleras of the Andes. It is a thick uninhabited forest, which, in the finest season, cannot be traversed in less than ten or twelve days. Not even a hut is to be seen, nor can any means of subsistence be found. Travellers, at all times of the year, furnish themselves with a month’s provision, since it often happens that, by the melting of the snows, and the sudden swell of the torrents, they find themselves so circumstanced that they can descend neither on the AND de side of Cartago nor that of Ibague. The highest point rv/0f the road, the Garito del Paramo, is 11,500 feet above the level of the sea. As the foot of the mountain, to¬ wards the banks of the Cauca, is only 3150 feet, the cli¬ mate there is generally mild and temperate. The path¬ way, which forms the passage of the Cordilleras, is only 12 or 15 inches in breadth, and has the appearance, in several places, of a gallery dug and left open to the sky. In this part of the Andes, as almost in every other, the rock is covered with a thick stratum of clay. The stream¬ lets which flow down the mountains have hollowed out gullies about 20 feet deep. Along these crevices, which are full of mud, the traveller is forced to grope his pas¬ sage, the darkness of which is increased by the thick vegetation that covers the opening above. The oxen, which are the beasts of burden commonly used in this country, can scarcely force their way through these gal¬ leries, some of which are more than a mile in length; and ,*. if perchance the traveller meets them in one of these passages, he finds no means of avoiding them but by turn¬ ing back and climbing the earthen wall which borders the crevice, and keeping himself suspended by laying hold of the roots which penetrate to this depth from the surface of the ground. “ We traversed the mountain of Quindiu in the month of October 1801, on foot, followed by twelve oxen, which carried our collections and instruments, amidst a deluge of rain, to which we were exposed during the last three or four days, in our descent on the western side of the Cor¬ dilleras. The road passes through a country full of bogs, and covered with bamboos. Our shoes were so torn by the prickles which shoot out from the roots of these gi¬ gantic gramina, that we were forced, like all other travel¬ lers who dislike being carried on men’s backs, to go bare¬ footed. This circumstance, the continual humidity, the length of the passage, the muscular force required to tread in a thick and muddy clay, the necessity of fording deep torrents of icy water, render this journey extremely fatiguing; but, however painful, it is accompanied by none ol those dangers with which the credulity of the people alarms travellers. The road is narrow, but the places where it skirts the precipices are very rare. “ When travellers reach Ibague, and prepare to cross the forests of Quindiu, they pluck, in the neighbouring mountains, several hundred leaves of the vijao, a plant of the family ol the bananas, which forms a genus approach¬ ing to the Thalia, and which must not be confounded with the Heiiconia Bibai. These leaves, which are membranous and silky, like those of the Musa, are of an oval form, two feet long and 16 inches broad. Their lower surface is a silvery white, and covered with a farinaceous substance, which falls oft in scales. This peculiar varnish enables them to resist the rain during a long time. In gathering these leaves, an incision is made in the middle rib, which is the continuation of the foot-stalk; and this serves as a hook to suspend them when the movable roof is formed. On taking it down, they are spread out, and carefully rol¬ led up in a cylindrical bundle. It requires about an hun¬ dredweight of leaves to cover a hut large enough to hold six or eight persons. When the travellers reach a spot in the midst ot the forests where the ground is dry, and where they propose to pass the night, the cargueros lop a few blanches from the trees, with which they make a tent. In a few minutes this slight timber-work is divided into squares, by the stalks of some climbing plant, or the threads of the agave placed in parallel lines 12 or 13 inches from each other. The vijao leaves meanwhile have sen unrolled, and are now spread over the above work, tl! aV° ?over .‘t the tiles of a house. These huts, us hastily built, are cool and commodious. If, during AND 119 the night, the traveller feels the rain, he points out the Andeuse spot where it enters, and a leaf is sufficient to obviate the II inconvenience. We passed several days in the valley of Andrada. Boquia, under one of those leafy tents, which was perfect- ly dry amidst violent and incessant rains.” For further information relative to the structure of the Andes, see the various sketches given by Humboldt, and particularly an abstract of his geological observations in¬ serted in the Journal de Physique, vol. liii. for 1801. See likewise, by the same author, a memoir on the Geographic and Geognostic labours of M. Pentland, in the Nouvelles Annates des Voyages et des Sciences Geographiques for October, November, and December, 1829. (b.) ANDEUSE, a city of Languedoc in France, situated in long. 3. 40. E. and lat. 43. 45. N. ANDORRE, or Andorra, a valley on the Spanish side of the Pyrenees, of about 190 square miles in extent, con¬ taining 6 parishes, and which has ever been considered as neutral. It is a hilly district, with some good pasture for sheep, and several mines of iron, which are worked by the numerous rapid streams that pass through the district in their course towards Catalonia. It is a kind of independent republic, governed by two deputies chosen in each of the six parishes, and by two syndics. The expenses of govern¬ ing are defrayed by a species of rent paid by owners of flocks, to the community, for the use of the pasture land. The king of France is called the protector in civil affairs, and the bishop of Urgel in ecclesiastical matters. It en¬ joys its peculiar criminal and civil lawrs and courts of justice. The chief judge is nominated alternately by the king of France and the bishop of Urgel, but there is no appeal to either France or Spain from his decisions. As the land does not produce sufficient corn for the supply of the inhabitants, a stipulated quantity is allowed annually to be introduced from France, for which permission a sub¬ sidy of 960 francs is granted to the king. The whole number of the inhabitants is about 15,000, who are neither French, nor Spaniards, nor Catalans, but speak a language containing a mixture of the three idioms. The council of state and the courts of justice assemble at the village of Andorra, and the former body retain the ancient Spanish title of cortes. ANDOVER, a market and borough town in the hun¬ dred of the same name, in Hampshire. It is on the river Ande, 63 miles from London, on the great road to the west of England. It is a clean, well-built town, carry¬ ing on some trade in shalloons, and more in malt. It is connected with the sea by means of a canal, navigable to Southampton. Two members are returned to parlia¬ ment, elected by the corporation, consisting of about 24 persons, who are said to be under the influence of Lord Portsmouth and Mr Etwall. Population in 1801, 3304 ; in 1811, 3701; and in 1821, 4123. ANDRA, or Andros, a Turkish paschaliac, including the island of that name and the neighbouring islands of Tine, Mitone, Delos, Syra or Syros, Thermia Zea, and Hydra. The inhabitants are estimated at 96,800, almost wholly Greeks; the most industrious, and, exclusive of the effects of the present war, the most flourishing people of the Archipelago. There are some smaller islands in¬ cluded in the paschaliac, whose whole extent is 474 square miles. The island of Andros contains 12,000 inhabitants, mostly Greeks, who raise much silk, wine, and oil, and suf¬ ficient corn for their own consumption. ANDRADA, Diego de Payva d’, or Andradius, a learned Portuguese, born at Coimbra in 1528, who dis¬ tinguished himself at the council of Trent, where King Sebastian sent him as one of his divines. He wrote seve¬ ral volumes of sermons, and other pieces ; one of which, De Conciliorum Autoritate, was highly esteemed at Rome 120 AND AND Andrapo- on account of the great extension of authority which it ANDRETTA, a town, with 4049 inhabitants, in the Ai, ^ disraus f^civc to the pope* He died, in l^TS* province Prmcipnto h/lteriore or the Kingdom of Nciplcs* | ,11. fe ANDR APODISMUS, in ancient writers, the selling of ANDREW, St, the apostle, born at Bethsaida in Ga- ' Andrehnus. sons for s]aves> Hence also andrapodistes, a dealer in lilee, brother to Simon Peter. He had been a disciple of^ ' slaves, more particularly a kidnapper, who steals men or John the Baptist, and followed Jesus, upon the testimony ' children to sell them,—a crime for which the Thessalians given of him by the Baptist. (John i. 35, 40, &c.) This were noted. was tlie first*disciple whom our Saviour received into his ANDRAPODOC APELI, in Antiquity, a kind of dealers train. Andrew introduced his brother Simon, and they in slaves. The Andrapodocapeli had a particular process passed a day with Christ, after which they went to the for taking off moles and the like disfigurements on the faces marriage in Cana (ibid, ii.), and at last returned to their of the staves they kept for sale, by rubbing them with ordinary occupation. Some months after, Jesus meeting bran. At Athens, several places in the forum were ap- them while they were both fishing together, called them pointed for the sale of slaves. Upon the first day of every to him, and promised to make them fishers of men. Im- month, the merchants called AvcSgawrodoxaOTjXo/ brought them mediately they left their nets, followed him (Matt. iv. 19), into the market and exposed them to sale: the crier, stand- and never afterwards separated from him. After our Sa- ing upon a stone erected for that purpose, called the viour’s ascension, his apostles having determined by lot people together. what parts of the world they should severally take, Scy- ANDREAS, John, a celebrated canonist of the 14th thia and the neighbouring countries fell to St Andrew, century, was born at Magello, near Florence ; and was pro- who, according to Eusebius, after he had planted the gos- lessor of the canon law at Padua, Pisa, and afterwards at pel in several places, came to Patrse in Achaia, where, Bologna. He had a beautiful daughter, named Novella, endeavouring to convert the proconsul fEgeas, he was by whom he is said to have instructed so well in all parts of that governor’s orders scourged, and then crucified. The learning, that when he was engaged in any affair which particular time of his suffering martyrdom is not known; hindered him from reading lectures to his scholars, he sent but all the ancient and modern martyrologies, both of the his daughter in his room ; and, lest her beauty should pre- Greeks and Latins, agree in celebrating his festival upon vent the attention of the hearers, she had a little curtain the 30th of November. drawn before her. To perpetuate the memory of this Andrew, or an order of knights, daughter, he entitled his commentary upon the Decretals more usually called the Order of the Thistle, of Gregory IX. the Novella. He married her to John Knights of St Andrew is also an order instituted by Calderinus, a learned canonist. The first work of Andreas Peter the Great of Muscovy in 1698, the badge of which was his Gloss upon the Sixth Book of the Decretals, which is a golden medal, on one side whereof is represented St he wrote when he was very young. He wrote also Glosses Andrew’s cross, with these words, Czar Pierre, monarque upon the Clementines ; and a Commentary in Regulas Scxti, de Unit le Kussie. This medal being fastened to a blue which he entitled Mercuriales, because he either engaged ribbon, is suspended from the right shoulder, in it on Wednesdays (diebus Mercuni), or because he in- St Andrew's Day, a festival of the Christian church, serted his Wednesday’s disputes in it. He enlarged the celebrated on the 30th of November in honour of the Speculum of Durant in the year 1347. He died of the apostle St Andrew. plague at Bologna in 1348, after he had been a professor ANDREWS, St, a city of Scotland, in the county of 45 years, and was buried in the church of Dominicans. Fife, pleasantly situated in a spacious bay of the German Andreas, St, a market-town on the Danube, in the Ocean, into which flow the river Eden, the small rivulet circle of Pilesch, and palatinate of Pest, in Hungary. It of Kinness, and several other streams. It was formerly contains one Catholic and seven Greek churches, 1040 a place of much greater extent and importance. At pre¬ houses, and 7980 inhabitants, who trade very extensively sent it is only about a mile and a half in circuit, and con- in the wine produced in the vicinity. It is also the name sists of three leading streets, intex*sected by a few consi- of an island in the Danube, opposite to the town, which is derable lanes. The principal street is well built, and is fourteen miles in length and one in breadth, and remark- straight and broad, and of late years its appearance has able for its great fertility. been much improved. The town contains many* interest- ANDREASBERG, a bailiwick in the pi'ovince of Gru- ing memorials of antiquity. Of the splendid cathedral, benhagen, in the kingdom of Hanover. It extends over which was founded by Bishop Arnold in 1159, and at- 87 square miles, or 55,680 acres, containing 4250 inhabit- tained to its highest magnificence in 1318, part of the ants, who are chiefly employed in mining. The soil is east and west ends, and of the south side, are all that very poor, and affords scarcely any produce ; but, like the now remain. The length of this edifice from east to west rest of the Hartz forest, it abounds with metallic sub- was 350 feet within the walls ; that of the transept 180 feet, stances. The capital of the bailiwick is of the same name. This whole pile of building, which it took 150 years to It is 1880 feet above the level of the sea, on the declivity complete, was in June 1559 demolished in a single day of the Hartz mountains, containing 400 houses, and 3206 by John Knox and his infuriated followers. About 40 inhabitants, who subsist chiefly by mining, whilst the fe- yards to the south-east is the chapel of St Regulus, the males are employed in making lace. Near it is the An- tower of which is a lofty square prism, the side of the base dreasberg silver mine, from which is produced annually being 20 feet, and the height 108. The chapel to the east about 40,000 ounces of silver, besides a large quantity of of the tower, which was the principal one, remains; but lead and copper, all of which are smelted in the town. of a small chapel to the west, which formerly existed, ANDRELINUS, Publius Faustus, born at Forli in there is now no trace. The arches of the windows Italy. He was a long time professor of poetry and philoso- and doors are round, and the figure of some of them phy in the university of Paris. Louis XII. of France made is more than half of the circle, which is an undoubted him his poet laureat, and Erasmus tells us that he was like- proof of their antiquity. The priory, which was founded wise poet to the queen. His pen was not wholly employ- by Robert, bishop of St Andrews, during the reign of ed in making verses, for he wrote also moral and prover- Alexander I. in 1120, was of great extent, and richly bial letters in prose, which were printed several times, endowed. The prior had precedence of all abbots and Flis poems, which are chiefly in Latin, are inserted in vol. i. priors, and on festival days had a right to wear a mitre of the Delicice Poetarum Italorum. He died in 1518. and all episcopal ornaments. The walls of the pre- AND AND 121 -c'‘, cinct are all that now remain to mark the vast extent ros- of this edifice. Part of the top of the great altar towards “^the east end of the cathedral was discovered between 30 and 40 years ago, in consequence of an excavation made in the hope of finding concealed treasure. At a much more recent period, the soil, which had for ages been permitted to accumulate, having been removed, part of the pavement was laid open, and several shafts of two rows of pillars parallel to each other, by which it is supposed that galleries in the inside of the walls had been supported. The other religious houses were, that of the Dominicans, founded in 1274 by Bishop Wishart; another of Observantines, founded by Bishop Kennedy, and finished by his successor Patrick Graham in 1478; and, according to some, the Carmelites had a fourth. Immediately above the harbour stood the collegiate church of Kirk-heugh, originally founded by Constantine III., who, retiring from the world, became here a Culdee. From its having been first built on a rock, it was styled Prcepositura Sanctce Marim de Rupe. On the east side of the city are the remains of the castle, on a rock overlooking the sea. This fortress was founded about the year 1200, by Roger, one of the bishops of St Andrews, and was repaired towards the end of the 14th century by Bishop Trail, who died in it in 1401. Pie was buried near the high altar of the cathedral, with this singular epitaph: Hie fuit ecclesise directa columna, fenestra Lucida, thuribulum redolens, campana sonora. The castle was the residence of Cardinal Beaton, who, after the cruel execution of the celebrated reformer George Wishart in front of it, was afraid of the fury of the people; and his knowledge of this, joined to his ap¬ prehension of an invasion from England, induced him to strengthen the fortifications, with a view of rendering the castle impregnable. In this fortress he was surprised and assassinated by Norman Lesley, aided by fifteen others. Early in the morning of May 29, 1546, they seized on the gate of the castle, which had been left open for the work¬ men who were finishing the fortifications; and having placed sentinels at the door of the cardinal’s apartment, they awakened his numerous domestics one by one, and, turning them out of the castle, without violence, tu¬ mult, or injury to any other person, inflicted on Beaton the death he justly merited. The conspirators were im¬ mediately besieged in this castle by the regent, earl of Arran; and although their strength consisted of only 150 men, they resisted his efforts for five months, owing more to the unskilfulness of the attack than the strength of the place, for in 1547 the castle was reduced and demo¬ lished, and its picturesque ruins serve as a land-mark to mariners. The entrance to the castle, and the window out of which it is said Cardinal Beaton leaned to witness the cruel martyrdom of George Wishart, are still pointed out. The parish church is a spacious structure, 162 feet in length by 63 in breadth, and is large enough to accommo¬ date 2500 persons. It contains a lofty monument of white marble, erected in honour of Archbishop Sharpe, w ho, in revenge for his oppressive conduct, was murdered by some of the exasperated reformers of that day. On this monu¬ ment is a rude piece of sculpture representing the tragical scene of the murder. To the north is situated the col- lege church, which belongs to the united college of St Salvator and St Leonard. It was founded in 1458 by Bishop Kennedy, and contains a beautiful tomb of its ounder, who died in 1466; wdiich is a fine specimen of ififtQ °^1*C ar<:llitecture ^iat period. About the year oo3, on opening this tomb, six highly ornamented silver vol. m. & J maces Were discovered, which had been concealed there St in times of trouble, three of which are still preserved in Andrews, the university, and three were sent to the other universi- ties of Scotland. On the top is represented our Saviour; around are angels with the instruments of his passion. With these are shown some silver arrows, with large sil¬ ver plates affixed to them, on which are inscribed the arms and names of those who were victors in the annual compe¬ titions of archery, which were regularly held until within these few years. Golf is now the reigning game. That sport, and foot-ball, were formerly prohibited by an act of parliament passed in the reign of James II. in 1457, as interfering too much with the acquisition of dexterity in archery, an accomplishment in those days of much conse¬ quence to the safety of the state. The statute has been long obsolete, and the inhabitants, and the students who attend the university, have full permission to enjoy this elegant amusement. The celebrated university of this city was founded in 1411 by Bishop Wardlaw. It consisted formerly of three colleges. 1. St Salvator’s was founded in 1458 by Bishop Kennedy. This was a handsome building, with a court or quadrangle, which is 230 feet long by 150 wide, and a gateway surmounted by a spire 156 feet high. On one side is the church; on another what was the library of St Salvator’s; the third contains apartments for students; the fourth is unfinished. The buildings connected with the college being in a state of great decay, a grant has been made by government for erecting a new structure. Part of this building will soon be finished, and the remainder, it is expected, will be immediately after commenced. When it is completed it will be in a high degree elegant and commodious. 2. St Leonard’s College was founded by Prior Hepburn in 1522. This is now united with the last, and the buildings sold, and converted into private houses. 3. New or St Mary’s College was established by Archbishop Hamilton in 1552; but the house was com¬ pleted by Archbishop Beaton. This is said to have been the site of a celebrated school long before the esta¬ blishment even of the university, where several eminent clergymen taught gratis the sciences and languages ; but it was called the New College, because of its late erection into a divinity college by the archbishop. The buildings of this college have been substantially repaired, and with great taste. The university is governed by a chancellor,—an office which it was originally intended should be permanently exercised by the archbishop of St Andrews. Subsequent to the Reformation, the chancellor has been elected by the two principals and the professors of both colleges. The rector is the next great officer, to whose care are committed the privileges, discipline, and statutes of the university. The colleges have their principals, and pro¬ fessors of different sciences, who are indefatigable in their attention to the instruction and the morals of the students. The place possesses, in its pure and salubrious air, and in its extensive grounds for exercise, very great advantages for the education of youth. In St Salvator’s College ai’e taught the languages, philosophy, and the sciences. St Mary’s, which stands in a different part of the town, is re¬ served exclusively for theology. The classes and disci¬ pline of the two colleges are quite distinct, each having its respective principal and professors. They have a com¬ mon library, which is entitled to a copy of every work en¬ tered in Stationers Hall, and which now (1830) contains upwards of 40,000 volumes. An addition has been in the course of last season made to the library, which was much required, in consequence of the great increase of the number of books. Seventy-five bursaries or endowments belong to the tt 122 AND St university, and are conferred on the students. Of these, Andrews. 58 belong to the United, and 17 to the New College. The number of students at St Andrews never exceeded 300 at any period. During the present year (1830) they amount to 190 at both colleges—149 at the United Col¬ lege, 41 at St Mary’s. The trade of St Andrews was once very considerable. So late as the reign of Charles I. it had thirty or forty trading vessels, and carried on a considerable herring and white fishery, by means of busses, in deep water, which had for ages been a most profitable branch of commerce, and a source of wealth. During the troubles which fol¬ lowed the death of this monarch, this whole coast, and St Andrews in particular, became a scene of murder and ra¬ pine ; and every town suffered in proportion to its magni¬ tude and opulence. St Andrews was required to pay a contribution of L.1000, which the inhabitants, after being plundered, were not able to raise : a composition of L.500 was accepted, which was raised by a loan at interest, and has remained a burden upon the corporation, it is believ¬ ed, ever since. The harbour of St Andrews is artificial: it is guarded by piers, and is safe and commodious; but it is difficult of access, having a narrow entrance, and being exposed to the east winds, which raise a heavy sea on the coast. The shore of the bay is low; and, in the storms of winter, vessels are frequently driven on it and lost. St Andrews had a manufactory of sail-cloth to some ex¬ tent, but it is now discontinued. The game of golf being much practised here, there is a manufactory of golf balls, which, after supplying the home consumption, sends about 9000 annually to other parts. The shipping of the port consists of about eleven vessels, which are employed in the coasting trade. St Andrews is a royal borough, uniting with Cupar, Perth, Dundee, and Forfar, in returning a member to parliament. According to early traditions, St Andrews owes its ori¬ gin to a Greek monk, Regulus, who having been warned in a vision to visit Albion, was shipwrecked in the bay about the end ot the fourth century. He was hospitably received, according to these ancient accounts, by the king, who presented him with his own palace, and built near it the church of St Regulus, the remains of which are still to be seen. At this time the place was styled Mucross, or the land of boars. All around was forest, and the lands bestowed on the saint were called Byrehid. The boars equalled in size the ancient Erymanthian ; as a proof of which, two tusks, each sixteen inches long and four thick, were chained to the altar of St Andrew. The king changed the name to Kilrymont, and established here the first Christian priests of the country, called Culdees. The church was supreme in the kingdom of the Piets, Hungus having granted to God and St Andrew that it should be the head and mother of all the churches in his dominion. He also directed that the cross of St Andrew should become the badge of the country. In 818, after the conquest of the Piets, he removed the episcopal see to St Andrews, and the bishop was styled maximus Sco- torum episcopus. In 1471 it was erected into an arch¬ bishopric by Sextus IV. at the intercession of James III. In 1606 the priory was suppressed; and in 1617 the power of election was transferred to eight bishops, the principal of St Leonard’s College, the archdeacon, the vicars of St Andrews, Leuchars, and Cupar. This see contained the greater part of the shire of Fife, with a part of the counties of Perth, Forfar, and Kincardine, and a great number of parishes, churches, and chapels in other dioceses. The town of St Andrews was erected into a royal bo¬ rough by David I. in the year 1140, and its privileges af- AND terwards confirmed. The charter of Malcolm II., written Am; on a small bit of parchment, is preserved in the tolbooth.^i Here also are kept the silver keys of the city, which, for form’s sake, are delivered to the king if he should visit : the place, or to a victorious enemy in token of submission. In this place, likewise, is to be seen the enormous axe with which, in 1646, Sir Robert Spotswood and other dis¬ tinguished loyalists were beheaded. The town under¬ went a siege in 1337, at which time it was possessed by the English and other partisans of Baliol; but the loyal¬ ists, under the earls of March and Fife, made themselves masters of it in three weeks, by the help of their batter¬ ing machines. It is 9 miles E. of Cupar, and 39 N. N. E. of Edinburgh. The population, according to the last cen¬ sus, was 4899. Long. 2. 50. W. Lat. 56. 19. 33. N. (f.) ANDREWS, James Pettit, a late English historian and miscellaneous writer, was the younger son of Joseph Andrews, Esq. of Shaw-house, near Newbury, Berks, where he was born in the year 1737. He was educated privately, and is said to have discovered an early taste for literature and the fine arts. He joined the Berkshire mi¬ litia when they were first called out, being then about 18 or 19, and held the rank of lieutenant in that regiment until it was disbanded. On the institution of the new system of London police, he was appointed one of the commissioners for the district ol Queen’s Square and St Margaret’s, Westminster, and discharged the duties of that office with great industry and integrity until his death, which took place at his house in London, on the 6th of August 1797, in the 60th year of his age. He married Miss Ann Penrose, daughter of the Rev. Mr Pen¬ rose, late rector of Newbury, by whom he had two sons and a daughter. He seems to have possessed a cheerful and social disposition, and enjoyed the conversation of a large circle of literary acquaintance, who frequently met at his house and experienced his hospitality. Mr Andrews appears to have devoted a considerable portion of his time to literary pursuits; and he is the au¬ thor of several works which are not undeserving of notice. The first publication upon which we find him employed is an edition of the poems of his friend and relation Penrose, in 1781; to which he prefixed an introduction, containing a short account of the life and character of the author. His first original production, so far as we have been able to ascertain, was a pamphlet in behalf of the chimney¬ sweepers’ apprentices in 1788, which is said to have led to the act of parliament passed not long afterwards for the purpose of ameliorating the condition of that unfortu¬ nate class of beings. In 1789 he published Anecdotes, An¬ cient and Modern, 8vo, a work of pleasantry, in the com¬ position of which he acknowledged having received assist¬ ance from the late laureat Mr Pye, the facetious antiquary Captain Grose, and others. To this volume he added a Supplement in 1790. The most extensive work undertaken by Mr Andrews was his History of Great Britain, connected with the Chro¬ nology of Europe; with Notes, &c. The first volume, which commences with Caesar’s invasion, and ends with the deposition and death of Richard II., was published in 1794 in 4to. A second volume, in which the history is continued from the deposition and death of Richard II. to the accession of Edward VI., appeared in 1795. The plan of this work is new, and in some respects singular; a cer¬ tain portion of the history of England is given on one page, and a corresponding portion of the contemporaneous history of Europe on the one opposite. The notes consist of a variety of curious and amusing particulars, not imme¬ diately connected with the main story. Appendixes are also added at proper intervals, containing an account ol the state of literature, science, religion, government, ma11’ AND tew ners, &c. at different periods. The author, however, did | not live to complete this curious and extensive work, no Uro- more of it having appeared than the two volumes above nes* mentioned. In 1796 he published a continuation of /^Henry’s History of Britain, in one volume 4to and two volumes 8vo. t „ 1 . The other productions of this author are, An Account oj Saxon Coins found in Kintbury Churchyard, Berks, print¬ ed in the seventh volume of the Archceologia; the Ac¬ count of Shaw, in Mr Mores’s Berkshire Collections; The Savages of Europe, a popular French novel, which he translated, and illustrated by prints from his own designs. Mr Andrews was also a frequent contributor to the Gentlemans Magazine. See the Gen. Biog. Diet, by Chalmers; Introduction to Poems by the Rev. Thomas Penrose, 1781; Gent. Mag. for 1797 and 1801; and Ly- sons’ Supplement to Environs of London, 1811. (k.) Andrews, Lancelot, bishop of Winchester, was born at London in 1555, and educated at Cambridge. After se¬ veral preferments he was made bishop, first of Chiches¬ ter, then of Ely, and in 1610 was raised to the see of Winchester. This very learned prelate, who was distin¬ guished by his piety, charity, and integrity, may be justly ranked with the best preachers and scholars of his age. He appeared, however, to much greater advantage in the pulpit than he does now in his works, which abound with Latin quotations and trivial witticisms. He died at Win¬ chester House in Southwark, September 27, 1626, in the 71st year of his age, and was buried in the parish church of St Saviour’s, where his executors erected to him a very fair monument of marble and alabaster, on which is an elegant inscription in Latin, written by one of his chaplains. His most popular works are, his Sermons, his Lectures on the Ten Commandments, and his Orphan Lec¬ tures, each forming a folio volume. There is a collection by Felix Kyngston, of some other pieces written by him, which was published in 4to in 1629. ANDRIA, in Grecian Antiquity, public entertainments first instituted by Minos of Crete, and, after his example, appointed by Lycurgus at Sparta, at which a whole city or a tribe assisted. They were managed with the utmost frugality, and persons of all ages were admitted. Andria, a city and a bishop’s see in the territory of Bari, in the kingdom of Naples. It is pretty large, well peopled, and seated in a spacious plain, four miles from the Adriatic coast. Long. 17. 4. E. Lat. 41. 15. N. ANDRISCUS, a man of mean extraction, who, pre¬ tending to be the son of Perseus, last king of Macedonia, took upon him the name of Philip, for which reason he was called Pseudo-Philippus, the False Philip. After a complete victory over Juventus, the Roman pretor who was sent against him, he assumed kingly power, but ex¬ ercised it with vast cruelty. At last the Romans obliged him to fly into Thrace, where he was betrayed and de¬ livered into the hands of Metellus. This victory placed Macedonia once more in the power of the Romans, and gained for Metellus the name of Macedonicus, but cost the Romans 25,000 men. Andriscus adorned the triumph of Metellus, walking in chains before the general’s chariot. ANDROGEUS, in Fabulous History, the son of Minos, king of Crete, was murdered by the Athenian youth and those of Megara, who envied his being always victor at the Attic games. But Minos having taken Athens and Megara, obliged the inhabitants to send him an annual tribute of seven young men and as many virgins, to be devoured by the Minotaur. From this tribute they were delivered by Theseus. androgynes, in Ancient Mythology, creatures of whom, according to the fable, each individual possessed the powers and characters of both sexes, having two heads, AND 123 four arms, and two feet. The word itself is compounded Andro- of two Greek radical words ; oo^g, in the genitive avSgog, a gynes male; and yuvTj, & female. Many of the rabbinical writers pretend that Adam was created double, one body being male, the other female, which in their origin not being essentially joined, God afterwards did nothing but sepa¬ rate them. Androgynes, in Natural History, a name given to those living creatures which, by a monstrous formation of their generative parts, seem (for it is only seeming) to unite in themselves the two sexes, that of the male and of the female. See Hermaphrodite. ANDROIDES, in Mechanics, a human figure, which, by certain springs or other movements, is capable of per¬ forming some of the natural motions of a living man. The motions of the human body are more complicated, and consequently more difficult to be imitated, than those of any other creature; whence the construction of an an- droides, in such a manner as to imitate any of these ac¬ tions with tolerable exactness, is justly supposed to indi¬ cate a greater skill in mechanics than any other piece of workmanship whatever. A very remarkable figure of this kind appeared in Pa¬ ris in the year 1738. It represented a flute-player, and wras capable of performing different pieces of music on the German flute; which, considering the difficulty of blowing that instrument, the different contractions of the lips necessary to produce the distinctions between the high and low notes, and the complicated motions of the fingers, must appear truly wonderful. This machine was the invention of M. Yaucanson, member of the Royal Academy of Sciences; and a par¬ ticular description of it was published in the Memoirs of the Academy for that year. The figure itself was about 51 feet in height, situated at the end of an artificial rock, and placed upon a square pedestal 41 feet high and 31 broad. The air entered the body by three pipes separated one from the other. It was conveyed to them by nine pairs of bellows, three of which were placed above and six below. These were made to expand and contract regularly in succession by means of an axis of steel turned round by some clock¬ work. On this axis were different protuberances at pro¬ per distances, to which were fixed cords thrown over pul¬ leys, and terminating in the upper boards of the bellows, so that, as the axis turned, these boards were alternately raised and let down. A contrivance was also used to prevent the disagreeable hissing fluttering noise usually attending the motion of bellows. This was by making the cord by which the bellows was moved press, in its descent, upon one end of a smaller lever, the other end of which ascending, forced open the small leathern valve that admitted the air, and kept it so till, the cord being relaxed by the descent of the upper board, the lever fell, and the air was forced out. Thus the bellows performed their functions constantly without the least hissing, or other noise by which it could be judged in what manner the air was conveyed to the machine. The upper boards of three of the pairs of bellows were pressed down by a weight of 41bs., those of three others by a weight of 21bs., and those of the three remaining ones by nothing but their own weight. The three tubes by which the air entered terminated in three small reservoirs in the trunk of the figure. There they united, and, ascending towards the throat, formed the cavity of the mouth, which terminated in two small lips adapted in some measure to perform their pro¬ per functions. Within this cavity also was a small mov¬ able tongue, which, by its play, at proper periods ad¬ mitted the air, or intercepted its passage to the flute. 124 AND AND Androides. The fingers, lips, and tongue, received their proper di- rections by means of a steel cylinder turned by clock¬ work. It was divided into 15 equal parts, which, by means of pegs pressing upon the ends of 15 different le¬ vers, caused the other extremities to ascend. Seven of these levers directed the fingers, having wires and chains affixed to their ascending extremities, which being at¬ tached to the fingers, caused them to ascend in proportion as the other extremity was pressed down by the motion of the cylinder, and vice versa. Thus the ascent or descent of one end of a lever produced a similar ascent or descent in the corresponding finger, by which one of the holes of the flute was occasionally opened or stopped, as by a living performer. Three of the levers served to regulate the ingress of the air, being contrived so as to open and shut, by means of valves, the three re¬ servoirs of air above mentioned, so that more or less strength might be given, and a higher or lower note pro¬ duced, as occasion required. The lips were, by a similar mechanism, directed by four levers, one of which opened them, to give the air a freer passage; the other contract¬ ed them; the third drew them backward; and the fourth pushed them forward. The lips were projected upon that part of the flute which receives the air; and, by the dif¬ ferent motions already mentioned, modified the tone in a proper manner. The remaining lever was employed in the direction of the tongue, which it easily moved so as to shut or open the mouth of the flute. Thus we see how all the motions necessary for a flute- player could be performed by this machine; but a con¬ siderable difficulty still remains, namely, how to regulate these motions properly, and make each of them follow in just succession. This, however, was effected by the fol¬ lowing simple method: the extremity of the axis of the cylinder was terminated on the right side by an endless screw, consisting of twelve threads, each placed at the distance of a line and a half from the other. Above this screw was fixed a piece of copper, and in it a steel pivot, which, falling in between the threads of the screw, ob¬ liged the cylinder to follow the threads, and instead of turning directly round, it was continually pushed to one side. Hence, if a lever was moved by a peg placed on the cylinder in any one revolution, it could not be moved by the same peg in the succeeding revolution, because the peg would be moved a line and a half beyond it by the lateral motion of the cylinder. Thus, by an artificial disposition of those pegs in different parts of the cylinder, the statue was made, by the successive elevation of the proper levers, to exhibit all the different motions of a flute-player, to the admiration of every one who saw it. The construction of machines capable of imitating even the mechanical actions of the human body shows exquisite skill; but what shall we say of one capable, not only of imitating actions of this kind, but of acting as external circumstances require, as though it were endued with life and reason ? This, nevertheless, has been done. M. de Kempelen, a gentleman of Presburg in Hungary, excited by the performances of M. de Vaucanson, at first endea¬ voured to imitate them, and at last far excelled them. This gentleman constructed an androides capable of play¬ ing at chess ! Every one who is in the least acquainted with this game must know that it is so far from being me¬ chanically performed, as to require a greater exertion of the judgment and rational faculties than is sufficient to accomplish many matters of greater importance. That this machine really was made, however, the public have had ocular demonstration. The inventor came over to Britain in 1783, where he remained above a year with his automaton. It is a figure as large as life, in a Turkish dress, sitting behind a table, with doors of three feet and a half in - length, two in depth, and two and a half in height. The chair on which it sits is fixed to the table, which runs i on four wheels. The automaton leans its right arm on ^ the table, and in its left hand holds a pipe: with this^1 hand it plays after the pipe is removed. A chess-board of 18 inches is fixed before it. This table, or rather cup. board, contains wheels, levers, cylinders, and other pieces of mechanism, all which are publicly displayed. The vest¬ ments of the automaton are then lifted over its head, and the body is seen full of similar wheels and levers. There is a little door in its thigh, which is likewise opened; and with this, and the table also open, and the automaton un¬ covered, the whole is wheeled about the room. The doors are then shut, and the automaton is ready to play; and it always takes the first move. At every motion the wheels are heard; the image moves its head, and looks over every part of the chess¬ board. When it checks the queen it shakes its head twice, and thrice in giving check to the king. It likewise shakes its head when a false move is made, replaces the piece, and makes its own move ; by which means the ad¬ versary loses one. M. de Kempelen remarks as the most surprising cir¬ cumstance attending his automaton, that it had been ex¬ hibited at Presburg, Vienna, Paris, and London, to thou¬ sands, many of whom were mathematicians and chess¬ players, and yet the secret by which he governed the mo¬ tion of its arm was never discovered. He prided himself solely in the construction of the mechanical powers, by which the arm could perform ten or twelve moves. It then required to be wound up like a watch, after which it was capable of continuing the same number of motions. The automaton could not play unless M. de Kempelen or his substitute was near it to direct its moves. A small square box, during the game, was frequently consulted by the exhibiter; and herein consisted the secret, which he said he could in a moment communicate. He who could beat M. de Kempelen was, of course, certain of con¬ quering the automaton. It was made in 1769. His own account of it was, “ (Test une bagatelle qui n’est pas sans merite du cote du mechanisme; mais les effets n’en pa* roissent si merveilleux que par la hardiesse de 1’idee, et par I’heureux choix des moyens employes pour faire il¬ lusion.” See Automaton. ANDROLEPSY, in Grecian Antiquity, an action al¬ lowed by the Athenians against such as protected persons guilty of murder. The relations of the deceased were empowered to seize three men in the city or house whither the malefactor had fled, till he should be either surren¬ dered, or satisfaction made some way or other for the murder. ANDROMACHE, the wife of Hector, the mother of Astyanax, and daughter of Eetion king of Thebes in Ci¬ licia. After the death of Hector and the destruction of Troy she married Pyrrhus, and afterwards Helenus the son of Priam, with whom she reigned over part of Epirus. ANDROMEDA, in Astronomy, a northern constella¬ tion, behind Pegasus, Cassiopeia, and Perseus. It re¬ presents the figure of a woman chained, and is fabled to have been formed in memory of Andromeda, daughter of Cepheus and Cassiopeia, and wife of Perseus, by whom she had been delivered from a sea monster, to which she had been exposed to be devoured for her mother’s pride. Minerva translated her into the heavens. ANDRON, in Grecian Antiquity, denotes the apartment in houses designed for the use of men; in which sense it stands opposed to Gynceceum.—The Greeks also gave their dining-rooms the title of andron, because the wo¬ men had no admittance to feasts with the men. AND AND 125 flilr a ANDRONA, in ancient writers, denotes a street or T|| public place where people met and conversed together, udrc- jn some writers androna is more expressly used for the ' ^ J space between two houses; in which sense the Greeks *v"" alSo use the term for the way or passage between two apartments. Androna is also used, in ecclesiastical writers, for that part in churches destined for the men. Anciently it was the custom for the men and women to have separate apart¬ ments in places of worship, where they performed their devotions asunder; which method is still religiously ob¬ served in the Greek church. The avdguv, or androna, was in the southern side of the church, and the women’s apart¬ ment on the northern. ANDRONICUS I. Comnenus, emperor of the East, was the son of Isaac, and grandson of Alexius Comnenus. Naturally active, martial, and eloquent, he shines as one of the most conspicuous characters of his age. Following the bent of his inclination, he attended the Roman army in their retreat; but in their march through Asia Minor, wan¬ dering into the mountains, he fell into the hands of some Turkish huntsmen, was carried to the sultan, and remained his prisoner; but regaining his liberty, both his virtues and vices soon recommended him to the favour of his cousin Manuel, the reigning emperor. The vicious heart of Andronicus manifested itself clearly in maintaining a licentious correspondence with Eudocia, the emperor’s niece, while the emperor himself lived in public incest with her sister Theodora. His martial spirit gained him a considerable command in Cilicia, where he laid siege to Mopsuestia; but by a successful sally of the enemy he was obliged to raise the siege, and retire in considerable disorder. Inflamed with a desire of revenging the infamy of their sister in his blood, the brothers of Eudocia made an unsuccessful attempt to assassinate Andronicus at mid¬ night in his tent; but being awakened, he defended him¬ self with surprising bravery, forced his way through his enemies, and escaped in safety. Afterwards engaging in a treasonable correspondence with the emperor of Ger¬ many and the king of Hungary, he was arrested and thrown into confinement. He remained in this state about twelve years, and after several repeated attempts to escape, he at last effected his purpose, and fled for re¬ fuge to the court of the great duke of Russia. The cun¬ ning of Andronicus soon found means to regain his favour with the emperor Manuel; for having exerted all his in¬ fluence, he succeeded in obtaining the Russian prince to engage to join his troops with those of Manuel, in the invasion of Hungary. Accordingly, on account of his important service, he obtained a free pardon from the em¬ peror, and after an expedition to the Danube returned with him to Constantinople. He again fell under the dis¬ pleasure of the emperor, by refusing to take an oath of allegiance to the prince of Hungary, his intended son-in- law, and consequently presumptive heir to the crown, and was thereupon returned to his former command in Cilicia. W Idle residing here, his powerful address captivated the heart of Philippa, daughter of the Latin prince of Antioch, and sister to the empress Maria; and in her company he spent his time in all the amusements that country could afford, till the emperor’s resentment put a stop to their correspondence. Thus circumstanced, he collected a hand of adventurers, and undertook a pilgrimage to the Holy Land, where, by his insinuating turn of mind, he so far succeeded in gaining the favour of the king and clergy, as to be invested with the lordship of Berytus, on the coast of Phoenicia. In this neighbourhood Theodora, the beau¬ tiful widow of Baldwin, king of Jerusalem, and nearly allied in blood to him, resided. The personal accomplish¬ ments and address of Andronicus captivated her heart, and she became the third victim to his artful seduction, and lived publicly as his concubine. Still pursued by the emperor with unabating resentment, he was forced to take refuge in Damascus, and then in several other places in the east, till at length he settled in Asia Minor. While residing here he made frequent incursions into the pro¬ vince of Trebizond, and seldom returned without success. After several occurrences Theodora was made captive by the governor of Trebizond, along with her two children, and sent to Constantinople; upon which Andronicus im¬ plored and obtained pardon. He acted the affected peni¬ tent in such a manner, that he again ingratiated himself into the favour both of the church and state; but was sent to dwell at G£noe, a town situated on the Euxine coast. In the year 1177 Manuel died, and was succeeded by his son Alexius II., a youth about twelve or fourteen years of age, without wisdom or experience, by which the ambition of Andronicus was again called into action. A civil war having been occasioned by the misconduct of the empress in Constantinople, the public mind was di¬ rected towards Andronicus, as the only person whose rank and accomplishments could restore the public tranquillity. Incited by the patriarchs and patricians, he marched to¬ wards Constantinople, which he entered, took possession of the palace, confined the empress, consigned her minis¬ ter to death, assumed the office of protector, put to death many persons of distinction, tried and executed the queen on a charge of corresponding with the king of Hungary, and vowed fidelity to the young emperor upon his corona¬ tion, at the same time teaching the necessity of an expe¬ rienced ruler, to assuage the evils that threatened the em¬ pire ; upon which his adherents called out, “ Long live Alexius and Andronicus, Roman emperors.” While he af¬ fected reluctance, he was elevated to a partnership in the empire. This conjunction of the royal power was soon dissolved by the murder of the unfortunate Alexius. The body of the deceased being brought into his presence, striking it with his foot, he said, “ Thy father was a knave, thy mother a whore, and thyself a fool.” Having arrived at the dignity of sole emperor, a. d. 1183, he con¬ tinued to sway the sceptre with a mixture of justice and bounty towards his subjects at large; but those whom he feared or hated he governed with the most cruel tyranny. The noble families that were either cut off or exiled by him were all allied to the Comneni. Some of these were engaged in revolt; and the public calamity was height¬ ened by an invasion of the Sicilians, in which they took and sacked Thessalonica. A rival without merit, and a people without arms, at last overturned his throne. A descendant from the first Alexius in the female line, named Isaac Angelus, being singled out by Andronicus as a victim to his cruelty, he with courage and resolution defended his life and liberty, slew his executioner, fled to the church of St Sophia, and there took refuge with se¬ veral of his friends. Isaac was instantly raised by the po¬ pulace from a sanctuary to a throne. When this event took place Andronicus was absent from Constantinople; but he no sooner heard of it than he with the utmost speed returned. Upon his arrival there he found himself deserted by all, and was seized and dragged in chains be¬ fore the new emperor. All the eloquence he displayed was of no avail; for Isaac delivered him into the hands of those whom he had injured, and for the space of three days he endured with uncommon patience all the insults and torments that were inflicted upon him. At last two friendly or furious Italians, plunging their swords into his body, put a period to his life. His death, in the 73d year of his age, terminated the dynasty of the Comneni. Andhonicus of Cyrrhus, an Athenian astronomer, built 126 AND Andro- at Athens an octagon tower, with figures carved on each phagi side, representing the eight principal winds. A brazen li Triton at the summit, with a rod in its hand, turned round Andry- t]ie w[n^ pointed to the quarter from whence it blew. this model is derived the custom of placing wea¬ thercocks on steeples. ANDROPHAGI, in Ancient Geography, the name of a nation whose country, according to Herodotus, was ad¬ jacent to Scythia. Their name, compounded of two Greek words, signifies man-eaters. See Anthropophagi. ANDROS, one of the ancient Cyclades, lying between Tenos and Euboea, being one mile distant from the former, and ten from the latter. The ancients gave it various names, viz. Cauros, Lasia, Nonagria, Epagris, Antandros, and Hydrusia. The name of Andros it received from one Andreus, appointed, according to Diodorus Siculus, by Rhadamanthus, one of the generals, to govern the Cy¬ clades after they had of their own accord submitted to him. It had formerly a city of great note, bearing the same name, and situated very advantageously on the brow of a hill which commanded the whole coast. In this city, according to Strabo and Pliny, stood a famous temple dedicated to Bacchus. Near this temple Mutianus, as quoted by Pliny, tells us there was a spring called the gift of Jupiter, the water of which had the taste of wine in the month of January, during the feast of Bacchus, which lasted seven days. The Andrians were the first of all the islanders who joined the Persians when Xerxes invaded Greece; and therefore Themistocles, after the victory at Salamis, resolved to attack the city of An¬ dros, and oblige the inhabitants to pay large contributions for the maintenance of his fleet. Having landed his men on the island, he sent heralds to the magistrates, acquaint¬ ing them that the Athenians were coming against them with two powerful divinities, Persuasion and Force, and therefore they must part with their money by fair means or foul. The Andrians replied that they likewise had two mighty deities, who were very fond of their island, viz. Poverty and Impossibility, and therefore could give no money. Themistocles, not satisfied with this answer, laid siege to the town, which he probably made himself mas¬ ter of and destroyed, as we are informed by Plutarch that Pericles, a few years after, sent thither a colony of 250 Athenians. It was, however, soon retaken by the Per¬ sians, and, on the overthrow of that empire by Alexander the Great, submitted to him, along with the other islands. On his death it sided with Antigonus, who was driven out by Ptolemy. The successors of the last-mentioned prince held it till the time of the Romans, when Attalus, king of Pergamus, besieged the metropolis at the head of a Roman army, and, having taken it, was by them put in possession of the whole island. Upon the death of Atta¬ ins the republic claimed this island, as well as his other dominions, in virtue of his last will. Andros is now subject to the Turks, and contains a town of the same name, with a great many villages. It is the most fruitful island in the Archipelago, and yields a great quantity of silk. Andros, when visited by Tour- nefort, contained about 4000 inhabitants. There were seven monasteries, a great number of churches, and a ca¬ thedral for the bishops of the Roman Catholic persuasion; but most of the inhabitants were of the Greek communion. It forms a part of the new Greek republic. It is about 23 miles long and 6 broad. Long. 24.50. E. Lat. 37.50. N. ANDRUM, a kind of hydrocele, to which the people of Malabar are very subject. It is said to be derived from the bad quality of the country waters, impregnated with certain salts. ANDRYCHOW, a city in the circle of Myslenicze, in the Austrian province of Galizia, on the river Wieprzow- A N G ka, with a castle, and 2805 inhabitants, amongst whom Ar. are several manufacturers of fine damask and other table- linen, who produce annually about 30,000 pieces. ANDUJAN, a city of Spain, in the province of Jaen, | in Andalusia. It is situated on the south side of the Sierra Morena, which defends it from the cold winds of the north, near the sources of the Jandula. There is a very fine bridge over the Guadalquivir at this place. The surrounding country is well watered, and yields abundant harvests of wheat, barley, oil, and wine; and numerous hives of bees furnish abundant supplies of honey and wax. From a whitish clay found here there are manufactured a vast quantity of jars called alcarrasas, which are highly esteemed for their property of keeping water cool in the hottest summer weather. It contains a castle, six churches, nine monasteries, a theatre, and 14,000 inhabitants. Long. 3. 28. 25. W. Lat. 38. 1. 32. N. ANDUZE, a town of France, in the department of the Gard, seated on the river Gardon. It carries on a consi¬ derable trade in serges and woollen cloth. Long. 3. 42. E. Lat. 43. 39. N. ANECDOTES (Anecdota), a term used by some authors for the titles of Secret Histories ; but it more properly de¬ notes a relation of detached and interesting particulars. The word is Greek, avwdora,, signifying things not yet known or hitherto kept secret. Procopius gives this title to a book which he published against Justinian and his wife Theodora. Anecdotes is also an appellation given to such works of the ancients as have not yet been published; in which sense Muratori gives the name Anecdota Grceca to several writings of the Greek fathers, found in the li¬ braries, and first published by him. Martene and Du¬ rand have given a Thesaurus novus Anecdotorum, in 5 vols. folio. ANEMOMETER, or Anemoscope, machines for mea¬ suring the force, and indicating the course, of the wind. See Wind-gage. ANEMUR, the most southern point of Asia Minor, on the south coast of Caramania. The castle of Anemur stands six miles east of the cape, on the edge of the sea, and extends about 800 feet by 300. Its citadel is placed on a small rocky eminence, and is in a ruinous state. Long. 32. 30. E. Lat. 36. 15. N. ANGAR, Angan, or Hindsham, a barren and unin¬ habited island on the Arabian shore of the Persian Gulf, on the south side of the island of Kishma, about 12 miles in circuit. It must have been formerly inhabited, as it contains the ruins of a considerable town, and many reser¬ voirs of water. It has also two wells, and a stream of good water, which unfortunately become dry in the hot wreather. It is covered with pits of salt and metallic ores, and a soft rocky substance resembling lava. The hills, which are overspread with shells of oysters and other fish, abound in wild goats, rabbits, and partridges. ANGARA, a river of Siberia, which has its rise in the Lake Baikal. It passes the town of Irkutsk with a rapid course, and receiving the Oka, it changes its name to Toungooska, and falls into the Yeneseior Jenesei after a course of 700 miles. It is navigable, and is noted for the clearness of its waters. ANGARI, or Angarii, in Antiquity, public couriers, appointed for the carrying of messages. The ancient Persians, Budaeus observes, had their ayyugeiov tyoyriya, which was a set of couriers on horseback, posted at cer¬ tain stages or distances, always in readiness to receive the dispatches from one, and forward them to another, with wonderful celerity, answering to what the moderns call posts (positi), as being posted at certain places or stages. The angari wrere also called by the Persians astandce; by the Greeks on account of the long journeys A N G A N G 127 they made in one day, which, according to Suidas, amount¬ ed to not less than 1500 stadia. ANGARIA, in Homan Antiquity, a kind of public service imposed on the provincials, which consisted in pro¬ viding horses and carriages for the conveyance of military stores and other public burdens. It is sometimes also used for a guard of soldiers, posted for the defence of a place. In a more general sense it is used for any kind of oppression, or services performed through compulsion. 1 ANGAZYA, one of the Comora Islands, lying between the north end of Madagascar and the coast of Zanguebar in Africa, from lat. 10. to 15. S. It is inhabited by Moors, who trade with divers parts of the continent, in cattle, fruits, and other commodities of the island, which they exchange for calicoes and other cotton cloths. ANGEIOTOMY, in Surgery, implies the opening of a vein or artery, as in bleeding; and consequently includes both arteriotomy and phlebotomy. ANGEL, a spiritual intelligent substance, the first in rank and dignity among created beings. The word angel is Greek, and signifies a messenger: the Hebrew “ixbn signifies the same thing. The angels are in Daniel, chap, iv. ver. 13, &c. called ow-, or watchers, from their vigi¬ lance : for the same reason they are, in the remains we have of the prophecy attributed to Enoch, named/fyrcyon; which word imports the same in Greek. The term Angel, therefore, in the proper signification of the word, does not import the nature of any being, but only an office; in which sense angels are called the minis¬ ters of God, and ministering spirits. That there are such beings, invisible and imperceptible to our senses, endued with understanding and power superior to those of human nature, created by God, and subject to him,—ministering to his divine providence in the government of the world,— are truths fully attested by Scripture. Nay, the existence of such invisible beings was generally acknowledged by the heathens, though under different appellations : the Greeks called them demons, and the Romans genii or lares. Epicurus seems to have been the only one among the ancient philosophers who absolutely rejected them. As to the nature of these beings, we are told that they are spirits; but whether pure spirits divested of all matter*, or united to some corporeal vehicles, has been a contro¬ versy of long standing. Not only the ancient philosophers, but some of the Christian fathers, were of opinion that angels were clothed with ethereal or fiery bodies, of the same nature with those which we shall one day have when we come to be equal to them. But the more general opi¬ nion, especially of later times, has been, that they are sub¬ stances entirely spiritual, though they can at any time as¬ sume bodies, and appear in human or other shapes. Besides their attendance on God, and their waiting and executing his commands, they are also presumed to be employed in taking care of mankind and their concerns: and that every man had such a tutelar or guardian angel, even from his birth, was a firm belief and tradition among the Jews; and our Saviour himself seems to have been of the same sentiment. The heathens were also of that per¬ suasion, and thought it a crime to neglect the admonitions of so divine a guide. The Romans thought the tutelar genii of those who attained the empire to be of an eminent order, on which account they had great honours shown them. Nations and cities also had their several genii. Ihe ancient Persians so firmly believed the ministry of angels, and their superintendence over human affairs, that they gave their names to their months, and the days of their months, and assigned them distinct offices and pro¬ vinces : and it is from them the Jews confess to have re¬ ceived the names of the months and angels, which they brought with them when they returned from the Babylo¬ nish captivity; after which, we find they also assigned Angelics charges to the angels, and in particular the patronage of II empires and nations; Michael being the prince of the Jews, ^nger* as Raphael is supposed to have been of the Persians. Although the angels were originally created perfect, Of the fal- good, and obedient to their Master’s will, yet some of themlen angels, sinned and kept not their first estate, but left their habi¬ tation, and so, from the most blessed and glorious, became the most vile and miserable, of all God’s creatures. They were expelled the regions of light, and cast down to hell, to be reserved in everlasting chains under darkness, until the day of judgment. With heaven they lost their heaven¬ ly disposition, which delighted once in doing good and praising God ; and fell into a settled rancour against the Deity, and malice against men. Their inward peace was gone ; all desire of doing good departed from them, and instead thereof, revengeful thoughts and despair took pos¬ session of them, and created an eternal hell within them. ANGELICS, Angelici, in Ecclesiastical History, an ancient sect of heretics, supposed by some to have got this appellation from their excessive veneration of angels; and by others, from their maintaining that the world was cre¬ ated by angels. Angelics is the name of an order of knights, instituted in 1191, by Angelus Flavius Comnenus, emperor of Con¬ stantinople : also of a congregation of nuns, founded at Milan in 1534, by Louisa Torelli, countess of Guastalla. They observe the rule of St Augustin. ANGELITES, in Ecclesiastical History, a sect of Christian heretics in the reign of the emperor Anastasius, and the pontificate of Symmachus, about the year 494 ; so called from Angelium, a place in the city of Alexandria where they held their first meetings. They were called likewise Severites, from one Severus, who was the head of their sect; as also Theodosians, from one among them named Theodosius, whom they made pope at Alexandria. They held that the persons of the Trinity are not the same ; that none of them exists of himself, and of his own nature, but that there is a common god or deity existing in them all, and that each is God, by a participation of this deity. ANGELO, Michael. See Buonaroti, Michael Angelo. Angelo, St, a small but strong town of Italy, in the Ca- pitanata. There are several other towns and castles of the same name in Italy, and particularly the castle of St Angelo at Rome. Long. 15. 56. E. Lat. 41. 43. N. ANGELOT, an ancient English gold coin, struck at Paris while under subjection to the English. It was thus called from the figure of an angel supporting the scutcheon of the arms of England and France. There was another coin of the same denomination struck under Philip de Valois. Angelot is also used in Commerce to denote a small, fat, rich sort of cheese brought from Normandy. Skinner supposes it to have been thus called from the name of the person who first made it up in that form, and perhaps stamped it with his own name. Menage supposes it to have been denominated from the resemblance it bears to the English coin called angelot. It is made chiefly in the Pays de Bray, whence it is also denominated angelot de Bray. It is commonly made in vats, either square or shaped like a heart. ANGER, a painful feeling of the mind, excited by the receipt of an injury or affront, and accompanied with a disposition to retaliate on the author of the injury. Bishop Butler observes that anger is far from being a selfish passion, since it is produced by injuries offered to others as well as to ourselves, and was designed by the Author of nature not only to excite us to act vigorously in defend¬ ing ourselves from evil, but to interest us in the defence or 128 A N G Angerburgrescue of the injured and helpless, and to raise us above H the fear of the proud and mighty oppressor. Angers. The same author makes an important distinction, as Dr Reid observes (Active Powers, Essay 3), “ between sudden anger or resentment, which is a blind impulse arising from our constitution, and that which is delibe¬ rate. The first may be raised by hurt of any kind; but the last can only be raised by injury, real or conceived. Both these kinds of anger or resentment are raised whether the hurt or injury be done to ourselves or to those we are interested in.” Physicians and naturalists have recorded instances of extraordinary cases produced by anger. Borrichius cured a woman of an inveterate tertian ague, which had baffled the art of physic, by putting the patient in a furious fit of anger. Valeriola made use of the same means, with the like success, in a quartan ague. The same passion has been equally salutary to paralytic, gouty, and even dumb persons ; to which last it has sometimes given the use of speech. Etmuller gives divers instances of very singular cures wrought by anger: among others, he mentions a person laid up by the gout, who being provoked by his physician, flew upon him, and was cured. It has often, on the other hand, been productive of fatal effects. We meet with several instances of princes to whom it has proved mortal, e. g. Valentinian the First, Wenceslaus, Matthias Corvinus, king of Hungary, and others. There are also instances where it produced epilepsy, jaundice, cholera morbus, diarrhoea, &c. ANGERBURG, a circle in the government of Gum- binnen, or province of East Prussia, formerly a part of Poland. It extends over 374 square miles, or 239,360 acres. It is watered by the Angerap, which rises in seve¬ ral lakes, with which the district abounds. The inhabit¬ ants are 21,172, who produce some corn and flax, and much fresh-water fish and fire-wood. The females are all employed in spinning linen yarn. The capital is of the same name, and contains 250 houses and 2619 inhabitants. ANGERMANLANDS-LAPMARK, or ^Esle-Lap- mark, a province in the northern part of Sweden, of the vast extent of 6560 square miles. The inhabitants are only 1200, partly Laplanders, with a few Swedes denomi¬ nated colonists. The chief productions to spare are butter and some iron wares. Aisle is the chief place, and be¬ sides it there are two parishes with churches. ANGERMUNDE, a circle in the government of Pots¬ dam, and province of Brandenburg, in Prussia. It extends over 503 square miles, or 321,920 acres ; and comprehends six cities, three market-towns, 109 villages, and 4201 dwellings, with 34,896 inhabitants. The river Oder washes its eastern boundary, and receives the several smaller streams by which it is watered. The borders of the rivers present some excellent meadow-land, on which many cat¬ tle are pastured. It produces good corn, tobacco, flax, and abundance of garden fruit. There is much wood land, and several lakes which yield fish in great plenty. The capital, of the same name, contains three churches, 291 houses, and 2654 inhabitants, whose chief occupation is making snuff and tobacco. ANGERONA, in Mythology, the name of a pagan deity whom the Romans prayed to for the cure of the quinsy, in Latin angina. Pliny calls her the goddess of silence and calmness of mind, who banishes all uneasiness and melancholy. She is represented with her mouth covered, to denote patience and refraining from complaints. Her statue was set up and sacrificed to in the temple of the goddess Volupia, to show that a patient enduring of affliction leads to pleasure. ANGERS, an arrondissement in the department of the Mayenne and Loire, in France, extending over 436 square A N G miles, or 279,040 acres. It is divided into seven cantons, and those again into 59 communes, containing 92,810 I' inhabitants. The chief city, of the same name, contains i| 33,100 inhabitants. M ANGHIERA, a town of Italy, in the duchy of Milan, and capital of a county of the same name. It is seated on the eastern side of the lake Maggiore, in lat. 45. 42. N. long. 8. 40. E. ANGLE. This term is, owing to the poverty of lan¬ guage, employed to signify very different things. In Plane Geometry, it means the opening or separation of two straight lines which meet in a point; but in Solid Geometry, it variously denotes the deviation of a straight line from a plane, the divergence of one plane from ano¬ ther at their line of junction, or even a cluster of plane angles terminating in a common summit. This diversified application of the same word is not likely, however, among mathematicians, to occasion any misconception. But it would be more perspicuous, and certainly more philo¬ sophical, to imitate the practice of naturalists in framing a set of cognate words to express the several transitions of meaning. The word angle was drawn from common discourse into the vocabulary of science. Its primitive sense, in all the languages in which it can be traced, is merely a nook or cor¬ ner ; but it has acquired a more precise and extensive appli¬ cation in its transfer to geometry. In its simplest form, it now denotes generally the divergence or difference of direc¬ tion between two concurring straight lines. Yet a learner still experiences some difficulty in seizing the correct idea of its nature, which has always baffled the attempts of authors to reduce to the terms of a strict definition. Apollonius, at once the most elegant and inventive of the Greek geometers, was satisfied with representing an angle as a collection of space about a point,—a description which is not only extremely loose, but which intimates quite a different conception. Euclid, the great compiler of the Elements, has defined an angle to be the xX/erate, but has a gnawing stomach that will not endure i I nuch fasting, but must observe hours, it troubleth the i nind and body, and loseth that delight which only mak- il :th pastime pleasing.” “ He must be of a well-settled 8 md constant belief, to enjoy the benefit of his expecta- (■ ion; for then to despair, it were better never to be put n practice: and he must ever think when the waters are Peasant, and any thing likely, that there the Creator of 1 til good things hath stored up much of plenty; and )■ hough your satisfaction be not as ready as your wishes, ( 1 fet you must hope still, that with perseverance you shall ; eap the fulness of your harvest with contentment. Then I r 16 must be full of love both to his pleasure and his neigh- xmr—to his pleasure, which otherwise will be irksome ' md tedious—and to his neighbour, that he never give of- ( ence in any particular, nor be guilty of any general de- t truction : then he must be exceeding patient, and neither f 'ex nor excruciate himself with losses or mischances, as 5 n losing the prey when it is almost in the hand, or by II waking his tools by ignorance or negligence; but with ^leased sufferance amend errors, and think mischances nstructions to better carefulness.” In regard to the antiquity of angling, it has been traced ) >y some to the time of Seth, who is asserted to have \ fght it to his sons ; and so highly have others esteemed li 16 *no'vledge of the art, as to maintain that its rules and 3 naxims were engraven on those pillars by which an ac¬ quaintance with music, the mathematics, and other branch¬ es of useful knowledge, was preserved by God’s appoint¬ ment from extinction in the days of Noah. It is fre¬ quently alluded to in the holy Scriptures; as in Isaiah, xix. 8, “ The fishers also shall mourn, and all they that cast angle into the brooks shall lament, and they that spread nets upon the waters shall languish;” so in the prophet Habbakuk, i. 15, “ They take up all of them with the angle, they catch them in their net, and gather them in their drag; therefore they rejoice and are glad.” We deem it unnecessary to multiply quotations from an¬ cient authors, whether sacred or profane; but shall rest satisfied with pointing out, at the close of this article, the principal works on angling which have appeared in our own language, and in relation to the practice of the art in British streams. As expert angling never was and never will be success¬ fully taught by rule, but is almost entirely the result of assiduous and long-continued practice, we purpose being very brief in our disquisition on the subject. We shall commence by stating our belief that fly-fishing, by far the most elegant and interesting branch of the art, ought not to be regarded exclusively as an art of imitation. It no doubt depends on deception, which usually proceeds on the principle of one thing being successfully substituted in the likeness of another ; but Bacon’s distinctive defini¬ tions of simulation and dissimulation place the subject in a truer light. As simulation consists in the adoption or affectation of what is not, while dissimulation consists in the careful concealment of what really is—the one being a positive, the other rather a negative act—so the great ob¬ ject of the fly-fisher is to dissimulate in such a manner as to prevent his expected prey from detecting the artificial nature of his lure, without troubling himself by a vain effort to simulate or assume, with his fly, the appearance of any individual or specific form of insect life. There is, in truth, little or no connection between the art of angling and the science of entomology ; and therefore the success of the angler, in by far the greater proportion of cases, does not depend on the resemblance which subsists be¬ tween his artificial fly and the natural insect. This state¬ ment is no doubt greatly at variance, as well with the principles as the practice of all who have deemed fishing worthy of consideration, from the days of Isaiah and Theo¬ critus, to those of Carrol and Bainbridge. But we are not the less decidedly of opinion, that in nine instances out of ten a fish seizes upon an artificial fly as upon an insect or moving creature sui generis, and not on account of its exact and successful resemblance to any accustomed and familiar object. It may be asked, upon what principle of imitative art the different varieties of salmon-fly can be supposed to bear the most distant resemblance to any species of dragon- ffy, to imitate which we are frequently told that they are intended ? Certainly no perceptible similarity in form or aspect exists between them, all the species of dragonfly, with the exception of one or two of the sub-genus Calep- terix, being characterized by very clear, lace-like, pellucid wings, entirely unadorned by those fantastic and gaudy colours, borrowed from the peacock and other “ birds of gayest plume,” which are made to distinguish the suppos¬ ed resemblance. Besides, the finest salmon-fishing is in mild weather during the colder seasons of the year, and in early spring, several months before any dragonfly has become visible on the face of the waters, as it is a sum¬ mer insect, and rarely makes its appearance in the perfect state till the month of June. If they bear no resemblance to each other in form or colour, how much more unlike must they be, when, instead of being swept down the cur¬ rent, as a real one would be, the artificial fly is seen cross- Angling. 134 ANGLING. Angling, ing and recrossing every stream and torrent, with the agility of an otter, and the strength of an alligator ? Now, as it is demonstrable that the artificial fly generally used for salmon bears no resemblance, except in size, to any living one ; that the only tribe which, from their respec¬ tive dimensions, it may be supposed to represent, does not exist in the winged state during the period when the imitation is most generally and most successfully practis¬ ed ; and if they did, that their habits and natural powers totally disenable them from being at any time seen under such circumstances as would give a colour to the supposi¬ tion of the one being ever mistaken for the other ; may we not fairly conclude that, in this instance at least, the fish proceed upon other grounds, and are deceived by an appearance of life and motion, rather than by a specific resemblance to any thing which they had previously been in the habit of capturing? What natural insect do the large flies, at which sea-trout rise so readily, resemble ? These, as well as gilse and salmon, frequently take the lure far within the bounds of the salt-water mark; and yet naturalists know that no such thing as a salt-water fly exists, or at least has ever been discovered by their re¬ searches. Indeed no true insect inhabits the sea. What species are imitated by the palmer, or by three fourths of the dressed flies in common use ? An artificial fly can, at the best, be considered only as the representation of a natu¬ ral one which has been drowned, as it is impossible to imitate the dancing or hovering flight of the real insect over the surface of the stream; and, even with that re¬ stricted idea of its resemblance to nature, the likeness must be scarcely perceptible, owing to the difference of motion, and the great variety of directions in which the angler drags his flies, according to the nature and special localities of the current, and the prevailing direction of the wind. The same observations apply, with almost equally few exceptions, to bait-fishing. The minnow is fastened upon swivels, which cause it to revolve upon its axis with such rapidity, that it loses every vestige of its original appear¬ ance ; and in angling with the par tail, one of the most killing lures for large trout, the bait consists of the nether half of a small fish, mangled and mis-shapen, and in every point of view divested of its natural form. Fly-fishing has been compared, though by a somewhat circuitous mode of reasoning, to sculpture. It proceeds upon a few simple principles, and the theory is easily ac¬ quired, although it may require long and severe labour to become a great master in the art. Yet it is needless to encompass it with difficulties which have no existence in reality, or to render a subject intricate and confused, which is in itself so plain and unencumbered. In truth, the ideas which at present prevail on the matter degrade it beneath its real dignity and importance. When Plato, speaking of painting, says that it is merely an art of imi¬ tation, and that our pleasure arises from the truth and accuracy of the likeness, he is surely wrong; for if it were so, where would be the superiority of the Homan and Bolognese over the Dutch and Flemish schools ? So also in regard to fishing: The accomplished angler does not condescend to imitate specifically, and in a servile manner, the detail of things; he attends, or ought to at¬ tend, only to the great and invariable ideas which are in¬ herent in universal nature. He throws his fly lightly and with elegance on the surface of the glittering wa¬ ters, because he knows that an insect with outspread gawzy wings would so fall; but he does not imitate (or if he does so, his practice proceeds upon an erroneous prin¬ ciple), either in the air or on his favourite element, the flight or the motion of a particular species, because he also knows that trouts are much less conversant in ento¬ mology than M. Latreille, and that their omnivorous pro- u pensities induce them, when inclined for food, to rise with Jr equal eagerness at every minute thing which creepeth ^ upon the earth or swimmeth in the waters. On this fact he generalizes,—and this is the philosophy of fishing. We are therefore of opinion that all, or a great pro. portion, of what has been so often and sometimes so well said about the great variety of flies necessary to an angler, —about the necessity of changing his tackle according to each particular month throughout the season,—about one fly being adapted solely to the morning, another to noon¬ day, and a third to the evening,—and about every river having its own particular flies, &c. is, if not erroneous, at least exaggerated and misconceived. That determinate relations exist between flies of a certain colour and parti¬ cular conditions of a river, is, we doubt not, true; but these are rather connected with angling as an artificial science, and have but little to do with any analogous re¬ lations in nature. The great object, by whatever means it is to be accomplished, is to render the fly deceptive; and this, in fact, we believe to be more frequently effect¬ ed when fishing with flies which differ in colour and ge¬ neral appearance from those which are upon the water. When a particular fly prevails upon a river, an artificial one in imitation of it will never resemble it so closely as to appear the same to those below (i. e. the fish): on the contrary, a certain degree of resemblance, without any thing like an exact similitude, will only render the finny tribe the more cautious through suspicion; while a differ¬ ent shape and colour, by exciting no minute or invidious comparisons, would probably have been swallowed with¬ out examination. Indeed, it seems sufficiently plain, that where means of comparison are allowed, and where exact imitation is at the same time impossible, it is much better to have recourse to a general idea, than to an awkward and bungling individual representation. How often has it been asserted, with all the gravity of sententious wis¬ dom, that the true mode of proceeding in fly-fishing is to busk your hook by the river-side, after beating the shrubs to see what colour of insect prevails. A very expert angler, who perhaps carried the opposite theory rather too far (although he always filled his pannier), wras in the habit of stirring the briars and willows to ascertain what manner of fly was not there, and with that he tempt¬ ed the fishes. It is admitted, that during mid-summer, when the wea¬ ther is calm, the sky cleai’, and the river low, and when what is called fine fishing is necessary, a close imitation both of the appearance and motions of the natural fly may frequently be tried with advantage; in which case the tackle may be allowed to drop gently down the stream: but it more usually happens, from the style of fishing practised during the vernal and autumnal states of a river, that the hook is not deceptive from its appearing like a winged fly which has fallen from its native element, but from its motion and aspect resembling that of some aquatic insect. When the end of the line first falls on the surface of the water, the fish may be deceived by the idea of a natural fly; and it is on that account that the angler should throw his tackle lightly and with accuracy, and it is on that account also that we would advise the more frequent throwing of the line: but so soon as the practitioner begins to describe his semicircle across the river, the character of the lure is changed, and the trout then seizes the bait, not as a drowning insect, but as a creature inhabiting its own element, which had ventured too far from the protection of the shallow shore or the sedgy bank. That this is the case, a subsidiary argument may also be drawn from the fact, that in most rivers the greater number and the finest fish are generally killed by ANGLING. 135 r he drag-fly, which, during the process of angling, swims LL inch or two under water. Nevertheless, as these opinions of ours may not accord dth the practice of other anglers, we shall proceed some¬ what more methodically to explain a few of the principles f the art as usually received and followed. The great secret in fly-fishing, after a person has ac- |uired the art of throwing a long and a light line, is perse- erance,—that is, constant and continuous exertion. Fish re whimsical creatures, even when the angler, with all ppliances and means to boot, is placed apparently under he most favourable circumstances. Let him, however, ommence his operations with flies which, upon general irinciples, he knows to be good,—for example, a water- nouse body and dark wing, hare-ear and moorfowl wing, ed hackle and teal or mallard wing. It may frequently mppen that for an hour, or even two hours, he will kill tothing; but then it will as often happen that for another ouple of hours he will pull them ashore with a most leasing celerity. Awake but one, and, lo, what myriads rise! Next comes a pause of another hour or more, during which little or nothing is obtained, so that if the inter- nediate period is frittered away, success is doubtful or mpossible. We believe that the appetites and motions if the finny tribes are regulated and directed by certain to us) almost imperceptible changes in the state of the tmosphere, with which, as they do not proceed from any leterminate or ascertained principles of meteorological cience, it is not easy for the angler to become acquaint- id; and therefore the only method to remedy the des- igrdment thus arising, is to fish without ceasing as long :is he remains by the “ pure element of waters.” The irt of angling, if worthily followed, and with an observant ye, will probably one day or other be the means of browing considerable light on the science of electricity, it present one of the most obscure, though at the same line the most important and influential, of all the subjects pf physical learning. The best natural flies, either to use fresh, or to serve is models for the artificial kinds, are, first, the different iorts of stoneflies (Phryganea and Limnephilus), which ire usually found by the water-side. Their common co- ours are various shades of brown ; they have pretty long feelers or antennae, which, in a state of repose, are bent >ver their shoulders and along their sides; their wings ire held decumbent, or close to the sides. They fly leavily, and are produced from aquatic larvae called caddis 3r case-worms, remarkable for their curious dwelling- places, which are hollow tubes composed of sand, small shells, and pieces of wood, agglutinated together, and made heavier or lighter, according to circumstances, that they may the more easily sink or swim. They are open at either end, and the worm crawls along the stones and gravel, by protruding its legs at the anterior extremity. They disencumber themselves from their aquatic habita¬ tions, and assume the winged state in spring and the ear¬ lier part of summer. Secondly, The different kinds of May Hies {Ephemerae), called green drakes, &c. are also pro¬ duced from larvae, which, for a long time previous to their appearance as perfect insects, have inhabited the waters. There are many species of this genus, all of which are Angling, greedily sought for by trout. They are easily known by' their tapering abdomens, veined wings, short antennae, and the long slender setae or hairs which terminate their bodies. They chiefly abound from May to mid-sum¬ mer. Thirdly, The small black or ant-fly, is the winged female of the common black ant, and occurs in the nests or hills of that insect during the summer and autumnal months. There is scarcely any season of the year, excepting the winter months, in which an experienced angler may not suc¬ cessfully ply his trade.1 In the mid-summer season, when the pools are very clear and shallow, and the streams al¬ most dried up, little can be done without a stirring breeze ; so also after a heavy summer flood, immediately ensuing a continuance of dry weather, when the mountain torrents are a sheet of dingy foam, and the crystal depths of the river are converted for a time into an opaque flow of muddy water, the fly-fisher’s occupation’s gone. But when the turmoil ceases, and the soft south wind begins to disperse or break in upon the dense array of clouds, so as to chequer the streams, and rocks, and “ pastoral melan¬ choly” of the green mountains with the enlivening beams of the returning sun, with what pleasure does the angler approach the banks of a favourite and accustomed river ! How various and delightful are his sensations! Custom cannot stale their infinite variety:—on the contrary, the longer and more assiduously the pleasure is pursued, the greater the immediate enjoyment, and the more extended the train of agreeable remembrances for after-days. How exciting the first cast into a breeze-ruffled pool, when the unwetted gut still lies in rebellious and unyielding circles on the surface, and yet almost at the same moment the sounding reel gives notice that these circles have been instantaneously stretched into a straight and tightened line ! Then comes the long and continuous vibration of rod and reel, indicating the secure hooking of a goodly fish ; or that sullen and pulse-like tug, by which a still good¬ lier one, when hooked in a deep pool, frequently manifests a desire to dig its way to the bottom ; or that more inter¬ rupted music which results from the fantastic leaps of some whimsical individual, which skims and flounders on the top of the water like a juvenile wild-duck. The ordinary rules for fly-fishing are, to be most assi¬ duous when the streams are somewhat disturbed and in¬ creased by rain,—when the day is cloudy, and the waters moved by a gentle breeze, especially from the south. If the river contains long placid pools, then a steady stirring breeze is very desirable, as angling in such situations re¬ sembles lake-fishing, where nothing can be achieved upon a glassy surface. If the wind is low and the weather clear, of course the best angling is in the swiftest streams, and in those curling and perturbed eddies which head the smoother depths. In fishing the smoother pools of no great depth, be careful that the shadows of neither rod nor angler come upon the surface; but if a person is skilful in other respects, he need not fear his own sha¬ dow in a broad river, but w'ade boldly down the centre of the stream, fishing its various depths and currents before him and on either side. In clear rivers the flies should be small and rather slender-winged; but when the waters are muddy or increased by rain, a larger lure may be 1 Although Izaak Walton, that “ great master in the art of angling,” informs us that no man should in honesty catch a trout till the middle of March, yet the grayling is in best condition during the winter season. “ I do assure you,” says Charles Cotton, in the second part of the Complete Angler, “ which I remember by a very remarkable token, I did once take, upon the sixth day of December, one, and only one, of the biggest graylings, and the best in season, that ever I yet saw or tasted ; and do usually take trouts too, and with a fly, not only before the middle of this month, but, almost every year, in February, unless it be a very ill spring indeed; and have sometimes in January, so early as new-year’s tide, and in frost and snow, taken grayling in a warm sun-shine day Lr an hour or two about noon; and to fish for him with a grub it is then the best time of all.” 136 ANGLING. Angling, made use of. When the streams are brown with rain, an orange-coloured fly is good; in very clear weather a light- coloured one ; and a dark fly is advisable for troubled wa¬ ters. Though a great deal no doubt depends on a quick eye and an active and delicate hand, we are no great advo¬ cates for what is called striking a fish. If a large trout rises in a deep pool, it may be of advantage so to do; and this will be sufficiently accomplished by inclining the rod quickly aside, so as to draw out a few inches of the line; for if the reel is not allowed to run, this operation is apt to snap the gut, or otherwise injure the tackle. But if a trout, whether great or small, rises in a current or rapid stream, the sudden change in its position, immediately after it has seized the fly, is generally quite sufficient to fix the barb, without any exertion on the part of the angler. A variable state of the atmosphere is bad for angling; but neither is a uniformly dull gloomy day the most fa¬ vourable. It is scarcely possible to lay down any general rules on this branch of the subject. We have half filled a pannier during an electric hail-storm, when “ sky low¬ ered and muttered thunder,” and the aspect of the day was such as to deter more experienced though less zealous sportsmen from leaving the shelter of their homes. If the river is not too low, we always prefer what in ordinary language might be called a fine cheerful day, more parti¬ cularly if there is a fresh breeze. And what we would more particularly press upon the notice of the angler, as soon as he becomes master of the line, is, that he should cast his flies more frequently than is the usual practice, and, generally speaking, fish rapidly. This should be more especially attended to in streams where the trout are numerous and not large. Before enumerating and describing the different kinds of artificial flies in greatest repute, we shall mention a few of the principal materials used by the fly-fisher. The articles which he employs in common with those who pro¬ secute the other branches of the trade, are of course rods, hair and gut lines, reels and hooks, panniers and land¬ ing-nets ; but in addition to these he must be provided with a great variety of feathers, such as the slender plumes called hackles, from the necks and backs of com¬ mon poultry, and the wings of a considerable number of birds, such as woodcocks, snipes, rails, plovers, ducks, grouse, partridges, and others. The furs of quadrupeds are also indispensable; and of these the most useful are hares, squirrels, moles, martens, mice, and water-rats. The most esteemed hackles are the duns. The red, striped down the centi-e with black, and the red with a blackish root, are likewise useful, and more easily ob¬ tained. Since the introduction of Spanish poultry (by which name are designated the black breed with white tops), black hackles are now more common than for¬ merly. The proper time for plucking hackles is about Christmas. The feathers of the ostrich and peacock are of frequent service; and for salmon and sea-trout the gaudy plumes of parrots and other brilliantly attired fo¬ reign species, however unlike the generality of our north¬ ern insects, ought to be collected by every fly-fisher. The silks commonly used by the angler are of three kinds:—Is#, Barbers’ silk, used double, for splicing the top-pieces of rods ; 2<%, a more delicate kind, for fasten¬ ing on the rings through which the reel-line runs; 3t%, fine netting silk for whipping hooks and dressing flies. When we mention a pair of small pliers, fine-pointed scis¬ sors, needles, and wax, we have noted the principal mate¬ rials for the angler’s trade. In regard to rods, their length and formation are so much matters of individual taste, that few general rules can be laid down upon the subject. According to Da-^,1 niel, the wood should be cut about Christmas, and allowed^1 rl to season for a twelvemonth. Hazel is very generally used, especially that from the cob-nut, which grows to a great length, and is for the most part very straight and taper. The but-end should rather exceed an inch in dia- meter, and the shoots for stocks, middle pieces, and tops, should be as free from knots as possible. The tops are made from the best rush ground shoots. All these pieces should be kept free from moisture till the ensuing autumn, when such as are required to form a rod are selected; and, after being warmed over a gentle fire, they are set as straight as possible, and laid aside for several days. They are then rubbed over, by means of a piece of flannel, with linseed oil, which produces a polish, and brings off the superfluous bark: they are then bound tight to a straight pole, and kept till next spring, by which time they will be seasoned for use. They are then matched together in due proportions, in two, three, or more parts, according to the desired length, or the opinion of the maker as to the num¬ ber of pieces of which a rod should be composed. A well- constructed spliced rod of no more than two pieces casts a line with fully as much force, neatness, and accuracy as any other; but it is inconvenient to a traveller, or to any one whose dwelling is not close upon a stream. If the pieces are not ferruled, they must be spliced so as to join each other with great exactness. The principal object to be kept in view in the formation of rods in general is, that they should taper gradually and bend regularly. A frequent defect is their bending too much in the middle, owing to that part not being sufficiently strong. We have said that the length of a rod is rather a mat¬ ter of taste than of established rule. It must, however, bear a relation to the size of the river and the nature of the expected capture. A trouting rod is usually made from 12 to 14 feet in length, though some prefer them of greater extent, as giving more command over lakes and spreading pools. It should be made as light as is con¬ sistent with strength and durability, as a heavy rod is cumbersome, fatiguing, and unwieldy ; and a light one gives a more ready power in casting under hollow banks, or among trees or bushes. For pike and barbel a proper length is 16 feet; for perch, chub, bream, carp, eels, and tench, a shorter rod may be used; and 8 or 10 feet is sufficient for dace, gudgeon, ruff, bleak, &c. The port¬ ability of a rod depends of course on the number of joints; but its excellence being almost in the inverse ra¬ tio, care must be taken not to sacrifice its goodness mere¬ ly for the sake of a convenient form. According to Mr Bainbridge, the best rods are made from ash, hickery, and lance-wood; ash for the bottom piece, hickery for the middle, and lance-wood for the top- joints. If real bamboo can be procured of good quality, it is preferable to lance-wood. Rose-wood and partridge- wood from the Brazils may also be used for the top-pieces. The extreme length of the top-piece is usually composed of a few inches of whalebone. The rings for the reel¬ line may be made by twisting a piece of soft brass-wire round a tobacco-pipe, and soldering the ends together. They ought to diminish in size as they are made to ap¬ proach the top, and must form a straight and regular line with each other when the rod is put up for use. In finishing a rod the usual varnish is copal varnish, or Indian rubber dissolved over a slow fire in linseed oil. h may be stained by a dilution of nitric acid or oil of vitriol. When rods are stored for the winter, after use, they ought to be rubbed over with tallow or salad oil. As few anglers require to make their own rods, we deem it unnecessary to enter into a full detail of a mechanical practice which can only be sufficiently executed by an individual of pro- ANGLING. 137 essional experience. We shall merely mention, that, in addition to the woods already named, elder, holly, yew, nountain ash, and briar, all of which are indigenous to his country, furnish materials to the makers of rods. As lines may be purchased from the tackle-makers at a ;heaper rate than they can be made by an amateur whose ;ime and labour are of value for any other purpose, we hall not here enter into a detail of their formation. The I jest hair is procured from the tail of a well-grown stallion. 