id sid tid token lemma pos 37602 1 1 _ _ NNP 37602 1 2 OCCASIONAL OCCASIONAL NNP 37602 1 3 PAPERS PAPERS NNPS 37602 1 4 _ _ NNP 37602 1 5 of of IN 37602 1 6 the the DT 37602 1 7 MUSEUM MUSEUM NNP 37602 1 8 OF of IN 37602 1 9 NATURAL NATURAL NNP 37602 1 10 HISTORY HISTORY NNP 37602 1 11 The the DT 37602 1 12 University University NNP 37602 1 13 of of IN 37602 1 14 Kansas Kansas NNP 37602 1 15 Lawrence Lawrence NNP 37602 1 16 , , , 37602 1 17 Kansas Kansas NNP 37602 1 18 --------------------------------------------------------------- --------------------------------------------------------------- : 37602 1 19 NUMBER number NN 37602 1 20 1 1 CD 37602 1 21 APRIL APRIL NNP 37602 1 22 29 29 CD 37602 1 23 , , , 37602 1 24 1971 1971 CD 37602 1 25 --------------------------------------------------------------- --------------------------------------------------------------- NFP 37602 1 26 A a NN 37602 1 27 SYNOPSIS synopsis NN 37602 1 28 OF of IN 37602 1 29 NEOTROPICAL NEOTROPICAL NNP 37602 1 30 HYLID HYLID NNP 37602 1 31 FROGS FROGS NNP 37602 1 32 , , , 37602 1 33 GENUS GENUS NNP 37602 1 34 _ _ NNP 37602 1 35 OSTEOCEPHALUS OSTEOCEPHALUS NNP 37602 1 36 _ _ NNP 37602 1 37 By by IN 37602 1 38 LINDA LINDA NNP 37602 1 39 TRUEB[1 TRUEB[1 NNP 37602 1 40 ] ] -RRB- 37602 1 41 AND and CC 37602 1 42 WILLIAM WILLIAM NNP 37602 1 43 E. E. NNP 37602 1 44 DUELLMAN[2 DUELLMAN[2 NNP 37602 1 45 ] ] -RRB- 37602 1 46 [ [ -LRB- 37602 1 47 1 1 CD 37602 1 48 ] ] -RRB- 37602 1 49 Research Research NNP 37602 1 50 Associate Associate NNP 37602 1 51 , , , 37602 1 52 Division Division NNP 37602 1 53 of of IN 37602 1 54 Herpetology Herpetology NNP 37602 1 55 , , , 37602 1 56 Museum Museum NNP 37602 1 57 of of IN 37602 1 58 Natural Natural NNP 37602 1 59 History History NNP 37602 1 60 , , , 37602 1 61 University University NNP 37602 1 62 of of IN 37602 1 63 Kansas Kansas NNP 37602 1 64 . . . 37602 2 1 [ [ -LRB- 37602 2 2 2 2 LS 37602 2 3 ] ] -RRB- 37602 2 4 Curator Curator NNP 37602 2 5 , , , 37602 2 6 Division Division NNP 37602 2 7 of of IN 37602 2 8 Herpetology Herpetology NNP 37602 2 9 , , , 37602 2 10 Museum Museum NNP 37602 2 11 of of IN 37602 2 12 Natural Natural NNP 37602 2 13 History History NNP 37602 2 14 , , , 37602 2 15 University University NNP 37602 2 16 of of IN 37602 2 17 Kansas Kansas NNP 37602 2 18 . . . 37602 3 1 When when WRB 37602 3 2 we -PRON- PRP 37602 3 3 initiated initiate VBD 37602 3 4 a a DT 37602 3 5 study study NN 37602 3 6 of of IN 37602 3 7 the the DT 37602 3 8 herpetofauna herpetofauna NN 37602 3 9 at at IN 37602 3 10 Santa Santa NNP 37602 3 11 Cecilia Cecilia NNP 37602 3 12 in in IN 37602 3 13 Amazonian Amazonian NNP 37602 3 14 Ecuador Ecuador NNP 37602 3 15 in in IN 37602 3 16 1966 1966 CD 37602 3 17 , , , 37602 3 18 we -PRON- PRP 37602 3 19 were be VBD 37602 3 20 immediately immediately RB 37602 3 21 confronted confront VBN 37602 3 22 with with IN 37602 3 23 many many JJ 37602 3 24 kinds kind NNS 37602 3 25 of of IN 37602 3 26 animals animal NNS 37602 3 27 that that IN 37602 3 28 we -PRON- PRP 37602 3 29 could could MD 37602 3 30 not not RB 37602 3 31 identify identify VB 37602 3 32 with with IN 37602 3 33 the the DT 37602 3 34 existing exist VBG 37602 3 35 literature literature NN 37602 3 36 . . . 37602 4 1 Comparisons comparison NNS 37602 4 2 of of IN 37602 4 3 our -PRON- PRP$ 37602 4 4 specimens specimen NNS 37602 4 5 with with IN 37602 4 6 those those DT 37602 4 7 preserved preserve VBN 37602 4 8 in in IN 37602 4 9 other other JJ 37602 4 10 museums museum NNS 37602 4 11 resolved resolve VBD 37602 4 12 some some DT 37602 4 13 of of IN 37602 4 14 the the DT 37602 4 15 problems problem NNS 37602 4 16 , , , 37602 4 17 but but CC 37602 4 18 many many JJ 37602 4 19 identifications identification NNS 37602 4 20 could could MD 37602 4 21 be be VB 37602 4 22 made make VBN 37602 4 23 only only RB 37602 4 24 after after IN 37602 4 25 study study NN 37602 4 26 of of IN 37602 4 27 type type NN 37602 4 28 specimens specimen NNS 37602 4 29 ; ; : 37602 4 30 even even RB 37602 4 31 then then RB 37602 4 32 some some DT 37602 4 33 determinations determination NNS 37602 4 34 remained remain VBD 37602 4 35 questionable questionable JJ 37602 4 36 . . . 37602 5 1 We -PRON- PRP 37602 5 2 now now RB 37602 5 3 find find VBP 37602 5 4 that that IN 37602 5 5 in in IN 37602 5 6 order order NN 37602 5 7 to to TO 37602 5 8 prepare prepare VB 37602 5 9 a a DT 37602 5 10 meaningful meaningful JJ 37602 5 11 account account NN 37602 5 12 of of IN 37602 5 13 the the DT 37602 5 14 herpetofauna herpetofauna NN 37602 5 15 of of IN 37602 5 16 Santa Santa NNP 37602 5 17 Cecilia Cecilia NNP 37602 5 18 , , , 37602 5 19 we -PRON- PRP 37602 5 20 must must MD 37602 5 21 complete complete VB 37602 5 22 several several JJ 37602 5 23 taxonomic taxonomic JJ 37602 5 24 studies study NNS 37602 5 25 , , , 37602 5 26 the the DT 37602 5 27 limits limit NNS 37602 5 28 of of IN 37602 5 29 which which WDT 37602 5 30 extend extend VBP 37602 5 31 far far RB 37602 5 32 beyond beyond IN 37602 5 33 eastern eastern JJ 37602 5 34 Ecuador Ecuador NNP 37602 5 35 . . . 37602 6 1 Because because IN 37602 6 2 of of IN 37602 6 3 our -PRON- PRP$ 37602 6 4 interests interest NNS 37602 6 5 in in IN 37602 6 6 hylids hylid NNS 37602 6 7 we -PRON- PRP 37602 6 8 have have VBP 37602 6 9 begun begin VBN 37602 6 10 our -PRON- PRP$ 37602 6 11 studies study NNS 37602 6 12 on on IN 37602 6 13 these these DT 37602 6 14 frogs frog NNS 37602 6 15 . . . 37602 7 1 One one CD 37602 7 2 of of IN 37602 7 3 us -PRON- PRP 37602 7 4 ( ( -LRB- 37602 7 5 Trueb Trueb NNP 37602 7 6 , , , 37602 7 7 1970a 1970a CD 37602 7 8 ) ) -RRB- 37602 7 9 studied study VBD 37602 7 10 the the DT 37602 7 11 cranial cranial JJ 37602 7 12 osteology osteology NN 37602 7 13 of of IN 37602 7 14 casque casque NN 37602 7 15 - - HYPH 37602 7 16 headed head VBN 37602 7 17 hylid hylid NN 37602 7 18 frogs frog NNS 37602 7 19 and and CC 37602 7 20 redefined redefine VBD 37602 7 21 the the DT 37602 7 22 genus genus NN 37602 7 23 Osteocephalus Osteocephalus NNP 37602 7 24 but but CC 37602 7 25 did do VBD 37602 7 26 not not RB 37602 7 27 determine determine VB 37602 7 28 the the DT 37602 7 29 number number NN 37602 7 30 of of IN 37602 7 31 species specie NNS 37602 7 32 in in IN 37602 7 33 the the DT 37602 7 34 genus genus NN 37602 7 35 . . . 37602 8 1 Our -PRON- PRP$ 37602 8 2 work work NN 37602 8 3 in in IN 37602 8 4 Amazonian amazonian JJ 37602 8 5 Ecuador Ecuador NNP 37602 8 6 resulted result VBD 37602 8 7 in in IN 37602 8 8 the the DT 37602 8 9 discovery discovery NN 37602 8 10 of of IN 37602 8 11 the the DT 37602 8 12 sympatric sympatric JJ 37602 8 13 occurrence occurrence NN 37602 8 14 of of IN 37602 8 15 three three CD 37602 8 16 species specie NNS 37602 8 17 at at IN 37602 8 18 each each DT 37602 8 19 of of IN 37602 8 20 two two CD 37602 8 21 localities locality NNS 37602 8 22 ; ; : 37602 8 23 one one CD 37602 8 24 of of IN 37602 8 25 these these DT 37602 8 26 species specie NNS 37602 8 27 was be VBD 37602 8 28 found find VBN 37602 8 29 with with IN 37602 8 30 a a DT 37602 8 31 fourth fourth JJ 37602 8 32 species species NN 37602 8 33 at at IN 37602 8 34 another another DT 37602 8 35 locality locality NN 37602 8 36 . . . 37602 9 1 Study study NN 37602 9 2 of of IN 37602 9 3 museum museum NN 37602 9 4 specimens specimen NNS 37602 9 5 confirmed confirm VBD 37602 9 6 the the DT 37602 9 7 recognition recognition NN 37602 9 8 of of IN 37602 9 9 these these DT 37602 9 10 four four CD 37602 9 11 species specie NNS 37602 9 12 in in IN 37602 9 13 the the DT 37602 9 14 Amazon Amazon NNP 37602 9 15 Basin Basin NNP 37602 9 16 and and CC 37602 9 17 lower low JJR 37602 9 18 Amazonian amazonian JJ 37602 9 19 slopes slope NNS 37602 9 20 of of IN 37602 9 21 the the DT 37602 9 22 Andes Andes NNP 37602 9 23 . . . 37602 10 1 A a DT 37602 10 2 fifth fifth JJ 37602 10 3 species species NN 37602 10 4 from from IN 37602 10 5 Bolivia Bolivia NNP 37602 10 6 and and CC 37602 10 7 Perú Perú NNP 37602 10 8 also also RB 37602 10 9 is be VBZ 37602 10 10 included include VBN 37602 10 11 in in IN 37602 10 12 the the DT 37602 10 13 genus genus NN 37602 10 14 . . . 37602 11 1 Examination examination NN 37602 11 2 of of IN 37602 11 3 museum museum NN 37602 11 4 specimens specimen NNS 37602 11 5 has have VBZ 37602 11 6 provided provide VBN 37602 11 7 data datum NNS 37602 11 8 on on IN 37602 11 9 the the DT 37602 11 10 geographic geographic JJ 37602 11 11 variation variation NN 37602 11 12 in in IN 37602 11 13 , , , 37602 11 14 and and CC 37602 11 15 distribution distribution NN 37602 11 16 of of IN 37602 11 17 , , , 37602 11 18 the the DT 37602 11 19 five five CD 37602 11 20 species specie NNS 37602 11 21 . . . 37602 12 1 However however RB 37602 12 2 , , , 37602 12 3 our -PRON- PRP$ 37602 12 4 conclusions conclusion NNS 37602 12 5 pertaining pertain VBG 37602 12 6 to to IN 37602 12 7 some some DT 37602 12 8 populations population NNS 37602 12 9 need need VB 37602 12 10 substantiation substantiation NN 37602 12 11 , , , 37602 12 12 because because IN 37602 12 13 we -PRON- PRP 37602 12 14 have have VBP 37602 12 15 been be VBN 37602 12 16 hampered hamper VBN 37602 12 17 by by IN 37602 12 18 inadequate inadequate JJ 37602 12 19 material material NN 37602 12 20 from from IN 37602 12 21 areas area NNS 37602 12 22 beyond beyond IN 37602 12 23 Ecuador Ecuador NNP 37602 12 24 . . . 37602 13 1 More More JJR 37602 13 2 than than IN 37602 13 3 half half NN 37602 13 4 of of IN 37602 13 5 the the DT 37602 13 6 905 905 CD 37602 13 7 specimens specimen NNS 37602 13 8 of of IN 37602 13 9 _ _ NNP 37602 13 10 Osteocephalus Osteocephalus NNP 37602 13 11 _ _ NNP 37602 13 12 are be VBP 37602 13 13 from from IN 37602 13 14 Ecuador Ecuador NNP 37602 13 15 , , , 37602 13 16 a a DT 37602 13 17 relatively relatively RB 37602 13 18 small small JJ 37602 13 19 part part NN 37602 13 20 of of IN 37602 13 21 the the DT 37602 13 22 total total JJ 37602 13 23 range range NN 37602 13 24 of of IN 37602 13 25 the the DT 37602 13 26 genus genus NN 37602 13 27 . . . 37602 14 1 In in IN 37602 14 2 this this DT 37602 14 3 paper paper NN 37602 14 4 we -PRON- PRP 37602 14 5 are be VBP 37602 14 6 presenting present VBG 37602 14 7 a a DT 37602 14 8 taxonomic taxonomic JJ 37602 14 9 review review NN 37602 14 10 of of IN 37602 14 11 the the DT 37602 14 12 genus genus NNP 37602 14 13 _ _ NNP 37602 14 14 Osteocephalus Osteocephalus NNP 37602 14 15 _ _ NNP 37602 14 16 ; ; : 37602 14 17 of of IN 37602 14 18 necessity necessity NN 37602 14 19 our -PRON- PRP$ 37602 14 20 study study NN 37602 14 21 has have VBZ 37602 14 22 been be VBN 37602 14 23 at at IN 37602 14 24 the the DT 37602 14 25 alpha alpha NN 37602 14 26 level level NN 37602 14 27 . . . 37602 15 1 We -PRON- PRP 37602 15 2 have have VBP 37602 15 3 utilized utilize VBN 37602 15 4 all all DT 37602 15 5 of of IN 37602 15 6 the the DT 37602 15 7 usual usual JJ 37602 15 8 external external JJ 37602 15 9 characters character NNS 37602 15 10 , , , 37602 15 11 as as RB 37602 15 12 well well RB 37602 15 13 as as IN 37602 15 14 osteological osteological JJ 37602 15 15 features feature NNS 37602 15 16 in in IN 37602 15 17 our -PRON- PRP$ 37602 15 18 definitions definition NNS 37602 15 19 of of IN 37602 15 20 the the DT 37602 15 21 species specie NNS 37602 15 22 . . . 37602 16 1 Tadpoles tadpole NNS 37602 16 2 and and CC 37602 16 3 mating mating NN 37602 16 4 calls call NNS 37602 16 5 are be VBP 37602 16 6 available available JJ 37602 16 7 for for IN 37602 16 8 only only RB 37602 16 9 one one CD 37602 16 10 species species NN 37602 16 11 , , , 37602 16 12 _ _ NNP 37602 16 13 O. O. NNP 37602 17 1 verrucigerus verrucigerus NNP 37602 17 2 _ _ NNP 37602 17 3 ( ( -LRB- 37602 17 4 Trueb Trueb NNP 37602 17 5 and and CC 37602 17 6 Duellman Duellman NNP 37602 17 7 , , , 37602 17 8 1970 1970 CD 37602 17 9 ) ) -RRB- 37602 17 10 ; ; : 37602 17 11 these these DT 37602 17 12 and and CC 37602 17 13 other other JJ 37602 17 14 important important JJ 37602 17 15 systematic systematic JJ 37602 17 16 characters character NNS 37602 17 17 , , , 37602 17 18 such such JJ 37602 17 19 as as IN 37602 17 20 karyotypes karyotype NNS 37602 17 21 , , , 37602 17 22 are be VBP 37602 17 23 not not RB 37602 17 24 available available JJ 37602 17 25 for for IN 37602 17 26 the the DT 37602 17 27 group group NN 37602 17 28 at at IN 37602 17 29 this this DT 37602 17 30 time time NN 37602 17 31 . . . 37602 18 1 Our -PRON- PRP$ 37602 18 2 tendency tendency NN 37602 18 3 has have VBZ 37602 18 4 been be VBN 37602 18 5 to to TO 37602 18 6 take take VB 37602 18 7 a a DT 37602 18 8 conservative conservative JJ 37602 18 9 view view NN 37602 18 10 of of IN 37602 18 11 species specie NNS 37602 18 12 ; ; : 37602 18 13 thus thus RB 37602 18 14 it -PRON- PRP 37602 18 15 is be VBZ 37602 18 16 doubtful doubtful JJ 37602 18 17 that that IN 37602 18 18 any any DT 37602 18 19 subsequent subsequent JJ 37602 18 20 worker worker NN 37602 18 21 will will MD 37602 18 22 recognize recognize VB 37602 18 23 fewer few JJR 37602 18 24 species specie NNS 37602 18 25 in in IN 37602 18 26 the the DT 37602 18 27 genus genus NN 37602 18 28 . . . 37602 19 1 Our -PRON- PRP$ 37602 19 2 observations observation NNS 37602 19 3 on on IN 37602 19 4 these these DT 37602 19 5 frogs frog NNS 37602 19 6 in in IN 37602 19 7 Amazonian Amazonian NNP 37602 19 8 Ecuador Ecuador NNP 37602 19 9 are be VBP 37602 19 10 presented present VBN 37602 19 11 in in IN 37602 19 12 a a DT 37602 19 13 final final JJ 37602 19 14 section section NN 37602 19 15 of of IN 37602 19 16 this this DT 37602 19 17 paper paper NN 37602 19 18 . . . 37602 20 1 ACKNOWLEDGMENTS acknowledgment NNS 37602 20 2 For for IN 37602 20 3 the the DT 37602 20 4 loan loan NN 37602 20 5 of of IN 37602 20 6 specimens specimen NNS 37602 20 7 or or CC 37602 20 8 for for IN 37602 20 9 the the DT 37602 20 10 provision provision NN 37602 20 11 of of IN 37602 20 12 working work VBG 37602 20 13 space space NN 37602 20 14 in in IN 37602 20 15 their -PRON- PRP$ 37602 20 16 respective respective JJ 37602 20 17 institutions institution NNS 37602 20 18 , , , 37602 20 19 we -PRON- PRP 37602 20 20 are be VBP 37602 20 21 indebted indebted JJ 37602 20 22 to to IN 37602 20 23 James James NNP 37602 20 24 E. E. NNP 37602 20 25 Böhlke Böhlke NNP 37602 20 26 , , , 37602 20 27 Werner Werner NNP 37602 20 28 C. C. NNP 37602 20 29 A. a. NN 37602 20 30 Bokermann Bokermann NNP 37602 20 31 , , , 37602 20 32 F. F. NNP 37602 20 33 W. W. NNP 37602 20 34 Braestrup Braestrup NNP 37602 20 35 , , , 37602 20 36 Nelly Nelly NNP 37602 20 37 Carrillo Carrillo NNP 37602 20 38 de de NNP 37602 20 39 Espinoza Espinoza NNP 37602 20 40 , , , 37602 20 41 Osvaldo Osvaldo NNP 37602 20 42 R. R. NNP 37602 20 43 da da NNP 37602 20 44 Cunha Cunha NNP 37602 20 45 , , , 37602 20 46 Josef Josef NNP 37602 20 47 Eiselt Eiselt NNP 37602 20 48 , , , 37602 20 49 M. M. NNP 37602 20 50 J. J. NNP 37602 20 51 Fouquette Fouquette NNP 37602 20 52 , , , 37602 20 53 Jr. Jr. NNP 37602 20 54 , , , 37602 20 55 Alice Alice NNP 37602 20 56 G. G. NNP 37602 20 57 C. C. NNP 37602 20 58 Grandison Grandison NNP 37602 20 59 , , , 37602 20 60 Jean Jean NNP 37602 20 61 Guibé Guibé NNP 37602 20 62 , , , 37602 20 63 Birgitta Birgitta NNP 37602 20 64 Hansson Hansson NNP 37602 20 65 , , , 37602 20 66 Walter Walter NNP 37602 20 67 Hellmich Hellmich NNP 37602 20 68 , , , 37602 20 69 M. M. NNP 37602 20 70 J. J. NNP 37602 20 71 Hoogmoed Hoogmoed NNP 37602 20 72 , , , 37602 20 73 Robert Robert NNP 37602 20 74 F. F. NNP 37602 20 75 Inger Inger NNP 37602 20 76 , , , 37602 20 77 Konrad Konrad NNP 37602 20 78 Klemmer Klemmer NNP 37602 20 79 , , , 37602 20 80 Jean Jean NNP 37602 20 81 Lescure Lescure NNP 37602 20 82 , , , 37602 20 83 Alan Alan NNP 37602 20 84 E. E. NNP 37602 20 85 Leviton Leviton NNP 37602 20 86 , , , 37602 20 87 Clarence Clarence NNP 37602 20 88 J. J. NNP 37602 20 89 McCoy McCoy NNP 37602 20 90 , , , 37602 20 91 Robert Robert NNP 37602 20 92 H. H. NNP 37602 20 93 Mount Mount NNP 37602 20 94 , , , 37602 20 95 Charles Charles NNP 37602 20 96 W. W. NNP 37602 20 97 Myers Myers NNP 37602 20 98 , , , 37602 20 99 Umberto Umberto NNP 37602 20 100 Parenti Parenti NNP 37602 20 101 , , , 37602 20 102 Günther Günther NNP 37602 20 103 Peters Peters NNP 37602 20 104 , , , 37602 20 105 James James NNP 37602 20 106 A. A. NNP 37602 20 107 Peters Peters NNP 37602 20 108 , , , 37602 20 109 William William NNP 37602 20 110 F. F. NNP 37602 20 111 Pyburn Pyburn NNP 37602 20 112 , , , 37602 20 113 Juan Juan NNP 37602 20 114 A. A. NNP 37602 20 115 Rivero Rivero NNP 37602 20 116 , , , 37602 20 117 Dorothy Dorothy NNP 37602 20 118 M. M. NNP 37602 20 119 Smith Smith NNP 37602 20 120 , , , 37602 20 121 Paulo Paulo NNP 37602 20 122 E. E. NNP 37602 20 123 Vanzolini Vanzolini NNP 37602 20 124 , , , 37602 20 125 Greta Greta NNP 37602 20 126 Vestergren Vestergren NNP 37602 20 127 , , , 37602 20 128 David David NNP 37602 20 129 B. B. NNP 37602 21 1 Wake Wake NNP 37602 21 2 , , , 37602 21 3 Charles Charles NNP 37602 21 4 F. F. NNP 37602 21 5 Walker Walker NNP 37602 21 6 , , , 37602 21 7 Ernest Ernest NNP 37602 21 8 E. E. NNP 37602 21 9 Williams Williams NNP 37602 21 10 , , , 37602 21 11 and and CC 37602 21 12 Richard Richard NNP 37602 21 13 G. G. NNP 37602 21 14 Zweifel Zweifel NNP 37602 21 15 . . . 37602 22 1 Study study NN 37602 22 2 of of IN 37602 22 3 specimens specimen NNS 37602 22 4 in in IN 37602 22 5 European european JJ 37602 22 6 museums museum NNS 37602 22 7 was be VBD 37602 22 8 made make VBN 37602 22 9 possible possible JJ 37602 22 10 by by IN 37602 22 11 a a DT 37602 22 12 grant grant NN 37602 22 13 ( ( -LRB- 37602 22 14 No no UH 37602 22 15 . . . 37602 23 1 5063 5063 LS 37602 23 2 ) ) -RRB- 37602 23 3 from from IN 37602 23 4 the the DT 37602 23 5 Penrose Penrose NNP 37602 23 6 Fund Fund NNP 37602 23 7 of of IN 37602 23 8 the the DT 37602 23 9 American American NNP 37602 23 10 Philosophical Philosophical NNP 37602 23 11 Society Society NNP 37602 23 12 . . . 37602 24 1 Field field NN 37602 24 2 work work NN 37602 24 3 in in IN 37602 24 4 Ecuador Ecuador NNP 37602 24 5 was be VBD 37602 24 6 partially partially RB 37602 24 7 supported support VBN 37602 24 8 by by IN 37602 24 9 grants grant NNS 37602 24 10 from from IN 37602 24 11 the the DT 37602 24 12 Watkins Watkins NNP 37602 24 13 Fund Fund NNP 37602 24 14 of of IN 37602 24 15 the the DT 37602 24 16 Museum Museum NNP 37602 24 17 of of IN 37602 24 18 Natural Natural NNP 37602 24 19 History History NNP 37602 24 20 , , , 37602 24 21 University University NNP 37602 24 22 of of IN 37602 24 23 Kansas Kansas NNP 37602 24 24 . . . 37602 25 1 At at IN 37602 25 2 our -PRON- PRP$ 37602 25 3 base base NN 37602 25 4 camp camp NN 37602 25 5 at at IN 37602 25 6 Santa Santa NNP 37602 25 7 Cecilia Cecilia NNP 37602 25 8 , , , 37602 25 9 Ecuador Ecuador NNP 37602 25 10 , , , 37602 25 11 we -PRON- PRP 37602 25 12 enjoyed enjoy VBD 37602 25 13 the the DT 37602 25 14 hospitality hospitality NN 37602 25 15 of of IN 37602 25 16 Ing Ing NNP 37602 25 17 . . . 37602 26 1 Ildefonso Ildefonso NNP 37602 26 2 Muñoz Muñoz NNP 37602 26 3 B. B. NNP 37602 27 1 Transportation transportation NN 37602 27 2 in in IN 37602 27 3 Ecuador Ecuador NNP 37602 27 4 was be VBD 37602 27 5 generously generously RB 37602 27 6 provided provide VBN 37602 27 7 by by IN 37602 27 8 the the DT 37602 27 9 Texaco Texaco NNP 37602 27 10 Petroleum Petroleum NNP 37602 27 11 Company Company NNP 37602 27 12 . . . 37602 28 1 During during IN 37602 28 2 the the DT 37602 28 3 course course NN 37602 28 4 of of IN 37602 28 5 our -PRON- PRP$ 37602 28 6 field field NN 37602 28 7 work work NN 37602 28 8 Stephen Stephen NNP 37602 28 9 R. R. NNP 37602 28 10 Edwards Edwards NNP 37602 28 11 and and CC 37602 28 12 Thomas Thomas NNP 37602 28 13 H. H. NNP 37602 28 14 Fritts Fritts NNP 37602 28 15 contributed contribute VBD 37602 28 16 directly directly RB 37602 28 17 to to IN 37602 28 18 our -PRON- PRP$ 37602 28 19 study study NN 37602 28 20 of of IN 37602 28 21 _ _ NNP 37602 28 22 Osteocephalus Osteocephalus NNP 37602 28 23 _ _ NNP 37602 28 24 . . . 37602 29 1 Michael Michael NNP 37602 29 2 J. J. NNP 37602 29 3 Tyler Tyler NNP 37602 29 4 of of IN 37602 29 5 the the DT 37602 29 6 South South NNP 37602 29 7 Australian Australian NNP 37602 29 8 Museum Museum NNP 37602 29 9 provided provide VBD 37602 29 10 information information NN 37602 29 11 on on IN 37602 29 12 the the DT 37602 29 13 vocal vocal JJ 37602 29 14 sac sac NN 37602 29 15 structure structure NN 37602 29 16 . . . 37602 30 1 We -PRON- PRP 37602 30 2 extend extend VBP 37602 30 3 our -PRON- PRP$ 37602 30 4 sincere sincere JJ 37602 30 5 thanks thank NNS 37602 30 6 to to IN 37602 30 7 all all DT 37602 30 8 of of IN 37602 30 9 these these DT 37602 30 10 persons person NNS 37602 30 11 for for IN 37602 30 12 their -PRON- PRP$ 37602 30 13 contributions contribution NNS 37602 30 14 to to IN 37602 30 15 our -PRON- PRP$ 37602 30 16 endeavors endeavor NNS 37602 30 17 . . . 37602 31 1 MATERIALS material NNS 37602 31 2 AND and CC 37602 31 3 METHODS methods JJ 37602 31 4 We -PRON- PRP 37602 31 5 have have VBP 37602 31 6 examined examine VBN 37602 31 7 893 893 CD 37602 31 8 preserved preserve VBN 37602 31 9 frogs frog NNS 37602 31 10 , , , 37602 31 11 including include VBG 37602 31 12 the the DT 37602 31 13 type type NN 37602 31 14 specimens specimen NNS 37602 31 15 of of IN 37602 31 16 all all DT 37602 31 17 included include VBN 37602 31 18 nominal nominal JJ 37602 31 19 taxa taxa NN 37602 31 20 , , , 37602 31 21 8 8 CD 37602 31 22 skeletons skeleton NNS 37602 31 23 , , , 37602 31 24 1 1 CD 37602 31 25 lot lot NN 37602 31 26 of of IN 37602 31 27 eggs egg NNS 37602 31 28 , , , 37602 31 29 and and CC 37602 31 30 3 3 CD 37602 31 31 lots lot NNS 37602 31 32 of of IN 37602 31 33 tadpoles tadpole NNS 37602 31 34 that that IN 37602 31 35 we -PRON- PRP 37602 31 36 refer refer VBP 37602 31 37 to to IN 37602 31 38 the the DT 37602 31 39 genus genus NNP 37602 31 40 _ _ NNP 37602 31 41 Osteocephalus Osteocephalus NNP 37602 31 42 _ _ NNP 37602 31 43 ; ; : 37602 31 44 in in IN 37602 31 45 addition addition NN 37602 31 46 skulls skull NNS 37602 31 47 were be VBD 37602 31 48 removed remove VBN 37602 31 49 from from IN 37602 31 50 five five CD 37602 31 51 preserved preserve VBN 37602 31 52 specimens specimen NNS 37602 31 53 , , , 37602 31 54 and and CC 37602 31 55 radiographs radiograph NNS 37602 31 56 were be VBD 37602 31 57 made make VBN 37602 31 58 of of IN 37602 31 59 12 12 CD 37602 31 60 other other JJ 37602 31 61 preserved preserve VBN 37602 31 62 specimens specimen NNS 37602 31 63 . . . 37602 32 1 We -PRON- PRP 37602 32 2 have have VBP 37602 32 3 been be VBN 37602 32 4 fortunate fortunate JJ 37602 32 5 in in IN 37602 32 6 seeing see VBG 37602 32 7 living living NN 37602 32 8 individuals individual NNS 37602 32 9 of of IN 37602 32 10 all all DT 37602 32 11 species specie NNS 37602 32 12 , , , 37602 32 13 except except IN 37602 32 14 _ _ NNP 37602 32 15 O. O. NNP 37602 33 1 pearsoni pearsoni NNP 37602 33 2 _ _ NNP 37602 33 3 , , , 37602 33 4 but but CC 37602 33 5 we -PRON- PRP 37602 33 6 have have VBP 37602 33 7 colored color VBN 37602 33 8 photographs photograph NNS 37602 33 9 of of IN 37602 33 10 a a DT 37602 33 11 living live VBG 37602 33 12 specimen speciman NNS 37602 33 13 of of IN 37602 33 14 that that DT 37602 33 15 species specie NNS 37602 33 16 . . . 37602 34 1 Figures figure NNS 37602 34 2 1 1 CD 37602 34 3 and and CC 37602 34 4 2 2 CD 37602 34 5 were be VBD 37602 34 6 drawn draw VBN 37602 34 7 from from IN 37602 34 8 projected project VBN 37602 34 9 colored colored JJ 37602 34 10 transparencies transparency NNS 37602 34 11 of of IN 37602 34 12 living living NN 37602 34 13 frogs frog NNS 37602 34 14 . . . 37602 35 1 Terminology terminology NN 37602 35 2 follows follow VBZ 37602 35 3 that that DT 37602 35 4 of of IN 37602 35 5 Duellman Duellman NNP 37602 35 6 ( ( -LRB- 37602 35 7 1970b 1970b CD 37602 35 8 ) ) -RRB- 37602 35 9 . . . 37602 36 1 On on IN 37602 36 2 the the DT 37602 36 3 distribution distribution NN 37602 36 4 maps map NNS 37602 36 5 solid solid JJ 37602 36 6 symbols symbol NNS 37602 36 7 indicate indicate VBP 37602 36 8 localities locality NNS 37602 36 9 from from IN 37602 36 10 which which WDT 37602 36 11 we -PRON- PRP 37602 36 12 have have VBP 37602 36 13 examined examine VBN 37602 36 14 specimens specimen NNS 37602 36 15 ; ; : 37602 36 16 open open JJ 37602 36 17 symbols symbol NNS 37602 36 18 represent represent VBP 37602 36 19 additional additional JJ 37602 36 20 locality locality NN 37602 36 21 records record NNS 37602 36 22 based base VBN 37602 36 23 on on IN 37602 36 24 the the DT 37602 36 25 literature literature NN 37602 36 26 . . . 37602 37 1 Throughout throughout IN 37602 37 2 the the DT 37602 37 3 text text NN 37602 37 4 specimens specimen NNS 37602 37 5 are be VBP 37602 37 6 listed list VBN 37602 37 7 by by IN 37602 37 8 their -PRON- PRP$ 37602 37 9 catalogue catalogue NN 37602 37 10 numbers number NNS 37602 37 11 preceded precede VBN 37602 37 12 by by IN 37602 37 13 the the DT 37602 37 14 appropriate appropriate NNP 37602 37 15 museum museum NN 37602 37 16 abbreviation abbreviation NN 37602 37 17 , , , 37602 37 18 as as IN 37602 37 19 follows follow VBZ 37602 37 20 : : : 37602 37 21 AMNH AMNH NNP 37602 37 22 American American NNP 37602 37 23 Museum Museum NNP 37602 37 24 of of IN 37602 37 25 Natural Natural NNP 37602 37 26 History History NNP 37602 37 27 ANSP ANSP NNP 37602 37 28 Academy Academy NNP 37602 37 29 of of IN 37602 37 30 Natural Natural NNP 37602 37 31 Sciences Sciences NNPS 37602 37 32 of of IN 37602 37 33 Philadelphia Philadelphia NNP 37602 37 34 ASU ASU NNP 37602 37 35 Arizona Arizona NNP 37602 37 36 State State NNP 37602 37 37 University University NNP 37602 37 38 AUM AUM NNP 37602 37 39 Auburn Auburn NNP 37602 37 40 University University NNP 37602 37 41 Museum Museum NNP 37602 37 42 BMNH BMNH NNP 37602 37 43 British British NNP 37602 37 44 Museum Museum NNP 37602 37 45 ( ( -LRB- 37602 37 46 Natural Natural NNP 37602 37 47 History History NNP 37602 37 48 ) ) -RRB- 37602 37 49 CAS CAS NNP 37602 37 50 California California NNP 37602 37 51 Academy Academy NNP 37602 37 52 of of IN 37602 37 53 Sciences Sciences NNPS 37602 37 54 CAS CAS NNP 37602 37 55 - - HYPH 37602 37 56 SU SU NNP 37602 37 57 Stanford Stanford NNP 37602 37 58 University University NNP 37602 37 59 Collection Collection NNP 37602 37 60 ( ( -LRB- 37602 37 61 In in IN 37602 37 62 California California NNP 37602 37 63 Academy Academy NNP 37602 37 64 of of IN 37602 37 65 Sciences Sciences NNPS 37602 37 66 ) ) -RRB- 37602 37 67 CM CM NNP 37602 37 68 Carnegie Carnegie NNP 37602 37 69 Museum Museum NNP 37602 37 70 FMNH FMNH NNP 37602 37 71 Field Field NNP 37602 37 72 Museum Museum NNP 37602 37 73 of of IN 37602 37 74 Natural Natural NNP 37602 37 75 History History NNP 37602 37 76 KU KU NNP 37602 37 77 University University NNP 37602 37 78 of of IN 37602 37 79 Kansas Kansas NNP 37602 37 80 Museum Museum NNP 37602 37 81 of of IN 37602 37 82 Natural Natural NNP 37602 37 83 History History NNP 37602 37 84 MCZ MCZ NNP 37602 37 85 Museum Museum NNP 37602 37 86 of of IN 37602 37 87 Comparative Comparative NNP 37602 37 88 Zoology Zoology NNP 37602 37 89 , , , 37602 37 90 Harvard Harvard NNP 37602 37 91 University University NNP 37602 37 92 MIZS MIZS NNPS 37602 37 93 Museo Museo NNP 37602 37 94 ed ed NNP 37602 37 95 Istituto Istituto NNP 37602 37 96 di di NNP 37602 37 97 Zoologi Zoologi NNP 37602 37 98 Sistematico Sistematico NNP 37602 37 99 , , , 37602 37 100 Università Università NNP 37602 37 101 di di NN 37602 37 102 Torino Torino NNP 37602 37 103 MJP MJP NNP 37602 37 104 Museo Museo NNP 37602 37 105 Javier Javier NNP 37602 37 106 Prado Prado NNP 37602 37 107 , , , 37602 37 108 Lima Lima NNP 37602 37 109 MNHN MNHN NNP 37602 37 110 Muséum Muséum NNP 37602 37 111 National National NNP 37602 37 112 d'Histoire d'Histoire NNP 37602 37 113 Naturelle Naturelle NNP 37602 37 114 , , , 37602 37 115 Paris Paris NNP 37602 37 116 MPEG MPEG NNP 37602 37 117 Museu Museu NNP 37602 37 118 Paraense Paraense NNP 37602 37 119 Emiliano Emiliano NNP 37602 37 120 Goeldi Goeldi NNP 37602 37 121 , , , 37602 37 122 Belém Belém NNP 37602 37 123 MVZ MVZ NNP 37602 37 124 Museum Museum NNP 37602 37 125 of of IN 37602 37 126 Vertebrate Vertebrate NNP 37602 37 127 Zoology Zoology NNP 37602 37 128 , , , 37602 37 129 University University NNP 37602 37 130 of of IN 37602 37 131 California California NNP 37602 37 132 , , , 37602 37 133 Berkeley Berkeley NNP 37602 37 134 MZUSP MZUSP NNP 37602 37 135 Museu Museu NNP 37602 37 136 de de NNP 37602 37 137 Zoología Zoología NNP 37602 37 138 , , , 37602 37 139 Universidade Universidade NNP 37602 37 140 da da NNP 37602 37 141 São São NNP 37602 37 142 Paulo Paulo NNP 37602 37 143 NHMG NHMG NNP 37602 37 144 Naturhistoriska Naturhistoriska NNP 37602 37 145 Museet Museet NNP 37602 37 146 Göteborg Göteborg NNP 37602 37 147 NHMW NHMW NNP 37602 37 148 Naturhistorisches Naturhistorisches NNP 37602 37 149 Museum Museum NNP 37602 37 150 , , , 37602 37 151 Wien Wien NNP 37602 37 152 NHRM NHRM NNP 37602 37 153 Naturhistoriska Naturhistoriska NNP 37602 37 154 Riksmuseet Riksmuseet NNP 37602 37 155 , , , 37602 37 156 Stockholm Stockholm NNP 37602 37 157 RMNH RMNH NNP 37602 37 158 Rijksmuseum Rijksmuseum NNP 37602 37 159 van van NNP 37602 37 160 Natuurlijke Natuurlijke NNP 37602 37 161 Histoire Histoire NNP 37602 37 162 , , , 37602 37 163 Leiden Leiden NNP 37602 37 164 SMF SMF NNP 37602 37 165 Senckenbergische Senckenbergische NNP 37602 37 166 Museum Museum NNP 37602 37 167 , , , 37602 37 168 Frankfurt Frankfurt NNP 37602 37 169 UIMNH UIMNH NNP 37602 37 170 University University NNP 37602 37 171 of of IN 37602 37 172 Illinois Illinois NNP 37602 37 173 , , , 37602 37 174 Museum Museum NNP 37602 37 175 of of IN 37602 37 176 Natural Natural NNP 37602 37 177 History History NNP 37602 37 178 UMMZ UMMZ NNP 37602 37 179 University University NNP 37602 37 180 of of IN 37602 37 181 Michigan Michigan NNP 37602 37 182 Museum Museum NNP 37602 37 183 of of IN 37602 37 184 Zoology Zoology NNP 37602 37 185 UP UP NNP 37602 37 186 Université Université NNP 37602 37 187 de de FW 37602 37 188 Paris Paris NNP 37602 37 189 UPR UPR NNP 37602 37 190 - - HYPH 37602 37 191 M M NNP 37602 37 192 University University NNP 37602 37 193 of of IN 37602 37 194 Puerto Puerto NNP 37602 37 195 Rico Rico NNP 37602 37 196 , , , 37602 37 197 Mayagüez Mayagüez NNP 37602 37 198 UTA UTA NNP 37602 37 199 University University NNP 37602 37 200 of of IN 37602 37 201 Texas Texas NNP 37602 37 202 , , , 37602 37 203 Arlington Arlington NNP 37602 37 204 USNM USNM NNP 37602 37 205 United United NNP 37602 37 206 States States NNP 37602 37 207 National National NNP 37602 37 208 Museum Museum NNP 37602 37 209 UZM UZM NNP 37602 37 210 Universitets Universitets NNP 37602 37 211 Zoologiske Zoologiske NNP 37602 37 212 Museum Museum NNP 37602 37 213 , , , 37602 37 214 Copenhagen Copenhagen NNP 37602 37 215 WCAB WCAB NNP 37602 37 216 Werner Werner NNP 37602 37 217 C. C. NNP 37602 37 218 A. a. NN 37602 37 219 Bokermann Bokermann NNP 37602 37 220 , , , 37602 37 221 São São NNP 37602 37 222 Paulo Paulo NNP 37602 37 223 , , , 37602 37 224 Brasil Brasil NNP 37602 37 225 ZMB ZMB NNP 37602 37 226 Zoologisches Zoologisches NNP 37602 37 227 Museum Museum NNP 37602 37 228 Berlin Berlin NNP 37602 37 229 ZSM ZSM NNP 37602 37 230 Zoologisches zoologische VBZ 37602 37 231 Sammlung Sammlung NNP 37602 37 232 München München NNP 37602 37 233 HISTORICAL HISTORICAL NNP 37602 37 234 RESUMÉ RESUMÉ NNP 37602 37 235 Because because IN 37602 37 236 of of IN 37602 37 237 the the DT 37602 37 238 taxonomic taxonomic JJ 37602 37 239 confusion confusion NN 37602 37 240 that that WDT 37602 37 241 has have VBZ 37602 37 242 surrounded surround VBN 37602 37 243 the the DT 37602 37 244 generic generic JJ 37602 37 245 name name NN 37602 37 246 _ _ NNP 37602 37 247 Osteocephalus Osteocephalus NNP 37602 37 248 _ _ NNP 37602 37 249 and and CC 37602 37 250 two two CD 37602 37 251 of of IN 37602 37 252 the the DT 37602 37 253 species specie NNS 37602 37 254 ( ( -LRB- 37602 37 255 and and CC 37602 37 256 their -PRON- PRP$ 37602 37 257 synonyms synonyms NN 37602 37 258 ) ) -RRB- 37602 37 259 , , , 37602 37 260 we -PRON- PRP 37602 37 261 present present VBP 37602 37 262 a a DT 37602 37 263 brief brief JJ 37602 37 264 resumé resumé NN 37602 37 265 of of IN 37602 37 266 the the DT 37602 37 267 taxonomic taxonomic JJ 37602 37 268 history history NN 37602 37 269 of of IN 37602 37 270 the the DT 37602 37 271 group group NN 37602 37 272 . . . 37602 38 1 Among among IN 37602 38 2 the the DT 37602 38 3 amphibians amphibian NNS 37602 38 4 sent send VBN 37602 38 5 to to IN 37602 38 6 the the DT 37602 38 7 Muséum Muséum NNP 37602 38 8 National National NNP 37602 38 9 d'Histoire d'Histoire NNP 37602 38 10 Naturelle Naturelle NNP 37602 38 11 in in IN 37602 38 12 Paris Paris NNP 37602 38 13 by by IN 37602 38 14 a a DT 37602 38 15 Monsieur Monsieur NNP 37602 38 16 Leprieur Leprieur NNP 37602 38 17 in in IN 37602 38 18 French French NNP 37602 38 19 Guiana Guiana NNP 37602 38 20 was be VBD 37602 38 21 a a DT 37602 38 22 single single JJ 37602 38 23 female female JJ 37602 38 24 specimen speciman NNS 37602 38 25 of of IN 37602 38 26 a a DT 37602 38 27 moderately moderately RB 37602 38 28 large large JJ 37602 38 29 hylid hylid NN 37602 38 30 having have VBG 37602 38 31 a a DT 37602 38 32 well well RB 37602 38 33 - - HYPH 37602 38 34 ossified ossify VBN 37602 38 35 skull skull NN 37602 38 36 and and CC 37602 38 37 smooth smooth JJ 37602 38 38 dorsal dorsal NN 37602 38 39 skin skin NN 37602 38 40 . . . 37602 39 1 This this DT 37602 39 2 specimen specimen VBZ 37602 39 3 escaped escape VBD 37602 39 4 from from IN 37602 39 5 the the DT 37602 39 6 covetous covetous JJ 37602 39 7 eyes eye NNS 37602 39 8 of of IN 37602 39 9 Johann Johann NNP 37602 39 10 Tschudi Tschudi NNP 37602 39 11 , , , 37602 39 12 who who WP 37602 39 13 prematurely prematurely RB 37602 39 14 named name VBD 37602 39 15 several several JJ 37602 39 16 species specie NNS 37602 39 17 on on IN 37602 39 18 the the DT 37602 39 19 basis basis NN 37602 39 20 of of IN 37602 39 21 specimens specimen NNS 37602 39 22 in in IN 37602 39 23 Paris Paris NNP 37602 39 24 , , , 37602 39 25 and and CC 37602 39 26 survived survive VBD 37602 39 27 without without IN 37602 39 28 an an DT 37602 39 29 epithet epithet NN 37602 39 30 until until IN 37602 39 31 Duméril Duméril NNP 37602 39 32 and and CC 37602 39 33 Bibron Bibron NNP 37602 39 34 ( ( -LRB- 37602 39 35 1841 1841 CD 37602 39 36 ) ) -RRB- 37602 39 37 proposed propose VBD 37602 39 38 for for IN 37602 39 39 it -PRON- PRP 37602 39 40 the the DT 37602 39 41 name name NN 37602 39 42 _ _ NNP 37602 39 43 Hyla Hyla NNP 37602 39 44 leprieurii leprieurii JJ 37602 39 45 _ _ NNP 37602 39 46 . . . 37602 40 1 The the DT 37602 40 2 description description NN 37602 40 3 of of IN 37602 40 4 the the DT 37602 40 5 species specie NNS 37602 40 6 is be VBZ 37602 40 7 fairly fairly RB 37602 40 8 detailed detailed JJ 37602 40 9 , , , 37602 40 10 but but CC 37602 40 11 the the DT 37602 40 12 specimen speciman NNS 37602 40 13 was be VBD 37602 40 14 not not RB 37602 40 15 illustrated illustrate VBN 37602 40 16 . . . 37602 41 1 This this DT 37602 41 2 is be VBZ 37602 41 3 the the DT 37602 41 4 earliest early JJS 37602 41 5 trivial trivial JJ 37602 41 6 name name NN 37602 41 7 now now RB 37602 41 8 associated associate VBN 37602 41 9 with with IN 37602 41 10 _ _ NNP 37602 41 11 Osteocephalus Osteocephalus NNP 37602 41 12 _ _ NNP 37602 41 13 . . . 37602 42 1 Fitzinger Fitzinger NNP 37602 42 2 ( ( -LRB- 37602 42 3 1843 1843 CD 37602 42 4 ) ) -RRB- 37602 42 5 in in IN 37602 42 6 his -PRON- PRP$ 37602 42 7 generic generic JJ 37602 42 8 synopsis synopsis NN 37602 42 9 of of IN 37602 42 10 amphibians amphibian NNS 37602 42 11 and and CC 37602 42 12 reptiles reptile NNS 37602 42 13 proposed propose VBD 37602 42 14 the the DT 37602 42 15 generic generic JJ 37602 42 16 name name NN 37602 42 17 _ _ NNP 37602 42 18 Osteocephalus Osteocephalus NNP 37602 42 19 _ _ NNP 37602 42 20 but but CC 37602 42 21 did do VBD 37602 42 22 not not RB 37602 42 23 associate associate VB 37602 42 24 a a DT 37602 42 25 specific specific JJ 37602 42 26 name name NN 37602 42 27 with with IN 37602 42 28 the the DT 37602 42 29 genus genus NN 37602 42 30 . . . 37602 43 1 Consequently consequently RB 37602 43 2 , , , 37602 43 3 _ _ NNP 37602 43 4 Osteocephalus Osteocephalus NNP 37602 43 5 _ _ NNP 37602 43 6 Fitzinger Fitzinger NNP 37602 43 7 , , , 37602 43 8 1843 1843 CD 37602 43 9 , , , 37602 43 10 is be VBZ 37602 43 11 a a DT 37602 43 12 _ _ NNP 37602 43 13 nomen nomen NNP 37602 43 14 nudum nudum NNP 37602 43 15 _ _ NNP 37602 43 16 . . . 37602 44 1 Franz Franz NNP 37602 44 2 Steindachner Steindachner NNP 37602 44 3 followed follow VBD 37602 44 4 Leopoldo Leopoldo NNP 37602 44 5 Fitzinger Fitzinger NNP 37602 44 6 at at IN 37602 44 7 the the DT 37602 44 8 Naturhistorisches Naturhistorisches NNP 37602 44 9 Museum Museum NNP 37602 44 10 in in IN 37602 44 11 Vienna Vienna NNP 37602 44 12 , , , 37602 44 13 where where WRB 37602 44 14 he -PRON- PRP 37602 44 15 had have VBD 37602 44 16 access access NN 37602 44 17 to to IN 37602 44 18 Fitzinger Fitzinger NNP 37602 44 19 's 's POS 37602 44 20 notes note NNS 37602 44 21 and and CC 37602 44 22 , , , 37602 44 23 of of IN 37602 44 24 course course NN 37602 44 25 , , , 37602 44 26 the the DT 37602 44 27 important important JJ 37602 44 28 collections collection NNS 37602 44 29 housed house VBN 37602 44 30 in in IN 37602 44 31 that that DT 37602 44 32 museum museum NN 37602 44 33 . . . 37602 45 1 Steindachner Steindachner NNP 37602 45 2 ( ( -LRB- 37602 45 3 1862 1862 CD 37602 45 4 ) ) -RRB- 37602 45 5 named name VBN 37602 45 6 two two CD 37602 45 7 species specie NNS 37602 45 8 of of IN 37602 45 9 _ _ NNP 37602 45 10 Osteocephalus Osteocephalus NNP 37602 45 11 _ _ NNP 37602 45 12 on on IN 37602 45 13 the the DT 37602 45 14 basis basis NN 37602 45 15 of of IN 37602 45 16 Brasilian brasilian JJ 37602 45 17 specimens specimen NNS 37602 45 18 collected collect VBN 37602 45 19 by by IN 37602 45 20 Johann Johann NNP 37602 45 21 Natterer Natterer NNP 37602 45 22 . . . 37602 46 1 Both both DT 37602 46 2 species specie NNS 37602 46 3 were be VBD 37602 46 4 named name VBN 37602 46 5 in in IN 37602 46 6 the the DT 37602 46 7 same same JJ 37602 46 8 publication publication NN 37602 46 9 ; ; : 37602 46 10 _ _ NNP 37602 46 11 O. O. NNP 37602 47 1 taurinus taurinus NNP 37602 47 2 _ _ NNP 37602 47 3 appeared appear VBD 37602 47 4 on on IN 37602 47 5 page page NN 37602 47 6 77 77 CD 37602 47 7 , , , 37602 47 8 and and CC 37602 47 9 _ _ NNP 37602 47 10 O. O. NNP 37602 48 1 favolineatus favolineatus NNP 37602 48 2 _ _ NNP 37602 48 3 , , , 37602 48 4 on on IN 37602 48 5 p. p. NN 37602 48 6 80 80 CD 37602 48 7 . . . 37602 49 1 This this DT 37602 49 2 is be VBZ 37602 49 3 the the DT 37602 49 4 earliest early JJS 37602 49 5 association association NN 37602 49 6 of of IN 37602 49 7 the the DT 37602 49 8 generic generic JJ 37602 49 9 name name NN 37602 49 10 _ _ NNP 37602 49 11 Osteocephalus Osteocephalus NNP 37602 49 12 _ _ NNP 37602 49 13 with with IN 37602 49 14 a a DT 37602 49 15 specific specific JJ 37602 49 16 name name NN 37602 49 17 and and CC 37602 49 18 a a DT 37602 49 19 description description NN 37602 49 20 , , , 37602 49 21 both both DT 37602 49 22 of of IN 37602 49 23 which which WDT 37602 49 24 satisfy satisfy NN 37602 49 25 the the DT 37602 49 26 Code Code NNP 37602 49 27 of of IN 37602 49 28 Zoological Zoological NNP 37602 49 29 Nomenclature Nomenclature NNP 37602 49 30 for for IN 37602 49 31 generic generic JJ 37602 49 32 availability availability NN 37602 49 33 . . . 37602 50 1 Therefore therefore RB 37602 50 2 , , , 37602 50 3 Steindachner Steindachner NNP 37602 50 4 is be VBZ 37602 50 5 the the DT 37602 50 6 authority authority NN 37602 50 7 for for IN 37602 50 8 the the DT 37602 50 9 generic generic JJ 37602 50 10 name name NN 37602 50 11 _ _ NNP 37602 50 12 Osteocephalus Osteocephalus NNP 37602 50 13 _ _ NNP 37602 50 14 , , , 37602 50 15 which which WDT 37602 50 16 has have VBZ 37602 50 17 _ _ NNP 37602 50 18 O. O. NNP 37602 51 1 taurinus taurinus NNP 37602 51 2 _ _ NNP 37602 51 3 as as IN 37602 51 4 the the DT 37602 51 5 type type NN 37602 51 6 species specie NNS 37602 51 7 by by IN 37602 51 8 original original JJ 37602 51 9 designation designation NN 37602 51 10 . . . 37602 52 1 It -PRON- PRP 37602 52 2 is be VBZ 37602 52 3 not not RB 37602 52 4 possible possible JJ 37602 52 5 to to TO 37602 52 6 determine determine VB 37602 52 7 whether whether IN 37602 52 8 or or CC 37602 52 9 not not RB 37602 52 10 Steindachner Steindachner NNP 37602 52 11 's 's POS 37602 52 12 usage usage NN 37602 52 13 of of IN 37602 52 14 _ _ NNP 37602 52 15 Osteocephalus Osteocephalus NNP 37602 52 16 _ _ NNP 37602 52 17 was be VBD 37602 52 18 the the DT 37602 52 19 same same JJ 37602 52 20 as as IN 37602 52 21 that that DT 37602 52 22 intended intend VBN 37602 52 23 by by IN 37602 52 24 Fitzinger Fitzinger NNP 37602 52 25 19 19 CD 37602 52 26 years year NNS 37602 52 27 earlier early RBR 37602 52 28 . . . 37602 53 1 Steindachner Steindachner NNP 37602 53 2 ( ( -LRB- 37602 53 3 1862 1862 CD 37602 53 4 ) ) -RRB- 37602 53 5 gave give VBD 37602 53 6 reasonably reasonably RB 37602 53 7 good good JJ 37602 53 8 descriptions description NNS 37602 53 9 of of IN 37602 53 10 his -PRON- PRP$ 37602 53 11 two two CD 37602 53 12 new new JJ 37602 53 13 species specie NNS 37602 53 14 and and CC 37602 53 15 provided provide VBD 37602 53 16 excellent excellent JJ 37602 53 17 illustrations illustration NNS 37602 53 18 of of IN 37602 53 19 the the DT 37602 53 20 two two CD 37602 53 21 specimens specimen NNS 37602 53 22 , , , 37602 53 23 both both CC 37602 53 24 large large JJ 37602 53 25 females female NNS 37602 53 26 . . . 37602 54 1 Apparently apparently RB 37602 54 2 impressed impress VBN 37602 54 3 by by IN 37602 54 4 the the DT 37602 54 5 similarities similarity NNS 37602 54 6 between between IN 37602 54 7 _ _ NNP 37602 54 8 Trachycephalus Trachycephalus NNP 37602 54 9 nigromaculatus nigromaculatus NN 37602 54 10 _ _ NNP 37602 54 11 Tschudi Tschudi NNP 37602 54 12 , , , 37602 54 13 1838 1838 CD 37602 54 14 , , , 37602 54 15 and and CC 37602 54 16 _ _ NNP 37602 54 17 Osteocephalus Osteocephalus NNP 37602 54 18 taurinus taurinus NN 37602 54 19 _ _ NNP 37602 54 20 , , , 37602 54 21 Steindachner Steindachner NNP 37602 54 22 ( ( -LRB- 37602 54 23 1867 1867 CD 37602 54 24 ) ) -RRB- 37602 54 25 used use VBD 37602 54 26 the the DT 37602 54 27 combination combination NN 37602 54 28 _ _ NNP 37602 54 29 Trachycephalus Trachycephalus NNP 37602 54 30 ( ( -LRB- 37602 54 31 Osteocephalus Osteocephalus NNP 37602 54 32 ) ) -RRB- 37602 54 33 taurinus taurinus NN 37602 54 34 _ _ NN 37602 54 35 . . . 37602 55 1 This this DT 37602 55 2 ambiguous ambiguous JJ 37602 55 3 usage usage NN 37602 55 4 for for IN 37602 55 5 the the DT 37602 55 6 1860 1860 CD 37602 55 7 's 's POS 37602 55 8 precludes preclude NNS 37602 55 9 our -PRON- PRP$ 37602 55 10 determining determine VBG 37602 55 11 if if IN 37602 55 12 Steindachner Steindachner NNP 37602 55 13 was be VBD 37602 55 14 in in IN 37602 55 15 effect effect NN 37602 55 16 synonymizing synonymize VBG 37602 55 17 _ _ NNP 37602 55 18 Osteocephalus Osteocephalus NNP 37602 55 19 _ _ NNP 37602 55 20 with with IN 37602 55 21 _ _ NNP 37602 55 22 Trachycephalus Trachycephalus NNP 37602 55 23 _ _ NNP 37602 55 24 or or CC 37602 55 25 whether whether IN 37602 55 26 he -PRON- PRP 37602 55 27 was be VBD 37602 55 28 placing place VBG 37602 55 29 _ _ NNP 37602 55 30 Osteocephalus Osteocephalus NNP 37602 55 31 _ _ NNP 37602 55 32 in in IN 37602 55 33 a a DT 37602 55 34 subgeneric subgeneric JJ 37602 55 35 status status NN 37602 55 36 . . . 37602 56 1 Steindachner Steindachner NNP 37602 56 2 ( ( -LRB- 37602 56 3 1867 1867 CD 37602 56 4 ) ) -RRB- 37602 56 5 did do VBD 37602 56 6 not not RB 37602 56 7 mention mention VB 37602 56 8 _ _ NNP 37602 56 9 O. O. NNP 37602 57 1 flavolineatus flavolineatus NNP 37602 57 2 _ _ NNP 37602 57 3 ; ; : 37602 57 4 perhaps perhaps RB 37602 57 5 by by IN 37602 57 6 that that DT 37602 57 7 time time NN 37602 57 8 he -PRON- PRP 37602 57 9 had have VBD 37602 57 10 concluded conclude VBN 37602 57 11 that that IN 37602 57 12 _ _ NNP 37602 57 13 flavolineatus flavolineatus NN 37602 57 14 _ _ NNP 37602 57 15 was be VBD 37602 57 16 only only RB 37602 57 17 a a DT 37602 57 18 color color NN 37602 57 19 morph morph NN 37602 57 20 of of IN 37602 57 21 _ _ NNP 37602 57 22 taurinus taurinus NNP 37602 57 23 _ _ NNP 37602 57 24 . . . 37602 58 1 Cope Cope NNP 37602 58 2 ( ( -LRB- 37602 58 3 1867 1867 CD 37602 58 4 ) ) -RRB- 37602 58 5 placed place VBD 37602 58 6 _ _ NNP 37602 58 7 Hyla Hyla NNP 37602 58 8 leprieurii leprieurii JJ 37602 58 9 _ _ NNP 37602 58 10 in in IN 37602 58 11 the the DT 37602 58 12 genus genus NN 37602 58 13 _ _ NNP 37602 58 14 Hypsiboas Hypsiboas NNP 37602 58 15 _ _ NNP 37602 58 16 Wagler Wagler NNP 37602 58 17 , , , 37602 58 18 1830 1830 CD 37602 58 19 . . . 37602 59 1 Cope Cope NNP 37602 59 2 ( ( -LRB- 37602 59 3 1874 1874 CD 37602 59 4 ) ) -RRB- 37602 59 5 named name VBN 37602 59 6 _ _ NNP 37602 59 7 Osteocephalus Osteocephalus NNP 37602 59 8 planiceps planicep NNS 37602 59 9 _ _ NNP 37602 59 10 from from IN 37602 59 11 Nauta Nauta NNP 37602 59 12 , , , 37602 59 13 Perú Perú NNP 37602 59 14 . . . 37602 60 1 The the DT 37602 60 2 single single JJ 37602 60 3 specimen speciman NNS 37602 60 4 was be VBD 37602 60 5 among among IN 37602 60 6 the the DT 37602 60 7 collections collection NNS 37602 60 8 made make VBN 37602 60 9 by by IN 37602 60 10 the the DT 37602 60 11 Orton Orton NNP 37602 60 12 Expedition Expedition NNP 37602 60 13 to to IN 37602 60 14 the the DT 37602 60 15 upper upper JJ 37602 60 16 Amazon Amazon NNP 37602 60 17 Basin Basin NNP 37602 60 18 and and CC 37602 60 19 was be VBD 37602 60 20 deposited deposit VBN 37602 60 21 in in IN 37602 60 22 the the DT 37602 60 23 Academy Academy NNP 37602 60 24 of of IN 37602 60 25 Natural Natural NNP 37602 60 26 Sciences Sciences NNPS 37602 60 27 in in IN 37602 60 28 Philadelphia Philadelphia NNP 37602 60 29 . . . 37602 61 1 Boulenger boulenger NN 37602 61 2 ( ( -LRB- 37602 61 3 1882 1882 CD 37602 61 4 ) ) -RRB- 37602 61 5 placed place VBD 37602 61 6 both both CC 37602 61 7 _ _ NNP 37602 61 8 Osteocephalus Osteocephalus NNP 37602 61 9 _ _ NNP 37602 61 10 and and CC 37602 61 11 _ _ NNP 37602 61 12 Trachycephalus Trachycephalus NNP 37602 61 13 _ _ NNP 37602 61 14 the the DT 37602 61 15 synonymy synonymy NN 37602 61 16 of of IN 37602 61 17 _ _ NNP 37602 61 18 Hyla Hyla NNP 37602 61 19 _ _ NNP 37602 61 20 ; ; : 37602 61 21 he -PRON- PRP 37602 61 22 recognized recognize VBD 37602 61 23 _ _ NNP 37602 61 24 Hyla Hyla NNP 37602 61 25 taurina taurina NNP 37602 61 26 _ _ NNP 37602 61 27 ( ( -LRB- 37602 61 28 with with IN 37602 61 29 _ _ NNP 37602 61 30 O. O. NNP 37602 62 1 flavolineatus flavolineatus NNP 37602 62 2 _ _ NNP 37602 62 3 as as IN 37602 62 4 a a DT 37602 62 5 synonym synonym NN 37602 62 6 ) ) -RRB- 37602 62 7 , , , 37602 62 8 _ _ NNP 37602 62 9 H. H. NNP 37602 63 1 leprieurii leprieurii JJ 37602 63 2 _ _ NNP 37602 63 3 , , , 37602 63 4 and and CC 37602 63 5 _ _ NNP 37602 63 6 H. H. NNP 37602 64 1 planiceps planiceps NNP 37602 64 2 _ _ NNP 37602 64 3 . . . 37602 65 1 In in IN 37602 65 2 the the DT 37602 65 3 same same JJ 37602 65 4 publication publication NN 37602 65 5 Boulenger Boulenger NNP 37602 65 6 named name VBN 37602 65 7 _ _ NNP 37602 65 8 Hyla Hyla NNP 37602 65 9 buckleyi buckleyi NN 37602 65 10 _ _ NNP 37602 65 11 on on IN 37602 65 12 the the DT 37602 65 13 basis basis NN 37602 65 14 of of IN 37602 65 15 10 10 CD 37602 65 16 specimens specimen NNS 37602 65 17 in in IN 37602 65 18 the the DT 37602 65 19 British British NNP 37602 65 20 Museum Museum NNP 37602 65 21 from from IN 37602 65 22 Ecuador Ecuador NNP 37602 65 23 ; ; : 37602 65 24 in in IN 37602 65 25 the the DT 37602 65 26 description description NN 37602 65 27 he -PRON- PRP 37602 65 28 stated state VBD 37602 65 29 that that IN 37602 65 30 _ _ NNP 37602 65 31 buckleyi buckleyi NN 37602 65 32 _ _ NNP 37602 65 33 was be VBD 37602 65 34 like like IN 37602 65 35 _ _ NNP 37602 65 36 leprieurii leprieurii JJ 37602 65 37 _ _ NNP 37602 65 38 and and CC 37602 65 39 _ _ NNP 37602 65 40 taurinus taurinus NN 37602 65 41 _ _ NNP 37602 65 42 in in IN 37602 65 43 having have VBG 37602 65 44 paired pair VBN 37602 65 45 lateral lateral JJ 37602 65 46 vocal vocal JJ 37602 65 47 sacs sac NNS 37602 65 48 . . . 37602 66 1 Boulenger boulenger NN 37602 66 2 held hold VBD 37602 66 3 a a DT 37602 66 4 lasting lasting JJ 37602 66 5 influence influence NN 37602 66 6 on on IN 37602 66 7 taxonomic taxonomic JJ 37602 66 8 herpetology herpetology NN 37602 66 9 , , , 37602 66 10 and and CC 37602 66 11 his -PRON- PRP$ 37602 66 12 generic generic JJ 37602 66 13 synonymy synonymy NN 37602 66 14 of of IN 37602 66 15 _ _ NNP 37602 66 16 Osteocephalus Osteocephalus NNP 37602 66 17 _ _ NNP 37602 66 18 was be VBD 37602 66 19 unchallenged unchallenged JJ 37602 66 20 until until IN 37602 66 21 only only RB 37602 66 22 a a DT 37602 66 23 decade decade NN 37602 66 24 ago ago RB 37602 66 25 . . . 37602 67 1 Goin go VB 37602 67 2 ( ( -LRB- 37602 67 3 1961 1961 CD 37602 67 4 ) ) -RRB- 37602 67 5 presented present VBD 37602 67 6 a a DT 37602 67 7 generic generic JJ 37602 67 8 synopsis synopsis NN 37602 67 9 of of IN 37602 67 10 the the DT 37602 67 11 genera genera NN 37602 67 12 of of IN 37602 67 13 hylid hylid NNP 37602 67 14 frogs frog NNS 37602 67 15 , , , 37602 67 16 in in IN 37602 67 17 which which WDT 37602 67 18 he -PRON- PRP 37602 67 19 recognized recognize VBD 37602 67 20 _ _ NNP 37602 67 21 Osteocephalus Osteocephalus NNP 37602 67 22 _ _ NNP 37602 67 23 and and CC 37602 67 24 stated state VBD 37602 67 25 : : : 37602 67 26 " " `` 37602 67 27 There there EX 37602 67 28 are be VBP 37602 67 29 perhaps perhaps RB 37602 67 30 eight eight CD 37602 67 31 or or CC 37602 67 32 ten ten CD 37602 67 33 species specie NNS 37602 67 34 of of IN 37602 67 35 this this DT 37602 67 36 genus genus NN 37602 67 37 in in IN 37602 67 38 South South NNP 37602 67 39 America America NNP 37602 67 40 . . . 37602 68 1 Certainly certainly RB 37602 68 2 _ _ NNP 37602 68 3 taurinus taurinus NN 37602 68 4 _ _ NNP 37602 68 5 , , , 37602 68 6 _ _ NNP 37602 68 7 britti britti NNP 37602 68 8 _ _ NNP 37602 68 9 , , , 37602 68 10 _ _ NNP 37602 68 11 leprieuri leprieuri NNP 37602 68 12 _ _ NNP 37602 68 13 , , , 37602 68 14 _ _ NNP 37602 68 15 buckleyi buckleyi NN 37602 68 16 _ _ NNP 37602 68 17 and and CC 37602 68 18 _ _ NNP 37602 68 19 pearsoni pearsoni NNP 37602 68 20 _ _ NNP 37602 68 21 belong belong VBP 37602 68 22 here here RB 37602 68 23 . . . 37602 69 1 _ _ NNP 37602 69 2 O. O. NNP 37602 70 1 planiceps planiceps NNP 37602 70 2 _ _ NNP 37602 70 3 is be VBZ 37602 70 4 surely surely RB 37602 70 5 a a DT 37602 70 6 synonym synonym NN 37602 70 7 of of IN 37602 70 8 _ _ NNP 37602 70 9 leprieuri leprieuri NN 37602 70 10 _ _ NNP 37602 70 11 and and CC 37602 70 12 I -PRON- PRP 37602 70 13 believe believe VBP 37602 70 14 that that IN 37602 70 15 _ _ NNP 37602 70 16 garbei garbei NNP 37602 70 17 _ _ NNP 37602 70 18 is be VBZ 37602 70 19 as as RB 37602 70 20 well well RB 37602 70 21 . . . 37602 71 1 The the DT 37602 71 2 status status NN 37602 71 3 of of IN 37602 71 4 such such JJ 37602 71 5 forms form NNS 37602 71 6 as as IN 37602 71 7 _ _ NNP 37602 71 8 macrotis macrotis NN 37602 71 9 _ _ NNP 37602 71 10 , , , 37602 71 11 _ _ NNP 37602 71 12 riopastazae riopastazae VBZ 37602 71 13 _ _ NNP 37602 71 14 , , , 37602 71 15 and and CC 37602 71 16 _ _ NNP 37602 71 17 depressa depressa NNP 37602 71 18 _ _ NNP 37602 71 19 has have VBZ 37602 71 20 not not RB 37602 71 21 yet yet RB 37602 71 22 been be VBN 37602 71 23 settled settle VBN 37602 71 24 . . . 37602 71 25 " " '' 37602 72 1 Goin go NNP 37602 72 2 defined define VBN 37602 72 3 _ _ NNP 37602 72 4 Osteocephalus Osteocephalus NNP 37602 72 5 _ _ NNP 37602 72 6 as as IN 37602 72 7 follows follow VBZ 37602 72 8 : : : 37602 72 9 " " `` 37602 72 10 Males male NNS 37602 72 11 with with IN 37602 72 12 paired pair VBN 37602 72 13 vocal vocal JJ 37602 72 14 pouches pouch NNS 37602 72 15 , , , 37602 72 16 one one CD 37602 72 17 at at IN 37602 72 18 each each DT 37602 72 19 angle angle NN 37602 72 20 of of IN 37602 72 21 the the DT 37602 72 22 jaw jaw NN 37602 72 23 ; ; : 37602 72 24 derm derm NN 37602 72 25 of of IN 37602 72 26 head head NN 37602 72 27 not not RB 37602 72 28 co co VB 37602 72 29 - - JJ 37602 72 30 ossified ossify VBN 37602 72 31 with with IN 37602 72 32 skull skull NN 37602 72 33 but but CC 37602 72 34 roof roof NNP 37602 72 35 of of IN 37602 72 36 skull skull NNP 37602 72 37 exostosed exostose VBD 37602 72 38 . . . 37602 72 39 " " '' 37602 73 1 Trueb Trueb NNP 37602 73 2 ( ( -LRB- 37602 73 3 1970a 1970a CD 37602 73 4 ) ) -RRB- 37602 73 5 elaborated elaborate VBN 37602 73 6 on on IN 37602 73 7 Goin go NNP 37602 73 8 's 's POS 37602 73 9 definition definition NN 37602 73 10 and and CC 37602 73 11 assuredly assuredly RB 37602 73 12 included include VBN 37602 73 13 only only RB 37602 73 14 _ _ NNP 37602 73 15 O. O. NNP 37602 74 1 taurinus taurinus NNP 37602 74 2 _ _ NNP 37602 74 3 and and CC 37602 74 4 _ _ NNP 37602 74 5 O. O. NNP 37602 75 1 leprieurii leprieurii JJ 37602 75 2 _ _ NNP 37602 75 3 in in IN 37602 75 4 the the DT 37602 75 5 genus genus NN 37602 75 6 . . . 37602 76 1 Goin go NNP 37602 76 2 's 's POS 37602 76 3 inclusion inclusion NN 37602 76 4 of of IN 37602 76 5 _ _ NNP 37602 76 6 buckleyi buckleyi NN 37602 76 7 _ _ NNP 37602 76 8 , , , 37602 76 9 _ _ NNP 37602 76 10 britti britti NNP 37602 76 11 _ _ NNP 37602 76 12 , , , 37602 76 13 and and CC 37602 76 14 _ _ NNP 37602 76 15 pearsoni pearsoni NNP 37602 76 16 _ _ NNP 37602 76 17 in in IN 37602 76 18 _ _ NNP 37602 76 19 Osteocephalus Osteocephalus NNP 37602 76 20 _ _ NNP 37602 76 21 was be VBD 37602 76 22 the the DT 37602 76 23 first first JJ 37602 76 24 association association NN 37602 76 25 of of IN 37602 76 26 any any DT 37602 76 27 of of IN 37602 76 28 these these DT 37602 76 29 names name NNS 37602 76 30 with with IN 37602 76 31 that that DT 37602 76 32 genus genus NN 37602 76 33 . . . 37602 77 1 Duellman duellman NN 37602 77 2 ( ( -LRB- 37602 77 3 1970a 1970a CD 37602 77 4 ) ) -RRB- 37602 77 5 demonstrated demonstrate VBD 37602 77 6 that that IN 37602 77 7 _ _ NNP 37602 77 8 Garbeana Garbeana NNP 37602 77 9 garbei garbei NN 37602 77 10 _ _ NNP 37602 77 11 Miranda Miranda NNP 37602 77 12 - - HYPH 37602 77 13 Ribeiro Ribeiro NNP 37602 77 14 , , , 37602 77 15 1926 1926 CD 37602 77 16 , , , 37602 77 17 was be VBD 37602 77 18 a a DT 37602 77 19 member member NN 37602 77 20 of of IN 37602 77 21 the the DT 37602 77 22 _ _ NNP 37602 77 23 Hyla Hyla NNP 37602 77 24 rubra rubra NN 37602 77 25 _ _ NNP 37602 77 26 group group NN 37602 77 27 . . . 37602 78 1 _ _ NNP 37602 78 2 Hyla Hyla NNP 37602 78 3 macrotis macrotis NN 37602 78 4 _ _ NNP 37602 78 5 Andersson Andersson NNP 37602 78 6 , , , 37602 78 7 1945 1945 CD 37602 78 8 , , , 37602 78 9 is be VBZ 37602 78 10 a a DT 37602 78 11 _ _ NNP 37602 78 12 Phrynohyas Phrynohyas NNP 37602 78 13 _ _ NNP 37602 78 14 . . . 37602 79 1 Trueb Trueb NNP 37602 79 2 and and CC 37602 79 3 Duellman Duellman NNP 37602 79 4 ( ( -LRB- 37602 79 5 1970 1970 CD 37602 79 6 ) ) -RRB- 37602 79 7 determined determine VBD 37602 79 8 that that IN 37602 79 9 _ _ NNP 37602 79 10 Hyla Hyla NNP 37602 79 11 verrucigera verrucigera NN 37602 79 12 _ _ NNP 37602 79 13 Werner Werner NNP 37602 79 14 , , , 37602 79 15 1901 1901 CD 37602 79 16 , , , 37602 79 17 is be VBZ 37602 79 18 the the DT 37602 79 19 earliest early JJS 37602 79 20 name name NN 37602 79 21 for for IN 37602 79 22 an an DT 37602 79 23 _ _ NNP 37602 79 24 Osteocephalus Osteocephalus NNP 37602 79 25 _ _ NNP 37602 79 26 displaying display VBG 37602 79 27 striking striking JJ 37602 79 28 sexual sexual JJ 37602 79 29 dimorphism dimorphism NN 37602 79 30 in in IN 37602 79 31 coloration coloration NN 37602 79 32 and and CC 37602 79 33 texture texture NN 37602 79 34 of of IN 37602 79 35 the the DT 37602 79 36 dorsal dorsal NN 37602 79 37 skin skin NN 37602 79 38 ; ; : 37602 79 39 _ _ NNP 37602 79 40 Hyla Hyla NNP 37602 79 41 riopastazae riopastazae VBZ 37602 79 42 _ _ NNP 37602 79 43 Andersson Andersson NNP 37602 79 44 , , , 37602 79 45 1945 1945 CD 37602 79 46 ( ( -LRB- 37602 79 47 female female JJ 37602 79 48 holotype holotype NN 37602 79 49 ) ) -RRB- 37602 79 50 , , , 37602 79 51 and and CC 37602 79 52 _ _ NNP 37602 79 53 Hyla Hyla NNP 37602 79 54 orcesi orcesi NN 37602 79 55 _ _ NNP 37602 79 56 Funkhouser Funkhouser NNP 37602 79 57 , , , 37602 79 58 1956 1956 CD 37602 79 59 ( ( -LRB- 37602 79 60 male male JJ 37602 79 61 holotype holotype NN 37602 79 62 ) ) -RRB- 37602 79 63 , , , 37602 79 64 were be VBD 37602 79 65 placed place VBN 37602 79 66 in in IN 37602 79 67 the the DT 37602 79 68 synonymy synonymy NN 37602 79 69 of of IN 37602 79 70 _ _ NNP 37602 79 71 Osteocephalus Osteocephalus NNP 37602 79 72 verrucigerus verrucigerus NN 37602 79 73 _ _ NNP 37602 79 74 . . . 37602 80 1 _ _ NNP 37602 80 2 Hyla Hyla NNP 37602 80 3 pearsoni pearsoni NN 37602 80 4 _ _ NNP 37602 80 5 Gaige Gaige NNP 37602 80 6 , , , 37602 80 7 1929 1929 CD 37602 80 8 , , , 37602 80 9 is be VBZ 37602 80 10 a a DT 37602 80 11 small small JJ 37602 80 12 species specie NNS 37602 80 13 of of IN 37602 80 14 _ _ NNP 37602 80 15 Osteocephalus Osteocephalus NNP 37602 80 16 _ _ NNP 37602 80 17 . . . 37602 81 1 Our -PRON- PRP$ 37602 81 2 findings finding NNS 37602 81 3 substantiate substantiate VBP 37602 81 4 Goin go NNP 37602 81 5 's 's POS 37602 81 6 suggestions suggestion NNS 37602 81 7 relative relative JJ 37602 81 8 to to IN 37602 81 9 two two CD 37602 81 10 other other JJ 37602 81 11 taxa taxa NN 37602 81 12 . . . 37602 82 1 _ _ NNP 37602 82 2 Hyla Hyla NNP 37602 82 3 leprieurii leprieurii JJ 37602 82 4 britti britti NNP 37602 82 5 _ _ NNP 37602 82 6 Melin Melin NNP 37602 82 7 , , , 37602 82 8 1941 1941 CD 37602 82 9 , , , 37602 82 10 from from IN 37602 82 11 the the DT 37602 82 12 Rio Rio NNP 37602 82 13 Uaupés Uaupés NNP 37602 82 14 , , , 37602 82 15 Brasil Brasil NNP 37602 82 16 , , , 37602 82 17 and and CC 37602 82 18 _ _ NNP 37602 82 19 Hyla Hyla NNP 37602 82 20 depressa depressa NN 37602 82 21 _ _ NNP 37602 82 22 Andersson Andersson NNP 37602 82 23 , , , 37602 82 24 1945 1945 CD 37602 82 25 , , , 37602 82 26 from from IN 37602 82 27 the the DT 37602 82 28 Río Río NNP 37602 82 29 Pastaza Pastaza NNP 37602 82 30 watershed watershe VBD 37602 82 31 , , , 37602 82 32 Ecuador Ecuador NNP 37602 82 33 , , , 37602 82 34 are be VBP 37602 82 35 members member NNS 37602 82 36 of of IN 37602 82 37 the the DT 37602 82 38 genus genus NNP 37602 82 39 _ _ NNP 37602 82 40 Osteocephalus Osteocephalus NNP 37602 82 41 _ _ NNP 37602 82 42 , , , 37602 82 43 but but CC 37602 82 44 both both DT 37602 82 45 are be VBP 37602 82 46 synonyms synonyms JJ 37602 82 47 of of IN 37602 82 48 earlier early JJR 37602 82 49 names--_leprieurii names--_leprieurii IN 37602 82 50 _ _ NNP 37602 82 51 and and CC 37602 82 52 _ _ NNP 37602 82 53 taurinus taurinus NN 37602 82 54 _ _ NNP 37602 82 55 , , , 37602 82 56 respectively respectively RB 37602 82 57 . . . 37602 83 1 Another another DT 37602 83 2 name name NN 37602 83 3 proposed propose VBN 37602 83 4 by by IN 37602 83 5 Melin Melin NNP 37602 83 6 ( ( -LRB- 37602 83 7 1941 1941 CD 37602 83 8 ) ) -RRB- 37602 83 9 , , , 37602 83 10 _ _ NNP 37602 83 11 Hyla Hyla NNP 37602 83 12 ( ( -LRB- 37602 83 13 Trachycephalus Trachycephalus NNP 37602 83 14 ) ) -RRB- 37602 83 15 vilarsi vilarsi NN 37602 83 16 _ _ NNP 37602 83 17 from from IN 37602 83 18 Taracuá Taracuá NNP 37602 83 19 , , , 37602 83 20 Brasil Brasil NNP 37602 83 21 , , , 37602 83 22 also also RB 37602 83 23 is be VBZ 37602 83 24 placed place VBN 37602 83 25 in in IN 37602 83 26 the the DT 37602 83 27 synonymy synonymy NN 37602 83 28 of of IN 37602 83 29 _ _ NNP 37602 83 30 O. O. NNP 37602 84 1 taurinus taurinus NNP 37602 84 2 _ _ NNP 37602 84 3 . . . 37602 85 1 Cochran Cochran NNP 37602 85 2 and and CC 37602 85 3 Goin go NNP 37602 85 4 ( ( -LRB- 37602 85 5 1970 1970 CD 37602 85 6 ) ) -RRB- 37602 85 7 were be VBD 37602 85 8 unaware unaware JJ 37602 85 9 of of IN 37602 85 10 the the DT 37602 85 11 identities identity NNS 37602 85 12 of of IN 37602 85 13 _ _ NNP 37602 85 14 Hyla Hyla NNP 37602 85 15 verrucigera verrucigera NN 37602 85 16 _ _ NNP 37602 85 17 and and CC 37602 85 18 _ _ NNP 37602 85 19 riopastazae riopastazae VBZ 37602 85 20 _ _ NNP 37602 85 21 ; ; : 37602 85 22 they -PRON- PRP 37602 85 23 used use VBD 37602 85 24 the the DT 37602 85 25 later later JJ 37602 85 26 name name NN 37602 85 27 _ _ NNP 37602 85 28 Osteocephalus Osteocephalus NNP 37602 85 29 orcesi orcesi NN 37602 85 30 _ _ NNP 37602 85 31 for for IN 37602 85 32 Colombian colombian JJ 37602 85 33 frogs frog NNS 37602 85 34 that that WDT 37602 85 35 are be VBP 37602 85 36 correctly correctly RB 37602 85 37 referred refer VBN 37602 85 38 to to IN 37602 85 39 _ _ NNP 37602 85 40 O. O. NNP 37602 86 1 verrucigerus verrucigerus NNP 37602 86 2 _ _ NNP 37602 86 3 . . . 37602 87 1 Although although IN 37602 87 2 Goin go NNP 37602 87 3 ( ( -LRB- 37602 87 4 1961 1961 CD 37602 87 5 ) ) -RRB- 37602 87 6 placed place VBD 37602 87 7 _ _ NNP 37602 87 8 Hyla Hyla NNP 37602 87 9 buckleyi buckleyi NN 37602 87 10 _ _ NNP 37602 87 11 and and CC 37602 87 12 _ _ NNP 37602 87 13 H. H. NNP 37602 88 1 pearsoni pearsoni NNP 37602 88 2 _ _ NNP 37602 88 3 in in IN 37602 88 4 _ _ NNP 37602 88 5 Osteocephalus Osteocephalus NNP 37602 88 6 _ _ NNP 37602 88 7 , , , 37602 88 8 Cochran Cochran NNP 37602 88 9 and and CC 37602 88 10 Goin go NNP 37602 88 11 ( ( -LRB- 37602 88 12 1970 1970 CD 37602 88 13 ) ) -RRB- 37602 88 14 recognized recognize VBD 37602 88 15 a a DT 37602 88 16 " " `` 37602 88 17 _ _ NNP 37602 88 18 buckleyi buckleyi NN 37602 88 19 _ _ NNP 37602 88 20 group group NN 37602 88 21 " " '' 37602 88 22 in in IN 37602 88 23 _ _ NNP 37602 88 24 Hyla Hyla NNP 37602 88 25 _ _ NNP 37602 88 26 that that WDT 37602 88 27 included include VBD 37602 88 28 these these DT 37602 88 29 two two CD 37602 88 30 species specie NNS 37602 88 31 plus plus CC 37602 88 32 a a DT 37602 88 33 new new JJ 37602 88 34 species specie NNS 37602 88 35 , , , 37602 88 36 _ _ NNP 37602 88 37 Hyla Hyla NNP 37602 88 38 cabrerai cabrerai NN 37602 88 39 _ _ NNP 37602 88 40 from from IN 37602 88 41 Amazonian Amazonian NNP 37602 88 42 Colombia Colombia NNP 37602 88 43 and and CC 37602 88 44 Brasil Brasil NNP 37602 88 45 ( ( -LRB- 37602 88 46 total total NN 37602 88 47 of of IN 37602 88 48 three three CD 37602 88 49 specimens specimen NNS 37602 88 50 ) ) -RRB- 37602 88 51 . . . 37602 89 1 Also also RB 37602 89 2 , , , 37602 89 3 these these DT 37602 89 4 authors author NNS 37602 89 5 named name VBN 37602 89 6 _ _ NNP 37602 89 7 Hyla Hyla NNP 37602 89 8 carri carri VBD 37602 89 9 _ _ NNP 37602 89 10 from from IN 37602 89 11 a a DT 37602 89 12 single single JJ 37602 89 13 Colombian colombian JJ 37602 89 14 specimen speciman NNS 37602 89 15 . . . 37602 90 1 Study study NN 37602 90 2 of of IN 37602 90 3 the the DT 37602 90 4 types type NNS 37602 90 5 of of IN 37602 90 6 _ _ NNP 37602 90 7 Hyla Hyla NNP 37602 90 8 cabrerai cabrerai NN 37602 90 9 _ _ NNP 37602 90 10 , , , 37602 90 11 _ _ NNP 37602 90 12 H. H. NNP 37602 91 1 carri carri NNP 37602 91 2 _ _ NNP 37602 91 3 , , , 37602 91 4 and and CC 37602 91 5 _ _ NNP 37602 91 6 H. H. NNP 37602 92 1 festae festae NNP 37602 92 2 _ _ NNP 37602 92 3 Peracca Peracca NNP 37602 92 4 , , , 37602 92 5 1904 1904 CD 37602 92 6 , , , 37602 92 7 from from IN 37602 92 8 Ecuador Ecuador NNP 37602 92 9 , , , 37602 92 10 reveal reveal VBP 37602 92 11 that that IN 37602 92 12 all all DT 37602 92 13 of of IN 37602 92 14 these these DT 37602 92 15 names name NNS 37602 92 16 are be VBP 37602 92 17 synonyms synonyms JJ 37602 92 18 of of IN 37602 92 19 _ _ NNP 37602 92 20 Osteocephalus Osteocephalus NNP 37602 92 21 buckleyi buckleyi NN 37602 92 22 _ _ NNP 37602 92 23 . . . 37602 93 1 Much much JJ 37602 93 2 of of IN 37602 93 3 the the DT 37602 93 4 taxonomic taxonomic JJ 37602 93 5 confusion confusion NN 37602 93 6 and and CC 37602 93 7 multiplicity multiplicity NN 37602 93 8 of of IN 37602 93 9 trivial trivial JJ 37602 93 10 names name NNS 37602 93 11 is be VBZ 37602 93 12 due due JJ 37602 93 13 to to IN 37602 93 14 the the DT 37602 93 15 great great JJ 37602 93 16 amount amount NN 37602 93 17 of of IN 37602 93 18 color color NN 37602 93 19 variation variation NN 37602 93 20 in in IN 37602 93 21 _ _ NNP 37602 93 22 taurinus taurinus NNP 37602 93 23 _ _ NNP 37602 93 24 and and CC 37602 93 25 to to IN 37602 93 26 the the DT 37602 93 27 sexual sexual JJ 37602 93 28 dimorphism dimorphism NN 37602 93 29 in in IN 37602 93 30 the the DT 37602 93 31 texture texture NN 37602 93 32 of of IN 37602 93 33 the the DT 37602 93 34 dorsal dorsal NN 37602 93 35 skin skin NN 37602 93 36 in in IN 37602 93 37 all all DT 37602 93 38 of of IN 37602 93 39 the the DT 37602 93 40 species specie NNS 37602 93 41 . . . 37602 94 1 The the DT 37602 94 2 details detail NNS 37602 94 3 of of IN 37602 94 4 variation variation NN 37602 94 5 in in IN 37602 94 6 these these DT 37602 94 7 and and CC 37602 94 8 other other JJ 37602 94 9 characters character NNS 37602 94 10 and and CC 37602 94 11 our -PRON- PRP$ 37602 94 12 justifications justification NNS 37602 94 13 for for IN 37602 94 14 the the DT 37602 94 15 synonymies synonymy NNS 37602 94 16 are be VBP 37602 94 17 given give VBN 37602 94 18 in in IN 37602 94 19 the the DT 37602 94 20 accounts account NNS 37602 94 21 of of IN 37602 94 22 the the DT 37602 94 23 species specie NNS 37602 94 24 . . . 37602 95 1 All all DT 37602 95 2 of of IN 37602 95 3 the the DT 37602 95 4 trivial trivial JJ 37602 95 5 names name NNS 37602 95 6 that that WDT 37602 95 7 apply apply VBP 37602 95 8 to to IN 37602 95 9 species species NNP 37602 95 10 herein herein NNP 37602 95 11 recognized recognize VBD 37602 95 12 as as IN 37602 95 13 members member NNS 37602 95 14 of of IN 37602 95 15 the the DT 37602 95 16 genus genus NNP 37602 95 17 _ _ NNP 37602 95 18 Osteocephalus Osteocephalus NNP 37602 95 19 _ _ NNP 37602 95 20 are be VBP 37602 95 21 listed list VBN 37602 95 22 in in IN 37602 95 23 table table NN 37602 95 24 1 1 CD 37602 95 25 . . . 37602 96 1 = = NFP 37602 96 2 Osteocephalus= Osteocephalus= NNP 37602 96 3 Steindachner Steindachner NNP 37602 96 4 , , , 37602 96 5 1862 1862 CD 37602 96 6 _ _ NNP 37602 96 7 Osteocephalus Osteocephalus NNP 37602 96 8 _ _ NNP 37602 96 9 Steindachner Steindachner NNP 37602 96 10 , , , 37602 96 11 1862:77 1862:77 CD 37602 96 12 [ [ -LRB- 37602 96 13 Type type NN 37602 96 14 species.--_Osteocephalus species.--_Osteocephalus NNP 37602 96 15 taurinus taurinus NN 37602 96 16 _ _ NNP 37602 96 17 Steindachner Steindachner NNP 37602 96 18 , , , 37602 96 19 1862 1862 CD 37602 96 20 , , , 37602 96 21 by by IN 37602 96 22 original original JJ 37602 96 23 designation designation NN 37602 96 24 ] ] -RRB- 37602 96 25 . . . 37602 97 1 Not not RB 37602 97 2 _ _ NNP 37602 97 3 Osteocephalus Osteocephalus NNP 37602 97 4 _ _ NNP 37602 97 5 Fitzinger Fitzinger NNP 37602 97 6 , , , 37602 97 7 1843:50 1843:50 NNP 37602 97 8 ( ( -LRB- 37602 97 9 _ _ NNP 37602 97 10 nomen noman NNS 37602 97 11 nudum nudum NNP 37602 97 12 _ _ NNP 37602 97 13 ) ) -RRB- 37602 97 14 . . . 37602 98 1 _ _ NNP 37602 98 2 Diagnostic Diagnostic NNP 37602 98 3 Definition._--1 Definition._--1 NNP 37602 98 4 ) ) -RRB- 37602 98 5 Skull Skull NNP 37602 98 6 broader broader RBR 37602 98 7 than than IN 37602 98 8 long long JJ 37602 98 9 ; ; : 37602 98 10 2 2 LS 37602 98 11 ) ) -RRB- 37602 98 12 dermal dermal JJ 37602 98 13 roofing roofing NN 37602 98 14 bones bone NNS 37602 98 15 of of IN 37602 98 16 skull skull NNP 37602 98 17 well well RB 37602 98 18 ossified ossify VBD 37602 98 19 , , , 37602 98 20 exostosed exostose VBN 37602 98 21 , , , 37602 98 22 and/or and/or CC 37602 98 23 co co JJ 37602 98 24 - - JJ 37602 98 25 ossified ossified JJ 37602 98 26 in in IN 37602 98 27 some some DT 37602 98 28 species specie NNS 37602 98 29 ; ; : 37602 98 30 3 3 LS 37602 98 31 ) ) -RRB- 37602 98 32 prenasal prenasal NN 37602 98 33 and and CC 37602 98 34 internasal internasal NN 37602 98 35 bones bone NNS 37602 98 36 absent absent JJ 37602 98 37 ; ; : 37602 98 38 4 4 LS 37602 98 39 ) ) -RRB- 37602 98 40 parasphenoid parasphenoid NN 37602 98 41 alae alae NNP 37602 98 42 posterolaterally posterolaterally RB 37602 98 43 oriented orient VBD 37602 98 44 ; ; : 37602 98 45 5 5 LS 37602 98 46 ) ) -RRB- 37602 98 47 dentigerous dentigerous JJ 37602 98 48 processes process NNS 37602 98 49 of of IN 37602 98 50 prevomers prevomer NNS 37602 98 51 angular angular JJ 37602 98 52 ( ( -LRB- 37602 98 53 /--\ /--\ NFP 37602 98 54 ) ) -RRB- 37602 98 55 ; ; : 37602 98 56 6 6 LS 37602 98 57 ) ) -RRB- 37602 98 58 vocal vocal NN 37602 98 59 sacs sac NNS 37602 98 60 paired pair VBN 37602 98 61 , , , 37602 98 62 posterior posterior NN 37602 98 63 , , , 37602 98 64 and and CC 37602 98 65 when when WRB 37602 98 66 inflated inflate VBN 37602 98 67 protruding protrude VBG 37602 98 68 posteroventral posteroventral JJ 37602 98 69 or or CC 37602 98 70 posterolateral posterolateral JJ 37602 98 71 to to IN 37602 98 72 angles angle NNS 37602 98 73 of of IN 37602 98 74 jaws jaw NNS 37602 98 75 ; ; : 37602 98 76 7 7 LS 37602 98 77 ) ) -RRB- 37602 98 78 submentalis submentalis NNP 37602 98 79 muscle muscle NN 37602 98 80 moderate moderate JJ 37602 98 81 in in IN 37602 98 82 size size NN 37602 98 83 and and CC 37602 98 84 araphic araphic NN 37602 98 85 ; ; : 37602 98 86 8) 8) CD 37602 98 87 intermandibularis intermandibularis NNP 37602 98 88 muscle muscle NN 37602 98 89 undifferentiated undifferentiate VBD 37602 98 90 and and CC 37602 98 91 bearing bear VBG 37602 98 92 an an DT 37602 98 93 elongate elongate NN 37602 98 94 median median JJ 37602 98 95 aponeurosis aponeurosis NN 37602 98 96 ; ; : 37602 98 97 9 9 LS 37602 98 98 ) ) -RRB- 37602 98 99 parotoid parotoid NN 37602 98 100 glands gland NNS 37602 98 101 absent absent JJ 37602 98 102 or or CC 37602 98 103 poorly poorly RB 37602 98 104 developed develop VBN 37602 98 105 , , , 37602 98 106 skin skin NN 37602 98 107 not not RB 37602 98 108 producing produce VBG 37602 98 109 viscous viscous JJ 37602 98 110 secretion secretion NN 37602 98 111 characteristic characteristic NN 37602 98 112 of of IN 37602 98 113 _ _ NNP 37602 98 114 Phrynohyas Phrynohyas NNP 37602 98 115 _ _ NNP 37602 98 116 ; ; : 37602 98 117 10 10 LS 37602 98 118 ) ) -RRB- 37602 98 119 skin skin NN 37602 98 120 on on IN 37602 98 121 dorsum dorsum NN 37602 98 122 tuberculate tuberculate NN 37602 98 123 in in IN 37602 98 124 males male NNS 37602 98 125 , , , 37602 98 126 smooth smooth JJ 37602 98 127 in in IN 37602 98 128 females female NNS 37602 98 129 ; ; : 37602 98 130 11 11 LS 37602 98 131 ) ) -RRB- 37602 98 132 tympanum tympanum VBD 37602 98 133 large large JJ 37602 98 134 , , , 37602 98 135 60 60 CD 37602 98 136 percent percent NN 37602 98 137 or or CC 37602 98 138 more more JJR 37602 98 139 of of IN 37602 98 140 diameter diameter NN 37602 98 141 of of IN 37602 98 142 eye eye NN 37602 98 143 ; ; : 37602 98 144 12 12 CD 37602 98 145 ) ) -RRB- 37602 98 146 fingers finger NNS 37602 98 147 about about IN 37602 98 148 one one CD 37602 98 149 - - HYPH 37602 98 150 third third NN 37602 98 151 , , , 37602 98 152 toes toe VBZ 37602 98 153 more more JJR 37602 98 154 than than IN 37602 98 155 three three CD 37602 98 156 - - HYPH 37602 98 157 fourths fourth NNS 37602 98 158 webbed web VBN 37602 98 159 ; ; : 37602 98 160 13 13 CD 37602 98 161 ) ) -RRB- 37602 98 162 discs disc NNS 37602 98 163 large large JJ 37602 98 164 , , , 37602 98 165 round round JJ 37602 98 166 ; ; : 37602 98 167 14 14 CD 37602 98 168 ) ) -RRB- 37602 98 169 nuptial nuptial JJ 37602 98 170 excrescences excrescence NNS 37602 98 171 present present JJ 37602 98 172 in in IN 37602 98 173 breeding breeding NN 37602 98 174 males male NNS 37602 98 175 ; ; : 37602 98 176 15 15 LS 37602 98 177 ) ) -RRB- 37602 98 178 inner inner JJ 37602 98 179 metatarsal metatarsal NN 37602 98 180 tubercle tubercle NN 37602 98 181 not not RB 37602 98 182 modified modify VBN 37602 98 183 for for IN 37602 98 184 digging dig VBG 37602 98 185 ; ; : 37602 98 186 16 16 LS 37602 98 187 ) ) -RRB- 37602 98 188 outer outer JJ 37602 98 189 metatarsal metatarsal NN 37602 98 190 tubercle tubercle NN 37602 98 191 absent absent NN 37602 98 192 ; ; : 37602 98 193 17 17 CD 37602 98 194 ) ) -RRB- 37602 98 195 tarsal tarsal NN 37602 98 196 fold fold VB 37602 98 197 weak weak JJ 37602 98 198 or or CC 37602 98 199 absent absent JJ 37602 98 200 ; ; : 37602 98 201 18 18 CD 37602 98 202 ) ) -RRB- 37602 98 203 pupil pupil NN 37602 98 204 horizontal horizontal NN 37602 98 205 ; ; : 37602 98 206 19 19 CD 37602 98 207 ) ) -RRB- 37602 98 208 palpebrum palpebrum NN 37602 98 209 clear clear JJ 37602 98 210 ; ; : 37602 98 211 20 20 CD 37602 98 212 ) ) -RRB- 37602 98 213 known know VBN 37602 98 214 tadpoles tadpole NNS 37602 98 215 having have VBG 37602 98 216 two two CD 37602 98 217 upper upper JJ 37602 98 218 and and CC 37602 98 219 five five CD 37602 98 220 lower low JJR 37602 98 221 rows row NNS 37602 98 222 of of IN 37602 98 223 teeth tooth NNS 37602 98 224 . . . 37602 99 1 TABLE TABLE NNP 37602 99 2 1.--Alphabetical 1.--alphabetical CD 37602 99 3 Synonymy Synonymy NNPS 37602 99 4 of of IN 37602 99 5 the the DT 37602 99 6 Species Species NNPS 37602 99 7 of of IN 37602 99 8 _ _ NNP 37602 99 9 Osteocephalus Osteocephalus NNP 37602 99 10 _ _ NNP 37602 99 11 . . . 37602 100 1 Trivial Trivial NNP 37602 100 2 Name Name NNP 37602 100 3 , , , 37602 100 4 Original Original NNP 37602 100 5 Generic Generic NNP 37602 100 6 Name Name NNP 37602 100 7 , , , 37602 100 8 Author author NN 37602 100 9 , , , 37602 100 10 Date Date NNP 37602 100 11 Current Current NNP 37602 100 12 Name Name NNP 37602 100 13 ---------------------------------------------------------------------- ---------------------------------------------------------------------- NFP 37602 100 14 _ _ NNP 37602 100 15 britti britti NNP 37602 100 16 ( ( -LRB- 37602 100 17 Hyla Hyla NNP 37602 100 18 leprieurii leprieurii JJ 37602 100 19 ) ) -RRB- 37602 100 20 _ _ NNP 37602 100 21 Melin Melin NNP 37602 100 22 , , , 37602 100 23 1941 1941 CD 37602 100 24 _ _ NNP 37602 100 25 O. O. NNP 37602 101 1 leprieurii leprieurii NNP 37602 101 2 _ _ NNP 37602 101 3 _ _ NNP 37602 101 4 buckleyi buckleyi NN 37602 101 5 ( ( -LRB- 37602 101 6 Hyla Hyla NNP 37602 101 7 ) ) -RRB- 37602 101 8 _ _ NNP 37602 101 9 Boulenger Boulenger NNP 37602 101 10 , , , 37602 101 11 1882 1882 CD 37602 101 12 _ _ NNP 37602 101 13 O. O. NNP 37602 102 1 buckleyi buckleyi NNP 37602 102 2 _ _ NNP 37602 102 3 _ _ NNP 37602 102 4 cabrerai cabrerai NN 37602 102 5 ( ( -LRB- 37602 102 6 Hyla Hyla NNP 37602 102 7 ) ) -RRB- 37602 102 8 _ _ NNP 37602 102 9 Cochran Cochran NNP 37602 102 10 and and CC 37602 102 11 Goin go NNP 37602 102 12 , , , 37602 102 13 1970 1970 CD 37602 102 14 _ _ NNP 37602 102 15 O. O. NNP 37602 103 1 buckleyi buckleyi NNP 37602 103 2 _ _ NNP 37602 103 3 _ _ NNP 37602 103 4 carri carri NN 37602 103 5 ( ( -LRB- 37602 103 6 Hyla Hyla NNP 37602 103 7 ) ) -RRB- 37602 103 8 _ _ NNP 37602 103 9 Cochran Cochran NNP 37602 103 10 and and CC 37602 103 11 Goin go NNP 37602 103 12 , , , 37602 103 13 1970 1970 CD 37602 103 14 _ _ NNP 37602 103 15 O. O. NNP 37602 104 1 buckleyi buckleyi NNP 37602 104 2 _ _ NNP 37602 104 3 _ _ NNP 37602 104 4 depressa depressa NNP 37602 104 5 ( ( -LRB- 37602 104 6 Hyla Hyla NNP 37602 104 7 ) ) -RRB- 37602 104 8 _ _ NNP 37602 104 9 Andersson Andersson NNP 37602 104 10 , , , 37602 104 11 1945 1945 CD 37602 104 12 _ _ NNP 37602 104 13 O. O. NNP 37602 105 1 taurinus taurinus NNP 37602 105 2 _ _ NNP 37602 105 3 _ _ NNP 37602 105 4 festae festae NN 37602 105 5 ( ( -LRB- 37602 105 6 Hyla Hyla NNP 37602 105 7 ) ) -RRB- 37602 105 8 _ _ NNP 37602 105 9 Peracca Peracca NNP 37602 105 10 , , , 37602 105 11 1904 1904 CD 37602 105 12 _ _ NNP 37602 105 13 O. O. NNP 37602 106 1 buckleyi buckleyi NNP 37602 106 2 _ _ NNP 37602 106 3 _ _ NNP 37602 106 4 flavolineatus flavolineatus NN 37602 106 5 ( ( -LRB- 37602 106 6 Osteocephalus osteocephalus NN 37602 106 7 ) ) -RRB- 37602 106 8 _ _ NNP 37602 106 9 Steindachner Steindachner NNP 37602 106 10 , , , 37602 106 11 1862 1862 CD 37602 106 12 _ _ NNP 37602 106 13 O. O. NNP 37602 107 1 taurinus taurinus NNP 37602 107 2 _ _ NNP 37602 107 3 _ _ NNP 37602 107 4 leprieurii leprieurii VBZ 37602 107 5 ( ( -LRB- 37602 107 6 Hyla Hyla NNP 37602 107 7 ) ) -RRB- 37602 107 8 _ _ NNP 37602 107 9 Duméril Duméril NNP 37602 107 10 and and CC 37602 107 11 Bibron Bibron NNP 37602 107 12 , , , 37602 107 13 1841 1841 CD 37602 107 14 _ _ NNP 37602 107 15 O. O. NNP 37602 108 1 leprieurii leprieurii NNP 37602 108 2 _ _ NNP 37602 108 3 _ _ NNP 37602 108 4 orcesi orcesi NN 37602 108 5 ( ( -LRB- 37602 108 6 Hyla Hyla NNP 37602 108 7 ) ) -RRB- 37602 108 8 _ _ NNP 37602 108 9 Funkhouser Funkhouser NNP 37602 108 10 , , , 37602 108 11 1956 1956 CD 37602 108 12 _ _ NNP 37602 108 13 O. O. NNP 37602 109 1 verrucigerus verrucigerus NNP 37602 109 2 _ _ NNP 37602 109 3 _ _ NNP 37602 109 4 pearsoni pearsoni NNP 37602 109 5 ( ( -LRB- 37602 109 6 Hyla Hyla NNP 37602 109 7 ) ) -RRB- 37602 109 8 _ _ NNP 37602 109 9 Gaige Gaige NNP 37602 109 10 , , , 37602 109 11 1929 1929 CD 37602 109 12 _ _ NNP 37602 109 13 O. O. NNP 37602 110 1 pearsoni pearsoni NNP 37602 110 2 _ _ NNP 37602 110 3 _ _ NNP 37602 110 4 planiceps planicep VBZ 37602 110 5 ( ( -LRB- 37602 110 6 Osteocephalus Osteocephalus NNP 37602 110 7 ) ) -RRB- 37602 110 8 _ _ NNP 37602 110 9 Cope Cope NNP 37602 110 10 , , , 37602 110 11 1874 1874 CD 37602 110 12 _ _ NNP 37602 110 13 O. O. NNP 37602 111 1 taurinus taurinus NNP 37602 111 2 _ _ NNP 37602 111 3 _ _ NNP 37602 111 4 riopastazae riopastazae VBZ 37602 111 5 ( ( -LRB- 37602 111 6 Hyla Hyla NNP 37602 111 7 ) ) -RRB- 37602 111 8 _ _ NNP 37602 111 9 Andersson Andersson NNP 37602 111 10 , , , 37602 111 11 1945 1945 CD 37602 111 12 _ _ NNP 37602 111 13 O. O. NNP 37602 112 1 verrucigerus verrucigerus NNP 37602 112 2 _ _ NNP 37602 112 3 _ _ NNP 37602 112 4 taurinus taurinus NN 37602 112 5 ( ( -LRB- 37602 112 6 Osteocephalus osteocephalus JJ 37602 112 7 ) ) -RRB- 37602 112 8 _ _ NNP 37602 112 9 Steindachner Steindachner NNP 37602 112 10 , , , 37602 112 11 1862 1862 CD 37602 112 12 _ _ NNP 37602 112 13 O. O. NNP 37602 113 1 taurinus taurinus NNP 37602 113 2 _ _ NNP 37602 113 3 _ _ NNP 37602 113 4 verrucigera verrucigera NN 37602 113 5 ( ( -LRB- 37602 113 6 Hyla Hyla NNP 37602 113 7 ) ) -RRB- 37602 113 8 _ _ NNP 37602 113 9 Werner Werner NNP 37602 113 10 , , , 37602 113 11 1901 1901 CD 37602 113 12 _ _ NNP 37602 113 13 O. O. NNP 37602 114 1 verrucigerus verrucigerus NNP 37602 114 2 _ _ NNP 37602 114 3 _ _ NNP 37602 114 4 vilarsi vilarsi NN 37602 114 5 ( ( -LRB- 37602 114 6 Hyla Hyla NNP 37602 114 7 ) ) -RRB- 37602 114 8 _ _ NNP 37602 114 9 Melin Melin NNP 37602 114 10 , , , 37602 114 11 1941 1941 CD 37602 114 12 _ _ NNP 37602 114 13 O. O. NNP 37602 115 1 taurinus taurinus NNP 37602 115 2 _ _ NNP 37602 115 3 _ _ NNP 37602 115 4 Content._--As content._--as NN 37602 115 5 defined define VBN 37602 115 6 here here RB 37602 115 7 , , , 37602 115 8 the the DT 37602 115 9 genus genus NN 37602 115 10 contains contain VBZ 37602 115 11 five five CD 37602 115 12 known know VBN 37602 115 13 species specie NNS 37602 115 14 : : : 37602 115 15 _ _ NNP 37602 115 16 O. O. NNP 37602 116 1 buckleyi buckleyi NNP 37602 116 2 _ _ NNP 37602 116 3 ( ( -LRB- 37602 116 4 Boulenger Boulenger NNP 37602 116 5 ) ) -RRB- 37602 116 6 , , , 37602 116 7 _ _ NNP 37602 116 8 O. O. NNP 37602 117 1 leprieurii leprieurii NNP 37602 117 2 _ _ NNP 37602 117 3 ( ( -LRB- 37602 117 4 Duméril Duméril NNP 37602 117 5 and and CC 37602 117 6 Bibron Bibron NNP 37602 117 7 ) ) -RRB- 37602 117 8 , , , 37602 117 9 _ _ NNP 37602 117 10 O. O. NNP 37602 118 1 pearsoni pearsoni NNP 37602 118 2 _ _ NNP 37602 118 3 ( ( -LRB- 37602 118 4 Gaige Gaige NNP 37602 118 5 ) ) -RRB- 37602 118 6 , , , 37602 118 7 _ _ NNP 37602 118 8 O. O. NNP 37602 119 1 taurinus taurinus NNP 37602 119 2 _ _ NNP 37602 119 3 Steindachner Steindachner NNP 37602 119 4 , , , 37602 119 5 and and CC 37602 119 6 _ _ NNP 37602 119 7 O. O. NNP 37602 120 1 verrucigerus verrucigerus NNP 37602 120 2 _ _ NNP 37602 120 3 ( ( -LRB- 37602 120 4 Werner Werner NNP 37602 120 5 ) ) -RRB- 37602 120 6 . . . 37602 121 1 _ _ NNP 37602 121 2 Distribution._--The Distribution._--The NNP 37602 121 3 Guianas Guianas NNP 37602 121 4 and and CC 37602 121 5 Amazon Amazon NNP 37602 121 6 Basin Basin NNP 37602 121 7 ; ; : 37602 121 8 also also RB 37602 121 9 in in IN 37602 121 10 the the DT 37602 121 11 upper upper JJ 37602 121 12 Orinoco Orinoco NNP 37602 121 13 and and CC 37602 121 14 Magdalena Magdalena NNP 37602 121 15 drainages drainage NNS 37602 121 16 . . . 37602 122 1 Most Most JJS 37602 122 2 localities locality NNS 37602 122 3 are be VBP 37602 122 4 at at IN 37602 122 5 elevations elevation NNS 37602 122 6 below below IN 37602 122 7 500 500 CD 37602 122 8 m m NNS 37602 122 9 , , , 37602 122 10 but but CC 37602 122 11 the the DT 37602 122 12 genus genus NN 37602 122 13 ascends ascend VBZ 37602 122 14 the the DT 37602 122 15 Amazonian amazonian JJ 37602 122 16 slopes slope NNS 37602 122 17 of of IN 37602 122 18 the the DT 37602 122 19 Andes Andes NNP 37602 122 20 to to IN 37602 122 21 elevations elevation NNS 37602 122 22 of of IN 37602 122 23 about about RB 37602 122 24 1800 1800 CD 37602 122 25 m. m. NN 37602 122 26 ANALYSIS ANALYSIS NNP 37602 122 27 OF of IN 37602 122 28 CHARACTERS character NNS 37602 122 29 _ _ NNP 37602 122 30 Size Size NNP 37602 122 31 and and CC 37602 122 32 Proportions._--Frogs proportions._--frog NNS 37602 122 33 of of IN 37602 122 34 the the DT 37602 122 35 genus genus NNP 37602 122 36 _ _ NNP 37602 122 37 Osteocephalus Osteocephalus NNP 37602 122 38 _ _ NNP 37602 122 39 are be VBP 37602 122 40 moderate moderate JJ 37602 122 41 to to IN 37602 122 42 large large JJ 37602 122 43 hylids hylid NNS 37602 122 44 . . . 37602 123 1 The the DT 37602 123 2 largest large JJS 37602 123 3 species species NN 37602 123 4 is be VBZ 37602 123 5 _ _ NNP 37602 123 6 taurinus taurinus NN 37602 123 7 _ _ NNP 37602 123 8 , , , 37602 123 9 attaining attain VBG 37602 123 10 a a DT 37602 123 11 snout snout NN 37602 123 12 - - HYPH 37602 123 13 vent vent NN 37602 123 14 length length NN 37602 123 15 of of IN 37602 123 16 103.1 103.1 CD 37602 123 17 mm mm NNS 37602 123 18 ; ; : 37602 123 19 the the DT 37602 123 20 smallest small JJS 37602 123 21 is be VBZ 37602 123 22 _ _ NNP 37602 123 23 pearsoni pearsoni NNP 37602 123 24 _ _ NNP 37602 123 25 , , , 37602 123 26 which which WDT 37602 123 27 attains attain VBZ 37602 123 28 a a DT 37602 123 29 length length NN 37602 123 30 of of IN 37602 123 31 54.7 54.7 CD 37602 123 32 mm mm NNS 37602 123 33 . . . 37602 124 1 Considerable considerable JJ 37602 124 2 intraspecific intraspecific JJ 37602 124 3 geographic geographic JJ 37602 124 4 variation variation NN 37602 124 5 occurs occur VBZ 37602 124 6 in in IN 37602 124 7 adult adult NN 37602 124 8 size size NN 37602 124 9 , , , 37602 124 10 especially especially RB 37602 124 11 in in IN 37602 124 12 _ _ NNP 37602 124 13 taurinus taurinus NNP 37602 124 14 _ _ NNP 37602 124 15 . . . 37602 125 1 Females female NNS 37602 125 2 of of IN 37602 125 3 all all DT 37602 125 4 species specie NNS 37602 125 5 attain attain VBP 37602 125 6 a a DT 37602 125 7 noticeably noticeably RB 37602 125 8 larger large JJR 37602 125 9 size size NN 37602 125 10 than than IN 37602 125 11 males male NNS 37602 125 12 , , , 37602 125 13 but but CC 37602 125 14 no no DT 37602 125 15 significant significant JJ 37602 125 16 differences difference NNS 37602 125 17 are be VBP 37602 125 18 apparent apparent JJ 37602 125 19 in in IN 37602 125 20 proportions proportion NNS 37602 125 21 ( ( -LRB- 37602 125 22 Table table NN 37602 125 23 2 2 CD 37602 125 24 ) ) -RRB- 37602 125 25 . . . 37602 126 1 TABLE TABLE NNP 37602 126 2 2.--Comparison 2.--comparison CD 37602 126 3 of of IN 37602 126 4 Size Size NNP 37602 126 5 and and CC 37602 126 6 Proportions Proportions NNPS 37602 126 7 in in IN 37602 126 8 the the DT 37602 126 9 Species Species NNPS 37602 126 10 of of IN 37602 126 11 _ _ NNP 37602 126 12 Osteocephalus Osteocephalus NNP 37602 126 13 _ _ NNP 37602 126 14 . . . 37602 127 1 ( ( -LRB- 37602 127 2 Means mean NNS 37602 127 3 are be VBP 37602 127 4 given give VBN 37602 127 5 in in IN 37602 127 6 parentheses parenthesis NNS 37602 127 7 below below IN 37602 127 8 observed observed JJ 37602 127 9 ranges range NNS 37602 127 10 ) ) -RRB- 37602 127 11 = = NFP 37602 127 12 = = NFP 37602 127 13 = = SYM 37602 127 14 = = SYM 37602 127 15 = = SYM 37602 127 16 = = SYM 37602 127 17 = = SYM 37602 127 18 = = SYM 37602 127 19 = = SYM 37602 127 20 = = SYM 37602 127 21 = = SYM 37602 127 22 = = SYM 37602 127 23 = = SYM 37602 127 24 = = SYM 37602 127 25 = = SYM 37602 127 26 = = SYM 37602 127 27 = = SYM 37602 127 28 = = SYM 37602 127 29 = = SYM 37602 127 30 = = SYM 37602 127 31 = = SYM 37602 127 32 = = SYM 37602 127 33 = = SYM 37602 127 34 = = SYM 37602 127 35 = = SYM 37602 127 36 = = SYM 37602 127 37 = = SYM 37602 127 38 = = SYM 37602 127 39 = = SYM 37602 127 40 = = SYM 37602 127 41 = = SYM 37602 127 42 = = SYM 37602 127 43 = = SYM 37602 127 44 = = SYM 37602 127 45 = = SYM 37602 127 46 = = SYM 37602 127 47 = = SYM 37602 127 48 = = SYM 37602 127 49 = = SYM 37602 127 50 = = SYM 37602 127 51 = = SYM 37602 127 52 = = SYM 37602 127 53 = = SYM 37602 127 54 = = SYM 37602 127 55 = = SYM 37602 127 56 = = SYM 37602 127 57 = = SYM 37602 127 58 = = SYM 37602 127 59 = = SYM 37602 127 60 = = SYM 37602 127 61 = = SYM 37602 127 62 = = SYM 37602 127 63 = = SYM 37602 127 64 = = SYM 37602 127 65 = = SYM 37602 127 66 = = SYM 37602 127 67 = = SYM 37602 127 68 = = SYM 37602 127 69 = = SYM 37602 127 70 = = SYM 37602 127 71 = = SYM 37602 127 72 = = SYM 37602 127 73 = = SYM 37602 127 74 = = SYM 37602 127 75 = = SYM 37602 127 76 = = SYM 37602 127 77 = = SYM 37602 127 78 = = SYM 37602 127 79 = = SYM 37602 127 80 = = SYM 37602 127 81 = = SYM 37602 127 82 = = SYM 37602 127 83 = = SYM 37602 127 84 = = SYM 37602 127 85 = = SYM 37602 127 86 = = SYM 37602 127 87 = = SYM 37602 127 88 = = SYM 37602 127 89 = = SYM 37602 127 90 = = SYM 37602 127 91 = = SYM 37602 127 92 = = SYM 37602 127 93 = = SYM 37602 127 94 = = SYM 37602 127 95 = = SYM 37602 127 96 = = SYM 37602 127 97 = = SYM 37602 127 98 = = SYM 37602 127 99 = = SYM 37602 127 100 = = SYM 37602 127 101 = = SYM 37602 127 102 = = SYM 37602 127 103 = = SYM 37602 127 104 = = SYM 37602 127 105 = = SYM 37602 127 106 = = SYM 37602 127 107 = = SYM 37602 127 108 = = SYM 37602 127 109 = = SYM 37602 127 110 = = SYM 37602 127 111 = = SYM 37602 127 112 = = SYM 37602 127 113 = = NFP 37602 127 114 = = NFP 37602 127 115 Species Species NNP 37602 127 116 N N NNP 37602 127 117 Snout Snout NNP 37602 127 118 - - HYPH 37602 127 119 vent vent NN 37602 127 120 Tibia Tibia NNP 37602 127 121 Length/ Length/ NNP 37602 127 122 Foot Foot NNP 37602 127 123 Length/ Length/ -RRB- 37602 127 124 Head Head NNP 37602 127 125 Length/ Length/ -RRB- 37602 127 126 Head Head NNP 37602 127 127 Width/ width/ ADD 37602 127 128 Tympanum/ tympanum/ PRP 37602 127 129 Length Length NNP 37602 127 130 S S NNP 37602 127 131 - - HYPH 37602 127 132 V V NNP 37602 127 133 L L NNP 37602 127 134 S S NNP 37602 127 135 - - HYPH 37602 127 136 V V NNP 37602 127 137 L L NNP 37602 127 138 S S NNP 37602 127 139 - - HYPH 37602 127 140 V V NNP 37602 127 141 L L NNP 37602 127 142 S S NNP 37602 127 143 - - HYPH 37602 127 144 V V NNP 37602 127 145 L L NNP 37602 127 146 Eye Eye NNP 37602 127 147 -------------------------------------------------------------------------------------------------------- -------------------------------------------------------------------------------------------------------- NNP 37602 127 148 _ _ NNP 37602 127 149 O. O. NNP 37602 128 1 buckleyi buckleyi NNP 37602 128 2 _ _ NNP 37602 128 3 [ [ -LRB- 37602 128 4 M M NNP 37602 128 5 ] ] -RRB- 37602 128 6 30 30 CD 37602 128 7 37.9 37.9 CD 37602 128 8 - - SYM 37602 128 9 48.1 48.1 CD 37602 128 10 0.478 0.478 CD 37602 128 11 - - SYM 37602 128 12 0.580 0.580 CD 37602 128 13 0.375 0.375 CD 37602 128 14 - - SYM 37602 128 15 0.444 0.444 CD 37602 128 16 0.319 0.319 CD 37602 128 17 - - HYPH 37602 128 18 0.357 0.357 CD 37602 128 19 0.329 0.329 CD 37602 128 20 - - HYPH 37602 128 21 0.368 0.368 CD 37602 128 22 0.608 0.608 CD 37602 128 23 - - HYPH 37602 128 24 0.820 0.820 CD 37602 128 25 ( ( -LRB- 37602 128 26 43.3 43.3 CD 37602 128 27 ) ) -RRB- 37602 128 28 ( ( -LRB- 37602 128 29 0.520 0.520 CD 37602 128 30 ) ) -RRB- 37602 128 31 ( ( -LRB- 37602 128 32 0.408 0.408 CD 37602 128 33 ) ) -RRB- 37602 128 34 ( ( -LRB- 37602 128 35 0.343 0.343 CD 37602 128 36 ) ) -RRB- 37602 128 37 ( ( -LRB- 37602 128 38 0.351 0.351 CD 37602 128 39 ) ) -RRB- 37602 128 40 ( ( -LRB- 37602 128 41 0.711 0.711 CD 37602 128 42 ) ) -RRB- 37602 128 43 [ [ -LRB- 37602 128 44 F f NN 37602 128 45 ] ] -RRB- 37602 128 46 31 31 CD 37602 128 47 48.6 48.6 CD 37602 128 48 - - SYM 37602 128 49 75.1 75.1 CD 37602 128 50 0.476 0.476 CD 37602 128 51 - - HYPH 37602 128 52 0.599 0.599 CD 37602 128 53 0.363 0.363 CD 37602 128 54 - - HYPH 37602 128 55 0.469 0.469 CD 37602 128 56 0.310 0.310 CD 37602 128 57 - - HYPH 37602 128 58 0.358 0.358 CD 37602 128 59 0.318 0.318 CD 37602 128 60 - - HYPH 37602 128 61 0.367 0.367 CD 37602 128 62 0.574 0.574 CD 37602 128 63 - - HYPH 37602 128 64 0.905 0.905 CD 37602 128 65 ( ( -LRB- 37602 128 66 61.7 61.7 CD 37602 128 67 ) ) -RRB- 37602 128 68 ( ( -LRB- 37602 128 69 0.553 0.553 CD 37602 128 70 ) ) -RRB- 37602 128 71 ( ( -LRB- 37602 128 72 0.428 0.428 CD 37602 128 73 ) ) -RRB- 37602 128 74 ( ( -LRB- 37602 128 75 0.333 0.333 CD 37602 128 76 ) ) -RRB- 37602 128 77 ( ( -LRB- 37602 128 78 0.348 0.348 CD 37602 128 79 ) ) -RRB- 37602 128 80 ( ( -LRB- 37602 128 81 0.734 0.734 CD 37602 128 82 ) ) -RRB- 37602 128 83 _ _ NNP 37602 128 84 O. O. NNP 37602 129 1 leprieurii leprieurii NNP 37602 129 2 _ _ NNP 37602 129 3 [ [ -LRB- 37602 129 4 M M NNP 37602 129 5 ] ] -RRB- 37602 129 6 21 21 CD 37602 129 7 41.2 41.2 CD 37602 129 8 - - SYM 37602 129 9 48.4 48.4 CD 37602 129 10 0.514 0.514 CD 37602 129 11 - - HYPH 37602 129 12 0.571 0.571 CD 37602 129 13 0.383 0.383 CD 37602 129 14 - - SYM 37602 129 15 0.430 0.430 CD 37602 129 16 0.308 0.308 CD 37602 129 17 - - HYPH 37602 129 18 0.357 0.357 CD 37602 129 19 0.326 0.326 CD 37602 129 20 - - HYPH 37602 129 21 0.368 0.368 CD 37602 129 22 0.652 0.652 CD 37602 129 23 - - HYPH 37602 129 24 0.884 0.884 CD 37602 129 25 ( ( -LRB- 37602 129 26 44.7 44.7 CD 37602 129 27 ) ) -RRB- 37602 129 28 ( ( -LRB- 37602 129 29 0.538 0.538 CD 37602 129 30 ) ) -RRB- 37602 129 31 ( ( -LRB- 37602 129 32 0.408 0.408 CD 37602 129 33 ) ) -RRB- 37602 129 34 ( ( -LRB- 37602 129 35 0.335 0.335 CD 37602 129 36 ) ) -RRB- 37602 129 37 ( ( -LRB- 37602 129 38 0.348 0.348 CD 37602 129 39 ) ) -RRB- 37602 129 40 ( ( -LRB- 37602 129 41 0.777 0.777 CD 37602 129 42 ) ) -RRB- 37602 129 43 [ [ -LRB- 37602 129 44 F f NN 37602 129 45 ] ] -RRB- 37602 129 46 21 21 CD 37602 129 47 46.6 46.6 CD 37602 129 48 - - HYPH 37602 129 49 61.5 61.5 CD 37602 129 50 0.516 0.516 CD 37602 129 51 - - SYM 37602 129 52 0.592 0.592 CD 37602 129 53 0.382 0.382 CD 37602 129 54 - - HYPH 37602 129 55 0.453 0.453 CD 37602 129 56 0.314 0.314 CD 37602 129 57 - - HYPH 37602 129 58 0.343 0.343 CD 37602 129 59 0.328 0.328 CD 37602 129 60 - - HYPH 37602 129 61 0.363 0.363 CD 37602 129 62 0.698 0.698 CD 37602 129 63 - - HYPH 37602 129 64 0.909 0.909 CD 37602 129 65 ( ( -LRB- 37602 129 66 57.1 57.1 CD 37602 129 67 ) ) -RRB- 37602 129 68 ( ( -LRB- 37602 129 69 0.539 0.539 CD 37602 129 70 ) ) -RRB- 37602 129 71 ( ( -LRB- 37602 129 72 0.404 0.404 CD 37602 129 73 ) ) -RRB- 37602 129 74 ( ( -LRB- 37602 129 75 0.329 0.329 CD 37602 129 76 ) ) -RRB- 37602 129 77 ( ( -LRB- 37602 129 78 0.349 0.349 CD 37602 129 79 ) ) -RRB- 37602 129 80 ( ( -LRB- 37602 129 81 0.785 0.785 CD 37602 129 82 ) ) -RRB- 37602 129 83 _ _ NNP 37602 129 84 O. O. NNP 37602 130 1 pearsoni pearsoni NNP 37602 130 2 _ _ NNP 37602 130 3 [ [ -LRB- 37602 130 4 M m NN 37602 130 5 ] ] -RRB- 37602 130 6 2 2 CD 37602 130 7 45.3 45.3 CD 37602 130 8 - - SYM 37602 130 9 46.2 46.2 CD 37602 130 10 0.481 0.481 CD 37602 130 11 - - SYM 37602 130 12 0.504 0.504 CD 37602 130 13 0.404 0.404 CD 37602 130 14 - - SYM 37602 130 15 0.437 0.437 CD 37602 130 16 0.322 0.322 CD 37602 130 17 - - HYPH 37602 130 18 0.335 0.335 CD 37602 130 19 0.327 0.327 CD 37602 130 20 - - HYPH 37602 130 21 0.342 0.342 CD 37602 130 22 0.660 0.660 CD 37602 130 23 - - HYPH 37602 130 24 0.673 0.673 CD 37602 130 25 ( ( -LRB- 37602 130 26 45.8 45.8 CD 37602 130 27 ) ) -RRB- 37602 130 28 ( ( -LRB- 37602 130 29 0.493 0.493 CD 37602 130 30 ) ) -RRB- 37602 130 31 ( ( -LRB- 37602 130 32 0.421 0.421 CD 37602 130 33 ) ) -RRB- 37602 130 34 ( ( -LRB- 37602 130 35 0.329 0.329 CD 37602 130 36 ) ) -RRB- 37602 130 37 ( ( -LRB- 37602 130 38 0.335 0.335 CD 37602 130 39 ) ) -RRB- 37602 130 40 ( ( -LRB- 37602 130 41 0.666 0.666 CD 37602 130 42 ) ) -RRB- 37602 130 43 [ [ -LRB- 37602 130 44 F f NN 37602 130 45 ] ] -RRB- 37602 130 46 1 1 CD 37602 130 47 54.7 54.7 CD 37602 130 48 0.521 0.521 CD 37602 130 49 0.405 0.405 CD 37602 130 50 0.318 0.318 CD 37602 130 51 0.346 0.346 CD 37602 130 52 0.862 0.862 CD 37602 130 53 _ _ NNP 37602 130 54 O. O. NNP 37602 131 1 taurinus taurinus NNP 37602 131 2 _ _ NNP 37602 131 3 [ [ -LRB- 37602 131 4 M M NNP 37602 131 5 ] ] -RRB- 37602 131 6 59 59 CD 37602 131 7 40.3 40.3 CD 37602 131 8 - - SYM 37602 131 9 84.6 84.6 CD 37602 131 10 0.512 0.512 CD 37602 131 11 - - SYM 37602 131 12 0.576 0.576 CD 37602 131 13 0.387 0.387 CD 37602 131 14 - - HYPH 37602 131 15 0.445 0.445 CD 37602 131 16 0.296 0.296 CD 37602 131 17 - - SYM 37602 131 18 0.345 0.345 CD 37602 131 19 0.301 0.301 CD 37602 131 20 - - SYM 37602 131 21 0.355 0.355 CD 37602 131 22 0.638 0.638 CD 37602 131 23 - - HYPH 37602 131 24 0.896 0.896 CD 37602 131 25 ( ( -LRB- 37602 131 26 66.3 66.3 CD 37602 131 27 ) ) -RRB- 37602 131 28 ( ( -LRB- 37602 131 29 0.541 0.541 CD 37602 131 30 ) ) -RRB- 37602 131 31 ( ( -LRB- 37602 131 32 0.416 0.416 CD 37602 131 33 ) ) -RRB- 37602 131 34 ( ( -LRB- 37602 131 35 0.318 0.318 CD 37602 131 36 ) ) -RRB- 37602 131 37 ( ( -LRB- 37602 131 38 0.324 0.324 CD 37602 131 39 ) ) -RRB- 37602 131 40 ( ( -LRB- 37602 131 41 0.752 0.752 CD 37602 131 42 ) ) -RRB- 37602 131 43 [ [ -LRB- 37602 131 44 F f NN 37602 131 45 ] ] -RRB- 37602 131 46 45 45 CD 37602 131 47 45.1 45.1 CD 37602 131 48 - - SYM 37602 131 49 103.1 103.1 CD 37602 131 50 0.520 0.520 CD 37602 131 51 - - HYPH 37602 131 52 0.577 0.577 CD 37602 131 53 0.391 0.391 CD 37602 131 54 - - HYPH 37602 131 55 0.448 0.448 CD 37602 131 56 0.306 0.306 CD 37602 131 57 - - SYM 37602 131 58 0.334 0.334 CD 37602 131 59 0.308 0.308 CD 37602 131 60 - - HYPH 37602 131 61 0.347 0.347 CD 37602 131 62 0.640 0.640 CD 37602 131 63 - - HYPH 37602 131 64 0.817 0.817 CD 37602 131 65 ( ( -LRB- 37602 131 66 75.8 75.8 CD 37602 131 67 ) ) -RRB- 37602 131 68 ( ( -LRB- 37602 131 69 0.542 0.542 CD 37602 131 70 ) ) -RRB- 37602 131 71 ( ( -LRB- 37602 131 72 0.420 0.420 CD 37602 131 73 ) ) -RRB- 37602 131 74 ( ( -LRB- 37602 131 75 0.321 0.321 CD 37602 131 76 ) ) -RRB- 37602 131 77 ( ( -LRB- 37602 131 78 0.327 0.327 CD 37602 131 79 ) ) -RRB- 37602 131 80 ( ( -LRB- 37602 131 81 0.758 0.758 CD 37602 131 82 ) ) -RRB- 37602 131 83 _ _ NNP 37602 131 84 O. O. NNP 37602 132 1 verrucigerus verrucigerus NNP 37602 132 2 _ _ NNP 37602 132 3 [ [ -LRB- 37602 132 4 M M NNP 37602 132 5 ] ] -RRB- 37602 132 6 11 11 CD 37602 132 7 50.4 50.4 CD 37602 132 8 - - HYPH 37602 132 9 54.3 54.3 CD 37602 132 10 0.494 0.494 CD 37602 132 11 - - HYPH 37602 132 12 0.552 0.552 CD 37602 132 13 0.409 0.409 CD 37602 132 14 - - HYPH 37602 132 15 0.442 0.442 CD 37602 132 16 0.322 0.322 CD 37602 132 17 - - HYPH 37602 132 18 0.346 0.346 CD 37602 132 19 0.328 0.328 CD 37602 132 20 - - HYPH 37602 132 21 0.344 0.344 CD 37602 132 22 0.623 0.623 CD 37602 132 23 - - HYPH 37602 132 24 0.804 0.804 CD 37602 132 25 ( ( -LRB- 37602 132 26 53.0 53.0 CD 37602 132 27 ) ) -RRB- 37602 132 28 ( ( -LRB- 37602 132 29 0.519 0.519 CD 37602 132 30 ) ) -RRB- 37602 132 31 ( ( -LRB- 37602 132 32 0.427 0.427 CD 37602 132 33 ) ) -RRB- 37602 132 34 ( ( -LRB- 37602 132 35 0.333 0.333 CD 37602 132 36 ) ) -RRB- 37602 132 37 ( ( -LRB- 37602 132 38 0.337 0.337 CD 37602 132 39 ) ) -RRB- 37602 132 40 ( ( -LRB- 37602 132 41 0.730 0.730 CD 37602 132 42 ) ) -RRB- 37602 132 43 [ [ -LRB- 37602 132 44 F f NN 37602 132 45 ] ] -RRB- 37602 132 46 3 3 CD 37602 132 47 63.1 63.1 CD 37602 132 48 - - HYPH 37602 132 49 65.8 65.8 CD 37602 132 50 0.532 0.532 CD 37602 132 51 - - HYPH 37602 132 52 0.561 0.561 CD 37602 132 53 0.435 0.435 CD 37602 132 54 - - SYM 37602 132 55 0.463 0.463 CD 37602 132 56 0.345 0.345 CD 37602 132 57 - - HYPH 37602 132 58 0.347 0.347 CD 37602 132 59 0.348 0.348 CD 37602 132 60 - - HYPH 37602 132 61 0.379 0.379 CD 37602 132 62 0.692 0.692 CD 37602 132 63 - - HYPH 37602 132 64 0.808 0.808 CD 37602 132 65 ( ( -LRB- 37602 132 66 64.5 64.5 CD 37602 132 67 ) ) -RRB- 37602 132 68 ( ( -LRB- 37602 132 69 0.545 0.545 CD 37602 132 70 ) ) -RRB- 37602 132 71 ( ( -LRB- 37602 132 72 0.448 0.448 CD 37602 132 73 ) ) -RRB- 37602 132 74 ( ( -LRB- 37602 132 75 0.346 0.346 CD 37602 132 76 ) ) -RRB- 37602 132 77 ( ( -LRB- 37602 132 78 0.358 0.358 CD 37602 132 79 ) ) -RRB- 37602 132 80 ( ( -LRB- 37602 132 81 0.731 0.731 CD 37602 132 82 ) ) -RRB- 37602 132 83 -------------------------------------------------------------------------------------------------------- -------------------------------------------------------------------------------------------------------- . 37602 132 84 _ _ NNP 37602 132 85 Coloration._--All Coloration._--All NNP 37602 132 86 _ _ NNP 37602 132 87 Osteocephalus Osteocephalus NNP 37602 132 88 _ _ NNP 37602 132 89 are be VBP 37602 132 90 predominantly predominantly RB 37602 132 91 brown brown JJ 37602 132 92 frogs frog NNS 37602 132 93 usually usually RB 37602 132 94 with with IN 37602 132 95 some some DT 37602 132 96 darker dark JJR 37602 132 97 dorsal dorsal NN 37602 132 98 markings marking NNS 37602 132 99 ( ( -LRB- 37602 132 100 Figs fig NNS 37602 132 101 . . . 37602 133 1 1 1 CD 37602 133 2 and and CC 37602 133 3 2 2 CD 37602 133 4 ) ) -RRB- 37602 133 5 . . . 37602 134 1 _ _ NNP 37602 134 2 Osteocephalus Osteocephalus NNP 37602 134 3 verrucigerus verrucigerus NN 37602 134 4 _ _ NNP 37602 134 5 has have VBZ 37602 134 6 a a DT 37602 134 7 nearly nearly RB 37602 134 8 uniform uniform JJ 37602 134 9 dark dark JJ 37602 134 10 brown brown JJ 37602 134 11 dorsum dorsum NN 37602 134 12 and and CC 37602 134 13 no no DT 37602 134 14 distinct distinct JJ 37602 134 15 transverse transverse JJ 37602 134 16 bars bar NNS 37602 134 17 on on IN 37602 134 18 the the DT 37602 134 19 limbs limb NNS 37602 134 20 , , , 37602 134 21 whereas whereas IN 37602 134 22 all all DT 37602 134 23 of of IN 37602 134 24 the the DT 37602 134 25 other other JJ 37602 134 26 species specie NNS 37602 134 27 have have VBP 37602 134 28 distinct distinct JJ 37602 134 29 bars bar NNS 37602 134 30 on on IN 37602 134 31 the the DT 37602 134 32 limbs limb NNS 37602 134 33 . . . 37602 135 1 The the DT 37602 135 2 dorsal dorsal NN 37602 135 3 markings marking NNS 37602 135 4 on on IN 37602 135 5 the the DT 37602 135 6 body body NN 37602 135 7 consist consist VB 37602 135 8 of of IN 37602 135 9 irregular irregular JJ 37602 135 10 blotches blotch NNS 37602 135 11 in in IN 37602 135 12 _ _ NNP 37602 135 13 buckleyi buckleyi NN 37602 135 14 _ _ NNP 37602 135 15 , , , 37602 135 16 _ _ NNP 37602 135 17 pearsoni pearsoni NNP 37602 135 18 _ _ NNP 37602 135 19 , , , 37602 135 20 and and CC 37602 135 21 _ _ NNP 37602 135 22 taurinus taurinus NN 37602 135 23 _ _ NNP 37602 135 24 but but CC 37602 135 25 are be VBP 37602 135 26 narrow narrow JJ 37602 135 27 transverse transverse JJ 37602 135 28 marks mark NNS 37602 135 29 in in IN 37602 135 30 _ _ NNP 37602 135 31 leprieurii leprieurii JJ 37602 135 32 _ _ NNP 37602 135 33 . . . 37602 136 1 A a DT 37602 136 2 narrow narrow JJ 37602 136 3 middorsal middorsal NN 37602 136 4 cream cream NN 37602 136 5 or or CC 37602 136 6 yellow yellow JJ 37602 136 7 stripe stripe NN 37602 136 8 is be VBZ 37602 136 9 present present JJ 37602 136 10 in in IN 37602 136 11 some some DT 37602 136 12 individuals individual NNS 37602 136 13 of of IN 37602 136 14 _ _ NNP 37602 136 15 buckleyi buckleyi NN 37602 136 16 _ _ NNP 37602 136 17 and and CC 37602 136 18 _ _ NNP 37602 136 19 taurinus taurinus NN 37602 136 20 _ _ NNP 37602 136 21 but but CC 37602 136 22 absent absent JJ 37602 136 23 in in IN 37602 136 24 all all DT 37602 136 25 individuals individual NNS 37602 136 26 of of IN 37602 136 27 the the DT 37602 136 28 other other JJ 37602 136 29 species specie NNS 37602 136 30 . . . 37602 137 1 The the DT 37602 137 2 flanks flank NNS 37602 137 3 are be VBP 37602 137 4 uniform uniform JJ 37602 137 5 pale pale JJ 37602 137 6 tan tan NNP 37602 137 7 in in IN 37602 137 8 _ _ NNP 37602 137 9 leprieurii leprieurii JJ 37602 137 10 _ _ NNP 37602 137 11 and and CC 37602 137 12 uniform uniform JJ 37602 137 13 reddish reddish JJ 37602 137 14 brown brown NNP 37602 137 15 in in IN 37602 137 16 _ _ NNP 37602 137 17 verrucigerus verrucigerus NNP 37602 137 18 _ _ NNP 37602 137 19 ; ; : 37602 137 20 in in IN 37602 137 21 the the DT 37602 137 22 other other JJ 37602 137 23 species specie NNS 37602 137 24 the the DT 37602 137 25 flanks flank NNS 37602 137 26 are be VBP 37602 137 27 cream cream NN 37602 137 28 to to TO 37602 137 29 brown brown VB 37602 137 30 with with IN 37602 137 31 dark dark JJ 37602 137 32 brown brown JJ 37602 137 33 or or CC 37602 137 34 black black JJ 37602 137 35 spots spot NNS 37602 137 36 ( ( -LRB- 37602 137 37 also also RB 37602 137 38 dark dark JJ 37602 137 39 diagonal diagonal JJ 37602 137 40 marks mark NNS 37602 137 41 in in IN 37602 137 42 some some DT 37602 137 43 _ _ NNP 37602 137 44 buckleyi buckleyi NN 37602 137 45 _ _ NNP 37602 137 46 ) ) -RRB- 37602 137 47 . . . 37602 138 1 A a DT 37602 138 2 creamy creamy JJ 37602 138 3 white white JJ 37602 138 4 anal anal JJ 37602 138 5 stripe stripe NN 37602 138 6 is be VBZ 37602 138 7 present present JJ 37602 138 8 in in IN 37602 138 9 some some DT 37602 138 10 specimens specimen NNS 37602 138 11 of of IN 37602 138 12 _ _ NNP 37602 138 13 leprieurii leprieurii JJ 37602 138 14 _ _ NNP 37602 138 15 but but CC 37602 138 16 absent absent JJ 37602 138 17 in in IN 37602 138 18 all all DT 37602 138 19 individuals individual NNS 37602 138 20 of of IN 37602 138 21 other other JJ 37602 138 22 species specie NNS 37602 138 23 . . . 37602 139 1 The the DT 37602 139 2 postocular postocular JJ 37602 139 3 region region NN 37602 139 4 , , , 37602 139 5 encompassing encompass VBG 37602 139 6 the the DT 37602 139 7 tympanum tympanum NN 37602 139 8 , , , 37602 139 9 is be VBZ 37602 139 10 dark dark JJ 37602 139 11 brown brown JJ 37602 139 12 in in IN 37602 139 13 most most JJS 37602 139 14 specimens specimen NNS 37602 139 15 . . . 37602 140 1 In in IN 37602 140 2 adults adult NNS 37602 140 3 of of IN 37602 140 4 _ _ NNP 37602 140 5 pearsoni pearsoni NNP 37602 140 6 _ _ NNP 37602 140 7 and and CC 37602 140 8 _ _ NNP 37602 140 9 taurinus taurinus NN 37602 140 10 _ _ NNP 37602 140 11 the the DT 37602 140 12 upper upper JJ 37602 140 13 lips lip NNS 37602 140 14 are be VBP 37602 140 15 dark dark JJ 37602 140 16 brown brown JJ 37602 140 17 . . . 37602 141 1 A a DT 37602 141 2 pale pale JJ 37602 141 3 cream cream NN 37602 141 4 or or CC 37602 141 5 tan tan NN 37602 141 6 suborbital suborbital JJ 37602 141 7 spot spot NN 37602 141 8 is be VBZ 37602 141 9 present present JJ 37602 141 10 in in IN 37602 141 11 _ _ NNP 37602 141 12 pearsoni pearsoni NNP 37602 141 13 _ _ NNP 37602 141 14 and and CC 37602 141 15 in in IN 37602 141 16 some some DT 37602 141 17 _ _ NNP 37602 141 18 taurinus taurinus NN 37602 141 19 _ _ NNP 37602 141 20 ; ; : 37602 141 21 in in IN 37602 141 22 some some DT 37602 141 23 specimens specimen NNS 37602 141 24 of of IN 37602 141 25 _ _ NNP 37602 141 26 taurinus taurinus NN 37602 141 27 _ _ NNP 37602 141 28 the the DT 37602 141 29 suborbital suborbital JJ 37602 141 30 spot spot NN 37602 141 31 is be VBZ 37602 141 32 expanded expand VBN 37602 141 33 posteriorly posteriorly RB 37602 141 34 forming form VBG 37602 141 35 a a DT 37602 141 36 labial labial JJ 37602 141 37 stripe stripe NN 37602 141 38 on on IN 37602 141 39 the the DT 37602 141 40 posterior posterior JJ 37602 141 41 part part NN 37602 141 42 of of IN 37602 141 43 the the DT 37602 141 44 lip lip NN 37602 141 45 . . . 37602 142 1 The the DT 37602 142 2 labial labial JJ 37602 142 3 markings marking NNS 37602 142 4 of of IN 37602 142 5 _ _ NNP 37602 142 6 verrucigerus verrucigerus NN 37602 142 7 _ _ NNP 37602 142 8 are be VBP 37602 142 9 similar similar JJ 37602 142 10 to to IN 37602 142 11 the the DT 37602 142 12 latter latter JJ 37602 142 13 pattern pattern NN 37602 142 14 , , , 37602 142 15 except except IN 37602 142 16 that that IN 37602 142 17 in in IN 37602 142 18 females female NNS 37602 142 19 a a DT 37602 142 20 distinct distinct JJ 37602 142 21 , , , 37602 142 22 light light JJ 37602 142 23 labial labial JJ 37602 142 24 stripe stripe NN 37602 142 25 extends extend VBZ 37602 142 26 the the DT 37602 142 27 length length NN 37602 142 28 of of IN 37602 142 29 the the DT 37602 142 30 lip lip NN 37602 142 31 . . . 37602 143 1 _ _ NNP 37602 143 2 Osteocephalus Osteocephalus NNP 37602 143 3 leprieurii leprieurii JJ 37602 143 4 _ _ NNP 37602 143 5 has have VBZ 37602 143 6 a a DT 37602 143 7 distinct distinct JJ 37602 143 8 , , , 37602 143 9 broad broad JJ 37602 143 10 , , , 37602 143 11 pale pale JJ 37602 143 12 labial labial JJ 37602 143 13 stripe stripe NN 37602 143 14 . . . 37602 144 1 The the DT 37602 144 2 lips lip NNS 37602 144 3 are be VBP 37602 144 4 barred bar VBN 37602 144 5 cream cream NN 37602 144 6 and and CC 37602 144 7 dark dark JJ 37602 144 8 brown brown NN 37602 144 9 in in IN 37602 144 10 _ _ NNP 37602 144 11 buckleyi buckleyi NN 37602 144 12 _ _ NNP 37602 144 13 . . . 37602 145 1 The the DT 37602 145 2 venter venter NN 37602 145 3 is be VBZ 37602 145 4 uniform uniform JJ 37602 145 5 creamy creamy JJ 37602 145 6 white white JJ 37602 145 7 or or CC 37602 145 8 pale pale JJ 37602 145 9 tan tan NN 37602 145 10 in in IN 37602 145 11 _ _ NNP 37602 145 12 leprieurii leprieurii JJ 37602 145 13 _ _ NNP 37602 145 14 , , , 37602 145 15 uniform uniform JJ 37602 145 16 white white JJ 37602 145 17 in in IN 37602 145 18 some some DT 37602 145 19 _ _ NNP 37602 145 20 buckleyi buckleyi NN 37602 145 21 _ _ NNP 37602 145 22 ( ( -LRB- 37602 145 23 most most JJS 37602 145 24 males male NNS 37602 145 25 ) ) -RRB- 37602 145 26 , , , 37602 145 27 and and CC 37602 145 28 uniform uniform JJ 37602 145 29 tan tan NNP 37602 145 30 in in IN 37602 145 31 some some DT 37602 145 32 _ _ NNP 37602 145 33 taurinus taurinus NN 37602 145 34 _ _ NNP 37602 145 35 . . . 37602 146 1 The the DT 37602 146 2 other other JJ 37602 146 3 species specie NNS 37602 146 4 and and CC 37602 146 5 some some DT 37602 146 6 individuals individual NNS 37602 146 7 of of IN 37602 146 8 _ _ NNP 37602 146 9 taurinus taurinus NNP 37602 146 10 _ _ NNP 37602 146 11 and and CC 37602 146 12 _ _ NNP 37602 146 13 buckleyi buckleyi NN 37602 146 14 _ _ NNP 37602 146 15 ( ( -LRB- 37602 146 16 most most JJS 37602 146 17 females female NNS 37602 146 18 ) ) -RRB- 37602 146 19 have have VBP 37602 146 20 dark dark JJ 37602 146 21 ventral ventral JJ 37602 146 22 markings marking NNS 37602 146 23 . . . 37602 147 1 These these DT 37602 147 2 markings marking NNS 37602 147 3 are be VBP 37602 147 4 most most RBS 37602 147 5 distinctive distinctive JJ 37602 147 6 in in IN 37602 147 7 _ _ NNP 37602 147 8 verrucigerus verrucigerus NN 37602 147 9 _ _ NNP 37602 147 10 , , , 37602 147 11 in in IN 37602 147 12 which which WDT 37602 147 13 the the DT 37602 147 14 venter venter NN 37602 147 15 is be VBZ 37602 147 16 white white JJ 37602 147 17 with with IN 37602 147 18 bold bold JJ 37602 147 19 black black JJ 37602 147 20 mottling mottling NN 37602 147 21 and and CC 37602 147 22 spots spot NNS 37602 147 23 ( ( -LRB- 37602 147 24 Fig fig NN 37602 147 25 . . . 37602 148 1 3c 3c LS 37602 148 2 ) ) -RRB- 37602 148 3 . . . 37602 149 1 Those those DT 37602 149 2 individuals individual NNS 37602 149 3 of of IN 37602 149 4 _ _ NNP 37602 149 5 taurinus taurinus NN 37602 149 6 _ _ NNP 37602 149 7 having have VBG 37602 149 8 ventral ventral JJ 37602 149 9 markings marking NNS 37602 149 10 usually usually RB 37602 149 11 have have VBP 37602 149 12 indistinct indistinct JJ 37602 149 13 , , , 37602 149 14 diffuse diffuse VB 37602 149 15 brown brown JJ 37602 149 16 spots spot NNS 37602 149 17 on on IN 37602 149 18 the the DT 37602 149 19 throat throat NN 37602 149 20 and and CC 37602 149 21 chest chest NN 37602 149 22 ( ( -LRB- 37602 149 23 Fig fig NN 37602 149 24 . . . 37602 150 1 3b 3b LS 37602 150 2 ) ) -RRB- 37602 150 3 . . . 37602 151 1 _ _ NNP 37602 151 2 Osteocephalus osteocephalus JJ 37602 151 3 pearsoni pearsoni NN 37602 151 4 _ _ NNP 37602 151 5 is be VBZ 37602 151 6 characterized characterize VBN 37602 151 7 by by IN 37602 151 8 a a DT 37602 151 9 fine fine JJ 37602 151 10 brown brown JJ 37602 151 11 reticulation reticulation NN 37602 151 12 on on IN 37602 151 13 the the DT 37602 151 14 venter venter NN 37602 151 15 and and CC 37602 151 16 on on IN 37602 151 17 the the DT 37602 151 18 anterior anterior NN 37602 151 19 and and CC 37602 151 20 posterior posterior NN 37602 151 21 surfaces surface NNS 37602 151 22 of of IN 37602 151 23 the the DT 37602 151 24 thighs thigh NNS 37602 151 25 in in IN 37602 151 26 adults adult NNS 37602 151 27 ( ( -LRB- 37602 151 28 Fig fig NN 37602 151 29 . . . 37602 152 1 3a 3a LS 37602 152 2 ) ) -RRB- 37602 152 3 . . . 37602 153 1 Individuals individual NNS 37602 153 2 of of IN 37602 153 3 _ _ NNP 37602 153 4 buckleyi buckleyi NN 37602 153 5 _ _ NNP 37602 153 6 that that WDT 37602 153 7 have have VBP 37602 153 8 ventral ventral JJ 37602 153 9 markings marking NNS 37602 153 10 vary vary VBP 37602 153 11 between between IN 37602 153 12 the the DT 37602 153 13 patterns pattern NNS 37602 153 14 illustrated illustrate VBN 37602 153 15 for for IN 37602 153 16 _ _ NNP 37602 153 17 pearsoni pearsoni NNP 37602 153 18 _ _ NNP 37602 153 19 and and CC 37602 153 20 _ _ NNP 37602 153 21 taurinus taurinus NN 37602 153 22 _ _ NNP 37602 153 23 ( ( -LRB- 37602 153 24 Figs Figs NNP 37602 153 25 . . . 37602 154 1 3b 3b NNP 37602 154 2 and and CC 37602 154 3 c c NNP 37602 154 4 ) ) -RRB- 37602 154 5 . . . 37602 155 1 Ontogenetic ontogenetic JJ 37602 155 2 change change NN 37602 155 3 in in IN 37602 155 4 coloration coloration NN 37602 155 5 is be VBZ 37602 155 6 slight slight JJ 37602 155 7 or or CC 37602 155 8 non non JJ 37602 155 9 - - JJ 37602 155 10 existent existent NN 37602 155 11 in in IN 37602 155 12 _ _ NNP 37602 155 13 buckleyi buckleyi NN 37602 155 14 _ _ NNP 37602 155 15 , , , 37602 155 16 _ _ NNP 37602 155 17 pearsoni pearsoni NNP 37602 155 18 _ _ NNP 37602 155 19 , , , 37602 155 20 and and CC 37602 155 21 _ _ NNP 37602 155 22 taurinus taurinus NN 37602 155 23 _ _ NNP 37602 155 24 , , , 37602 155 25 except except IN 37602 155 26 that that DT 37602 155 27 juveniles juvenile NNS 37602 155 28 lack lack VBP 37602 155 29 ventral ventral JJ 37602 155 30 markings marking NNS 37602 155 31 . . . 37602 156 1 A a DT 37602 156 2 dark dark JJ 37602 156 3 blotch blotch NN 37602 156 4 on on IN 37602 156 5 the the DT 37602 156 6 back back JJ 37602 156 7 and and CC 37602 156 8 distinct distinct JJ 37602 156 9 transverse transverse JJ 37602 156 10 bars bar NNS 37602 156 11 on on IN 37602 156 12 the the DT 37602 156 13 limbs limb NNS 37602 156 14 are be VBP 37602 156 15 evident evident JJ 37602 156 16 in in IN 37602 156 17 juveniles juvenile NNS 37602 156 18 of of IN 37602 156 19 _ _ NNP 37602 156 20 verrucigerus verrucigerus NNP 37602 156 21 _ _ NNP 37602 156 22 ; ; : 37602 156 23 these these DT 37602 156 24 markings marking NNS 37602 156 25 are be VBP 37602 156 26 obscured obscure VBN 37602 156 27 in in IN 37602 156 28 the the DT 37602 156 29 adults adult NNS 37602 156 30 . . . 37602 157 1 Juveniles Juveniles NNP 37602 157 2 of of IN 37602 157 3 _ _ NNP 37602 157 4 leprieurii leprieurii JJ 37602 157 5 _ _ NNP 37602 157 6 are be VBP 37602 157 7 olive olive JJ 37602 157 8 - - HYPH 37602 157 9 brown brown JJ 37602 157 10 with with IN 37602 157 11 yellow yellow JJ 37602 157 12 dorsolateral dorsolateral JJ 37602 157 13 stripes stripe NNS 37602 157 14 ; ; : 37602 157 15 the the DT 37602 157 16 transverse transverse JJ 37602 157 17 dark dark JJ 37602 157 18 marks mark NNS 37602 157 19 , , , 37602 157 20 characteristic characteristic JJ 37602 157 21 of of IN 37602 157 22 the the DT 37602 157 23 adults adult NNS 37602 157 24 , , , 37602 157 25 appear appear VB 37602 157 26 before before IN 37602 157 27 the the DT 37602 157 28 stripes stripe NNS 37602 157 29 are be VBP 37602 157 30 lost lose VBN 37602 157 31 . . . 37602 158 1 [ [ -LRB- 37602 158 2 Illustration illustration NN 37602 158 3 : : : 37602 158 4 FIG FIG NNP 37602 158 5 . . . 37602 159 1 1 1 LS 37602 159 2 . . . 37602 160 1 Species Species NNP 37602 160 2 of of IN 37602 160 3 _ _ NNP 37602 160 4 Osteocephalus Osteocephalus NNP 37602 160 5 _ _ NNP 37602 160 6 : : : 37602 160 7 Top Top NNP 37602 160 8 . . . 37602 161 1 _ _ NNP 37602 161 2 O. O. NNP 37602 162 1 pearsoni pearsoni NNP 37602 162 2 _ _ NNP 37602 162 3 , , , 37602 162 4 KU KU NNP 37602 162 5 136312 136312 CD 37602 162 6 , , , 37602 162 7 [ [ -LRB- 37602 162 8 M m NN 37602 162 9 ] ] -RRB- 37602 162 10 ; ; : 37602 162 11 Middle Middle NNP 37602 162 12 . . . 37602 163 1 _ _ NNP 37602 163 2 O. O. NNP 37602 164 1 buckleyi buckleyi NNP 37602 164 2 _ _ NNP 37602 164 3 , , , 37602 164 4 KU KU NNP 37602 164 5 123172 123172 CD 37602 164 6 , , , 37602 164 7 [ [ -LRB- 37602 164 8 M m NN 37602 164 9 ] ] -RRB- 37602 164 10 ; ; : 37602 164 11 Bottom bottom NN 37602 164 12 . . . 37602 165 1 _ _ NNP 37602 165 2 O. O. NNP 37602 166 1 verrucigerus verrucigerus NNP 37602 166 2 _ _ NNP 37602 166 3 , , , 37602 166 4 KU KU NNP 37602 166 5 123177 123177 CD 37602 166 6 , , , 37602 166 7 [ [ -LRB- 37602 166 8 M m NN 37602 166 9 ] ] -RRB- 37602 166 10 . . . 37602 167 1 ×1.5 ×1.5 UH 37602 167 2 . . . 37602 167 3 ] ] -RRB- 37602 168 1 [ [ -LRB- 37602 168 2 Illustration illustration NN 37602 168 3 : : : 37602 168 4 FIG FIG NNP 37602 168 5 . . . 37602 169 1 2 2 LS 37602 169 2 . . . 37602 170 1 Species Species NNP 37602 170 2 of of IN 37602 170 3 _ _ NNP 37602 170 4 Osteocephalus Osteocephalus NNP 37602 170 5 _ _ NNP 37602 170 6 : : : 37602 170 7 Top Top NNP 37602 170 8 . . . 37602 171 1 _ _ NNP 37602 171 2 O. O. NNP 37602 172 1 leprieurii leprieurii NNP 37602 172 2 _ _ NNP 37602 172 3 , , , 37602 172 4 KU KU NNP 37602 172 5 126611 126611 CD 37602 172 6 , , , 37602 172 7 [ [ -LRB- 37602 172 8 F f NN 37602 172 9 ] ] -RRB- 37602 172 10 ; ; : 37602 172 11 Bottom bottom NN 37602 172 12 . . . 37602 173 1 _ _ NNP 37602 173 2 O. O. NNP 37602 174 1 taurinus taurinus NNP 37602 174 2 _ _ NNP 37602 174 3 , , , 37602 174 4 KU KU NNP 37602 174 5 126648 126648 CD 37602 174 6 , , , 37602 174 7 [ [ -LRB- 37602 174 8 M m NN 37602 174 9 ] ] -RRB- 37602 174 10 . . . 37602 175 1 ×1 ×1 RB 37602 175 2 . . . 37602 175 3 ] ] -RRB- 37602 176 1 [ [ -LRB- 37602 176 2 Illustration illustration NN 37602 176 3 : : : 37602 176 4 FIG FIG NNP 37602 176 5 . . . 37602 177 1 3 3 LS 37602 177 2 . . . 37602 178 1 Diagrammatic diagrammatic JJ 37602 178 2 views view NNS 37602 178 3 of of IN 37602 178 4 ventral ventral JJ 37602 178 5 color color NN 37602 178 6 patterns pattern NNS 37602 178 7 in in IN 37602 178 8 _ _ NNP 37602 178 9 Osteocephalus Osteocephalus NNP 37602 178 10 _ _ NNP 37602 178 11 : : : 37602 178 12 a. a. NN 37602 179 1 _ _ NNP 37602 179 2 O. O. NNP 37602 180 1 pearsoni pearsoni NNP 37602 180 2 _ _ NNP 37602 180 3 , , , 37602 180 4 UMMZ UMMZ NNP 37602 180 5 57533 57533 CD 37602 180 6 , , , 37602 180 7 [ [ -LRB- 37602 180 8 M m NN 37602 180 9 ] ] -RRB- 37602 180 10 ; ; : 37602 180 11 b. b. NNP 37602 181 1 _ _ NNP 37602 181 2 O. O. NNP 37602 182 1 taurinus taurinus NNP 37602 182 2 _ _ NNP 37602 182 3 , , , 37602 182 4 USNM USNM NNP 37602 182 5 166037 166037 CD 37602 182 6 , , , 37602 182 7 [ [ -LRB- 37602 182 8 M M NNP 37602 182 9 ] ] -RRB- 37602 182 10 ; ; : 37602 182 11 c. c. NNP 37602 182 12 _ _ NNP 37602 182 13 O. O. NNP 37602 183 1 verrucigerus verrucigerus NNP 37602 183 2 _ _ NNP 37602 183 3 , , , 37602 183 4 KU KU NNP 37602 183 5 123185 123185 CD 37602 183 6 , , , 37602 183 7 [ [ -LRB- 37602 183 8 F f NN 37602 183 9 ] ] -RRB- 37602 183 10 . . . 37602 183 11 ] ] -RRB- 37602 184 1 _ _ NNP 37602 184 2 Skin._--The skin._--the JJ 37602 184 3 dorsal dorsal NN 37602 184 4 skin skin NN 37602 184 5 of of IN 37602 184 6 all all DT 37602 184 7 males male NNS 37602 184 8 of of IN 37602 184 9 _ _ NNP 37602 184 10 Osteocephalus Osteocephalus NNP 37602 184 11 _ _ NNP 37602 184 12 is be VBZ 37602 184 13 tuberculate tuberculate NN 37602 184 14 to to IN 37602 184 15 varying vary VBG 37602 184 16 degrees degree NNS 37602 184 17 , , , 37602 184 18 whereas whereas IN 37602 184 19 the the DT 37602 184 20 dorsal dorsal NN 37602 184 21 skin skin NN 37602 184 22 of of IN 37602 184 23 females female NNS 37602 184 24 is be VBZ 37602 184 25 smooth smooth JJ 37602 184 26 , , , 37602 184 27 or or CC 37602 184 28 nearly nearly RB 37602 184 29 so so RB 37602 184 30 ( ( -LRB- 37602 184 31 Fig fig NN 37602 184 32 . . . 37602 185 1 4 4 LS 37602 185 2 ) ) -RRB- 37602 185 3 . . . 37602 186 1 _ _ NNP 37602 186 2 Osteocephalus Osteocephalus NNP 37602 186 3 verrucigerus verrucigerus NN 37602 186 4 _ _ NNP 37602 186 5 differs differ VBZ 37602 186 6 from from IN 37602 186 7 other other JJ 37602 186 8 members member NNS 37602 186 9 of of IN 37602 186 10 the the DT 37602 186 11 genus genus NN 37602 186 12 by by IN 37602 186 13 the the DT 37602 186 14 presence presence NN 37602 186 15 of of IN 37602 186 16 numerous numerous JJ 37602 186 17 , , , 37602 186 18 large large JJ 37602 186 19 tubercles tubercle NNS 37602 186 20 bearing bear VBG 37602 186 21 keratinized keratinize VBN 37602 186 22 tips tip NNS 37602 186 23 . . . 37602 187 1 The the DT 37602 187 2 tubercles tubercle NNS 37602 187 3 of of IN 37602 187 4 _ _ NNP 37602 187 5 leprieurii leprieurii JJ 37602 187 6 _ _ NNP 37602 187 7 are be VBP 37602 187 8 numerous numerous JJ 37602 187 9 and and CC 37602 187 10 spinous spinous JJ 37602 187 11 but but CC 37602 187 12 much much RB 37602 187 13 smaller small JJR 37602 187 14 than than IN 37602 187 15 those those DT 37602 187 16 of of IN 37602 187 17 _ _ NNP 37602 187 18 verrucigerus verrucigerus NNP 37602 187 19 _ _ NNP 37602 187 20 ; ; : 37602 187 21 those those DT 37602 187 22 of of IN 37602 187 23 _ _ NNP 37602 187 24 taurinus taurinus NNP 37602 187 25 _ _ NNP 37602 187 26 are be VBP 37602 187 27 spinous spinous JJ 37602 187 28 but but CC 37602 187 29 less less RBR 37602 187 30 numerous numerous JJ 37602 187 31 than than IN 37602 187 32 in in IN 37602 187 33 _ _ NNP 37602 187 34 leprieurii leprieurii JJ 37602 187 35 _ _ NNP 37602 187 36 . . . 37602 188 1 _ _ NNP 37602 188 2 Osteocephalus osteocephalus JJ 37602 188 3 buckleyi buckleyi NN 37602 188 4 _ _ NNP 37602 188 5 has have VBZ 37602 188 6 a a DT 37602 188 7 mixture mixture NN 37602 188 8 of of IN 37602 188 9 large large JJ 37602 188 10 and and CC 37602 188 11 small small JJ 37602 188 12 , , , 37602 188 13 non non JJ 37602 188 14 - - JJ 37602 188 15 spinous spinous JJ 37602 188 16 tubercles tubercle NNS 37602 188 17 , , , 37602 188 18 and and CC 37602 188 19 _ _ NNP 37602 188 20 pearsoni pearsoni NNP 37602 188 21 _ _ NNP 37602 188 22 has have VBZ 37602 188 23 only only RB 37602 188 24 a a DT 37602 188 25 few few JJ 37602 188 26 , , , 37602 188 27 small small JJ 37602 188 28 , , , 37602 188 29 scattered scatter VBN 37602 188 30 , , , 37602 188 31 non non JJ 37602 188 32 - - JJ 37602 188 33 spinous spinous JJ 37602 188 34 tubercles tubercle NNS 37602 188 35 . . . 37602 189 1 Fleshy Fleshy NNP 37602 189 2 tubercles tubercle NNS 37602 189 3 occur occur VBP 37602 189 4 on on IN 37602 189 5 the the DT 37602 189 6 eyelids eyelid NNS 37602 189 7 and and CC 37602 189 8 supratympanic supratympanic NNP 37602 189 9 fold fold VBP 37602 189 10 in in IN 37602 189 11 females female NNS 37602 189 12 of of IN 37602 189 13 _ _ NNP 37602 189 14 buckleyi buckleyi NN 37602 189 15 _ _ NNP 37602 189 16 ; ; : 37602 189 17 a a DT 37602 189 18 few few JJ 37602 189 19 small small JJ 37602 189 20 tubercles tubercle NNS 37602 189 21 are be VBP 37602 189 22 present present JJ 37602 189 23 on on IN 37602 189 24 the the DT 37602 189 25 back back NN 37602 189 26 of of IN 37602 189 27 females female NNS 37602 189 28 of of IN 37602 189 29 _ _ NNP 37602 189 30 pearsoni pearsoni NNP 37602 189 31 _ _ NNP 37602 189 32 , , , 37602 189 33 whereas whereas IN 37602 189 34 the the DT 37602 189 35 dorsal dorsal NN 37602 189 36 skin skin NN 37602 189 37 in in IN 37602 189 38 females female NNS 37602 189 39 of of IN 37602 189 40 the the DT 37602 189 41 other other JJ 37602 189 42 species specie NNS 37602 189 43 is be VBZ 37602 189 44 smooth smooth JJ 37602 189 45 . . . 37602 190 1 The the DT 37602 190 2 skin skin NN 37602 190 3 on on IN 37602 190 4 the the DT 37602 190 5 flanks flank NNS 37602 190 6 of of IN 37602 190 7 both both DT 37602 190 8 sexes sex NNS 37602 190 9 of of IN 37602 190 10 _ _ NNP 37602 190 11 buckleyi buckleyi NN 37602 190 12 _ _ NNP 37602 190 13 is be VBZ 37602 190 14 weakly weakly IN 37602 190 15 areolate areolate NN 37602 190 16 ; ; : 37602 190 17 in in IN 37602 190 18 the the DT 37602 190 19 other other JJ 37602 190 20 species specie NNS 37602 190 21 the the DT 37602 190 22 flanks flank NNS 37602 190 23 are be VBP 37602 190 24 smooth smooth JJ 37602 190 25 . . . 37602 191 1 The the DT 37602 191 2 skin skin NN 37602 191 3 on on IN 37602 191 4 the the DT 37602 191 5 top top NN 37602 191 6 of of IN 37602 191 7 the the DT 37602 191 8 head head NN 37602 191 9 in in IN 37602 191 10 _ _ NNP 37602 191 11 taurinus taurinus NNP 37602 191 12 _ _ NNP 37602 191 13 is be VBZ 37602 191 14 rugose rugose NN 37602 191 15 as as IN 37602 191 16 a a DT 37602 191 17 consequence consequence NN 37602 191 18 of of IN 37602 191 19 co co NN 37602 191 20 - - NN 37602 191 21 ossification ossification NN 37602 191 22 . . . 37602 192 1 In in IN 37602 192 2 all all DT 37602 192 3 species specie NNS 37602 192 4 the the DT 37602 192 5 anal anal JJ 37602 192 6 opening opening NN 37602 192 7 is be VBZ 37602 192 8 directed direct VBN 37602 192 9 posteriorly posteriorly RB 37602 192 10 at at IN 37602 192 11 the the DT 37602 192 12 upper upper JJ 37602 192 13 level level NN 37602 192 14 of of IN 37602 192 15 the the DT 37602 192 16 thighs thigh NNS 37602 192 17 . . . 37602 193 1 _ _ NNP 37602 193 2 Hands Hands NNPS 37602 193 3 and and CC 37602 193 4 Feet._--The Feet._--The NNP 37602 193 5 feet foot NNS 37602 193 6 of of IN 37602 193 7 _ _ NNP 37602 193 8 Osteocephalus Osteocephalus NNP 37602 193 9 _ _ NNP 37602 193 10 are be VBP 37602 193 11 fully fully RB 37602 193 12 webbed web VBN 37602 193 13 or or CC 37602 193 14 nearly nearly RB 37602 193 15 so so RB 37602 193 16 . . . 37602 194 1 Webbing web VBG 37602 194 2 between between IN 37602 194 3 fingers finger NNS 37602 194 4 one one CD 37602 194 5 and and CC 37602 194 6 two two CD 37602 194 7 is be VBZ 37602 194 8 basal basal NN 37602 194 9 in in IN 37602 194 10 all all DT 37602 194 11 species specie NNS 37602 194 12 . . . 37602 195 1 Webbing web VBG 37602 195 2 between between IN 37602 195 3 fingers finger NNS 37602 195 4 two two CD 37602 195 5 , , , 37602 195 6 three three CD 37602 195 7 , , , 37602 195 8 and and CC 37602 195 9 four four CD 37602 195 10 is be VBZ 37602 195 11 most most RBS 37602 195 12 extensive extensive JJ 37602 195 13 in in IN 37602 195 14 _ _ NNP 37602 195 15 taurinus taurinus NN 37602 195 16 _ _ NNP 37602 195 17 , , , 37602 195 18 in in IN 37602 195 19 which which WDT 37602 195 20 the the DT 37602 195 21 three three CD 37602 195 22 fingers finger NNS 37602 195 23 are be VBP 37602 195 24 about about RB 37602 195 25 one one CD 37602 195 26 - - HYPH 37602 195 27 half half NN 37602 195 28 webbed web VBN 37602 195 29 ( ( -LRB- 37602 195 30 Fig fig NN 37602 195 31 . . . 37602 196 1 5 5 LS 37602 196 2 ) ) -RRB- 37602 196 3 . . . 37602 197 1 _ _ NNP 37602 197 2 Osteocephalus osteocephalus JJ 37602 197 3 buckleyi buckleyi NN 37602 197 4 _ _ NNP 37602 197 5 , , , 37602 197 6 _ _ NNP 37602 197 7 pearsoni pearsoni NNP 37602 197 8 _ _ NNP 37602 197 9 , , , 37602 197 10 and and CC 37602 197 11 _ _ NNP 37602 197 12 verrucigerus verrucigerus NN 37602 197 13 _ _ NNP 37602 197 14 have have VBP 37602 197 15 reduced reduce VBN 37602 197 16 webbing webbing NN 37602 197 17 between between IN 37602 197 18 fingers finger NNS 37602 197 19 two two CD 37602 197 20 and and CC 37602 197 21 three three CD 37602 197 22 , , , 37602 197 23 and and CC 37602 197 24 _ _ NNP 37602 197 25 leprieurii leprieurii JJ 37602 197 26 _ _ NNP 37602 197 27 has have VBZ 37602 197 28 reduced reduce VBN 37602 197 29 webbing webbing NN 37602 197 30 between between IN 37602 197 31 fingers finger NNS 37602 197 32 two two CD 37602 197 33 , , , 37602 197 34 three three CD 37602 197 35 , , , 37602 197 36 and and CC 37602 197 37 four four CD 37602 197 38 . . . 37602 198 1 All all DT 37602 198 2 members member NNS 37602 198 3 of of IN 37602 198 4 the the DT 37602 198 5 genus genus NNPS 37602 198 6 have have VBP 37602 198 7 well well RB 37602 198 8 - - HYPH 37602 198 9 developed develop VBN 37602 198 10 subconical subconical JJ 37602 198 11 subarticular subarticular JJ 37602 198 12 tubercles tubercle NNS 37602 198 13 on on IN 37602 198 14 the the DT 37602 198 15 fingers finger NNS 37602 198 16 and and CC 37602 198 17 toes toe NNS 37602 198 18 ; ; : 37602 198 19 there there EX 37602 198 20 is be VBZ 37602 198 21 a a DT 37602 198 22 tendency tendency NN 37602 198 23 for for IN 37602 198 24 the the DT 37602 198 25 distal distal JJ 37602 198 26 tubercle tubercle NN 37602 198 27 on on IN 37602 198 28 the the DT 37602 198 29 fourth fourth JJ 37602 198 30 finger finger NN 37602 198 31 to to TO 37602 198 32 be be VB 37602 198 33 weakly weakly RB 37602 198 34 bifid bifid VB 37602 198 35 . . . 