1 dlack hair is generally strong, but the colour is not very serviceable. Transparent and almost colourless hair is he most approved; and it ought to be round, regular, and I ree from blemishes. In the formation of Fines each hair n a link should be equal, round, and even, which propor- ions the strength and prevents single hairs from break- ng, and thus weakening the others. Chesnut or brown- ■ boloured hairs are best for ground angling, especially in nuddy water. Some anglers stain their lines a pale green f or fishing in weedy waters. Black is occasionally used in ■ (streams which flow from mosses, and are themselves of an jnusually dingy hue. The following are some of the methods used by anglers or dyeing their lines, whether of hair or gut. For a pale watery green.—To a pint of strong ale add lalf a pound of soot, a small quantity of walnut leaves, ind a little powdered alum; boil these materials for half J jr three quarters of an hour, and when the mixture is cold steep the gut or hair in it for ten or twelve hours. For a brown.—Boil some powdered alum till it is dis- • solved; add a pound of walnut-tree bark from the branches when the sap is in them, or from the buds, or the unripe | :ruit. Let the liquid stand till nearly cool, and skim it; , then put in the gut or hair, and stir it round for about a minute, or till it appears to have imbibed the desired tint. It ought not to be strongly tinctured, as it is apt to rot ivhen too dark. For a bluish watery tint the above ingre- t dients are also used, with the substitution of logwood in¬ stead of walnut. For a yelloiv.—The inner bark of a crab-tree boiled in water, with some alum, makes a good yellow, excellent for staining tackle used among decayed weeds, the colour of which it closely resembles. A tawny hue is obtained by steeping hair among lime and water for four or five hours, and then allowing it to soak for a day in a tan-pit. In the absence of other ingre¬ dients, both gut and hair may be easily stained by being left for twenty-four hours in strong tea, either with or without a few log-wood scrapings. The hair to be dyed ought to be selected from the best white. Silken or hempen lines may be tinted by a decoc¬ tion of oak bark, which is said to add to the durability of these materials. We shall now proceed to consider the subject in relation to the different species of fish which form the principal | objects of the angler’s art. The Salmon. (Salmo Salar.) As the natural history, classification, and characteristic distinctions of fishes will be given at length under the ar¬ ticle Ichthyology in this work, we shall not here enter into any description of either the structure or habits of the class, but confine ourselves for the present almost i, entirely to such points as are most essential to the prac¬ tical knowledge of the angler. 'I his fine fish delights in large and rapid rivers. It bites best from six in the morning till eleven in the forenoon, and from three in the afternoon till sunset. A moderate Angling, breeze is of advantage; and the best months are March, April, May, and June. The salmon is justly regarded by the angler as the king of fish; and when we consider that they occasionally measure four feet in length, and weigh upwards of 70 pounds, we may conceive how difficult a capture and how valuable a prize they sometimes prove. The most successful bait, as well as the most agreeable in the usage, is the artificial fly. This is made in imitation both of dragonflies and butterflies of various kinds; but the principles which we have already endeavoured to es¬ tablish at the commencement of this article make it un¬ necessary to describe the natural species. Even those who most warmly advocate the necessity of imitating ex¬ isting insects in the formation of their lures, admit that the salmon is so capricious as frequently to rise at an ar¬ tificial fly which bears no resemblance to any natural form of insect life. The following are the descriptions of six artificial flies which have been found very successful in raising salmon. No. 1 (see Plate XLV. fig. 1) is recommended as a spring fly, and is composed of the following materials: Wings of the dark mottled brown or blackish feather of a turkey; body of orange camlet mixed with a little mohair; and a dusky red or bright brown cock’s hackle, plucked from the back where the fibres are longest, for the legs. The hook should be of the same size as represented in the plate ; and it has been observed that all large salmon- flies should be dressed upon two or three lengths of gut twisted together, and that the silk in dressing be brought beyond the shank of the hook, and wrapped four or five times round the gut, so that it may not be speedily cut by the sharpness of the steel.1 This same fly, dressed with the wings of a somewhat brighter shade, and with the addition of a little gold wire or thread wrapped round the body at equal distances, will also serve for a more ad¬ vanced season of the year. No. 2 (see Plate XLV. fig. 2) is of smaller size, and may sometimes be dressed upon very strong single gut. Any feather of a coppery or dingy yellow colour, if not too coarse in the fibres, will be suitable for the wings; the body is of lemon-coloured mo¬ hair, mixed with a small portion of light brown fur or camlet, with a pale dusky ginger hackle over the whole. The chief object to be attended to in dressing this fly is to produce that uniform hue, devoid of gaudy colouring, from which it has received the name of the quaker fly. Of No. 3 (see Plate XLV. fig. 3) the wings are made fi'om the plumes of a cormorant, or from the mottled fea¬ thers of a dark mallard : the body is of dark sable, ribbed with gold wire, over which a dusky red hackle is thickly wound: the mottled feathers of a drake are used for the tail; and previous to fastening off, a small portion of flos silk should be unravelled and fastened at the extremity of the hook. This fly, though, like the preceding, of a some¬ what sombre cast, is frequently used with success in summer. No. 4 (see Plate XLV. fig. 4) belongs to the gaudy class of lures, “ which,” says Mr Bainbridge, “ however fanciful or varied in shade or materials, will frequently raise fish when all the imitations of nature have proved unsuccessful; indeed so fastidious and whim¬ sical are the salmon at times, that the more brilliant and extravagant the fly, the more certain is the angler of di¬ version.” In this, of course, we perfectly agree. The wings of the fly in question are formed of the extreme end of a Guinea fowl’s feather, not stripped, but having the fibres remaining on both sides of the middle stem. A blood- red hackle is fastened on with the wings, and so arranged vol. in. 1 Bftinbridge’s Fly-F'uher's Guide, p. 96. s 138 ANGLING. Angling, as to extend beyond them: the dyed feathers used by military men will suit, if another showy biped, the scarlet maccaw, is not accessible. The green feather which forms the eye of the peacock’s tail should be fastened at the head, and left hanging downwards, so as to cover the body for the space of half an inch ; and a few filaments of the same part of the feather may be fastened at the tail. No. 5 has the wings formed from the darkish brown speckled portion of a bittei-n’s wing stripped off from the stem: the head ought to be of the same colours as the body, which is formed of the reddish brown part of a hare’s fur, and deep copper-coloured mohair; a bittern’s hackle is put over the body for legs, and a forked tail is added, made of a pair of single filaments of the same fea¬ ther as the wings. Of No. 6 the wings are formed of the mottled feathers of a peacock’s wing, intermixed with any fine plain dusky red ; the best mixture for the body is the light brown inner hair from a bear’s skin, sable fur, and gold-coloured mohair; gold twist, a large black cock’s hackle, and a red one a little larger, with a bit of deep red mohair for the head. In addition to these, we might enumerate the brown fly, the blue fly, the king-fisher, the prime dun, the great palmer, the golden pheasant, the gray mallard, and many others; but such as are above de¬ scribed will suffice for the purposes of the present trea¬ tise. Fig. 6 of Plate XLV. represents an excellent spring fly; and an approved summer kind is shown on the same plate, fig. 5. It may be stated at once, that so far from imitating na¬ ture, the maker of salmon flies can scarcely form them in too unnatural and extravagant a manner. Let him call in the aid of fancy at all times and places, at least in this country; for the cold and cloudy clime of Scotland assur¬ edly furnishes nothing resembling the lures most frequent¬ ly and most successfully used. The superabundant use of gold and silver wire ought, however, to be avoided, as it not only causes the fly to sink too much in the water, but prevents its being neatly or lightly thrown. Spring flies for salmon are usually made of a larger size, though not so gaudily dressed as those of summer. A salmon rod is generally proportioned to the size of the river which the angler frequents; but it ought not to be less than 15 feet in length. The reel ought to be large enough to contain 80 or 90 yards, so as to admit of abund¬ ance of line being given out when required; for many fish, when struck, run out to a great distance, and with such immense rapidity as to prevent the possibility of the angler’s moving in the proper direction with sufficient quickness. A salmon, for the most part, darts violently up the stream; and, as the command and direction of the fish is more easily kept with a short than a long line, it is advisable to prevent his getting too far ahead, by keeping the rod well back in the opposite direction, and by run¬ ning towards him along the margin of the stream. When he gains the head of the current, a salmon frequently throws himself several times out of the water, on which occasions the angler must yield him freely a little of the line ; but during his general and less violent manoeuvring, he will of course be the sooner exhausted the more firmly he is held. When he appears to be making for some safe haunt or secret sheltering place, the great object is to turn him towards safer ground, either by relying on the soundness of the tackle, or, if he proves very powerful as well as very obstinate, then a pebble or two may be thrown, so as to fall a little in advance of his position, and he will probably turn himself round. Some fish become very sulky, and will lie after being hooked, for a long time, motionless near the bottom. In this case also the pebbles must be had recourse to; for the more a fish is kept in motion, the sooner he becomes exhausted. When he begins to show his side, and exhibits other unequivocal % symptoms of exhaustion, a favourable landing-place should\ be looked for; and when the proper time arrives, which can only be learned by the (sometimes dearly bought) lessons of experience, then is he to be drawn by degrees to the gravel bank, or the gaff applied, and the prize secured. When feeding, salmon are usually found at the foot of a strong stream, terminating in an eddy or whirlpool. “ And first,” says our father Walton, “ you shall observe that usually he stays not long in a place, as trouts will, but, as I said, covets still to go nearer the spring-head; and that he does not, as the trout and many other fish, lie near the water side or bank, or roots of trees, but swims in the deep and broad parts of the water, and usually in the middle, and near the ground; and that there you are to fish for him; and that he is to be caught as the trout is, with a worm, a minnow, which some call a penk, or with a fly.” When the water is either too much discoloured for the use of the artificial fly, or, running into the opposite ex¬ treme, becomes (especially in very still weather) too clear and bright, salmon may be successfully angled for with the worm. In this case trolling tackle is sometimes used. In trolling with minnow or other small fish, the foot lengths ought to be about three yards long, and furnished with one or two swivels, to prevent the line from twisting, as well as to enable the bait to play freely. A lead or shot proportioned to the strength of the stream should be fastened to the line, about a foot above the bait. The top of the rod should be stiffer than that used for fly-fishing; and when the hook is baited, it ought to be thrown first across, and then drawn up the stream. The salmon fry leave the spawning beds early in spring, and ere long make their way towards the sea. In a few months they again seek their native streams, having, du¬ ring a very short period, attained to the weight of several pounds. Without entering into the merits of the much- disputed question (still in some points of difficult solution) as to the difference or the identity of samlets, salmon fry, salmon trout, white trout, gilse, and salmon, suffice it to say, that all the migratory kinds of trout, usually charac¬ terized by their small number of spots, their silvery co¬ lour, and the superiority of their flesh, may be angled for with much larger and more gaudily plumed flies than those used in the capture of river-trout. Flies dressed upon the same principles as those used for salmon, but of smaller size, will be found well adapted for whitings, and the different kinds of sea or migratory trouts above men¬ tioned. The Common Fresh-water or Fiver Trout. (Salmo Fario.) This species varies greatly in size and colour, in accord¬ ance probably with the nature and abundance of its food, the strength and depth of the river in which it occurs, and the physical properties of soil and climate. Fish seem, more than most animals, to depend on peculiar and unappreciable circumstances, for the full and charac¬ teristic development of their characters ; and they conse¬ quently exhibit great contrariety of aspect among indivi¬ duals of the same species. If a canal or reservoir, or any other great accumulation of water, is formed by the hand of man, where the hand of nature had from time imme¬ morial recognised only some small and solitary streamlet, the lapse of a very few years produces large and heavy fish, where none but trouts of the most trifling size had ever been seen before. The writer of these observations kept a minnow little more than half an inch long in a glass tumbler for a period of two years, during which time there was no perceptible increase in its dimensions. ANGLING. n*. lad it continued in its native stream, subjected to the -w ittening influence of a continuous flow of water, and a 11 f onsequent increase in the quantity and variety of its 0 jod, its cubic dimensions would probably have been 20 I lines greater; yet it must have attained, prior to the lapse fa couple of years, to the usual period of the adult state, n regard to birds and quadrupeds, the individuals of the II ame species are not distinguishable from each other by r ny peculiarities either of form or colour, at least within e he limits of a restricted locality; but it appears to be 1 therwise with several species of fish, more especially 1 1 routs. Those of the Clyde and Tweed, although both I ivers draw their primary sources almost from the same > , icuntain, present a constant and well-marked difference > i their external aspect; and a corresponding dissimilarity xists among the characteristic varieties of almost every < iver and lake in Scotland; “ which I tell you that you 1 nay the better believe that I am certain, if I catch a II rout in one meadow he shall be white and faint, and very :( ke to be lousy; and as certainly, if I catch a trout in 2 he next meadow, he shall be strong, and red, and lusty, ( , nd much better meat. Trust me, scholar, I have caught ii viany a trout in a particular meadow, that the very shape d nd the enamelled colour of him hath been such as hath i . )yed me to look on him; and I have then with much ! Measure concluded with Solomon, ‘ Every thing is beau- 11 iful in his season.’ ” • No fish affords the angler more varied or more constant ) port than the trout. For nine months in the year, under " . avourable circumstances, fly-fishing may be practised for i t with success. Westerly and southerly winds are the 1 nost favourable, especially in spring; but during warm t K nd cloudy summer weather, the point from which the ” . find blows is of slight consequence. “ He who considers e he wind,” says Solomon, “ will never sow;” and the same » emark is not inapplicable to angling. Whoever desires I ! o become a successful practitioner in the art, must angle i n all weathers, and under every variety of circumstances, ' towever unpropitious the prospect may be. Trout are i generally supposed to rise more freely during a dark and 5 owering day, following a clear bright night, as brilliant i) noonshine detains them in their lurking-places; and on I he ensuing day they are consequently more inclined for (1 bod. On the other hand, after a gloomy or darkish night ii hey are less easily tempted, having glutted themselves i I vith moths and other nocturnal insects, which, during the i 1 summer months, are abundant on the waters. In throw- i| | Jig the line the angler should endeavour to make his e ‘ ^ear fall as lightly as possible on the surface, and his flies i f should drop opposite, or somewhat above his own position, a ind then be played gently and neatly downwards and i icross the stream. When a trout is seen to rise at a na- ] tural fly or other insect, the artificial one should be of- i’ered him by being thrown, not directly over him, but !) t ibout a yard higher up the stream; and, if he is inclined o to rise again, he will probably meet it half-way. When a E * hsh, on being hooked, descends beneath the surface, and t | struggles in the depths below, it is a pretty sure sign that i f he is well secured; but when he flounders on the surface, » 1 or leaps occasionally into the air, more care is necessary, ii as in that case the hook will frequently be found to be i] I only skin-deep. In playing and landing a large trout the ii same precautions are necessary as in salmon-fishing, al¬ ii though in regard to smaller fish, if the angler is standing » in the centre of a stream, and finds it inconvenient to « wade frequently ashore, a few additional turns will ex¬ haust the capture, which may then be drawn rapidly and 139 steadily to the hand, and secured by a firm grasp behind Angling, the gills. We have frequently practised a summary me-'v-<<~v->^' thod of landing even tolerably sized fish, which, though it cannot always be effected, is, when possible, a great saving of time. If, from the moment the trout is struck, he is prevented from redescending in such a manner that the upper part of his head and eyes are retained above or on a level with the surface, he will for the space of a good many seconds be so much astonished as to be incapable of any active exertions, and will frequently allow himself to be drawn in that position, and without resistance, straight ashore. The following flies are in repute among anglers. The black gnat appears about the end of April. The body is formed of a black strip from an ostrich feather, and ought to be dressed thick and short; the wings of a pale star¬ ling’s feather, or dressed as a hackle with a pale dun. The March brown or dun drake is frequently visible by the middle of March. The wings are made from the mottled feathers of a partridge’s tail, and the body of hare- ear fur, intermixed with a little yellow worsted; a grizzled hackle for legs. The hazel fly is of a round form, and difficult of imita¬ tion. It is a killing fly in May and June, especially where bushes abound. The body is composed of ostrich harl of two colours, black and purple twisted together; the wings of the sandy-coloured feathers from under the w ings of a thrush, or the reddish plumes of a partridge’s tail; a bluish hackle, twisted pretty full, serves both for the under wings and legs. The great dark dun., according to Mr Bainbridge, is one of the earliest flies which appear upon the water, and may be used in February, if the weather is mild. The wings are formed from the dun feathers of a mallard’s wing; the body of mole-fur, mixed with a little dark brown mohair; a dark grizzled hackle for legs. Salmon frequently rise at this fly, which may be used with success early in the morning during the whole fishing season. The wrens tail has no wings: the body is of sable fur, with a little gold-coloured mohair, and a feather from the tail of a wren. The grouse hackle is also wingless: the mixture for the body is dark olive, dusky yellow, and a little gold-coloured mohair. It is formed of a fine mottled grouse’s feather of a reddish brown, running a little dusky towards the but- end of the stem, and the downy portion, if any, plucked away. The stonefly is found along the edges of streams, and is a favourite article of food among trout. It is a species of Phryganea, and springs from a caddis or aquatic larva. The wings lie flat, and are imitated by the mottled fea¬ ther of a hen-pheasant or peahen. The body is com¬ posed of dark brown fur from a bear’s skin, or the deeper part of a hare’s ear, mixed with yellow camlet or mohair; a longish grizzled hackle is wrapped under the wings. The mealy brown or fern fly is excellent for grayling in May. Its wings should be formed from the under part of a throstle or fieldfare’s wing, especially from such feathers as have a yellowish tinge. Its body is of a dusky orange, from the light brown fur of a fox’s breast, with a pale dun hackle for legs. The orange fly has four wings made of the blue feather of a mallard-teal. The head is of the dark fur from a hare’s ear; the body gold-coloured mohair mixed with orange-camlet and brown fur, a small blue cock’s hackle for legs. This is said to be an alluring fly to young salmon. 1 Walton’s Complete Angler, p. 128, edit. 1823. 140 ANGLING. Angling. The hares ear is chiefly used as a drop-fly. The wings -^v^/are from the light part of a starling’s wing feather, and the body of dark hare-ear fur. According to Daniel, when the streams are deep, the same body winged with a rail¬ wing’s feather and a red hackle is very killing during the summer season. The yellow dun is used in the morning and evening during the months of April and May, and again in Sep¬ tember. The body, is made of yellow yarn unravelled, or with marten’s fur, and mixed with a little pale ash-colour¬ ed fox-cub fur ; the wings are formed upright, from the under part of a snipe’s wing, with a pale dun hackle for legs. The hawthorn fly is in use from the middle of April to the end of May, from ten o’clock till three. It has trans¬ parent wings, which may be imitated with the palest fea¬ ther of a snipe or mallard’s wing; horn shavings, or the membranous substance found in the core of an apple, serve the same purpose ; the body is of black ostrich harl, with a black hackle for legs. The summer dun has a thicker form than most of the dun flies, and is dressed upon a short-shanked hook. Mole-fur ribbed with ash-coloured silk is employed for the body; the wings are from the wood-pigeon, with an ash-coloured hackle for legs. The black-hackle fly is an approved lure during warm weather, early in a summer morning. The body is form¬ ed of a thin-dressed ostrich harl, cut close; the wings, four in number, are from the pale feather of the starling’s wing. The red spinner is used as a dropper. The wings are formed of the grayish feather of a drake, tinged with reddish yellow; the body a red hackle, with a twist of gold. This fly is eagerly taken by chub in the evenings of July. The little yellow May or willow fly resembles the green drake, on a small scale. The body is formed of yellow fur from the marten’s neck, or of yellow worsted unravelled, and mixed with a very small portion of hare-ear fur; the wings are of mallard’s feather dyed yellow. This fly ap¬ pears early in May, and may be used till the appearance of the green drake, of which it is the usual precursor. The brown dun is made of otter’s fur mixed with le¬ mon-coloured mohair; the wings are from the fieldfare, with a ginger hackle for legs. This is an excellent fly towards the approach of twilight. The green drake or May fly appears about the second or third week of May, and continues about a month. The body is made of hog’s down, or light bear’s hair, inter¬ mixed with yellow mohair; or of barbers’ yellow silk only, warped with pale flos silk, and a small strip of peacock’s harl for the head : a bittern’s hackle is the best imitation of the legs and dark stripes of the body; with the long hairs of the sable or polecat for the tail. The rayed feathers of a wild mallard, dyed of a greenish yellow, suf¬ fice for wings. The blue blow is a very small fly used during the sum¬ mer months, and for the first fortnight in August. It is made of a lapwing’s top, or any dark blue fur, dressed on a very small hook. The wings are of thistle-down or bluish- white hackle. The black midge is also a small species, the body of which is dressed with brownish-black silk, and a blue cock’s hackle. It is taken freely after a shower in the summer evenings. The gray drake follows the green of the same name, although they sometimes occur together. It is an excel¬ lent afternoon fly for large trouts. The body is formed of a dirty-white ostrich harl, dressed with flesh-coloured silk, and ribbed with a dark-grizzled cock’s hackle; the head is made of peacock’s harl, like that of the green drake; the wings from a mottled mallard feather, or thatC^J ''l of a mallard-teal, and the tail of sable or polecat’s hair. The cinnamon fly has four wings, large in proportion to the body. They are made from the pale reddish-brown feathers of a hen, dressed full; the body of dark brown fur, with a ginger hackle for legs. This fly, according to Mr Bainbriclge, is excellent for the Welsh rivers during the months of August and September. The sandfly forms an excellent lure, and may be very generally used from April to September. The wings are formed from the sandy-coloured feathers of the landrail’s wing, with a ginger hackle for legs ; and the bright sandy, coloured fur from the neck of a hare, mixed with a little orange mohair for the body. If dressed as a hackle, the feathers from under the thrush’s wing resemble the na¬ tural hue of the wings of the insect. The great black ant makes its appearance in sultry weather, from the middle of June to the latter end of Au¬ gust. The wings are made of the pale-blue feathers from beneath a snipe’s wing, or from a tomtit's tail. The body is of black ostrich harl, made thick towards the tail and be¬ neath the but of the wings; the legs of a reddish-brown hackle. The great red ant is nearly contemporaneous with the preceding, which it resembles in size and form. The wings are made of a light starling’s feathers ; the body of gold-coloured mohair, or copper-coloured peacock’s harl, with a ginger hackle for legs. Among the preceding flies will be found some which will assuredly suit for any river, or for any period of the fishing season. The angler who places implicit confidence in the generally received opinion, that in every stream, and at each season, there is one particular fly in much more special request than any other, will do well to pre¬ pare for an unknown river, by making ready a couple of lines, each with three flies all of different kinds. For example, a March brown at the end of the line, a dun hackle, with a lighter or darker body to suit the weather or complexion of the stream, for the first dropper, and a red hackle with peacock body for the second dropper; or, 2dly, a sand fly at the end, with a grouse hackle or wren’s tail, with orange body, for the first dropper; and a pale yellow or cream-coloured hackle over a bluish body, or one of the ant flies, as the second dropper. These are promising flies for most seasons of the year, though, like the others, they require to be changed according to the circumstances of time and place, or the varying caprices of the finny tribes. As night fishing is a favourite amusement with many anglers, we shall describe a few of the flies in most repute for the practice of nocturnal sport. The white moth, with wings made from the feathers of a white owl, the body of white ostrich harl, with a white cock’s hackle over it. The brown moth is winged from the feathers of a brown owl, or the back feathers of a brown hen; the body is of dark bear’s hair, covered with a brown cock’s hackle. The mealy cream is made from the tawny feathers of a white owj for the wings, with some soft fur of the same colour, and a pale yellow hackle for the body. The following selection has been recommended by an experienced angler, in relation to the alleged succession of flies during several of the principal months of the fishing season. Ltf, For March, a dun fly, made of dun wool, and the feathers of a partridge wing; or the body made of black wool, and the feathers of a black drake: 2d, For April, a stonefly, the body made of dark wool, dyed yellow under the wings and tail: 3r/, For the beginning of May, a ruddy fly, made of red wool, and bound about A N G L 'ith black silk, with the feathers of a black cock hanging •angling on his sides next his tail: 4/4, For June, a green- jh fly, the body made of black wool, with a yellow list n either side, the wings taken off the wing of a buzzard, ound with black broken hemp : 5/4, The moorish fly, the ody made of dusky wool, and the wings of the blackish mil of a drake: 6/4, The tawny fly, in great repute till ie middle of June ; the body made of tawny wool, the ings contrary, one against another, composed of the hitish mail of a white drake: 7/4, For July, the wasp y, the body made of black wool, cast about with yellow Ik, and the wings of drakes’ feathers: 8/4, The steel fly, jproved in the middle of July; the body made with reenish wool, cast about with the feathers of a peacock’s lil, and the wings made of those of a buzzard: 9/4, For aigust, the drake fly, the body made with black wool ist about with black silk, the wings of the mail of a lack drake, with a black head. When rivers are very low and clear, from a long conti- uance of summer drought, it has been recommended to se a pair of wings made from the feather of a landrail, i r the mottled feather of a teal, with a well-cleaned gen- e fixed upon the hook. During a similar condition of ie water, even when no wind is stirring, and the sun fining in its greatest lustre, trouts may be taken with a nail wrens tail, grouse, smoky dun, or black hackles, the igler fishing straight down the water, by the sides of reams and banks, and keeping well out of sight, with as Iing a line as can be neatly managed, and the foot-lengths iry fine. At these times the fish may be often seen with leir dorsal fins above water, and with skilful manage- icnt may be made to snap at the above-named flies, /hen one is hooked, the rest dart off; but if the angler 3eps concealed, they will return again in a very short me; and thus several fish may be taken even in summer om the clearest pools. Another plan has also been re- | ommended as likely to prove successful when the wea- icr is bright and the water low: Take a line of about a ard in length, and fix it to a short, stiff rod, and having lited the hook with a natural fly, such as the stonefly, 1 r the gray or green drake (Ephemeree), drop it between ushes over steep hollow banks, or under the projecting )ots of trees. In fishing a river with which the angler has no previous ’! cquaintance, the most approved practice is to try the ddies which are frequent at the corners of streams, and here the circular movement of the current throws out a '• 'equent sustenance for the finny race. There the larger •out often lie; and it must consist with the experience 1 every angler, that an excellent capture is sometimes lade repeatedly from some small spot behind or beside a articular stone, where from day to day one well-sized sh seems to succeed another in the favourite feeding round. In this knowledge of peculiar localities consists ie chief advantage of a previous acquaintance with the ater. The smaller fish are found in most abundance in ie widely spread and shallow streams, as well as in the i xtended parts of pools of no great depth. As a general ule, the angler may be advised to fish with the wind on is back and the sun in front, which not .only gives him greater command of his line, but prevents himself or is shadow from being so distinctly perceived. A strict dherence, however, to this plan is by no means advisable, s the angler’s position in relation to sun and wind must equently vary with the natural course of the river, the Instruction of overhanging wood, and the greater or less ommand of pool and stream presented by the varying ^ nrm of the adjoining shore. As bait-fishing for trout, though regularly followed by ome> less generally admired and practised than the I N G. 141 more elegant use of the artificial fly, fewer words will suf- Angling, fice. When the streams are swollen and discoloured, fine^^v^^ trout may be taken with a running line without float, and so leaded that it shall touch the ground without resisting the force of the stream. The lead should be fixed about eight or ten inches above the hook; and the best baits are well-scoured earthworms. The dew, the garden, and the lob worms are one and the same in species, although they vary considerably in form, size, and colour, according to age, and season, and the nature of the soils. The lob¬ worm, according to Daniel, is of two sorts, the old, knotted, the young, without knots, which some for distinction call maiden lobs, and others red worms. The latter kind, with a red head, a streak down the back, and a broad tail, are the most esteemed. By some they are called squirrel- tails. These and other worms, it need scarcely be ob¬ served, are easily obtained in fields and gardens, espe¬ cially where there has been any recent mixture of veget¬ able or animal remains. They may be preserved for a considerable period, and even improved in their texture and condition, by being kept in damp moss, changed from time to time, and occasionally wetted with a little new milk. In dry weather, when worms are difficult to be obtained, they may be procured by emptying a few buckets of water in situations where they were known to occur during a moister season. The brandling worm is streaked from head to tail in alternate circles of a red and yellow hue, and is darker at its anterior than posterior portion. They occur in old dunghills, in heaps of inch vegetable mould, and among rotten tan bark. They have this advantage over the others, that they may be used without preparation or scouring. Though the choice of wrorms does not seem a very suitable subject for poetry, it has been thus versified by Mr Gay in the Rural Sports : You must not every worm promiscuous use, Judgment will tell thee proper baits to choose : The worm that draws a long immod’rate size, The trout abhors, and the rank morsel flies; And if too small, the naked fraud’s in sight, And fear forbids, while hunger does invite. Those baits will best reward the fisher’s pains. Whose polish’d tails a shining yellow stains; Cleanse them from filth, to give a tempting gloss Cherish the sullied reptile race with moss ; Amid the verdant bed they twine, they toil, And from their bodies wipe their native soil. The preceding rhymes apply chiefly to the kind called gilt-tails. Gentles are the larvae of different kinds of car¬ nivorous winged flies. They may be kept in a mixture of oatmeal and bran, and are readily produced in a piece of liver, or any other flesh or fish, exposed in an earthen vessel to prevent their escape when grown to a proper size. All kinds of maggots, as well as those called gen¬ tles, serve admirably for the more delicate kinds of bait¬ fishing. The caddis worms, before alluded to as the larvae of the Phryganea or stonefly, when taken out of their cases, are a favourite bait for trout; and different kinds of grasshoppers are likewise used with great success. The creeper or water cricket, an aquatic - larva, found under stones within the water-mark, ought also to be attended to by the natural bait-fisher. The palmer worms or wool beds are the hairy catei'pil- lars of certain nocturnal moths. Though refused by al¬ most all birds except the cuckoo, they are swallowred by trouts, and may be preserved alive for many weeks in a box with damp earth, strewed over with the leaves of the tree or bush on which the species was observed naturally to feed. The young brood of wasps and bees are useful to the angler; and for eight or ten days after their first appear¬ ance in summer there is no better or more killing bait 142 ANGLING. Angling, than a small reddish beetle called the bracken clock in the —north of England, the Melolontha horticola of naturalists. Salmon roe is greatly lauded by Barker, who appears to have been the first to discover its merits. “ I have found an experience of late which you may angle with, and take great store of this kind of fish. First, it is the best bait for trout that I have seen in all my time, and will take great store, and not fail if they be there. Secondly, it is a special bait for dace or dare, good for chub, or bottlin, or grayling. The bait is a roe of a salmon or trout. If it be a large trout, that the spawns be any thing great, you must angle for the trout with this bait as you angle with the brandling, taking a pair of scissors and cut so much as a large hazel-nut, and bait your hook; so fall to your sport,—there is no doubt of pleasure. If I had known it but twenty years ago, I would have gained a hundred pounds only with that bait: I am bound in duty to di¬ vulge it to your honour, and not to carry it to my grave with me. I do desire that men of quality should have it, that delight in that pleasure. The greedy angler will murmur at me, but for that I care not.” Many kinds of pastes are prized by the bait-fisher. They may be used for chub, carp, and bream in Septem¬ ber and during all the winter months, and may be made up about the size of a hazel-nut; if foj1 2 roach and dace, the bigness of a pea will suffice. All pastes are improved by being mixed up in the making with a little cotton wool, which makes them firmer and more tenacious, and hang better on the hook. They suit well for fishing in quiet places, with a small hook and quill float. We shall here subjoin a few recipes for the making of fishing pastes, which, although we introduce them under the head of the river-trout, may be regarded as equally efficacious in the capture of other kinds of fish.1 Minnow-fishing for trout is a favourite pastime with many anglers, and the process is one by which very large fish are frequently captured. The tackle used resembles that for salmon, but is lighter and finer, with a single line of gut at the bottom. The hooks vary in size according to the general dimensions of the trout angled tor; and the middle- sized and whitest minnows are the most esteemed. The following were Walton’s directions for baiting, with a view to this department of the sport. Put your hook in at his mouth and out at his gill; then having drawn your hook two A or three inches beyond or through his gill, put it againO n into his mouth, and the point and beard out at his tail, ^ and then tie the hook and his tail about very neatly with a white thread, which will make it the apter to turn quick in the water : that done, pull back that part of your line which was slack when you did put your hook into the minnow the second time; I say, says Walton, pull that part of your line back so that it shall fasten the head, so that the body of the minnow shall be almost straight on your hook : this done, try how it will turn by drawing it across the water or against a stream ; and if it do not turn nimbly, then turn the tail a little to the right or left hand, and try again, till it turn quick. We may add, that the practice of fishing with the artificial minnow is justly dis¬ carded by all judicious anglers. For representations of the minnow tackle, and hooks baited, see Plate XLV., fig. 12, 13, and 19. The Great Lake Trout. (Salmo Ferox.'f It appears that the gray or great trout of the British fresh-water lakes, though never described or characterized as a distinct species, has at various times excited the at¬ tention of ichthyologists. Trout of enormous dimensions are mentioned by Pennant as occurring in the Welsh lakes; and Donovan gives Loch Neagh in Ireland as an¬ other locality. Very large trout have been killed in Ulls- water in Cumberland, and still larger in Loch Awe in Argyllshire. The late Mr Morrison of Glasgow claimed the merit of discovering these fish in the last-named lo¬ cality about 40 years ago; and the largest recorded to have been killed there weighed 25 pounds. Mr Lascelles, a Liverpool gentleman, has also of late years been equally assiduous and successful in their capture; and it appears that any persevering sportsman is almost certain, with the proper tackle, to obtain specimens in Loch Awe, of this gi'eat fish, weighing from 10 to 20 pounds. The largest we have lately heard of weighed 191 pounds. It is said to be by far the most powerful of our fresh-water fishes, exceeding the salmon in actual strength, though not in activity. The most general size caught by trolling ranges from 3 to 15 pounds: beyond that weight they are of uncommon occurrence. If hooked upon tackle of mode- 1 Red paste may be made with a large spoonful of fine wheat-flour, moistened with the white of an egg, and worked with the hands until tough. A small quantity of honey or loaf-sugar finely powdered must be added, together with some cotton-wool spread equally over the paste when pressed flat in the hand; it must be well kneaded, to mix the cotton thoroughly; colour it with a little vermi¬ lion. A small piece ot fresh butter will prevent it from becoming hard, and it will keep good a week. White paste may be composed of the same ingredients, omitting the vermilion; and yellow paste in like manner, with the addition of turmeric. Salmon paste.— lake one pound of salmon-spawn in September or October; boil it about 15 minutes, then beat it in a mortar until sufficiently mixed, with an ounce of salt and a quarter of an ounce of saltpetre; carefully pick out the membrane in which the spawn is contained, as it is disengaged from it; when beat to a proper consistence, put it into gallipots, and cover it over with bladders tied down close, and it will keep good for many months. Various oils were formerly in great repute among anglers for rubbing over their baits, but as we believe their beneficial effects were entirely imaginary, we shall not occupy our pages by their repetition. A single extract from Izaak Walton will suffice. “ And now I shall tell you that which may be called a secret: I have been a-fishing with old Oliver Henley, now with God, a noted fisher both for trout and salmon, and have observed that he would usually take three or four worms out of his bag, and put them into a little box in his pocket, where he would usually let them continue half an hour or more before he would bait his hook with them. I have asked him his reason, and he has replied, ‘ he did but pick the best out, to be in readiness against he baited his hook the next time. But he has been observed, both by others and myself, to catch more fish than I or any other body that has ever gone a-fishing With him could do, and especially salmons; and I have been told lately, by one of his most intimate and secret friends, that the box in which he put those worms was anointed with a drop, or two or three, of the oil of ivy-herries made by expression or infusion; and told that by the worms remaining in that box an hour, or a like time, they had incorporated a kind of smell, that was irresistibly at- tractive, enough to force any fish within the smell of them to bite.” We need scarcely remind the reader of the “ Complete Angler,” that that admirable work is of higher value for the manner in which the subject is discussed, and the beautiful accessories of pure style, poetical sentiment, and picturesque illustration, than for the amount of direct practical information which it conveys. The simplicity and goodness of Izaak alton’s nature seem to have induced a greater degree of credulity than was always consistent with an accurate perception of the truth, and hence every chapter abounds with statements which could not pass current in these more critical days. As a useful work in relation to the mere angler, it cannot be said to hold a high rank, although it must ever delight the general reader, and all who desire to refresh themselves by “ the pure well of English undefiled ” 2 We are indebted for the principal materials of the following account of this interesting fish to a manuscript of Sir William Jar- dine s, with which we have been kindly favoured by the author. It forms part of a series of Memoirs on British Fishes, which that assiduous naturalist has been for some time past preparing for the press. ANGLING. ling,irate strength, they afford excellent sport; but the gene- /■^■ral method of fishing for them is almost as well adapted for catching sharks as trout; the angler being apparently more anxious to have it in his power to state that he had ^ught a fish of such a size, than to enjoy the pleasure of die sport itself. However, to the credit of both parties, it nay be stated, that the very strongest tackle is sometimes mapped in two by its first tremendous springs. The or- linary method of fishing for this king of trouts is with a powerful rod, from a boat rowing at the rate of from three ;o four miles an hour, the lure a common trout from three ;o ten inches in length, baited upon six or eight salmon looks, tied back to back upon strong gimp, assisted by wo swivels, and the wheel-line strong whip-cord. Yet ill this, in the first impetuous efforts of the fish to regain ts liberty, is frequently carried away for ever into the crystal depths of Loch Awe ! When in their highest health and condition, and in- leed during the whole of the time in which they are not •mployed in the operation of spawning, these fish will carcely ever rise at a fly. At these periods they appear o be almost entirely piscivorous; so, with the exception of light lines, baited also with trout, trolling is the only ad- isable mode of angling for them. The young, however, ise very freely at ordinary lake-trout flies, and are general- Y caught in this way from one to one and a half pound /eight. They occur abundantly near the outlet of the lake. About the middle of August, and during the three fol¬ ding months, the parent fish retire, for the purpose of pawning, to the deep banks of the lake in the neighbour- ood of the gorge, and into the gorge of the lake itself, /here it empties its immense waters, forming the river Awe. "hey arc said to remain engaged in this operation for two r three months, and at this time their instinctive tenden- ies are so far changed, that they will rise eagerly at irge and gaudily dressed salmon-flies, and may be either ngled for from the banks, or trolled with a cross line, 'here the outlet of the lake is narrow. They do not ap- ear either to ascend the rivers which enter the loch, or ) descend the Awe to any extent, though an occasional traggler has been taken some way down the river. Their I pawning places are exclusively on the banks, or at . ie gorge of the loch, and they never attempt to seek ie salt-water.. When in good season, and in their rongest condition, they appear to roam indiscriminately nough every part of the loch, though there are certain >ots which may be more depended upon than others, ■ ipd where an experienced angler will have little difficulty i hooking one of these fine fish. To their great strength o may observe that they add unequalled rapacity; and tor attaining to the weight of three or four pounds, they fj ipear to feed almost exclusively on smaller fish, and do | at spare even their own young. A small trout of this >ecies, not weighing more than 11 pound, will often dash ■ ; a, bait not much inferior to itself in size ; and instances e recorded of larger fish following with eager eye, and - tempting to seize upon others of their own kind after l'ey Had been hooked and were in the act of being land- y t ie angler. It is probably on account of this ng manifestation of a more than usually" predaceous 143 ]mlmofoTx Sir WiUiam Jardine has named the sPecies Angling. When in perfect season, and full-grown, it is a very handsome fish, though the head is always too large and long to be in accordance with our ideas of perfect sym¬ metry in a trout. The body is deep and thickly formed, and all the members seem conducive to the exercise of great strength. The colours are deep purplish brown on the upper parts, changing into reddish gray, and thence into fine orange-yellow on the breast and belly. The whole body, when the fish is newly caught, appears as if glazed over with a thin tint of rich lake-colour, which fades away as the fish dies, and so rapidly, that the pro¬ gressive changes of colour are easily perceived by an at¬ tentive eye. The gill-covers are marked with large dark spots; and the whole body is covered with markings of different sizes, and varying in amount in different indivi¬ duals. In some these markings are few, scattered, and of a laige size ; in others they are thickly set, and of small¬ er dimensions. Each spot is surrounded byr a paler rin0-, which sometimes assumes a reddish hue; and the spots become more distant from each other as they descend beneath the lateral line. The lower parts of these fish are spotless. All the fins are broad, muscular, and ex¬ tremely powerful; and it is from the number of their bony rays that the specific characters which distinguish this species from the common trout (salmofario) are the most easily and accurately evolved. The dorsal fin is of the same colour with the upper part of the fish; it is marked with large dark spots, and contains fifteen rays, which number exceeds by three that which characterizes the common trout. The caudal fin is much larger and more fleshy. The pectoral, ventral, and anal fins, are very muscular on their anterior edges, and of a rich yel¬ lowish-green colour, darker towards their extremities. They contain respectively 14, 10, and 12 rays, whereas the numbers in the corresponding fins of the common trout are 13, 9, and 11. The tail is remarkable for its breadth and consequent power. In adults it is perfectly square, or might even be described as slightly rounded at its extremity: in the young it is slightly forked, and appears to fill up gradually as the fish advances in age. In the common trout, on the contrary, the forked shape of the tail is a permanent characteristic. The flavour of this great lacustrine species is coarse and indifferent. The colour of the flesh is orange-yellow, not the rich salmon-colour of a fine common trout in good season. The stomach is very capacious, and on dissec¬ tion (differing singularly in this respect from the salmon) is almost always found gorged with fish. W e have bestowed a somewhat lengthened notice on the species above described ; but we doubt not that the novelty of the subject will not only plead our excuse, but render the information now given highly acceptable both to the sportsman and the naturalist. It is certain that a more attentive examination of the finny tribes which in¬ habit our lakes and rivers would bring to light several new species, and more clearly illustrate the history of others which are still involved in obscurity.1 The gigantic species of the Swiss lakes, one of which, ith the great troits of Loch Awe In , r T " ' '^'7 ,12, Poundf. each* These were probably of the same species -xles Grassmere and Rvdal we HdnWl chain of English lakes of which Windermere is the chief, and which in- Hed, of these beautiful 2ers when f t PmeS /^"onymous with the salmo ferox of Jardine. The lake trouts, properly so ken of the weight of ^ t0 three t0 ^pounds, and have in one or two rare instances been indermere chain corresnond more iP • n •S' l08?,0* ^ hwater, again, which does not belong to what we have denominated the e would particularly reifommend t fu °ther characters with the species found in the lakes of the Scotish Highlands, nietiines called This is rl a i u 16 attentl,on of T01'tsmcn the great fresh-water river trout, or bull trout as we believe it is hers, especially when fmimHn Z,8?"16 anSlers as an agedt overgrown individual of the ordinary kind (mlmo fario); and by curacy of the latter onininn ; - i utumlh as a lake trout which had left its more usual haunts for the purpose of spawning. The p mon is, however, interfered with by the occasional occurrence of this variety in such unambiguous situa- 144 Angling, killed in the lake of Geneva, weighed 67 pounds, are probably identical with those of Loch Awe, Ullswater, &c. Though equal to salmon, both in size and strength, they differ in their habits from those fish, and do not appear at any time to seek the waters of the ocean. In¬ deed their existence in the lake of Constance, the avail¬ able communication of which with the sea is cut oft by the falls of Schaffhausen, demonstrates their independence of saline waters. It does not appear, from any informa¬ tion which has reached us on the subject, that these great continental lake-trouts ever rise at the artificial fly. In Loch Ard there is a finely formed and beautiful va¬ riety of the common trout, varying in its matured condi¬ tion from one and a half to three pounds in weight. It affords excellent sport to the angler fishing from a boat about 50 yards off the deeper parts of the shores. Ihey rise freely to small salmon or sea-trout flies, dressed after the model of a gray or green drake. ANGLING. The Grayling. (Salmo Thymdllns.) This fish loves the clear streams of mountainous coun¬ it is common in Lapland, where its intestine is usod i , I. !„ A wiillr ifno Tnrma + inr* tries. al ia r-; . . „ . as rennet, along with rein-deer milk, in the formation of cheese. It is a bold and sportive fish, but more tender in the mouth than the trout. It rises well to the camlet fly, and to several of the other small-sized trout-flies. We killed it readily in Switzerland with a moorfowl wing and hare-ear body. They may also be taken with the caddis worm and other ground baits. According to the Rev. Mr Low, the grayling is frequent in the streams of the Orkney Islands, though very rare in the rivers of the mainland of Scotland. The Pike. (Esox Lucius.) This “ fell tyrant of the liquid plain” is not regarded as indigenous to the waters of Britain, but is said to have been introduced in the time of Henry VIII. That it was well known in England at an earlier period is how- ever evident, both from the book of St Alban’s, printed by Wynken de Worde in 1496, and from the account of the great feast given by George Nevil, archbishop of York, in the year 1466. There is in truth no evidence either of its non-existence in this country at a remote peiiod, or of its importation during comparatively recent times. The voracity of this fish is almost unexampled, even in a class remarkable for their omnivorous propensities. Goslings, young ducks, and coots, water-rats, kittens, and the young of its own species, besides every kind of fresh¬ water fish, have been found in the stomach of the pike. The rich and varied supply of all kinds of tackle, which may be obtained in the shops of the principal dealers m our larger cities, induces us to abstain from any descrip¬ tion of "the different hooks employed in minnow and other bait-fishing, as such details are not very intelligible with¬ out the aid of numerous engravings. More knowledge will be gained by a few minutes inspection of the articles themselves in the hands of an intelligent workman, than can be conveyed by the most elaborate treatise on the Aaving described the characteristic modes of angling surpasfaU the^others yielded to’lhe ftTslidlo contend with the otter for its prey, and h> amder we shall now proceed to a briefer consideration of been known to pull a mule into die water by t e no e. Lme of the remaking subjects of our sport. This fish is in season from May to February, and is angled some ot tne remaning j for by trolUng with a strong-topped rod. I he hooks are The Char or Case Char. (Salmo Alpinus.) generally fastened to a bit of brass wire for a few inches The Torgoch or Bed Char. (Sahno Salvelinus.) from the shaft, to prevent the line from being snapped. These fish which in a culinary point of view, are deserv- Different methods are used in angling for pike. Troll- edly the most highly prized of all the permanently fresh- ing, in the more limited sense of the word, signifies ca.c i- miter species, ar? scarcely attainable by the angler's skill, ing fish with the gorge hook which .s com^o edof Of late Tears they have risen more freely than m former or what is called * times in the meres of the north of England ; but the cap- practised with the aid of a floated hne mid tore of a char by rod and line is still regarded as an uncom- consists in the use ot large hooks, so baited as . mon occurrence. They appear to retire during the warmer the angler to strike the fish the moment he feels it bite, months to the deepest of the still waters, as the fisher- immediately after which he d™f ® men engaged in throwing their nets for pike, perch, and Trolling for pike may be practised during the want trout, over the very grounds where, during the colder months, when trout fishing has ceased, and t e season of the year, the char abound, never catch any of season of the year is in fact more convenient for the sp these fish in summer. Although a good deal has been owing to the decay or diminution of the weeds wh ch written upon the subject, it does not appear that the dis- usually surround their favourite ^aunts; ^1^ t^ ^ tinctive history of the two species above named has been tion of chub and dace, which bite pretty flee y as yet made out. Both the case char and the red char bottom all winter, scarcely any other fish can be are1 found in Windermere; and the principal distinc- upon for sport during the more inclement portion tion in their habits and history consists in this, that the year. To bait a gorge hook take a baiting needle, and ho former ascends the rivers, where it spawns about Michael- the curved end to the loop of the gimp, to which the hook, mas; whereas the latter deposits its ova along the shores tied; see Plate XL V. fig. 20; then introduce the point oDh of the lake, and not till the end of December or the be- needle into a dead bait s mouth, and bring it out at the miu sinning of the year. In angling for char the same flies may die of the fork of the tail, by which means the piece o be used as those best adapted for the smaller-sized lake- which covers the shank of the hook, and partof the conne trouts; and as the latter occur wherever the former is found, ing wire, will lie concealed m the interior ot the bait. ' the sportsman has the better chance of making amends shank will be in the inside of its mouth, and the ha for the probable disappointment which will attend his pur- the outside, turning upwards. See 1 late L^ V. hg. suit of the one, by a more successful capture of the other, keep the bait steady on the hook, fasten the tai pa J lions as the Clyde above the falls, the waters of which have no communication with any lake. Ihis species sometimes attains weight of 8, 10, or even 12 pounds; hut it differs from the Loch Awe trout n being generally, it not exclusively, tound i Those of many of the lesser Highland lakes, such as Loch Ard and Loch Chon, ascend the mountain streams in the autumn to p and in the ordinary practice of angling (with the. artificial fly) are rarely caught above the weight of three pounds. W e lastyeart ^ received some very singular trouts from a small loch called Lochdow, near Pit main, in Inverness-shire. Their heads were si round and their upper jaws were truncated like that of a bull-dog. They do not occur in any of the neighbouring lochs, an not been observed above the weight of half a pound. Trouts of the ordinary shape likewise occur in Lochdow. ANGLING. : linn above the fork to the gimp, with a silk or cotton thread; 5^.0r a neater method is, to pass the needle and thread through the side of the bait, about half an inch above the tail so as to encircle the gimp in the interior. The baits used vary in weight from one to four ounces, and the hooks must be proportioned to the size of the fish with which they are baited. The barbs of the hook ought not to pro¬ ject much beyond the sides of the mouth, because, as the pike generally seizes his prey crosswise, and turns it be¬ fore it is pouched or swallowed, if he feels the points of the hook he may cast it out entirely. In trolling for pike, it is advised to keep as far from the water as possible, and to commence casting close by the near shore, with the wind blowing from behind. When the water is clear and the weather bright, some prefer to fish against the wind. “ After trying closely,” says Mr Salter, “ make your next throw further in the water, and draw and sink the baited hook, drawing it straight up¬ wards near to the surface of the water, and also to right and left, searching carefully every foot of water; and draw your bait with the stream, because you must know that jack and pike lay in wait for food with their heads and eyes pointing up the stream, to catch what may be com¬ ing down; therefore experienced trollers fish a river or stream down, or obliquely across ; but the inconsiderate as frequently troll against the stream, which is improper, because they then draw their baited hook behind either I jack or pike when they are stationary, instead of bringing it before his eyes and mouth to tempt him. Note—Be particularly careful, in drawing up or taking the baited hook out of the water, not to do it too hastily, because you will find by experience that the jack and pike strike or seize your bait more frequently when you are drawing it upwards than when it is sinking. And also further observe, that when drawing your bait upwards, if you occasionally shake the rod, it will cause the bait to spin and twist about, which is very likely to attract either jack or pike.”1 These fish are partial to the bends of rivers and the bays of lakes, where the water is shallow, and abounding in weeds, reeds, water lilies, &c. In fishing with the gorge hook, and the angler feels a run, he ought not to strike for several minutes after the fish has become stationary, lest he pull the bait away before it is fairly pouched. If a pike makes a very short run, then remains stationary for about a minute, and again makes one or two short runs, he is probably merely retiring to some quiet haunt before he swallows the bait; but if, after remaining still for three or four minutes, he begins to shake the line and move about, the inference is that he has pouched the bait, and feels some annoyance from the hook within, then such part of the line as has been slackened may be wound up, and the fish struck. It is an unsafe practice to lay down the rod during the interval between a run and the sup¬ posed pouching of the bait, because it not unfrequently happens that a heavy fish, when he first feels the hooks in his interior, will make a sudden and most violent rush up the river or along the lake, and the line is either instantly broken, or is carried, together with both the rod and reel, for ever beyond the angler’s reach. “ When the pike cometh,” says Colonel Venables, “ you may see the water move, at least you may feel him; then slack your line and give him length enough to run away to his hould, whither he will go directly, and there pouch it, ever be¬ ginning (as you may observe) with the head, swallowing that first. Thus let him lye untill you see the line move in the water, and then you may certainly conclude he hath Angl' pouched your bait, and rangeth about for more; then^-^v- with your trowl wind up your line till you think you have it almost streight, then with a smart jerk hook him, and make your pleasure to your content.”2 The fresher and cleaner the bait is kept, whether for trolling, live-bait, or snap-fishing, the greater is the chance of success. As pike, notwithstanding their usual voracity, are some¬ times, as the anglers phrase it, more on the play than the feed, they will occasionally seize the bait across the body, and, instead of swallowing it, blow it from them re¬ peatedly and then take no further notice of it. The skil¬ ful and wily angler must instantly convert his gorge into a snap, and strike him in the lips or jaws when he next attempts such dangerous amusement. The dead snap may be made either with two or four hooks. (See Plate XLV. fig. 21.) Take about twelve inches of stout gimp, make a loop at one end, at the other tie a hook (size No. 2), and about an inch farther up the gimp tie another hook of the same dimensions; then pass the loop of the gimp into the gill of a dead bait-fish, and out at its mouth, and draw the gimp till the hook at the bottom comes just behind the back fin of the bait, and the point and barb are made to pierce slightly through its skin, which keeps the whole steady; now pass the ring of a drop-bead lead over theloop of the gimp,fix the leadinsidethebait’smouth, and sew the mouth up. (See Plate XLV. fig. 22.) This will suffice for the snap with a couple of hooks. If the four- hooked snap is desired (and it is very killing), take a piece of stout gimp about four inches long, and making a loop at one end, tie a couple of hooks of the same size, and in the same manner as those before described. After the first two and the lead are in their places, and previous to the sewing up of the mouth, pass the loop of the shorter gimp through the opposite gill, and out at the mouth of the bait; then draw up the hooks till they occupy a position correspond¬ ing to those of the other side: next pass the loop of the longer piece of gimp through that of the shorter, and pull all straight: finally, tie the two pieces of gimp together close to the fish’s mouth, and sew the latter up. Some anglers prefer fishing for pike with a floated line and a live bait. When a single hook is used for this pur¬ pose, it is baited in one or other of the two following ways: Either pass the point and barb of the hook through the lips of the bait, towards the side of the mouth, or through beneath the base of the anterior portion of the dorsal fin. (See Plate XLV. fig. 25.) W’hen a double hook is used take a baiting needle, hook its curved end into the loop of the gimp, and pass its point beneath the skin of the bait from behind the gills upwards in a sloping direction, bring¬ ing it out behind the extremity of the dorsal fin; then draw the gimp till the bend of the hooks are brought to the place where the needle entered, and attach the loop to the trolling line. (See Plate XLV. fig. 23.) Unless a kind of snap-fishing is intended, the hooks for the above purpose should be of such a size as that neither the points nor the barbs project beyond either the shoulder or the belly of the bait. . Snap-fishing is certainly a less scientific method of ang¬ ling for pike than that with the gorge or live bait; for when the hooks are baited the angler casts in search, draws, raises, and sinks his bait, until he feels a bite. He then strikes with violence, and drags or throws his victim on shore; for there is little fear of his tackle giving way, 1 The Troller'i Guide, by T. F. Salter, Lond. 1820. In the work above quoted will be found a full account of the necessary imple¬ ments, and the most approved practice, in this department of the art. The Experienced Angler, p. 36. Third edit. Lond. 1668. VOL. III. T 14G A N G Angling, as that used in snap-fishing is of the largest and strongest kind. “This hurried and unsportsman-like way of taking fish,” it is observed in the Trailers Guide, “ can only please those who value the game more than the sport af¬ forded by killing a jack or pike with tackle which gives the fish a chance of escaping, and excites the angler’s skill and patience, mixed with a certain pleasing anxiety, and the reward of his hopes. Neither has the snap-fisher so good a chance of success, unless he angles in a pond or piece of water where the jack or pike are very numerous or half starved, and will hazard their lives for almost any thing that comes in their way. But in rivers where they are well fed, worth killing, and rather scarce, the coarse snap-tackle, large hooks, &c. generally alarm them. On the whole, I think it is two to one against the snap in most rivers; and if there are many weeds in the water, the large hooks of the snap, by standing rank, are continu¬ ally getting foul, damaging the bait, and causing much trouble and loss of time.” Pike sometimes rise at an artificial fly, especially in dark, windy days. The fly ought to be dressed upon a double hook, and composed of very gaudy materials. The head is formed of a little fur, some gold twist, and (if the angler’s taste inclines that way, for it is probably a matter of indifference to the fish) two small black or blue beads for eyes. The body is framed rough, full, and round, the wings not parted, but made to stand upright on the back, with some small feathers continued down the back to the end of the tail, so that when finished they may exceed the length of the hook. The whole should be about the bulk of a wren. The largest pike ever killed in Britain was taken with a peacock-feather fly in Loch Ken, near New Galloway. It weighed 72 pounds.1 During clear and calm wreather in summer and autumn, pike take most freely about three in the afternoon: in winter they may be angled for with equal chances of suc¬ cess during the whole day: early in the morning and late in the evening are the periods best adapted for the spring. The Carp. (Cyprinus Carpio.') This fish, like the preceding, is asserted to have been introduced into England by Leonard Mascal, a gentleman of Sussex, early in the 16th century; and in good com¬ pany, if there is truth in the old distich, Turkies, carps, hops, pickerell, and beer, Came into England all in one year. The carp is, however, mentioned as a dayntous fysshe, though scarce, by Juliana Barnes, in the year 1496. It attains to a prodigious size in the waters of the south of Europe, and in the Lake of Como is said sometimes to weigh 200 pounds. It breeds more freely in ponds than in rivers, although those of the latter are more esteemed. Angling for carp requires, according to Walton, “ a very large measure of patience.” The haunts of this fish in the winter months are the broadest and least disturbed parts of rivers, where the bottom is soft and muddy; but in summer it usually lies in deep holes, near some scour, under roots of trees, and beneath hollow banks, or in the neighbourhood of beds of aquatic weeds. In ponds they thrive best in a rich marl or clayey soil, where they have the benefit of shade from an overhanging grove of trees. Small carp hite eagerly, but the larger and more experi¬ enced fish are deceived with difficulty. The rod should be of good length, the line strong, furnished with a quill float, and ending in a few lengths of the best silk-worm gut. The hook is proportioned to the size of the bait, and a single shot is fixed about 12 inches above it. “ Three LING. rods,” says Daniel, “ may be employed; one with the baiti at mid-water, another a foot or less from the bottom, and® the third to lie upon it when the line and lead are not * discovered, as in the two former ; the places intended to be fished in should, the night before, be ground-baited with grains, blood, and broken worms, incorporated together with clay, the hook baits should be red worms taken out of tan, flag or marsh-worms, green peas so boiled as to soften, but not to break the skin, and throwing some in now and then. When this bait is used (which should be with one on the hook to swim a foot from the ground), in case of a bite, strike immediately; a large carp, upon taking the bait, directly steers for the opposite side of the river or pond.”2 During hot weather, when these fish are about to spawn, and whilst lying among the weeds near the sur¬ face, they may be angled for with a fine line, without either sink or float. The hook may be baited with a red worm, a pair of gentles, a caterpillar, or a cad bait, and thrown lightly as in fly-fishing, and then drawn towards the angler. If it can be made to fall first upon the leaf of! some water plant, and then dropped upon the surface, the chance of success will be increased. The best months are May, June, and July, and the most advisable times of the day are from sunrise to eight in the morning, and from sunset during the continuance of twilight, and onwards through the night. It is the opinion of many, though we cannot trace the origin of the idea, no doubt an erroneous one, that the 10th of April is a fatal day for carp. The Bream. {Cyprinus Brama.) This fish breeds both in deep, slow-running rivers, and in ponds. It prefers the latter. The most enticing bait is a well-conditioned earthworm, although the angler also uses paste made of bread and honey, wasp grubs, grass¬ hoppers, &c. Boiled wheat serves well for ground-baiting the spot on the preceding nights, and some fasten a num¬ ber of worms to a piece of turf, and sink it to the bottom. When the ground has been thus prepared, and the tackle put in order, the angler should commence his labours by three or four in the morning. Let him approach the place with caution, so as not to be perceived by the fish, and cast his hook neatly baited with a live and mov¬ ing worm, in such manner that the lead may lie about the centre of the prepared ground. The bream is a strong fish, and runs smartly wdien first struck; but after a few turns he falls over on his side, and allows the angler to land him without much trouble. He is by no means so lively as the carp. The best hours for bream are from four till eight in the morning, and from four in the after¬ noon till eight in the evening. In the river Trent, near Newark, there are two kinds of bream. The common species is that called the carp bream, from its yellow colour; and it sometimes attains the weight of eight pounds. The other species or variety, regarded by Mr Revett Shepherd as a nondescript, never exceeds a pound in weight. It is of a silvery hue, and is known by the name of white bream.3 The bream, though rare in Scotland, occurs in Loch Maben. The Tench. {Cyprinus Tinea.) This species is a lover of still waters, and his haunts in rivers are among weeds, or pools well screened by bushes. Tench are found spawning from June till Septem¬ ber, and they are in the best condition from the latter month till the end of May. The tackle should be strong, with a swan or goose-quill float for ponds, and a piece of cork for rivers. The hook (in size from No. 4 to 6) should 1 Daniel’s Rural Sports, vol. ii. p. 270. “ Ibid. p. 257* 3 Linn. Trans, vol. xiv. p. 037* A N G I ,r be whint to sound silk-worm gut, with two or three shot «^xed to it at the distance of a foot. The bait should float about a couple of feet from the surface, and should be drawn occasionally gently upwards, and allowed slowly to sink again. Small marsh worms, middle-sized lobs, or the red species found in rotten tan, are to be recommended. « He will bite,” says Walton, “ at a paste made of brown bread and honey, or at a marsh-worm, or a lob-worm : he inclines very much to any paste with which tan is mixed, and he will bite also at a smaller worm with his head nipped off, and a sod-worm put on the hook before that worm ; and I doubt not but that he will also in the three hot months, for in the nine colder he stirs not much, bite at a flag-worm, or at a green gentle: but can positively say no more of the tench, he being a fish that I have not often angled for; but I wish my honest scholar may, and be ever fortunate when he fishes.” The Barbel. ( Cyprinus Barbus.) In a culinary point of view this is one of the worst of the fresh-water fishes. It is gregarious, and roots among the soft banks with its nose, like a sow. The angling season commences in May, and continues till September. The most approved hours are from daylight till ten in the morning, and from four in the afternoon till about sunset. The line should be strong and rather heavily leaded, so that the bait may float about half an inch from the ground. Considerable caution is required in playing this fish, as he is apt to run off when struck, with great violence, to¬ wards some stronghold, and in so doing sometimes breaks both rod and line. He is rather nice in his baits, which must be kept clean and sweet, and untainted by musty moss. “ One caution,” says Mr Daniel, “ in angling for barbel, will bear repetition: never throw in the bait far¬ ther than enabled by a gentle cast of the rod, letting the plumb fall into the water with the least possible noise. It is an error to think that large fish are in the middle of the river: experience teaches the fallacy of this opinion : they naturally seek their food near the banks, and agitating the waters by an injudicious management of the plumb will certainly drive them away. It is incredible the quan¬ tities of barbel sometimes caught by this method. Per¬ sons of great veracity have asserted that upwards of one hundredweight have been taken in one morning.” The Chub. (Cyprinus Cephalus.) The rivers of England are stored with a much finer variety of fresh-water fish than those of Scotland. The chub occurs in the Annan. It is however a fish but lightly esteemed, either for sport or the table. He is a dull fish on the hook, bites eagerly, and is soon tired. Caution is requisite on the part of the angler, as the chub is naturally fearful, and sinks towards the bottom of the stream on the slightest alarm. The baits used are mag¬ gots, beetles, grasshoppers, salmon-roe, &c. Black and dun flies gaudily dressed, and ribbed with gold or silver twist, are well adapted to take them in streams. They likewise rise at the red-spinner. But perhaps the best way to secure this fish is by dibbing with a grasshopper. As chub are often seen in some favourite haunt lying near the surface of the pool, the angler, concealing himself as much as possible, ought to move his rod cautiously over the spot, and drop his bait gently upon the water, a few inches in advance of the fish’s head. The landing net is particularly necessary in angling for chub, as the best spots are generally encumbered by trees or bushes, which prevent the fish from being drawn to hand, or pulled ashore. . I N G. 147 The Dace. (Cyprinus Leuciscus.) Angling. This fish is of gregarious habits, and haunts the deeper waters near the piles of bridges, shady pools, and beneath the masses of collected foam caused by eddies. In the warmer months of the year they also congregate in the shallows. They rise at a variety of flies, and are likewise angled for with red worms, brandlings, &c. Above Rich¬ mond, as soon as the weeds begin to rot, a grasshopper used as an artificial fly is found very successful in hot weather among the shallows. This mode can only be practised in a boat, with a heavy stone to serve as an an¬ chor, fastened to a few yards of rope. The boat drifts gently down the stream, and the stone is dropped when¬ ever the angler considers himself in the neighbourhood of a likely place. Standing in the stern, he first throws directly down the stream, and then to the right and left; and after trying for about a quarter of an hour in one spot, he again weighs anchor, and proceeds to another station. The Roach. ( Cyprinus Rutilus.) The carp has been named the “ water-fox” on account of his subtlety, and the roach the “ water-sheep,” by rea¬ son of his silliness. This fish makes good soup, though very bony, and otherwise not much esteemed. The sea¬ son for roach fishing in the Thames, where it attains to a larger size than elsewhere, commences about the end of August. “ Next let me tell you,” says Walton, “ you shall fish for this roach in winter with paste or gentles, in April with worms or caddis, in the very hot months with little white snails, or with flies under water; for he sel¬ dom takes them at the top, though the dace will. In many of the hot months roaches may be also caught thus :—Take a May-fly or ant-fly; sink him with a little lead to the bottom, near the piles or posts of a bridge, or near to any posts of a wear—I mean any deep place where roaches lie quietly—and then pull your fly up very leisurely, and usually a roach will follow your bait to the very top of the water, and gaze on it there, and run at it, and take it, lest the fly should fly away from him.”1 Vast shoals of this species ascend the streams in the parish of Killearn, from Loch Lomond, and are caught by nets in thousands. Their emigration from the loch, however, continues only for the space of three or four days towards the end of May.2 The Bleak. ( Cyprinus Alburnus.) This small and active fish may be angled for with what is called a pater noster line, which consists of half a dozen fine hooks fastened about 6 or 8 inches from each other. These may be baited with gentles, or more variously, to increase the temptation, with a gentle, a small red woi'm, a fly, &c. and thus several fish may be hooked at the same time. In angling for bleak the tackle must be very fine. In fresh streams they rise well at the black gnat, or any other small sad-coloured fly. The Gudgeon. (Cyprinus Gobio.) Gudgeons are angled for near the ground with a small red worm. They frequent the shallows during the hot months, and retire before winter to the stiller and deeper waters. As an article of food they are highly esteemed. The Minnow. ( Cyprinus Phoxinus.) This is the fish by means of which most youthful ang¬ lers commence their experience of the art. “ He is a sharp biter,” says Walton, “ at a small worm, and in hot 1 Complete Angler, p. 218. 2 Statistical Account of Scotland, vol. xvi. p. 100. 148 ANGLING. Angling, weather makes excellent sport for young anglers, or boys, or women that love that recreation; and in the spring they make of them excellent minnow-tansies ; for being washed well in salt, and their heads and tails cut olf, and their guts taken out, and not washed after, they prove excellent for that use; that is, being fried with yolks of eggs, the flower of cowslips and of primroses, and a little tansie. Thus used, they make a dainty dish of meat.” The Loach. ( Cobitis Barbatula.) The loach is entirely a ground fish,living in clear and gra¬ velly streams. It is an excellent bait for eels, and is also a nutritious food for man, though of a slimy and somewhat forbidding aspect. The Eel. (Anguilla Vulgaris.) This well-known and snake-like species has its favourite haunts in the muddy bottom of the bays of lakes, among weeds, under large stones, and in the clefts of the banks of rivers. The habits of the eel are nocturnal, and the largest and finest are usually caught with night-lines. They are troublesome fish, from their great tenacity of life, and the tortuous motions by which, in their endea¬ vours to disengage themselves, they entangle or destroy the angler’s tackle. They afford little amusement to those accustomed to the more elegant branches of the art. The isle of Ely, according to some authorities, was so called in consequence of its being the place from whence the kings of England were anciently supplied with eels. Indeed Cambridgeshire is still famous for this fish.1 The Perch. (Perea Fluviatilis.) This gregarious fish is angled for with a worm or min¬ now. It is a bold biter during the warm months of the year, though very abstemious in the winter season. When a shoal is met with, great sport is frequently obtained. A small cork float is used, and the bait is hung at various depths, according to circumstances, a knowledge of which can only be obtained by practice. In angling near the bottom, the bait should be frequently raised nearly to the surface, and then allowed gently to sink again. When the weather is cool and cloudy, with a ruffling breeze from the south, perch will bite during the whole day. The best hours towards the end of spring are from seven to eleven in the morning, and from two to six in the af¬ ternoon. In warm and bright summer weather, excellent times are from sunrise till six or seven in the morning, and from six in the evening till sunset. The first printed work on angling in the English lan¬ guage is by Dame Juliana Barnes or Berners ( The Trea- tyse of Fgsshinge with an Angle), and forms part of the JBooh of St Albans, emprented at Westmestre by Wynken de Worde, in 1496. Of this book there are various old editions, and it has, we believe, been twice reprinted in modern times. It is less useful to the angler than curious in the eyes of the bibliographer. Hawking, Hunting, Fouling, and Fishing, with the true measures of Blowing, &c. now newly collected by W. G. Faukener. 4to, Lond. 1596. A Book of Fishing with Hooke and Line, and of all other Instruments thereunto belonginge, made by L. M. 4to, Lond. 1590. This work contains remarks on the pre¬ servation of fish in pools, and some improvements on the directions of the “ religious sportswoman” Juliana Barnes. L. M. signifies Leonard Mascall. A Neu Book of good Husbandry, very pleasaunt, and of great profite, both for gentlemen and yeomen ; conteining the Order and Maner of Making of Fish-pondes, with the Breeding, Preseruing, and Multiplyinge of the Carpe, 1 ench, Pike, and Troute, and diverse kindes of other Fresh Fish. Written in La- tine by Janus Dubrauius, and translated into English at the speciale request of George Churchey, Fellow of Lion’s Inne, the 9th Februarie 1599. 4to, Lond. 1599. Certain Experiments concerning Fish and Fruit, practised by John Taverner, Gentleman, and by him published for the benefit of others. 4to, Lond. 1600. The Secrets of Ang- ling; teaching the Choicest Tooles, Baytes, and.Seasons for the taking of any Fish in Pond or River: practised and 'familiarly opened in Three Bookes. By J. D., Esquire. 8vo, Lond. 1613. The author of this work is named in the third edition of Walton’s Angler as one Jo. Davors; but, from an entry in the books of Stationers Hall, as given in the second volume of “ British Bibliography,” p. 355, he is mentioned as John Dennys, Esquire. Large ex¬ tracts from this work are given by Sir Egerton Bridges, in the last volume of his Censura Literaria. The poetry, of which several passages are quoted by Walton, is remark¬ able for its beauty. As the volume is rare, we shall here present the reader with a few stanzas. You nymphs that in the springs and waters sweet Your dwelling have, of every hill and dale, And oft amid the meadows green do meet To sport and play, and hear the nightingale, And in the rivers fresh do wash your feet, While Progne’s sister tells her wofull tale ; Such ayd and power unto my verses lend As may suffice this little work to end. And thou, sweet Boyd, that with thy wat’ry sway Dost wash the cliffes of Deignton and of Week, And through their rocks with crooked winding way Thy mother Avon runnest soft to seek; In whose fair streams the speckled trout doth play, The roach, the dace, the gudgin, and the bleike; Teach me the skill, with slender line and hook, To take each fish of river, pond, and brook. In comparing the amusement of angling with the ex¬ citement to be derived from gaming and other pleasures, he adds— O let me rather on the pleasant brinke Of Tyne and Trent possess some dwelling place, Where I may see my quill and corke down sinke. With eager bite of barbel, bleike, or dace ; And on the world and his Creatour thinke. While they proud Thais painted sheet embrace ; And with the fume of strong tobacco smoke And quaffing round are ready for to choke. Let them that list these pastimes then pursue, And on their pleasing fancies feed their fill; So I the fields and meadows green may view, And by the rivers fresh may walke at wille, Among the dazies and the violets blue, Bed hyacinth and yellow daffodil, Purple narcissus like the morning rayes, Pale ganderglas, and azore culverkayes. I count it better pleasure to behold The goodly compasse of the lofty skie; And in the midst thereof, like burning gold. The flaming chariot of the world’s great eye; 1 “ Here I hope I shall not tresspass upon gravity, in mentioning a passage observed by the reverend professor of Oxford, Doc*®1 Prideaux, referring the reader to him for the author’s attesting the same. When the priests in this part of the country would st retain their wives, in despight of whatever the pope or monks would doe to the contrary, their wives and children were miraculous} turned all into eels (surely the great into Congers, the less into Greggs), whence it had the name of Eely. I understand him, a Li - of Eels.” (Fuller’s Worthies. Cambridgeshire.) ANGLING. 149 The wat’ry clouds that in the ayre uproPd With sundry kinds of painted colours flie ; And faire Aurora lifting up her head, All blushing rise from old Tithonus bed. The hills and mountains raised from the plains, The plains extended levell with the ground, The ground divided into sundry vains, The vains enclos’d with running rivers round, The rivers making way thro’ nature’s chains, With headlong course into the sea profound; The surging sea beneath the vallies low, The vallies sweet, and lakes that lovely flow. Then follow the two stanzas quoted at the beginning of this article. Our next work on angling is The Pleasures of Princes, or Good Men’s Recreations; containing a Dis¬ course of the General Art of Fishing with the Angle, or otherwise, and of all the hidden secrets belonging tfiere- unto; together with the Choyce, Ordering, Breeding, and Dyetting of the Fighting Cocke ; being a worke never in that nature handled by any former author. Lond. 1614, 4to. This work forms part of the second book of the English Husbandman, by G. M. (Gervais Markham.) A Briefe Treatise of Fishing; with the Art of Angling. Lond. 1614, 4to. This work is little else than a reprint from a portion of the Book of St Alban’s, and forms part of the Jewell for Gentrie, by T. S. Cheap and Good Husbandry, by Gervais Markham. 4to, Lond. 1616. This work contains a chapter on Fish and Fish-Ponds. Country Contentments; or the Husbandman’s Recreations, bjr J. M. In the fifth and sixth editions of this volume (4to, Lond. 1633 and 1639), will be found, the Whole Art of Angling, as it was written in a small treatise in rime, and now, for the better understanding of the reader, put into prose, and adorned and enlarged. This work is a prose version, with additions, of Davors’ Secrets of Angling. The Country Gentleman’s Companion, 2 vols. 12mo, Lond. 1753, is a reprint, without acknowledgement, of Markham’s work. The Art of Angling ; wherein are discovered many rare secrets very necessary to be known by all that delight in that recreation, written by Thomas Barker, an antient practitioner in the said art. 12mo, Lond. 1651. In an epistle to the reader, prefixed to the first edition, and in the dedication of the two last to Edward Lord Montague, Barker speaks of himself as having practised angling for more than half a century. He also says he was born and educated at Bracemeall, in the liberty of Sa¬ lop, being a freeman and burgesse of the same city; adding, “ if any noble or gentle angler, of whatever de¬ gree soever he be, have a mind to discourse of any of these wayes and experiments, I live in Henry the 7th’s Gifts, the next doore to the Gatehouse, in Westm. My name is Barker, where I shall be ready, as long as please God, to satisfye them, and maintain my art during life, which is not like to be long.” See British Bibliography, by Sir Eg. Bridges and Joseph Haselwood, vol. ii. p. 356. I he Compleat Angler, or the Contemplative Man’s Recrea¬ tion ; being a discourse of Fish and Fishing, not unworthy the perusal of most anglers. 12mo, Lond. 1653. This is the first edition of Izaak Walton’s celebrated work. It werrt through five editions during the author’s lifetime; and in the course of its republication was enlarged and improved. I he fifth edition forms the first part of the niversal Angler, by Walton, Cotton, and Venables, 12mo, °nd. 1676 ; and is accompanied hy & second part (written J Gotten), which treats more fully of fly-fishing. The I'- Ji' an(^ seventh editions were published in 1750 and ‘J' ’ Moses Browne, author of the Piscatory Eclogues M ° i\er " ?r^s; The eighth edition was published by Sir ‘Oin Hawkins in 1760, and has been succeeded by many er® 81nce that period, of which the most recent and °s eautifully adorned is that by John Major, with an introductory essay and illustrative notes. 8vo, Lond. Angling. 1823. The third edition of the Compleat Gentleman, by'^v^. Henry Peacham, 4to, Lond. 1661, contains a chapter concerning Fishing. The Experienced Angler, or Ang¬ ling Improved ; being a general discourse of Angling. 8vo, Lond. 1662. This work, of which there are seve¬ ral editions, is by Colonel Robert Venables. Its fourth edition forms the third part of the Universal Angler. Angling Improved to Spiritual Uses, forms part of an octavo volume entitled Occasional Reflections upon se¬ veral Subjects, by the Hon. Robert Boyle. 8vo, Lond. 1665. In a volume called The Epitome of the Art of Husbandry, by J. B. Gent. 12mo, Lond. 1669, are brief experimental directions for the right use of the angle. The author’s name was Blagrave. The Angler’s Delight; containing the whole Art of Neat and Clean Angling; wherein is taught the readiest way to take all sorts of Fish, from the Pike to the Minnow, together with their proper baits, haunts, and time of fishing for them, whether in mere, pond, or river. As also the method of fishing in Hackney River, and the names of all the best stands there; with the manner of making all sorts of good „ tackle fit for any water whatsoever. The like never be¬ fore in print. By William Gilbert, Gent. 12mo, Lond. 1676. The Compleat Troller, or the Art of Trolling, by Robert Nobbes. 8vo, Lond. 1682. There are several editions of this work, of which the third and fourth are appended to the Angler’s Pocket-Book. Gentleman’s Recreations ; treating of the Art of Horsemanship, Hunt¬ ing, Fowling, Fishing, and Agriculture. Fol. Lond. 1686. The Gentleman’s Recreation, in four parts, viz. Hunting, Hawking, Fowling, Fishing. 8vo, Lond. 1674. (By Ni¬ cholas Cox.) The Angler’s Vade Mecum, or a compen¬ dious yet full Discourse of Angling. By T. Cheetham. 8vo, Lond. 1681. Northern Memoirs, calculated for the meridian of Scotland; wherein most or all of the Cities, Citadels, Seaports, Castles, Forts, Fortresses, Rivers, and Rivulets, are compendiously described; to which is added, the Contemplative and Practical Angler, by way of diver¬ sion ; with a Narrative of that dextrous and mysterious Art experimented in England, and perfected in more remote and solitary parts of Scotland; by way of Dia¬ logue : writ in the year 1658, but not till now made publick. By Richard Franck, Philanthropus. 8vo, Lond. 1694. Of this curious volume a reprint was published of late years. The Gentleman Fisher; or the Whole Art of Angling. 8vo, Lond., second edition, 1727. The True Art of Angling, by J. S. 24to, Lond. 1696. The Compleat Fisher, or the True Art of Angling, by J. S., third edition, 1704. The preceding work, revised and corrected by W. Wright and other experienced anglers, was republished in 1740. The Compleat Fisherman; being a large and particular account of all the several ways of Fishing now practised in Europe ; by James Saunders, Esq. of Newton-Awbery, upon Trent. 12mo, Lond. 1724. The Genteel Recrea¬ tion, or the Pleasure of Angling; a Poem: with a dia¬ logue between Piscator and Corydon. By John Whitney, a lover of the Angle. 8vo, Lond. 1700. The School of Recreation, or a Guide to the most ingenious Exercises; by R. H. 8vo, Lond. 1701. The Secrets of Angling, by C. G. 12mo, Lond. 1705. The Angler’s Sure Guide, or Angling Improved and Methodically Digested, by R. H. Esq. 8vo, Lond. 1706. The Innocent Epicure, or the Art of Angling; a Poem. 8vo, Lond. 1697. The whole Art of Fishing, being a Collection and Improvement of all that has been written on this subject; with many new ex¬ periments. 12mo, Lond. 1714. The second edition of this work is entitled The Gentleman Fisher, or the whole Art of Angling. 8vo, Lond. 1727. A Discourse of Fish and Fish-Ponds-, by a Person of Honour. 8vo, London. ANGLING. The author of this work was the Hon. Roger North. A subsequent edition (of which there were more than one} bears the date of 1713. It was also published as an ap¬ pendage to the Gentleman Farmer. 8vo, Lond. 1 /26. The Country Gentleman’s Vade Mecum, by G. Jacob, Gent. 8vo, Lond. 1717; and the Compleat Sportsman, by the same author (1718), of which the 3d part relates to Fish and Fishing. England’s Interest, or the Gentleman and Farmer’s Friend, by Sir John Moore. 8vo, Lond. 1721. The Gentleman Angler, Lond. 1726. Piscatory Eclogues (by Moses Browne). 8vo, Lond. 1729. Of this work there are several editions. Sportsman’s Dictionary, or the Gentle¬ man’s Companion in all Rural Recreations. 2 vols. 8vo, 1735. The British Angler, or a Pocket Companion for Gen- tlemenFisherSjby John Williamson, Gent. 8vo,Lond. 1740. The Art of Angling, Rock and Sea Fishing, with a Natu¬ ral History of River, Pond, and Sea Fish, by R. Brookes. 8vo, Lond. 1740. Of this treatise there have been various reprints, at different periods, up to the year 1807. Ang¬ ling, a Poem. 12mo, Lond. 1741, 2d edit. The Art of Angling improved in all its Parts, especially Fly-fishing, by Richard Bowlker. 12mo, Worcester. Published some time preceding the year 1759. There is a recent edition (1806) by Charles Bowlker, Ludlow. The Angler’s Ma¬ gazine, or Necessary and Delightful Store House ; where¬ in every thing proper to be known relating to his art is digested in such a method as to assist his knowledge and practice upon bare inspection; being the completest manual ever published upon the subject, largely treating of all things relating to Fish and Fishing, and whereby the angler may acquire his experience without the help of a master. By a Lover of that innocent and healthful diversion. 12mo, Lond. 1754. The Angler’s Eight Dia¬ logues, in Verse. 8vo, Lond. 1758. The Art of Angling, eight Dialogues, in Verse. 8vo. The Universal Angler, or that art Improved in all its Parts, especially in Fly-fishing. 8vo, Lond. 1766. The Complete Sportsman, or Country Gentleman’s Recreation, by Thomas Fairfax. 8vo, London. The Complete Fisherman, or Universal Angler. 8vo, Lond. (2d edit.) Lond. 1778. The Angler’s Complete Assistant, being an Epitome of the whole Art of Angling. 4th edit. 4to, London. The True Art of Angling. 12mo, Lond. 1770. Translation of a Letter from the Hanover Magazine, No. 23, March 21, 1763; giving an account of a method to breed fish to advantage. 8vo, Lond. 1778. The Angler’s Museum, or the whole Art of Float and Fly Fishing, by Thomas Shirley. 12mo, Lond. 1784. The Fisherman, or Art of Angling made Easy, by Guiniad Charfey, Esq. 8vo, London. The North Country Ang¬ ler, or the Art of Angling as practised in the Northern Counties of England. 8vo, Lond. 1786. A Concise Trea¬ tise on the Art of Angling, by Thomas Best, Gent. 8vo, Lond. 1787. Of this work there have been published many editions, of which the 9th is dated 1810. An Essay on the Right of Angling in the River Thames, and in all the other public Navigable Rivers. 8vo, Reading. A Letter to a Proprietor of a Fishery in the River Thames; in which an attempt is made to show in whom the right of Fishing in public streams now resides. 2d edit. 8vo, Reading, 1787. The Natural History of Fishes and Ser¬ pents, by R. Brookes; to which is added, an Appendix, containing the whole Art of Float and Fly Fishing. 8vo, Lond. 1790. The Young Angler’s Pocket Companion, by Ralph Cole, Gent. 12mo, Lond. 1795. The Modern Angler, being a practical Treatise on the Art of Fishing, &c., in a series of Letters to a Friend; by Robert Salter, Esq. 12mo, London. Angling in all its branches reduced to a complete Science, in three parts, by Samuel Taylor, Gent. 8vo, London, 1800. Practical Observations on Angling in the River Trent. 8vo, Newark, 1801. Every Man his own Fisherman, by Thomas Smith. 24to, Lon- Atj don. The Driffield Angler, in two parts, by Alexander^, Mackintosh of Great Driffield, Yorkshire. 8vo, Gains¬ borough. The Angler’s Pocket Book; to which is pre- fixed, Nobbes’ celebrated Treatise on the Art of Trolling. 8vo, Norw. The New and Complete Angler, or Univer- sal Fisherman, by Richard Pollard, Esq. of Clapton, Mid¬ dlesex. 8vo, Lond. 1802. Rural Sports, by W. B. Da¬ niel. 4to, Lond. 1802. Part of vol. ii. relates to fly-fish¬ ing, and the other kinds of angling. The Kentish Ang- ler, or the Young Fisherman’s Instructor; showing the nature and properties of Fish which are angled for in Kent. 12mo, Canterb. 1804. The Complete Angler’s Vade Me¬ cum, being a perfect Code of Instruction on the above pleasing Science, &c., by Captain T. Williamson. 8vo, London, 1808. The Angler’s Manual, or concise Les¬ sons of Experience, which the proficient in the delightful recreation of Angling will not despise, and the Learners will find the advantage of practising; containing useful Instructions on every approved method of Angling, and particularly on the management of the Hand and Rodin each method. 4to, Liverpool, 1808. The Fisher s Boy, a Poem, by W. H. Ireland. 8vo, 1808. The Angler’s Manual, or concise Lessons of Experience, &c. 8vo, 1809. Practical Observations on Angling in the River Trent. 12mo, 1812. Daniel’s Rural Sports. Royal 8vo, 1812. Howitt’s Foreign Field Sports, Fisheries, &c. 4to, 1814. The Secrets of Angling, by J. D. (Davors); augmented by W. Lawson. 8vo, 1814. The Angler’s Guide, by T. F. Salter. 8vo, 1815. Art of Angling, by Charles Bowl¬ ker. 12mo, 1815. The Fly-Fisher’s Guide, by G. C. Bainbridge. 8vo, 1816. W. H. Scot’s British Field Sports. Royal 8vo, 1818. The Angler’s Vade Mecum, by W. Carroll. 12mo, 1818. Sportsman’s Repository, by J. Scott, 1820. The Troller’s Guide, a new and com¬ plete Practical Treatise on the Art of Trolling for Jack and Pike ; to which is added, the Best Method of Baiting and Laying Lines for large Eels. By T. F. Salter, author of the Angler’s Guide. Small 8vo, Lond. 1820. Instruc¬ tions to Young Sportsmen, by Lieutenant-Colonel Haw¬ ker. Royal 8vo, 1824. Salmonia, or Days of Fly-Fishing. By an Angler (the late Sir Humphry Davy). 12mo, Lond. 1828. The preceding extensive list will probably suffice for the instruction and guidance of the most studious angler. Those who are curious in regard to bibliographical de¬ tails concerning the different editions of the earlier works may consult a Catalogue of Books on Angling, 8vo, 1811, published by Mr Ellis of the British Museum, and ori¬ ginally printed in the British Bibliographer, (t.) A Explanation of Plate XLV- Fig. 1. Salmon-fly for spring. 2. Ditto ditto. 3. Ditto ditto. 4. Ditto for summer. 5. Ditto ditto. 6. Ditto for spring. 7. May fly. 8. Red-brown fly. 9. Green-drake fly. 10. Ditto. 11. Dun cut fly. 12.1 Minnow-tackle 13. J hooks, baited. 14. Moth fly. 15. Palmer fly, 16. Ditto. Jif Fig. 17. Ant fly. 18. Hawthorn fly. 19. Minnow-tackle. 20. Gorge hook andbait* ing-needle. 21. Dead-snap with four hooks. 22. Dead-snap with two hooks, baited. 23. Live bait double hook, baited. 24. Gorge hook, baited. 