37602 199 1 Palmar Palmar NNP 37602 199 2 and and CC 37602 199 3 plantar plantar NNP 37602 199 4 supernumerary supernumerary JJ 37602 199 5 tubercles tubercle NNS 37602 199 6 are be VBP 37602 199 7 well well RB 37602 199 8 developed developed JJ 37602 199 9 in in IN 37602 199 10 _ _ NNP 37602 199 11 taurinus taurinus NNP 37602 199 12 _ _ NNP 37602 199 13 , , , 37602 199 14 moderately moderately RB 37602 199 15 developed develop VBD 37602 199 16 in in IN 37602 199 17 _ _ NNP 37602 199 18 buckleyi buckleyi NN 37602 199 19 _ _ NNP 37602 199 20 , , , 37602 199 21 _ _ NNP 37602 199 22 leprieurii leprieurii JJ 37602 199 23 _ _ NNP 37602 199 24 , , , 37602 199 25 and and CC 37602 199 26 _ _ NNP 37602 199 27 pearsoni pearsoni NNP 37602 199 28 _ _ NNP 37602 199 29 , , , 37602 199 30 and and CC 37602 199 31 barely barely RB 37602 199 32 evident evident JJ 37602 199 33 in in IN 37602 199 34 _ _ NNP 37602 199 35 verrucigerus verrucigerus NNP 37602 199 36 _ _ NNP 37602 199 37 . . . 37602 200 1 All all DT 37602 200 2 of of IN 37602 200 3 the the DT 37602 200 4 species specie NNS 37602 200 5 have have VBP 37602 200 6 a a DT 37602 200 7 noticeable noticeable JJ 37602 200 8 fold fold VB 37602 200 9 on on IN 37602 200 10 the the DT 37602 200 11 wrist wrist NN 37602 200 12 and and CC 37602 200 13 enlarged enlarged JJ 37602 200 14 prepollices prepollice NNS 37602 200 15 , , , 37602 200 16 bearing bear VBG 37602 200 17 horny horny JJ 37602 200 18 nuptial nuptial JJ 37602 200 19 excrescences excrescence NNS 37602 200 20 in in IN 37602 200 21 breeding breeding NN 37602 200 22 males male NNS 37602 200 23 . . . 37602 201 1 The the DT 37602 201 2 prepollex prepollex NN 37602 201 3 is be VBZ 37602 201 4 least least RBS 37602 201 5 enlarged enlarged JJ 37602 201 6 in in IN 37602 201 7 _ _ NNP 37602 201 8 buckleyi buckleyi NN 37602 201 9 _ _ NNP 37602 201 10 . . . 37602 202 1 Outer outer JJ 37602 202 2 metatarsal metatarsal NN 37602 202 3 tubercles tubercle NNS 37602 202 4 are be VBP 37602 202 5 absent absent JJ 37602 202 6 . . . 37602 203 1 The the DT 37602 203 2 inner inner JJ 37602 203 3 metatarsal metatarsal NN 37602 203 4 tubercle tubercle NN 37602 203 5 is be VBZ 37602 203 6 moderately moderately RB 37602 203 7 well well RB 37602 203 8 developed developed JJ 37602 203 9 and and CC 37602 203 10 ovoid ovoid JJ 37602 203 11 in in IN 37602 203 12 _ _ NNP 37602 203 13 leprieurii leprieurii JJ 37602 203 14 _ _ NNP 37602 203 15 and and CC 37602 203 16 _ _ NNP 37602 203 17 pearsoni pearsoni NNP 37602 203 18 _ _ NNP 37602 203 19 ; ; : 37602 203 20 it -PRON- PRP 37602 203 21 is be VBZ 37602 203 22 elliptical elliptical JJ 37602 203 23 and and CC 37602 203 24 flat flat JJ 37602 203 25 in in IN 37602 203 26 the the DT 37602 203 27 other other JJ 37602 203 28 species specie NNS 37602 203 29 . . . 37602 204 1 Tarsal tarsal NN 37602 204 2 folds fold NNS 37602 204 3 are be VBP 37602 204 4 absent absent JJ 37602 204 5 in in IN 37602 204 6 all all DT 37602 204 7 species specie NNS 37602 204 8 except except IN 37602 204 9 _ _ NNP 37602 204 10 verrucigerus verrucigerus NNP 37602 204 11 _ _ NNP 37602 204 12 , , , 37602 204 13 in in IN 37602 204 14 which which WDT 37602 204 15 the the DT 37602 204 16 folds fold NNS 37602 204 17 are be VBP 37602 204 18 barely barely RB 37602 204 19 evident evident JJ 37602 204 20 . . . 37602 205 1 [ [ -LRB- 37602 205 2 Illustration illustration NN 37602 205 3 : : : 37602 205 4 FIG FIG NNP 37602 205 5 . . . 37602 206 1 4 4 LS 37602 206 2 . . . 37602 207 1 Segments segment NNS 37602 207 2 of of IN 37602 207 3 dorsal dorsal NN 37602 207 4 skin skin NN 37602 207 5 of of IN 37602 207 6 males male NNS 37602 207 7 of of IN 37602 207 8 _ _ NNP 37602 207 9 Osteocephalus Osteocephalus NNP 37602 207 10 _ _ NNP 37602 207 11 showing show VBG 37602 207 12 size size NN 37602 207 13 and and CC 37602 207 14 arrangement arrangement NN 37602 207 15 of of IN 37602 207 16 tubercles tubercle NNS 37602 207 17 : : : 37602 207 18 a. a. NN 37602 208 1 _ _ NNP 37602 208 2 O. O. NNP 37602 209 1 verrucigerus verrucigerus NNP 37602 209 2 _ _ NNP 37602 209 3 , , , 37602 209 4 KU KU NNP 37602 209 5 123183 123183 CD 37602 209 6 ; ; : 37602 209 7 b. b. NNP 37602 210 1 _ _ NNP 37602 210 2 O. O. NNP 37602 211 1 taurinus taurinus NNP 37602 211 2 _ _ NNP 37602 211 3 , , , 37602 211 4 USNM USNM NNP 37602 211 5 166033 166033 CD 37602 211 6 ; ; : 37602 211 7 c. c. NNP 37602 211 8 _ _ NNP 37602 211 9 O. O. NNP 37602 212 1 leprieurii leprieurii NNP 37602 212 2 _ _ NNP 37602 212 3 , , , 37602 212 4 KU KU NNP 37602 212 5 126616 126616 CD 37602 212 6 ; ; : 37602 212 7 d. d. NNP 37602 212 8 _ _ NNP 37602 212 9 O. O. NNP 37602 213 1 buckleyi buckleyi NNP 37602 213 2 _ _ NNP 37602 213 3 , , , 37602 213 4 USNM USNM NNP 37602 213 5 165999 165999 CD 37602 213 6 . . . 37602 214 1 Each each DT 37602 214 2 square square NN 37602 214 3 = = SYM 37602 214 4 1 1 CD 37602 214 5 sq sq NN 37602 214 6 . . . 37602 215 1 cm cm NNP 37602 215 2 . . . 37602 215 3 ] ] -RRB- 37602 216 1 _ _ NNP 37602 216 2 Cranium._--As Cranium._--As NNP 37602 216 3 a a DT 37602 216 4 genus genus NN 37602 216 5 , , , 37602 216 6 the the DT 37602 216 7 cranial cranial JJ 37602 216 8 structure structure NN 37602 216 9 is be VBZ 37602 216 10 remarkably remarkably RB 37602 216 11 uniform uniform JJ 37602 216 12 and and CC 37602 216 13 quite quite RB 37602 216 14 generalized generalize VBN 37602 216 15 when when WRB 37602 216 16 viewed view VBN 37602 216 17 in in IN 37602 216 18 the the DT 37602 216 19 context context NN 37602 216 20 of of IN 37602 216 21 the the DT 37602 216 22 family family NN 37602 216 23 Hylidae Hylidae NNP 37602 216 24 . . . 37602 217 1 The the DT 37602 217 2 skulls skull NNS 37602 217 3 are be VBP 37602 217 4 broad broad JJ 37602 217 5 and and CC 37602 217 6 relatively relatively RB 37602 217 7 flat flat JJ 37602 217 8 , , , 37602 217 9 each each DT 37602 217 10 being be VBG 37602 217 11 only only RB 37602 217 12 slightly slightly RB 37602 217 13 more more RBR 37602 217 14 broad broad JJ 37602 217 15 than than IN 37602 217 16 long long JJ 37602 217 17 and and CC 37602 217 18 about about RB 37602 217 19 one one CD 37602 217 20 - - HYPH 37602 217 21 third third NN 37602 217 22 as as RB 37602 217 23 high high JJ 37602 217 24 as as RB 37602 217 25 long long RB 37602 217 26 . . . 37602 218 1 In in IN 37602 218 2 dorsal dorsal NN 37602 218 3 aspect aspect NN 37602 218 4 the the DT 37602 218 5 snouts snout NNS 37602 218 6 are be VBP 37602 218 7 broadly broadly RB 37602 218 8 rounded round VBN 37602 218 9 ; ; : 37602 218 10 the the DT 37602 218 11 snout snout NN 37602 218 12 of of IN 37602 218 13 _ _ NNP 37602 218 14 buckleyi buckleyi NN 37602 218 15 _ _ NNP 37602 218 16 is be VBZ 37602 218 17 somewhat somewhat RB 37602 218 18 less less RBR 37602 218 19 rounded rounded JJ 37602 218 20 and and CC 37602 218 21 appears appear VBZ 37602 218 22 to to TO 37602 218 23 be be VB 37602 218 24 slightly slightly RB 37602 218 25 longer long JJR 37602 218 26 than than IN 37602 218 27 the the DT 37602 218 28 snouts snout NNS 37602 218 29 of of IN 37602 218 30 other other JJ 37602 218 31 species specie NNS 37602 218 32 . . . 37602 219 1 This this DT 37602 219 2 subtle subtle JJ 37602 219 3 difference difference NN 37602 219 4 relates relate VBZ 37602 219 5 to to IN 37602 219 6 the the DT 37602 219 7 relative relative JJ 37602 219 8 narrowness narrowness NN 37602 219 9 of of IN 37602 219 10 the the DT 37602 219 11 premaxillaries premaxillarie NNS 37602 219 12 in in IN 37602 219 13 _ _ NNP 37602 219 14 buckleyi buckleyi NN 37602 219 15 _ _ NNP 37602 219 16 . . . 37602 220 1 [ [ -LRB- 37602 220 2 Illustration illustration NN 37602 220 3 : : : 37602 220 4 FIG FIG NNP 37602 220 5 . . . 37602 221 1 5 5 CD 37602 221 2 . . . 37602 222 1 Palmar Palmar NNP 37602 222 2 views view NNS 37602 222 3 of of IN 37602 222 4 hands hand NNS 37602 222 5 of of IN 37602 222 6 males male NNS 37602 222 7 of of IN 37602 222 8 _ _ NNP 37602 222 9 Osteocephalus Osteocephalus NNP 37602 222 10 _ _ NNP 37602 222 11 : : : 37602 222 12 a. a. NN 37602 223 1 _ _ NNP 37602 223 2 O. O. NNP 37602 224 1 buckleyi buckleyi NNP 37602 224 2 _ _ NNP 37602 224 3 , , , 37602 224 4 KU KU NNP 37602 224 5 109506 109506 CD 37602 224 6 ; ; : 37602 224 7 b. b. NNP 37602 225 1 _ _ NNP 37602 225 2 O. O. NNP 37602 226 1 leprieurii leprieurii NNP 37602 226 2 _ _ NNP 37602 226 3 , , , 37602 226 4 KU KU NNP 37602 226 5 126627 126627 CD 37602 226 6 ; ; : 37602 226 7 c. c. NNP 37602 226 8 _ _ NNP 37602 226 9 O. O. NNP 37602 227 1 pearsoni pearsoni NNP 37602 227 2 _ _ NNP 37602 227 3 , , , 37602 227 4 MCZ MCZ NNP 37602 227 5 15565 15565 CD 37602 227 6 ; ; : 37602 227 7 d. d. NNP 37602 227 8 _ _ NNP 37602 227 9 O. O. NNP 37602 228 1 taurinus taurinus NNP 37602 228 2 _ _ NNP 37602 228 3 , , , 37602 228 4 KU KU NNP 37602 228 5 126653 126653 CD 37602 228 6 ; ; : 37602 228 7 e. e. NNP 37602 228 8 _ _ NNP 37602 228 9 O. O. NNP 37602 229 1 verrucigerus verrucigerus NNP 37602 229 2 _ _ NNP 37602 229 3 , , , 37602 229 4 KU KU NNP 37602 229 5 123177 123177 CD 37602 229 6 . . . 37602 230 1 ×4 ×4 NFP 37602 230 2 . . . 37602 230 3 ] ] -RRB- 37602 231 1 [ [ -LRB- 37602 231 2 Illustration illustration NN 37602 231 3 : : : 37602 231 4 FIG FIG NNP 37602 231 5 . . . 37602 232 1 6 6 CD 37602 232 2 . . . 37602 233 1 Skulls skull NNS 37602 233 2 of of IN 37602 233 3 two two CD 37602 233 4 species specie NNS 37602 233 5 of of IN 37602 233 6 _ _ NNP 37602 233 7 Osteocephalus Osteocephalus NNP 37602 233 8 _ _ NNP 37602 233 9 : : : 37602 233 10 a a NNP 37602 233 11 and and CC 37602 233 12 b. b. NN 37602 234 1 _ _ NNP 37602 234 2 O. O. NNP 37602 235 1 leprieurii leprieurii NNP 37602 235 2 _ _ NNP 37602 235 3 , , , 37602 235 4 KU KU NNP 37602 235 5 125961 125961 CD 37602 235 6 ; ; : 37602 235 7 c c NNP 37602 235 8 and and CC 37602 235 9 d. d. NNP 37602 235 10 _ _ NNP 37602 235 11 O. O. NNP 37602 236 1 pearsoni pearsoni NNP 37602 236 2 _ _ NNP 37602 236 3 , , , 37602 236 4 UMMZ UMMZ NNP 37602 236 5 67465 67465 CD 37602 236 6 . . . 37602 237 1 ×3 ×3 NNP 37602 237 2 . . . 37602 237 3 ] ] -RRB- 37602 238 1 The the DT 37602 238 2 genus genus NN 37602 238 3 is be VBZ 37602 238 4 characterized characterize VBN 37602 238 5 by by IN 37602 238 6 well well RB 37602 238 7 - - HYPH 37602 238 8 developed develop VBN 37602 238 9 dermal dermal JJ 37602 238 10 roofing roofing NN 37602 238 11 bones bone NNS 37602 238 12 and and CC 37602 238 13 an an DT 37602 238 14 unusually unusually RB 37602 238 15 large large JJ 37602 238 16 exposure exposure NN 37602 238 17 of of IN 37602 238 18 the the DT 37602 238 19 sphenethmoid sphenethmoid NN 37602 238 20 dorsally dorsally RB 37602 238 21 ( ( -LRB- 37602 238 22 Fig fig NN 37602 238 23 . . . 37602 239 1 6 6 LS 37602 239 2 ) ) -RRB- 37602 239 3 . . . 37602 240 1 The the DT 37602 240 2 conformation conformation NN 37602 240 3 of of IN 37602 240 4 the the DT 37602 240 5 sphenethmoid sphenethmoid NN 37602 240 6 exposed expose VBN 37602 240 7 dorsally dorsally RB 37602 240 8 is be VBZ 37602 240 9 determined determine VBN 37602 240 10 by by IN 37602 240 11 the the DT 37602 240 12 marginal marginal JJ 37602 240 13 positions position NNS 37602 240 14 of of IN 37602 240 15 the the DT 37602 240 16 adjacent adjacent JJ 37602 240 17 , , , 37602 240 18 overlapping overlap VBG 37602 240 19 elements element NNS 37602 240 20 -- -- : 37602 240 21 the the DT 37602 240 22 nasals nasal NNS 37602 240 23 and and CC 37602 240 24 frontoparietals frontoparietal NNS 37602 240 25 . . . 37602 241 1 Medially medially RB 37602 241 2 the the DT 37602 241 3 nasals nasal NNS 37602 241 4 overlap overlap VBP 37602 241 5 the the DT 37602 241 6 lateral lateral JJ 37602 241 7 margins margin NNS 37602 241 8 of of IN 37602 241 9 the the DT 37602 241 10 sphenethmoid sphenethmoid NN 37602 241 11 . . . 37602 242 1 Anteromedially anteromedially RB 37602 242 2 , , , 37602 242 3 the the DT 37602 242 4 nasals nasal NNS 37602 242 5 converge converge VBP 37602 242 6 in in IN 37602 242 7 _ _ NNP 37602 242 8 leprieurii leprieurii JJ 37602 242 9 _ _ NNP 37602 242 10 and and CC 37602 242 11 _ _ NNP 37602 242 12 taurinus taurinus NN 37602 242 13 _ _ NNP 37602 242 14 , , , 37602 242 15 are be VBP 37602 242 16 narrowly narrowly RB 37602 242 17 separated separate VBN 37602 242 18 in in IN 37602 242 19 _ _ NNP 37602 242 20 buckleyi buckleyi NN 37602 242 21 _ _ NNP 37602 242 22 and and CC 37602 242 23 _ _ NNP 37602 242 24 pearsoni pearsoni NNP 37602 242 25 _ _ NNP 37602 242 26 , , , 37602 242 27 or or CC 37602 242 28 are be VBP 37602 242 29 more more RBR 37602 242 30 widely widely RB 37602 242 31 separated separate VBN 37602 242 32 in in IN 37602 242 33 _ _ NNP 37602 242 34 verrucigerus verrucigerus NNP 37602 242 35 _ _ NNP 37602 242 36 . . . 37602 243 1 In in IN 37602 243 2 all all DT 37602 243 3 species specie NNS 37602 243 4 the the DT 37602 243 5 nasals nasal NNS 37602 243 6 terminate terminate VBP 37602 243 7 at at IN 37602 243 8 the the DT 37602 243 9 anterodorsal anterodorsal NNP 37602 243 10 corner corner NN 37602 243 11 of of IN 37602 243 12 the the DT 37602 243 13 orbit orbit NN 37602 243 14 . . . 37602 244 1 The the DT 37602 244 2 frontoparietals frontoparietal NNS 37602 244 3 of of IN 37602 244 4 _ _ NNP 37602 244 5 buckleyi buckleyi NN 37602 244 6 _ _ NNP 37602 244 7 , , , 37602 244 8 _ _ NNP 37602 244 9 leprieurii leprieurii JJ 37602 244 10 _ _ NNP 37602 244 11 , , , 37602 244 12 and and CC 37602 244 13 _ _ NNP 37602 244 14 taurinus taurinus NNP 37602 244 15 _ _ NNP 37602 244 16 have have VBP 37602 244 17 an an DT 37602 244 18 anterolateral anterolateral JJ 37602 244 19 extension extension NN 37602 244 20 , , , 37602 244 21 which which WDT 37602 244 22 marginally marginally RB 37602 244 23 overlaps overlap VBZ 37602 244 24 the the DT 37602 244 25 dorsolateral dorsolateral JJ 37602 244 26 edge edge NN 37602 244 27 of of IN 37602 244 28 the the DT 37602 244 29 sphenethmoid sphenethmoid NN 37602 244 30 and and CC 37602 244 31 articulates articulate VBZ 37602 244 32 with with IN 37602 244 33 the the DT 37602 244 34 posterodorsal posterodorsal NN 37602 244 35 corner corner NN 37602 244 36 of of IN 37602 244 37 the the DT 37602 244 38 nasal nasal NN 37602 244 39 in in IN 37602 244 40 _ _ NNP 37602 244 41 buckleyi buckleyi NN 37602 244 42 _ _ NNP 37602 244 43 and and CC 37602 244 44 _ _ NNP 37602 244 45 taurinus taurinus NNP 37602 244 46 _ _ NNP 37602 244 47 ; ; : 37602 244 48 the the DT 37602 244 49 bones bone NNS 37602 244 50 are be VBP 37602 244 51 narrowly narrowly RB 37602 244 52 separated separate VBN 37602 244 53 in in IN 37602 244 54 _ _ NNP 37602 244 55 leprieurii leprieurii JJ 37602 244 56 _ _ NNP 37602 244 57 . . . 37602 245 1 The the DT 37602 245 2 frontoparietals frontoparietal NNS 37602 245 3 of of IN 37602 245 4 _ _ NNP 37602 245 5 pearsoni pearsoni NNP 37602 245 6 _ _ NNP 37602 245 7 and and CC 37602 245 8 _ _ NNP 37602 245 9 verrucigerus verrucigerus NN 37602 245 10 _ _ NNP 37602 245 11 have have VBP 37602 245 12 more more RBR 37602 245 13 extensive extensive JJ 37602 245 14 median median JJ 37602 245 15 ossification ossification NN 37602 245 16 and and CC 37602 245 17 less less RBR 37602 245 18 extensive extensive JJ 37602 245 19 anterolateral anterolateral JJ 37602 245 20 ossification ossification NN 37602 245 21 . . . 37602 246 1 Thus thus RB 37602 246 2 , , , 37602 246 3 in in IN 37602 246 4 those those DT 37602 246 5 species specie NNS 37602 246 6 the the DT 37602 246 7 posteromedian posteromedian JJ 37602 246 8 portion portion NN 37602 246 9 of of IN 37602 246 10 the the DT 37602 246 11 sphenethmoid sphenethmoid NN 37602 246 12 is be VBZ 37602 246 13 obscured obscure VBN 37602 246 14 , , , 37602 246 15 and and CC 37602 246 16 the the DT 37602 246 17 lateral lateral JJ 37602 246 18 margins margin NNS 37602 246 19 are be VBP 37602 246 20 partly partly RB 37602 246 21 exposed expose VBN 37602 246 22 . . . 37602 247 1 The the DT 37602 247 2 frontoparietal frontoparietal JJ 37602 247 3 fontanelle fontanelle NN 37602 247 4 is be VBZ 37602 247 5 completely completely RB 37602 247 6 covered cover VBN 37602 247 7 in in IN 37602 247 8 all all DT 37602 247 9 species specie NNS 37602 247 10 , , , 37602 247 11 except except IN 37602 247 12 _ _ NNP 37602 247 13 buckleyi buckleyi NN 37602 247 14 _ _ NNP 37602 247 15 and and CC 37602 247 16 _ _ NNP 37602 247 17 leprieurii leprieurii JJ 37602 247 18 _ _ NNP 37602 247 19 , , , 37602 247 20 in in IN 37602 247 21 which which WDT 37602 247 22 an an DT 37602 247 23 irregular irregular JJ 37602 247 24 , , , 37602 247 25 median median JJ 37602 247 26 separation separation NN 37602 247 27 of of IN 37602 247 28 the the DT 37602 247 29 frontoparietals frontoparietal NNS 37602 247 30 exposes expose VBZ 37602 247 31 a a DT 37602 247 32 small small JJ 37602 247 33 portion portion NN 37602 247 34 of of IN 37602 247 35 the the DT 37602 247 36 fontanelle fontanelle NN 37602 247 37 . . . 37602 248 1 The the DT 37602 248 2 posterolateral posterolateral JJ 37602 248 3 margins margin NNS 37602 248 4 of of IN 37602 248 5 the the DT 37602 248 6 frontoparietals frontoparietal NNS 37602 248 7 lie lie VBP 37602 248 8 medial medial JJ 37602 248 9 to to IN 37602 248 10 the the DT 37602 248 11 epiotic epiotic JJ 37602 248 12 eminences eminence NNS 37602 248 13 . . . 37602 249 1 Dermal dermal JJ 37602 249 2 ornamentation ornamentation NN 37602 249 3 , , , 37602 249 4 involving involve VBG 37602 249 5 the the DT 37602 249 6 nasals nasal NNS 37602 249 7 , , , 37602 249 8 frontoparietals frontoparietal NNS 37602 249 9 , , , 37602 249 10 and and CC 37602 249 11 sphenethmoid sphenethmoid NN 37602 249 12 , , , 37602 249 13 occurs occur VBZ 37602 249 14 in in IN 37602 249 15 _ _ NNP 37602 249 16 taurinus taurinus NNP 37602 249 17 _ _ NNP 37602 249 18 and and CC 37602 249 19 , , , 37602 249 20 to to IN 37602 249 21 a a DT 37602 249 22 limited limited JJ 37602 249 23 extent extent NN 37602 249 24 , , , 37602 249 25 in in IN 37602 249 26 _ _ NNP 37602 249 27 pearsoni pearsoni NNP 37602 249 28 _ _ NNP 37602 249 29 . . . 37602 250 1 In in IN 37602 250 2 the the DT 37602 250 3 latter latter JJ 37602 250 4 species species NN 37602 250 5 marginal marginal JJ 37602 250 6 portions portion NNS 37602 250 7 of of IN 37602 250 8 the the DT 37602 250 9 frontoparietals frontoparietal NNS 37602 250 10 , , , 37602 250 11 the the DT 37602 250 12 dorsal dorsal NN 37602 250 13 surfaces surface NNS 37602 250 14 of of IN 37602 250 15 the the DT 37602 250 16 nasals nasal NNS 37602 250 17 , , , 37602 250 18 and and CC 37602 250 19 the the DT 37602 250 20 posteromedial posteromedial JJ 37602 250 21 part part NN 37602 250 22 of of IN 37602 250 23 the the DT 37602 250 24 exposed expose VBN 37602 250 25 sphenethmoid sphenethmoid NN 37602 250 26 are be VBP 37602 250 27 slightly slightly RB 37602 250 28 exostosed exostose VBN 37602 250 29 , , , 37602 250 30 resulting result VBG 37602 250 31 in in IN 37602 250 32 an an DT 37602 250 33 open open JJ 37602 250 34 , , , 37602 250 35 reticulate reticulate JJ 37602 250 36 pattern pattern NN 37602 250 37 of of IN 37602 250 38 dermal dermal JJ 37602 250 39 sculpturing sculpturing NN 37602 250 40 of of IN 37602 250 41 exceedingly exceedingly RB 37602 250 42 low low JJ 37602 250 43 relief relief NN 37602 250 44 ( ( -LRB- 37602 250 45 Fig fig NN 37602 250 46 . . . 37602 251 1 6c 6c NNS 37602 251 2 ) ) -RRB- 37602 251 3 . . . 37602 252 1 _ _ NNP 37602 252 2 Osteocephalus osteocephalus JJ 37602 252 3 taurinus taurinus NN 37602 252 4 _ _ NNP 37602 252 5 is be VBZ 37602 252 6 characterized characterize VBN 37602 252 7 by by IN 37602 252 8 casquing casque VBG 37602 252 9 , , , 37602 252 10 co co NN 37602 252 11 - - NN 37602 252 12 ossification ossification NN 37602 252 13 , , , 37602 252 14 and and CC 37602 252 15 a a DT 37602 252 16 rather rather RB 37602 252 17 intricate intricate JJ 37602 252 18 pattern pattern NN 37602 252 19 of of IN 37602 252 20 dermal dermal JJ 37602 252 21 sculpturing sculpturing NN 37602 252 22 , , , 37602 252 23 which which WDT 37602 252 24 was be VBD 37602 252 25 described describe VBN 37602 252 26 in in IN 37602 252 27 detail detail NN 37602 252 28 and and CC 37602 252 29 illustrated illustrate VBN 37602 252 30 by by IN 37602 252 31 Trueb Trueb NNP 37602 252 32 ( ( -LRB- 37602 252 33 1970a 1970a CD 37602 252 34 ) ) -RRB- 37602 252 35 . . . 37602 253 1 The the DT 37602 253 2 squamosals squamosal NNS 37602 253 3 of of IN 37602 253 4 all all DT 37602 253 5 species specie NNS 37602 253 6 are be VBP 37602 253 7 moderately moderately RB 37602 253 8 large large JJ 37602 253 9 structures structure NNS 37602 253 10 having have VBG 37602 253 11 otic otic JJ 37602 253 12 plates plate NNS 37602 253 13 that that WDT 37602 253 14 overlie overlie VBP 37602 253 15 the the DT 37602 253 16 lateral lateral JJ 37602 253 17 portion portion NN 37602 253 18 of of IN 37602 253 19 the the DT 37602 253 20 cristae cristae NN 37602 253 21 paroticae paroticae NN 37602 253 22 . . . 37602 254 1 The the DT 37602 254 2 posterior posterior NN 37602 254 3 rami rami NN 37602 254 4 are be VBP 37602 254 5 short short JJ 37602 254 6 ; ; : 37602 254 7 the the DT 37602 254 8 zygomatic zygomatic JJ 37602 254 9 rami rami NN 37602 254 10 of of IN 37602 254 11 all all DT 37602 254 12 species specie NNS 37602 254 13 , , , 37602 254 14 except except IN 37602 254 15 _ _ NNP 37602 254 16 taurinus taurinus NN 37602 254 17 _ _ NNP 37602 254 18 , , , 37602 254 19 extend extend VB 37602 254 20 slightly slightly RB 37602 254 21 more more JJR 37602 254 22 than than IN 37602 254 23 one one CD 37602 254 24 - - HYPH 37602 254 25 half half NN 37602 254 26 of of IN 37602 254 27 the the DT 37602 254 28 distance distance NN 37602 254 29 to to IN 37602 254 30 the the DT 37602 254 31 maxillary maxillary NN 37602 254 32 . . . 37602 255 1 In in IN 37602 255 2 _ _ NNP 37602 255 3 taurinus taurinus NN 37602 255 4 _ _ IN 37602 255 5 the the DT 37602 255 6 zygomatic zygomatic JJ 37602 255 7 ramus ramus NN 37602 255 8 extends extend VBZ 37602 255 9 nearly nearly RB 37602 255 10 to to TO 37602 255 11 , , , 37602 255 12 or or CC 37602 255 13 articulates articulate VBZ 37602 255 14 with with IN 37602 255 15 , , , 37602 255 16 the the DT 37602 255 17 maxillary maxillary NN 37602 255 18 . . . 37602 256 1 The the DT 37602 256 2 maxillary maxillary JJ 37602 256 3 arches arch NNS 37602 256 4 are be VBP 37602 256 5 complete complete JJ 37602 256 6 and and CC 37602 256 7 relatively relatively RB 37602 256 8 robust robust JJ 37602 256 9 . . . 37602 257 1 The the DT 37602 257 2 alary alary JJ 37602 257 3 processes process NNS 37602 257 4 of of IN 37602 257 5 the the DT 37602 257 6 premaxillaries premaxillarie NNS 37602 257 7 are be VBP 37602 257 8 vertically vertically RB 37602 257 9 oriented orient VBN 37602 257 10 in in IN 37602 257 11 _ _ NNP 37602 257 12 leprieurii leprieurii JJ 37602 257 13 _ _ NNP 37602 257 14 , , , 37602 257 15 _ _ NNP 37602 257 16 pearsoni pearsoni NNP 37602 257 17 _ _ NNP 37602 257 18 , , , 37602 257 19 and and CC 37602 257 20 _ _ NNP 37602 257 21 taurinus taurinus NN 37602 257 22 _ _ NNP 37602 257 23 and and CC 37602 257 24 very very RB 37602 257 25 slightly slightly RB 37602 257 26 inclined incline VBN 37602 257 27 posteriorly posteriorly RB 37602 257 28 in in IN 37602 257 29 _ _ NNP 37602 257 30 buckleyi buckleyi NN 37602 257 31 _ _ NNP 37602 257 32 and and CC 37602 257 33 _ _ NNP 37602 257 34 verrucigerus verrucigerus NNP 37602 257 35 _ _ NNP 37602 257 36 . . . 37602 258 1 The the DT 37602 258 2 maxillaries maxillary NNS 37602 258 3 are be VBP 37602 258 4 uniformly uniformly RB 37602 258 5 characterized characterize VBN 37602 258 6 by by IN 37602 258 7 the the DT 37602 258 8 absence absence NN 37602 258 9 of of IN 37602 258 10 postorbital postorbital JJ 37602 258 11 processes process NNS 37602 258 12 and and CC 37602 258 13 by by IN 37602 258 14 the the DT 37602 258 15 presence presence NN 37602 258 16 of of IN 37602 258 17 preorbital preorbital JJ 37602 258 18 processes process NNS 37602 258 19 that that WDT 37602 258 20 articulate articulate VBP 37602 258 21 with with IN 37602 258 22 the the DT 37602 258 23 maxillary maxillary JJ 37602 258 24 processes process NNS 37602 258 25 of of IN 37602 258 26 the the DT 37602 258 27 nasals nasal NNS 37602 258 28 . . . 37602 259 1 The the DT 37602 259 2 partes parte NNS 37602 259 3 facialae facialae RB 37602 259 4 of of IN 37602 259 5 the the DT 37602 259 6 maxillaries maxillary NNS 37602 259 7 are be VBP 37602 259 8 moderately moderately RB 37602 259 9 developed develop VBN 37602 259 10 in in IN 37602 259 11 all all DT 37602 259 12 species specie NNS 37602 259 13 , , , 37602 259 14 except except IN 37602 259 15 _ _ NNP 37602 259 16 taurinus taurinus NN 37602 259 17 _ _ NNP 37602 259 18 , , , 37602 259 19 in in IN 37602 259 20 which which WDT 37602 259 21 the the DT 37602 259 22 pars par NNS 37602 259 23 fascialis fascialis NNP 37602 259 24 tends tend VBZ 37602 259 25 to to TO 37602 259 26 articulate articulate VB 37602 259 27 with with IN 37602 259 28 the the DT 37602 259 29 lateral lateral JJ 37602 259 30 margin margin NN 37602 259 31 of of IN 37602 259 32 the the DT 37602 259 33 nasal nasal NN 37602 259 34 in in IN 37602 259 35 well well RB 37602 259 36 - - HYPH 37602 259 37 ossified ossify VBN 37602 259 38 individuals individual NNS 37602 259 39 . . . 37602 260 1 The the DT 37602 260 2 partes parte NNS 37602 260 3 palatinae palatinae RB 37602 260 4 are be VBP 37602 260 5 poorly poorly RB 37602 260 6 developed develop VBN 37602 260 7 in in IN 37602 260 8 all all DT 37602 260 9 species specie NNS 37602 260 10 , , , 37602 260 11 except except IN 37602 260 12 _ _ NNP 37602 260 13 buckleyi buckleyi NN 37602 260 14 _ _ NNP 37602 260 15 , , , 37602 260 16 in in IN 37602 260 17 which which WDT 37602 260 18 the the DT 37602 260 19 pars par NNS 37602 260 20 palatina palatina NN 37602 260 21 of of IN 37602 260 22 the the DT 37602 260 23 premaxillary premaxillary NN 37602 260 24 is be VBZ 37602 260 25 moderately moderately RB 37602 260 26 robust robust JJ 37602 260 27 . . . 37602 261 1 The the DT 37602 261 2 pterygoids pterygoid NNS 37602 261 3 are be VBP 37602 261 4 uniformly uniformly RB 37602 261 5 tri tri JJ 37602 261 6 - - HYPH 37602 261 7 radiate radiate JJ 37602 261 8 structures structure NNS 37602 261 9 . . . 37602 262 1 The the DT 37602 262 2 anterior anterior NNP 37602 262 3 rami rami NN 37602 262 4 terminate terminate NN 37602 262 5 at at IN 37602 262 6 about about IN 37602 262 7 the the DT 37602 262 8 mid mid JJ 37602 262 9 - - NN 37602 262 10 level level NN 37602 262 11 of of IN 37602 262 12 the the DT 37602 262 13 orbit orbit NN 37602 262 14 , , , 37602 262 15 and and CC 37602 262 16 the the DT 37602 262 17 medial medial NNP 37602 262 18 rami rami NNP 37602 262 19 articulate articulate NN 37602 262 20 firmly firmly RB 37602 262 21 with with IN 37602 262 22 the the DT 37602 262 23 anterolateral anterolateral JJ 37602 262 24 corner corner NN 37602 262 25 of of IN 37602 262 26 the the DT 37602 262 27 otic otic JJ 37602 262 28 capsule capsule NN 37602 262 29 . . . 37602 263 1 The the DT 37602 263 2 palatines palatine NNS 37602 263 3 are be VBP 37602 263 4 well well RB 37602 263 5 - - HYPH 37602 263 6 developed develop VBN 37602 263 7 elements element NNS 37602 263 8 bearing bear VBG 37602 263 9 ventral ventral JJ 37602 263 10 ridges ridge NNS 37602 263 11 ; ; : 37602 263 12 the the DT 37602 263 13 ridges ridge NNS 37602 263 14 are be VBP 37602 263 15 somewhat somewhat RB 37602 263 16 irregular irregular JJ 37602 263 17 in in IN 37602 263 18 _ _ NNP 37602 263 19 buckleyi buckleyi NN 37602 263 20 _ _ NNP 37602 263 21 , , , 37602 263 22 _ _ NNP 37602 263 23 taurinus taurinus NN 37602 263 24 _ _ NNP 37602 263 25 , , , 37602 263 26 and and CC 37602 263 27 _ _ NNP 37602 263 28 verrucigerus verrucigerus NNP 37602 263 29 _ _ NNP 37602 263 30 but but CC 37602 263 31 smooth smooth VB 37602 263 32 in in IN 37602 263 33 _ _ NNP 37602 263 34 leprieurii leprieurii JJ 37602 263 35 _ _ NNP 37602 263 36 and and CC 37602 263 37 _ _ NNP 37602 263 38 pearsoni pearsoni NNP 37602 263 39 _ _ NNP 37602 263 40 . . . 37602 264 1 The the DT 37602 264 2 parasphenoids parasphenoid NNS 37602 264 3 are be VBP 37602 264 4 large large JJ 37602 264 5 elements element NNS 37602 264 6 characterized characterize VBN 37602 264 7 by by IN 37602 264 8 acuminate acuminate NN 37602 264 9 cultriform cultriform NN 37602 264 10 processes process NNS 37602 264 11 and and CC 37602 264 12 posterolaterally posterolaterally RB 37602 264 13 inclined inclined JJ 37602 264 14 alary alary JJ 37602 264 15 processes process NNS 37602 264 16 . . . 37602 265 1 The the DT 37602 265 2 basal basal NN 37602 265 3 areas area NNS 37602 265 4 of of IN 37602 265 5 the the DT 37602 265 6 cultriform cultriform NN 37602 265 7 processes process NNS 37602 265 8 bear bear VBP 37602 265 9 small small JJ 37602 265 10 odontoid odontoid JJ 37602 265 11 protuberances protuberance NNS 37602 265 12 in in IN 37602 265 13 _ _ NNP 37602 265 14 buckleyi buckleyi NN 37602 265 15 _ _ NNP 37602 265 16 , , , 37602 265 17 _ _ NNP 37602 265 18 taurinus taurinus NN 37602 265 19 _ _ NNP 37602 265 20 , , , 37602 265 21 and and CC 37602 265 22 _ _ NNP 37602 265 23 verrucigerus verrucigerus NNP 37602 265 24 _ _ NNP 37602 265 25 , , , 37602 265 26 whereas whereas IN 37602 265 27 they -PRON- PRP 37602 265 28 are be VBP 37602 265 29 smooth smooth JJ 37602 265 30 in in IN 37602 265 31 _ _ NNP 37602 265 32 leprieurii leprieurii JJ 37602 265 33 _ _ NNP 37602 265 34 and and CC 37602 265 35 _ _ NNP 37602 265 36 pearsoni pearsoni NNP 37602 265 37 _ _ NNP 37602 265 38 . . . 37602 266 1 The the DT 37602 266 2 prevomers prevomer NNS 37602 266 3 are be VBP 37602 266 4 remarkably remarkably RB 37602 266 5 uniform uniform JJ 37602 266 6 , , , 37602 266 7 moderately moderately RB 37602 266 8 well well RB 37602 266 9 - - HYPH 37602 266 10 developed develop VBN 37602 266 11 structures structure NNS 37602 266 12 . . . 37602 267 1 In in IN 37602 267 2 each each DT 37602 267 3 species specie NNS 37602 267 4 the the DT 37602 267 5 anterior anterior NNP 37602 267 6 ramus ramus NNP 37602 267 7 lies lie VBZ 37602 267 8 adjacent adjacent JJ 37602 267 9 to to IN 37602 267 10 the the DT 37602 267 11 premaxillary premaxillary NN 37602 267 12 , , , 37602 267 13 and and CC 37602 267 14 the the DT 37602 267 15 lateral lateral JJ 37602 267 16 wings wing NNS 37602 267 17 form form VBP 37602 267 18 the the DT 37602 267 19 anterior anterior NNP 37602 267 20 , , , 37602 267 21 medial medial NNP 37602 267 22 , , , 37602 267 23 and and CC 37602 267 24 posteromedial posteromedial JJ 37602 267 25 margins margin NNS 37602 267 26 of of IN 37602 267 27 the the DT 37602 267 28 internal internal JJ 37602 267 29 nares naris NNS 37602 267 30 . . . 37602 268 1 The the DT 37602 268 2 dentigerous dentigerous JJ 37602 268 3 processes process NNS 37602 268 4 are be VBP 37602 268 5 characteristically characteristically RB 37602 268 6 large large JJ 37602 268 7 and and CC 37602 268 8 angular angular JJ 37602 268 9 and and CC 37602 268 10 bear bear VB 37602 268 11 numerous numerous JJ 37602 268 12 teeth tooth NNS 37602 268 13 . . . 37602 269 1 The the DT 37602 269 2 sphenethmoid sphenethmoid NN 37602 269 3 and and CC 37602 269 4 otoccipitals otoccipital NNS 37602 269 5 are be VBP 37602 269 6 well well RB 37602 269 7 ossified ossified JJ 37602 269 8 ; ; : 37602 269 9 a a DT 37602 269 10 dermal dermal JJ 37602 269 11 sphenethmoid sphenethmoid NN 37602 269 12 is be VBZ 37602 269 13 present present JJ 37602 269 14 only only RB 37602 269 15 in in IN 37602 269 16 _ _ NNP 37602 269 17 taurinus taurinus NN 37602 269 18 _ _ NNP 37602 269 19 . . . 37602 270 1 [ [ -LRB- 37602 270 2 Illustration illustration NN 37602 270 3 : : : 37602 270 4 FIG FIG NNP 37602 270 5 . . . 37602 271 1 7 7 LS 37602 271 2 . . . 37602 272 1 Dorsal dorsal NN 37602 272 2 views view NNS 37602 272 3 of of IN 37602 272 4 vertebral vertebral JJ 37602 272 5 columns column NNS 37602 272 6 of of IN 37602 272 7 two two CD 37602 272 8 species specie NNS 37602 272 9 of of IN 37602 272 10 _ _ NNP 37602 272 11 Osteocephalus Osteocephalus NNP 37602 272 12 _ _ NNP 37602 272 13 : : : 37602 272 14 a. a. NN 37602 273 1 _ _ NNP 37602 273 2 O. O. NNP 37602 274 1 leprieurii leprieurii NNP 37602 274 2 _ _ NNP 37602 274 3 , , , 37602 274 4 KU KU NNP 37602 274 5 125962 125962 CD 37602 274 6 , , , 37602 274 7 [ [ -LRB- 37602 274 8 F F NNP 37602 274 9 ] ] -RRB- 37602 274 10 ; ; : 37602 274 11 b. b. NNP 37602 275 1 _ _ NNP 37602 275 2 O. O. NNP 37602 276 1 buckleyi buckleyi NNP 37602 276 2 _ _ NNP 37602 276 3 , , , 37602 276 4 USNM USNM NNP 37602 276 5 165997 165997 CD 37602 276 6 , , , 37602 276 7 [ [ -LRB- 37602 276 8 F f NN 37602 276 9 ] ] -RRB- 37602 276 10 . . . 37602 277 1 ×2 ×2 LS 37602 277 2 . . . 37602 277 3 ] ] -RRB- 37602 278 1 _ _ NNP 37602 278 2 Vertebral Vertebral NNP 37602 278 3 Column._--The Column._--The NNP 37602 278 4 cervical cervical JJ 37602 278 5 cotyles cotyle NNS 37602 278 6 are be VBP 37602 278 7 uniformly uniformly RB 37602 278 8 widely widely RB 37602 278 9 displaced displace VBN 37602 278 10 . . . 37602 279 1 The the DT 37602 279 2 neural neural JJ 37602 279 3 arches arch NNS 37602 279 4 are be VBP 37602 279 5 low low JJ 37602 279 6 and and CC 37602 279 7 non non JJ 37602 279 8 - - JJ 37602 279 9 imbricate imbricate JJ 37602 279 10 . . . 37602 280 1 The the DT 37602 280 2 transverse transverse JJ 37602 280 3 processes process NNS 37602 280 4 of of IN 37602 280 5 the the DT 37602 280 6 third third JJ 37602 280 7 presacral presacral JJ 37602 280 8 vertebrae vertebra NNS 37602 280 9 are be VBP 37602 280 10 approximately approximately RB 37602 280 11 equal equal JJ 37602 280 12 in in IN 37602 280 13 width width NN 37602 280 14 to to IN 37602 280 15 the the DT 37602 280 16 sacral sacral JJ 37602 280 17 diapophyses diapophysis NNS 37602 280 18 in in IN 37602 280 19 all all DT 37602 280 20 species specie NNS 37602 280 21 , , , 37602 280 22 except except IN 37602 280 23 _ _ NNP 37602 280 24 buckleyi buckleyi NN 37602 280 25 _ _ NNP 37602 280 26 , , , 37602 280 27 in in IN 37602 280 28 which which WDT 37602 280 29 the the DT 37602 280 30 processes process NNS 37602 280 31 of of IN 37602 280 32 the the DT 37602 280 33 third third JJ 37602 280 34 presacral presacral NN 37602 280 35 are be VBP 37602 280 36 slightly slightly RB 37602 280 37 narrower narrow JJR 37602 280 38 than than IN 37602 280 39 the the DT 37602 280 40 diapophyses diapophysis NNS 37602 280 41 . . . 37602 281 1 _ _ NNP 37602 281 2 Osteocephalus osteocephalus JJ 37602 281 3 buckleyi buckleyi NN 37602 281 4 _ _ NNP 37602 281 5 is be VBZ 37602 281 6 further further RB 37602 281 7 distinguished distinguish VBN 37602 281 8 by by IN 37602 281 9 the the DT 37602 281 10 presence presence NN 37602 281 11 of of IN 37602 281 12 narrow narrow JJ 37602 281 13 transverse transverse JJ 37602 281 14 processes process NNS 37602 281 15 on on IN 37602 281 16 presacrals presacral NNS 37602 281 17 five five CD 37602 281 18 through through IN 37602 281 19 eight eight CD 37602 281 20 ( ( -LRB- 37602 281 21 Fig fig NN 37602 281 22 . . . 37602 282 1 7b 7b NNP 37602 282 2 ) ) -RRB- 37602 282 3 ; ; : 37602 282 4 males male NNS 37602 282 5 have have VBP 37602 282 6 narrower narrow JJR 37602 282 7 processes process NNS 37602 282 8 than than IN 37602 282 9 do do VBP 37602 282 10 females female NNS 37602 282 11 . . . 37602 283 1 The the DT 37602 283 2 processes process NNS 37602 283 3 are be VBP 37602 283 4 moderately moderately RB 37602 283 5 wide wide JJ 37602 283 6 but but CC 37602 283 7 subequal subequal JJ 37602 283 8 in in IN 37602 283 9 width width NN 37602 283 10 in in IN 37602 283 11 _ _ NNP 37602 283 12 pearsoni pearsoni NNP 37602 283 13 _ _ NNP 37602 283 14 , , , 37602 283 15 _ _ NNP 37602 283 16 taurinus taurinus NN 37602 283 17 _ _ NNP 37602 283 18 , , , 37602 283 19 and and CC 37602 283 20 _ _ NNP 37602 283 21 verrucigerus verrucigerus NNP 37602 283 22 _ _ NNP 37602 283 23 , , , 37602 283 24 whereas whereas IN 37602 283 25 they -PRON- PRP 37602 283 26 are be VBP 37602 283 27 nearly nearly RB 37602 283 28 equivalent equivalent JJ 37602 283 29 in in IN 37602 283 30 width width NN 37602 283 31 to to IN 37602 283 32 one one CD 37602 283 33 another another DT 37602 283 34 and and CC 37602 283 35 to to IN 37602 283 36 the the DT 37602 283 37 sacral sacral JJ 37602 283 38 diapophyses diapophysis NNS 37602 283 39 in in IN 37602 283 40 _ _ NNP 37602 283 41 leprieurii leprieurii JJ 37602 283 42 _ _ NNP 37602 283 43 ( ( -LRB- 37602 283 44 Fig Fig NNP 37602 283 45 . . . 37602 284 1 7a 7a NN 37602 284 2 ) ) -RRB- 37602 284 3 . . . 37602 285 1 The the DT 37602 285 2 sacral sacral JJ 37602 285 3 diapophyses diapophysis NNS 37602 285 4 of of IN 37602 285 5 all all DT 37602 285 6 species specie NNS 37602 285 7 are be VBP 37602 285 8 moderately moderately RB 37602 285 9 dilated dilate VBN 37602 285 10 and and CC 37602 285 11 posterolaterally posterolaterally RB 37602 285 12 inclined inclined JJ 37602 285 13 . . . 37602 286 1 The the DT 37602 286 2 coccyx coccyx NN 37602 286 3 bears bear VBZ 37602 286 4 a a DT 37602 286 5 distinct distinct JJ 37602 286 6 dorsal dorsal NN 37602 286 7 ridge ridge NN 37602 286 8 and and CC 37602 286 9 has have VBZ 37602 286 10 a a DT 37602 286 11 bicondylar bicondylar JJ 37602 286 12 articulation articulation NN 37602 286 13 with with IN 37602 286 14 the the DT 37602 286 15 sacrum sacrum NN 37602 286 16 . . . 37602 287 1 _ _ NNP 37602 287 2 Pectoral Pectoral NNP 37602 287 3 Girdle._--The Girdle._--The NNP 37602 287 4 pectoral pectoral JJ 37602 287 5 girdles girdle NNS 37602 287 6 are be VBP 37602 287 7 fully fully RB 37602 287 8 arciferal arciferal JJ 37602 287 9 and and CC 37602 287 10 bear bear VBP 37602 287 11 small small JJ 37602 287 12 , , , 37602 287 13 cartilaginous cartilaginous JJ 37602 287 14 omosterna omosterna NNP 37602 287 15 and and CC 37602 287 16 moderately moderately RB 37602 287 17 large large JJ 37602 287 18 cartilaginous cartilaginous JJ 37602 287 19 sterna sterna NNP 37602 287 20 . . . 37602 288 1 The the DT 37602 288 2 coracoids coracoid NNS 37602 288 3 are be VBP 37602 288 4 robust robust JJ 37602 288 5 , , , 37602 288 6 and and CC 37602 288 7 the the DT 37602 288 8 clavicles clavicle NNS 37602 288 9 are be VBP 37602 288 10 strongly strongly RB 37602 288 11 arched arch VBN 37602 288 12 . . . 37602 289 1 Procoracoid procoracoid JJ 37602 289 2 cartilage cartilage NN 37602 289 3 seems seem VBZ 37602 289 4 to to TO 37602 289 5 be be VB 37602 289 6 absent absent JJ 37602 289 7 . . . 37602 290 1 The the DT 37602 290 2 scapulae scapula NNS 37602 290 3 are be VBP 37602 290 4 large large JJ 37602 290 5 , , , 37602 290 6 longer long JJR 37602 290 7 than than IN 37602 290 8 the the DT 37602 290 9 clavicles clavicle NNS 37602 290 10 , , , 37602 290 11 and and CC 37602 290 12 bicapitate bicapitate VB 37602 290 13 proximally proximally RB 37602 290 14 . . . 37602 291 1 The the DT 37602 291 2 suprascapulae suprascapulae NNS 37602 291 3 are be VBP 37602 291 4 moderately moderately RB 37602 291 5 large large JJ 37602 291 6 and and CC 37602 291 7 well well RB 37602 291 8 ossified ossify VBN 37602 291 9 in in IN 37602 291 10 _ _ NNP 37602 291 11 leprieurii leprieurii JJ 37602 291 12 _ _ NNP 37602 291 13 and and CC 37602 291 14 _ _ NNP 37602 291 15 taurinus taurinus NN 37602 291 16 _ _ NNP 37602 291 17 . . . 37602 292 1 The the DT 37602 292 2 suprascapula suprascapula NN 37602 292 3 of of IN 37602 292 4 _ _ NNP 37602 292 5 verrucigerus verrucigerus NNP 37602 292 6 _ _ NNP 37602 292 7 is be VBZ 37602 292 8 poorly poorly RB 37602 292 9 ossified ossify VBN 37602 292 10 , , , 37602 292 11 and and CC 37602 292 12 that that DT 37602 292 13 of of IN 37602 292 14 _ _ NNP 37602 292 15 buckleyi buckleyi NN 37602 292 16 _ _ NNP 37602 292 17 is be VBZ 37602 292 18 not not RB 37602 292 19 ossified ossify VBN 37602 292 20 . . . 37602 293 1 _ _ NNP 37602 293 2 Pelvic Pelvic NNP 37602 293 3 Girdle._--The Girdle._--The NNP 37602 293 4 ilia ilia NN 37602 293 5 of of IN 37602 293 6 _ _ NNP 37602 293 7 buckleyi buckleyi NN 37602 293 8 _ _ NNP 37602 293 9 , , , 37602 293 10 _ _ NNP 37602 293 11 taurinus taurinus NN 37602 293 12 _ _ NNP 37602 293 13 , , , 37602 293 14 and and CC 37602 293 15 _ _ NNP 37602 293 16 verrucigerus verrucigerus NN 37602 293 17 _ _ NNP 37602 293 18 lack lack NN 37602 293 19 any any DT 37602 293 20 indication indication NN 37602 293 21 of of IN 37602 293 22 a a DT 37602 293 23 crest crest NN 37602 293 24 on on IN 37602 293 25 the the DT 37602 293 26 shaft shaft NN 37602 293 27 , , , 37602 293 28 whereas whereas IN 37602 293 29 _ _ NNP 37602 293 30 leprieurii leprieurii JJ 37602 293 31 _ _ NNP 37602 293 32 has have VBZ 37602 293 33 a a DT 37602 293 34 low low JJ 37602 293 35 crest crest NN 37602 293 36 . . . 37602 294 1 The the DT 37602 294 2 dorsal dorsal NN 37602 294 3 acetabular acetabular JJ 37602 294 4 expansion expansion NN 37602 294 5 of of IN 37602 294 6 the the DT 37602 294 7 ilia ilia NNP 37602 294 8 is be VBZ 37602 294 9 moderately moderately RB 37602 294 10 low low JJ 37602 294 11 in in IN 37602 294 12 _ _ NNP 37602 294 13 taurinus taurinus NNP 37602 294 14 _ _ NNP 37602 294 15 and and CC 37602 294 16 _ _ NNP 37602 294 17 verrucigerus verrucigerus NNP 37602 294 18 _ _ NNP 37602 294 19 , , , 37602 294 20 but but CC 37602 294 21 distinctly distinctly RB 37602 294 22 lower low JJR 37602 294 23 in in IN 37602 294 24 _ _ NNP 37602 294 25 buckleyi buckleyi NN 37602 294 26 _ _ NNP 37602 294 27 and and CC 37602 294 28 _ _ NNP 37602 294 29 leprieurii leprieurii JJ 37602 294 30 _ _ NNP 37602 294 31 . . . 37602 295 1 The the DT 37602 295 2 ilia ilia NN 37602 295 3 of of IN 37602 295 4 all all DT 37602 295 5 species specie NNS 37602 295 6 bear bear VBP 37602 295 7 low low JJ 37602 295 8 dorsal dorsal NN 37602 295 9 protuberances protuberance NNS 37602 295 10 . . . 37602 296 1 The the DT 37602 296 2 ischia ischia NN 37602 296 3 of of IN 37602 296 4 _ _ NNP 37602 296 5 leprieurii leprieurii JJ 37602 296 6 _ _ NNP 37602 296 7 , , , 37602 296 8 _ _ NNP 37602 296 9 taurinus taurinus NN 37602 296 10 _ _ NNP 37602 296 11 , , , 37602 296 12 and and CC 37602 296 13 _ _ NNP 37602 296 14 verrucigerus verrucigerus NN 37602 296 15 _ _ NNP 37602 296 16 are be VBP 37602 296 17 moderately moderately RB 37602 296 18 expanded expand VBN 37602 296 19 ; ; : 37602 296 20 that that DT 37602 296 21 of of IN 37602 296 22 _ _ NNP 37602 296 23 buckleyi buckleyi NN 37602 296 24 _ _ NNP 37602 296 25 is be VBZ 37602 296 26 somewhat somewhat RB 37602 296 27 less less RBR 37602 296 28 expanded expand VBN 37602 296 29 dorsally dorsally RB 37602 296 30 . . . 37602 297 1 The the DT 37602 297 2 pubis pubis NN 37602 297 3 of of IN 37602 297 4 _ _ NNP 37602 297 5 leprieurii leprieurii JJ 37602 297 6 _ _ NNP 37602 297 7 , , , 37602 297 8 _ _ NNP 37602 297 9 taurinus taurinus NN 37602 297 10 _ _ NNP 37602 297 11 , , , 37602 297 12 and and CC 37602 297 13 _ _ NNP 37602 297 14 verrucigerus verrucigerus NN 37602 297 15 _ _ NNP 37602 297 16 are be VBP 37602 297 17 calcified calcify VBN 37602 297 18 , , , 37602 297 19 whereas whereas IN 37602 297 20 that that DT 37602 297 21 of of IN 37602 297 22 _ _ NNP 37602 297 23 buckleyi buckleyi NN 37602 297 24 _ _ NNP 37602 297 25 remains remain VBZ 37602 297 26 cartilaginous cartilaginous JJ 37602 297 27 . . . 37602 298 1 _ _ NNP 37602 298 2 Throat Throat NNP 37602 298 3 Musculature Musculature NNP 37602 298 4 and and CC 37602 298 5 Vocal Vocal NNP 37602 298 6 Sac Sac NNP 37602 298 7 Structure._--Tyler structure._--tyler CD 37602 298 8 ( ( -LRB- 37602 298 9 1971 1971 CD 37602 298 10 ) ) -RRB- 37602 298 11 described describe VBD 37602 298 12 the the DT 37602 298 13 throat throat NN 37602 298 14 myology myology NN 37602 298 15 of of IN 37602 298 16 _ _ NNP 37602 298 17 Osteocephalus Osteocephalus NNP 37602 298 18 _ _ NNP 37602 298 19 . . . 37602 299 1 The the DT 37602 299 2 genus genus NN 37602 299 3 is be VBZ 37602 299 4 characterized characterize VBN 37602 299 5 by by IN 37602 299 6 a a DT 37602 299 7 moderate moderate JJ 37602 299 8 - - HYPH 37602 299 9 sized sized JJ 37602 299 10 araphic araphic JJ 37602 299 11 submentalis submentalis NNP 37602 299 12 muscle muscle NNP 37602 299 13 and and CC 37602 299 14 an an DT 37602 299 15 undifferentiated undifferentiated JJ 37602 299 16 intermandibularis intermandibularis NNP 37602 299 17 having have VBG 37602 299 18 an an DT 37602 299 19 elongate elongate NN 37602 299 20 median median JJ 37602 299 21 aponeurosis aponeurosis NN 37602 299 22 . . . 37602 300 1 The the DT 37602 300 2 intermandibularis intermandibularis NNP 37602 300 3 and and CC 37602 300 4 submentalis submentalis NNP 37602 300 5 are be VBP 37602 300 6 completely completely RB 37602 300 7 independent independent JJ 37602 300 8 in in IN 37602 300 9 _ _ NNP 37602 300 10 buckleyi buckleyi NN 37602 300 11 _ _ NNP 37602 300 12 , , , 37602 300 13 whereas whereas IN 37602 300 14 in in IN 37602 300 15 the the DT 37602 300 16 other other JJ 37602 300 17 species specie NNS 37602 300 18 there there EX 37602 300 19 is be VBZ 37602 300 20 a a DT 37602 300 21 small small JJ 37602 300 22 attachment attachment NN 37602 300 23 between between IN 37602 300 24 these these DT 37602 300 25 muscles muscle NNS 37602 300 26 . . . 37602 301 1 According accord VBG 37602 301 2 to to IN 37602 301 3 Tyler Tyler NNP 37602 301 4 ( ( -LRB- 37602 301 5 pers per NNS 37602 301 6 . . . 37602 302 1 com com NN 37602 302 2 . . . 37602 303 1 ) ) -RRB- 37602 303 2 , , , 37602 303 3 _ _ NNP 37602 303 4 Osteocephalus Osteocephalus NNP 37602 303 5 _ _ NNP 37602 303 6 has have VBZ 37602 303 7 three three CD 37602 303 8 distinctive distinctive JJ 37602 303 9 types type NNS 37602 303 10 of of IN 37602 303 11 vocal vocal JJ 37602 303 12 sac sac NN 37602 303 13 structure structure NN 37602 303 14 which which WDT 37602 303 15 result result VBP 37602 303 16 from from IN 37602 303 17 differences difference NNS 37602 303 18 in in IN 37602 303 19 the the DT 37602 303 20 development development NN 37602 303 21 of of IN 37602 303 22 the the DT 37602 303 23 interhyoideus interhyoideus NNP 37602 303 24 muscle muscle NN 37602 303 25 and and CC 37602 303 26 the the DT 37602 303 27 overlying overlying NN 37602 303 28 skin skin NN 37602 303 29 . . . 37602 304 1 In in IN 37602 304 2 _ _ NNP 37602 304 3 leprieurii leprieurii JJ 37602 304 4 _ _ NNP 37602 304 5 and and CC 37602 304 6 _ _ NNP 37602 304 7 verrucigerus verrucigerus NN 37602 304 8 _ _ NNP 37602 304 9 the the DT 37602 304 10 supramandibular supramandibular JJ 37602 304 11 portions portion NNS 37602 304 12 of of IN 37602 304 13 the the DT 37602 304 14 interhyoideus interhyoideu NNS 37602 304 15 form form NN 37602 304 16 a a DT 37602 304 17 simple simple JJ 37602 304 18 tubular tubular NN 37602 304 19 , , , 37602 304 20 posterolateral posterolateral JJ 37602 304 21 extension extension NN 37602 304 22 ; ; : 37602 304 23 there there EX 37602 304 24 is be VBZ 37602 304 25 no no DT 37602 304 26 modification modification NN 37602 304 27 of of IN 37602 304 28 the the DT 37602 304 29 associated associated JJ 37602 304 30 skin skin NN 37602 304 31 . . . 37602 305 1 _ _ NNP 37602 305 2 Osteocephalus osteocephalus JJ 37602 305 3 buckleyi buckleyi NN 37602 305 4 _ _ NNP 37602 305 5 and and CC 37602 305 6 _ _ NNP 37602 305 7 pearsoni pearsoni NNP 37602 305 8 _ _ NNP 37602 305 9 have have VBP 37602 305 10 more more RBR 37602 305 11 extensive extensive JJ 37602 305 12 development development NN 37602 305 13 of of IN 37602 305 14 the the DT 37602 305 15 supramandibular supramandibular JJ 37602 305 16 portions portion NNS 37602 305 17 of of IN 37602 305 18 the the DT 37602 305 19 interhyoideus interhyoideu NNS 37602 305 20 ; ; : 37602 305 21 furthermore furthermore RB 37602 305 22 , , , 37602 305 23 the the DT 37602 305 24 associated associated JJ 37602 305 25 skin skin NN 37602 305 26 forms form VBZ 37602 305 27 a a DT 37602 305 28 broad broad JJ 37602 305 29 , , , 37602 305 30 loose loose JJ 37602 305 31 fold fold JJ 37602 305 32 extending extend VBG 37602 305 33 from from IN 37602 305 34 the the DT 37602 305 35 ventromedial ventromedial JJ 37602 305 36 surface surface NN 37602 305 37 of of IN 37602 305 38 the the DT 37602 305 39 throat throat NN 37602 305 40 dorsolaterally dorsolaterally RB 37602 305 41 to to IN 37602 305 42 the the DT 37602 305 43 base base NN 37602 305 44 of of IN 37602 305 45 the the DT 37602 305 46 supratympanic supratympanic JJ 37602 305 47 fold fold NN 37602 305 48 . . . 37602 306 1 Like like IN 37602 306 2 _ _ NNP 37602 306 3 buckleyi buckleyi NN 37602 306 4 _ _ NNP 37602 306 5 and and CC 37602 306 6 _ _ NNP 37602 306 7 pearsoni pearsoni NNP 37602 306 8 _ _ NNP 37602 306 9 , , , 37602 306 10 the the DT 37602 306 11 supramandibular supramandibular JJ 37602 306 12 portion portion NN 37602 306 13 of of IN 37602 306 14 the the DT 37602 306 15 interhyoideus interhyoideu NNS 37602 306 16 is be VBZ 37602 306 17 much much RB 37602 306 18 expanded expand VBN 37602 306 19 in in IN 37602 306 20 _ _ NNP 37602 306 21 taurinus taurinus NNP 37602 306 22 _ _ NNP 37602 306 23 . . . 37602 307 1 The the DT 37602 307 2 vocal vocal JJ 37602 307 3 sac sac NN 37602 307 4 structure structure NN 37602 307 5 of of IN 37602 307 6 _ _ NNP 37602 307 7 taurinus taurinus NN 37602 307 8 _ _ NNP 37602 307 9 differs differ VBZ 37602 307 10 from from IN 37602 307 11 that that DT 37602 307 12 of of IN 37602 307 13 other other JJ 37602 307 14 members member NNS 37602 307 15 of of IN 37602 307 16 the the DT 37602 307 17 genus genus NN 37602 307 18 in in IN 37602 307 19 that that IN 37602 307 20 the the DT 37602 307 21 skin skin NN 37602 307 22 of of IN 37602 307 23 _ _ NNP 37602 307 24 taurinus taurinus NNP 37602 307 25 _ _ NNP 37602 307 26 forms form VBZ 37602 307 27 an an DT 37602 307 28 everted evert VBN 37602 307 29 pouch pouch NN 37602 307 30 , , , 37602 307 31 which which WDT 37602 307 32 dangles dangle VBZ 37602 307 33 loosely loosely RB 37602 307 34 beneath beneath IN 37602 307 35 the the DT 37602 307 36 supratympanic supratympanic JJ 37602 307 37 fold fold NN 37602 307 38 . . . 37602 308 1 KEY key NN 37602 308 2 TO to IN 37602 308 3 THE the DT 37602 308 4 SPECIES specie NNS 37602 308 5 OF of IN 37602 308 6 _ _ NNP 37602 308 7 Osteocephalus Osteocephalus NNP 37602 308 8 _ _ NNP 37602 308 9 1 1 CD 37602 308 10 . . . 37602 309 1 Inner inner JJ 37602 309 2 edge edge NN 37602 309 3 of of IN 37602 309 4 third third JJ 37602 309 5 finger finger NN 37602 309 6 webbed web VBN 37602 309 7 to to IN 37602 309 8 mid mid NN 37602 309 9 - - NN 37602 309 10 length length NN 37602 309 11 of of IN 37602 309 12 antepenultimate antepenultimate NN 37602 309 13 phalange phalange NN 37602 309 14 ; ; : 37602 309 15 dorsum dorsum NN 37602 309 16 brown brown NNP 37602 309 17 with with IN 37602 309 18 dark dark JJ 37602 309 19 brown brown JJ 37602 309 20 spots spot NNS 37602 309 21 or or CC 37602 309 22 median median JJ 37602 309 23 blotch blotch NN 37602 309 24 ; ; : 37602 309 25 skull skull VBG 37602 309 26 in in IN 37602 309 27 adults adult NNS 37602 309 28 casqued casque VBN 37602 309 29 and and CC 37602 309 30 co co VBN 37602 309 31 - - VBN 37602 309 32 ossified ossified JJ 37602 309 33 with with IN 37602 309 34 prominent prominent JJ 37602 309 35 supraorbital supraorbital JJ 37602 309 36 flanges flange NNS 37602 309 37 _ _ NNP 37602 309 38 O. O. NNP 37602 310 1 taurinus taurinus NNP 37602 310 2 _ _ NNP 37602 310 3 Inner inner JJ 37602 310 4 edge edge NN 37602 310 5 of of IN 37602 310 6 third third JJ 37602 310 7 finger finger NN 37602 310 8 webbed web VBN 37602 310 9 to to IN 37602 310 10 base base NN 37602 310 11 of of IN 37602 310 12 antepenultimate antepenultimate NN 37602 310 13 phalange phalange NN 37602 310 14 ; ; : 37602 310 15 dorsum dorsum NN 37602 310 16 plain plain JJ 37602 310 17 or or CC 37602 310 18 marked mark VBN 37602 310 19 with with IN 37602 310 20 dark dark JJ 37602 310 21 blotches blotch NNS 37602 310 22 or or CC 37602 310 23 transverse transverse JJ 37602 310 24 bars bar NNS 37602 310 25 ; ; : 37602 310 26 skull skull VBG 37602 310 27 in in IN 37602 310 28 adults adult NNS 37602 310 29 smooth smooth VBP 37602 310 30 or or CC 37602 310 31 slightly slightly RB 37602 310 32 exostosed exostose VBN 37602 310 33 , , , 37602 310 34 lacking lack VBG 37602 310 35 supraorbital supraorbital JJ 37602 310 36 flanges flange NNS 37602 310 37 2 2 CD 37602 310 38 2 2 CD 37602 310 39 . . . 37602 311 1 Skin skin NN 37602 311 2 on on IN 37602 311 3 flanks flank NNS 37602 311 4 areolate areolate NN 37602 311 5 ; ; , 37602 311 6 dorsum dorsum NN 37602 311 7 in in IN 37602 311 8 males male NNS 37602 311 9 bearing bear VBG 37602 311 10 a a DT 37602 311 11 mixture mixture NN 37602 311 12 of of IN 37602 311 13 large large JJ 37602 311 14 and and CC 37602 311 15 small small JJ 37602 311 16 non non JJ 37602 311 17 - - JJ 37602 311 18 spinous spinous JJ 37602 311 19 tubercles tubercle NNS 37602 311 20 ; ; : 37602 311 21 lips lip NNS 37602 311 22 distinctly distinctly RB 37602 311 23 barred bar VBN 37602 311 24 _ _ NNP 37602 311 25 O. O. NNP 37602 312 1 buckleyi buckleyi NNP 37602 312 2 _ _ NNP 37602 312 3 Skin Skin NNP 37602 312 4 on on IN 37602 312 5 flanks flank NNS 37602 312 6 smooth smooth VBP 37602 312 7 ; ; : 37602 312 8 dorsum dorsum NN 37602 312 9 in in IN 37602 312 10 males male NNS 37602 312 11 bearing bear VBG 37602 312 12 tubercles tubercle NNS 37602 312 13 of of IN 37602 312 14 uniform uniform JJ 37602 312 15 size size NN 37602 312 16 ; ; , 37602 312 17 lips lip NNS 37602 312 18 not not RB 37602 312 19 barred bar VBN 37602 312 20 3 3 CD 37602 312 21 3 3 CD 37602 312 22 . . . 37602 313 1 Dorsal dorsal NN 37602 313 2 pattern pattern NN 37602 313 3 consisting consist VBG 37602 313 4 of of IN 37602 313 5 narrow narrow JJ 37602 313 6 transverse transverse JJ 37602 313 7 dark dark JJ 37602 313 8 bars bar NNS 37602 313 9 ; ; , 37602 313 10 dorsum dorsum NN 37602 313 11 in in IN 37602 313 12 males male NNS 37602 313 13 bearing bear VBG 37602 313 14 numerous numerous JJ 37602 313 15 small small JJ 37602 313 16 spinous spinous JJ 37602 313 17 tubercles tubercle NNS 37602 313 18 _ _ NNP 37602 313 19 O. O. NNP 37602 314 1 leprieurii leprieurii JJ 37602 314 2 _ _ NNP 37602 314 3 Dorsal Dorsal NNP 37602 314 4 pattern pattern NN 37602 314 5 not not RB 37602 314 6 consisting consist VBG 37602 314 7 of of IN 37602 314 8 transverse transverse JJ 37602 314 9 bars bar NNS 37602 314 10 ; ; : 37602 314 11 dorsal dorsal NN 37602 314 12 tubercles tubercle NNS 37602 314 13 large large JJ 37602 314 14 or or CC 37602 314 15 few few JJ 37602 314 16 in in IN 37602 314 17 number number NN 37602 314 18 4 4 CD 37602 314 19 4 4 CD 37602 314 20 . . . 37602 315 1 Dorsum dorsum NN 37602 315 2 uniformly uniformly JJ 37602 315 3 dark dark JJ 37602 315 4 brown brown NN 37602 315 5 ; ; : 37602 315 6 venter venter NN 37602 315 7 heavily heavily RB 37602 315 8 mottled mottle VBN 37602 315 9 with with IN 37602 315 10 black black JJ 37602 315 11 , , , 37602 315 12 especially especially RB 37602 315 13 in in IN 37602 315 14 females female NNS 37602 315 15 ; ; : 37602 315 16 dorsum dorsum NN 37602 315 17 in in IN 37602 315 18 males male NNS 37602 315 19 bearing bear VBG 37602 315 20 large large JJ 37602 315 21 , , , 37602 315 22 keratinized keratinize VBN 37602 315 23 tubercles tubercle NNS 37602 315 24 _ _ NNP 37602 315 25 O. O. NNP 37602 316 1 verrucigerus verrucigerus NNP 37602 316 2 _ _ NNP 37602 316 3 Dorsum Dorsum NNP 37602 316 4 tan tan NN 37602 316 5 with with IN 37602 316 6 irregular irregular JJ 37602 316 7 dark dark JJ 37602 316 8 brown brown JJ 37602 316 9 blotches blotch NNS 37602 316 10 ; ; : 37602 316 11 venter venter NN 37602 316 12 cream cream NN 37602 316 13 with with IN 37602 316 14 fine fine JJ 37602 316 15 brown brown JJ 37602 316 16 reticulations reticulation NNS 37602 316 17 ; ; , 37602 316 18 dorsum dorsum NN 37602 316 19 in in IN 37602 316 20 males male NNS 37602 316 21 bearing bear VBG 37602 316 22 few few JJ 37602 316 23 , , , 37602 316 24 small small JJ 37602 316 25 non non JJ 37602 316 26 - - JJ 37602 316 27 spinous spinous JJ 37602 316 28 tubercles tubercle NNS 37602 316 29 _ _ NNP 37602 316 30 O. O. NNP 37602 317 1 pearsoni pearsoni NNP 37602 317 2 _ _ NNP 37602 317 3 ACCOUNTS ACCOUNTS NNP 37602 317 4 OF of IN 37602 317 5 SPECIES SPECIES NNP 37602 317 6 = = SYM 37602 317 7 Osteocephalus osteocephalus NN 37602 317 8 buckleyi= buckleyi= NN 37602 317 9 ( ( -LRB- 37602 317 10 Boulenger boulenger NN 37602 317 11 ) ) -RRB- 37602 317 12 _ _ NNP 37602 317 13 Hyla Hyla NNP 37602 317 14 buckleyi buckleyi NN 37602 317 15 _ _ NNP 37602 317 16 Boulenger Boulenger NNP 37602 317 17 , , , 37602 317 18 1882:362 1882:362 CD 37602 317 19 [ [ -LRB- 37602 317 20 Syntypes.--BMNH Syntypes.--BMNH NNP 37602 317 21 1947.2.13.36 1947.2.13.36 CD 37602 317 22 - - SYM 37602 317 23 39 39 CD 37602 317 24 from from IN 37602 317 25 Sarayacu Sarayacu NNP 37602 317 26 , , , 37602 317 27 Provincia Provincia NNP 37602 317 28 Pastaza Pastaza NNP 37602 317 29 , , , 37602 317 30 Ecuador Ecuador NNP 37602 317 31 ; ; : 37602 317 32 BMNH BMNH NNP 37602 317 33 1947.2.13.40 1947.2.13.40 CD 37602 317 34 - - SYM 37602 317 35 41 41 CD 37602 317 36 , , , 37602 317 37 1947.2.13 1947.2.13 CD 37602 317 38 . . . 37602 318 1 43 43 CD 37602 318 2 - - SYM 37602 318 3 45 45 CD 37602 318 4 from from IN 37602 318 5 Canelos Canelos NNP 37602 318 6 , , , 37602 318 7 Provincia Provincia NNP 37602 318 8 Pastaza Pastaza NNP 37602 318 9 , , , 37602 318 10 Ecuador Ecuador NNP 37602 318 11 ; ; : 37602 318 12 BMNH BMNH NNP 37602 318 13 1947.2.13.46 1947.2.13.46 CD 37602 318 14 from from IN 37602 318 15 " " `` 37602 318 16 Paitanga Paitanga NNP 37602 318 17 " " '' 37602 318 18 (= (= NNP 37602 318 19 Pallatanga Pallatanga NNP 37602 318 20 ) ) -RRB- 37602 318 21 , , , 37602 318 22 Provincia Provincia NNP 37602 318 23 Chimborazo Chimborazo NNP 37602 318 24 , , , 37602 318 25 Ecuador Ecuador NNP 37602 318 26 ( ( -LRB- 37602 318 27 in in IN 37602 318 28 error error NN 37602 318 29 ) ) -RRB- 37602 318 30 ; ; : 37602 318 31 Mr. Mr. NNP 37602 318 32 Buckley Buckley NNP 37602 318 33 collector collector NN 37602 318 34 ; ; : 37602 318 35 BMNH BMNH NNP 37602 318 36 1947.2.13.44 1947.2.13.44 NNS 37602 318 37 here here RB 37602 318 38 designated designate VBN 37602 318 39 as as IN 37602 318 40 lectotype lectotype NNP 37602 318 41 ] ] -RRB- 37602 318 42 . . . 37602 319 1 _ _ NNP 37602 319 2 Hyla Hyla NNP 37602 319 3 festae festae VBD 37602 319 4 _ _ NNP 37602 319 5 Peracca Peracca NNP 37602 319 6 , , , 37602 319 7 1904:39 1904:39 CD 37602 319 8 [ [ -LRB- 37602 319 9 Holotype.--MIZS holotype.--miz VBZ 37602 319 10 2950 2950 CD 37602 319 11 from from IN 37602 319 12 " " `` 37602 319 13 Valle Valle NNP 37602 319 14 de de NNP 37602 319 15 Santiago Santiago NNP 37602 319 16 " " '' 37602 319 17 (= (= NNP 37602 319 18 lower low JJR 37602 319 19 Río Río NNP 37602 319 20 Zamora Zamora NNP 37602 319 21 ) ) -RRB- 37602 319 22 , , , 37602 319 23 Provincia Provincia NNP 37602 319 24 Morona Morona NNP 37602 319 25 - - HYPH 37602 319 26 Santiago Santiago NNP 37602 319 27 , , , 37602 319 28 Ecuador Ecuador NNP 37602 319 29 ; ; : 37602 319 30 Enrico Enrico NNP 37602 319 31 Festa Festa NNP 37602 319 32 collector collector NN 37602 319 33 ] ] -RRB- 37602 319 34 . . . 37602 320 1 New new JJ 37602 320 2 synonymy synonymy NNP 37602 320 3 . . . 37602 321 1 _ _ NNP 37602 321 2 Osteocephalus Osteocephalus NNP 37602 321 3 buckleyi_--Goin buckleyi_--Goin NNP 37602 321 4 , , , 37602 321 5 1961:13 1961:13 CD 37602 321 6 . . . 37602 322 1 _ _ NNP 37602 322 2 Hyla Hyla NNP 37602 322 3 carri carri VBD 37602 322 4 _ _ NNP 37602 322 5 Cochran Cochran NNP 37602 322 6 and and CC 37602 322 7 Goin go NNP 37602 322 8 , , , 37602 322 9 1970:211 1970:211 CD 37602 322 10 [ [ -LRB- 37602 322 11 Holotype.--FMNH holotype.--fmnh NN 37602 322 12 69702 69702 CD 37602 322 13 from from IN 37602 322 14 Acevedo Acevedo NNP 37602 322 15 , , , 37602 322 16 Río Río NNP 37602 322 17 Suaza Suaza NNP 37602 322 18 , , , 37602 322 19 Departamento Departamento NNP 37602 322 20 Huila Huila NNP 37602 322 21 , , , 37602 322 22 Colombia Colombia NNP 37602 322 23 ; ; : 37602 322 24 Philip Philip NNP 37602 322 25 collector collector NN 37602 322 26 ] ] -RRB- 37602 322 27 . . . 37602 323 1 New new JJ 37602 323 2 synonymy synonymy NNP 37602 323 3 . . . 37602 324 1 _ _ NNP 37602 324 2 Hyla Hyla NNP 37602 324 3 cabrerai cabrerai NN 37602 324 4 _ _ NNP 37602 324 5 Cochran Cochran NNP 37602 324 6 and and CC 37602 324 7 Goin go NNP 37602 324 8 , , , 37602 324 9 1970:215 1970:215 CD 37602 324 10 [ [ -LRB- 37602 324 11 Holotype.--USNM holotype.--usnm NN 37602 324 12 152759 152759 CD 37602 324 13 from from IN 37602 324 14 Caño Caño NNP 37602 324 15 Guacayá Guacayá NNP 37602 324 16 , , , 37602 324 17 tributary tributary NN 37602 324 18 of of IN 37602 324 19 lower low JJR 37602 324 20 Río Río NNP 37602 324 21 Apoporis Apoporis NNP 37602 324 22 , , , 37602 324 23 Comisaria Comisaria NNP 37602 324 24 Amazonas Amazonas NNP 37602 324 25 , , , 37602 324 26 Colombia Colombia NNP 37602 324 27 ; ; : 37602 324 28 Isadore Isadore NNP 37602 324 29 Cabrera Cabrera NNP 37602 324 30 collector collector NN 37602 324 31 ] ] -RRB- 37602 324 32 . . . 37602 325 1 New new JJ 37602 325 2 synonymy synonymy NNP 37602 325 3 . . . 37602 326 1 _ _ NNP 37602 326 2 Justification Justification NNP 37602 326 3 of of IN 37602 326 4 Synonymy._--Boulenger Synonymy._--Boulenger NNP 37602 326 5 ( ( -LRB- 37602 326 6 1882:362 1882:362 CD 37602 326 7 ) ) -RRB- 37602 326 8 listed list VBN 37602 326 9 11 11 CD 37602 326 10 specimens specimen NNS 37602 326 11 in in IN 37602 326 12 his -PRON- PRP$ 37602 326 13 description description NN 37602 326 14 of of IN 37602 326 15 _ _ NNP 37602 326 16 Hyla Hyla NNP 37602 326 17 buckleyi buckleyi NN 37602 326 18 _ _ NNP 37602 326 19 . . . 37602 327 1 We -PRON- PRP 37602 327 2 have have VBP 37602 327 3 examined examine VBN 37602 327 4 all all DT 37602 327 5 of of IN 37602 327 6 these these DT 37602 327 7 and and CC 37602 327 8 conclude conclude VBP 37602 327 9 that that IN 37602 327 10 one one NN 37602 327 11 ( ( -LRB- 37602 327 12 BMNH BMNH NNP 37602 327 13 1947.2.13.42 1947.2.13.42 NNS 37602 327 14 ) ) -RRB- 37602 327 15 is be VBZ 37602 327 16 _ _ NNP 37602 327 17 O. O. NNP 37602 328 1 leprieurii leprieurii NNP 37602 328 2 _ _ NNP 37602 328 3 . . . 37602 329 1 Cochran Cochran NNP 37602 329 2 and and CC 37602 329 3 Goin go NNP 37602 329 4 ( ( -LRB- 37602 329 5 1970:213 1970:213 CD 37602 329 6 ) ) -RRB- 37602 329 7 restricted restrict VBD 37602 329 8 the the DT 37602 329 9 type type NN 37602 329 10 locality locality NN 37602 329 11 to to IN 37602 329 12 Canelos Canelos NNP 37602 329 13 , , , 37602 329 14 Provincia Provincia NNP 37602 329 15 Pastaza Pastaza NNP 37602 329 16 , , , 37602 329 17 Ecuador Ecuador NNP 37602 329 18 ; ; : 37602 329 19 we -PRON- PRP 37602 329 20 here here RB 37602 329 21 select select VBP 37602 329 22 BMNH BMNH NNS 37602 329 23 1947.2.13.44 1947.2.13.44 NNS 37602 329 24 from from IN 37602 329 25 that that DT 37602 329 26 locality locality NN 37602 329 27 as as IN 37602 329 28 the the DT 37602 329 29 lectotype lectotype NN 37602 329 30 . . . 