25. Live bait singlehook, baited accordingto two different me¬ thods. A N G to. ANGLO-CALVINISTS, a name given by some writers Ipujo the members of the church of England, as agreeing 'B pith the other Calvinists in most points except church fP’ 'overnment. AxGLO-Saxon, an appellation given to the language * % poken by the English Saxons, in contradistinction to the 11 rue Saxon, as well as to the modern English. ANGLUS, Thomas. See White, Thomas. ANGOLA, an extensive coast and territory of Western Vfrica, extending to the south oi the river Congo or 1R iaire. According to Degrandpre, the name is given to he whole coast, from Cape Lopez Gonsalvo, in 0. 44. to St f I Felipe de Benguela, in lat. 12. 14. S., a range of nearly 12 l|| agrees, or about 800 miles. The country strictly called 4 Angola, however, occupies only the middle part of this ! § pace between Congo on the north and Benguela on the ;( 5 outh, being watered by the large rivers Coanza and E)ando, to the former of which especially, the Portuguese n iaps assign a very long course through the interior; but 1,1 • either of these rivers has been ascended to any consi- •1' | erable height. From the early Portuguese accounts, it would appear hat Angola or Ngola was a title assumed by the kings, !■ nd transmitted by one to another from the original T )under of their dynasty. The first Ngola was reported tySti y tradition to have been raised to distinction by the skill if i; dth which he exercised the occupation of a smith ; and til hough this mode of elevation may be somewhat foreign to It 3 our ideas, yet this trade is in its products so important tup .3 a warlike people, and in the infancy of art is attended w it dth such difficulty, that among the nations of Africa ge- erally it is held in peculiar honour. Ngola seems also top 3 have secured the attachment of his countrymen by va- mf ious benefits, particularly by forming a store of grain, •om which, during periods of scarcity, he supplied their Kfl ecessities. Among his successors, several were fierce aiif; nd warlike; one, in particular, named Chilivagni Angola, ready extended the boundaries of the kingdom, making it p . comprehend Matamba and great part of Benguela. His uccessor Bandi Angola, however, being hard pressed 1 y the Giagas, a fierce and wandering tribe from the ill iterior, applied for aid to the king of Congo, and was n ^ ided by a body of Portuguese, who were then established in | i that territory. Through their valour and discipline ii iandi was enabled to repel the invaders, and was thus iif iduced for some time to lavish on the strangers the li lighest honours. Becoming, however, jealous of their Pp ower, and perhaps disgusted with their haughty deport- i| lent, he formed a scheme to cut them off, which they scaped only partially, and by a very hasty flight. Re- til li ning to Portugal, they recommended very strongly An- w ola as a theatre both of commerce and conquest,—an n| tlvice highly accordant with the enterprising and ambi- ous views by which that court was then actuated. Ihe river Congo had in 1484 been discovered, and in ; ime degree explored, by Diego Cam ; but his successors, i artholomew Diaz and Vasco de Gama, when sailing J long the coast to the southward, were solely intent on ' B 'aching and passing the dangerous cape which formed l ie utmost boundary of the African continent. After the ilendid discoveries and settlements attempted to be •rmed on the coast of India, the Almeidas and the Al- aquerques pushed onwards to that region without allow- *g themselves to be detained by undertaking any settle- icnts on this barbarous coast. The king, however, lout the middle of the 16th century, sent out an arma- icnt under Paulo de Diaz, a descendant of Bartholo- icvv, constituting him viceroy of all the territories which e should conquer in this part of Africa. Diaz landed, K established the Portuguese headquarters at a place A N G which he called Loando St Paul, near the mouth of the Coanza; then proceeded with an armament up that river, and erected a fort on its banks, whence his troops began to spread over the surrounding country. The king, un¬ able to brook such rivals to his power, assembled an army, which, with absurd exaggeration, is represented as exceeding 100,000 men. It was probalbly, indeed, much superior in numbers to the handful which the Portuguese could oppose to it; but such was their superiority in arms and discipline, that both the king, and his son, who soon succeeded him, were beaten in successive encounters. At length there arrived, as ambassadress at the camp of the Portuguese, no less a person than the king’s sister, Zingha Bandi, whose manners and address struck them with surprise and a sort of admiration. They relate, that on being introduced to Don John, who had succeeded to the rank of viceroy, and seeing no chair provided for her, without making any complaint, she caused her attendant to bend down on her hands and knees, and made use of her back as a seat. A demand being made of vassalage and tribute, she rejected it with indignation; and it was finally agreed that both parties should remain in their present position, and a mutual exchange of prisoners take place,—terms which leave much room to doubt if the suc¬ cesses of the Portuguese were so decided as they chose to represent. Zingha, on the whole highly pleased with her reception, remained with them a considerable time, during which she became a convert to Christianity, and finally parted on the most friendly terms. The Portuguese were not long in receiving very un¬ favourable tidings of their new convert. By the murder of her brother and his son, she paved her way to the sovereignty; and then began, equally with her predeces¬ sors, to feel deep dissatisfaction at the great extent of valuable territory possessed by this foreign nation. Un¬ able to hope for its recovery by fair means, she form¬ ed alliances with the neighbouring states, and even with the ferocious and formerly hostile tribe of Giagas; after which she commenced war, and for many years carried it on with the utmost fury, putting to death, amid the most cruel tortures, all Europeans who fell into her hands. The Portuguese boast of their numerous vic¬ tories over Zingha; yet admit that they were often very hard pressed, and had at one time their whole force cooped up and closely besieged on an island in the Coanza. They were at the same time involved in war with the Dutch, who in 1641 took their principal settlement of St Paul de Loando, which was recovered, however, by an expedition sent thither on the 15th August 1648. As Zingha reject¬ ed all terms of peace which did not include the entire restoration of the conquered country, the Portuguese en¬ deavoured to set up in opposition to her two successive members of the royal family, whom they called John I. and II.; but these phantom monarchs never enjoyed any weight with the people in general. At length the queen, having attained to an advanced age, being no longer seconded by the Dutch, and finding all her efforts to ex¬ pel the Portuguese abortive, listened to overtures of peace. She indignantly rejected any demand of homage, or of the most trifling tribute; and the treaty was concluded alto¬ gether on equal terms, except that she paid a high ransom for one of her sisters, who had fallen into the hands of the Portuguese. Permission was then obtained to send mis¬ sionaries to her coast, who persuaded her to resume the profession of Christianity, which she had renounced dur¬ ing her long period of enmity; and, on the 15th July 1662, her union with the church of Rome was celebrated with extraordinary pomp. Zingha died at the age of eighty, and the crown then devolved on her sister’s husband Mona Zingha, who com- 152 A N G Angola, menced a violent war against the Europeans, which was in a few years terminated by his death. Since that time the af¬ fairs of the Portuguese have been involved in that mystery which they studiously throw over all their colonial trans¬ actions ; but it is not believed that they now occupy more than a very limited range of territory around their ports of St Paul de Loando and St Phelipe de. Benguela, which they maintain for the purpose of carrying on the slave-trade. The whole of this coast, according to Degrandpre, to whom we are indebted for the only recent account of. it, has been singularly favoured by nature. 1 he soil consists generally of a rich black loam, fitted for yielding.in the utmost luxuriance all the productions of tropical climates. In its present uncultivated state, it is covered with dense and varied vegetation. The forests reach to the top of the hills, and their trunks are often washed by the waves. This vegetation bears generally the same character as in other parts of tropical Africa. Some of the trees, under the influence of heat and moisture on a rich soil, arrive at the most extraordinary dimensions. Mention is made of the mapou, which seems to be the same species with the calabash of Adanson, and whose trunk it has required a boat’s crew, with arms extended, to embrace. This huge production of nature is unfortunately incapable of being applied to any use whatever : the wood, moist and spongy, is fitted neither for carpentry nor even fuel; the fruit is small and unpalatable ; the foliage too scanty to afford any grateful shade. On the other hand, the family of the palms are profusely distributed, and minister to the most important uses of human life. The cocoa-palm, in particular, serves almost every economical purpose.. Its juices afford both food and drink ; its wood is hard and durable ; its filaments form strong ropes; the leaves afford a roofing which resists the rain ; even the rind of the fruit is made into vessels for domestic purposes. The date- palm exists, but the soil is too moist to produce it in abundance. The culinary vegetables of Europe, when planted, grow to an extraordinary size, but soon degene¬ rate in a soil which appears foreign to their nature. Cul¬ tivation is confined to small spots resembling gardens, in the immediate vicinity of the towns and villages, leaving all the open country in a state of nature. Its efforts are chiefly bestowed on the manioc, plantations of which sur¬ round all the villages; and the root, ground down by a species of mill, and then dried in little furnaces, affords the bread generally used in the country. The animal races are numerous, all wild, and of the same species which generally prevail throughout Africa. Elephants are less numerous than in some other quarters; but hyaenas, ounces, and tiger-cats people the forests; and there are all the various species of antelope. Apes and mon¬ keys of various and often peculiar species fill all the woods; and some are distinguished by a remarkable share of intelli¬ gence. It seems chiefly in this quarter of the continent that the orang-outang, called by the natives kimpezey, sometimes, though rarely, makes his appearance. De¬ grandpre saw one on board a slave-vessel four feet two inches high, of a mixed red and black colour, and which seemed almost to make an approach to the intellect as well as the form of humanity. It assisted the sailors in drawing the ropes, and was employed to heat the oven, giving notice to the baker exactly at the time when it was fit to receive the bread. Here also are found in great numbers those remarkable insects, the termites or white ants, with their conical structures raised three feet from the ground, and which often destroy in a short time the slight habitations of the natives, with every thing they contain. The inhabitants of this coast rank below those of the rest of the continent, in not having succeeded in A N G taming any description of domestic animals. They have W neither horses, asses, oxen, nor sheep, of which last two'J [ species such large herds are in the possession of the most ! barbarous among the other African tribes. The Portu- guese reared them in great numbers round their settle¬ ment of Loando, whence individuals occasionally strayed into the bordering districts; but no care having been taken to preserve them, they were soon either killed by the natives, or devoured by wild beasts. The mineral character of this region seems not without interest, though very little explored. Little scope is in¬ deed afforded for geological observation, the surface being everywhere covered with a deep stratum of clay, from be¬ neath which, even on the highest hills, no trace of rock is discoverable; nor have petrifactions or marine re¬ mains been anywhere observed. The metallic products, however, are supposed to be of importance, though neither examined nor turned to account. The indications of iron are very widely diffused; but the Angolans are supplied by European powers with all they require. Copper, to the utmost extent of their wants, is found on the surface of the ground. The Portuguese are said to work gold¬ mines of considerable value in the vicinity of Loando; but the nature and amount of these is studiously concealed, and perhaps exaggerated. The nations on the coast of Angola seem to rank lower in the scale of improvement than almost any other native African race, except the Hottentots. Their habitations are formed merely of straw, or rather dried leafy plants, cemented by a frame-work of wooden stakes. Containing no aperture for the admission of light, they form not so properly houses, as dark dens for sleeping in, while the ten¬ ants spend the clay and receive company in an open space in front, covered with a slight roof. The abodes of the grandees are in no respect superior, except that they con¬ sist of a number of these hovels grouped together, and in¬ closed by a hedge or earthen wall. A village or town con¬ sists merely of a cluster of these inclosures, separated by narrow and winding footpaths, and leaving extensive open spaces, which serve for markets or for scenes of recrea¬ tion. Loango, which seems to be the largest native town, though four leagues in circuit, is not supposed to contain above 15,000 inhabitants. A town in this country at a little distance resembles a wood, from the multitude of trees with which it is filled; but on a near approach its nature is soon detected by the fetid odour exhaling from its precincts. The people on this coast, like most rude nations in a tropical climate, wear scarcely any clothes, yet studiously load their persons with ornaments. Even the rich wear only a cotton cloth round the middle, hanging down to the knee ; but their legs and arms are profusely covered with rings of iron or copper, while strings of beads or coral are fastened round the neck, or hang down the breast. Peculiar pride is felt when they can procure a cast-off European suit, covered with gaudy colours and tarnished embroidery. This they display in triumph for several days, when, becoming tired, they bestow it on one of their inferiors. The natives here, like other unenlightened tribes, are deeply addicted to superstition; and it is remarked as a singular circumstance, that their idols do not present at all the negro visage, but one more nearly approaching to the European. From the slight description that is given, we should suspect the face to be Copt; nor does it seem improbable that the superstitions of Egypt may have found their way throughout the continent. The priests pretend to bestow rain, favourable winds, and various other bless¬ ings, upon those who have propitiated them by libera gifts. Much use is made in criminal cases, of what our A N G A N G 153 ancestors called “ the judgments of God.” The accused is made to swallow poison, to plunge into water, or to take in his hand burning coals, and, unless he escapes unhurt ';from these trials, is at once pronounced guilty. It seems reasonably concluded, that the priests who administer these tests contrive to secure immunity to their favourites, or to those who bestow upon them liberal donations. M. Deorandpre considers it as marvellous that such a coin¬ cidence should exist between the customs of Africa and those of Europe during the middle ages ; but both are de¬ rived from the same principle in human nature; and in¬ deed nothing can appear more probable to the uncultivat¬ ed mind, which has made little observation on the actual course of nature, than the idea that a special interposi¬ tion will take place in favour of innocence. The government of these countries seems established oh a species of feudal polity. Each towm or village has a sovereign of its own, in whose family the dignity is here¬ ditary; but a number of these princes pay homage and perform certain services to a general head of the nation. Most of the great capitals are situated at some distance in the interior. The succession is transmitted exclusively in the female line. The son of a prince has no dignity above the rest of the nation. The son of a princess alone is a prince by birth, or capable of succeeding to the crown. This is a custom not without example in rude states; and the dissolute manners prevalent and sanctioned in this country seem to afford otherwise no security that the off¬ spring will be of royal blood. The only employment carried on with any activity throughout these countries seems to be the trade in slaves. The personage next in dignity to the sovereign is the Masook, who conducts all the royal sales, levies the du¬ ties, and regulates all the transactions of the private mer¬ chants. The engagement entered into by Portugal not to practise this odious traffic to the north of the line does not extend to this coast, where it appears to be car¬ ried on still on an immense scale. The slaves imported into Brazil in the year 1828 are said to have amounted to 46,000, who, with the exception of 3000 or 4000 from Mosambique, must all have been drawn from Angola. St Paul de Loando is supposed by Malte-Brun to contain 3000 white inhabitants, besides a much greater number of negroes. An abundant supply of provisions is drawn from the surrounding country, but the water is bad. The Portuguese hold also St Phelipe de Benguela. Vessels destined for the coast of Angola, after reaching Cape Verde, have two routes by which they may proceed. 1 hey may take the short route by steering directly along the African coast, through the Gulf of Benin. If favoured by winds and currents, they may make this voyage very speedily; but in the event of these circumstances proving adverse, they are liable to great detention; and the na¬ vigation has even occupied eleven months. The other, called the long route, is performed by proceeding due south, and even south-west, till they pass the 20th degree ot latitude, when a favourable wind and tide carries them directly eastward to the African coast. This route neces¬ sarily occupies a considerable time; but it is liable to no vicissitude, and the period may be calculated almost to a day. ANGON, in the ancient military art, a kind of javelin used by the^ French. They darted it at a considerable distance. I he iron head of this weapon resembled a eur-de-luce. It is the opinion of some writers that the aims of France are not fleurs-de-luce, but the iron point ° A \ ?n^on or jave^n °f the ancient French. NGORA, Angura, or Ankora, a city of Asia in natoha, the ancient Ancyra, situated on a stream called t a una. It is large and neatly built, and its streets are vol. in. J causewayed with large blocks of granite, but have no Angot footpaths. Its position is lofty and imposing, surrounded II by mountains, with numerous gardens which yield the finest fruits. The city was celebrated in the ancient world for its fine edifices; and it still contains the remains of its former, grandeur. In its streets are to be seen pil¬ lars, some of porphyry, and jasper, and fine marbles. The city has been encompassed by a substantial wall, appa¬ rently double in some places, formed of durable stone. Some of the gates exhibit Greek inscriptions, and the masonry is intermixed with pillars, architraves, capitals, and other ancient ornaments. The vestiges of an amphitheatre are still to be seen, and the ruins of a magnificent coria or senate-house, of Corinthian architecture, with inscriptions complimentary to Augustus, in whose reign Angora was built. The inhabitants are distinguished in Natolia for their polished manners: they consist of Mahometans and Christians, there being 9000 of the latter, who are subjected to grievous tyranny and exactions from the for¬ mer. The Christians have a Greek and an Arminian archbishop, and seven churches. Although the trade and population of this place have declined, a great quantity of yarn, Angora stuffs, and shawls, are still manufactured. The shawls, which rival those of Cashmere, are fabricat¬ ed from the hair of the Angora goat, which is of a fine silken texture. The soil of the adjacent territory is en¬ tirely employed in rearing these animals; and the city is supplied with grain from a distance. This city was con¬ stituted the capital of Galatia by Nero. It was taken by the Saracens, and afterwards by Tamerlane in 1402, after the battle in which Bajazet was overthrown. The popu¬ lation, which was formerly 80,000, does not now exceed 20,000. 212 miles E. S. E. of Constantinople. Long. 33. 18. E. Lat. 40. 4. N. ANGOT, a considerable province in the southern part of the kingdom of Abyssinia, which Alvarez, who visited it in the sixteenth century, describes as an extremely rich and beautiful territory. It has not since been reach¬ ed by any European traveller, and is entirely in the pos¬ session of the Galla, who have overrun all the central provinces of Abyssinia. The tribe now in possession of Angot is called the Betzuma Galla. ANGOU, a small province in the kingdom of Congo, of which, nearly two centuries ago, it rendered itself inde¬ pendent. It extends along the northern bank of the Congo or Zaire, from its junction with the sea upwards. A great proportion of the surface is covered with forests and swamps: cultivation has made little progress, and the po¬ pulation is thin. Captain Tuckey’s expedition must have sailed along the coast of Angou, though it did not recog¬ nise it under that name, which was given by the old Por¬ tuguese writers. Bomangoi, in the interior, is stated to be the capital; but Cabenda, near the mouth of the Zaire, is the seat of trade, which consists chiefly in slaves, and was considerable previous to the prohibition of that traf¬ fic by the British government. See Congo. ANGOULEME, an arrondissement in the department of the Charente in France, extending over 556 square miles, or 355,840 acres. It is divided into nine cantons, and these again into 144 communes, with 118,871 inhabi¬ tants. The chief city, of the same name, contains 2101 houses, 15,011 inhabitants, a cathedral, an hospital, and seven churches. ANGOUMOIS, formerly a province of France, now a district, bounded on the north by Poitou, on the east by Limosin and Marche, on the south by Perigord, and on the west by Saintonge. Through this province run the rivers Touvre and Charente. This last is full of excel¬ lent fish ; and though it often overflows its banks, it is so far from doing any damage, that it greatly enriches the v 154 A N H A N H Angra II Anhalt- Bernburg. soil. The Touvre is full of trouts. The air is generally warmer than at Paris, though the country is hilly, 'live soil produces plenty of wheat, rye, oats, Spanish corn, saffron, grapes, and all sorts of fruits. Here are several iron mines, which yield a very good sort of iron. ANGRA, a city of Tercera, one of the Azores; the capital, not only of that island, but of all the group, and the residence of the governor. It is seated on the south¬ ern shore, and the harbour is the only tolerable one in the whole island. It is in the form of a crescent, the ex¬ tremities of which are defended by two high rocks, that run so far into the sea as to render the entrance narrow, and easily covered by the batteries on each side. From this harbour the town is said to derive its name, the word Angra signifying a creek, bay, or station for ships. Here ships may ride in great safety during the summer; but as soon as the winter begins, the storms are so furi¬ ous, that the only safety is in putting to sea with all pos¬ sible expedition. Happily these storms are preceded by infallible signs, with which experience has made the in¬ habitants perfectly acquainted. The town is well built and populous, and forms the see of a bishop, under the jurisdiction of the archbishop of Lisbon. It contains five parishes, a cathedral, four mo¬ nasteries, as many nunneries, besides a bishop’s court, which extends its jurisdiction over all the Azores. The fortifications are strong, but not in good repair. At Angra are kept the royal magazines for anchors, cables, sails, and other stores for the royal navy, and occasionally for merchant vessels in great distress. All maritime affairs are under the inspection of an officer, Desembergrador, who has subordinate officers and pilots for conducting ships into the harbour, or to proper water¬ ing places. The English, French, and Dutch, have each a consul residing at Angra, though the commerce of any of these nations with the Azores is very inconsiderable. Long. 27. 14. W. Lat. 38. 38. N. ANGRI, a town with 4426 inhabitants, in the province of Principato Citeriore, in the kingdom of Naples. ANGUILLIFORM, an appellation given by zoologists, not only to the different species of eels, but to other ani¬ mals resembling them in shape. ANGUINUM Ovum, a fabulous kind of egg, said to be produced by the saliva of a cluster of serpents, and pos¬ sessed of certain magical virtues. The superstition in respect to these was very prevalent among the ancient Britons, and there still remains a strong tradition of it in Wales. This wondrous egg seems to be nothing more than a bead of glass, used by the Druids as a charm to impose on the vulgar, whom they taught to believe that the possessor would be fortunate in all his attempts, and that it would gain the favour of the great. ANGUS, a district of the county of Forfar, in Scotland. It was an earldom belonging to the Douglases, now extinct. ANGUSTICLAVIA, in Raman Antiquity, a tunica embroidered with little purple studs. It was worn by the Roman knights, as the laticlavia was by the senators. ANHALT-BERNBURG, one of those small indepen¬ dent sovereignties still left in Germany, and one of the monuments of its ancient state. The sovereign bears the title of duke. His territories are much scattered between the Hartz forest and the rivers Saale and Elbe. Their whole extent is nearly 242 square miles, or 154,880 Eng¬ lish acres. It is divided into two parts, each of which is denominated a duchy. The upper duchy lies on the side and at the foot of the Hartz Mountains, and, though hilly and woody, has in it some beautiful valleys. The lower duchy, on the Elbe, is a flat marshy district, but the part which touches the Saale is dry and healthy. In the lower duchy the climate is temperate and mild> but in the upper it is cold and raw, the harvests are late, and the ^ fruits rarely come to perfect ripeness. The principal pro- I, ducts are corn, flax, rapeseed, tobacco, garden fruits, and i some wine. The country abounds with game, and the ^ lakes and rivers with fish. There are some mines, which J)' yield silver, copper, iron, sulphur, alum, and gypsum. The inhabitants in 1817 were 37,046, in 7 cities, 54 villages, and 33 hamlets, and are all of the Protestant profession. The chief employment is agriculture, the next is mining, and preparing the metals; but some little woollen cloth and paper are manufactured. The sovereign rules without limitation. The revenue of the state is about L.45,000 annually, of which L.9000 is derived from the royalties of the forests and mines. The remainder is chiefly de¬ rived from the patrimonial lands of the prince, and a part from taxes imposed on the subjects, who are mostly in poor circumstances. The military force required for the contingent is 370 men, of whom 120 are kept up, but the landsturm, a kind of militia, including every man be¬ tween 20 and 30, amounts to 7000, only 30 of whom have fire-arms. (G-) ANHALT-DESSAU, an independent sovereignty in Germany, with the title of duchy. It takes the first name from the family that governs it, and the second from the chief place in the territory. The greater part lies in a compact manner on the left bank of the Elbe, and on both sides of the Mulde. The other parts are scattered and se¬ parated from each other by the dominions of other princes. It is in general a level country, and the compact part on the left bank of the Elbe is cultivated with great care, and yields abundant produce; and though, in the scattered parts on the right bank of that river, there are some heaths and a poorer soil, yet the whole produces sufficient necessaries for its dense population. The whole duchy is divided into 15 amts or bailiwicks. Its extent is 363 square miles, or 232,320 English acres. The number of inhabitants in 1817 amounted to 52,947, living in 8 cities, 2 market- towns, and 115 villages and hamlets. The almost exclu¬ sive employment is husbandry, except that the females spin both linen and woollen yarn. All kinds of grain are raised. Flax, rapeseed and its oil, hops, madder, fruit, especially apples, wood, butter, cheese, game, and fish, are conveyed to the great markets of Berlin and Leipsic, and consti¬ tute the external trade. With the exception of 1200 Jews and two Catholic congregations, the whole of the inhabi¬ tants are Protestants, with 32 reformed and 21 Lutheran churches. The revenue of the state amounts to L.70,000 annually, of which the personal domains of the duke pro¬ duce L.20,000; the remainder arises from his feudal dues, and from some moderate taxes; It has no debt, and is the most favoured part of Germany. Though it suffered dread¬ fully by the war in the year 1813, it was enabled very speedily to recover from its losses, by the productive qualities of its soil, and the parsimonious industry of its cultivators. The whole military force is nominally 600 men, but scarcely an eighth are mustered at any one time. - yiov’S n (G-) ANHALT-KOTHEN, an independent duchy in Ger¬ many. It consists of four fragments, intermingled with the territories of surrounding princes. The most compact portion are the three bailiwicks around the capital. The duchy is on both sides of the river Elbe, though only one of its towns, Roslau, comes in contact with that stream, the others reaching it by the rivers Ziethe, the Saale, the Ruthe, or the Wipper. The land is generally a level pla®j in some parts with a sandy soil, but for the most part cultivated, especially on the left side of the Elbe, and very productive. The extent of the dominion is 324 square miles, or 140,800 acres. It is divided into nine amts or bailiwicks, and comprehends four;cities, one market-town, 1 I A N I and 106 villages and hamlets. The inhabitants in 1817 were 32,454, nearly all Protestants, the Lutherans having 19, and the reformed 28 churches. Husbandry is almost the exclusive occupation, and there is no other trade than that of exchanging its surplus productions for foreign commodities. The chief of that surplus consists of corn, wool, and fruit, especially apples, with which the duchy supplies Berlin to a great extent. The revenue amounts to about L.23,000 sterling, of which the ducal patrimony in Silesia forms L.9000. The state has a debt which in 1816 amounted to L.160,000; but, by means of a sinking fund, which was then established, it has been regularly diminishing. There are no military beyond a small body¬ guard. A militia is established, but not called out. The duke, besides this land, is also in possession of the princi¬ pality of Plesse, in the Prussian province of Silesia, and, in right of it, is a member of the states of that country, (g.) ANHOLT, a Danish island in the Cattegat, 32 miles north-east of Greenae, and 36 south-west from Lassoe. It is a barren spot, scarcely affording subsistence to 100 in¬ habitants, who from their language, which is Erse mixed with a few Danish words, are supposed to be emigrants from the Highlands of Scotland. Their chief pursuit is fishing. It is surrounded by dangerous banks of sand, for which reason a light-house is built on it, the occupation of which was a subject of contest with England during the war. It is in lat. 56. 32. 2. N. and long. 11. 49. 17. E. ANJAR, a fortified town of Hindostan, and capital of a district of the same name, in the province of Cutch, which was ceded to the British government in 1816. The country is dry and sandy, and depends entirely on irriga- A N I 155 tion by means of wells, the expense of sinking which is Anjengo great. The town is situated nearly 10 miles from the .11 gulf of Cutch. In 1816 it surrendered to the British. T^nil?aV It suffered severely from an earthquake in 1819, about 3000 houses being either destroyed or rendered uninhabi¬ table, and 165 persons having lost their lives; while the other half of the town, situated on low rocky ridges, suf¬ fered comparatively nothing. In 1820 the population was estimated at 10,000. The port named Toonea is fronted by a creek from the gulf. Long. 70. 11. E. Lat. 23. 3. N. ANIELLO. See Massaniello. ANJENGO, a small seaport town and fortress of Hin¬ dostan, in Travancore, nearly encircled by a deep and broad river, at the mouth of which it is situated. The fort was built by the English in 1684, and it was retained till 1813, when the factory was abolished on account of the useless expense attending it. Anjengo is infested with snakes, scorpions, and centipedes; those animals finding shelter in the matted leaves of the cocoa-tree, with which the houses are mostly thatched. Here and at Cochin are manufactured, of the fibres of the Lucadine cocoa-nut, the best coir cables on the Malabar coast. The exports are pepper, coarse piece goods, coir, and some drugs. ANIMAL. See Animal Kingdom. Animal-Flower, a name applied to certain species of the genus Actinia, remarkable for the brilliant colours which ornament their tentacula, and which produce an appearance resembling the variegated corolla of a plant. The name is of vicious construction, and is falling into disuse. See Zoophytes. ANIMAL KINGDOM. We embrace the earliest opportunity afforded by our alphabetical arrangement to present a few introductory observations in illustration of a science unrivalled in inte¬ rest, and not greatly surpassed in importance by any de¬ partment of human knowledge. Zoology, or the science which treats of the nature and history of animals (from <£wov, an animal, and Xoyo;, a discourse), embraces so vast a field of observation, that although it cannot be regarded other¬ wise than as a single and beautifully connected science, its great extent of general doctrine, and multiplied va¬ riety of details, render necessary a subdivision into many branches, each of which, if worthily followed, is in truth more than sufficient to occupy the undivided attention of the most zealous votary. If in voluminous works exclu¬ sively devoted to natural history a single department of the science usually engrosses the entire attention of an author, it is evident that, in the present publication, it would not only be in vain to attempt a complete exposition of the sub- iect under a single head, but that even the various treatises under which its different branches will be exhibited must I presented in a very abridged and compendious form, t shall be our endeavour, however, in the course of our natural history treatises, so to select and methodize the ?reat leading facts of the science, as to enable our readers to orm an accurate idea of the present state of zoological nowledge, even though certain minuter details, hitherto orming too conspicuous a feature of the subject, yet not essential to its truthful representation, should in some measure be curtailed. With a view to obviate to a certain extent the inconve¬ nience arising from the disconnection of the different parts of wkich the nature of an alphabetic encyclopaedia zlm,6880/uvier Rhisostoma many more; to say nothing of the in- and convey the air to the interior of their bodies. “ No usorial tribes, most of which have no mouth at all, but animals,” says Cuvier, “respire by a particular organ, except lerive their nourishment by imbibition through the me- such as have a real circulation; because in them the blood lium of external pores. Nutrition, or the power of de- coming from one common source, the heart, to which it iving nourishment from other bodies, is common alike to constantly returns, the vessels that contain it are so dis- •lants and animals, and effects for organized and living posed that it cannot arrive at the other parts until it has >odies that increase of bulk which inorganic or disorgan- passed through the lungs. This, however, cannot take place zed substances can only attain through the medium of an in vegetables, or in those animals in which this fluid is dfinity of particles, or by mechanical aggregation. The everywhere diffused in a uniform manner, without being unctions of nutrition, however, as manifested in the ani- contained in vessels.” Pulmonary or branchial respira- nal and vegetable kingdoms, are very differently perform- tion is therefore an animal function of a third order, in- •d in each, fixed for ever to the soil which gave them variably connected with circulation, and one degree re- >11 th, plants are rendered incapable of* searching after moved from such faculties as are essential to animal life, lounshment by a voluntary change of place, but derive When a vegetable dies of old age, it begins to decay in len chief support from roots, the pores of which absorb the centre. We frequently see ancient willow-trees en- ic nutritive portions from the humid soil, and by a uni- tirely dead, except in a few slender twigs, or in small por- orin and continuous action, which is only interrupted by tions of their superficial bark. An animal, on the con- m absence of the necessary moisture. The generality of trary, first dies in its extremities and circumference, whilst aninals, on the contrary, being possessed of the power of the heart or central portion continues for a time to per- ocomotion, are also endowed with the capacity of trans- form, however languidly, its accustomed actions. xji ting with themselves a supply of necessary nourish- Among plants both sexes usually occur in the same iitnt, for which purpose they are provided with an in- individual, or even on the same flower; but in a far ciua cavity or stomach, the inward surfaces of which greater proportion of animals the two sexes are repre- ire provided with absorbing pores, which Boerhaave ex- sented by separate individuals. There is indeed no •ressive y named internal roots. “ The magnitude of this genuine hermaphrodital union among mammiferous ani- the facts of nature. “ It is certain by revelation,” says Buffon, “ that all creatures have equally participated in the favours of creation; that the first two of each species were formed by the hands of the Creator; and we ought to believe that they were then nearly such as they appear at present in their descendants. We must also consider, that although nature proceeds by gradual, and frequently by imperceptible degrees, the intervals are not always the same. The more exalted the species, the fewer they are in number; and the shades by which they are separated are more conspicuous. The smaller species, on the con¬ trary, are very numerous, and have more affinity to each other, so that we are the more tempted to confound them together in the same family. But we should not forget that these families are our own works; that we have made them for the ease of our memories; and that if we cannot comprehend the real relations of all beings, it is ourselves, not nature, that are in fault; for she in truth knows not our pretended families, and recognises indivi¬ duals alone.” Omnipotence, the first, the greatest, and indeed the only truly creative powder, formed the species of animals ; and the influence of man and of physical agents has pro¬ duced the varieties. But it is only superficial characters which either the one or the other of these ulterior causes has the power of modifying. The basis of organization, or real specific mould, remains unalterable, though a thousand circumstances constantly tend to produce varia¬ tions in the external forms. Of these circumstances the most powerful is no doubt climate; under which name it is necessary to comprehend the differences of local situa¬ tion and temperature, the nature of the soil and its pro¬ ductions. It is climate, in the first place, which chiefly de¬ termines the geographical position of animals, and thus commences the action of the modifying powers. The na¬ ture of their food is also highly influential; and as it de¬ pends so immediately on the qualities of temperature and soil, the climate is still the modifying cause. If the same animals usually accompany the same vegetables, it is be¬ cause the constitution of both demands similar influences, ' and because through each other they are both dependent upon the same support. Certain animals are leagued with certain plants, and these again with certain soils and cli¬ mates ; and a careful observance of these mutual depend¬ encies exhibits one of the finest and most beautiful har¬ monies of nature. This, howrever, is not the place in which to discuss the intricate and important subject of the geographical distribution of animals. Certain original forms have been continued since the creation ot organized beings, and all the individuals which represent or belong to one of these forms constitute what is called a species. The slighter differences which occa¬ sionally prevail among the individuals themselves, inde- pendent ot the customary distinctions of age and sex, are railed varieties. Such varieties are seldom permanent, nid are usually lost by the progeny re-assuming the ordi- mry and characteristic form or colour, except in some emarkable instances, such as the horse, dog, and other ong-domesticated species, of which man has so thorough- y altered the original condition, as to have impressed lem with a second and more pliant nature. An indivi- ' I ’ according to Buffon, is a separate detached being, i . nothing in common with other beings, excepting . )at it resembles, or rather differs from them. All simi- ai individuals which exist on the earth are considered as 161 composing the species of those individuals. Yet it is nei- Animal ther the number nor collection of similar individuals which Kingdom, constitutes them ; for a being which existed for ever would not be a species. “ Species then is an abstract and ge¬ neral term, the meaning of which can only be determined by considering nature in the succession of time, and in the constant destruction and renewal of beings;” and it is by comparing the present state of nature with the past, and actual individuals with their predecessors in kind, that we come to attain a clear idea of what is called spe¬ cies ; for a comparison of the number or resemblance of individuals is only an accessory idea, and frequently inde¬ pendent of the first. The ass resembles the horse more than the barbet the greyhound; yet the latter are but one species, since they produce a fertile progeny; but the horse and the ass are certainly of different species, “ since they produce together vicious and unfertile young.” It is indeed difficult to define the term species, other¬ wise than as an assemblage of individuals descended from common parents, which bear as great a resemblance to them as they do to each other. Species then are distin¬ guished by fixed forms, which, though to a certain extent alterable, and for a limited time, by external or accidental causes, are yet handed down unimpaired from generation to generation ; and although certain species seem to have disappeared entirely from the earth, in consequence of the great natural catastrophes which have taken place in an¬ cient times, and the local distribution of many still exist¬ ing races has been modified or changed by the influence of man no less than by the accidents of nature, there is no reason to believe that any one species has sprung from the gradual alteration of another, or that the circumstances under which an individual may have been at first casually placed were sufficient to develope both form and function, without an impress from a higher and more powerful hand, by which it was fitted to perform its part (pre-ordained) in the great theatre of the world. Animals which by their union produce^/erft’/e individuals, are generally reputed of the same species. This law of nature, as it was formerly called, having been found to admit of certain though rare exceptions, is not now so broadly insisted on as a test of specific identity as it was in preceding times. But it appears, from the result of nu¬ merous experiments, that the generality of animals pro¬ duced from a cross between even the most nearly allied species, are either altogether incapable of reproduction, or fertile in so imperfect a degree, that their descendants speedily become entirely sterile. It has been said that birds alone were unsubjected to this rule, and that hence has arisen the wonderful variety which that beautiful class exhibits. There is no doubt of the occasional fertility of their hybrids, as in the case of those mule birds produced between the goldfinch and canary ; but as it has not been proved that such unions of distinct kinds ever take place when uncontrolled by the depraving influence of domesti¬ cation, there is no reason to attribute the origin of any of those species or varieties which are known to exist in a wild state to any such improbable alliance. It is known that a productive union may take place be¬ tween animals of a different species, provided such species belong to what naturalists call the same natural family. Thus the ass and the mare, or the horse and the female ass, produce the well-known animals called mules: the zebra also produces both with the horse and the ass; but in order to deceive the female zebra, it is said to be neces¬ sary to paint the hides of the former with those bizarre colours which adorn her accustomed mate.1 It is probable, VOL. Ml. 1 Diet. Class. d'Hist. Nat. tome i. p. Hi. X ANIMAL KINGDOM. 162 Animal however, that all these unions are so far forced and unnatu- Kingdom. ral, that they never take place except through the influence of man, when domestication, and the numerous changes consequent upon it, have altered or impaired their natui'al instincts; for it has been observed that, however educa¬ tion may perfect certain special qualities, which man has the art to render subservient to his own convenience, yet a more extended view of the effects of domestication will convince us, that it is almost always to the disadvantage of their natural capacities that the brute creation are made to borrow the mask of human intelligence. Buffon appears to have adopted from Ray a rule which many now regard as inaccurate and artificial, but which he made use of to determine the identity of animal species, viz. “ any two animals that can procreate together, and whose issue can also procreate, are specifically the same.” In this, however, it has been observed that he contradicts himself by afterwards admitting that the sheep and goat are of difl'erent species, at the same time that he asserts that the he-goat and the ewe produce a mixed breed which continue fertile for ever. Dr John Hunter (a great authority) was also of opinion that the true distinction between different species of animals must ultimately be gathered from their incapacity of propagating with each other an offspring capable of again continuing the kind. Thus the horse and ass beget a mule capable of copulation, but incapable of begetting or producing offspring. The acci¬ dent of a mule breeding, according to the same authority, even if it were proved, would only show that as many per¬ fect animals of true species and distinct sexes are incapable of breeding at all,—(thus showing that nature, even in her greatest perfection, sometimes deviates from general prin- . ciples),—so it may occasionally happen that a mule shall breed from the circumstance of its being “ a monster re¬ specting mules.” The doctrine of equivocal generation has received no support from any recent investigations. All that is known decidedly leads to the opposite conclusion ; and if certain mysterious or unaccountable phenomena have perplexed the physiologist, the only legitimate deduction is, that he has met with something which he cannot comprehend; for those aberrations (if such they really are) from the usual laws of nature are not so much exceptions to the general rule, as additional instances of effects in nature, the re¬ gulating causes of which we are as yet incapable of de¬ monstrating. The rules of philosophizing lead us to reject the admission of more causes than are sufficient for the explanation of phenomena ; and if, for example, mites, and “ such small deer,” derived their origin solely from the caseous and other substances in which they are generally found, the sexual distinctions which prevail among other animals would in them be unnecessary, and would not therefore be observable. But we know that distinct sexes do exist among these minute creatures—that they propa¬ gate their kinds after the accustomed mode—and we hence fairly infer that fortuitous generation does not take place among them. May we not therefore conclude that the origin of those first observed is similar to that of the thousands which we afterwards see 'produced according to the usual process ? Nature does nothing in vain, and it is not consonant with her usual practice to suppose that she would authorize two distinct modes of creation in the same animal, where one of these is evidently perfect and self- sufficing. It may therefore be laid down as a general rule, that all living beings proceed from others of a similar na¬ ture, either by generation, offset, or some other means; and that, in all instances of apparently spontaneous pro¬ duction, there has existed some minute or imperceptible a germs, which deceive us by their sudden development Kin ,1 when placed under favourable circumstances. “ In fact,”^ says Cuvier, “ however feeble and minute the parts of an /f embryo or the seed of a plant may be at the moment we are first capable of perceiving them, they then enjoy a real life, and possess the germ of all the phenomena which that life may afterwards develope. These observations, extended to all the classes of living bodies, lead to this ge¬ neral fact, that there are none of those bodies which have not heretofore formed part of a body similar to itself, from which it has been detached.” “ It is from them (their parents) they have received the vital impulse ; and hence it is evident that, in the actual state of things, life pro¬ ceeds only from life, and that there exists no other, except that which has been transmitted from one living body to another by an uninterrupted succession.” “ Origin by ' generation, growth by nutrition, termination by death, are the general and common characteristics of all organized beings.1” At the same time it must be admitted, that the origin of many infusory and intestinal animals is sufficiently ob¬ scure. Of the latter class, one of the most extraordinary is a monstrous worm, as it may be called, which at distant intervals, and in parts of the world far removed from each other, has been found to inhabit the liver of the human race. The means by which it is bred, or the circumstances favourable to its production, are quite unknown; and some of the most philosophical inquirers of the present day have been unable to account for its origin, otherwise than by supposing that the viscus called the liver becomes, under certain circumstances, endowed with the power of actually secreting a substance capable of assuming and presenting the phenomena of distinct animal existence, and which, prior to the period of its being observed, had in fact become a specific animal;—at least its existence, unlike that of mites, flies, &c. which so often misled the ancient naturalists (though “in limo non ex limo” is Ovid’s more accurate expression), cannot be accounted for in any other way. “ There are,” Mr W. S. Macleay observes, in his excellent and ingenious Horce Entomologies, “ many circumstances which might be adduced to support the belief that, whether from disease or other causes, there are periods when other parts of a body besides the ova¬ rium may produce living germs, and demonstrate thus the polype nature of the cellular substance.” A polypus has been sometimes described as an anima- lized tube, capable of digestion, and possessed of a certain power of motion and reproduction; and these few words may be said to contain almost all that we know of its es¬ sential nature and attributes. As we advance in the scale of creation, we find a more complicated system of organs, with more varied powers of action, and a higher develop¬ ment of those accommodating instincts which, though circumscribed within certain impassable boundaries, yet seem at times to form such an approach to reason, as to connect the unvarying mechanical actions of the most simple zoophytical tribes with the conscious self-regulat¬ ing power which has its final and most perfect develop¬ ment in the human race. The excellence of man, how¬ ever, physically considered, consists more in the balance of various powers than in any one bodily superiority; for there is in fact no single sense in which he is not excelled by one or other of the brute creation. Materialists, who regard the intellectual superiority of man as the result ot hisphysical structure, must also, for the sake of consistency, maintain his excellence as a machine to be infinitely he‘ 1 Lectures on Qomp. Amt. voi. i. ANIMAL KINGDOM. 163 U1 ore that of every other’animal. But as his indubitable m superiority depends on something totally immaterial, which ,< tiirows him out of the group of animals, and makes him in insulated being,” there is the less necessity for either laturalists or metaphysical inquirers endeavouring to de- nonstrate the superexcellence of his physical condition. Vlan is in all cases “ wonderfully,” and in many cases < fearfully made;” but in the powers of sight, touch, hear- iv, and smell, he is greatly surpassed by many animals ; aid even in the sense of taste, in which his practice is nore varied and extensive, he is at least equalled by the Teat majority. It is indeed somewhat remarkable that lie last-named sense, the only one in regard to which the mman race can lay claim to the possession of equal excel- ence with the brutes that perish, should be the grossest md least intellectual of the whole. Although it is by no means strictly true that the brain, ■onsidered in relation to the size of the body, decreases as ve descend from man to the lower tribes, because in many mail birds, such as sparrows and finches, the brain is re- atively much larger than in the human race, yet, exclu¬ de of some such exceptions, it may be asserted that that Important organ becomes less as we descend from mam- niferous animals and birds, to reptiles, fishes, and other ower forms. Yet no very accurate conclusion can be de- luced regarding the degrees of intelligence in different inimals, from the proportion which the quantity of brain iears to the mass of the whole body, particularly when we :ome to compare animals of the same class with each ither. It is in fact by no means easy to ascertain with :ccuracy what that proportion really is, because the weight if the brain is supposed to remain the same, while that if the animal varies according to its temporary condition, n this way only can we account for the contradictions rhich appear in the tabular views which have been given if these proportions by physiologists. Upon the whole, he smaller animals pertaining to the higher classes ap- iear to have proportionally the largest brain, though man s in this respect seldom surpassed. The proportional veight of the brain to the body of man varies from l-22d o l-35th part, that of the gibbon or long-armed ape is is 1 to 40, that of the young malbrouk (Simia Faunus) as l to 24, that of the fox as 1 to 205, that of the beaver as 1 to 290, that of the field-mouse as 1 to 31, that of the dephant as 1 to 500, that of the horse as 1 to 400, that >f the eagle as 1 to 160, that of the sparrow as 1 to 25, hat of the canary-bird as 1 to 14, that of the cock as 1 o 25, that of the goose as 1 to 360, that of the land- ortoise as 1 to 2240, and that of the sea-tortoise as 1 to >688. Soemmering, Ebell, Vicq-d’Azir, Gall, and Tiedemann, upposed that every thing depended on the volume of the irain. But as Button and Daubenton had proved that the sapajous have the brain proportionally larger than man, without surpassing their congeners in intelligence, it has >een maintained by others that the volume alone was not i condition of superiority. Now the Sapajous in question lave no convolutions to the brain, so that the surface of hat organ is represented by that of the interior of the iramum, and exceeds it in other cases in proportion as its olds are numerous and profound; and as there appears o be a constant relation among mammiferous animals be- ween the diminution of the cerebral surfaces and their in¬ effectual degradation, whilst no such relation can be traced ictween the degrees of degradation and the variations of he brain in respect to size, it has hence been inferred by >ome that the extent of surface, and not the volume of the brain, ought to be regarded as co-relative with the in- Animal tellectual faculties.1 Kingdom. In proportion, however, as the superior portion called the brain decreases in size, the medullary matter appears to collect in other parts of the body, or in the cords which emanate from the brain; so that many animals with much smaller brains have nerves more voluminous in pro¬ portion to their bodies than those of man. This medul¬ lary substance, the medium of sensation^is, in the human race especially, collected into one principal mass as the engine of thought and reflection, the intellectual attributes by which man is characterized ; but it becomes dispersed in the inferior animals, or ramified over the whole body in the form of ganglions or nervous chords, without any preponderating superior brain. It is owing to this dis¬ persion of the nervous system into these small separate centres in the polypus and other tribes, that almost every portion of the body, wdien separated from the rest, is cap¬ able of becoming a distinct animal, and of assuming an in¬ dependent existence. In the lowest tribes of all, in which the nervous system has not yet been demonstrated, it pro¬ bably consists of molecules of inconceivable minuteness, disseminated through the pulpy or gelatinous masses of which the bodies of many radiated and infusory animals are composed. Singular effects result from the dispersion of the brain into so many small and separate centres; and this class of phenomena also illustrates the analogy wrhich exists be¬ tween the lower animals and the vegetable world. Among the superior creatures no reproduction takes place except of the fluids, and of whatever partakes of the nature of the epidermis. Injury is repaired and superficial parts renewed, but nothing resembling regeneration of impor¬ tant organs ever takes place. But it is otherwise with the inferior orders. The tentacula of the polypus and of many molluscous animals, the rays of the star-fish, the external members of the salamander, and the entire head, with the eyes and antennae of the snail, when cut off, are speedily renewed. There are also animals, such as the planaria, which reproduce by offsets after the manner of plants; and a polypus may be divided into many portions, each of which becomes perfect according to its kind;—thus in a manner realizing what the ancient poets have feigned re¬ garding the hydi’a of the Lernean marshes. If the head of a mammiferous quadruped, or of a bird, is cut off, the consequences are of course fatal. But the most dreadful wounds that imagination can figure or cruel¬ ty inflict have scarcely any destructive influence on the vital functions of many of the inferior creatures. Riboud stuck different beetles through with pins, and cut and la¬ cerated others in the severest manner, without greatly ac¬ celerating death. Leeuwenhoeck had a mite which lived eleven weeks transfixed on a point for microscopical in¬ vestigation. Vaillant caught a locust at the Cape of Good Hope, and after excavating the intestines, he filled the abdomen with cotton, and stuck a stout pin through the thorax, yet the feet and antennae were in full play after the lapse of five months. In the beginning of November, Redi opened the scull of a land-tortoise, and removed the entire brain. A fleshy integument was observed to form over the opening, and the animal lived for six months. Spal¬ lanzani cut the heart out of three newts, which immedi¬ ately took to flight, leapt, swam, and executed their usual functions for 48 hours. M. Virey informs us, “ Nous avons vu une salamandre vivant depuis deux mois, quoique decapitee au moyen d’une ligature serree du cou.” A de¬ capitated beetle will advance over a table, and recognise 1 See Desmoulins, Rcch. Anat. ct Phys., and the Journ. Compt. du Diet, des Sciences Med. Septembre 1822. 164 ANIMAL KINGDOM. Animal a precipice on approaching the edge. Redi cut off the Kingdom, head of a tortoise, which survived 18 days. Colonel Pringle decapitated several libellulse or dragonflies, one of which afterwards lived for four months, and another for six ; and, which seems rather odd, he could never keep alive those with their heads on above a few days.1 Some curious particulars connected with the great te¬ nacity of life in the lower animals are mentioned by Mr Fothergill.2 A friend being employed one day in the pur¬ suit of insects, caught a large yellow dragonfly (Libellula varia), and had actually fastened it down in his insect box, by thrusting a pin through the thorax, before he perceiv¬ ed that the voracious creature held a small fly, which still struggled for liberty, in its jaws. The dragonfly continu¬ ed devouring its victim with great deliberation, and with¬ out expressing either pain or constraint, and seemed total¬ ly unconscious of being pinned down to the cork, till its prey was devoured, after which it made several desperate efforts to regain its liberty. A common flesh-fly was then presented to it, when it immediately became quiet, and ate the fly with greediness: when its repast was over it renewed its efforts to escape. This fact being mentioned to Mr Haworth, the well-known English entomologist, he confirmed the truth of the remarkable insensibility to pain manifested by insects, by narrating an additional cir¬ cumstance. Being in a garden with a friend who firmly believed in the delicate susceptibility of these creatures, he struck down a large dragonfly, and in so doing unfor¬ tunately severed its long abdomen from the rest of the body. He caught a small fly, which he presented to the mutilated insect, by which it was instantly seized and de¬ voured ; and a second was treated in the same manner. Mr Haworth then contrived to form a false abdomen, by means of a slender portion of a geranium; and after this operation was performed the dragonfly devoured another small insect as greedily as before. When set at liberty, it flew away with as much apparent glee as if it had received no injury. It is a fact well known to practical entomolo¬ gists, that lai'ge moths found asleep during the daytime may be pinned to the trunks of trees without their appear¬ ing to suffer such a degree of pain as even to awake them. It is only on the approach of the evening twilight that they seek to free themselves from what they must no doubt regard as an inconvenient situation. The cruelty of zoological, especially of entomological pursuits, has too often been stated as an objection to the practical parts of the study of natural history. When a noble aristocrat (who thinks it sport to shoot a shep¬ herd’s dog) slaughters 100 brace of grouse in a single day, we hear nothing of such an objection, possibly because the flavour of moor-game is very exquisite ; and the reason of defence is good. But the tastes of men differ, and fortunately, as all have not the means of an equal gratifi¬ cation from the same source. “ Cruelty,” say Messrs Kirby and Spence, “ is an unnecessary infliction of suffering, when a person is fond of torturing or destroying God’s creatures from mere wantonness, with no useful end in view; or when, if their death be useful and lawful, he has recourse to circuitous modes of killing them, where direct ones would answer equally well. This is cruelty, and this with you we abominate; but not the infliction of death when a just occasion calls for it. They who see no cruelty in the sports of the field, as they are called, can never, of course, consistently allege such a charge against the entomologist; the tortures of wounded birds, of fish that swallow the hook and break the line, or of the hunted hare, being beyond comparison greater than those of in¬ sects destroyed in the usual mode. With respect to uti- lity, the sportsman, who, though he adds indeed to the general stock of food, makes amusement his primary ob. ject, must surely yield the palm to the entomologist, who adds to the general stock of mental food, often supplies hints for useful improvement in the arts and sciences, and the objects of whose pursuit, unlike that of the for! mer, are preserved, and may be applied to use for many years. But in the view of those even who think inhuma¬ nity chargeable upon the sportsman, it will be easy to place considerations which may secure the entomologist from such reproof. It is well known, that in proportion as we descend in the scale of being, the sensibility of the objects that constitute it diminishes. The tortoise walks about after losing its head ; and the polypus, so far from being injured by the application of the knife, thereby ac¬ quires an extension of existence. Insensibility almost equally great may be found in the insect world. This, indeed, might be inferred a priori, since providence seems to have been more prodigal of insect life than of that of any other order of creatures, animalcula perhaps alone excepted. No part of the creation is exposed to the at¬ tack of so many enemies, or subject to so many disasters; so that the few individuals of each kind which enrich the valued museum of the entomologist, many of which are dearer to him than gold or gems, are snatched from the ravenous maw of some bird or fish, or rapacious insect, would have been driven by the winds into the waters and drowned, or trodden under foot by man or beasts; for it is not easy in some parts of the year to set foot to the ground without crushing these minute animals ; and thus also, instead of being buried in oblivion, they have a kind of immortality conferred upon them. Can it be believed that the beneficent Creator, whose tender mercies are over all his works, would expose these helpless beings to such innumerable enemies and injuries, wTere they endued with the same sense of pain and irritability of nerve with the higher orders of animals?”3 Instead, therefore, of believing, and being grieved by the belief, that the insect we tread upon, In corporal sufferance finds a pang as great As when a giant dies, the very converse is nearer the truth. “ Had a giant lost an arm or a leg,” continue the authors just quoted, “or were a sword or spear run through his body, he would feel no great inclination for running about, dancing, or eating. Yet a tipula will leave half its legs in the hands of an unlucky boy who has endeavoured to catch it, and will fly here and there with as much agility and uncon¬ cern as if nothing had happened to it; and an insect im¬ paled upon a pin will often devour its prey with as much avidity as when at liberty. Were a giant eviscerated, his body divided in the middle, or his head cut off, it would be all over with him ; he would move no more ; he would be dead to the calls of hunger, or the emotions of fear, anger, or love. Not so our insects : I have seen the com¬ mon cockchafer walk about with apparent indifference after some bird had nearly emptied its body of its vis¬ cera ; a humble bee will eat honey with greediness though deprived of its abdomen; and I myself lately saw an ant, which had been brought out of the nest by its comrades, walk when deprived of its head. The head of a wasp will attempt to bite after it is separated from the rest of the body ; and the abdomen, under similar circumstances, if the finger be moved to it, will attempt to sting.” Query, which part felt conscious of being the original wasp? That the acuteness of bodily suffering, even among the 1 See the observations prefixed to the translation of Sj/allanzani's Tracts, by John Graham Dalyell, Esq. * Essay on Natural History. a Introduction to Entomology, vol. L p. 56. ANIMAL KINGDOM. i Ml lirfier classes of the brute creation, is in some manner f En. .rovidentially subdued, and rendered so much less acute pkL,.s not to be a fit subject of comparison with the suffering ,f the human race, is indeed evident from various pheno- aena, whatever the cause may be. The writer of this rticle has seen a turtle-dove (Columba risoria) which ^as so severely lacerated by a cat, that the contents of Ml ts stomach were torn out. The painfully excited sym- I ,athy of those who had long cherished the gentle crea- f ure was, however, in a great measure allayed by see- ' il ng the bird immediately afterwards proceed to pick up lj5 he fresh grains of barley which (till the aid of the sur- | eon was called in) continued to fall from its wounded I aunch. , , . . , Considerations of the nature glanced at in the preced- i >i .ig paragraphs can never, of course, be so misconstrued i sDto afford any palliation to wanton or inconsiderate M| ruelty to the brute creation. The judges of the Areo- [ iagus who condemned to death the child whose amuse- i tient it had been to pluck out the eyes of quails, were i egulated in their determination by the motives imput- ( d to the young criminal, and which they deemed ex- [ iressive of so cruel and pernicious a character, that in «5f, fter-times he would assuredly offend the state. “ Nec i nihi videntur Areopagitae,” says Quintilian, “ cum dam- taverunt puerum coturnicum oculos eruentem, aliud ju- r lecessity of having recourse to a somewhat similar opera- ion, the case would have assumed another character, and hi he most sentimental philanthropist must have applauded i W he practice of the philosopher. So it is in a great mea- il are with the pursuits of the naturalist. If the wonderful tructure of the lower orders of creation cannot be studied ir understood, or their infinitely varied forms held in re- nembrance, without hastening by a few days or hours the ennination of that brief career which in truth scarcely wer meets with a strictly natural end, then is the student >f nature, following out the principles of an elevating and ntellectual pursuit, as well entitled to command a por- ion of animal life as he who, to pamper the refined gross- less of a sensual appetite, bleeds his turkeys to death )y cutting the roots of their tongues, boils crabs and obsters alive, and swallows unsuspecting oysters by the >core. The more perfect the nervous system, the greater is the legree of intelligence. Indeed, were it not that no trace >f that system has yet been discovered in many zoophytes, ve might almost assert that the presence of nerves con¬ stitutes animal life, and that their absence in organized natter reduces it to the vegetable state. The greater ffie extent of brain in proportion to the size of the body, tlie greater in general the degree of sensibility. A French matomist, in dissecting a horse of which he had admired the noble qualities, exclaimed, “ J’ai long temps doute si nous avions le droit de monter sur ton dos ; mais en voyant la petite capacite de ton cerveau, je n’en doute plus main- tenant ; tu n’est qu’une bete.” The most perfect animals ire such as are provided with a head which serves as the centre of their sentiments and sensations, and with a mouth for the reception of their nourishment. Their forms lire symmetrical, or composed of two equal parts; they 165 change their place by a voluntary act; their sexes are Animal distinct, and separately incapable of continuing the spe- Kingdom, cies; they are provided with five senses, and endowed with the perception of pleasure and of pain. The inferior tribes, on the contrary, which present so many analogies to the vegetable kingdom, have no distinct head, or single organ of life; they are not symmetrical or composed ex¬ actly of two equal parts, but rather affect the circular or radiated form; and for the most part they either remain fixed to the place which gave them birth, or with difficul¬ ty change their places of abode. The sexes are frequent¬ ly united in the same individual, and their senses are limited to such as are necessary to a very confined and almost vegetative existence. Though no animal has more than five senses, a great many are much more sparingly endowed. The only uni¬ versal sense seems that of touch. The next to be deve¬ loped is probably that of taste, then sight, hearing, and, lastly, smell. In the human race the senses are more equally balanced than in the brute creation, among the different tribes of which we find many animals as remark¬ able for their extreme acuteness in certain senses, as for their obtuseness in respect to others. The sense of smell in the dog, excepting some artificial varieties, such as the greyhound, prevails over every other; birds of prey are re¬ markable for their keenness of sight; the sense of hearing is strong in the hare; that of touch in the trunk of the elephant; that of taste in the lord of the creation. It follows as a consequence that the dog is by nature a hunt¬ ing animal; that the eagle, upborne upon resplendent wing, describes its magnificent circles in the air, “ saga¬ cious of its quarry from afar;” that the hare couches se¬ curely among the long dewy grass, with its head so low that its eyes must be almost useless, but trusting to its quicker ears, which warn it of an approaching foe; that the elephant examines the exact nature of all objects by touching them with the fleshy finger of its proboscis; and that Mrs Rundell’s work on cookery has run through nine¬ teen editions. All insects in the perfect state, and the greater proportion of their larvae, a part only of the mollus¬ cous tribes (such as the inhabitants of univalve shells), crustaceous animals, such as crabs and lobsters, and all fishes, reptiles, birds, and quadrupeds, enjoy the sense of sight; and all these classes (with the exception of in¬ sects and many of the mollusca) are also furnished with the organs of hearing. That the latter sense, however, also exists in insects, may be fairly inferred from the fre¬ quent and varied sounds which they are capable of pro¬ ducing, although the seat of the faculty has not yet been ascertained. Many zoophytical tribes, which have no special organs of sight, appear to become sensible to the presence and action of light, through a delicate perception of the sense of touch. According to Buffon, the sense which has the strongest affinity to thought is that of touch ; and he regards it as being enjoyed by man in greater per¬ fection than by animals. That which has the strongest affinity to instinct and appetite is smell,—a sense in which man must acknowledge an infinite inferiority. Hence, according to the Frenchman, man has the greatest ten¬ dency to knowledge, and the brute to appetite. There is no doubt that in man and the different species of monkeys the sense of touch is highly discriminating; but it is as¬ suredly a false view of the subject which has led Helve- tius and others to attach such an extraordinary degree of importance to the hand, as the medium of intellectual su¬ periority in the human race. Whatever exhibits the phenomena of either animal or 1 Dc Inst. Of at. lib. v. cap. ix. de Signis. 166 ANIMAL KINGDOM. Animal vegetable life, advances towards the perfect development Kingdom. 0f its parts through the medium of aliment. This name appiie(i to the numerous and diversified substances which, when introduced into the system of an organized body, have the power of identifying themselves in part with that system, in such a manner as to effect its nourishment, reparation, and increase. “ Nourishment,” says Bacon, “ ought to bee of an inferior nature and more simple sub¬ stance than the thing nourished. Plants are nourished with the earth and water, living creatures with plants, man with living creatures. There are also certain creatures feeding upon flesh ; and man himself takes plants into a part of his nourishment: but man and creatures feeding upon flesh are scarcely nourished with plants alone. Perhaps fruits or graines, baked or boyled, may with long use nourish them, but leaves of plants or herbs will not doe it; as the order of the Foliatanes showed by experience living creatures are nourished by the mouth; plants by the root; young ones in the womb by the navill; birds for a while are nourished with the yolke in the egge, whereof some is found in their crops after they are hatched. All nourishment moveth from the centre to the circumference, or from the inward to the outward: yet it is to be noted, that in trees and plants the nourishment passeth rather by the barke and outward parts than by the pith and in¬ ward parts ; for if the barke be pilled off, though but for a small bredth round, they live no more : and the blood in the veines of living creatures doth no lesse nourish the flesh beneath it than the flesh above it. Vegetables as¬ similate their nourishment simply without excerning; for gums and teares of trees are rather exuberances than ex¬ crements ; and knots or knobs are nothing but diseases. But the substance of living creatures is more perceptible of the like; and therefore it is conjoyned with a kinde of disdaine, whereby it rejecteth the bad and assimilateth the good. It is a strange thing of the stalkes of fruits, that all the nourishment which produceth sometimes such great fruits should be forced to passe thorow so narrow necks, for the fruit is never joyn’d to the stock without some stalkes. It is to be noted, that the seeds of living creatures will not be fruitful but when they are new; but the seeds of plants will be fruitful a long time after they are gathered ; yet the slips or cions of trees will not grow unlesse they be grafted green, neither will the roots keepe long fresh unlesse they be covered with earth.”1 Nutritive substances of course vary according to the nature of the bodies which consume or absorb them. Plants derive their chief nourishment from air and water, the former of which must contain carbonic acid gas, the latter the dissolved remains of animal or vegetable sub¬ stances. It is, however, possible to produce vegetable growth from pure wTater, assisted by warmth and air. Vegetables, again, serve as food to the greater proportion of animals, and these in their turn are devoured by the carnivorous few. It is thus that the productions of na¬ ture are connected together in one great circle, and are reciprocally dependent on each other. Without water there could be neither plants nor herbivorous animals, and without herbivorous animals there could be no carni¬ vorous ones; therefore, without water there could be no life. Inorganic matter furnishes the first and most sim¬ ple materials of existence ; organic bodies perish and be¬ come decomposed, and thus adding to the mass of inor¬ ganic matter which they had for a short period abandon¬ ed, they enter again as elements into the composition of other and more complex forms. Indeed, according to Mr W. S. Macleay, organized matter is nothing but a pecu¬ liar modification of brute matter acted upon by the vital 1 .«4 principle ; but this form of expression probably throws 1? 5C no new light upon the subject. 0 /'* We may here present a remark not unworthy of atten¬ tion regarding the nature of the flesh in the different classes of the animal kingdom. Considered as a viand, the flesh of animals becomes less substantially nourishing as we descend in the scale. For example, the flesh of a quadruped contains a greater quantity of nourishment in proportion to size and weight than that of a bird, while the latter exceeds in that respect both reptiles and fishes. Hence in catholic countries the latter are justly regarded as meagre, and form an exclusive food during the frequent days of abstinence by which it is sought to mortify the * flesh. Shell-fish and Crustacea, and a fortiori the zoo- phytical tribes, yield a still smaller proportion of nutritious matter. A revolting conclusion has been drawn from this alleged relation between the flesh of a highly organ¬ ized animal and the power and excellence of its nutritive qualities; viz. that cannibalism, or the habit of anthropopha¬ gous nations, opens up to those unnatural tribes a pleasure connected w ith the indulgence of the sense of taste greatly surpassing what is enjoyed by those who confine their mastication to the brute creation; because, in accordance with the rule supposed, the organic perfection and highly animalized nature of man is productive of a higher degree of nutrition, and of a greater capability of direct assimila¬ tion, when the substance of which he is composed is used as food by his fellow-mortals. But the scale of alimentary substances may rather be said to commence with air and water, and to terminate with the herbivorous animals; for the flesh of carnivorous kinds is, with very few exceptions, of a nature inadequate to the healthy sustenance of life. It is of a quality too putrescent, and decomposes with too great rapidity, as if the organization of matter could make no further progress, but passing rapidly from one extreme to another, hastened to throw off even the semblance of life, to assume again the simplest elementary form. “ Over-great affinity,” says Bacon, “ or consubstantiality of the nourishment to the thing nourished, proveth not well; for creatures feeding upon herbs touch no flesh; and of creatures feeding upon flesh few of them eat their owne kinde. As for men which are cannibals, they feed not ordinarily upon men’s flesh, but reserve it as a dainty either to serve their revenge upon their enemies, or tosa- tisfie their appetite at some times. So the ground is best sowne with seed growing elsewhere; and men do not use to graft or innoculate upon the same stocke.” Mineral bodies are still more unfit for the purposes of nutrition. They furnish both medicines and poisons in abundance, but never aliments. The difference between these objects may be shortly stated as follows: Aliments are substances alterable by the action of the organs which appropriate them; medicines act on the organs, of which they alter or modify the action; poisons attack and ex¬ tinguish life itself. But according to the specific nature of different animals, and various other circumstances, the qualities of these agents are convertible, so that aliments become poisonous, and poisons alimentary. Thus opium, which among European nations is a medicine, and too frequently a poison, has become, according to the prac¬ tice of several eastern nations, an alimentary substance. Aloes, which are simply^ medicinal for the human race, are a destructive poison to many carnivorous animals. On the other hand, according to Pallas, hedgehogs eat abun¬ dantly of cantharides without being in the slightest de¬ gree incommoded by them; and bees are known to feed History, Naturall and Experimental, of Life and Death. English translation, 1638. ANIMAL KINGDOM. 167 t pon and form their honey from the secretions of many iernicious and even poisonous plants. The caterpillar of - certain sphinx moth is highly delighted with the acrid nd venomous fluid of a tithymalis. The more that animals enjoy the qualities of youth, trength, and activity, the greater is the increase and de- elopment of their parts, and the greater the necessity for n abundant supply of food. Of many individuals exposed 3 an absolute abstinence of many days, the young are [ways the first to perish. Of this the history of war and bipwreck offers in all ages too many frightful examples, 'here are several instances on record of an almost total bstinence from food for an extraordinary length of time, laptain Bligh, of the Bounty, sailed nearly 4-000 miles in a open boat, with occasionally a single small bird not lany ounces in weight for the daily sustenance of 17 eople; and it is even alleged, that 14 men and women of ic Juno, having suffered shipwreck on the coast of Arra- an, lived 23 days without any food. Two people first ied of want on the fifth day. In the opinion of Rhedi, nimals support want much longer than is generally be- eved. A civet cat lived 10 days without food, an ante- >pe 20, and a very large wild cat also 20; an eagle sur- ived 28 days, a badger one month, and several dogs 36 ays. In the memoirs of the Academy of Sciences there an account of a bitch, which having been accidentally nit up alone in a country-house, existed for 40 days ithout any other nourishment than the stuff on the wool :'a mattress which she had torn to pieces. A crocodile ill live two months without food, a scorpion three, a bear x, a chameleon eight, and aviper ten. Vaillant had aspider lat lived nearly a year without food, and was so far from eing weakened by abstinence, that it immediately killed lother large spider, equally vigorous but not so hungry, Inch was put in along with it. John Hunter inclosed a >ad between two stone flower-pots, and found it as lively > ever after 14 months. Land-tortoises have lived with- it food for 18 months; and Baker is known to have kept a aetle in a state of total abstinence for three years. It af- i’wards made its escape.1 Dr Shaw gives an account of vo serpents which lived in a bottle without any food for ve years. The necessity of aliment becomes less vividly felt du¬ ng sleep, and certain other periods of prolonged repose, here are several animals which hybernate, or go into win- r quarters for six months in the year, during which period any of them require no food, but are maintained solely / that excellence of bodily condition which they had ac- tired during a prior period of activity and good cheer, his leads us naturally to consider what is called the hy- rnation of animals. Many creatures are so constituted that the activity of eir functions is greatly impaired by a comparatively ight reduction of temperature. Naturalists and anato- ists have alike sought in vain for either external or in- nial characters of general application, by which they ight distinguish, a priori, the species subjected to this range though well-ordered lethargy. They belong to rious genera and tribes, many of which have few cha- cters in common, as will be perceived when we name well-known instances the dormouse, the hedgehog, 'd the bat. It influences both warm and cold-blooded mnals. I he former of these, at certain advanced periods the autumn, according to the species, seek out places repose, either in the earth, among old walls, in caverns, unks of trees, or bushes; which retreats they usually ie with dried herbs, grasses, leaves, or moss. The bat tno»es caverns, churches, barns, and other situations where the temperature is milder than that of the open Animal air; and, contrary to the usual practice, it suspends it- Kingdom, self by the hooked claws of its binder extremities. It'^^v^s^ is the practice of other hybernating animals to contract themselves into a ball, in such a manner as to expose the smallest possible surface to the action of the air. When discovered in their retreats they are generally thus rolled up, cold to the touch, their limbs stiffened, their eyes closed, their respiration slow, interrupted, sometimes even imperceptible, and their insensibility so great that they may be removed, rolled about, and otherwise maltreated, without showing any further signs of life. It has been observed that the temperature of these ani¬ mals gradually lowers itself as the season declines. Their respiration also becomes slower, their motions less lively, and their appetite diminishes; but sensation and the power of locomotion still continue. This intermediate state between the perfect performance of the vital func¬ tions and confirmed torpidity endures for several weeks; the degree of temperature at which different animals be¬ come entirely overpowered varying, of course, according to the species. The propensity has been observed, in the following well-known animals, to correspond to a scale of descending temperature, according to the following or¬ der:—Is#, The bat; 2dly, the hedgehog; Mly, the dor¬ mouse ; ^thly, the marmot; othly, the hamster. Although many other animals are subject to the same law, it is only among those just enumerated that an exact comparison has been instituted. A complete state of hybernation consists in the sus¬ pension of sensation and voluntary motion, in addition to a great decrease in the temperature of the body, and in the frequency of respiration. Its different degrees of in¬ tensity are well ascertained by the number of respirations in a given time, or, in its most perfect state, by the total suspension of all respiratory movements. The different species of the bat tribe are those of which the torpidity is the least profound; and the marmot probably experiences the greatest degree of vital suspension. The temperature of these animals during their lethargy depends in a great measure upon that of the external air, and is consequent¬ ly variable. It is in general, however, superior to it by several degrees. It may descend to within a few degrees of the freezing point, but is not susceptible of reduction to that point, without producing either re-action of the vital functions or death. There is, therefore, contrary to the opinion of some of the older naturalists, a degree of external cold incompatible with the torpidity or existence of these animals. The species most easily rendered torpid, such as the bat, the hedgehog, the dormouse, the lerot, and the muscardine, cannot support a cold of 14° of Fahrenheit. A warmth of from 50° to 53° brings them again to life. Sundry mechanical means, such as different degrees of motion, serve to restore several of the last-named species without any increase of temperature; but to preserve them in a state of prolonged activity, a gentle warmth must be applied and continued. It is evident, from these and other obseiwations, that the sleep of mammiferous animals is not characterized by a uniform and constant duration. As it is dependent on the variations of the atmosphere, it will commence at an earlier, continue a longer, or be interrupted after a shorter period, according to the difference in the seasons of par¬ ticular years, the skill which the animals may have exhibit¬ ed in the choice of a protecting habitation, or the pecu¬ liar constitution of the species, or even of the individuals. The habit of storing up a supply of winter provisions also depends upon their greater or less degree of exposure to 1 See the observations prefixed to the translation of Spallanzani's Tracts, before referred to. 168 ANIMAL KINGDOM. Animal the power of awakening influences. The hedgehog, for Kingdom, example, has been observed to form several separate ma- gazines, to which it has recourse during the winter season ; and the marks of its little feet have sometimes been traced on the surface of the surrounding snow the cares of the family. This is usually the case among i the various tribes of birds, and also among carnivorous Kj quadrupeds; whilst the males of such as feed on vege.'- tables, and which consequently find almost everywhere an abundant and easy nourishment, abandon to the mo. The 'hybernatlon"'of the” swallow is a point on which then the rearing and education of their young. I, i • • *1 *i V*/-»/-»tv nlrrn 1"Pfl. T1 HOW very dissimilar opinions have been promulgated. It now ap pears to be the prevailing belief that these birds migrate on the approach of winter to other and more genial climes, but that cases do occur in which such individuals as are prevented by circumstances from joining the “ marshalled array,” are enabled to survive the rigours of our northern winters by the power which their constitution possesses of assuming the torpid state; at least the occunence of torpid swallows, however rare, is too well authenticated to be a matter of doubt. It is said that the tanrec, a species of hedgehog found in Madagascar, becomes torpid for some months in the year. If this assertion is well founded, it affords the only know n instance of torpidity in a mammiferous quadruped of a warm climate. Many cold-blooded animals may be regarded as of the hybernating kind. Indeed the greater proportion of rep- insects, molluscous animals, &c. inhabiting cold tilt es, countries, are very lethargic during the winter season, which they usually pass without food. They appear sub¬ ject to the influence of this feeling even in warm cli¬ mates; at least Humboldt describes certain reptiles in South America which pass a portion of the year buried in the earth, and which are only aroused by the occur¬ rence of rainy weather or the excitement of violent means. “ The manners of animals,” says this enlightened ob¬ server, “ vary in the same species, according to local cir¬ cumstances difficult to investigate. We were shown a hut, or rather a kind of shed, in which our host ol Cala- bozo, Don Miguel Cousin, had witnessed a very extraor¬ dinary scene. Sleeping with one of his friends on a bench covered with leather, Don Miguel was awakened early in the morning by violent shakes and a horrible noise. Clods of earth were thrown into the middle of the hut. Presently a young crocodile, two or three feet long, issued from under the bed, darted at a dog that lay on the thresh¬ old of the door, and, missing him in the impetuosity of his spring, ran toward the beach to attain the river. On examining the spot where the barbacon or bedstead was placed, the cause of this strange adventure’was easily dis¬ covered. The ground wras disturbed to a considerable depth. It wyas dried mud, that had covered the crocodile in that state of lethargy, or summer sleep, in which many of the species lie, during the absence of the rains, amid the llanos. The noise of men and horses, perhaps the smell of the dog, had awakened the crocodile. The hut being placed at the edge of the pool, and inundated during part of the year, the crocodile had no doubt entered, at the time of the inundation of the savannahs, by the same opening by which Mr Pozo saw it go out. The Indians also been observed, that among such birds as feed on liv. ing prey, the male is very assiduous in assisting hig mate to procure a sufficient supply. But naturalists have erred in assigning the polygamous habit as a general charac- teristic of the gallinaceous kinds. The instinct to pair, or the habit of monogamy, is no doubt only bestowed on those species to which it is necessary for the rearing of their offspring, and differs considerably in the nature and permanence of the attachment, according to the position of the nest, i. e. whether it is built upon or above the sur- face of the ground. All birds which build on trees, as was long ago remarked by Lord Kames, are hatched blind, and almost without feathers, and consequently require the sedulous care of both parents. But the generality even of gallinaceous birds, which breed upon the ground, do likewise pair, though the hatching of the eggs is entirely confined to the female, who completes her task by leading the young to their proper food, which they are able im¬ mediately to pick up for themselves, being active and well feathered from their birth. The male, at the same time, continues to manifest a certain degree of paternal solicitude, by uttering the alarm-note on the approach of birds of prey, or other dangerous foes. Black game and wood¬ grouse, however, do not appear to pair at all; but in the spring a male bird assembles a certain number of females about him, which afterwards deposit their eggs, and rear their young altogether independent of the male parent They are therefore polygamous in the proper acceptatiqn of tire term. Even among herbivorous quadrupeds pair¬ ing is rare, because the female can suckle her young while she herself is feeding ; but the monogamous habit proba¬ bly obtains among most carnivorous quadrupeds, and cer¬ tainly among all carnivorous birds, because incubation 1 -* y-v -f 1 y. yi y, i ■» v-» 4“ 4" 1 vv% 4" /"\ V ITA 4” 1 /A A* 1 /A /A rl ^ often find enormous boas, which they call ttji, or water- serpents, in the same lethargic state. To re-animate them, they must be irritated, or wetted with water.”1 Upon the whole, naturalists seem to be of opinion that no species of animal is condemned to torpidity by any in¬ herent property of its nature. It is a provisional faculty, dependent on external circumstances, and may be inter- rupted, postponed, or altogether prevented, by regulating the conditions under which the animal is placed. The disposition of animals in relation to other indivi¬ duals of the same species differs considerably. There are some which unite in couples and divide between them leaves the female no sufficient time to hunt for food,5 because young birds cannot bear a long fast, and therefore require the assistance of both parents, while unable to pro¬ vide for themselves. The association or fellowship of birds is either annual or for life; the former bond is the more usual, though eagles, crows, and several other species afford examples of a long-continued attachment. Many birds assemble in autumn, winter, and early spring, into flocks, but as soon as the pairing season has commenced they again separate into pairs. Others again appear to be more gregarious during the breeding season than at any other period of the year, for example the gan- net or soland-goose (P. bassanus) ; but this arises not so much from a love of fellowship with their kind, as from the accident of there being few places fitted for the pur¬ poses of nidification and the rearing of the young. We have said that pairing is rare among such quad¬ rupeds as feed on grass, because the female can feed her¬ self at the same time that she is suckling her young. The roe-deer, however, among herbivorous quadrupeds. forms an exception to the general rule. On the other hand, there are several carnivorous quadrupeds which do not pair, but the young of which are left entirely depen¬ dent on the mother; that is to say, the latter is obliged both to capture her own food and to suckle her offspring- Among gregarious quadrupeds which usually store up food for winter, pairing is probably necessary to prevent discord, and in this respect beavers are said to resemble those birds which place their nests upon the ground As I 1 Personal Narrative, vol. iv. p. 300. 3 See Karnes’s Sketches. ANIMAL KINGDOM. 169 I ,oon as the young are produced, the males abandon their Honjstock of food to their mates, and live at large, but return ^-frequently to visit them while they are suckling their young. Hedgehogs and most of the monkey kind pair. Seals are polygamous, and turtles leave their young to be hatched by the heated sand. Earwigs, spiders, bees, and ivoodlice (onisci), are amongst the few of the insect tribes which pay any attention either to their eggs or offspring. The young of the greater proportion of animals is produced in spring, when the supply of food is the most abundant, and when the long period which intervenes before the ap¬ proach of winter enables them to acquire strength to sup¬ port the rigours of that inclement season. Though the period of gestation varies considerably in the Efferent quadrupeds which feed on grass, yet the females ire regularly delivered in spring, or early in summer, when the herbage is nutritive and abundant. The mare com :eives in summer, carries 11 months, and brings forth in May. The same is nearly the case with the cow. The dieep and the goat are usually in season in November; they carry five months, and produce when grass has begun to spring. They love short, close herbage, upon which a horse or cow would barely thrive. The ass is in season iboutthebeginningof summer, but she bears twelvemonths, md consequently brings forth likewise early in summer. Wolves and foxes copulate in December, but as they only bear five months, they bring forth in April, when the sea- ion has assumed a genial aspect, and animal food is as abundant as at any other season. If we were to guess what would probably be the rutting season of animals, we would say summer, especially in a northern country; and yet, to quadrupeds which carry their young only for four or five months, such economy would be injurious and im¬ provident, as it would bring the time of delivery at an un¬ due season, both for warmth and food.1 There are a few exceptions to the above rule, which, however, in them¬ selves, belong to an equally beautiful system of providen¬ tial ordinances. Some gregarious and stoi'e-collecting animals, for example, bring forth in January, when their granary of provisions is still abundantly filled. The season of pairing, or of production, among wild ani¬ mals, usually takes place only once a year, and at a fixed period; but those which man has rendered domestic are observed to couple at all seasons. The species of warmer climates, when transported into colder regions, usually cease to pair, or at least their union is unproductive; and the same consequence generally follows a state of capti¬ vity. Among such species, however, as man has fairly reduced to a state of satisfied domestication, the indivi¬ duals become much more prolific than in the wild state. fhe season of love varies greatly among mammiferous animals. The greater proportion pair in spring and sum¬ mer ; but the wolf pairs in winter, the stag in autumn, and many domesticated animals at diversified periods throughout the year. Prolific union takes place among varieties of the same species; and it is by paying attention to these that the finer races of our domestic animals are maintained and continued. As the climate of northern countries causes several of our most valuable animals to degenerate (as it is called), it has been customary to ob¬ tain from time to time a male animal of a pure and noble lace, which, when paired with an ordinary female, produces a breed scarcely inferior to the male parent; for it has been observed that, with few exceptions, the new produce assumes the characteristics of the father. Thus, in uniting a. £CP °* an ordinary kind with a ram of the Merino race, t e first generation almost equals the father in beauty, hat is frequently called deterioration in animals is, more properly speaking, their natural assumption of those Animal peculiar attributes which fit them for the inclemencies of Kingdom, climates uncongenial to their original nature. A Lap-' lander is no more a deteriorated Asiatic of the Mongolian or Caucasian line, than a Georgian or Circassian is a highly refined Laplander; neither is the Shetland pony a deteriorated Arabian courser, any more than the steed of Araby is a thorough-bred shelty. Each has been enabled by a wise provision of nature to assimilate its character and constitution to the qualities of the climate in which it was destined to exist; and had it been incompetent to effect or undergo such assimilation, it would then indeed have deteriorated—that is to say, it would have died. If we admire the slim smooth elegance of the Italian grey¬ hound, and regard the rough shaggy coat of the dog of Nova-Zembla as a deterioration, let us remember that that which is the beauty of the one would be the bane of the other; and what would then become of that forlorn agriculturist, whose business it is to drill the ice and to furrow the snow ? The small stature and peculiar habits of the northern pony would have been as little fitted to sustain the fiery breath or the shifting sands of an eastern desert, as the graceful Arabian to withstand the cold and cloudy clime, and the rugged and precipitous mountains, of Lapland or Thule. Therefore, instead of being dete¬ riorated, each ought rather to be said to exist in the best and most improved condition, according to the nature of its particular calling. Using the word, however, in its more usual acceptation, it may be stated that an animal seldom degenerates in its native country, but more fre¬ quently in those for the climate of which its constitution is not adapted. Each species appears to have a certain extent or circle of natural distribution, in the centre of which it not only most abounds, but also there shows it¬ self in its finest and most characteristic proportions. As the places of its occurrence diverge from this centre of dominion, it becomes rarer, and exhibits a variation or considerable departure, at least in its external characters, from the primitive model. Thus the horses of Arabia and Barbary degenerate in Britain; and, to preserve the breed in purity, they must be frequently crossed by the original; but the Arabs themselves are very careful to pre¬ vent any mixture in the blood of their native and noble kinds, and would deem them deteriorated by such alliance. In the tabular or abridged views of classification which we here present, it is our intention merely to exhibit the great primary divisions of the animal kingdom called classes. The secondary divisions into orders, and the fur¬ ther dismemberment of these into minor groups called fa¬ milies, genera, &c., will be illustrated when we come to treat of each class in particular under its proper head. Neither do we intend to trace the progress of classifica¬ tion from the earliest ages of sciehtific record; because, as the object of the naturalist is rather to ascertain the nature and relationship of things as they are, than as they were supposed to be, there is the less necessity for leaving our direct x’oute, to trace either the origin or the progress of error. We shall proceed, after a few observations, at once to the system of Linnaeus, which is in fact the basis of all that have succeeded, and without a knowledge of which it is impossible to understand either the merits or defects of more recent systems. Indeed, with the excep¬ tion of the purely artificial classification of Klein, and the multiplied orders of Brisson and Vicq-d’Azir, all the sys¬ tems which have appeared since the middle of last cen¬ tury are indebted more or less to the labours of the im¬ mortal Swede, and may be valued almost exactly in pro- vol. in. Karnes’s Sketches. 170 ANIMAL Animal portion to their share in the lucidus ordo of the Linnaean Kingdom. SyStem. For example, the Systerna Regni Animalis of Erxleben (1777) is nothing more than a revised edition of the Systema Naturae, in which are engrafted, with no lack of skill, many additions both of species and genera; the whole being presented in a consecutive series, without the accustomed subdivision into the primary groups called orders. The Prodromus Methodi Animalium of Storr (1780) does not differ radically from the Linnaean system. The Elenchus Animalium of Bodaert (1787) is allied to it still more closely in every thing except its accuracy. Of Gmelin’s work (1789) we need not speak, as it is a pro¬ fessed revisal (being the 13th) of the Systema Naturae. And although no one will deny the merit of profound and original inquiry to the investigations of Blumenbach, most readily will those who are best acquainted with his la¬ bours admit, that in zoology he has wisely followed the footsteps of Linnaeus. The six classes into which the German naturalist divides the animated creation, viz. Mammalia, Aves, Amphibia, Pisces, Insecta, and Vermes, correspond with those of the Linnaean arrangement, al¬ though their orders and genera are in some respects dif¬ ferently combined. His motto appears to have been Malta jiunt eadem, sed aliter. (Quintilian.) The signal benefits conferred on natural history, in all its branches, by the learning and genius of Baron Cuvier, are known wherever the science has obtained a zealous and successful cultivator; and it cannot have escaped the notice of the critical observer, that after 30 years of profound and philosophical research into the mysteries of the animal kingdom, the most enlightened zoologist of the age should have finally reverted to a closer approximation to the Linnaean system, than had characterized his views at any former period of his brilliant career. When he first made known (in 1797), conjointly with M. Geoffrey, his new classification of mammiferous animals, his nume¬ rous genera were contained under no less than 14 differ¬ ent orders. Thirty years afterwards (in 1817) he pub¬ lished his Regne Animal, with many improvements in the composition and arrangement of the minor divisions, and with the addition of the order of which he is himself so bright an ornament, but otherwise composed (we speak at present of the mammalia alone) of primary divisions ex¬ actly the same in number, and nearly the same in nature, as those finally divulged and established by Linnaeus him¬ self just 60 years before. Latreille, Dumeril, Desmarest, and Frederick Cuvier, are followers or coadjutors of the Baron, and with him are partakers in the modification and amendment of the Lin¬ naean system. I he venerable Lamarck has greatly signa¬ lized himself in a field which, it must be confessed, was obscure to the eye of Linnaeus—that of the molluscous animals—which, under the name of Vermes Testacea, were but indifferently treated in the Systema Naturae. The error appears to have lain in the greater attention which was bestowed on the shells themselves, or testace¬ ous coverings, than on the animal inhabitants ; and the con¬ sequence has been, that the conchologist of the old school ranks with the collector of china, whether old or new. The names above enumerated are certainly among the foremost in the annals of modern science; and although, in addition to these, many more might be mentioned with honour as having contributed, by monographs or other par¬ tial though highly prized contributions, to the increase of knowledge, yet we are not aware that more than three systematists of acknowledged and wide-spread influence, or of what may be termed universal celebrity, remain unnoticed, of those who have essentially influenced the present condition of zoological science ; we mean Fa- biicius, Illiger, and iVl. de Blainville. We have no hesita- KINGDOM. tion in asserting, that as the writers first mentioned owe much of the success which has attended their labours to Kit their having judiciously engrafted their own improvements^ on the original stock of the Linnaean system, so the au¬ thors last named, though not less highly gifted, have in a great measure sacrificed the utility of many original and enlightened views to the fond conceit of a new, and in some instances an incomprehensible, nomenclature. The skill of Fabricius as an entomologist has never been surpassed, and it is therefore the more to be regretted that he should have been influenced in the formation of his system by other motives than a desire to perceive and point out the truth. But it is known that he was swayed as much by the ambitious hope of founding a new doc¬ trine, of which he destined himself to be the oracle, as by the desire of proceeding directly in the path of nature. Hence his avowed enmity to the eclectic system of La¬ treille, which, during the opening career of that celebrated entomologist, he declared it to be his intention utterly to destroy. Yet the system of Latreille not only stands, but, when viewed in relation to the application of its general principles, has in a great measure superseded that of Fa¬ bricius. At the same time, the accurate discrimination and extensive knowledge of the latter, and the wide circle which his system embraces in detail, render it still indis¬ pensable for a knowledge of the species. Illiger died young. Flis talents were such as to raise among his compatriots the highest hopes of his future eminence, and his death was a subject of just regret to all who knew what he had achieved so well at an early age, and who the more gladly lent themselves to the anticipa¬ tion of what he would afterwards have accomplished had his life been prolonged. Of his classification it has been written by a competent judge : “ Neque apud veterem, neque apud recentiorem quendam auctorem ullum systema inyenerim, quod, tam sua perspicuitate, quam accuratione, Illigeriano magis commendari mihi videatur.” Many of his genera are indeed remarkable for their felicitous con¬ struction and consonance with the natural arrangement. They have in consequence been readily adopted by his more fortunate fellow-labourers in the same field, in whose works they will remain, and be handed down in ample at¬ testation of the author’s genius; but the system itself will suffer a partition, and ere long cease to be practically known under the form in which it was originally promul¬ gated, and this mainly in consequence of his having adopt¬ ed so many new names. M. de Blainville is still alive, and the longer he lives the better for the sciences ef anatomy and physiology, neither of which contains in its modern annals the name of a more accomplished or enlightened expounder of its mysteries than his own ; but in the character of a natural¬ ist, and in connection with the subject of nomenclature, he unfortunately sins more than all his predecessors. He really miscalls the objects of zoology most sadly, yet his knowledge of the essential bases of the science is no doubt too profound to admit of his applying it without new and important results. Hence the pity that these should not at all times be stated in such terms as not only to amalgamate more closely with the kindred labours of his contemporaries, but to fall rather more clearly within the comprehension of ordinary minds. As it is not our intention in the present rapid sketch to enter into the distribution of the animal kingdom beyond the greater divisions called classes, we shall not exhibit the systems of the two first-named authors further than to say, that the former attended almost exclusively to entomology, the latter chiefly to the mammalia and birds. W hen we come to the divisions of our subject under their separate heads, tabular views or more detailed ana- ANIMAL KINGDOM. 171 ai lyses will be presented of the labours of these and other >m ingenious inquirers of the past and present times. _ We have said that we regarded the system of Linnmus is the basis of all those by which it has been succeeded, ind that without a knowledge of his classification it •vould be impossible to understand either the merits or lefects of more recent systems. We shall therefore here iresent the classes into which the great Swedish natural- st divides the animal kingdom. Division I. 4 heart with two auricles and two ventricles ; blood warm and red. ^lass I.—Viviparous animals, or such as suckle their young; commonly called quadrupeds, but includ¬ ing also the cetacea or whales, Mammalia. 3lass II.—Oviparous animals, or birds Ayes. Division II. 4 heart with one auricle and one ventricle ; blood cold and red. ’^lass III.—Animals breathing arbitrarily through lungs Amphibia. Hlass IV.—Fishes, or animals with gills... Pisces. Division III. 4 heart with one ventricle, no auricle; blood cold and white. Aass V.—With antennae ; undergoing transformations. Insects Insecta. Hlass VI.—With tentacula, and undergo- ingno transformations. Worms, Vermes. It may be observed, that the deservedly popular system >f Linnaeus, though it does not profess to be a natural nethod of classification, actually is so in many of its parts; lor can it be denied that, on the whole, it usually brings together as many groups of natural genera as occur in Animal most systems that have been promulgated, especially if Kingdom, we take into consideration the period at which it was'^^v->^' composed, and the comparatively scanty materials within his reach. Linnaeus was probably aware of the extreme difficulty, we might say at once of the utter impossibility, of a perfectly natural arrangement; for he confesses, in his Philosophia Botanica, his inability to define the great divisions called orders, on account of their being so con¬ nected with each other by various points of affinity, as to form a map rather than a linear series; and the observa¬ tion may be applied with equal truth to the subjects of the animal kingdom. In regard to the excellence of his genera themselves, their consonance with nature is ren¬ dered still more evident, by the great proportion of those which Cuvier and Latreille have retained as leading ge¬ neric divisions in their recent works,—certainly the most skilful approaches which have yet been made in the es¬ tablishment of a natural system. It has been asserted, and we believe with truth, that such naturalists as are perpetually intent on the abstract theory of classification, rarely attain the highest excellence in the discrimination or definition of the species,—the only distinctions possibly which have a real foundation in nature, and upon an ac¬ curate and extensive knowledge of which alone their theoretical systems can be substantially and permanently built. At all events, it is admitted that Linnaeus is a guide almost infallible, in as far as concerns his wonderful facility in discovering the minor natural groups. If he could have combined these as well as he has defined them, his possession of the sceptre would have been still undoubted.1 M. Virey, in the first edition (1803) of the Nouveau Dictionnaire d’Histoire Naturelle, divides the animal king¬ dom into three great tribes, in accordance with the nature and distribution of the nervous system. As he appears to have been among the first to attribute a due degree of importance to that system in the classification of animals, we shall here exhibit a view of his general arrangement. Wimals possessed of two nervous systems, the cerebro-spinal and the ganglionic Tribe Vertebrata. Heart with two ventricles and two auricles; blood warm, lungs cellular Heart with one ventricle and one auricle ; blood cold JMan and Mammalia. | Birds. \ Reptiles and Fishes. Tribe Invertebrata. \nimals possessed of a single nerv¬ ous system surrounding the (Eso¬ phagus, with ganglia and branch¬ es ; the sympathetic A heart; branchiae for respiration, mostly aquatic No heart; some vessels; tracheae for air or water f Mollusca. J Cirrhipedes. f Crustacea. f Arachnides and Aptera. ) Insects, winged, hexapod, y Annelides and Helminthides. C Intestinal worms. Tribe Zoophyta. Nervous system composed of mo¬ lecules more or less perceptible ; no distinction of sexes Ascidia, inclosed in a tunic Radiated animals ; composed of rays parting from a centre United in Polypiers, or stony masses ; coral- ligenous Microscopical Botrylli, &c. Echinodermata. Hydra and Polypus. Corals and Ceratophytes. Madrepores and Sponges. Infusory Animals. ... ^16 following summary will serve to illustrate M. neys views of the nature and characteristics of these .ree groat divisions. We commence with the zoophy- ical tribe. 1 J 1st, Zoophytes are distinguished by an organic tissue of | ' cry soft and pulpy nature, more or less diaphanous, n very contractile, though we cannot readily perceive its muscular fibres. Its fundamental character consists in the extreme division of its nervous molecules throughout the flesh of these animals. Except in the Echinodermata and some other radiated classes, we can scarcely assert the existence of a nervous system amongst them (on which account they are named apathiques by Lamarck). Each portion of the body having its nervous molecule and its par- 1 Horce Entumologicw, part ii. p. 428. 172 Animal Kingdom. ANIMAL KINGDOM. ticular source of vitality, there is no common centre of ■ sensation ; thus division and generation are almost syno- ^ nymous, and when individual parts are mutilated they ai e speedily reproduced. With these tribes the production of the species is in fact nothing more than a simple sprouting of a bud or offset, which separates itself from the maternal stalk. Zoophytical animals are of no sex, and thus resemble the agamous vegetables. Ihe mouth is usually placed in the centre of the body, and is fre¬ quently surrounded by a species of unarticulated arms, x'adiating from a centre like the petals of many flowers. Several genera have only a single opening for the recep¬ tion and rejection of the aliment, f hey have no visceia (excepting ccBca in certain species) ; no heart, not aiterial nor venous vessels; no true circulation; no apparent organs of respiration. They are all aquatic, and water seems indeed the only fluid which pervades their economy. They may be called the cryptogamia of the animal king¬ dom. The sense of touch, and perhaps that of taste, seem the only ones enjoyed by these animals. 2<%, The Invertebrata present a greater complication of organs. Their principal character consists in a nervous system, extending itself, especially in the intestinal cavity, by numerous ramifications. In all the families of this great tribe, the nervous trunks surround the oesophagus, pass beneath the belly, and are furnished with many ganglions which supply branches to the different organs. That which is regarded as the brain in this tribe (named sensible by Lamarck) is nothing more than one or two ganglia situated above the oesophagus ; but the particular distribution of the two nervous branches which spring from the collar of the gullet, and extend themselves over the body, gives rise to the divisions of molluscous and ar¬ ticulated animals established by Cuvier. The last-named naturalist has observed, that in the mol- lusca the nervous system is composed of many ganglionic masses, dispersed throughout the organization, but con¬ nected by means of nervous filaments ; and that the chief of these masses constitutes a kind of brain above the oesophagus. The mollusca have no skeleton ; their muscles are attached to the skin, a soft contractile envelope, in which in many species are produced shells, or stony bodies of calcareous carbonate, formed by exudation or superimposed concretion. Besides the sense of touch, common to all animals, the mollusca are gifted with that of taste, and sometimes of sight; but the sense of hearing has not been remarked, except among the Cephalopoda or cuttle-fish. Their systems of digestion and secretion are rather complex; they are provided with a liver, and possess a circulating or vascular system, through which flows a humour or whitish sanies in place of blood. Their respiration is effected by aquatic or aerial branchiae. The' sexual organs are frequently united in the same in¬ dividual. Among the articulated classes (such as the Crustacea, the arachnides, and insects) the nervous system consists of a double chord, extending from the head to the posterior extremity, and bearing knots or ganglia which correspond to the segments of the animal’s body. The first ganglion above the oesophagus takes the place of what we call the brain in the higher animals, but it is not voluminous in proportion. All these animals are composed of segments or annular divisions, and their forms are elongated, and more or less cylindrical. Their skin or outer covering, always of a somewhat solid texture, becomes in many fa¬ milies hard, corneous, or even stony; and the muscles are attached to its interior. The greater number have arti¬ culated members, feet, wings, pincers, palpi, &c. Many j of these animals have closed vessels; and the Crustacea Kij have a heart and branchiae. Others, according to Cuvier,^ are nourished by simple imbibition. Those insects which undergo metamorphoses are furnished with tracheae or air-vessels for respiration, dispersed over their bodies. The organs of the sense of hearing are not discernible except among the Crustacea; taste is universal, and also sight, except among the worms. Their jaws always ply laterally. The sexual organs are usually separate. 3£%, The Vertebrata comprehend all those animals which have a nervous system composed of ganglia, called syra- pathetic, for the functions of the internal life ; and another symmetrical nervous system, of which the principal por- tions are inclosed in the cranium and spinal column, and which sends off chords for the functions of the external life. These are the most perfect and most highly en¬ dowed of all animals ; they are named intelligens by La¬ marck, and they are always endowed with five senses, of which never fewer than four are situated in the head. They possess a heart, red blood, a liver, lungs in the spe¬ cies which live in air, and branchiae in those which live in water. An articulated, bony, symmetrical skeleton, placed in the interior of the body, gives support and so¬ lidity to the different parts. Such are man, mammiferous animals, and birds, which have warm blood, and respire by cellular lungs; such also are reptiles and fishes, of which the blood is cold. In all, the mouth has two hori- zontal jaws, and the members are never more than four in number. The preceding are M. Virey’s view's of the distribution and general characteristics of the different classes of the animal kingdom.1 They contain a sound exposition of several of the substantial relations which exist between the different systems of the animal economy, and we pre¬ sent them to the reader even at the risk of afterwards repeating in part several essentialities of his doctrine, when we come to promulgate the views of his celebrated countryman and contemporary Baron Cuvier. We shall next present a tabular view of the general dis¬ tribution and primary divisions of animals according to the system of Lamarck. It wdll be observed that this author commences with the lowest tribes. Animals without Vertcbrce. * Apathetic Animals. (Apathiques.) | Characters. No brain nor elongated medullary mass; no special senses; rarely articulated. 1. Infusoria. 2. Polypi. 3. Radiata. 4. Vermes. (Epizoaria^) * * Sensitive Animals. forms various ( Sensibles.) 5. Insecta. r Characters. No vertebral column; a 6. Arachnides. brain, and generally an elongated 7. Crustacea. medullary mass; some distinct or 8. Annelides. j special senses; organs of movement 9. Cirrhipedes. attached beneath the skin ; formed 10. Mollusca. [_ symmetrically of equal parts. Animals loith Vertcbrce. * * * Intelligent Animals. (Intelligens?) Characters. A vertebral column, a brain and spinal marrow; distinct senses ; organs of movement attach¬ ed to parts of an interior skeleton; formed symmetrically of equal parts. 11. Pisces. 12. Reptilia. 13. Aves. 14. Mammalia. Mceurs et Instinct dcs Animaux, tome i. p. 130. ANIMAL KINGDOM. The author of the preceding arrangement has entered Hit- pon the discussion of certain preliminary points connect¬ ed with the subject of classification, which maybe regard- 1 as composing the art rather than the science of Zoology, le inquires (in his introduction to the Hist. Nat. des 'nim. sans Vert.) what are the operations to be perform- 1 for the execution of a good distribution of animals, and tr the establishment of the necessary divisions of that istribution? These operations he states to be as fol- ■ws: 1st, To assemble animals together according to a dnciple which is not arbitrary, and so as to form a gene- il series, whether simple or ramified: 2dli/, To divide fis general series into different kinds of lesser divisions, ' which the one shall be subordinate to the other; and ith that view to make choice of and submit to certain utable and convenient principles (principes de conce¬ rned) : 3«7y, To fix the rank of each sort of division after le general principle, previously established; as, for ex- nple, The rank of each primary group in the total series; That of the classical divisions of each primary group ; That of the orders or families in each class ; That of the genera in each family ; and That of the species in each genus. The execution of these three sorts of operations is re- iirded by Lamarck as indispensable ; and that it is so has :en long felt by naturalists, almost all of whom have more less occupied themselves in the attempt, but always in i arbitrary manner—that is to say, without the previous tablishment of principles deserving of general consent, he first of the operations alluded to—that which con- rns the bringing together of species in such a manner as form a general series—is an essential preparation, which ight to precede the other operations, and without which deed the latter could not be executed. It tends, more- er, to enable us to discover the order of nature, with hich it is so highly important that we should become quainted. Although nature has necessarily followed a rtain order in the formation of organized beings, and pecially of animals, yet as she has now dispersed these dmals, and commingled all the different races over the I rface of the solid globe, or through the wide depth liquid waters, the original order of formation is to a cer- in extent disfigured, and so far imperceptible. We are erefore obliged, with a view to its ascertainment, to arch after some means by which we may attain to that scovery, and to work out some solid principles to lessen e chance of error. In regard to this, the most impor- nt step has been already attained, when we acknowledge e interest inspired by affinities or relations, and the ne- ssity of understanding these, with a view to submit to em, as to a test, the various parts of our general distri- ition. It may thus be perceived, that in order to esta- ish a good distribution of animals, in such a manner that > solidity shall run no risk of being enfeebled by the ar- trary nature of opinion, it is necessary first of all to as- 1 mble our species according to well-determined affinities ; toi which we may, without inconvenience, trace out the ics of demarcation which separate the groups called isses, and those other subordinate groups of which the tablishment is so advantageous, provided the natural unions are in nowise compromised by their formation, may perhaps be proper shortly to inquire into the na- re ^lese relations, their different degrees, and the act uses which it becomes us to make of such as we certain and acknowledge. We shall then be enabled ' greater facility to determine the principles which it ht to establish. delations, according to Lamarck, are those traits of tm dnce or analogy which nature has bestowed, as 173 well on her different productions when compared among Animal themselves, as on the different parts of those same pro- Kingdom, ductions when compared with each other. These traitsv ’ of resemblance and affinity are so necessary to be known and understood, that no methodical distribution can be established on a sure foundation, if the objects which it embraces are not arranged according to the law which they prescribe. Relations are of different orders, some being very general, others less so, and many altogether special or particular. Moreover, although, in general, relations belong to nature, all are not the direct result of her operations in regard to her productions ; for among the relations which we perceive between the compared parts of different beings, there are many which result merely from a modifying cause. Thus the relations of exterior form, which are so apparent between the cetacea and fishes, can only be attributed to a property resulting from the dense medium which each inhabits, and not to any direct plan in the operations of nature in regard to both. It is necessary then carefully to distinguish those obvious and acknowledged relations which pertain to the direct operations of nature in the progressive organization of animal life, from certain others, equally obvious and ac¬ knowledged, which result from the influence of local cir¬ cumstances, or from the peculiar habits which different races have in some instances been forced to acquire. Relations of the last-named nature, though certainly of greatly inferior value to the former, are by no means limited in their influence and exhibition to external cha¬ racters alone ; for it may be demonstrated that the exter¬ nal cause which possesses the power to modify the direct operations of nature frequently exercises an obvious in¬ fluence on several internal organs. It becomes the more necessary then to establish certain rules, devoid of arbi¬ trary qualities, to enable us justly to appreciate the nature and value of these relations; and it may be established as a principle in zoology, that it is from the interior or¬ ganization that the most essential are to be obtained. This principle is well founded if it expresses the pre-emi¬ nence which ought to be accorded to general considera¬ tions, gathered from the interior organization, over those derived from external parts. But if, instead of using it in this manner, we apply it to particular cases of our own choosing, and without pre-established rule, it is capable of great abuse; and we shall arbitrarily assign to relations presented by such or such system of internal organs, a preference over certain others, although the relations of the latter may in reality be of greater importance. By this means, sufficiently convenient for the changeable views of individual authors, we admit into various parts of our distribution inversions in every way contrary to the order of nature. It is true, as has been already observed, that a cause which modifies organization not only acts on the exterior parts of animals, but also produces various modifications in their internal structure. It follows, that it is incorrect to suppose that the relations which exist between the races, and especially between the genera, the families, the orders, or even in certain cases the classes of animals, can always be decided merely from the isolated conside¬ ration of any internal organ, arbitrarily selected. On the contrary, Lamarck is of opinion that the relations of which we speak cannot be suitably determined except by a consideration and comparison of the whole of the inte¬ rior organization, and, auxiliarly, by that of certain special internal organs which assured principles have demon¬ strated to be the most preferable and important. The second question proposed by M. Lamarck is the following: What are the principles by which we ought to be guided in our operations, so as to exclude from these 174 ANIMAL KING I) o M. Animal whatever is arbitrary ? Our author is of opinion that it is Kingdom, by the precise discrimination of each sort ot relations up, '^^’^and by aid of a determination, substantial and explained, of the preference which ought to be accorded to one kind over another, that we shall discover principles proper to regulate all the parts of our general distribution of ani¬ mals. It is necessary then to determine the principal kinds of relations which ought to be employed to attain this end, and then to fix the superiority in value which one kind possesses over another. The following is the classification of Lamarck in further illustration of this sub¬ ject. * Relations between Comparative Organizations, deduced from the whole of their parts. These relations, though general, manifest themselves in different degrees, according as we seek for them among races compared in themselves, or among groups of ani¬ mals of different races compared with each other. It is necessary then to distinguish several kinds. First kind of general relations.—These seem immedi¬ ately to connect races or species with each other. I hey are of necessity the first, because it is they which furnish the greatest of the relations between animals which differ from each other. The zoologist who determines it, taking into consideration all the parts of organization, as well in¬ terior as exterior, admits not of this sort of relation, un¬ less when it presents the smallest and least important difference. We know that animals which resemble each other perfectly, both in their internal and external organi¬ zation, can be nothing more than individuals of the same species. In this case the relation is not considered, as such animals offer no distinction. But those which present among themselves a difference, tangible, constant, and at the same time the smallest possible, are connected by the greatest and most immediate of relations, if they pre¬ sent elsewhere a great resemblance in all the parts of their interior organization, as well as in the greater pro¬ portion of their external features. And this sort of rela¬ tion does not demand a consideration of the degree of composition or relative perfection of the animal organiza¬ tion, for it determines itself in all the ranks of the scale. It is so easy to seize, that every one acknowledges it at first sight; and it is by employing it that naturalists have formed those smaller portions of the general series called genera, notwithstanding the arbitrary nature of their li¬ mits. Thus, in this first kind of relation, which may be called the relation of species, the difference between the objects compared is the smallest possible, and need only be sought for in the particularities of form or of the ex¬ ternal parts of individuals. Second hind of general relations.—This embraces the agreements which exist between groups of different ani¬ mals when compared together. It may be named the re¬ lation of groups; and, to acquire a knowledge of its nature, we must no longer occupy ourselves essentially with the particulars of the general form, nor with those of the ex¬ ternal parts, but almost solely with the interior organiza¬ tion, considered in all its parts. It is this kind chiefly which ought to furnish the differences by which we dis¬ tinguish the groups from each other; and it is inferior in one or more degrees to the first kind in the quantity of resemblances which exist between the compared objects. It serves to form the families, by connecting the genera with each other; to institute the orders, or the sections of the orders, by uniting several families ; and, lastly, it deter¬ mines the classical groups into which we ought to parti¬ tion the general series. The relations of which we now speak cannot, however, be employed to determine the rank of the great masses of that series, but only to form & diverse combinations for establishing and distinguishing these masses. # From the consideration of these relations, the two folA|A lowing principles may be deduced. First principle.—The general relations of the second kind do not require a perfect resemblance in the interior organization of the compared animals. “ Ils exigent seule- ment que les masses rapprochees se ressemblent plug entr’elles, sous ce point de vue, qu’elles ne le pourraient avec aucune autre.” (Anim. sans Vert, tome i. p. 355.) Second principle.—The greater and more general the compared masses, the more will such masses differ in their internal structure. Thus the families present a less difference in the interior organization of the animals by which they are constituted, than the orders or classes. Third hind of general relations.—We may denominate relations of rank those which serve to determine the posi¬ tion in the great series, and which, proceeding from a fixed point of comparison, effectively show among the compared objects a relation, whether great or small, in the composition or perfection of the organization. This kind is obtained by comparing any organization whatever, taken in the totality of its parts, with any other given or¬ ganization which may be presented as a point of depar¬ ture or of comparison. It is then determined, by the re¬ semblance, greater or less, which is found between the two compared structures, to what extent that which we com¬ pare departs from or approaches to that which is given as a point of comparison. \V e shall see that this sort of rela¬ tion is the only one which ought to serve for regulating the rank of all those important primary masses into which we divide the animal kingdom. If we consider the question concerning the choice of a particular organization, from or towards which to remove or approximate other organizations successively, accord¬ ing to their greater or less resemblance, it becomes evi¬ dent that the selection ought to fall on one or other ex¬ tremity of the animal kingdom (as in that case there would be no uncertain balancing), and the best known extremity should have the preference. Thus, in setting out from the most perfect and highly finished structure, we should, in the determination of all the ranks, proceed from the most composite to the most simple, and should close the series by the most imperfect and least organized of the whole. Of all forms of structure, that of man, considered in its totality, is at once the most composite and complete. From this M. Lamarck concludes that the more any ani¬ mal organization approaches that of the human race, the more it advances towards comparative perfection and its own completion. This being the case, the organiza¬ tion of man is with that author the point of comparison and departure from which to judge of the relation, whe¬ ther near or distant, of every form of animal structure, and by which we are to determine the rank which those forms, or the groups which they constitute, ought to oc¬ cupy in the general series. The organization alluded to, considered in the totality of its parts, furnishes the means by which to judge of the degree of composition and per¬ fection of other animal structures also regarded in their totality. And in doubtful cases it is not difficult to rid ourselves of uncertainty and embarrassment, by having re¬ course to the fourth kind of relations, or to those princi¬ ples which concern the comparison of the different organs separately considered, and establish a predominating value and influence among certain of those organs when com¬ pared with others. Thus, our point of departure or com¬ parison being found, the rank of all the divisions may be assigned with facility by the aid of the principles which follow. £ ANIMAL KINGDOM. 175 Fint principle.—For the determination of the rank of , ach mass in the series,—the most complicated and com- ■lete of animal organizations being selected as the point f comparison,—the more another form of animal struc- jre considered in the whole of its parts, resembles that f the fore-chosen, the more it will approximate towards by its relations; and reciprocally in the converse cases. Second principle.—Among the organizations of which ie plans are different from that which comprehends the articular structure selected as the point of comparison, lose which offer one or more systems of organs similar r analogous to such as form a portion of that with which ley are compared, shall rank superior to those possessed fa smaller number of these organs, and, a fortiori, to those i which they are wanting. With the assistance of the three kinds of relations above idicated, and the principles deducible from them, M. La- larck regards it as easy to determine the distinctions f species, and those of the various larger groups which iccies form ; and to decide, in a manner by no means ar- itrary, the rank and station of each group in the great iries. If this be true, the science will cease to be as va- jillating in its onward march as it has hitherto proved. But our efforts would be incomplete, and would still ■ave too large a field for the exercise of arbitrary opi- ions, if no attempt were made to establish and define the due of particular relations,—that is to say, of those which ;-e obtained by the comparison of particular internal Gr¬ ins, considered in an isolated manner in different animals. * Relations between similar or analogous parts talien se¬ parately in the organization of dijf h'ent animals, and compared ivith each other. The fourth hind of general relations merely embraces irticular relations between unmodified parts. It is drawn om the comparison of parts separately considered, and liich, in the system of organization to which they belong, fer no real anomaly. The consideration of this kind is imetimes of great consequence in assisting to decide in Dubtful cases, when we are anxious to determine, among Ttain compared groups, to which the superiority' of rank ight to be assigned. Such cases sometimesi occur where ic whole of the parts of the interior organization present ) means of deciding, in an unarbitrary manner, to which ' two organizations the superiority belongs. It is espe- ally in the formation and position of the orders, sections, milies, and even genera, of each class, and consequently the assignment of the rank of all these inferior groups, lat the employment of this fourth kind of relations is of Ivantage; because, in regard to such groups, the prin- ple of the third sort of relations is frequently very diffi- ilt of application ; and thus arbitrary modes of arrange- ent are introduced, most baneful to the science, by ex- ising the works of naturalists to a continual variation in ie determination of the relations which ought to fix the imposition of the groups, and their order of position, i fact, as many animals, really connected by the general I unacters of their class, present remarkable differences in Ttain of their interior organs, and yet at the same time dlibit equally striking resemblances in others, we feel lat, in order to appreciate the degree of importance pos- issed by the relations which exist between particular or- ms, we must have recourse to certain regulating prin- ples in our determinations, to avoid arbitrary conclusions, be two following principles are proposed by Lamarck, to nible us to appreciate the relations observable between irticular internal organs in different animals compared ith each other. First principle.—Between two internal organs, or syrs- ms of internal organs, separately considered and com¬ pared, that of which nature has made the most general Animal employment ought to have the pre-eminence assigned it Kingdom, in the value of the relations which it presents. Accord-v-^v^^1 ing to this principle, the following is the order of import¬ ance which we ought to attribute to the particular organs which nature has employed in the interior organization of animals The organs of digestion ; The organs of respiration ; The organs of movement; The organs of production; The organs of sensation ; The organs of circulation. Thiis, when we take into view the greatest generality of employment of the particular organs of which nature makes use in the interior organization of animals, we per¬ ceive that the organs of digestion occupy the foremost rank,—those of circulation the last. We have thus an or¬ der of value or precedence, in regard to these important organs, capable of regulating, in doubtful cases, the pre¬ ference which one relation merits over another. Second principle.—Between two different modes of the same organ, or system of organs, that which is most ana¬ logous to the mode employed in a superior or more com¬ posite and complete organization, merits the preference in the relations which it exhibits. If, for example, we desire to employ a relation afforded by the organs of respiration,— to judge of the preference which it deserves over that of¬ fered by other organs,—we are obliged, according to the principle above established, to keep in view the following consideration:—Although the system of organs provided for respiration is very widely employed in the organization of animals, since, with the exception of the polyped and infusorial classes, all the rest possess a respiratory sys¬ tem, yet the mode of that system not being the same in all the classes by which it is exercised, we assign a higher value to the true lung than to the branchia, to the latter than to the aeriferous tracheae, and to these than to the aquiferous tracheae. According to this view, we may judge whether the mode of respiratory organs of which we wish to employ the relation, is sufficiently high in value to per mit our yielding to it a preference over a relation deduced from some other kind of organs. The fifth hind of relations embraces the particidar rela¬ tions observable between the modified parts. It requires that, among the parts compared, we should discriminate between that which is due to the real plan of nature, and that which pertains to the modifications which that plan has experienced from accidental causes. Thus this class of relations is derived from parts which, considered sepa¬ rately in different animals, are not in the state in which they ought to be, according to the plan of organization to which they belong. To judge of the degree of import¬ ance which ought to be accorded to a relation, and the preference which it deserves over another, it is a matter of no slight consequence to distinguish if the form, the in¬ creased or diminished development, or even the entire disappearance of the particular organs under consideration, belongs to the plan of organization of the animals subject¬ ed to such modifications; or whether the state of these organs is not rather produced by a modifying and deter¬ minable cause, which has altered or annihilated that which nature had executed, and would have maintained, but for the influence of that later cause. “ For example,” says' Lamarck, “ it would have been impossible for nature to furnish a head to the infusoria, the polypi, or the radiata, &c.; for the condition of these bodies, and the degree of their organization, did not permit it; and it was only on arriving at the class of insects that a genuine head could be supplied.” Now, as nature never retrogrades in her operations, we naturally expect, that when once arrived at the formation of insects, and consequently of heads, the recipients of the special senses, all animal organizations 176 ANIMAL KINGDOM. Animal superior in composition to that of insects will also exhibit Kingdom, these organs. This, however, is not universally the case; because no distinct head is observable among the annehdes, the cirrhipedes, and many mollusca, all of which are gene¬ rally regarded as superior to the class of insects. Une cause etrangere a la nature, en un mot, une cause modih- ante et determinable, s’est done opposee a ce que les am- maux cites soient pourvus d une veritable tetc. ( nun. sans Vert, tome i. p. 363.) And that cause appears to have operated sometimes by hindering to a greater or less ex¬ tent the development of that part of the body, and at other times by effecting its complete destiuction. e find the same thing in regard to eyes and teeth, and to various other parts both of the internal and external struc¬ ture; because a modifying cause has had the power to alter, enlarge, diminish, or even to effect the total disappeai- ance of these organs. We may perceive, then, that the relations obtained from the consideration of changed or modified parts must be of very inferior value to those furnished by the same parts existing in a state conform¬ able to the plan of nature. Hence results the following Principle.—Whatever nature has directly formed de¬ serves a pre-eminence in value over that produced by a fortuitous cause, which has modified the work of nature; and in the choice of a relation to be employed, we should assign the preference to every organ, or system of organs, which we find existing as it ought to do according to the plan of organization of which it forms a part, over that or¬ gan, or that system of organs, of which either the condi¬ tion or existence has resulted from a modifying cause, ex¬ traneous to original nature. When two different organs, between which a choice is to be made, are both found to be changed or altered by a modifying cause, the preference should be given to that which is least removed by such a modification from the condition in which it would have existed according to the plan of organization of which it formed a part. The third question proposed by M. Lamarck is as fol¬ lows :—What disposition or mode of arrangement should be given to the general distribution of animals, so as to render it conformable to the order followed by nature in the production of these beings ? To resolve this question, we must also endeavour to deduce some principle from nature herself, with a view to such conformity; for if we were to determine the general distribution of animals ac¬ cording to the progression which exists in the animal or¬ ganization, it appears that we might, in that progression, proceed with as much reason from the most composite to the most simple, as from the most simple to the most com¬ posite. Such a proceeding, however, could not rest on a proper basis; for we shall find that nature, consulted in the order of her operations in regard to animals, indicates the following principle, which excludes all arbitrary se¬ lection :—Nature always operating in a gradual manner, and consequently never producing animals otherwise than suc¬ cessively, has obviously proceeded in such a production from the most simple towards the most complex. We ought, therefore, in our general distribution, to imitate the order which nature herself has followed. “ J’ai, en effet, montre,” says Lamarck, “ dans ma Philosophic Zoology j (tome i. p. 269), que, pour rendre la distribution generaje K des animaux conforme a fordre qua suivi la mature en^ produisant toutes les races qui existent, d fallait proceder du plus simple vers le plus compose,—e’est-a-dire, qu’il etait necessaire de commencer cette distribution par les plus imparfaits des animaux, et les plus simples en orga- nisation, afin de la terminer par les plus parfaits, par ceux qui ont 1’organisation la plus composee.” He further ob¬ serves, in his Anim. sans Vert. “ Get ordre est le seul qui soit naturel, instructif pour nous, favorable a nos etudes de la nature; et qui puisse, en outre, nous faire connoitre la marche de cette derniere, ses moyens, et les lois qui regissent ses operations a leur egard.” Although we may find it less pleasant or conformable to our habitual taste to present at the head of the animal kingdom crea¬ tures of the most limited perceptions, excessively minute in size, and of scarcely any consistence in their parts,- yet as in all things it is necessary to consider the end in view, and the means which conduct towards it, Lamarck is of opinion that the arrangement established by usage in the distribution of animals is precisely that which leads us away from the point in view, as well as the least favourable for our instruction, and that it opposes the greatest number of obstacles in the way of our perceiving the plan, the order, and the means employed by nature in her operations concerning the animal world. If in the study and examination of living bodies we had no other object in view than to distinguish the one from the other by characters deduced from their external forms—and if we were not desirous to regard their various and wonderful faculties otherwise than as simple matters of amusement, not altogether unfitted to excite the cu¬ riosity of a leisure hour—then the most ordinary and arti¬ ficial system would suffice ; for in that case it would be useless to occupy ourselves with researches concerning the affinities of animals, or to study their internal structure. But all naturalists are now agreed regarding the high im¬ portance of these affinities, and the necessity of holding them ever in view in our general arrangement of the ani¬ mal kingdom. The bat is no longer classed with birds merely because, like them, it wings its way -through the air; nor are seals or whales regarded as fishes because the dense nature of the medium which they inhabit re¬ quires a somewhat analogous form; neither are cuttle¬ fish and polypi confounded together, though the mouth of each is surrounded by numerous arms. We have dwelt at greater length than we intended on the system of Lamarck, or rather on the views by which he seeks to illustrate the principles on which his system professes to be built. Though occasionally prolix, and sometimes rather obscure, his observations, on the whole, are well deserving of an attentive consideration. Like most of his countrymen, he is unfortunately more re¬ gardful of secondary causes, and more anxious to illus¬ trate their fitness and sufficiency, than he is ready to ac¬ knowledge the source from which they spring, or to ad¬ mire the wisdom and beneficence of their providential in¬ stitution.1 1 ,U The doctrine of Epicurus, that the Deity concerns not himself with the affairs of the world or its inhabitants, which, as Cicero has judiciously observed (De Nat. Dear. lib. i. ad calcem), while it acknowledges a God in words, denies him in reality, has furnished the original stock upon which most of these bitter fruits of modern infidelity are grafted. Nature, in the eyes of a large proportion of the enemies of revelation, occupies the place, and does the work, of its great Author. Thus Hume, when he writes against mi¬ racles, appears to think that the Deity has delegated some or all of his powers to nature, and will not interfere with that trust (Essays, xi. 75); and, to name^ no more, Lamarck, treading in some measure in the steps of Robinet (who supposes that all theh ^ of the animal kingdom, in which nature gradually ascends from low to high, were experiments in her progress towards her great am ultimate aim, the foimation of man—Darclay cm Organization, &c» 263), thus states his opinion:—u La nature, dans toutes ses opeft* tions, ne pouvant proceder que graduellement, n’a pu produire tous les animaux a la fois ; elle d’abord n’a forme que les plus simple*' et passant de ceux-ci jusques aux plus composes, elle a etabli successivement en eux differens systemes d’organes particuliers, k5,i al We shall now proceed to the system of another natu™ collection of the oral demonstrations of his great master Animal jom. alist, also highly accomplished in the science, M. Du- we were originally indebted for the publication of the Kingdom. * .Ticri]} the friend and pupil of Baron Cuvier, and to whose Lemons d’Anatomic Comparee. ANIMAL KINGDOM. 177 Table of the Classification of Animals, according to the System of Dumeril. nternally articulated; vertebrated; •Ixternally articulated; invertebrated; \Tot articulated; oviparous; articulated members 'with mammae ; viviparous 1. Mammifeile. 1 without mammae ; filings; /feathers, wings 2. Birds. ’ & ^neither leathers nor wings 3. Reptiles no lungs; branchiae 4. Fishes. /tracheae 5. Insects. /branchiae 6. Crustacea. ^no articulated members 7. Worms. distinct respiratory organs ; vessels 8. Mollusc a. no respiratory organs; no vessels 9. Zoophytes. The ensuing tabular view exhibits the classification His nomenclature is no doubt intimately connected with imposed by M. de Blainville. We shall leave the reader his views of the structure and physiology of animals, and o judge for himself of the propriety of introducing so is highly approved of and adopted by many competent uany new appellations for groups constructed long ago. judges of the science. Synoptical Table of the Primary, Secondary, Tertiary, and Quaternary Divisions of the Animal Kingdom, denominated Sub-Kingdoms, Types, Sub-Types, and Classes, by M. de Blainville. From the Principes dAnatomic Comparee of that author. ANIMAL KINGDOM. /. Sub-Kingdom. tegular or Autio- morfhous Ani¬ mals. Art i ozo aires. articulated. Type /. interiorly. OSTEOZO AIRES. Type II. exteriorly. Entomozoaires, or articulated ani¬ mals. With appendages, I. Sub- Type. ’ Provided with mammae and ...hairs.. Viviparous. - II. Sub-Type. Without mammae. Oviparous, ffeathers Provided with j naked skin..!' ffins f 3 pair 4 pair..... f articulated to 1 the number of I not articulated [none 5 pair. variable 7 pair = the segments Classes. I. Piliferes or Mammalia. 2. Penxiferes or Birds. 3. Squammiferes or Reptiles. 4. Nudipelliferes—Amphibia. 5. PlNNIFERES or FlSHES. 6. Hexapodes. 7. OcTOPODES. 8. Decapodes. 9. Heteropodes. 10. Tetradecapodes. 11. Myriapodes. 12. Chitopodes. 13. Apodes. sub-articulated.. Type III. Malentozoaires or Molluscarticules. Type IV. not articulated Malacozoaires The head -f f‘*s/1li1.ch" Molluscous animals. II Sub-Kingdom. Radiated or Actinomorphous Animals. Actinozoaires. - ( not distinct... j 14. Nematopodes. (15. Polyplaxiphores. IfJ. Cephalopiiores. 17- Acephalophores. sub-radiated 18. Annelidaires. '19. Ceratodermaires normal or true.. 20. Arachnodermaires. - 21. XoANTHAIRES. 22. Poi.YPIAIRES. ^23. ZoOPlIYTAIRES. | II/. Sub-Kingdom. • , „, _ « 24. Sfongiaires. Irregular or Heteromorpiious Animals. Heterozoaires. 25. Monadaires or Moleculaires. I 26. Dendrolitiiaxres. multiplies, en a augmente de plus en plus I’energie, et les cumulant dans les plus parfaits, elle a fait exister tous les animaux connus ivec 1 organisation et les facultes que nous leur observons.” (Anim. sans Vertebr. i. 123.) Thus denying to the Creator the glory of omung those works of creation, the animal and vegetable kingdom (for he assigns to both the same origin, ibid. 83), in which his glo- nous attributes are most conspicuously manifested; and ascribing them to nature, or a certain order of things, as he defines it (214)—■ s urnd power, that operates necessarily (311), which he admits, however, to be the product of the will of the Supreme Being (216). ^remarkable, that in his earlier works, in which he broaches a similar opinion, we find no mention of a Supreme Being. (See his . ys unc des Animaux sans Vertebra^ Discours d'Ouvcrturc.) Thus we may say, that, like his forerunner Epicurus, Re tollit, dum oralkme rttnquit Dcum. But though he ascribes all to nature, yet, as the immediate cause of all the animal forms, he refers to the local circum- s .Hues, wants, and habits of individual animals themselves : these he regards as the modifiers of their organization and structure (162). VOL. HI. 2 kingdoms, viz. 1. Acrita, composed of the infusoria, the “polypi, the corallines, the tcenice, and the least organized of the intes¬ tinal worms. 2. Radiata, containing medusa:, star-fish, sea-urchins or echini, and others. 3. Annulosa, consisting of insecta, arachnida, and Crustacea. scripture. Metaphysicians enumerate seven principal ope¬ rations of the mind, musicians seven principal tones, and opticians seven primary colours. In scripture, the abstract idea of this number is, completion, fullness, perfection. Mr Kirby seems to think that Mr Macleay’s quinaries maybe found resolvable into septenaries, in consequence of future investigations.2 We shall enter at greater length into a detailed expo- Fo show the absurd nonplus to which this his favourite theory has reduced him, it will onlv he necessary to mention the individual instances which, in different works, he adduces to exemplify it. In his Systeme he supposes that the webfooted birds (Anseres) acquir¬ ed their natatory feet by frequently separating their toes as far as possible from each other in their efforts to swim. Thus the skin that unites these toes at their base contracted a habit of stretching itself; and thus in time the web-foot of the duck and the goose was produced. I he waders (Grallce), which, in order to procure their food, must stand in the water, but do not love to swim, from their constant efforts to keep their bodies from submersion, were in the habit of always stretching their legs with this view, till they giew Ion" enough to spare them the trouble! ! ! (13). How the poor birds escaped drowning before they had got their web-feet a*)( long legs the author does not inform us. In another work, which I have not now by me, I recollect he attributes the long neck of the camelopard to its efforts to reach the boughs of the mimosa, which, after the lapse of a few thousand years, it at length ac¬ complished ... In Ins last work he selects as an example one of the Moluscac, which, as it moved along, felt an inclination to ex¬ plore by means of touch the bodies in its path : for this purpose it caused the nervous and other fluids to move in masses successive¬ ly to certain points of its head, and thus in process of time it acquired its horns or tentacula ! ! (A nim. sans Verte.br. i. 188.) Hb grievous that this eminent zoologist, who in other respects stands at the head of his science, should patronize notions so confessedly absurd and childish. (Introduction to Entomology, by Kirby and Spence, vol. iii. p. 349 ) 1 See Horae Entomologies, and Kirby & Spence’s Introduction to Entomology vol iii p 12 - Introduction to Entomology, vol. iii. p. 15. The quinary system, in its application 'to insects and other annulose animals, is pretty fully developed by its ingenious author in his Horae Entomologicas, already more than once referred to. An excellent paper by Mr Vigors, on the classification of birds, m accordance with the same system, will be found in the 14th volume of tlie Transactions of the Juiiuicbcm society. ANIMAL KINGDOM. i jaal jition of the quinary system, of arrangement under the f:»irarticles Entomology and Ornithology. Cuvier divides the animal kingdom into four principal tranches. Setting aside all accessory and artificial cha- •acters, he proceeds upon the consideration of the essen- ial structure of animals, and thus deduces four great groups or separate types of form, to one or other of which ill the minor divisions may be ultimately referred. In the first of these forms the brain, and the great cen- ral trunk of the nervous system called the spinal marrow, ire protected by strong bony coverings—the cranium and /ertebral column. To the sides of that column are at- ached the ribs and the bones of the anterior and poste- •ior members. All the classes of this primary division are irovided with red blood, a muscular heart, a mouth with wo horizontal jaws, and special organs of vision, hearing, aste, and smell, placed in the head or upper and anterior portion. They have never more than four members ; their sexes are always separate; and they nearly resemble each >ther in the distribution of their medullary masses, and he principal branches of their nervous system. On ex- tmining more narrowly the constituent parts of the classes vhich compose this great assemblage, it is easy to disco- /er many striking analogies both of form and structure, ;ven in those groups which are most distantly related to ;ach other; and from the human species to the last of ;he fishes there exists an obvious conformity to the same general plan. The name of Vertebrated Animals is lestowed on this division, on account of their being pos¬ sessed of a vertebral column or back-bone. The following ire the principal groups or classes into which it is further livisible. FIRST PRIMARY DIVISION: ANIMALIA VERTEBRATA. Class ls£. Man, mammiferous land-animals, and ce- 'tacea. Class 2d. Birds. Class 3r?. Reptiles. Class kth. Fishes. The second great division possesses no skeleton. The nuscles are attached to the skin, which forms a soft con- ractile envelope; and many of the species are protected iy hard coverings, commonly called shells, which are sup¬ posed to occupy in the cutaneous system of this form of animal life the same station as the mucous membrane of the preceding division. The nervous system is contained along with the viscera within this general envelope, and is composed of many dispersed portions, of which the principal, placed above the oesophagus, may be regarded as representing the brain. Of the four special senses it is impossible to discover the organs of more than two, taste and sight; and even of these the last is frequently want¬ ing. The organs of hearing are visible only in one family. The system of circulation is however complete; there are particular organs for the performance of respiration ; and the functions of digestion and secretion are almost as complicated as in the vertebrated classes. The subdivi¬ sions of this second form are called Molluscous Ani¬ mals ; and although the external configuration of their parts does not exhibit the same agreement as that of the vertebrated classes, there is always a corresponding re¬ semblance in their essential structure and functions. The following are the classes of this branch of the animal kingdom. SECOND PRIMARY DIVISION: ANIMALIA MOLLUSCA. Class Itf. Cephalopoda, e. g. cuttle-fish, nautili, belem- nites, argonauts, &c. 179 Class 2d. Pteropoda. Genus Clio, &c. Animal Class 3d. Gasteropoda. Slugs, snails, and numerous Kingdom, groups of turbinated shells. Class kth. Acephala. Oysters, mussels, and other bi¬ valve shells, &c. Class 5th. Brachiopoda. Terebratulae, &c. Class &th. Cirrhopodes. Barnacle shells, &c. Lepas and Triton. (Linnaeus.) The third great preponderating form is represented by insects and other analogous classes. Their nervous sys¬ tem consists of two long ventral or sublateral chords, which swell out at intervals into knots or ganglia. The first of these ganglia, placed above the oesophagus, is analogous to the brain, although it does not exceed in size and scarcely in importance the ganglia of the lengthened cords, with which it communicates by means of a ring which embraces the oesophagus like a collar. The general covering of the body in this division is sometimes hard, sometimes soft, and is divided into segments by a certain number of transverse incisions. The muscles are always attached to the interior, and the body is usually, though not universally, provided with articulated members. It is among the classes of this form that we begin to perceive the passage from a system of circulation in closed vessels called arteries and veins, to nutrition derived from imbibi¬ tion ; and a corresponding passage from respiration in cir¬ cumscribed organs to that performed by tracheae or air- vessels, distributed over the whole body, is likewise ob¬ servable. The organs of taste and sight are the most distinct in this branch ; the organ of hearing is apparent only in a single family, although we can scarcely doubt that the sense exists in others in which the organ has not been ascertained. The following classes are ranked under this great form. THIRD PRIMARY DIVISION: ANIMALIA ARTICULATA. Class 1st. Annelides. Serpulse, nereids, leeches, earth¬ worms, the hair-eel, &c. Class 2d. Crustacea. Crabs, lobsters, shrimps, &c. Class 3d. Arachnides. Spiders, scorpions, mites, &c. Class Uh. Insecta. Beetles, flies, butterflies, &c. In the three preceding divisions, the organs of move¬ ment and of sensation are disposed symmetrically on both sides of an axis, with an anterior and posterior portion dif¬ fering from each other. Among the zoophytes, which form the last great division, the organs are usually disposed in a radiated form. They approach the nature of plants in the extreme simplicity of their structure. They have no distinct nervous system, nor organs of the special senses; and it is barely possible to detect in a few of the species some slight vestige of the circulating system. Their res¬ piratory organs are almost always on the surface of their bodies. The greater proportion of the classes exhibit no other intestine than a sack or caecum, and the composition of the last groups of all presents only a homogeneous pulpy mass, sensible, and endowed with motion. From a consideration of their most usual forms, the classes of this order are named Radiated Animals. They are as follows:— FOURTH PRIMARY DIVISION: ANIMALIA RADI AT A. Class ls£. Echinodermata. Star-fish, sea-urchins, &c. Class 2d. Entozoa. Intestinal worms. Class 3d. Acalephae. Sea-nettles, actiniae, medusae, &c. Class \th. Polypi. Corals, madrepores, sponges, &c. Class 5th. Infusoria. Infusory and other microscopi¬ cal animals. 180 ANIMAL KINGDOM. Animal Such are the great outlines of a system which, consi- Kingdom dered in its generality, is certainly the most satisfactory II which has vet appeared. Particular departments may Animal- - - . 1 - - • ’ 1 cule. have been filled up, modified, and perhaps improved by , ingenious observers, sedulous within a limited sphere (and of these ameliorations we shall be careful to avail our¬ selves when we come to enter upon a detailed view o each of the classes of the animal kingdom) ; but the con¬ struction and position of the principal groups, their leal as well as relative characters, are developed in the system of the great French anatomist, in a manner more clear and accordant with nature than in any other yet promulgated. We shall therefore in the course of this work adhere, with some slight transpositions, the reasons for which will be stated in their proper place, to the classes of Baron Cuvier. The greater extent and importance of some of these, in comparison with others, will induce us to bestow more attention and a larger space to their illustration; and as certain of the primary divisions, such as the Mol- lusca and Radiata, contain a greater number of classes, if not of less importance, at least by no means so strongly characterized or contradistinguished from each other as are those of the vertebrated tribes, we shall, in presenting the history and nomenclature of such classes, group them together in such a manner as to exhibit them to the reader either under one of the great primary divisions, or as an intermediate subdivision, containing one or more classes. For example, the article Mollusca of this work will present consecutively under a single head the history and classification of the six classes contained in the se¬ cond primary division so named;—but the four classes of vertebrated animals will be each discussed in a separate treatise. Thus mammiferous animals, birds, reptiles, and fishes, will form the articles Mammalia, Ornithology, Reptilia, and Ichthyology. The Classes of the third primary division, viz. Annelides, Crustacea, Arachnides, and Insecta, will (with the exception of the first, referred to Helminthology) likewise be treated of under distinct heads, in the alphabetical order, of the following terms Crustacea, Arachnides, and Entomology. In regard to the fourth primary division, that of the radiated ani- mals, commonly called Zoophytes, the first class, named Echinodermata, will be treated of separately under its own title; the second class, Entozoa, which contains the intestinal worms, will be grouped with the Annelides or red-blooded worms (as above excepted from the third primary division) ; and these two classes will be treated of together under the article Helminthology. The re¬ maining classes of the Animalia Radiata, that is to say, the Acalephae, the Polypi, and part of the Infusoria, as they form the last links of the animal kingdom, will come to be discussed with greater propriety at the concluding stage of this work, under the head of Zoophytes. Finally, that portion of the infusorial class which we have excepted in the above distribution will be found described in the pre¬ sent volume under the word Animalcule. This com¬ pletes the exposition of our zoological system. The following enumeration exhibits a view of the terms under which the principal subjects of zoology will be ex¬ plained and illustrated in the course of this work. Systematic Arrangement. Mammalia. Ornithology. Reptilia. Ichthyology. Mollusca. Crustacea. Arachnides. Entomology. Echinodermata. Helminthology. Zoophytes. Animalcules. Alphabetical Arrangement Animalcules. Arachnides. Crustacea. Echinodermata. Entomology. Helminthology. Ichthyology. Mammalia. Mollusca. Ornithology. Reptilia. Zoophytes. (t.) Animal Magnetism. See Magnetism, Animal. ANIMALCULE. Animalcule, a diminutive term (from the word ani¬ mal), applied by naturalists to those minute beings which become apparent in various fluids when subjected to the microscope. They were named infusory animals {Infuso¬ ria) by Muller, one of the most celebrated observers in this department of zoology; and the appellation, however inapplicable, now occurs in the majority of scientific pub¬ lications. Of course it applies with propriety only to such species as are developed through the medium of infused substances. Now we know, that of 400 species of Infuso¬ ria (commonly so called) described by Muller himself, not a sixth part were observed in any kind of infusions ; whilst the remainder inhabited the most translucent waters, and speedily died when placed in impure or corrupted liquids. Even the word animalcule (or little animal) does not con¬ vey a positive or sufficiently restricted idea in relation to this particular class; because mites and certain polypi are extremely minute in their dimensions, and equally re¬ quire the aid of microscopical investigation; and thus the term microscopies {microscopiques), recently proposed by M. Bory de St Vincent, is not less faulty than its prede¬ cessors. The size of an animal, in fact, bears no essential relation to the other conditions of its organization; and therefore we cannot infer its nature with any certainty from a knowledge of its dimensions. At the same time it must be admitted, that the most simply organized, both of plants and animals, are also the most minute; and thus the Infusoria may be regarded as possessed of certain cha¬ racters in common. We here adopt the word Animalcule, chiefly because it is the most familiar to the English reader. The subjects of our present observations may be thus defined :—Animals invisible to the naked eye ;1 more or less translucent; unprovided with members (the caudal, and other appendices, with which certain species are furnished, being scarcely regardable as such); no perceptible eyes; contractile in whole or in part; endowed with the sense of touch ; deriving nourishment by absorption; generation (when not apparently spontaneous, and consequently in¬ comprehensible) effected by division, or by the emissim oj gemmules or oviform bodies ; inhabitants of liquids. They are the smallest and most simple of living creatures, but not less perfect than the other tribes; for though they possess the fewest faculties, their means are in every way adequate to their wants, and their vital energies propor¬ tioned to their sphere of enjoyment. Among microscopical animals we find many species which, in their aspect and structure, present no analogy 1 Hie Volvox gldbator, and a few others, which are just discernible without the aid of a microscope, form exceptions to the above character. ' ’ ANIMALCULE. 181 0 other forms of animal life: they are merely moving mlecules of the simplest organization, the exact nature of vhich it is sometimes difficult to determine, and which ivolve in deeper obscurity the mysterious line of demar- ation by which we so often seek in vain to separate the nimal from the vegetable kingdom. If, however^ the •ue distinction between plants and animals consists chiefly i the irritability and power of contraction possessed by he latter, then the Infusoria, which are strongly endowed -ith these attributes, are indeed so far removed from the e^etable kingdom, that the name of Zoophytes, or animal lants, is inapplicable to the class to which they belong, a the extreme simplicity of their structure, they no doubt resent some analogy to the least complicated tribes of lants, such as the algae and others; but it is a mere ana- )e not now classed among the animalcular tribes. He divides the Class of Animalcula Infusoria as fol- Animal- lows : cule. Order I.—No External Organs. * Thickened. Genus 1. Monas; body punctiform. 10 species. 2. Proteus; body variable. 2 species. 3. Volvox; body spherical. 12 species. 4. Enchelis; body cylindrical. 27 species. 5. Vibrio; body elongated. 31 species. * * Membranous. 6. Cyclidium; body oval. 10 species. 7. Paramccdum; body oblong. 5 species. 8. Kolpoda; body sinuous. 16 species. 9. Gonium; body angular. 5 species. 10. Eursaria; body excavated. 5 species. Order II.—External Organs. * Naked. 11. Cercaria; smooth, tailed. 22 species. 12. Trichoda; haired or ciliated. 89 species. 13. Kerona; with horny appendices. 14 species. 14. Himantopus; with cirrated appendices. 7 spe¬ cies. 15. Leucophra; the entire surface ciliated. 26 species. 16. Vorticella; orifice ciliated. 75 species. * * Furnished with a Shell. 17. Brachionus; orifice ciliated. 22 species. In the year 1815 Lamarck published the first part of his Animaux sans Vertebres, a work which forms an epoch in the history of the inferior tribes. In this signal publication the author adopts a different course from that usually followed by systematic writers; and pursuing an ascending rather than a descending scale, he commences with the lowest and least complex species, viz. the Infu¬ soria. From this class, however, he rejects all those spe¬ cies in which any kind of complication of organs is appa¬ rent. All the genera so distinguished are referred by him to the first order of the second class of the animal king¬ dom, called Polypi, under the title of Polypi ciliati; and the true and restricted Infusoria are thus defined : Micro¬ scopical animals, gelatinous, transparent, polymorphous, con¬ tractile ; having no distinct mouth, nor constant, determin¬ able, internal organs ; generation jissiparous or sub-gemmi- parous. The genus Kerona, it may be further remarked, is in this system united to the Himantopus of Muller, while the genus Cercaria of that author is divided into two. Thus the class Infusoria of Lamarck, composed of two great sections, the naked and the appendiculated, may be said to correspond to the first 14 genera of the Danish naturalists. Cuvier, in the Eegne Animal (1817), places the Infuso¬ ria as a part of his fourth great division, the zoophytical or radiated animals. The term radiated was originally bestowed on a numerous tribe of animals, such as the Asterias and others, on account of their arms or tentacula being extended in a radiated or star-like form; but it certainly applies unfitly to the true Infusoria of Lamarck, which possess nothing resembling a radiated structure. It cannot, however, be always expected that a general term of wide import should apply with etymological accuracy to every part of the extensive series which it is intended to embrace. Cuvier then divides his Infusoria into two orders, Les Rotiferes, and Les Infusoires homogenes, the former of which correspond to the ciliated polypi of La- [g2 A N I M A Animal- marck, the latter to the Infusoria properly so called of cule. that author. , T „ According to M. de Blainville, the class Infusoria can scarcely be regarded as established upon a natural foun¬ dation. The organization of its component tribes is so various as to lead to the belief that a more precise knowledge would show that several of those tribes belong to different types of the animal kingdom. Some, as the genus Brachionus, are symmetrically formed both as re¬ gards their bodies and appendages, and are protected by a horny or crustaceous covering. Others, as Vibno, 1 a- rammeium, &c. have the body elongated,^ depressed, vei - miform, and without appendages. A third division ex¬ hibit a radiated structure, as for example the Vorticellae, which, however, we have already stated, are now seldom classed among the Infusoria. Many genera, such as Pio- teus, Volvox, Monas, are amorphous, or without deter¬ minate form, and cannot be referred to any other known type of the animal kingdom. They are regarded by many as the elementary molecules of all animal life, and in their structure no other than the cellular tissue is observable. They may be said to be dependent on external circum¬ stances, instead of being able, like, other animals, to mo¬ dify or control them ; and their usually spherical form is the necessary result of an equal pressure of w'ater on all sides of a frail and yielding texture. M. de Blainville considers the genera Brachionus, Urcelaria, Cercaria, Fur- cularia, Kerona, Trichocerca, and Himantopus, as belong¬ ing to the type of Enlomozoaires or articulated animals, and especially to the class Heteropoda, order Entomos- traca. Many species of Vibrio he regards as Apodes, as well as ParamcEcium and Kolpoda. Other species of the genera Vibrio and Cyclidium ought rather to be ranged with the Blanaria; and in the genus Leucophra there is even a species which M. de Blainville is inclined to look upon as an Ascidia ! Finally, the genera Gonium, Proteus, Volvox, and Monas, if they are really animals, appear to form a distinct type, which may be called Amorphes or Agastraires; so named from the circumstance of their hav¬ ing neither determinate form nor reduplication of the ex¬ ternal envelope for the formation of a stomach, as in all other true animals. Such is a brief exposition of the views of one of the most distinguished physiological inquirers of the present day. It may serve, if for nothing more, at least to show the unsettled state of opinion concerning these extraor¬ dinary creatures. In regard to this, however, we may rest assured that, in the future progress of science, the class Infusoria, as at present constituted, will suffer an entire dismemberment, and its component parts will be referred to various groups of the animal kingdom, some of them widely distant from each other. In the year 1826 a full and most elaborate classification of microscopical animals was given to the world by M. Bory de St Vincent. As it is the singular mode of ex¬ istence of animalcular beings, their general economy in the field of nature, the actual conditions of their organi¬ zation, and the state of their limited faculties so far as these can be ascertained, with which we are chiefly in¬ terested—so, in our systematic view of this extraordinary class, we shall merely present to our readers the characters of the principal genera, and of a few of the most remark¬ able species which they contain. But, as some may be desirous to possess at least a sketch of the full extent and condition of this intricate subject, we have constructed the accompanying tabular scheme of the orders, families, and genera of microscopical animals, according to the views of M. Bory de St Vincent, the latest and most assiduous writer on this department wuth whose labours we are ac¬ quainted. We have thought it advisable to retain the L c u L E. terms of the original language, lest, by inadvertence or - misconception on the part of the translator, any additional obscurity should rise around a subject already sufficiently^- encumbered. {See Tabular View on the opposite page.) The order Gymnodes of Bory de St Vincent nearly con responds to the entire class Infusoria of Lamarck; and although the observations by which he illustrates his ar¬ rangement partake of the accustomed defects of the French philosophy, the facts which he details, if not the theore- tical views which he inculcates, are worthy of an atten- tive consideration. These mysterious creatures are ob¬ served to swim with astonishing rapidity; and although their bodies are usually diaphanous, it has hitherto proved impossible, even by the aid of the most powerful glasses, to ascertain by what natural mechanism these movements are effected. They direct their courses by a discretion¬ ary power, in one direction rather than another, avoiding and turning round opposing obstacles, according to the necessities of the case—discerning, as the process of eva- poration proceeds, the points in which they may prolong their existence, and flocking in crowds to those places where they are best screened from the overpowering bril- liancy of the reflecting mirror. They thus appear to pos¬ sess volition, which we are accustomed to regard as a result dependent on the faculties of perception and com¬ parison. The principal obstacle to our understanding the essen¬ tial nature of animalcules results from their want of a nervous system, which, in ourselves, and in all the inter¬ mediate classes of the nature of whose' consciousness we have even a vague idea, we regard as the sine qua non of sensation and intelligence. Voluntary motion without muscular action is also a circumstance which we cannot very clearly comprehend. But as there may be “ more things in heaven and earth than are dream’t of in our philosophy,” we must not reject facts, that is to say ap¬ pearances which present themselves under the same de¬ terminate and uniform aspect to various unprejudiced ob¬ servers, merely because they do not coincide, or may pos¬ sibly controvert or interfere, with a previous hypothesis. On the other hand, the extreme softness of texture, and excessive minuteness, of most of the animalcular species, render anatomical investigation almost impossible; and naturalists may have erred in supposing the absence of what they are merely unable to perceive and demon¬ strate. It is in truth impossible to discover any traces of the nervous system, even among several tribes of animals in other respects much more highly organized than the sub¬ jects of our present inquiry. Trembley’s examination of the Polypus threw no positive light upon the matter;nor did Gade’s dissections of the larger Medusae enable him to discover either muscular or nervous fibres. According to M. Bory de St Vincent, the nervous system is one of the last to be developed. To the perfect simplicity of the Monads, the first perceptible addition is that of a central cavity, or rudimentary intestinal sac, which we find to occur even before the existence of a mouth. Next appears an opening to this canal, which serves both for the reception of nourishment, and the rejection of excrementitious parts where such exist. The hairs andcirrated appendages which ensue in still more complicated species have been regard¬ ed as the early rudiments of the respiratory system; and even a heart, or central organ of a circulating fluid, is partly developed before the appearance of any nervous chords. The earliest, most general, and perhaps the only indispensable function of animal life, is that of nutrition. But the materials of nutrition are so different, and thejf modes of reception so various, that the exercise of this 184 animalcule. Animal¬ cule. function by no means necessitates the existence of a mouth, a stomach, or an alimentary canal; for an increase ’ of parts may be effected even through the medium ot imponderable or elastic fluids, and by imperceptible and superficial pores. . The exterior of an infusory animal may be compared to the interior of one of the higher classes, in which nutri¬ tion is carried on by the reception of the chyle by the absorbent pores. These pores are external among t le Infusoria, and the process of absorption is with them ana¬ logous to that of plants, in which there is a direct reception and appropriation of fluids from the earth and air, wit i- out any previous preparation in a central cavity or sto¬ mach. Zoophytes in general have indeed been called the cryptogamia of the animal kingdom. According to Cams, the Infusoria ought to be regarded merely as little cells, partially filled with lymph, and possessed of the powers of nutrition and locomotion; and thus the infinite changes and variations perceptible in their forms may be supposed to be produced by the various degrees in which this fluid is collected at one or other of the points of theii bodies. In the opinion of that anatomist, a more complete deve¬ lopment of the organs of motion, and indeed of the whole organization, is inseparably united with the appearance of a distinct nervous system. This may be true as a gene¬ ral rule, but not as a universal principle ; for the Medusa has more apparent voluntary motion than the Asterias, though the former is destitute of those nerves which in the latter make their first appearance in the shape of a pale thread-like ring surrounding the oesophagus. It is this ring around the upper extremity of the alimentary canal which, in the molluscous and articulated classes, we shall afterwards find to constitute the most uniform and most essential portion of the nervous system. The Me¬ dusae, just referred to, being almost of the same specific gravity with water, are easily carried by currents, and moved about from place to place by the action of the waves, or even (as in the case of Holothuria physalis) by the winds; but Carus and other writers have assuredly erred in doubting that they execute a voluntary locomo¬ tion ; for that they do so in a very decided and graceful manner must be obvious to all who have attended to these animals in their native haunts along the shores, or among the land-locked waters of the beautiful firths of Scotland. The mysteries revealed by the glasses of Leeuwenhoeck were at first regarded as beyond belief. The uncertainty of microscopical investigations, in consequence of which so much was supposed to depend on the imagination of the beholder, was alleged against them; and even at an after-period, when men of sober judgment and the most industrious application had confirmed the experience of the indefatigable Dutchman, the wit of Voltaire did not disdain to throw its cutting sarcasm over the disciples of the “ anguilles de la pate et du vinaigre.” We hope it is now admitted, that however frequently those who en¬ deavour to expound the mysteries of nature may fail in their attempts at elucidation, yet that there is nothing in the manifold works of Omnipotent Wisdom which, if duly studied and rightly understood, would not conduce to our wellbeing and happiness; and that a single square inch of water, with its many millions of animalcular atoms, is in truth as wonderful a work of divine intelligence, and as interesting a field for human investigation, as the starry galaxy of heaven. In his tam parvis, atque tam nullis, qua> ratio ! Quanta vis ! quam inextricabilis perfectio ! And if a heathen philosopher (Pliny) has so expressed his almost reverential admiration, is it not to be deplored that A those whose labours might be carried on under the influ. I ence of a purer light, seem as often degraded as exalted^' by the contemplation of their Creator’s works; and, re- ferring all to the powers of nature, or some other inde¬ finite abstraction, refuse to recognise, amid so many won¬ ders, the “ Good Supreme” from whorn these and other mightier wonders have proceeded? It is in the study of the subject with which we are now engaged, and the ana¬ logous pursuits of physiology, where the completion of the most perfect design and the happiest results of super- human forethought are so constantly manifested, that we frequently meet, where we should least expect it, with the sneer of the sceptic, or the impious ridicule of the unbe¬ liever. ‘ How different are the sentiments -of one who combines the piety of the Christian with the genius of the poet and philosopher. “ But about the time of its invention (the invention of the telescope), another instru¬ ment was formed, which laid open a scene no less wonder¬ ful, and rewarded the inquisitive spirit of man with a dis¬ covery which serves to neutralize the whole of this argu¬ ment. This was the microscope. The one led me to see a system in every star ; the other leads me to see a world in every atom. The one taught me that this mighty globe, with the whole burden of its people and of its coun¬ tries, is but a grain of sand on the high field of immensity; the other teaches me that every grain of sand may har¬ bour within it the tribes and the families of a busy popu¬ lation. The one told me of the insignificance of the world I tread upon; the other redeems it from all its in¬ significance ; for it tells me, that in the leaves of every forest, and in the flowers of every garden, and in the wa¬ ters of every rivulet, there are worlds teeming with life, and numberless as are the glories of the firmament. The one has suggested to me, that beyond and above all that is visible to man, there may lie fields of creation which sweep immeasurably along, and carry the impress of the Almighty’s hand to the remotest scenes of the universe; the other suggests to me, that within and beneath all that minuteness which the aided eye of man has been able to explore, there may be a region of invisibles; and that, could we draw aside the mysterious curtain which shrouds it from our senses, we might then see a theatre of as many wonders as astronomy has unfolded, a universe within the compass of a point so small as to elude all the powers of the microscope, but where the wonder-working God finds room for the exercise of all his attributes, where he can raise another mechanism of worlds, and fill and animate them all wdth the evidence of his glory.1 Although we cannot hope to derive the same amuse¬ ment or advantage from the study of each of the animal¬ cular species considered separately, as we do from the consideration of the history of many of the higher animals, yet, in a philosophical point of view, a knowledge of the general attributes of the class presents several highly iro- portant objects; and their obscure origin, their singular organization, and more singular mode of existence, cannot fail to excite our unfeigned wonder and admiration. They can scarcely be described otherwise than by a negation of all those characters which constitute the life, power, and activity of other beings ; they have no head, no eyes, no muscles, no blood-vessels, no nerves, no determinate or¬ gans for respiration, generation, or digestion—and yd they are endowed with life. The animal nature of the Infusoria has indeed been de¬ nied by many; but such is the regular gradation from the most simply organized of the monadal forms to the muc 1 Washnexs's Astronomical Discourses, V). 112. ANIMALCULE. 185 Hi. .more complicated structure of the Polypi, which present, ^ under a remarkable aspect, such unequivocal characters ■of animality, that it is impossible to draw the line of de¬ marcation ; and if we admit the life of the one we can scarcely doubt that of the other. Yet many of the Infu¬ soria appear to present the very lowest conceivable point to which animal life can be reduced. The structure of an animal, the individual existence of vhich is preserved by the absorption of a circumambient luid, and the continuance of whose species is effected by he division or separation of a part of its own body, might, i priori, be supposed to be of the most simple kind. “ We nay rest assured,” observes Lamarck, “ that whenever m organic function is itself unnecessary, the special or- ran by which it is usually performed will not be found to :xist.” It is indeed by considering the nature of the Infu¬ soria that we are enabled to form a proper idea of the amplest condition of animal life; and the invention of he telescope was not of higher importance to the astro- lomer, than that of the microscope to the physiological laturalist. There are few subjects of reflection more in- eresting than the uses which philosophers of an enlight- -ned age have deduced and matured from the scanty snowledge of a barbarous people. Glass, a material known it an early period to the Asiatic nations, and once esti- nated at its weight in gold, has become in the hands of Europeans of more value than the finest gold. Whoever jolished the first lens may be said to have laid the foun- lation of an instrument destined to discover thousands of :elestial worlds above and around us, and an unseen world >f wonders beneath our feet. “ Indeed,” says Cuvier, ! it could not be expected that those Phoenician sailors vho saw the sand of the shores of Boetica transformed by ire into a transparent glass, should have at once foreseen hat this new substance would prolong the pleasures of ight to the old; that it would one day assist the astro- lomer in penetrating the depths of the heavens, and in lumbering the stars of the milky way; that it would lay •pen to the naturalist a miniature world as populous, as ich in wonders, as that which alone seemed to have been ganted to his senses and his contemplation; in fine, that he most simple and direct use of it would enable the in- labitants of the coast of the Baltic Sea to build palaces nore magnificent than those of Tyre and Memphis, and o cultivate, almost under the frost of the polar circle, the nost delicious fruits of the torrid zone.”1 The faculties of the most simple infusory animals, it las been observed, are reduced to such as are common to ill living beings, and to that irritability which results i'om their animal nature; and their bodies are destitute •1 special organs, precisely because their extremely limit¬ 'd faculties neither require nor admit of such organs •eing exercised. The chief interest to be derived from he study ot this class of beings results, according to La- narck, from the view with which such study presents us •1 the ultimate point to which the organization of an ani- nal is capable of being reduced; and, among all the renders of the creation, he regards as the most surpris- ng the existence of animal life in such inconceivably fail and simple bodies as the least complicated of the inimalcular species. It is not, however, to be said that nature was incapable of forming special organs from the materials of these frail gelatinous bodies,” but rather that ie all-wise Author and Ruler of Nature has seen fit to orm them in what to us may appear a destitute and in- -omplete condition, merely because their structure does >ot fulfil those other conditions which, erroneously, we have sought to establish as the indispensable bases of Animal- animal life. They truly show how confined a knowledge cule. our limited faculties enable us to gain of His unlimited powder ; for they not only present no analogy to other more accustomed forms of life, but almost seem to exist in di¬ rect opposition to those laws in accordance with which we “ live, move, and have our being.” Infusory animals, commonly so called, are found in the fresh and saline waters of all countries. They occur both naturally, if we may use the term in a contradistinctive sense, and as the apparent result of infused animal and vegetable substances. According to Leeuwenhoeck, the milt of a cod-fish contains more animalcules than there are individuals of the human race existing on the face of the earth ; and he calculated that 10,000 might be held within the bulk of a grain of sand. The size of many bears the same relation to that of a mite as the dimen¬ sions of a bee do to those of an elephant; and the most powerful microscopes frequently discover nothing more than merely perceptible points in motion. Flour and water made to the consistence of book-binders’ paste, ex¬ posed in an open vessel, and frequently stirred to prevent the surface from growing hard, will in a few days be found to contain millions of animalcules. The thin pel¬ licle which grows on the surface of an infusion of com¬ mon black pepper also produces an innumerable congre¬ gation of minute beings. Of these and others the real origin is still exceedingly obscure; and both Muller and Spallanzani maintained the improbable opinion that they fell from the air. Their subsequent increase or multi¬ plication is obviously effected in different and very sin¬ gular ways. Such as are spherical are usually propa¬ gated by minute portions, which, though they burst from the anterior of the animalcule itself, cannot be called eggs; and such as are of a depressed or flatten¬ ed form continue their race by cuttings or divisions of their own bodies. We first observe a line or groove, longitudinal or transverse, according to the species; and ere long a notch or incision is perceptible at one or other or both of the ends of that apparent line. This notch increases across or longitudinally, till at last a portion is separated or cut off, or the original animal is divided into two, and each assumes the form and nature of their mu¬ tual predecessor. These new objects retain for some time their natural shape, and then in their turn give rise to one or more individuals by a similar separation of parts. Lamarck seems to be of opinion (Philosophic Zoologique, tome ii. p. 120 and 150) that this multiplica¬ tion by division, and that by the emission of gemmules or oviform portions, are modifications of one and the same natural process ;—that substantially each is the result of an extension and separation of parts, which take place when the parent individual has reached the final term of its increase. It is in fact the same excess of nourishment and growth of particular parts that, even in the higher classes, give rise to the germ of separate life, physically considered, although in regard to these the additional process of fecundation is required. It is the new light which may be gathered from the observation of the mi¬ nutest of the animalcular tribes that renders their study both interesting and important to the physiologist; and it is the belief of some, that a persevering study of the history of microscopical animals will one day enable us to withdraw the mysterious veil which still conceals from our view the most important secrets of nature. The systematic arrangement of animalcules which we propose to follow in this place is nearly that of Lamarck, vox., in. 2 A 1 Reflexions sur la Marche actuclle des Sciences, &c. read to the Institute of France in April 181C. 186 Animal¬ cule. A N I M A which is itself founded on the systems of Muller and Bruguiere. The French naturalist includes in his system only such species of the same class described by Muller as are destitute of a mouth. Order I.—Naked Infusoria. Body extremely simple, apparently homogeneous, unprovided with organs or external appendages. The naked Infusoria are the smallest and simplest of those animals which are cognizable by the senses of uian. When we expose w^ater to air and light, especially ir it contains an infusion of animal or vegetable remains, we speedily perceive, by the assistance of the microscope, a variety of animalcules. These are divided into two sec¬ tions. Section I.—Body Thick. Of this section the bodies present such a perceptible degree of thickness as removes them from the simply membranous state. Genus Monas.—Body extremely small, of the simplest construction, transparent, punctiform. The Monads are the smallest and least organized of liv¬ ing creatures. We have indeed scarcely any proof of their animal life, except that they are moving corpuscles, allied to the genus Volvox, the animality of which is un¬ doubted. They have neither mouth nor alimentary canal, nor any apparent organs. They live by absorption, and are found in tranquil, but rarely in limpid waters. They are produced in infusions of animal and vegetable sub¬ stances. Sp. 1. Monas termo.—An extremely minute gelatinous point, scarcely perceptible even with the aid of a power¬ ful lens, and frequently disappearing under a strong light in consequence of its perfect transparency. This species is common in ditch-water, and in numerous infusions. See Plate XLVI. fig. 1. These minute creatures being the earliest discernible evidence of animal life, and the last result to which the higher and more perfect forms can be reduced by infusion, have been called the alpha and omega of all organized existence. Their history has given rise to many curious views, and more absurd spe¬ culations. Sp. 2. Monas atomus.—White, with a variable dark- coloured spot, which appears to change its position. This species was found in sea-water wdiich had been kept an entire winter, but was not very fetid. See Plate XLVI. fig. 2 and 3. Sp. 3. Monas punctum.—Nearly black, of a sub-cy¬ lindrical form. Found in the infusion of the pulp of a pear. Sp. 4. Monas lens.—Hyaline, of an ovoid form. Found in all kinds of waters. Multiplies by spontaneous incision. Sp. 5. Monas pulvisculus.