37602 330 1 This this DT 37602 330 2 specimen specimen NN 37602 330 3 is be VBZ 37602 330 4 a a DT 37602 330 5 male male NN 37602 330 6 having have VBG 37602 330 7 a a DT 37602 330 8 snout snout NN 37602 330 9 - - HYPH 37602 330 10 vent vent NN 37602 330 11 length length NN 37602 330 12 of of IN 37602 330 13 37.9 37.9 CD 37602 330 14 mm mm NNP 37602 330 15 ; ; : 37602 330 16 the the DT 37602 330 17 diameter diameter NN 37602 330 18 of of IN 37602 330 19 the the DT 37602 330 20 tympanum tympanum NN 37602 330 21 is be VBZ 37602 330 22 3.5 3.5 CD 37602 330 23 mm mm CD 37602 330 24 , , , 37602 330 25 81.4 81.4 CD 37602 330 26 percent percent NN 37602 330 27 of of IN 37602 330 28 the the DT 37602 330 29 diameter diameter NN 37602 330 30 of of IN 37602 330 31 the the DT 37602 330 32 eye eye NN 37602 330 33 . . . 37602 331 1 The the DT 37602 331 2 type type NN 37602 331 3 series series NN 37602 331 4 , , , 37602 331 5 exclusive exclusive JJ 37602 331 6 of of IN 37602 331 7 BMNH BMNH NNP 37602 331 8 1947.2.13.42 1947.2.13.42 NN 37602 331 9 ( ( -LRB- 37602 331 10 = = NFP 37602 331 11 _ _ NNP 37602 331 12 O. O. NNP 37602 332 1 leprieurii leprieurii NNP 37602 332 2 _ _ NNP 37602 332 3 ) ) -RRB- 37602 332 4 consists consist VBZ 37602 332 5 of of IN 37602 332 6 six six CD 37602 332 7 males male NNS 37602 332 8 having have VBG 37602 332 9 snout snout NN 37602 332 10 - - HYPH 37602 332 11 vent vent NN 37602 332 12 lengths length NNS 37602 332 13 of of IN 37602 332 14 37.9 37.9 CD 37602 332 15 - - HYPH 37602 332 16 44.6 44.6 CD 37602 332 17 ( ( -LRB- 37602 332 18 mean mean NNP 37602 332 19 40.4 40.4 CD 37602 332 20 ) ) -RRB- 37602 332 21 mm mm NNP 37602 332 22 , , , 37602 332 23 and and CC 37602 332 24 four four CD 37602 332 25 females female NNS 37602 332 26 having have VBG 37602 332 27 snout snout NN 37602 332 28 - - HYPH 37602 332 29 vent vent NN 37602 332 30 lengths length NNS 37602 332 31 of of IN 37602 332 32 50.0 50.0 CD 37602 332 33 - - HYPH 37602 332 34 53.9 53.9 CD 37602 332 35 ( ( -LRB- 37602 332 36 mean mean NNP 37602 332 37 51.5 51.5 CD 37602 332 38 ) ) -RRB- 37602 332 39 mm mm NNP 37602 332 40 . . . 37602 333 1 The the DT 37602 333 2 dorsum dorsum NN 37602 333 3 in in IN 37602 333 4 the the DT 37602 333 5 males male NNS 37602 333 6 bears bear VBZ 37602 333 7 a a DT 37602 333 8 mixture mixture NN 37602 333 9 of of IN 37602 333 10 large large JJ 37602 333 11 and and CC 37602 333 12 small small JJ 37602 333 13 tubercles tubercle NNS 37602 333 14 , , , 37602 333 15 whereas whereas IN 37602 333 16 the the DT 37602 333 17 dorsum dorsum NN 37602 333 18 in in IN 37602 333 19 females female NNS 37602 333 20 is be VBZ 37602 333 21 nearly nearly RB 37602 333 22 smooth smooth JJ 37602 333 23 . . . 37602 334 1 The the DT 37602 334 2 skin skin NN 37602 334 3 on on IN 37602 334 4 the the DT 37602 334 5 flanks flank NNS 37602 334 6 , , , 37602 334 7 especially especially RB 37602 334 8 the the DT 37602 334 9 axilla axilla NN 37602 334 10 , , , 37602 334 11 is be VBZ 37602 334 12 areolate areolate NN 37602 334 13 . . . 37602 335 1 The the DT 37602 335 2 coloration coloration NN 37602 335 3 consists consist VBZ 37602 335 4 of of IN 37602 335 5 a a DT 37602 335 6 creamy creamy JJ 37602 335 7 tan tan NN 37602 335 8 ground ground NN 37602 335 9 color color NN 37602 335 10 with with IN 37602 335 11 irregular irregular JJ 37602 335 12 reddish reddish JJ 37602 335 13 brown brown JJ 37602 335 14 markings marking NNS 37602 335 15 on on IN 37602 335 16 the the DT 37602 335 17 back back JJ 37602 335 18 and and CC 37602 335 19 broad broad JJ 37602 335 20 transverse transverse JJ 37602 335 21 bars bar NNS 37602 335 22 on on IN 37602 335 23 the the DT 37602 335 24 limbs limb NNS 37602 335 25 . . . 37602 336 1 The the DT 37602 336 2 dorsal dorsal NN 37602 336 3 markings marking NNS 37602 336 4 are be VBP 37602 336 5 narrowly narrowly RB 37602 336 6 bordered border VBN 37602 336 7 by by IN 37602 336 8 creamy creamy JJ 37602 336 9 white white NNP 37602 336 10 ; ; : 37602 336 11 those those DT 37602 336 12 on on IN 37602 336 13 the the DT 37602 336 14 back back JJ 37602 336 15 consist consist NN 37602 336 16 of of IN 37602 336 17 an an DT 37602 336 18 interorbital interorbital JJ 37602 336 19 bar bar NN 37602 336 20 and and CC 37602 336 21 a a DT 37602 336 22 pair pair NN 37602 336 23 of of IN 37602 336 24 longitudinal longitudinal JJ 37602 336 25 marks mark NNS 37602 336 26 beginning begin VBG 37602 336 27 in in IN 37602 336 28 the the DT 37602 336 29 scapular scapular JJ 37602 336 30 region region NN 37602 336 31 and and CC 37602 336 32 usually usually RB 37602 336 33 diverging diverge VBG 37602 336 34 posteriorly posteriorly RB 37602 336 35 in in IN 37602 336 36 the the DT 37602 336 37 sacral sacral JJ 37602 336 38 region region NN 37602 336 39 or or CC 37602 336 40 converging converge VBG 37602 336 41 into into IN 37602 336 42 a a DT 37602 336 43 broad broad JJ 37602 336 44 median median JJ 37602 336 45 blotch blotch NN 37602 336 46 . . . 37602 337 1 One one CD 37602 337 2 specimen speciman NNS 37602 337 3 has have VBZ 37602 337 4 a a DT 37602 337 5 middorsal middorsal NN 37602 337 6 creamy creamy JJ 37602 337 7 white white JJ 37602 337 8 stripe stripe NN 37602 337 9 from from IN 37602 337 10 the the DT 37602 337 11 tip tip NN 37602 337 12 of of IN 37602 337 13 the the DT 37602 337 14 snout snout NN 37602 337 15 to to IN 37602 337 16 the the DT 37602 337 17 vent vent NN 37602 337 18 . . . 37602 338 1 In in IN 37602 338 2 all all DT 37602 338 3 of of IN 37602 338 4 the the DT 37602 338 5 types type NNS 37602 338 6 large large JJ 37602 338 7 dark dark JJ 37602 338 8 brown brown JJ 37602 338 9 spots spot NNS 37602 338 10 are be VBP 37602 338 11 present present JJ 37602 338 12 on on IN 37602 338 13 the the DT 37602 338 14 flanks flank NNS 37602 338 15 and and CC 37602 338 16 posterior posterior NN 37602 338 17 surfaces surface NNS 37602 338 18 of of IN 37602 338 19 the the DT 37602 338 20 thighs thigh NNS 37602 338 21 . . . 37602 339 1 The the DT 37602 339 2 ventral ventral JJ 37602 339 3 surfaces surface NNS 37602 339 4 are be VBP 37602 339 5 pale pale JJ 37602 339 6 creamy creamy JJ 37602 339 7 tan tan NN 37602 339 8 with with IN 37602 339 9 or or CC 37602 339 10 without without IN 37602 339 11 diffuse diffuse NN 37602 339 12 brown brown JJ 37602 339 13 spots spot NNS 37602 339 14 on on IN 37602 339 15 the the DT 37602 339 16 throat throat NN 37602 339 17 and and CC 37602 339 18 chest chest NN 37602 339 19 . . . 37602 340 1 The the DT 37602 340 2 holotype holotype NN 37602 340 3 of of IN 37602 340 4 _ _ NNP 37602 340 5 Hyla Hyla NNP 37602 340 6 festae festae VBD 37602 340 7 _ _ NNP 37602 340 8 is be VBZ 37602 340 9 a a DT 37602 340 10 female female NN 37602 340 11 having have VBG 37602 340 12 a a DT 37602 340 13 snout snout NN 37602 340 14 - - HYPH 37602 340 15 vent vent NN 37602 340 16 length length NN 37602 340 17 of of IN 37602 340 18 75.0 75.0 CD 37602 340 19 mm mm NNS 37602 340 20 ; ; : 37602 340 21 the the DT 37602 340 22 diameter diameter NN 37602 340 23 of of IN 37602 340 24 the the DT 37602 340 25 tympanum tympanum NN 37602 340 26 is be VBZ 37602 340 27 3.9 3.9 CD 37602 340 28 mm mm CD 37602 340 29 , , , 37602 340 30 57.4 57.4 CD 37602 340 31 percent percent NN 37602 340 32 of of IN 37602 340 33 the the DT 37602 340 34 diameter diameter NN 37602 340 35 of of IN 37602 340 36 the the DT 37602 340 37 eye eye NN 37602 340 38 . . . 37602 341 1 The the DT 37602 341 2 skin skin NN 37602 341 3 is be VBZ 37602 341 4 smooth smooth JJ 37602 341 5 on on IN 37602 341 6 the the DT 37602 341 7 dorsum dorsum NN 37602 341 8 and and CC 37602 341 9 areolate areolate NN 37602 341 10 on on IN 37602 341 11 the the DT 37602 341 12 anterior anterior JJ 37602 341 13 part part NN 37602 341 14 of of IN 37602 341 15 the the DT 37602 341 16 flanks flank NNS 37602 341 17 . . . 37602 342 1 The the DT 37602 342 2 dorsum dorsum NN 37602 342 3 is be VBZ 37602 342 4 pale pale JJ 37602 342 5 brown brown JJ 37602 342 6 with with IN 37602 342 7 a a DT 37602 342 8 large large JJ 37602 342 9 median median JJ 37602 342 10 longitudinal longitudinal JJ 37602 342 11 dark dark JJ 37602 342 12 brown brown JJ 37602 342 13 blotch blotch NN 37602 342 14 on on IN 37602 342 15 the the DT 37602 342 16 back back JJ 37602 342 17 and and CC 37602 342 18 broad broad JJ 37602 342 19 transverse transverse JJ 37602 342 20 bars bar NNS 37602 342 21 , , , 37602 342 22 narrowly narrowly RB 37602 342 23 outlined outline VBN 37602 342 24 by by IN 37602 342 25 cream cream NN 37602 342 26 , , , 37602 342 27 on on IN 37602 342 28 the the DT 37602 342 29 limbs limb NNS 37602 342 30 . . . 37602 343 1 Dark dark JJ 37602 343 2 brown brown JJ 37602 343 3 spots spot NNS 37602 343 4 are be VBP 37602 343 5 present present JJ 37602 343 6 on on IN 37602 343 7 the the DT 37602 343 8 flanks flank NNS 37602 343 9 ; ; : 37602 343 10 the the DT 37602 343 11 posterior posterior NN 37602 343 12 surfaces surface NNS 37602 343 13 of of IN 37602 343 14 the the DT 37602 343 15 thighs thigh NNS 37602 343 16 are be VBP 37602 343 17 dark dark JJ 37602 343 18 brown brown JJ 37602 343 19 . . . 37602 344 1 The the DT 37602 344 2 throat throat NN 37602 344 3 and and CC 37602 344 4 belly belly NN 37602 344 5 are be VBP 37602 344 6 grayish grayish NN 37602 344 7 white white JJ 37602 344 8 with with IN 37602 344 9 irregular irregular JJ 37602 344 10 dark dark JJ 37602 344 11 brown brown JJ 37602 344 12 spots spot NNS 37602 344 13 . . . 37602 345 1 The the DT 37602 345 2 holotype holotype NN 37602 345 3 of of IN 37602 345 4 _ _ NNP 37602 345 5 Hyla Hyla NNP 37602 345 6 carri carri VBD 37602 345 7 _ _ NNP 37602 345 8 is be VBZ 37602 345 9 a a DT 37602 345 10 female female NN 37602 345 11 having have VBG 37602 345 12 a a DT 37602 345 13 snout snout NN 37602 345 14 - - HYPH 37602 345 15 vent vent NN 37602 345 16 length length NN 37602 345 17 of of IN 37602 345 18 66.1 66.1 CD 37602 345 19 mm mm NNS 37602 345 20 ; ; : 37602 345 21 the the DT 37602 345 22 diameter diameter NN 37602 345 23 of of IN 37602 345 24 the the DT 37602 345 25 tympanum tympanum NN 37602 345 26 is be VBZ 37602 345 27 4.7 4.7 CD 37602 345 28 mm mm CD 37602 345 29 , , , 37602 345 30 81.0 81.0 CD 37602 345 31 percent percent NN 37602 345 32 of of IN 37602 345 33 the the DT 37602 345 34 diameter diameter NN 37602 345 35 of of IN 37602 345 36 the the DT 37602 345 37 eye eye NN 37602 345 38 . . . 37602 346 1 The the DT 37602 346 2 skin skin NN 37602 346 3 on on IN 37602 346 4 the the DT 37602 346 5 dorsum dorsum NN 37602 346 6 is be VBZ 37602 346 7 smooth smooth JJ 37602 346 8 with with IN 37602 346 9 scattered scatter VBN 37602 346 10 small small JJ 37602 346 11 tubercles tubercle NNS 37602 346 12 and and CC 37602 346 13 areolate areolate NN 37602 346 14 on on IN 37602 346 15 the the DT 37602 346 16 anterior anterior JJ 37602 346 17 part part NN 37602 346 18 of of IN 37602 346 19 the the DT 37602 346 20 flanks flank NNS 37602 346 21 . . . 37602 347 1 The the DT 37602 347 2 dorsum dorsum NN 37602 347 3 is be VBZ 37602 347 4 tan tan NNP 37602 347 5 with with IN 37602 347 6 irregular irregular JJ 37602 347 7 dark dark JJ 37602 347 8 brown brown JJ 37602 347 9 blotches blotch NNS 37602 347 10 on on IN 37602 347 11 the the DT 37602 347 12 back back NN 37602 347 13 and and CC 37602 347 14 transverse transverse JJ 37602 347 15 bars bar NNS 37602 347 16 on on IN 37602 347 17 the the DT 37602 347 18 limbs limb NNS 37602 347 19 ; ; : 37602 347 20 all all DT 37602 347 21 dorsal dorsal NN 37602 347 22 markings marking NNS 37602 347 23 are be VBP 37602 347 24 narrowly narrowly RB 37602 347 25 outlined outline VBN 37602 347 26 by by IN 37602 347 27 creamy creamy JJ 37602 347 28 white white NNP 37602 347 29 . . . 37602 348 1 Dark dark JJ 37602 348 2 brown brown JJ 37602 348 3 spots spot NNS 37602 348 4 are be VBP 37602 348 5 present present JJ 37602 348 6 on on IN 37602 348 7 the the DT 37602 348 8 flanks flank NNS 37602 348 9 ; ; : 37602 348 10 the the DT 37602 348 11 venter venter NN 37602 348 12 and and CC 37602 348 13 posterior posterior NN 37602 348 14 surfaces surface NNS 37602 348 15 of of IN 37602 348 16 the the DT 37602 348 17 thighs thigh NNS 37602 348 18 are be VBP 37602 348 19 tan tan JJ 37602 348 20 without without IN 37602 348 21 dark dark JJ 37602 348 22 spots spot NNS 37602 348 23 . . . 37602 349 1 The the DT 37602 349 2 holotype holotype NN 37602 349 3 of of IN 37602 349 4 _ _ NNP 37602 349 5 Hyla Hyla NNP 37602 349 6 cabrerai cabrerai NN 37602 349 7 _ _ NNP 37602 349 8 is be VBZ 37602 349 9 a a DT 37602 349 10 female female NN 37602 349 11 having have VBG 37602 349 12 a a DT 37602 349 13 snout snout NN 37602 349 14 - - HYPH 37602 349 15 vent vent NN 37602 349 16 length length NN 37602 349 17 of of IN 37602 349 18 52.7 52.7 CD 37602 349 19 mm mm NNS 37602 349 20 ; ; : 37602 349 21 the the DT 37602 349 22 diameter diameter NN 37602 349 23 of of IN 37602 349 24 the the DT 37602 349 25 tympanum tympanum NN 37602 349 26 is be VBZ 37602 349 27 4.0 4.0 CD 37602 349 28 mm mm NNS 37602 349 29 , , , 37602 349 30 76.9 76.9 CD 37602 349 31 percent percent NN 37602 349 32 of of IN 37602 349 33 the the DT 37602 349 34 diameter diameter NN 37602 349 35 of of IN 37602 349 36 the the DT 37602 349 37 eye eye NN 37602 349 38 . . . 37602 350 1 The the DT 37602 350 2 skin skin NN 37602 350 3 on on IN 37602 350 4 the the DT 37602 350 5 dorsum dorsum NN 37602 350 6 is be VBZ 37602 350 7 weakly weakly IN 37602 350 8 tuberculate tuberculate NN 37602 350 9 and and CC 37602 350 10 that that IN 37602 350 11 on on IN 37602 350 12 the the DT 37602 350 13 anterior anterior JJ 37602 350 14 part part NN 37602 350 15 of of IN 37602 350 16 the the DT 37602 350 17 flanks flank NNS 37602 350 18 is be VBZ 37602 350 19 areolate areolate NN 37602 350 20 . . . 37602 351 1 The the DT 37602 351 2 dorsum dorsum NN 37602 351 3 is be VBZ 37602 351 4 creamy creamy JJ 37602 351 5 tan tan NNP 37602 351 6 with with IN 37602 351 7 dark dark JJ 37602 351 8 brown brown JJ 37602 351 9 markings marking NNS 37602 351 10 ( ( -LRB- 37602 351 11 interorbital interorbital NNP 37602 351 12 bar bar NN 37602 351 13 , , , 37602 351 14 reticulations reticulation NNS 37602 351 15 on on IN 37602 351 16 occiput occiput NN 37602 351 17 , , , 37602 351 18 three three CD 37602 351 19 longitudinal longitudinal JJ 37602 351 20 streaks streak NNS 37602 351 21 on on IN 37602 351 22 back back NN 37602 351 23 , , , 37602 351 24 and and CC 37602 351 25 broad broad JJ 37602 351 26 transverse transverse JJ 37602 351 27 bars bar NNS 37602 351 28 on on IN 37602 351 29 limbs limb NNS 37602 351 30 ) ) -RRB- 37602 351 31 . . . 37602 352 1 Irregular irregular JJ 37602 352 2 dark dark JJ 37602 352 3 brown brown JJ 37602 352 4 spots spot NNS 37602 352 5 are be VBP 37602 352 6 present present JJ 37602 352 7 on on IN 37602 352 8 the the DT 37602 352 9 flanks flank NNS 37602 352 10 . . . 37602 353 1 The the DT 37602 353 2 venter venter NN 37602 353 3 is be VBZ 37602 353 4 pinkish pinkish NNP 37602 353 5 tan tan NNP 37602 353 6 with with IN 37602 353 7 small small JJ 37602 353 8 reddish reddish JJ 37602 353 9 brown brown JJ 37602 353 10 spots spot NNS 37602 353 11 on on IN 37602 353 12 the the DT 37602 353 13 throat throat NN 37602 353 14 and and CC 37602 353 15 darker dark JJR 37602 353 16 brown brown JJ 37602 353 17 spots spot NNS 37602 353 18 on on IN 37602 353 19 the the DT 37602 353 20 chest chest NN 37602 353 21 and and CC 37602 353 22 belly belly NN 37602 353 23 . . . 37602 354 1 In in IN 37602 354 2 their -PRON- PRP$ 37602 354 3 description description NN 37602 354 4 of of IN 37602 354 5 _ _ NNP 37602 354 6 Hyla Hyla NNP 37602 354 7 cabrerai cabrerai NN 37602 354 8 _ _ NNP 37602 354 9 , , , 37602 354 10 Cochran Cochran NNP 37602 354 11 and and CC 37602 354 12 Goin go NNP 37602 354 13 ( ( -LRB- 37602 354 14 1970:217 1970:217 CD 37602 354 15 ) ) -RRB- 37602 354 16 stated state VBD 37602 354 17 : : : 37602 354 18 " " `` 37602 354 19 This this DT 37602 354 20 species species NN 37602 354 21 , , , 37602 354 22 together together RB 37602 354 23 with with IN 37602 354 24 _ _ NNP 37602 354 25 buckleyi buckleyi NN 37602 354 26 _ _ NNP 37602 354 27 and and CC 37602 354 28 _ _ NNP 37602 354 29 pearsoni pearsoni NN 37602 354 30 _ _ NNP 37602 354 31 certainly certainly RB 37602 354 32 make make VBP 37602 354 33 a a DT 37602 354 34 closely closely RB 37602 354 35 knit knit VBN 37602 354 36 group group NN 37602 354 37 .... .... . 37602 355 1 Both both DT 37602 355 2 _ _ NNP 37602 355 3 buckleyi buckleyi NN 37602 355 4 _ _ NNP 37602 355 5 and and CC 37602 355 6 _ _ NNP 37602 355 7 cabrerai cabrerai NN 37602 355 8 _ _ NNP 37602 355 9 have have VBP 37602 355 10 long long JJ 37602 355 11 hind hind NN 37602 355 12 legs leg NNS 37602 355 13 , , , 37602 355 14 with with IN 37602 355 15 the the DT 37602 355 16 extended extended JJ 37602 355 17 heel heel NN 37602 355 18 reaching reach VBG 37602 355 19 to to IN 37602 355 20 the the DT 37602 355 21 tip tip NN 37602 355 22 of of IN 37602 355 23 the the DT 37602 355 24 snout snout NN 37602 355 25 , , , 37602 355 26 while while IN 37602 355 27 in in IN 37602 355 28 _ _ NNP 37602 355 29 pearsoni pearsoni NNP 37602 355 30 _ _ NNP 37602 355 31 the the DT 37602 355 32 extended extended JJ 37602 355 33 heel heel NN 37602 355 34 reaches reach VBZ 37602 355 35 only only RB 37602 355 36 to to IN 37602 355 37 the the DT 37602 355 38 eye eye NN 37602 355 39 . . . 37602 356 1 _ _ NNP 37602 356 2 H. H. NNP 37602 357 1 buckleyi buckleyi NNP 37602 357 2 _ _ NNP 37602 357 3 has have VBZ 37602 357 4 the the DT 37602 357 5 belly belly NN 37602 357 6 dusky dusky JJ 37602 357 7 , , , 37602 357 8 while while IN 37602 357 9 it -PRON- PRP 37602 357 10 is be VBZ 37602 357 11 heavily heavily RB 37602 357 12 spotted spot VBN 37602 357 13 in in IN 37602 357 14 _ _ NNP 37602 357 15 cabrerai cabrerai NN 37602 357 16 _ _ NNP 37602 357 17 and and CC 37602 357 18 is be VBZ 37602 357 19 reticulated reticulate VBN 37602 357 20 in in IN 37602 357 21 _ _ NNP 37602 357 22 pearsoni pearsoni NNP 37602 357 23 _ _ NNP 37602 357 24 . . . 37602 358 1 _ _ NNP 37602 358 2 H. H. NNP 37602 359 1 cabrerai cabrerai NNP 37602 359 2 _ _ NNP 37602 359 3 seems seem VBZ 37602 359 4 to to TO 37602 359 5 have have VB 37602 359 6 the the DT 37602 359 7 heaviest heavy JJS 37602 359 8 hands hand NNS 37602 359 9 with with IN 37602 359 10 the the DT 37602 359 11 most most JJS 37602 359 12 webbing webbing NN 37602 359 13 between between IN 37602 359 14 the the DT 37602 359 15 fingers finger NNS 37602 359 16 ; ; : 37602 359 17 the the DT 37602 359 18 other other JJ 37602 359 19 two two CD 37602 359 20 species specie NNS 37602 359 21 have have VBP 37602 359 22 the the DT 37602 359 23 webbing webbing NN 37602 359 24 reduced reduce VBN 37602 359 25 between between IN 37602 359 26 the the DT 37602 359 27 fingers finger NNS 37602 359 28 . . . 37602 359 29 " " '' 37602 360 1 The the DT 37602 360 2 description description NN 37602 360 3 of of IN 37602 360 4 _ _ NNP 37602 360 5 Hyla Hyla NNP 37602 360 6 cabrerai cabrerai NN 37602 360 7 _ _ NNP 37602 360 8 was be VBD 37602 360 9 based base VBN 37602 360 10 on on IN 37602 360 11 three three CD 37602 360 12 specimens specimen NNS 37602 360 13 . . . 37602 361 1 We -PRON- PRP 37602 361 2 have have VBP 37602 361 3 examined examine VBN 37602 361 4 the the DT 37602 361 5 holotype holotype NN 37602 361 6 and and CC 37602 361 7 one one CD 37602 361 8 paratype paratype NN 37602 361 9 ( ( -LRB- 37602 361 10 WCAB wcab NN 37602 361 11 13284 13284 CD 37602 361 12 from from IN 37602 361 13 Territorio Territorio NNP 37602 361 14 do do VBP 37602 361 15 Amapá Amapá NNP 37602 361 16 , , , 37602 361 17 Brasil Brasil NNP 37602 361 18 ) ) -RRB- 37602 361 19 . . . 37602 362 1 Another another DT 37602 362 2 paratype paratype NN 37602 362 3 in in IN 37602 362 4 the the DT 37602 362 5 private private JJ 37602 362 6 collection collection NN 37602 362 7 of of IN 37602 362 8 C. C. NNP 37602 362 9 J. J. NNP 37602 362 10 Goin go NNP 37602 362 11 from from IN 37602 362 12 Caño Caño NNP 37602 362 13 Tuí Tuí NNP 37602 362 14 , , , 37602 362 15 between between IN 37602 362 16 Mitú Mitú NNP 37602 362 17 and and CC 37602 362 18 Raudal Raudal NNP 37602 362 19 de de IN 37602 362 20 Yurupari Yurupari NNP 37602 362 21 , , , 37602 362 22 Comisaria Comisaria NNP 37602 362 23 de de NNP 37602 362 24 Vaupés Vaupés NNP 37602 362 25 , , , 37602 362 26 Colombia Colombia NNP 37602 362 27 , , , 37602 362 28 was be VBD 37602 362 29 not not RB 37602 362 30 examined examine VBN 37602 362 31 . . . 37602 363 1 Cochran Cochran NNP 37602 363 2 and and CC 37602 363 3 Goin go NNP 37602 363 4 ( ( -LRB- 37602 363 5 1970:211 1970:211 CD 37602 363 6 ) ) -RRB- 37602 363 7 based base VBN 37602 363 8 their -PRON- PRP$ 37602 363 9 description description NN 37602 363 10 of of IN 37602 363 11 _ _ NNP 37602 363 12 Hyla Hyla NNP 37602 363 13 carri carri VBD 37602 363 14 _ _ NNP 37602 363 15 on on IN 37602 363 16 one one CD 37602 363 17 gravid gravid JJ 37602 363 18 female female NN 37602 363 19 and and CC 37602 363 20 stated state VBD 37602 363 21 : : : 37602 363 22 " " `` 37602 363 23 A a DT 37602 363 24 large large JJ 37602 363 25 _ _ NNP 37602 363 26 Hyla Hyla NNP 37602 363 27 _ _ NNP 37602 363 28 with with IN 37602 363 29 the the DT 37602 363 30 vomerine vomerine NN 37602 363 31 teeth tooth NNS 37602 363 32 in in IN 37602 363 33 two two CD 37602 363 34 ^^ ^^ VBD 37602 363 35 shaped shaped JJ 37602 363 36 patches patch NNS 37602 363 37 between between IN 37602 363 38 the the DT 37602 363 39 somewhat somewhat RB 37602 363 40 squarish squarish JJ 37602 363 41 choanae choanae NN 37602 363 42 ; ; , 37602 363 43 reduced reduce VBN 37602 363 44 webs webs NN 37602 363 45 between between IN 37602 363 46 the the DT 37602 363 47 fingers finger NNS 37602 363 48 ; ; : 37602 363 49 and and CC 37602 363 50 a a DT 37602 363 51 pattern pattern NN 37602 363 52 of of IN 37602 363 53 dorsal dorsal NN 37602 363 54 dark dark JJ 37602 363 55 blotches blotch NNS 37602 363 56 bordered border VBN 37602 363 57 by by IN 37602 363 58 light light JJ 37602 363 59 margins margin NNS 37602 363 60 . . . 37602 364 1 The the DT 37602 364 2 species species NN 37602 364 3 is be VBZ 37602 364 4 not not RB 37602 364 5 similar similar JJ 37602 364 6 to to IN 37602 364 7 any any DT 37602 364 8 other other JJ 37602 364 9 species specie NNS 37602 364 10 known know VBN 37602 364 11 in in IN 37602 364 12 Colombia Colombia NNP 37602 364 13 . . . 37602 365 1 It -PRON- PRP 37602 365 2 is be VBZ 37602 365 3 perhaps perhaps RB 37602 365 4 most most RBS 37602 365 5 closely closely RB 37602 365 6 related relate VBN 37602 365 7 to to IN 37602 365 8 _ _ NNP 37602 365 9 Hyla Hyla NNP 37602 365 10 claresignata claresignata NN 37602 365 11 _ _ NNP 37602 365 12 of of IN 37602 365 13 Brazil Brazil NNP 37602 365 14 , , , 37602 365 15 from from IN 37602 365 16 which which WDT 37602 365 17 it -PRON- PRP 37602 365 18 can can MD 37602 365 19 be be VB 37602 365 20 differentiated differentiate VBN 37602 365 21 by by IN 37602 365 22 its -PRON- PRP$ 37602 365 23 more more RBR 37602 365 24 heavily heavily RB 37602 365 25 spotted spot VBN 37602 365 26 dorsum dorsum NN 37602 365 27 , , , 37602 365 28 larger large JJR 37602 365 29 tympanum tympanum NN 37602 365 30 , , , 37602 365 31 and and CC 37602 365 32 lack lack NN 37602 365 33 of of IN 37602 365 34 dark dark JJ 37602 365 35 anal anal JJ 37602 365 36 spots spot NNS 37602 365 37 . . . 37602 365 38 " " '' 37602 366 1 Except except IN 37602 366 2 for for IN 37602 366 3 the the DT 37602 366 4 inclusion inclusion NN 37602 366 5 of of IN 37602 366 6 the the DT 37602 366 7 name name NN 37602 366 8 in in IN 37602 366 9 checklists checklists NNP 37602 366 10 , , , 37602 366 11 _ _ NNP 37602 366 12 Hyla Hyla NNP 37602 366 13 festae festae VBD 37602 366 14 _ _ NNP 37602 366 15 has have VBZ 37602 366 16 not not RB 37602 366 17 been be VBN 37602 366 18 mentioned mention VBN 37602 366 19 in in IN 37602 366 20 the the DT 37602 366 21 literature literature NN 37602 366 22 since since IN 37602 366 23 the the DT 37602 366 24 original original JJ 37602 366 25 description description NN 37602 366 26 . . . 37602 367 1 The the DT 37602 367 2 wholesale wholesale JJ 37602 367 3 synonymization synonymization NN 37602 367 4 of of IN 37602 367 5 names name NNS 37602 367 6 , , , 37602 367 7 which which WDT 37602 367 8 , , , 37602 367 9 on on IN 37602 367 10 the the DT 37602 367 11 bases basis NNS 37602 367 12 of of IN 37602 367 13 their -PRON- PRP$ 37602 367 14 published publish VBN 37602 367 15 diagnoses diagnosis NNS 37602 367 16 , , , 37602 367 17 seem seem VBP 37602 367 18 to to TO 37602 367 19 apply apply VB 37602 367 20 to to IN 37602 367 21 distinctly distinctly RB 37602 367 22 different different JJ 37602 367 23 species specie NNS 37602 367 24 , , , 37602 367 25 is be VBZ 37602 367 26 possible possible JJ 37602 367 27 with with IN 37602 367 28 the the DT 37602 367 29 application application NN 37602 367 30 of of IN 37602 367 31 uniform uniform JJ 37602 367 32 criteria criterion NNS 37602 367 33 to to IN 37602 367 34 the the DT 37602 367 35 types type NNS 37602 367 36 and and CC 37602 367 37 series series NN 37602 367 38 of of IN 37602 367 39 other other JJ 37602 367 40 specimens specimen NNS 37602 367 41 . . . 37602 368 1 In in IN 37602 368 2 measurements measurement NNS 37602 368 3 and and CC 37602 368 4 proportions proportion NNS 37602 368 5 the the DT 37602 368 6 type type NN 37602 368 7 specimens specimen NNS 37602 368 8 of of IN 37602 368 9 the the DT 37602 368 10 nominal nominal JJ 37602 368 11 taxa taxa NN 37602 368 12 all all DT 37602 368 13 fall fall VBP 37602 368 14 within within IN 37602 368 15 the the DT 37602 368 16 range range NN 37602 368 17 of of IN 37602 368 18 variation variation NN 37602 368 19 exhibited exhibit VBN 37602 368 20 by by IN 37602 368 21 a a DT 37602 368 22 series series NN 37602 368 23 of of IN 37602 368 24 18 18 CD 37602 368 25 males male NNS 37602 368 26 and and CC 37602 368 27 15 15 CD 37602 368 28 females female NNS 37602 368 29 from from IN 37602 368 30 Provincia Provincia NNP 37602 368 31 Pastaza Pastaza NNP 37602 368 32 , , , 37602 368 33 Ecuador Ecuador NNP 37602 368 34 , , , 37602 368 35 except except IN 37602 368 36 the the DT 37602 368 37 ratio ratio NN 37602 368 38 of of IN 37602 368 39 the the DT 37602 368 40 diameter diameter NN 37602 368 41 of of IN 37602 368 42 the the DT 37602 368 43 tympanum tympanum NN 37602 368 44 to to IN 37602 368 45 that that DT 37602 368 46 of of IN 37602 368 47 the the DT 37602 368 48 eye eye NN 37602 368 49 in in IN 37602 368 50 the the DT 37602 368 51 female female JJ 37602 368 52 holotype holotype NN 37602 368 53 of of IN 37602 368 54 _ _ NNP 37602 368 55 Hyla Hyla NNP 37602 368 56 festae festae VBD 37602 368 57 _ _ NNP 37602 368 58 . . . 37602 369 1 In in IN 37602 369 2 that that DT 37602 369 3 specimen specimen VBZ 37602 369 4 the the DT 37602 369 5 ratio ratio NN 37602 369 6 is be VBZ 37602 369 7 0.574 0.574 CD 37602 369 8 , , , 37602 369 9 whereas whereas IN 37602 369 10 the the DT 37602 369 11 ratio ratio NN 37602 369 12 in in IN 37602 369 13 the the DT 37602 369 14 15 15 CD 37602 369 15 females female NNS 37602 369 16 from from IN 37602 369 17 Provincia Provincia NNP 37602 369 18 Pastaza Pastaza NNP 37602 369 19 is be VBZ 37602 369 20 0.587 0.587 CD 37602 369 21 - - SYM 37602 369 22 0.905 0.905 CD 37602 369 23 ( ( -LRB- 37602 369 24 mean mean NNP 37602 369 25 0.736 0.736 CD 37602 369 26 ) ) -RRB- 37602 369 27 . . . 37602 370 1 Ventral ventral JJ 37602 370 2 coloration coloration NN 37602 370 3 is be VBZ 37602 370 4 the the DT 37602 370 5 most most RBS 37602 370 6 variable variable JJ 37602 370 7 character character NN 37602 370 8 among among IN 37602 370 9 the the DT 37602 370 10 types type NNS 37602 370 11 . . . 37602 371 1 The the DT 37602 371 2 venter venter NN 37602 371 3 in in IN 37602 371 4 the the DT 37602 371 5 type type NN 37602 371 6 of of IN 37602 371 7 _ _ NNP 37602 371 8 Hyla Hyla NNP 37602 371 9 festae festae VBD 37602 371 10 _ _ NNP 37602 371 11 is be VBZ 37602 371 12 boldly boldly RB 37602 371 13 spotted spot VBN 37602 371 14 ; ; : 37602 371 15 it -PRON- PRP 37602 371 16 is be VBZ 37602 371 17 distinctly distinctly RB 37602 371 18 spotted spot VBN 37602 371 19 in in IN 37602 371 20 _ _ NNP 37602 371 21 cabrerai cabrerai NN 37602 371 22 _ _ NNP 37602 371 23 , , , 37602 371 24 uniform uniform JJ 37602 371 25 tan tan NNP 37602 371 26 in in IN 37602 371 27 _ _ NNP 37602 371 28 carri carri NNP 37602 371 29 _ _ NNP 37602 371 30 , , , 37602 371 31 and and CC 37602 371 32 tan tan NNP 37602 371 33 , , , 37602 371 34 flecked fleck VBN 37602 371 35 , , , 37602 371 36 or or CC 37602 371 37 spotted spot VBN 37602 371 38 in in IN 37602 371 39 the the DT 37602 371 40 type type NN 37602 371 41 series series NN 37602 371 42 of of IN 37602 371 43 _ _ NNP 37602 371 44 buckleyi buckleyi NN 37602 371 45 _ _ NNP 37602 371 46 . . . 37602 372 1 The the DT 37602 372 2 ventral ventral JJ 37602 372 3 coloration coloration NN 37602 372 4 in in IN 37602 372 5 series series NN 37602 372 6 of of IN 37602 372 7 specimens specimen NNS 37602 372 8 from from IN 37602 372 9 Amazonian Amazonian NNP 37602 372 10 Ecuador Ecuador NNP 37602 372 11 encompasses encompass VBZ 37602 372 12 that that IN 37602 372 13 observed observe VBD 37602 372 14 in in IN 37602 372 15 all all DT 37602 372 16 of of IN 37602 372 17 the the DT 37602 372 18 types type NNS 37602 372 19 , , , 37602 372 20 except except IN 37602 372 21 that that DT 37602 372 22 of of IN 37602 372 23 _ _ NNP 37602 372 24 festae festae NNP 37602 372 25 _ _ NNP 37602 372 26 , , , 37602 372 27 which which WDT 37602 372 28 has have VBZ 37602 372 29 more more JJR 37602 372 30 ventral ventral JJ 37602 372 31 spotting spotting NN 37602 372 32 than than IN 37602 372 33 any any DT 37602 372 34 other other JJ 37602 372 35 individual individual NN 37602 372 36 . . . 37602 373 1 The the DT 37602 373 2 webbing webbing NN 37602 373 3 on on IN 37602 373 4 the the DT 37602 373 5 hand hand NN 37602 373 6 usually usually RB 37602 373 7 excludes exclude VBZ 37602 373 8 the the DT 37602 373 9 penultimate penultimate JJ 37602 373 10 phalanges phalanx NNS 37602 373 11 of of IN 37602 373 12 the the DT 37602 373 13 fingers finger NNS 37602 373 14 , , , 37602 373 15 but but CC 37602 373 16 in in IN 37602 373 17 some some DT 37602 373 18 specimens specimen NNS 37602 373 19 from from IN 37602 373 20 Amazonian Amazonian NNP 37602 373 21 Ecuador Ecuador NNP 37602 373 22 the the DT 37602 373 23 webbing webbing NN 37602 373 24 encompasses encompass VBZ 37602 373 25 the the DT 37602 373 26 proximal proximal JJ 37602 373 27 parts part NNS 37602 373 28 of of IN 37602 373 29 the the DT 37602 373 30 penultimate penultimate JJ 37602 373 31 phalanges phalanx NNS 37602 373 32 of of IN 37602 373 33 the the DT 37602 373 34 fingers finger NNS 37602 373 35 . . . 37602 374 1 In in IN 37602 374 2 a a DT 37602 374 3 few few JJ 37602 374 4 of of IN 37602 374 5 these these DT 37602 374 6 specimens specimen NNS 37602 374 7 , , , 37602 374 8 the the DT 37602 374 9 holotype holotype NN 37602 374 10 of of IN 37602 374 11 _ _ NNP 37602 374 12 festae festae NNP 37602 374 13 _ _ NNP 37602 374 14 , , , 37602 374 15 and and CC 37602 374 16 one one CD 37602 374 17 paratype paratype NN 37602 374 18 of of IN 37602 374 19 _ _ NNP 37602 374 20 cabrerai cabrerai NN 37602 374 21 _ _ NNP 37602 374 22 the the DT 37602 374 23 webbing webbing NN 37602 374 24 extends extend VBZ 37602 374 25 to to IN 37602 374 26 the the DT 37602 374 27 middle middle NN 37602 374 28 of of IN 37602 374 29 the the DT 37602 374 30 penultimate penultimate JJ 37602 374 31 phalanges phalanx NNS 37602 374 32 of of IN 37602 374 33 the the DT 37602 374 34 third third JJ 37602 374 35 and and CC 37602 374 36 fourth fourth JJ 37602 374 37 fingers finger NNS 37602 374 38 . . . 37602 375 1 In in IN 37602 375 2 the the DT 37602 375 3 holotype holotype NN 37602 375 4 of of IN 37602 375 5 _ _ NNP 37602 375 6 cabrerai cabrerai NN 37602 375 7 _ _ NNP 37602 375 8 the the DT 37602 375 9 webbing webbing NN 37602 375 10 extends extend VBZ 37602 375 11 to to IN 37602 375 12 the the DT 37602 375 13 middle middle NN 37602 375 14 of of IN 37602 375 15 the the DT 37602 375 16 penultimate penultimate JJ 37602 375 17 phalanges phalanx NNS 37602 375 18 of of IN 37602 375 19 the the DT 37602 375 20 third third JJ 37602 375 21 and and CC 37602 375 22 fourth fourth JJ 37602 375 23 fingers finger NNS 37602 375 24 and and CC 37602 375 25 to to IN 37602 375 26 the the DT 37602 375 27 base base NN 37602 375 28 of of IN 37602 375 29 the the DT 37602 375 30 disc disc NN 37602 375 31 of of IN 37602 375 32 the the DT 37602 375 33 second second JJ 37602 375 34 finger finger NN 37602 375 35 . . . 37602 376 1 The the DT 37602 376 2 types type NNS 37602 376 3 of of IN 37602 376 4 the the DT 37602 376 5 nominal nominal JJ 37602 376 6 taxa taxa NNP 37602 376 7 and and CC 37602 376 8 series series NN 37602 376 9 of of IN 37602 376 10 specimens specimen NNS 37602 376 11 from from IN 37602 376 12 Guyana Guyana NNP 37602 376 13 and and CC 37602 376 14 Amazonian Amazonian NNP 37602 376 15 Ecuador Ecuador NNP 37602 376 16 display display NN 37602 376 17 noticeable noticeable JJ 37602 376 18 variation variation NN 37602 376 19 in in IN 37602 376 20 dorsal dorsal NN 37602 376 21 coloration coloration NN 37602 376 22 . . . 37602 377 1 The the DT 37602 377 2 variety variety NN 37602 377 3 of of IN 37602 377 4 dorsal dorsal NN 37602 377 5 patterns pattern NNS 37602 377 6 of of IN 37602 377 7 all all DT 37602 377 8 of of IN 37602 377 9 the the DT 37602 377 10 types type NNS 37602 377 11 is be VBZ 37602 377 12 included include VBN 37602 377 13 in in IN 37602 377 14 the the DT 37602 377 15 variation variation NN 37602 377 16 displayed display VBN 37602 377 17 by by IN 37602 377 18 the the DT 37602 377 19 other other JJ 37602 377 20 specimens specimen NNS 37602 377 21 . . . 37602 378 1 All all DT 37602 378 2 specimens specimen NNS 37602 378 3 have have VBP 37602 378 4 some some DT 37602 378 5 amount amount NN 37602 378 6 of of IN 37602 378 7 dark dark JJ 37602 378 8 spotting spotting NN 37602 378 9 on on IN 37602 378 10 the the DT 37602 378 11 flanks flank NNS 37602 378 12 ; ; : 37602 378 13 all all DT 37602 378 14 have have VBP 37602 378 15 vertically vertically RB 37602 378 16 barred bar VBN 37602 378 17 lips lip NNS 37602 378 18 , , , 37602 378 19 on on IN 37602 378 20 which which WDT 37602 378 21 a a DT 37602 378 22 pale pale JJ 37602 378 23 subocular subocular JJ 37602 378 24 spot spot NN 37602 378 25 usually usually RB 37602 378 26 is be VBZ 37602 378 27 evident evident JJ 37602 378 28 . . . 37602 379 1 Probably probably RB 37602 379 2 the the DT 37602 379 3 most most RBS 37602 379 4 unifying unifying JJ 37602 379 5 physical physical JJ 37602 379 6 characteristic characteristic NN 37602 379 7 of of IN 37602 379 8 all all DT 37602 379 9 of of IN 37602 379 10 the the DT 37602 379 11 specimens specimen NNS 37602 379 12 is be VBZ 37602 379 13 the the DT 37602 379 14 nature nature NN 37602 379 15 of of IN 37602 379 16 the the DT 37602 379 17 skin skin NN 37602 379 18 on on IN 37602 379 19 the the DT 37602 379 20 anterior anterior JJ 37602 379 21 part part NN 37602 379 22 of of IN 37602 379 23 the the DT 37602 379 24 flank flank NN 37602 379 25 . . . 37602 380 1 The the DT 37602 380 2 skin skin NN 37602 380 3 is be VBZ 37602 380 4 elevated elevate VBN 37602 380 5 amidst amidst IN 37602 380 6 an an DT 37602 380 7 irregular irregular JJ 37602 380 8 network network NN 37602 380 9 of of IN 37602 380 10 depressions depression NNS 37602 380 11 . . . 37602 381 1 This this DT 37602 381 2 areolate areolate NN 37602 381 3 dermal dermal JJ 37602 381 4 condition condition NN 37602 381 5 is be VBZ 37602 381 6 present present JJ 37602 381 7 in in IN 37602 381 8 all all DT 37602 381 9 specimens specimen NNS 37602 381 10 and and CC 37602 381 11 does do VBZ 37602 381 12 not not RB 37602 381 13 occur occur VB 37602 381 14 in in IN 37602 381 15 other other JJ 37602 381 16 species specie NNS 37602 381 17 of of IN 37602 381 18 _ _ NNP 37602 381 19 Osteocephalus Osteocephalus NNP 37602 381 20 _ _ NNP 37602 381 21 . . . 37602 382 1 The the DT 37602 382 2 degree degree NN 37602 382 3 of of IN 37602 382 4 tubercularity tubercularity NN 37602 382 5 of of IN 37602 382 6 the the DT 37602 382 7 skin skin NN 37602 382 8 on on IN 37602 382 9 the the DT 37602 382 10 dorsum dorsum NN 37602 382 11 is be VBZ 37602 382 12 variable variable JJ 37602 382 13 and and CC 37602 382 14 sexually sexually RB 37602 382 15 dimorphic dimorphic JJ 37602 382 16 . . . 37602 383 1 All all DT 37602 383 2 males male NNS 37602 383 3 are be VBP 37602 383 4 tubercular tubercular JJ 37602 383 5 , , , 37602 383 6 whereas whereas IN 37602 383 7 small small JJ 37602 383 8 females female NNS 37602 383 9 are be VBP 37602 383 10 smooth smooth JJ 37602 383 11 or or CC 37602 383 12 have have VBP 37602 383 13 only only RB 37602 383 14 a a DT 37602 383 15 few few JJ 37602 383 16 scattered scatter VBN 37602 383 17 tubercles tubercle NNS 37602 383 18 . . . 37602 384 1 Large large JJ 37602 384 2 females female NNS 37602 384 3 usually usually RB 37602 384 4 have have VBP 37602 384 5 pronounced pronounce VBN 37602 384 6 tubercles tubercle NNS 37602 384 7 on on IN 37602 384 8 the the DT 37602 384 9 eyelids eyelid NNS 37602 384 10 and and CC 37602 384 11 supratympanic supratympanic NNP 37602 384 12 fold fold NNP 37602 384 13 . . . 37602 385 1 In in IN 37602 385 2 their -PRON- PRP$ 37602 385 3 description description NN 37602 385 4 of of IN 37602 385 5 _ _ NNP 37602 385 6 Hyla Hyla NNP 37602 385 7 carri carri VBD 37602 385 8 _ _ NNP 37602 385 9 , , , 37602 385 10 Cochran Cochran NNP 37602 385 11 and and CC 37602 385 12 Goin go NNP 37602 385 13 ( ( -LRB- 37602 385 14 1970:211 1970:211 CD 37602 385 15 ) ) -RRB- 37602 385 16 misrepresented misrepresent VBD 37602 385 17 the the DT 37602 385 18 nature nature NN 37602 385 19 of of IN 37602 385 20 the the DT 37602 385 21 dentigerous dentigerous JJ 37602 385 22 processes process NNS 37602 385 23 of of IN 37602 385 24 the the DT 37602 385 25 prevomers prevomer NNS 37602 385 26 , , , 37602 385 27 which which WDT 37602 385 28 are be VBP 37602 385 29 angular angular JJ 37602 385 30 , , , 37602 385 31 not not RB 37602 385 32 ^-shaped ^-shaped -RRB- 37602 385 33 . . . 37602 386 1 Their -PRON- PRP$ 37602 386 2 suggestion suggestion NN 37602 386 3 that that IN 37602 386 4 the the DT 37602 386 5 Colombian Colombian NNP 37602 386 6 _ _ NNP 37602 386 7 Hyla Hyla NNP 37602 386 8 carri carri VBD 37602 386 9 _ _ NNP 37602 386 10 is be VBZ 37602 386 11 related relate VBN 37602 386 12 to to IN 37602 386 13 _ _ NNP 37602 386 14 Hyla Hyla NNP 37602 386 15 claresignata claresignata NN 37602 386 16 _ _ NNP 37602 386 17 in in IN 37602 386 18 southeastern southeastern JJ 37602 386 19 Brasil Brasil NNP 37602 386 20 is be VBZ 37602 386 21 unfounded unfounded JJ 37602 386 22 . . . 37602 387 1 The the DT 37602 387 2 latter latter JJ 37602 387 3 species species NN 37602 387 4 is be VBZ 37602 387 5 smaller small JJR 37602 387 6 ( ( -LRB- 37602 387 7 40 40 CD 37602 387 8 mm mm CD 37602 387 9 ) ) -RRB- 37602 387 10 , , , 37602 387 11 has have VBZ 37602 387 12 a a DT 37602 387 13 yellow yellow JJ 37602 387 14 dorsum dorsum NN 37602 387 15 and and CC 37602 387 16 venter venter NN 37602 387 17 , , , 37602 387 18 dark dark JJ 37602 387 19 brown brown JJ 37602 387 20 spots spot NNS 37602 387 21 dorsolaterally dorsolaterally RB 37602 387 22 , , , 37602 387 23 oblique oblique JJ 37602 387 24 dentigerous dentigerous JJ 37602 387 25 processes process NNS 37602 387 26 of of IN 37602 387 27 the the DT 37602 387 28 prevomers prevomer NNS 37602 387 29 , , , 37602 387 30 small small JJ 37602 387 31 tympanum tympanum NN 37602 387 32 , , , 37602 387 33 and and CC 37602 387 34 smooth smooth JJ 37602 387 35 skin skin NN 37602 387 36 dorsally dorsally RB 37602 387 37 . . . 37602 388 1 The the DT 37602 388 2 ventral ventral JJ 37602 388 3 coloration coloration NN 37602 388 4 of of IN 37602 388 5 the the DT 37602 388 6 type type NN 37602 388 7 of of IN 37602 388 8 _ _ NNP 37602 388 9 Hyla Hyla NNP 37602 388 10 festae festae VBD 37602 388 11 _ _ NNP 37602 388 12 resembles resemble VBZ 37602 388 13 that that IN 37602 388 14 of of IN 37602 388 15 _ _ NNP 37602 388 16 Osteocephalus Osteocephalus NNP 37602 388 17 verrucigerus verrucigerus NN 37602 388 18 _ _ NNP 37602 388 19 , , , 37602 388 20 but but CC 37602 388 21 the the DT 37602 388 22 type type NN 37602 388 23 differs differ VBZ 37602 388 24 from from IN 37602 388 25 _ _ NNP 37602 388 26 verrucigerus verrucigerus NN 37602 388 27 _ _ NNP 37602 388 28 by by IN 37602 388 29 having have VBG 37602 388 30 areolate areolate NN 37602 388 31 skin skin NN 37602 388 32 on on IN 37602 388 33 the the DT 37602 388 34 flanks flank NNS 37602 388 35 and and CC 37602 388 36 distinct distinct JJ 37602 388 37 dark dark JJ 37602 388 38 markings marking NNS 37602 388 39 on on IN 37602 388 40 the the DT 37602 388 41 dorsum dorsum NN 37602 388 42 . . . 37602 389 1 In in IN 37602 389 2 _ _ NNP 37602 389 3 verrucigerus verrucigerus NN 37602 389 4 _ _ IN 37602 389 5 the the DT 37602 389 6 skin skin NN 37602 389 7 on on IN 37602 389 8 the the DT 37602 389 9 flanks flank NNS 37602 389 10 is be VBZ 37602 389 11 smooth smooth JJ 37602 389 12 , , , 37602 389 13 and and CC 37602 389 14 the the DT 37602 389 15 dorsum dorsum NN 37602 389 16 is be VBZ 37602 389 17 uniform uniform JJ 37602 389 18 dark dark JJ 37602 389 19 brown brown NNP 37602 389 20 , , , 37602 389 21 except except IN 37602 389 22 for for IN 37602 389 23 a a DT 37602 389 24 tan tan JJ 37602 389 25 snout snout NN 37602 389 26 in in IN 37602 389 27 females female NNS 37602 389 28 . . . 37602 390 1 Comparisons comparison NNS 37602 390 2 of of IN 37602 390 3 the the DT 37602 390 4 types type NNS 37602 390 5 of of IN 37602 390 6 the the DT 37602 390 7 nominal nominal JJ 37602 390 8 species specie NNS 37602 390 9 with with IN 37602 390 10 series series NN 37602 390 11 of of IN 37602 390 12 specimens specimen NNS 37602 390 13 from from IN 37602 390 14 Guyana Guyana NNP 37602 390 15 , , , 37602 390 16 Colombia Colombia NNP 37602 390 17 , , , 37602 390 18 Ecuador Ecuador NNP 37602 390 19 , , , 37602 390 20 and and CC 37602 390 21 Perú Perú NNP 37602 390 22 suggest suggest VBP 37602 390 23 strongly strongly RB 37602 390 24 that that IN 37602 390 25 the the DT 37602 390 26 types type NNS 37602 390 27 are be VBP 37602 390 28 representative representative JJ 37602 390 29 of of IN 37602 390 30 one one CD 37602 390 31 taxon taxon NNP 37602 390 32 , , , 37602 390 33 the the DT 37602 390 34 oldest old JJS 37602 390 35 name name NN 37602 390 36 for for IN 37602 390 37 which which WDT 37602 390 38 is be VBZ 37602 390 39 _ _ NNP 37602 390 40 Hyla Hyla NNP 37602 390 41 buckleyi buckleyi NN 37602 390 42 _ _ NNP 37602 390 43 Boulenger Boulenger NNP 37602 390 44 , , , 37602 390 45 1882 1882 CD 37602 390 46 . . . 37602 391 1 Consequently consequently RB 37602 391 2 , , , 37602 391 3 we -PRON- PRP 37602 391 4 place place VBP 37602 391 5 _ _ NNP 37602 391 6 Hyla Hyla NNP 37602 391 7 festae festae VBD 37602 391 8 _ _ NNP 37602 391 9 Peracca Peracca NNP 37602 391 10 , , , 37602 391 11 1904 1904 CD 37602 391 12 , , , 37602 391 13 _ _ NNP 37602 391 14 Hyla Hyla NNP 37602 391 15 carri carri VBD 37602 391 16 _ _ NNP 37602 391 17 Cochran Cochran NNP 37602 391 18 and and CC 37602 391 19 Goin go NNP 37602 391 20 , , , 37602 391 21 1970 1970 CD 37602 391 22 , , , 37602 391 23 and and CC 37602 391 24 _ _ NNP 37602 391 25 Hyla Hyla NNP 37602 391 26 cabrerai cabrerai NN 37602 391 27 _ _ NNP 37602 391 28 Cochran Cochran NNP 37602 391 29 and and CC 37602 391 30 Goin go NNP 37602 391 31 , , , 37602 391 32 1970 1970 CD 37602 391 33 , , , 37602 391 34 as as IN 37602 391 35 junior junior JJ 37602 391 36 synonyms synonyms NN 37602 391 37 of of IN 37602 391 38 _ _ NNP 37602 391 39 Hyla Hyla NNP 37602 391 40 buckleyi buckleyi NN 37602 391 41 _ _ NNP 37602 391 42 Boulenger Boulenger NNP 37602 391 43 , , , 37602 391 44 1882 1882 CD 37602 391 45 . . . 37602 392 1 _ _ NNP 37602 392 2 Diagnosis._--1 Diagnosis._--1 NNP 37602 392 3 ) ) -RRB- 37602 392 4 Size size NN 37602 392 5 moderate moderate JJ 37602 392 6 , , , 37602 392 7 sexual sexual JJ 37602 392 8 dimorphism dimorphism NN 37602 392 9 extreme extreme NN 37602 392 10 ; ; , 37602 392 11 maximum maximum JJ 37602 392 12 observed observed JJ 37602 392 13 snout snout NN 37602 392 14 - - HYPH 37602 392 15 vent vent NN 37602 392 16 length length NN 37602 392 17 in in IN 37602 392 18 males male NNS 37602 392 19 48.1 48.1 CD 37602 392 20 mm mm CD 37602 392 21 , , , 37602 392 22 in in IN 37602 392 23 females females NNP 37602 392 24 75.1 75.1 CD 37602 392 25 mm mm CD 37602 392 26 ; ; : 37602 392 27 2 2 LS 37602 392 28 ) ) -RRB- 37602 392 29 skin skin NN 37602 392 30 on on IN 37602 392 31 dorsum dorsum NN 37602 392 32 in in IN 37602 392 33 males male NNS 37602 392 34 bearing bear VBG 37602 392 35 a a DT 37602 392 36 mixture mixture NN 37602 392 37 of of IN 37602 392 38 large large JJ 37602 392 39 and and CC 37602 392 40 small small JJ 37602 392 41 non non JJ 37602 392 42 - - JJ 37602 392 43 spinous spinous JJ 37602 392 44 tubercles tubercle NNS 37602 392 45 ; ; : 37602 392 46 3 3 LS 37602 392 47 ) ) -RRB- 37602 392 48 skin skin NN 37602 392 49 on on IN 37602 392 50 flanks flank NNS 37602 392 51 , , , 37602 392 52 especially especially RB 37602 392 53 anteriorly anteriorly RB 37602 392 54 , , , 37602 392 55 areolate areolate NN 37602 392 56 ; ; : 37602 392 57 4 4 LS 37602 392 58 ) ) -RRB- 37602 392 59 web web NN 37602 392 60 usually usually RB 37602 392 61 extending extend VBG 37602 392 62 only only RB 37602 392 63 to to IN 37602 392 64 base base NN 37602 392 65 of of IN 37602 392 66 antepenultimate antepenultimate NN 37602 392 67 phalange phalange NN 37602 392 68 on on IN 37602 392 69 inner inner JJ 37602 392 70 edge edge NN 37602 392 71 of of IN 37602 392 72 third third JJ 37602 392 73 finger finger NN 37602 392 74 ; ; : 37602 392 75 5 5 LS 37602 392 76 ) ) -RRB- 37602 392 77 dorsum dorsum NN 37602 392 78 pale pale JJ 37602 392 79 tan tan NN 37602 392 80 or or CC 37602 392 81 green green JJ 37602 392 82 with with IN 37602 392 83 irregular irregular JJ 37602 392 84 , , , 37602 392 85 longitudinal longitudinal JJ 37602 392 86 , , , 37602 392 87 dark dark JJ 37602 392 88 brown brown JJ 37602 392 89 blotches blotch NNS 37602 392 90 , , , 37602 392 91 usually usually RB 37602 392 92 narrowly narrowly RB 37602 392 93 outlined outline VBN 37602 392 94 with with IN 37602 392 95 cream cream NN 37602 392 96 ; ; : 37602 392 97 6 6 LS 37602 392 98 ) ) -RRB- 37602 392 99 venter venter NN 37602 392 100 cream cream NN 37602 392 101 or or CC 37602 392 102 tan tan NN 37602 392 103 , , , 37602 392 104 suffused suffuse VBN 37602 392 105 with with IN 37602 392 106 brown brown JJ 37602 392 107 or or CC 37602 392 108 marked mark VBN 37602 392 109 with with IN 37602 392 110 brown brown JJ 37602 392 111 spots spot NNS 37602 392 112 in in IN 37602 392 113 some some DT 37602 392 114 specimens specimen NNS 37602 392 115 ; ; : 37602 392 116 7 7 LS 37602 392 117 ) ) -RRB- 37602 392 118 lips lip NNS 37602 392 119 marked mark VBN 37602 392 120 with with IN 37602 392 121 vertical vertical JJ 37602 392 122 brown brown NN 37602 392 123 and and CC 37602 392 124 cream cream NN 37602 392 125 bars bar NNS 37602 392 126 ; ; : 37602 392 127 8) 8) CD 37602 392 128 flanks flank VBZ 37602 392 129 creamy creamy NNP 37602 392 130 tan tan NNP 37602 392 131 with with IN 37602 392 132 irregular irregular JJ 37602 392 133 brown brown JJ 37602 392 134 spots spot NNS 37602 392 135 and/or and/or CC 37602 392 136 diagonal diagonal JJ 37602 392 137 marks mark NNS 37602 392 138 ; ; : 37602 392 139 9 9 LS 37602 392 140 ) ) -RRB- 37602 392 141 dermal dermal JJ 37602 392 142 roofing roofing NN 37602 392 143 bones bone NNS 37602 392 144 of of IN 37602 392 145 skull skull NN 37602 392 146 lacking lacking NN 37602 392 147 exostosis exostosis NN 37602 392 148 ; ; : 37602 392 149 10 10 LS 37602 392 150 ) ) -RRB- 37602 392 151 dermal dermal JJ 37602 392 152 sphenethmoid sphenethmoid JJ 37602 392 153 absent absent NN 37602 392 154 ; ; : 37602 392 155 11 11 CD 37602 392 156 ) ) -RRB- 37602 392 157 nasals nasal NNS 37602 392 158 widely widely RB 37602 392 159 separated separate VBD 37602 392 160 medially medially RB 37602 392 161 ; ; : 37602 392 162 12 12 CD 37602 392 163 ) ) -RRB- 37602 392 164 anteromedial anteromedial JJ 37602 392 165 margin margin NN 37602 392 166 of of IN 37602 392 167 frontoparietal frontoparietal JJ 37602 392 168 at at IN 37602 392 169 mid mid JJ 37602 392 170 - - NN 37602 392 171 level level NN 37602 392 172 of of IN 37602 392 173 orbit orbit NN 37602 392 174 ; ; : 37602 392 175 13 13 CD 37602 392 176 ) ) -RRB- 37602 392 177 frontoparietal frontoparietal JJ 37602 392 178 fontanelle fontanelle NN 37602 392 179 partially partially RB 37602 392 180 exposed expose VBN 37602 392 181 ; ; : 37602 392 182 14 14 CD 37602 392 183 ) ) -RRB- 37602 392 184 palatine palatine NNP 37602 392 185 serrate serrate NN 37602 392 186 ; ; : 37602 392 187 15 15 CD 37602 392 188 ) ) -RRB- 37602 392 189 parasphenoid parasphenoid NN 37602 392 190 bearing bearing NN 37602 392 191 odontoids odontoid NNS 37602 392 192 ; ; : 37602 392 193 16 16 LS 37602 392 194 ) ) -RRB- 37602 392 195 zygomatic zygomatic JJ 37602 392 196 ramus ramus NN 37602 392 197 of of IN 37602 392 198 squamosal squamosal NNP 37602 392 199 extending extend VBG 37602 392 200 approximately approximately RB 37602 392 201 one one CD 37602 392 202 - - HYPH 37602 392 203 half half NN 37602 392 204 of of IN 37602 392 205 distance distance NN 37602 392 206 to to IN 37602 392 207 maxillary maxillary NNP 37602 392 208 arch arch NNP 37602 392 209 ; ; : 37602 392 210 17 17 CD 37602 392 211 ) ) -RRB- 37602 392 212 transverse transverse JJ 37602 392 213 processes process NNS 37602 392 214 of of IN 37602 392 215 third third JJ 37602 392 216 presacral presacral JJ 37602 392 217 vertebra vertebra NN 37602 392 218 narrower narrower RBR 37602 392 219 than than IN 37602 392 220 sacral sacral JJ 37602 392 221 diapophyses diapophysis NNS 37602 392 222 ; ; : 37602 392 223 transverse transverse JJ 37602 392 224 processes process NNS 37602 392 225 of of IN 37602 392 226 presacral presacral JJ 37602 392 227 vertebrae vertebra NNS 37602 392 228 3 3 CD 37602 392 229 - - SYM 37602 392 230 8 8 CD 37602 392 231 subequal subequal JJ 37602 392 232 in in IN 37602 392 233 width width NN 37602 392 234 and and CC 37602 392 235 narrower narrow JJR 37602 392 236 in in IN 37602 392 237 males male NNS 37602 392 238 than than IN 37602 392 239 in in IN 37602 392 240 females female NNS 37602 392 241 ; ; : 37602 392 242 18 18 CD 37602 392 243 ) ) -RRB- 37602 392 244 intermandibularis intermandibularis NNP 37602 392 245 and and CC 37602 392 246 submentalis submentalis NNP 37602 392 247 muscles muscles NNP 37602 392 248 independent independent JJ 37602 392 249 ; ; : 37602 392 250 19 19 CD 37602 392 251 ) ) -RRB- 37602 392 252 supramandibular supramandibular JJ 37602 392 253 portion portion NN 37602 392 254 of of IN 37602 392 255 interhyoideus interhyoideus NNP 37602 392 256 extensively extensively RB 37602 392 257 developed developed JJ 37602 392 258 ; ; : 37602 392 259 associated associate VBN 37602 392 260 skin skin NN 37602 392 261 forming form VBG 37602 392 262 broad broad RB 37602 392 263 loose loose JJ 37602 392 264 fold fold NN 37602 392 265 . . . 37602 393 1 _ _ NNP 37602 393 2 Osteocephalus osteocephalus JJ 37602 393 3 buckleyi buckleyi NN 37602 393 4 _ _ NNP 37602 393 5 can can MD 37602 393 6 be be VB 37602 393 7 distinguished distinguish VBN 37602 393 8 readily readily RB 37602 393 9 from from IN 37602 393 10 all all DT 37602 393 11 other other JJ 37602 393 12 species specie NNS 37602 393 13 in in IN 37602 393 14 the the DT 37602 393 15 genus genus NN 37602 393 16 by by IN 37602 393 17 the the DT 37602 393 18 presence presence NN 37602 393 19 of of IN 37602 393 20 areolate areolate NN 37602 393 21 skin skin NN 37602 393 22 anteriorly anteriorly RB 37602 393 23 on on IN 37602 393 24 the the DT 37602 393 25 flanks flank NNS 37602 393 26 and and CC 37602 393 27 by by IN 37602 393 28 the the DT 37602 393 29 rather rather RB 37602 393 30 boldly boldly RB 37602 393 31 contrasting contrast VBG 37602 393 32 dorsal dorsal NN 37602 393 33 pattern pattern NN 37602 393 34 . . . 37602 394 1 Furthermore furthermore RB 37602 394 2 , , , 37602 394 3 females female NNS 37602 394 4 are be VBP 37602 394 5 distinctive distinctive JJ 37602 394 6 in in IN 37602 394 7 having have VBG 37602 394 8 tubercles tubercle NNS 37602 394 9 on on IN 37602 394 10 the the DT 37602 394 11 eyelids eyelid NNS 37602 394 12 and and CC 37602 394 13 supratympanic supratympanic JJ 37602 394 14 folds fold NNS 37602 394 15 . . . 37602 395 1 _ _ NNP 37602 395 2 Distribution._--The Distribution._--The NNP 37602 395 3 periphery periphery NN 37602 395 4 of of IN 37602 395 5 the the DT 37602 395 6 Amazon Amazon NNP 37602 395 7 Basin Basin NNP 37602 395 8 , , , 37602 395 9 in in IN 37602 395 10 the the DT 37602 395 11 Guianas Guianas NNP 37602 395 12 and and CC 37602 395 13 Territorio Territorio NNP 37602 395 14 do do VBP 37602 395 15 Amapá Amapá NNP 37602 395 16 in in IN 37602 395 17 northeastern northeastern JJ 37602 395 18 Brasil Brasil NNP 37602 395 19 ; ; : 37602 395 20 the the DT 37602 395 21 upper upper JJ 37602 395 22 Amazon Amazon NNP 37602 395 23 Basin Basin NNP 37602 395 24 from from IN 37602 395 25 southern southern JJ 37602 395 26 Colombia Colombia NNP 37602 395 27 to to TO 37602 395 28 east east JJ 37602 395 29 - - HYPH 37602 395 30 central central JJ 37602 395 31 Bolivia Bolivia NNP 37602 395 32 ; ; : 37602 395 33 one one CD 37602 395 34 locality locality NN 37602 395 35 ( ( -LRB- 37602 395 36 Acevedo Acevedo NNP 37602 395 37 ) ) -RRB- 37602 395 38 in in IN 37602 395 39 upper upper JJ 37602 395 40 Río Río NNP 37602 395 41 Magdalena Magdalena NNP 37602 395 42 drainage drainage NN 37602 395 43 in in IN 37602 395 44 Colombia Colombia NNP 37602 395 45 ( ( -LRB- 37602 395 46 Fig Fig NNP 37602 395 47 . . . 37602 396 1 8) 8) CD 37602 396 2 . . . 37602 397 1 All all DT 37602 397 2 localities locality NNS 37602 397 3 are be VBP 37602 397 4 at at IN 37602 397 5 elevations elevation NNS 37602 397 6 of of IN 37602 397 7 less less JJR 37602 397 8 than than IN 37602 397 9 700 700 CD 37602 397 10 m. m. NN 37602 397 11 Records Records NNP 37602 397 12 for for IN 37602 397 13 Pallatanga Pallatanga NNP 37602 397 14 and and CC 37602 397 15 Santiago Santiago NNP 37602 397 16 in in IN 37602 397 17 Provincia Provincia NNP 37602 397 18 Chimborazo Chimborazo NNP 37602 397 19 , , , 37602 397 20 Ecuador Ecuador NNP 37602 397 21 ( ( -LRB- 37602 397 22 high high RB 37602 397 23 on on IN 37602 397 24 the the DT 37602 397 25 Pacific Pacific NNP 37602 397 26 slopes slope NNS 37602 397 27 of of IN 37602 397 28 the the DT 37602 397 29 Andes Andes NNP 37602 397 30 ) ) -RRB- 37602 397 31 , , , 37602 397 32 are be VBP 37602 397 33 considered consider VBN 37602 397 34 to to TO 37602 397 35 be be VB 37602 397 36 erroneous erroneous JJ 37602 397 37 . . . 37602 398 1 78 78 CD 37602 398 2 specimens specimen NNS 37602 398 3 from from IN 37602 398 4 40 40 CD 37602 398 5 localities locality NNS 37602 398 6 . . . 37602 399 1 [ [ -LRB- 37602 399 2 Illustration illustration NN 37602 399 3 : : : 37602 399 4 FIG FIG NNP 37602 399 5 . . . 37602 400 1 8 8 LS 37602 400 2 . . . 37602 401 1 Distribution distribution NN 37602 401 2 of of IN 37602 401 3 _ _ NNP 37602 401 4 Osteocephalus Osteocephalus NNP 37602 401 5 buckleyi buckleyi NN 37602 401 6 _ _ NNP 37602 401 7 ( ( -LRB- 37602 401 8 circles circle NNS 37602 401 9 ) ) -RRB- 37602 401 10 and and CC 37602 401 11 _ _ NNP 37602 401 12 O. O. NNP 37602 402 1 pearsoni pearsoni NNP 37602 402 2 _ _ NNP 37602 402 3 ( ( -LRB- 37602 402 4 triangles triangle NNS 37602 402 5 ) ) -RRB- 37602 402 6 . . . 37602 402 7 ] ] -RRB- 37602 403 1 _ _ NNP 37602 403 2 Remarks._--In Remarks._--In NNP 37602 403 3 life life NN 37602 403 4 the the DT 37602 403 5 dorsum dorsum NN 37602 403 6 is be VBZ 37602 403 7 green green JJ 37602 403 8 with with IN 37602 403 9 dark dark JJ 37602 403 10 markings marking NNS 37602 403 11 . . . 37602 404 1 A a DT 37602 404 2 male male NN 37602 404 3 ( ( -LRB- 37602 404 4 KU KU NNP 37602 404 5 123171 123171 CD 37602 404 6 ) ) -RRB- 37602 404 7 from from IN 37602 404 8 Santa Santa NNP 37602 404 9 Cecilia Cecilia NNP 37602 404 10 , , , 37602 404 11 Ecuador Ecuador NNP 37602 404 12 , , , 37602 404 13 was be VBD 37602 404 14 : : : 37602 404 15 " " `` 37602 404 16 Dorsum dorsum NN 37602 404 17 green green NN 37602 404 18 with with IN 37602 404 19 dark dark JJ 37602 404 20 brown brown JJ 37602 404 21 blotches blotch NNS 37602 404 22 . . . 37602 405 1 Anterior Anterior NNP 37602 405 2 and and CC 37602 405 3 posterior posterior NN 37602 405 4 surfaces surface NNS 37602 405 5 of of IN 37602 405 6 thighs thigh NNS 37602 405 7 dull dull JJ 37602 405 8 blue blue NNP 37602 405 9 . . . 37602 406 1 Venter Venter NNP 37602 406 2 brown brown NNP 37602 406 3 , , , 37602 406 4 flecked fleck VBN 37602 406 5 with with IN 37602 406 6 white white NNP 37602 406 7 . . . 37602 407 1 Iris iris JJ 37602 407 2 greenish greenish JJ 37602 407 3 bronze bronze NN 37602 407 4 with with IN 37602 407 5 brown brown JJ 37602 407 6 horizontal horizontal JJ 37602 407 7 triangles triangle NNS 37602 407 8 and and CC 37602 407 9 ventromedian ventromedian JJ 37602 407 10 brown brown JJ 37602 407 11 line line NN 37602 407 12 . . . 37602 407 13 " " '' 37602 408 1 ( ( -LRB- 37602 408 2 W. W. NNP 37602 408 3 E. E. NNP 37602 408 4 Duellman Duellman NNP 37602 408 5 , , , 37602 408 6 field field NN 37602 408 7 notes note NNS 37602 408 8 , , , 37602 408 9 16 16 CD 37602 408 10 June June NNP 37602 408 11 1968 1968 CD 37602 408 12 . . . 37602 408 13 ) ) -RRB- 37602 409 1 A a DT 37602 409 2 female female NN 37602 409 3 ( ( -LRB- 37602 409 4 KU KU NNP 37602 409 5 126646 126646 CD 37602 409 6 ) ) -RRB- 37602 409 7 from from IN 37602 409 8 Lago Lago NNP 37602 409 9 Agrio Agrio NNP 37602 409 10 , , , 37602 409 11 Ecuador Ecuador NNP 37602 409 12 , , , 37602 409 13 was be VBD 37602 409 14 : : : 37602 409 15 " " `` 37602 409 16 Dorsum dorsum NN 37602 409 17 pale pale JJ 37602 409 18 green green NN 37602 409 19 with with IN 37602 409 20 darker dark JJR 37602 409 21 green green JJ 37602 409 22 blotches blotch NNS 37602 409 23 and and CC 37602 409 24 creamy creamy VB 37602 409 25 yellow yellow JJ 37602 409 26 middorsal middorsal NN 37602 409 27 stripe stripe NN 37602 409 28 . . . 37602 410 1 Lateral lateral JJ 37602 410 2 blotches blotch NNS 37602 410 3 bronze bronze NN 37602 410 4 - - HYPH 37602 410 5 tan tan NN 37602 410 6 . . . 37602 411 1 Flanks flank NNS 37602 411 2 tan tan VBP 37602 411 3 with with IN 37602 411 4 black black JJ 37602 411 5 blotches blotch NNS 37602 411 6 . . . 37602 412 1 Anterior Anterior NNP 37602 412 2 surfaces surface NNS 37602 412 3 of of IN 37602 412 4 thighs thigh NNS 37602 412 5 dark dark JJ 37602 412 6 brown brown NNP 37602 412 7 . . . 37602 413 1 Dorsal dorsal NN 37602 413 2 and and CC 37602 413 3 posterior posterior NN 37602 413 4 surfaces surface NNS 37602 413 5 of of IN 37602 413 6 thighs thigh NNS 37602 413 7 and and CC 37602 413 8 shanks shank VBZ 37602 413 9 tan tan NNP 37602 413 10 with with IN 37602 413 11 dark dark JJ 37602 413 12 brown brown JJ 37602 413 13 blotches blotch NNS 37602 413 14 . . . 37602 414 1 Webbing web VBG 37602 414 2 brown brown JJ 37602 414 3 . . . 37602 415 1 Suborbital suborbital JJ 37602 415 2 spot spot NN 37602 415 3 green green JJ 37602 415 4 . . . 37602 416 1 Postorbital postorbital JJ 37602 416 2 bar bar NN 37602 416 3 black black NN 37602 416 4 . . . 37602 417 1 Belly Belly NNP 37602 417 2 grayish grayish NN 37602 417 3 brown brown JJ 37602 417 4 in in IN 37602 417 5 appearance appearance NN 37602 417 6 -- -- : 37602 417 7 tips tip NNS 37602 417 8 of of IN 37602 417 9 granules granule NNS 37602 417 10 white white JJ 37602 417 11 ; ; , 37602 417 12 intergranular intergranular JJ 37602 417 13 spaces space NNS 37602 417 14 brown brown NNP 37602 417 15 . . . 37602 418 1 Iris iris JJ 37602 418 2 golden golden JJ 37602 418 3 bronze bronze NN 37602 418 4 with with IN 37602 418 5 black black JJ 37602 418 6 flecks fleck NNS 37602 418 7 peripherally peripherally RB 37602 418 8 and and CC 37602 418 9 median median JJ 37602 418 10 , , , 37602 418 11 horizontal horizontal JJ 37602 418 12 , , , 37602 418 13 reddish reddish JJ 37602 418 14 brown brown JJ 37602 418 15 streak streak NN 37602 418 16 . . . 37602 418 17 " " '' 37602 419 1 ( ( -LRB- 37602 419 2 W. W. NNP 37602 419 3 E. E. NNP 37602 419 4 Duellman Duellman NNP 37602 419 5 , , , 37602 419 6 field field NN 37602 419 7 notes note NNS 37602 419 8 , , , 37602 419 9 12 12 CD 37602 419 10 May May NNP 37602 419 11 1969 1969 CD 37602 419 12 . . . 37602 419 13 ) ) -RRB- 37602 420 1 No no DT 37602 420 2 ontogenetic ontogenetic JJ 37602 420 3 change change NN 37602 420 4 in in IN 37602 420 5 coloration coloration NN 37602 420 6 has have VBZ 37602 420 7 been be VBN 37602 420 8 noted note VBN 37602 420 9 . . . 37602 421 1 = = NFP 37602 421 2 Osteocephalus osteocephalus JJ 37602 421 3 leprieurii= leprieurii= NNS 37602 421 4 ( ( -LRB- 37602 421 5 Duméril Duméril NNP 37602 421 6 and and CC 37602 421 7 Bibron Bibron NNP 37602 421 8 ) ) -RRB- 37602 421 9 _ _ NNP 37602 421 10 Hyla Hyla NNP 37602 421 11 leprieurii leprieurii VBZ 37602 421 12 _ _ NNP 37602 421 13 Duméril Duméril NNP 37602 421 14 and and CC 37602 421 15 Bibron Bibron NNP 37602 421 16 , , , 37602 421 17 1841:553 1841:553 CD 37602 421 18 [ [ -LRB- 37602 421 19 Holotype.--MNHN holotype.--mnhn NN 37602 421 20 4629 4629 CD 37602 421 21 from from IN 37602 421 22 " " `` 37602 421 23 Cayenne Cayenne NNP 37602 421 24 " " '' 37602 421 25 ; ; : 37602 421 26 Mons mon NNS 37602 421 27 . . . 37602 422 1 Leprieur Leprieur NNP 37602 422 2 collector collector NNP 37602 422 3 ] ] -RRB- 37602 422 4 . . . 37602 423 1 _ _ NNP 37602 423 2 Hypsiboas Hypsiboas NNP 37602 423 3 leprieurii_--Cope leprieurii_--Cope NNP 37602 423 4 , , , 37602 423 5 1867:200 1867:200 CD 37602 423 6 . . . 37602 424 1 _ _ NNP 37602 424 2 Hyla Hyla NNP 37602 424 3 leprieurii leprieurii JJ 37602 424 4 britti britti NNP 37602 424 5 _ _ NNP 37602 424 6 Melin Melin NNP 37602 424 7 , , , 37602 424 8 1941:42 1941:42 CD 37602 424 9 [ [ -LRB- 37602 424 10 Holotype.--NHMG holotype.--nhmg RB 37602 424 11 489 489 CD 37602 424 12 from from IN 37602 424 13 the the DT 37602 424 14 Rio Rio NNP 37602 424 15 Uaupés Uaupés NNP 37602 424 16 , , , 37602 424 17 north north NN 37602 424 18 of of IN 37602 424 19 the the DT 37602 424 20 Rio Rio NNP 37602 424 21 Japu Japu NNP 37602 424 22 , , , 37602 424 23 Territorio Territorio NNP 37602 424 24 do do VBP 37602 424 25 Amazonas Amazonas NNP 37602 424 26 , , , 37602 424 27 Brasil Brasil NNP 37602 424 28 ; ; : 37602 424 29 Douglas Douglas NNP 37602 424 30 Melin Melin NNP 37602 424 31 collector collector NN 37602 424 32 ] ] -RRB- 37602 424 33 . . . 37602 425 1 New new JJ 37602 425 2 synonymy synonymy NNP 37602 425 3 . . . 37602 426 1 _ _ NNP 37602 426 2 Hyla Hyla NNP 37602 426 3 leprieurii leprieurii JJ 37602 426 4 leprieurii_--Melin leprieurii_--Melin NNP 37602 426 5 , , , 37602 426 6 1941:42 1941:42 CD 37602 426 7 . . . 37602 427 1 _ _ NNP 37602 427 2 Osteocephalus Osteocephalus NNP 37602 427 3 britti_--Goin britti_--Goin NNP 37602 427 4 , , , 37602 427 5 1961:13 1961:13 CD 37602 427 6 . . . 37602 428 1 _ _ NNP 37602 428 2 Osteocephalus Osteocephalus NNP 37602 428 3 leprieurii_--Goin leprieurii_--Goin NNP 37602 428 4 , , , 37602 428 5 1961:13 1961:13 CD 37602 428 6 . . . 