—Hyaline, with a greenish margin. Found in the waters of marshes. This species has been lately regarded as an enchelis. Indeed, so great is the difficulty of microscopical investigation, and such indefatigable patience is required in order to see things as they really are, that the species and genera of this class of beings are frequently transposed and altered in their relative position and arrangement, in consequence of the very dissimilar views which different observers have taken of the same object. As it would be inconsistent with our present limits to describe more than a few species of each genus, we shall content ourselves with the preceding Monads. “ How many kinds,” observes Mr Adams, “ there may be of these invisibles, is yet unknown, as they are discerned of all sizes, from those which are barely invisible to the naked L C u L E. eye, to such as resist the force of the microscope, as the k fixed stars do that of the telescope, and, with the greatest t! powers hitherto invented, appear only as so many movingG points.” Genus Volvox.—Body very, simple, transparent, sphe. rfcal or ovoid, revolving on itself as on an axis. With the exception of one species ( V. globator) the vol- voces are invisible to the naked eye. Under the micro- scope they assume the aspect of small gelatinous masses, which sometimes present an oval form. In some the body seems composed of numerous smaller globules united in one common mass. I here is reason to suppose that these interior bodies are afterwards propelled outwardly, and become separate individuals. The species occur both in fresh and salt waters, and in vegetable infusions. They derive their generic name from the manner in which they turn or revolve upon their axis. Leeuwenhoeck describes an animalcule “ a thousand times smaller than a louse’s eye, which exceeded all the rest in briskness,” and turned itself round as it were upon a point, with the celerity of a spinning-top. The genus is divisible into two sections. * Interior of the body apparently simple and homogeneous. Sp. 1. Volvox punctum.—Spherical, blackish, with a central lucid point. Of this species many are sometimes seen together in their passage through the water. They occasionally move as if subjected to the influence of a whirlpool, and then separate. Numerous in fetid sea¬ water. Sp. 2. Volvox granulum.—Spherical, green, periphery hyaline. Dwells in the water of marshes. Sp. 3. Volvox globulus.—Globular, sub-obscure behind. This species is ten times larger than the Monas lens. It occurs in most vegetable infusions, and moves with a slow fluttering motion. Plate XLVI. fig. 4. * * Interior of the body exhibiting smaller corpuscles. Sp. 4. Volvox pilula.—Spherical, with greenish inter¬ nal globules. In those pure waters which nourish the Lenina minor. Plate XLVI. fig. 5. Sp. 5. Volvox lunula.—Roundish and transparent, and composed of an innumerable assemblage of homogeneous crescent-shaped molecules, without any common margin. Its movements are of two kinds, that of the molecules among themselves, and that of the whole revolving mass. It occurs in marshy places in the early spring. Plate XLVI. fig. 6. Sp. 6. Volvox globator.—Commonly called the globe- animal. Spherical, membranous, the internal globules dis¬ tant or scattered. Abundant in the infusions of hemp and tremella, and in stagnant pools during spring and summer. The following is an account of it by Mr Baker. “ There is no appearance of either head, tail, or fins, and yet it moves in every direction, backwards, forwards, up or down, rolling over and over like a bowl, spinning horizontally like a top, or gliding along smoothly without turning itself at all; sometimes its motions are very slow, sometimes very swift; and, when it pleases, it can turn round as upon an axis very nimbly, without moving out of its place- The body is transparent, except ■where the circular spots are placed, which are probably its young.” Another au¬ thority states that this species is at first very small, but increases to such a size that it may be discerned by the naked eye, and that its interior is filled with small glo¬ bules, which are smaller animalcules, each of which con¬ tains within itself a still smaller generation, all percep¬ tible by means of powerful glasses. The lesser globules may be seen escaping from the parent, and increasing w size. A N I M A i m\ Jen-us Proteus.—Body very small, simple, transparent, | | of varying form, changing itself instantaneously into f different lobated shapes. This genus is more obviously contractile than the pre- I e(]ing. bit is seldom seen above a minute under the same * mn, but is continually passing from a simple oval or ob- ? m-r’to an irregular or sinuated shape, and vice versa. L species described by Iloesel is so remarkable for this iculty, that it has been compared to a drop of water : iroWn upon oil. Hence also the generic name. Sp. 1. Proteus diffluens.—Body diverging into branches. W tccurs in the water of marshes. Plate XLVI. fig. 7, 8, lind9. , Sp. 2. Proteus tenax.—Body prolonged to a fine point. | iccurs in rivers and in sea-water. There are only two hi lecies described as belonging to this genus. hi Ienus Enchelis.—Body very small, simple, oblong, cylindrical, slightly variable. There is a marked analogy between this genus and the | <1 illowing. The Enchelides are, however, short and thick |<1 Compared with the Vibriones, which are slender and lingthened. To the genus now under consideration be¬ ing those animals which, if the recorded observations on ic subject have been accurately made and faithfully re- orted, more than any other confound our preconceived leas regarding the distinction between animal and vege- ible life. The species alluded to are named Zoocarpes by I. Boryde St Vincent, or animated seeds, which appear re- iprocally to give rise to and proceed from certain aquatic lants of the conferva1 kind. They are formed in a bulb- iis-shaped part or swelling of the plant, are ejected when pe, swim about for some time with a voluntary motion, irow out a root and a branch, become genuine vegetables, roduce living seeds, and give birth to animals which, af- ;r a similar change of form, speedily return again to the Dgetable state. These facts are vouched for by M. Bory de t Vincent, and are credited and confirmed by M.Dutrochet nd several other continental inquirers, some of whom eciare that they kept so watchful an eye upon the same tdividual as never to lose sight of it for a moment till * ley had witnessed the singular transformation above f lent ioned. We recommend it to our readers’ consideration. Sp. 1. Enchelis viridis.—Subcylindrical, obliquely trun- iated anteriorly. This species has an obtuse tail or ter- linal part. It continually varies its motion, turning from ght to left. Occurs in long-kept water. Sp. 2. Enchelis punctifera.—Subcylindrical, green, ob- ise anteriorly, pointed posteriorly. This species is opaque, ith a small pellucid spot in the fore part, in which two lack points are seen, and a kind of double band crosses ie middle of the body. It occurs in marshes. Plate XVI. fig. 10, 11. Sp. 3. Enchelis pulvisculus.—This species bears a great ^semblance to the Monas pulviscidus of Muller, which i the E. monadina of Bory de St Vincent. It is, how- I iver, double the size, deeper tinted, and more ovoid, t is found in the waters of marshes, and accumulates round the sides of jars or vases in which confervae have een kept. It forms on the surface of water a slight , chicle ot a delicate green colour, which is supposed to ave been erroneously regarded by many botanists as a ogetable production, and described under the name of p/ssusflos aqua;. On dying it becomes more lengthened nd pellucid, or at least retains only a slight central spot 1 green. Plate XLVI. fig. 12. Sp. 4. Enchelis amcena.—This is a new species, of a vely green colour, discovered by Bory de St Vincent. In " mmi‘ng it appears to elongate itself, and advances with L C U L E. 187 the more slender end foremost. Two individuals are Animai- sometimes observed to unite and form one animal, of a cule- perfectly spherical form, and similar in aspect to a Volvox. Sp. 5. Enchelis tiresias.—This species was also disco¬ vered by the above-named writer, and led to his peculiar views regarding those apparently animated seeds which he has named Zoocarpes. He asserts that he has seen this animalcule formed in the articulations of a true con¬ ferva ; that it burst from its vegetable envelope with a gy¬ ration or circular movement; that it soon produced a translucent prolongation of its body, which may be called anterior, as it then swam in the direction of that new organ, which, with the body itself, became visibly longer, till the creature finally acquired the exact form of the Enchelis deses of Muller. The chief difference seemed to be that it always moved with the slender end foremost, whereas the species just mentioned swims with its blunt end in advance. It is described by recent French writers as an “ animal extraordinaire qui n’est certainement que la grain vivante d’un vegetal.” (See Diet. Class, d'Hist. Nat. tome vi. p. 156.) Sp. 6. Enchelis deses.—This species is of an obscure green, much elongated, and moves with the thick end anteriorly. “ Celui-ci (the obtuse portion) parait comme tronque dans certains aspects ; et en examinant attentive- ment cette sorte de troncature, on la reconnait formee par un cercle en forme de disque moins fonce que le reste de I’animal. La pointe posterieure est parfaitement hyaline. Dans la pensee ou nous sommes que les Enchelides vertes ne sont que des Zoocarpes, ou propagules animes de quelques genres d’Arthrodiees, nous croyons que le disque obscurement transparent de la partie anterieure n’est que la marque du point sur lequel doit se developper 1’article par lequel doit s’allonger en filament confervoide le Zoocarpe, lorsque, arrive au terme de sa carriere ani- male, il doit se fixer et prendre racine par le point hyalin de la partie posterieure.” (Xoc. cit. p. 157.) We present the above passage to our readers without note or com¬ ment, as we do not ourselves understand the zoocarpal nature of an Enchelis. Genus Vibrio.—Body very small, simple, cylindrical, elongated. Animalcules have been described as constituents of this genus, which probably do not at all belong to it, being too complicated in their structure. If the V. aceti, for example, commonly called the vinegar eel, is furnished with a mouth, lips, and alimentary canal, it does not even pertain to the class Infusoria, however small its dimensions. But many of the species are undoubtedly of the simplest construction; and although they may present some ap¬ pearance of an internal cavity or sac, they yet exhibit neither mouth nor other external orifice of any kind. Sp. 1. Vibrio lineola.—Body linear, extremely minute. Occurs in many vegetable infusions in such numbers as ap¬ parently to occupy their entire space. It is so small, that with the best magnifiers little more can be discerned than an obscure tremulous motion. It is supposed to exceed even the Monas termo in tenuity. Plate XLVI. fig. 13. Sp. 2. Vibrio spirillum.—Filiform, and twisted spirally, which seems to be its natural shape, as it is never observed * to unbend, but moves forwards with a vibratory motion at both ends. Found in an infusion of Sonchus arvensis. Plate XLVI. fig. 14. Sp. 3. Vibrio vermiculus.—Presents a milky aspect, with a blunt apex, and moves with a languid vermicular motion. It has been found in marshy water in November, but is seldom seen. It agrees with the animal mentioned by Leeuwenhoeck as occurring in the dung of frogs. Sp. 4. Vibrio paxillifer.—“ Animalculum, ’ says Miil- Igg A N I M A Animal- ler, “ vel congeries animalculorum mirabilis. Pluries m cule. guttulis aquae marinae vidi corpuscula linearia flavescentia (solitaria paleas, in quadrangula disposita scobes refere- bant), granulaque seminalia qualiscunque vegetabdis diu credidi; demum nocte inter 6 et 7 Octobrem 1/81 as- spectu fili flavescentis, sese in longum producentis et in breve contrahentis, ac ex bis paxillis compositi, obstupe- factus, novoque phaenomeno gavisus, ejusdem variis evo- lutionibus incubui.” A salt-water species, abundant in ulva latissima. Observed during the months of September and October. Plate XLYI. fig. 15, 16, 17, 18, 19. Sp. 5. Vibrio serpens.—Slender and gelatinous, with obtuse windings or flexures, resembling a serpentine line. It is rare, and occurs in river-water. Section II.—Body Membranaceous. Of scarcehj perceptible thickness, whether fiat or concave. The organization of the animalcules of this section is scarcely less simple than that of the preceding; but their form, being in a small degree resistant, is less subject to variation from the pressure or other action of the sur¬ rounding fluids, which has been regarded as the proof of a certain progress or advance in the scale of structure. Genus Gonium.—Body very small and simple, flattened, short, angular. Some species of this genus appear to be composed of several corpuscles united together un¬ der a common membrane. This appearance probably results from their cellular tissue, or from certain lines which are the rudiments of those spontaneous divisions formerly mentioned, by which their propagation is af¬ fected. Their movements are oscillatory. Sp. 1. Gonium pectorale.—Quadrangular and pellucid, with sixteen globules of a greenish colour set in a quadran¬ gular membrane, “ like jewels in the breast-plate of the high-priest, reflecting light on both sides.” Occurs in pure waters. Plate XLVI. fig. 20. Sp. 2. Gonium pulvinatum.—Quadrangular and opaque. Found in dunghills. Sp. 3. Gonium corrugatum.—Sub-quadrangular, whitish, marked by a longitudinal line. This species is found in various infusions, particularly that of the pear. Sp. 4'. Gonium truncatum.—Internal molecules dark green. Anterior extremity forming a straight line, with which the sides produce an obtuse angle, terminated poste¬ riorly by a curved line. This species exhibits a languid motion. It is much larger than the preceding, and oc¬ curs, though rarely, in pure water. Genus Cyclidium.—Body very small and simple, trans- parent, flattened, orbicular or oval. The motions of this genus are oscillatory, circulatory, or demi-circulatory, more or less interrupted, and languid or lively, according to the species. Sp. 1. Cyclidium bulla.—Orbicular and hyaline. General appearance pale and pellucid, with the edges somewhat daikei than the rest. It moves slowly in a semicircular direction, and occurs in the infusion of hay. Sp. 2. Cyclidium hyalinum.—Oval, depressed, perfectly ti ansparent, terminated by a tail-like elongation. This species is very common, and is produced in many infu¬ sions, particularly in those of the cerealia. It swims in a vacillating manner, and as if continually trembling. Plate XLVI. fig. 21. b Sp- o. Cyclidium Nucleus.—Of a brownish tinge, deeper behind, and shaped exactly like an apple pippin. Genus Paramecium.—Body very small, simple, transpa¬ rent, membranous, oblong. . ^he species of this genus, according to Lamarck, scarce¬ ly differ from those of the preceding, except in their more L C U L E. lengthened forms and a slight increase of animal deve- j lopment. They appear to vary instantaneously accord- ! ing to their position in relation to the eye of the observer X' but their real form is tolerably determinate. The mode of increasing the species by transverse and longitudinal divisions, or natural cuttings, is very obvious in this genus. They are nearly related to the following, but are less sinii- ous and irregular. Their movements are for the most part slow and indefinite. They swim horizontally on one of their flattened surfaces, after the manner of flounders. Sp. 1. Paramcecium aurelia.—Body compressed, with a kind of plait or fold towards the apex, acute behind. Very common in water where confervae grow. Plate XLVl. fig. 22, 23, 24. Sp. 2. Paramcecium chrysalis.—Plicated anteriorly, ob¬ tuse behind. Occurs during the autumn in sea-water. Sp. 3. Paramcecium versutum.—Cylindrical, thickened posteriorly, obtuse at both ends. Found in ditches. Genus Kolpoda.—Body very small, simple, flattened, oblong, sinuous, irregular, transparent. This genus is nearly allied to the preceding, and differs from it chiefly in its more varied forms. It is also less subject to the influence of pressure by the medium in which it lives. An Italian naturalist of the name of Losana has lately published a monograph on Kolpoda ; but his figures are somewhat exaggerated, and not very naturally ex¬ pressed. Sp. 1. Kolpoda lamella.—Elongated, membranaceous, curved anteriorly. This species is seldom met with. It has a singular vacillatory mode of movement, and advances on its sharp edge, instead of its flattened side, the more usual position. Sp. 2. Kolpoda gallinula.—Oblong, the anterior por. tion of the back membranaceous and hyaline. In cor- rupted sea-water. Sp. 3. Kolpoda crassa.—Yellow, thickish, somewhat opaque, curved a little in the centre, kidney-shaped. This species has a quick vacillatory motion, and becomes appa¬ rent in the infusion of hay generally in about 13 hours. When the water is nearly evaporated it assumes an oval form, becomes compressed, and bursts. Sp. 4. Kolpoda rostrum.—Oblong, hooked anteriorly. The movements of this species are slow and horizontal. It is found, but not frequently, in water where the lemna grows. Sp. 5. Kolpoda cucullus.—Ovate, ventricose, the top bent into a kind of beak, and an oblique incision beneath the apex. This species is found in vegetable infusions, and in fetid hay, and usually moves with great velocity. It is pellucid, and appears as if filled with little bright vesicles, which differ in size. Some have supposed them to be lesser animalcules which the Kolpoda has swallowed; but as it has no mouth wherewith to swallow, Miiller is more probably right in regarding them in the light of germs. When about to die in consequence of evaporation, it protrudes its contents, whether food or offspring, with great violence. Plate XLVII. fig. 25, 26, 27, 28. Genus Bursaria.—Body simple, membranaceous, con¬ cave. This genus occurs in fresh, saline, and stagnant waters. It appears to vary its form beneath the eye of the obser¬ ver, and, from a rounded flattened shape, assumes a con¬ cave or somewhat purse-shaped aspect. Sp. 1. Bursaria truncatella.—Follicular, with a trun¬ cated apex. This species is visible to the naked eye. It moves from left to right, and from right to left; ascends to the surface in a straight line, and sometimes rolls about as it descends. ANIMALCULE. 189 tSp. 2. Bursaria bulina.—Boat-shaped, labiated ante- iorly. This species is pellucid and crystalline, with ■rilliant globules within it. It is concave on the upper de, and convex below. Sp. 3. Bursaria hirundella.—With two small wing-like rejections, which give it somewhat of the appearance of bird. It is invisible to the naked eye, but appears under ie microscope like a pellucid hollow membrane. Recent licrographical observers doubt if this species really per- lins to the present genus. Plate XLVII. fig. 29. Order II.—Appendiculated Infusoria. Furnished exteriorly with projecting parts. The animalcules of this second principal order or divi- on, though still infinitely small, gelatinous, and trans- arent, are so far less simply organized than their pre- ecessors, inasmuch as they are furnished with salient arts, such as hairs, horns, or tail-like appendages, or at ;ast with such projecting organs as, for want of more ap- ropriate terms, we are obliged to designate by those ames. They multiply both by division and by the burst- ig forth of internal germs. Though their essential fluids, nd the living tissue which contains them, are probably of more compound nature than those of the naked Infusoria, icy have not yet reached that point of organization in hich special organs are developed for the performance f particular functions; and it is not till we reach the igher class of Polypi that these are distinctly percepti- le. “ II parait,” observes Lamarck, “ par les nombreuses speces deja connues et publiees, que les Infusoires de cet rdre sont bien plus nombreux dans la nature que les In- ' isoires nus. Cela doit etre ainsi d’apres les principes ue je me suis cru fonde a etablir. “ En effet, dans les Infusoires nus, forigine encore trop 3cente des races qui proviennent de celles, en petit nom- re, qui furent generees spontanement, n’a permis a la uree de la vie et aux circonstances qui ont influe sur ces ices, qu’une diversite peu considerable. Mais a mesure ue la duree de la vie, que sa transmission dans les indi- idus qui se sont succedes en se multipliant, et que les irconstances ont eu plus de temps pour exercer leurs in- uences, les races se sont diversifiees de plus en plus, et )nt devenues plus nombreuses. “ Cet ordre des choses, qu’il est facile de reconnoitre our celui meme de la nature, nous fait sentir pourquoi is Infusoires sont bien moins diversifies et moins nom- reux que les Polypes. Effectivement, quoique nous ne onnaissions pas probablement tous les Infusoires, et que ous connaissions bien moins encore tous les Polypes, ce ui est deja connu de part et d’autre indique que la diver- te des Polypes est considerablement plus grande que es Infusoires. Aussi les Polypes sont plus eloignes de ■ur origine que les Infusoires.” (Animaux sans Vertebres, >me i. p. 433.) I he first genus of this order (Trichoda), as constituted ) y Muller, contains several species which manifest the -Kliments of a mouth and the commencement of an ali- lentary canal: these, according to the negative charac- ;rs of the class, do not belong to the Infusoria. rENUs Trichoda.—Body very small, transparent, di¬ versiform, without caudal appendage, but garnished with soft hairs either on the whole or on part of its sur¬ face. According to the views of Lamarck, this genus contains ot only a great proportion of the genus Trichoda of u ler, but also the whole of the genus Leucophra of the amsh author. It is distinguished from Kerona by the want of the long, stiff, distant, corniform hairs which cha- Animal- racterize the latter. cirle. A. Body ciliated over its entire surface. (Leucophra of Muller.) Sp. 1. Trichoda viridiscens.—Greenish, cylindrical, opaque, thicker posteriorly. Found in sea-water. Sp. 2. Trichoda dilata.—Body flattened, variable, with sinuated margins. Inhabits sea-water, and swims like a Planaria. It scarcely differs from the genus Kolpoda, ex¬ cept in being ciliated. Sp. 3. Trichoda scintillans.—Of a green colour, oval, slender, and opaque. Occurs in stagnant water. A doubt¬ ful species, closely allied to Volvox. Sp. 4. Trichoda acuta.—Ovate, with a sharpened point. Colour yellowish. Of this species the form is very vari¬ able. It occurs in sea-water among ulvse. Plate XLVII. fig. 30, 31, 32. Sp. 5. Trichoda signata.—Oblong, sub-depressed, with a blackish margin. Common in sea-water. This species is distinguished by a curved line in its centre, shaped like the letter S, one end of which sometimes assumes a spiral form. Sp. 6. Trichoda mammilla.—Spherical, opaque, with an exsertile papilla. Occurs in the waters of marshes. It is of a dark colour, its short hairs are curved inwards, and it occasionally appears to project and draw in a little white protuberance. B. Body ciliated.) or covered with short hair only over a part of its surface. (The greater proportion of the genus Trichoda of Muller.) Sp. 7. Trichoda grandinella.—Spherical, pellucid, haired above. The minute ciliary appendages of this species are not easily discovered, as it seems to possess the power of withdrawing them at pleasure and instantaneously. It oc¬ curs both in pure water and that of infusions. Sp. 8. Trichoda cometa.—Spherical, ciliated anteriorly, with one or more globular appendages behind. Found in pure water in the autumnal season. Plate XLVII. fig. 33, 34. Sp. 9. Trichoda Solaris.—Spherical and crystalline, its edges beset with diverging rays, which exceed in length the diameter of the body. This animalcule contracts and dilates, but is stationary in the same spot. In marine in¬ fusions. Plate XLVII. fig. 35. Sp. 10. Trichoda pubes.—Oval oblong, gibbous, de¬ pressed anteriorly. The apex of this species is furnished with hairs, which are seldom visible till it is about to ex¬ pire, when it protrudes and extends them vehemently, as if in a vain attempt to secure and detain a remaining par¬ ticle of water. Sp. 11. Trichodaproteus.—Oval, obtuse behind, with an elongated retractile neck. Apex haired. This species, ac¬ cording to Muller and Lamarck, is found in river water. It appears, however, to agree in general character and appearance with the Pi-oteus described by Mr Baker, which usually occurs in the slimy matter adhering to the sides of vessels in which either animal or vegetable substances have been some time kept. That of which an account is given by Mr Adams was found in the slime produced by water containing small fishes, snails, &c. The body was something similar to that of a snail, but pointed at one end, while from the other proceeded a long, slender, “ and finely proportioned neck, of a size suitable to the rest of the animal.” If we credit Mr Baker, this animalcule, though its eyes are not discernible, plainly demonstrates by its actions that it can see; for though multitudes swim about in the same water, and its own progressive motion is 190 ANIMALCULE. Animal- very swift, it never strikes against its neighbours, but cb- cule. :«■" utLL q rlovi-pvitv “ wbollv unaccountable reels its course with a dexterity “ wholly unaccountable should we suppose it destitute of sight. ^ Its entire shape bears a resemblance to that of a swan. See Plate XL v 11. fig. 86 and 37. When alarmed, it draws in its supposed neck, becomes more opaque, and moves about slow ly with the large end foremost. See fig. 38. After continuing for some time under this form, it will put forth a kind of wheel machinery, the motions of which are alleged to draw a current of water towards it from a considerable distance. After frequently pushing out and pulling in this shorter head, sometimes with and sometimes without the wheel-work, it will remain motionless, as if wearied or worn out; and then its long head and neck or apex will be_ again slowly protruded, after which it generally resumes its ac¬ customed agility. Genus Kerona.—Body very small, diversiform, without tail-like prolongation, and furnished with scattered, stiff, corniform hairs on some parts of its body. Sp. 1. Kerona rostdlum.—Orbicular and membrana¬ ceous ; one side angulated, the other furnished with a se¬ ries of triple horns. Inhabits sea and river water. This species is alleged by Bory de St Vincent to be entirely destitute of hairs and cirri, and he therefore proposes to remove it to the order Gymnodes, which corresponds with our first order, the naked Infusoria. It is sometimes diffi¬ cult to identify species in this department, or to ascertain, in a doubtful or contested case, that the same animalcule has been the subject of observation by two or more dis¬ putants. Plate XLVII. fig. 39. Sp. 2. Kerona cypris.—Somewhat pear-shaped, com¬ pressed, the front furnished with hairs or vibrating points, inserted beneath the edge, shorter behind, and partly ex¬ tended straight forward, partly bent downwards. Motion retrograde. Inhabits fresh water. Sp. 3. Kerona India.—Smooth, pellucid, full of small points, the fore part clubbed and a little bent, the hinder part narrow ; the base obliquely truncated, and terminating in a tail stretched out transversely. The top of that part which maybe called the head, and the centre of the back, are furnished with long hairs. When this animalcule is at rest, its tail is curled; when in motion it is drawn tight and extended upwards. The movements of this species are lively and diverting. Genus Cercaria.—Body very small, transparent, di¬ versiform, furnished with a distinct but very simple tail. This genus, as constituted by Muller, contains many species which bear no natural relation to each other; but his characters are precise and definite, and strictly appli¬ cable to those species which now form the genus as limit¬ ed by modern observers. They occur more rarely among animal and vegetable infusions than in running streams and the waters of marshes. Their movements are for the most part circular and very rapid. With the exception of a well-marked tail, their organization is in every other respect extremely simple. If a mouth and the rudiment To this genus Lamarck has united the Himantopus of Muller. The species are rare. They seldom occur in infusions, and are most frequent in the purer kinds of fresh and salt waters. T1rL?1LlC!!1nnpnm,t!,eotiff-charaCterS ^ the ffenus ^osperma, as recently established: “ Corps non contractile, oybide, tres-com- cie Ce "enre dnnt nmfc 011 J)eaucouP plus longue, implantee a la partie posterieure, qui est peu ou point annn- L™ ;;; ^ P ^ °ns u" trfs-Srand "ombre d’especes, se compose d’animaux spermatiques.” (Dkt Clans. d'Hist-^ tnirm in tv 3o6.) I he production and existence of these animals, their nature and tome in. p teries of nature. spermatiques.” {Diet. _ uses, are still among the many inscrutable mis' of a stomach or alimentary canal exist in any of these a animals, such characters would remove them not only 1 iiif 1 from the genus Cercaria, but from the class Infusoria, as'"' defined at the commencement of this treatise. Aforti^ 1 the existence of eyes (a fact assumed by some inaccurate observers) in any of the animalcular species, would en- tirely alter their position and arrangement in the animal kingdom. We cannot do better than report the observa. tions of the venerable Lamarck (himself unfortunately now deprived of sight) on this obscure subjectIci, comme dans le genre suivant, Ton est expose, d’apres la petitesse extreme des individus, a rapporter a la classe des Infusoires des animaux qui, par leur organisation, appar. tiennent a d’autres points de 1 echelle animale.” “Une bouche, quoique d’abord inaper^ue, et consequem- ment febauche d'un sac alimentaire, peuvent exister dans certains de ces animaux, et des-lors ils appartiennent au premier ordre des Polypes; mais des yeux, comme on en a suppose dans certains Cercaires, cela est impossible. “ Avant de dire que le fait lui-meme vaut mieuxquela raisonnement, il faut, \mo, constater que les points que Ton a pris pour des yeux, en sent reellement, et qu’ils ont chacun un nerf optique qui se rend a une masse medul- laire, centre de rapports pour des sensations; 2do, il faut ensuite etablir positivement que des animalcules reelle¬ ment pourvus cl’yeux, sont neanmoins, par leur organisa¬ tion, de la meme classe que les Infusoires.” (Animam sans Vertebres, tome i. p. 444.) This genus forms the nucleus of the new family of In¬ fusoria proposed by M. Bory de St Vincent under the name of Cercariees, and which contains in all seven genera, the names and nature of which will be seen by referring to the tabular view. Muller, who was not practically ac¬ quainted with the spermatic animalcules, wras attracted by the resemblance which some of the Cercariae bore to the figures of those organic molecules in the works of his pre¬ decessors. He did not, however, assert their identity; and probably perceived that, although in their general as¬ pect and mode of movement they resembled each other, their peculiar and very different localities, and even the details of their structure, rendered it advisable that they should be assigned to separate genera.1 Sp. 1. Cercaria inquieta.—Changeable, convex, with a slender tail. This species occurs in salt water, and is re¬ markable for assuming a variety of different forms. It is sometimes oval, sometimes cylindrical, sometimes shaped like a sphere. Plate XLVII. fig. 40. Sp. 2. Cercaria gyrinus.-—Body of a rounded form, with an acuminated tail. In swimming, this animalcule moves its tail like a tadpole. Sp. 3. Cercaria lemna.—Changeable, sub-depressed, with an annulated tail. Tire C. lemna varies the form ot its body in a manner almost as singular as that exhibited by the Proteus, already described. The body is triangular,or oblong, or kidney-shaped. Its tail is at times thick, short, annulated; at others it is long, cylindrical, and without rings; and when stretched out it sometimes vibrates with such velocity as to appear double. A small pellucid gb* bule, which Muller regarded as the mouth, is perceptible near the apex ; and there are also two excessively minute black points, which, whatever they may really be, are by some called eyes. It advances slowly by a few steps or movements at a time, and frequently shakes and bends its tail, in which position it bears a great resemblance to a ANIMALCULE. 191 U ; Tina leaf in. miniature. This animalcule exhibits an ad- " | need organization. Plate XLVII. fig. 41, 42, 43. (snus Furcocerca.—Body very small, transparent, rarely ciliated, furnished with a bipartite tail. This terminal genus, according to the views of Lamarck, (nducts us to the limits of the infusorial class, and we Icome thus more liable to deception in regard to the r n-existence of a mouth than in the preceding genera. 1 is a dismemberment of the Cercaria of Muller, and pro- t bly contains many species which will be placed else- iere when future and more continuous observation s ill have thrown additional light upon their nature and tributes. Sp. 1. Furcocerca podura.—Cylindrical, acuminated pteriorly. This species is pellucid, and seems to con¬ st of a head, trunk, and tail, the first of which, in the \ ;w of some observers, “ resembles that of a herring.” It rns round as if upon an axis when it moves, and is i jally found in the months of November and December, i places where the lenma abounds. The tail frequently, t not always, appears to be divided into two. One of iller’s figures of this species is probably erroneous. He rSresents it as covered with short hairs; whereas, to nre recent observers, it appears perfectly smooth. Plate ivil. fig.‘44, 45. Sp. 2. Furcocerca viridis.—Cylindrical, variable, divided a, 1 acuminated behind. Occurs in spring in ditches and s liding pools. It frequently contracts its anterior and psterior portions, so as to assume a spherical form. It is cricult to determine the genus to which this species be- l gs. Lamarck is supposed to have erred in placing it , vi ere it now stands. In truth, the genus appears to have I'm rather established provisionally, than upon an assured a 1 natural foundation. The varying forms of the species u V ich it contains render it extremely difficult either to I d cern or describe them with precision. Plate XLVII. t 46, 47, 48. We have now endeavoured to present a general view, pi I al a systematic exposition, of the principal features of “ 11[: animalcular world; and if our statements have been 1 s explicit, and our arrangement less complete and me- i-Btndical, than accords with the reader’s expectation, these yBqjects must in part be attributed to the uncertainty h lwich still prevails regarding a subject of which many of iV t; essential characters scarcely lie within the limits of 1 man intelligence. The observations and experiments I'lothe English microscopical observers of last century, ■ tough they might amuse the general student, are too ni Vjpie and fanciful to be now regarded as parts of the as- Pis’ed history of animalcules. The ultra-analogical rea- s ling on subjects of natural science with which we have 1 m lately favoured by such men as Oken and Geoffroy ' ' Hilaire, are tame in comparison with the inferences > i luced by some of our older observers, who describe v :h minuteness the head, eyes, mouth, jaws, throat, sto- ! Lch, intestines, and other parts of animalcules, which t; improved glasses of modern times do not reveal to the v 1011 °f not less patient inquirers. The recent observa- t ns on the motions of the pollen of plants, which have I zzled the modern philosophers, would have opposed but f ble barriers in the way of our predecessors. “ To dis- 1 >'er, says Buffon, “ whether all the parts of animals, * I all the seeds of plants, contained moving organic par- 1 ks, I made infusions of the flesh of different animals, ^ I of the seeds of more than twenty different species of ' potables; and after remaining some days in close glasses, mu the pleasure of seeing organic moving particles in of them. In some they appeared sooner, in others later; some preserved their motions for months, and Animal- others soon lost it. Some at first produced large moving cule. globules resembling animals, which changed their figure, split, and became gradually smaller; others produced only small globules, whose motions were extremely rapid ; and others produced filaments, which grew longer, seemed to vegetate, and then swelled and poured forth torrents of moving globules.” It was from these and similar obser¬ vations that the theory arose proposed by Baron Mun¬ chausen (an ominous name !). The Baron perceiving that these moving globules, after taking a little exercise, began again to vegetate, drew the conclusion that they were first animals and then plants ; thus anticipating by more than half a century the supposed discoveries of some mo¬ dern physiologists. Which of them was first in error it is perhaps of little consequence to inquire; and we allude to the subject here rather in connection with some singu¬ lar observations by Mr Ellis, recorded in the 59th volume of the Philosophical Transactions, than from its own in¬ trinsic importance. His object was to overturn Mun¬ chausen’s hypothesis, by showing that the supposed Zoo- carpes were nothing more than “ the seeds of that genus of fungi called mucor or mouldiness” and that their mo¬ tions were caused by the attacks of myriads of animal¬ cules ! “ Having at the request of Dr Linnaeus made se¬ veral experiments on the infusion of mushrooms in water, in order to prove the theory of Baron Munchausen, that these seeds are first animals and then plants, it appeared evidently that the seeds were put in motion by very mi¬ nute animalcules, which proceeded from the putrefaction of the mushroom: for by pecking at these seeds, which are reddish, light, round bodies, they moved them about with great agility in a variety of directions; while the little .animals themselves were scarcely visible till the food they had eaten had discovered them. The satisfac¬ tion I received from clearing up this point led me into many other curious and interesting experiments. The in¬ genious Mr Needham supposes these little transparent ra¬ mified filaments, and jointed or coralloid bodies, which the microscope discovers to us on the surface of most animal and vegetable infusions when they become putrid, to be zoophytes, or branched animals; but to me they appear, after a careful scrutiny with the best glasses, to be of that genus of fungi called mucor or mouldiness, many of which Michellius has figured, and Linnaeus has accurately de¬ scribed. Their vegetation is so amazingly quick, that they may be perceived in the microscope even to grow and feed under the eye of the observer. Mr Needham has pointed out to us a species that is very remarkable for its parts of fructification (See Phil. Trans, vol. xlv. tab. 5, fig. 3, a, A). This, he says, proceeds from an infusion of bruised wheat. I have seen the same species proceed from the body of a dead fly, which was become putrid by lying floating for some time in a glass of water where some flowers had been, in the month of August 1768. This species of mucor sends forth a mass of transparent filamentous roots; from whence arise hollow stems, that support little oblong oval seed-vessels, with a hole on the top of each. From these I could plainly see minute globu¬ lar seeds issue forth in great abundance, with an elastic force, and turn about in the water as if they were ani¬ mated. Continuing to view them with some attention, I could just discover that the putrid water which surround¬ ed them was full of the minutest animaleula; and that these little creatures began to attack the seeds of the mu¬ cor for food, as I have observed before in the experiment on the seeds of the larger kind of fungi or mushrooms. This new motion continued the appearance of their being alive for some time longer; but soon after many of them arose to the surface of the water, remaining there without animalcule. 192 Animal, motion; and a succession of them afterwards coming up, cule. they united together in little thin masses, and floated to the edge of the water, remaining there quite inactive du¬ ring the time of observation.” In like manner, the move¬ ments of the jointed coralloid bodies which Mr Needham has named chaplets and pearl necklaces are attributed to the attacks of their animalcular enemies. “ M hen a small portion of these branches and seeds are put into a drop ot the same putrid water upon which the scum floats, many of these millions of little animalcula with which it abounds immediately seize them as food, and turn them abou with a variety of motions, as in the experiments on the seeds of the common mushrooms, either singly, or two or three seeds connected together ; answering exactly to Mr Needham’s description, but evidently without^any motion of their own, and consequently not animated !” We shall conclude this subject with a short notice of another view of the matter, which has resulted fiom some recent experiments and observations by our celebrated botanist Robert Brown. While engaged in some inqui¬ ries regarding the structure of the pollen of plants, and its mode of action on the pistillum of phsenogamous tribes, that accurate observer had occasion to immerse in water some particles taken from the full-grown anthers (previous to bursting) of Clarkia pulchella. Of these, he perceived by the microscope that many were evidently in motion, and that their motion consisted not only of a change of place in the fluid, but of a change of form in themselves; that is to say, a contraction or curvature about the middle of one side, accompanied by a corre¬ sponding enlargement or convexity on the other, frequent¬ ly occurred. The particles were seen, in a few instances, to turn on their longer axis ; and their general motions were of such a nature as to produce the conviction in Mr Brown’s mind that they did not arise either from currents or evaporation of the fluid, but were proper to the particles themselves. Having ascertained that mo¬ tion existed in the pollen of all the living plants which he examined, he next inquired whether, and for what length of time, this singular property was retained after the death of the plant. Specimens were experimented on, which had been dried and preserved in an herbarium for 100 years, and the moving molecules or small spheri¬ cal bodies were still perceived in considerable numbers. “ The very unexpected fact,” says Mr Brown, “ of seem¬ ing vitality retained by those minute particles so long after the death of the plant, would not perhaps have ma¬ terially lessened my confidence in the supposed peculia¬ rity ; but I at the same time observed, that on bruising the ovula or seeds of Equisetum, which at first happened accidentally, I so greatly increased the number of moving particles, that the source of the added quantity could not be doubted. I found also, on bruising first the floral •leaves of mosses, and then all other parts of those plants, that I readily obtained similar particles, not in equal quantity indeed, but equally in motion. My supposed test of the male organ was therefore necessarily abandon¬ ed. Reflecting on all the facts with which I had now be¬ come acquainted, I was disposed to believe that the mi¬ nute spherical particles or molecules of apparently uni¬ form size, first seen in the advanced state of the pollen of Onagrariae, and most other phaenogamous plants,—then in the antherae of mosses, and on the surfaces of the bo¬ dies regarded as the stamina of Equisetum,—and, lastly, in bruised portions of other parts of the same plants,—were in reality the supposed constituent or elementary mole¬ cules of organic bodies, first so considered by Button and Needham, then by Wrisberg with greater precision, soon after and still more particularly by Muller, and very re¬ cently by Dr Milne Edwards, who has revived the doc¬ trine, and supported it with much interesting detail. I A! now, therefore, expected to find these molecules in all organic bodies ; and accordingly, on examining the variM ous animal and vegetable tissues, whether living or dead, they were always found to exist; and merely by bruis¬ ing these substances in water, I never failed to disen¬ gage the molecules in sufficient numbers to ascertain their apparent identity in size, form, and motion, with the smaller particles of the grains of the pollen. I ex¬ amined also various products of organic bodies, parti- cularly the gum raisins, and substances of vegetable ori¬ gin, extending my inquiry even to pit-coal; and in all these bodies molecules were found in abundance. I re¬ mark here also, partly as a caution to those who may hereafter engage in the same inquiry, that the duct or soot deposited on all bodies in such quantity, especially in London, is entirely composed of these molecules. One of the substances examined was a specimen of fossil wood, found in Wiltshire oolite, in a state to burn with flame; and as I found these molecules abundantly and in motion in this specimen, I supposed that their existence, though in smaller quantity, might be ascertained in mineralized vegetable remains. With this view, a minute portion of sili- cified wood, which exhibited the structure of conifers, was bruised, and spherical particles, or molecules in all respects like those so frequently mentioned, were readily obtained from it; in such quantity, however, that the whole sub¬ stance of the petrifaction seemed to be formed of them. But hence I inferred that these molecules were not li¬ mited to organic bodies, nor even to their products. To establish the correctness of the inference, and to ascertain to what extent the molecules existed in mineral bodies, became the next object of inquiry. The first substance examined was a minute fragment of window-glass, from which, when merely bruised on the stage of the micro¬ scope, I readily and copiously obtained molecules, agree¬ ing in size, form, and motion, with those which I had already seen. I then proceeded to examine, and with similar results, such minerals as I either had at hand or could readily obtain, including several of the simple earths and metals, with many of their combinations. Rocks of all ages, including those in which organic remains have never been found, yielded the molecules in abundance. Their existence was ascertained in each of the constituent minerals of granite, a fragment of the sphinx being one of the specimens examined. To mention all the mineral sub¬ stances in which I have found these molecules would be tedious; and I shall confine myself, in this summary, to an enumeration of a few of the most remarkable. These were both of aqueous and igneous origin, as travertine, stalactites, lava, obsidian, pumice, volcanic ashes, and me¬ teorites from various localities. Of metals I may mention manganese, nickel, plumbago, bismuth, antimony, and arsenic. In a word, in every mineral which I could re¬ duce to powder sufficiently fine to be temporarily sus¬ pended in water, I found these molecules more or less co¬ piously ; and in some cases, more particularly in siliceous crystals, the whole bodyr submitted to examination ap¬ peared to be composed of them.” There were three points of importance which Mr Brown was anxious to ascertain regarding these molecules, viz- their form, whether they were of uniform size, and their absolute magnitude. He seems, however, not to have been entirely satisfied with his determination on any of these points. As to form, he states the molecules to be spherical. • His manner of estimating the absolute nragn1' tude and uniformity of size of the molecules found in the various bodies submitted to examination, ^yas by plac>nb’ them on a micrometer divided to five thousandths of an inch, the lines of which were very distinct; or, more rare- A N J A N K 193 ■ on one divided to ten thousandths, with fainter lines,not ctile is of uniform size, though, as existing in various sub- Anker ■adily visible without the application of plumbago, as em- stances, and examined in circumstances more or less oyed by Dr Wollaston, but which in this case was inad- favourable, he regards it necessary to state that its dia-( issible. The results can only be regarded as approxi- meter appeared to vary from -jy^oth to 20000th of an' Anklam. ate, but Mr Brown is disposed to believe that the mole- inch.1 (t.) ANIMATED, or Animate, in a general sense, denotes tmething endowed with animal life. ANIMATION signifies the communication of life to an limal body. The different hypotheses of physicians and lilosophers concerning die time of animation have had ieir influence on the penal laws made against artificial >ortions; it having been made capital to procure mi scar- age in the one state, while in the other it was only deem- 1 a venial crime. The emperor Charles V., by a consti- ition published in 1532, put the matter on another foot- g : instead of the distinction of an animated and unani- ated foetus, he introduced that of a vital and non-vital ;tus, as a thing of more obvious and easy decision, and it depending on any system either of creation, traduc- pn, or infusion. Accordingly a foetus was formerly said, a legal sense, to be animated when it was perceived to ir in the womb ; but this doctrine is exploded, animation }ing now dated from the moment of conception. ANIME, in Heraldry, a term used when the eyes of a pacious creature are borne of a different tincture from ie creature itself. Anime, a resin exuding from the trunk of a large Ame- can tree, called by the Indians courbaril (a species of ymen;ea). This resin is of a transparent amber colour, light agreeable smell, and little or no taste. It dis- ilves entirely, but not very readily, in rectified spirit ' wine; the impurities, which are very often in large lantity, remaining behind. The Brazilians are said to nploy anime in fumigations for pains and aches proceed- ig from cold: with us, it is rarely, if ever, made use of r any medicinal purpose. ANIMETTA, among ecclesiastical writers, denotes the oth wherewith the cup of the eucharist is covered. ANINGA, in Commerce, a root which grows in the ntilles Islands, and is pretty much like the China plant. . is used by sugar-bakers for refining the sugar. ANJOU, a province and duchy of France before the evolution, bounded on the east by Touraine, on the south y Poitou, on the west by Bretagne, and on the north by Maine. It is now included under the Maine and Loire and the Sarthe and Mayenne. It is 70 miles in length, and in breadth 60. Through this province run five navi¬ gable rivers; the Loire, which divides it into two parts; the Vienne, the Toue, the Mayenne, and the Sarthe. The air is temperate, and the country agreeably diver¬ sified with hills and meadows. There are 33 forests of oak-trees mixed with beech. The country produces white wine, wheat, barley, rye, oats, peas, beans, flax, hemp, walnuts, and some chesnuts. In Lower Anjou they make cider. There are fruit-trees of all kinds, and pasture proper for horses. The greatest riches of the province consist in cows, oxen, and sheep. There are several coal and iron mines, and yet there are but two forges in the whole province. There are quarries of marble and of slate, as well as quarries of white stone, proper for building, on the side of the river Loire. Here are also several salt¬ petre works, and some glass-houses. The principal towns, besides Angers the capital, are, Saumur, Brisac, Pont de Ce, La Fleche, and Beaufort. ANKER, a liquid measure at Amsterdam. It contains about 32 gallons English measure. ANKLAM, a circle in the government of Stettin and the Prussian province of Pomerania. It extends over 566 square miles, or 363,240 English acres. There are within it 4 cities, 6 towns, and 20l villages. The inhabitants are 30,856. The land is a plain, with extensive woods, and about twenty fresh-water lakes, the largest of which is the Ahlbeckr. The feeding of cattle and growing of corn are the chief objects of agriculture ; besides which, some hops and tobacco, and much flax, are grown. The woods afford much profitable employment, and furnish charcoal to the iron-works in Pomerania. The capital of the circle is of the same name. It is situated on the river Peene, and is now without fortifications. It contains 3 churches, 3 hospitals, 599 houses, and 5833 inhabitants. By means of the river, which is navigable, it carries on some trade; and it has manufactures of cloth, hosiery, tobacco, snuff, and leather. It is in long. 24. 1. 59. E. lat. 53. 49. 15. N. 1 The following summary from the pen of Mr Brown contains the renewed expression of that gentleman’s opinion, matured by >me recent experiments on the subject of active molecules. “ That extremely minute particles of solid matter, whether obtained om organic or inorganic substances, when suspended in pure water or in some other aqueous fluids, exhibit motions for which I am nable to account, and which, from their irregularity and seeming independence, resemble in a remarkable degree the less rapid mo¬ ons of some of the simplest animalcules of infusions. That the smallest moving particles observed, and which I have termed Ac- ve Molecules, appear to be spherical, or nearly so, and to be between EC5S5th and 3550 oth of an inch in diameter; and that other irticles of considerably greater and various size, and either of similar or of very different figure, also present analogous mo- i ons in like circumstances. I have formerly stated my belief that these motions of the particles neither arose from currents in the uid containing them, nor depended on that intestine motion which may be supposed to accompany its evaporation. These causes j motion, however, either singly or combined with others,—as the attractions and repulsions among the particles themselves, their nstable equilibrium in the fluid in which they are suspended, their hygrometrical or capillary action, and in some cases the disengage- icnt of volatile matter, or of minute air-bubbles,—have been considered by several writers as sufficiently accounting for the appear- ices. Some of the alleged causes here stated, with others which I have considered it unnecessary to mention, are not likely to be cerlooked, or to deceive observers of any experience in microscopical researches ; and the insufficiency of those enumerated may, I nnk, be satisfactorily shown by means of a very simple experiment. This experiment consists in reducing the drop of water con- umng the particles to microscopic minuteness, and prolonging its existence by immersing it in a transparent fluid of inferior specific ravity, with which it is not miscible, and in which evaporation is extremely slow. If to almond oil, which is a fluid having these roperties, a considerably smaller proportion of water, duly impregnated with particles, be added, and the two fluids shaken or tritu- ited together, drops of water of various sizes, from j’^th to S!jl55th of an inch in diameter, will be immediately produced. Of these, ie most minute necessarily contain but few particles, and some may be occasionally observed with one particle only. In this man- er minute drops, which, if exposed to the air, would be dissipated in less than a minute, may be retained for more than an hour. ’Ut in all the drops thus formed and protected, the motion of the particles takes place with undiminished activity, while the nncipal causes assigned for that motion, namely, evaporation and their mutual attraction and repulsion, are either materially re- ■iced or absolutely null.” VOL. in. 2 B 194 ANN ANN Ann Anna. ANN, or Annat, in Scotish Law, is half a year’s stipend, which the act 1672, c. 13, gives to the executors of mi¬ nisters of the church of Scotland, oyer and above what ;was due to the minister himself for his incumbency. s it is a mere gratuity given by the law to those whom, it is presumed, the deceased could not sufficiently provide for, so it is neither assignable by him during his life, nor a - tachable by his creditors after his death. ANNA, Ana, or Anah, a town of Arabian Iiac, pachalic of Bagdad, which extends five or six miles along the western bank of the Euphrates. It consists of a single street built on both sides. The houses are of stone, two stories high, and separated from each other, as m other eastern towns, by beautiful gardens, filled with fruit-trees, bearing lemons, oranges, citrons, quinces, figs, dates, pome¬ granates, and olives. It is an open and defenceless place; and in 1807 it was attacked by the Wahabees, who gave it up to plunder, and perpetrated the most horrible cruel¬ ties, massacring the greater part of the inhabitants, and setting the town on fire; after which they retreated with their plunder, carrying into captivity many women and children. The inhabitants are said, previous to this ca¬ lamity, to have been more polished than those in the neighbourhood, and to have consisted chiefly of Arabians, who were, however, addicted to their usual^ vocation ot robbery when any opportunity offered. Population about 3000. 260 miles east of Damascus; 220 south-east ot Aleppo. Long. 41. 15. E. Eat. 34. N. . Anna Comnena, daughter of the emperor Alexius Lom- nenus I., was not less distinguished by her elevated rank than by her mental qualifications. Her superiority of mind began early to display itself. Despising the effemi¬ nacy and voluptuousness of the court in which she was educated, she directed her attention to literary pursuits. Indulging her favourite studies, she solicited the acquaint¬ ance of the more eminent philosophers of that period. But the pursuits of literature did not induce her en¬ tirely to abandon society; she gave her hand to Nicepho- rus Briennius, a young nobleman of a respectable family. This accomplished woman was, however, actuated with unjustifiable ambition; and, during the last illness of her father, she united with the empress Irene in attempting to prevail upon that monarch to disinherit his own son, and give the crown to her husband. The affection and virtue of the father prevailed over female address and intrigue. But the ambition of Comnena was not diminished ; for she entered into a conspiracy to depose her brother ; and w*hen her husband displayed a timidity and hesitation in this unjust enterprise, she exclaimed that “ nature had mistaken their sexes, for he ought to have been the woman.” Either through the vigilance of her brother, or the ti¬ midity of her husband, the treasonable plot was discover¬ ed, and Anna punished with the confiscation of all her property. But generosity has an opportunity of display¬ ing its real nature when an enemy is vanquished; thus was the -generosity of her brother displayed on the pre¬ sent occasion, by returning all her property. Ashamed, however, of her base conduct, she retired from court, and never more possessed any influence there. Disappointed ambition took shelter among the walks of literature, and she employed herself in her solitude in writing the his¬ tory of her father’s reign. This production of her pen is still extant, and composes a part of the celebrated collec¬ tion of the Byzantine Historians. The stores of rhetoric are ransacked to embellish this work, and every effort made to enrich it with science; but the general com¬ plexion of it is rather like an apology than an impartial narrative. It must, however, be acknowledged that she is not more partial than many other Latin historians, and that her history contains many valuable facts and obser-, vations. , ANNABERG, a city on the mountains m the bailiwick Am ^ of Wolkenstein, in the circle of Erzgebirg, in the kingdom^' i/'J of Saxony. It is 2820 feet above the level of the sea, in the midst of a mining district, which yields silver, tin, and cobalt. It contains 592 houses, and 4500 inhabitants, who are employed in manufacturing tapes, lace, sewing-silk, and many other articles. It is in latitude 50. 35.8. N. The longitude is not exactly ascertained. ANNACHNAN, a small island of Ireland, on the south- west coast of the county of Galway, 22 miles from Galway, ANNAGH, an island about 5 miles in circumference, on the west coast of Ireland, between the isle of Achil and the mainland of the county of Mayo. Long. 9.39. W. Lat. 53. 58. N. There is a small village of the same name in the county of Cork, 5 miles from Charleville. ANNALS, in matters of literature, a species of history which relates events in the chronological order wherein they happened. They differ from perfect history in this, that annals are but a bare relation of what passes every year, as a journal is of what passes every day: whereas history relates not only the transactions themselves, but also the causes, motives, and springs of actions. Annals require nothing but brevity; history demands ornament. Cicero gives the following account of the origin of annals, To preserve the memory of events, the Pontifex Maximus, says he, wrote what passed each year, and exposed it on tables in his own house, where every one was at liberty to read: this they called annales maximi ; and hence the writers who imitated this simple method of narrating facts were called annalists. ANNAN, a royal borough and parish of Scotland, in the county of Dumfries, situated on the river of the same name, about two miles above its junction with the Solway Frith. An elegant new bridge of three arches has been built within these three years. It has a good harbour; the highest tides rise 21 feet; 23 vessels belonged to the port in 1818, registering 1025 tons; at present (1830) there are 39 vessels, registering 3054 tons. These vessels are chiefly employed in the coasting trade. A cotton spin¬ ning manufactory has been long established in the town, giving employment to between 80 and 100 persons of different ages. The salmon fishery, which in former years was so very productive, has fallen off very considerably. The town has recently been much improved by the addition of several new streets and public buildings: among the latter is a handsome new academy, built and endowed by the heritors and borough-council; it is conducted by a rec¬ tor and two masters, and is in a very flourishing state. The town unites with Dumfries, Kirkcudbright, Sanquhar, and Lochmaben, in sending a representative to parliament. The environs are very inviting; and few places are more beautiful than the river and its finely wooded banks, for eight or ten miles above the town. There was formerly a castle, built by the Bruces after they became lords ot Annandale. The population of the parish is 4500, of the town 3000. The latter is 16 miles from Dumfries, and 78 from Edinburgh. . . The river Annan, on which the town stands, rises in the county of Peebles, and flowing through Dumfriesshire, falls into the Solway Frith after a course of 30 miles, abounds with trout and salmon. The stewartry or dis¬ trict of Annandale, of which Lochmaben castle was t e chief fortalice, is a fertile vale, 24 miles long and abou 14 miles broad. From its vicinity to England, and t e continual incursions and predatory wars of the borderers, the greater part of it was uncultivated and common; u since the beginning of the last century all these was and commons have been subdivided and brought in ANN ANN 195 nd ' oils jlture, and the country has assumed a new appearance, hich may be ascribed not only to the division of the com- • ions, but likewise to the improvement made on the roads. ' Annandale formed a part of the Roman province of Va- mtia; and Severus’s wall ending at Bowness on the op- osite coast of the Solway, it abounds with Roman sta- ons and antiquities. The RdVnan camps at Birrens in Middlebie, on the hill of irrenswark, and at Torwood Muir in Dryfesdale, are still early entire, and their form is preserved; and the traces id remains of a military road are yet visible in different irts of the country. The ruins of the house or castle of uchincass, in the neighbourhood of Moffat, once the :at of that potent baron Thomas Randolph, earl of lurray, lord of Annandale, and regent of Scotland dur- g the minority of David II., covers above an acre of •ound, and even now conveys an idea of the plan and rength of the building. The ancient castle of Com- ngan, formerly belonging to the Murrays, earls of An- indale, and now to the earl of Mansfield, is still in a tole- ible state of preservation ; but except this castle, and that ■ Hoddam, most of the other old fortalices and towers are dw taken down or in ruins. ANNAND, William, dean of Edinburgh, the son of Villiam Annand, minister of Ayr, was born at Ayr in 333. Five years after, his father was obliged to quit cotland with his family, on account of their loyalty to ic king, and adherence to the episcopal government itablished by law in that country. In 1651 young An- ind was admitted a scholar in University College, Ox- rd; and though he was put under the care of a pres- jterian tutor, yet he took all occasions to be present I; the sermons preached by the loyal divines in and jar Oxford. In 1656, being then bachelor of arts, j received holy orders from the hands of Dr Thomas ulwar, bishop of Ardfert or Kerry in Ireland, and was ipointed preacher at Weston on the Green, near Bices- r in Oxfordshire. After he had taken the degree of aster of arts, he was presented to the vicarage of eighton-Buzzard in Bedfordshire, where he distinguished imself by his edifying manner of preaching till 1662, hen he went into Scotland in quality of chaplain to )hn earl of Middleton, the king’s high commissioner to le church of that kingdom. In the latter end of the sar 1663 he was instituted to the Tolbooth Church at dinburgh, and from thence was removed, some years ifter, to the Tron Church of that city. In April 1676 e was nominated by the king to the deanery of Edin- urgh; and in 1685 he commenced doctor of divinity in le university of St Andrews. He wrote, 1. Fides Ca- dica, or the Doctrine of the Catholic Church. Lond. 561-2, 4to. 2. Solutions of many proper and profitable uestions; printed with the Fides Catholica. 3. Panem }mtidianum, or a short Discourse tending to prove ie legality, decency, and expediency of set forms of rayers in the churches of Christ; with a particular De- nce of the Book of Common Prayer of the Church of ngland. Lond. 1661, 4to. 4. Pater Noster, or the ord’s Prayer explained. Lond. 1670, 8vo. 5. Myste- um Pietatis, or the Mystery of Godliness. Lond. 1672, \ro. 6. Doxologia, or Glory to the Father, the Church’s lymn, reduced to glorifying the Trinity. Lond. 1672, vo. 7. Dualitas, or a twofold subject displayed and pened, conducible to godliness and peace in order: irst, Lex Loquens, the honour and dignity of magistracy; econdly, Duorum Unitas, or the agreement of magis- acy and ministry. Edinb. 1674, 4to. Dr Annand died n the 13th of June 1689, and was interred in the Grey- "lars Church, Edinburgh. ANNAPOLIS, the chief town in Maryland, in North America. It stands upon a sort of peninsula on the west side of the Chesapeak, and is a small town, but well built. It contained 2260 inhabitants in 1820. ANNE, queen of Great Britain, second daughter of King James II. by his first wife, Anne Hyde, was born in 1664. In 1683 she married George prince of Den¬ mark, by whom she had several children, but none of them arrived at the age of maturity. On the death of King William she ascended the throne, a. d. 1702, and her reign comprehends one of the most illustrious periods of English history. Possessed, however, of a very feeble character, which did not permit her to act for herself, this period is the reign of her counsellors and favourites; and she exhibited no decided inclination which could in¬ fluence state affairs, except a strong passion for tory principles, both in church and state. She died in Au¬ gust 1714, of a dropsy, in the 50th year of her age and 13th of her reign. ANNE Boleyn, queen of Henry VIII. king of Eng¬ land, daughter of Sir Thomas Boleyn, a nobleman of a powerful family and numerous alliances. The daughter of the duke of Norfolk was her mother, and during the reign of the former king her father had been honoured with several embassies. Mary, the king’s sister, who mar¬ ried Louis XII. king of France, carried over this lady with her at an early age, where she imbibed the freedom, the vivacity, and the openness of manners of that nation. After the death of Louis, that queen returned to Eng¬ land, and Anne continued to attend her royal mistress. Having some time after left her service, she was intro¬ duced into the family of the duchess of Alen^on. In addition to all her acquired accomplishments, she pos¬ sessed the greatest personal elegance, and was highly famed in that age. History does not explicitly mention whether or not it was on her account, but upon her return to England the king expressed his scruples concerning his union with Catharine of Arragon. Enamoured, however, of Anne, he expressed his attachment to her ; but she was possessed of too much virtue and policy to confer any improper fa¬ vours. This prudent and virtuous restraint only increased the passion of Henry; and placing her at court, he dis¬ tinguished her by many marks of royal favour. The im¬ petuous king at length came to the resolution to divorce his queen, to make way for his favourite Anne. In this instance the injury done to that queen proved the cause of the final separation of England from the dominion of the pope. Various delays and difficulties occurring to the divorce, Henry privately married Anne during the month of November 1532; and in April following he publicly de¬ clared her queen of England. The famous Queen Eliza¬ beth was the first-fruits of this marriage, who was born in September following. For some time Anne enjoyed a considerable share of the royal favour, and she made use of that influence in subduing the haughty prelate Wolsey, and widening the breach between the king and the pope. But this favour was not of long continuance; for the king, ever varying in his temper, allowed jealousy to enter his bosom, which her thoughtless demeanour tended in some measure to excuse. She was accused of adultery with several of the household officers, and even with her own brother Lord Rochford; and having been tried on a charge of high treason, was condemned on very scanty proof to be beheaded ; which sentence was execut¬ ed in May 1536. She to the last resolutely denied any serious guilt. St Anne’s Day, a festival of the Christian church, ce¬ lebrated by the Latins on the 26th of July, but by the Greeks on the 9th of December. It is kept in honour of Anne or Anna, mother of the Virgin Mary. Anne St Anne’s Day. 196 Annealing. Glass. ANN ANNEALING, by the workmen called nealing, is a process used in glass-making, and in the manufactui e of certain metals. In glass-making it consists in placing the bottles, &c. whilst hot, in a kind of oven or furnace, where they are suffered to cool gradually. They would other¬ wise be too brittle for use. dhe difference between un¬ annealed and annealed glass, with respect to brittleness, is very remarkable. When an unannealed glass vessel is broken, it often flies into a small powder, with a violence seemingly very unproportioned to the stroke it has re¬ ceived. In general it is in greater danger of breaking from a very slight stroke than from one of some consider¬ able force. One of these vessels will often resist the ef¬ fects of a pistol bullet dropt into it from the height of two or three feet; yet a grain of sand falling into it will make it burst into small fragments. This takes place sometimes immediately on dropping the sand into it; but often the vessel will stand for several minutes after, seemingly se¬ cure ; and then, without any new injury, it will fly to pieces. If the vessel be very thin, it does not break in this manner, but seems to possess all the properties of annealed glass. The same phenomena are still more strikingly seen in glass drops or tears. They are globular at one end, and taper to a small tail at the other. They are the drops which fall from the melted mass of glass on the rods on which the bottles are made. They drop into the tubs of water which are used in the work; the greater part of them burst immediately in the water. When those that remain entire are examined, they discover all the proper¬ ties of unannealed glass in the highest degree. They will bear a smart stroke on the thick end without breaking; but if the small tail be broken, they burst into small powder with a loud explosion. They appear to burst with more violence, and the powder is smaller, in an exhausted receiver, than in the open air. When they are annealed they lose these properties. Glass is one of those bodies which increase in bulk when passing from a fluid to a solid state. W hen it is al¬ lowed to crystallize regularly, the particles are so arranged that it has a fibrous texture. It is elastic, and suscep¬ tible of long-continued vibrations; but when a mass of melted glass is suddenly exposed to the cold, the surface crystallizes, and forms a solid shell round the interior fluid parts. This prevents them from expanding when they become solid. They therefore have not the opportunity of a regular crystallization, but are compressed together with little mutual cohesion. On the contrary, they press outward to occupy more space, but are prevented by the external crust. In consequence of the effort of expansion in the internal parts, the greater number of glass drops burst in cooling; and those which remain entire are not regularly crystallized. A smart stroke upon them com¬ municates a vibration to the whole mass, which is nearly synchronous in every part; and therefore the effort of ex¬ pansion has little more effect than if the body were at rest; but the small tail and the surface only are regularly crystallized. If the tail be broken, this communicates a vibration along the crystallized surface, without reaching the internal parts. By this they are allowed some ex¬ pansion ; and overcoming the cohesion of the thin outer shell, they burst it, and are dispersed in powder. In an unannealed glass vessel the same thing takes place. Sometimes the vibration may continue for a con¬ siderable time before the internal parts overcome the re¬ sistance. If the vessel be very thin, the regular crystal¬ lization extends through the whole thickness; or at least the quantity of compressed matter in the middle is so incon¬ siderable as to be incapable of bursting the external plate. By the process of annealing the glass is kept for some time in a state approaching to fluidity; the heat increases ANN the bulk of the crystallized part, and renders it so soften, that the internal parts have the opportunity of expanding J and forming a regular crystallization. In the manufactures in which the malleable metals are j, employed, annealing is used to soften a metal after it has been rendered hard by the hammer; and also to soften cast-iron, which is rendered very hard and brittle by ra¬ pid cooling. In the manufacture of steel goods, which are first form¬ ed by the hammer, and require to be filed or otherwise treated, and in which softness and flexibility are essential to the change, annealing is absolutely necessary. This is particularly the case in making files and scissors, that the metal may be left sufficiently soft for cutting the teeth, and for filing off those parts which cannot be ground. An¬ nealing is not less necessary in the drawing of wire, whe¬ ther iron, copper, brass, silver, or gold. The operation of drawing soon gives the wire a degree of hardness and elasticity which, if not removed from time to time by an¬ nealing, would prevent the extension of the wire, and ren- der it extremely brittle. The same operation is also ne¬ cessary in rolling or flatting those metals which are in a cold state, such as brass, silver, gold, &c. The brazier who forms vessels of copper and brass by the hammer, can work upon it only for a little time before he is obliged to anneal it. The common methods employed for annealing iron and steel are very injudicious, and materially injure the latter when it is used for making cutting instruments. After they have been formed by the hammer, they are generally piled up in an open fire, slowly raised to red heat, and then allowed as gradually to cool. By this method the surface of the steel will be found considerably scaled, from the action of the oxygen of the atmosphere. When it is re¬ membered that steel consists of iron joined to carbon, it will be evident that the steel immediately under the scaly oxide will be deprived of its carbon, which has been car¬ ried off by the attraction of the oxygen; and, in conse¬ quence, will lose the property of acquiring that degree of hardness necessary to a cutting instrument. Nothing, therefore, can be more obvious, than that steel particularly should be annealed in close vessels, to prevent that effect. For this purpose the goods should be placed in a trough or recess made of fire-stone or fire-brick, and stratified with ashes or clean sand, and finally covered with a thick stratum of the same; but if the size of the vessel be small, it may have a cover of its own materials. This oven or trough must now be heated by the flame of a furnace passing under and round it, till the whole is of a red heat. It must then be suffered to cool, without let¬ ting in the air. The goods so treated will be much softer than by the common method. The surface, instead of becoming scaled, will have acquired a metallic whiteness, from the presence of a small quantity of carbonaceous matter contained in the ashes in which they were imbed¬ ded. They will become so flexible also, as to allow them to bend considerably without breaking, which is very far from being the case before the operation. The fracture, before annealing, will be smooth and short; but afterwards it will be rough, exhibiting bright parts, of a crystalline ap¬ pearance. Wire, especially that of iron and steel, should be treated in a similar way when it is annealed, h’6 wire used for some purposes requires to be soft, and is sold in that state. If the wire, after finishing, when it i> bright and clean, were to be annealed in contact with oxy¬ gen, it would not only lose all its lustre and smoothness, but much of its tenacity. The process above mentione will therefore be particularly necessary in annealing nms * ed wire, as well as in softening it from time to time during the drawing. ANN ANN 197 ey. Conner and brass suffer much less than iron and steel ' om annealing in the open air and do not require to be noted above a low red heat. If, however, the lustre is to „ preserved, a close vessel would be desirable. The lat- >r metals, after annealing, although much discoloured v the oxygen of the atmosphere, may be cleansed by nmersion in a hot liquor composed of water and a small uantity of sulphuric or nitric acid. Very small brass or inper wire is frequently annealed by exposing it to the ame of hay or straw. In casting minute pieces of pig- on, which is generally done in wet sand, the metal pos- >sses the property of steel to such a degree as to assume, y the rapid cooling, a degree of hardness equal to hard- led steel; at the same time that the articles are so brittle ; to break by falling on the ground. When, however, icse goods are treated in the way above directed, they -.quire a degree of softness which renders them pene- able by the file, and at the same time capable of bend- ig. In this state they are much less tenacious than steel, ut still so much so as to have been sold in the form of jtlery for steel. The change which metals undergo by annealing is not ct thoroughly understood. Most of the malleable metals »re susceptible of two distinct forms, one called the crys- illine form, which they assume by slow cooling; and the ther the fibrous, which is acquired by hammering or roll- ig. When this, however, is carried beyond a certain oint, the metal becomes so hard that it is not capable of eing bent far without breaking. All the malleable me¬ tis in the ingot or in their cast state are brittle, and ex- ibit a crystalline fracture. By hammering or rolling icy become more tenacious, and break with difficulty, xhibiting what is called a fibrous fracture. At the same me they become stiffer and more elastic. They lose the itter properties by annealing, but become more malleable. . f the annealing, however, be long continued, the mal- ;ability diminishes, and they again have a crystalline •acture. Zinc by wire-drawing becomes very flexible, nd possesses a degree of tenacity not inferior to that of opper; but, if it be kept in boiling water for a length of me, it will resume its original brittleness, and show a rystalline appearance when broken. This proves that ie particles of metals can change their arrangement 'ithout losing their solid form ; which is still more strong- f confirmed by the fact, that brass wire loses its tenacity y exposure to the fumes of acids, and even by the pre- ence of a damp atmosphere. This is not caused by the noisture, but by the action of air upon the moistened sur- ace. The manufacturers of common pins are obliged to eep their wire in a dry atmosphere, or immersed in /ater. If the wire be first moistened, and then exposed o the air, it will assume the brittle state much sooner, n this condition it breaks with a crystalline fracture, si- ailar to that exhibited by an ingot. When a steel plate, uch as a watch-spring, has been once tempered, the ope- ation of simply rubbing it bright will render it soft and lastic. The same change is brought about by slightly lamrnering it. It, however, resumes its elastic state by icing carefully heated till it becomes of a blue colour, f the heat be continued to redness, particularly in a close 'essel, it becomes perfectly annealed. ANNECY, a city in the kingdom of Sardinia, the capi- al of the province of Genevois, in the duchy of Savoy. It is •t the foot of the mountain Semina, on the banks of the ake of that name. It is the most industrious place in iavoy, having manufactures of cotton goods, of hats, glass, aid earthenware, and several distilleries and tanneries, aid near it some iron-works. It contains a cathedral, a diurch, five monasteries, and the same number of nun- icries. The relics of St Francis de Sales, preserved in St Mary’s Church, draw a vast number of pilgrims annu- Annesley ally to the city. The population in 1816 amounted to II 5467 persons. It is in long. 5. 9. E. and lat. 45. 53. N. Annobon. ANNESLEY, Arthur, earl of Anglesea, and lord privy seal in the reign of King Charles II., was the son of Sir Francis Annesley, baronet, Lord Mount-Norris, and Viscount Valentia, in Ireland, and was born at Dublin on the 10th of July 1614. He was for some time at the uni¬ versity of Oxford, and afterwards studied the law at Lin¬ coln’s Inn. He had a considerable share in public trans¬ actions, for in the beginning of the civil war he sat in the parliament held at Oxford; but afterwards became recon¬ ciled to the opposite party, and was sent commissioner to Ulster, to oppose the designs of the rebel Owen Roe O’Neal. He engaged in several other affairs with great success. He was president of the council of state after the death of Oliver, and was principally concerned in bringing about the Restoration, soon after which King Charles II. raised him to the dignity of a baron, by the title of Lord Annesley of Newport-Pagnel, Bucks; and a short time after he was made earl of Anglesea. Du¬ ring that reign he was employed in some very important affairs, was made treasurer of the navy, and for some time held the office of lord privy seal. He was a person of great abilities, of very extensive learning, and was well ac¬ quainted with the constitution and laws of England. He died in April 1686, in the 73d year of his age. In his lifetime he published the following pieces:—1. Truth un¬ veiled, in behalf of the Church of England; being a Vin¬ dication of Mr John Standish’s Sermon, preached be¬ fore the king, and published by his Majesty’s command. 1676, 4to. To which is added, A short Treatise on the subject of Transubstantiation. 2. A Letter from a Person of Honour in the Country, written to the Earl of Castle- haven ; being observations and reflections on his Lord¬ ship’s Memoirs concerning the Wars of Ireland. 1681, 8vo. 3. A true Account of the whole Proceedings between James Duke of Ormond and Arthur Earl of Anglesea, before the king and his council, &c. 1682, folio. 4. A Letter of Re¬ marks upon Jovian. 1683, 4to. Besides these, he wrote several other works, some of which were published after his decease ; as, 5. The Privileges of the House of Lords and Commons, argued and stated in two conferences be-r tween both Houses, April 19 and 22, 1671: To which is added, A Discourse, wherein the Rights of the House of Lords are truly asserted; with remarks on the seeming arguments and pretended precedents offered at that time against their lordships. 6. The King’s Right of Indulgence in Spiritual Matters, with the Equity thereof, asserted. 1688, 4to. 7. Memoirs, intermixt with moral, political, and historical observations, by way of discourse, in a let¬ ter to Sir Peter Pett. 1693, 8vo. ANNIVERSARY, the annual return of any remarkable day. Anniversary days, in old times, more particularly denoted those days in which an office was yearly perform¬ ed for the souls of the deceased, or the martyrdom of the saints was yearly celebrated in the church. ANNOBON, a small island in Africa, on the east coast of Loango, belonging to the Portuguese. It lies in long. 5. 30. E. lat. 1. 32. S., and receives its name from being discovered on the new year’s day. According to Pyrard, it is about five or six French leagues in circuit; Braud- raud makes it ten leagues. It contains two high moun¬ tains, having their tops continually enveloped in clouds, and thus occasioning frequent rains. Off the south-east of the island are two rocks, one of which is low, and upon a level with the surface of the sea; the other higher and larger, but both dangerous to shipping in the night: between them the channel is deep and clear. On the same side of the island is a convenient watering place; but the road 198 ANN Annobon to the north-west side is difficult and dangerous, though II the one most frequented by ships that have no intention Annuities. touching upon the continent. In either place it is dif- ficult to take in a sufficient quantity of water, on account of the violent surf. The best road for ships lies on the north-east side, where they may anchor in 7, 10, 13, or 16 fathoms, on a fine sand, opposite to the Negro village. The climate is wholesome, and the air clear and serene for the greater part of the year. The island contains a number of fertile valleys, which produce Turkey corn, rice, millets, yams, potatoes, and afford pasture for abundance of cattle and sheep. Poultry and fish also abound; but the only mercantile production is cotton, in small quantity, but esteemed equal in quality to any produced in India. In the year 1605 the Dutch admiral Matelief found 200 negroes and two Portuguese on Annobon, most of them able to bear arms. The inhabitants are subject to the Portuguese governor, who is the chief person in the island; at the same time the negroes have their own chief, subordinate to him. They are all rigid catholics, having been either compelled or persuaded by the Portuguese to embrace that religion; and, like all other converts, they are bigoted in proportion to their ignorance. ANN ANNONA, in Roman Antiquity, denotes provision for a year of all sorts, as of flesh, wine, &c. but especially 0f \ corn. Annona is likewise the allowance of oil, salt, bread M flesh, corn, wine, hay, and straw, which was annually pr0\ vided by the contractors for the maintenance of an army, ANNONiE Pr^fectus, in Antiquity, an extraordinary magistrate, whose business it was to prevent a scarcity of provisions, and to regulate the weight and fineness of bread. ANNONAY, a small town of France, in the depart- ment of Ardeche, formerly Upper Vivarais, seated on the river Deume. Long. 4. 52. E. Lat. 45. 15. N. ANNOT, a small city on the mountains of Provence in France. Long. 7. 0. E. Lat. 44. 4. N. ANNOVER, a Spanish town not far from the Tagus, in the province of Toledo, containing 400 houses, 2000 inhabitants, and a saltpetre manufactory. ANNUALRENT, in Scotish Law, denotes the yearly interest or profit due by a debtor in a sum of money to a creditor for the use of it.—A Right of Annualrent was the original method in Scotland of burdening lands with a yearly payment for the loan of money, before the tahimr of interest was allowed. ANNUITIES.1 The doctrine of Compound Interest and Annuities-cer- tain is too simple ever to have occupied much of the at¬ tention of mathematicians : inquiries into the values of in¬ terests dependent upon the continuance or the failure of human life, being more interesting and difficult, have oc¬ cupied them more, but yet not so much as their import¬ ance would seem to demand ; the discoveries both in Pure Mathematics and Physics, especially those of Newton, which distinguished the close of the seventeenth century, having provided them with ample employment of a more interesting kind, ever since the subjects of this article were submitted to calculation. Fermat, Pascal, and Huygens, by laying the foundation of the doctrine of probabilities, about the middle of that century, first opened the way to the solution of problems of this kind. The earliest mathematical publication on probabilities, the little tract of Huygens, De Ratiociniis in Ludo Alecs, appeared in 1658; and in 1671 his celebrated countryman John de Witt published a treatise on Life- Annuities in Dutch. (Montucla, Hist, des Math, tome iii. p. 407.) This, however, appears to have been very little known or read, and to have had no sensible influence on the subsequent progress of the science, the origin of which may be properly dated from the publication of Dr Halley’s paper on the subject, in the Philosophical Transactions for the year 1693 (No. 196). That celebrated mathemati¬ cian there first gave a table of mortality, which he had constructed from observations made at Breslaw, and showed how the probabilities of life and death, and the values of annuities and assurances on lives, might be de¬ termined by such tables; which, he informs us, had till then been only done by an imaginary valuation. Besides his algebraical reasonings, he illustrated the subject by the properties of parallelograms and parallelopipedons: there are, perhaps, no other mathematical inquiries, in the prosecution of which algebra is entitled to so decided a preference to the elementary geometry as in these, and this example of the application of geometry has not been followed by any of the succeeding writers. In the year 1724 M. de Moivre published the first edi¬ tion of his tract entitled Annuities on Lives. In order to shorten the calculation of the values of such annuities, he assumed the annual decrements of life to be equal; that is, that out of a given number of persons living at any age, an equal number die every year until they are all extinct; and upon that hypothesis he gave a general theorem, by which the values of annuities on single lives might be easily determined. This approximation, when the utmost limit of life was supposed to be 86 years, agreed very well with the true values between 30 and 70 years of age, as deduced from Dr Halley’s table; and the method was of great use at the time, as no tables of the true values of i-PirW 'lionirfV ernis ^re .ma^e use °fIn article which may properly be considered technical. But since it is desirable that the the hi«tnnVnl etu ‘ C an^.wefl*defined ideas of the terms that are employed, in the demonstrative part, which follow cal order with thp nn^;6 th m thi? ParaffraPbs where they are first introduced; and we here give those terms in al"^4!. cal order, with the numbers of the paragraphs in which their definitions are given Term. Annuity Annuity, Certain **. Annuity, Deferred Annuity, Life Annuity, Temporary Life Annuity on any Life or Lives. Assurance on any Life or Lives Mortality, Table of. Years Purchase, N o. of, that an Annuity is worth. Paragraph. 3 4 20 31 58 62 77 32 6 ANNUITIES. 199 a nuities had then been calculated, except a very con- Ot cted one inserted by Dr Halley in the paper mentioned a ive. But, upon the whole, this hypothesis of De Moivre fejhs probably contributed to retard the progress of the sc. sence, by turning the attention of mathematicians from f t ■ investigation of the true law of mortality, and the best jif mthods of constructing tables of the real values of an- ca fl i ■. Pi llfl n tki tici a: t;: O'! tjl ' • til O'! r till h le tie i: |!;r lil, tk p k : 1R rtities. The same distinguished analyst also endeavoured to ap- piximate the values of joint lives; but it has since been f end that the formulae he gave for that purpose are too iuorrect for use. Mr Thomas Simpson published his l.idrine of Annuities and Reversions in the year 1742, in wich the subject is treated in a manner much more ge¬ nial and perspicuous than it had been previously. His fctnulae are adapted to any table of mortality; and, in the scenth corollary to his first problem, he gave the theorem d nonstrated in the 149th number of this article, to which w owe all the best tables of the values of life-annuities tli t have since been published. n the same work he also gave a table of mortality de- deed from the London observations, and four others cal- cjited from it, of the values of annuities on lives, each at tliee rates of interest; the first for single lives, the three o ers for two and three equal joint lives, and for the kgest of two or of three lives. These were the first tables of the values of joint lives tit had been calculated; for although Dr Halley had sl wn, half a century before, how such tables might be onputed, and had taken considerable pains to facili¬ ty! the work, the necessary calculations by the methods k wn previous to the publication of Mr Simpson’s treatise we so very laborious that no one had had the courage ^undertake them. And unfortunately the mortality ac- c ding to the London table was so much above the com- n:i average, that the values of annuities in Mr Simpson’s f iles were much too small for general use. n the year 1746 M. Deparcieux published his Essai sur letProbabilites de la Duree de la Vie Humaine, in which h igave several valuable tables of mortality deduced from tl mortuary registers of different religious houses, and fin the lists of the nominees in the French tontines; a t a table of the values of annuities on single lives, at the rates of interest, calculated from his table of morta- li for the tontine annuitants. These tables were a great ai uisition to the science, as, before their publication, tl re were only two extant that gave tolerably exact re- )) mentations of the true law of mortality—Dr Halley’s fuBreslaw, and one constructed but a short time before h iM. Kersseboom, principally from registers of Dutch anuitants. Those of M. Deparcieux for the monks and n s were the first ever constructed for the two sexes se irately; and by them the greater longevity of females w made evident. he work commences with an algebraical theory of an- l!‘ 11 -ies-certain; but the principal essay, On the Probabili- ( tb of the Duration of Human Life, is perfectly intelligible 1 tf hose who have not studied mathematics. It is written *' i great judgment and perspicuity, but contains very 1 i e more than the explanation of the construction of his 1 rfff’. s°me °f which relate to tontines; and he did not in il himself to the extent he might have done, of the ex- rl c< tract of Thomas Simpson. his work, however, appears to have been more read J| ^t11. e Continent, and to have contributed more to the i ision of this kind of information there, than all the other F P ^le su^jecj* The article Viagercs in the ne i Encyclopedic is acknowledged to have been taken m rely from it, as was also the article Vie, duree de la ; ‘ ai* t icse are proofs, among many others that might be produced, how little M. d’Alembert and the principal ma- History, thematicians his contemporaries attended to the subject, In the year 1752 Mr Simpson published, in his Select Exercises, a supplement to his doctrine of Annuities; wherein he gave new tables of the values of annuities on two joint lives, and on the survivor of two lives, much more copious than those he had inserted in the principal work; but these also were calculated from his London table of mortality. The celebrated Euler, in a paper inserted in the Me¬ moirs of the Royal Academy of Sciences at Berlin for the year 1760, gave a formula by which the value of an annuity on a single life of any age may be derived from that of an annuity on a life one year older; which formula was included in that given by Mr Simpson 18 years be¬ fore for effecting the same purpose in the case of any number of joint lives; and by this compendious method M. Euler calculated a table of the values of single lives from M. Kersseboom’s table of mortality. The first edition of Dr Price’s Observations on Rever¬ sionary Payments was published in 1770, and its chief object was, to give information to persons desirous of form¬ ing themselves into societies for the purpose of making provision for themselves in old age, or for their widows. When tables of the values of single lives, and of two joint lives, are given, the methods of determining the terms on which such provisions can be made with safety to all the parties concerned are very simple, and were at that time well understood in theory by the mathematicians who had studied the subject; but, for want of the requisite tables, the algebraical formulae had till then been of little prac¬ tical utility. In the prosecution of this laudable design, Dr Price was obliged to have recourse to approximations. He in¬ forms us, that by following M. de Moivre too implicitly in his rules for determining the value of two joint lives, he was led into difficulties which convinced him that they were not only useless but dangerous: he therefore calcu¬ lated a table of these values upon M. de Moivre’s hypo¬ thesis of the decrements of life being equal, and its utmost limit 86 years, from a correct formula given by Mr Simp¬ son in his doctrine of Annuities (Cor. 5, Prob. 1). By this, and a table of the values of single lives, calculated by Mr Dodson on M. de Moivre’s hypothesis, he was enabled to give answers tolerably near the truth, to some of the most interesting questions of this kind, and to show that the plans of several of the societies then recently established, were quite inadequate; and instead of the benefits they promised, could only, in the end, produce disappointment and distress, unless they either dissolved or reformed themselves. The work also contained instructive dissertations on the probabilities and expectations of life, and on the mean duration of marriage and of widowhood; besides accounts of some of the principal societies which had then been formed for the benefit of old age and of widows, with ob¬ servations on the method of forming tables of mortality for towns, and two new tables of that kind constructed from registers kept at Norwich and Northampton. Mr Morgan’s Doctrine of Annuities and Assurances was published in 1779, containing tables of the values of single lives, of two equal joint lives, and of two lives differing in age by 60 years, calculated from the Northampton table of mor¬ tality. And in the same year M. de Saint-Cyran pub¬ lished his Calcul des Rentes Viageres sur Une et sur Plu- sieurs Tetes, wherein the valuation of annuities on lives is treated algebraically, but in a manner much inferior in all respects to that of Mr Simpson; and six tables are given of the values of annuities—on single lives, on the survivor of two lives, and on the last survivor of three, calculated 200 History. annuities. from M. Kersseboom’s table of mortality. Although the values in the cases of two and of three lives were only de¬ termined by approximation, these tables were, just then, a valuable acquisition to the science; but their use was entirely superseded only four years after, y p tion of others much more valuable. The fourth edition of Dr Price’s Observations on Rever- sionary Payments appeared m 1783. One of the best effects of the preceding editions on the progress of the science had been, to direct the public atten.tlonh‘° ^ innuiries bv showing their important uses in the attairs oflife; amHo procure the requisite for forming tables of mortality, that should illustrate the laws according to which human life wastes under different circumstances, by exciting the curiosity of intelligent men who had the ne¬ cessary leisure and means of information. The ingenious author had accordingly been furnished with the neces¬ sary abstracts of mortuary registers which had been kept with these views, by Dr Haygarth at Chester, Dr Aikin at Warrington, and the Rev. Mr Gorsuch at Holy-Cross, near Shrewsbury, since the publication of the first edition; also by Mr Wargentin, with the mean numbers both ot the living and the annual deaths in all Sweden and 1 in- land for 21 successive years; in all of which the sexes were distinguished; and from these data he constructe tables of mortality that threw great light on the subject. He also inserted in this edition an improved table of mor¬ tality for Northampton; and, what had been so long want¬ ed, a complete set of tables of the values of annuities on single lives at six rates of interest, and on two joint lives at four, all calculated from the new Northampton table. The combinations of joint lives were sufficiently numerous to admit of all the values not included being easily inter¬ polated. Besides these, he also gave tables of the values of annuities on single lives from the Swedish observations, both with and without distinction of the sexes, and on two joint lives without that distinction. The values given in these tables are too low for the general average of lives at all ages under 60; but in the treatise of Mr Baron Maseres on the Principles of the Doc¬ trine of Life Annuities, which was published in the same year (1783), others were given, calculated from the table of mortality which M. Deparcieux constructed from the lists of the nominees in the French tontines. The tables for single lives are calculated at twelve different rates of interest from 2 to 10 per cent, but those for joint lives only at 31 and 41 per cent.; and the combinations they include are only those of ages that are equal, or that differ by 5 or 10 years, and the multiples of 10. There is reason to believe that the values in these tables, at all ages under 75 or 80 years, are nearer the truth, for the average of this country, than any others then extant; but certainly for the average of lives on which annuities and reversions depend. After that period of life, however, they are too small; and, in most cases, it is difficult to derive the values of joint lives from them with sufficient accuracy, on account of the contracted scale they have been calculated upon. It was not Dr Price’s object to deliver the elements of the science systematically ; but he treated most parts of it with great judgment, enriched it with a vast collection of valuable facts and observations, and corrected several errors into which some of the most eminent writers upon it had fallen. The mathematical demonstrations (which are given in the notes) are much inferior to the rest of the work. The values of reversionary sums and annuities, which depend upon some of the lives involved failing according to assigned orders of precedency, had been approximated by Mr Simpson in his Select Exercises, and by Mr Morgan in his Doctrine of Annuities; but the latter gentleman first gave accurate solutions of problems of this kind, in theGi Philosophical Transactions for the years 1788,1789,1791 1794, and 1799. Mr Baily’s Doctrine of Life Annuities and Assurances was published in 1810. In it the whole subject is treated, except the construction of tables of mortality, on which the practical application of all the rest depends. In con- sequence of the author having adopted Mr Simpson’s no- tation, this work presented a more perspicuous exposition of the whole theory, especially of the improvements made in it between the time when Mr Simpson wrote and the date of its publication, than had previously appeared. And in an appendix to it, published in 1813, principally for the purpose of explaining the construction and uses of tables for determining the values of life-annuities, cal¬ culated at a vast ‘sacrifice of time and labour by Mr George Barrett, since deceased, formulae were given for calculating from tables of that kind the values of tempo¬ rary and deferred life-annuities and assurances, and also for determining the values of annuities and assurances when the annuity or the sum assured, instead of remain¬ ing always the same, increases or decreases from year to year by equal differences, with considerably greater fa¬ cility and expedition than the same things could have been effected with by the tables and methods of calculation in previous use. Except by these improvements, and the solution of the problems above stated to have been first given by Mr Morgan, which were severely criticised and given anew, with some amendments besides the important one of the notation in Mr Baily’s work, the science had not been materially advanced, during a period of more than 30 years, which had elapsed since the appearance of the fourth edition of Dr Price’s observations, when Mr Milne published his Treatise on the Valuation of Annuities ad Assurances on Lives and Survivorships, in the year 1815, The work consists of two volumes ; the first is mathe¬ matical, the second entirely popular, except the notes and a few of the tables. Ihe algebraical part of this article is merely a short abstract ot the first volume, and may serve as a specimen of the manner in which the sub¬ ject has been treated there ; but the construction of tables of mortality, which forms the subject ot the third chap¬ ter, has not been noticed here; neither is the valuation of reversionary sums or annuities depending upon assigned orders of survivorship treated in the present article; and these are parts of the work which will not be found the least interesting to mathematicians. The second volume contains upwards of 50 new tables, with a few others that had been published before, hut have been reprinted either on account of their value or scarcity, or both. Four of the new ones are tables oi mortality constructed by the author, from registers ep at Carlisle and Montpellier, and in all Sweden and tin- land, since the period of the observations Dr Price ma e use of: the sexes are distinguished in the tables foi den and Montpellier, but not in that for Carlisle. 115 last is the only table, besides those for Sweden and tin- land, applicable to the mass of the people, that ^as formed from the necessary data,—enumerations o living, as well as registers of the deaths, in every in e val of age. , t Twenty-one of these tables, being the seventeen ^ the thirty-seventh inclusive, in the collection at of the work, render it easy to apply the algebraical or lae to practical purposes, and numerous examples o s applications are given. They have all been ca‘cUlj ; from the Carlisle table of mortality; those of .^VU of life-annuities on the same extensive scale with & jit’l' ANNU L , jici! Dr Price derived from the Northampton table. It Hw i the author’s opinion that the values of interests de- |'W indent upon the continuance or the failure of life may ;1> I 1 derived from them more correctly than from any others tl| ten extant, and he has taken considerable pains to assist l» h readers in judging of this for themselves. Besides the tables, the principal contents of the second ',(1 \ lume are explanations of their construction and uses. Ill jany of them relate to the progress of population,—the pi se ^ years; that is, 20 times the mean numbers of — I T I E S. 201 them in the several intervals during the same term. But History, neither of Mr Morgan’s explanations of them would ad- mit of that construction.1 The present article was first published in the Supple¬ ment to this Encyclopaedia in 1816. In the article on the Law of Mortality in that Supple¬ ment, which appeared in 1822, it was shown that, accord¬ ing to Mr Morgan’s statement, made at a general court of the Equitable Society in the year 1800, of the mortality which had taken place among the lives insured by that society as compared with the Northampton table, the mortality among those lives in each decade of age from 10 years to 50 was very nearly the same as in the Car¬ lisle table of mortality; also that, above 50 years of age, the difference, upon the average, was not great. And early in 1826 were published Tables of Life Contingencies, by Mr Davies, and A Comparative View of the various In¬ stitutions for the Assurance of Lives, by Mr Babbage ; in each of which works was given a table of the mortality which had prevailed among the lives insured in the Equit¬ able Society at all ages above 10 years, constructed from that statement of Mr Morgan. Mr Babbage gave also a table of the values of annuities on single lives of all ages above ten years, derived from his table of mortality above mentioned; and the inde¬ fatigable Mr Davies gave tables of the values of an¬ nuities on single and joint lives, calculated both from his table of mortality above mentioned and from the Northampton table, rather fuller and more complete than any that had previously been published, except that those derived from the law of mortality in the Equitable So¬ ciety necessarily included no ages under 10 years. The values according to the Northampton table were given only at the rates of 3 and 4 per cent, interest; but Mr Davies, not content to take them on Dr Price’s authority, has, like Mr Barrett, calculated them anew, and, as well as the other values of annuities, has carried them to four places of decimals. This author’s comprehensive little book also contains many other tables of the values of an¬ nuities and assurances on lives and survivorships, with one of the values of policies of assurance, calculated from the Northampton table of mortality at 3 per cent, inte¬ rest, which, as well as many another single table contained in it, must have cost him much time and labour; and those derived from the Northampton table of mortality must be very valuable to such of the assurance companies as take that table for their guide in transacting business. Mr Babbage and Mr Davies also gave formula; and tables similar to, or not materially different from, those of Mr Barrett above mentioned, for determining the va¬ lues of temporary and deferred, as well as increasing or decreasing annuities and assurances on single lives; the tables of Mr Babbage being derived from his table of mortality in the Equitable Society, and from the Carlisle table; that of Mr Davies from the Northampton table of mortality alone. Mr Morgan, in the statement on which those tables of Mr Babbage and Mr Davies were founded, contented him¬ self with the use of two simple digits only, to express the proportion of the mortality in the Equitable Society to that in the Northampton table in each decade of age; and although that gentleman for twenty-five years after 1800, when that statement of the thirty years’ previous observations was made, continued, in Notes on Dr Price’s Observations on Reversionary Payments, and in his ad¬ dresses to the general courts of the Equitable Society, to state that the proportions still remained the same, the vol. in. 1 See the note at the end of this Historical Introduction. 2 c annuities. observations extending over a period of fifty-five years, he never gave any statement more full or distinct. These were but scanty materials certainly to construct table of mortality from; and yet the agreement between the tables formed from them, and the best of other ex ing tables of mortality, is very remarkable. _ A committee of the House of Commons on friend y socie¬ ties having been appointed in 1827, chiefly for the pur¬ pose of inquiring into the law of mortality, and the values of life-annuities and assurances in this country, the re¬ port of that committee, by bringing the subject prominent¬ ly before the public, and exciting attention to well-esta¬ blished but much neglected results of inquiries into it, had the effect of correcting to a considerable extent opi¬ nions upon it, taken upon trust without due examination, and generally diffused. The establishment of many new assurance companies, and the increasing prevalence of life-assurance for 15 or 20 years before, by exciting c is- cussion and examination of their rates, had also contribut¬ ed to produce that effect. At length the members of the Equitable Assurance Society, as the period of the decennial investigation of their affairs in 1829 approached, express¬ ed a desire to avail themselves of the information respect- ing the law of mortality in the society which the office books might afford, for estimating and dividing their pro¬ fits ; and in 1828 Mr Morgan published a pamphlet en¬ titled A View of the Rise and Progress of the Equitable Society, in which (p. 42) he gave the following “ table of the decrements of life in the society during the preced¬ ing 12 years.” given at the end of Mr Morgan’s work on annuities (2d: edit. 1821)—having also some doubts about the statement made in 1800—wrote to Mr Morgan, and requested the desired information respecting them, when that gentleman forwarded him the following: 1. That the second column should have, been headed, “ Number of persons living at the beginning of each year during the last 12 years and the fourth, “ the num¬ ber which should have died, according to the North¬ ampton table.” 2. That as to the table at the end of his treatise on an- nuities, he knew the numbers of deaths by the different diseases to be correct; but “ the numbers in the last line of that table were those of the policies, which he had since found greatly to exceed the numbers of the members.” 3. Upon his statement of 1800 Mr Morgan made no ob¬ servation. From the first part of the information thus obtained, it appears that the number in the second column on any line in table (a), is twelve times the mean number of lives in the interval of age set against it, on which the Equitable Society had policies in force during these 12 years; so that, in a society similar in all other respects, in which the law of mortality w'as constantly the same, and the number of lives in each interval of age was al¬ ways just 12 times as great as in the Equitable during these 12 years, the population and mortality would be as in columns 2 and 3 of the following table (b). V' -- TABLE (1)). TABLE (a). Age. 20 to 30 30 to 40 40 to 50 50 to 60 60 to 70 70 to 80 80 to 95 Number. 4,720 15,951 27,072 23,307 14,705 5,056 701 Died. 29 106 201 339 426 289 99 Should have died. 68 243 506 545 502 290 94 In the same place, the author informs his readers that “ in his former statement (without mentioning which of them) he was not aware of the great number of instances in which there were several policies on one and the same life.” Also that “ the present is, in fact, the only correct table of the decrements of life in the society.” In the same pamphlet the author has given a table to show the law of mortality among the lives insured in the Equitable Society, founded upon his last statement; also a table of the expectations of life, and one of the values of annuities on single lives at three per cent, per annum interest, both derived from that table of mortality, none of them including any age under 20 years. The author of the present article, when preparing this historical sketch, not clearly understanding the import of the numbers in the second column of this table (a), nor the last line in the table of deaths by the different diseases, Between the Ages of 20& 30 30 & 40 40 & 50 50 & 60 60 & 70 70 & 80 80& 95 Number constantly living. Totals 4,720 15,951 27,072 23,307 14,705 5,056 701 Of whom die annually. 29 106 201 339 426 289 99 91,512 1489 6 Of whom would die annually accord¬ ing to Our calculations from Mr Morgan's calculations from Mr Morgan’s Table of M ortality in Equitable Society. 37 150 337 536 664 460 146 2330 The Northampton Table of Mortality. 74 299 648 780 726 461 147 3135 243 506 545 502 290 94 2248 If our inferences in the 4th and 5th columns of this table from Mr Morgan’s data in the 2d and 3d be correct, neither Mr Morgan’s in the 6th column, nor his table o mortality, nor consequently those of the expectations an values of lives above mentioned, can have been accurate) derived from the same data. But even with these data we have reason to believe that Mr Morgan is not well satisfied, and that that gent e- man has since been able to form a table from much oof® unexceptionable documents, which shows the probabilities 1 It may be proper to show here, how our numbers in columns 4 and 5 have been determined. For this purpose we give asane* ample the manner in which the 2d in col. 5 has been calculated. = { ^o’oQfil number living above Expectation of life at | ^ ^ j number annually attaining that age \ 83,896 J The differences are the numbers between 30 and 40 ; 750 dying annually, and 40,068 constantly living. And 40,068 : 750 :: 15,951 : 299. See article on Law of Moiitality. r *pa t ei I ) ANNUITIES. \ oltife, especially in the latter periods, to be much nearer ijti probabilities of life among mankind in general than he h; previously imagined. t is satisfactory to find Mr Morgan at last coming into j nippinion, of the truth of which, evidence had previously ee b( n adduced sufficient to convince almost all who at- 111 teded to it; and, trusting that he will publish the results obis latest inquiries on the subject, we shall take no fur- tlr notice here of the table of the values of annuities intiis pamphlet above mentioned, which, otherwise, we cdd not with propriety have avoided. ^ Ir Morgan (Pamph. p. 42 and 43) asserts that a com- piiison of columns 3 and 6 of this table (b) affords a st King proof of the accuracy of the Northampton table ; bu to judge of that, we consider it is not column 6, but 5, /hich should be compared with column 3. I i the Transactions of the Cambridge Philosophical So- ci before; in which registers the exact ages at which annuitants were nominated, and those at which they l were stated. 203 • ms the data not otherwise accessible being provided, ,1 !e jOur lessened by the number of calculators em- p o '(1 the expense also being defrayed by the public— P e end of ten years, viz. in March 1829, Mr Finlaison a . a icport to the Lords of the Treasury, which was all1; e^oc ?.r(^er t^le House of Commons, and, in tables v I ’ „ 0 10 P?8es> shows the rates of mortality, and the din S ^nnultles 011 s*ngle lives at all ages, among many the ^niclas?es °f annuitants, both separate and combined; . . cllIA1UiLclul'®5 Dorn separate and cornbinecl 5 t exes being generally distinguished both in exhibiting me uv of mortaliMr r- ° pi ' c <=> O UUtil 111 e r v 0 mortality and the values of annuities, mo va’ Vi'11 tJle number and accuracy of the data, are pro mi uV 6 1 lan, ,an^ th*nS of the same kind that had ties mb-' i)-C)n Pu.)Isled; but it is the values of annui- «es toy which we have to notice here. The lives on which annuities depend will generally be History, somewhat better (by which we here mean, will attain to' ^ — greater longevity) than the general average of the popu¬ lation, though probably not nearly so much better as many believe them to be. The prevailing error in the popular estimate on this subject appears to have arisen in great measure from comparing the mortality among annuitants and assured lives, with that represented to take place by tables of mortality erroneously considered to correspond with the general average of the people; while, from be¬ ing constructed on erroneous principles, and from insuffi¬ cient data, or else being derived from observations made where the mortality was and is much greater than in Bri¬ tain, the mortality according to these tables was consider- ably greater than that which actually prevails among the bulk of the people here. Proofs of this will be found un¬ der the article Law of Mortality. That the lives on which annuities and assurances de¬ pend cannot be so very select or so much better than the common average as has generally been supposed, might reasonably be expected on these grounds :— 1. As to annuitants. The lives are not all chosen on account of their pre¬ sumed goodness ; for many persons who have no occasion to provide for others who may survive them, purchase an¬ nuities on their own lives, only that they may themselves enjoy the whole benefit of the purchase-money, both principal and interest, during their lives. And the greatest recommendation of these lives seems to be, that they are generally prudent persons, of tempe¬ rate and regular habits. Many other persons, especially females, spendthrifts, and faithful servants, enjoy annuities bequeathed to them by their deceased relatives, masters, or mistresses, as the most eligible provision for their future comfort and secu¬ rity from want; and there seems little ground to sup¬ pose them to be better lives than the common average of the same age and sex. 2. In such cases as tontines, where most of the lives are selected for their presumed goodness, the best crite¬ rion probably is, hereditary longevity in the family of the nominee ; but partiality for their own friends or kindred often has considerable influence in biassing the judgment of those who select them. That they will generally be persons of good constitu¬ tions and regular habits when selected, is all that is like¬ ly to be obtained under these circumstances; and that is also the case with the average of the population in com¬ fortable circumstances. Whatever the constitutions and habits of annuitants may be, the annuities held by them during their own lives, by protecting them from many of the wants, cares, and anxie¬ ties which the bulk of the people are exposed to, no doubt contribute to their longevity. But where powerful motives to raise money by the sale of an annuity on a per¬ son’s own life exist, it is extremely difficult to prevent him from parting with it, whatever precautions may have been taken with that view; and with it, he also loses that help to longevity. 3. Insured lives are also generally supposed to be much better than the average of the population, as it is incum¬ bent upon the insurance offices to be cautious in select¬ ing them. But bad lives, by the failure of which persons interested in them would sustain loss, are most likely to be offered, and are continually offered, for insurance; and there is reason to believe that all the caution in selection which the offices in general can exercise, is necessary to keep the lives insured up to the average goodness of the bulk of the population;—supposing always that people in gene- 204 ANNUITIES. History, ral of the industrious classes are in prosperous, or at least in comfortable circumstances. When that is not the case, as for some years previous to 1830 there is reason to ap¬ prehend it was not in this country, there will be a corre¬ sponding increase in the general mortality, which will not sensibly affect the general mass of persons on whose lives annuities and reversions or assurances depend. In many cases of insurance, the managers who choose the lives know very little more about them than is stated by themselves or the medical attendants they refer to, and frequently they depend much on the reports of their agents in the country; but where they are in daily ha¬ bits of intercourse with most of the persons whose lives they insure, or else in habits of intimacy with some who His are so, they have the means of making a much better se-^i lection. That is stated to be the case with one of the London offices, and in which, accordingly, there is said to have been observed a remarkably small mortality. For tbe reasons assigned above, it appears, that al¬ though lives carefully selected, with the advantage of the necessary facilities, and unmixed with others, may be con- siderably better than the general average, those on which annuities and assurances are granted are not, on the ave¬ rage, likely to be much better. This part of the subject is also adverted to at the end of the article on the Law of Mortality. The two following Tables, A and B, show the number of Years’ Purchase Annuities on Lives of different ages are worth in present money, according to More Correct Tables of Mortality. TABLE A. Without distinction of Sex. Observations began.. Ended Years’ duration RATE OF INTEREST FOUR PER CENT. PER ANNUM. Column Description of Lives. Greatest number of j lives J Mean number Number of deaths... Table of Mortality! published in J Constructed by a Nominees in the French Tontines. Population of Carlisle and 1690 1742 50 9260 + 5293=1=: 7933 + 1746. Deparcieux Age. 10 15 20 25 30 35 40 45 50 55 60 65 70 75 19-008 18-502 17-938 17-420 16-810 16-084 15-133 13-904 12-526 11-173 9-713 8-039 6-394 4-945 1779 1787 9 Equitable Assurance Society. / Nominees in English Tontines. Selected by Contributors. 10,517 + 8177 1840 1815. Milne. 19-585 18-956 18-363 17-645 16-852 16-041 15-074 14-104 12-869 11-300 9-663 8-307 6-709 5-239 1770 1800 30 6344 + 2522 =fc 1220=+= 1827. Davies. 19-647 18-944 18-242 17-494 16-701 15-867 14-939 13-845 12-599 11-349 10-052 8-635 7-167 5-670 1789 1826 37 3518 2860=+= 1315 Chosen by Lot. 1789 1826 37 4831 3920: 1823 The two last together. 1789 1826 37 8349 6780: 3138 Various combined. 1773 1826 53 18,798 15,459=+= 6,679 1829. Finlaison. 19-167 18-475 18-011 17-526 16-889 16-098 15-195 14-061 12-671 19-068 18-422 17-946 17-530 16-925 16-099 15-124 13-985 12-528 The numbers for the two classes sepa¬ rately were deemed insufficient at these ages. 19-118 18-448 17-979 17-528 16-907 16-099 15-160 14-023 12-599 11-163 9-772 8-308 6-729 5-122 19-242 18-532 17-954 17-534 16-995 16-314 15-516 14-533 13-295 11-915 10-491 8-896 7-316 5-837 h Population of Sweden and Finland. 1755 1776 21 4,051,116 2,310,160 1,401,989 1783. Price. 18-891 18-336 17-603 16-839 16-006 15-138 14-034 12-959 11-658 10-320 8-789 7-328 5-783 4-534 First English Tontine. 1693 1783 90 1002 601: 1002 1829. Finlaison, 17-128 16-207 15-349 14-976 14-624 14-023 13-193 12-199 11-183 10-141 8-836 7-342 5-823 4-456 Histy. ■ " ANNUITIES. TABLE B. The Sexes distinguished. RATE OF INTEREST FOUR PER CENT. Age. 10 15 20 25 30 35 40 45 50 55 60 65 70 75 Sweden and Finland, 1755-1766. Dr Price. d f British Tontines and Annuities, 1773-1826. Mr Finlaison. Male. 18-674 18-105 17-335 16-592 15-751 14-812 13-668 12-535 11-267 9-998 8-540 7-090 5-670 4-487 Female. 19-109 18-568 17-872 17-087 16-261 15-465 14-401 13-383 12-049 10-642 9-039 7-566 5-897 4-582 Excess in Value of a Female above a Male Life. 0-435 0-463 0-537 0-495 0-510 0-653 0-733 0-848 0-782 0-644 0-499 0-476 0-227 0-095 0-919 1-055 1-318 1-187 1-102 1-131 1-281 1-471 1-731 1-751 1-540 1-361 1-083 0-854 Male. 18-782 18-004 17-295 16-940 16-444 15-749 14-875 13-798 12-430 11-039 9-721 8-216 6-775 5-410 Female. 19-701 19-059 18-613 18-127 17-546 16-880 16-156 15-269 14-161 12-790 11-261 9-577 7-858 6-264 Age. 10 15 20 25 30 35 40 45 50 55 60 65 70 75 The three following Tables, M, N, and O, show the Values of Annuities on Lives, according to Less Correct Tables of Mortality.—All without Distinction of Sex. M. N. O. RATE OF INTEREST FOUR PER CENT. ge. a London Bills, 1742. Simpson. 16-4 15-8 14-8 14-0 13-1 12-3 11-5 10-8 10-1 9-3 8-4 7-5 6-5 5-4 Demoivre’s Hypothesis, 1753. Dodson. 16-882 16-410 15-891 15-318 14-684 13-979 13-196 12-322 11-344 10-248 9-017 7-631 6-065 4-293 Northampton Table, 1783. Price. 17-523 16-791 16-033 15-438 14-781 14-039 13-197 12-283 11-264 10-201 9-039 7-761 6-361 4-962 d Mort. Itegrs. in France, 1806. Duvillard. 17-882 17-154 16-442 15-747 15-021 14-214 13-286 J2-218 11-026 9-709 8-342 6-965 5-635 4-420 Age. 10 15 20 25 30 35 40 45 50 55 60 65 70 75 FIVE PER CENT. 12 country and 3 Parisian Parishes, 1779- Duprd and Age. St Cyran. 10 15 20 25 30 35 40 45 50 55 60 65 70 75 15-40 14-82 14-15 13-59 12-96 12-32 11-72 11-05 10-15 8-93 7-73 6-73 5-74 4-59 Northampton Table, 1783. Dr Price. 15.139 14-588 14-007 13-567 13-072 12-502 11-837 11-105 10-269 9-382 8-392 7-276 6-023 4-744 SIX PER CENT. a Breslaw Bills, 1693. Halley. 13-44 13-33 12-78 12-27 11-72 11-12 10-57 9-91 9-21 8-51 7-60 6-54 5-32 Demoivre’s Hypothesis, 1753. Dodson. 12-839 12-586 12-301 11-978 11-610 11-189 10-704 10-144 9-492 8-729 7-831 6-770 5-508 4-000 Northampton Table, 1783. Price. 13-285 12-857 12-398 12-063 11-682 11-236 10-705 10-110 9-417 8-670 7-820 6-841 5-716 4-542 Age. 10 15 20 25 30 35 40 45 50 55 60 65 70 75 ANNUITIES. 206 History. Explanation of Table A. The sign (+) after the number of lives and of deaths in column a signifies that the real number was greater than is there stated. Those stated were the numbers of the nominees in the two tontines which commenced in 1689 and 1696, the particulars of which have been given by M. Deparcieux; but he also made all the use he could of the tontine which commenced in 1734 (less than eight years before his observations terminated), without stating any of the numbers in his essay. And whatever may have been the whole number of nominees, or of their deaths, which he availed himself of in this tontine, M. Deparcieux’s observations were actually made on so many more than 9260 nominees, and 7933 deaths, among them. The number of persons living in the two Carlisle pa¬ rishes at the end of the observations was 8677; but be¬ sides them, the observations were made upon the 1840 persons who died in the place in the term of nine years during which they were continued; and these numbers together amount to 10,517, the greatest number stated in column b. But the real number the observations were made upon was greater still, by the number who left the place and did not return during the observations, which is the reason of the mark ( + ) being put after 10,517 in column b. For information respecting the number of lives insured, and of deaths, in the Equitable Assurance Society, given at the head of column c, the reader is referred to the note at the end of this historical introduction. The better to enable the reader to judge of the compa¬ rative extent of the observations made upon the nominees in tontines, and other annuitants, by M. Deparcieux and Mr hinlaison, and of those made upon the population of the two Carlisle parishes, the lives insured in the Equi¬ table Society, and the population of Sweden and Finland, the mean number of living annuitants has been assumed to have been an arithmetical mean proportional between the numbers of them at the commencement and at the end of the term, which can only be precisely true if they died olf by equal numbers in equal times; and that is the reason why the double sign (=ir) has been placed after the mean number of the nominees or other annuitants in each column. Thus 2860:±= in column d shows that the mean number of living nominees of that description was 2860 more or less. The deviation from precision in this case is of no importance. The values in column a have been taken from the work on Annuities of Mr Baron Maseres; those in columns d and e from Mr Finlaison’s report (obs. 4 and 5); that in column f at each age to 50 inclusive is a mean between those in columns d and e /—after 50 they are taken from Mi Finlaison s 5th observation. The value in column g at each age is a mean between the two against the same age in columns e andy*of table B; the values in column i are from Mr Finlaison’s first observation. Of Table B. The values in columns e and f have been taken from the 20th and 13th observations respectively in Mr Fin- laison s report, and were calculated from the rates of moi tality for the two sexes, which have been adopted for use by government. 1 hey were deduced from observations on the mortality among the nominees in the three Irish tontines v/hich commenced in 1773, 1775, and 1778 respectively, on the tontine of 1789, and those of the sinking fund from 1808 to 1822. It will be observed that the excess of the value of an annuity on a female life above that of a male is, according to the table for Sweden, in many cases not half, and in H some less than one third as much as according to MrkJ Finlaison’s, derived from the government annuitants. The 1 cause of this cannot but be an object of interest, and de¬ serves further investigation. It may arise in great mea¬ sure from the ages of many females being stated below the truth in the Swedish returns, while they were accu¬ rately ascertained among the government annuitants. Of Tables M, N, and O. The values according to Demoivre’s hypothesis were taken from Dodson’s Mathematical Repository (vol. ii. p. 169) ; those in column d of table M, founded upon Duvil- lard’s table of mortality for France before the Revolution, published in his work on the Mortality from Small-Pox (4to, Paris, 1806), were taken from The Doctrine of Com¬ pound Interest by M. Corbaux (8vo, London, 1825); the values according to Dupre and St Cyran from the Cal- culs des Rentes Viageres of the latter. The authorities for the rest appear sufficiently from the preceding historical sketch. The values of annuities according to M. Kersseboom’s table of mortality are not given here, that table being of doubtful character, as he neither published the whole of the data from which he formed it, nor explained the man¬ ner of its construction. It would have been desirable to include the values ac¬ cording to the tables of Dr Halley for Breslaw, and Dupre de St Maur for the Parisian and French country parishes in table M; but as the values of annuities have not been calculated from these tables at four per cent, we have added tables N and O, and have given in each of them the values from the Northampton table, with the view of fa¬ cilitating the comparison of the values in N and O with those in M and A. Observations on the above Tables. All the tables of mortality from which the values of an¬ nuities in tables M, N, and O, have been deduced, were calculated from bills of mortality alone of places where the population was variable, and the numbers of the people at the different ages were not ascertained. And therefore, notwithstanding the attempts to supply their defects, which were made by the eminent mathematicians who constructed them, none of them represented truly the laws of mortality in the places where the respective obser¬ vations were made ; as will be evident to those who un¬ derstand the article on the Law of Mortality in this work, and pay the necessary attention to the materials and man¬ ner of construction of those tables. Consequently the va¬ lues of annuities derived from them cannot be correct, but will in general be considerably less than the truth, even for the general average of the whole population of the places in which the observations were made. But those values of annuities are also objectionable on this ground—that the places they were intended for, and understood to be adapted to, were generally populous towns, containing a large proportion of poor persons de¬ pendent upon their daily labour for their supply of food from day to day, often with little forethought, and many of them engaged in unwholesome employments, amongst whom great distress is often endured by the comparatively high prices of bread and potatoes, or the low' rate of wages, when the unwholesome and scanty food they are reduced to produces typhus fever, and sometimes the dysentery among them, which carry them off in great numbers. And these visitations were much more common at the times when the observations were made from which most of those tables were constructed, than they have been of late years. ANNUITIES. ^tiy None of those causes of mortality operate sensibly upon >v-^the general average of those persons upon whose lives leases or annuities, and reversions or assurances, depend, they being generally in the higher and middle classes. Neither do they produce much effect among the more de¬ serving persons in the lower class, such as the members of friendly societies, and others who are both industrious and frugal enough to live within their incomes; nor in¬ deed upon any who are in comfortable circumstances. Hence it follows that the values of life-annuities, and, consequently, those of any pecuniary interests dependent upon the continuance or the failure of human life, cannot be correctly determined from observations made on a whole population similar to those of the places these tables were constructed from. But this was not distinctly seen till of late years, and appears to be very imperfectly understood at present (in the year 1830), even by some who might be expected to possess correct information on the subject. The tables constructed by Dr Price, both from the Swedish observations, and those made by Dr Haygorth at Chester, threw valuable light on this subject. But defi¬ cient crops in Sweden operate powerfully in raising the mortality there, in comparison with the more fruitful parts of Europe; therefore, the values of annuities copied into table B, and col. h of table A, from Dr Price’s work, must only be understood as sufficiently correct for the period and place in which the observations were made. And the Chester table is in some degree liable to the same objec¬ tions as the others above mentioned. On Table A. For nearly 70 years after its publication, M. Depar- cieux’s table, from which the values given in col. a of table A were derived, was the only one from fthich the values of life-interests and of reversions depending upon lives could be determined with considerable accuracy. But the comparatively high values of annuities according to that table were always supposed to arise from the care¬ ful selection of the lives ; notwithstanding that they were almost all inhabitants of Paris and its environs.1 At that time (1689-1696) the Parisians were much worse lives than during the last 50 years, and a judicious selection was much less likely to be made then than now. It is to be regretted that the Carlisle observations were only continued nine years, commencing with 1779; but the less so since Mr Milne has shown in his work on an¬ nuities (p. 429), that during the term of 22 years com¬ mencing with 1779, the proportion of the annual average number of deaths to the mean number of the people was the same as in these first nine years, viz. that of 1 to 40. In comparing the values in column c of table A with the rest, it should be borne in mind that a great majority of insured lives are males, on which account the values are somewhat lower, especially from 15 to 55 years of age, than they would have been had there been nearly equal • numbers of both sexes. Columns d and e are very instructive. In the session of 1789, an act (29 Geo. III. cap. 41) was passed for raising the sum of L.1,002,500 by the sale of shares in a tontine; but the scheme did not succeed, the persons who in the first instance had taken the whole of the shares with the expectation of selling them at a profit not having been able to dispose of half of them ; and to afford those persons re¬ lief, an act (30 Geo. III. cap. 45) was passed in the next session, allowing them, before the 20th September of that year (1790), to exchange each of the tontine shares they 207 had not been able to dispose of, for an annuity-certain History, payable during 69^ years. And in order that those whov— had taken shares, and fixed upon their nominees in the tontine, might be placed in the same situation with regard to the benefit of survivorship as if the scheme had com¬ pletely succeeded, this act empowered the commissioners of the treasury to select tontine nominees for the ex¬ changed shares, from the Peers of Great Britain and Ireland, their children or grandchildren ; baronets, lords of manors, justices of the peace in England and Wales, or their chil¬ dren ; the dignitaries of the church or beneficed clergy, fel- loios of colleges, the governors of the Charter House, the Foundling Hospital, or Christ's Hospital, and those who were registered in the books of the Amicable Life Assurance Society. The commissioners of the treasury were to distribute their nominees into six classes according to their ages, in proportion to the nominees of the contributors in the same classes. Tickets with the names of the nominees were then to be put into six boxes, set apart for the respective classes, and drawn out till a sufficient number to complete the tontine was obtained for each class. All of which was performed accordingly. The values of annuities on the lives chosen by the con¬ tributors are given in col. d, on those thus drawn by lot in col. e, and the two combined in col.^i Thus it appears that, of the lives in col. e, there was no selection except their being taken from the descriptions of persons above mentioned, who were all in the upper or middle classes of society. But, as might have been anti¬ cipated, a considerable majority of female lives were chosen by the contributors, and a considerable majority of males were drawn by the commissioners of the treasury, the descriptions of persons they were restricted to, consisting principally of males. It will be observed that the values in columns d and e agree very nearly:—they would probably have agreed better still had the proportion of the two sexes been the same in both. And this shows how little advantage the contributors derived from choosing their nominees, beyond what was secured to them by the classes of society they were se¬ lected from. The values of annuities in column i are much less than in any other of table A; they are even less than those de¬ rived from the Northampton table of mortality. But it would be equally precipitate and unphilosophical to conclude from thence, without further investigation, that the bulk of the people of England 100 years ago were so much shorter lived than they are now. That the prolongation of life among the bulk of the population, from and after every age, has been very con¬ siderable during the last century, no unprejudiced person who has paid sufficient attention to the subject to qualify himself for judging of it can entertain a doubt; or that it has also been somewhat lengthened among the upper and middle classes of society; but not nearly to the extent which a comparison of columns g and i of table A would seem to imply. There is certainly no rational ground for supposing that the physical constitution of man has altered: any change that has taken place can only have been produced by changes in the habits of the people, and the circumstances in which they have been placed. All this might have been reasonably concluded in the absence of further information; but an examination of the circumstances under which the tontine of 1693 was 1 As M. Deparcieux states in his Essai, p. 62. 208 ANNUITIES. History- formed, and of the first observation in Mr Finlaison’s re- port, from which the values in col. i have been deduced, will confirm it. In that year, the same in which Dr Halley’s paper on the Breslaw Bills was published in the Philosophical Transactions, this branch of knowledge was in its infancy. By the 22d section of the act (4 William and Mary, cap. 3) for raising a million of money by the sale of shares in this tontine, it was enacted that, if the sale of the tontine shares did not produce the whole sum wanted by the first of May 1693, then between that day and the 29th Sep¬ tember following, for any sum contributed towards the completion of the million, the contributor should receive 14 per cent, per annum on such sum during the life of any person he might choose to nominate; the common interest of money at that time being § per cent.per annum. And by an act passed in the next session (5 William and Mary, cap. 5), the term for granting annuities on these terms, and for the same purpose, was extended to the 1st of May 1694. This was selling annuities at half their true value. The age of a nominee is never mentioned in either of these acts, and those in the tontine were not distinguish¬ ed into classes. These things were, at the same time, managed better in France. Even Dr Halley was not aware of the greater mortality of males, and the consequent greater proportion of females in the population, as appears by his paper above mention¬ ed ; for after calculating from his table of mortality the number of inhabitants in Breslaw between 18 and 56 years of age to be 18,053, he says, “ at least one half of these are males.” The contributors to the tontine could not be expected to be better informed on these subjects than the parlia¬ ment and Dr Halley; and, with respect to ages and sexes, the following appears by Mr Finlaison’s statement to have been their selection of nominees. ations, and upon their moral conduct: all of which must His be very uncertain, and difficult to judge of at such early^ ages. It is not improbable, too, that, on account of their beauty and healthy appearance, many children of scrofu- lous constitutions were selected; and they, on an average, would be short-lived. It is probable that a great majority of the contributors and therefore of their nominees, resided in London or other large and crowded towns, which have always been peculiarly unfavourable to the health of children, but were much more so then than they are now. All circumstances considered, there appears sufficient evidence to show that the mortality among these nomi¬ nees must have been much greater than among the general average of selected lives, or the general average of the people in comfortable circumstances at that time ; and that, if the nominees in the English tontine of 1693 had been distributed into classes according to their ages, and a larger annuity for the same purchase-money had been allowed to the older classes, always with benefit of survivorship, the lives would have been more judiciously chosen, and would not probably have differed materially from the no¬ minees in the French tontines of 1689 and 1696, which M. Deparcieux’s observations were made upon,—they, it has been shown, were but little inferior In goodness to our present annuitants and insured lives. The values of annuities on single lives given in Mr Fin¬ laison’s report are only specimens at one rate of interest of the results of calculations made at several rates. And extensive calculations have also been made at the Go¬ vernment Life-Annuity Office, of the values of annuities on tw'o and on three joint lives, with distinction of the sexes. But none of these have been published, nor do they now (in July 1830) at that office grant any annuity depending upon more than one life, nor expect to do so for two or three years to come. Aged. Under 6 Between 6 and 11. Under 11 Between 11 and 16. Under 16 Between 16 and 21. Number of Males. 178 178 356 119 475 49 Females. 113 118 231 96 327 39 Both. 291 296 587 215 802 88 Under 21 Above that age, 524 ' 70 Total. 594 366 42 408 890 112 1002 In the absence of better information, it would seem reasonable to conclude that the younger a life was, the longer it would be likely to last; and accordingly we find that more than half the lives were under 11 years of a^e at the time of their nomination. And as males are more robust than females, it was also natural to conclude that a less rate of mortality would prevail among them; and accordingly three fifths of the nominees were males. . But subsequent observations have shown that, in both instances, the contributors made a bad choice. Besides, of all the nominees, only one ninth had com¬ pleted their 21st year at the time of their nomination. Not only the constitutions of these young nominees were not then fully formed or developed, but the mortality among them would depend greatly upon their destiny in after-life, or the circumstances of their respective situ- In this country, cases are continually occurring, in which an equitable adjustment of the rights and interests of different parties in property depending upon the conti¬ nuance or the failure of human life, is of great impor¬ tance ; and in many cases it cannot be made with suffi¬ cient accuracy without tables of the values of all possible combinations of two and even of three joint lives of differ¬ ent ages and of both sexes. But not unfrequently, espe¬ cially in cases of contingent reversions, we have been hi¬ therto, and are still, obliged to use approximations to the values of all the possible combinations of the lives involved, not only with each other, but also of them with others one year younger than each of them respectively. In these cases it sometimes happens that the whole value sought, being but a small part of a year’s purchase, is less than the probable error of several of these approximations considered separately; and then it is very difficult to give with confidence even a near approximation to the value sought. But in a great many other cases of frequent occurrence and less difficulty, the want of a complete set of good tables of this kind is much felt. It is therefore highly desirable that, numerous calculations for them having been made at the expense of the public, they should be completed at the public expense, and rendered accessible to persons having occasion to use them; by printing, if the expense would not be too great; otherwise, by hav¬ ing several manuscript copies accurately made, and depo¬ sited in convenient places for inspection, upon payment ofa small fee. ’ 1 A few writers on these subjects, of late years, have employed the differential and integral calculus in their in- ANN U f. vestigations. We have not yet seen.any fruits of this -^application of the calculus which appear to us of much va¬ lue, nor are we at all sanguine in expecting any. Although Lambert and Duvillard had made some efforts in this way before, Laplace (in his Theor. Anal, des Probabilites, No. 40) was the principal writer who thus treated the subject, and that very shortly, merely touch¬ ing upon the elements. He arrived in the usual manner at the same formulae that are given in the elementary al¬ gebraic method, and are here demonstrated by common arithmetic; only expressed in the manner of the higher calculus, in terms of the absciss and ordinate of the curve of mortality, both considered as variable quantities. He judiciously observed that the integral might be obtained in every case by calculating all its terms from a table of mortality, and taking their sum; and that in this manner tables of the values of annuities on single and joint lives might be calculated; which is only reverting to the usual method. But he also observed that the same thing might be effected by describing a parabolic curve through the ver¬ tices of the two extreme and several intermediate ordi¬ nates of the curve of mortality, and even that a few of these would be sufficient, since the differences between the different tables of mortality would justify us in con¬ sidering that method to be equally exact with those tables themselves. And in this we should entirely concur with that profound mathematician, provided we could admit that those tables, or most of them, had equal titles to our confidence, which he appears tacitly to assume. But here it is that M. Laplace appears to us to have fallen into the same error as most others respecting those tables of mortality, from not having paid sufficient atten¬ tion to the data they were constructed from, and the man¬ ner of their construction. After what has been advanced in this article, and in that on the Law of Mortality in this work, we think it quite I T I E S. 209 unnecessary to say more here, than that we consider it an History, established truth, that tables of mortality well construct- ed from proper data,for determining the values of annuities and reversions, do not differ materially from each other. If imperfect data for constructing a table of mortality be obtained, and any one already constructed, or the mean of several of them, be taken as a pattern or standard, to which it is desired that the new table should approach, it will not be difficult, by the known methods of approxi¬ mation and interpolation, so to construct such new table that it shall not differ much from the standard; but such new table, being in a certain degree hypothetical, can be of little or no value. According to the usual methods of treating these sub¬ jects, and constructing accurate tables, we never depart from the observations, but are supported by them at every step: our clear and simple methods of reasoning and cal¬ culation are much superior to the data we can obtain: proper data are alone wanting to further the science at present; government only can effectually supply them, and all who take any interest in these subjects must be grieved to find that there is little or no hope of assistance from that quarter. Even if a wiser course be adopted in future, 20 years more must elapse before we can reap the benefit of it. This is not the proper place to enter further into that part of the subject; but to those who take an interest in it, we would recommend the perusal of the minutes of evidence taken before the committee on the population bill, ordered by the House of Commons to be printed 11th May 1830 ; and the minutes of the committee on the re¬ committed bill, printed 26th May 1830; especially, in the latter, Mr Milne’s letter to Mr Davies Gilbert, the chair¬ man, in answer to an application made to him for his opi¬ nion, with Mr Rickman’s marginal notes on that letter, and his observations on it in his letter to the chairman, which Mr Milne knew nothing of till the bill was passed. NOTE REFERRED TO IN TWO PLACES ABOVE. According to Mr Morgan’s statements in the places here referred to, the number of members, or of assurances, or of policies, found to be in the Equitable Assurance Society, was— At the end of the year 1768 564 Policies 1770 490 Policies 1772 500 + Members... 1773 734 Members.... 1776 913 Policies 1783 1608 Members..., 1786 2100 -J- Members... 1792 4640 Assurances. 1799 5124 Members... From which we infer, that if, at the end of each year, beginning with 1770, and ending with 1799, the number had been taken, the sum of all the 30 would have been 75,664, and the mean number during these 30 years 2522. In a note on p. 443, vol. ii. of Dr Price’s Obs. on Rev. Paym. (7th edit.), Mr Morgan states, that during 33 years, from January 1768 to January 1801, the number of as¬ surances on single lives had been 83,201; but this great number can only be the sum of the 33 annual numbers as above mentioned, and the mean of these will be 2521. What we wish to know is, the mean number of lives insured on which policies were in force during the obser¬ vations ; but that Mr Morgan never mentions. As more policies than one are not unfrequently granted for so many distinct assurances on the same life, neither the number of policies nor of assurances will answer our purpose; neither Reference to Mr Morgan’s Statement. .View of Rise and Progress, &c. p. 10. Ditto ditto. Ditto p. 27. Address of 7th March 1793 p. 118.1 Ditto of 24th April 1800 p. 140. .Ditto of 7th March 1793 p. 118. .Ditto of 24th April 1800 p. 140. .View of Rise and Progress, &c. p. 24. .Ditto p. 26. will the number of members, for if a policy be granted to A for insurance on the life of B, A is the member of the society, and not B, who is only the life assured, and several other members besides A may insure the life of B, while A may also hold more policies than one insuring B’s life. As has been already observed, Mr Morgan has repeated¬ ly stated that the rate of mortality in the Equitable Society has always continued the same. And by tables a and b it appears, that out of 91,512 living persons in a similar society above 20 years of age, 1489 would die annually ; also, in the table at the end of Mr Morgan’s annuities (2d ed.), it is stated, that during the first 20 years of the 19th century, 1923 of the lives assured in the society died above 20 years of age; but 1489 :1923 :: 91,512:118,185, so that this last is the number of lives in a similar society, out of which these 1923 deaths would happen in one year. ' 1 hese addresses are printed at the end of the deed of settlement of the society, for the use of the members: the copy quoted from was printed in 1801. .n . rv i VOL. Ill, g jj 210 A N N U History. Then supposing, what is probably near the truth, that under 10 years of age the number of policies was the same as that of the lives insured, we shall have, by the state¬ ment last mentioned, Number of Annual Deaths. Between 10 and 20 years of age Above 20 Above 10 years of age , 119,679 And Number of Lives. 1,494 118,185 119,679 7 1923 1930 20 5984 is the mean number above 10. That is, a population falling short of 6000, instead of ex¬ ceeding 150,000, as stated by Mr Morgan. But by the table above mentioned in Mr Morgan’s work on annuities, and the explanations of it given above, it ap¬ pears that by 151,754 policies in force in the society, 119,679 lives were insured ; and 119,679:151,754 :: 7 : 9 nearly (more nearly:: 11:14). So that the number of lives was to that of the policies as 7 to 9 nearly, during the twenty years ending with 1820. I T I E S. On p. 61 of his pamphlet On the Rise and Progress of Hist the Equitable Society, Mr Morgan states the proportion inW 1827 to have been that of 3 to 4 nearly; and from an¬ other statement of his in a note on p. 42 of the same pamphlet, it would appear to have been that of 5 to 6 during the 12 years ending with 1827. The proportion above stated is nearly a mean between these two. Taking the mean number of lives on which assurances were in force in the Equitable Society during the last 30 years of the eighteenth century to have been 2522, as de¬ termined above ; since 119,679 : 2522 : : 1930 : 40*671, this last is the annual average number of deaths in the society during these 30 years; therefore the whole num¬ ber of them must have been 1220, which being added to 5124, the greatest number living at any one time during the term (that is, at the end of 1799, as stated above), the sum is 6344. And the greatest number of lives on which Mr Morgan’s observations were made during that term must have exceeded this, by the number who went out of the society by sale or forfeiture of the assurances, or by the expiration of the limited terms some of them were granted for, and did not enter it again; wherefore the sign (+) is added as before in table A. In treating of Annuities, we think that, it maybe useful in a work of this kind to address ourselves as well to those readers who have not, as to those who have, an acquaintance with Algebra ; and we shall accordingly divide what follows into two Parts, corresponding to these two views of the subject. PART I. We shall in this Part demonstrate all that is most use¬ ful and important in the doctrine of annuities and assur¬ ances on lives, without using algebra or introducing the idea of probability ; but the reader is of course supposed to understand common arithmetic. In the first 30 num¬ bers of this Part, compound interest and annuities-certain are treated of; from the 31st to the 76th the doctrine of annuities on lives is delivered ; and that of assurances on lives from thence to the 108th, where the popular view terminates. What is demonstrated in this Part will be sufficient to give the reader clear and scientific views of the subjects treated, and, with the assistance of the necessary tables, will enable him to solve the more common and simple pro¬ blems respecting the values of annuities and assurances. He will also understand clearly the general principles on which problems of greater difficulty are resolved; but these he cannot undertake with propriety when the ob¬ ject is to make a fair valuation of any claims or interests, with a view to an equitable distribution of property, un¬ less he has studied the subject carefully, wfith the assist¬ ance of algebra; for intricate problems of this kind can hardly be solved without it; and those who are not much exercised in such inquiries often think they have arrived at a complete solution, while they have overlooked some circumstance or event, or some possible combination of events or circumstances, which materially affects the value sought. Eminent mathematicians have in this way fallen into considerable errors, and it can hardly be doubted that those who are not mathematicians must (cceteris pa* ribus) be more liable to them. I.—On Annuities-Certain. No. 1. When the rate is 5 per cent., L.l improved at simple interest during one year will amount to L.H>5; which, improved in the same manner during the second year, will be augmented in the same ratio of 1 to 1*05: the amount then will therefore be 1*05 X 1*05, or fPOhV2 - 1*1025. V 7 In the same manner it appears that this last amount, improved at interest during the third year, vnll be increas¬ ed to (1*05)3 = 1*157625 ; at the end of the fourth year it will be (1*05)4; at the end of the fifth (1*05)5, and so on ; the amount at the end of any number of years being always determined by raising the number which expresses the amount at the end of the first year to the power of which the exponent is the number of years. So that when the rate of interest is 5 per cent., L.l improved at com¬ pound interest will in seven years amount to (1*05)7, and in 21 years to (1*05)21. But if the rate of interest were only 3 per cent., these amounts would only be (1*03)7 and (1*03)21 respectively. 2. The present value of L.l to be received certainly at the end of any assigned term, is such a less sum as, be¬ ing improved at compound interest during the term, will just amount to one pound. It must therefore be less than L.l, in the same ratio as L.l is less than its amount in that time; but in three years at 5 per cent. L.l will amount to L.(l*05)3 (1). And (1*05)3 : 1 :: 1 : 80 that (1-05)3 = L157625 = 0‘863838 is the Present value of L.l to be received at the expiration of three years. In the same manner it appears that, at 4 per cent, inter¬ est, the present value of L.l to be received at the end of a year is 0*961538; and if it were not to be re¬ ceived until the expiration of 21 years, its present value WOUld be (I-04)2i ~ (°'961538)21 = 0*438834. Hence it appears, that if unit be divided by the amount of L.l improved at compound interest during any num¬ ber ol years, the quotient will be the present value of L.l to be received at the expiration of the term; which may also be obtained by raising the number which expresses the present value of L.l receivable at the expiration of a year, to the power of which the exponent is the number of years in the term. ANNUITIES. 211 Popta- 3. When a certain sum of money is receivable annual- Vie ly, it is called an Annuity, and its quantum is expressed ■^/w/by saying it is an annuity of so much; thus, according as the annual payment is L.l, L.10, or L.100, it is called an annuity of L.l, of L.10, or of L.100. 4. When the annual payment does not depend upon any contingent event, but is to be made certainly, either in perpetuity or during an assigned term, it is called an Annuity-certain. 5. In calculating the value of an annuity, the first pay¬ ment is always considered to be made at the end of the first year from the time of the valuation, unless the con¬ trary be expressly stated. 6. Tlie whole number, and part or parts of one annual payment of an annuity, which all the future payments are worth in present money, is called the number of years purchase the annuity is worth, and, being the sum of the present values of all the future payments, is also the sum which, being put out and improved at compound interest, will just suffice for the payment of the annuity. (2.) 7. Hence it follows, that when the annuity is L.l, the number of years’ purchase and parts of a year is the same as the number of pounds and parts of a pound in its pre¬ sent value. And throughout this article, whenever the quantum of an annuity is not mentioned, it is to be understood to be 10. The calculation must begin with Table III., the first number in which should evidently be 1*05, the amount of L.l improved at interest during one year, which' being multiplied by 1*05, the product is L025, the second number. This second number being multiplied by 1-05, the product is 1*157625, the amount at the end of three years. And so the calculation proceeds throughout the whole of the column; each number after the first being the product of the multiplication of the preceding number, by the amount of L.l in a year. (1.) 11. The number against any year in Table I. is found by dividing unit by the number against the same year in Table III. (2); thus, the number against the term of six years in Table I. is = *746215. All the numbers J 1*340096 in that table after the first may also be found by multi¬ plying that first number continually into itself. (2.) 12. The number against any year in Table II., being the sum of the numbers against that and all the preceding years in Table I., is found by adding the number against that year in Table I. to the number against the preceding year in Table II.; thus, the number against four years in Table II. being the sum of 0*822702 and 2*723248 Popular View. L.l. 8. The sum of which the simple interest for one year is L.l, is evidently that which, being put out at interest, will just suffice for the payment of L.l at the end of every year, without any augmentation or diminution of the prin¬ cipal, and, being equivalent to the title to L.l per annum for ever, is called the value of the perpetuity, or the num¬ ber of years’ purchase the perpetuity is worth. But while the rate remains the same, the annual inte¬ rests produced by any two sums are to each other as the principals which produce them; therefore, since 5 : 1 :: 100 : 100 “IT 20, when the rate of interest is b per cent., the value of the perpetuity is 20 years’ purchase. In the same manner it appears, that according as the rate may be 3 or 6 per cent., the value of the perpetuity will be -g- = 33^, or =r 16§ years’ purchase; and may be found in every case, by dividing any sum by its interest for a year. 9. All the most common and useful questions in the doctrines of compound interest and annuities-certain may be easily resolved by means of the first four tables at the end of this article. Their construction may be explained by the following specimen, rate of interest 5 per cent. CONSTRUCTION OF Term. Table IV. Table III. Table I. Table II. Amount of L. 1 jtcr an¬ num Amount of L.l improved at Interest until Present value of L.l to be received at Present va¬ lue of L.l per annum, to be receiv¬ ed until Term. the Expiration of the Term. 1 yr. 2 yrs, 3 4 5 6 7 1-000000 2*050000 3*152500 4*310125 5*525631 6*801913 8*142009 1*050000 1*102500 1*157625 1*215506 1*276282 1*340096 1*407100 •952381 •907029 •863838 •822702 •783526 •746215 •710681 0*952381 1*859410 2*723248 3*545950 4*329476 5*075691 5*786372 1 yr. 2 yrs. 3 4 5 6 7 is 3*545950. 13. If each payment of an annuity of L.l be put out as it becomes due, and improved at compound interest du¬ ring the remainder of the term, it is evident that at the expiration of the term the payment then due will be but L.l, having received no improvement at interest. That received one year before will be augmented to the amount of L.l in a year; that received two years before will be augmented to the amount of L.l in two years; that re¬ ceived three years before to the amount of L.l in three years; and so on until the first payment, which will be augmented to the amount of L.l in a term one year less than that of the annuity. Hence it is manifest that the number against any year in Table IV. will be unit added to the sum of all those against the preceding years in Table III. And therefore that the number against any year in Table IV. is the sum of those in Tables III. and IV. against the next preceding year. Thus, the number against seven years in Table IV. be¬ ing the sum of 1*340096 and 6*801913 is 8*142009. 14. The method of construction is obviously the same at any other rate of interest. 15. All the amounts and values which are the objects of this inquiry evidently depend upon the improvement of money at compound interest; it is therefore, that the first, second, and fourth tables, all depend upon the third. But every pound, and every part of a pound, when put out at interest, is improved in the same manner as any single pound considered separately. Whence it is ob¬ vious, that while the term and the rate of interest remain the same, both the amount and the present value, either of any sum or of any annuity, will be the same multiple, and part or parts of the amount or the present value found against the same term, and under the same rate of interest in these tables, as the sum or the annuity pro¬ posed is of L.L So that to find the amount or the present value of any sum or annuity for a given term and rate of interest, we have only to multiply the corresponding tabular value by 212 ANNUITIES. Popular the sum or the annuity proposed; the product will be the View, amount or the value sought, according as the case may be. 16. Example 1. To what sum will L.100 amount when improved at compound interest during 20 years, the rate of interest being 4 per cent, per annum ? By Table III. it appears that L.l so improved would, at the expiration of the term, amount to L.2'191123 ; therefore L.100 would amount to 100 times as much, that is, to L.219T123, or L.219. 2s. 3d. 17. Ex. 2. What is the present value of L.400, which is not to be received until the expiration of 14 years, when the rate of interest is 5 per cent. ? The present value of L.l to be received then will be found by Table I. to be L.0’505068 : L.400 to be received at the same time will therefore be worth, in present money, 400 times as much, or L.202‘0272, that is, L.202. 0s. 6|d. 18. Ex. 3. Required the present value of an annuity of L.50 for 21 years, when the rate of interest is 5per cent. Table II. shows the value of an annuity of L.l for the same term to be L.12,8212; the required value must therefore be 50 times as much, or L.64L06, that is L.641. Is. 2Jd. 19. Ex. 4. What will an annuity of L.10. 10s. or L.10-5 for thirty years amount to, when each payment is put out as it becomes due, and improved at compound interest until the end of the term; the rate of interest being 4 per cent. ? The amount of an annuity of L.l so improved would be L.56-084938, as appears by Table IV.; the amount requir¬ ed will therefore be 10*5 times this, or L.588*89185, that is L.588. 17s. lOd. 20. When the.interval between the time of the pur¬ chase of an annuity and the first payment thereof exceeds that which is interposed between each two immediately successive payments, such annuity is said to be deferred for a time equal to that excess, and to be entered upon at the expiration of that time. 21. If two persons, A and B, purchase an annuity be¬ tween them, which A is to enter upon immediately, and to enjoy during a certain part of the term, and B or his heirs or assigns for the remainder of it, the present value of B's interest will evidently be the excess of the value of the annuity for the whole of the term from this time, above the value of the interest of A. So that when the entrance on an annuity is deferred for a certain term, its present value will be the excess of the value of the annuity for the term of delay and continuance together, above the value of an equal annuity for the term of delay only. 22. Example 1. Required the value of a perpetual an¬ nuity of L.120, which is not to be entered upon until the expiration of 14 years from this time, reckoning interest at 3 per cent. The perpetuity, with immediate possession, would be worth 33t years’ purchase (8); and an annuity for the term of delay is worth 11-2961 (Table II.) From 33-3333 subtract 11-2961, and multiply the remainder 22-0372 by 120 the product, 2644-464 = L.2644. 9s. 34d., is the required value. 2^. Ex. 2. Allowing interest at 5 per cent., what sum should be paid down now, for the renewal of 14 years lapsed in a lease for 21 years of an estate producing L.300 per annum, clear of all deductions ? I his is the price of an annuity for 14 years, to be en¬ tered upon 7 years hence ; the term of delay, therefore, is 7 years, and that of the delay and continuance together 21 years. By Table II. it appears, that the present value of an annuity for 21 years, is 12-8212^ for 7 J,ears’ 5,78(54 > years’ purchase. Value of the deferred annuity, 7-0348 J Multiply by 300 ! The product, L.2110-44, or L.2110. 8s. 9jd., is the price required. 24. Hitherto we have proceeded upon the supposition of the annuity being payable, and the interest convertible into principal, which shall reproduce interest, only once a year. But annuities are generally payable half yearly, and sometimes quarterly; and the same circumstances that render it desirable for an annuitant to receive his annual sum in equal half-yearly or quarterly portions, also give occasion to the interest of money being paid in the same manner. But whatever has been advanced above concerning the present value or the amount of an annuity, when both that and the interest of money were only payable once a year, will evidently be true when applied to half the annuity and half the interest paid twice as often, on the supposition of half-yearly payments; or to a quarter of the annuity and a quarter of the interest paid four times as often, when the payments are made quarterly. 25. Half-yearly payments are, however, by far the most common; and these four tables will also enable us to answer the most useful questions concerning them. For we have only to extract the present value, or the amount, from the table, against twice the number of years in the term, at half the annual rate of interest, and, in the case of an annuity, to multiply the number so extracted by half the annuity proposed. 26. Ex. 1. To what sum will L.100 amount in 20 years, when the interest at the rate of 4 per cent, per annum is convertible into principal half-yearly? This being the amount in 40 half years at 2 per cent. interest for every half year, will be the same as the amount in 40 years at 2 per cent, per annum, which, by Table III. will be found to be 220-804, or L.220. 16s. Id.; and is only L.L 13s.TOd. more than it would amount to if the interest were not convertible more than once a year. (iO.) 27. Ex. 2. W hat is the present value of an annuity of L.50 for 21 years, receivable in equal half-yearly pay¬ ments, when money yields an interest of 2\per cent, every half year ? By Table II. it appears, that an annuity of L.l for 42 years, when the interest of money is 21 per cent, per an- num, will be worth L.25-8206 (25) ; 25 times this sum, or L.645. 10s. 3id., is therefore the required value, and ex¬ ceeds the value when the interest and the annuity are only payable once a year, by L.4. 9s. Id. (18.) 28. I he excess of an annuity-certain above the interest of the purchase-money, is the sum which, being put out at the time of each payment becoming due, and improved at compound interest until the expiration of the term, will just amount to the purchase-money ox-iginally paid. But, while every thing else remains the same, the longer the term of the annuity is, the less must its excess above the interest of the purchase-money be, because a less an¬ nuity will suffice for raising the same sum within the term. Therefore, the proportion of that excess to the annual in¬ terest of the purchase-money continually diminishes as ANNUITIES. 213 the term is extended; and when the annuity is a perpe¬ tuity, there is no such excess. (8.) 29. The reason why the value of an annuity is increas¬ ed by that and the interest being both payable more than once in the year, is, that the granter loses and the pur¬ chaser gains the interest produced by that part of each payment which is in excess above the interest then due upon the purchase-money, from the time of such payment being made until the expiration of the year. Hence it is obvious, that the less this excess is, that is, the longer the term of the annuity is (28), the less must the increase of value be. And when the annuity is a perpetuity, its value will be the same, whether it and the interest of money be both payable several times in the year, or once only. 30. When the annuity is not payable at the same inter¬ vals at which the interest is convertible into principal, its value will depend upon the frequencies both of payment and conversion; but its investigation without algebra would be too long, and of too little use, to be worth pro¬ secuting here. II.—Of Annuities on Lives. 31. When the payment of an annuity depends upon the existence of some life or lives, it is called a Life-Annuity. 32. The values of such annuities are calculated by means of tables of mortality, which show, out of a consi¬ derable number of individuals born, how many upon an average have lived to complete each year of their age, and, consequently, what proportion of those who attained to any one age have survived any greater age. The fifth table at the end of this article is one of that kind which has been taken from Mr Milne’s Treatise on Annuities, and was constructed from accurate observations made at Carlisle by Dr Heysham, during a period of nine years, ending with 1787. 33. By this table it appears, that during the period in which these observations were made, out of 10,000 chil¬ dren born, 3203 died under five years of age, and the re¬ maining 6797 completed their fifth year. Also, that out of 6797 children who attained to five years of age, 6460 survived their tenth year. But the mortality under ten years of age has been great¬ ly reduced since then by the practice of vaccination. This table also shows, that of 6460 individuals who attained to 10 years of age, 6047 survived 21; and that of 5075 who attained to 40, only 3643 survived their 60th year. 34. There is good reason to believe (as has been shown in another place) that the general law of mortality, that is, the average proportion of persons attaining to any one age, who survive any greater age, remains much the same now among the entire mass of the people throughout Eng¬ land, as it was found to be at Carlisle during the period of these observations, except among children under ten years of age, as was noticed above. (33.) If this be so, it will follow, that of 6460 children now 10 years of age, just 6047 will attain to 21; or rather, that if any great number be taken in several instances, this will be the average proportion of them that will survive the period. And if 6460 children were to be taken indiscriminately from the general mass of the population at 10 years of age, and an office or company were to engage to pay L.l eleven years hence for each of them that might then be living, this engagement would be equivalent to that which should bind them to pay L.6047 certainly at the expira¬ tion of the term. Therefore the office, in order that it might neither gain nor lose by the engagement, should, upon entering into it, be paid for the whole, the present value of L.6047 to be received at the expiration of eleven years ; and for each life the -g^goth part of it, that is, the ffto^ Part t^ie Present value of L. 1, to be received then. But when the rate of interest is 5 per cent., the present value of L.l, to be received at the expiration of 11 years, is L.0-584679; therefore, at that rate of interest, there Popular View,. should be paid for each life 6047 X 0-584679 6460 L.0-5473. And the present value of L.100, to be received upon a life now 10 years of age attaining to 21, will be L.54'73, or L.54. 14s. 7d. In the same manner it will be found, that reckoning interest at 4/>er cent, the value would be L.60. 16s. Id. 35. This is the method of calculating the present values of endowments for children of given ages ; and the values of annuities on lives may be computed in the same manner. For, from the above reasoning it is manifest, that if the present value of L.l, to be received certainly at the ex¬ piration of a given term, be multiplied by the number in the table of mortality against the age, greater than that of any proposed life by the number of years in the term, and the product be divided by the number in the same table, against the present age of that life; the quotient will be the present value of L.l, to be received at the ex¬ piration of the term, provided that the life survive it. And if, in this manner, the value be determined of L.l, to be received upon any proposed life, surviving each of the years in its greatest possible continuance, according to the table of mortality adapted to it; that is, according to the Carlisle table, upon its surviving every age greater than its present, to that of 104 years inclusive; then, the sum of all these values will evidently be the present value of an annuity on the proposed life. 36. If 5642 lives at 30 years of age be proposed, and 5075 at the age of 40; since each of the 5642 younger lives may be combined with every one of the 5075 that are 10 years older, the number of different pairs, or dif¬ ferent combinations of two lives differing in age by 10 years, that may be formed out of the proposed lives, is 5642 times 5075. But at the expiration of 15 years the survivors of the lives now 30 and 40 years of age, being then of the re¬ spective ages of 45 and 55, will be reduced to the num¬ bers of 4727 and 4073 respectively; and the number of pairs, or combinations of two, differing in age by 10 years, that can be formed out of them, will be reduced from 5642 X 5075 to 4727 X 4073. So that L.l to be paid at the expiration of 15 years for each of these 5642 X 5075 pairs or combinations of two now existing, which may survive the term, will be of the same value in present money as 4727 times L.4073 to be received certainly at the same time. Now, let A be any one of these lives of 30 years of age, and B any one of those aged 40; and, from what has been advanced, it will be evident that the present value of L.l, to be received upon the two lives in this particular com¬ bination jointly surviving the term, will be the same as that of the sum L.4727X 4073 5642 X 5075 to be then received certainly. But, when the rate of interest is 5 per cent, L.l to be received certainly at the expiration of 15 years, is equiva¬ lent to L.0-481017 in present money. (Table I.) Therefore, at that rate of interest, and according to the Carlisle table of mortality, the present value of L.l to be received upon A and B, now aged 30 and 40 years re¬ spectively, jointly surviving the term of 15 years, will be 4727 X 4073 X L.0-481017 5642 X 5075 214 ANNUITIES. Popular View. 37. Hence It Is sufficiently evident how the present value of L.l to be received upon the same two lives jointly surviving any other year may be found. And if that value for each year from this time until the eldest life attain to the limit of the table of mortality be calculated, the sum of all these will be the present value of an annuity of L.l dependent upon their joint continuance. In this manner, it is obvious that the value of an annuity on the joint continuance of any other two lives might be determined. 38. If, besides the 5642 lives at 30 years of age, and the 5075 at 40 (mentioned in No. 36), there be also pro¬ posed 3643 at 60 years of age, each of these 3643 at 60 may be combined with every one of the 5642 X 5075 dif¬ ferent combinations of a life of 30, with one of 40 years of age; and, therefore, out of these three classes of lives 5642 X 5075 X 3643 different combinations may be formed, each containing a life of 30 years of age, another of 40, and a third of 60. But at the expiration of 15 years the numbers of lives in these three classes will, according to the table of mor¬ tality, be reduced to 4727, 4073, and 1675 respectively; the respective ages of the survivors in the several classes being then 45, 55, and 75 years; and the number of dif¬ ferent combinations of three lives (each of a different class from either of the other two) that can be formed out of them, will be reduced to 4727 X 4073 X 1675. Hence, by reasoning as in No. 36, it will be found, that if A, B, and C be three such lives, now aged 30, 40, and 60 years, the present value of L.l to be received upon these three jointly surviving the term of 15 years from this time, will be 4727 X 4073 X1675 5642 X 5075 X 3643 XL.0'481017 : inte¬ rest being reckoned at 5 per cent. Thus it is shown how the present value of an annuity dependent upon the joint continuance of these three lives might be calculated, that being the sum of the present values thus determined, of the rents for all the years which, according to the table of mortality, the eldest life can survive. 39. But it is easy to see that the same method of rea¬ soning may be used in the case of four, five, or six lives, and so on without limit. Whence this inference is obvious. The present value of L.l to be received at the expira¬ tion of a given term, provided that any given number of lives all survive it, may be found by multiplying the pre¬ sent value of L.l to be received certainly at the end of the term, by the continual product of the numbers in the table of mortality against the ages greater respectively by the number of years in the term than the ages of the lives proposed, and dividing the last result of these ope¬ rations by the continual product of the numbers in the table of mortality against the present ages of the proposed lives. And by a series of similar operations, the present value of an annuity on the joint continuance of all these lives might be determined. But it should be observed, that, in calculating the value of a life-annuity in this way, the denominator of the frac¬ tions expressing the values of the several years’ rents, that is, the divisor used in each of the operations, remains al¬ ways the same: the division should, therefore, be left till the sum of the numerators is determined; and one opera¬ tion of that kind will suffice. 40. Enough has been said to show that these methods of constructing tables of the values of annuities on lives are practicable, though excessively laborious; and, in fact, all the early tables of this kind were constructed in that manner. We proceed now to show how such tables may be calculated with much greater facility. 41. By the method of No. 34, it' will be found that, reckoning interest at 5 per cent., the present value of V«l L.l to be received at the expiration of a year, provided'-^ that a life, now 89 years of age, survived till then, is 142 X 0 95~38 ^ ^ ^ge 0f that jjf€ wjji tjlen ^ 181 years, and the proprietor of an annuity of L.1 now depend¬ ing upon it will, in that event, receive his annual payment of L.l then due; therefore, if the value then of all the subsequent payments, that is, the value of an annuity on a life of 90, be 2‘339 years’ purchase, the present value of what the title to this annuity may produce to the proprie¬ tor at the end of the year will be the same as that of L.3'339, to be received then, if the life be still subsisting, or *•-—-■ X L.3 339 , wniclij tlicrc- lol fore, will be the present value of an annuity of L.l on a life of 89 years of age. That is to say, an annuity on that life will now be worth 2-495 years’ purchase. (7.) 42. In the same manner it appears generally, that if unit be added to the number of years’ purchase that an annuity on any life is worth, and the sum be multiplied by the present value of L.l to be received at the end of a year, provided that a life one year younger survive till then, the product will be the number of years’ purchase an annuity on that younger life is worth in present money. 43. But, according to the table of mortality, an annuity on the eldest life in it is worth nothing; therefore, the present value of L.l to be received at the end of a year provided that the eldest life but one in the table survive till then, is the total present value of an annuity of L.l on that life; which being obtained, the value of an annuity on a life one year younger than it may be found by the preceding number; and so on for every younger life suc¬ cessively. Exemplification. Rate of Interest 5 per cent. Age of Life. 104 103 102 101 100 99 98 97 96 95 Value of an Annuity on that Life, increased by Unit, 1-000 1-317 1- 753 2- 192 2- 624 3- 045 3-278 3-428 3-555 3-596 Which be¬ ing multi¬ plied bv 0-952381, and the Product by tV •H rt If Produces the value of an Annuity on the next younger Life, 0-317 0-753 T192 1- 624 2- 045 2-278 2-428 2-555 2-596 2-569 Its Age being 103 102 101 100 99 98 97 96 95 94 44. Proceeding as in No. 36, it will be found that, at 5 per cent, interest, and according to the Carlisle table of mortality, the present value of L.l to be received at the expiration of a year provided that a person now 89 years of age, and another now 99, be then living, is 142 X 9 X L.0-952381 , rsTxTI ’ t*lere*ore> “ the present value ot an annuity of L.l on the joint continuance of two lives, now aged 90 and 100 years respectively, be L.0-950, by reasoning as in No. 41, it will be found that the present value of an annuity on the joint continuance of two lives, ANNUITIES. 215 ;j of the respective ages of 89 and 99 years, will be worth ^142Xj9X0;952381 x 1.930_1.192 years’ purchase. 45. In this manner, commencing with the two oldest lives in the table that differ in age by ten years, and pro¬ ceeding according to No. 43, the values of annuities on all the other combinations of two lives of the same difference of age may be determined. The method is exemplified in the following specimen. Ages of two Lives. Value of an Annuity on their joint continuance, increased by Unit, Which be¬ ing multi¬ plied bv 0-952381, and the Pro. duct by Produces the value of an Annuity on the two joint Lives one year younger re¬ spectively, Their Ages being 94 & 104 93 & 103 92 & 102 91 & 101 90 & 100 89 & 99 1-000 1-235 1-508 1-733 1- 950 2- 192 1X40 3X54 3X54 5X75 5X75 7X 105 7X105 9X 142 9X142 11X181 11X181 14X232 0-235 0-508 0-733 0-950 1-192 1-280 93 & 103 92 & 102 91 & 101 90 & 100 89 & 99 88 & 98 46. Hence, and by what has been advanced in the 39th number of this article, it is sufficiently evident how a table may be computed of the values of annuities on the joint continuance of the lives in every combination of three, or any greater number; the differences between the ages of the lives in each combination remaining al¬ ways the same in the same series of operations, while the calculation proceeds back from the combination in which the oldest life is the oldest in the table, to that in which the youngest is a child just born. 47. But when there are more than two lives in each combination, the calculations are so very laborious, on ac¬ count principally of the great number of combinations, that no tables of that kind have yet been published, ex¬ cept three or four for three lives. And in the books that contain tables of the values of two joint lives, methods are given of approximating to¬ wards the values of such combinations of two and of three lives as have not yet been calculated. Therefore, assuming the values of annuities on single fives, and on the joint continuance of two or of three lives, to be given, we have next to show how the most useful problems respecting the values of any interests that de¬ pend upon the continuance or the failure of life may be resolved by them. 48. Proposition 1. The value of an annuity on the survivor of two lives, A and J3, is equal to the excess of the sum of the values of annuities on the two single lives above the value of an annuity on their joint continuance. 49. Demo?istration. If annuities on each of the two lives were granted to P, during their joint continuance he would have two annuities; but if jP were only to re¬ ceive these upon condition that, during the joint lives of A and B, he should pay one annuity to Q, then there would only remain one to be enjoyed by him or his heirs or assigns, until the lives both of A and B were extinct; whence the truth of the proposition is manifest. 50. Prop. 2. The value of an annuity on the joint con- Popular tinuance of the two last survivors out of three lives, A, View. B, and C, is equal to the excess of the sum of the values of annuities on the three combinations of two lives {A with B, A with C, and B with C) that can be formed out of them, above twice the value of an annuity on the joint continuance of all the three lives. 51. Bern. If one annuity were granted to P on the joint continuance of the two lives A and B, another on the joint continuance of A and £7, and a third on the joint continuance of B and C} during the joint continuance of all the lives he would have three annuities. But if he were only to receive these upon condition that he should pay two annuities to Q during the joint continuance of all the three lives, then there would only remain to himself one annuity during the joint existence of the last two survivors out of the three lives. And the truth of the proposition is manifest. 52. Prop. 3. The value of an annuity on the last sur¬ vivor of three lives, A, B, and C, is equal to the excess of the sum of the values of annuities on each of the three, single lives, together with the value of an annuity on the joint continuance of all the three, above the sum of the values of three other annuities; the first dependent upon the joint continuance of A and B, the second on that of A and C, and the third on B and C. 53. Bern. If annuities on each of the three single lives were granted to R, during the joint continuance of all the three he would have three annuities; and from the time of the extinction of the first life that failed, till the ex¬ tinction of the second, he would have two. So that he would have two annuities during the joint existence of the two last survivors out of the three lives; and besides these, a third annuity during the joint conti¬ nuance of all the three. Therefore, if out of these R were to pay one annuity to P during the joint continuance of the last two survir vors out of the three lives, and another to Q during the joint continuance of all the three, he would only have left one annuity, which would be receivable during the life of the last survivor of the thx-ee. But in the demonstration of the last proposition (51) it was shown, that the value of what he paid to .P would fall short of the sum of the values of annuities dependent respectively on the joint continuance of A and B, of A and C, and of B and C, by two annuities on the joint continuance of all the three lives. Whence it is evident that the value of the annuities he paid both to P and Q would fall short of the sum of these three values of joint lives, only by the value of one annuity on the joint conti¬ nuance of all the three lives. Wherefore, if from the sum of the values of all the three single lives, the sum of the values of the three com¬ binations of two that can be formed out of them were ta¬ ken, there would remain less than the value of an annuity on the last survivor, by that of an annuity on the joint continuance of the three lives. But if, to the sum of the values of the three single lives A, B, and C, there be added that of an annuity on the joint continuance of the three, and from the sum of these four values the sum of the values of the three combina¬ tions A with B, A with C, and B with C be subtracted, then the remainder will be the value of an annuity on the last survivor of the three lives ^ which was to be demon¬ strated. 54. Prop. 4. Problem. The law of mortality and the values of single lives at all ages being given, to determine the present value of an annuity on any proposed life, de¬ ferred for any assigned term. 55. Solution. Find the present value of an annuity on 216 ANNU Popular View. a life older than the proposed, by the number of years during which the other annuity is deferred; multiply this by the present value of L.l to be received upon the pro¬ posed life surviving the term, and the product will be the value sought. 56. Dem. Upon the proposed life surviving the term, the annuity dependent upon it will be worth the same sum that an annuity on a life so much older is now worth ; therefore it is evident that the deferred annuity is of the same present value as that sum to be received at the ex¬ piration of the term, provided the life survive it. 57. Corollary. In the same manner it appears, that the present value of an annuity on the joint continuance of any number of lives, deferred for a given term, may be found by multiplying the present value of an annuity on the joint continuance of the same number of lives, older respectively than the proposed by the number of years in the term, by the present value of L.l to be received upon the proposed lives all surviving it. 58. A temporary annuity on any single life, or on the joint continuance of any number of lives, that is, an an¬ nuity which is to be paid during a certain term, provided that the single life or the other lives jointly subsist so long, together with an annuity on the same life or lives deferred for the same term, is evidently equivalent to an annuity on the whole duration of the same life or lives. So that the value of an annuity on any life, or on the joint continuance of any number of lives, for an assigned term, is equal to the excess of the value of an annuity on their whole duration, with immediate possession, above the value of air annuity on them deferred for the term. 59. Whatever has been advanced from No. 48 to 53 inclusive, respecting the values of annuities for the whole duration of the lives whereon they depend, will apply equally to those which are either deferred or temporary; and therefore, to determine the value of any deferred or temporary annuity dependent upon the last survivor of two or of three lives, or upon the joint continuance of the last two survivors out of three lives, we have only to substi¬ tute temporary or deferred annuities, as the case may re¬ quire, for annuities on the whole duration of the lives; and the result will accordingly be the value of a tempo- rary or deferred annuity on the life of the last survivor, or on the joint lives of the two last survivors. 60. Prop. 5. A and B being any two proposed lives now in existence, the present value of an annuity to be payable only while A survives L?, is equal to the excess of the value of an annuity on the life of A above that of an annuity on the joint existence of both the lives. 61. Dem. If an annuity on the life of ^4, and to be entered upon immediately, were now granted to P upon condition that he should pay it to B during the joint lives of and _Z?, it is evident that there would only remain to P an annuity on the life of A after the decease of B; whence the truth of the proposition is manifest. y When any thing is affirmed or demonstrated any life or lives, it is to be understood as applying equally to any proposed single life, or to the joint continuance of the whole of any number of lives that may be proposed to¬ gether, or to that of any assigned number of the last sur¬ vivors of them, or to the last surviving life of the whole. 63. Prop. 6. The present value of the reversion of'a perpetual annuity after the failure of any life or lives, is equal.to the excess of the present value of the perpetuity, with immediate possession, above the present value of an annuity on the same life or lives. 64. Dem. If a perpetual annuity with immediate pos¬ session were granted to P, upon condition that he should pay the annual produce to another individual during the existence of the life or lives proposed, it is evident that I T I E S. there would only remain to P the reversion after the Po failure of such life or lives; and the present value of that VI reversion would manifestly be as stated above. ^ , 65. The 6th, 7th, and 8th tables at the end of this article, which have been extracted from the 19th, 21st, and 22d respectively, in Mr Milne’s Treatise on Annuities, will serve to illustrate the application of these propositions to the solution of questions in numbers. In all the following examples, we suppose the lives to be such as the general average of those from which the Carlisle table of mortality was constructed, and the rate of interest to be hper cent. 66. Ex. 1. What is the present value of an annuity on the joint lives, and the life of the survivor of two persons now aged 40 and 50 years respectively ? According to No. 48, the process is as follows: Value of a single life of 40 50 sum Subtract the value of their | joint lives, J 13-390 11-660 | (by Table VI.) 25-050 9-984 (Table VIII.) remains 15-066 years’purchase, the required value. And if the annuity be L.200, its present value will be L.3013-2, or L.3013. 4s. 67. Ex. 2. The lives A, B, and C, being now aged 50, 55, and 60 years respectively, an annuity on the joint con¬ tinuance of all the three is worth 6-289 years’ purchase, what is the value of an annuity on the joint existence of the last two survivors of them ? According to No. 50, the process is thus: Ages. Values. 50 & 55 55 & 60 50 & 60 mog} (Table VIL> 7-601 (Table VIII.) sum 23-235 Subtract 2 X 6-289 =12-578 remains 10-657 years’ purchase, the required value. Therefore, if the annuity were L.300, it would be worth L.3197. 2s. in present money. 68. Ex. 3. Required the value of an annuity on the last survivor of the three lives in the last example. Proceeding according to No. 52, we have Ages. Values. 50 55 60 50, 55, & 60 11-660 ) 10-347 } (Table VI.) 8-940 ) 6-289 (No. 67). sum 37-236 Subtract the sum of the values! of annuities on the three com- >- 23*235 (No. 67.) binations of two lives, J remains 14-001 years’ purchase, the required value. And if the annuity were L.300, it would now be worth L.4200. 6s. 69. Ex. 4. What is the present value of an annuity on a life now 45 years of age, which is not to be entered upon until the expiration of ten years ; the first payment there¬ of being to be made at the expiration of eleven years from this time, if the life survive till then ? ANNUITIES. 217 popu,1 Vie'* Solution. fK" . The present value of an annuity on a life of 55 is 10-347 (Table VI.), and the present value of L.l to be received upon the proposed life attaining to the age of 55, is X 0-613913 ; therefore, by No. 55, the required value is 4073 X 0-613913 X 10-347 -5.4,73 s> purciiase; so 4727 ^ 1 that if the annuity were L.200, its present value would be L.1094. 12s. 70. Ex. 5. Required the present value of an annuity to be received for the next ten years, provided that a person now 45 years of age shall so long live. Solution. The present value of an annuity on a life of 45, to be entered upon immediately, is 12-648 (Table VI.) Subtract that of an annuity onl the same life deferred 10 years, j 5-473 ^ the remainder 7-175 is the required number of years’ purchase. And if the annuity were L.200, its present value would be L.1435. 71. Ex. 6. An annuity on a life of 45, deferred 10 years, was shown in No. 69 to be worth 5-473 years’ purchase in present money; let it be required to determine the equiva¬ lent annual payment for the same, to be made at the end of each of the next 10 years, but subject to failure upon the life failing in the term. Solution. The present value of L.l per annum on the proposed life for the next 10 years has just been shown to be L.7-175, and this, multiplied by the required annual pay¬ ment, must produce L.5-473; that payment must, there- fore, be 77^ = 0-76279. And since the annual pay¬ ment for the deferred annuity of L.l per annum would be L.0-76279, that for an annuity of L.200 must be L.152. 11s. 2d. 72. Ex. 7. Let the present value be required of an an¬ nuity on a life now 40 years of age, to be payable only while that life survives another now of the age of 50 years. Frnfo of40resent va'ue °f“113'390 (Table VL) Subtract that of the two joint | g<984 (Table yjjj) 75. Ex. 8. Let it be required to determine the present Popular value of the reversion of a perpetual annuity after the View, failure of a life now 50 years of age. Solution. The value of the perpetuity is 20 years’ purchase. (8.) Subtract that of an annuity onl the life of 50, j 11-660 (Table VI.) the remainder, 3-406 years’ purchase, is the required value. (60.) Therefore, if the annuity were L.100, it would be worth L.340. 12s. in present money. 73. If the annuity in the last example were to be paid for by a constant annual premium at the end of each year while both the lives survived; by reasoning as in No. 71, it will be found, that such annual premium for an annuity of L.l should be = L.0-341146 ; for an annuity of L.100 it should therefore be L.34. 2s. 3^d. 74. Rut if one of the equal premiums for this annuity is to be paid down now, and another at the end of each year while both the lives survive, the number of years’ purchase the whole of these premiums are worth will evidently be increased by unit; on account of the payment made now, it will therefore be 10-984 ; and each premium for an annuity of L.l must, in this case, be -f, = lU-yo4i L.0-310087; for an annuity of L.100 it should therefore he L.31. 0s. 2d. vol. in. remains 8-34 years’ purchase, the required value of the reversion. (63.) So that if the annuity were L.300, its present value would be L.2502. 76. In the same manner it will be found, by the 68th number and those referred to in the last example, that the reversion of a perpetuity, after the failure of the last survivor of three lives, now aged 50, 55, and 60 years re¬ spectively, is worth 5-999 years’purchase in present money; therefore, if it were L.100 per annum, its present value would be L.599. 18s. III.—Of Assurances on Lives. 77. An assurance upon a life or lives is a contract by which the office or underwriter, in consideration of a stipulated premium, engages to pay a certain sum upon such life or lives failing within the term for which the as¬ surance is effected. 78. If the term of the assurance be the whole duration of the life or lives assured, the sum must necessarily be paid whenever the failure happens ; and, in what follows, that payment is always supposed to be made at the end of the year in which the event assured against takes place; the anniversary of the assurance, or the day of the date of the policy, being accounted the beginning of each year. 79. At the end of the year in which any proposed life or lives may fail, the proprietor of the reversion of a per¬ petual annuity of L.l after their failure will receive the pound then due, and will at the same time enter upon the perpetuity ; therefore, the present value of the rever¬ sion is the same as that of L.l added to the money a per¬ petual annuity of L.l would cost, supposing this sum not to be receivable until the expiration of the year in which the failure of the life or lives might happen. 