37602 429 1 _ _ NNP 37602 429 2 Justification Justification NNP 37602 429 3 of of IN 37602 429 4 Synonymy._--The synonymy._--the CD 37602 429 5 holotype holotype NN 37602 429 6 of of IN 37602 429 7 _ _ NNP 37602 429 8 Hyla Hyla NNP 37602 429 9 leprieurii leprieurii JJ 37602 429 10 _ _ NNP 37602 429 11 is be VBZ 37602 429 12 a a DT 37602 429 13 female female NN 37602 429 14 having have VBG 37602 429 15 a a DT 37602 429 16 snout snout NN 37602 429 17 - - HYPH 37602 429 18 vent vent NN 37602 429 19 length length NN 37602 429 20 of of IN 37602 429 21 46.6 46.6 CD 37602 429 22 mm mm NNS 37602 429 23 . . . 37602 430 1 The the DT 37602 430 2 diameter diameter NN 37602 430 3 of of IN 37602 430 4 the the DT 37602 430 5 tympanum tympanum NN 37602 430 6 is be VBZ 37602 430 7 3.7 3.7 CD 37602 430 8 mm mm CD 37602 430 9 , , , 37602 430 10 69.8 69.8 CD 37602 430 11 percent percent NN 37602 430 12 of of IN 37602 430 13 the the DT 37602 430 14 diameter diameter NN 37602 430 15 of of IN 37602 430 16 the the DT 37602 430 17 eye eye NN 37602 430 18 . . . 37602 431 1 The the DT 37602 431 2 dorsal dorsal NN 37602 431 3 roofing roofing NN 37602 431 4 bones bone NNS 37602 431 5 are be VBP 37602 431 6 smooth smooth JJ 37602 431 7 , , , 37602 431 8 and and CC 37602 431 9 the the DT 37602 431 10 skin skin NN 37602 431 11 on on IN 37602 431 12 the the DT 37602 431 13 dorsum dorsum NN 37602 431 14 is be VBZ 37602 431 15 smooth smooth JJ 37602 431 16 . . . 37602 432 1 The the DT 37602 432 2 penultimate penultimate JJ 37602 432 3 phalanges phalanx NNS 37602 432 4 of of IN 37602 432 5 the the DT 37602 432 6 fingers finger NNS 37602 432 7 are be VBP 37602 432 8 not not RB 37602 432 9 included include VBN 37602 432 10 in in IN 37602 432 11 the the DT 37602 432 12 webbing webbing NN 37602 432 13 . . . 37602 433 1 When when WRB 37602 433 2 we -PRON- PRP 37602 433 3 examined examine VBD 37602 433 4 the the DT 37602 433 5 specimen speciman NNS 37602 433 6 on on IN 37602 433 7 2 2 CD 37602 433 8 July July NNP 37602 433 9 1969 1969 CD 37602 433 10 , , , 37602 433 11 it -PRON- PRP 37602 433 12 was be VBD 37602 433 13 slightly slightly RB 37602 433 14 soft soft JJ 37602 433 15 and and CC 37602 433 16 somewhat somewhat RB 37602 433 17 faded fade VBD 37602 433 18 to to IN 37602 433 19 a a DT 37602 433 20 peculiar peculiar JJ 37602 433 21 grayish grayish NN 37602 433 22 green green JJ 37602 433 23 color color NN 37602 433 24 with with IN 37602 433 25 faint faint JJ 37602 433 26 darker dark JJR 37602 433 27 transverse transverse JJ 37602 433 28 bars bar NNS 37602 433 29 on on IN 37602 433 30 the the DT 37602 433 31 limbs limb NNS 37602 433 32 . . . 37602 434 1 Duméril Duméril NNP 37602 434 2 and and CC 37602 434 3 Bibron Bibron NNP 37602 434 4 ( ( -LRB- 37602 434 5 1841:554 1841:554 CD 37602 434 6 ) ) -RRB- 37602 434 7 described describe VBD 37602 434 8 the the DT 37602 434 9 coloration coloration NN 37602 434 10 , , , 37602 434 11 as as IN 37602 434 12 follows follow VBZ 37602 434 13 : : : 37602 434 14 " " `` 37602 434 15 The the DT 37602 434 16 loreal loreal NN 37602 434 17 region region NN 37602 434 18 is be VBZ 37602 434 19 black black JJ 37602 434 20 . . . 37602 435 1 A a DT 37602 435 2 stripe stripe NN 37602 435 3 of of IN 37602 435 4 the the DT 37602 435 5 same same JJ 37602 435 6 color color NN 37602 435 7 extends extend VBZ 37602 435 8 from from IN 37602 435 9 the the DT 37602 435 10 posterior posterior JJ 37602 435 11 border border NN 37602 435 12 of of IN 37602 435 13 the the DT 37602 435 14 orbit orbit NN 37602 435 15 to to IN 37602 435 16 the the DT 37602 435 17 corner corner NN 37602 435 18 of of IN 37602 435 19 the the DT 37602 435 20 mouth mouth NN 37602 435 21 , , , 37602 435 22 passing pass VBG 37602 435 23 through through IN 37602 435 24 the the DT 37602 435 25 tympanum tympanum NN 37602 435 26 . . . 37602 436 1 All all DT 37602 436 2 of of IN 37602 436 3 the the DT 37602 436 4 dorsal dorsal NN 37602 436 5 parts part NNS 37602 436 6 are be VBP 37602 436 7 grayish grayish NN 37602 436 8 white white JJ 37602 436 9 with with IN 37602 436 10 large large JJ 37602 436 11 transverse transverse JJR 37602 436 12 brown brown JJ 37602 436 13 bands band NNS 37602 436 14 , , , 37602 436 15 which which WDT 37602 436 16 are be VBP 37602 436 17 more more RBR 37602 436 18 expanded expanded JJ 37602 436 19 and and CC 37602 436 20 less less RBR 37602 436 21 regularly regularly RB 37602 436 22 outlined outline VBN 37602 436 23 on on IN 37602 436 24 the the DT 37602 436 25 back back NN 37602 436 26 than than IN 37602 436 27 on on IN 37602 436 28 the the DT 37602 436 29 limbs limb NNS 37602 436 30 . . . 37602 437 1 There there EX 37602 437 2 is be VBZ 37602 437 3 one one CD 37602 437 4 of of IN 37602 437 5 these these DT 37602 437 6 on on IN 37602 437 7 the the DT 37602 437 8 occiput occiput NN 37602 437 9 that that WDT 37602 437 10 is be VBZ 37602 437 11 in in IN 37602 437 12 a a DT 37602 437 13 triangular triangular JJ 37602 437 14 shape shape NN 37602 437 15 . . . 37602 438 1 All all DT 37602 438 2 of of IN 37602 438 3 the the DT 37602 438 4 venter venter NN 37602 438 5 is be VBZ 37602 438 6 white white JJ 37602 438 7 . . . 37602 438 8 " " '' 37602 439 1 ( ( -LRB- 37602 439 2 Free free JJ 37602 439 3 translation translation NN 37602 439 4 from from IN 37602 439 5 French French NNP 37602 439 6 . . . 37602 439 7 ) ) -RRB- 37602 440 1 The the DT 37602 440 2 holotype holotype NN 37602 440 3 of of IN 37602 440 4 _ _ NNP 37602 440 5 Hyla Hyla NNP 37602 440 6 leprieurii leprieurii JJ 37602 440 7 britti britti NNP 37602 440 8 _ _ NNP 37602 440 9 is be VBZ 37602 440 10 a a DT 37602 440 11 male male NN 37602 440 12 having have VBG 37602 440 13 a a DT 37602 440 14 snout snout NN 37602 440 15 - - HYPH 37602 440 16 vent vent NN 37602 440 17 length length NN 37602 440 18 of of IN 37602 440 19 48.1 48.1 CD 37602 440 20 mm mm NNS 37602 440 21 . . . 37602 441 1 The the DT 37602 441 2 diameter diameter NN 37602 441 3 of of IN 37602 441 4 the the DT 37602 441 5 tympanum tympanum NN 37602 441 6 is be VBZ 37602 441 7 3.6 3.6 CD 37602 441 8 mm mm CD 37602 441 9 , , , 37602 441 10 65.5 65.5 CD 37602 441 11 percent percent NN 37602 441 12 of of IN 37602 441 13 the the DT 37602 441 14 diameter diameter NN 37602 441 15 of of IN 37602 441 16 the the DT 37602 441 17 eye eye NN 37602 441 18 . . . 37602 442 1 The the DT 37602 442 2 skin skin NN 37602 442 3 on on IN 37602 442 4 the the DT 37602 442 5 dorsum dorsum NN 37602 442 6 is be VBZ 37602 442 7 tubercular tubercular JJ 37602 442 8 ; ; : 37602 442 9 the the DT 37602 442 10 tubercles tubercle NNS 37602 442 11 are be VBP 37602 442 12 small small JJ 37602 442 13 on on IN 37602 442 14 head head NN 37602 442 15 and and CC 37602 442 16 on on IN 37602 442 17 the the DT 37602 442 18 dorsal dorsal NN 37602 442 19 surfaces surface NNS 37602 442 20 of of IN 37602 442 21 the the DT 37602 442 22 limbs limb NNS 37602 442 23 and and CC 37602 442 24 slightly slightly RB 37602 442 25 larger large JJR 37602 442 26 on on IN 37602 442 27 the the DT 37602 442 28 back back NN 37602 442 29 . . . 37602 443 1 The the DT 37602 443 2 penultimate penultimate JJ 37602 443 3 phalanges phalanx NNS 37602 443 4 of of IN 37602 443 5 the the DT 37602 443 6 fingers finger NNS 37602 443 7 are be VBP 37602 443 8 not not RB 37602 443 9 included include VBN 37602 443 10 in in IN 37602 443 11 the the DT 37602 443 12 webbing webbing NN 37602 443 13 . . . 37602 444 1 Melin Melin NNP 37602 444 2 ( ( -LRB- 37602 444 3 1941:43 1941:43 CD 37602 444 4 ) ) -RRB- 37602 444 5 stated state VBD 37602 444 6 : : : 37602 444 7 " " `` 37602 444 8 Above above IN 37602 444 9 blackish blackish NNP 37602 444 10 brown brown JJ 37602 444 11 with with IN 37602 444 12 a a DT 37602 444 13 very very RB 37602 444 14 indistinct indistinct JJ 37602 444 15 band band NN 37602 444 16 between between IN 37602 444 17 the the DT 37602 444 18 eyes eye NNS 37602 444 19 ; ; , 37602 444 20 iris iris NNP 37602 444 21 with with IN 37602 444 22 mottle mottle NNP 37602 444 23 of of IN 37602 444 24 metallic metallic NNP 37602 444 25 lustre lustre NNP 37602 444 26 ; ; : 37602 444 27 hinder hinder NN 37602 444 28 parts part NNS 37602 444 29 of of IN 37602 444 30 upper upper JJ 37602 444 31 jaw jaw NN 37602 444 32 whitish whitish NN 37602 444 33 ; ; : 37602 444 34 sides side NNS 37602 444 35 of of IN 37602 444 36 body body NN 37602 444 37 mottled mottle VBN 37602 444 38 with with IN 37602 444 39 blackish blackish NNP 37602 444 40 brown brown NNP 37602 444 41 ; ; : 37602 444 42 hind hind NNP 37602 444 43 limbs limb NNS 37602 444 44 ( ( -LRB- 37602 444 45 especially especially RB 37602 444 46 tibiae tibia NNS 37602 444 47 and and CC 37602 444 48 tarsi tarsi NNP 37602 444 49 ) ) -RRB- 37602 444 50 with with IN 37602 444 51 narrow narrow JJ 37602 444 52 , , , 37602 444 53 diffuse diffuse VB 37602 444 54 cross cross NN 37602 444 55 bars bar NNS 37602 444 56 ; ; : 37602 444 57 beneath beneath IN 37602 444 58 whitish whitish VBP 37602 444 59 with with IN 37602 444 60 slight slight JJ 37602 444 61 brown brown JJ 37602 444 62 mottle mottle NN 37602 444 63 along along IN 37602 444 64 jaw jaw NN 37602 444 65 . . . 37602 444 66 " " '' 37602 445 1 We -PRON- PRP 37602 445 2 examined examine VBD 37602 445 3 the the DT 37602 445 4 type type NN 37602 445 5 on on IN 37602 445 6 17 17 CD 37602 445 7 February February NNP 37602 445 8 1969 1969 CD 37602 445 9 ; ; : 37602 445 10 at at IN 37602 445 11 that that DT 37602 445 12 time time NN 37602 445 13 it -PRON- PRP 37602 445 14 was be VBD 37602 445 15 dull dull JJ 37602 445 16 brown brown NNP 37602 445 17 above above RB 37602 445 18 with with IN 37602 445 19 faint faint JJ 37602 445 20 , , , 37602 445 21 narrow narrow JJ 37602 445 22 , , , 37602 445 23 dark dark JJ 37602 445 24 brown brown JJ 37602 445 25 , , , 37602 445 26 transverse transverse JJ 37602 445 27 bars bar NNS 37602 445 28 on on IN 37602 445 29 the the DT 37602 445 30 back back NN 37602 445 31 and and CC 37602 445 32 dorsal dorsal NN 37602 445 33 surfaces surface NNS 37602 445 34 of of IN 37602 445 35 the the DT 37602 445 36 limbs limb NNS 37602 445 37 . . . 37602 446 1 A a DT 37602 446 2 cream cream NN 37602 446 3 subocular subocular JJ 37602 446 4 spot spot NN 37602 446 5 was be VBD 37602 446 6 evident evident JJ 37602 446 7 , , , 37602 446 8 and and CC 37602 446 9 the the DT 37602 446 10 venter venter NN 37602 446 11 was be VBD 37602 446 12 creamy creamy JJ 37602 446 13 white white JJ 37602 446 14 . . . 37602 447 1 Melin Melin NNP 37602 447 2 ( ( -LRB- 37602 447 3 1941:42 1941:42 CD 37602 447 4 ) ) -RRB- 37602 447 5 stated state VBD 37602 447 6 that that IN 37602 447 7 the the DT 37602 447 8 holotype holotype NN 37602 447 9 of of IN 37602 447 10 _ _ NNP 37602 447 11 Hyla Hyla NNP 37602 447 12 leprieurii leprieurii JJ 37602 447 13 britti britti NNP 37602 447 14 _ _ NNP 37602 447 15 " " '' 37602 447 16 ... ... . 37602 447 17 resembles resemble VBZ 37602 447 18 a a DT 37602 447 19 good good JJ 37602 447 20 deal deal NN 37602 447 21 _ _ NNP 37602 447 22 H. H. NNP 37602 448 1 leprieurii leprieurii JJ 37602 448 2 _ _ NNP 37602 448 3 Dum Dum NNP 37602 448 4 . . . 37602 449 1 & & CC 37602 449 2 Bibr Bibr NNP 37602 449 3 . . . 37602 450 1 As as IN 37602 450 2 , , , 37602 450 3 however however RB 37602 450 4 , , , 37602 450 5 it -PRON- PRP 37602 450 6 differs differ VBZ 37602 450 7 from from IN 37602 450 8 the the DT 37602 450 9 latter latter JJ 37602 450 10 species specie NNS 37602 450 11 by by IN 37602 450 12 its -PRON- PRP$ 37602 450 13 very very JJ 37602 450 14 concave concave NNP 37602 450 15 loreal loreal NN 37602 450 16 region region NN 37602 450 17 , , , 37602 450 18 small small JJ 37602 450 19 tympanum tympanum NN 37602 450 20 , , , 37602 450 21 and and CC 37602 450 22 almost almost RB 37602 450 23 uniformly uniformly RB 37602 450 24 brownish brownish JJ 37602 450 25 colour colour NN 37602 450 26 , , , 37602 450 27 it -PRON- PRP 37602 450 28 may may MD 37602 450 29 at at IN 37602 450 30 least least JJS 37602 450 31 form form VB 37602 450 32 a a DT 37602 450 33 subspecies subspecie NNS 37602 450 34 of of IN 37602 450 35 _ _ NNP 37602 450 36 leprieurii leprieurii JJ 37602 450 37 _ _ NNP 37602 450 38 .... .... NFP 37602 450 39 " " `` 37602 450 40 The the DT 37602 450 41 pattern pattern NN 37602 450 42 of of IN 37602 450 43 narrow narrow JJ 37602 450 44 transverse transverse JJ 37602 450 45 bars bar NNS 37602 450 46 on on IN 37602 450 47 the the DT 37602 450 48 backs back NNS 37602 450 49 of of IN 37602 450 50 the the DT 37602 450 51 holotypes holotype NNS 37602 450 52 of of IN 37602 450 53 _ _ NNP 37602 450 54 H. H. NNP 37602 451 1 leprieurii leprieurii NNP 37602 451 2 _ _ NNP 37602 451 3 and and CC 37602 451 4 _ _ NNP 37602 451 5 H. H. NNP 37602 452 1 britti britti NNP 37602 452 2 _ _ NNP 37602 452 3 is be VBZ 37602 452 4 a a DT 37602 452 5 condition condition NN 37602 452 6 shared share VBN 37602 452 7 only only RB 37602 452 8 by by IN 37602 452 9 these these DT 37602 452 10 two two CD 37602 452 11 nominal nominal JJ 37602 452 12 taxa taxa NN 37602 452 13 that that WDT 37602 452 14 are be VBP 37602 452 15 placed place VBN 37602 452 16 in in IN 37602 452 17 _ _ NNP 37602 452 18 Osteocephalus Osteocephalus NNP 37602 452 19 _ _ NNP 37602 452 20 . . . 37602 453 1 Melin Melin NNP 37602 453 2 noted note VBD 37602 453 3 that that IN 37602 453 4 _ _ NNP 37602 453 5 britti britti NNP 37602 453 6 _ _ NNP 37602 453 7 differed differ VBD 37602 453 8 from from IN 37602 453 9 _ _ NNP 37602 453 10 leprieurii leprieurii JJ 37602 453 11 _ _ NNP 37602 453 12 in in IN 37602 453 13 the the DT 37602 453 14 depth depth NN 37602 453 15 of of IN 37602 453 16 the the DT 37602 453 17 loreal loreal NN 37602 453 18 concavity concavity NN 37602 453 19 and and CC 37602 453 20 in in IN 37602 453 21 the the DT 37602 453 22 size size NN 37602 453 23 of of IN 37602 453 24 the the DT 37602 453 25 tympanum tympanum NN 37602 453 26 . . . 37602 454 1 Neither neither DT 37602 454 2 of of IN 37602 454 3 these these DT 37602 454 4 differences difference NNS 37602 454 5 is be VBZ 37602 454 6 noteworthy noteworthy JJ 37602 454 7 in in IN 37602 454 8 comparison comparison NN 37602 454 9 with with IN 37602 454 10 series series NN 37602 454 11 of of IN 37602 454 12 specimens specimens NNP 37602 454 13 . . . 37602 455 1 The the DT 37602 455 2 depth depth NN 37602 455 3 of of IN 37602 455 4 the the DT 37602 455 5 loreal loreal NN 37602 455 6 concavity concavity NN 37602 455 7 is be VBZ 37602 455 8 a a DT 37602 455 9 highly highly RB 37602 455 10 subjective subjective JJ 37602 455 11 character character NN 37602 455 12 , , , 37602 455 13 and and CC 37602 455 14 we -PRON- PRP 37602 455 15 note note VBP 37602 455 16 no no DT 37602 455 17 differences difference NNS 37602 455 18 between between IN 37602 455 19 the the DT 37602 455 20 types type NNS 37602 455 21 . . . 37602 456 1 The the DT 37602 456 2 ratio ratio NN 37602 456 3 of of IN 37602 456 4 the the DT 37602 456 5 diameter diameter NN 37602 456 6 of of IN 37602 456 7 the the DT 37602 456 8 tympanum tympanum NN 37602 456 9 to to IN 37602 456 10 the the DT 37602 456 11 diameter diameter NN 37602 456 12 of of IN 37602 456 13 the the DT 37602 456 14 eye eye NN 37602 456 15 is be VBZ 37602 456 16 relatively relatively RB 37602 456 17 smaller small JJR 37602 456 18 in in IN 37602 456 19 both both DT 37602 456 20 holotypes holotype NNS 37602 456 21 ( ( -LRB- 37602 456 22 0.698 0.698 CD 37602 456 23 in in IN 37602 456 24 _ _ NNP 37602 456 25 leprieurii_--[F leprieurii_--[F NNP 37602 456 26 ] ] -RRB- 37602 456 27 ; ; : 37602 456 28 0.655 0.655 CD 37602 456 29 in in IN 37602 456 30 _ _ NNP 37602 456 31 britti_--[M britti_--[M NNP 37602 456 32 ] ] -RRB- 37602 456 33 ) ) -RRB- 37602 456 34 than than IN 37602 456 35 in in IN 37602 456 36 series series NN 37602 456 37 of of IN 37602 456 38 fresh fresh JJ 37602 456 39 specimens specimen NNS 37602 456 40 from from IN 37602 456 41 Lago Lago NNP 37602 456 42 Agrio Agrio NNP 37602 456 43 , , , 37602 456 44 Ecuador Ecuador NNP 37602 456 45 ( ( -LRB- 37602 456 46 0.652 0.652 CD 37602 456 47 - - HYPH 37602 456 48 0.884 0.884 CD 37602 456 49 , , , 37602 456 50 mean mean VB 37602 456 51 0.785 0.785 CD 37602 456 52 in in IN 37602 456 53 17 17 CD 37602 456 54 males male NNS 37602 456 55 ; ; : 37602 456 56 0.700 0.700 CD 37602 456 57 - - SYM 37602 456 58 0.909 0.909 CD 37602 456 59 , , , 37602 456 60 mean mean VB 37602 456 61 0.790 0.790 CD 37602 456 62 in in IN 37602 456 63 20 20 CD 37602 456 64 females female NNS 37602 456 65 ) ) -RRB- 37602 456 66 . . . 37602 457 1 The the DT 37602 457 2 smaller small JJR 37602 457 3 proportions proportion NNS 37602 457 4 in in IN 37602 457 5 the the DT 37602 457 6 types type NNS 37602 457 7 may may MD 37602 457 8 be be VB 37602 457 9 due due IN 37602 457 10 to to IN 37602 457 11 geographic geographic JJ 37602 457 12 variation variation NN 37602 457 13 or or CC 37602 457 14 to to TO 37602 457 15 shrinkage shrinkage VB 37602 457 16 as as IN 37602 457 17 a a DT 37602 457 18 result result NN 37602 457 19 of of IN 37602 457 20 many many JJ 37602 457 21 years year NNS 37602 457 22 in in IN 37602 457 23 preservative preservative JJ 37602 457 24 ( ( -LRB- 37602 457 25 130 130 CD 37602 457 26 + + CD 37602 457 27 years year NNS 37602 457 28 for for IN 37602 457 29 _ _ NNP 37602 457 30 leprieurii leprieurii JJ 37602 457 31 _ _ NNP 37602 457 32 ; ; : 37602 457 33 45 45 CD 37602 457 34 for for IN 37602 457 35 _ _ NNP 37602 457 36 britti britti NNP 37602 457 37 _ _ NNP 37602 457 38 ) ) -RRB- 37602 457 39 . . . 37602 458 1 Comparisons comparison NNS 37602 458 2 of of IN 37602 458 3 the the DT 37602 458 4 holotypes holotype NNS 37602 458 5 with with IN 37602 458 6 series series NN 37602 458 7 of of IN 37602 458 8 specimens specimen NNS 37602 458 9 from from IN 37602 458 10 Ecuador Ecuador NNP 37602 458 11 , , , 37602 458 12 Guyana Guyana NNP 37602 458 13 , , , 37602 458 14 and and CC 37602 458 15 Surinam Surinam NNP 37602 458 16 indicate indicate VBP 37602 458 17 that that IN 37602 458 18 one one CD 37602 458 19 morphological morphological JJ 37602 458 20 species specie NNS 37602 458 21 occurs occur VBZ 37602 458 22 throughout throughout IN 37602 458 23 the the DT 37602 458 24 upper upper JJ 37602 458 25 Amazon Amazon NNP 37602 458 26 Basin Basin NNP 37602 458 27 and and CC 37602 458 28 the the DT 37602 458 29 Guianas Guianas NNPS 37602 458 30 and and CC 37602 458 31 that that IN 37602 458 32 both both DT 37602 458 33 type type NN 37602 458 34 specimens specimen NNS 37602 458 35 are be VBP 37602 458 36 representatives representative NNS 37602 458 37 of of IN 37602 458 38 one one CD 37602 458 39 species species NN 37602 458 40 . . . 37602 459 1 Consequently consequently RB 37602 459 2 , , , 37602 459 3 we -PRON- PRP 37602 459 4 consider consider VBP 37602 459 5 _ _ NNP 37602 459 6 Hyla Hyla NNP 37602 459 7 leprieurii leprieurii JJ 37602 459 8 _ _ NNP 37602 459 9 Duméril Duméril NNP 37602 459 10 and and CC 37602 459 11 Bibron Bibron NNP 37602 459 12 , , , 37602 459 13 1841 1841 CD 37602 459 14 , , , 37602 459 15 to to TO 37602 459 16 be be VB 37602 459 17 a a DT 37602 459 18 monotypic monotypic JJ 37602 459 19 species specie NNS 37602 459 20 with with IN 37602 459 21 _ _ NNP 37602 459 22 Hyla Hyla NNP 37602 459 23 leprieurii leprieurii JJ 37602 459 24 britti britti NNP 37602 459 25 _ _ NNP 37602 459 26 Melin Melin NNP 37602 459 27 , , , 37602 459 28 1941 1941 CD 37602 459 29 , , , 37602 459 30 as as IN 37602 459 31 a a DT 37602 459 32 junior junior JJ 37602 459 33 synonym synonym NN 37602 459 34 . . . 37602 460 1 In in IN 37602 460 2 their -PRON- PRP$ 37602 460 3 account account NN 37602 460 4 of of IN 37602 460 5 _ _ NNP 37602 460 6 Osteocephalus Osteocephalus NNP 37602 460 7 leprieurii leprieurii JJ 37602 460 8 _ _ NNP 37602 460 9 , , , 37602 460 10 Cochran Cochran NNP 37602 460 11 and and CC 37602 460 12 Goin go NNP 37602 460 13 ( ( -LRB- 37602 460 14 1970:323 1970:323 CD 37602 460 15 ) ) -RRB- 37602 460 16 stated state VBD 37602 460 17 : : : 37602 460 18 " " `` 37602 460 19 The the DT 37602 460 20 specimen speciman NNS 37602 460 21 described describe VBD 37602 460 22 and and CC 37602 460 23 illustrated illustrate VBN 37602 460 24 ( ( -LRB- 37602 460 25 MCZ MCZ NNP 37602 460 26 28042 28042 CD 37602 460 27 ) ) -RRB- 37602 460 28 has have VBZ 37602 460 29 been be VBN 37602 460 30 directly directly RB 37602 460 31 compared compare VBN 37602 460 32 with with IN 37602 460 33 the the DT 37602 460 34 types type NNS 37602 460 35 of of IN 37602 460 36 _ _ NNP 37602 460 37 leprieurii leprieurii JJ 37602 460 38 _ _ NNP 37602 460 39 , , , 37602 460 40 _ _ NNP 37602 460 41 planiceps planicep NNS 37602 460 42 _ _ NNP 37602 460 43 , , , 37602 460 44 and and CC 37602 460 45 _ _ NNP 37602 460 46 vilarsi vilarsi NN 37602 460 47 _ _ NNP 37602 460 48 by by IN 37602 460 49 the the DT 37602 460 50 junior junior JJ 37602 460 51 author author NN 37602 460 52 and and CC 37602 460 53 there there EX 37602 460 54 seems seem VBZ 37602 460 55 to to TO 37602 460 56 be be VB 37602 460 57 no no DT 37602 460 58 doubt doubt NN 37602 460 59 that that IN 37602 460 60 all all DT 37602 460 61 are be VBP 37602 460 62 conspecific conspecific NN 37602 460 63 . . . 37602 461 1 Another another DT 37602 461 2 specimen speciman NNS 37602 461 3 ( ( -LRB- 37602 461 4 CNHM CNHM NNP 37602 461 5 69716 69716 CD 37602 461 6 ) ) -RRB- 37602 461 7 has have VBZ 37602 461 8 been be VBN 37602 461 9 directly directly RB 37602 461 10 compared compare VBN 37602 461 11 with with IN 37602 461 12 the the DT 37602 461 13 types type NNS 37602 461 14 of of IN 37602 461 15 _ _ NNP 37602 461 16 planiceps planicep NNS 37602 461 17 _ _ NNP 37602 461 18 and and CC 37602 461 19 _ _ NNP 37602 461 20 vilarsi vilarsi NN 37602 461 21 _ _ NNP 37602 461 22 and and CC 37602 461 23 these these DT 37602 461 24 , , , 37602 461 25 likewise likewise RB 37602 461 26 , , , 37602 461 27 are be VBP 37602 461 28 considered consider VBN 37602 461 29 conspecific conspecific NN 37602 461 30 . . . 37602 461 31 " " '' 37602 462 1 With with IN 37602 462 2 this this DT 37602 462 3 justification justification NN 37602 462 4 Cochran Cochran NNP 37602 462 5 and and CC 37602 462 6 Goin go NNP 37602 462 7 ( ( -LRB- 37602 462 8 1970:322 1970:322 CD 37602 462 9 ) ) -RRB- 37602 462 10 included include VBD 37602 462 11 _ _ NNP 37602 462 12 Osteocephalus Osteocephalus NNP 37602 462 13 planiceps planicep NNS 37602 462 14 _ _ NNP 37602 462 15 Cope Cope NNP 37602 462 16 , , , 37602 462 17 1874 1874 CD 37602 462 18 , , , 37602 462 19 and and CC 37602 462 20 _ _ NNP 37602 462 21 Hyla Hyla NNP 37602 462 22 vilarsi vilarsi NN 37602 462 23 _ _ NNP 37602 462 24 Melin Melin NNP 37602 462 25 , , , 37602 462 26 1941 1941 CD 37602 462 27 , , , 37602 462 28 in in IN 37602 462 29 the the DT 37602 462 30 synonymy synonymy NN 37602 462 31 of of IN 37602 462 32 _ _ NNP 37602 462 33 Osteocephalus Osteocephalus NNP 37602 462 34 leprieurii leprieurii JJ 37602 462 35 _ _ NNP 37602 462 36 . . . 37602 463 1 We -PRON- PRP 37602 463 2 do do VBP 37602 463 3 not not RB 37602 463 4 concur concur VB 37602 463 5 with with IN 37602 463 6 Cochran Cochran NNP 37602 463 7 and and CC 37602 463 8 Goin go NNP 37602 463 9 's 's POS 37602 463 10 synonymy synonymy NN 37602 463 11 and and CC 37602 463 12 contend contend VBP 37602 463 13 that that IN 37602 463 14 _ _ NNP 37602 463 15 planiceps planicep NNS 37602 463 16 _ _ NNP 37602 463 17 and and CC 37602 463 18 _ _ NNP 37602 463 19 vilarsi vilarsi NN 37602 463 20 _ _ NNP 37602 463 21 are be VBP 37602 463 22 synonyms synonyms NN 37602 463 23 of of IN 37602 463 24 _ _ NNP 37602 463 25 Osteocephalus Osteocephalus NNP 37602 463 26 taurinus taurinus NNP 37602 463 27 _ _ NNP 37602 463 28 ; ; : 37602 463 29 we -PRON- PRP 37602 463 30 give give VBP 37602 463 31 our -PRON- PRP$ 37602 463 32 reasons reason NNS 37602 463 33 in in IN 37602 463 34 the the DT 37602 463 35 account account NN 37602 463 36 of of IN 37602 463 37 that that DT 37602 463 38 species specie NNS 37602 463 39 . . . 37602 464 1 We -PRON- PRP 37602 464 2 have have VBP 37602 464 3 examined examine VBN 37602 464 4 the the DT 37602 464 5 specimens specimen NNS 37602 464 6 listed list VBN 37602 464 7 as as IN 37602 464 8 _ _ NNP 37602 464 9 O. O. NNP 37602 465 1 leprieurii leprieurii JJ 37602 465 2 _ _ NNP 37602 465 3 by by IN 37602 465 4 Cochran Cochran NNP 37602 465 5 and and CC 37602 465 6 Goin go NNP 37602 465 7 ; ; : 37602 465 8 several several JJ 37602 465 9 of of IN 37602 465 10 them -PRON- PRP 37602 465 11 , , , 37602 465 12 including include VBG 37602 465 13 CNHM cnhm NN 37602 465 14 (= (= NNP 37602 465 15 FMNH FMNH NNP 37602 465 16 ) ) -RRB- 37602 465 17 69716 69716 CD 37602 465 18 , , , 37602 465 19 are be VBP 37602 465 20 _ _ NNP 37602 465 21 taurinus taurinus NN 37602 465 22 _ _ NNP 37602 465 23 . . . 37602 466 1 Thus thus RB 37602 466 2 , , , 37602 466 3 due due IN 37602 466 4 to to IN 37602 466 5 Cochran Cochran NNP 37602 466 6 and and CC 37602 466 7 Goin go NNP 37602 466 8 's 's POS 37602 466 9 confusion confusion NN 37602 466 10 of of IN 37602 466 11 two two CD 37602 466 12 taxa taxa NN 37602 466 13 , , , 37602 466 14 their -PRON- PRP$ 37602 466 15 comparisons comparison NNS 37602 466 16 of of IN 37602 466 17 certain certain JJ 37602 466 18 specimens specimen NNS 37602 466 19 with with IN 37602 466 20 types type NNS 37602 466 21 has have VBZ 37602 466 22 little little JJ 37602 466 23 meaning meaning NN 37602 466 24 . . . 37602 467 1 Cochran Cochran NNP 37602 467 2 and and CC 37602 467 3 Goin go NNP 37602 467 4 did do VBD 37602 467 5 not not RB 37602 467 6 include include VB 37602 467 7 _ _ NNP 37602 467 8 Hyla Hyla NNP 37602 467 9 leprieurii leprieurii JJ 37602 467 10 britti britti NNP 37602 467 11 _ _ NNP 37602 467 12 in in IN 37602 467 13 their -PRON- PRP$ 37602 467 14 synonymy synonymy NN 37602 467 15 of of IN 37602 467 16 _ _ NNP 37602 467 17 Osteocephalus Osteocephalus NNP 37602 467 18 leprieurii leprieurii JJ 37602 467 19 _ _ NNP 37602 467 20 but but CC 37602 467 21 did do VBD 37602 467 22 discuss discuss VB 37602 467 23 the the DT 37602 467 24 name name NN 37602 467 25 in in IN 37602 467 26 their -PRON- PRP$ 37602 467 27 account account NN 37602 467 28 of of IN 37602 467 29 _ _ NNP 37602 467 30 Osteocephalus Osteocephalus NNP 37602 467 31 orcesi orcesi NN 37602 467 32 _ _ NNP 37602 467 33 (= (= NNP 37602 467 34 _ _ NNP 37602 467 35 O. O. NNP 37602 468 1 verrucigerus verrucigerus NNP 37602 468 2 _ _ NNP 37602 468 3 ) ) -RRB- 37602 468 4 , , , 37602 468 5 as as IN 37602 468 6 follows follow VBZ 37602 468 7 ( ( -LRB- 37602 468 8 1970:319 1970:319 CD 37602 468 9 ) ) -RRB- 37602 468 10 : : : 37602 468 11 " " `` 37602 468 12 When when WRB 37602 468 13 we -PRON- PRP 37602 468 14 first first RB 37602 468 15 examined examine VBD 37602 468 16 one one CD 37602 468 17 of of IN 37602 468 18 the the DT 37602 468 19 specimens specimen NNS 37602 468 20 we -PRON- PRP 37602 468 21 felt feel VBD 37602 468 22 sure sure JJ 37602 468 23 that that IN 37602 468 24 we -PRON- PRP 37602 468 25 had have VBD 37602 468 26 Melin Melin NNP 37602 468 27 's 's POS 37602 468 28 _ _ NNP 37602 468 29 Hyla Hyla NNP 37602 468 30 britti britti NNP 37602 468 31 _ _ NNP 37602 468 32 at at IN 37602 468 33 hand hand NN 37602 468 34 , , , 37602 468 35 but but CC 37602 468 36 on on IN 37602 468 37 direct direct JJ 37602 468 38 comparison comparison NN 37602 468 39 with with IN 37602 468 40 the the DT 37602 468 41 type type NN 37602 468 42 of of IN 37602 468 43 _ _ NNP 37602 468 44 britti britti NNP 37602 468 45 _ _ NNP 37602 468 46 the the DT 37602 468 47 two two CD 37602 468 48 proved prove VBD 37602 468 49 to to TO 37602 468 50 be be VB 37602 468 51 different different JJ 37602 468 52 . . . 37602 469 1 After after IN 37602 469 2 studying study VBG 37602 469 3 the the DT 37602 469 4 type type NN 37602 469 5 of of IN 37602 469 6 _ _ NNP 37602 469 7 orcesi orcesi NNP 37602 469 8 _ _ NNP 37602 469 9 ( ( -LRB- 37602 469 10 SUNHM SUNHM NNP 37602 469 11 13150 13150 CD 37602 469 12 ) ) -RRB- 37602 469 13 we -PRON- PRP 37602 469 14 have have VBP 37602 469 15 no no DT 37602 469 16 doubt doubt NN 37602 469 17 that that IN 37602 469 18 the the DT 37602 469 19 specimens specimen NNS 37602 469 20 at at IN 37602 469 21 hand hand NN 37602 469 22 are be VBP 37602 469 23 _ _ NNP 37602 469 24 orcesi orcesi NNP 37602 469 25 _ _ NNP 37602 469 26 and and CC 37602 469 27 that that IN 37602 469 28 _ _ NNP 37602 469 29 britti britti NNP 37602 469 30 _ _ NNP 37602 469 31 is be VBZ 37602 469 32 a a DT 37602 469 33 different different JJ 37602 469 34 , , , 37602 469 35 probably probably RB 37602 469 36 valid valid JJ 37602 469 37 species specie NNS 37602 469 38 . . . 37602 469 39 " " '' 37602 470 1 _ _ NNP 37602 470 2 Diagnosis._--1 Diagnosis._--1 NNP 37602 470 3 ) ) -RRB- 37602 470 4 Size size NN 37602 470 5 moderate moderate JJ 37602 470 6 , , , 37602 470 7 sexual sexual JJ 37602 470 8 dimorphism dimorphism NN 37602 470 9 evident evident NN 37602 470 10 ; ; : 37602 470 11 maximum maximum JJ 37602 470 12 observed observed JJ 37602 470 13 snout snout NN 37602 470 14 - - HYPH 37602 470 15 vent vent NN 37602 470 16 length length NN 37602 470 17 in in IN 37602 470 18 males male NNS 37602 470 19 48.4 48.4 CD 37602 470 20 mm mm NNS 37602 470 21 , , , 37602 470 22 in in IN 37602 470 23 females female NNS 37602 470 24 , , , 37602 470 25 61.5 61.5 CD 37602 470 26 mm mm CD 37602 470 27 ; ; : 37602 470 28 2 2 LS 37602 470 29 ) ) -RRB- 37602 470 30 skin skin NN 37602 470 31 on on IN 37602 470 32 dorsum dorsum NN 37602 470 33 in in IN 37602 470 34 males male NNS 37602 470 35 bearing bear VBG 37602 470 36 numerous numerous JJ 37602 470 37 , , , 37602 470 38 minute minute NN 37602 470 39 , , , 37602 470 40 spinous spinous JJ 37602 470 41 tubercles tubercle NNS 37602 470 42 ; ; : 37602 470 43 3 3 LS 37602 470 44 ) ) -RRB- 37602 470 45 skin skin NN 37602 470 46 on on IN 37602 470 47 flanks flank NNS 37602 470 48 smooth smooth VBP 37602 470 49 ; ; : 37602 470 50 4 4 LS 37602 470 51 ) ) -RRB- 37602 470 52 web web NN 37602 470 53 extending extend VBG 37602 470 54 to to IN 37602 470 55 base base NN 37602 470 56 of of IN 37602 470 57 antepenultimate antepenultimate NN 37602 470 58 phalange phalange NN 37602 470 59 on on IN 37602 470 60 inner inner JJ 37602 470 61 edge edge NN 37602 470 62 of of IN 37602 470 63 third third JJ 37602 470 64 finger finger NN 37602 470 65 ; ; : 37602 470 66 5 5 LS 37602 470 67 ) ) -RRB- 37602 470 68 dorsum dorsum NN 37602 470 69 tan tan NN 37602 470 70 or or CC 37602 470 71 olive olive NN 37602 470 72 - - HYPH 37602 470 73 brown brown JJ 37602 470 74 with with IN 37602 470 75 transverse transverse NNP 37602 470 76 brown brown JJ 37602 470 77 or or CC 37602 470 78 olive olive NN 37602 470 79 bars bar NNS 37602 470 80 ; ; : 37602 470 81 6 6 LS 37602 470 82 ) ) -RRB- 37602 470 83 venter venter NN 37602 470 84 creamy creamy JJ 37602 470 85 white white JJ 37602 470 86 or or CC 37602 470 87 pale pale JJ 37602 470 88 tan tan NN 37602 470 89 without without IN 37602 470 90 markings marking NNS 37602 470 91 ; ; : 37602 470 92 7 7 LS 37602 470 93 ) ) -RRB- 37602 470 94 lips lip NNS 37602 470 95 marked mark VBN 37602 470 96 with with IN 37602 470 97 creamy creamy NNP 37602 470 98 tan tan NNP 37602 470 99 labial labial NNP 37602 470 100 stripe stripe NN 37602 470 101 and and CC 37602 470 102 suborbital suborbital JJ 37602 470 103 spot spot NN 37602 470 104 ; ; : 37602 470 105 8) 8) CD 37602 470 106 flanks flank VBZ 37602 470 107 pale pale JJ 37602 470 108 tan tan NNP 37602 470 109 with with IN 37602 470 110 no no DT 37602 470 111 markings marking NNS 37602 470 112 ; ; : 37602 470 113 9 9 LS 37602 470 114 ) ) -RRB- 37602 470 115 dermal dermal JJ 37602 470 116 roofing roofing NN 37602 470 117 bones bone NNS 37602 470 118 of of IN 37602 470 119 skull skull NN 37602 470 120 lacking lacking NN 37602 470 121 exostosis exostosis NN 37602 470 122 ; ; : 37602 470 123 10 10 LS 37602 470 124 ) ) -RRB- 37602 470 125 dermal dermal JJ 37602 470 126 sphenethmoid sphenethmoid JJ 37602 470 127 absent absent NN 37602 470 128 ; ; : 37602 470 129 11 11 CD 37602 470 130 ) ) -RRB- 37602 470 131 nasals nasal NNS 37602 470 132 juxtaposed juxtapose VBD 37602 470 133 medially medially RB 37602 470 134 ; ; : 37602 470 135 12 12 CD 37602 470 136 ) ) -RRB- 37602 470 137 anteromedial anteromedial JJ 37602 470 138 margin margin NN 37602 470 139 of of IN 37602 470 140 frontoparietal frontoparietal NN 37602 470 141 between between IN 37602 470 142 mid- mid- NN 37602 470 143 and and CC 37602 470 144 anterior anterior NN 37602 470 145 levels level NNS 37602 470 146 of of IN 37602 470 147 orbit orbit NN 37602 470 148 ; ; : 37602 470 149 13 13 CD 37602 470 150 ) ) -RRB- 37602 470 151 frontoparietal frontoparietal JJ 37602 470 152 fontanelle fontanelle NN 37602 470 153 partially partially RB 37602 470 154 exposed expose VBN 37602 470 155 ; ; : 37602 470 156 14 14 CD 37602 470 157 ) ) -RRB- 37602 470 158 palatine palatine NN 37602 470 159 not not RB 37602 470 160 serrate serrate NN 37602 470 161 ; ; : 37602 470 162 15 15 CD 37602 470 163 ) ) -RRB- 37602 470 164 parasphenoid parasphenoid NN 37602 470 165 lacking lacking NN 37602 470 166 odontoids odontoid NNS 37602 470 167 ; ; : 37602 470 168 16 16 LS 37602 470 169 ) ) -RRB- 37602 470 170 zygomatic zygomatic JJ 37602 470 171 ramus ramus NN 37602 470 172 of of IN 37602 470 173 squamosal squamosal NNP 37602 470 174 extending extend VBG 37602 470 175 about about RB 37602 470 176 one one CD 37602 470 177 - - HYPH 37602 470 178 half half NN 37602 470 179 of of IN 37602 470 180 distance distance NN 37602 470 181 to to IN 37602 470 182 maxillary maxillary NNP 37602 470 183 arch arch NNP 37602 470 184 ; ; : 37602 470 185 17 17 CD 37602 470 186 ) ) -RRB- 37602 470 187 transverse transverse JJ 37602 470 188 processes process NNS 37602 470 189 of of IN 37602 470 190 presacral presacral JJ 37602 470 191 vertebrae vertebra NNS 37602 470 192 3 3 CD 37602 470 193 - - SYM 37602 470 194 8 8 CD 37602 470 195 about about RB 37602 470 196 equal equal JJ 37602 470 197 in in IN 37602 470 198 width width NN 37602 470 199 to to IN 37602 470 200 one one CD 37602 470 201 another another DT 37602 470 202 and and CC 37602 470 203 to to IN 37602 470 204 sacral sacral JJ 37602 470 205 diapophyses diapophysis NNS 37602 470 206 ; ; : 37602 470 207 18 18 CD 37602 470 208 ) ) -RRB- 37602 470 209 intermandibularis intermandibularis NNP 37602 470 210 and and CC 37602 470 211 submentalis submentalis NNP 37602 470 212 muscles muscle NNS 37602 470 213 connected connect VBD 37602 470 214 ; ; : 37602 470 215 19 19 CD 37602 470 216 ) ) -RRB- 37602 470 217 supramandibular supramandibular JJ 37602 470 218 portion portion NN 37602 470 219 of of IN 37602 470 220 interhyoideus interhyoideu NNS 37602 470 221 forming form VBG 37602 470 222 simple simple JJ 37602 470 223 tubular tubular NNP 37602 470 224 posterolateral posterolateral JJ 37602 470 225 extension extension NN 37602 470 226 ; ; , 37602 470 227 associated associate VBN 37602 470 228 skin skin NN 37602 470 229 unmodified unmodifie VBD 37602 470 230 . . . 37602 471 1 _ _ NNP 37602 471 2 Osteocephalus Osteocephalus NNP 37602 471 3 leprieurii leprieurii VBZ 37602 471 4 _ _ NNP 37602 471 5 differs differ VBZ 37602 471 6 from from IN 37602 471 7 all all DT 37602 471 8 other other JJ 37602 471 9 members member NNS 37602 471 10 of of IN 37602 471 11 the the DT 37602 471 12 genus genus NN 37602 471 13 by by IN 37602 471 14 having have VBG 37602 471 15 transverse transverse JJ 37602 471 16 dark dark JJ 37602 471 17 bars bar NNS 37602 471 18 on on IN 37602 471 19 the the DT 37602 471 20 back back NN 37602 471 21 . . . 37602 472 1 Two two CD 37602 472 2 other other JJ 37602 472 3 hylids hylid NNS 37602 472 4 ( ( -LRB- 37602 472 5 _ _ NNP 37602 472 6 Hyla Hyla NNP 37602 472 7 lanciformis lanciformis NN 37602 472 8 _ _ NNP 37602 472 9 and and CC 37602 472 10 _ _ NNP 37602 472 11 multifasciata multifasciata NN 37602 472 12 _ _ NNP 37602 472 13 ) ) -RRB- 37602 472 14 in in IN 37602 472 15 the the DT 37602 472 16 Amazon Amazon NNP 37602 472 17 Basin Basin NNP 37602 472 18 have have VBP 37602 472 19 transverse transverse JJ 37602 472 20 dark dark JJ 37602 472 21 marks mark NNS 37602 472 22 on on IN 37602 472 23 the the DT 37602 472 24 dorsum dorsum NN 37602 472 25 . . . 37602 473 1 Both both DT 37602 473 2 of of IN 37602 473 3 these these DT 37602 473 4 differ differ VBP 37602 473 5 from from IN 37602 473 6 _ _ NNP 37602 473 7 leprieurii leprieurii JJ 37602 473 8 _ _ NNP 37602 473 9 by by IN 37602 473 10 having have VBG 37602 473 11 pointed point VBN 37602 473 12 snouts snout NNS 37602 473 13 , , , 37602 473 14 much much RB 37602 473 15 longer long RBR 37602 473 16 hind hind VBP 37602 473 17 limbs limb NNS 37602 473 18 , , , 37602 473 19 and and CC 37602 473 20 smooth smooth JJ 37602 473 21 skin skin NN 37602 473 22 dorsally dorsally RB 37602 473 23 . . . 37602 474 1 _ _ NNP 37602 474 2 Distribution._--The Distribution._--The NNP 37602 474 3 periphery periphery NN 37602 474 4 of of IN 37602 474 5 the the DT 37602 474 6 Amazon Amazon NNP 37602 474 7 Basin Basin NNP 37602 474 8 , , , 37602 474 9 in in IN 37602 474 10 the the DT 37602 474 11 Guianas Guianas NNP 37602 474 12 and and CC 37602 474 13 the the DT 37602 474 14 upper upper JJ 37602 474 15 part part NN 37602 474 16 of of IN 37602 474 17 the the DT 37602 474 18 basin basin NN 37602 474 19 in in IN 37602 474 20 southern southern JJ 37602 474 21 Colombia Colombia NNP 37602 474 22 , , , 37602 474 23 Ecuador Ecuador NNP 37602 474 24 , , , 37602 474 25 Perú Perú NNP 37602 474 26 , , , 37602 474 27 and and CC 37602 474 28 extreme extreme JJ 37602 474 29 western western JJ 37602 474 30 Brasil Brasil NNP 37602 474 31 ( ( -LRB- 37602 474 32 Fig fig NN 37602 474 33 . . . 37602 475 1 9 9 LS 37602 475 2 ) ) -RRB- 37602 475 3 . . . 37602 476 1 Most Most JJS 37602 476 2 localities locality NNS 37602 476 3 are be VBP 37602 476 4 at at IN 37602 476 5 elevations elevation NNS 37602 476 6 of of IN 37602 476 7 less less JJR 37602 476 8 than than IN 37602 476 9 500 500 CD 37602 476 10 m m NN 37602 476 11 , , , 37602 476 12 but but CC 37602 476 13 the the DT 37602 476 14 species species NN 37602 476 15 ascends ascend VBZ 37602 476 16 the the DT 37602 476 17 lower low JJR 37602 476 18 Andean andean JJ 37602 476 19 slopes slope NNS 37602 476 20 to to IN 37602 476 21 elevations elevation NNS 37602 476 22 of of IN 37602 476 23 1100 1100 CD 37602 476 24 m. m. NN 37602 476 25 265 265 CD 37602 476 26 specimens specimen NNS 37602 476 27 from from IN 37602 476 28 31 31 CD 37602 476 29 localities locality NNS 37602 476 30 . . . 37602 477 1 [ [ -LRB- 37602 477 2 Illustration illustration NN 37602 477 3 : : : 37602 477 4 FIG FIG NNP 37602 477 5 . . . 37602 478 1 9 9 CD 37602 478 2 . . . 37602 479 1 Distribution distribution NN 37602 479 2 of of IN 37602 479 3 _ _ NNP 37602 479 4 Osteocephalus Osteocephalus NNP 37602 479 5 leprieurii leprieurii JJ 37602 479 6 _ _ NNP 37602 479 7 ( ( -LRB- 37602 479 8 circles circles NNP 37602 479 9 ) ) -RRB- 37602 479 10 and and CC 37602 479 11 _ _ NNP 37602 479 12 O. O. NNP 37602 480 1 verrucigerus verrucigerus NNP 37602 480 2 _ _ NNP 37602 480 3 ( ( -LRB- 37602 480 4 triangles triangle NNS 37602 480 5 ) ) -RRB- 37602 480 6 . . . 37602 480 7 ] ] -RRB- 37602 481 1 _ _ NNP 37602 481 2 Remarks._--Most remarks._--most CD 37602 481 3 adults adult NNS 37602 481 4 of of IN 37602 481 5 _ _ NNP 37602 481 6 leprieurii leprieurii JJ 37602 481 7 _ _ NNP 37602 481 8 have have VBP 37602 481 9 distinct distinct JJ 37602 481 10 transverse transverse JJ 37602 481 11 markings marking NNS 37602 481 12 on on IN 37602 481 13 the the DT 37602 481 14 back back NN 37602 481 15 ; ; : 37602 481 16 these these DT 37602 481 17 are be VBP 37602 481 18 variable variable JJ 37602 481 19 in in IN 37602 481 20 width width NN 37602 481 21 , , , 37602 481 22 extent extent NN 37602 481 23 , , , 37602 481 24 and and CC 37602 481 25 arrangement arrangement NN 37602 481 26 . . . 37602 482 1 In in IN 37602 482 2 some some DT 37602 482 3 specimens specimen NNS 37602 482 4 , , , 37602 482 5 such such JJ 37602 482 6 as as IN 37602 482 7 USNM USNM NNP 37602 482 8 166557 166557 CD 37602 482 9 , , , 37602 482 10 some some DT 37602 482 11 of of IN 37602 482 12 the the DT 37602 482 13 transverse transverse JJ 37602 482 14 bars bar NNS 37602 482 15 are be VBP 37602 482 16 fragmented fragment VBN 37602 482 17 into into IN 37602 482 18 spots spot NNS 37602 482 19 ; ; : 37602 482 20 in in IN 37602 482 21 a a DT 37602 482 22 few few JJ 37602 482 23 specimens specimen NNS 37602 482 24 the the DT 37602 482 25 dorsal dorsal NN 37602 482 26 pattern pattern NN 37602 482 27 consists consist VBZ 37602 482 28 solely solely RB 37602 482 29 of of IN 37602 482 30 small small JJ 37602 482 31 dark dark JJ 37602 482 32 spots spot NNS 37602 482 33 arranged arrange VBN 37602 482 34 in in IN 37602 482 35 transverse transverse JJ 37602 482 36 rows row NNS 37602 482 37 . . . 37602 483 1 Such such JJ 37602 483 2 specimens specimen NNS 37602 483 3 have have VBP 37602 483 4 a a DT 37602 483 5 dorsal dorsal NN 37602 483 6 pattern pattern NN 37602 483 7 resembling resembling NN 37602 483 8 that that DT 37602 483 9 of of IN 37602 483 10 some some DT 37602 483 11 _ _ NNP 37602 483 12 taurinus taurinus NN 37602 483 13 _ _ NNP 37602 483 14 . . . 37602 484 1 The the DT 37602 484 2 transverse transverse JJ 37602 484 3 nature nature NN 37602 484 4 of of IN 37602 484 5 the the DT 37602 484 6 dorsal dorsal NN 37602 484 7 markings marking NNS 37602 484 8 is be VBZ 37602 484 9 further further RB 37602 484 10 modified modify VBN 37602 484 11 in in IN 37602 484 12 some some DT 37602 484 13 specimens specimen NNS 37602 484 14 , , , 37602 484 15 such such JJ 37602 484 16 as as IN 37602 484 17 USNM USNM NNP 37602 484 18 166555 166555 CD 37602 484 19 , , , 37602 484 20 in in IN 37602 484 21 which which WDT 37602 484 22 the the DT 37602 484 23 dark dark JJ 37602 484 24 bars bar NNS 37602 484 25 are be VBP 37602 484 26 fragmented fragmented JJ 37602 484 27 and and CC 37602 484 28 oblique oblique JJ 37602 484 29 . . . 37602 485 1 Extreme extreme JJ 37602 485 2 ontogenetic ontogenetic JJ 37602 485 3 change change NN 37602 485 4 in in IN 37602 485 5 color color NN 37602 485 6 pattern pattern NN 37602 485 7 is be VBZ 37602 485 8 exhibited exhibit VBN 37602 485 9 by by IN 37602 485 10 this this DT 37602 485 11 species species NN 37602 485 12 ( ( -LRB- 37602 485 13 Fig fig NN 37602 485 14 . . . 37602 486 1 10 10 LS 37602 486 2 ) ) -RRB- 37602 486 3 . . . 37602 487 1 Juveniles juvenile NNS 37602 487 2 having have VBG 37602 487 3 snout snout NN 37602 487 4 - - HYPH 37602 487 5 vent vent NN 37602 487 6 lengths length NNS 37602 487 7 of of IN 37602 487 8 less less JJR 37602 487 9 than than IN 37602 487 10 28 28 CD 37602 487 11 mm mm NNS 37602 487 12 have have VBP 37602 487 13 an an DT 37602 487 14 olive olive JJ 37602 487 15 - - HYPH 37602 487 16 brown brown JJ 37602 487 17 dorsum dorsum NN 37602 487 18 with with IN 37602 487 19 a a DT 37602 487 20 pale pale JJ 37602 487 21 cream cream NN 37602 487 22 stripe stripe NN 37602 487 23 across across IN 37602 487 24 the the DT 37602 487 25 head head NN 37602 487 26 and and CC 37602 487 27 broad broad JJ 37602 487 28 , , , 37602 487 29 cream cream NN 37602 487 30 , , , 37602 487 31 dorsolateral dorsolateral JJ 37602 487 32 stripes stripe NNS 37602 487 33 ; ; : 37602 487 34 transverse transverse JJ 37602 487 35 dark dark JJ 37602 487 36 bars bar NNS 37602 487 37 are be VBP 37602 487 38 absent absent JJ 37602 487 39 on on IN 37602 487 40 the the DT 37602 487 41 body body NN 37602 487 42 and and CC 37602 487 43 limbs limb NNS 37602 487 44 . . . 37602 488 1 Individuals individual NNS 37602 488 2 having have VBG 37602 488 3 snout snout NN 37602 488 4 - - HYPH 37602 488 5 vent vent NN 37602 488 6 lengths length NNS 37602 488 7 of of IN 37602 488 8 30 30 CD 37602 488 9 - - SYM 37602 488 10 35 35 CD 37602 488 11 mm mm CD 37602 488 12 have have VBP 37602 488 13 dark dark JJ 37602 488 14 brown brown JJ 37602 488 15 transverse transverse JJ 37602 488 16 bars bar NNS 37602 488 17 on on IN 37602 488 18 the the DT 37602 488 19 back back NN 37602 488 20 and and CC 37602 488 21 limbs limb NNS 37602 488 22 but but CC 37602 488 23 still still RB 37602 488 24 retain retain VB 37602 488 25 the the DT 37602 488 26 light light JJ 37602 488 27 dorsolateral dorsolateral JJ 37602 488 28 stripes stripe NNS 37602 488 29 , , , 37602 488 30 whereas whereas IN 37602 488 31 the the DT 37602 488 32 stripes stripe NNS 37602 488 33 are be VBP 37602 488 34 lost lose VBN 37602 488 35 in in IN 37602 488 36 larger large JJR 37602 488 37 individuals individual NNS 37602 488 38 . . . 37602 489 1 [ [ -LRB- 37602 489 2 Illustration illustration NN 37602 489 3 : : : 37602 489 4 FIG FIG NNP 37602 489 5 . . . 37602 490 1 10 10 CD 37602 490 2 . . . 37602 491 1 Ontogenetic ontogenetic JJ 37602 491 2 change change NN 37602 491 3 in in IN 37602 491 4 color color NN 37602 491 5 pattern pattern NN 37602 491 6 in in IN 37602 491 7 _ _ NNP 37602 491 8 Osteocephalus Osteocephalus NNP 37602 491 9 leprieurii leprieurii JJ 37602 491 10 _ _ NNP 37602 491 11 : : : 37602 491 12 a. a. NN 37602 491 13 KU KU NNP 37602 491 14 126644 126644 CD 37602 491 15 ; ; : 37602 491 16 b. b. NNP 37602 491 17 KU KU NNP 37602 491 18 126640 126640 CD 37602 491 19 ; ; : 37602 491 20 c. c. NNP 37602 491 21 KU KU NNP 37602 491 22 126625 126625 CD 37602 491 23 . . . 37602 492 1 ×2 ×2 LS 37602 492 2 . . . 37602 492 3 ] ] -RRB- 37602 493 1 Coloration coloration NN 37602 493 2 in in IN 37602 493 3 life life NN 37602 493 4 of of IN 37602 493 5 specimens specimen NNS 37602 493 6 from from IN 37602 493 7 Lago Lago NNP 37602 493 8 Agrio Agrio NNP 37602 493 9 , , , 37602 493 10 Ecuador Ecuador NNP 37602 493 11 : : : 37602 493 12 " " `` 37602 493 13 In in IN 37602 493 14 males male NNS 37602 493 15 the the DT 37602 493 16 dorsal dorsal NN 37602 493 17 ground ground NN 37602 493 18 color color NN 37602 493 19 varies vary VBZ 37602 493 20 from from IN 37602 493 21 dark dark JJ 37602 493 22 brown brown NNP 37602 493 23 to to IN 37602 493 24 ochre ochre NNP 37602 493 25 - - HYPH 37602 493 26 tan tan NNP 37602 493 27 ; ; , 37602 493 28 dorsal dorsal NN 37602 493 29 markings marking NNS 37602 493 30 uniformly uniformly RB 37602 493 31 dark dark RB 37602 493 32 brown brown JJ 37602 493 33 . . . 37602 494 1 Most Most JJS 37602 494 2 specimens specimen NNS 37602 494 3 have have VBP 37602 494 4 dark dark JJ 37602 494 5 brown brown NN 37602 494 6 and and CC 37602 494 7 cream cream NN 37602 494 8 anal anal JJ 37602 494 9 stripes stripe NNS 37602 494 10 ; ; , 37602 494 11 labial labial JJ 37602 494 12 area area NN 37602 494 13 cream cream NN 37602 494 14 - - HYPH 37602 494 15 colored color VBN 37602 494 16 . . . 37602 495 1 Flanks flank NNS 37602 495 2 vary vary VBP 37602 495 3 from from IN 37602 495 4 tan tan NNP 37602 495 5 to to IN 37602 495 6 white white NNP 37602 495 7 . . . 37602 496 1 Ventral ventral JJ 37602 496 2 coloration coloration NN 37602 496 3 varies vary VBZ 37602 496 4 from from IN 37602 496 5 salmon salmon NN 37602 496 6 to to IN 37602 496 7 tan tan NNP 37602 496 8 to to IN 37602 496 9 white white NNP 37602 496 10 . . . 37602 497 1 The the DT 37602 497 2 iris iris NN 37602 497 3 is be VBZ 37602 497 4 bronze bronze NN 37602 497 5 with with IN 37602 497 6 a a DT 37602 497 7 greenish greenish JJ 37602 497 8 cast cast NN 37602 497 9 and and CC 37602 497 10 black black JJ 37602 497 11 reticulations reticulation NNS 37602 497 12 . . . 37602 498 1 In in IN 37602 498 2 females female NNS 37602 498 3 the the DT 37602 498 4 dorsal dorsal NN 37602 498 5 coloration coloration NN 37602 498 6 is be VBZ 37602 498 7 the the DT 37602 498 8 same same JJ 37602 498 9 as as IN 37602 498 10 in in IN 37602 498 11 males male NNS 37602 498 12 , , , 37602 498 13 except except IN 37602 498 14 that that DT 37602 498 15 dark dark JJ 37602 498 16 marks mark NNS 37602 498 17 tend tend VBP 37602 498 18 to to TO 37602 498 19 be be VB 37602 498 20 outlined outline VBN 37602 498 21 with with IN 37602 498 22 cream cream NN 37602 498 23 ; ; : 37602 498 24 venter venter NN 37602 498 25 tannish tannish JJ 37602 498 26 salmon salmon NN 37602 498 27 . . . 37602 498 28 " " '' 37602 499 1 ( ( -LRB- 37602 499 2 W. W. NNP 37602 499 3 E. E. NNP 37602 499 4 Duellman Duellman NNP 37602 499 5 , , , 37602 499 6 field field NN 37602 499 7 notes note NNS 37602 499 8 , , , 37602 499 9 12 12 CD 37602 499 10 May May NNP 37602 499 11 1969 1969 CD 37602 499 12 ) ) -RRB- 37602 499 13 . . . 37602 500 1 = = NFP 37602 500 2 Osteocephalus osteocephalus JJ 37602 500 3 pearsoni= pearsoni= NN 37602 500 4 ( ( -LRB- 37602 500 5 Gaige Gaige NNP 37602 500 6 ) ) -RRB- 37602 500 7 _ _ NNP 37602 500 8 Hyla Hyla NNP 37602 500 9 pearsoni pearsoni NN 37602 500 10 _ _ NNP 37602 500 11 Gaige Gaige NNP 37602 500 12 , , , 37602 500 13 1929:3 1929:3 CD 37602 500 14 [ [ -LRB- 37602 500 15 Holotype.--UMMZ holotype.--ummz NN 37602 500 16 57548 57548 CD 37602 500 17 from from IN 37602 500 18 the the DT 37602 500 19 upper upper JJ 37602 500 20 Río Río NNP 37602 500 21 Beni Beni NNP 37602 500 22 , , , 37602 500 23 below below IN 37602 500 24 mouth mouth NN 37602 500 25 of of IN 37602 500 26 Río Río NNP 37602 500 27 Mapiri Mapiri NNP 37602 500 28 , , , 37602 500 29 Departamento Departamento NNP 37602 500 30 El El NNP 37602 500 31 Beni Beni NNP 37602 500 32 , , , 37602 500 33 Bolivia Bolivia NNP 37602 500 34 ; ; : 37602 500 35 N. N. NNP 37602 500 36 E. E. NNP 37602 500 37 Pearson Pearson NNP 37602 500 38 collector collector NN 37602 500 39 ] ] -RRB- 37602 500 40 . . . 37602 501 1 _ _ NNP 37602 501 2 Osteocephalus Osteocephalus NNP 37602 501 3 pearsoni_--Goin pearsoni_--Goin NNP 37602 501 4 , , , 37602 501 5 1961:13 1961:13 CD 37602 501 6 . . . 37602 502 1 _ _ NNP 37602 502 2 Justification Justification NNP 37602 502 3 of of IN 37602 502 4 Synonymy._--Goin Synonymy._--Goin NNP 37602 502 5 ( ( -LRB- 37602 502 6 1961:13 1961:13 CD 37602 502 7 ) ) -RRB- 37602 502 8 suggested suggest VBD 37602 502 9 that that IN 37602 502 10 _ _ NNP 37602 502 11 Hyla Hyla NNP 37602 502 12 pearsoni pearsoni NN 37602 502 13 _ _ NNP 37602 502 14 Gaige Gaige NNP 37602 502 15 was be VBD 37602 502 16 an an DT 37602 502 17 _ _ NNP 37602 502 18 Osteocephalus Osteocephalus NNP 37602 502 19 _ _ NNP 37602 502 20 , , , 37602 502 21 but but CC 37602 502 22 Cochran Cochran NNP 37602 502 23 and and CC 37602 502 24 Goin go NNP 37602 502 25 ( ( -LRB- 37602 502 26 1970:217 1970:217 CD 37602 502 27 ) ) -RRB- 37602 502 28 considered consider VBD 37602 502 29 _ _ NNP 37602 502 30 pearsoni pearsoni NN 37602 502 31 _ _ NNP 37602 502 32 to to TO 37602 502 33 be be VB 37602 502 34 a a DT 37602 502 35 _ _ NNP 37602 502 36 Hyla Hyla NNP 37602 502 37 _ _ NNP 37602 502 38 . . . 37602 503 1 The the DT 37602 503 2 presence presence NN 37602 503 3 of of IN 37602 503 4 exostosed exostose VBN 37602 503 5 dermal dermal JJ 37602 503 6 roofing roofing NN 37602 503 7 bones bone NNS 37602 503 8 , , , 37602 503 9 angulate angulate VBP 37602 503 10 prevomerine prevomerine JJ 37602 503 11 dentigerous dentigerous JJ 37602 503 12 processes process NNS 37602 503 13 , , , 37602 503 14 and and CC 37602 503 15 the the DT 37602 503 16 structure structure NN 37602 503 17 of of IN 37602 503 18 the the DT 37602 503 19 vocal vocal JJ 37602 503 20 sacs sacs NN 37602 503 21 are be VBP 37602 503 22 characters character NNS 37602 503 23 which which WDT 37602 503 24 place place VBP 37602 503 25 the the DT 37602 503 26 species specie NNS 37602 503 27 in in IN 37602 503 28 _ _ NNP 37602 503 29 Osteocephalus Osteocephalus NNP 37602 503 30 _ _ NNP 37602 503 31 . . . 37602 504 1 _ _ NNP 37602 504 2 Diagnosis._--1 Diagnosis._--1 NNP 37602 504 3 ) ) -RRB- 37602 504 4 Size size NN 37602 504 5 moderate moderate JJ 37602 504 6 , , , 37602 504 7 sexual sexual JJ 37602 504 8 dimorphism dimorphism NN 37602 504 9 evident evident NN 37602 504 10 ; ; : 37602 504 11 maximum maximum JJ 37602 504 12 observed observed JJ 37602 504 13 snout snout NN 37602 504 14 - - HYPH 37602 504 15 vent vent NN 37602 504 16 length length NN 37602 504 17 in in IN 37602 504 18 males male NNS 37602 504 19 46.2 46.2 CD 37602 504 20 mm mm NNS 37602 504 21 , , , 37602 504 22 in in IN 37602 504 23 females females NNP 37602 504 24 54.7 54.7 CD 37602 504 25 mm mm NNS 37602 504 26 ; ; : 37602 504 27 2 2 LS 37602 504 28 ) ) -RRB- 37602 504 29 skin skin NN 37602 504 30 on on IN 37602 504 31 dorsum dorsum NN 37602 504 32 in in IN 37602 504 33 males male NNS 37602 504 34 bearing bear VBG 37602 504 35 a a DT 37602 504 36 few few JJ 37602 504 37 , , , 37602 504 38 small small JJ 37602 504 39 , , , 37602 504 40 scattered scatter VBN 37602 504 41 non non JJ 37602 504 42 - - JJ 37602 504 43 spinous spinous JJ 37602 504 44 tubercles tubercle NNS 37602 504 45 ; ; : 37602 504 46 3 3 LS 37602 504 47 ) ) -RRB- 37602 504 48 skin skin NN 37602 504 49 on on IN 37602 504 50 flanks flank NNS 37602 504 51 smooth smooth VBP 37602 504 52 ; ; : 37602 504 53 4 4 LS 37602 504 54 ) ) -RRB- 37602 504 55 web web NN 37602 504 56 extending extend VBG 37602 504 57 to to IN 37602 504 58 base base NN 37602 504 59 of of IN 37602 504 60 antepenultimate antepenultimate NN 37602 504 61 phalange phalange NN 37602 504 62 on on IN 37602 504 63 inner inner JJ 37602 504 64 edge edge NN 37602 504 65 of of IN 37602 504 66 third third JJ 37602 504 67 finger finger NN 37602 504 68 ; ; : 37602 504 69 5 5 LS 37602 504 70 ) ) -RRB- 37602 504 71 dorsum dorsum NN 37602 504 72 tan tan NN 37602 504 73 with with IN 37602 504 74 irregular irregular JJ 37602 504 75 brown brown JJ 37602 504 76 blotches blotch NNS 37602 504 77 ; ; : 37602 504 78 6 6 LS 37602 504 79 ) ) -RRB- 37602 504 80 venter venter NN 37602 504 81 cream cream NN 37602 504 82 with with IN 37602 504 83 fine fine JJ 37602 504 84 brown brown JJ 37602 504 85 reticulations reticulation NNS 37602 504 86 ; ; : 37602 504 87 7 7 LS 37602 504 88 ) ) -RRB- 37602 504 89 lips lip VBZ 37602 504 90 dark dark JJ 37602 504 91 with with IN 37602 504 92 pale pale JJ 37602 504 93 vertical vertical JJ 37602 504 94 bar bar NN 37602 504 95 below below IN 37602 504 96 eye eye NN 37602 504 97 ; ; : 37602 504 98 8) 8) CD 37602 504 99 flanks flank VBZ 37602 504 100 pale pale JJ 37602 504 101 tan tan NNP 37602 504 102 with with IN 37602 504 103 round round JJ 37602 504 104 , , , 37602 504 105 brown brown JJ 37602 504 106 spots spot NNS 37602 504 107 ; ; : 37602 504 108 9 9 LS 37602 504 109 ) ) -RRB- 37602 504 110 dermal dermal JJ 37602 504 111 roofing roofing NN 37602 504 112 bones bone NNS 37602 504 113 of of IN 37602 504 114 skull skull NN 37602 504 115 slightly slightly RB 37602 504 116 exostosed exostose VBD 37602 504 117 ; ; : 37602 504 118 10 10 LS 37602 504 119 ) ) -RRB- 37602 504 120 dermal dermal JJ 37602 504 121 sphenethmoid sphenethmoid JJ 37602 504 122 absent absent NN 37602 504 123 ; ; : 37602 504 124 11 11 CD 37602 504 125 ) ) -RRB- 37602 504 126 nasals nasal NNS 37602 504 127 narrowly narrowly RB 37602 504 128 separated separate VBD 37602 504 129 medially medially RB 37602 504 130 ; ; : 37602 504 131 12 12 CD 37602 504 132 ) ) -RRB- 37602 504 133 anteromedial anteromedial JJ 37602 504 134 margin margin NN 37602 504 135 of of IN 37602 504 136 frontoparietal frontoparietal NN 37602 504 137 between between IN 37602 504 138 mid- mid- NN 37602 504 139 and and CC 37602 504 140 anterior anterior NN 37602 504 141 levels level NNS 37602 504 142 of of IN 37602 504 143 orbit orbit NN 37602 504 144 ; ; : 37602 504 145 13 13 CD 37602 504 146 ) ) -RRB- 37602 504 147 frontoparietal frontoparietal JJ 37602 504 148 fontanelle fontanelle NN 37602 504 149 covered cover VBN 37602 504 150 ; ; : 37602 504 151 14 14 CD 37602 504 152 ) ) -RRB- 37602 504 153 palatine palatine NN 37602 504 154 not not RB 37602 504 155 serrate serrate NN 37602 504 156 ; ; : 37602 504 157 15 15 CD 37602 504 158 ) ) -RRB- 37602 504 159 parasphenoid parasphenoid NN 37602 504 160 lacking lacking NN 37602 504 161 odontoids odontoid NNS 37602 504 162 ; ; : 37602 504 163 16 16 LS 37602 504 164 ) ) -RRB- 37602 504 165 zygomatic zygomatic JJ 37602 504 166 ramus ramus NN 37602 504 167 of of IN 37602 504 168 squamosal squamosal NNP 37602 504 169 extending extend VBG 37602 504 170 about about RB 37602 504 171 one one CD 37602 504 172 - - HYPH 37602 504 173 half half NN 37602 504 174 distance distance NN 37602 504 175 to to IN 37602 504 176 maxillary maxillary JJ 37602 504 177 arch arch NNP 37602 504 178 ; ; : 37602 504 179 17 17 CD 37602 504 180 ) ) -RRB- 37602 504 181 transverse transverse JJ 37602 504 182 processes process NNS 37602 504 183 of of IN 37602 504 184 third third JJ 37602 504 185 presacral presacral JJ 37602 504 186 vertebra vertebra NNS 37602 504 187 approximately approximately RB 37602 504 188 equal equal JJ 37602 504 189 in in IN 37602 504 190 width width NN 37602 504 191 to to IN 37602 504 192 sacral sacral JJ 37602 504 193 diapophyses diapophysis NNS 37602 504 194 ; ; : 37602 504 195 transverse transverse JJ 37602 504 196 processes process NNS 37602 504 197 of of IN 37602 504 198 presacral presacral JJ 37602 504 199 vertebrae vertebra NNS 37602 504 200 3 3 CD 37602 504 201 - - SYM 37602 504 202 8 8 CD 37602 504 203 subequal subequal JJ 37602 504 204 in in IN 37602 504 205 width width NN 37602 504 206 ; ; : 37602 504 207 18 18 CD 37602 504 208 ) ) -RRB- 37602 504 209 intermandibularis intermandibularis NNP 37602 504 210 and and CC 37602 504 211 submentalis submentalis NNP 37602 504 212 muscles muscle NNS 37602 504 213 connected connect VBD 37602 504 214 ; ; : 37602 504 215 19 19 CD 37602 504 216 ) ) -RRB- 37602 504 217 supramandibular supramandibular JJ 37602 504 218 portion portion NN 37602 504 219 of of IN 37602 504 220 interhyoideus interhyoideus NNP 37602 504 221 extensively extensively RB 37602 504 222 developed developed JJ 37602 504 223 ; ; : 37602 504 224 associated associate VBN 37602 504 225 skin skin NN 37602 504 226 forming form VBG 37602 504 227 broad broad RB 37602 504 228 loose loose JJ 37602 504 229 fold fold NN 37602 504 230 . . . 37602 505 1 _ _ NNP 37602 505 2 Osteocephalus osteocephalus JJ 37602 505 3 pearsoni pearsoni NN 37602 505 4 _ _ NNP 37602 505 5 can can MD 37602 505 6 be be VB 37602 505 7 distinguished distinguish VBN 37602 505 8 most most RBS 37602 505 9 readily readily RB 37602 505 10 from from IN 37602 505 11 other other JJ 37602 505 12 members member NNS 37602 505 13 of of IN 37602 505 14 the the DT 37602 505 15 genus genus NN 37602 505 16 by by IN 37602 505 17 the the DT 37602 505 18 brown brown JJ 37602 505 19 reticulate reticulate NN 37602 505 20 pattern pattern NN 37602 505 21 on on IN 37602 505 22 the the DT 37602 505 23 venter venter NN 37602 505 24 , , , 37602 505 25 round round JJ 37602 505 26 brown brown JJ 37602 505 27 spots spot NNS 37602 505 28 on on IN 37602 505 29 the the DT 37602 505 30 flanks flank NNS 37602 505 31 , , , 37602 505 32 and and CC 37602 505 33 smooth smooth VB 37602 505 34 skin skin NN 37602 505 35 on on IN 37602 505 36 the the DT 37602 505 37 flanks flank NNS 37602 505 38 . . . 37602 506 1 Also also RB 37602 506 2 , , , 37602 506 3 it -PRON- PRP 37602 506 4 is be VBZ 37602 506 5 the the DT 37602 506 6 least least JJS 37602 506 7 tuberculate tuberculate NN 37602 506 8 species specie NNS 37602 506 9 in in IN 37602 506 10 the the DT 37602 506 11 genus genus NN 37602 506 12 . . . 37602 507 1 _ _ NNP 37602 507 2 Distribution._--Upper Distribution._--Upper NNP 37602 507 3 Amazon Amazon NNP 37602 507 4 Basin Basin NNP 37602 507 5 and and CC 37602 507 6 Amazonian amazonian JJ 37602 507 7 slopes slope NNS 37602 507 8 of of IN 37602 507 9 the the DT 37602 507 10 Andes Andes NNP 37602 507 11 in in IN 37602 507 12 central central JJ 37602 507 13 Perú Perú NNP 37602 507 14 ( ( -LRB- 37602 507 15 1620 1620 CD 37602 507 16 m m NNP 37602 507 17 in in IN 37602 507 18 Río Río NNP 37602 507 19 Ucayali Ucayali NNP 37602 507 20 drainage drainage NN 37602 507 21 ) ) -RRB- 37602 507 22 and and CC 37602 507 23 northern northern JJ 37602 507 24 Bolivia Bolivia NNP 37602 507 25 ( ( -LRB- 37602 507 26 less less JJR 37602 507 27 than than IN 37602 507 28 500 500 CD 37602 507 29 m m NN 37602 507 30 in in IN 37602 507 31 Río Río NNP 37602 507 32 Beni Beni NNP 37602 507 33 drainage drainage NN 37602 507 34 ) ) -RRB- 37602 507 35 ( ( -LRB- 37602 507 36 Fig fig NN 37602 507 37 . . . 37602 508 1 8) 8) CD 37602 508 2 . . . 37602 509 1 6 6 CD 37602 509 2 specimens specimen NNS 37602 509 3 from from IN 37602 509 4 3 3 CD 37602 509 5 localities locality NNS 37602 509 6 . . . 37602 510 1 _ _ NNP 37602 510 2 Remarks._--The Remarks._--The NNP 37602 510 3 specimen speciman NNS 37602 510 4 from from IN 37602 510 5 Yaupi Yaupi NNP 37602 510 6 , , , 37602 510 7 Perú Perú NNP 37602 510 8 ( ( -LRB- 37602 510 9 KU KU NNP 37602 510 10 136312 136312 CD 37602 510 11 ) ) -RRB- 37602 510 12 is be VBZ 37602 510 13 a a DT 37602 510 14 subadult subadult NN 37602 510 15 female female NN 37602 510 16 having have VBG 37602 510 17 a a DT 37602 510 18 snout snout NN 37602 510 19 - - HYPH 37602 510 20 vent vent NN 37602 510 21 length length NN 37602 510 22 of of IN 37602 510 23 39.8 39.8 CD 37602 510 24 mm mm NNS 37602 510 25 . . . 37602 511 1 In in IN 37602 511 2 life life NN 37602 511 3 the the DT 37602 511 4 coloration coloration NN 37602 511 5 was be VBD 37602 511 6 : : : 37602 511 7 " " `` 37602 511 8 Dorsum Dorsum NNP 37602 511 9 light light JJ 37602 511 10 pinkish pinkish NN 37602 511 11 brown brown NN 37602 511 12 with with IN 37602 511 13 large large JJ 37602 511 14 rich rich JJ 37602 511 15 chocolate chocolate NN 37602 511 16 brown brown JJ 37602 511 17 blotch blotch NN 37602 511 18 from from IN 37602 511 19 eyes eye NNS 37602 511 20 to to IN 37602 511 21 anterior anterior JJ 37602 511 22 tips tip NNS 37602 511 23 of of IN 37602 511 24 ilia ilia NNP 37602 511 25 ; ; : 37602 511 26 numerous numerous JJ 37602 511 27 small small JJ 37602 511 28 chocolate chocolate NN 37602 511 29 blotches blotch NNS 37602 511 30 on on IN 37602 511 31 flanks flank NNS 37602 511 32 ; ; : 37602 511 33 dorsal dorsal NN 37602 511 34 surfaces surface NNS 37602 511 35 of of IN 37602 511 36 thighs thigh NNS 37602 511 37 and and CC 37602 511 38 shanks shank NNS 37602 511 39 , , , 37602 511 40 canthus canthus NNP 37602 511 41 , , , 37602 511 42 and and CC 37602 511 43 supraorbital supraorbital JJ 37602 511 44 region region NN 37602 511 45 to to IN 37602 511 46 insertion insertion NN 37602 511 47 of of IN 37602 511 48 forearm forearm NN 37602 511 49 chocolate chocolate NNP 37602 511 50 brown brown NNP 37602 511 51 ; ; , 37602 511 52 supralabial supralabial JJ 37602 511 53 border border NN 37602 511 54 and and CC 37602 511 55 short short JJ 37602 511 56 bar bar NN 37602 511 57 from from IN 37602 511 58 eye eye NN 37602 511 59 to to TO 37602 511 60 lip lip VB 37602 511 61 bronze bronze NN 37602 511 62 - - HYPH 37602 511 63 white white JJ 37602 511 64 ; ; : 37602 511 65 venter venter NN 37602 511 66 bronze bronze NN 37602 511 67 - - HYPH 37602 511 68 white white JJ 37602 511 69 with with IN 37602 511 70 numerous numerous JJ 37602 511 71 tiny tiny JJ 37602 511 72 chocolate chocolate NN 37602 511 73 brown brown NN 37602 511 74 flecks fleck NNS 37602 511 75 [ [ -LRB- 37602 511 76 tending tend VBG 37602 511 77 to to TO 37602 511 78 form form VB 37602 511 79 reticulations reticulation NNS 37602 511 80 on on IN 37602 511 81 throat throat NN 37602 511 82 and and CC 37602 511 83 chest chest NN 37602 511 84 ] ] -RRB- 37602 511 85 ; ; : 37602 511 86 anterior anterior NNP 37602 511 87 and and CC 37602 511 88 posterior posterior NN 37602 511 89 surfaces surface NNS 37602 511 90 of of IN 37602 511 91 thighs thigh NNS 37602 511 92 light light NNP 37602 511 93 olive olive JJ 37602 511 94 - - HYPH 37602 511 95 brown brown JJ 37602 511 96 ; ; : 37602 511 97 iris iris NNP 37602 511 98 largely largely RB 37602 511 99 black black JJ 37602 511 100 with with IN 37602 511 101 gold gold JJ 37602 511 102 flecks fleck NNS 37602 511 103 . . . 37602 511 104 " " '' 37602 512 1 ( ( -LRB- 37602 512 2 Thomas Thomas NNP 37602 512 3 H. H. NNP 37602 512 4 Fritts Fritts NNP 37602 512 5 , , , 37602 512 6 field field NN 37602 512 7 notes note NNS 37602 512 8 , , , 37602 512 9 23 23 CD 37602 512 10 March March NNP 37602 512 11 1970 1970 CD 37602 512 12 . . . 37602 512 13 ) ) -RRB- 37602 513 1 On on IN 37602 513 2 the the DT 37602 513 3 basis basis NN 37602 513 4 of of IN 37602 513 5 this this DT 37602 513 6 one one CD 37602 513 7 subadult subadult NN 37602 513 8 , , , 37602 513 9 it -PRON- PRP 37602 513 10 seems seem VBZ 37602 513 11 likely likely JJ 37602 513 12 that that IN 37602 513 13 reticulations reticulation NNS 37602 513 14 on on IN 37602 513 15 the the DT 37602 513 16 venter venter NN 37602 513 17 develop develop NN 37602 513 18 with with IN 37602 513 19 age age NN 37602 513 20 . . . 37602 514 1 = = NFP 37602 514 2 Osteocephalus osteocephalus JJ 37602 514 3 taurinus= taurinus= NN 37602 514 4 Steindachner Steindachner NNP 37602 514 5 _ _ NNP 37602 514 6 Osteocephalus osteocephalus JJ 37602 514 7 taurinus taurinus NN 37602 514 8 _ _ NNP 37602 514 9 Steindachner Steindachner NNP 37602 514 10 , , , 37602 514 11 1862:77 1862:77 CD 37602 514 12 [ [ -LRB- 37602 514 13 Holotype.--NHMW holotype.--nhmw NN 37602 514 14 16492 16492 CD 37602 514 15 from from IN 37602 514 16 Barra Barra NNP 37602 514 17 do do VBP 37602 514 18 Río Río NNP 37602 514 19 Negro Negro NNP 37602 514 20 , , , 37602 514 21 Manáus Manáus NNP 37602 514 22 , , , 37602 514 23 Territorio Territorio NNP 37602 514 24 do do VBP 37602 514 25 Amazonas Amazonas NNP 37602 514 26 , , , 37602 514 27 Brasil Brasil NNP 37602 514 28 ; ; : 37602 514 29 Johann Johann NNP 37602 514 30 Natterer Natterer NNP 37602 514 31 collector collector NN 37602 514 32 ] ] -RRB- 37602 514 33 . . . 37602 515 1 _ _ NNP 37602 515 2 Osteocephalus osteocephalus JJ 37602 515 3 flavolineatus flavolineatus NN 37602 515 4 _ _ NNP 37602 515 5 Steindachner Steindachner NNP 37602 515 6 , , , 37602 515 7 1862:80 1862:80 CD 37602 515 8 [ [ -LRB- 37602 515 9 Holotype.--NHMW holotype.--nhmw NN 37602 515 10 16495 16495 CD 37602 515 11 from from IN 37602 515 12 Cucuí Cucuí NNP 37602 515 13 , , , 37602 515 14 Territorio Territorio NNP 37602 515 15 do do VBP 37602 515 16 Amazonas Amazonas NNP 37602 515 17 , , , 37602 515 18 Brasil Brasil NNP 37602 515 19 ; ; : 37602 515 20 Johann Johann NNP 37602 515 21 Natterer Natterer NNP 37602 515 22 collector collector NN 37602 515 23 ] ] -RRB- 37602 515 24 . . . 37602 516 1 _ _ NNP 37602 516 2 Trachycephalus Trachycephalus NNP 37602 516 3 ( ( -LRB- 37602 516 4 Osteocephalus Osteocephalus NNP 37602 516 5 ) ) -RRB- 37602 516 6 taurinus taurinus NN 37602 516 7 _ _ NNP 37602 516 8 Steindachner Steindachner NNP 37602 516 9 , , , 37602 516 10 1867:64 1867:64 CD 37602 516 11 . . . 37602 517 1 _ _ NNP 37602 517 2 Osteocephalus osteocephalus JJ 37602 517 3 planiceps planicep NNS 37602 517 4 _ _ NNP 37602 517 5 Cope Cope NNP 37602 517 6 . . . 37602 518 1 1874:122 1874:122 CD 37602 518 2 [ [ -LRB- 37602 518 3 Holotype.--ANSP Holotype.--ANSP NNP 37602 518 4 11399 11399 CD 37602 518 5 from from IN 37602 518 6 Nauta Nauta NNP 37602 518 7 , , , 37602 518 8 Departamento Departamento NNP 37602 518 9 de de IN 37602 518 10 Loreto Loreto NNP 37602 518 11 , , , 37602 518 12 Perú Perú NNP 37602 518 13 ; ; : 37602 518 14 James James NNP 37602 518 15 Orton Orton NNP 37602 518 16 collector collector NN 37602 518 17 ] ] -RRB- 37602 518 18 . . . 37602 519 1 New new JJ 37602 519 2 synonymy synonymy NNP 37602 519 3 . . . 37602 520 1 _ _ NNP 37602 520 2 Hyla Hyla NNP 37602 520 3 taurina_--Boulenger taurina_--Boulenger NNP 37602 520 4 , , , 37602 520 5 1882:363 1882:363 CD 37602 520 6 [ [ -LRB- 37602 520 7 synonymized synonymize VBN 37602 520 8 _ _ NNP 37602 520 9 Osteocephalus osteocephalus JJ 37602 520 10 flavolineatus flavolineatus NN 37602 520 11 _ _ NNP 37602 520 12 Steindachner Steindachner NNP 37602 520 13 , , , 37602 520 14 1862 1862 CD 37602 520 15 , , , 37602 520 16 with with IN 37602 520 17 _ _ NNP 37602 520 18 O. O. NNP 37602 521 1 taurinus taurinus NNP 37602 521 2 _ _ NNP 37602 521 3 Steindachner Steindachner NNP 37602 521 4 , , , 37602 521 5 1862 1862 CD 37602 521 6 ] ] -RRB- 37602 521 7 . . . 37602 522 1 _ _ NNP 37602 522 2 Hyla Hyla NNP 37602 522 3 planiceps_--Boulenger planiceps_--Boulenger NNP 37602 522 4 , , , 37602 522 5 1882:364 1882:364 CD 37602 522 6 . . . 37602 523 1 _ _ NNP 37602 523 2 Hyla Hyla NNP 37602 523 3 ( ( -LRB- 37602 523 4 Trachycephalus Trachycephalus NNP 37602 523 5 ) ) -RRB- 37602 523 6 vilarsi vilarsi NN 37602 523 7 _ _ NNP 37602 523 8 Melin Melin NNP 37602 523 9 , , , 37602 523 10 1941:40 1941:40 CD 37602 523 11 [ [ -LRB- 37602 523 12 Holotype.--NHMG holotype.--nhmg RB 37602 523 13 488 488 CD 37602 523 14 from from IN 37602 523 15 Taracuá Taracuá NNP 37602 523 16 , , , 37602 523 17 Río Río NNP 37602 523 18 Uaupés Uaupés NNP 37602 523 19 , , , 37602 523 20 Territorio Territorio NNP 37602 523 21 do do VBP 37602 523 22 Amazonas Amazonas NNP 37602 523 23 , , , 37602 523 24 Brasil Brasil NNP 37602 523 25 ; ; : 37602 523 26 Douglas Douglas NNP 37602 523 27 Melin Melin NNP 37602 523 28 collector collector NN 37602 523 29 ] ] -RRB- 37602 523 30 . . . 37602 524 1 ( ( -LRB- 37602 524 2 _ _ NNP 37602 524 3 fide fide NN 37602 524 4 _ _ NNP 37602 524 5 Bokermann Bokermann NNP 37602 524 6 , , , 37602 524 7 1966:64 1966:64 NNP 37602 524 8 . . . 37602 524 9 ) ) -RRB- 37602 525 1 _ _ NNP 37602 525 2 Hyla Hyla NNP 37602 525 3 depressa depressa NN 37602 525 4 _ _ NNP 37602 525 5 Andersson Andersson NNP 37602 525 6 , , , 37602 525 7 1945:73 1945:73 CD 37602 525 8 [ [ -LRB- 37602 525 9 Holotype.--NHRM holotype.--nhrm VBP 37602 525 10 1966 1966 CD 37602 525 11 from from IN 37602 525 12 the the DT 37602 525 13 Río Río NNP 37602 525 14 Pastaza Pastaza NNP 37602 525 15 watershed watershe VBD 37602 525 16 ( ( -LRB- 37602 525 17 ? ? . 37602 526 1 Provincia Provincia NNP 37602 526 2 Pastaza Pastaza NNP 37602 526 3 ) ) -RRB- 37602 526 4 , , , 37602 526 5 Ecuador Ecuador NNP 37602 526 6 ; ; : 37602 526 7 William William NNP 37602 526 8 Clarke Clarke NNP 37602 526 9 - - HYPH 37602 526 10 MacIntyre MacIntyre NNP 37602 526 11 collector collector NN 37602 526 12 ] ] -RRB- 37602 526 13 . . . 37602 527 1 New new JJ 37602 527 2 synonymy synonymy NNP 37602 527 3 . . . 37602 528 1 _ _ NNP 37602 528 2 Justification Justification NNP 37602 528 3 of of IN 37602 528 4 Synonymy._--The synonymy._--the CD 37602 528 5 holotype holotype NN 37602 528 6 of of IN 37602 528 7 _ _ NNP 37602 528 8 Osteocephalus Osteocephalus NNP 37602 528 9 taurinus taurinus NN 37602 528 10 _ _ NNP 37602 528 11 is be VBZ 37602 528 12 a a DT 37602 528 13 female female NN 37602 528 14 having have VBG 37602 528 15 a a DT 37602 528 16 snout snout NN 37602 528 17 - - HYPH 37602 528 18 vent vent NN 37602 528 19 length length NN 37602 528 20 of of IN 37602 528 21 103.9 103.9 CD 37602 528 22 mm mm NNS 37602 528 23 . . . 37602 529 1 The the DT 37602 529 2 diameter diameter NN 37602 529 3 of of IN 37602 529 4 the the DT 37602 529 5 tympanum tympanum NN 37602 529 6 is be VBZ 37602 529 7 6.8 6.8 CD 37602 529 8 mm mm CD 37602 529 9 , , , 37602 529 10 77.3 77.3 CD 37602 529 11 percent percent NN 37602 529 12 of of IN 37602 529 13 the the DT 37602 529 14 diameter diameter NN 37602 529 15 of of IN 37602 529 16 the the DT 37602 529 17 eye eye NN 37602 529 18 . . . 37602 530 1 The the DT 37602 530 2 skull skull NN 37602 530 3 is be VBZ 37602 530 4 strongly strongly RB 37602 530 5 exostosed exostose VBN 37602 530 6 , , , 37602 530 7 and and CC 37602 530 8 the the DT 37602 530 9 lateral lateral JJ 37602 530 10 edges edge NNS 37602 530 11 of of IN 37602 530 12 the the DT 37602 530 13 frontoparietals frontoparietal NNS 37602 530 14 are be VBP 37602 530 15 elevated elevate VBN 37602 530 16 so so IN 37602 530 17 as as IN 37602 530 18 to to TO 37602 530 19 form form VB 37602 530 20 distinct distinct JJ 37602 530 21 ridges ridge NNS 37602 530 22 . . . 37602 531 1 The the DT 37602 531 2 skin skin NN 37602 531 3 on on IN 37602 531 4 the the DT 37602 531 5 dorsum dorsum NN 37602 531 6 is be VBZ 37602 531 7 smooth smooth JJ 37602 531 8 . . . 37602 532 1 When when WRB 37602 532 2 we -PRON- PRP 37602 532 3 examined examine VBD 37602 532 4 the the DT 37602 532 5 type type NN 37602 532 6 on on IN 37602 532 7 5 5 CD 37602 532 8 August August NNP 37602 532 9 1969 1969 CD 37602 532 10 , , , 37602 532 11 the the DT 37602 532 12 specimen speciman NNS 37602 532 13 was be VBD 37602 532 14 soft soft JJ 37602 532 15 and and CC 37602 532 16 badly badly RB 37602 532 17 faded fade VBD 37602 532 18 to to IN 37602 532 19 a a DT 37602 532 20 pale pale JJ 37602 532 21 creamy creamy JJ 37602 532 22 tan tan NN 37602 532 23 with with IN 37602 532 24 pale pale JJ 37602 532 25 brown brown JJ 37602 532 26 transverse transverse JJ 37602 532 27 bars bar NNS 37602 532 28 on on IN 37602 532 29 the the DT 37602 532 30 hind hind NN 37602 532 31 limbs limb NNS 37602 532 32 and and CC 37602 532 33 spots spot NNS 37602 532 34 on on IN 37602 532 35 the the DT 37602 532 36 flanks flank NNS 37602 532 37 . . . 37602 533 1 Steindachner Steindachner NNP 37602 533 2 ( ( -LRB- 37602 533 3 1862:79 1862:79 CD 37602 533 4 ) ) -RRB- 37602 533 5 described describe VBD 37602 533 6 the the DT 37602 533 7 coloration coloration NN 37602 533 8 of of IN 37602 533 9 the the DT 37602 533 10 type type NN 37602 533 11 : : : 37602 533 12 " " `` 37602 533 13 In in IN 37602 533 14 the the DT 37602 533 15 preserved preserve VBN 37602 533 16 specimen specimen VBZ 37602 533 17 the the DT 37602 533 18 dorsum dorsum NN 37602 533 19 of of IN 37602 533 20 the the DT 37602 533 21 entire entire JJ 37602 533 22 body body NN 37602 533 23 , , , 37602 533 24 including include VBG 37602 533 25 fore fore NN 37602 533 26 and and CC 37602 533 27 hind hind NN 37602 533 28 limbs limb NNS 37602 533 29 , , , 37602 533 30 is be VBZ 37602 533 31 a a DT 37602 533 32 light light JJ 37602 533 33 yellow yellow JJ 37602 533 34 - - HYPH 37602 533 35 brown brown JJ 37602 533 36 color color NN 37602 533 37 , , , 37602 533 38 which which WDT 37602 533 39 becomes become VBZ 37602 533 40 lighter light JJR 37602 533 41 towards towards IN 37602 533 42 the the DT 37602 533 43 venter venter NN 37602 533 44 . . . 37602 534 1 The the DT 37602 534 2 belly belly NN 37602 534 3 is be VBZ 37602 534 4 whitish whitish JJ 37602 534 5 , , , 37602 534 6 as as IN 37602 534 7 are be VBP 37602 534 8 the the DT 37602 534 9 undersides underside NNS 37602 534 10 of of IN 37602 534 11 the the DT 37602 534 12 arms arm NNS 37602 534 13 and and CC 37602 534 14 legs leg NNS 37602 534 15 . . . 37602 535 1 The the DT 37602 535 2 throat throat NN 37602 535 3 is be VBZ 37602 535 4 indistinctly indistinctly RB 37602 535 5 marbled marble VBN 37602 535 6 with with IN 37602 535 7 brown brown NNP 37602 535 8 . . . 37602 536 1 Roundish roundish JJ 37602 536 2 dark dark JJ 37602 536 3 brown brown JJ 37602 536 4 flecks fleck NNS 37602 536 5 are be VBP 37602 536 6 randomly randomly RB 37602 536 7 distributed distribute VBN 37602 536 8 in in IN 37602 536 9 a a DT 37602 536 10 considerable considerable JJ 37602 536 11 number number NN 37602 536 12 along along IN 37602 536 13 the the DT 37602 536 14 side side NN 37602 536 15 of of IN 37602 536 16 the the DT 37602 536 17 body body NN 37602 536 18 up up RP 37602 536 19 to to IN 37602 536 20 the the DT 37602 536 21 eye eye NN 37602 536 22 ; ; : 37602 536 23 the the DT 37602 536 24 tympanum tympanum NN 37602 536 25 is be VBZ 37602 536 26 more more RBR 37602 536 27 or or CC 37602 536 28 less less RBR 37602 536 29 fully fully RB 37602 536 30 surrounded surround VBN 37602 536 31 by by IN 37602 536 32 brown brown NNP 37602 536 33 . . . 37602 537 1 A a DT 37602 537 2 few few JJ 37602 537 3 discrete discrete JJ 37602 537 4 spots spot NNS 37602 537 5 , , , 37602 537 6 always always RB 37602 537 7 more more RBR 37602 537 8 or or CC 37602 537 9 less less RBR 37602 537 10 drawn draw VBN 37602 537 11 out out RP 37602 537 12 in in IN 37602 537 13 length length NN 37602 537 14 , , , 37602 537 15 on on IN 37602 537 16 the the DT 37602 537 17 sides side NNS 37602 537 18 of of IN 37602 537 19 the the DT 37602 537 20 body body NN 37602 537 21 , , , 37602 537 22 are be VBP 37602 537 23 also also RB 37602 537 24 found find VBN 37602 537 25 on on IN 37602 537 26 the the DT 37602 537 27 posterior posterior JJ 37602 537 28 part part NN 37602 537 29 of of IN 37602 537 30 the the DT 37602 537 31 back back NN 37602 537 32 . . . 37602 538 1 The the DT 37602 538 2 dorsal dorsal NN 37602 538 3 surfaces surface NNS 37602 538 4 of of IN 37602 538 5 the the DT 37602 538 6 fore fore NN 37602 538 7 and and CC 37602 538 8 hind hind NN 37602 538 9 feet foot NNS 37602 538 10 are be VBP 37602 538 11 marked mark VBN 37602 538 12 with with IN 37602 538 13 somewhat somewhat RB 37602 538 14 obliquely obliquely RB 37602 538 15 arranged arrange VBN 37602 538 16 brown brown JJ 37602 538 17 transverse transverse NN 37602 538 18 bands band NNS 37602 538 19 , , , 37602 538 20 which which WDT 37602 538 21 are be VBP 37602 538 22 more more RBR 37602 538 23 intensively intensively RB 37602 538 24 colored color VBN 37602 538 25 near near IN 37602 538 26 the the DT 37602 538 27 margin margin NN 37602 538 28 than than IN 37602 538 29 in in IN 37602 538 30 the the DT 37602 538 31 middle middle NN 37602 538 32 of of IN 37602 538 33 the the DT 37602 538 34 band band NN 37602 538 35 . . . 37602 538 36 " " '' 37602 539 1 ( ( -LRB- 37602 539 2 free free JJ 37602 539 3 translation translation NN 37602 539 4 from from IN 37602 539 5 German German NNP 37602 539 6 . . . 37602 539 7 ) ) -RRB- 37602 540 1 The the DT 37602 540 2 holotype holotype NN 37602 540 3 of of IN 37602 540 4 _ _ NNP 37602 540 5 Osteocephalus Osteocephalus NNP 37602 540 6 flavolineatus flavolineatus NN 37602 540 7 _ _ NNP 37602 540 8 is be VBZ 37602 540 9 a a DT 37602 540 10 female female NN 37602 540 11 having have VBG 37602 540 12 a a DT 37602 540 13 snout snout NN 37602 540 14 - - HYPH 37602 540 15 vent vent NN 37602 540 16 length length NN 37602 540 17 of of IN 37602 540 18 81.8 81.8 CD 37602 540 19 mm mm NNS 37602 540 20 . . . 37602 541 1 The the DT 37602 541 2 diameter diameter NN 37602 541 3 of of IN 37602 541 4 the the DT 37602 541 5 tympanum tympanum NN 37602 541 6 is be VBZ 37602 541 7 6.0 6.0 CD 37602 541 8 mm mm CD 37602 541 9 , , , 37602 541 10 71.4 71.4 CD 37602 541 11 percent percent NN 37602 541 12 of of IN 37602 541 13 the the DT 37602 541 14 diameter diameter NN 37602 541 15 of of IN 37602 541 16 the the DT 37602 541 17 eye eye NN 37602 541 18 . . . 37602 542 1 The the DT 37602 542 2 skull skull NN 37602 542 3 is be VBZ 37602 542 4 strongly strongly RB 37602 542 5 exostosed exostose VBN 37602 542 6 , , , 37602 542 7 and and CC 37602 542 8 the the DT 37602 542 9 lateral lateral JJ 37602 542 10 edges edge NNS 37602 542 11 of of IN 37602 542 12 the the DT 37602 542 13 frontoparietals frontoparietal NNS 37602 542 14 are be VBP 37602 542 15 elevated elevate VBN 37602 542 16 so so IN 37602 542 17 as as IN 37602 542 18 to to TO 37602 542 19 form form VB 37602 542 20 a a DT 37602 542 21 ridge ridge NN 37602 542 22 on on IN 37602 542 23 each each DT 37602 542 24 side side NN 37602 542 25 . . . 37602 543 1 The the DT 37602 543 2 skin skin NN 37602 543 3 on on IN 37602 543 4 the the DT 37602 543 5 dorsum dorsum NN 37602 543 6 is be VBZ 37602 543 7 very very RB 37602 543 8 weakly weakly JJ 37602 543 9 tuberculate tuberculate NN 37602 543 10 . . . 37602 544 1 We -PRON- PRP 37602 544 2 examined examine VBD 37602 544 3 the the DT 37602 544 4 type type NN 37602 544 5 on on IN 37602 544 6 9 9 CD 37602 544 7 August August NNP 37602 544 8 1969 1969 CD 37602 544 9 and and CC 37602 544 10 found find VBD 37602 544 11 it -PRON- PRP 37602 544 12 to to TO 37602 544 13 be be VB 37602 544 14 in in IN 37602 544 15 excellent excellent JJ 37602 544 16 condition condition NN 37602 544 17 . . . 37602 545 1 The the DT 37602 545 2 color color NN 37602 545 3 pattern pattern NN 37602 545 4 is be VBZ 37602 545 5 unchanged unchanged JJ 37602 545 6 from from IN 37602 545 7 that that DT 37602 545 8 described describe VBN 37602 545 9 by by IN 37602 545 10 Steindachner Steindachner NNP 37602 545 11 ( ( -LRB- 37602 545 12 1862:81 1862:81 CD 37602 545 13 ) ) -RRB- 37602 545 14 . . . 37602 546 1 The the DT 37602 546 2 dorsum dorsum NN 37602 546 3 is be VBZ 37602 546 4 tan tan NNP 37602 546 5 with with IN 37602 546 6 irregular irregular JJ 37602 546 7 brown brown JJ 37602 546 8 blotches blotch NNS 37602 546 9 on on IN 37602 546 10 the the DT 37602 546 11 back back NN 37602 546 12 , , , 37602 546 13 spots spot NNS 37602 546 14 on on IN 37602 546 15 the the DT 37602 546 16 flanks flank NNS 37602 546 17 , , , 37602 546 18 and and CC 37602 546 19 transverse transverse JJ 37602 546 20 bars bar NNS 37602 546 21 on on IN 37602 546 22 the the DT 37602 546 23 limbs limb NNS 37602 546 24 . . . 37602 547 1 A a DT 37602 547 2 narrow narrow JJ 37602 547 3 creamy creamy JJ 37602 547 4 white white NNP 37602 547 5 , , , 37602 547 6 middorsal middorsal NNP 37602 547 7 stripe stripe NN 37602 547 8 extends extend VBZ 37602 547 9 from from IN 37602 547 10 the the DT 37602 547 11 snout snout NN 37602 547 12 to to IN 37602 547 13 the the DT 37602 547 14 vent vent NN 37602 547 15 . . . 37602 548 1 The the DT 37602 548 2 subocular subocular JJ 37602 548 3 area area NN 37602 548 4 is be VBZ 37602 548 5 creamy creamy JJ 37602 548 6 tan tan NNP 37602 548 7 , , , 37602 548 8 and and CC 37602 548 9 the the DT 37602 548 10 venter venter NN 37602 548 11 is be VBZ 37602 548 12 tan tan NNP 37602 548 13 . . . 37602 549 1 Boulenger boulenger NN 37602 549 2 ( ( -LRB- 37602 549 3 1882:363 1882:363 NN 37602 549 4 ) ) -RRB- 37602 549 5 questionably questionably RB 37602 549 6 synonymized synonymize VBN 37602 549 7 _ _ NNP 37602 549 8 flavolineatus flavolineatus NN 37602 549 9 _ _ NNP 37602 549 10 with with IN 37602 549 11 _ _ NNP 37602 549 12 taurinus taurinus NNP 37602 549 13 _ _ NNP 37602 549 14 . . . 37602 550 1 We -PRON- PRP 37602 550 2 have have VBP 37602 550 3 observed observe VBN 37602 550 4 that that IN 37602 550 5 a a DT 37602 550 6 middorsal middorsal NN 37602 550 7 cream cream NN 37602 550 8 stripe stripe NN 37602 550 9 occurs occur VBZ 37602 550 10 in in IN 37602 550 11 about about RB 37602 550 12 10 10 CD 37602 550 13 percent percent NN 37602 550 14 of of IN 37602 550 15 the the DT 37602 550 16 specimens specimen NNS 37602 550 17 of of IN 37602 550 18 _ _ NNP 37602 550 19 taurinus taurinus NNP 37602 550 20 _ _ NNP 37602 550 21 and and CC 37602 550 22 in in IN 37602 550 23 some some DT 37602 550 24 specimens specimen NNS 37602 550 25 of of IN 37602 550 26 _ _ NNP 37602 550 27 buckleyi buckleyi NN 37602 550 28 _ _ NNP 37602 550 29 . . . 37602 551 1 This this DT 37602 551 2 is be VBZ 37602 551 3 a a DT 37602 551 4 common common JJ 37602 551 5 color color NN 37602 551 6 morph morph NN 37602 551 7 in in IN 37602 551 8 many many JJ 37602 551 9 species specie NNS 37602 551 10 of of IN 37602 551 11 _ _ NNP 37602 551 12 Eleutherodactylus Eleutherodactylus NNP 37602 551 13 _ _ NNP 37602 551 14 . . . 37602 552 1 In in IN 37602 552 2 the the DT 37602 552 3 absence absence NN 37602 552 4 of of IN 37602 552 5 distinguishing distinguish VBG 37602 552 6 morphological morphological JJ 37602 552 7 characteristics characteristic NNS 37602 552 8 we -PRON- PRP 37602 552 9 can can MD 37602 552 10 only only RB 37602 552 11 conclude conclude VB 37602 552 12 that that IN 37602 552 13 the the DT 37602 552 14 middorsal middorsal NN 37602 552 15 stripe stripe NN 37602 552 16 is be VBZ 37602 552 17 a a DT 37602 552 18 pattern pattern NN 37602 552 19 variant variant NN 37602 552 20 and and CC 37602 552 21 that that IN 37602 552 22 Boulenger Boulenger NNP 37602 552 23 was be VBD 37602 552 24 correct correct JJ 37602 552 25 in in IN 37602 552 26 synonymizing synonymize VBG 37602 552 27 _ _ NNP 37602 552 28 flavolineatus flavolineatus NN 37602 552 29 _ _ NNP 37602 552 30 with with IN 37602 552 31 _ _ NNP 37602 552 32 taurinus taurinus NNP 37602 552 33 _ _ NNP 37602 552 34 . . . 37602 553 1 The the DT 37602 553 2 holotype holotype NN 37602 553 3 of of IN 37602 553 4 _ _ NNP 37602 553 5 Osteocephalus Osteocephalus NNP 37602 553 6 planiceps planicep NNS 37602 553 7 _ _ NNP 37602 553 8 is be VBZ 37602 553 9 a a DT 37602 553 10 male male NN 37602 553 11 having have VBG 37602 553 12 a a DT 37602 553 13 snout snout NN 37602 553 14 - - HYPH 37602 553 15 vent vent NN 37602 553 16 length length NN 37602 553 17 of of IN 37602 553 18 58.5 58.5 CD 37602 553 19 mm mm NNS 37602 553 20 . . . 37602 554 1 The the DT 37602 554 2 diameter diameter NN 37602 554 3 of of IN 37602 554 4 the the DT 37602 554 5 tympanum tympanum NN 37602 554 6 is be VBZ 37602 554 7 4.9 4.9 CD 37602 554 8 mm mm CD 37602 554 9 , , , 37602 554 10 77.8 77.8 CD 37602 554 11 percent percent NN 37602 554 12 of of IN 37602 554 13 the the DT 37602 554 14 diameter diameter NN 37602 554 15 of of IN 37602 554 16 the the DT 37602 554 17 eye eye NN 37602 554 18 . . . 37602 555 1 The the DT 37602 555 2 skull skull NN 37602 555 3 is be VBZ 37602 555 4 moderately moderately RB 37602 555 5 exostosed exostose VBN 37602 555 6 , , , 37602 555 7 and and CC 37602 555 8 the the DT 37602 555 9 lateral lateral JJ 37602 555 10 edges edge NNS 37602 555 11 of of IN 37602 555 12 the the DT 37602 555 13 frontoparietals frontoparietal NNS 37602 555 14 are be VBP 37602 555 15 distinctly distinctly RB 37602 555 16 elevated elevate VBN 37602 555 17 . . . 37602 556 1 The the DT 37602 556 2 skin skin NN 37602 556 3 on on IN 37602 556 4 the the DT 37602 556 5 dorsum dorsum NN 37602 556 6 is be VBZ 37602 556 7 tuberculate tuberculate JJ 37602 556 8 . . . 37602 557 1 Cope Cope NNP 37602 557 2 ( ( -LRB- 37602 557 3 1874:122 1874:122 CD 37602 557 4 ) ) -RRB- 37602 557 5 described describe VBD 37602 557 6 the the DT 37602 557 7 coloration coloration NN 37602 557 8 of of IN 37602 557 9 the the DT 37602 557 10 type type NN 37602 557 11 as as IN 37602 557 12 follows follow VBZ 37602 557 13 : : : 37602 557 14 " " `` 37602 557 15 Color color NN 37602 557 16 above above IN 37602 557 17 uniform uniform VBP 37602 557 18 dark dark JJ 37602 557 19 brown brown NNP 37602 557 20 , , , 37602 557 21 concealed conceal VBD 37602 557 22 surfaces surface NNS 37602 557 23 on on IN 37602 557 24 the the DT 37602 557 25 limbs limb NNS 37602 557 26 similar similar JJ 37602 557 27 and and CC 37602 557 28 without without IN 37602 557 29 any any DT 37602 557 30 markings marking NNS 37602 557 31 . . . 37602 558 1 Sides side NNS 37602 558 2 a a DT 37602 558 3 little little JJ 37602 558 4 varied varied JJ 37602 558 5 with with IN 37602 558 6 the the DT 37602 558 7 white white NN 37602 558 8 of of IN 37602 558 9 the the DT 37602 558 10 belt belt NN 37602 558 11 . . . 37602 559 1 A a DT 37602 559 2 light light JJ 37602 559 3 border border NN 37602 559 4 to to IN 37602 559 5 the the DT 37602 559 6 upper upper JJ 37602 559 7 lip lip NN 37602 559 8 , , , 37602 559 9 and and CC 37602 559 10 lighter light JJR 37602 559 11 line line NN 37602 559 12 from from IN 37602 559 13 the the DT 37602 559 14 orbit orbit NN 37602 559 15 to to IN 37602 559 16 the the DT 37602 559 17 angle angle NN 37602 559 18 of of IN 37602 559 19 the the DT 37602 559 20 mouth mouth NN 37602 559 21 ; ; , 37602 559 22 dermal dermal JJ 37602 559 23 scapular scapular JJ 37602 559 24 fold fold VBP 37602 559 25 pale pale NN 37602 559 26 edged edge VBN 37602 559 27 . . . 37602 560 1 Femur femur NN 37602 560 2 and and CC 37602 560 3 tibia tibia NNP 37602 560 4 with with IN 37602 560 5 dark dark JJ 37602 560 6 crossbands crossband NNS 37602 560 7 on on IN 37602 560 8 the the DT 37602 560 9 exposed expose VBN 37602 560 10 surfaces surface NNS 37602 560 11 . . . 37602 560 12 " " '' 37602 561 1 We -PRON- PRP 37602 561 2 examined examine VBD 37602 561 3 the the DT 37602 561 4 holotype holotype NN 37602 561 5 on on IN 37602 561 6 25 25 CD 37602 561 7 September September NNP 37602 561 8 1969 1969 CD 37602 561 9 , , , 37602 561 10 and and CC 37602 561 11 found find VBD 37602 561 12 it -PRON- PRP 37602 561 13 to to TO 37602 561 14 be be VB 37602 561 15 soft soft JJ 37602 561 16 and and CC 37602 561 17 rubbed rub VBN 37602 561 18 . . . 37602 562 1 The the DT 37602 562 2 coloration coloration NN 37602 562 3 remains remain VBZ 37602 562 4 much much RB 37602 562 5 the the DT 37602 562 6 same same JJ 37602 562 7 as as IN 37602 562 8 described describe VBN 37602 562 9 by by IN 37602 562 10 Cope Cope NNP 37602 562 11 , , , 37602 562 12 who who WP 37602 562 13 provided provide VBD 37602 562 14 no no DT 37602 562 15 means means NN 37602 562 16 of of IN 37602 562 17 distinguishing distinguish VBG 37602 562 18 _ _ NNP 37602 562 19 planiceps planicep NNS 37602 562 20 _ _ NNP 37602 562 21 from from IN 37602 562 22 _ _ NNP 37602 562 23 taurinus taurinus NNP 37602 562 24 _ _ NNP 37602 562 25 . . . 37602 563 1 The the DT 37602 563 2 coloration coloration NN 37602 563 3 and and CC 37602 563 4 morphometric morphometric JJ 37602 563 5 and and CC 37602 563 6 structural structural JJ 37602 563 7 characters character NNS 37602 563 8 of of IN 37602 563 9 the the DT 37602 563 10 type type NN 37602 563 11 of of IN 37602 563 12 _ _ NNP 37602 563 13 planiceps planicep NNS 37602 563 14 _ _ NNP 37602 563 15 all all DT 37602 563 16 fall fall VBP 37602 563 17 within within IN 37602 563 18 the the DT 37602 563 19 range range NN 37602 563 20 of of IN 37602 563 21 variation variation NN 37602 563 22 displayed display VBN 37602 563 23 by by IN 37602 563 24 series series NN 37602 563 25 of of IN 37602 563 26 _ _ NNP 37602 563 27 O. O. NNP 37602 564 1 taurinus taurinus NNP 37602 564 2 _ _ NNP 37602 564 3 from from IN 37602 564 4 the the DT 37602 564 5 upper upper JJ 37602 564 6 Amazon Amazon NNP 37602 564 7 Basin Basin NNP 37602 564 8 . . . 37602 565 1 The the DT 37602 565 2 type type NN 37602 565 3 of of IN 37602 565 4 _ _ NNP 37602 565 5 Hyla Hyla NNP 37602 565 6 vilarsi vilarsi NN 37602 565 7 _ _ NNP 37602 565 8 is be VBZ 37602 565 9 a a DT 37602 565 10 gravid gravid JJ 37602 565 11 female female NN 37602 565 12 having have VBG 37602 565 13 a a DT 37602 565 14 snout snout NN 37602 565 15 - - HYPH 37602 565 16 vent vent NN 37602 565 17 length length NN 37602 565 18 of of IN 37602 565 19 62.7 62.7 CD 37602 565 20 mm mm NNS 37602 565 21 . . . 37602 566 1 The the DT 37602 566 2 diameter diameter NN 37602 566 3 of of IN 37602 566 4 the the DT 37602 566 5 tympanum tympanum NN 37602 566 6 is be VBZ 37602 566 7 4.8 4.8 CD 37602 566 8 mm mm CD 37602 566 9 , , , 37602 566 10 73.8 73.8 CD 37602 566 11 percent percent NN 37602 566 12 of of IN 37602 566 13 the the DT 37602 566 14 diameter diameter NN 37602 566 15 of of IN 37602 566 16 the the DT 37602 566 17 eye eye NN 37602 566 18 . . . 37602 567 1 The the DT 37602 567 2 dorsal dorsal NN 37602 567 3 roofing roofing NN 37602 567 4 bones bone NNS 37602 567 5 of of IN 37602 567 6 the the DT 37602 567 7 skull skull NN 37602 567 8 are be VBP 37602 567 9 moderately moderately RB 37602 567 10 exostosed exostose VBN 37602 567 11 , , , 37602 567 12 and and CC 37602 567 13 the the DT 37602 567 14 lateral lateral JJ 37602 567 15 edges edge NNS 37602 567 16 of of IN 37602 567 17 the the DT 37602 567 18 frontoparietals frontoparietal NNS 37602 567 19 are be VBP 37602 567 20 distinctly distinctly RB 37602 567 21 elevated elevate VBN 37602 567 22 . . . 37602 568 1 The the DT 37602 568 2 skin skin NN 37602 568 3 on on IN 37602 568 4 the the DT 37602 568 5 dorsum dorsum NN 37602 568 6 is be VBZ 37602 568 7 smooth smooth JJ 37602 568 8 . . . 37602 569 1 Melin Melin NNP 37602 569 2 ( ( -LRB- 37602 569 3 1941:42 1941:42 CD 37602 569 4 ) ) -RRB- 37602 569 5 described describe VBD 37602 569 6 the the DT 37602 569 7 coloration coloration NN 37602 569 8 of of IN 37602 569 9 the the DT 37602 569 10 holotype holotype NN 37602 569 11 as as IN 37602 569 12 follows follow VBZ 37602 569 13 : : : 37602 569 14 " " `` 37602 569 15 Above above IN 37602 569 16 uniform uniform JJ 37602 569 17 reddish reddish JJ 37602 569 18 brown brown NN 37602 569 19 ; ; : 37602 569 20 upper upper JJ 37602 569 21 eyelids eyelid NNS 37602 569 22 and and CC 37602 569 23 sides side NNS 37602 569 24 of of IN 37602 569 25 head head NN 37602 569 26 darkish darkish NNP 37602 569 27 brown brown NNP 37602 569 28 ; ; : 37602 569 29 below below IN 37602 569 30 the the DT 37602 569 31 rostral rostral JJ 37602 569 32 edge edge NN 37602 569 33 a a DT 37602 569 34 narrow narrow JJ 37602 569 35 dark dark JJ 37602 569 36 band band NN 37602 569 37 , , , 37602 569 38 continuing continue VBG 37602 569 39 as as IN 37602 569 40 a a DT 37602 569 41 broader broad JJR 37602 569 42 light light NN 37602 569 43 - - HYPH 37602 569 44 edged edge VBN 37602 569 45 one one CD 37602 569 46 through through IN 37602 569 47 the the DT 37602 569 48 eye eye NN 37602 569 49 and and CC 37602 569 50 tympanum tympanum NN 37602 569 51 towards towards IN 37602 569 52 the the DT 37602 569 53 base base NN 37602 569 54 of of IN 37602 569 55 the the DT 37602 569 56 forelimb forelimb NN 37602 569 57 and and CC 37602 569 58 then then RB 37602 569 59 farther farther RB 37602 569 60 on on RB 37602 569 61 continuing continue VBG 37602 569 62 along along IN 37602 569 63 the the DT 37602 569 64 sides side NNS 37602 569 65 as as IN 37602 569 66 a a DT 37602 569 67 line line NN 37602 569 68 of of IN 37602 569 69 black black JJ 37602 569 70 spots spot NNS 37602 569 71 ; ; : 37602 569 72 sides side NNS 37602 569 73 of of IN 37602 569 74 upper upper JJ 37602 569 75 jaw jaw NN 37602 569 76 whitish whitish VB 37602 569 77 with with IN 37602 569 78 traces trace NNS 37602 569 79 of of IN 37602 569 80 dark dark JJ 37602 569 81 cross cross NN 37602 569 82 bars bar NNS 37602 569 83 ( ( -LRB- 37602 569 84 one one CD 37602 569 85 distinct distinct JJ 37602 569 86 under under IN 37602 569 87 the the DT 37602 569 88 eye eye NN 37602 569 89 ) ) -RRB- 37602 569 90 ; ; : 37602 569 91 sides side NNS 37602 569 92 of of IN 37602 569 93 body body NN 37602 569 94 darkish darkish VBP 37602 569 95 with with IN 37602 569 96 black black JJ 37602 569 97 spots spot NNS 37602 569 98 and and CC 37602 569 99 marble marble NN 37602 569 100 , , , 37602 569 101 often often RB 37602 569 102 on on IN 37602 569 103 a a DT 37602 569 104 whitish whitish JJ 37602 569 105 ground ground NN 37602 569 106 ; ; : 37602 569 107 thighs thigh NNS 37602 569 108 , , , 37602 569 109 tibiae tibia NNS 37602 569 110 , , , 37602 569 111 and and CC 37602 569 112 tarsi tarsi VB 37602 569 113 each each DT 37602 569 114 with with IN 37602 569 115 two two CD 37602 569 116 broad broad JJ 37602 569 117 light light NN 37602 569 118 - - HYPH 37602 569 119 edged edged JJ 37602 569 120 , , , 37602 569 121 dark dark JJ 37602 569 122 cross cross NN 37602 569 123 bars bar NNS 37602 569 124 on on IN 37602 569 125 a a DT 37602 569 126 brownish brownish JJ 37602 569 127 ground ground NN 37602 569 128 ( ( -LRB- 37602 569 129 less less RBR 37602 569 130 distinct distinct JJ 37602 569 131 on on IN 37602 569 132 thighs thigh NNS 37602 569 133 ) ) -RRB- 37602 569 134 ; ; : 37602 569 135 sides side NNS 37602 569 136 of of IN 37602 569 137 thighs thigh NNS 37602 569 138 finely finely RB 37602 569 139 mottled mottle VBD 37602 569 140 with with IN 37602 569 141 brown brown NNP 37602 569 142 ; ; : 37602 569 143 beneath beneath IN 37602 569 144 whitish whitish VBP 37602 569 145 with with IN 37602 569 146 small small JJ 37602 569 147 , , , 37602 569 148 sparse sparse JJ 37602 569 149 spots spot NNS 37602 569 150 along along IN 37602 569 151 jaw jaw NN 37602 569 152 , , , 37602 569 153 on on IN 37602 569 154 the the DT 37602 569 155 chest chest NN 37602 569 156 and and CC 37602 569 157 sides side NNS 37602 569 158 . . . 37602 569 159 " " '' 37602 570 1 We -PRON- PRP 37602 570 2 examined examine VBD 37602 570 3 the the DT 37602 570 4 type type NN 37602 570 5 on on IN 37602 570 6 17 17 CD 37602 570 7 February February NNP 37602 570 8 1969 1969 CD 37602 570 9 , , , 37602 570 10 at at IN 37602 570 11 which which WDT 37602 570 12 time time NN 37602 570 13 the the DT 37602 570 14 specimen speciman NNS 37602 570 15 was be VBD 37602 570 16 somewhat somewhat RB 37602 570 17 desiccated desiccate VBN 37602 570 18 , , , 37602 570 19 especially especially RB 37602 570 20 the the DT 37602 570 21 hands hand NNS 37602 570 22 and and CC 37602 570 23 feet foot NNS 37602 570 24 . . . 37602 571 1 The the DT 37602 571 2 coloration coloration NN 37602 571 3 remains remain VBZ 37602 571 4 much much RB 37602 571 5 the the DT 37602 571 6 same same JJ 37602 571 7 as as IN 37602 571 8 described describe VBN 37602 571 9 by by IN 37602 571 10 Melin Melin NNP 37602 571 11 , , , 37602 571 12 except except IN 37602 571 13 that that IN 37602 571 14 he -PRON- PRP 37602 571 15 failed fail VBD 37602 571 16 to to TO 37602 571 17 note note VB 37602 571 18 the the DT 37602 571 19 presence presence NN 37602 571 20 of of IN 37602 571 21 four four CD 37602 571 22 elongate elongate NN 37602 571 23 spots spot NNS 37602 571 24 on on IN 37602 571 25 the the DT 37602 571 26 back back NN 37602 571 27 . . . 37602 572 1 The the DT 37602 572 2 status status NN 37602 572 3 of of IN 37602 572 4 the the DT 37602 572 5 names name NNS 37602 572 6 _ _ NNP 37602 572 7 Osteocephalus Osteocephalus NNP 37602 572 8 planiceps planicep NNS 37602 572 9 _ _ NNP 37602 572 10 Cope Cope NNP 37602 572 11 and and CC 37602 572 12 _ _ NNP 37602 572 13 Hyla Hyla NNP 37602 572 14 vilarsi vilarsi NN 37602 572 15 _ _ NNP 37602 572 16 Melin Melin NNP 37602 572 17 was be VBD 37602 572 18 confused confuse VBN 37602 572 19 by by IN 37602 572 20 Cochran Cochran NNP 37602 572 21 and and CC 37602 572 22 Goin go NNP 37602 572 23 ( ( -LRB- 37602 572 24 1970:322 1970:322 CD 37602 572 25 ) ) -RRB- 37602 572 26 , , , 37602 572 27 who who WP 37602 572 28 assigned assign VBD 37602 572 29 these these DT 37602 572 30 names name NNS 37602 572 31 to to IN 37602 572 32 the the DT 37602 572 33 synonymy synonymy NN 37602 572 34 of of IN 37602 572 35 _ _ NNP 37602 572 36 O. O. NNP 37602 573 1 leprieurii leprieurii NNP 37602 573 2 _ _ NNP 37602 573 3 . . . 37602 574 1 Bokermann Bokermann NNP 37602 574 2 ( ( -LRB- 37602 574 3 1966:64 1966:64 CD 37602 574 4 ) ) -RRB- 37602 574 5 placed place VBD 37602 574 6 _ _ NNP 37602 574 7 Hyla Hyla NNP 37602 574 8 vilarsi vilarsi NN 37602 574 9 _ _ NNP 37602 574 10 in in IN 37602 574 11 the the DT 37602 574 12 synonymy synonymy NN 37602 574 13 of of IN 37602 574 14 _ _ NNP 37602 574 15 Osteocephalus Osteocephalus NNP 37602 574 16 taurinus taurinus NN 37602 574 17 _ _ NNP 37602 574 18 without without IN 37602 574 19 justification justification NN 37602 574 20 . . . 37602 575 1 The the DT 37602 575 2 type type NN 37602 575 3 specimens specimen NNS 37602 575 4 of of IN 37602 575 5 both both DT 37602 575 6 _ _ NNP 37602 575 7 planiceps planicep NNS 37602 575 8 _ _ NNP 37602 575 9 and and CC 37602 575 10 _ _ NNP 37602 575 11 vilarsi vilarsi NN 37602 575 12 _ _ NNP 37602 575 13 have have VBP 37602 575 14 moderately moderately RB 37602 575 15 exostosed exostose VBN 37602 575 16 dermal dermal JJ 37602 575 17 roofing roofing NN 37602 575 18 bones bone NNS 37602 575 19 and and CC 37602 575 20 distinct distinct JJ 37602 575 21 cranial cranial JJ 37602 575 22 ridges ridge NNS 37602 575 23 . . . 37602 576 1 The the DT 37602 576 2 type type NN 37602 576 3 of of IN 37602 576 4 _ _ NNP 37602 576 5 planiceps planicep NNS 37602 576 6 _ _ NNP 37602 576 7 has have VBZ 37602 576 8 moderately moderately RB 37602 576 9 large large JJ 37602 576 10 tubercles tubercle NNS 37602 576 11 on on IN 37602 576 12 the the DT 37602 576 13 dorsum dorsum NN 37602 576 14 , , , 37602 576 15 and and CC 37602 576 16 the the DT 37602 576 17 type type NN 37602 576 18 of of IN 37602 576 19 _ _ NNP 37602 576 20 vilarsi vilarsi NN 37602 576 21 _ _ NNP 37602 576 22 has have VBZ 37602 576 23 spots spot NNS 37602 576 24 on on IN 37602 576 25 the the DT 37602 576 26 throat throat NN 37602 576 27 , , , 37602 576 28 chest chest NN 37602 576 29 , , , 37602 576 30 and and CC 37602 576 31 flanks flank NNS 37602 576 32 and and CC 37602 576 33 longitudinal longitudinal JJ 37602 576 34 markings marking NNS 37602 576 35 on on IN 37602 576 36 the the DT 37602 576 37 back back NN 37602 576 38 . . . 37602 577 1 All all DT 37602 577 2 of of IN 37602 577 3 these these DT 37602 577 4 features feature NNS 37602 577 5 are be VBP 37602 577 6 characteristic characteristic JJ 37602 577 7 of of IN 37602 577 8 _ _ NNP 37602 577 9 taurinus taurinus NNP 37602 577 10 _ _ NNP 37602 577 11 and and CC 37602 577 12 not not RB 37602 577 13 of of IN 37602 577 14 _ _ NNP 37602 577 15 leprieurii leprieurii JJ 37602 577 16 _ _ NNP 37602 577 17 , , , 37602 577 18 which which WDT 37602 577 19 lacks lack VBZ 37602 577 20 exostosis exostosis NN 37602 577 21 and and CC 37602 577 22 cranial cranial JJ 37602 577 23 ridges ridge NNS 37602 577 24 and and CC 37602 577 25 has have VBZ 37602 577 26 transverse transverse JJ 37602 577 27 markings marking NNS 37602 577 28 on on IN 37602 577 29 the the DT 37602 577 30 back back NN 37602 577 31 , , , 37602 577 32 no no DT 37602 577 33 spots spot NNS 37602 577 34 on on IN 37602 577 35 the the DT 37602 577 36 throat throat NN 37602 577 37 , , , 37602 577 38 chest chest NN 37602 577 39 , , , 37602 577 40 and and CC 37602 577 41 flanks flank NNS 37602 577 42 , , , 37602 577 43 and and CC 37602 577 44 in in IN 37602 577 45 males male NNS 37602 577 46 has have VBZ 37602 577 47 small small JJ 37602 577 48 dorsal dorsal NN 37602 577 49 tubercles tubercle NNS 37602 577 50 . . . 37602 578 1 The the DT 37602 578 2 type type NN 37602 578 3 of of IN 37602 578 4 _ _ NNP 37602 578 5 Hyla Hyla NNP 37602 578 6 depressa depressa NNP 37602 578 7 _ _ NNP 37602 578 8 is be VBZ 37602 578 9 a a DT 37602 578 10 male male NN 37602 578 11 having have VBG 37602 578 12 a a DT 37602 578 13 snout snout NN 37602 578 14 - - HYPH 37602 578 15 vent vent NN 37602 578 16 length length NN 37602 578 17 of of IN 37602 578 18 69.8 69.8 CD 37602 578 19 mm mm NNS 37602 578 20 . . . 37602 579 1 The the DT 37602 579 2 diameter diameter NN 37602 579 3 of of IN 37602 579 4 the the DT 37602 579 5 tympanum tympanum NN 37602 579 6 is be VBZ 37602 579 7 5.2 5.2 CD 37602 579 8 mm mm CD 37602 579 9 , , , 37602 579 10 77.6 77.6 CD 37602 579 11 percent percent NN 37602 579 12 of of IN 37602 579 13 the the DT 37602 579 14 diameter diameter NN 37602 579 15 of of IN 37602 579 16 the the DT 37602 579 17 eye eye NN 37602 579 18 . . . 37602 580 1 The the DT 37602 580 2 dorsal dorsal NN 37602 580 3 roofing roofing NN 37602 580 4 bones bone NNS 37602 580 5 of of IN 37602 580 6 the the DT 37602 580 7 skull skull NN 37602 580 8 are be VBP 37602 580 9 moderately moderately RB 37602 580 10 exostosed exostose VBN 37602 580 11 , , , 37602 580 12 and and CC 37602 580 13 the the DT 37602 580 14 lateral lateral JJ 37602 580 15 edges edge NNS 37602 580 16 of of IN 37602 580 17 the the DT 37602 580 18 frontoparietals frontoparietal NNS 37602 580 19 are be VBP 37602 580 20 elevated elevate VBN 37602 580 21 . . . 37602 581 1 The the DT 37602 581 2 skin skin NN 37602 581 3 on on IN 37602 581 4 the the DT 37602 581 5 dorsum dorsum NN 37602 581 6 is be VBZ 37602 581 7 tuberculate tuberculate JJ 37602 581 8 . . . 37602 582 1 The the DT 37602 582 2 dorsum dorsum NN 37602 582 3 is be VBZ 37602 582 4 dull dull JJ 37602 582 5 brown brown JJ 37602 582 6 with with IN 37602 582 7 a a DT 37602 582 8 broad broad JJ 37602 582 9 darker dark JJR 37602 582 10 brown brown JJ 37602 582 11 longitudinal longitudinal JJ 37602 582 12 mark mark NNP 37602 582 13 having have VBG 37602 582 14 indistinct indistinct JJ 37602 582 15 lateral lateral JJ 37602 582 16 edges edge NNS 37602 582 17 from from IN 37602 582 18 the the DT 37602 582 19 snout snout NN 37602 582 20 to to IN 37602 582 21 the the DT 37602 582 22 post post JJ 37602 582 23 - - JJ 37602 582 24 sacral sacral JJ 37602 582 25 area area NN 37602 582 26 . . . 37602 583 1 A a DT 37602 583 2 narrow narrow JJ 37602 583 3 cream cream NN 37602 583 4 middorsal middorsal NN 37602 583 5 line line NN 37602 583 6 extends extend VBZ 37602 583 7 from from IN 37602 583 8 the the DT 37602 583 9 snout snout NN 37602 583 10 to to IN 37602 583 11 the the DT 37602 583 12 vent vent NN 37602 583 13 . . . 37602 584 1 The the DT 37602 584 2 side side NN 37602 584 3 of of IN 37602 584 4 the the DT 37602 584 5 head head NN 37602 584 6 is be VBZ 37602 584 7 dark dark JJ 37602 584 8 brown brown JJ 37602 584 9 , , , 37602 584 10 palest pale JJS 37602 584 11 posteroventral posteroventral NN 37602 584 12 to to IN 37602 584 13 the the DT 37602 584 14 orbit orbit NN 37602 584 15 . . . 37602 585 1 The the DT 37602 585 2 posterior posterior NN 37602 585 3 surfaces surface NNS 37602 585 4 of of IN 37602 585 5 the the DT 37602 585 6 thighs thigh NNS 37602 585 7 are be VBP 37602 585 8 dull dull JJ 37602 585 9 brown brown NNP 37602 585 10 ; ; : 37602 585 11 the the DT 37602 585 12 flanks flank NNS 37602 585 13 are be VBP 37602 585 14 pale pale JJ 37602 585 15 brown brown JJ 37602 585 16 , , , 37602 585 17 and and CC 37602 585 18 the the DT 37602 585 19 ventral ventral JJ 37602 585 20 surfaces surface NNS 37602 585 21 are be VBP 37602 585 22 pale pale JJ 37602 585 23 creamy creamy JJ 37602 585 24 tan tan NN 37602 585 25 . . . 37602 586 1 Dark dark JJ 37602 586 2 brown brown JJ 37602 586 3 transverse transverse JJ 37602 586 4 bars bar NNS 37602 586 5 are be VBP 37602 586 6 present present JJ 37602 586 7 on on IN 37602 586 8 the the DT 37602 586 9 limbs limb NNS 37602 586 10 . . . 37602 587 1 When when WRB 37602 587 2 we -PRON- PRP 37602 587 3 examined examine VBD 37602 587 4 the the DT 37602 587 5 type type NN 37602 587 6 on on IN 37602 587 7 3 3 CD 37602 587 8 January January NNP 37602 587 9 1969 1969 CD 37602 587 10 , , , 37602 587 11 it -PRON- PRP 37602 587 12 was be VBD 37602 587 13 in in IN 37602 587 14 excellent excellent JJ 37602 587 15 condition condition NN 37602 587 16 . . . 37602 588 1 Andersson Andersson NNP 37602 588 2 ( ( -LRB- 37602 588 3 1945:75 1945:75 NN 37602 588 4 ) ) -RRB- 37602 588 5 contrasted contrast VBD 37602 588 6 the the DT 37602 588 7 type type NN 37602 588 8 of of IN 37602 588 9 _ _ NNP 37602 588 10 Hyla Hyla NNP 37602 588 11 depressa depressa NNP 37602 588 12 _ _ NNP 37602 588 13 with with IN 37602 588 14 _ _ NNP 37602 588 15 leprieurii leprieurii JJ 37602 588 16 _ _ NNP 37602 588 17 and and CC 37602 588 18 _ _ NNP 37602 588 19 buckleyi buckleyi NN 37602 588 20 _ _ NNP 37602 588 21 , , , 37602 588 22 but but CC 37602 588 23 he -PRON- PRP 37602 588 24 did do VBD 37602 588 25 not not RB 37602 588 26 compare compare VB 37602 588 27 his -PRON- PRP$ 37602 588 28 specimen speciman NNS 37602 588 29 with with IN 37602 588 30 _ _ NNP 37602 588 31 taurinus taurinus NNP 37602 588 32 _ _ NNP 37602 588 33 , , , 37602 588 34 from from IN 37602 588 35 which which WDT 37602 588 36 it -PRON- PRP 37602 588 37 exhibits exhibit VBZ 37602 588 38 no no DT 37602 588 39 distinguishing distinguishing NN 37602 588 40 features feature NNS 37602 588 41 . . . 37602 589 1 _ _ NNP 37602 589 2 Osteocephalus osteocephalus JJ 37602 589 3 taurinus taurinus NN 37602 589 4 _ _ NNP 37602 589 5 is be VBZ 37602 589 6 a a DT 37602 589 7 widespread widespread JJ 37602 589 8 and and CC 37602 589 9 variable variable JJ 37602 589 10 species specie NNS 37602 589 11 , , , 37602 589 12 and and CC 37602 589 13 it -PRON- PRP 37602 589 14 has have VBZ 37602 589 15 received receive VBN 37602 589 16 several several JJ 37602 589 17 specific specific JJ 37602 589 18 names name NNS 37602 589 19 . . . 37602 590 1 We -PRON- PRP 37602 590 2 are be VBP 37602 590 3 convinced convinced JJ 37602 590 4 that that IN 37602 590 5 _ _ NNP 37602 590 6 Osteocephalus osteocephalus JJ 37602 590 7 taurinus taurinus NN 37602 590 8 _ _ NNP 37602 590 9 Steindachner Steindachner NNP 37602 590 10 , , , 37602 590 11 1862 1862 CD 37602 590 12 , , , 37602 590 13 is be VBZ 37602 590 14 the the DT 37602 590 15 oldest old JJS 37602 590 16 available available JJ 37602 590 17 name name NN 37602 590 18 for for IN 37602 590 19 this this DT 37602 590 20 large large JJ 37602 590 21 Amazonian amazonian JJ 37602 590 22 species specie NNS 37602 590 23 . . . 37602 591 1 The the DT 37602 591 2 following follow VBG 37602 591 3 names name NNS 37602 591 4 are be VBP 37602 591 5 junior junior JJ 37602 591 6 synonyms synonyms NN 37602 591 7 : : : 37602 591 8 _ _ NNP 37602 591 9 Osteocephalus osteocephalus JJ 37602 591 10 flavolineatus flavolineatus NN 37602 591 11 _ _ NNP 37602 591 12 Steindachner Steindachner NNP 37602 591 13 , , , 37602 591 14 1862 1862 CD 37602 591 15 ; ; : 37602 591 16 _ _ NNP 37602 591 17 Osteocephalus Osteocephalus NNP 37602 591 18 planiceps planicep NNS 37602 591 19 _ _ NNP 37602 591 20 Cope Cope NNP 37602 591 21 , , , 37602 591 22 1874 1874 CD 37602 591 23 ; ; : 37602 591 24 _ _ NNP 37602 591 25 Hyla Hyla NNP 37602 591 26 ( ( -LRB- 37602 591 27 Trachycephalus Trachycephalus NNP 37602 591 28 ) ) -RRB- 37602 591 29 vilarsi vilarsi NN 37602 591 30 _ _ NNP 37602 591 31 Melin Melin NNP 37602 591 32 , , , 37602 591 33 1941 1941 CD 37602 591 34 ; ; : 37602 591 35 _ _ NNP 37602 591 36 Hyla Hyla NNP 37602 591 37 depressa depressa NN 37602 591 38 _ _ NNP 37602 591 39 Andersson Andersson NNP 37602 591 40 , , , 37602 591 41 1945 1945 CD 37602 591 42 . . . 37602 592 1 _ _ NNP 37602 592 2 Diagnosis._--1 Diagnosis._--1 NNP 37602 592 3 ) ) -RRB- 37602 592 4 Size size NN 37602 592 5 large large JJ 37602 592 6 ; ; : 37602 592 7 sexual sexual JJ 37602 592 8 dimorphism dimorphism NN 37602 592 9 evident evident NN 37602 592 10 ; ; : 37602 592 11 maximum maximum JJ 37602 592 12 observed observed JJ 37602 592 13 snout snout NN 37602 592 14 - - HYPH 37602 592 15 vent vent NN 37602 592 16 length length NN 37602 592 17 in in IN 37602 592 18 males males NNP 37602 592 19 84.6 84.6 CD 37602 592 20 mm mm NNS 37602 592 21 , , , 37602 592 22 in in IN 37602 592 23 females females NNP 37602 592 24 104 104 CD 37602 592 25 mm mm CD 37602 592 26 ; ; : 37602 592 27 2 2 LS 37602 592 28 ) ) -RRB- 37602 592 29 skin skin NN 37602 592 30 on on IN 37602 592 31 dorsum dorsum NN 37602 592 32 in in IN 37602 592 33 males male NNS 37602 592 34 bearing bear VBG 37602 592 35 many many JJ 37602 592 36 moderately moderately RB 37602 592 37 large large JJ 37602 592 38 , , , 37602 592 39 spinous spinous JJ 37602 592 40 tubercles tubercle NNS 37602 592 41 ; ; : 37602 592 42 3 3 LS 37602 592 43 ) ) -RRB- 37602 592 44 skin skin NN 37602 592 45 on on IN 37602 592 46 flanks flank NNS 37602 592 47 smooth smooth VBP 37602 592 48 ; ; : 37602 592 49 4 4 LS 37602 592 50 ) ) -RRB- 37602 592 51 web web NN 37602 592 52 extending extending NN 37602 592 53 to to IN 37602 592 54 middle middle NN 37602 592 55 of of IN 37602 592 56 antepenultimate antepenultimate NN 37602 592 57 phalange phalange NN 37602 592 58 on on IN 37602 592 59 inner inner JJ 37602 592 60 edge edge NN 37602 592 61 of of IN 37602 592 62 third third JJ 37602 592 63 finger finger NN 37602 592 64 ; ; : 37602 592 65 5 5 LS 37602 592 66 ) ) -RRB- 37602 592 67 dorsum dorsum NN 37602 592 68 brown brown NN 37602 592 69 usually usually RB 37602 592 70 with with IN 37602 592 71 a a DT 37602 592 72 large large JJ 37602 592 73 medial medial JJ 37602 592 74 dark dark JJ 37602 592 75 brown brown JJ 37602 592 76 blotch blotch NN 37602 592 77 or or CC 37602 592 78 , , , 37602 592 79 less less RBR 37602 592 80 frequently frequently RB 37602 592 81 , , , 37602 592 82 several several JJ 37602 592 83 dark dark JJ 37602 592 84 spots spot NNS 37602 592 85 ; ; , 37602 592 86 narrow narrow JJ 37602 592 87 middorsal middorsal NNP 37602 592 88 yellow yellow NNP 37602 592 89 line line NN 37602 592 90 present present JJ 37602 592 91 in in IN 37602 592 92 some some DT 37602 592 93 ; ; : 37602 592 94 6 6 LS 37602 592 95 ) ) -RRB- 37602 592 96 venter venter NN 37602 592 97 cream cream NN 37602 592 98 or or CC 37602 592 99 tan tan NN 37602 592 100 with with IN 37602 592 101 or or CC 37602 592 102 without without IN 37602 592 103 small small JJ 37602 592 104 , , , 37602 592 105 irregular irregular JJ 37602 592 106 brown brown JJ 37602 592 107 flecks fleck NNS 37602 592 108 ; ; : 37602 592 109 7 7 LS 37602 592 110 ) ) -RRB- 37602 592 111 lips lip NNS 37602 592 112 brown brown JJ 37602 592 113 with with IN 37602 592 114 vertical vertical JJ 37602 592 115 cream cream NN 37602 592 116 bar bar NN 37602 592 117 below below IN 37602 592 118 eye eye NN 37602 592 119 in in IN 37602 592 120 some some DT 37602 592 121 , , , 37602 592 122 expanded expand VBN 37602 592 123 into into IN 37602 592 124 pale pale JJ 37602 592 125 labial labial JJ 37602 592 126 stripe stripe NN 37602 592 127 posteriorly posteriorly RB 37602 592 128 in in IN 37602 592 129 some some DT 37602 592 130 females female NNS 37602 592 131 ; ; : 37602 592 132 8) 8) CD 37602 592 133 flanks flank VBZ 37602 592 134 tan tan NNP 37602 592 135 or or CC 37602 592 136 cream cream NN 37602 592 137 with with IN 37602 592 138 or or CC 37602 592 139 without without IN 37602 592 140 small small JJ 37602 592 141 , , , 37602 592 142 irregular irregular JJ 37602 592 143 brown brown JJ 37602 592 144 spots spot NNS 37602 592 145 ; ; : 37602 592 146 9 9 LS 37602 592 147 ) ) -RRB- 37602 592 148 dermal dermal JJ 37602 592 149 roofing roofing NN 37602 592 150 bones bone NNS 37602 592 151 of of IN 37602 592 152 skull skull NN 37602 592 153 exostosed exostose VBD 37602 592 154 , , , 37602 592 155 casqued casque VBD 37602 592 156 , , , 37602 592 157 and and CC 37602 592 158 co co JJ 37602 592 159 - - JJ 37602 592 160 ossified ossified JJ 37602 592 161 ( ( -LRB- 37602 592 162 in in IN 37602 592 163 large large JJ 37602 592 164 adults adult NNS 37602 592 165 ) ) -RRB- 37602 592 166 ; ; : 37602 592 167 10 10 LS 37602 592 168 ) ) -RRB- 37602 592 169 dermal dermal JJ 37602 592 170 sphenethmoid sphenethmoid JJ 37602 592 171 present present NN 37602 592 172 ; ; : 37602 592 173 11 11 CD 37602 592 174 ) ) -RRB- 37602 592 175 nasals nasal NNS 37602 592 176 juxtaposed juxtapose VBD 37602 592 177 medially medially RB 37602 592 178 ; ; : 37602 592 179 12 12 CD 37602 592 180 ) ) -RRB- 37602 592 181 anteromedial anteromedial JJ 37602 592 182 margin margin NN 37602 592 183 of of IN 37602 592 184 frontoparietals frontoparietal NNS 37602 592 185 at at IN 37602 592 186 mid mid NN 37602 592 187 - - NN 37602 592 188 level level NN 37602 592 189 of of IN 37602 592 190 orbit orbit NN 37602 592 191 ; ; : 37602 592 192 13 13 CD 37602 592 193 ) ) -RRB- 37602 592 194 frontoparietal frontoparietal JJ 37602 592 195 fontanelle fontanelle NN 37602 592 196 covered cover VBN 37602 592 197 ; ; : 37602 592 198 14 14 CD 37602 592 199 ) ) -RRB- 37602 592 200 palatine palatine NNP 37602 592 201 serrate serrate NN 37602 592 202 ; ; : 37602 592 203 15 15 CD 37602 592 204 ) ) -RRB- 37602 592 205 parasphenoid parasphenoid NN 37602 592 206 bearing bearing NN 37602 592 207 odontoids odontoid NNS 37602 592 208 ; ; : 37602 592 209 16 16 LS 37602 592 210 ) ) -RRB- 37602 592 211 zygomatic zygomatic JJ 37602 592 212 ramus ramus NN 37602 592 213 of of IN 37602 592 214 squamosal squamosal NNP 37602 592 215 usually usually RB 37602 592 216 articulating articulate VBG 37602 592 217 with with IN 37602 592 218 maxillary maxillary JJ 37602 592 219 arch arch NN 37602 592 220 ; ; : 37602 592 221 17 17 CD 37602 592 222 ) ) -RRB- 37602 592 223 transverse transverse JJ 37602 592 224 processes process NNS 37602 592 225 of of IN 37602 592 226 third third JJ 37602 592 227 presacral presacral JJ 37602 592 228 vertebra vertebra NNS 37602 592 229 approximately approximately RB 37602 592 230 equal equal JJ 37602 592 231 in in IN 37602 592 232 width width NN 37602 592 233 to to IN 37602 592 234 sacral sacral JJ 37602 592 235 diapophyses diapophysis NNS 37602 592 236 ; ; : 37602 592 237 transverse transverse JJ 37602 592 238 processes process NNS 37602 592 239 of of IN 37602 592 240 presacral presacral JJ 37602 592 241 vertebrae vertebra NNS 37602 592 242 3 3 CD 37602 592 243 - - SYM 37602 592 244 8 8 CD 37602 592 245 subequal subequal JJ 37602 592 246 in in IN 37602 592 247 width width NN 37602 592 248 ; ; : 37602 592 249 18 18 CD 37602 592 250 ) ) -RRB- 37602 592 251 intermandibularis intermandibularis NNP 37602 592 252 and and CC 37602 592 253 submentalis submentalis NNP 37602 592 254 muscles muscle NNS 37602 592 255 connected connect VBD 37602 592 256 ; ; : 37602 592 257 19 19 CD 37602 592 258 ) ) -RRB- 37602 592 259 supramandibular supramandibular JJ 37602 592 260 portion portion NN 37602 592 261 of of IN 37602 592 262 interhyoideus interhyoideus NNP 37602 592 263 extensively extensively RB 37602 592 264 developed developed JJ 37602 592 265 ; ; : 37602 592 266 associated associate VBN 37602 592 267 skin skin NN 37602 592 268 forming form VBG 37602 592 269 everted evert VBN 37602 592 270 pouch pouch JJ 37602 592 271 . . . 37602 593 1 The the DT 37602 593 2 moderately moderately RB 37602 593 3 rugose rugose JJ 37602 593 4 dorsum dorsum NN 37602 593 5 ( ( -LRB- 37602 593 6 in in IN 37602 593 7 males male NNS 37602 593 8 ) ) -RRB- 37602 593 9 , , , 37602 593 10 large large JJ 37602 593 11 size size NN 37602 593 12 , , , 37602 593 13 extensive extensive JJ 37602 593 14 webbing webbing NN 37602 593 15 on on IN 37602 593 16 the the DT 37602 593 17 hand hand NN 37602 593 18 , , , 37602 593 19 and and CC 37602 593 20 frontoparietal frontoparietal JJ 37602 593 21 flanges flange NNS 37602 593 22 in in IN 37602 593 23 adults adult NNS 37602 593 24 serve serve VBP 37602 593 25 to to TO 37602 593 26 distinguish distinguish VB 37602 593 27 _ _ NNP 37602 593 28 taurinus taurinus NN 37602 593 29 _ _ NNP 37602 593 30 from from IN 37602 593 31 other other JJ 37602 593 32 members member NNS 37602 593 33 of of IN 37602 593 34 the the DT 37602 593 35 genus genus NN 37602 593 36 . . . 37602 594 1 _ _ NNP 37602 594 2 Distribution._--The Distribution._--The NNP 37602 594 3 Amazon Amazon NNP 37602 594 4 Basin Basin NNP 37602 594 5 , , , 37602 594 6 the the DT 37602 594 7 upper upper JJ 37602 594 8 Orinoco Orinoco NNP 37602 594 9 Basin Basin NNP 37602 594 10 , , , 37602 594 11 and and CC 37602 594 12 the the DT 37602 594 13 Guianas Guianas NNPS 37602 594 14 . . . 37602 595 1 Most Most JJS 37602 595 2 localities locality NNS 37602 595 3 are be VBP 37602 595 4 below below IN 37602 595 5 500 500 CD 37602 595 6 m m NNS 37602 595 7 , , , 37602 595 8 but but CC 37602 595 9 the the DT 37602 595 10 species species NN 37602 595 11 ascends ascend VBZ 37602 595 12 the the DT 37602 595 13 lower low JJR 37602 595 14 Amazonian amazonian JJ 37602 595 15 slopes slope NNS 37602 595 16 of of IN 37602 595 17 the the DT 37602 595 18 Andes Andes NNP 37602 595 19 to to IN 37602 595 20 elevations elevation NNS 37602 595 21 of of IN 37602 595 22 about about RB 37602 595 23 1000 1000 CD 37602 595 24 m m NN 37602 595 25 ( ( -LRB- 37602 595 26 Fig fig NN 37602 595 27 . . . 37602 596 1 11 11 CD 37602 596 2 ) ) -RRB- 37602 596 3 . . . 37602 597 1 A a DT 37602 597 2 record record NN 37602 597 3 from from IN 37602 597 4 Caracas Caracas NNP 37602 597 5 , , , 37602 597 6 Venezuela Venezuela NNP 37602 597 7 , , , 37602 597 8 and and CC 37602 597 9 those those DT 37602 597 10 from from IN 37602 597 11 Provincia Provincia NNP 37602 597 12 Carchi Carchi NNP 37602 597 13 and and CC 37602 597 14 Provincia Provincia NNP 37602 597 15 Esmeraldas Esmeraldas NNP 37602 597 16 , , , 37602 597 17 Ecuador Ecuador NNP 37602 597 18 , , , 37602 597 19 are be VBP 37602 597 20 considered consider VBN 37602 597 21 to to TO 37602 597 22 be be VB 37602 597 23 erroneous erroneous JJ 37602 597 24 . . . 37602 598 1 The the DT 37602 598 2 latter latter JJ 37602 598 3 specimens specimen NNS 37602 598 4 were be VBD 37602 598 5 included include VBN 37602 598 6 in in IN 37602 598 7 a a DT 37602 598 8 collection collection NN 37602 598 9 sold sell VBN 37602 598 10 to to IN 37602 598 11 the the DT 37602 598 12 University University NNP 37602 598 13 of of IN 37602 598 14 Illinois Illinois NNP 37602 598 15 ; ; : 37602 598 16 contained contain VBN 37602 598 17 in in IN 37602 598 18 the the DT 37602 598 19 collection collection NN 37602 598 20 are be VBP 37602 598 21 many many JJ 37602 598 22 common common JJ 37602 598 23 Amazonian amazonian JJ 37602 598 24 species specie NNS 37602 598 25 unknown unknown JJ 37602 598 26 from from IN 37602 598 27 the the DT 37602 598 28 Pacific Pacific NNP 37602 598 29 lowlands lowland NNS 37602 598 30 . . . 37602 599 1 516 516 CD 37602 599 2 specimens specimen NNS 37602 599 3 from from IN 37602 599 4 151 151 CD 37602 599 5 localities locality NNS 37602 599 6 . . . 37602 600 1 [ [ -LRB- 37602 600 2 Illustration illustration NN 37602 600 3 : : : 37602 600 4 FIG FIG NNP 37602 600 5 . . . 37602 601 1 11 11 CD 37602 601 2 . . . 37602 602 1 Distribution distribution NN 37602 602 2 of of IN 37602 602 3 _ _ NNP 37602 602 4 Osteocephalus Osteocephalus NNP 37602 602 5 taurinus taurinus NNP 37602 602 6 _ _ NNP 37602 602 7 . . . 37602 602 8 ] ] -RRB- 37602 603 1 _ _ NNP 37602 603 2 Remarks._--This remarks._--this CD 37602 603 3 widespread widespread JJ 37602 603 4 species species NN 37602 603 5 is be VBZ 37602 603 6 highly highly RB 37602 603 7 variable variable JJ 37602 603 8 in in IN 37602 603 9 size size NN 37602 603 10 and and CC 37602 603 11 coloration coloration NN 37602 603 12 . . . 37602 604 1 Striking strike VBG 37602 604 2 differences difference NNS 37602 604 3 in in IN 37602 604 4 snout snout NN 37602 604 5 - - HYPH 37602 604 6 vent vent NN 37602 604 7 length length NN 37602 604 8 are be VBP 37602 604 9 evident evident JJ 37602 604 10 in in IN 37602 604 11 series series NN 37602 604 12 from from IN 37602 604 13 various various JJ 37602 604 14 parts part NNS 37602 604 15 of of IN 37602 604 16 the the DT 37602 604 17 range range NN 37602 604 18 . . . 37602 605 1 The the DT 37602 605 2 smallest small JJS 37602 605 3 calling calling NN 37602 605 4 males male NNS 37602 605 5 ( ( -LRB- 37602 605 6 CAS CAS NNP 37602 605 7 - - HYPH 37602 605 8 SU SU NNP 37602 605 9 12351 12351 CD 37602 605 10 - - SYM 37602 605 11 6 6 CD 37602 605 12 from from IN 37602 605 13 Rio Rio NNP 37602 605 14 Tapirapé Tapirapé NNP 37602 605 15 , , , 37602 605 16 Brasil Brasil NNP 37602 605 17 ) ) -RRB- 37602 605 18 have have VBP 37602 605 19 snout snout VBN 37602 605 20 - - HYPH 37602 605 21 vent vent NN 37602 605 22 lengths length NNS 37602 605 23 of of IN 37602 605 24 46.5 46.5 CD 37602 605 25 - - SYM 37602 605 26 60.3 60.3 CD 37602 605 27 ( ( -LRB- 37602 605 28 mean mean NNP 37602 605 29 53.3 53.3 CD 37602 605 30 ) ) -RRB- 37602 605 31 mm mm NNP 37602 605 32 , , , 37602 605 33 whereas whereas IN 37602 605 34 the the DT 37602 605 35 largest large JJS 37602 605 36 ( ( -LRB- 37602 605 37 FMNH FMNH NNP 37602 605 38 140254 140254 CD 37602 605 39 , , , 37602 605 40 KU KU NNP 37602 605 41 92243 92243 CD 37602 605 42 - - SYM 37602 605 43 6 6 CD 37602 605 44 , , , 37602 605 45 WCAB wcab NN 37602 605 46 9997 9997 CD 37602 605 47 , , , 37602 605 48 10001 10001 CD 37602 605 49 , , , 37602 605 50 10003 10003 CD 37602 605 51 - - SYM 37602 605 52 4 4 CD 37602 605 53 from from IN 37602 605 54 Igarapé Igarapé NNP 37602 605 55 Marmelo Marmelo NNP 37602 605 56 , , , 37602 605 57 Brasil Brasil NNP 37602 605 58 ) ) -RRB- 37602 605 59 have have VBP 37602 605 60 snout snout VBN 37602 605 61 - - HYPH 37602 605 62 vent vent NN 37602 605 63 lengths length NNS 37602 605 64 of of IN 37602 605 65 71.5 71.5 CD 37602 605 66 - - HYPH 37602 605 67 84.6 84.6 CD 37602 605 68 ( ( -LRB- 37602 605 69 mean mean NNP 37602 605 70 77.6 77.6 CD 37602 605 71 ) ) -RRB- 37602 605 72 mm mm NNP 37602 605 73 . . . 37602 606 1 Mean mean JJ 37602 606 2 values value NNS 37602 606 3 of of IN 37602 606 4 snout snout NN 37602 606 5 - - HYPH 37602 606 6 vent vent NN 37602 606 7 lengths length NNS 37602 606 8 of of IN 37602 606 9 males male NNS 37602 606 10 from from IN 37602 606 11 other other JJ 37602 606 12 localities locality NNS 37602 606 13 are be VBP 37602 606 14 : : : 37602 606 15 Río Río NNP 37602 606 16 Pastaza Pastaza NNP 37602 606 17 drainage drainage NN 37602 606 18 , , , 37602 606 19 Ecuador Ecuador NNP 37602 606 20 73.8 73.8 CD 37602 606 21 mm mm NN 37602 606 22 , , , 37602 606 23 Surinam Surinam NNP 37602 606 24 67.7 67.7 CD 37602 606 25 mm mm NNP 37602 606 26 , , , 37602 606 27 Río Río NNP 37602 606 28 Ucayali Ucayali NNP 37602 606 29 drainage drainage NN 37602 606 30 , , , 37602 606 31 Perú Perú NNP 37602 606 32 57.6 57.6 CD 37602 606 33 mm mm NNS 37602 606 34 , , , 37602 606 35 and and CC 37602 606 36 Guyana Guyana NNP 37602 606 37 55.5 55.5 CD 37602 606 38 mm mm NNP 37602 606 39 . . . 37602 607 1 Although although IN 37602 607 2 the the DT 37602 607 3 difference difference NN 37602 607 4 between between IN 37602 607 5 the the DT 37602 607 6 smallest small JJS 37602 607 7 and and CC 37602 607 8 largest large JJS 37602 607 9 adults adult NNS 37602 607 10 is be VBZ 37602 607 11 highly highly RB 37602 607 12 significant significant JJ 37602 607 13 , , , 37602 607 14 populations population NNS 37602 607 15 bridging bridge VBG 37602 607 16 the the DT 37602 607 17 gap gap NN 37602 607 18 do do VBP 37602 607 19 exist exist VB 37602 607 20 . . . 37602 608 1 Furthermore furthermore RB 37602 608 2 , , , 37602 608 3 the the DT 37602 608 4 geographic geographic JJ 37602 608 5 arrangement arrangement NN 37602 608 6 of of IN 37602 608 7 small small JJ 37602 608 8 versus versus IN 37602 608 9 large large JJ 37602 608 10 frogs frog NNS 37602 608 11 is be VBZ 37602 608 12 a a DT 37602 608 13 confusing confusing JJ 37602 608 14 mosaic mosaic NNP 37602 608 15 . . . 37602 609 1 We -PRON- PRP 37602 609 2 have have VBP 37602 609 3 entertained entertain VBN 37602 609 4 the the DT 37602 609 5 thought thought NN 37602 609 6 that that IN 37602 609 7 we -PRON- PRP 37602 609 8 have have VBP 37602 609 9 included include VBN 37602 609 10 more more JJR 37602 609 11 than than IN 37602 609 12 one one CD 37602 609 13 species species NN 37602 609 14 in in IN 37602 609 15 _ _ NNP 37602 609 16 taurinus taurinus NN 37602 609 17 _ _ NNP 37602 609 18 , , , 37602 609 19 but but CC 37602 609 20 on on IN 37602 609 21 the the DT 37602 609 22 basis basis NN 37602 609 23 of of IN 37602 609 24 preserved preserve VBN 37602 609 25 specimens specimen NNS 37602 609 26 we -PRON- PRP 37602 609 27 are be VBP 37602 609 28 unable unable JJ 37602 609 29 to to TO 37602 609 30 detect detect VB 37602 609 31 consistent consistent JJ 37602 609 32 differences difference NNS 37602 609 33 distinguishing distinguish VBG 37602 609 34 two two CD 37602 609 35 or or CC 37602 609 36 more more JJR 37602 609 37 taxa taxa NN 37602 609 38 . . . 