80. Hence we have this proportion. As the value of a perpetuity increased by unit is to L.l, so is the present value of the reversion of a perpetual annuity of L.l, after the failure of any life or lives, to the present value of L.1, receivable at the end of the year in which such failure shall take place. 81. Therefore, if the value of an annuity of one pound on any life or lives be subtracted from that of the perpetuity, and the remainder be divided by the value of the perpetuity increased by unit, the quotient will be the value, in present money, of the assurance of one pound on the same life or lives. (63.) 82. Ex. 1. What is the present value of L.l to be re¬ ceived at the end of the year in which a life now 50 years of age may fail ? The rate of interest being 5 per cent., the value of the perpetuity is 20 years’ purchase, and that of the life 11-66; , , , . 20—11-66 8-34 T norvt^,0 the answer therefore is ■ 20 -f-1— =: ~2\ == L-.0-397143. And if the sum assured were L.1000, the present value of the assurance would be L.397. 2s. lOd. 83. When the term of a life-assurance exceeds one year, its whole value is hardly ever paid down at the time that the contract is entered into; but in the instrument (called a policy) whereby the assurance is effected, an equivalent annual premium is stipulated for, payable at the commence- 2 E 218 ANNUITIES. Popular ment of each year during the term, but subject to failure View. with the life or lives assured. 84. But by reasoning as in No. 74, it will be found that an annual premium, payable at the commencement of each year in the whole duration of the life or lives assured, will be worth one year’s purchase more than an annuity on them payable at the end of each year; and, conse¬ quently, that if the value in present money of an assur¬ ance on any life or lives be divided by the number of years’ purchase an annuity on the same life or lives is worth, increased by unit, the quotient will be the equiva¬ lent annual premium for the same assurance. 85. Ex. 2. Required the annual premium for the assur¬ ance of L.l on a life of 50 years of age. In No. 82 the single premium for that assurance was shown to be 0-397143, and the value of an annuity on the life is 11-66; therefore, by the preceding number, the re¬ quired annual premium will be —- = -0313699 for the assurance of L.l; and for the assurance of L.1000 it will be L.31. 7s. 5d. 86. Ex. 3. Let both the single payment in present money, and the equivalent annual premium, be required for the assurance of L.l, on the joint continuance of two lives of the respective ages of 45 and 50 years. The value of an annuity of L.l on the joint continu¬ ance of these two lives appears by Table VII. to be t ^ 20—9-737 10-263 . L.9-737, therefore ,■ ^— — L.0-488714 is 20+1 21 0-488714 the single premium, and — L.0-0455168 the equivalent annual one, for the assurance of L.l to be paid at the end of the year in which that life becomes extinct which may happen to fail the first of the two. Therefore, if the sum assured were L.500, the total present value of the assurance would be L.244. 7s. 2d., and the equivalent annual premium L.22. 15s. 2d. 87. Ex. 4. Let both the single and the equivalent an¬ nual premium be required for the assurance of L.l on the life of the survivor of two persons now aged 40 and 50 years respectively. The value of an annuity on the survivor of these two lives was shown in No. 66 to be 15-066, therefore, by No. 81, the single premium will be —^—— —gj— = L.0-234952; and, by No. 84. the annual one will be L.0-234952 16-066 “ ’ ’ 46~4~ That is, for the assurance of L.l to be received at the end of the year in which the last surviving life of the two becomes extinct. Therefore, for the assurance of L.5000, the single pre¬ mium will be L.l 174. 15s. 2d., the equivalent annual one L.73. 2s. 5d. 88. Ex. b. What should the single and equivalent an¬ nual premiums be for an assurance on the last survivor of three lives of the respective ages of 50, 55, and 60 years ? The value of an annuity on the last survivor of them was shown in No. 68 to be 14-001, the single premium , ,, r- u 20—14-001 5-999 should therefore be - ,— = —— L.0-285666, 20+1 L.0-285666 21 and the annual —^ ^— = L.0-0190431, for the assur¬ ance of L.l to be received at the end of the year in which the last surviving life of the three may fail. For the assurance of L.2000, the single premium would therefore be L.57L 6s. 8cL, the annual one L.38. Is. 9d. 89. Lemma. If an annuity be payable at the commence- Por, ment of each year which some assigned life or lives may vi| enter upon in a given term, the number of years’ purchased in its present value will exceed by unit the number of years’ purchase in the present value of an annuity on the same life or lives for one year less than the given term, but payable, as annuities generally are, at the end of each year. Demonstration. Since the proposed life or lives can only enter upon any year after the first by surviving the year that precedes it, the receipt of each of the pay¬ ments after the first, that are to be made at the com¬ mencement of the year, will take place at the same time and upon the same conditions as that of the rent for the year then expired of the life-annuity, for a term one year less than the term proposed : this last-mentioned annuity will therefore be worth, in present money, just the same number of years’ purchase as all the payments subsequent to the first which may be made at the commencements of the several years. And since the first of these is to be made immediate¬ ly, the present value of the whole of them will evidently exceed tlje number of years’ purchase last mentioned by unit; which was to be demonstrated. 90. If, while the rest remains the same, the payment of the annuity which depends upon the assigned life or lives entering upon any year is not to be made until the end of that year ; as the condition upon which every payment is to be made will remain the same, but each of them will be one year later; the only alteration in the value of the whole will arise from this increase in the remoteness of payment, by which it will be reduced in the ratio of L.l to the present value of L.l receivable at the end of a year. (2.) 91. When the value of an annuity on any proposed life or lives for an assigned term is given, it is evident that the value of an annuity on the same life or lives for one year less may be found, by subtracting from the given value the present value of the rent to be received upon the proposed life or lives surviving the term assigned. 92. Proposition. The present value of an assurance on any proposed life or lives for a given term is equal to the excess of the value of an annuity to be paid at the end of each year which the life or lives proposed may enter upon in the term, above the value of an annuity on them for the same term, but dependent, as usual, upon their surviving each year. Demonstration. If an annuity, payable at the end of each year which the proposed lifo or lives may enter up¬ on during the given term, be granted to P upon condition that he shall pay over what he receives to Q at the end of each year which the same life or lives may survive, it is manifest that there will only remain to P the rent for the year in which the proposed life or lives may fail; that is, the assurance of that sum thereon for the given term (77) ; which was to be demonstrated. 93. From the last four numbers (89-92) we derive the following Rule for determining the present value of an assurance on any life or lives for a given term. Add unit to the value of an annuity on the proposed life or lives for the given term, and from the sum subtract the present value of one pound to be received upon the same life or lives surviving the term ; multiply the remainder by the present value of L.l to be received certainly at the end of a year, and from the product subtract the present value of an annuity on the proposed life or lives for the term. This last remainder will be the value in. present money ANNUITIES. Bopwr of the assurance of L.l during the same term on the life Vie or lives proposed. sv-J 94 It has been shown above (34-39) how the pre¬ sent value of L.l receivable upon any single or joint lives surviving an assigned term may be found. And all that was demonstrated from No. 48 to 53 inclusive, respect¬ ing annuities on the last survivor of two or of three lives, or on the joint continuance of the two last survivors out of three lives, is equally true of any particular year’s rent of those annuities. Hence it is evident how the present value of L.l, to be received upon the last survivor of two or of three lives, or upon the last two survivors out of three lives, surviving any assigned term, may be found. 95. Example. Required the present value of L.l, to be received at the end of the year in which a life now forty-five years of age may fail, provided that such fail¬ ure happen before the expiration of ten years. Here the present value of L.l, to be received on the ‘ life surviving the term, will be found to be L.0528976, and the value of an annuity on the proposed life for the I term is 7-175. (70.) From 8-175 subtract 0-528976 the remainder 7-646024 being multiplied by 0-952381 produces 7-28193 from this subtract 7*17500 remains L.0-10693, the required va¬ lue of the assurance; and if the sum assured were L.3000, the value of the assurance in present money would be L.320. 15s. 7d. 96. By numbers 89, 91, and 95, it appears that an an¬ nuity, payable at the commencement of each of the next ten years that a life of 45 enters upon, is worth 7*646 0-10693 Xrt„lono„ years purchase; therefore, — L.0-013985 will be the annual premium for the assurance of L.l for ten years on that life. For the assurance of L.3000, it will therefore be L.41. 19s. Id. 97. When the term of the assurance is the whole duration of the life or lives assured, L.l to be received upon their surviving the term is worth nothing; and an annuity on the lives for the term is also for their whole duration. Therefore from No. 93 we derive the following Rule for determining the present value of an assurance on any life or lives. Add unit to the value of an annuity on the proposed life or lives ; multiply the sum by the present value of L.l to be received certainly at the end of a year, and from the product subtract the value of an annuity on the same life or lives. The remainder will be the value of the assurance in present money. 98. Example. Required the present value of L.l to be received at the end of the year in which the survivor of two lives may fail, their ages now being 40 and 50 years respectively. The value of an annuity on these lives was shown in No. 66 to be 15-066. Multiply 16-066 by 0-952381, from the product 15-3009 subtract 15-066, the remainder L.0-2349 is the required value, agreeably to No. 87. And, in all other cases, the values determined by the rule in the preceding number will be found to agree with those obtained by the method of No. 81. 99. When an assurance On any life or lives has been 219 effected at a constant annual premium, and kept up for Popular some time by the regular payment of that premium, the View, annual premium required for a new assurance of the same sum on the same life or lives will, on account of the in¬ crease of age, be greater than that at which the first as¬ surance was effected: Therefore the present value of all these greater annual premiums, that is, the total present value of the new assurance, will exceed the present value of all the premiums that may hereafter be received under the existing policy. And the excess will evidently be the value of the policy, supposing the life or lives to be still insurable ; that being the only advantage which can now be derived from the premiums already paid. So that if the present value of all the future annual premiums to be paid under an existing policy, for the as¬ surance of a certain sum upon any life or lives, be subtract¬ ed from the present value of the assurance of the same sum on the same life or lives, the remainder will be the value of the policy. 100. Example. L.1000 has been assured some years on a life now 50 years of age, for its whole duration, at the annual premium of L.20, one of which has just now been paid. What is the value of the policy ? The present value of the assurance of L.1000 on that life has been shown in No. 82 to be L.397. 2s. lOd.; and an annuity on the life being worth 11-66 years’ purchase (Table VI.), the present value of all the premiums to be paid in future under the existing policy is 11-66 X L.20 i= L.233. 4s.; the value of the policy, therefore, is L.163. 18s. lOd. Immediately before the payment of the premium the value of the policy would evidently have been less by the premium then due. 101. In our investigations of the values of annuities on lives, we have hitherto assumed that no part of the rent is to be received for the year in which the life wherewith the annuity may terminate fails. But if a part of the annuity is to be received at the end of that year, proportional to the part of the year which may have elapsed at the time of such failure; as, in a great number of such cases, some of the lives wherewith the annuity terminates will fail in every part of the year, and as many, or very nearly so, in any one part of it as in any other: we may assume that, upon an average, half a year’s rent will be received at the end of the year in which such failure happens; and therefore, that by the title to what may be received after the failure of the life or lives where¬ on the annuity depends, the present value of that annuity will be increased by the present value of the assurance of half a year’s rent on the same life or lives. 102. Thus, for example, the present value of the as¬ surance of L.l on a life of 50 years of age having in No. 82 been shown to be L.0-397143, the value of an annuity of L.l on that life, when payable till the last moment of its existence, will exceed L.l 1-66, its value if only pay¬ able until the expiration of the last year it survives, by (L0'397143 \ ^ L.0-199; it will therefore be L.ll-859. 103. If at the end of the year in which an assigned life A may fail, Q or his heirs are to receive L.l, and are at the same time to enter upon an annuity of L.l, to be en¬ joyed during another life P, to be then fixed upon; the present value of (fs interest will evidently be the same as that of the assurance on the life of A, of a number of pounds, exceeding by unit the number of years’ purchase in the value of an annuity on the life of P, at the time of nomination. 104. What is the present value of the next presentation to a living of the clear annual value of L.500, A, the pre¬ sent incumbent, being now 50 years of age; supposing 220 annuities:. Popular the age of the clerk presented to be 25 at the end of the View, year in which the present incumbent dies; also, that he takes the whole produce of the living for that year ? By Table VI. it will be found that the value of an an¬ nuity of L.l on a life of 25 is L.15'303; and in No. 82 it has been shown, that the present value of the assurance of L.l on a life of 50 is L.0‘397143. Hence, and by the last number, it appears that if the annual produce of the living were but L.l, the present value of the next presen¬ tation would be L.16'303 X 0-397143. L.6'47467. Ihe required value, therefore, is L.3237. 6s. 9d. 105. If to the value of the succeeding life, determined according to No. 103, the value of the present be added, the sum of these will evidently be the present value of both the lives in succession; and, in the case of the pre¬ ceding number, will be 6’475 -f- 11‘66 — 18,135 years’ purchase. 106. In No. 103 we proceeded upon the supposition that the annuity on the present life is only payable up to the expiration of the last year it survives, and, conse¬ quently, that the succeeding life takes the whole rent for the year in which the present fails. But if the succeeding life is only to take a part of that rent, in the same proportion to the whole as the portion of the year which intervenes between the failure of the present life and the end of the year is to the whole year, p™ then, by reasoning as in No. 101, it will be found that the % portion of that rent which the succeeding life will received may be properly assumed to be one half. And, in* stead of increasing the number of years’ purchase the an¬ nuity on the succeeding life will be worth at the end of the year in which the other fails by unit, w'e must only add one half to that number, in order that the present value of the assurance of the sum on the existing life may be the number of years’ purchase which all that may be received during the succeeding life is now worth. 107. The value of the succeeding life, in the case of No. 104, will, upon this hypothesis, be 15-803 X 0-397143 — 6-27605 years’ purchase. And this appears to be the most correct way of calcu¬ lating the value of an annuity on a succeeding life, al¬ though that of No. 103 proceeds upon the principle on which life-interests are generally valued. 108. But the value of two lives in succession will be the same on both hypotheses ; the rent for the year in which the.first may fail being, in the one case, given entirely to the successor; and, in the other, divided equally between the two. This is also true of any greater number of successive lives. PART II. 109. We now proceed to treat the subject of annuities Algebraically. are to enjoy during the first n years, and B and his heirs or assigns for ever after. Since the value of the perpe¬ tuity to be entered upon immediately has just been shown I.—On Annuities-Certain. to be -, the present value of B's share, that is, the present Let r denote the simple interest of L.l for one year, s, any sum put out at interest, w, the number of years for which it is lent.. m, its amount in that time. «, an annuity for the same time. (3 and 4.) m, the amount to which that annuity will increase when each payment is laid up as it becomes due, and improved at compound interest until the end of the term. v, the present value of the same annuity. (6.) 110. Then, reasoning as in the first number of this ar¬ ticle, it will be found that w = s (1 + r)n. And by No. 2 it appears that the present value of s pounds to be re¬ ceived certainly at the expiration of n years, is s -y^ ^ or s(l + r)-n. 111. The amount of L.l in n years being (1 + r)n, its increase in that time will be (1 + r)” — 1; and when it value of the same perpetuity when the entrance thereon is deferred until the expiration n years, will be - (/• +1)-”) (110) ; and the value of the share of A will be thus much less than that of the whole perpetuity (21), therefore equal to^ |^1 —(1 + 7’)_,l^j = v, the present value of an annuity of a pounds for the term of n years certain. 114. If the annuity is not to be entered upon until the expiration of d years, but is then to continue n years, its value at the time of entering upon it will be ^j^l —(1 + as has just been shown ; therefore its present value will be ^[a+’-r (!+'■)' ^rzV. (110.) is considered that this increase arises entirely from the simple interest (r) of L.l being laid up at the end of each year, and improved at compound interest during the re¬ mainder of the term, it must be obvious that (1 + r)n — 1 is the amount of an annuity of r pounds in that time ;#but 115. In the same manner it appears that, when the entrance on a perpetuity of a pounds is deferred d years, its present value will be -(1 + rf^. (110 and 112.) :: (1 + r)n — 1 : -j^(l + r)n — l,~j which, there¬ fore, is equal to m, the amount of an annuity of a pounds in n years. 112. Reasoning as in No. 8, it will be found, that since r : \ a : the present value of a perpetual annuity of 116. q being any number whatever, whole, fractionator mixed, let "Kq denote its logarithm, and xq the arithmeti¬ cal complement of that logarithm ; so that these equations may obtain, ^ = xq. Then, for the resolution of the principal questions of this kind by logarithms, we shall have these formulae. a pounds is -. 1. Amount of a stem improved at interest. Xm = wX (1 + r) + Xs. (110.) 113. If two persons, A and B, purchase a perpetuity of a pounds between them, which A and his heirs or assigns 2. Amount of an annuity when each payment is laid up us it becomes due, and improved at interest until the expi¬ ration of the term. rebraid Views 'S/I'V'' 3. X^X[(1 + r)n — 1~J + +x; Value of a lease or an annuity. Xv=7.[l-(l + r)^+Xa + ANN UI T IE S. (in.) (113.) 4 Value of a deferred annuity, or the renewal of any num¬ ber of years lapsed in the. term of a lease. *v = x[(l + r)-d—(\ + r)-<<* + "f]+Xa+xr. (114.) 5. Value of a deferred perpetuity, or the reversion of an estate in fee-simple, after an assigned term. Xv z= ?.a + xr + cfo (1 + r). (115.) By means of each of these equations, it is manifest that any one of the quantities involved in it may be found when the rest are given, 117. If the interest be convertible into principal v times in the year, at v equal intervals; since the interest of L.l for one of these intervals will be - (109), and the number of conversions of interest into principal in n years m ; to adapt the formula in No. 110 to this case, we have only to substitute - for r, and m for n, in the equation = s(l + r)n there given, whereby it will be transform- / tA m ed to this, m = s (1 + -J . 118. According as v is equal to 1, 2, 4, or is infinite, that is, according as the interest is convertible into prin¬ cipal yearly, half-yearly, quarterly, or continually, let rn be equal to y, h, q, or c ; so shall Y= s(l + r)n, n=s(\ + rT, «=»Kr. and c ~ s. n ; n being the number of which nr is the hyperbolic lo¬ garithm, and nr x 0*43429448 its logarithm in Briggs’ System, and the common tables. 119. From No. 117 and 110, it follows that the present value of s pounds to be received at the end of n years, when the interest is convertible into principal at v equal # # # / yri intervals in each year, is s fl + - J 120. When the present values and the amounts of an¬ nuities are desired, let the interest be convertible into principal at v equal intervals in the year, while the annuity is payable at w intervals therein, the amount of each pay¬ ment being ft 121. Case 1. y, being any whole number not greater than v, let - ^ so that the interest may be convertible into principal [i times in each of the intervals between the payments of the annuity. Then will the amount of L.l at the expiration of the Peri°d -be ^1 (117), and the interest of L.l for the same time will be 1J whence the present va- 1 - a lue of the perpetuity will be —- (8), and the va¬ lue of the same deferred n years, will be - (119), therefore the present value of the annuity to be en¬ tered upon immediately, and to continue n years, will be 1 * K)'-' 122, Case 2. y, being any whole number greater than t, let ~ so that the annuity may be payable y* times in each of the intervals between the payments of interest, or the conversion thereof into principal. Then, at the expiration of the -th of a year, when the interest on the purchase-money is first payable or contro¬ vertible, the interest on all the p—1 payments of the an¬ nuity previously made will be + 2) + 0a—3)+ + 3+ 2+l^j ; to which adding the /x payments of ^ each (including the one only then due), the sum ^ j^/M. 4~^n*:eres^ value of the perpetuity should yield at the expiration of each vth part of a year, in order to supply the deficiency (both of principal and interest) that would be occasioned during each of those periods, in any fund out of which the several payments of the annuity might be taken, as they re • i , , , . r aY~ . ryj(y,—1)“ spectively became due; and since^•r ] :£iL+re(|=±n= = l, the values of v given in the two preceding cases, will be found to coincide with this. 124. According as v and tt are each equal to 1, 2, 4, or are infinite; that is, according as the interest and the an¬ nuity are each payable yearly, half-yearly, quarterly, or continually, let v be equal to y, h, g, or c, then will ]= V. (i+rrMJ> 222 ANNUITIES. Algebraical View. h=a- r and c—~ r ■(1 + i)' i-ci+zr4” being as in No. 118. nuity, we have (>+;)' • ('+;) ■=M. (117, 1 (> +;)’”V="Q+ - [(>+;)’”- ■] —M. (117, 122, and 125.) And, in the 3c? Case, when the annuity is always pay¬ able at the same time that the interest is convertible, (r\v» nr r vn ~\ 1 + H v — ^I (1 + -) —1 —m.(117, 123, and 125.) 127. According as v and or are each equal to 1, 2,4, or are infinite ; that is, according as the interest and the annuity are each payable yearly, half-yearly, quarterly, or conti¬ nually, let m be denoted by y', h', q', or c'; then will y'= h' — - r a q=- r 0 + r)”-l _(1+; ’ V and c = - (n —1 ); n being as in No. 118. 128. Example 1. What will L.320 amount to, when im¬ proved at compound interest during 40 years, the rate of interest being 4 per cent, per annum ? By the first formula in No. 116, the operation will be as follows: 1 X r= 1-04X0-01703334 + » = 40 (1 + r)" AO-6813336 s = 320 A 2-5051500 m = 1536-327 A 3.1864836 And the answer is L.1536. 6s. 6^d. 129. Ex. 2. If the interest were convertible into princi¬ pal every half-year, the operation, according to No. 117, would be thus: 1 + J= 1-02 A 0-00860017 A X m— 80 ■ fi O-y 0-6880136 5 = 320 A 2-5051500 125. The amount of an annuity is equal to the sum to which the purchase-money would amount if it were put out and improved at interest during the whole term. For, from the time of the purchase of the annuity, what¬ ever part of the money that was paid for it may be in the hands of the granter, he must improve thus to provide for the payments thereof; and if the purchaser also improve in the same manner all he receives, the original purchase- money must evidently receive the same improvement du¬ ring the term, as if it had been laid up at interest at its commencement. 126. The periods of conversion of interest into princi¬ pal, and of the payment of the annuity being still desig¬ nated as in No. 120; since in n years the number of periods of conversion will be m, in the Is* Case, where the interest is convertible y. times in each of the intervals between the payments of the an- (i+T—i a \ v) m = 1560-14 A 3-1931636 So that in this case the amount would be L.1560. 2s.91d. 130. Ex. 3. Required the present value of an annuity of L.250 for 30 years, reckoning interest at 5 per cent. By the third formula in No. 116, the operation will be thus: _ A (1 -fr)-1 = x 1-05 = 1-9788107 Xra = 30 (1 + 7-)-”= -23137704 A 1-3643210 1_(1 _f_r)-» = -76862296 A 1-8857133 a = 250 A 2-3979400 r = -05 x 1-3010300 121, and 125.) In the 2rtf Case, when the annuity is pay¬ able y times, in each interval between the conversions of interest, [i— v =3843-114X3-5846833 And the required value is L.3843. 2s. 3^d. 131. Ex. 4. The rest being still the same, if the annuity in the last example be payable half-yearly, in the formula of No. 122 v will be equal to 1, t — 2, and /*, = 2; that formula will therefore become . |^1—(1+r)“"^J = v; and the operation will be thus : 1 — (1+ 7-)“” aT-88571331 n 130 a = 250 A 2-3979400] 1 0‘ -+t = 20-25 A 1-3064250 v = 3891-15 A 3-5900783 The value of the annuity will therefore in this case be L.3891. 3s. . * 132. Ex. 5. To what sum will an annuity of L.120 for 20 years amount, when each payment is improved at com¬ pound interest from the time of its becoming due until the expiration of the term, the rate of interest being 6 per cent. ? The operation by the second formula in No. 116 is thus: l+r= 1-06 A 0-025305865 X 77 = 20 (1 + 7-)” = 3-207135 A 0-5061173 (1 + /•)” — 1 = 2-207135 A 0-3438289 a= 120X2-0791812 r= -06 x 1-2218487 M = 4414-27 A 3-6448588 And the amount required is L.4414. 5s. 5d. 133. Ex. 6. The rest being the same as in the last ex¬ ample ; if both the interest and the annuity be payable half-yearly, the amount will be determined by the second of the formulae given in No. 127, which in this case will become-^|^(1-OS)40—l"l, and the operation will be as follows: 1-03 AO-01283723 X 40 (1-03)40 = 3-26204 A 0-5134892 (1-03)40—1=2-26204 A0-3545003 A 2-0791812 •06x1-2218487 m = 4524-08 A 3-6555302 So that the amount in this case would be L.4524. Is. 7*d. ebrai l ,'iew ANNUITIES. II.—On the Probabilities of Life. 134. Any persons, A, B, C, &c. being proposed, let the numbers which tables of mortality (32) adapted to them represent to attain to their respective ages be denoted by the symbols a, b, c, &c.; while lives n years older than those respectively are denoted thus, WA, nB, nC, &c. and the numbers that attain to their ages, by the symbols na, ”b, nc, &c.; also let lives w years younger than B, C, &c. be denoted thus, An, Bn, Cn, &c., while the numbers which the tables show to attain to those younger ages are designated by the symbols an, bn, cn, &c. Then, if ^4 be 21 years of age, and we use the Carlisle table, we shall have a — 6047, and 14a == 5362, the num¬ ber that attain to the age of 35, or that live to be 14 years older than A. Hence the number that are represented by the table to die in n years from the age of A will be a — na, that is, in 14 years, a — 14a ; and, by the Carlisle table, in 14 years from the age of 21, that is, between 21 and 35, it will be 6047 — 5362 = 685. 135. Problem. To determine the probability that a pro¬ posed life A will survive n years. Solution, a being the number of lives in the table of mortality attaining the age of that which is proposed, conceive that number of lives to be so selected (of which A must be one), that they may each have the same pro¬ spect with regard to longevity as the proposed life and those in the table, or the average of those from which it was constructed; then will the number of them that sur¬ vive the term be na. (134.) So that the number of ways all equally probable, or the number of equal chances for the happening of the event in question, is na ; and the whole number for its either happening or failing is a ; therefore, according to the first principles of the doctrine of probabilities, the probability of the event happening, that is, of A surviving the term, . na is —. a If the age of A be 14, the probability of that life sur¬ viving 7 years, or the age of 21, will, according to the „ , ,, , 70 6047 Carlisle table, be — = or 0’95454. a 6335 136. Since the number that die in n years from the age of J is a — na (134), it appears, in the same manner, that d the probability of that life failing in w years will be —— na . : . \ '7 = 1 — ; which probability, when the life, term, and table of mortality are the same as in the last No., will be 0-04546. 137. If two lives A and B be proposed, since the pro- n(Z bability of A surviving n years will be —, and that of B nb a nb n(ab) .. r- or ■ / will be the a b ab surviving the same term will be it appears from the doctrine of probabilities that measure of the probability that these lives will both sur¬ vive n years. In the same manner it may be shown, that the proba¬ bility of the three lives A> B, and C all surviving n years, ... , rif. nU nr n1)r\ will be measured by — • —r • — or -—j-A. And, universal- J a b c, abc ly, that any number of lives A, B, C, &c. will jointly sur¬ vive n years, the probability is ^ abc, &c. 138. Let „ _ 7 ”c „ c i i .Algebraical — — nb, — — nc, &c.; also let vipw. "a a o " c n(abc &c.) abc Ac ~ ::::: &C’) ’ 80 t^iat ^le probabilities of A, B, C, &c. surviving n years may be denoted by na, nb, nc, &c. respectively; and that of all these lives jointly sur¬ viving that term by n(abc, &c.) Then will the probability that none of these lives will survive n years be (1 — na) . (1 — „&).(! — nc) . &c. 139. But the probability that some one or more of these lives will survive n years will be just what the pro¬ bability last mentioned is deficient of certainty: its mea¬ sure, therefore, being just what the measure of that pro¬ bability is deficient of unit, will be 1 — (1 — ««) • (1 — J>): (1 — nc) • Ac. 140. Carol. 1. When there is only one life A, this will be na. 141. Carol. 2. When there are two lives A and B, it becomes na + — n(ab)- 142. Carol. 3. When there are three lives A, B, and C, it becomes na + n5 + nc — n(«5) — n(ac) — »(^c) + n(abc). 143. When three lives A, B, and C are proposed, that at the expiration of n years there will be living dead the probability is ABC none AB C AC B BC A And the sum of these four n(«5) + n(ac) + n(^c) 2n(abc), is the measure of the probability that some two at the least out of these three lives will survive the term. + n(abc) .(ab) .(\ — nc)— n(ab) — n(abc) i(ac) . (1 —nb) = n(ac) — n(abc) .(be) .(I — na) = n(bc) — n(abc) III.—Of Annuities on Lives. 144. Let the number of years’ purchase that an an¬ nuity on the life of A is worth, that is, the present value of L.l to be received at the end of every year during the continuance of that life, be denoted by a ; while the pre¬ sent value of an annuity on any number of joint lives A, By C, &c. that is, of an annuity which is to continue du¬ ring the joint existence of all the lives, but to cease with the first that fails, is denoted by abc, &c. Then will the value of an annuity-on the joint conti¬ nuance of the three lives A, B, and C, be denoted by abc. And on the joint continuance of the two A and B, by ab. 145. Also let fA and Af denote the values of annuities on lives respectively older and younger than A, by £ years; while ‘(abc, &c.) designates the value of an an¬ nuity on the joint continuance of lives t years older than A, B, C, &c. respectively; and (abc, &c.)f that of an annuity on the same number of joint lives, as many years younger than these respectively. 146. Let r—r—> the present value of L.l to be received 1+7- certainly at the expiration of a year, be denoted by «?. Then will vn be the present value of that sum certain to be received at the expiration of n years. But if its receipt at the end of that time be dependent upon an assigned life A surviving the term, its present value will, by that condition, be reduced in the ratio of certainty to the probability of A surviving the term, that is, in the ratio of unit to na, and will therefore be navn. In the same manner it appears, that if the receipt of the money at the expiration of the term be dependent upon any assigned lives, as A, B, C, &c. jointly surviving that period, its present value will be n(abc, &c.)vn. 224 ANNUITIES. Algebraical 147. Let us denote the sum of any series, as ^1^+ View. Q(abc)v'2 + 5(abc)v3 + &c. thus, Sn(abc)vn, by prefixing the Italic capital S to the general term thereof. Then, from what has just been advanced, it will be evident that ABC, &c. = Sn(abc, &c.)vn. , „ , . , When there are but three lives, A, B, and C, this be¬ comes abc = Sn(abc)vn. When there are but two, A and B, it becomes ab = Sn(ab)vn. . And in the same manner it appears, that tor a single life A, a — jSnavn. 148. n(abc, &C')vn = (138)» where the de_ nominator (abc, &c.) is constant, while the numerator va¬ ries with the variable exponent n. And the most obvious method of finding the value of an annuity on any assigned single or joint lives, is to calculate the numerical value of the term n(abc, &c.)vn for each value of n, and then to divide Sn(abc9 the sum of all these values by abc, &c.; for — = Sn(abc, &c.)vn = abc, &c. In calculating a table of the values of annuities on lives in that manner, for every combination of joint lives it would be necessary to calculate the term n(abc, &c.)vn for as many years as there might be between the age of the oldest life involved and the oldest in the table; and the same number of the terms navn for any single life of the same age. But this labour may be greatly abridged as follows: ning with the oldest in the table, and proceeding regular-Alg ly age by age to the youngest. Also a table of the values of any number of joint lives, the lives in each succeeding combination, in any one se ries of operations (according to the retrograde order of the ages in which they are computed), being one year younger respectively than those in the preceding combi¬ nation. And, if a table of single lives be computed first, then of two joint lives, next of three joint lives, and so on, the calculations made for the preceding tables will be of great use for the succeeding. 155. Having shown how to compute tables of the values of annuities on single and joint lives, we shall, in what follows, always suppose those values to be given. 156. Let the value of an annuity on the joint continu¬ ance of any number of lives, A, B, C, &c. that is not to be entered upon until the expiration of t years, be denot¬ ed by —|t(ABC, &c.). Then, if it were certain that all the lives would survive the term, since the value of the annuity at the expiration of the term would be ‘(abc, &c.) (145), its present value would be . ‘(abc, &c.) (146). But the measure of the probability that all the lives will survive the term is t(abc, &c.), therefore —j*(abc, &c.) =; t(abc, &c.) v*. ‘(abc, &c.). In the same manner, it appears, that for a single life ^4, ■, a ,— ^av^n ‘a. ■Ale A Prob. I. 149. Given '(abc, &c.), the value of an annuity on any number of joint lives, to determine abc, &c. that of an annuity on the same number of joint lives respectively one year younger than they. Solution. 157. Let an annuity for the term of £ years only, depen¬ dent upon the joint continuance of any number of lives, A, B, C, &c. be denoted by —(abc, &c.) ; and, since this If it were certain that the lives abc, &c. would all sur¬ vive one year, the proprietor of an annuity of L.l depen¬ dent upon their joint continuance would, at the expira¬ tion of a year, be in possession of L.l (the first year’s rent), and an annuity on the same number of lives one year older respectively than abc, &c. ; therefore, in that case, the required present value of the annuity would be r?[l +'(abc, &c.)]. (146.) But the probability of the lives A, B, C, &c. jointly surviving one year is less than certainty in the ratio of t(abc, &c.) to unit; therefore abc, &c. = ^abc, &c.) v[l + '(abc, &c.)]. 150. Corol. 1. When there are but three lives, A, B, and C, this becomes abc — t(abc)v\\ + '(abc)]. 151. Corol. 2. When there are only two, A and B, ab == /(ai)tj[l + '(AB)]* 152. Corol. 3. And for a single life A, it appears, in the same manner, that a =■ tav(\ + 'a). 153. Hence, in logarithms, we have these equations, 'Ka~'Kv Xta + X(1 + 'a), X ab = Xr + X/z + X> + X [1 + '(ab)], X abc = Xv + X/«-j-X/5 + X/c + X[l + '(abc)], &c. &c. &c. Upon which it may be observed, that X v + X/z, the sum of the first two logarithms that are employed in de¬ termining a from 'a, also enters the operation whereby ab is determined from '(ab). And that X v + X;a + XA the sum of the first three logarithms that serve to deter¬ mine ab from '(ab), is also required to determine abc from '(abc) ; which observation may be extended in a si¬ milar manner to any greater number of joint lives. 154. By these means it is easy to complete a table of the values of annuities on single lives of all ages, begin- temporary annuity, together with an annuity on the joint continuance of the same lives deferred for the same term, will evidently be of the same value as an annuity to be entered upon immediately, and enjoyed during their whole joint continuance, we have —;(abc, &c.) + ^ (abc, &c.) = abc, &c.; whence, — ,(abc, &c.) = ABC, &C. 1 (ABC, &C.). And for a single life A, —A = A if A# 158. Prob. II. To determine the present value of an annuity on the survivor of the two lives A and B (155) ; which we designate thus, ab. Solution. The probability that the survivor of these two lives will outlive the term of n years was shown in No. 141 to be Ma + — „(«&); therefore, reasoning as in No. 146, it will be found that the present value of the nth. year’s rent of this annuity is [wa +5 — „(«&)]v", and the value of all the rents thereof will be + —n(a or Snavn -(- Snbvn — iSn(ab) vn; so that ab = A -{- b — ab (147), agreeably to No. 48. Prob. III. 159. To determine the present value of an annuity on the last survivor of three lives, A,B, and C (155); which we denote thus, abc. Solution. ■' The present value of the nth year’s rent is [n« + + „c — n(ab) —- n(ac) — n(bc) + n(afc)>B (142. and 146); whence, it appears, as in the preceding number, that abc = a+ b + c-t-ab-— ac — bc + abc, agreeably to No, 52. - . . ANNUITIES. i»l Prob. IV. 160. To determine the present value of an annuity on ie joint existence of the last two survivors out of three J t 2 lives, A, B, C, (155); which we denote thus, abc. Solution. The present value of the nth year’s rent is \_n(ab) + n(ac) + n(bc) — 2n{abc)~\vn (143 and 146); whence, reasoning as in the two preceding numbers, we 2 ml jnferj that = ab + ac + bc — 2 abc, as was de- ® monstrated otherwise in No. 51. 161. Since the solutions of the last three problems were all obtained by showing each year’s rent (as for instance the nth) of the annuity in question to be of the same value with the aggregate of the rents for the same year of all the annuities (taken with their proper signs) on the single and joint lives exhibited in the resulting formula ; if any term of years be assigned, it is manifest that the value of such annuity for the term must be the same as that of the aggregate of the annuities above mentioned, each for the same term. Prob. V. 162. A and B being any two proposed lives now both existing, to determine the present value of an annuity re¬ ceivable only while A survives B. Solution. A rent of this annuity will only be payable at the end of the nth year, provided that B be then dead and A living; but the probability of B being then dead is 1 — nb, and that of ^4 being then living na, and these two events are independent; therefore, the probability of their both happening, or that of the rent being received, is (1 — nb) nat=.na — n(ab) ; the present value of that rent is, there¬ fore, [na — n(aby\vn; whence it follows, that the re¬ quired value of the annuity on the life of A after that of B is a — ab, agreeably to No. 60. 163. If the payment for the annuity which was the sub¬ ject of the last problem is not to be made in present money, but by a constant annual premium p at the end of each year, while both the lives survive ; since ab is the m I number of years’ purchase (6) that an annuity on the joint continuance of those lives is worth, the value of p will be determined by this equation, p . ab = A — ab, whence we have p r= 1. AB 164. But if one premium p is to be paid down now, and an equal premium at the end of each year while both the lives survive, we shall have jt?. (1 + ab) = a — ab, and a — ab P ~ ThFab* 165. For numerical examples illustrative of the formulae given from No. 158 to the present, see Nos. 66-74. Prob. VI. 166. A and B are in possession of an annuity on the life of the survivor of them, which, if either of them die before a third person C, is then to be divided equally be¬ tween C and the survivor during their joint lives ; to de¬ termine the value of (Js interest. That at the end of the «th year there will be dead A B living BC AC VOL. III. na) . n(bc) = A ln(bc) —Jobe)] ) = | [»(«c) — n(abc)1; H1 2 (1 —— nb) . n(pc and the sum of these being n(ac) + 1 n(bc) - the value of C’s interest is ac + ^ bc — abc. Prob. VII. 167. An annuity, after the decease of A, is to be equal¬ ly divided between B and C during their joint lives, and is then to go entirely to the last survivor for his life ; it is proposed to find the value of B's interest therein. Solution. That at the end of the nth year there will be 225 n(abc), Algebraical View. dead A AC living BC B the probability, multiplied by li's proportion of the annuity in that circumstance, is of these being nb — of B's interest is b A" 2m(^c) \n(abc) - n(ab) — n{b() + n{ahc); and the sum n(ab) — \n(bc) + \n(abc\ the value — AB ^ BC +1 ABC. Prob. VIII. 168. A, B, and C purchase an annuity on the life of the last survivor of them, which is to be divided equally at the end of every year among such of them as may then be living; what should A contribute towards the purchase of this annuity ? Solution. That at the end of n years there will be dead none C B BC livin;; ABC AB AC A the probability, multiplied by A's proportion of the annuity in that circumstance, is -j- }^n(cibc) ...+ i„(«6) —%n(abc) + i«(«c) — \n(abc) xa—n{ab)—n(ac) -f- Jahc) ; and the sum of these being na — \n(ab) — l„(ac) + ^ra(aic), the requir¬ ed value of A’s interest is a—^ ab — ^ AC-f-^ abc. Prob. IX. 169. As soon as any two of the three lives, A, B, and C, are extinct, Z) or his heirs are to enter upon an annui¬ ty, which they are to enjoy during the remainder of the survivor’s life; to determine the value of D’s interest therein. Solution. That at the end of n years there will be dead living The probability is Solution. the probability, multiplied by C's proportion of the annuity in that circumstance, is AB C nc—n(ac)—„(&c)+M(«&c) AC B nb—n(ab)—n(bc) + n(abc) BC A na—n(ab)-—n(«c) + ?1(«6c) ; and the sum of all these being na-\-nb-\-nc—2n(ab)—2n(ac)—2n(bc) ~[-3n(abc), the value of B’s interest is A + B -f- C. 2 AB 2 AC 2 BC-j-3 ABC. 170. The last four may be sufficient to show the me¬ thod of proceeding with any similar problems. 171. Let t(abc, &c.) denote the probability that the last m survivors out of m + lives A, B, C, &c. will jointly survive the term of t years. And when /i=0, the expres¬ sion will become t(abc, &c.), the probability that the lives will all survive the term. (138.) t(aic, &c.), the measure of 2 F When vn zz\ it will become 226 ANNUITIES. Algebraical the probability that the last survivor of them will outlive greatest joint continuance of any m of the proposed aw the term, which it will be better to write thus, t(abc, &c.), hves, according to the tables of mortality adapted to them, H retaining the vinculum, but omitting the unit over it, as (HI \ ™^ \abc. &c./— 0, in the notation of powers. -•abc, &c. in that case, Also let abc, &c. denote the value of an annuity on the joint continuance of the same number of last survivors out of the same lives. Then, if ^ be equal to 0, it will be and *|abc, &c. = abc, &c.; therefore, the general formula of No. 172 becomes — abc, &c.; that is, the excess of the value of an abc, &c. the value of an annuity on the joint continuance annuity certain for the whole term above that of an of all the lives; when «.= 1, it will be abcT&c. the value of on .the w',oIe duratlon of-ioint contin,la”ce of the an annuity on the last survivor of them. The values of an- as. I'l sl*rv'vl‘1S. l^s‘ » . , nuities on the last survivor of two and of three lives will ." nt ^ ’ y1 ie case P™P0Se. ^ be denoted as in Nos. 158 and 159 respectively; and that annuit^ certain a PerPetulty> as ln No. 173, the formula of an annuity on the joint continuance of the last two sur- 1 ———p— vivors out of three lives, as in No. 160. will become— abc, &c. the excess of the value of the Ihe value of an annuity on the last m survivors out of perpetuity above the value of an annuity on the ioint lives pco llTTPC. n rl 1 n rr oc if 1C limif/^rl fr\ f li 4-rwr-Y-% r-i-f r* i • i t ^ these w + ^u. lives, according as it is limited to the term of t years, or deferred during that term, will also be denoted by—abc, &c. or —,fABC, &c. (156 and 157.) Prob. X. 172. An annuity certain for the term of < + years is to be enjoyed by P and his heirs during the joint exist¬ ence of the last m survivors out of ^ lives, A, B, C, &c.; and if that joint existence fail before the expiration of t years, the annuity is to go to Q and his heirs for the remainder of the term ; to determine the value of Q’s in¬ terest in that annuity. Solution. of the last m survivors ; agreeably to No. 63. 176. Example 1. Required the present value of the ab¬ solute reversion of an estate in fee simple, after the ex¬ tinction of the last survivor of three lives, A, B, C, now aged 50, 55, and 60 years respectively; reckoning inter¬ est at 5 per cent. The general algebraical expression of this value has 1 just been shown to be - — abc. But - — 0-5 = 20-000 And abc, = 14-001 (68.) Qs expectation may be distinguished into two parts: Therefore 5-999 years’ purchase is the value required. And if the annual produce of the estate, l^, ThaTof enjoying thelnnuTty dudn^the term of C.lear °f Reductions, were L.100, the title to the rever- ® sion would now be worth L.599. 18s. —, agreeably to No. 76. 177. Ex. 2. An annuity for the term of 70 years cer¬ tain (from this time) is to revert to Q and his heirs at the failure of a life A, now 45 years of age ; what is the present value of Q’s interest therein, reckoning the inter¬ est of money at 5 per cent. ? In No. 174 the algebraical expression of the required value is shown to be ^ A. t years. 2d, That of enjoying it after the expiration of that term. The sum of the present values of the interests of P and Q together in the annuity for the term of t years, is manifestly equal to the whole present value of the annuity certain for that term ; that is, equal to 1 — (113 and 146); and the value of P’s interest for the term of t years 1S __abc, &c. (171) ; therefore the value of Q's in- But t.v terest for the same term is 1 — v* x 1-05 = 1-9788107 X 70 tABC, &C. The present value of the annuity certain for v years af¬ ter t years is v((l—vv) (114 and 146); and Q and his heirs will receive this annuity if the joint continuance of the last m survivors above mentioned fail before the piration of t years ; but the probability of their joint is 1—X «;70_ .^7° — ■ #70 032866X2-5167490 •967134 967134 •05 = 19-34268 Subtract A = 12-64800 (Tab. VI.) ex- con- tmuance failing in the term is 1—^abc, &c./; therefore the value of Q s interest in the annuity to be received after t years, is [‘ —j\abc, &c whole value of Qs interest is >] ?/(l—Vs) and the if, ,+t (m • ~ ™ - I v —- r'( 1 v”) • &e./| — —abc, 1/3. Carol. 1. When the whole annuity certain is a perpetuity, and the value of Q's interest is &c. remains 6-69468 years’ purchase; so that if the annuity were L.1000, the value of the rever¬ sion would be L.6694. 13s. 7d. 178. Ex. 3. An annuity for the term of 70 years cer¬ tain from this time is to revert to Q and his heirs at the extinction of the survivor of two lives, A and B, now aged 40 and 50 years respectively, the interest of money being 6 per cent.; it is required to determine the value of Q's interest in this annuity. The algebraical expression of the value is, 1—^70 r — ab (174 and 171). 174. Coral. 2. When the term But by the last example 1 — 1-70 cc. •s -f- 5 ,881 ami by No. 66, ab = t is not less than the so that the required value is = 19-34268 15-06600 4*27668 years' nfcaourchase; and if the annuity be L.1000, the present value ^ >f the reversion will be L.4276. 13s. 7d. IV.—Of Assurances on Lives. 179. Let the present value of the assurance (77 and ?8) of L.l on the life of A be denoted by the old English capital and that of an assurance on the joint continu- mce of any number of lives A, B, C, &c. by &3SC, &c. \l.so, let the value of an assurance on the joint continu- mce of any m of them out of the whole number m + /i ANNUITIES. 227 ™ I” ^ ™j Algebraical © &c. |_1—t(abc, &c.) v* A" j ABC, View. Cm ~| m m 1—((aic, &c.)v*_J—(1—v)—abc, &c.~ 1—t(abc, &c.)?/ • D —(i—• L_1 — t(abc, &c.)«t + —jAbc we have , &c. ^|, whence >e denoted by &UC, &c. 180. And, in every case, let us designate the annual premium (83) for an assurance, by prefixing the charac- er O to the symbol for the single premium; so that O ^ nay denote the annual premium for an assurance on the ife of A; ©^33C, &c. the same for an assurance on the oint continuance of all the lives, A, B, C, &c.; and 0 JJC, &c.= 1 — f(a£c, &c.) v* + 1—t(a6c, &c.)wf+—abc, &c. v— 1. 184. Corol. 1. When (i), the term of the assurance, is not less than the greatest possible joint duration of any m 3 &c. the annual premium for an assurance en the oint continuance of the last m survivors out of the whole tlumber »i + /4 of those lives. 181. Then will .—%. and O —— &33C, &c. and t\ t\ t\ of the proposed lives, t(abc, &c.) = 0, —^abc, &c. = abc, &c., and the general formulae of the two preceding numbers become respectively &c. and © __ & 23 C, &c. 3_t3St+ ABC \-v — I respec¬ tively. Where <(a6) z= f« + tb — t(ab) (141), and t(abc) — ta + tb + tc — Q(«&) + t(ac) + t(bc)~\ + t(abc). (142.) For the values of ab, abc, ab> —jAbc, see num¬ bers 157-159, and 161. 189. Corol. 6. When the assurance is on the joint con¬ tinuance of the two last survivors out of the three lives A, B, C, the formulae of No. 184 become respectively g 2 = v — (1 — v) abc’ 2 1 and © &33C — + v— 1. 1 + ABC Those of numbers 182 and 183, =*[i -f*) J -a—) -^Tc, _2 1 — ((abc^v* and O fBJC = *1 1 — tiabc^v* —tAbc — + V 1. in No. 81; - being the value of the perpetuity. (112.) 193. Examples of the determination of the single pre¬ miums for assurances, and of the derivation of the annual premiums from them, have been given in No. 82-88, also in 95 and 96; but by the algebraical formulae given here, the annual premiums may be determined directly, without first finding the total present values of the assur¬ ances. 194. Example 1. Required the annual premium for an assurance on the life +, now 50 years of age, interest 5 per cent. According to No. 185, the operation is thus, 1 + A = 12-660 A 4 _ [. 2-8975663 —= =-0789890 x ) 1 + A J adding v — -9523809, and subtracting unit, we have © & -0313699, agreeably to No. 85. 195. Ex. 2. What should the annual premium be for an assurance on the last survivor of three lives +, B, and C, now aged 50, 55, and 60 years respectively, rate of in¬ terest 5 per cent. ? Operation by No. 188. 1 + abc — 15-001 A 1 2-8238798 (68) rmc" 'o666622 v — -9523809 rh Where t(abc) = ^ab) + t(ac) + t(bc) — 2 t(abc\ (143.) V ' 2 _2 For the values of abc and abc 160, and 161. Lm ~j m 1— t{abc, &c.)^J —(1 —v) -tfAbc, value of an assurance on any life or lives for the term of t years, which was given in No. 182, may also be express¬ ed thus: ©S13SC = *0190431, agreeably to No. 88. 196. Ex. 3. Required the annual premium for an assur¬ ance for 10 years only, on a life now 45 years of age, in- see numbers 157, terest 5 per cent. Operation accordingjo No. 186. v10 = -613913 A 1-7881068 l0a z=. 4073 A 3-6099144 a = 4727 x 4-3254144 &c. the ! 0av10 = -528976 A 1-7234356 1 + 10A = 11-347 A 1-0548811 D + —(abc, &c.) — t(abc, &c.) v*~\v — ABC, &C. ^ I And this, in words at length, is the rule given in No. 93. 191. When t is not less than the greatest possible joint duration ot any m of the proposed lives, the last expres¬ sion becomes / \ m \1 + ABC, &X.J ^ ABC, &C. which is also equivalent to the first in No. 184; and, in words at length, is the rule given in No. 97 for determin¬ ing the value of an assurance on any life or lives for their whole duration. Subtract 6-002 A 0-7783167 from 1 + A 13-648 remains 7-646 x 1-1165657 1 — io^io — -471024 A1-6730430 •061604 A 2 7896087 v — -952381 © —| ^ — -013985, agreeably to No. 96. What has been advanced from number 99 to 109 needs no algebraical illustration. ANNUITIES, TABLE I. Showing the present Value of One Pound to be received at the End of any Number of Years not exceeding 50. (See No. 9 12 of the preceding Article.) Years. 2 per Cent. per Cent. 3 per Cent. 4 per Cent. 5 per Cent. 6 per Cent. 7 per Cent. 8 per Cent. 9 per Cent. Years. 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 43 44 45 46 47 48 49 50 ■980392 ■961169 ■942322 ■923845 ■905731 ■887971 ■870560 •853490 ■836755 •820348 ■804263 •788493 •773033 ■757875 •743015 •728446 •714163 •700159 •686431 •672971 •659776 •646839 •634156 •621721 •609531 •597579 *585862 •574375 •563112 •552071 •541246 •530633 •520229 •510028 •500028 •490223 •480611 •471187 •461948 •452890 •444010 •435304 •426769 •418401 •410197 •402154 •394268 •386538 •378958 •371528 975610 ■951814 ■928599 ■905951 •883854 •862297 •841265 •820747 •800728 •781198 •762145 •743556 •725420 •707727 •690466 •673625 •657195 •641166 •625528 •610271 •595386 •580865 •566697 •552875 •539391 •526235 •513400 •500878 •488661 •476743 •465115 •453771 •442703 •431905 •421371 •411094 •401067 •391285 •381741 •372431 •363347 •354485 •345839 •337404 •329174 •321146 •313313 •305671 •298216 •290942 •970874 ■942596 ■915142 •888487 ■862609 ■837484 •813092 •789409 •766417 •744094 •722421 •701380 •680951 •661118 •641862 •623167 •605016 •587395 •570286 •553676 •537549 •521893 •506692 •491934 •477606 •463695 •450189 •437077 •424346 •411987 •399987 •388337 •377026 •366045 •355383 •345032 •334983 •325226 •315754 •306557 •297628 •288959 •280543 •272372 •264439 •256737 •249259 •241999 •234950 •228107 ■961538 ■924556 •888996 ■854804 •821927 •790315 ■759918 ■730690 •702587 •675564 •649581 •624597 •600574 •577475 •555265 •533908 •513373 •493628 •474642 •456387 •438834 •421955 •405726 •390121 •375117 •360689 •346817 •333477 •320651 •308319 •296460 •285058 •274094 •263552 •253415 •243669 •234297 •225285 •216621 •208289 •200278 •192575 •185168 •178046 •171198 •164614 •158283 •152195 •146341 •140713 •952381 ■907029 ■863838 ■822702 ■783526 •746215 •710681 •676839 •644609 •613913 •584679 •556837 •530321 •505068 •481017 •458112 •436297 •415521 •395734 •376889 •358942 •341850 •325571 •310068 •295303 •281241 •267848 •255094 •242946 •231377 *220359 •209866 •199873 T90355 •181290 T72657 T64436 •156605 •149148 •142046 •135282 •128840 •122704 •116861 •111297 •105997 •100949 •096142 •091564 •087204 •943396 •889996 •839619 •792094 •747258 •704961 •665057 •627412 •591898 •558395 •526788 •496969 •468839 •442301 •417265 •393646 •371364 •350344 •330513 •311805 •294155 •277505 •261797 •246979 •232999 •219810 •207368 •195630 •184557 •174110 •164255 •154957 •146186 •137912 •130105 •122741 •115793 •109239 •103056 •097222 •091719 •086527 •081630 •077009 •072650 •068538 •064658 •060998 •057546 •054288 ■934579 •873439 •816298 •762895 •712986 ■666342 •622750 •582009 •543934 •508349 •475093 •444012 •414964 •387817 •362446 •338735 •316574 •295864 •276508 •258419 •241513 •225713 •210947 •197147 •184249 •172195 •160930 •150402 •140563 •131367 •122773 •114741 •107235 •100219 •093663 •087535 •081809 •076457 •071455 •066780 •062412 •058329 •054513 •050946 •047613 •044499 •041587 •038867 •036324 •033948 •925926 •857339 •793832 •735030 •680583 •630170 •583490 •540269 •500249 •463193 •428883 •397114 •367698 •340461 •315242 •291890 •270269 •250249 •231712 •214548 •198656 •183941 •170315 •157699 •146018 •135202 •125187 •115914 •107328 •099377 •092016 •085200 •078889 •073045 •067635 •062625 •057986 •053690 •049713 •046031 •042621 •039464 •036541 •033834 •031328 •029007 •026859 •024869 •023027 •021321 •917431 •841680 •772183 •708425 •649931 •596267 •547034 •501866 •460428 •422411 •387533 •355535 •326179 •299246 •274538 •251870 •231073 •211994 •194490 •178431 •163698 •150182 •137781 •126405 •115968 •106393 •097608 •089548 •082155 •075371 •069148 •063438 •058200 •053395 •048986 •044941 •041231 •037826 •034703 •031838 •029209 •026797 •024584 •022555 •020692 •018984 •017416 •015978 •014659 •013449 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 43 44 45 46 47 48 49 50 230 ANNUITIES, TABLE II. Showing the present Value of an Annuity of One Pound for any Number of Years not exceeding 50. No. 9 12. Years. 2 per Cent, per Cent. 3 per Cent. 4 per Cent. 5 per Cent. 6 per Cent 7 per Cent. 8 per Cent. 9 per Cent. Years 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 43 44 45 46 47 48 49 50 •9804 1- 9416 2- 8839 3- 8077 4- 7135 5- 6014 6- 4720 7- 3255 8- 1622 8- 9826 9- 7868 10- 5753 11- 3484 12- 1062 12- 8493 13- 5777 14- 2919 14- 9920 15- 6785 16- 3514 17- 0112 17- 6580 18- 2922 18- 9139 19- 5235 20- 1210 20- 7069 21- 2813 21- 8444 22- 3965 22- 9377 23- 4683 23- 9886 24- 4986 24- 9986 25- 4888 25- 9695 26- 4406 26- 9026 27- 3555 27- 7995 28- 2348 28-6616 29- 0800 29-4902 29- 8923 30- 2866 30- 6731 31- 0521 31-4236 •9756 1- 9274 2- 8560 3- 7620 4- 6458 5- 5081 6- 3494 7- 1701 7- 9709 8- 7521 9- 5142 10-2578 10- 9832 11- 6909 12- 3814 13- 0550 13- 7122 14- 3534 14- 9789 15- 5892 16- 1845 16- 7654 17- 3321 17- 8850 18- 4244 18- 9506 19- 4640 19- 9649 20- 4535 20- 9303 21- 3954 21- 8492 22- 2919 22- 7238 23- 1452 23-5563 23- 9573 24- 3486 24- 7303 25- 1028 25-4661 25- 8206 26- 1664 26-5038 26- 8330 27- 1542 27-4675 27- 7732 28- 0714 28-3623 Perp. •9709 1- 9135 2- 8286 3- 7171 4- 5797 5- 4172 6- 2303 7- 0197 7- 7861 8- 5302 9- 2526 9-9540 10- 6350 11- 2961 11- 9379 12- 5611 13- 1661 13- 7535 14- 3238 14- 8775 15- 4150 15- 9369 16- 4436 16- 9355 17- 4131 17- 8768 18- 3270 18- 7641 19- 1885 19- 6004 20- 0004 20-3888 20- 7658 21- 1318 21-4872 21- 8323 22- 1672 22- 4925 22-8082 23- 1148 23-4124 23-7014 23- 9819 24- 2543 24-5187 24- 7754 25- 0247 25-2667 25-5017 25-7298 •9615 1-8861 2- 7751 3- 6299 4- 4518 5- 2421 6- 0021 6- 7327 7- 4353 8- 1109 8- 7605 9- 3851 9-9856 10- 5631 11- 1184 11- 6523 12- 1657 12- 6593 13- 1339 13- 5903 14- 0292 14-4511 14- 8568 15- 2470 15-6221 15- 9828 16- 3296 16-6631 16- 9837 17- 2920 17-5885 17- 8736 18- 1476 18-4112 18-6646 18- 9083 19- 1426 19-3679 19-5845 19-7928 19- 9931 20- 1856 20-3708 20-5488 20-7200 20- 8847 21- 0429 21-1951 21-3415 21-4822 50-0000 40-0000 33-3333 25-0000 •9524 1- 8594 2- 7232 3- 5460 4- 3295 5- 0757 5- 7864 6- 4632 7- 1078 7- 7217 8- 3064 8- 8633 9- 3936 9-8986 10-3797 10- 8378 11- 2741 11- 6896 12- 0853 12- 4622 12-8212 13- 1630 13-4886 13- 7986 14- 0939 14-3752 14-6430 14- 8981 15- 1411 15-3725 15-5928 15- 8027 16- 0025 16-1929 16-3742 16-5469 16-7113 16- 8679 17- 0170 17-1591 17-2944 17-4232 17-5459 17-6628 17-7741 17-8801 17- 9810 18- 0772 18-1687 18-2559 20-0000 •9434 1- 8334 2- 6730 3- 4651 4- 2124 4- 9173 5- 5824 6- 2098 6- 8017 7- 3601 7- 8869 8- 3838 8- 8527 9- 2950 9-7122 10-1059 10-4773 10- 8276 11- 1581 11-4699 11- 7641 12- 0416 12-3034 12-5504 12- 7834 13- 0032 13-2105 13-4062 13-5907 13-7648 13- 9291 14- 0840 14-2302 14-3681 14-4982 14-6210 14-7368 14-8460 14- 9491 15- 0463 15-1380 15-2245 15-3062 15-3832 15-4558 15*5244 15-5890 15-6500 15-7076 15-7619 •9346 1-8080 2- 6243 3- 3872 4- 1002 4- 7665 5- 3893 5- 9713 6- 5152 7- 0236 7-4987 7- 9427 8- 3577 8- 7455 9- 1079 9-4466 9-7632 10-0591 10-3356 10-5940 10- 8355 11- 0612 11-2722 11-4693 11-6536 11-8258 11- 9867 12- 1371 12-2777 12-4090 12-5318 12-6466 12-7538 12-8540 12- 9477 13- 0352 13-1170 13-1935 13-2649 13-3317 13-3941 13-4524 13-5070 13-5579 13-6055 13-6500 13*6916 13-7305 13-7668 13-8007 16-6667 14-2857 •9259 1- 7833 2- 5771 3- 3121 3- 9927 4- 6229 5- 2064 5- 7466 6- 2469 6- 7101 7- 1390 7-5361 7- 9038 8- 2442 8-5595 8- 8514 9- 1216 9-3719 9-6036 9-8181 10-0168 10-2007 10-3711 10-5288 10-6748 10-8100 10- 9352 11- 0511 11-1584 11-2578 11-3498 11-4350 11-5139 11-5869 11-6546 11-7172 11-7752 11-8289 11-8786 11-9246 11- 9672 12- 0067 12-0432 12-0771 12-1084 12-1374 12-1643 12-1891 12-2122 12-2335 12-5000 •9174 1- 7591 2- 5313 3- 2397 3- 8897 4- 4859 5- 0330 5-5348 5- 9952 6- 4177 6- 8052 7- 1607 7-4869 7- 7862 8- 0607 8-3126 8-5436 8-7556 8- 9501 9- 1285 9-2922 9-4424 9-5802 9-7066 9-8226 9-9290 10-0266 10-1161 10-1983 10-2737 10-3428 10-4062 10-4644 10-5178 10-5668 10-6118 10-6530 10-6908 10-7255 10-7574 10-7866 10-8134 10-8380 10-8605 10-8812 10-9002 10-9176 10-9336 10-9482 10-9617 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 43 44 45 46 47 48 49 50 11-1111 Perp. ANNUITIES 231 TABLE III. Showing the Sum to which One Pound will increase when improved at Compound Interest during any . Number of Years not exceeding 50. (No. 9 12.) Years. 2 per Cent. 21- per Cent. 3 per Cent. 4 per Cent. 5 per Cent. 6 per Cent. 7 per Cent. 8 per Cent. Years. 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 43 44 45 46 47 48 49 50 1-02000 1-04040 1-06121 1-08243 1-10408 1-12616 1-14869 1-17166 1-19509 1-21899 1-24337 1-26824 1-29361 1-31948 1-34587 1-37279 1-40024 1-42825 1-45681 1-48595 1-51567 1-54598 1-57690 1-60844 1-64061 1-67342 1-70689 1-74102 1-77584 1-81136 1-84759 1-88454 1-92223 1-96068 1- 99989 2- 03989 2-08069 2-12230 2-16474 2-20804 2-25220 2-29724 2-34319 2-39005 2-43785 2-48661 2-53634 2-58707 2-63881 2-69159 1-02500 1-05063 1-07689 1-10381 1-13141 1-15969 1-18869 1-21840 1-24886 1-28008 1*31209 1-34489 1-37851 1-41297 1-44830 1-48451 1-52162 1-55966 1-59865 1-63862 1-67958 1-72157 1-76461 1-80873 1-85394 1-90029 1-94780 1- 99650 2- 04641 2-09757 2-15001 2*20376 2-25885 2-31532 2-37321 2-43254 2-49335 2-55568 2-61957 2-68506 2-75219 2-82100 2-89152 2- 96381 3- 03790 3-11385 3-19170 3-27149 3-35328 3-43711 1-030000 1-060900 1-092727 1-125509 1-159274 1-194052 1-229874 1-266770 1-304773 1-343916 1-384234 1-425761 1-468534 1-512590 1-557967 1-604706 1-652848 1-702433 1-753506 1-806111 1-860295 1-916103 1- 973587 2- 032794 2-093778 2-156591 2*221289 2-287928 2-356566 2-427262 2-500080 2-575083 2-652335 2-731905 2-813862 2-898278 2- 985227 3- 074783 3-167027 3-262038 3-359899 3-460696 3-564517 3-671452 3- 781596 3*895044 4- 011895 4-132252 4*256219 4-383906 1-040000 1-081600 1-124864 1-169859 1-216653 1-265319 1-315932 1-368569 1-423312 1-480244 1-539454 1-601032 1-665074 1-731676 1-800944 1-872981 1- 947901 2- 025817 2-106849 2-191123 2-278768 2-369919 2-464716 2-563304 2-665836 2-772470 2-883369 2- 998703 3- 118651 3-243398 3-373133 3*508059 3-648381 3-794316 3- 946089 4- 103933 4-268090 4-438813 4-616366 4-801021 4- 993061 5- 192784 5-4G0495 5-616515 5- 841176 6- 074823 6-317816 6-570528 6- 833349 7- 106683 1-050000 1-102500 1-157625 1-215506 1-276282 1-340096 1-407100- 1-477455 1-551328 1-628895 1-710339 1-795856 1-885649 1- 979932 2- 078928 2-182875 2-292018 2-406619 2-526950 2-653298 2-785963 2- 925261 3- 071524 3-225100 3-386355 3-555673 3-733456 3- 920129 4- 116136 4-321942 4-538039 4- 764941 5- 003189 5-253348 5-516015 5- 791816 6- 081407 6-385477 6- 704751 7- 039989 7-391988 7- 761588 8- 149667 8-557150 8- 985008 9- 434258 9-905971 10-401270 10- 921333 11- 467400 1-060000 1-123600 1-191016 1-262477 1-338226 1-418519 1-503630 1-593848 1-689479 1-790848 1- 898299 2- 012196 2-132928 2-260904 2-396558 2-540352 2-692773 2- 854339 3- 025600 3-207135 3-399564 3-603537 3- 819750 4- 048935 4-291871 4-549383 4- 822346 5- 111687 5-418388 5- 743491 6- 088101 6-453387 6- 840590 7- 251025 7- 686087 8- 147252 8- 636087 9- 154252 9-703507 10-285718 10- 902861 11- 557033 12- 250455 12- 985482 13- 764611 14- 590487 15- 465917 16- 393872 17- 377504 18- 420154 1-070000 1-144900 1-225043 1-310796 1-402552 1-500730 1-605781 1-718186 1-838459 1- 967151 2- 104852 2-252192 2-409845 2-578534 2-759032 2- 952164 3- 158815 3-379932 3-616528 3- 869684 4- 140562 4-430402 4- 740530 5- 072367 5-427433 5- 807353 6- 213868 6- 648838 7- 114257 7- 612255 8- 145113 8- 715271 9- 325340 9-978114 10- 676581 11- 423942 12- 223618 13- 079271 13- 994820 14- 974458 16- 022670 17- 144257 18- 344355 19- 628460 21- 002452 22- 472623 24- 045707 25- 728907 27-529930 29-457025 1-080000 1-166400 1-259712 1-360489 1-469328 1-586874 1-713824 1-850930 1- 999005 2- 158925 2-331639 2-518170 2-719624 2- 937194 3- 172169 3-425943 3-700018 3- 996020 4- 315701 4- 660957 5- 033834 5-436540 5- 871464 6- 341181 6- 848475 7- 396353 7- 988061 8- 627106 9- 317275 10-062657 10- 867669 11- 737083 12- 676050 13- 690134 14- 785344 15- 968172 17*245626 18-625276 20- 115298 21- 724522 23-462483 25-339482 27-366640 29-555972 31-920449 34-474085 37-232012 40-210573 43-427419 46-901613 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 SO 31 32 33 34 35 36 37 38 39 40 41 42 43 44 45 46. 47 48 49 50 00 232 ANNUITIES, TABLE IV. Showing the Amount to which One Pound per Annum will increase at Compound Interest in any Number of Years not exceeding 50. (No. 9 12.) Years. 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 43 44 45 46 47 48 49 50 2 per Cent. 1-0000 2-0200 3- 0604 4- 1216 5- 2040 6- 3081 7- 4343 8- 5830 9- 7546 10-9497 12- 1687 13- 4121 14- 6803 15- 9739 17- 2934 18- 6393 20-0121 21- 4123 22- 8406 24- 2974 25- 7833 27- 2990 28- 8450 30-4219 32- 0303 33- 6709 35-3443 37- 0512 38- 7922 40-5681 42-3794 44-2270 46-1116 48- 0338 49- 9946 51-9944 54-0343 56-1149 58-2372 60-4020 62-6100 64-8622 67-1595 69-5027 71-8927 74-3308 76-8172 79-3535 81-9406 84-5794 21 per Cent. 1-0000 2- 0250 3- 0756 4- 1525 5- 2563 6- 3877 7- 5474 8- 7361 9- 9545 11- 2034 12- 4835 13- 7956 15- 1404 16- 5190 17- 9319 19- 3802 20- 8647 22- 3863 23- 9460 25-5447 27- 1833 28- 8629 30-5844 32-3490 34-1578 36- 0117 37- 9120 39-8598 41-8563 43-9027 46-0003 48-1503 50-3540 52-6129 54-9282 57-3014 59-7339 62-2273 64-7830 67-4026 70-0876 72-8398 75-6608 78-5523 81-5161 84-5540 87-6679 90-8596 94-1311 97-4843 3 per Cent. 1-000000 2- 030000 3- 090900 4- 183627 5- 309136 6- 468410 7- 662462 8- 892336 10- 159106 11- 463879 12- 807796 14- 192030 15- 617790 17-086324 • 18-598914 20- 156881 21- 761588 23-414435 25- 116868 26- 870374 28-676486 30-536780 32-452884 34-426470 36-459264 38-553042 40-709634 42-930923 45-218850 47-575416 50-002678 52-502759 55-077841 57-730177 60-462082 63-275944 66-174223 69-159449 72-234233 75-401260 78-663298 82-023196 85-483892 89-048409 92-719861 96-501457 100-396501 104-408396 108-540648 112-796867 4 per Cent. 1-000000 2- 040000 3- 121600 4- 246464 5- 416323 6- 632975 7- 898294 9-214226 10-582795 12- 006107 13- 486351 15- 025805 16- 626838 18-291911 20- 023588 21- 824531 23-697512 25-645413 27-671229 29-778079 31-969202 34-247970 36-617889 39-082604 41-645908 44-311745 47-084214 49-967583 52-966286 56-084938 59-328335 62-701469 66-209527 69-857909 73-652225 77-598314 81-702246 85-970336 90-409150 95-025516 99-826536 104-819598 110-012382 115-412877 121-029392 126-870568 132-945390 139-263206 145-833734 152-667084 5 per Cent. 1-000000 2- 050000 3- 152500 4- 310125 5- 525631 6- 801913 8- 142008 9- 549109 11- 026564 12- 577893 14- 206787 15- 917127 17-712983 19-598632 21-578564 23-657492 25-840366 28-132385 30-539004 33-065954 35-719252 38-505214 41-430475 44-501999 47-727099 51-113454 54-669126 58-402583 62-322712 66-438848 70-760790 75-298829 80-063771 85-066959 90-320307 95-836323 101-628139 107-709546 114-095023 120-799774 127-839763 135-231751 142-993339 151-143006 159-700156 168-685164 178-119422 188-025393 198-426663 209-347996 6 per Cent. 1-000000 2-060000 3- 183600 4- 374616 5- 637093 6- 975319 8- 393838 9- 897468 11-491316 13- 180795 14- 971643 16-869941 18-882138 21-015066 23-275970 25-672528 28-212880 30-905653 33-759992 36-785591 39-992727 43-392290 46-995828 50-815577 54-864512 59-156383 63-705766 68-528112 73-639798 79-058186 84-801677 90-889778 97-343165 104-183755 111-434780 119-120867 127-268119 135-904206 145-058458 154-761966 165-047684 175-950545 187-507577 199-758032 212-743514 226-508125 241-098612 256-564529 272-958401 290-335905 7 per Cent. 1-000000 2- 070000 3- 214900 4- 439943 5- 750739 7- 153291 8- 654021 10- 259803 11- 977989 13-816448 15-783599 17-888451 20-140643 22-550488 25-129022 27-888054 30-840217 33-999033 37-378965 40-995492 44-865177 49-005739 53-436141 58-176671 63-249038 68-676470 74-483823 80-697691 87-346529 94-460786 102-073041 110-218154 118-933425 128-258765 138-236878 148-913460 160-337402 172-561020 185-640292 199-635112 214-609570 230-632240 247-776496 266-120851 285-749311 306-751763 329-224386 353-270093 378-999000 406-528929 Years. 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 43 44 45 46 47 48 49 50 0*£ tff! ■&8 ue-a ANNUITIES. TABLE V. Exhibiting the Law of Mortality at Carlisle. (No. 32.) Age. Number who complete that Age. 0 1 Month 2 Months 1 Year 2 Years 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 10000 9467 9313 9226 8970 8715 8461 7779 7274 6998 6797 6676 6594 6536 6493 6460 6431 6400 6368 6335 6300 6261 6219 6176 6133 6090 6047 6005 5963 5921 5879 5836 5793 5748 5698 5642 5585 die in the next interval. 533 154 87 256 255 254 682 505 276 201 121 82 58 43 33 29 31 32 33 35 39 42 43 43 43 43 42 42 42 42 43 43 45 50 56 57 57 Age. completel die in that their next 32 33 34 35 36 37 38 39 40 41 42 43 44 45 46 47 48 49 50 51 52 53 54 55 56 57 58 59 60 61 62 63 64 65 66 67 68 Number who Year. 5528 5472 5417 5362 5307 5251 5194 5136 5075 5009 4940 4869 4798 4727 4657 4588 4521 4458 4397 4338 4276 4211 4143 4073 4000 3924 3842 3749 3643 3521 3395 3268 3143 3018 2894 2771 2648 56 55 55 55 56 57 58 61 66 69 71 71 71 70 69 67 63 61 59 62 65 68 70 73 76 82 93 106 122 126 127 125 125 124 123 123 123 Age. 69 70 71 72 73 74 75 76 77 78 79 80 81 82 83 84 85 86 87 88 89 90 91 92 93 94 95 96 97 98 99 100 101 102 103 104 Number who complete that die in their next Year. 2525 2401 2277 2143 1997 1841 1675 1515 1359 1213 1081 953 837 725 623 529 445 367 296 232 181 142 105 75 54 40 30 23 18 14 11 9 7 5 3 1 124 124 134 146 156 166 160 156 146 132 128 116 112 102 94 84 78 71 64 51 39 37 30 21 14 10 7 5 4 3 2 2 2 2 2 1 TABLE VI. Showing the Value of an Annuity on a Single Life at every Age, according to the Carlisle Table of Mortality, when the Rate of Interest is Five per Cent. (No. 65.) Age. 0 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 Value. 12- 083 13- 995 14- 983 15- 824 16- 271 16-590 16-735 16-790 16-786 16-742 16-669 16-581 16-494 16-406 16-316 16-227 16-144 16-066 Age. 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 Value. 15-987 15-904 15-817 15-726 15-628 15-525 15-417 15-303 15-187 15-065 14-942 14-827 14-723 14-617 14-506 14-387 14-260 Age. 35 36 37 38 39 40 41 42 43 44 45 46 47 48 49 50 51 Value. 14-127 13-987 13-843 13-695 13-542 13-390 13-245 13-101 12-957 12-806 12-648 12-480 12-301 12-107 11-892 11-660 11-410 Age. 52 53 54 55 56 57 58 59 60 61 62 63 64 65 66 67 68 Value. 154 892 624 347 063 771 478 199 940 712 487 258 016 765 503 ■227 941 Age. 69 70 71 72 73 74 75 76 77 78 79 80 81 82 83 84 85 Value. 6-643 6-336 6-015 5-711 5-435 5-190 4-989 4-792 4-609 4-422 4-210 4-015 3-799 3-606 3-406 3-211 3-009 Age. 86 87 88 89 90 91 92 93 94 95 96 97 98 99 100 101 102 103 Value. 2-830 2-685 2-597 2-495 2-339 2-321 2:412 2-518 2-569 2-596 2-555 2-428 2-278 2-045 1-624 1-192 0-753 0-317 2 G 233 VOL. III. 234 ANNUITIES. TABLE VII. Ages. 0 & 1 & 2 & 3 & 4 & 5 & 10 6 & 11 7 & 12 8 & 13 9 & 14 10 & 15 11 & 16 12 & 17 13 & 18 14 & 19 15 & 20 16 & 21 17 & 22 18 & 23 19 & 24 Value. 10-551 12- 331 13- 258 14- 019 14-402 14-649 14-731 14-736 14-689 14-606 14-500 14-389 14-284 14-178 14-069 13-959 13-853 13-746 13-636 13-520 Ages. 20 & 25 21 & 26 22 & 27 23 & 28 24 & 29 25 & 30 26 & 31 27 & 32 28 & 33 29 & 34 30 & 35 31 & 36 32 & 37 33 & 38 34 & 39 35 & 40 36 & 41 37 & 42 38 & 43 39 & 44 Value. 13-398 13-272 13-137 13-000 12-867 12-742 12-615 12-482 12-344 12-208 12-078 11-944 11-806 11-661 11-508 11-354 11-204 11-056 10-907 10-753 Ages. 40 & 45 41 & 46 42 & 47 43 & 48 44 & 49 45 & 50 46 & 51 47 & 52 48 & 53 49 & 54 50 & 55 51 & 56 52 & 57 53 & 58 54 & 59 55 & 60 56 & 61 57 & 62 58 & 63 Value. 10-598 10-444 10-287 10-121 9-937 9-737 9-519 9-292 9-054 8-799 8-528 8-242 7-950 7-657 7-375 7-106 6-860 6-615 6-370 Ages. 59 & 64 60 & 65 61 & 66 62 & 67 63 & 68 64 & 69 65 & 70 66 & 71 67 & 72 68 & 73 69 & 74 70 & 75 71 & 76 72 & 77 73 & 78 74 & 79 75 & 80 76 & 81 77 & 82 78 & 83 Value. 6-127 5-895 5-678 5-458 5-230 4-988 4-737 4-469 4-207 3-961 3-731 3-528 3-319 3-127 2-948 2-767 2-623 2-467 2-333 2-194 Ages. 79 & 80 & 81 & 82 & 83 & 84 & 85 & 86 & 87 & 88 & 89 & 90 & 91 & 92 & 93 & 94 & 95 & 100 96 & 101 97 & 102 98 & 103 84 85 86 87 88 89 90 91 92 93 94 95 96 97 98 99 Value. 2-045 1-895 1-747 1-626 1-535 1-433 1-279 1-203 1-192 1-219 1-214 1-167 1-161 1-181 1-215 1-191 1-038 0-828 0-555 0-249 TABLE VIII. Showing the Value of an Annuity on the Joint Continuance of Two Lives according to the Carlisle Table of Mortality, when the difference of their Ages is ten years, and the Rate of Interest Five per Cent. (No. 65.) Ages. 0 & 10 1 & 11 2 & 12 3 & 13 4 & 14 5 & 15 6.& 16 7 & 17 8 & 18 9 & 19 10 & 20 11 & 21 12 & 22 13 & 23 14 & 24 15 & 25 16 & 26 17 & 27 18 & 28 Value. 10-649 12- 275 13- 087 13- 769 14- 106 14-334 14-419 14-432 14-395 14-321 14-221 14-106 13-987 13-864 13-737 13-608 13-483 13-359 13-235 Ages. 19 & 29 20 & 30 21 & 31 22 & 32 23 & 33 24 & 34 25 & 35 26 & 36 27 & 37 28 & 38 29 & 39 30 & 40 31 & 41 32 & 42 33 & 43 34 & 44 35 & 45 36 & 46 37 & 47 Value. 13-117 13-008 12-896 12-776 12-648 12-510 12-365 12-214 12-058 11-900 11-747 11-607 11-474 11-342 11-207 11-063 10-912 10-750 10-579 Ages. 38 & 48 39 & 49 40 & 50 41 & 51 42 & 52 43 & 53 44 & 54 45 & 55 46 & 56 47 & 57 48 & 58 49 & 59 50 & 60 51 & 61 52 & 62 53 & 63 54 & 64 55 & 65 Value. 10-396 10-195 9-984 9-766 9-548 9-329 9-104 8-870 8-626 8-372 8-111 7-851 7-601 7-370 7-142 6-911 6-669 6-418 Ages. 56 & 66 57 & 67 58 & 68 59 & 69 60 & 70 61 & 71 62 & 72 63 & 73 64 & 74 65 & 75 66 & 76 67 & 77 68 & 78 69 & 79 70 & 80 71 & 81 72 & 82 73 & 83 74 & 84 Value. 6-156 5-881 5-600 5-319 5-044 4-779 4-529 4-302 4-094 3-921 3-746 3-580 3-407 3-210 3-020 2-807 2-616 2-430 2-260 Ages. 75 & 76 & 77 & 78 & 79 & 80 & 81 & 82 & 83 & 84 & 85 & 86 & 87 & 88 & 89 & 90 & 100 91 & 101 92 & 102 93 & 103 Value. 2-100 1-956 1-838 1-759 1-657 1-515 1-450 1-460 1-479 1-468 1-443 1-397 1-324 1-280 1-192 0-950 0-733 0-508 0-235 Showing the Value of an Annuity on the Joint Continuance of Two Lives according to the Carlisle Table of Mortality, when the difference of their ages is five years, and the Rate of Interest Five per Cent. (No. 65.) (A. A.) j AND r Acuities, Borroimng upon. See Debt, National. ||U ANNULAR, in a general sense, something in the form lyn)i-of or resembling a ring. It is also a peculiar denomina- lesi tjon 0f the fourth finger, commonly called the ring finger. ANNULET, a little circle, borne as a charge in coats of arms, as also added to them as a difference. Among the Romans it represented liberty and nobility. It also denotes strength and eternity, by reason of its circular form. ANNULOSA (from annulus, a ring or segment), a term in zoology applied by naturalists to a great division of the animal kingdom. It contains five classes, viz. Crustacea, Myriopoda, Arachnides, Insecta, and Vermes. These classes will be treated of separately in this work, as explained under the words Animal King¬ dom. See the concluding part of that article. ANNUNCIADA, Annuntiada, or Annunciata, an order of knighthood in Savoy, first instituted by Ama¬ deus I. in the year 1409. Their collar was of 15 links, in¬ terwoven one with another, in form of a true lover’s knot; and the motto F. E. R. T., signifying Fortitudo ejus Rho- dum tenuit. Amadeus VIII. gave the name Annunciada to this order, which was formerly known by that of the knot of love; changing at the same time the image of St Maurice, patron of Savoy, which hung at the collar, for that of the Virgin Mary; and, instead of the motto above mentioned, substituting the words of the angel’s salu¬ tation. ANNUNCIATION, the tidings brought by the angel Gabriel to the Virgin Mary, of the incarnation of Christ. —Annunciation is also a festival kept by the church on the 25th of March, in commemoration of these tidings. This festival appears to be of very great antiquity. There is mention made of it in a sermon which goes under the name of Athanasius. Others carry it up to the time of Gregory Thaumaturgus, because there is a sermon like¬ wise attributed to him upon the same subject. But the best critics reject both these writings as spurious. How¬ ever, it is certain that this festival was observed before the time of the council of Trullo, in which there is a canon forbidding the celebration of all festivals in Lent, except¬ ing the Lord’s day and the feast of the annunciation; so that we may date its original from the 7th century. In the Romish church, on this feast, the pope performs the ceremony of marrying or cloistering a certain number of maidens, who are presented to him in the church, clothed in white serge, and muffled up from head to foot: an officer stands by, with purses containing notes of 50 crowns for those who make choice of marriage, and notes of 100 for those who choose to veil. Annunciation is likewise a title given by the Jews to a part of the ceremony of the passover. ANODYNE (from a privative, and ohvvaw, doleo; or a negative, and odvi/Tj, pain), a term applied to medicines which ease pain and procure sleep. ANOINTERS, a religious sect in some parts of Eng¬ land, so called from the ceremony they used of anointing all persons before they admitted them into their church. This ceremony they founded upon James v. 14, 15. ANOLYMPIADES, in Antiquity, a name given by the Elians to those Olympic games which had been cele¬ brated under the direction of the Pisaeans and Arcadians. The Elians claimed the sole right of managing the Olympic games, in which they sometimes met with com¬ petitors. The hundred and fourth Olympiad was cele¬ brated by order of the Arcadians, by whom the Elians were at that time reduced very low. This, as well as those managed by the inhabitants of Pisa, they called avoXu/x- muduf, that is, unlawful Olympiads; and left them out of their annals, wherein the names of their victors and remark¬ able occurrences were registered. A N 0 235 ANOMALISTICAL Year, in Astronomy, the time that Anomalis- the earth takes to pass through her orbit: it is also called Leal the Periodical Year. Theispace of time belonging to this N year is greater than the tropical year, on account of the precession of the equinoxes. ANOMALOUS, a term applied to whatever is irregu¬ lar, or deviates from the rule observed by other things of the like nature. Anomalous Verbs, in Grammar, such as are not con¬ jugated conformably to the paradigm of their conjuga¬ tion. They are found in all languages. In Latin, the verb lego is the paradigm of the third conjRation, and runs thus, lego, legis, legit: by the same rule it should be fero, feris, ferit; but we say, fero, firs, fert: fero, then, is an anomalous verb. In English the irregularity relates often to the preter tense and passive participle; for ex¬ ample, give, were it formed according to rule, would make gived in the preter tense and passive participle ; whereas, in the former, it makes gave, and in the latter given. ANOMALY, in Astronomy, an irregularity in the mo¬ tions of the planets, whereby they deviate from the aphe¬ lion or apogee. ANOMGEANS, in Ecclesiastical History, the name by which the pure Arians were called in the 4th century, in contradistinction to the Semi-Arians. The word is de¬ rived from the Greek avogoiog, different, dissimilar : for the pure Arians asserted that the Son was of a nature dif¬ ferent from, and in nothing like, that of the Father; whereas the Semi-Arians acknowledged a likeness of na¬ ture in the Son, at the same time that they denied, with the pure Arians, the consubstantiality of the Word. The Semi-Arians condemned the Anomceans in the council of Seleucia; and the Anomteans in their turn condemned the Semi-Arians in the councils of Constantinople and Antioch, erasing the word ogoiog, like, out of the formula of Rimini and that of Constantinople. ANOMORHOMBOIDIA, in Natural History, the name of a genus of spars. The word is derived from the Greek amiLahrig, irregular, and %ofLZoubr\g, a rhomboidal figure. The bodies of this genus are pellucid, crystalline spars, of no determinate or regular external form, but al¬ ways breaking into regularly rhomboidal masses; easily fissile, and composed of plates running both horizontally and perpendicularly through the masses, but cleaving more readily and evenly in a horizontal than in a perpen¬ dicular direction, the plates being ever composed of irre¬ gular arrangements of rhomboidal concretions. ANONYMOUS, something that is nameless, or of which the name is concealed. It is a term equally ap¬ plied to books which do not express the author’s name, or to authors whose names are unknown. See Biblio¬ graphy. ANOREXIA, Anorexy (from a negative, and ogsg/f, appetite), a want of appetite, or a loathing of food. ANOSSI, a province of the island of Madagascar, lying between lat. 23. 18. and 26. 0. S. See Madagascar. ANOTTA, or Arnotta, in dyeing, an elegant red colour, formed from the pellicles or pulp of the seeds of the bixa, a tree common in South America. It is also called Terra Orleana, and Roucou. The manner of making anotta is as follows: The red seeds, cleared from the pods, are steeped in water for seven or eight days, or longer, till the liquor begins to fer¬ ment; then strongly stirred, stamped with wooden paddles and beaters, to promote the separation of the red skins: this process is repeated several times, till the seeds are left white. The liquor, passed through close cane sieves, is pretty thick, of a deep red colour, and an offensive smell; in boiling, it throws up its colouring matter to the surface in form of scum, which is afterwards boiled down by itself 233 A N Q Anout to a due consistence, and made up while soft into balls. II The anotta commonly met with among us is moderately hard and dry, of a brown colour on the outside, and a dull red within. It is difficultly acted upon by water, and tinges the liquor only of a pale brownish yellow colour. It veiy readily dissolves in rectified spirit of wine, and commu¬ nicates a high orange or yellowish red. Hence it is used as an ingredient in varnishes, for giving more or less of an orange cast to the simple yellows. Alkaline salts render it perfectly soluble in boiling water, without altering its colour. Wool or silk boiled in the solution acquires a deep, but not a very durable, orange dye. Its colour is not changed by alum or by acids, any more than by alka¬ lies ; but when imbibed in cloth, it is discharged by soap, and destroyed by exposure to the air. It is said to be an antidote to the poisonous juice of manioc or cassava. ANOUT, a small island in the Schagerrack, or that part of the sea of Denmark which has Norway on the north, Jutland on the west, and the isle of Zealand on the south. It lies in long. 13. 0. E. and lat. 56. 36. N. ANQUETIL, Lewis Peter, a French historian, was born at Paris on the 21st of January 1723. At the age of 17 he entered the congregation of St Genevieve, where he taught theology and literature with ability and success. He afterwards became director of the academy at Rheims ; and, in 1759, he was appointed prior of the abbey de la Roe, in Anjou. Soon after this he was sent, in the capacity of director, to the college of Senlis. In 1766 he obtained the curacy or priory of Chateau-Renard, near Montargis, which he exchanged, at the commencement of the Revo¬ lution, for the curacy of La Villette, in the neighbourhood of Paris. During the reign of terror he was imprisoned at St Lazare. On the establishment of the National Insti¬ tute, he was elected a member of the second class, and was soon afterwards employed in the office of the minister for foreign affairs. Endowed with a robust constitution, which was preserved by a natural equality of temper, and general moderation in diet, Anquetil was capable of very laborious exertions, and is said to have passed ten hours every day regularly in study. When upwards of eighty he still meditated extensive literary undertakings; but he was carried off by death, in the midst of his projects and researches, on the 6th of September 1808, in the 86th year of his age. On the evening previous to this event, he is reported to have said to one of his friends, “ Come find see a man who is dying full of life.” As an author, M. Anquetil does not stand very high in the ranks of literature. He possessed more industry in research than ability or judgment in execution. His style is censurable in many respects, and he appears to have been almost entirely destitute of the critical discernment and philosophical sagacity which are requisite to form the character of a good historian. The following is a list of his principal works. Histoire Civile et Politicpie de la Ville de Reims. 1756—57, 3 vols. 12mo. I he history is brought no farther down than 1657: a fourth volume should have been added, but it never appeared. Anquetil is said to have written this work in concert with one Felix de la Salle, and it is, perhaps, the best of all his productions. 2. Almanack de Remis. 1754, 24to 3. LEsprit de la Ligue, ou His¬ tone Politique des Troubles de France pendant les 16 et 17 $iecks. 1767, 3 vols. 12mo. This work has been frequent¬ ly reprinted. 4. Intrigue du Cabinet sous Henri 1V. et sous Louis XIII. terminee par la Fronde. 1780, 4 vols. 12mo. 5. Louis XIV. sa Coin et le Regent. 1789, 4 vols IJmo preprinted in 1794, 5 vols. 12mo. 6. Vie du Marechal de Iillars, ecnte par lui-mhne, suivie du Journal dela Cour de 1724 a 1734. Paris, 1787, 4 vols, 12mo; re¬ printed in 1792. 7. Precis de l'Histoire Universelle. 1797, A N Q 9 vbls. 12mo; reprinted in 1801 and 1807, in 12 vols. 12mo. This work has been translated into English, Spa¬ nish, and Italian. 8. Motifs des Guerres et des Traites de Paix de la France pendant les regnes de Louis XIV., Louis ^li XV., et Louis XVI. 1798, 8vo. 9. Histoire de France^ depuis les Gaules jusqu a la jin de la Monarchic. 1805 et seqq. 14 vols. 12mo. This work was composed in haste and is of no great value. 10. Notice sur la Vie deM. An- quetil du Perron. M. Anquetil likewise wrote several papers in the Memoirs of the Institute. See Biographk Universelle. ANQUETIL DU PERRON (Abraham Hyacinth), brother of the subject of the preceding article, was born at Paris on the 7th of December 1731. Having distin¬ guished himself as a student at the university of that city, and acquired a considerable knowledge of the Hebrew language, he was invited to Auxerre by M. de Caylus, then the bishop of that diocese. This prelate made him study theology, first at the academy of his diocese, and afterwards at that of Amersfort, near Utrecht; but An¬ quetil had no desire to embrace the ecclesiastical voca¬ tion, and devoted himself with ardour to the study of the different dialects of the Hebrew, and of the Arabic and Persian. Neither the solicitations of M. de Caylus, nor the hopes of rapid preferment, had the power to detain him at Amersfort after he thought he had acquired every thing that was to be learnt there. He returned to Paris, where his diligent attendance at the Royal Library, and his ardour in the prosecution of his favourite studies, at¬ tracted the attention of the Abbe Sallier, keeper of the manuscripts, who introduced him to the acquaintance of his associates and friends, whose united exertions pro¬ cured for him a small salary, as student of the oriental languages. He had scarcely received this appointment, when, having accidentally laid his hands on some manu¬ scripts in the Zend, he formed the project of a voyage to India, with the view of discovering the works of Zoroaster. At this period an expedition was preparing at the port of L’Orient, which was destined for India. M. du Perron, however, applied in vain, through his protectors, for a passage ; and seeing no other means of accomplishing his plan, he enlisted as a common soldier, and set out from Paris,^with a knapsack on his back, on the 7th of Novem¬ ber 1/54. His friends procured his discharge; and the minister, affected by this romantic zeal for science, grant¬ ed him a free passage, a seat at the captain’s table, and a salary, the amount of which was to be fixed by the gover- nor of the French settlements in India. After a passage of nine months, Anquetil landed, on the 10th of August 1755, at Pondicherry. Here he remained no longer than was necessary to make himself master of the modern Per¬ sian, and then hastened to Chandernagore, where he thought to acquire the Sanscrit. But in this he was de¬ ceived ; and he was on the point of returning, when a seiious complaint threatened his life. He had scarcely escaped from this danger, when war was declared between I ranee and England; Chandernagore was taken; and Anquetil resolved to return to Pondicherry by land. After a journey of one hundred days, in the course of which he encountered many adventures, and suffered many hard¬ ships, he arrived at Pondicherry. Here he found one of his brothers who had arrived from France, and embarked with him for Surat; but, with the view of exploring the country, he landed at Mahe, and proceeded on foot. It was at Surat that he succeeded, by perseverance and ad- dressm his intercoursewith the native priests, in acquiring a sufficient know ledge of the languages to enable him to translate the Dictionary called the Vedidad-Sade, and some other works, hrom thence he proposed going to Benares, to study the languages, antiquities, and sacred laws of A N Q the Hindoos; but the capture of Pondicherry obliged him felon, to return to France. He accordingly embarked on board Sv.'-'an English vessel, and landed at Portsmouth in the month of November 1761. After spending some time in London, and visiting Oxford, he set out for Paris, where he arrived on the 4th of May 1762, without fortune or the desire of acquiring any, but esteeming himself rich in the posses¬ sion of one hundred and eighty oriental manuscripts, be¬ sides other curiosities. The Abbe Barthelemy, and his other friends, procured for him a pension, with the title and appointments of Interpreter for the oriental languages at the royal library. In 1763 the Academy of the Belles Lettres received him among the number of its associates; and from that period he devoted himself to the arrange¬ ment and publication of the materials he had collected during his eastern travels. In 1771 he published a work in three volumes 4to, under the title of Zend-Avesta, contain¬ ing collections from the sacred writings of the Persians, among which are fragments of works ascribed to Zoro¬ aster ; and he accompanied this work with an account of the life of that sage. This publication must be consider¬ ed as constituting a very important accession to our stores of oriental literature. A recent historian, and very com¬ petent judge, refers to the Zend-Avesta, as certainly the most authentic source from which we can derive informa¬ tion regarding the religion and institutions of the great Persian legislator. (Sir John Malcolm’s Hist, of Persia, vol. i. p. 193, note.) To the Zend-Avesta M. du Perron prefixed a discourse, in which he treated the university of Oxford, and some of its learned members, with ridicule and disrespect. Mr (afterwards Sir William) Jones re¬ plied to these invectives in an anonymous letter, address¬ ed to the author, written in French, with uncommon force and correctness of style, but at the same time with a de¬ gree of asperity which could only be justified by the pe¬ tulance of M. du Perron. In 1778 he published his Le¬ gislation Orientale, in 4to, a work in which he controverts the system of Montesquieu, and endeavours to prove that the nature of oriental despotism has been misrepresented by most authors; that in the empires of Turkey, Persia, and Hindostan, there are codes of written law which equally bind the prince and subject; and that in these three empires the inhabitants possess both movable and immovable property, which they enjoy with perfect se¬ curity. His llecherches Historiques et Geographiques sur Z’/wefe appeared in 1786, and formed part of Thieffenthaler’s Geography of India. They were followed in 1789 by his treatise De la Dignite du Commerce et de I'Etat du Com- mer^ant. The Revolution seems to have greatly affected him. During that period he abandoned society, shut himself up in his study, and devoted himself entirely to literary seclusion. In 1798 he published L'Inde en Rap¬ port avec lEurope, &c. in 2 vols. 8vo; a work which is more remarkable for its virulent invectives against the English, and for its numerous misrepresentations, than for the information which it contains, or the soundness of the reflections which it conveys. The spirit of the work, indeed, may be ascertained from the summary of its con¬ tents stated in the title-page, in which the author pro¬ fesses to give a detailed, accurate, and terrific picture of the English Machiavelism in India; and lie addresses his work in a ranting bombastic dedication to the Manes of Dupleix and Labourdonnais. In 1804 he published a La¬ tin translation from the Persian of the Oupneh'hat or Upa- nischada, i. e. Secrets which must not be revealed, in 2 vols. 4to, On the re-organization of the Institute, M. Anquetil was elected a member, but soon afterwards gave in his resignation. He died at Paris on the 17th of January 1805., , osf obno/R nrnn i .idhow isdio Besides the works we have already enumerated, M, A N /S 237 Anquetil read to the academy several memoirs on subjects Ansje connected with the history and antiquities of the East. 11 At the time of his death he was engaged in revising a Anselm^ translation of the Travels of Father Paulin de St Barihe- lemy in India, which work was continued by M. Silvestre de Sacy, and published in 1808, in 3 vols. 8vo. He also left behind him a great number of manuscripts, among which his biographers particularly notice the translation of a Latin treatise on the Church, by Doctor Legros, in 4 vols. 4to. From the preceding narrative our readers will be ena¬ bled to form some notion of the character of Anquetil du Perron. Among his countrymen he is regarded as one of the most learned men of the 18th century. He certainly distinguished himself by a very ardent and disinterested zeal in the prosecution of those studies to which he dedi¬ cated the labours of a long life; but the lustre of his lite¬ rary character was obscured by a very absurd vanity, and the most inveterate prejudices. In a Discourse addressed to the Asiatic Society at Calcutta, in 1789, Sir William Jones speaks of him as “ having had the merit of under¬ taking a voyage to India, in his earliest youth, with no other view than to recover the writings of Zeratust (Zo¬ roaster), and who would have acquired a brilliant reputa¬ tion in France, if he had not sullied it by his immoderate vanity and virulence of temper, which alienated the good¬ will even of his own countrymen.” In the same Discourse he affirms that M. Anquetil most certainly had no know¬ ledge of the Sanscrit.—See Biographic Universelle; Month¬ ly Revieiv, vol. Ixi.; Lord Teignmouth’s Life of Sir Wil¬ liam Jones. (k.) ANSfE, in Astronomy, implies the parts of Saturn’s ring projecting beyond the disk of the planet. The word is Latin, and properly signifies handles; these parts of the ring appearing like handles to the body of the planet. ANSARIANS, a people of Syria, so called in the country, but styled in Delisle’s maps Ensarians, and in those of D’Anville Nassaris. The territory occupied by the Ansarians is that chain of mountains which extends from Antakia to the rivulet called Nahr-el-Kahir, or the Great River. They are divided into several tribes or sects, among which are distinguished the Shamsia, or adorers of the sun; the Kelbia, or worshippers of the dog ; and the Kadmousia, who are said to pay a particu¬ lar homage to that part in women which corresponds to the priapus. ANSBACH, a bailiwick in the circle of Rezat, in the kingdom of Bavaria, extending over 124 square miles, or 79,360 acres, with 2 market-towns and 79 villages. It is an undulating plain, watered by the rivers Rezat, Zenn, and Bidert. It contains 11,800 inhabitants, who draw from its fruitful soil excellent corn and tobacco, and keep many horses and cattle. There is an alabaster and a gyp¬ sum quarry worked in the bailiwick. The capital, of the same name, is about 30 miles south-west of Nuremberg, and contains 950 inhabitants. ANSE, an ancient town of France, in the Lyonnois, on the river Saone. It is the head of a canton in the depart¬ ment of the Rhone, and formerly bore the title of barony. Population 1640. 131 miles north of Lyons. Long. 6. 55. W. Lat. 45.55. N. ANSELM, archbishop of Canterbury, in the reigns of William Rufus and Henry I. He was born in the year 1033, at Aost, a town in Savoy, at the foot of the Alps. He became a monk in the abbey of Bee in Normandy, of which he was afterwards chosen prior, and then abbot. In the year 1092 he was invited over to England by Hugh earl of Chester; and in the year following was prevailed on, as we are told, with great difficulty, to accept the archbishopric of Canterbury. He enjoined celibacy on the 238 A N 8 Ansiko clergy, for which he was banished by King Rufus; but he II was recalled by Henry on his coming to the crown. In con- Anslo. ^ formity to Pope Urban’s decree, he refused to consecrate such bishops as were invested by the king, denying it to be the royal prerogative: for this he was banished again, till, the pope and king agreeing, he was recalled in 1107. In short, from the day of his consecration to that of his death, he was continually employed in defending the pre¬ rogative of the church against the encroachments of the crown ; and for that purpose spent much of his time in tra- vellingbackwardsandforwardsbetween England and Rome, for the advice and direction of his holiness. At the coun¬ cil of Bari, in the kingdom of Naples, the pope being puzzled by the arguments of the Greeks against the Holy Ghost’s proceeding from the Father, called upon Anselm, who was present, and discussed their objections with great applause. Priests call him a resolute saint; to other peo¬ ple he appears to have been an obstinate and insolent priest. He wrought many miracles, if we may believe the au¬ thor of his life, both before and after his death, which happened at Canterbury in the 76th year of his age, anno 1109. He was canonized in the reign of Henry VII. Anselm, though we may disregard him as a saint, deserves to be remembered as one of the principal revivers of lite¬ rature, after three centuries of profound ignorance. His works have been often reprinted. The best edition is that of Father Gerberon, printed at Paris in 1675, in 2 vols. folio. ANSIKO, a kingdom in the interior of western Africa, bounded on the west by the river Umbre, which runs into the Zaire, the kingdom of Wangua, and the Amboes, who border on Loango; and on the south by Songo and Sunda, provinces of Congo. It contains a great variety of wild beasts, as lions, rhinoceroses, &c. and many copper-mines. The king of Ansiko, or the great Micoco, is said to command thirteen kingdoms, and is esteemed the most powerful mo¬ narch in this part of Africa. Our knowledge of the country, however, is very imperfect, being derived solely from the hearsay reports of Lopez and Merolla, who visited Congo in the 16th century. According to them, the people are brave, swift, active, but savage and cruel in the most frightful degree, human flesh being not only eaten, but openly sold in the markets, and the subjects offering them¬ selves to the sovereign for the gratification of his palate. These reports, which are not very credible, are contrast¬ ed with others that represent them as an industrious peo¬ ple, manufacturing cloths from the fibres of the palm-tree, and carrying on an extensive trade both with Congo and interior Africa. Their language is barbarous, and difficult to be learned, even by the inhabitants of Congo. The most distinguished among them wear red and black caps of Portuguese velvet; the lower ranks go naked from the waist upwards; and, to preserve their health, anoint their bodies with a composition of pounded sandal-wood and palm-oil. Their arms are battle-axes, and small but very strong bows, adorned with serpents’ skins. Their strings are made of pliant and tender shoots of trees, that will not break; and their arrows of hard and light wood. They kill birds flying, and shoot with surprising swiftness. With equal dexterity they manage their battle-axes, one end of which is sharpened and cuts like a wedge, while the other is flattened like a mallet, with a handle set be¬ tween, about half the length of the iron, rounded at the end like an apple, and covered with the skin of a serpent. The current money in this country is the zimbis or shell, which passes among several African nations. They wor¬ ship the sun as their chief deity, representing him by a male figure, and the moon by a female. They have also an in¬ finite number of inferior deities, each individual having a particulai idol. They seem on the whole to be a people A N S respecting whom it might be desirable to obtain further Ansi j and more recent information. ANSLO, a seaport town of Norway, in the province of Aggerhuus, with a bishop’s see. The supreme court of, tll£ u justice is held here for Norway. It is seated on a bay ot^ the same name. Long. 10. 14. E. Eat. 50. 24. N. ANSON, George, a gentleman whose merit and good fortune as a naval commander exalted him to the rank of nobility. He was the son of William Anson, Esq. of Huckborough, in Staffordshire ; and, showing an early in, clination for the sea, received a suitable education. The first command he enjoyed was that of the Weasel sloop in 1722; but the most memorable action of his life, and the foundation of his future good fortune, took place on his receiving the command of five ships, a sloop, and two victuallers, equipped to annoy the Spaniards in the South Seas, and to co-operate with Admiral Vernon across the Isthmus of Darien ; an expedition the principal object of which failed by the unaccountable delay in fitting him out. He sailed, however, in September 1740; doubled Cape Horn in a dangerous season ; lost most of his men by the scurvy ; and with only one remaining ship, the Centurion, crossed the great Pacific Ocean. If no considerable na¬ tional advantage resulted from this voyage, Commodore Anson made his own fortune, and enriched his surviving companions, by the capture of a rich galleon on her pas¬ sage from Acapulco to Manilla, with which he returned home round the Cape of Good Hope. If he was lucky in meeting this galleon, he was no less fortunate in escaping j a French fleet then cruising in the Channel, by sailing through it during a fog. He arrived at Spithead in June 1744. In a short time after his return he was appointed rear-admiral of the blue, and one of the lords of the ad¬ miralty. In April 1745 he was made rear-admiral of the white, and the following year vice-admiral of the blue, at which time he was chosen to represent the borough of Heydon in parliament. In 1747, being on board the Prince George of 90 guns, in company with Admiral W^arren and 12 other ships, he intercepted, off Cape Finisterre, a powerful fleet, bound from France to the East and West Indies ; when, by his valour and conduct, he again enriched himself and his officers, and at the same time strengthened the British navy, by taking six men of war and four East Indiamen, not one of them es¬ caping. The French admiral, M. Jonquiere, on present¬ ing his sword to the conqueror, said, Monsieur, vous aves vaincu VInvincible, et la Gloire vous suit—“ Sir, you have conquered the Invincible, and Glory follows you point¬ ing to the ships named the Invincible and the Glory, which he had taken. For his signal services he was created baron of Soberton in Hants. The same year he was appointed vice-admiral of the red; and, on the death of Sir John Norris, was made vice-admiral of England. In 1748 he was made admiral of the blue : he was afterwards appoint¬ ed first lord of the admiralty, and was at length made ad¬ miral and commander-in-chief of his majesty’s fleet, in which rank he continued, with a very short interval, until his death ; and the last service he performed was to con¬ vey Queen Charlotte to England. He died in June 1762. No performance ever met with a more favourable recep¬ tion than the account of Anson’s voyage round the world. Though it is printed under the name of his chaplain, it was composed under his lordship’s own inspection, and from the materials he himself furnished, by Mr Benjamin Robins. ANSTRUTHER, Easter, a royal borough and parish of Scotland, in the county of Fife. It is situated on the north shore of the Frith of Forth, and possesses an ex¬ cellent harbour. Population in 1821, 1090. 10 miles S. of St Andrews. ANT fu er Anstruther, Wester, a parish and small seaport of H Scotland, situated near Easter Anstruther, with which, nf and the towns of Crail, Pittenweem, and Kilrenny^ it ^unites in returning a representative to parliament. The parish includes the Isle of May. There is a weekly packet-boat to Leith; and the steam-boat that plies be¬ tween Edinburgh and Aberdeen touches at Wester as well as at Easter Anstruther. Population 429. 23 miles north-east of Edinburgh. Long. 2. 44. W. Lat. 56. 12. N. ANT. The history of a tribe of insects so long cele¬ brated for their industry and frugality, and for the dis¬ play of that sagacity which characterizes some of the higher orders of animals, is peculiarly calculated to oc¬ cupy the attention of modern naturalists. The ancients, indeed, had often noticed the habits and economy of the ant; but their accounts, at all times deficient in accuracy from the want of precise definitions and logical arrange¬ ment of the objects they describe, are in this instance so mixed up with fanciful notions and chimerical doc¬ trines, and so coloured by the vivid imagination and cre¬ dulity of the narrators, as to have retarded rather than advanced the progress of real knowledge. Aristotle and Pliny report, for instance, that the labours of ants are in a great measure regulated by the phases of the moon; and the latter mentions a species found in the northern parts of India, whose size was said to equal that of the wolves of Egypt, whose colour was the same as that of a cat, and whose occupation in winter consisted in digging up gold from the bowels of the earth; while the inhabi¬ tants in the summer robbed them of their treasures, after having decoyed them, by stratagem, from their nests. Great mistakes have prevailed, even in later times, from the circumstance of the larvae of ants bearing a resem¬ blance to grains of corn, which it was supposed these in¬ sects hoarded up as a provision for winter consumption. The form of the eggs and of the larvae, and the attention paid to them by the ants, were described by Dr King in the 23d number of the Philosophical Transactions; but Leeuwenhoeck was the first who distinguished with precision the different forms which the insect assumes in the several stages of its growth. He traced the succes¬ sive changes from the egg to the larva, the nymph, and the perfect insect. Swammerdam pursued his scrutiny into these successive developments with greater minute¬ ness ; and, unrivalled in the art of microscopic dissection, discovered the wonderful encasement of all the parts of the future ant at every preceding stage, and showed that it appears under such different forms only from the nature of its envelopes, each of which, at the proper period, is in its turn cast off’. Linnaeus (Memoirs of the Royal Acade- my °f Sciences at Stockholm, vol. ii.) ascertained some of the leading facts with regard to the distinction between the sexes, and determined that the ants which are fur¬ nished with wings are the only individuals that exercise the sexual functions. Several particulars with regard to the economy of the ants were published by Mr Gould, in a book entitled An Account of English Ants, of which an abstract is given in the Philosophical Transactions for 1747, by the Rev. Dr Miles. The facts are there stated with tolerable correctness; but some errors have been committed by following too closely the analogy with bees. Geoffrey (Histoire des Insectes qui se trouvent aux Environs de Paris), though a good naturalist on other topics, is a bad authority on the subject of ants. The most complete series ot observations on the natural history of these in¬ sects, is that for which we are indebted to the celebrated Swedish entomologist De Geer (Memoires pour servir d l Histoire des Insectes), an observer on whose fidelity the most implicit reliance may be placed. In the Encyclopedic Methodique, under the article ANT 239 Fourmi, Olivier has drawn up an able statement of all Ant. the material facts that had been established by preceding naturalists ; without, however, adding any original obser¬ vations of his own, excepting the description of five or six undescribed species. A full account of the habits of those ants which for a long period infested the island of Mar¬ tinique, is contained in some of the earlier numbers of the Journal de Physique (vols. ix. and x.) The author of these memoirs, M. Barboteau, has given many curious details on this subject, and has cited a number of facts on various authorities; and the account might now be swell¬ ed by the reports of subsequent travellers in different parts of the world; but these statements are often made upon slender authority, and are too much tinctured with the marvellous to admit of much credit being attached to them. The narrative given to us by Bonnet in the second volume of his Observations sur les Insectes, of the proceed¬ ings of a colony of ants which had established itself in the head of a large thistle, and which he transported into his house, is highly interesting; but it elucidates only a few points of their economy, and leaves us to regret that so patient and indefatigable an observer had not bestowed more of his attention to the study of this tribe of insects. In the Philosophical Transactions for 1790 we find an in¬ teresting memoir on the sugar-ant, a species which, for a period of ten years, committed dreadful ravages in the sugar plantations throughout the whole island of Grenada. The most methodical account of this tribe of insects that has yet appeared is that of Latreille, in his Histoire Na- turelle des Fourmis, published at Paris in 1802, a work which alone would have secured the reputation of the author as an able and scientific naturalist. His merit is particularly conspicuous in the clearness and accuracy of his descriptions of each species, and the luminous method of arrangement which he has adopted in their classifica¬ tion. He gives an account of one hundred species which he had himself observed, and of twenty-four which he has described from the reports of others : these he distributes into nine natural families, according to the situation and structure of the antennae, and the form of the abdominal scales. But the work which contains the most copious collection of facts relative to the habits and economy of ants is that of Mr P. Pluber of Geneva, entitled Trade des Mceurs des Fourmis Indigenes, published in 1810. By means of an apparatus which he contrived so as to admit of his obtaining a view whenever he pleased of the inmost recesses of their habitation, he was enabled to observe what was going on in the interior of the nest, and to in¬ vestigate with success some of the most important and in¬ teresting features of their history. The results of his re¬ searches, as they are reported in his work, are highly curious and instructive, and open a wide field of specula¬ tion and inquiry to the philosophical entomologist. They have not only elucidated many obscure points with regard to one tribe of insects, but have disclosed some general views of the instincts and faculties of this order of the creation, which are totally new, and must tend, in a con¬ siderable degree, to exalt our conceptions of the inex¬ haustible powers and resources of nature. Having thus pointed out the principal sources of infor-Economy mation in this department of entomology, we shall proceed ancl policy to give an outline of the leading facts that have been0*ants' ascertained relative to the economy and domestic policy of these remarkable insects. In common with many tribes of hymenopterous insects, Functions ants present the remarkable peculiarity of a threefold dis-ofthe neu- tinction of sex among the individuals of the same species ;ters* a circumstance which is met with in no other order of the animal kingdom, and which appears, as far as obser¬ vation has extended, to be totally excluded from the plan ANT. of the vegetable creation. Besides males and females, there exists an apparently intermediate order of neuters, which are also denominated labouring or working ants. The neuters, thus exempted from every sexual function, exercise, on the other hand, all the other offices neces¬ sary for the existence and welfare of the community to which they belong. It is they who collect supplies of food, who explore the country for this purpose, and seize upon every animal substance, whether living or dead, which they can lay hold of, and transport to their nest. It is they who construct every part of their dwelling- place, who attend to the hatching of the eggs, to the feeding of the larvae, and to their removal, as occasion may require, to different situations favourable to their growth and development; and who, both as aggressors and as defenders, fight all the battles of the common¬ wealth, and provide for the safety of their weaker and more passive companions. Thus all the laborious and perilous duties of the state are performed solely by this description of ants, who act the part of helots in these singularly constituted republics of insects. We find, how¬ ever, on closer examination, that, in all probability, this anomaly in point of sex is more apparent than real; and that, however different in external conformation from the productive females, they nevertheless originally and es¬ sentially belong to the same sex. There is every reason to believe that the development of the sexual organs in the former is the consequence of some difference in the cir¬ cumstances in which the larva is placed during its growth. That such is the case with bees, is now perfectly well established; and the analogy of bees with ants, in many points of physiology, must be admitted as a strong argu¬ ment in corroboration of this theory. In all the essential features of internal structure, the supposed neuters agree with the female, and differ from the male of the same species. In all hymenopterous insects, which are armed with stings, a difference exists in the two sexes, as to the number of articulations composing the antennae; those of the female consisting of fewer pieces than those of the male. The accurate observations of Mr Kirby {Mono- graphia Apum) have determined that in the bee the an¬ tennae of the male have fifteen articulations, while those of the female and the neuter have only fourteen. In the ant, likewise, we find thirteen articulations in the male, and only twelve in the female; and likewise only twelve in the neuter. In the male ant the abdomen has seven rings, in the female and neuter only six. In the two lat¬ ter classes the head is broader, and the mandibles very large and powerful compared with those of the male, and are furnished with serrated edges, and a sharp and often hooked point. The external sexual organs of the female and of the neuter are so nearly similar in appearance, that Latreille declares he was unable to perceive the least difference between them. On the other hand, it is to be observed, that in the neuter the principal deviation from the model of the female consists in the absence of wings; a circumstance which, as it regards the organs of locomotion only, is one of subordinate importance in the economy ; and .their presence may, without difficulty, be conceived to be connected with a certain condition of the sexual organs, as are the horns of the deer, and the beard in the human species. But although of so little consequence in a physiological point of view, it is a cir¬ cumstance materially affecting their external condition. It dooms them to severe toil and exertion in traversing the ground, and in climbing up the steep paths that may he in then loute; while their more luxurious and favoured associates are fluttering in the spacious realms of air in search of amusement, and wafted to the objects of their gratification on the light breezes of the summer. Ants appear to be endowed with a greater share of ^ muscular strength than almost any other insect of the^, same size. Of this we have sufficient proofs in the viva-Sen! city of their movements, the incessant toil which manypow undergo, the great loads which they are seen to carry,^ often exceeding ten or twelve times their own weight, and the agility which they exert in making their escape from danger. This high degree of irritability is conjoined, apparently, with a corresponding share of the power of sensation; a power which is manifested in their suscepti- bility to a variety of impressions capable of affecting the organs of sense. They have a quick perception of all changes of temperature, as well as of other conditions of the atmosphere ; and are readily and disagreeably affect¬ ed by moisture. In the perfection of the sense of sight they seem to be nearly on a level with other insects; and the males and females are provided with both the descrip¬ tions of eyes peculiar to this class, namely, the-composite and the simple eyes. The labouring ants, indeed, who never fly, are frequently destitute of the latter kind; a circumstance which appears to confirm the suspicion that has often been entertained, that the simple eyes are chiefly instrumental in the vision of distant objects. Latreille describes two species of ants, in which he could not dis¬ cover the least appearance whatsoever of eyes, although he employed a high magnifying power in examining them. One of these (the Formica ccecd) is a foreign species, in¬ habiting the forests of Guiana, and of which the history is therefore little known. The other (the Formica con- tractd) is met with in the vicinity of Paris. It always conceals itself during the day under stones, or in obscure recesses, where no light can penetrate ; and emerges from its retreat only during the night. It is much less social in its habits than other ants, collecting in groups only of about a dozen individuals, and appears to be far inferior in sagacity to the rest of the tribe. Ants possess a considerable acuteness of smell, a sense ' which appears to be useful not only in directing them to their food, but also, as Bonnet first remarked, in enabling them to follow by the scent the track of their companions. If the end of the finger be passed two or three times across the line of their march, so as to brush off the odo¬ rous particles with which the ants who had already passed that way may have impregnated the track, those who fol¬ low immediately stop on arriving at the place where the experiment has been made, and afterwards direct their course irregularly, till they have passed over the space touched by the finger, when they soon find the path, and proceed with the same confidence as before. Bonnet re¬ peated this experiment frequently, and always with the same result. Latreille has endeavoured to discover the seat of smell, which had long been suspected to reside in the antennae. He, with this view, deprived several la¬ bouring ants of these organs, and replaced them near their nests. When thus mutilated, they wandered to and fro in all directions, as if they were delirious, and utterly uncon¬ scious of where they were going. Some of their compa¬ nions were seen to notice their distress, and, approaching them with apparent compassion, applied their tongues to the bleeding wounds of the sufferers, and anointed them with a liquor which they caused to flow from their own mouths. This trait of sensibility was repeatedly witnessed by Latreille, while he was observing their actions with a magnifying lens. It is indeed evident that, in all insects, the antennae are organs of the greatest utility in conveying impressions from external objects. But in the ant, independently of their importance as organs of touch, they appear to be of still greater consequence to the welfare of the individual, and of the community to which it belongs, by being the ANT. 241 chief instruments which enable them to communicate to ■'Vw'one another intelligence in which they are mutually inte¬ rested, and on which they are called upon immediately to act. Mr Huber, to whom we are indebted for a variety of curious observations on this subject, has given the name of Language Antennal to this species of intercourse. The situation of the antennae, which are placed in front of the head, their great mobility, their peculiar mecha¬ nism, which presents a series of phalanges having great freedom of play, and endowed with exquisite sensibility, conspire to fit them admirably for the function which he assigns to them,—that of producing a variety of different impressions, when applied in different ways to the anten¬ nae or other parts of those ants with which they come in contact. Thus the signal of danger, which consists in the ant which gives the alarm striking its head against the corslet of the other, is propagated from ant to ant with astonishing quickness, throughout the whole society. For a few minutes a general ferment prevails, as if they were deliberating what measures to pursue; but their re¬ solution is soon formed, and they are ready to rush in a body against the enemy. Any small animal that is disco¬ vered to have insolently invaded their repose is certain of falling a victim to their resentment, unless he can make a precipitate retreat, which he seldom effects without being covered with the bites of these furious insects. They are not, however, equally jealous of the intrusion of every kind of insect, for woodlice are often found in the interior of the nest, to whom, according to Latreille, they offer no molestation. Ants appear to be incapable of emit¬ ting sounds, so as to communicate with one another at a distance ; and there is, indeed, no evidence that they pos¬ sess the sense of hearing. The consideration of the sense of taste naturally comprehends that of their food, to which we shall therefore next proceed, eii Very erroneous opinions were prevalent with regard to r stone, and set on the architrave. ANTEPENULTIMA, in Grammar, the third syllable )f a word from the end, or the last syllable but two. ANTEPILANI, in the Roman armies, a name given to he hastati and principes, because they marched next be- ore the triarii, who were called pilani. ANTEPILEPTICS, among physicians, medicines es- .eemed good in the epilepsy. ANTEPOSITION, a grammatical figure, whereby a vord, which by the ordinary rules of syntax ought to fol- ow another, comes before it; as when, in the Latin, the t| idjective is put before the substantive, the verb before -he nominative case, &c. ANTEPREDICAMENTS, among logicians, certain * areliminary questions which illustrate the doctrine of pre- licaments and categories. ANTEQUERA, a commandery in the province of Granada in Spain, extending over 165 square miles, be¬ tween the cities of Granada, Seville, and Cordova, in a fertile soil, comprehending the city of the same name, and eight towns, besides several villages, having in the whole i population of about 40,000. Antequeha, a city of Spain, in the province of Seville, , I in Andalusia. It is situate in a beautiful and fertile valley, covered with olive-trees, around which, in all directions, lofty mountains rise to a great height, which contain valu¬ able quarries of marble of every description. Near it is i saline lake, which supplies the inhabitants of the district with salt for all their culinary purposes. As long as the Moors possessed the kingdom of Granada, this city, from being the point which commands the eastern entrance to the plain, was a military post of vast importance, and its possession was perpetually contested by the rival monarchs on both sides of it. An ancient castle still remains, in which are preserved specimens of the various military weapons and accoutrements both of the Moors and the Christians of the 14th century. It contains about 14,000 inhabitants, employed in cultivation and in the manufac- Anteros tures of cloth and leather. II ANTEROS, in Mythology, one of the two Cupids who Antheste- wrere the chief of the number. They are placed at the foot of the Venus de’ Medici. This is represented with a heavy and sullen look, agreeably to the poetical descrip¬ tion of him, as the cause of love’s ceasing. The other was called Eros. ANTESIGNANI, in the Roman armies, soldiers placed before the standards, in order to defend them, according to Lipsius; but Caesar and Livy mention the antesignani as the first line or first body of heavy armed troops. The velites, who used to skirmish before the army, were like¬ wise called antesignani. ANTESTARI, in Roman Antiquity, signifies to bear witness against any one who refused to make his appear¬ ance in the Roman courts of judicature, on the day ap¬ pointed, and according to the tenor of his bail. The plain¬ tiff finding the defendant after such a breach of his en¬ gagement, was allowred to carry him into court by force, having first asked any of the persons present to bear wit¬ ness. The person asked to bear witness in this case expressed his consent by turning his right ear, which was instantly taken hold of by the plaintiff; and this was to answer the end of a subpoena. The ear was touched upon this occasion, says Pliny, as being the seat of memory, and therefore the ceremony was a sort of caution to the party to remember his engagement. ANTESTATURE, in Fortification, a small intrench- ment made of palisadoes, or sacks of earth, with a view to dispute with an enemy the remainder of a piece of ground. ANTHEM, a church song performed in cathedral ser¬ vice by choristers, who sung alternately. It was used to denote both psalms and hymns when performed in this manner; but at present anthem is used in a more con¬ fined sense, being applied to certain passages taken out of the Scriptures, and adapted to a particular solemnity. Anthems were first introduced, in ttfe reformed service of the English church, in the beginning of the reign of Queen Elizabeth. ANTHESPHORIA, in Antiquity, a Sicilian festival instituted in honour of Proserpine. The word is derived from the Greek av^oi, flower, and I carry ; because that goddess was forced away by Pluto when she was gathering flowers in the fields. Yet Festus does not ascribe the feast to Proserpine, but says it was thus called because ears of corn were carried on this day to the temples. Anthesphoria seems to be the same thing with the florifertum of the Latins, and answers to the harvest- home among us. ANTHESTERIA, in Antiquity, was a feast celebrated by the Athenians in honour of Bacchus. The most natu¬ ral derivation of the word is from the Greek ai&oj (flos), a flower, it being the custom at this feast to offer garlands of flowers to Bacchus. The anthesteria lasted three days, the 11th, 12th, and 13th of the month, each of which had a name suited to the proper office of the day. The first day of the feast was called ftfooiyiu, i. e. opening xoj the vessels, because on this day they tapped the vessels and tasted the wine. The second day they called %oos, congius, the name of a measure containing the weight of 10 pounds : on this they drank the wine prepared the day before. The third day they called ypr^ot, kettles : on this day they boiled all sorts of pulse in kettles; which however they were not allowed to taste, as being offered to Mercury. ANTHESTERION, in ancient chronology, the sixth month of the Athenian year. It contained 29 days, and answered to the latter part of our November and begin- 248 ANT ANT Antholo- gion Anthrax. rung of December. The Macedonians called it dasion or desion. It had its name from the festival anthesteria kept in it. ANTHOLOGION, the title of the service-book used in the Greek church. It is divided into 12 months, contain¬ ing the offices sung throughout the year, on the festivals of our Saviour, the Virgin, and other remarkable saints. ANTHOLOGY, a discourse of flowers, or a selection of beautiful passages from various authors. It is also the name given to a collection of epigrams taken from several Greek poets. ANTHONY, Saint, was born in Egypt in 251, and in¬ herited a large fortune, which he distributed among his neighbours and the poor, retired into solitude, founded a religious order, built many monasteries, and died anno 356. Many ridiculous stories are told of his conflicts with the devil, and of his miracles. There are seven epistles extant attributed to him. St Anthony is sometimes represented with a fire by his side, signifying that he relieves persons from the inflam¬ mation called after his name ; but always accompanied by a hog, on account of his having been a swine-herd, and curing all disorders in that animal. To do him the greater honour, the Romanists in several places keep at common charges a hog denominated St Anthony's hog, for which they have great veneration. Some will have St Anthony’s picture on the walls of their houses, hoping by that to be preserved from the plague ; and the Italians, who do not know the true signification of the fire painted at the side of their saint, concluding that he preserves houses from being burnt, invoke him on such occasions. Both painters and poets have made very free with this saint and his followers ; the former by the many ludicrous pictures of his temptation, and the latter by divers epigrams on his disciples or friars. Anthony, or Knights of St Anthony, a military order instituted by Albert duke of Bavaria, Holland, and Zea¬ land, when he designed to make war against the Turks in 1382. The knights wore a collar of gold made in form of a hermit’s girdle, from which hung a stick cut like a crutch, with a little bell, as they are represented in St Anthony’s pictures. St Anthony also gives the denomination to a religious order founded in France about the year 1095, to take care of those afflicted with St Anthony’s fire (see the next ar¬ ticle). It is said that in some places these monks assume to themselves a power of giving, as well as removing, the ignis sacer, or erysipelas; a power which stands them in great stead, for keeping the poor people in subjection, and extorting alms. St Anthony's Fire, a name properly given to the ery¬ sipelas. Apparently it took this denomination, as those afflicted with it made their peculiar application to St An¬ thony of Padua for a cure. It is known that, anciently, particular diseases had their peculiar saints. Thus, in the ophthalmia, persons had recourse to St Lucia; in the toothache, to St Apollonia ; in the hydrophobia, to St Hu¬ bert, &c. ANTHORISMUS, in Khetoric, denotes a contrary de¬ scription or definition of a thing from that given by the adverse party. Thus, if the plaintiff urge, that to take any thing away from another without his knowledge or consent, is a theft; this is called o^o;, or definition. If the defendant reply, that to take a thing away from another without his knowledge or consent, provided it be done with design to return it to him again, is not theft; this is an avdo^idfjMf. ANTHR AX, a Greek term, literally signifying a burn¬ ing coal, used by the ancients to denote a gem, as well as a disease more generally known by the name of carbuncle. ANTHROPOGLOTTUS, among zoologists, an appel-An; of1'' lation given to such animals as have tongues resembling g! I that of mankind, particularly to the parrot kind. | I t ANTHROPOLATRiE, in Ecclesiastical History, an ap-^ T'c pellation given to the Nestorians, on account of their wor-' shipping Christ, notwithstanding that they believed him c to be a mere man. 8 ANTHROPOLATRIA, the paying of divine honours to a man : supposed to be the most ancient kind of idolatry. 11 ANTHROPOLITES, a term denoting petrifactions of the human body, as those of quadrupeds are called zoo- lites. See Petrifaction. ANTHROPOLOGY, a discourse upon human nature, j ^ It is sometimes applied to designate the speculations and inquiries that have obtained concerning the varieties of the human race. See Man. Anthropology, among divines, denotes that manner of expression by which the inspired writers attribute hu¬ man parts and passions to God. * ANTHROPOMANCY, a species of divination, per¬ formed by inspecting the entrails of a human creature. ANTHROPOMORPHA, a term formerly given to the primates of that class of animals which have the greatest resemblance to the human kind. J ANTHROPOMORPHISM, among ecclesiastical wri- * ters, denotes the heresy or error of the Anthropomorphites. i Sgg the next article* ^ ANTHROPOMORPHITES, in Ecclesiastical History, a sect of ancient heretics, who, taking every thing spoken of God in Scripture in a literal sense, particularly that passage of Genesis in which it is said God made man after his oum c image, maintained that God had a human shape. They are likewise called Audeans, from Audeus their leader. ANTHROPOMORPHOUS, something that bears the figure or resemblance of a man. Naturalists give instances of anthropomorphous plants, anthropomorphous minerals, 1 &c. These generally come under the class of what are called lusus natures, or monsters. 1 ANTHROPOPATHY, a figure or expression by which some passion is ascribed to God, which properly belongs only to man. ANTHROPOPHAGI (of av&gurrog, a man, and payw, to eat, Men-Eaters). That there have been, in almost all ages of the world, nations who have followed this barba- 1 rous practice, we have abundance of testimonies. The Cyclops, the Lestrygons, and Scylla, are all repre¬ sented in Homer as Anthropophagi, or man-eaters; and the female phantoms, Circe and the Sirens, first bewitch¬ ed with a show of pleasure, and then destroyed. This, like the other parts of Homer’s poetry, had a foundation in the manners of the times preceding his own. Accord¬ ing to Herodotus, among the Essedonian Scythians, when a man’s father died, the neighbours brought several beasts, which they killed, mixed up their flesh with that of the deceased, and made a feast. Among the Massagetae, when any person grew old, they killed him and ate his flesh ; but if he died of sickness they buried him, esteem¬ ing him unhappy. The same author also assures us that several nations in the Indies killed all their old people and their sick, to feed on their flesh. He adds, that persons in health were sometimes accused of being sick, to afford a pretence for devouring them. According to Sextus Empiricus, the first laws that were made were for. the pre¬ venting of this barbarous practice, which the Greek wri¬ ters represent as universal before the time of Orpheus. Of the practice of anthropophagy in later times, we have the testimonies of all the Romish missionaries who have visited the interior parts of Africa, and even some parts of Asia. When America was discovered, this prac¬ tice was found to be almost universal, insomuch that se- A N T 'tbroi* veral authors have supposed it to be occasioned through a ’ cufflia Want of other food, or through the indolence of the people iS t0 seek for it; though others ascribe its origin to a spirit o tichr i< 0£ revenge. ) 'V^' It appears pretty certain, from Dr Hawkesworth s ac- c ' count of the voyages to the South Seas, that the inhabit- * ants of the island of New Zealand, a country unfurnished ^ with the necessaries of life, eat the bodies of their ene- 11 mies. It appears also to be very probable, that both the ^ wars and anthropophagy of these savages take their rise ft from irresistible necessity, and owe their continuance to d the dreadful alternative of destroying each other by vio- if lence, or of perishing by hunger. Mr Marsden informs us that this horrible custom is P practised by the Battas, a people in the island of Sumatra. “ << They do not eat human flesh,” says he, “ as a means of s satisfying the cravings of nature, owing to a deficiency of 8 other food; nor is it sought after as a gluttonous delicacy, ® as it would seem among the New Zealanders. The Bat- 11 tas eat it as a species of ceremony; as a mode of showing ^ their detestation of crimes, by an ignominious punishment; * and as a horrid indication of revenge and insult to their un- fortunate enemies. The objects of this barbarous repast J are the prisoners taken in war, and oftenders convicted » and condemned for capital crimes.” It may be said, that whether the dead body of an e enemy be eaten or buried, is a matter perfectly indiffer- « ent. But whatever the practice of eating human flesh ” may be in itself, it certainly is relatively, and in its con- * sequences, most pernicious. It manifestly tends to eradi- ( cate a principle which is the chief security of human life, « and more frequently restrains the hand of the murderer than the sense of duty or the dread of punishment. Even i if this horrid practice originates from hunger, still it must be perpetuated from revenge. Death must lose much of its horror among those who are accustomed to eat the < dead; and where there is little horror at the sight of death, there must be less repugnance to murder. ANTHROPOSCOPIA (from avtiguiroe, and 30,000 persons lost their lives. In 634 it fell into the hands of the Saracens, who kept possession of it till the J f year 858, when it was surprised by one Burtzas, and again » annexed to the Roman empire. The Romans continued i masters of it for some time, till the civil dissensions in the ( empire gave the Turks an opportunity of mastering it, as well as the whole kingdom of Syria. From them it was i again taken by the crusaders in 1098. In 1262 it was i > again taken by Bibaris, sultan of Egypt, who put a final | ’ period to its glory. Antioch, called by the Arabs Antahia, is now a ruinous ; town, whose houses, built with mud and straw, and narrow i and miry streets, exhibit every appearance of misery and wretchedness. These houses are situated on the south¬ ern bank of the Orontes, at the extremity of an old de¬ cayed bridge: they are covered to the south by a moun¬ tain, upon the slope of which is a wall built by the cru¬ saders. The distance between the present town and this mountain may be about 400 yards, which space is occu¬ pied by gardens and heaps of rubbish, but presents nothing interesting. Antioch was better calculated than Aleppo to be the emporium of the Europeans. By clearing the mouth of the Orontes, which is six leagues lower down, )oats might have been towed up that river, though they could not have sailed up, as Pococke has asserted, its current being too rapid. The natives, who never knew ANT 253 the name Orontes, call it, on account of the swiftness of Antiochian its stream, El Aasi, that is, the rebel. Its breadth at An- Ij tioch is about forty paces. Seven leagues above that town Antipas. it passes by a lake abounding in fish, and especially in^^^^^ eels. A great quantity of these are salted every year, but not sufficient for the numerous fasts of the Greek Christians. The plain of Antioch, though the soil of it is excellent, is uncultivated, and abandoned to the Turco¬ mans ; but the hills on the side of the Orontes, particu¬ larly opposite Serkin, abound in plantations of figs and olives, vines and mulberry trees. Seleucus Nicator, who founded Antioch, built also at the mouth of the Orontes, on the northern bank, a large and well-fortified city, which bore his name, but of which at present not a single habi¬ tation remains; nothing is to be seen but heaps of rub¬ bish, and works in the adjacent rock, which prove that this was once a place of very considerable importance. In the sea also may be perceived the traces of two piers, which are indications of an ancient port, now choked up. The inhabitants of the country go thither to fish, and call the place Souaidia. Antioch is situate in long. 37. 5. E. and lat. 36. 20. N. ANTIOCHIAN Sect or Academy, a name given to what was called the fifth academy. It took the deno¬ mination from its being founded by Antiochus, a philo¬ sopher contemporary with Cicero. The Antiochian aca¬ demy succeeded the Philonian. As to doctrine, the philo¬ sophers of this sect appear to have restored that of the ancient academy, except in the article of the criterion of truth. Antiochus was really a Stoic, and only nominally an Academic. Antiochian Epocha, a method of computing time from the proclamation of liberty granted to the city of Antioch about the time of the battle of Pharsalia. ANTIOCHUS of Ascalon, a celebrated philosopher, the disciple of Philo of Larissa, the master of Cicero and the friend of Lucullus and Brutus. He was founder of a fifth academy. See Antiochian Sect. ANTIOPE, in fabulous history, the wife of Licus, king of Thebes, who being deflowered by Jupiter in the form of a satyr, brought forth Amphion and Zethus. Another Antiope was queen of the Amazons, and, with the assist¬ ance of the Scythians, invaded the Athenians, and was vanquished by Theseus. J ANTIPJEDOBAPTISTS, (derived from am, against, Kaii, ‘jraiooi, child, and iSwirri^u, I baptize, whence jSaftriGrris), is a distinguishing denomination given to those who ob¬ ject to the baptism of infants; because they say infants are incapable of being instructed, and of making that pro¬ fession of faith which entitles them to this ordinance, and an admission into church communion. ANTIPAROS, an island in the Archipelago, opposite to Paros, from which it is separated by a strait about seven miles over. It is the Oleares or Oliaros mentioned by Strabo, Pliny, Virgil, Ovid, &c.; and was, according to Heraclides Ponticus, as quoted by Stephanus, first peopled by a Phoenician colony from Sidon. According to Mr Tournefort’s account, it is about 16 miles in circumference, and tolerably fertile. This island is remarkable for a subterraneous cavern or grotto, accounted one of the greatest natural curiosi¬ ties in the world. It is 300 fathoms below the surface of the earth, and appears to be about 40 fathoms high and 50 wide, penetrating far into the bosom of the island. It is full of large and beautiful stalactites. There have been many descriptions of this celebrated grotto, of which that by Tournefort is supposed to be very complete and exact. (Relation d’un Voyage du Levant.) ANTIPAS Herod, or Herod-Antipas, the son of Herod the Great, by one of his wives called Cleopatra, a 254 ANT ANT Antipas native of Jerusalem. Herod the Great, in his first will, . II appointed Antipas his successor in the kingdom; but af- Antipater. terwar(js altering that will, he named his son Archelaus his successor, giving to Antipas the title only of tetrarch of Galilee and Peraea. Antipas took a great deal of pains in adorning and for¬ tifying the principal places of his dominions. He married the daughter of Aretas king of Arabia, whom he divorced about the year of Christ 33, to marry his sister-in-law He¬ redias, wife to his brother Philip, who was still living. St John the Baptist exclaiming continually against this in¬ cest, was, by order of Antipas, imprisoned in the castle of Machserus. Josephus says that Antipas caused St John to be laid hold of because he drew too great a concourse of people after him; and that he was afraid lest he should make use of the authority which he had acquired over the minds and affections of the people to induce them to revolt. But the evangelists, who were better informed than Josephus, as being eye-witnesses of what passed, and acquainted in a particular manner with St John and his disciples, assure us that the true reason of imprison¬ ing St John was the aversion which Herod and Herodias had conceived against him for the liberty he had used in censuring their scandalous marriage. The virtue and ho¬ liness of St John were such that even Herod feared and respected him; but his passion for Herodias would have induced him to kill that prophet, had he not been re¬ strained by his apprehensions of the people, who esteem¬ ed John the Baptist as a prophet. (Matt. xiv. 5, 6.) One day, however, while the king was celebrating the festival of his birth with the principal persons of his court, the daughter of Herodias danced before him, and pleased him so well, that he promised with an oath to give her whatever she should ask of him. By her mother’s advice she asked the head of John the Baptist; upon which the king commanded John to be beheaded in prison, and the head to be given her. Aretas, king of Arabia, to revenge the affront which Herod had offered to his daughter, de¬ clared war against him, and overcame him in a very ob¬ stinate engagement. Herod being afterwards detected as a party in Sejanus’s conspiracy, was banished by the emperor Caius to Lyons in Gaul, whither Herodias accom¬ panied him. The time when he died is not known ; how¬ ever, it is certain he died in exile, as well as Herodias. ANTIPATER, the disciple of Aristotle, and one of Alexander the Great’s generals, was a man of great abili¬ ties, and a lover of the sciences ; but was accused of poi¬ soning Alexander. He subdued the revolted Thracians, relieved Megalopolis, and overthrew the Spartans there. He died 321 years before the Christian era. Antipater, an Idumean of illustrious birth, and pos¬ sessed of great riches and abilities, taking advantage of the confusion into which the two brothers Hyrcanus and Aristobulus plunged Judea by their contest for the office of high priest, took such measures as to gain for Hyrcanus that office, and under his government to obtain the abso¬ lute direction of all affairs; while his great abilities and application to business made him so considerable, that he was honoured as much as if he had been invested with the royal authority in form: but he was at last poisoned by a Jew named Malachus, 43 years before the Christian era. He left, among his other children, the famous Herod, king of the Jews. Antipater, Ccdius, a Roman historian, who wrote a history of the Punic war, much valued by Cicero. The emperor Adrian preferred him to Sallust. Antipater of Sidon, a Stoic philosopher, and likewise a poet, commended by Cicero and Seneca. He flourished about the 171st Olympiad. We have several of his epi¬ grams in the Anthologia. ANTIPATHY, in Physiology, is formed from the two Ant Greek words, am, contrary, and naioi, passion. LiterallyCrl jj taken, the word signifies incompatibility: but for the most part the term antipathy is not used to signify such incom¬ patibilities as are merely physical; it is reserved to ex¬ press the aversion which an animated or sensitive being feels at the real or ideal presence of particular objects. In this point of view, which is the light in which we at present consider the term, antipathy, in common language, signifies a natural and involuntary aversion which a sensi¬ tive being feels for some other object, whatever it is, though the person who feels this abhorrence is entirely ignorant of its cause, and can by no means account for it. Such is, they say, the natural and reciprocal hostility be¬ tween the toad and the weasel, between sheep and wolves. Such is the invincible aversion of particular persons against cats, mice, spiders, &c., a prepossession which is sometimes so violent as to make them faint at the sight of these animals. To explore the matter with¬ out prejudice, we shall find it necessary to abstract from this disquisition all such antipathies as are not ascertain¬ ed, as that which is supposed to be felt between the sala¬ mander and tortoise, and between the weasel and the toad. We must abstract those which can be extinguished or resumed at pleasure; those fictitious aversions which only certain persons feel or pretend to feel. When we abstract aversions, the causes of which are known and evident, we shall be surprised, after deduction of pretend¬ ed antipathies, how small, how inconsiderable, is the quantity of those which are conformable to our definition. Will any one pretend to call by the name of antipathy those real, innate, and incontestable aversions which pre¬ vail between sheep and wolves? Their cause is obvious: the wolf devours the sheep, and subsists upon his victims; and every animal naturally flies with terror from pain or destruction: sheep ought therefore to regard wolves with horror, which, for their nutrition, tear and mangle the unresisting prey. From principles similar to this arises that aversion which numbers of people feel against serpents, against small animals, such as reptiles in gene¬ ral, and the greater number of insects. During the cre¬ dulous and susceptible period of infancy, pains have been taken to impress on our minds the frightful idea that they are venomous; that their bite is mortal; that their sting is dangerous, productive of tormenting inflammations or tumours, and sometimes fatal: they have been represent¬ ed to us as ugly and sordid ; as being for that reason per¬ nicious to those who touch them; as poisoning those who have the misfortune to swallow them. Is it then wonder¬ ful (if our false impressions as to this subject have been corrected neither by future reflections nor experiments) that we should entertain during our whole lives an aver¬ sion for these objects, even when we have forgot the ad¬ monitions, the conversations, and examples, which have taught us to believe and apprehend them as noxious be¬ ings ? To explain these facts, is it necessary to fly to the exploded subterfuge of occult qualities inherent in bodies, to latent relations productive of antipathies, of which no person could ever form an idea ? To what then are those antipathies of which we have heard so much reducible? Either to legendary tales, or to aversions against objects which we believe dangerous, or to a childish terror of imaginary perils, or to a disre¬ lish of which the cause is disguised, or to an infirmity oi the stomach,—in a word, to a real or pretended reluctance for things which are either invested or supposed to be invested with qualities hurtful to us. Too much care cannot be taken in preventing or regulating the antipathy of children; in familiarizing them with objects of every kind; in discovering to them, without emotion, such as ANT 'ipith are dangerous; and in teaching them the means of de- T fence and security, or the methods of escaping their iqaitidnoxious influence. v^,‘ Antipathy, in Ethics, hatred, aversion, repugnancy. R Hatred is entertained against persons, aversion and anti- f indiscriminately against persons or things, and repug- ,l( nancy against actions alone. ANTIPELARGIA, among the ancients, a law where- i)] by children were obliged to furnish necessaries to their ag aged parents. The ciconia, or stork, is a bird famous for ^ the care it takes of its parents when grown old ; hence, in some Latin writers, this is rendered lex ciconiaria, or D the stork’s law. ANTIPHONARY, Antiphonarium, a service-book, fl which contained all the invitatories, responsories, collects, M and whatever else was sung or said in the choir, except ^ the lessons. This is otherwise called responsorium, from ^ the responses contained therein. The author of the Ro- 1,1 man antiphonary was Pope Gregory the Great. We also in find mention of nocturnal and diurnal antiphonaries, for ^ the use of the daily and nightly offices ; summer and win- tf ter antiphonaries ; also antiphonaries for country churches, &c. By the provincial constitutions of Archbishop Win- d chelsea, made at Merton A. d. 1305, it is required that ® one of these should be found in every church within the F province of Canterbury. The use of these and many other f popish books was forbid by the 3d and 4th of Edward VI. t c. 10. ANTIPHONY, the answer made by one choir to an- *1 . other, when the psalm or anthem is sung between two. Antiphony sometimes denotes a species of psalmody, * wherein the congregation being divided into two parts, ft repeat the psalms, verse for verse, alternately. In this * sense antiphony stands contradistinguished from sym- phony, where the whole congregation sing together. Antiphony differs from responsorium, because in this ■; latter the verse is only spoken by one person, whereas in the ' former the verses are sung by the two choirs alternately. The original of antiphonal singing in the western churches if is referred to the time of St Ambrose, about the year s 374. That father is said to have first introduced it into the church of Milan, in imitation of the custom of the s eastern church, where it appears to be of greater anti- \ quity, though as to the time of its institution authors are not agreed. It was probably introduced at Antioch be¬ lli tween the years of Christ 347 and 356. Antiphony is also used to denote the words given out a" at the beginning of the psalm, to which both the choirs a are to accommodate their singing. Antiphony, in a more modern sense, denotes a kind of c| composition made of several verses extracted out of dif- ■i ferent psalms, adapted to express the mystery solemnized o on the occasion. ANTIPODES, in Geography, a name given to those in¬ habitants of the globe that live diametrically opposite to !l each other. The word is Greek, and compounded of am, opposite, and wous, a foot, because their feet are opposite to each other. Plato is esteemed the first who thought it 1 possible that antipodes subsisted, and is looked upon as ANT 255 the inventoi of the word. As this philosopher apprehend- Antiquare ed the earth to be spherical, he had only one step to make II to conclude the existence of the antipodes. The ancients Antiquities- in general treated this opinion with the highest contempt, never being able to conceive how men and trees could subsist suspended in the air with their feet upwards, for so they apprehended they must be in the other hemi¬ sphere. They never reflected that these terms upwards and downwards are merely relative, and signify only near¬ er to, or farther from, the centre of the earth, the common centre to which all heavy bodies gravitate; and that, therefore, our antipodes have not their feet upwards and head downwards any more than ourselves, because they, like us, have their feet nearer the centre of the earth, and their heads farther from it. To have the head downwards and feet upwards, is to place the body in a direction of gravity tending from the feet to the head; but this can¬ not be supposed with regard to the antipodes, for they, like us, tend toward the centre of the earth in a direction from head to foot. ANTIQUARE, among Roman lawyers, properly de¬ notes the rejecting of a new law, or refusing to pass it. In which sense antiquating differs from abrogating, as the latter imports the annulling of an old law, the former the rejecting of a new one. ANTIQUARII, a name given to copiers of old books. After the decline of learning among the Romans, and when many religious houses were erected, learning was chiefly in the hands of the clergy, the greater number of whom were regulars, and lived in monasteries. In these houses were many industrious men, who were con¬ tinually employed in making new copies of old books, either for the use of the monastery or for their own emo¬ lument. These writing monks were distinguished by the name of Antiquarii. ANTIQUARY, a person who studies and searches after monuments and remains of antiquity, as old medals, books, statues, sculptures, and inscriptions, and, in general, all curious pieces that may afford any light into antiquity. In the chief cities of Greece and Italy there were persons of distinction called antiquaries, whose business it was to show strangers the antiquities of the place, to explain the ancient inscriptions, and to give them all the assistance they could in this way of learning. Pausanias calls these antiquaries E^yproc/. The Sicilians call them mystagogi. Antiquary is also’ used by ancient writers for the keeper of the antiquarium or cabinet of antiquities. The officer is otherwise called archceota, or antiquary of a king, a prince, a state, or the like. Henry VIII. gave John Leland the title of his antiquary. ANTIQUE, in a general sense, something that is an¬ cient ; but the term is chiefly used by sculptors, painters, and architects, to denote such pieces of their different arts as wrere made by the ancient Greeks and Romans. Thus we say, an antique bust, an antique statue, &c. Antique is something contradistinguished from ancient, which signifies a less degree of antiquity. Thus antique architecture is frequently distinguished from ancient ar¬ chitecture. ANTIQUITIES. Etymologists derive the word antiquities, and many other kindred words, from an obvious source, the Latin preposition ante, before. The great arbiter of language, custom, has ordained that it should mark out and relate ,° Periods of time, not immediately, but long before the buys ot those who make use of it. The early records and older monuments of every na¬ tion, of whatever kind they may be, may properly be named antiquities; but this term ordinarily calls up in the mind the one or the other of the two grand divisions into which the practice of the learned has long continued to separate so much of the wide field as has hitherto 256 ANTIQUITIES. Antiquities.befin carefully cultivated. It either awakens the recol- sion, from the miscellaneous composition of the publica' lections of the Greeks and the Romans, who are com- tion in which it finds a place, it is necessarily confined monly known by the designation of the ancients, and of Not only are the remains themselves numerous and al some other nations, who are remembered chiefly on ac- most infinite, but the works in which they have been re. count of their connection with them, and points out the presented in figures, or described in words, are so bulk} antiquities of the scholar, which are often described by and numberless, that it is not only impossible to give £ the Greek appellation, archeology; or it reminds us of detailed account of them, but it would be vain to en- the history of our ancestors, and of the other European deavour to enumerate each individual. We must content people in the middle ages, which forms the antiquities of ourselves, therefore, with arranging the vast subject un- the antiquary. We are anxious to mark strongly the dis- der several heads, with treating briefly and in general i4 # tinction between archaeology, or the study of the Grecian and Roman antiquities, and the study of the antiquary,— the antiquities of the middle ages and of times subsequent. The former pursuit it is scarcely possible to carry to an improper excess, because we may derive advantage and improvement from almost every department of the stu¬ dies of the archaeologist,—so admirable were the institu¬ tions of Greece and Rome, and to such a wonderful pitch did the cultivation of the human intellect attain, especially in Greece. But in the middle ages, on the contrary, there was little peculiar of which human nature ought not to be ashamed ; and whatever we would not wish to unlearn as to those barbarous times, we would only consent to re¬ member for the purpose of avoiding it in future. It is not to be denied, however, that even in the darkest and rudest of those days of barbarism, several useful and agreeable institutions existed, the loss of which we have abundant cause to deplore, and to regret that they were swept away by ignorant and bigoted men, who, under the specious pretence of reformation, destroyed much that was valuable ; yet it is certain—and to those who are well read terms of each, and with allowing ourselves to dwell £ short time, as an indulgence, on one or two matters thal appear in our eyes to be of paramount importance. If it were possible to explain fully and in a satisfactorj manner all the monuments of antiquity that are at pre¬ sent known, the subject would not be exhausted. Everj year brings important additions to the large stock of an¬ tiques ; and the exposition of them, like the study of na¬ ture, in consequence of new discoveries, is infinite ant inexhaustible. The learned Montfaucon, after much ex¬ perience, expresses, in these remarkable words, his senst of the labours of an archaeologist, which appear to ter¬ minate only that they may commence again: “ II n’esl guere de matiere si vaste que celle des monumens dt 1’antiquite. On voit tous les jours sortir de fobscuritt quelque chose de singulier, et qui n’avoit pas encore etf remarque. Lorsqu’on croit finir ses recueils, on est sou vent oblige de recommencer sur nouveaux frais : a pein< a-t-on acheve un ouvrage, que des materiaux se pre sentent pour un autre. La terre en cache une infinitt que le pur hazard fait decouvrir. On en deterre dans lei in the history of classical antiquity the proofs of the as- champs,^ on en trouve dans les yilles: quand on se flattt d’avoir epuise tous les cabinets, il en sort encore de nou veaux, inconnus quelquefois meme a ceux qui les pos sedent. Presque toutes les parties de 1’Europe en four nissent: le Levant et 1’Egypte nous en envoient tres souvent et de fort curieux ; et ce qui surprend davantage il s’en est rencontre, qui exposes depuis plusieurs siecle a la vue de tout le monde, ont demeure aussi inconnus the worship of Greece and Rome Men of learning, after the most laborious investigation and study of the ancient religion, have usually come t * the conclusion that it is a profound and impenetrabl ' mystery, and that such it was in the time of Cicero, an as far back as the earliest records extend. All the ex sertion are sufficiently familiar—that the institutions which shed a light (if it were but a dim one, it was still a com¬ fort and a guide in a period of darkness, and the ab¬ sence of it has left a void in society) were not the off¬ spring of the middle ages, but were precious relics still remaining, vestiges still unobliterated, ruins still standing, of older and better times—the creations of the wisdom and virtue of the ancient world. The antiquities of the que s’ils avoient 6te caches en terre, jusqu a ce que \t middle ages, so far as they are the peculiar productions of reflexion nous les a fait estimer ce qu’ils valent.’’ those ages, are not wanting in good taste alone—that is a T-he productions ot the fine arts occupy a distinguishei defect comparatively of trifling importance, and it might place among the remains of antiquity; and as these ge more easily be pardoned; but they are deficient in a good nerally are closely connected with the pagan religion, i spirit also. This is undeniably a more serious charge, a will be necessary to premise a few observations respectm more grievous deficiency: they are devoid of a patriotic, a benevolent, a humane intention; they are manifestly without the purpose of benefiting the many, of bringing profit to the people at large, of working the good of man¬ kind in general; they seek to aggrandize some feudal ty¬ rant, it may be sometimes a petty tyrant, but still a ty- rant—to fasten the yoke upon the neck of the vassal—to planations that have hitherto been given are equally ur add another link to his chain ; not, like the happier insti- satisfactory, the astronomical not less so than the res^ tutions of earlier and better times, to instruct and glad- Meanwhile the beauty of the mythology is undoubted den the citizen of a free state. It is chiefly on account 6f this important and remarkable distinction that we give the preference to the study of archaeology over the more vulgar and more modern study of what are most commonly denominated antiquities. We will not affect, , — _ - y. however, to be insensible to the superior allurements of beauty, and was admirably adapted to the wants 01 - the former pursuit, or to be ignorant of the intrinsic ex- imagination, and to satisfy the craving of the fancy ' cellence in all other respects of the relics of ages of ex- human beings of every age and rank. To admit thi traordinary refinement. all of its creations are allegorical, is equally impossifi The consideration of these precious relics is not to be as to deny that some are. This solution indeed oug t undertaken without difficulty. Their number and variety be applied sparingly: a good allegory is no doubt sati would perplex any one who entered upon the delicate factory, but an indifferent one is scarcely tolerable, and ni office of explaining them, even if their nature and uses thing is more tiresome and offensive than allegory or were as obvious in all instances as they are obscure and sustained or frequently repeated. Theorists have use mysterious in many, and if the space that was afforded this key with much confidence, and have sought to Jf for explanation were as unlimited as on the present occa- open by its assistance the sense of the ancient mystene it is like nature, for whose wonderful operations the me dern theories will not account better than the ancien while the excellence of her works cannot be denied ( questioned. It was a collection of fables from all com tries, but it had one characteristic unity, the unity P ANTIQUITIES. 257 ijiti Each department of mythology has been exposed in its turn ^Wto their fruitless attempts, and the origin of the twelve ' signs of the zodiac has been a favourite point of inquiry. Every speculator has found in them a confirmation of his own views, a theory after his own heart, plainly written in the heavens in legible and indelible characters, to unfold to mankind matters of very different natures, and of very unequal importance;—a complete development, as some assert, of the foundation of every mode of religious faith, or a plan of diet, as others affirm, so salubrious as to obvi¬ ate the possibility of disease in the human frame, or of moral evil and consequent misery in the mind of man. We may enumerate amongst mythic curiosities the m French notion, version, or recension, of the ancient my¬ thology, which we see exemplified in their poetry, in their If i til staiyi statues and paintings, and in innumerable engravings, but perhaps most perfectly and fully on the French stage, and especially in the ballets which follow the opera. It is a fantastic, affected, insipid, and strange thing, perfectly un¬ like the original, of which, with a marvellous credulity, an % acute and sceptical nation firmly believes that it is a faithful imitation. The French have the merit of afford¬ ing great encouragements to the study of antiquity, but unhappily they have no feeling for the true antique : they want every quality that is required for comprehending it, and possess all those that can impede the pursuit. They want, in short, nature, ease, grace, and simplicity. The pagan religion was addressed to the imagination and the feelings, and not to the understanding, from which no submission was required: it sought only to amuse and delight, not to convince. If the forms of the gods were sometimes whimsical,—if strange actions were some¬ times attributed to them,—we are told that the object was to attract attention, which ever has been, and ever will be, not less difficult than desirable. The modern practice of endeavouring to obtain hearers by scolding has been found, and ought to be acknowledged, as equally unplea¬ sant and inefficient. Innumerable passages will occur to the scholar, especially in the writings of Cicero, to show that the influence of superstition did not cramp the understanding of the philosopher,—that it was either a plaything or an engine of state, and a part of the powers of government. There is evidence also, that in much earlier times it was not considered necessary to manifest a very lively faith. Valerius Maximus relates, that in the first Punic war, in days of pristine simplicity and primitive piety, when Publius Claudius was consulting the augurs, on being told that the sacred chickens refused to eat, “ Then throw them into the sea,” he said, “ that if they will not eat, they may drink.” If the sacred chickens eat the corn that was placed before them, it was a favourable omen; if they devoured it so greedily as to drop a part, success was certain; if, on the contrary, they refused it, there was mischief in the wind; but if they attempted to fly away on quitting the ivory coop, inevitable destruction was impending. The consultation respected the event of a naval engagement; the profane remark of the consul, therefore, was more apposite : “ P. Claudius, bello Punico pnmo, cum praelium navale committere vellet, auspi- eiaque, more majorum, petiisset, et pullarius non exire cavea pullos nuntiasset, abjici eos in mare jussit, dicens, Quia esse nolunt, bibant.” Notwithstanding the slight influence which the pagan religion possessed over their thoughts and actions, the higher classes were attached to it; great men were glad to be deified, and their relations considered it an honour and a happiness. It was esteemed formerly as advanta¬ geous to have a god, as in Catholic countries it still is to have a saint, in the family. It was not only an addi¬ tion to the splendour of an illustrious house, but pos- vol. in. sibly it also gratified the spirit of monopoly, as nothing Antiquities, was required to go out of the family, not even prayers and sacrifices ; and, moreover, every kind of favour might be reasonably expected from an uncle a god, or an aunt a goddess. Whatever parts of this religion were mys¬ terious, it is at least clear and indisputable that it was essentially and fundamentally of a popular character, and was totally and entirely designed for the public amusement and gratification : it was a political institution for the edu¬ cation of the people. We cannot be surprised, therefore, that it captivated the multitude, who were well aware that it might have been difficult to bestow so much upon them in any other form; and that it was used as a pretext to cover, not taxes and severe exactions, but large, fre¬ quent, and liberal donations from the wealthy to the poor. Among the cheerful ancients, as well as among graver people, the immortal gods were often compelled to furnish a decent excuse to enable men to do without censure whatever they wished: there was this important difference, however, that the religious hypocrisy of the former sought commonly to shelter under the awful sanction of celes¬ tial patronage, objects, not of a narrow and selfish, but of a liberal and patriotic nature. Works of art being for the most part public property, were frequently vested in a god or a goddess, as a trustee for the people ; religion sup¬ plied the means of enjoying the only species of property that can advantageously be enjoyed in common. The temples were in truth the museums in which works of art were preserved and exhibited, and the ancient usage still lingers in countries that have not been reformed, where the churches, although not avowedly, are substan¬ tially" galleries for the display of the productions of the fine arts. Wherever well-stored libraries and rich collec¬ tions of paintings and statues, and similar objects of virtu, are accessible at all hours to the public, the wants of private persons are greatly" diminished. It was not only through these exhibitions, which were constantly present to their eyes, that the people were gratified by the an¬ cient religion,—the occasional and periodical festivals were of frequent recurrence, and were a source of joy and happiness for many ages to all ranks. The subject is an agreeable one: it is pleasant to go back to the en¬ joyments of the ancient world; and we regret that on the present occasion we are only permitted to allude to this interesting topic. M. de Pauw speaks sensibly of the excellent effects of these festive institutions, in the countries where they were most prevalent. “ Au reste, si tous ces petits etats de 1’Attique avoient leurs vices, ils avoient aussi leurs vertus ; et Dicearque lui-meme se loue de 1’humanite et de la politesse avec laquelle il avoit ete re^u sur toutes les grandes routes de cette contree. En- suite il assure que nulle part au monde on ne pouvoit vivre plus agreablement qu’a Athenes; soit qu’on eut de 1’argent, soit qu’on n’en eut absolument pas. Les riches, dit-il, peuvent s’y procurer tous les agremens imaginables, et il-y-a, ajoute-t-il, tant de spectacles, tant de fetes, tant de jeux, tant de divertissemens, que les citoyens indigens n’y pensent jamais a leur pauvrete.” (liecherc/icsPhilosnphiques sur les Grecs.) All who have experienced the oppo¬ site condition, who have resided in countries where there are no public entertainments, no festivals, no popular amusements, no holydays, are able to bear testimony, that there wealth can procure for the rich few pleasures, and that the many are reminded of, and never see cause to forget, their indigence. Festivals are very useful to the lower classes, especially in country places, as they tend to check the selfishness of individuals, by which their minds are apt to be cramped, and make them feel that they form a part of a great whole : humanity and politeness are want¬ ing wherever they are discouraged, and an absence of 2 K ANTIQUITIES. Antiquities, amusements is sure to generate a brutal and sullen inci- vilitj. The cheerfulness and courtesy of the Athenians were the fair fruits of a system of reasonable indulgence; and intellectual culture being generally diffused, caused an extreme toleration of all pursuits and opinions. If the people are enabled to judge of works of art, their critical powers will be engaged and exercised more usefully and agreeably than in scanning the actions of their superiors, which they are unable to estimate correctly. Where no congenial occupation is supplied, every low ami ignorant fellow may grow into a sour bigot, and, it may be, into an oracle and a prophet, and in that capacity may ima¬ gine that it is his duty to seek to shape the conduct and creed of wiser men according to the misconceptions of his wayward fancy. It is very desirable that some competent person should undertake to give a plain and faithful account of a religion which has never met with fair treatment from its histo¬ rians. It indicates a bad taste, to say the least of it, to imitate and repeat the absurd and vehement declamations of the fathers of the Christian church against the gods of the ancients. In their times those deities were danger¬ ous rivals; there is certainly no cause to fear them now. No one can see any reason to apprehend that the world will return again to the old worship. The pope and the archbishop of Canterbury may rest in equal security that they will never receive a short notice either to sacrifice a black heifer to Proserpine, or to vacate their respective palaces and offices; the unpleasant option will never be offered to either. It is a mere waste of words, there¬ fore, to persist in calling the heathen, of whom we have learned so much, blind, and to rail at their gods. A religion which was exempt from sordid rites, and of which the end and object were the gratification and cultiva¬ tion of the people, at least deserves to have its story told and its fortunes related without scurrility, passion, or prejudice. Persons of good taste will be equally of¬ fended, on the other hand, at the attempts that^are often made by writers, of more zeal than discretion, to wrest the pagan mythology, and to compel it to seem to con¬ sent with the mysteries of the Christian religion. Lord Bacon wisely says, in allusion to such unholy and of¬ fensive practices, “ But I have interdicted my pen all liberties of this kind, lest I should use strange fire at the altar of the Lord.” Nor should that excessive dread of idolatry, which characterizes a barbarous people, find a place in such a work: it is time to discard the absurd no¬ tion, that it is impossible to see and admire a beautiful statue without worshipping it—that true piety and a taste for the fine arts are of necessity incompatible. The admirable Heyne complains of the unnatural and forced hypotheses of mythists, and of their disposition to torture the meaning of the ancient fables, and to pervert them to signify matters that were diametrically opposed to the spirit and sense of antiquity. He at the same time lays down a golden rule, to guide the future historian of mytholpgy; and happy will he be who shall have the good sense and the good fortune to be governed by it. “ To compose a plain narrative, representing the primitive form under which each fable has been handed down to us by the first poets and the first artists, and afterwards to show the changes and additions that have been made by later poets and artists.” “ II nous manque egalement pour 1’etude dont il est question (the study of antiquity), des traites sur les connoissances accessoires, et surtout, un bon ouvrage sur la fable. Nous avons a la verite beaucoup d’ecrits sur la mythologie; mais je ne sais quel mauvais genie se saisit de tous ceux qui veulent traiter cette matiere. Ils commencent par etablir quelque hypo- these, d’apres laquelle ils cherchent a denaturer et a forcer le sens des anciennes fables; et aucune do ces hypo-Anty.# theses ne se trouve etablie sur 1’esprit meme de I’anti-^vy % quite. II nous faut une mythologie qui ne consiste qu’en un simple recit, qui nous presente la forme primitive sous laquelle chaque fable nous a ete transmise par les pre¬ miers poetes et par les premiers artistes. Nous devons connoitre aussi les changemens et les additions que les artistes et les poetes posterieurs y ont faits. La meilleure explication est celle qu’on pent tirer de cette methode et de cette maniere d’exposer les choses, en partant des terns de la premiere origine des fables, et en les suivant dans les diiferens changemens qu’elles ont eprouvees.” ■ The inexhaustible quarries of fine marble at Athens,^, and in other parts of Greece, furnished materials for innu-ofGi, merable statues and other sculptures: many of the mostandu celebrated works were executed in ivory. ihese, ofniail> course, have entirely perished; but of those in marble we are so fortunate as to possess some specimens from the hands of the most renowned artists, and it is universally allowed, that even in their present state, all of them hav¬ ing received more or less injury, they are fully worthy of the fame that has attended on the memory of the great men by whom they were fashioned. It has been asserted, that the most lovely of all things are the Greek statues and the Greek tragedy; and ingenious, and perhaps somewhat fanciful persons, have imagined that they per¬ ceived a certain resemblance in the peculiar style ol the sculptors and tragedians of Athens. The sentiment of repose is strongly inculcated by the best productions of the Grecian chisel, and critics are sometimes dissatisfied with the small amount of action and the little apparent exertion ; and they affirm that the expression is occasionally even that of listlessness. In inferior productions, on the contrary, we observe an ex¬ cess of action, and a straining, that are painful to the spec¬ tator, especially if he regards them for a long time; we find the same sobriety, frugality, and even parsimony, in the other compositions of the ancients, especially in their writings, which must seem cold and languid to the admirers of the exaggerated expressions of modern authors. In classical works, the more exalted the personage, the more calm and dignified are his emotions; but the hero of a modern drama or other work of the imagination often seems, by his violence and unnatural contortions, to be convulsed by the powerful action of a Galvanic battery, rather than impelled and agitated by mere human feel¬ ings ; unless we may adopt a more obvious and simple solution of the difficulty, and suppose him to be a maniac. We presume to speak on this subject with great diffidence and distrust, but we must confess that the ancients have sometimes carried the sentiment of repose to an excess, especially in has reliefs, in which the figures, although they are engaged in pursuits demanding the utmost enei- gy, and of intense interest to all concerned, appear fre¬ quently to be more than half-asleep. It cannot be de¬ nied, however, that an excess of tranquillity is less dis¬ pleasing than a display of unnecessary and outrageous violence. • c I The nakedness and full draperies of ancient times form a striking contrast with the close, strict, and scanty clothing of modern days, and show us symbolically the former copiousness anil pristine candour, in opposition to the disingenuous concealments and narrow poverty o degenerate souls. Statues and other marbles and sculp¬ tures may be so easily, cheaply, and perfectly copied, by means of plaster-casts, that no considerable city ought to be without a public museum, containing a copy of all the best pieces of this kind in existence. If a few churhs persons should for a time refuse permission to take a cast, a drawing or some other imitation of the sculpture tha * ANTIQUITIES. 259 qnkifiWas wanting might occupy the place of the cast, and an .inscription might inform the visitor where the original was to be found, and give the name of its unworthy possessor, who refused to the lovers of art the facilities which others had cheerfully granted. We possess many excellent an¬ tiques in bronze. If the colour be less agreeable than that of marble, the superior toughness of the material ad¬ mits of greater spirit and freedom of execution; and of this advantage the ancient workmen knew how to avail themselves to the utmost, as any good collection of these remains will fully demonstrate, especially the many ad¬ mirable specimens that are exhibited in the Studii at Naples. ... The most remarkable, and perhaps the most inimitable, excellence of ancient art, i& the perfection that was dis¬ played in cutting gems, whether the figure be sunk be¬ neath the plane surface of the stone, as in the intaglio, or projected above it, as in the cameo. If, on the one hand, gems are imperishable, and almost indestructible, on the other they are easily lost by reason of their small size. The prodigious number of engraved stones already known to the curious, and which is continually- augmented by the discovery of new and beautiful speci¬ mens, proves that a vast amount of labour and of exquisite skill was formerly employed on these minute and delicate works. Rings, which were originally composed of coarse materials, and were used as seals, tokens, and for other purposes of necessity, in progress of time gradually grew into articles of immense luxury and cost. The taste for these ornaments had become universal in the civilized world in the time of Alexander the Great, who was fond of having his portrait cut on gems; and men were equally fond of wearing his image, not through a servile adulation, for the fashion or passion continued for several centuries after his death, but from a notion that it was lucky, which was indeed the chief motive in the choice of subjects for rings. In modern times we waste our gems by engraving upon them the ugliest device of a barbarous age—coat- armour ; in like manner much valuable material and much precious labour were expended anciently, in the days of waning liberty, to preserve the worthless portrait of some dull sample of a Jove-descended king. The public acts of free states had sometimes been sealed with the likeness of an individual who had been a benefactor to the com¬ munity, as a lasting and delicate compliment, that might flatter, but could not wound, the generous modesty of a disinterested patriot. The emperors of Rome, with that more than Aristophanic genius for burlesque and carica¬ ture for which they were remarkable, and of which the comic effect was the more striking and humorous, be¬ cause the masters of the world were perfectly unconscious that they were heaping ridicule upon their own sacred heads, adopted the usage with many aggravations and ad¬ ditions for their own glory. The patriotic Tiberius ex¬ tended the laws of treason—laws by which it was declared to be high treason to treat with disrespect the statues of the emperor—in order to protect from insult, by the last and highest penalties, his own image on a ring. We de¬ sire to confine our censure to the abuse of portraits. Many of the antique likenesses of individuals, who are remem¬ bered on account of their virtues or their talents, are highly interesting: there are several valuable collections of this kind in other places, but the Capitoline Museum at Rome is the grand repertory of ancient portraits. We read that in ancient times there were large collections of engraved gems, both glyphs and anaglyphs, as in our own days; and we have moreover innumerable books, contain¬ ing representations of the most celebrated Dactyliothecce, ot very various degrees of merit, to enumerate which would be tedious. The amulets and astrological charac¬ ters on rings have given occasion to the display of much Antiquities, curious learning. Superstition has long been familiar with the importance of rings: Apollonius of Tyana deemed them so essential to quackery, that he had seven—for each day of the week a different one, marked with the planet of the day. The scenic mask, whether tragic or comic, but especially the latter, was a favourite device : we have seen many gems that exhibit beautiful and sur¬ prising examples of that wonderful creation of the fancy. It affords one instance also of the many that might be cited to show that ancient forms are often preserved in a whimsical manner, and applied to purposes very remote from those for which they were originally designed. We have often observed an antique comic mask carved in stone on the end of a spout on our churches, and the exagge¬ rated mouth from which jests not less grotesque than it¬ self, but well seasoned with Attic salt, were used to issue, giving free vent to mere rain-water: so universal and in¬ destructible is the influence of antiquity. As a class, coins are perhaps the least interesting of antiques; for, v/ith few exceptions, they serve to illustrate nothing but the succession of a family or dynasty of kings. They have, however, long been favourite objects with collectors, partly on this account, and partly because persons who can¬ not attain to the comprehension of any other part of the study of antiquities, can at least understand the formation of an unbroken series, and that it is often difficult to com¬ plete it. We may concede also in favour of this branch of the science, that the Numismatists have sometimes rendered assistance to chronology, and have even cleared up certain points of geography, and therefore in subordi¬ nate departments have aided the historian. Certain eminent artists and accomplished critics have given it as their decided opinion, that enough remains of the ancient painting to demonstrate that the ancients were as much superior to the moderns in this art, as they are admitted to have been in sculpture. It seems proba¬ ble that the first efforts of design were mere shadows or outlines ; of this early style the fictile vases afford exam¬ ples. We have, however, many specimens of finished works in fresco, which are preserved at Rome and in Naples. Of the former, the Roman remains, some idea, although, it must be confessed, a very inadequate one, may be de¬ rived from the engravings in Turnbull’s Treatise on Ancient Painting. Of the latter, the Pitturedi Ercolano are more faithful and satisfactory representations. The style of the ancient painters, so far as our imperfect materials will permit us to judge, rather resembled that of Perugino and Raphael in grace, beauty, and sweetness, than the subli¬ mity and graphic bavorrn of the angelic and immortal Michael. Michel piu che mortal, Angiol divino. Of the celebrated encaustic painting we know nothing, but that the ancients have left us some brief and vague descriptions, and the moderns have made some fruitless attempts to revive or re-discover it. Of the Mosaic painting we have some very lovely ves¬ tiges, and some of them are as fresh and as fair as when they were first laid; with patient industry they may be imitated, either with stones, with morsels of glass, or with small tiles coloured and glazed. This kind of work is of bewitching beauty : the eyes of Homer had been so cap¬ tivated with it, that he continued after his blindness to describe heaven by its pavement; at least if the critics will permit us to give this sense to the word bumbov. We learn from the remains of ancient painting, that it was usual to adhere scrupulously to particular colours for the draperies of certain gods and heroes; and we may trace the same practice in the works of many of the Italian masters, who assign invariably the appropriated colours to 260 ANTIQUITIES. Antiquities, the clothing of saints and personages of note in Christian In countries where fuel was scarce the process of burn¬ ing bricks was expensive ; unburnt bricks were therefore much used. In consequence of the greater heat of the sun they were hardened in southern climates more perfectly than the very limited portion of sunshine that northern regions enjoy could possibly effect; and we read that the desiccation was continued for several years before they wrere consigned to the builder. If the rain, which in the south at certain seasons falls in torrents, were warded off, wherever the air is generally dry such structures would last for a long period (the perpetual humidity of our at¬ mosphere, even if we were able to prepare them properly, would speedily decompose them) ; but, even under the most favourable circumstances, a pile of unburnt bricks is a perishable structure, and many celebrated edifices have accordingly perished. More durable and more ornamen¬ tal materials, however, have been fortunately very plenti¬ ful in those countries where invention and a pure taste in architecture were most prevalent; and some of the most celebrated of the productions of antiquity have re¬ sisted the gradual attacks of time, and the sijdden vio¬ lence of barbarians. In a few instances they are nearly entire ; in many more, enough remains to enable us to re¬ store the building to its original state, or to build another exactly similar to the former. Nor have architects been wanting in diligence in this respect; very accurate sur¬ veys, and exact and minute admeasurements and drawings, models and descriptions of every kind, have been exe¬ cuted by competent persons, to the great benefit of their art. We should be enabled to judge very correctly of the effect of the ancient temples and other edifices of celebri¬ ty, many of which have been admirably restored in small drawings and engravings by architects, if the restorations were painted in fresco on the walls of our public buildings in various points of view, and of the full size of the origi¬ nals. Since Greece, and especially Athens, has been so carefully explored, and the results of much valuable labour made public, our taste in architecture has been greatly improved. The various members, at least, and the details of the buildings that have been lately erected in Great Britain, are more elegant than they were formerly. Un¬ fortunately, however, we do not seem to possess an archi¬ tect capable of combining the beautiful parts into one har¬ monious whole. It should appear, therefore, that a more profound and enlarged study of antiquity is required to make the artist in this line a master of general effect. A literary work may be faultless in its details, and yet it may be inefficient as a whole ; it may be impossible to fix blame upon any single page or paragraph, yet the entire history of Greece, or of Rome, may be unworthy of the events which it undertakes to record. The narration, al¬ though the style be chaste and correct, may be lifeless, spiritless, and uninspiring. So is it in architecture; and such is the present state of that art in Great Britain. Our architects seem equally incapable of producing a whole, either in the Grecian or the Gothic style, although the parts may sometimes in themselves have merit. A sil¬ houette, shade, or profile of the object in question, espe¬ cially if it be taken in various points of view, will afford the most simple, ready, and satisfactory means of deter¬ mining, and is the most certain and conclusive test whe¬ ther the effect as a whole be good. If our latest erec¬ tions be tried in this manner, the contours will invariably prove to be insipid and uninteresting, and often ill propor¬ tioned and ugly. The Romans surpassed the rest of mankind less in the arts of government than in that remarkable art which Se¬ neca aptly compares to the constitution of civil society, the !#* than it is possible to convey by verbal description alone, in his great work entitled Histoire de l'Art par les Mo- namens depuis sa Decadence d Ame Siecle jusqud son Renouvellement d If)mg. Paris, 1823. The same disposition of stones, and the same artifices, which would serve to support more securely the roof of a temple of masonry, would be equally effectual in diminish¬ ing the tendency of the roof to fall in a cavern or excavat¬ ed temple; and we see, in fact, that they were actually employed in the latter case, and with results equally sa¬ tisfactory. The ordinary configuration of the artificial roof of a temple is well adapted to lighten the natural roof of a cavern, especially if the peculiarities of forma¬ tion be carried to a greater extent than we usually find in edifices. The part that would fall in first, and bring down the rest with it, is the middle, it being most distant from the supports: this is cut away and formed into hol¬ low pannels, deepening in gradual succession. The Etrus¬ can catacombs supply remarkable instances of all these contrivances. We even find the hole in the centre of the cupola, which is designed to lighten the living roof, where it is least able to hang in the air, and which also serves to admit the day. Of the two problems to support a natural and an artificial roof, the one is precisely the converse of the other; it being necessary in the former to hew away whatever portion of space would not be occupied by the structure in the latter. Certain architectural critics at¬ tribute, we know not how correctly, the undercrofts or crypts which are commonly found beneath cathedral churches, to the catacombs; of which, they assert, they are an imitation and memorial; because the catacombs were, it is said, the first places of worship used by the early Christians. It is certain that the catacombs at Rome serve as a crypt to the church of St Sebastian, and it is also certain that the sepulchral chambers of the Pagans, building of arches and vaulted roofs of masonry. TheAmUi astonishing and colossal relics of their mastery in this use-^Jy ful and difficult department of architecture have received much Jess attention than many works of very inferior beauty and utility. The most vulnerable part of a building is the wooden frame of the roof, which is always in danger of being destroyed by fire. It seems, however, to be unnecessary in the construction of edifices of some kinds to use wood at all. The tiles or other external covering might be support¬ ed, as they were in some of the ancient temples, by light arches of masonry, which, as the weight they would have to bear would be insignificant, might be built so slightly as not to oppress the parts beneath that sustain them. Many of the arches in the circus of Caracalla, near Rome, are composed of large earthen vessels walled together, instead of bricks or stones; and of this contrivance there are other examples at Rome. The superior lightness is manifest; the strength of an earthen vessel, and its power of resisting mere pressure, is considerable. That such structures were sufficiently solid, is demonstrated by their standing to this day. The expedient is worthy of imi¬ tation, and of more attention than it has hitherto re¬ ceived. The cupola of the church of San Vitale at Ra¬ venna is the most remarkable specimen of this sort of building, being composed entirely of cylindrical earthen vessels, which are placed in a horizontal position, and are so arranged as to form one spiral coil, the end of the one vessel being always inserted into the mouth of the other, like the pipes that convey water through our streets. This very curious structure is exactly coeval with the body of the civil law, being of the age of the emperor Justinian ; and it is said to be in equally good repair, and as likely to last for some ages longer, as the Corpus Juris. M. d’Agincourt has fully explained this simple but inge- contrivance, and has expressed in a plate, more ANTIQUITIES. 261 ^Lultieiwhieh are far more ancient than the Christian worship, al- rv,though they were never built under the floor of temples, closely resemble crypts. The tombs of the ancients are very interesting to the archaeologist, because, besides a multitude of pots and pans, and some vases of merit, they have furnished a countless host of lamps, that have formed the subject of many amusing volumes, and some excellent paintings in fresco, and other objects of considerable importance. The ancients sought to alleviate as much as possible, by calling up ideas of cheerfulness, grace, and beauty, the heavy burthen of death. The Greeks were emi¬ nently distinguished by the warmth and the strength of the domestic affections. Their tragedies present many lovely pictures of the vigorous and luxuriant growth of all the more tender charities, and every part of the history of their rites of sepulture brings fresh proof of the power and prevalence of the most amiable feelings among this ingenious and cultivated people. The crea¬ tion of families, and the maintenance of them in the closest union and intimacy, is undoubtedly the most desirable object of public and private institutions ; for it is from his family in childhood and in youth, in manhood and in old age, that man’s happiness is mainly derived. We must be careful, however, not to mistake the means for the end, and we must always remember that laws, institu¬ tions, and principles, that were designed to advance an end, are only valuable so far as they serve the purpose for which they were designed. Men who do not steadily look to final causes are apt to gild and hang garlands on a scaffolding pole, and, in their insane worship of their wooden idol, to forget the marble palace, for the sake of which alone the unsightly pole was erected. The great success of the Greeks in the cultivation of these affec¬ tions may be principally attributed to their extraordinary toleration and liberality; for, in other countries, where there is more prudish precision than mild and temperate forbearance, by drawing the silken bonds too tight, they are often broken. If those rules of decorum which are to be observed only with a certain moderation, and ought often to be considerably relaxed with a wise equity, are enforced by the arbiters of society with a rigid and literal severity, they will infallibly create heartlessness, hypo¬ crisy, and disgust, and there will be but little domestic happiness or affection. Genius is in its nature eccentric ; and if no allowances be made for its aberrations, men of talent will be converted into enemies,—and they are most formidable ones. If the treatment that may not be un¬ suitable to the humble and patient ass be attempted to¬ wards the generous steed, it will rouse a spirit of resist¬ ance and revenge not unworthy of the lion. Many writers have discoursed at great length of the ancient sepulchral rites. The folio of Mark Anthony Boldetti, entitled Os- servazioni sopra i Cimiterj de Santi Martiri, ed antichi Cristiani di Roma, contains a great body of curious infor¬ mation, and many valuable inscriptions ; and the compila¬ tion is made palatable by a certain simplicity and amusing credulity. It is difficult to believe, if we reason from analogy, that were it possible to restore the ancient system of music, it would not richly repay the labour that might be bestowed on the task. It is not from the contemplation of the remains of anti¬ quity in the fine arts alone that we may hope to derive benefit; in many of the mechanical arts the ancients were decidedly our superiors; and by assiduous study of the specimens of various articles, and of the descriptions of certain processes, we may hope that ingenious practical men will learn to restore lost arts, and to amend and im-Antiquities, prove those which we possess. The ancient locks, forv-^^v-y example, were of an admirable and very various construc¬ tion. Bramah’s justly celebrated inventions are merely, as is commonly known, an adoption of as many of the de¬ vices of the ancient locksmiths as modern artists are at present able to understand. In the manufacture of glass, the ancients were as much our superiors as in several other arts. This is demonstrated by the accuracy, variety, and delicacy of the forms into which they have moulded ves¬ sels of this substance. Among the other excellencies of this manufacture, we may mention the gems formed of paste in imitation of stones, to which we owe the preser¬ vation of some of the most beautiful antique engravings, the originals having been lost: many of these are of a large size. The manufacturers of porcelain have not hitherto been very successful in impressing their works with the stamp of genius and true taste ; the Chinese origin is still manifest in the shapes of even the best pieces of crockery: herein let them pray the aid of the ancients. The con¬ struction of the antique chariots is perhaps not altogether unworthy of our attention, for even the smallest and light¬ est of our carriages appear to be unnecessarily large and heavy. Among the minor difficulties that were master¬ ed by the skill of antiquity, was that of driving and ma¬ naging many horses abreast. We see on a gem in the work of Count Caylus {Recueil d'Antiquites, tome i. Plate LX. fig. 4) a chariot with 20 horses yoked in this manner. Three horses abreast are often used on the Continent, and with a good effect. We have heard that an equestrian amateur tried the experiment in England successfully, but we do not remember to what extent. The Grecian breed of horses, it is believed, was much smaller than the English carriage-horses. If a London drayman, therefore, would be surprised to meet an Athenian team rapidly ad¬ vancing in a single line, the Attic charioteer would scarcely be less astonished at the immense bulk of the animals that slowly march in a long file. This branch of the subject, however, would lead us too far from our original design. It would be dangerous to enter upon the disputed ground of Etruscan art; for even the most experienced and confident critics confess that it is sometimes very diffi¬ cult, if not impossible, to distinguish Etruscan monuments from those of the early ages of Greece. It is certain, how¬ ever, that this ancient nation was much addicted to super¬ stition and the fine arts. The lax erudition of Dempster,1 the learning and critical skill of Lanzi,2 and the labours of a myriad of antiquaries, and particularly of the estim¬ able Gori,3 are ready to conspire in instructing and per¬ plexing the student. Archaeological critics are by no means agreed as to the Remains of degree of merit that is to be ascribed to the Egyptian Egyptian works of art. Artists unanimously admit that the mechani-31,1, cal execution is very admirable ; and travellers assert with one voice, that we cannot safely and correctly judge of their peculiar character or general effect from the ordi¬ nary engravings. We are required, moreover, in order to arrive at sound and solid conclusions on this difficult ques¬ tion, to distinguish most carefully between the original and pure Egyptian works, and the modern and spurious imi¬ tations that were manufactured when the rites of Isis were fashionable in Rome ; and generally to be on our guard against whatever was executed whilst Egypt was under the sway of the Romans, and to esteem it as very inferior to the genuine productions of the country. A still further degree of caution may perhaps be necessary: the degradation of the old style probably was gradually pre- 1 De Etruria liegali, 2 vols. fol. Saggio di Lingua Etrusca, 3 vols. 8vo. s Museum Etruscum, 3 vols. fol. 262 ANTIQUITIES. Antiquities, pared, and advancod slowly during the period of more than words are confined to some particular temple, which he A three centuries, when Egypt was under the Grecian domi- criticises : they seem, however, to. admit ot a general apA| nion of the Ptolemies, before it passed by conquest into plication, and they express the opinion of many. Effr; cn the hands of the Romans. Plato indeed informs us, at the ns KoXvaroXos or/.og xa^tx-ig sv Me/xSe/, fiagfiagiitriv iyuv ry^ commencement of his second book De Legibus, that the x-aratfxsuTjv n'hnv ya% tcj gzyaXm umi, xa; xoXkuv, xa/ coXu, Egyptian sculptors and painters were forbidden by law to enyon,