37602 610 1 The the DT 37602 610 2 coloration coloration NN 37602 610 3 and and CC 37602 610 4 pattern pattern NN 37602 610 5 of of IN 37602 610 6 _ _ NNP 37602 610 7 taurinus taurinus NNP 37602 610 8 _ _ NNP 37602 610 9 are be VBP 37602 610 10 so so RB 37602 610 11 variable variable JJ 37602 610 12 that that IN 37602 610 13 no no DT 37602 610 14 one one CD 37602 610 15 series series NN 37602 610 16 of of IN 37602 610 17 statements statement NNS 37602 610 18 can can MD 37602 610 19 describe describe VB 37602 610 20 samples sample NNS 37602 610 21 drawn draw VBN 37602 610 22 from from IN 37602 610 23 the the DT 37602 610 24 entire entire JJ 37602 610 25 range range NN 37602 610 26 of of IN 37602 610 27 species specie NNS 37602 610 28 . . . 37602 611 1 We -PRON- PRP 37602 611 2 have have VBP 37602 611 3 been be VBN 37602 611 4 unable unable JJ 37602 611 5 to to TO 37602 611 6 determine determine VB 37602 611 7 geographic geographic JJ 37602 611 8 trends trend NNS 37602 611 9 in in IN 37602 611 10 color color NN 37602 611 11 pattern pattern NN 37602 611 12 ; ; : 37602 611 13 instead instead RB 37602 611 14 the the DT 37602 611 15 variation variation NN 37602 611 16 within within IN 37602 611 17 a a DT 37602 611 18 given give VBN 37602 611 19 sample sample NN 37602 611 20 can can MD 37602 611 21 encompass encompass VB 37602 611 22 the the DT 37602 611 23 variety variety NN 37602 611 24 known know VBN 37602 611 25 in in IN 37602 611 26 most most JJS 37602 611 27 other other JJ 37602 611 28 samples sample NNS 37602 611 29 . . . 37602 612 1 Two two CD 37602 612 2 minor minor JJ 37602 612 3 exceptions exception NNS 37602 612 4 do do VBP 37602 612 5 exist exist VB 37602 612 6 . . . 37602 613 1 A a DT 37602 613 2 narrow narrow JJ 37602 613 3 middorsal middorsal NN 37602 613 4 light light NN 37602 613 5 stripe stripe NN 37602 613 6 is be VBZ 37602 613 7 present present JJ 37602 613 8 in in IN 37602 613 9 some some DT 37602 613 10 individuals individual NNS 37602 613 11 from from IN 37602 613 12 throughout throughout IN 37602 613 13 the the DT 37602 613 14 range range NN 37602 613 15 , , , 37602 613 16 but but CC 37602 613 17 striped stripe VBN 37602 613 18 specimens specimen NNS 37602 613 19 are be VBP 37602 613 20 most most RBS 37602 613 21 common common JJ 37602 613 22 in in IN 37602 613 23 the the DT 37602 613 24 upper upper JJ 37602 613 25 Amazon Amazon NNP 37602 613 26 Basin Basin NNP 37602 613 27 . . . 37602 614 1 The the DT 37602 614 2 absence absence NN 37602 614 3 of of IN 37602 614 4 dorsal dorsal NN 37602 614 5 markings marking NNS 37602 614 6 is be VBZ 37602 614 7 uncommon uncommon JJ 37602 614 8 in in IN 37602 614 9 the the DT 37602 614 10 entire entire JJ 37602 614 11 species species NN 37602 614 12 , , , 37602 614 13 but but CC 37602 614 14 it -PRON- PRP 37602 614 15 is be VBZ 37602 614 16 most most RBS 37602 614 17 frequent frequent JJ 37602 614 18 in in IN 37602 614 19 individuals individual NNS 37602 614 20 from from IN 37602 614 21 the the DT 37602 614 22 Guianas Guianas NNPS 37602 614 23 . . . 37602 615 1 A a DT 37602 615 2 few few JJ 37602 615 3 individuals individual NNS 37602 615 4 , , , 37602 615 5 such such JJ 37602 615 6 as as IN 37602 615 7 KU KU NNP 37602 615 8 105230 105230 CD 37602 615 9 , , , 37602 615 10 have have VBP 37602 615 11 scattered scatter VBN 37602 615 12 white white JJ 37602 615 13 spots spot NNS 37602 615 14 on on IN 37602 615 15 the the DT 37602 615 16 dorsum dorsum NN 37602 615 17 . . . 37602 616 1 The the DT 37602 616 2 coloration coloration NN 37602 616 3 of of IN 37602 616 4 four four CD 37602 616 5 males male NNS 37602 616 6 in in IN 37602 616 7 life life NN 37602 616 8 from from IN 37602 616 9 Lago Lago NNP 37602 616 10 Agrio Agrio NNP 37602 616 11 , , , 37602 616 12 Ecuador Ecuador NNP 37602 616 13 ( ( -LRB- 37602 616 14 KU KU NNP 37602 616 15 126652 126652 CD 37602 616 16 - - SYM 37602 616 17 5 5 CD 37602 616 18 ) ) -RRB- 37602 616 19 was be VBD 37602 616 20 : : : 37602 616 21 " " `` 37602 616 22 Dorsal dorsal NN 37602 616 23 ground ground NN 37602 616 24 color color NN 37602 616 25 tan tan NN 37602 616 26 to to IN 37602 616 27 dark dark JJ 37602 616 28 brown brown NNP 37602 616 29 with with IN 37602 616 30 darker dark JJR 37602 616 31 brown brown JJ 37602 616 32 markings marking NNS 37602 616 33 . . . 37602 617 1 Flanks flank NNS 37602 617 2 creamy creamy VBP 37602 617 3 tan tan NN 37602 617 4 to to IN 37602 617 5 yellow yellow NN 37602 617 6 with with IN 37602 617 7 brown brown JJ 37602 617 8 or or CC 37602 617 9 black black JJ 37602 617 10 flecks fleck NNS 37602 617 11 or or CC 37602 617 12 mottling mottling NN 37602 617 13 . . . 37602 618 1 Venter venter JJ 37602 618 2 uniform uniform NN 37602 618 3 creamy creamy NN 37602 618 4 yellow yellow NN 37602 618 5 or or CC 37602 618 6 yellow yellow JJ 37602 618 7 with with IN 37602 618 8 brown brown JJ 37602 618 9 spots spot NNS 37602 618 10 or or CC 37602 618 11 reticulations reticulation NNS 37602 618 12 . . . 37602 619 1 Iris Iris NNPS 37602 619 2 greenish greenish JJ 37602 619 3 yellow yellow NN 37602 619 4 with with IN 37602 619 5 radiating radiate VBG 37602 619 6 black black JJ 37602 619 7 streaks streak NNS 37602 619 8 and and CC 37602 619 9 a a DT 37602 619 10 median median JJ 37602 619 11 , , , 37602 619 12 horizontal horizontal JJ 37602 619 13 reddish reddish JJ 37602 619 14 brown brown JJ 37602 619 15 streak streak NN 37602 619 16 . . . 37602 619 17 " " '' 37602 620 1 ( ( -LRB- 37602 620 2 W. W. NNP 37602 620 3 E. E. NNP 37602 620 4 Duellman Duellman NNP 37602 620 5 , , , 37602 620 6 field field NN 37602 620 7 notes note NNS 37602 620 8 , , , 37602 620 9 12 12 CD 37602 620 10 May May NNP 37602 620 11 1969 1969 CD 37602 620 12 . . . 37602 620 13 ) ) -RRB- 37602 621 1 A a DT 37602 621 2 female female NN 37602 621 3 from from IN 37602 621 4 Santa Santa NNP 37602 621 5 Cecilia Cecilia NNP 37602 621 6 , , , 37602 621 7 Ecuador Ecuador NNP 37602 621 8 ( ( -LRB- 37602 621 9 KU KU NNP 37602 621 10 123173 123173 CD 37602 621 11 ) ) -RRB- 37602 621 12 , , , 37602 621 13 was be VBD 37602 621 14 : : : 37602 621 15 " " `` 37602 621 16 Dorsum dorsum NN 37602 621 17 mottled mottle VBN 37602 621 18 olive olive NN 37602 621 19 - - HYPH 37602 621 20 green green JJ 37602 621 21 and and CC 37602 621 22 tan tan NN 37602 621 23 . . . 37602 622 1 Flanks flank NNS 37602 622 2 tan tan VBP 37602 622 3 with with IN 37602 622 4 brown brown JJ 37602 622 5 spots spot NNS 37602 622 6 . . . 37602 623 1 Belly belly NN 37602 623 2 and and CC 37602 623 3 throat throat NN 37602 623 4 creamy creamy JJ 37602 623 5 white white JJ 37602 623 6 , , , 37602 623 7 becoming become VBG 37602 623 8 tan tan NNP 37602 623 9 posteriorly posteriorly RB 37602 623 10 . . . 37602 624 1 Edge edge NN 37602 624 2 of of IN 37602 624 3 upper upper JJ 37602 624 4 jaw jaw NN 37602 624 5 olive olive NN 37602 624 6 - - HYPH 37602 624 7 green green JJ 37602 624 8 . . . 37602 624 9 " " '' 37602 625 1 ( ( -LRB- 37602 625 2 W. W. NNP 37602 625 3 E. E. NNP 37602 625 4 Duellman Duellman NNP 37602 625 5 , , , 37602 625 6 field field NN 37602 625 7 notes note NNS 37602 625 8 , , , 37602 625 9 16 16 CD 37602 625 10 June June NNP 37602 625 11 1968 1968 CD 37602 625 12 . . . 37602 625 13 ) ) -RRB- 37602 626 1 Another another DT 37602 626 2 female female NN 37602 626 3 from from IN 37602 626 4 Santa Santa NNP 37602 626 5 Cecilia Cecilia NNP 37602 626 6 ( ( -LRB- 37602 626 7 KU KU NNP 37602 626 8 123175 123175 CD 37602 626 9 ) ) -RRB- 37602 626 10 , , , 37602 626 11 was be VBD 37602 626 12 : : : 37602 626 13 " " `` 37602 626 14 Brown Brown NNP 37602 626 15 dorsally dorsally RB 37602 626 16 with with IN 37602 626 17 cream cream NN 37602 626 18 - - HYPH 37602 626 19 colored color VBN 37602 626 20 mottling mottling NN 37602 626 21 . . . 37602 627 1 Transverse Transverse NNP 37602 627 2 bars bar NNS 37602 627 3 on on IN 37602 627 4 legs leg NNS 37602 627 5 darker darker RB 37602 627 6 brown brown JJ 37602 627 7 with with IN 37602 627 8 cream cream NN 37602 627 9 - - HYPH 37602 627 10 colored color VBN 37602 627 11 edges edge NNS 37602 627 12 . . . 37602 628 1 Margin margin NN 37602 628 2 of of IN 37602 628 3 upper upper JJ 37602 628 4 lip lip NN 37602 628 5 creamy creamy NNP 37602 628 6 yellow yellow NN 37602 628 7 . . . 37602 629 1 Anterior Anterior NNP 37602 629 2 and and CC 37602 629 3 posterior posterior NN 37602 629 4 surfaces surface NNS 37602 629 5 of of IN 37602 629 6 thighs thigh NNS 37602 629 7 tan tan NNP 37602 629 8 . . . 37602 630 1 Flanks flank NNS 37602 630 2 white white JJ 37602 630 3 with with IN 37602 630 4 brown brown JJ 37602 630 5 spots spot NNS 37602 630 6 . . . 37602 631 1 Venter venter JJ 37602 631 2 creamy creamy JJ 37602 631 3 white white NN 37602 631 4 . . . 37602 632 1 Iris iris JJ 37602 632 2 greenish greenish JJ 37602 632 3 bronze bronze NN 37602 632 4 with with IN 37602 632 5 heavy heavy JJ 37602 632 6 radiating radiating NN 37602 632 7 reticulations reticulation NNS 37602 632 8 of of IN 37602 632 9 black black NN 37602 632 10 . . . 37602 632 11 " " '' 37602 633 1 ( ( -LRB- 37602 633 2 W. W. NNP 37602 633 3 E. E. NNP 37602 633 4 Duellman Duellman NNP 37602 633 5 , , , 37602 633 6 field field NN 37602 633 7 notes note NNS 37602 633 8 , , , 37602 633 9 22 22 CD 37602 633 10 July July NNP 37602 633 11 1968 1968 CD 37602 633 12 . . . 37602 633 13 ) ) -RRB- 37602 634 1 The the DT 37602 634 2 tendency tendency NN 37602 634 3 for for IN 37602 634 4 females female NNS 37602 634 5 to to TO 37602 634 6 have have VB 37602 634 7 a a DT 37602 634 8 labial labial JJ 37602 634 9 stripe stripe NN 37602 634 10 posteriorly posteriorly RB 37602 634 11 and and CC 37602 634 12 the the DT 37602 634 13 absence absence NN 37602 634 14 of of IN 37602 634 15 dorsal dorsal NN 37602 634 16 tubercles tubercle NNS 37602 634 17 in in IN 37602 634 18 females female NNS 37602 634 19 has have VBZ 37602 634 20 resulted result VBN 37602 634 21 in in IN 37602 634 22 the the DT 37602 634 23 identification identification NN 37602 634 24 of of IN 37602 634 25 many many JJ 37602 634 26 such such JJ 37602 634 27 specimens specimen NNS 37602 634 28 as as IN 37602 634 29 _ _ NNP 37602 634 30 O. O. NNP 37602 635 1 leprieurii leprieurii NNP 37602 635 2 _ _ NNP 37602 635 3 . . . 37602 636 1 Ontogenetic ontogenetic JJ 37602 636 2 change change NN 37602 636 3 in in IN 37602 636 4 coloration coloration NN 37602 636 5 is be VBZ 37602 636 6 slight slight JJ 37602 636 7 in in IN 37602 636 8 _ _ NNP 37602 636 9 taurinus taurinus NN 37602 636 10 _ _ NNP 37602 636 11 . . . 37602 637 1 Most Most JJS 37602 637 2 juveniles juvenile NNS 37602 637 3 ( ( -LRB- 37602 637 4 less less JJR 37602 637 5 than than IN 37602 637 6 40 40 CD 37602 637 7 mm mm NNS 37602 637 8 in in IN 37602 637 9 snout snout NN 37602 637 10 - - HYPH 37602 637 11 vent vent NN 37602 637 12 length length NN 37602 637 13 ) ) -RRB- 37602 637 14 can can MD 37602 637 15 be be VB 37602 637 16 identified identify VBN 37602 637 17 readily readily RB 37602 637 18 . . . 37602 638 1 There there EX 37602 638 2 is be VBZ 37602 638 3 a a DT 37602 638 4 tendency tendency NN 37602 638 5 for for IN 37602 638 6 the the DT 37602 638 7 dorsal dorsal NN 37602 638 8 markings marking NNS 37602 638 9 of of IN 37602 638 10 juveniles juvenile NNS 37602 638 11 to to TO 37602 638 12 consist consist VB 37602 638 13 of of IN 37602 638 14 several several JJ 37602 638 15 small small JJ 37602 638 16 spots spot NNS 37602 638 17 . . . 37602 639 1 Apparently apparently RB 37602 639 2 with with IN 37602 639 3 growth growth NN 37602 639 4 the the DT 37602 639 5 spots spot NNS 37602 639 6 usually usually RB 37602 639 7 coalesce coalesce VBP 37602 639 8 , , , 37602 639 9 forming form VBG 37602 639 10 a a DT 37602 639 11 large large JJ 37602 639 12 median median JJ 37602 639 13 blotch blotch NN 37602 639 14 , , , 37602 639 15 but but CC 37602 639 16 some some DT 37602 639 17 adults adult NNS 37602 639 18 retain retain VBP 37602 639 19 the the DT 37602 639 20 juvenile juvenile JJ 37602 639 21 pattern pattern NN 37602 639 22 . . . 37602 640 1 Cochran Cochran NNP 37602 640 2 and and CC 37602 640 3 Goin go NNP 37602 640 4 ( ( -LRB- 37602 640 5 1970:251 1970:251 CD 37602 640 6 ) ) -RRB- 37602 640 7 erroneously erroneously RB 37602 640 8 identified identify VBD 37602 640 9 several several JJ 37602 640 10 juveniles juvenile NNS 37602 640 11 from from IN 37602 640 12 Colombia Colombia NNP 37602 640 13 as as IN 37602 640 14 _ _ NNP 37602 640 15 Hyla Hyla NNP 37602 640 16 palpebrogranulata palpebrogranulata NN 37602 640 17 _ _ NNP 37602 640 18 Andersson Andersson NNP 37602 640 19 . . . 37602 641 1 = = NFP 37602 641 2 Osteocephalus osteocephalus JJ 37602 641 3 verrucigerus= verrucigerus= NN 37602 641 4 ( ( -LRB- 37602 641 5 Werner Werner NNP 37602 641 6 ) ) -RRB- 37602 641 7 _ _ NNP 37602 641 8 Hyla Hyla NNP 37602 641 9 verrucigera verrucigera NN 37602 641 10 _ _ NNP 37602 641 11 Werner Werner NNP 37602 641 12 , , , 37602 641 13 1901:601 1901:601 CD 37602 641 14 [ [ -LRB- 37602 641 15 Holotype.--ZMB holotype.--zmb NN 37602 641 16 16589 16589 CD 37602 641 17 from from IN 37602 641 18 " " `` 37602 641 19 Ecuador Ecuador NNP 37602 641 20 " " '' 37602 641 21 ; ; : 37602 641 22 Richard Richard NNP 37602 641 23 Haensch Haensch NNP 37602 641 24 collector collector NN 37602 641 25 ] ] -RRB- 37602 641 26 . . . 37602 642 1 _ _ NNP 37602 642 2 Hyla Hyla NNP 37602 642 3 riopastazae riopastazae VBZ 37602 642 4 _ _ NNP 37602 642 5 Andersson Andersson NNP 37602 642 6 , , , 37602 642 7 1945:72 1945:72 CD 37602 642 8 [ [ -LRB- 37602 642 9 Holotype.--NHRM Holotype.--NHRM VBD 37602 642 10 1960 1960 CD 37602 642 11 from from IN 37602 642 12 Baños Baños NNP 37602 642 13 , , , 37602 642 14 Río Río NNP 37602 642 15 Pastaza Pastaza NNP 37602 642 16 , , , 37602 642 17 Provincia Provincia NNP 37602 642 18 Tungurahua Tungurahua NNP 37602 642 19 , , , 37602 642 20 Ecuador Ecuador NNP 37602 642 21 ; ; : 37602 642 22 William William NNP 37602 642 23 Clarke Clarke NNP 37602 642 24 - - HYPH 37602 642 25 MacIntyre MacIntyre NNP 37602 642 26 collector collector NN 37602 642 27 ] ] -RRB- 37602 642 28 . . . 37602 643 1 _ _ NNP 37602 643 2 Hyla Hyla NNP 37602 643 3 orcesi orcesi NN 37602 643 4 _ _ NNP 37602 643 5 Funkhouser Funkhouser NNP 37602 643 6 , , , 37602 643 7 1956:78 1956:78 CD 37602 643 8 [ [ -LRB- 37602 643 9 Holotype.--CAS Holotype.--CAS NNP 37602 643 10 - - HYPH 37602 643 11 SU SU NNP 37602 643 12 13150 13150 CD 37602 643 13 from from IN 37602 643 14 Río Río NNP 37602 643 15 Pacayacu Pacayacu NNP 37602 643 16 , , , 37602 643 17 tributary tributary NN 37602 643 18 of of IN 37602 643 19 Río Río NNP 37602 643 20 Cotapino Cotapino NNP 37602 643 21 , , , 37602 643 22 Provincia Provincia NNP 37602 643 23 Napo Napo NNP 37602 643 24 , , , 37602 643 25 Ecuador Ecuador NNP 37602 643 26 ; ; : 37602 643 27 collector collector NN 37602 643 28 unknown unknown NN 37602 643 29 ] ] -RRB- 37602 643 30 . . . 37602 644 1 _ _ NNP 37602 644 2 Osteocephalus Osteocephalus NNP 37602 644 3 orcesi_--Cochran orcesi_--Cochran NNP 37602 644 4 and and CC 37602 644 5 Goin go NNP 37602 644 6 , , , 37602 644 7 1970:317 1970:317 CD 37602 644 8 . . . 37602 645 1 _ _ NNP 37602 645 2 Osteocephalus Osteocephalus NNP 37602 645 3 verrucigerus_--Trueb verrucigerus_--Trueb NNP 37602 645 4 and and CC 37602 645 5 Duellman Duellman NNP 37602 645 6 , , , 37602 645 7 1970:601 1970:601 CD 37602 645 8 [ [ -LRB- 37602 645 9 Synonymized Synonymized NNP 37602 645 10 _ _ NNP 37602 645 11 Hyla Hyla NNP 37602 645 12 riopastazae riopastazae NNS 37602 645 13 _ _ NNP 37602 645 14 Andersson Andersson NNP 37602 645 15 , , , 37602 645 16 1945 1945 CD 37602 645 17 , , , 37602 645 18 and and CC 37602 645 19 _ _ NNP 37602 645 20 Hyla Hyla NNP 37602 645 21 orcesi orcesi NN 37602 645 22 _ _ NNP 37602 645 23 Funkhouser Funkhouser NNP 37602 645 24 , , , 37602 645 25 1956 1956 CD 37602 645 26 , , , 37602 645 27 with with IN 37602 645 28 _ _ NNP 37602 645 29 Hyla Hyla NNP 37602 645 30 verrucigera verrucigera NN 37602 645 31 _ _ NNP 37602 645 32 Werner Werner NNP 37602 645 33 , , , 37602 645 34 1901 1901 CD 37602 645 35 ] ] -RRB- 37602 645 36 . . . 37602 646 1 _ _ NNP 37602 646 2 Justification Justification NNP 37602 646 3 of of IN 37602 646 4 Synonymy._--Trueb Synonymy._--Trueb NNP 37602 646 5 and and CC 37602 646 6 Duellman Duellman NNP 37602 646 7 ( ( -LRB- 37602 646 8 1970:605 1970:605 CD 37602 646 9 ) ) -RRB- 37602 646 10 discussed discuss VBD 37602 646 11 the the DT 37602 646 12 assignment assignment NN 37602 646 13 of of IN 37602 646 14 the the DT 37602 646 15 names name NNS 37602 646 16 in in IN 37602 646 17 the the DT 37602 646 18 synonymy synonymy NN 37602 646 19 of of IN 37602 646 20 _ _ NNP 37602 646 21 O. O. NNP 37602 647 1 verrucigerus verrucigerus NNP 37602 647 2 _ _ NNP 37602 647 3 ; ; : 37602 647 4 only only RB 37602 647 5 a a DT 37602 647 6 brief brief JJ 37602 647 7 resumé resumé NN 37602 647 8 is be VBZ 37602 647 9 given give VBN 37602 647 10 here here RB 37602 647 11 . . . 37602 648 1 The the DT 37602 648 2 extant extant JJ 37602 648 3 type type NN 37602 648 4 of of IN 37602 648 5 _ _ NNP 37602 648 6 Hyla Hyla NNP 37602 648 7 verrucigera verrucigera NN 37602 648 8 _ _ NNP 37602 648 9 is be VBZ 37602 648 10 a a DT 37602 648 11 juvenile juvenile JJ 37602 648 12 male male NN 37602 648 13 having have VBG 37602 648 14 a a DT 37602 648 15 snout snout NN 37602 648 16 - - HYPH 37602 648 17 vent vent NN 37602 648 18 length length NN 37602 648 19 of of IN 37602 648 20 32.0 32.0 CD 37602 648 21 mm mm NNS 37602 648 22 . . . 37602 649 1 The the DT 37602 649 2 dorsum dorsum NN 37602 649 3 is be VBZ 37602 649 4 smooth smooth JJ 37602 649 5 except except IN 37602 649 6 for for IN 37602 649 7 tubercles tubercle NNS 37602 649 8 on on IN 37602 649 9 the the DT 37602 649 10 eyelids eyelid NNS 37602 649 11 ; ; : 37602 649 12 the the DT 37602 649 13 skin skin NN 37602 649 14 is be VBZ 37602 649 15 loose loose JJ 37602 649 16 , , , 37602 649 17 and and CC 37602 649 18 the the DT 37602 649 19 body body NN 37602 649 20 is be VBZ 37602 649 21 soft soft JJ 37602 649 22 . . . 37602 650 1 The the DT 37602 650 2 specimen speciman NNS 37602 650 3 is be VBZ 37602 650 4 faded fade VBN 37602 650 5 to to IN 37602 650 6 a a DT 37602 650 7 pale pale JJ 37602 650 8 brown brown NN 37602 650 9 ; ; : 37602 650 10 indistinct indistinct JJ 37602 650 11 dark dark JJ 37602 650 12 spots spot NNS 37602 650 13 are be VBP 37602 650 14 present present JJ 37602 650 15 on on IN 37602 650 16 the the DT 37602 650 17 back back NN 37602 650 18 , , , 37602 650 19 and and CC 37602 650 20 transverse transverse JJ 37602 650 21 bars bar NNS 37602 650 22 are be VBP 37602 650 23 evident evident JJ 37602 650 24 on on IN 37602 650 25 the the DT 37602 650 26 limbs limb NNS 37602 650 27 . . . 37602 651 1 The the DT 37602 651 2 holotype holotype NN 37602 651 3 of of IN 37602 651 4 _ _ NNP 37602 651 5 Hyla Hyla NNP 37602 651 6 riopastazae riopastazae VBZ 37602 651 7 _ _ NNP 37602 651 8 is be VBZ 37602 651 9 a a DT 37602 651 10 gravid gravid JJ 37602 651 11 female female NN 37602 651 12 having have VBG 37602 651 13 a a DT 37602 651 14 snout snout NN 37602 651 15 - - HYPH 37602 651 16 vent vent NN 37602 651 17 length length NN 37602 651 18 of of IN 37602 651 19 64.7 64.7 CD 37602 651 20 mm mm NNS 37602 651 21 . . . 37602 652 1 The the DT 37602 652 2 dorsum dorsum NN 37602 652 3 is be VBZ 37602 652 4 smooth smooth JJ 37602 652 5 . . . 37602 653 1 The the DT 37602 653 2 dorsal dorsal NN 37602 653 3 ground ground NN 37602 653 4 color color NN 37602 653 5 is be VBZ 37602 653 6 pale pale JJ 37602 653 7 brown brown JJ 37602 653 8 with with IN 37602 653 9 indistinct indistinct NNP 37602 653 10 brown brown JJ 37602 653 11 transverse transverse NNP 37602 653 12 bars bar NNS 37602 653 13 on on IN 37602 653 14 the the DT 37602 653 15 limbs limb NNS 37602 653 16 . . . 37602 654 1 The the DT 37602 654 2 throat throat NN 37602 654 3 , , , 37602 654 4 chest chest NN 37602 654 5 , , , 37602 654 6 and and CC 37602 654 7 belly belly NN 37602 654 8 are be VBP 37602 654 9 cream cream NN 37602 654 10 with with IN 37602 654 11 brown brown JJ 37602 654 12 spots spot NNS 37602 654 13 and and CC 37602 654 14 mottling mottling NN 37602 654 15 . . . 37602 655 1 The the DT 37602 655 2 holotype holotype NN 37602 655 3 of of IN 37602 655 4 _ _ NNP 37602 655 5 Hyla Hyla NNP 37602 655 6 orcesi orcesi NN 37602 655 7 _ _ NNP 37602 655 8 is be VBZ 37602 655 9 an an DT 37602 655 10 adult adult NN 37602 655 11 male male NN 37602 655 12 having have VBG 37602 655 13 a a DT 37602 655 14 snout snout NN 37602 655 15 - - HYPH 37602 655 16 vent vent NN 37602 655 17 length length NN 37602 655 18 of of IN 37602 655 19 52.6 52.6 CD 37602 655 20 mm mm NNS 37602 655 21 . . . 37602 656 1 The the DT 37602 656 2 dorsum dorsum NN 37602 656 3 is be VBZ 37602 656 4 heavily heavily RB 37602 656 5 tuberculate tuberculate JJ 37602 656 6 . . . 37602 657 1 The the DT 37602 657 2 dorsum dorsum NN 37602 657 3 is be VBZ 37602 657 4 dark dark JJ 37602 657 5 brown brown JJ 37602 657 6 with with IN 37602 657 7 faint faint JJ 37602 657 8 transverse transverse JJ 37602 657 9 bars bar NNS 37602 657 10 on on IN 37602 657 11 the the DT 37602 657 12 forearms forearm NNS 37602 657 13 and and CC 37602 657 14 feet foot NNS 37602 657 15 ; ; , 37602 657 16 the the DT 37602 657 17 ventral ventral JJ 37602 657 18 surfaces surface NNS 37602 657 19 are be VBP 37602 657 20 creamy creamy JJ 37602 657 21 brown brown JJ 37602 657 22 . . . 37602 658 1 Trueb Trueb NNP 37602 658 2 and and CC 37602 658 3 Duellman Duellman NNP 37602 658 4 ( ( -LRB- 37602 658 5 1970 1970 CD 37602 658 6 ) ) -RRB- 37602 658 7 provided provide VBD 37602 658 8 conclusive conclusive JJ 37602 658 9 evidence evidence NN 37602 658 10 that that IN 37602 658 11 the the DT 37602 658 12 types type NNS 37602 658 13 of of IN 37602 658 14 _ _ NNP 37602 658 15 H. H. NNP 37602 659 1 verrucigera verrucigera NNP 37602 659 2 _ _ NNP 37602 659 3 , , , 37602 659 4 _ _ NNP 37602 659 5 riopastazae riopastazae VBZ 37602 659 6 _ _ NNP 37602 659 7 , , , 37602 659 8 and and CC 37602 659 9 _ _ NNP 37602 659 10 orcesi orcesi NN 37602 659 11 _ _ NNP 37602 659 12 are be VBP 37602 659 13 a a DT 37602 659 14 juvenile juvenile JJ 37602 659 15 , , , 37602 659 16 adult adult NN 37602 659 17 female female NN 37602 659 18 , , , 37602 659 19 and and CC 37602 659 20 adult adult NN 37602 659 21 male male NN 37602 659 22 , , , 37602 659 23 respectively respectively RB 37602 659 24 , , , 37602 659 25 of of IN 37602 659 26 one one CD 37602 659 27 species species NN 37602 659 28 , , , 37602 659 29 the the DT 37602 659 30 earliest early JJS 37602 659 31 available available JJ 37602 659 32 name name NN 37602 659 33 for for IN 37602 659 34 which which WDT 37602 659 35 is be VBZ 37602 659 36 _ _ NNP 37602 659 37 Hyla Hyla NNP 37602 659 38 verrucigera verrucigera NN 37602 659 39 _ _ NNP 37602 659 40 Werner Werner NNP 37602 659 41 , , , 37602 659 42 1901 1901 CD 37602 659 43 . . . 37602 660 1 _ _ NNP 37602 660 2 Diagnosis._--1 Diagnosis._--1 NNP 37602 660 3 ) ) -RRB- 37602 660 4 Size size NN 37602 660 5 moderate moderate JJ 37602 660 6 , , , 37602 660 7 sexual sexual JJ 37602 660 8 dimorphism dimorphism NN 37602 660 9 evident evident NN 37602 660 10 ; ; : 37602 660 11 maximum maximum JJ 37602 660 12 observed observed JJ 37602 660 13 snout snout NN 37602 660 14 - - HYPH 37602 660 15 vent vent NN 37602 660 16 length length NN 37602 660 17 in in IN 37602 660 18 males male NNS 37602 660 19 54.3 54.3 CD 37602 660 20 mm mm NNP 37602 660 21 , , , 37602 660 22 in in IN 37602 660 23 females female NNS 37602 660 24 65.8 65.8 CD 37602 660 25 mm mm CD 37602 660 26 ; ; : 37602 660 27 2 2 LS 37602 660 28 ) ) -RRB- 37602 660 29 skin skin NN 37602 660 30 on on IN 37602 660 31 dorsum dorsum NN 37602 660 32 in in IN 37602 660 33 males male NNS 37602 660 34 bearing bear VBG 37602 660 35 large large JJ 37602 660 36 , , , 37602 660 37 keratinized keratinized JJ 37602 660 38 tubercles tubercle NNS 37602 660 39 ; ; : 37602 660 40 3 3 LS 37602 660 41 ) ) -RRB- 37602 660 42 skin skin NN 37602 660 43 on on IN 37602 660 44 flanks flank NNS 37602 660 45 smooth smooth VBP 37602 660 46 ; ; : 37602 660 47 4 4 LS 37602 660 48 ) ) -RRB- 37602 660 49 web web NN 37602 660 50 extending extend VBG 37602 660 51 to to IN 37602 660 52 base base NN 37602 660 53 of of IN 37602 660 54 antepenultimate antepenultimate NN 37602 660 55 phalange phalange NN 37602 660 56 on on IN 37602 660 57 inner inner JJ 37602 660 58 edge edge NN 37602 660 59 of of IN 37602 660 60 third third JJ 37602 660 61 finger finger NN 37602 660 62 ; ; : 37602 660 63 5 5 LS 37602 660 64 ) ) -RRB- 37602 660 65 dorsum dorsum NN 37602 660 66 uniformly uniformly JJ 37602 660 67 dark dark JJ 37602 660 68 brown brown JJ 37602 660 69 or or CC 37602 660 70 black black JJ 37602 660 71 , , , 37602 660 72 with with IN 37602 660 73 tan tan NNP 37602 660 74 snout snout NN 37602 660 75 in in IN 37602 660 76 females female NNS 37602 660 77 ; ; : 37602 660 78 6 6 LS 37602 660 79 ) ) -RRB- 37602 660 80 venter venter NN 37602 660 81 creamy creamy JJ 37602 660 82 white white JJ 37602 660 83 , , , 37602 660 84 heavily heavily RB 37602 660 85 mottled mottle VBN 37602 660 86 with with IN 37602 660 87 black black JJ 37602 660 88 or or CC 37602 660 89 dark dark JJ 37602 660 90 brown brown NN 37602 660 91 , , , 37602 660 92 especially especially RB 37602 660 93 in in IN 37602 660 94 females female NNS 37602 660 95 ; ; : 37602 660 96 7 7 LS 37602 660 97 ) ) -RRB- 37602 660 98 lips lip NNS 37602 660 99 marked mark VBN 37602 660 100 with with IN 37602 660 101 pale pale JJ 37602 660 102 tan tan NNP 37602 660 103 labial labial NNP 37602 660 104 stripe stripe NNP 37602 660 105 and and CC 37602 660 106 suborbital suborbital JJ 37602 660 107 bar bar NN 37602 660 108 ; ; : 37602 660 109 8) 8) CD 37602 660 110 flanks flank VBZ 37602 660 111 dull dull JJ 37602 660 112 reddish reddish JJ 37602 660 113 brown brown NN 37602 660 114 ; ; : 37602 660 115 9 9 LS 37602 660 116 ) ) -RRB- 37602 660 117 dermal dermal JJ 37602 660 118 roofing roofing NN 37602 660 119 bones bone NNS 37602 660 120 of of IN 37602 660 121 skull skull NN 37602 660 122 lacking lacking NN 37602 660 123 exostosis exostosis NN 37602 660 124 ; ; : 37602 660 125 10 10 LS 37602 660 126 ) ) -RRB- 37602 660 127 dermal dermal JJ 37602 660 128 sphenethmoid sphenethmoid JJ 37602 660 129 absent absent NN 37602 660 130 ; ; : 37602 660 131 11 11 CD 37602 660 132 ) ) -RRB- 37602 660 133 nasals nasal NNS 37602 660 134 widely widely RB 37602 660 135 separated separate VBD 37602 660 136 medially medially RB 37602 660 137 ; ; : 37602 660 138 12 12 CD 37602 660 139 ) ) -RRB- 37602 660 140 anteromedial anteromedial JJ 37602 660 141 margin margin NN 37602 660 142 of of IN 37602 660 143 frontoparietals frontoparietal NNS 37602 660 144 at at IN 37602 660 145 anterior anterior NNP 37602 660 146 border border NN 37602 660 147 of of IN 37602 660 148 orbit orbit NN 37602 660 149 ; ; : 37602 660 150 13 13 CD 37602 660 151 ) ) -RRB- 37602 660 152 frontoparietal frontoparietal JJ 37602 660 153 fontanelle fontanelle NN 37602 660 154 covered cover VBN 37602 660 155 ; ; : 37602 660 156 14 14 CD 37602 660 157 ) ) -RRB- 37602 660 158 palatine palatine NNP 37602 660 159 serrate serrate NN 37602 660 160 ; ; : 37602 660 161 15 15 CD 37602 660 162 ) ) -RRB- 37602 660 163 parasphenoid parasphenoid NN 37602 660 164 bearing bearing NN 37602 660 165 odontoids odontoid NNS 37602 660 166 ; ; : 37602 660 167 16 16 LS 37602 660 168 ) ) -RRB- 37602 660 169 zygomatic zygomatic JJ 37602 660 170 ramus ramus NN 37602 660 171 of of IN 37602 660 172 squamosal squamosal NNP 37602 660 173 extending extend VBG 37602 660 174 approximately approximately RB 37602 660 175 one one CD 37602 660 176 - - HYPH 37602 660 177 half half NN 37602 660 178 of of IN 37602 660 179 distance distance NN 37602 660 180 to to IN 37602 660 181 maxillary maxillary NNP 37602 660 182 arch arch NNP 37602 660 183 ; ; : 37602 660 184 17 17 CD 37602 660 185 ) ) -RRB- 37602 660 186 transverse transverse JJ 37602 660 187 processes process NNS 37602 660 188 of of IN 37602 660 189 third third JJ 37602 660 190 presacral presacral JJ 37602 660 191 vertebra vertebra NNS 37602 660 192 approximately approximately RB 37602 660 193 equal equal JJ 37602 660 194 in in IN 37602 660 195 width width NN 37602 660 196 to to IN 37602 660 197 sacral sacral JJ 37602 660 198 diapophyses diapophysis NNS 37602 660 199 ; ; : 37602 660 200 transverse transverse JJ 37602 660 201 processes process NNS 37602 660 202 of of IN 37602 660 203 presacral presacral JJ 37602 660 204 vertebrae vertebra NNS 37602 660 205 3 3 CD 37602 660 206 - - SYM 37602 660 207 8 8 CD 37602 660 208 subequal subequal JJ 37602 660 209 in in IN 37602 660 210 width width NN 37602 660 211 ; ; : 37602 660 212 18 18 CD 37602 660 213 ) ) -RRB- 37602 660 214 intermandibularis intermandibularis NNP 37602 660 215 and and CC 37602 660 216 submentalis submentalis NNP 37602 660 217 muscles muscle NNS 37602 660 218 connected connect VBD 37602 660 219 ; ; : 37602 660 220 19 19 CD 37602 660 221 ) ) -RRB- 37602 660 222 supramandibular supramandibular JJ 37602 660 223 portion portion NN 37602 660 224 of of IN 37602 660 225 interhyoideus interhyoideu NNS 37602 660 226 forming form VBG 37602 660 227 simple simple JJ 37602 660 228 , , , 37602 660 229 tubular tubular JJ 37602 660 230 , , , 37602 660 231 posterolateral posterolateral JJ 37602 660 232 extension extension NN 37602 660 233 ; ; , 37602 660 234 associated associate VBN 37602 660 235 skin skin NN 37602 660 236 unmodified unmodifie VBD 37602 660 237 . . . 37602 661 1 _ _ NNP 37602 661 2 Osteocephalus Osteocephalus NNP 37602 661 3 verrucigerus verrucigerus NN 37602 661 4 _ _ NNP 37602 661 5 can can MD 37602 661 6 be be VB 37602 661 7 distinguished distinguish VBN 37602 661 8 from from IN 37602 661 9 other other JJ 37602 661 10 members member NNS 37602 661 11 of of IN 37602 661 12 the the DT 37602 661 13 genus genus NN 37602 661 14 by by IN 37602 661 15 its -PRON- PRP$ 37602 661 16 uniformly uniformly JJ 37602 661 17 dark dark JJ 37602 661 18 dorsum dorsum NN 37602 661 19 , , , 37602 661 20 heavily heavily RB 37602 661 21 mottled mottle VBN 37602 661 22 venter venter NN 37602 661 23 , , , 37602 661 24 and and CC 37602 661 25 large large JJ 37602 661 26 , , , 37602 661 27 spinous spinous JJ 37602 661 28 tubercles tubercle NNS 37602 661 29 on on IN 37602 661 30 the the DT 37602 661 31 dorsum dorsum NN 37602 661 32 in in IN 37602 661 33 males male NNS 37602 661 34 . . . 37602 662 1 _ _ NNP 37602 662 2 Distribution._--Lower distribution._--low JJR 37602 662 3 Amazonian amazonian JJ 37602 662 4 slopes slope NNS 37602 662 5 ( ( -LRB- 37602 662 6 500 500 CD 37602 662 7 - - SYM 37602 662 8 1840 1840 CD 37602 662 9 m m NNP 37602 662 10 ) ) -RRB- 37602 662 11 of of IN 37602 662 12 the the DT 37602 662 13 Andes Andes NNP 37602 662 14 and and CC 37602 662 15 on on IN 37602 662 16 the the DT 37602 662 17 western western JJ 37602 662 18 fringe fringe NN 37602 662 19 of of IN 37602 662 20 the the DT 37602 662 21 Amazon Amazon NNP 37602 662 22 Basin Basin NNP 37602 662 23 in in IN 37602 662 24 Ecuador Ecuador NNP 37602 662 25 and and CC 37602 662 26 Perú Perú NNP 37602 662 27 ; ; : 37602 662 28 one one CD 37602 662 29 locality locality NN 37602 662 30 ( ( -LRB- 37602 662 31 Acevedo Acevedo NNP 37602 662 32 ) ) -RRB- 37602 662 33 in in IN 37602 662 34 upper upper JJ 37602 662 35 Río Río NNP 37602 662 36 Magdalena Magdalena NNP 37602 662 37 drainage drainage NN 37602 662 38 in in IN 37602 662 39 Colombia Colombia NNP 37602 662 40 ( ( -LRB- 37602 662 41 Fig Fig NNP 37602 662 42 . . . 37602 663 1 9 9 LS 37602 663 2 ) ) -RRB- 37602 663 3 . . . 37602 664 1 40 40 CD 37602 664 2 specimens specimen NNS 37602 664 3 from from IN 37602 664 4 13 13 CD 37602 664 5 localities locality NNS 37602 664 6 . . . 37602 665 1 _ _ NNP 37602 665 2 Remarks._--In Remarks._--In NNP 37602 665 3 life life NN 37602 665 4 the the DT 37602 665 5 dorsum dorsum NN 37602 665 6 in in IN 37602 665 7 males male NNS 37602 665 8 is be VBZ 37602 665 9 dull dull JJ 37602 665 10 olive olive JJ 37602 665 11 - - HYPH 37602 665 12 green green JJ 37602 665 13 ; ; : 37602 665 14 the the DT 37602 665 15 groin groin NN 37602 665 16 , , , 37602 665 17 anterior anterior NNP 37602 665 18 and and CC 37602 665 19 posterior posterior NN 37602 665 20 surfaces surface NNS 37602 665 21 of of IN 37602 665 22 the the DT 37602 665 23 thighs thigh NNS 37602 665 24 , , , 37602 665 25 inner inner JJ 37602 665 26 surfaces surface NNS 37602 665 27 of of IN 37602 665 28 limbs limb NNS 37602 665 29 , , , 37602 665 30 and and CC 37602 665 31 upper upper JJ 37602 665 32 arms arm NNS 37602 665 33 are be VBP 37602 665 34 dark dark JJ 37602 665 35 brown brown JJ 37602 665 36 . . . 37602 666 1 The the DT 37602 666 2 ventral ventral JJ 37602 666 3 surfaces surface NNS 37602 666 4 of of IN 37602 666 5 the the DT 37602 666 6 limbs limb NNS 37602 666 7 are be VBP 37602 666 8 pinkish pinkish NNP 37602 666 9 tan tan NNP 37602 666 10 ; ; : 37602 666 11 the the DT 37602 666 12 other other JJ 37602 666 13 ventral ventral JJ 37602 666 14 surfaces surface NNS 37602 666 15 are be VBP 37602 666 16 pale pale JJ 37602 666 17 creamy creamy JJ 37602 666 18 tan tan NNP 37602 666 19 with with IN 37602 666 20 reddish reddish JJ 37602 666 21 brown brown JJ 37602 666 22 spots spot NNS 37602 666 23 . . . 37602 667 1 The the DT 37602 667 2 suborbital suborbital JJ 37602 667 3 spot spot NN 37602 667 4 is be VBZ 37602 667 5 pale pale JJ 37602 667 6 greenish greenish JJ 37602 667 7 tan tan NNP 37602 667 8 , , , 37602 667 9 and and CC 37602 667 10 the the DT 37602 667 11 iris iris NN 37602 667 12 is be VBZ 37602 667 13 deep deep RB 37602 667 14 reddish reddish JJ 37602 667 15 brown brown NN 37602 667 16 . . . 37602 668 1 In in IN 37602 668 2 females female NNS 37602 668 3 the the DT 37602 668 4 dorsum dorsum NN 37602 668 5 is be VBZ 37602 668 6 dull dull JJ 37602 668 7 olive olive JJ 37602 668 8 - - HYPH 37602 668 9 brown brown JJ 37602 668 10 ; ; : 37602 668 11 the the DT 37602 668 12 anterior anterior JJ 37602 668 13 part part NN 37602 668 14 of of IN 37602 668 15 the the DT 37602 668 16 head head NN 37602 668 17 is be VBZ 37602 668 18 tan tan NNP 37602 668 19 , , , 37602 668 20 and and CC 37602 668 21 the the DT 37602 668 22 suborbital suborbital JJ 37602 668 23 spot spot NN 37602 668 24 is be VBZ 37602 668 25 yellowish yellowish JJ 37602 668 26 tan tan NNP 37602 668 27 . . . 37602 669 1 The the DT 37602 669 2 groin groin NN 37602 669 3 and and CC 37602 669 4 hidden hidden JJ 37602 669 5 surfaces surface NNS 37602 669 6 of of IN 37602 669 7 the the DT 37602 669 8 limbs limb NNS 37602 669 9 are be VBP 37602 669 10 dark dark JJ 37602 669 11 reddish reddish JJ 37602 669 12 brown brown NN 37602 669 13 . . . 37602 670 1 The the DT 37602 670 2 ventral ventral JJ 37602 670 3 surfaces surface NNS 37602 670 4 of of IN 37602 670 5 the the DT 37602 670 6 limbs limb NNS 37602 670 7 are be VBP 37602 670 8 brown brown JJ 37602 670 9 ; ; : 37602 670 10 the the DT 37602 670 11 throat throat NN 37602 670 12 and and CC 37602 670 13 chest chest NN 37602 670 14 are be VBP 37602 670 15 creamy creamy JJ 37602 670 16 white white JJ 37602 670 17 , , , 37602 670 18 and and CC 37602 670 19 the the DT 37602 670 20 belly belly NN 37602 670 21 is be VBZ 37602 670 22 reddish reddish JJ 37602 670 23 tan tan NN 37602 670 24 , , , 37602 670 25 both both CC 37602 670 26 with with IN 37602 670 27 dark dark JJ 37602 670 28 brown brown JJ 37602 670 29 mottling mottling NN 37602 670 30 . . . 37602 671 1 Considerable considerable JJ 37602 671 2 ontogenetic ontogenetic JJ 37602 671 3 change change NN 37602 671 4 occurs occur VBZ 37602 671 5 in in IN 37602 671 6 coloration coloration NN 37602 671 7 . . . 37602 672 1 Juveniles juvenile NNS 37602 672 2 are be VBP 37602 672 3 pale pale JJ 37602 672 4 above above RB 37602 672 5 with with IN 37602 672 6 a a DT 37602 672 7 dark dark JJ 37602 672 8 median median JJ 37602 672 9 dorsal dorsal NN 37602 672 10 blotch blotch NN 37602 672 11 and and CC 37602 672 12 dark dark JJ 37602 672 13 transverse transverse JJ 37602 672 14 bars bar NNS 37602 672 15 on on IN 37602 672 16 the the DT 37602 672 17 limbs limb NNS 37602 672 18 . . . 37602 673 1 The the DT 37602 673 2 venter venter NN 37602 673 3 is be VBZ 37602 673 4 white white JJ 37602 673 5 . . . 37602 674 1 The the DT 37602 674 2 change change NN 37602 674 3 consists consist VBZ 37602 674 4 principally principally RB 37602 674 5 of of IN 37602 674 6 an an DT 37602 674 7 increase increase NN 37602 674 8 in in IN 37602 674 9 dark dark JJ 37602 674 10 pigment pigment NN 37602 674 11 and and CC 37602 674 12 subsequent subsequent JJ 37602 674 13 obliteration obliteration NN 37602 674 14 of of IN 37602 674 15 the the DT 37602 674 16 juvenile juvenile JJ 37602 674 17 pattern pattern NN 37602 674 18 . . . 37602 675 1 Tadpoles tadpole NNS 37602 675 2 of of IN 37602 675 3 this this DT 37602 675 4 species specie NNS 37602 675 5 have have VBP 37602 675 6 moderately moderately RB 37602 675 7 long long JJ 37602 675 8 tails tail NNS 37602 675 9 with with IN 37602 675 10 low low JJ 37602 675 11 fins fin NNS 37602 675 12 , , , 37602 675 13 robust robust JJ 37602 675 14 bodies body NNS 37602 675 15 , , , 37602 675 16 two two CD 37602 675 17 rows row NNS 37602 675 18 of of IN 37602 675 19 labial labial JJ 37602 675 20 papillae papilla NNS 37602 675 21 with with IN 37602 675 22 median median JJ 37602 675 23 part part NN 37602 675 24 of of IN 37602 675 25 the the DT 37602 675 26 upper upper JJ 37602 675 27 lip lip NN 37602 675 28 bare bare NNP 37602 675 29 , , , 37602 675 30 and and CC 37602 675 31 two two CD 37602 675 32 upper upper JJ 37602 675 33 and and CC 37602 675 34 five five CD 37602 675 35 lower low JJR 37602 675 36 rows row NNS 37602 675 37 of of IN 37602 675 38 teeth tooth NNS 37602 675 39 . . . 37602 676 1 Trueb Trueb NNP 37602 676 2 and and CC 37602 676 3 Duellman Duellman NNP 37602 676 4 ( ( -LRB- 37602 676 5 1970 1970 CD 37602 676 6 ) ) -RRB- 37602 676 7 described describe VBD 37602 676 8 the the DT 37602 676 9 eggs egg NNS 37602 676 10 , , , 37602 676 11 tadpoles tadpole NNS 37602 676 12 , , , 37602 676 13 mating mate VBG 37602 676 14 call call NN 37602 676 15 , , , 37602 676 16 and and CC 37602 676 17 variation variation NN 37602 676 18 in in IN 37602 676 19 the the DT 37602 676 20 adults adult NNS 37602 676 21 . . . 37602 677 1 GENERIC generic NN 37602 677 2 RELATIONSHIPS relationships NN 37602 677 3 Among among IN 37602 677 4 the the DT 37602 677 5 33 33 CD 37602 677 6 genera genera NN 37602 677 7 currently currently RB 37602 677 8 recognized recognize VBN 37602 677 9 in in IN 37602 677 10 the the DT 37602 677 11 family family NN 37602 677 12 Hylidae hylidae NN 37602 677 13 , , , 37602 677 14 there there EX 37602 677 15 are be VBP 37602 677 16 two two CD 37602 677 17 basic basic JJ 37602 677 18 types type NNS 37602 677 19 of of IN 37602 677 20 vocal vocal JJ 37602 677 21 sac sac NN 37602 677 22 structure structure NN 37602 677 23 ( ( -LRB- 37602 677 24 Duellman Duellman NNP 37602 677 25 , , , 37602 677 26 1970b 1970b CD 37602 677 27 ) ) -RRB- 37602 677 28 , , , 37602 677 29 namely namely RB 37602 677 30 the the DT 37602 677 31 subgular subgular JJ 37602 677 32 type type NN 37602 677 33 and and CC 37602 677 34 the the DT 37602 677 35 lateral lateral JJ 37602 677 36 type type NN 37602 677 37 . . . 37602 678 1 Only only RB 37602 678 2 four four CD 37602 678 3 hylid hylid NN 37602 678 4 genera genera NN 37602 678 5 , , , 37602 678 6 all all DT 37602 678 7 Neotropical neotropical JJ 37602 678 8 lowland lowland NN 37602 678 9 groups group NNS 37602 678 10 , , , 37602 678 11 are be VBP 37602 678 12 known know VBN 37602 678 13 to to TO 37602 678 14 possess possess VB 37602 678 15 paired pair VBN 37602 678 16 lateral lateral JJ 37602 678 17 vocal vocal JJ 37602 678 18 sacs sac NNS 37602 678 19 ; ; : 37602 678 20 these these DT 37602 678 21 are be VBP 37602 678 22 _ _ NNP 37602 678 23 Osteocephalus Osteocephalus NNP 37602 678 24 _ _ NNP 37602 678 25 , , , 37602 678 26 _ _ NNP 37602 678 27 Argenteohyla Argenteohyla NNP 37602 678 28 _ _ NNP 37602 678 29 , , , 37602 678 30 _ _ NNP 37602 678 31 Phrynohyas Phrynohyas NNP 37602 678 32 _ _ NNP 37602 678 33 , , , 37602 678 34 and and CC 37602 678 35 _ _ NNP 37602 678 36 Trachycephalus Trachycephalus NNP 37602 678 37 _ _ NNP 37602 678 38 . . . 37602 679 1 The the DT 37602 679 2 geographical geographical JJ 37602 679 3 distributions distribution NNS 37602 679 4 and and CC 37602 679 5 morphological morphological JJ 37602 679 6 characteristics characteristic NNS 37602 679 7 of of IN 37602 679 8 these these DT 37602 679 9 four four CD 37602 679 10 genera genus NNS 37602 679 11 suggest suggest VBP 37602 679 12 that that IN 37602 679 13 they -PRON- PRP 37602 679 14 are be VBP 37602 679 15 more more RBR 37602 679 16 closely closely RB 37602 679 17 related related JJ 37602 679 18 to to IN 37602 679 19 one one CD 37602 679 20 another another DT 37602 679 21 than than IN 37602 679 22 with with IN 37602 679 23 any any DT 37602 679 24 other other JJ 37602 679 25 hylid hylid NN 37602 679 26 genera genera NN 37602 679 27 . . . 37602 680 1 Of of IN 37602 680 2 the the DT 37602 680 3 four four CD 37602 680 4 genera genera NN 37602 680 5 , , , 37602 680 6 _ _ NNP 37602 680 7 Osteocephalus Osteocephalus NNP 37602 680 8 _ _ NNP 37602 680 9 is be VBZ 37602 680 10 the the DT 37602 680 11 most most RBS 37602 680 12 generalized generalize VBN 37602 680 13 in in IN 37602 680 14 morphology morphology NN 37602 680 15 , , , 37602 680 16 and and CC 37602 680 17 , , , 37602 680 18 like like IN 37602 680 19 _ _ NNP 37602 680 20 Phrynohyas Phrynohyas NNP 37602 680 21 _ _ NNP 37602 680 22 , , , 37602 680 23 has have VBZ 37602 680 24 no no DT 37602 680 25 specialized specialized JJ 37602 680 26 habits habit NNS 37602 680 27 . . . 37602 681 1 _ _ NNP 37602 681 2 Osteocephalus Osteocephalus NNP 37602 681 3 _ _ NNP 37602 681 4 and and CC 37602 681 5 _ _ NNP 37602 681 6 Argenteohyla Argenteohyla NNP 37602 681 7 _ _ NNP 37602 681 8 are be VBP 37602 681 9 similarly similarly RB 37602 681 10 distinguished distinguish VBN 37602 681 11 from from IN 37602 681 12 _ _ NNP 37602 681 13 Phrynohyas Phrynohyas NNP 37602 681 14 _ _ NNP 37602 681 15 and and CC 37602 681 16 _ _ NNP 37602 681 17 Trachycephalus Trachycephalus NNP 37602 681 18 _ _ NNP 37602 681 19 on on IN 37602 681 20 the the DT 37602 681 21 basis basis NN 37602 681 22 of of IN 37602 681 23 vocal vocal JJ 37602 681 24 sac sac NN 37602 681 25 structure structure NN 37602 681 26 . . . 37602 682 1 The the DT 37602 682 2 vocal vocal JJ 37602 682 3 sacs sacs NN 37602 682 4 of of IN 37602 682 5 _ _ NNP 37602 682 6 Osteocephalus Osteocephalus NNP 37602 682 7 _ _ NNP 37602 682 8 and and CC 37602 682 9 _ _ NNP 37602 682 10 Argenteohyla Argenteohyla NNP 37602 682 11 _ _ NNP 37602 682 12 are be VBP 37602 682 13 posterior posterior NN 37602 682 14 and and CC 37602 682 15 protrude protrude NN 37602 682 16 posterolateral posterolateral NN 37602 682 17 to to IN 37602 682 18 the the DT 37602 682 19 angles angle NNS 37602 682 20 of of IN 37602 682 21 the the DT 37602 682 22 jaws jaw NNS 37602 682 23 when when WRB 37602 682 24 they -PRON- PRP 37602 682 25 are be VBP 37602 682 26 inflated inflate VBN 37602 682 27 , , , 37602 682 28 whereas whereas IN 37602 682 29 those those DT 37602 682 30 of of IN 37602 682 31 _ _ NNP 37602 682 32 Phrynohyas Phrynohyas NNP 37602 682 33 _ _ NNP 37602 682 34 and and CC 37602 682 35 _ _ NNP 37602 682 36 Trachycephalus Trachycephalus NNP 37602 682 37 _ _ NNP 37602 682 38 are be VBP 37602 682 39 more more RBR 37602 682 40 lateral lateral JJ 37602 682 41 and and CC 37602 682 42 protrude protrude NN 37602 682 43 posterior posterior NN 37602 682 44 to to IN 37602 682 45 the the DT 37602 682 46 angles angle NNS 37602 682 47 of of IN 37602 682 48 the the DT 37602 682 49 jaws jaw NNS 37602 682 50 when when WRB 37602 682 51 inflated inflate VBN 37602 682 52 . . . 37602 683 1 Although although IN 37602 683 2 _ _ NNP 37602 683 3 Osteocephalus Osteocephalus NNP 37602 683 4 _ _ NNP 37602 683 5 and and CC 37602 683 6 _ _ NNP 37602 683 7 Argenteohyla Argenteohyla NNP 37602 683 8 _ _ NNP 37602 683 9 have have VBP 37602 683 10 similar similar JJ 37602 683 11 vocal vocal JJ 37602 683 12 sac sac NN 37602 683 13 structure structure NN 37602 683 14 , , , 37602 683 15 they -PRON- PRP 37602 683 16 are be VBP 37602 683 17 obviously obviously RB 37602 683 18 distinct distinct JJ 37602 683 19 . . . 37602 684 1 The the DT 37602 684 2 monotypic monotypic JJ 37602 684 3 _ _ NNP 37602 684 4 Argenteohyla Argenteohyla NNP 37602 684 5 _ _ NNP 37602 684 6 is be VBZ 37602 684 7 a a DT 37602 684 8 rather rather RB 37602 684 9 specialized specialized JJ 37602 684 10 , , , 37602 684 11 semifossorial semifossorial JJ 37602 684 12 frog frog NN 37602 684 13 ( ( -LRB- 37602 684 14 Trueb Trueb NNP 37602 684 15 , , , 37602 684 16 1970b 1970b CD 37602 684 17 ) ) -RRB- 37602 684 18 , , , 37602 684 19 characterized characterize VBN 37602 684 20 by by IN 37602 684 21 smooth smooth JJ 37602 684 22 skin skin NN 37602 684 23 , , , 37602 684 24 moderate moderate JJ 37602 684 25 - - HYPH 37602 684 26 sized sized JJ 37602 684 27 digital digital JJ 37602 684 28 discs disc NNS 37602 684 29 , , , 37602 684 30 and and CC 37602 684 31 a a DT 37602 684 32 large large JJ 37602 684 33 inner inner JJ 37602 684 34 metatarsal metatarsal NN 37602 684 35 tubercle tubercle NN 37602 684 36 . . . 37602 685 1 The the DT 37602 685 2 general general JJ 37602 685 3 architecture architecture NN 37602 685 4 of of IN 37602 685 5 the the DT 37602 685 6 skull skull NN 37602 685 7 is be VBZ 37602 685 8 not not RB 37602 685 9 unlike unlike IN 37602 685 10 that that DT 37602 685 11 of of IN 37602 685 12 _ _ NNP 37602 685 13 Osteocephalus Osteocephalus NNP 37602 685 14 _ _ NNP 37602 685 15 ; ; : 37602 685 16 the the DT 37602 685 17 skulls skull NNS 37602 685 18 of of IN 37602 685 19 both both DT 37602 685 20 are be VBP 37602 685 21 well well RB 37602 685 22 roofed roof VBN 37602 685 23 , , , 37602 685 24 broader broad JJR 37602 685 25 than than IN 37602 685 26 long long JJ 37602 685 27 , , , 37602 685 28 and and CC 37602 685 29 characterized characterize VBN 37602 685 30 by by IN 37602 685 31 posterolaterally posterolaterally RB 37602 685 32 oriented orient VBN 37602 685 33 parasphenoid parasphenoid NN 37602 685 34 alae alae NNP 37602 685 35 . . . 37602 686 1 _ _ NNP 37602 686 2 Argenteohyla Argenteohyla NNP 37602 686 3 _ _ NNP 37602 686 4 bears bear VBZ 37602 686 5 small small JJ 37602 686 6 , , , 37602 686 7 slightly slightly RB 37602 686 8 curved curved JJ 37602 686 9 prevomerine prevomerine JJ 37602 686 10 dentigerous dentigerous JJ 37602 686 11 processes process NNS 37602 686 12 in in IN 37602 686 13 contrast contrast NN 37602 686 14 to to IN 37602 686 15 the the DT 37602 686 16 large large JJ 37602 686 17 , , , 37602 686 18 angular angular JJ 37602 686 19 processes process NNS 37602 686 20 of of IN 37602 686 21 _ _ NNP 37602 686 22 Osteocephalus Osteocephalus NNP 37602 686 23 _ _ NNP 37602 686 24 . . . 37602 687 1 The the DT 37602 687 2 skull skull NN 37602 687 3 of of IN 37602 687 4 _ _ NNP 37602 687 5 Argenteohyla Argenteohyla NNP 37602 687 6 _ _ NNP 37602 687 7 shows show VBZ 37602 687 8 specializations specialization NNS 37602 687 9 , , , 37602 687 10 apparently apparently RB 37602 687 11 adaptations adaptation NNS 37602 687 12 to to IN 37602 687 13 its -PRON- PRP$ 37602 687 14 semifossorial semifossorial JJ 37602 687 15 mode mode NN 37602 687 16 of of IN 37602 687 17 existence existence NN 37602 687 18 , , , 37602 687 19 which which WDT 37602 687 20 further further RB 37602 687 21 distinguish distinguish VBP 37602 687 22 the the DT 37602 687 23 genus genus NN 37602 687 24 from from IN 37602 687 25 _ _ NNP 37602 687 26 Osteocephalus Osteocephalus NNP 37602 687 27 _ _ NNP 37602 687 28 . . . 37602 688 1 In in IN 37602 688 2 comparison comparison NN 37602 688 3 with with IN 37602 688 4 _ _ NNP 37602 688 5 Osteocephalus Osteocephalus NNP 37602 688 6 _ _ NNP 37602 688 7 , , , 37602 688 8 the the DT 37602 688 9 cranium cranium NN 37602 688 10 of of IN 37602 688 11 _ _ NNP 37602 688 12 Argenteohyla Argenteohyla NNP 37602 688 13 _ _ NNP 37602 688 14 is be VBZ 37602 688 15 slightly slightly RB 37602 688 16 depressed depress VBN 37602 688 17 anteriorly anteriorly RB 37602 688 18 , , , 37602 688 19 the the DT 37602 688 20 roofing roofing NN 37602 688 21 bones bone NNS 37602 688 22 extensively extensively RB 37602 688 23 casqued casque VBN 37602 688 24 , , , 37602 688 25 and and CC 37602 688 26 the the DT 37602 688 27 palatines palatine NNS 37602 688 28 robust robust JJ 37602 688 29 . . . 37602 689 1 Osteologically osteologically RB 37602 689 2 , , , 37602 689 3 _ _ NNP 37602 689 4 Osteocephalus Osteocephalus NNP 37602 689 5 _ _ NNP 37602 689 6 more more RBR 37602 689 7 closely closely RB 37602 689 8 resembles resemble VBZ 37602 689 9 _ _ NNP 37602 689 10 Phrynohyas Phrynohyas NNP 37602 689 11 _ _ NNP 37602 689 12 than than IN 37602 689 13 either either DT 37602 689 14 of of IN 37602 689 15 the the DT 37602 689 16 other other JJ 37602 689 17 two two CD 37602 689 18 genera genera NN 37602 689 19 , , , 37602 689 20 but but CC 37602 689 21 _ _ NNP 37602 689 22 Osteocephalus Osteocephalus NNP 37602 689 23 _ _ NNP 37602 689 24 and and CC 37602 689 25 _ _ NNP 37602 689 26 Phrynohyas Phrynohyas NNP 37602 689 27 _ _ NNP 37602 689 28 are be VBP 37602 689 29 clearly clearly RB 37602 689 30 distinct distinct JJ 37602 689 31 on on IN 37602 689 32 the the DT 37602 689 33 basis basis NN 37602 689 34 of of IN 37602 689 35 their -PRON- PRP$ 37602 689 36 respective respective JJ 37602 689 37 vocal vocal JJ 37602 689 38 sac sac NN 37602 689 39 structure structure NN 37602 689 40 . . . 37602 690 1 Like like IN 37602 690 2 _ _ NNP 37602 690 3 Osteocephalus Osteocephalus NNP 37602 690 4 _ _ NNP 37602 690 5 , , , 37602 690 6 skulls skull NNS 37602 690 7 of of IN 37602 690 8 the the DT 37602 690 9 members member NNS 37602 690 10 of of IN 37602 690 11 the the DT 37602 690 12 genus genus NNP 37602 690 13 _ _ NNP 37602 690 14 Phrynohyas Phrynohyas NNP 37602 690 15 _ _ NNP 37602 690 16 are be VBP 37602 690 17 broader broad JJR 37602 690 18 than than IN 37602 690 19 long long JJ 37602 690 20 , , , 37602 690 21 have have VBP 37602 690 22 extensive extensive JJ 37602 690 23 dermal dermal JJ 37602 690 24 roofing roofing NN 37602 690 25 bones bone NNS 37602 690 26 , , , 37602 690 27 and and CC 37602 690 28 have have VBP 37602 690 29 posterolaterally posterolaterally RB 37602 690 30 oriented orient VBN 37602 690 31 parasphenoid parasphenoid NN 37602 690 32 alae alae NNP 37602 690 33 . . . 37602 691 1 In in IN 37602 691 2 contrast contrast NN 37602 691 3 to to IN 37602 691 4 _ _ NNP 37602 691 5 Osteocephalus Osteocephalus NNP 37602 691 6 _ _ NNP 37602 691 7 , , , 37602 691 8 the the DT 37602 691 9 dentigerous dentigerous JJ 37602 691 10 processes process NNS 37602 691 11 of of IN 37602 691 12 the the DT 37602 691 13 prevomers prevomer NNS 37602 691 14 are be VBP 37602 691 15 curved curved JJ 37602 691 16 , , , 37602 691 17 rather rather RB 37602 691 18 than than IN 37602 691 19 angular angular JJ 37602 691 20 in in IN 37602 691 21 _ _ NNP 37602 691 22 Phrynohyas Phrynohyas NNP 37602 691 23 _ _ NNP 37602 691 24 . . . 37602 692 1 Furthermore furthermore RB 37602 692 2 , , , 37602 692 3 the the DT 37602 692 4 latter latter JJ 37602 692 5 genus genus NN 37602 692 6 is be VBZ 37602 692 7 singularly singularly RB 37602 692 8 distinguished distinguish VBN 37602 692 9 from from IN 37602 692 10 _ _ NNP 37602 692 11 Osteocephalus Osteocephalus NNP 37602 692 12 _ _ NNP 37602 692 13 , , , 37602 692 14 _ _ NNP 37602 692 15 Argenteohyla Argenteohyla NNP 37602 692 16 _ _ NNP 37602 692 17 , , , 37602 692 18 and and CC 37602 692 19 _ _ NNP 37602 692 20 Trachycephalus Trachycephalus NNP 37602 692 21 _ _ NNP 37602 692 22 by by IN 37602 692 23 having have VBG 37602 692 24 extensively extensively RB 37602 692 25 developed develop VBN 37602 692 26 parotoid parotoid NN 37602 692 27 glands gland NNS 37602 692 28 that that WDT 37602 692 29 produce produce VBP 37602 692 30 a a DT 37602 692 31 viscous viscous JJ 37602 692 32 , , , 37602 692 33 milky milky NNP 37602 692 34 volatile volatile JJ 37602 692 35 secretion secretion NN 37602 692 36 . . . 37602 693 1 _ _ NNP 37602 693 2 Trachycephalus Trachycephalus NNP 37602 693 3 _ _ NNP 37602 693 4 is be VBZ 37602 693 5 the the DT 37602 693 6 most most RBS 37602 693 7 readily readily RB 37602 693 8 identifiable identifiable JJ 37602 693 9 of of IN 37602 693 10 the the DT 37602 693 11 four four CD 37602 693 12 genera genera NN 37602 693 13 under under IN 37602 693 14 discussion discussion NN 37602 693 15 . . . 37602 694 1 Members member NNS 37602 694 2 of of IN 37602 694 3 this this DT 37602 694 4 genus genus NN 37602 694 5 are be VBP 37602 694 6 large large JJ 37602 694 7 frogs frog NNS 37602 694 8 with with IN 37602 694 9 heavily heavily RB 37602 694 10 casqued casque VBN 37602 694 11 and and CC 37602 694 12 co co JJ 37602 694 13 - - JJ 37602 694 14 ossified ossified JJ 37602 694 15 skulls skull NNS 37602 694 16 ( ( -LRB- 37602 694 17 Trueb Trueb NNP 37602 694 18 , , , 37602 694 19 1970a 1970a CD 37602 694 20 ) ) -RRB- 37602 694 21 . . . 37602 695 1 The the DT 37602 695 2 dermal dermal JJ 37602 695 3 roofing roofing NN 37602 695 4 bones bone NNS 37602 695 5 bear bear VBP 37602 695 6 ornate ornate JJ 37602 695 7 and and CC 37602 695 8 characteristic characteristic JJ 37602 695 9 patterns pattern NNS 37602 695 10 of of IN 37602 695 11 sculpturing sculpturing NN 37602 695 12 . . . 37602 696 1 The the DT 37602 696 2 medial medial JJ 37602 696 3 ramus ramus NN 37602 696 4 of of IN 37602 696 5 the the DT 37602 696 6 pterygoid pterygoid NN 37602 696 7 does do VBZ 37602 696 8 not not RB 37602 696 9 articulate articulate VB 37602 696 10 with with IN 37602 696 11 the the DT 37602 696 12 otic otic JJ 37602 696 13 capsule capsule NN 37602 696 14 , , , 37602 696 15 and and CC 37602 696 16 the the DT 37602 696 17 parasphenoid parasphenoid NN 37602 696 18 alae alae NNP 37602 696 19 are be VBP 37602 696 20 laterally laterally RB 37602 696 21 , , , 37602 696 22 rather rather RB 37602 696 23 than than IN 37602 696 24 posterolaterally posterolaterally RB 37602 696 25 , , , 37602 696 26 oriented orient VBN 37602 696 27 . . . 37602 697 1 A a DT 37602 697 2 dermal dermal JJ 37602 697 3 sphenethmoid sphenethmoid NN 37602 697 4 is be VBZ 37602 697 5 present present JJ 37602 697 6 , , , 37602 697 7 and and CC 37602 697 8 the the DT 37602 697 9 parasphenoid parasphenoid NN 37602 697 10 bears bear VBZ 37602 697 11 odontoids odontoid NNS 37602 697 12 . . . 37602 698 1 The the DT 37602 698 2 basic basic JJ 37602 698 3 structure structure NN 37602 698 4 of of IN 37602 698 5 the the DT 37602 698 6 skull skull NN 37602 698 7 has have VBZ 37602 698 8 many many JJ 37602 698 9 characters character NNS 37602 698 10 in in IN 37602 698 11 common common NN 37602 698 12 with with IN 37602 698 13 both both DT 37602 698 14 _ _ NNP 37602 698 15 Osteocephalus Osteocephalus NNP 37602 698 16 _ _ NNP 37602 698 17 and and CC 37602 698 18 _ _ NNP 37602 698 19 Phrynohyas Phrynohyas NNP 37602 698 20 _ _ NNP 37602 698 21 . . . 37602 699 1 The the DT 37602 699 2 obvious obvious JJ 37602 699 3 modifications modification NNS 37602 699 4 of of IN 37602 699 5 dermal dermal JJ 37602 699 6 roofing roofing NN 37602 699 7 bones bone NNS 37602 699 8 and and CC 37602 699 9 of of IN 37602 699 10 palatal palatal JJ 37602 699 11 and and CC 37602 699 12 suspensory suspensory JJ 37602 699 13 elements element NNS 37602 699 14 seem seem VBP 37602 699 15 to to TO 37602 699 16 be be VB 37602 699 17 specializations specialization NNS 37602 699 18 adapting adapt VBG 37602 699 19 members member NNS 37602 699 20 of of IN 37602 699 21 the the DT 37602 699 22 genus genus NNP 37602 699 23 _ _ NNP 37602 699 24 Trachycephalus Trachycephalus NNP 37602 699 25 _ _ NNP 37602 699 26 to to IN 37602 699 27 their -PRON- PRP$ 37602 699 28 peculiar peculiar JJ 37602 699 29 phragmotic phragmotic JJ 37602 699 30 habits habit NNS 37602 699 31 . . . 37602 700 1 The the DT 37602 700 2 vocal vocal JJ 37602 700 3 sac sac NN 37602 700 4 structure structure NN 37602 700 5 of of IN 37602 700 6 _ _ NNP 37602 700 7 Trachycephalus Trachycephalus NNP 37602 700 8 _ _ NNP 37602 700 9 is be VBZ 37602 700 10 like like IN 37602 700 11 that that DT 37602 700 12 of of IN 37602 700 13 _ _ NNP 37602 700 14 Phrynohyas Phrynohyas NNP 37602 700 15 _ _ NNP 37602 700 16 and and CC 37602 700 17 therefore therefore RB 37602 700 18 further further RB 37602 700 19 distinguishes distinguish VBZ 37602 700 20 it -PRON- PRP 37602 700 21 from from IN 37602 700 22 _ _ NNP 37602 700 23 Osteocephalus Osteocephalus NNP 37602 700 24 _ _ NNP 37602 700 25 . . . 37602 701 1 Morphologically morphologically RB 37602 701 2 , , , 37602 701 3 _ _ NNP 37602 701 4 Osteocephalus Osteocephalus NNP 37602 701 5 _ _ NNP 37602 701 6 seems seem VBZ 37602 701 7 to to TO 37602 701 8 be be VB 37602 701 9 sufficiently sufficiently RB 37602 701 10 diverse diverse JJ 37602 701 11 and and CC 37602 701 12 generalized generalize VBD 37602 701 13 so so IN 37602 701 14 as as IN 37602 701 15 to to TO 37602 701 16 represent represent VB 37602 701 17 a a DT 37602 701 18 modern modern JJ 37602 701 19 derivative derivative NN 37602 701 20 of of IN 37602 701 21 an an DT 37602 701 22 ancestral ancestral JJ 37602 701 23 type type NN 37602 701 24 which which WDT 37602 701 25 might may MD 37602 701 26 have have VB 37602 701 27 given give VBN 37602 701 28 rise rise NN 37602 701 29 to to IN 37602 701 30 _ _ NNP 37602 701 31 Phrynohyas Phrynohyas NNP 37602 701 32 _ _ NNP 37602 701 33 , , , 37602 701 34 _ _ NNP 37602 701 35 Trachycephalus Trachycephalus NNP 37602 701 36 _ _ NNP 37602 701 37 , , , 37602 701 38 and and CC 37602 701 39 _ _ NNP 37602 701 40 Argenteohyla Argenteohyla NNP 37602 701 41 _ _ NNP 37602 701 42 . . . 37602 702 1 The the DT 37602 702 2 specialized specialized JJ 37602 702 3 vocal vocal JJ 37602 702 4 sac sac NN 37602 702 5 structure structure NN 37602 702 6 in in IN 37602 702 7 _ _ NNP 37602 702 8 Phrynohyas Phrynohyas NNP 37602 702 9 _ _ NNP 37602 702 10 and and CC 37602 702 11 _ _ NNP 37602 702 12 Trachycephalus Trachycephalus NNP 37602 702 13 _ _ NNP 37602 702 14 suggests suggest VBZ 37602 702 15 that that IN 37602 702 16 these these DT 37602 702 17 two two CD 37602 702 18 genera genera NN 37602 702 19 may may MD 37602 702 20 be be VB 37602 702 21 rather rather RB 37602 702 22 closely closely RB 37602 702 23 allied ally VBN 37602 702 24 and and CC 37602 702 25 represent represent VB 37602 702 26 a a DT 37602 702 27 single single JJ 37602 702 28 phyletic phyletic JJ 37602 702 29 line line NN 37602 702 30 from from IN 37602 702 31 an an DT 37602 702 32 ancestral ancestral JJ 37602 702 33 stock stock NN 37602 702 34 similar similar JJ 37602 702 35 to to IN 37602 702 36 _ _ NNP 37602 702 37 Osteocephalus Osteocephalus NNP 37602 702 38 _ _ NNP 37602 702 39 . . . 37602 703 1 _ _ NNP 37602 703 2 Argenteohyla Argenteohyla NNP 37602 703 3 _ _ NNP 37602 703 4 is be VBZ 37602 703 5 quite quite RB 37602 703 6 distinct distinct JJ 37602 703 7 from from IN 37602 703 8 _ _ NNP 37602 703 9 Phrynohyas Phrynohyas NNP 37602 703 10 _ _ NNP 37602 703 11 and and CC 37602 703 12 _ _ NNP 37602 703 13 Trachycephalus Trachycephalus NNP 37602 703 14 _ _ NNP 37602 703 15 and and CC 37602 703 16 apparently apparently RB 37602 703 17 represents represent VBZ 37602 703 18 a a DT 37602 703 19 distinct distinct JJ 37602 703 20 phyletic phyletic JJ 37602 703 21 line line NN 37602 703 22 from from IN 37602 703 23 the the DT 37602 703 24 ancestral ancestral JJ 37602 703 25 stock stock NN 37602 703 26 . . . 37602 704 1 OCCURRENCE OCCURRENCE NNP 37602 704 2 OF of IN 37602 704 3 _ _ NNP 37602 704 4 OSTEOCEPHALUS OSTEOCEPHALUS NNP 37602 704 5 _ _ XX 37602 704 6 IN in IN 37602 704 7 AMAZONIAN AMAZONIAN NNP 37602 704 8 ECUADOR ECUADOR NNP 37602 704 9 All all DT 37602 704 10 of of IN 37602 704 11 our -PRON- PRP$ 37602 704 12 observations observation NNS 37602 704 13 on on IN 37602 704 14 members member NNS 37602 704 15 of of IN 37602 704 16 this this DT 37602 704 17 genus genus NN 37602 704 18 have have VBP 37602 704 19 been be VBN 37602 704 20 made make VBN 37602 704 21 at at IN 37602 704 22 four four CD 37602 704 23 localities locality NNS 37602 704 24 : : : 37602 704 25 1 1 LS 37602 704 26 ) ) -RRB- 37602 704 27 Santa Santa NNP 37602 704 28 Cecilia Cecilia NNP 37602 704 29 at at IN 37602 704 30 an an DT 37602 704 31 elevation elevation NN 37602 704 32 of of IN 37602 704 33 340 340 CD 37602 704 34 meters meter NNS 37602 704 35 on on IN 37602 704 36 the the DT 37602 704 37 Río Río NNP 37602 704 38 Aguarico Aguarico NNP 37602 704 39 , , , 37602 704 40 a a DT 37602 704 41 tributary tributary NN 37602 704 42 of of IN 37602 704 43 the the DT 37602 704 44 Río Río NNP 37602 704 45 Napo Napo NNP 37602 704 46 , , , 37602 704 47 2 2 LS 37602 704 48 ) ) -RRB- 37602 704 49 Lago Lago NNP 37602 704 50 Agrio Agrio NNP 37602 704 51 , , , 37602 704 52 330 330 CD 37602 704 53 meters meter NNS 37602 704 54 , , , 37602 704 55 about about RB 37602 704 56 14 14 CD 37602 704 57 kilometers kilometer NNS 37602 704 58 east east RB 37602 704 59 of of IN 37602 704 60 Santa Santa NNP 37602 704 61 Cecilia Cecilia NNP 37602 704 62 , , , 37602 704 63 3 3 LS 37602 704 64 ) ) -RRB- 37602 704 65 Puerto Puerto NNP 37602 704 66 Libre Libre NNP 37602 704 67 , , , 37602 704 68 570 570 CD 37602 704 69 meters meter NNS 37602 704 70 , , , 37602 704 71 on on IN 37602 704 72 the the DT 37602 704 73 Río Río NNP 37602 704 74 Aguarico Aguarico NNP 37602 704 75 just just RB 37602 704 76 east east RB 37602 704 77 of of IN 37602 704 78 its -PRON- PRP$ 37602 704 79 formation formation NN 37602 704 80 by by IN 37602 704 81 the the DT 37602 704 82 confluence confluence NN 37602 704 83 of of IN 37602 704 84 the the DT 37602 704 85 Río Río NNP 37602 704 86 Cofanes Cofanes NNP 37602 704 87 and and CC 37602 704 88 Río Río NNP 37602 704 89 Chingua Chingua NNP 37602 704 90 , , , 37602 704 91 and and CC 37602 704 92 4 4 LS 37602 704 93 ) ) -RRB- 37602 704 94 south south JJ 37602 704 95 slope slope NN 37602 704 96 of of IN 37602 704 97 the the DT 37602 704 98 Cordillera Cordillera NNP 37602 704 99 del del NNP 37602 704 100 Dué Dué NNP 37602 704 101 , , , 37602 704 102 above above IN 37602 704 103 the the DT 37602 704 104 Río Río NNP 37602 704 105 Coca Coca NNP 37602 704 106 , , , 37602 704 107 1150 1150 CD 37602 704 108 meters meter NNS 37602 704 109 . . . 37602 705 1 _ _ NNP 37602 705 2 Osteocephalus Osteocephalus NNP 37602 705 3 leprieurii leprieurii JJ 37602 705 4 _ _ NNP 37602 705 5 was be VBD 37602 705 6 found find VBN 37602 705 7 at at IN 37602 705 8 all all RB 37602 705 9 four four CD 37602 705 10 localities locality NNS 37602 705 11 , , , 37602 705 12 and and CC 37602 705 13 _ _ NNP 37602 705 14 buckleyi buckleyi NN 37602 705 15 _ _ NNP 37602 705 16 was be VBD 37602 705 17 found find VBN 37602 705 18 at at RB 37602 705 19 all all RB 37602 705 20 but but IN 37602 705 21 the the DT 37602 705 22 last last JJ 37602 705 23 ; ; : 37602 705 24 _ _ NNP 37602 705 25 taurinus taurinus NN 37602 705 26 _ _ NNP 37602 705 27 was be VBD 37602 705 28 found find VBN 37602 705 29 at at IN 37602 705 30 Santa Santa NNP 37602 705 31 Cecilia Cecilia NNP 37602 705 32 and and CC 37602 705 33 Lago Lago NNP 37602 705 34 Agrio Agrio NNP 37602 705 35 , , , 37602 705 36 and and CC 37602 705 37 _ _ NNP 37602 705 38 verrucigerus verrucigerus NN 37602 705 39 _ _ NNP 37602 705 40 was be VBD 37602 705 41 found find VBN 37602 705 42 only only RB 37602 705 43 in in IN 37602 705 44 the the DT 37602 705 45 Cordillera Cordillera NNP 37602 705 46 del del NNP 37602 705 47 Dué Dué NNP 37602 705 48 . . . 37602 706 1 Our -PRON- PRP$ 37602 706 2 data datum NNS 37602 706 3 are be VBP 37602 706 4 based base VBN 37602 706 5 on on IN 37602 706 6 collections collection NNS 37602 706 7 of of IN 37602 706 8 113 113 CD 37602 706 9 frogs frog NNS 37602 706 10 and and CC 37602 706 11 three three CD 37602 706 12 lots lot NNS 37602 706 13 of of IN 37602 706 14 tadpoles tadpole NNS 37602 706 15 , , , 37602 706 16 as as RB 37602 706 17 well well RB 37602 706 18 as as IN 37602 706 19 observations observation NNS 37602 706 20 on on IN 37602 706 21 calling call VBG 37602 706 22 sites site NNS 37602 706 23 and and CC 37602 706 24 young young JJ 37602 706 25 . . . 37602 707 1 The the DT 37602 707 2 observations observation NNS 37602 707 3 are be VBP 37602 707 4 summarized summarize VBN 37602 707 5 by by IN 37602 707 6 species specie NNS 37602 707 7 , , , 37602 707 8 as as IN 37602 707 9 follows follow VBZ 37602 707 10 : : : 37602 707 11 _ _ NNP 37602 707 12 Osteocephalus Osteocephalus NNP 37602 707 13 buckleyi._--No buckleyi._--no NN 37602 707 14 breeding breeding NN 37602 707 15 activity activity NN 37602 707 16 was be VBD 37602 707 17 observed observe VBN 37602 707 18 . . . 37602 708 1 Males male NNS 37602 708 2 were be VBD 37602 708 3 found find VBN 37602 708 4 only only RB 37602 708 5 at at IN 37602 708 6 night night NN 37602 708 7 in in IN 37602 708 8 March March NNP 37602 708 9 , , , 37602 708 10 June June NNP 37602 708 11 , , , 37602 708 12 and and CC 37602 708 13 July July NNP 37602 708 14 . . . 37602 709 1 One one CD 37602 709 2 was be VBD 37602 709 3 perched perch VBN 37602 709 4 on on IN 37602 709 5 a a DT 37602 709 6 _ _ NNP 37602 709 7 Heliconia Heliconia NNP 37602 709 8 _ _ NNP 37602 709 9 leaf leaf NN 37602 709 10 in in IN 37602 709 11 a a DT 37602 709 12 swamp swamp NN 37602 709 13 at at IN 37602 709 14 Puerto Puerto NNP 37602 709 15 Libre Libre NNP 37602 709 16 , , , 37602 709 17 and and CC 37602 709 18 two two CD 37602 709 19 were be VBD 37602 709 20 on on IN 37602 709 21 bushes bush NNS 37602 709 22 in in IN 37602 709 23 the the DT 37602 709 24 forest forest NN 37602 709 25 at at IN 37602 709 26 Santa Santa NNP 37602 709 27 Cecilia Cecilia NNP 37602 709 28 . . . 37602 710 1 A a DT 37602 710 2 gravid gravid JJ 37602 710 3 female female NN 37602 710 4 was be VBD 37602 710 5 found find VBN 37602 710 6 on on IN 37602 710 7 a a DT 37602 710 8 recently recently RB 37602 710 9 felled fell VBN 37602 710 10 tree tree NN 37602 710 11 at at IN 37602 710 12 Lago Lago NNP 37602 710 13 Agrio Agrio NNP 37602 710 14 on on IN 37602 710 15 the the DT 37602 710 16 night night NN 37602 710 17 of of IN 37602 710 18 12 12 CD 37602 710 19 May May NNP 37602 710 20 1969 1969 CD 37602 710 21 . . . 37602 711 1 _ _ NNP 37602 711 2 Osteocephalus Osteocephalus NNP 37602 711 3 leprieurii._--Males leprieurii._--male NNS 37602 711 4 were be VBD 37602 711 5 heard hear VBN 37602 711 6 calling call VBG 37602 711 7 sporadically sporadically RB 37602 711 8 at at IN 37602 711 9 Puerto Puerto NNP 37602 711 10 Libre Libre NNP 37602 711 11 in in IN 37602 711 12 July July NNP 37602 711 13 1968 1968 CD 37602 711 14 , , , 37602 711 15 and and CC 37602 711 16 at at IN 37602 711 17 Santa Santa NNP 37602 711 18 Cecilia Cecilia NNP 37602 711 19 in in IN 37602 711 20 May May NNP 37602 711 21 1969 1969 CD 37602 711 22 . . . 37602 712 1 A a DT 37602 712 2 small small JJ 37602 712 3 chorus chorus NN 37602 712 4 was be VBD 37602 712 5 found find VBN 37602 712 6 on on IN 37602 712 7 the the DT 37602 712 8 night night NN 37602 712 9 of of IN 37602 712 10 12 12 CD 37602 712 11 May May NNP 37602 712 12 1969 1969 CD 37602 712 13 at at IN 37602 712 14 Lago Lago NNP 37602 712 15 Agrio Agrio NNP 37602 712 16 , , , 37602 712 17 where where WRB 37602 712 18 the the DT 37602 712 19 frogs frog NNS 37602 712 20 were be VBD 37602 712 21 perched perch VBN 37602 712 22 on on IN 37602 712 23 branches branch NNS 37602 712 24 of of IN 37602 712 25 fallen fall VBN 37602 712 26 trees tree NNS 37602 712 27 over over IN 37602 712 28 a a DT 37602 712 29 temporary temporary JJ 37602 712 30 pool pool NN 37602 712 31 . . . 37602 713 1 The the DT 37602 713 2 call call NN 37602 713 3 is be VBZ 37602 713 4 a a DT 37602 713 5 soft soft JJ 37602 713 6 rattling rattle VBG 37602 713 7 chuckle chuckle NN 37602 713 8 . . . 37602 714 1 In in IN 37602 714 2 late late JJ 37602 714 3 April April NNP 37602 714 4 and and CC 37602 714 5 May May NNP 37602 714 6 many many JJ 37602 714 7 gravid gravid JJ 37602 714 8 females female NNS 37602 714 9 and and CC 37602 714 10 males male NNS 37602 714 11 with with IN 37602 714 12 well well RB 37602 714 13 - - HYPH 37602 714 14 developed develop VBN 37602 714 15 nuptial nuptial JJ 37602 714 16 excrescences excrescence NNS 37602 714 17 were be VBD 37602 714 18 obtained obtain VBN 37602 714 19 from from IN 37602 714 20 trees tree NNS 37602 714 21 as as IN 37602 714 22 they -PRON- PRP 37602 714 23 were be VBD 37602 714 24 felled fell VBN 37602 714 25 at at IN 37602 714 26 Lago Lago NNP 37602 714 27 Agrio Agrio NNP 37602 714 28 . . . 37602 715 1 The the DT 37602 715 2 reproductive reproductive JJ 37602 715 3 condition condition NN 37602 715 4 of of IN 37602 715 5 the the DT 37602 715 6 frogs frog NNS 37602 715 7 indicates indicate VBZ 37602 715 8 that that IN 37602 715 9 they -PRON- PRP 37602 715 10 probably probably RB 37602 715 11 breed breed VBP 37602 715 12 in in IN 37602 715 13 May. May NNP 37602 716 1 One one CD 37602 716 2 individual individual NN 37602 716 3 called call VBD 37602 716 4 nearly nearly RB 37602 716 5 every every DT 37602 716 6 night night NN 37602 716 7 from from IN 37602 716 8 a a DT 37602 716 9 large large JJ 37602 716 10 tree tree NN 37602 716 11 at at IN 37602 716 12 Puerto Puerto NNP 37602 716 13 Libre Libre NNP 37602 716 14 between between IN 37602 716 15 4 4 CD 37602 716 16 - - SYM 37602 716 17 17 17 CD 37602 716 18 July July NNP 37602 716 19 1968 1968 CD 37602 716 20 . . . 37602 717 1 The the DT 37602 717 2 tree tree NN 37602 717 3 was be VBD 37602 717 4 felled fell VBN 37602 717 5 on on IN 37602 717 6 the the DT 37602 717 7 latter latter JJ 37602 717 8 date date NN 37602 717 9 , , , 37602 717 10 but but CC 37602 717 11 no no DT 37602 717 12 frog frog NN 37602 717 13 was be VBD 37602 717 14 found find VBN 37602 717 15 . . . 37602 718 1 Two two CD 37602 718 2 nights night NNS 37602 718 3 later later RB 37602 718 4 apparently apparently RB 37602 718 5 the the DT 37602 718 6 same same JJ 37602 718 7 individual individual NN 37602 718 8 called call VBD 37602 718 9 from from IN 37602 718 10 a a DT 37602 718 11 bromeliad bromeliad NN 37602 718 12 at at IN 37602 718 13 a a DT 37602 718 14 height height NN 37602 718 15 of of IN 37602 718 16 about about IN 37602 718 17 10 10 CD 37602 718 18 m m NNS 37602 718 19 on on IN 37602 718 20 a a DT 37602 718 21 large large JJ 37602 718 22 bamboo bamboo NN 37602 718 23 adjacent adjacent JJ 37602 718 24 to to IN 37602 718 25 the the DT 37602 718 26 felled fell VBN 37602 718 27 tree tree NN 37602 718 28 ; ; : 37602 718 29 the the DT 37602 718 30 frog frog NN 37602 718 31 was be VBD 37602 718 32 collected collect VBN 37602 718 33 when when WRB 37602 718 34 the the DT 37602 718 35 bamboo bamboo NN 37602 718 36 was be VBD 37602 718 37 cut cut VBN 37602 718 38 down down RP 37602 718 39 . . . 37602 719 1 Throughout throughout IN 37602 719 2 the the DT 37602 719 3 rainy rainy JJ 37602 719 4 months month NNS 37602 719 5 that that WDT 37602 719 6 we -PRON- PRP 37602 719 7 have have VBP 37602 719 8 worked work VBN 37602 719 9 in in IN 37602 719 10 Ecuador Ecuador NNP 37602 719 11 ( ( -LRB- 37602 719 12 April April NNP 37602 719 13 - - HYPH 37602 719 14 August August NNP 37602 719 15 ) ) -RRB- 37602 719 16 we -PRON- PRP 37602 719 17 have have VBP 37602 719 18 found find VBN 37602 719 19 occasional occasional JJ 37602 719 20 individuals individual NNS 37602 719 21 perched perch VBN 37602 719 22 on on IN 37602 719 23 bushes bush NNS 37602 719 24 or or CC 37602 719 25 low low JJ 37602 719 26 trees tree NNS 37602 719 27 at at IN 37602 719 28 night night NN 37602 719 29 . . . 37602 720 1 Large large JJ 37602 720 2 numbers number NNS 37602 720 3 of of IN 37602 720 4 adults adult NNS 37602 720 5 were be VBD 37602 720 6 observed observe VBN 37602 720 7 only only RB 37602 720 8 during during IN 37602 720 9 a a DT 37602 720 10 clearing clearing NN 37602 720 11 operation operation NN 37602 720 12 which which WDT 37602 720 13 resulted result VBD 37602 720 14 in in IN 37602 720 15 the the DT 37602 720 16 felling felling NN 37602 720 17 of of IN 37602 720 18 many many JJ 37602 720 19 large large JJ 37602 720 20 trees tree NNS 37602 720 21 . . . 37602 721 1 Thus thus RB 37602 721 2 , , , 37602 721 3 it -PRON- PRP 37602 721 4 seems seem VBZ 37602 721 5 likely likely JJ 37602 721 6 that that IN 37602 721 7 _ _ NNP 37602 721 8 leprieurii leprieurii JJ 37602 721 9 _ _ NNP 37602 721 10 is be VBZ 37602 721 11 a a DT 37602 721 12 tree tree NN 37602 721 13 - - HYPH 37602 721 14 top top NN 37602 721 15 inhabitant inhabitant NN 37602 721 16 . . . 37602 722 1 A a DT 37602 722 2 partially partially RB 37602 722 3 digested digested JJ 37602 722 4 adult adult NN 37602 722 5 male male NN 37602 722 6 was be VBD 37602 722 7 removed remove VBN 37602 722 8 from from IN 37602 722 9 the the DT 37602 722 10 stomach stomach NN 37602 722 11 of of IN 37602 722 12 a a DT 37602 722 13 _ _ NNP 37602 722 14 Hemiphractus Hemiphractus NNP 37602 722 15 proboscideus proboscideus NN 37602 722 16 _ _ NNP 37602 722 17 . . . 37602 723 1 At at IN 37602 723 2 Santa Santa NNP 37602 723 3 Cecilia Cecilia NNP 37602 723 4 many many JJ 37602 723 5 recently recently RB 37602 723 6 metamorphosed metamorphose VBN 37602 723 7 young young JJ 37602 723 8 and and CC 37602 723 9 juveniles juvenile NNS 37602 723 10 were be VBD 37602 723 11 found find VBN 37602 723 12 in in IN 37602 723 13 June June NNP 37602 723 14 and and CC 37602 723 15 July July NNP 37602 723 16 1968 1968 CD 37602 723 17 . . . 37602 724 1 Most Most JJS 37602 724 2 of of IN 37602 724 3 these these DT 37602 724 4 were be VBD 37602 724 5 on on IN 37602 724 6 low low JJ 37602 724 7 bushes bush NNS 37602 724 8 or or CC 37602 724 9 herbs herb NNS 37602 724 10 in in IN 37602 724 11 swamp swamp NN 37602 724 12 forest forest NN 37602 724 13 at at IN 37602 724 14 night night NN 37602 724 15 ; ; : 37602 724 16 some some DT 37602 724 17 were be VBD 37602 724 18 found find VBN 37602 724 19 in in IN 37602 724 20 unfolded unfolded JJ 37602 724 21 _ _ NNP 37602 724 22 Heliconia Heliconia NNP 37602 724 23 _ _ NNP 37602 724 24 leaves leave VBZ 37602 724 25 by by IN 37602 724 26 day day NN 37602 724 27 , , , 37602 724 28 and and CC 37602 724 29 one one CD 37602 724 30 was be VBD 37602 724 31 observed observe VBN 37602 724 32 on on IN 37602 724 33 the the DT 37602 724 34 forest forest NN 37602 724 35 floor floor NN 37602 724 36 by by IN 37602 724 37 day day NN 37602 724 38 . . . 37602 725 1 Snout snout NN 37602 725 2 - - HYPH 37602 725 3 vent vent NN 37602 725 4 lengths length NNS 37602 725 5 of of IN 37602 725 6 18 18 CD 37602 725 7 specimens specimen NNS 37602 725 8 are be VBP 37602 725 9 12.3 12.3 CD 37602 725 10 - - SYM 37602 725 11 17.0 17.0 CD 37602 725 12 ( ( -LRB- 37602 725 13 mean mean NNP 37602 725 14 15.1 15.1 CD 37602 725 15 ) ) -RRB- 37602 725 16 mm mm NNP 37602 725 17 . . . 37602 726 1 The the DT 37602 726 2 smaller small JJR 37602 726 3 frogs frog NNS 37602 726 4 were be VBD 37602 726 5 recently recently RB 37602 726 6 metamorphosed metamorphose VBN 37602 726 7 as as IN 37602 726 8 evidenced evidence VBN 37602 726 9 by by IN 37602 726 10 the the DT 37602 726 11 melanophore melanophore NN 37602 726 12 deposits deposit NNS 37602 726 13 above above IN 37602 726 14 the the DT 37602 726 15 vent vent NN 37602 726 16 . . . 37602 727 1 The the DT 37602 727 2 coloration coloration NN 37602 727 3 of of IN 37602 727 4 the the DT 37602 727 5 young young JJ 37602 727 6 is be VBZ 37602 727 7 strikingly strikingly RB 37602 727 8 different different JJ 37602 727 9 from from IN 37602 727 10 that that DT 37602 727 11 of of IN 37602 727 12 the the DT 37602 727 13 adults adult NNS 37602 727 14 ( ( -LRB- 37602 727 15 see see VB 37602 727 16 account account NN 37602 727 17 of of IN 37602 727 18 _ _ NNP 37602 727 19 O. O. NNP 37602 728 1 leprieurii leprieurii NNP 37602 728 2 _ _ NNP 37602 728 3 ) ) -RRB- 37602 728 4 , , , 37602 728 5 so so RB 37602 728 6 the the DT 37602 728 7 association association NN 37602 728 8 of of IN 37602 728 9 the the DT 37602 728 10 young young JJ 37602 728 11 and and CC 37602 728 12 adults adult NNS 37602 728 13 was be VBD 37602 728 14 not not RB 37602 728 15 made make VBN 37602 728 16 until until IN 37602 728 17 individuals individual NNS 37602 728 18 with with IN 37602 728 19 intermediate intermediate JJ 37602 728 20 patterns pattern NNS 37602 728 21 were be VBD 37602 728 22 obtained obtain VBN 37602 728 23 at at IN 37602 728 24 Lago Lago NNP 37602 728 25 Agrio Agrio NNP 37602 728 26 in in IN 37602 728 27 May May NNP 37602 728 28 1969 1969 CD 37602 728 29 . . . 37602 729 1 Probably probably RB 37602 729 2 juveniles juvenile NNS 37602 729 3 obtained obtain VBN 37602 729 4 in in IN 37602 729 5 June June NNP 37602 729 6 and and CC 37602 729 7 July July NNP 37602 729 8 are be VBP 37602 729 9 the the DT 37602 729 10 offspring offspring NN 37602 729 11 of of IN 37602 729 12 an an DT 37602 729 13 April April NNP 37602 729 14 or or CC 37602 729 15 May May NNP 37602 729 16 breeding breeding NN 37602 729 17 . . . 37602 730 1 We -PRON- PRP 37602 730 2 have have VBP 37602 730 3 been be VBN 37602 730 4 unable unable JJ 37602 730 5 to to TO 37602 730 6 associate associate VB 37602 730 7 tadpoles tadpole NNS 37602 730 8 with with IN 37602 730 9 this this DT 37602 730 10 species specie NNS 37602 730 11 . . . 37602 731 1 _ _ NNP 37602 731 2 Osteocephalus osteocephalus JJ 37602 731 3 taurinus._--A taurinus._--A NNP 37602 731 4 small small JJ 37602 731 5 chorus chorus NN 37602 731 6 occurred occur VBD 37602 731 7 at at IN 37602 731 8 Lago Lago NNP 37602 731 9 Agrio Agrio NNP 37602 731 10 on on IN 37602 731 11 12 12 CD 37602 731 12 May May NNP 37602 731 13 1969 1969 CD 37602 731 14 . . . 37602 732 1 Males male NNS 37602 732 2 were be VBD 37602 732 3 calling call VBG 37602 732 4 from from IN 37602 732 5 the the DT 37602 732 6 ground ground NN 37602 732 7 adjacent adjacent JJ 37602 732 8 to to IN 37602 732 9 a a DT 37602 732 10 small small JJ 37602 732 11 pool pool NN 37602 732 12 amidst amidst IN 37602 732 13 recently recently RB 37602 732 14 felled fell VBD 37602 732 15 trees tree NNS 37602 732 16 . . . 37602 733 1 The the DT 37602 733 2 males male NNS 37602 733 3 were be VBD 37602 733 4 very very RB 37602 733 5 wary wary JJ 37602 733 6 and and CC 37602 733 7 , , , 37602 733 8 when when WRB 37602 733 9 approached approach VBN 37602 733 10 , , , 37602 733 11 jumped jump VBD 37602 733 12 onto onto IN 37602 733 13 limbs limb NNS 37602 733 14 and and CC 37602 733 15 ran run VBD 37602 733 16 up up RP 37602 733 17 branches branch NNS 37602 733 18 ; ; : 37602 733 19 this this DT 37602 733 20 behavior behavior NN 37602 733 21 was be VBD 37602 733 22 noted note VBN 37602 733 23 by by IN 37602 733 24 Bokermann Bokermann NNP 37602 733 25 ( ( -LRB- 37602 733 26 1964 1964 CD 37602 733 27 ) ) -RRB- 37602 733 28 . . . 37602 734 1 The the DT 37602 734 2 call call NN 37602 734 3 consists consist VBZ 37602 734 4 of of IN 37602 734 5 a a DT 37602 734 6 series series NN 37602 734 7 of of IN 37602 734 8 low low RB 37602 734 9 - - HYPH 37602 734 10 pitched pitched JJ 37602 734 11 , , , 37602 734 12 short short JJ 37602 734 13 notes note NNS 37602 734 14 -- -- : 37602 734 15 like like IN 37602 734 16 a a DT 37602 734 17 slow slow JJ 37602 734 18 trill trill NN 37602 734 19 -- -- : 37602 734 20 four four CD 37602 734 21 to to TO 37602 734 22 six six CD 37602 734 23 notes note NNS 37602 734 24 per per IN 37602 734 25 call call NN 37602 734 26 group group NN 37602 734 27 . . . 37602 735 1 Call call NN 37602 735 2 groups group NNS 37602 735 3 are be VBP 37602 735 4 repeated repeat VBN 37602 735 5 two two CD 37602 735 6 , , , 37602 735 7 three three CD 37602 735 8 , , , 37602 735 9 or or CC 37602 735 10 four four CD 37602 735 11 times time NNS 37602 735 12 followed follow VBN 37602 735 13 by by IN 37602 735 14 a a DT 37602 735 15 lapse lapse NN 37602 735 16 of of IN 37602 735 17 several several JJ 37602 735 18 minutes minute NNS 37602 735 19 . . . 37602 736 1 Although although IN 37602 736 2 no no DT 37602 736 3 amplectant amplectant JJ 37602 736 4 pairs pair NNS 37602 736 5 were be VBD 37602 736 6 found find VBN 37602 736 7 , , , 37602 736 8 several several JJ 37602 736 9 gravid gravid JJ 37602 736 10 females female NNS 37602 736 11 were be VBD 37602 736 12 collected collect VBN 37602 736 13 at at IN 37602 736 14 Lago Lago NNP 37602 736 15 Agrio Agrio NNP 37602 736 16 in in IN 37602 736 17 May May NNP 37602 736 18 , , , 37602 736 19 so so IN 37602 736 20 it -PRON- PRP 37602 736 21 can can MD 37602 736 22 be be VB 37602 736 23 safely safely RB 37602 736 24 assumed assume VBN 37602 736 25 that that IN 37602 736 26 the the DT 37602 736 27 species species NN 37602 736 28 breeds breed NNS 37602 736 29 in in IN 37602 736 30 May. May NNP 37602 737 1 From from IN 37602 737 2 April April NNP 37602 737 3 through through IN 37602 737 4 July July NNP 37602 737 5 occasional occasional JJ 37602 737 6 individuals individual NNS 37602 737 7 were be VBD 37602 737 8 observed observe VBN 37602 737 9 on on IN 37602 737 10 bushes bush NNS 37602 737 11 and and CC 37602 737 12 trees tree NNS 37602 737 13 at at IN 37602 737 14 night night NN 37602 737 15 . . . 37602 738 1 During during IN 37602 738 2 clearing clearing NN 37602 738 3 operations operation NNS 37602 738 4 at at IN 37602 738 5 Lago Lago NNP 37602 738 6 Agrio Agrio NNP 37602 738 7 several several JJ 37602 738 8 individuals individual NNS 37602 738 9 were be VBD 37602 738 10 obtained obtain VBN 37602 738 11 from from IN 37602 738 12 the the DT 37602 738 13 tops top NNS 37602 738 14 of of IN 37602 738 15 trees tree NNS 37602 738 16 as as IN 37602 738 17 they -PRON- PRP 37602 738 18 were be VBD 37602 738 19 felled fell VBN 37602 738 20 . . . 37602 739 1 _ _ NNP 37602 739 2 Osteocephalus Osteocephalus NNP 37602 739 3 verrucigerus._--Observations verrucigerus._--observation NNS 37602 739 4 were be VBD 37602 739 5 made make VBN 37602 739 6 in in IN 37602 739 7 a a DT 37602 739 8 broad broad JJ 37602 739 9 , , , 37602 739 10 shallow shallow JJ 37602 739 11 ravine ravine NN 37602 739 12 , , , 37602 739 13 in in IN 37602 739 14 which which WDT 37602 739 15 there there EX 37602 739 16 was be VBD 37602 739 17 a a DT 37602 739 18 small small JJ 37602 739 19 stream stream NN 37602 739 20 . . . 37602 740 1 On on IN 37602 740 2 2 2 CD 37602 740 3 - - SYM 37602 740 4 4 4 CD 37602 740 5 August August NNP 37602 740 6 1968 1968 CD 37602 740 7 , , , 37602 740 8 males male NNS 37602 740 9 were be VBD 37602 740 10 observed observe VBN 37602 740 11 calling call VBG 37602 740 12 from from IN 37602 740 13 low low JJ 37602 740 14 bushes bush NNS 37602 740 15 and and CC 37602 740 16 rocks rock NNS 37602 740 17 at at IN 37602 740 18 the the DT 37602 740 19 edge edge NN 37602 740 20 of of IN 37602 740 21 a a DT 37602 740 22 quiet quiet JJ 37602 740 23 pool pool NN 37602 740 24 in in IN 37602 740 25 the the DT 37602 740 26 stream stream NN 37602 740 27 . . . 37602 741 1 The the DT 37602 741 2 call call NN 37602 741 3 consists consist VBZ 37602 741 4 of of IN 37602 741 5 a a DT 37602 741 6 series series NN 37602 741 7 of of IN 37602 741 8 well well RB 37602 741 9 - - HYPH 37602 741 10 pulsed pulse VBN 37602 741 11 , , , 37602 741 12 low low RB 37602 741 13 - - HYPH 37602 741 14 pitched pitched JJ 37602 741 15 , , , 37602 741 16 guttural guttural JJ 37602 741 17 notes note NNS 37602 741 18 produced produce VBN 37602 741 19 at at IN 37602 741 20 the the DT 37602 741 21 rate rate NN 37602 741 22 of of IN 37602 741 23 5 5 CD 37602 741 24 - - SYM 37602 741 25 10 10 CD 37602 741 26 per per IN 37602 741 27 minute minute NN 37602 741 28 . . . 37602 742 1 One one CD 37602 742 2 amplectant amplectant JJ 37602 742 3 pair pair NN 37602 742 4 was be VBD 37602 742 5 found find VBN 37602 742 6 at at IN 37602 742 7 the the DT 37602 742 8 base base NN 37602 742 9 of of IN 37602 742 10 a a DT 37602 742 11 bush bush NNP 37602 742 12 adjacent adjacent JJ 37602 742 13 to to IN 37602 742 14 the the DT 37602 742 15 pool pool NN 37602 742 16 on on IN 37602 742 17 3 3 CD 37602 742 18 August August NNP 37602 742 19 . . . 37602 743 1 Another another DT 37602 743 2 female female NN 37602 743 3 was be VBD 37602 743 4 found find VBN 37602 743 5 on on IN 37602 743 6 a a DT 37602 743 7 branch branch NN 37602 743 8 of of IN 37602 743 9 a a DT 37602 743 10 tree tree NN 37602 743 11 2 2 CD 37602 743 12 m m CD 37602 743 13 above above IN 37602 743 14 the the DT 37602 743 15 ground ground NN 37602 743 16 and and CC 37602 743 17 10 10 CD 37602 743 18 m m NN 37602 743 19 from from IN 37602 743 20 the the DT 37602 743 21 stream stream NN 37602 743 22 . . . 37602 744 1 Tadpoles tadpole NNS 37602 744 2 of of IN 37602 744 3 this this DT 37602 744 4 species specie NNS 37602 744 5 were be VBD 37602 744 6 found find VBN 37602 744 7 in in IN 37602 744 8 the the DT 37602 744 9 quiet quiet JJ 37602 744 10 silt silt NN 37602 744 11 - - HYPH 37602 744 12 bottomed bottom VBN 37602 744 13 pool pool NN 37602 744 14 . . . 37602 745 1 SPECIMENS SPECIMENS NNP 37602 745 2 EXAMINED examine VBD 37602 745 3 The the DT 37602 745 4 localities locality NNS 37602 745 5 for for IN 37602 745 6 each each DT 37602 745 7 of of IN 37602 745 8 the the DT 37602 745 9 specimens specimen NNS 37602 745 10 examined examine VBN 37602 745 11 are be VBP 37602 745 12 given give VBN 37602 745 13 in in IN 37602 745 14 the the DT 37602 745 15 following follow VBG 37602 745 16 paragraphs paragraph NNS 37602 745 17 . . . 37602 746 1 The the DT 37602 746 2 arrangement arrangement NN 37602 746 3 of of IN 37602 746 4 the the DT 37602 746 5 data datum NNS 37602 746 6 is be VBZ 37602 746 7 as as IN 37602 746 8 follows follow VBZ 37602 746 9 : : : 37602 746 10 alphabetically alphabetically RB 37602 746 11 by by IN 37602 746 12 country country NN 37602 746 13 , , , 37602 746 14 state state NN 37602 746 15 ( ( -LRB- 37602 746 16 department department NN 37602 746 17 or or CC 37602 746 18 province province NNP 37602 746 19 ) ) -RRB- 37602 746 20 , , , 37602 746 21 and and CC 37602 746 22 locality locality NN 37602 746 23 ; ; : 37602 746 24 alphabetically alphabetically RB 37602 746 25 by by IN 37602 746 26 the the DT 37602 746 27 first first JJ 37602 746 28 letter letter NN 37602 746 29 in in IN 37602 746 30 the the DT 37602 746 31 abbreviations abbreviation NNS 37602 746 32 for for IN 37602 746 33 the the DT 37602 746 34 museums museum NNS 37602 746 35 , , , 37602 746 36 and and CC 37602 746 37 numerically numerically RB 37602 746 38 after after IN 37602 746 39 each each DT 37602 746 40 museum museum NN 37602 746 41 abbreviation abbreviation NN 37602 746 42 . . . 37602 747 1 Specimens specimen NNS 37602 747 2 lacking lack VBG 37602 747 3 precise precise JJ 37602 747 4 locality locality NN 37602 747 5 data datum NNS 37602 747 6 are be VBP 37602 747 7 listed list VBN 37602 747 8 first first RB 37602 747 9 in in IN 37602 747 10 the the DT 37602 747 11 most most RBS 37602 747 12 restricted restrict VBN 37602 747 13 political political JJ 37602 747 14 unit unit NN 37602 747 15 possible possible JJ 37602 747 16 ; ; : 37602 747 17 localities locality NNS 37602 747 18 which which WDT 37602 747 19 have have VBP 37602 747 20 not not RB 37602 747 21 been be VBN 37602 747 22 found find VBN 37602 747 23 on on IN 37602 747 24 maps map NNS 37602 747 25 or or CC 37602 747 26 the the DT 37602 747 27 positions position NNS 37602 747 28 of of IN 37602 747 29 which which WDT 37602 747 30 are be VBP 37602 747 31 not not RB 37602 747 32 known know VBN 37602 747 33 to to IN 37602 747 34 us -PRON- PRP 37602 747 35 are be VBP 37602 747 36 given give VBN 37602 747 37 in in IN 37602 747 38 quotation quotation NN 37602 747 39 marks mark NNS 37602 747 40 . . . 37602 748 1 Where where WRB 37602 748 2 more more JJR 37602 748 3 than than IN 37602 748 4 one one CD 37602 748 5 specimen speciman NNS 37602 748 6 is be VBZ 37602 748 7 included include VBN 37602 748 8 under under IN 37602 748 9 one one CD 37602 748 10 museum museum NN 37602 748 11 number number NN 37602 748 12 , , , 37602 748 13 the the DT 37602 748 14 number number NN 37602 748 15 of of IN 37602 748 16 specimens specimen NNS 37602 748 17 is be VBZ 37602 748 18 given give VBN 37602 748 19 in in IN 37602 748 20 parentheses parenthesis NNS 37602 748 21 after after IN 37602 748 22 the the DT 37602 748 23 museum museum NN 37602 748 24 number number NN 37602 748 25 . . . 37602 749 1 Unless unless IN 37602 749 2 noted note VBN 37602 749 3 otherwise otherwise RB 37602 749 4 , , , 37602 749 5 all all DT 37602 749 6 specimens specimen NNS 37602 749 7 are be VBP 37602 749 8 alcoholics alcoholic NNS 37602 749 9 . . . 37602 750 1 _ _ NNP 37602 750 2 Osteocephalus osteocephalus JJ 37602 750 3 buckleyi buckleyi NN 37602 750 4 _ _ NNP 37602 750 5 BOLIVIA BOLIVIA NNS 37602 750 6 : : : 37602 750 7 _ _ NNP 37602 750 8 El El NNP 37602 750 9 Beni Beni NNP 37602 750 10 _ _ NNP 37602 750 11 : : : 37602 750 12 Ivón Ivón NNPS 37602 750 13 , , , 37602 750 14 BMNH BMNH NNP 37602 750 15 1967.2070 1967.2070 CD 37602 750 16 - - SYM 37602 750 17 1 1 CD 37602 750 18 . . . 37602 751 1 _ _ NNP 37602 751 2 Santa Santa NNP 37602 751 3 Cruz Cruz NNP 37602 751 4 _ _ NNP 37602 751 5 : : : 37602 751 6 Buenavista Buenavista NNP 37602 751 7 , , , 37602 751 8 CM CM NNP 37602 751 9 4333 4333 CD 37602 751 10 , , , 37602 751 11 4339 4339 CD 37602 751 12 , , , 37602 751 13 UMMZ UMMZ NNP 37602 751 14 66563 66563 CD 37602 751 15 - - SYM 37602 751 16 5 5 CD 37602 751 17 . . . 37602 752 1 BRASIL BRASIL NNP 37602 752 2 : : : 37602 752 3 _ _ NNP 37602 752 4 Amapá Amapá NNP 37602 752 5 _ _ NNP 37602 752 6 : : : 37602 752 7 No no DT 37602 752 8 specific specific JJ 37602 752 9 locality locality NN 37602 752 10 , , , 37602 752 11 WCAB wcab NN 37602 752 12 13284 13284 CD 37602 752 13 . . . 37602 753 1 COLOMBIA COLOMBIA NNP 37602 753 2 : : : 37602 753 3 _ _ NNP 37602 753 4 Amazonas Amazonas NNP 37602 753 5 _ _ NNP 37602 753 6 : : : 37602 753 7 Río Río NNP 37602 753 8 Guacaya Guacaya NNP 37602 753 9 , , , 37602 753 10 USNM USNM NNP 37602 753 11 152759 152759 CD 37602 753 12 . . . 37602 754 1 _ _ NNP 37602 754 2 Huila Huila NNP 37602 754 3 _ _ NNP 37602 754 4 : : : 37602 754 5 Acevedo Acevedo NNP 37602 754 6 , , , 37602 754 7 Río Río NNP 37602 754 8 Suaza Suaza NNP 37602 754 9 , , , 37602 754 10 FMNH FMNH NNP 37602 754 11 69702 69702 CD 37602 754 12 . . . 37602 755 1 _ _ NNP 37602 755 2 Nariño Nariño NNP 37602 755 3 _ _ NNP 37602 755 4 : : : 37602 755 5 Rumiyacu Rumiyacu NNP 37602 755 6 , , , 37602 755 7 FMNH FMNH NNP 37602 755 8 54756 54756 CD 37602 755 9 . . . 37602 756 1 _ _ NNP 37602 756 2 Meta Meta NNP 37602 756 3 _ _ NNP 37602 756 4 : : : 37602 756 5 Río Río NNP 37602 756 6 Guejar Guejar NNP 37602 756 7 , , , 37602 756 8 Campamento Campamento NNP 37602 756 9 La La NNP 37602 756 10 Macarena Macarena NNP 37602 756 11 , , , 37602 756 12 USNM USNM NNP 37602 756 13 152199 152199 CD 37602 756 14 . . . 37602 757 1 ECUADOR ECUADOR NNP 37602 757 2 : : : 37602 757 3 No no DT 37602 757 4 specific specific JJ 37602 757 5 locality locality NN 37602 757 6 , , , 37602 757 7 NHMW NHMW NNP 37602 757 8 6209 6209 CD 37602 757 9 , , , 37602 757 10 WCAB WCAB NNP 37602 757 11 35499 35499 CD 37602 757 12 . . . 37602 758 1 _ _ NNP 37602 758 2 Chimborazo Chimborazo NNP 37602 758 3 _ _ NNP 37602 758 4 : : : 37602 758 5 Pallatanga Pallatanga NNP 37602 758 6 , , , 37602 758 7 BMNH BMNH NNP 37602 758 8 1947.2.13.46 1947.2.13.46 CD 37602 758 9 ; ; : 37602 758 10 Santiago Santiago NNP 37602 758 11 , , , 37602 758 12 FMNH FMNH NNP 37602 758 13 42529 42529 CD 37602 758 14 . . . 37602 759 1 _ _ NNP 37602 759 2 Morona Morona NNP 37602 759 3 - - HYPH 37602 759 4 Santiago Santiago NNP 37602 759 5 _ _ NNP 37602 759 6 : : : 37602 759 7 " " `` 37602 759 8 Río Río NNP 37602 759 9 Santiago Santiago NNP 37602 759 10 " " '' 37602 759 11 (= (= NNP 37602 759 12 Río Río NNP 37602 759 13 Zamora Zamora NNP 37602 759 14 ) ) -RRB- 37602 759 15 , , , 37602 759 16 MIZS mizs NN 37602 759 17 2950 2950 CD 37602 759 18 . . . 37602 760 1 _ _ NNP 37602 760 2 Napo Napo NNP 37602 760 3 _ _ NNP 37602 760 4 : : : 37602 760 5 Lago Lago NNP 37602 760 6 Agrio Agrio NNP 37602 760 7 , , , 37602 760 8 KU KU NNP 37602 760 9 126646 126646 CD 37602 760 10 ; ; : 37602 760 11 Puerto Puerto NNP 37602 760 12 Libre Libre NNP 37602 760 13 , , , 37602 760 14 Río Río NNP 37602 760 15 Aguarico Aguarico NNP 37602 760 16 , , , 37602 760 17 KU KU NNP 37602 760 18 123172 123172 CD 37602 760 19 ; ; : 37602 760 20 Santa Santa NNP 37602 760 21 Cecilia Cecilia NNP 37602 760 22 , , , 37602 760 23 AUM AUM NNP 37602 760 24 8138 8138 CD 37602 760 25 , , , 37602 760 26 KU KU NNP 37602 760 27 105208 105208 CD 37602 760 28 - - SYM 37602 760 29 9 9 CD 37602 760 30 , , , 37602 760 31 109506 109506 CD 37602 760 32 , , , 37602 760 33 123171 123171 CD 37602 760 34 . . . 37602 761 1 _ _ NNP 37602 761 2 Pastaza Pastaza NNP 37602 761 3 _ _ NNP 37602 761 4 : : : 37602 761 5 Alpayacu Alpayacu NNP 37602 761 6 , , , 37602 761 7 BMNH BMNH NNP 37602 761 8 1912.11.1.64 1912.11.1.64 CD 37602 761 9 ; ; : 37602 761 10 Canelos Canelos NNP 37602 761 11 , , , 37602 761 12 BMNH BMNH NNP 37602 761 13 1947.2.13.40 1947.2.13.40 CD 37602 761 14 - - SYM 37602 761 15 1 1 CD 37602 761 16 , , , 37602 761 17 1947.2.13.43 1947.2.13.43 NNS 37602 761 18 - - SYM 37602 761 19 5 5 CD 37602 761 20 ; ; : 37602 761 21 Colonia Colonia NNP 37602 761 22 Mena Mena NNP 37602 761 23 , , , 37602 761 24 Río Río NNP 37602 761 25 Conambo Conambo NNP 37602 761 26 , , , 37602 761 27 ZSM ZSM NNP 37602 761 28 33/1962 33/1962 CD 37602 761 29 ; ; : 37602 761 30 Don Don NNP 37602 761 31 Tomás Tomás NNP 37602 761 32 , , , 37602 761 33 USNM USNM NNP 37602 761 34 166014 166014 CD 37602 761 35 ; ; : 37602 761 36 Guaché Guaché NNP 37602 761 37 , , , 37602 761 38 Río Río NNP 37602 761 39 Pastaza Pastaza NNP 37602 761 40 , , , 37602 761 41 AMNH AMNH NNP 37602 761 42 79986 79986 CD 37602 761 43 ; ; : 37602 761 44 Río Río NNP 37602 761 45 Bobonaza Bobonaza NNP 37602 761 46 , , , 37602 761 47 USNM USNM NNP 37602 761 48 166005 166005 CD 37602 761 49 ; ; : 37602 761 50 Río Río NNP 37602 761 51 Capahuari Capahuari NNP 37602 761 52 , , , 37602 761 53 USNM USNM NNP 37602 761 54 166554 166554 CD 37602 761 55 ; ; : 37602 761 56 Río Río NNP 37602 761 57 Conambo Conambo NNP 37602 761 58 at at IN 37602 761 59 Río Río NNP 37602 761 60 Shiona Shiona NNP 37602 761 61 - - HYPH 37602 761 62 yacu yacu NNP 37602 761 63 , , , 37602 761 64 USNM USNM NNP 37602 761 65 166018 166018 CD 37602 761 66 ; ; : 37602 761 67 Río Río NNP 37602 761 68 Copataza Copataza NNP 37602 761 69 , , , 37602 761 70 upper upper JJ 37602 761 71 Río Río NNP 37602 761 72 Pastaza Pastaza NNP 37602 761 73 , , , 37602 761 74 USNM USNM NNP 37602 761 75 166007 166007 CD 37602 761 76 - - SYM 37602 761 77 13 13 CD 37602 761 78 ; ; : 37602 761 79 Río Río NNP 37602 761 80 Pastaza Pastaza NNP 37602 761 81 , , , 37602 761 82 NHRM nhrm NN 37602 761 83 1946 1946 CD 37602 761 84 ; ; : 37602 761 85 Río Río NNP 37602 761 86 Pucyacu Pucyacu NNP 37602 761 87 , , , 37602 761 88 USNM USNM NNP 37602 761 89 165997 165997 CD 37602 761 90 ( ( -LRB- 37602 761 91 skeleton skeleton NNP 37602 761 92 ) ) -RRB- 37602 761 93 , , , 37602 761 94 165998 165998 CD 37602 761 95 - - SYM 37602 761 96 6001 6001 CD 37602 761 97 ; ; : 37602 761 98 Río Río NNP 37602 761 99 Rutuno Rutuno NNP 37602 761 100 , , , 37602 761 101 USNM USNM NNP 37602 761 102 166006 166006 CD 37602 761 103 ; ; : 37602 761 104 Río Río NNP 37602 761 105 Villano Villano NNP 37602 761 106 , , , 37602 761 107 USNM USNM NNP 37602 761 108 166002 166002 CD 37602 761 109 - - SYM 37602 761 110 4 4 CD 37602 761 111 ; ; : 37602 761 112 Sarayacu Sarayacu NNP 37602 761 113 , , , 37602 761 114 BMNH BMNH NNP 37602 761 115 1947.2.13.36 1947.2.13.36 NNS 37602 761 116 - - SYM 37602 761 117 9 9 CD 37602 761 118 , , , 37602 761 119 MCZ MCZ NNP 37602 761 120 26090 26090 CD 37602 761 121 , , , 37602 761 122 ZMB ZMB NNP 37602 761 123 10166 10166 CD 37602 761 124 . . . 37602 762 1 GUYANA guyana NN 37602 762 2 : : : 37602 762 3 _ _ NNP 37602 762 4 Mazaruni Mazaruni NNP 37602 762 5 - - HYPH 37602 762 6 Potaro Potaro NNP 37602 762 7 _ _ NNP 37602 762 8 : : : 37602 762 9 Kartabo Kartabo NNP 37602 762 10 , , , 37602 762 11 AMNH AMNH NNP 37602 762 12 70971 70971 CD 37602 762 13 ; ; : 37602 762 14 Membaru Membaru NNP 37602 762 15 River River NNP 37602 762 16 , , , 37602 762 17 upper upper JJ 37602 762 18 Mazaruni Mazaruni NNP 37602 762 19 River River NNP 37602 762 20 , , , 37602 762 21 UMMZ UMMZ NNP 37602 762 22 85168 85168 CD 37602 762 23 ; ; : 37602 762 24 Oko Oko NNP 37602 762 25 Mountains Mountains NNPS 37602 762 26 , , , 37602 762 27 FMNH FMNH NNP 37602 762 28 26722 26722 CD 37602 762 29 - - SYM 37602 762 30 3 3 CD 37602 762 31 . . . 37602 763 1 _ _ NNP 37602 763 2 North North NNP 37602 763 3 West West NNP 37602 763 4 _ _ NNP 37602 763 5 : : : 37602 763 6 Amakura Amakura NNP 37602 763 7 River River NNP 37602 763 8 , , , 37602 763 9 Haulover Haulover NNP 37602 763 10 , , , 37602 763 11 UMMZ UMMZ NNP 37602 763 12 83558 83558 CD 37602 763 13 - - SYM 37602 763 14 9 9 CD 37602 763 15 . . . 37602 764 1 _ _ NNP 37602 764 2 Rupununi Rupununi NNP 37602 764 3 _ _ NNP 37602 764 4 : : : 37602 764 5 Marudi Marudi NNP 37602 764 6 River River NNP 37602 764 7 , , , 37602 764 8 AMNH AMNH NNP 37602 764 9 46233 46233 CD 37602 764 10 ; ; : 37602 764 11 Shudi Shudi NNP 37602 764 12 - - HYPH 37602 764 13 kar kar NNP 37602 764 14 - - HYPH 37602 764 15 wau wau NNP 37602 764 16 , , , 37602 764 17 AMNH AMNH NNP 37602 764 18 49252 49252 CD 37602 764 19 . . . 37602 765 1 _ _ NNP 37602 765 2 West West NNP 37602 765 3 Demerara Demerara NNP 37602 765 4 _ _ NNP 37602 765 5 : : : 37602 765 6 Dunoon Dunoon NNP 37602 765 7 , , , 37602 765 8 UMMZ UMMZ NNP 37602 765 9 52449 52449 CD 37602 765 10 , , , 37602 765 11 52508 52508 CD 37602 765 12 . . . 37602 766 1 PERÚ PERÚ NNP 37602 766 2 : : : 37602 766 3 _ _ NNP 37602 766 4 Junín Junín NNP 37602 766 5 _ _ NNP 37602 766 6 : : : 37602 766 7 Chanchamayo Chanchamayo NNP 37602 766 8 , , , 37602 766 9 BMNH BMNH NNP 37602 766 10 1911.12.13.79 1911.12.13.79 CD 37602 766 11 - - SYM 37602 766 12 80 80 CD 37602 766 13 . . . 37602 767 1 _ _ NNP 37602 767 2 Loreto Loreto NNP 37602 767 3 _ _ NNP 37602 767 4 : : : 37602 767 5 Andoas Andoas NNP 37602 767 6 , , , 37602 767 7 AMNH AMNH NNP 37602 767 8 79984 79984 CD 37602 767 9 - - SYM 37602 767 10 5 5 CD 37602 767 11 ; ; : 37602 767 12 Cashiboya Cashiboya NNP 37602 767 13 , , , 37602 767 14 AMNH AMNH NNP 37602 767 15 43454 43454 CD 37602 767 16 ; ; : 37602 767 17 San San NNP 37602 767 18 Antonio Antonio NNP 37602 767 19 , , , 37602 767 20 Río Río NNP 37602 767 21 Itaya Itaya NNP 37602 767 22 , , , 37602 767 23 AMNH AMNH NNP 37602 767 24 43218 43218 CD 37602 767 25 . . . 37602 768 1 _ _ NNP 37602 768 2 Puno Puno NNP 37602 768 3 _ _ NNP 37602 768 4 : : : 37602 768 5 Yahuaramayo Yahuaramayo NNP 37602 768 6 , , , 37602 768 7 BMNH BMNH VBZ 37602 768 8 1913.2.25.7 1913.2.25.7 CD 37602 768 9 . . . 37602 769 1 SURINAM SURINAM NNP 37602 769 2 : : : 37602 769 3 _ _ NNP 37602 769 4 Suriname Suriname NNP 37602 769 5 _ _ NNP 37602 769 6 : : : 37602 769 7 Powakka Powakka NNP 37602 769 8 , , , 37602 769 9 CM CM NNP 37602 769 10 44217 44217 CD 37602 769 11 . . . 37602 770 1 SOUTH south JJ 37602 770 2 AMERICA AMERICA NNP 37602 770 3 : : : 37602 770 4 No no DT 37602 770 5 specific specific JJ 37602 770 6 locality locality NN 37602 770 7 , , , 37602 770 8 NHMW NHMW NNP 37602 770 9 6208 6208 CD 37602 770 10 . . . 37602 771 1 _ _ NNP 37602 771 2 Osteocephalus Osteocephalus NNP 37602 771 3 leprieurii leprieurii VBZ 37602 771 4 _ _ NNP 37602 771 5 BRASIL BRASIL NNP 37602 771 6 : : : 37602 771 7 _ _ NNP 37602 771 8 Acre Acre NNP 37602 771 9 _ _ NNP 37602 771 10 : : : 37602 771 11 Tarauacá Tarauacá NNP 37602 771 12 , , , 37602 771 13 FMNH FMNH NNP 37602 771 14 83247 83247 CD 37602 771 15 . . . 37602 772 1 _ _ NNP 37602 772 2 Amazonas Amazonas NNP 37602 772 3 _ _ NNP 37602 772 4 : : : 37602 772 5 Rio Rio NNP 37602 772 6 Javarí Javarí NNP 37602 772 7 , , , 37602 772 8 Benjamin Benjamin NNP 37602 772 9 Constant Constant NNP 37602 772 10 , , , 37602 772 11 CAS CAS NNP 37602 772 12 - - HYPH 37602 772 13 SU SU NNP 37602 772 14 12620 12620 CD 37602 772 15 ; ; : 37602 772 16 Río Río NNP 37602 772 17 Uaupés Uaupés NNP 37602 772 18 , , , 37602 772 19 north north RB 37602 772 20 of of IN 37602 772 21 Rio Rio NNP 37602 772 22 Japú Japú NNP 37602 772 23 , , , 37602 772 24 NHMG NHMG NNP 37602 772 25 489 489 CD 37602 772 26 . . . 37602 773 1 COLOMBIA COLOMBIA NNP 37602 773 2 : : : 37602 773 3 _ _ NNP 37602 773 4 Amazonas Amazonas NNP 37602 773 5 _ _ NNP 37602 773 6 : : : 37602 773 7 Gino Gino NNP 37602 773 8 - - HYPH 37602 773 9 goje goje NNP 37602 773 10 , , , 37602 773 11 lower low JJR 37602 773 12 Río Río NNP 37602 773 13 Apoporis Apoporis NNP 37602 773 14 , , , 37602 773 15 MCZ MCZ NNP 37602 773 16 28038 28038 CD 37602 773 17 , , , 37602 773 18 28040 28040 CD 37602 773 19 - - SYM 37602 773 20 2 2 CD 37602 773 21 , , , 37602 773 22 28044 28044 CD 37602 773 23 , , , 37602 773 24 USNM USNM NNP 37602 773 25 152136 152136 CD 37602 773 26 - - SYM 37602 773 27 8 8 CD 37602 773 28 . . . 37602 774 1 ECUADOR ECUADOR NNP 37602 774 2 : : : 37602 774 3 No no DT 37602 774 4 specific specific JJ 37602 774 5 locality locality NN 37602 774 6 , , , 37602 774 7 WCAB WCAB NNP 37602 774 8 35452 35452 CD 37602 774 9 - - SYM 37602 774 10 3 3 CD 37602 774 11 ; ; : 37602 774 12 " " `` 37602 774 13 Napo Napo NNP 37602 774 14 - - HYPH 37602 774 15 Pastaza Pastaza NNP 37602 774 16 , , , 37602 774 17 " " '' 37602 774 18 USNM USNM NNP 37602 774 19 166571 166571 CD 37602 774 20 . . . 37602 775 1 _ _ NNP 37602 775 2 Napo Napo NNP 37602 775 3 _ _ NNP 37602 775 4 : : : 37602 775 5 Avila Avila NNP 37602 775 6 , , , 37602 775 7 UMMZ UMMZ NNP 37602 775 8 92093 92093 CD 37602 775 9 ; ; : 37602 775 10 south south JJ 37602 775 11 slope slope NN 37602 775 12 Cordillera Cordillera NNP 37602 775 13 del del NNP 37602 775 14 Dué Dué NNP 37602 775 15 , , , 37602 775 16 KU KU NNP 37602 775 17 123170 123170 CD 37602 775 18 ; ; : 37602 775 19 Lago Lago NNP 37602 775 20 Agrio Agrio NNP 37602 775 21 , , , 37602 775 22 KU KU NNP 37602 775 23 125961 125961 CD 37602 775 24 - - SYM 37602 775 25 2 2 CD 37602 775 26 ( ( -LRB- 37602 775 27 skeletons skeleton NNS 37602 775 28 ) ) -RRB- 37602 775 29 , , , 37602 775 30 126611 126611 CD 37602 775 31 - - SYM 37602 775 32 44 44 CD 37602 775 33 , , , 37602 775 34 UMMZ UMMZ NNP 37602 775 35 129326 129326 CD 37602 775 36 ( ( -LRB- 37602 775 37 2 2 CD 37602 775 38 ) ) -RRB- 37602 775 39 ; ; : 37602 775 40 Limón Limón NNP 37602 775 41 Cocha Cocha NNP 37602 775 42 , , , 37602 775 43 Río Río NNP 37602 775 44 Napo Napo NNP 37602 775 45 , , , 37602 775 46 KU KU NNP 37602 775 47 99210 99210 CD 37602 775 48 - - SYM 37602 775 49 6 6 CD 37602 775 50 , , , 37602 775 51 UIMNH UIMNH NNP 37602 775 52 63087 63087 CD 37602 775 53 - - SYM 37602 775 54 9 9 CD 37602 775 55 , , , 37602 775 56 63098 63098 CD 37602 775 57 , , , 37602 775 58 63106 63106 CD 37602 775 59 - - SYM 37602 775 60 9 9 CD 37602 775 61 , , , 37602 775 62 63118 63118 CD 37602 775 63 - - SYM 37602 775 64 9 9 CD 37602 775 65 , , , 37602 775 66 64802 64802 CD 37602 775 67 - - SYM 37602 775 68 4 4 CD 37602 775 69 , , , 37602 775 70 64858 64858 CD 37602 775 71 , , , 37602 775 72 87998 87998 CD 37602 775 73 - - SYM 37602 775 74 9 9 CD 37602 775 75 , , , 37602 775 76 88001 88001 CD 37602 775 77 - - SYM 37602 775 78 30 30 CD 37602 775 79 , , , 37602 775 80 88437 88437 CD 37602 775 81 - - SYM 37602 775 82 8 8 CD 37602 775 83 , , , 37602 775 84 88580 88580 CD 37602 775 85 , , , 37602 775 86 88604 88604 CD 37602 775 87 - - SYM 37602 775 88 5 5 CD 37602 775 89 , , , 37602 775 90 89852 89852 CD 37602 775 91 - - SYM 37602 775 92 97 97 CD 37602 775 93 , , , 37602 775 94 89999 89999 CD 37602 775 95 - - SYM 37602 775 96 90000 90000 CD 37602 775 97 ; ; : 37602 775 98 Loreto Loreto NNP 37602 775 99 , , , 37602 775 100 CAS CAS NNP 37602 775 101 - - HYPH 37602 775 102 SU SU NNP 37602 775 103 11439 11439 CD 37602 775 104 , , , 37602 775 105 WCAB wcab NN 37602 775 106 36526 36526 CD 37602 775 107 ; ; : 37602 775 108 Puerto Puerto NNP 37602 775 109 Libre Libre NNP 37602 775 110 , , , 37602 775 111 Río Río NNP 37602 775 112 Aguarico Aguarico NNP 37602 775 113 , , , 37602 775 114 KU KU NNP 37602 775 115 123190 123190 CD 37602 775 116 - - SYM 37602 775 117 1 1 CD 37602 775 118 ; ; : 37602 775 119 Puerto Puerto NNP 37602 775 120 Napo Napo NNP 37602 775 121 , , , 37602 775 122 UIMNH UIMNH NNP 37602 775 123 55818 55818 CD 37602 775 124 - - SYM 37602 775 125 20 20 CD 37602 775 126 ; ; : 37602 775 127 Río Río NNP 37602 775 128 Cotapino Cotapino NNP 37602 775 129 , , , 37602 775 130 UMMZ UMMZ NNP 37602 775 131 92094 92094 CD 37602 775 132 ; ; : 37602 775 133 Río Río NNP 37602 775 134 Napo Napo NNP 37602 775 135 , , , 37602 775 136 UMMZ UMMZ NNP 37602 775 137 92078 92078 CD 37602 775 138 ; ; : 37602 775 139 Santa Santa NNP 37602 775 140 Cecilia Cecilia NNP 37602 775 141 , , , 37602 775 142 AUM AUM NNP 37602 775 143 8099 8099 CD 37602 775 144 , , , 37602 775 145 8102 8102 CD 37602 775 146 , , , 37602 775 147 8113 8113 CD 37602 775 148 - - SYM 37602 775 149 5 5 CD 37602 775 150 , , , 37602 775 151 8127 8127 CD 37602 775 152 - - SYM 37602 775 153 9 9 CD 37602 775 154 , , , 37602 775 155 8131 8131 CD 37602 775 156 , , , 37602 775 157 8137 8137 CD 37602 775 158 , , , 37602 775 159 8139 8139 CD 37602 775 160 - - SYM 37602 775 161 46 46 CD 37602 775 162 , , , 37602 775 163 8148 8148 CD 37602 775 164 , , , 37602 775 165 KU KU NNP 37602 775 166 105210 105210 CD 37602 775 167 - - SYM 37602 775 168 20 20 CD 37602 775 169 , , , 37602 775 170 109509 109509 CD 37602 775 171 - - SYM 37602 775 172 11 11 CD 37602 775 173 , , , 37602 775 174 111971 111971 CD 37602 775 175 , , , 37602 775 176 122964 122964 CD 37602 775 177 - - SYM 37602 775 178 87 87 CD 37602 775 179 , , , 37602 775 180 123169 123169 CD 37602 775 181 , , , 37602 775 182 126645 126645 CD 37602 775 183 . . . 37602 776 1 _ _ NNP 37602 776 2 Pastaza Pastaza NNP 37602 776 3 _ _ NNP 37602 776 4 : : : 37602 776 5 Canelos Canelos NNP 37602 776 6 , , , 37602 776 7 BMNH BMNH NNP 37602 776 8 1947.2.13.42 1947.2.13.42 NN 37602 776 9 , , , 37602 776 10 KU KU NNP 37602 776 11 120915 120915 CD 37602 776 12 ; ; : 37602 776 13 Río Río NNP 37602 776 14 Alpayacu Alpayacu NNP 37602 776 15 , , , 37602 776 16 UMMZ UMMZ NNP 37602 776 17 92079 92079 CD 37602 776 18 ; ; : 37602 776 19 Río Río NNP 37602 776 20 Arajuno Arajuno NNP 37602 776 21 , , , 37602 776 22 USNM USNM NNP 37602 776 23 166560 166560 CD 37602 776 24 - - SYM 37602 776 25 2 2 CD 37602 776 26 , , , 37602 776 27 WCAB WCAB NNP 37602 776 28 40176 40176 CD 37602 776 29 ; ; : 37602 776 30 Río Río NNP 37602 776 31 Oglán Oglán NNP 37602 776 32 , , , 37602 776 33 USNM USNM NNP 37602 776 34 16655203 16655203 CD 37602 776 35 , , , 37602 776 36 166558 166558 CD 37602 776 37 ; ; : 37602 776 38 Río Río NNP 37602 776 39 Rutuno Rutuno NNP 37602 776 40 , , , 37602 776 41 USNM USNM NNP 37602 776 42 166559 166559 CD 37602 776 43 ; ; : 37602 776 44 Río Río NNP 37602 776 45 Shilcayacu Shilcayacu NNP 37602 776 46 , , , 37602 776 47 below below IN 37602 776 48 Puyo Puyo NNP 37602 776 49 , , , 37602 776 50 USNM USNM NNP 37602 776 51 166557 166557 CD 37602 776 52 ; ; : 37602 776 53 Río Río NNP 37602 776 54 Villano Villano NNP 37602 776 55 , , , 37602 776 56 USNM USNM NNP 37602 776 57 166551 166551 CD 37602 776 58 . . . 37602 777 1 FRENCH french JJ 37602 777 2 GUIANA guiana NN 37602 777 3 : : : 37602 777 4 No no DT 37602 777 5 specific specific JJ 37602 777 6 locality locality NN 37602 777 7 , , , 37602 777 8 MNHN MNHN NNP 37602 777 9 4629 4629 CD 37602 777 10 . . . 37602 778 1 _ _ NNP 37602 778 2 Inini Inini NNP 37602 778 3 _ _ NNP 37602 778 4 : : : 37602 778 5 Lunier Lunier NNP 37602 778 6 River River NNP 37602 778 7 , , , 37602 778 8 MNHN MNHN NNP 37602 778 9 98/217 98/217 CD 37602 778 10 . . . 37602 779 1 GUYANA guyana NN 37602 779 2 : : : 37602 779 3 _ _ NNP 37602 779 4 Mazaruni Mazaruni NNP 37602 779 5 - - HYPH 37602 779 6 Potaro Potaro NNP 37602 779 7 _ _ NNP 37602 779 8 : : : 37602 779 9 Kartabo Kartabo NNP 37602 779 10 , , , 37602 779 11 AMNH AMNH NNP 37602 779 12 70967 70967 CD 37602 779 13 - - HYPH 37602 779 14 8 8 CD 37602 779 15 , , , 37602 779 16 70972 70972 CD 37602 779 17 , , , 37602 779 18 70976 70976 CD 37602 779 19 . . . 37602 780 1 _ _ NNP 37602 780 2 Rupununi Rupununi NNP 37602 780 3 _ _ NNP 37602 780 4 : : : 37602 780 5 Shudi Shudi NNP 37602 780 6 - - HYPH 37602 780 7 kar kar NNP 37602 780 8 - - HYPH 37602 780 9 wau wau NNP 37602 780 10 , , , 37602 780 11 AMNH AMNH NNP 37602 780 12 49255 49255 CD 37602 780 13 . . . 37602 781 1 _ _ NNP 37602 781 2 West West NNP 37602 781 3 Demerara Demerara NNP 37602 781 4 _ _ NNP 37602 781 5 : : : 37602 781 6 Demerara Demerara NNP 37602 781 7 Falls Falls NNP 37602 781 8 , , , 37602 781 9 BMNH BMNH NNP 37602 781 10 72.10.16.23 72.10.16.23 CD 37602 781 11 , , , 37602 781 12 72.10.16.37 72.10.16.37 CD 37602 781 13 - - HYPH 37602 781 14 8 8 CD 37602 781 15 . . . 37602 782 1 PERÚ PERÚ NNP 37602 782 2 : : : 37602 782 3 _ _ NNP 37602 782 4 Loreto Loreto NNP 37602 782 5 _ _ NNP 37602 782 6 : : : 37602 782 7 Estirón Estirón NNP 37602 782 8 , , , 37602 782 9 Río Río NNP 37602 782 10 Ampiyacu Ampiyacu NNP 37602 782 11 , , , 37602 782 12 MZUSP MZUSP NNP 37602 782 13 31033 31033 CD 37602 782 14 - - SYM 37602 782 15 4 4 CD 37602 782 16 ; ; : 37602 782 17 Pebas Pebas NNP 37602 782 18 , , , 37602 782 19 CAS CAS NNP 37602 782 20 - - HYPH 37602 782 21 SU SU NNP 37602 782 22 3158 3158 CD 37602 782 23 , , , 37602 782 24 3160 3160 CD 37602 782 25 ; ; : 37602 782 26 Roaboya Roaboya NNP 37602 782 27 , , , 37602 782 28 AMNH AMNH NNP 37602 782 29 43064 43064 CD 37602 782 30 . . . 37602 783 1 SURINAM SURINAM NNP 37602 783 2 : : : 37602 783 3 No no DT 37602 783 4 specific specific JJ 37602 783 5 locality locality NN 37602 783 6 , , , 37602 783 7 MCZ MCZ NNP 37602 783 8 2036 2036 CD 37602 783 9 , , , 37602 783 10 RMNH RMNH NNP 37602 783 11 11468 11468 CD 37602 783 12 . . . 37602 784 1 _ _ NNP 37602 784 2 Marowijne Marowijne NNP 37602 784 3 _ _ NNP 37602 784 4 : : : 37602 784 5 Camp Camp NNP 37602 784 6 3 3 CD 37602 784 7 , , , 37602 784 8 RMNH RMNH NNP 37602 784 9 13045 13045 CD 37602 784 10 - - SYM 37602 784 11 6 6 CD 37602 784 12 ; ; : 37602 784 13 Wane Wane NNP 37602 784 14 Creek Creek NNP 37602 784 15 North North NNP 37602 784 16 , , , 37602 784 17 RMNH RMNH NNP 37602 784 18 11469 11469 CD 37602 784 19 - - SYM 37602 784 20 70 70 CD 37602 784 21 . . . 37602 785 1 _ _ NNP 37602 785 2 Saramacca Saramacca NNP 37602 785 3 _ _ NNP 37602 785 4 : : : 37602 785 5 Right Right NNP 37602 785 6 Coppename Coppename NNP 37602 785 7 River River NNP 37602 785 8 , , , 37602 785 9 RMNH RMNH NNP 37602 785 10 11467 11467 CD 37602 785 11 . . . 37602 786 1 _ _ NNP 37602 786 2 Osteocephalus osteocephalus JJ 37602 786 3 pearsoni pearsoni NN 37602 786 4 _ _ NNP 37602 786 5 BOLIVIA BOLIVIA NNS 37602 786 6 : : : 37602 786 7 _ _ NNP 37602 786 8 El El NNP 37602 786 9 Beni Beni NNP 37602 786 10 _ _ NNP 37602 786 11 : : : 37602 786 12 upper upper NNP 37602 786 13 Río Río NNP 37602 786 14 Beni Beni NNP 37602 786 15 , , , 37602 786 16 below below IN 37602 786 17 mouth mouth NN 37602 786 18 of of IN 37602 786 19 Río Río NNP 37602 786 20 Mapiri Mapiri NNP 37602 786 21 , , , 37602 786 22 MCZ MCZ NNP 37602 786 23 15565 15565 CD 37602 786 24 , , , 37602 786 25 UMMZ UMMZ NNP 37602 786 26 57548 57548 CD 37602 786 27 , , , 37602 786 28 67464 67464 CD 37602 786 29 - - SYM 37602 786 30 5 5 CD 37602 786 31 ; ; : 37602 786 32 Rurrenbaque Rurrenbaque NNP 37602 786 33 , , , 37602 786 34 UMMZ UMMZ NNP 37602 786 35 57533 57533 CD 37602 786 36 . . . 37602 787 1 PERÚ PERÚ NNP 37602 787 2 : : : 37602 787 3 _ _ NNP 37602 787 4 Pasco Pasco NNP 37602 787 5 _ _ NNP 37602 787 6 : : : 37602 787 7 Yaupi Yaupi NNP 37602 787 8 , , , 37602 787 9 KU KU NNP 37602 787 10 136312 136312 CD 37602 787 11 . . . 37602 788 1 _ _ NNP 37602 788 2 Osteocephalus osteocephalus JJ 37602 788 3 taurinus taurinus NN 37602 788 4 _ _ NNP 37602 788 5 BOLIVIA BOLIVIA NNS 37602 788 6 : : : 37602 788 7 _ _ NNP 37602 788 8 El El NNP 37602 788 9 Beni Beni NNP 37602 788 10 _ _ NNP 37602 788 11 : : : 37602 788 12 Ivón Ivón NNPS 37602 788 13 , , , 37602 788 14 BMNH BMNH NNP 37602 788 15 1967.2040 1967.2040 CD 37602 788 16 ; ; : 37602 788 17 Reyes Reyes NNP 37602 788 18 , , , 37602 788 19 UMMZ UMMZ NNP 37602 788 20 57532 57532 CD 37602 788 21 . . . 37602 789 1 _ _ NNP 37602 789 2 La La NNP 37602 789 3 Paz Paz NNP 37602 789 4 _ _ NNP 37602 789 5 : : : 37602 789 6 San San NNP 37602 789 7 Ernesto Ernesto NNP 37602 789 8 , , , 37602 789 9 Mapiri Mapiri NNP 37602 789 10 District District NNP 37602 789 11 , , , 37602 789 12 BMNH BMNH NNP 37602 789 13 1901.8.2.54 1901.8.2.54 CD 37602 789 14 . . . 37602 790 1 _ _ NNP 37602 790 2 Santa Santa NNP 37602 790 3 Cruz Cruz NNP 37602 790 4 _ _ NNP 37602 790 5 : : : 37602 790 6 Buenavista Buenavista NNP 37602 790 7 , , , 37602 790 8 AMNH AMNH NNP 37602 790 9 33951 33951 CD 37602 790 10 - - SYM 37602 790 11 2 2 CD 37602 790 12 , , , 37602 790 13 33958 33958 CD 37602 790 14 , , , 37602 790 15 BMNH BMNH NNP 37602 790 16 1927.8.1.19 1927.8.1.19 CD 37602 790 17 , , , 37602 790 18 1927.8.1.118 1927.8.1.118 CD 37602 790 19 , , , 37602 790 20 FMNH FMNH NNP 37602 790 21 27091 27091 CD 37602 790 22 , , , 37602 790 23 UMMZ UMMZ NNP 37602 790 24 63319 63319 CD 37602 790 25 - - SYM 37602 790 26 21 21 CD 37602 790 27 , , , 37602 790 28 63959(2 63959(2 CD 37602 790 29 ) ) -RRB- 37602 790 30 , , , 37602 790 31 63961(2 63961(2 CD 37602 790 32 ) ) -RRB- 37602 790 33 , , , 37602 790 34 66566(2 66566(2 CD 37602 790 35 ) ) -RRB- 37602 790 36 , , , 37602 790 37 66567 66567 CD 37602 790 38 , , , 37602 790 39 66568(2 66568(2 CD 37602 790 40 ) ) -RRB- 37602 790 41 , , , 37602 790 42 66569 66569 CD 37602 790 43 ( ( -LRB- 37602 790 44 2 2 CD 37602 790 45 ) ) -RRB- 37602 790 46 , , , 37602 790 47 66570 66570 CD 37602 790 48 , , , 37602 790 49 66571(2 66571(2 CD 37602 790 50 ) ) -RRB- 37602 790 51 , , , 37602 790 52 66575 66575 CD 37602 790 53 - - SYM 37602 790 54 6 6 CD 37602 790 55 , , , 37602 790 56 68196 68196 CD 37602 790 57 ; ; : 37602 790 58 Río Río NNP 37602 790 59 Mamore Mamore NNP 37602 790 60 , , , 37602 790 61 2 2 CD 37602 790 62 km km NN 37602 790 63 N N NNP 37602 790 64 Boca Boca NNP 37602 790 65 Chaparé Chaparé NNP 37602 790 66 , , , 37602 790 67 AMNH AMNH NNP 37602 790 68 79324 79324 CD 37602 790 69 ; ; : 37602 790 70 Sara Sara NNP 37602 790 71 , , , 37602 790 72 CM CM NNP 37602 790 73 3840 3840 CD 37602 790 74 - - SYM 37602 790 75 1 1 CD 37602 790 76 ; ; : 37602 790 77 Surutu Surutu NNP 37602 790 78 , , , 37602 790 79 CM CM NNP 37602 790 80 3814 3814 CD 37602 790 81 - - SYM 37602 790 82 5 5 CD 37602 790 83 . . . 37602 791 1 BRASIL BRASIL NNP 37602 791 2 : : : 37602 791 3 No no DT 37602 791 4 specific specific JJ 37602 791 5 locality locality NN 37602 791 6 : : : 37602 791 7 " " `` 37602 791 8 Interior Interior NNP 37602 791 9 , , , 37602 791 10 " " '' 37602 791 11 BMNH BMNH NNP 37602 791 12 74.7.16.8 74.7.16.8 CD 37602 791 13 - - SYM 37602 791 14 9 9 CD 37602 791 15 . . . 37602 792 1 _ _ NNP 37602 792 2 Acre Acre NNP 37602 792 3 _ _ NNP 37602 792 4 : : : 37602 792 5 Plácido Plácido NNP 37602 792 6 de de FW 37602 792 7 Castro Castro NNP 37602 792 8 , , , 37602 792 9 MZUSP MZUSP NNP 37602 792 10 6518 6518 CD 37602 792 11 ; ; : 37602 792 12 Tarauacá Tarauacá NNP 37602 792 13 , , , 37602 792 14 WCAB WCAB NNP 37602 792 15 2496 2496 CD 37602 792 16 . . . 37602 793 1 _ _ NNP 37602 793 2 Amazonas Amazonas NNP 37602 793 3 _ _ NNP 37602 793 4 : : : 37602 793 5 Cucuí Cucuí NNP 37602 793 6 , , , 37602 793 7 NHMW NHMW NNP 37602 793 8 16495 16495 CD 37602 793 9 ; ; : 37602 793 10 Manacapurú Manacapurú NNP 37602 793 11 , , , 37602 793 12 ZMB ZMB NNP 37602 793 13 28492 28492 CD 37602 793 14 , , , 37602 793 15 ZSM ZSM NNP 37602 793 16 278/1925 278/1925 CD 37602 793 17 ; ; : 37602 793 18 Manáus Manáus NNP 37602 793 19 , , , 37602 793 20 MCZ MCZ NNP 37602 793 21 56281 56281 CD 37602 793 22 , , , 37602 793 23 NHMW NHMW NNP 37602 793 24 16492 16492 CD 37602 793 25 ; ; : 37602 793 26 Maués Maués NNP 37602 793 27 , , , 37602 793 28 AMNH AMNH NNP 37602 793 29 69623 69623 CD 37602 793 30 , , , 37602 793 31 76177 76177 CD 37602 793 32 ; ; : 37602 793 33 Taracuá Taracuá NNP 37602 793 34 , , , 37602 793 35 NHMG NHMG NNP 37602 793 36 488 488 CD 37602 793 37 , , , 37602 793 38 WCAB WCAB NNP 37602 793 39 18463 18463 CD 37602 793 40 - - SYM 37602 793 41 4 4 CD 37602 793 42 . . . 37602 794 1 _ _ NNP 37602 794 2 Mato Mato NNP 37602 794 3 Grosso Grosso NNP 37602 794 4 _ _ NNP 37602 794 5 : : : 37602 794 6 Mabuca Mabuca NNP 37602 794 7 , , , 37602 794 8 MZUSP MZUSP NNP 37602 794 9 4272 4272 CD 37602 794 10 ; ; : 37602 794 11 Posto Posto NNP 37602 794 12 Coluene Coluene NNP 37602 794 13 , , , 37602 794 14 Rio Rio NNP 37602 794 15 Xingú Xingú NNP 37602 794 16 , , , 37602 794 17 WCAB WCAB NNP 37602 794 18 812 812 CD 37602 794 19 ; ; : 37602 794 20 " " `` 37602 794 21 Puerto Puerto NNP 37602 794 22 Cabello Cabello NNP 37602 794 23 , , , 37602 794 24 " " '' 37602 794 25 AMNH AMNH NNP 37602 794 26 3154 3154 CD 37602 794 27 ; ; : 37602 794 28 Tapirapé Tapirapé NNP 37602 794 29 , , , 37602 794 30 AMNH AMNH NNP 37602 794 31 73647 73647 CD 37602 794 32 - - SYM 37602 794 33 62 62 CD 37602 794 34 , , , 37602 794 35 CAS CAS NNP 37602 794 36 - - HYPH 37602 794 37 SU SU NNP 37602 794 38 12351 12351 CD 37602 794 39 - - SYM 37602 794 40 6 6 CD 37602 794 41 , , , 37602 794 42 MNHN MNHN NNP 37602 794 43 46/324 46/324 CD 37602 794 44 . . . 37602 795 1 _ _ NNP 37602 795 2 Pará Pará NNP 37602 795 3 _ _ NNP 37602 795 4 : : : 37602 795 5 No no DT 37602 795 6 specific specific JJ 37602 795 7 locality locality NN 37602 795 8 , , , 37602 795 9 MPEG MPEG NNP 37602 795 10 623 623 CD 37602 795 11 - - SYM 37602 795 12 6 6 CD 37602 795 13 ; ; : 37602 795 14 Belém Belém NNP 37602 795 15 , , , 37602 795 16 KU KU NNP 37602 795 17 129866 129866 CD 37602 795 18 ; ; : 37602 795 19 Cachimbo cachimbo NN 37602 795 20 , , , 37602 795 21 FMNH FMNH NNP 37602 795 22 175876 175876 CD 37602 795 23 , , , 37602 795 24 UIMNH UIMNH NNP 37602 795 25 42149 42149 CD 37602 795 26 , , , 37602 795 27 WCAB WCAB NNP 37602 795 28 813 813 CD 37602 795 29 ; ; : 37602 795 30 Cametá Cametá NNP 37602 795 31 , , , 37602 795 32 NHMW NHMW NNP 37602 795 33 15892 15892 CD 37602 795 34 ; ; : 37602 795 35 Gurupá Gurupá NNP 37602 795 36 , , , 37602 795 37 BMNH BMNH VBZ 37602 795 38 96.6.29.13 96.6.29.13 CD 37602 795 39 ; ; : 37602 795 40 Ilha Ilha NNP 37602 795 41 de de FW 37602 795 42 Marajó Marajó NNP 37602 795 43 , , , 37602 795 44 BMNH BMNH NNP 37602 795 45 1923.11.9.20 1923.11.9.20 CD 37602 795 46 - - SYM 37602 795 47 4 4 CD 37602 795 48 ; ; : 37602 795 49 Ilha Ilha NNP 37602 795 50 Mexicana Mexicana NNP 37602 795 51 , , , 37602 795 52 ZSM ZSM NNP 37602 795 53 111/1911 111/1911 CD 37602 795 54 , , , 37602 795 55 112/1912 112/1912 CD 37602 795 56 ; ; : 37602 795 57 " " `` 37602 795 58 Ponto Ponto NNP 37602 795 59 Dois Dois NNP 37602 795 60 Indios Indios NNPS 37602 795 61 , , , 37602 795 62 " " '' 37602 795 63 BMNH BMNH NNP 37602 795 64 1939.1.5.5 1939.1.5.5 CD 37602 795 65 ; ; : 37602 795 66 Santarém Santarém NNP 37602 795 67 , , , 37602 795 68 BMNH BMNH NNP 37602 795 69 75.10.22.1 75.10.22.1 NN 37602 795 70 - - SYM 37602 795 71 4 4 CD 37602 795 72 , , , 37602 795 73 MCZ MCZ NNP 37602 795 74 354 354 CD 37602 795 75 . . . 37602 796 1 _ _ NNP 37602 796 2 Rondonia Rondonia NNP 37602 796 3 _ _ NNP 37602 796 4 : : : 37602 796 5 Abuná Abuná NNP 37602 796 6 , , , 37602 796 7 CAS CAS NNP 37602 796 8 49773 49773 CD 37602 796 9 - - SYM 37602 796 10 4 4 CD 37602 796 11 , , , 37602 796 12 FMNH FMNH NNP 37602 796 13 64239 64239 CD 37602 796 14 ; ; : 37602 796 15 Forte Forte NNP 37602 796 16 Principe Principe NNP 37602 796 17 da da NNP 37602 796 18 Beira Beira NNP 37602 796 19 , , , 37602 796 20 WCAB WCAB NNP 37602 796 21 10230 10230 CD 37602 796 22 ; ; : 37602 796 23 Igarapé Igarapé NNP 37602 796 24 Marmelo Marmelo NNP 37602 796 25 , , , 37602 796 26 FMNH FMNH NNP 37602 796 27 140254 140254 CD 37602 796 28 , , , 37602 796 29 KU KU NNP 37602 796 30 84725 84725 CD 37602 796 31 ( ( -LRB- 37602 796 32 skeleton skeleton NNP 37602 796 33 ) ) -RRB- 37602 796 34 , , , 37602 796 35 92243 92243 CD 37602 796 36 - - SYM 37602 796 37 6 6 CD 37602 796 38 , , , 37602 796 39 92247 92247 CD 37602 796 40 - - SYM 37602 796 41 8 8 CD 37602 796 42 ( ( -LRB- 37602 796 43 skeletons skeleton NNS 37602 796 44 ) ) -RRB- 37602 796 45 , , , 37602 796 46 WCAB wcab NN 37602 796 47 9997 9997 CD 37602 796 48 , , , 37602 796 49 10001 10001 CD 37602 796 50 , , , 37602 796 51 10003 10003 CD 37602 796 52 - - SYM 37602 796 53 4 4 CD 37602 796 54 ; ; : 37602 796 55 Porto Porto NNP 37602 796 56 Velho Velho NNP 37602 796 57 , , , 37602 796 58 MZUSP MZUSP NNP 37602 796 59 16343 16343 CD 37602 796 60 . . . 37602 797 1 COLOMBIA COLOMBIA NNP 37602 797 2 : : : 37602 797 3 _ _ NNP 37602 797 4 Amazonas Amazonas NNP 37602 797 5 _ _ NNP 37602 797 6 : : : 37602 797 7 Gino Gino NNP 37602 797 8 - - HYPH 37602 797 9 goje goje NNP 37602 797 10 , , , 37602 797 11 lower low JJR 37602 797 12 Río Río NNP 37602 797 13 Apoporis Apoporis NNP 37602 797 14 , , , 37602 797 15 USNM USNM NNP 37602 797 16 152139 152139 CD 37602 797 17 ; ; : 37602 797 18 Leticia Leticia NNP 37602 797 19 , , , 37602 797 20 USNM USNM NNP 37602 797 21 152010 152010 CD 37602 797 22 - - SYM 37602 797 23 1 1 CD 37602 797 24 ; ; : 37602 797 25 Raudal Raudal NNP 37602 797 26 de de NNP 37602 797 27 la la NNP 37602 797 28 Playa Playa NNP 37602 797 29 , , , 37602 797 30 lower low JJR 37602 797 31 Río Río NNP 37602 797 32 Apoporis Apoporis NNP 37602 797 33 , , , 37602 797 34 MCZ MCZ NNP 37602 797 35 28050 28050 CD 37602 797 36 ; ; : 37602 797 37 Río Río NNP 37602 797 38 Apoporis Apoporis NNP 37602 797 39 , , , 37602 797 40 MCZ MCZ NNP 37602 797 41 28060 28060 CD 37602 797 42 . . . 37602 798 1 _ _ NNP 37602 798 2 Boyacá Boyacá NNP 37602 798 3 _ _ NNP 37602 798 4 : : : 37602 798 5 Sutatenza Sutatenza NNP 37602 798 6 , , , 37602 798 7 USNM USNM NNP 37602 798 8 152054 152054 CD 37602 798 9 - - SYM 37602 798 10 6 6 CD 37602 798 11 . . . 37602 799 1 _ _ NNP 37602 799 2 Cundinamarca Cundinamarca NNP 37602 799 3 _ _ NNP 37602 799 4 : : : 37602 799 5 Medina Medina NNP 37602 799 6 , , , 37602 799 7 MCZ MCZ NNP 37602 799 8 16269 16269 CD 37602 799 9 - - SYM 37602 799 10 71 71 CD 37602 799 11 , , , 37602 799 12 USNM USNM NNP 37602 799 13 152089 152089 CD 37602 799 14 - - SYM 37602 799 15 90 90 CD 37602 799 16 , , , 37602 799 17 152092 152092 CD 37602 799 18 - - SYM 37602 799 19 7 7 CD 37602 799 20 , , , 37602 799 21 152757 152757 CD 37602 799 22 . . . 37602 800 1 _ _ NNP 37602 800 2 Meta Meta NNP 37602 800 3 _ _ NNP 37602 800 4 : : : 37602 800 5 El El NNP 37602 800 6 Mico Mico NNP 37602 800 7 , , , 37602 800 8 Río Río NNP 37602 800 9 Guejar Guejar NNP 37602 800 10 , , , 37602 800 11 USNM USNM NNP 37602 800 12 152203 152203 CD 37602 800 13 ; ; : 37602 800 14 Río Río NNP 37602 800 15 Duda Duda NNP 37602 800 16 , , , 37602 800 17 Sierra Sierra NNP 37602 800 18 de de NNP 37602 800 19 Macarena Macarena NNP 37602 800 20 , , , 37602 800 21 AMNH AMNH NNP 37602 800 22 79914 79914 CD 37602 800 23 ; ; : 37602 800 24 Río Río NNP 37602 800 25 Guapaya Guapaya NNP 37602 800 26 , , , 37602 800 27 Sierra Sierra NNP 37602 800 28 de de NNP 37602 800 29 Macarena Macarena NNP 37602 800 30 , , , 37602 800 31 FMNH FMNH NNP 37602 800 32 81332 81332 CD 37602 800 33 ; ; : 37602 800 34 Río Río NNP 37602 800 35 Guaviari Guaviari NNP 37602 800 36 , , , 37602 800 37 Casa Casa NNP 37602 800 38 de de NNP 37602 800 39 Piedra Piedra NNP 37602 800 40 , , , 37602 800 41 UTA UTA NNP 37602 800 42 No no DT 37602 800 43 number number NN 37602 800 44 . . . 37602 801 1 _ _ NNP 37602 801 2 Putumayo Putumayo NNP 37602 801 3 _ _ NNP 37602 801 4 : : : 37602 801 5 Río Río NNP 37602 801 6 Mecaya Mecaya NNP 37602 801 7 , , , 37602 801 8 FMNH FMNH NNP 37602 801 9 69711 69711 CD 37602 801 10 - - SYM 37602 801 11 4 4 CD 37602 801 12 , , , 37602 801 13 69716 69716 CD 37602 801 14 . . . 37602 802 1 _ _ NNP 37602 802 2 Vaupés Vaupés NNP 37602 802 3 _ _ NNP 37602 802 4 : : : 37602 802 5 Gomogoje Gomogoje NNP 37602 802 6 , , , 37602 802 7 lower low JJR 37602 802 8 Río Río NNP 37602 802 9 Apoporis Apoporis NNP 37602 802 10 , , , 37602 802 11 MCZ MCZ NNP 37602 802 12 28048 28048 CD 37602 802 13 . . . 37602 803 1 ECUADOR ECUADOR NNP 37602 803 2 : : : 37602 803 3 No no DT 37602 803 4 specific specific JJ 37602 803 5 locality locality NN 37602 803 6 , , , 37602 803 7 WCAB wcab NN 37602 803 8 35451 35451 CD 37602 803 9 , , , 37602 803 10 35785 35785 CD 37602 803 11 ; ; : 37602 803 12 " " `` 37602 803 13 Oriente Oriente NNP 37602 803 14 , , , 37602 803 15 " " '' 37602 803 16 UMMZ UMMZ NNP 37602 803 17 90418 90418 CD 37602 803 18 . . . 37602 804 1 _ _ NNP 37602 804 2 Carchí Carchí NNP 37602 804 3 _ _ NNP 37602 804 4 : : : 37602 804 5 below below IN 37602 804 6 Salinas Salinas NNP 37602 804 7 , , , 37602 804 8 USNM USNM NNP 37602 804 9 166059 166059 CD 37602 804 10 . . . 37602 805 1 _ _ NNP 37602 805 2 Esmeraldas Esmeraldas NNP 37602 805 3 _ _ NNP 37602 805 4 : : : 37602 805 5 Carondelet Carondelet NNP 37602 805 6 , , , 37602 805 7 UIMNH UIMNH NNP 37602 805 8 53560 53560 CD 37602 805 9 - - SYM 37602 805 10 9 9 CD 37602 805 11 ; ; : 37602 805 12 Lagartera Lagartera NNP 37602 805 13 , , , 37602 805 14 Río Río NNP 37602 805 15 Caoni Caoni NNP 37602 805 16 , , , 37602 805 17 UIMNH UIMNH NNP 37602 805 18 53441 53441 CD 37602 805 19 , , , 37602 805 20 53458 53458 CD 37602 805 21 - - SYM 37602 805 22 79 79 CD 37602 805 23 . . . 37602 806 1 _ _ NNP 37602 806 2 Morona Morona NNP 37602 806 3 - - HYPH 37602 806 4 Santiago Santiago NNP 37602 806 5 _ _ NNP 37602 806 6 : : : 37602 806 7 Macuma Macuma NNP 37602 806 8 , , , 37602 806 9 UIMNH UIMNH NNP 37602 806 10 63142 63142 CD 37602 806 11 - - SYM 37602 806 12 3 3 CD 37602 806 13 , , , 37602 806 14 63145 63145 CD 37602 806 15 , , , 37602 806 16 63147 63147 CD 37602 806 17 , , , 37602 806 18 63151 63151 CD 37602 806 19 , , , 37602 806 20 63154 63154 CD 37602 806 21 , , , 37602 806 22 63157 63157 CD 37602 806 23 , , , 37602 806 24 USNM USNM NNP 37602 806 25 166060 166060 CD 37602 806 26 . . . 37602 807 1 _ _ NNP 37602 807 2 Napo Napo NNP 37602 807 3 _ _ NNP 37602 807 4 : : : 37602 807 5 Avila Avila NNP 37602 807 6 , , , 37602 807 7 UMMZ UMMZ NNP 37602 807 8 92077 92077 CD 37602 807 9 ; ; : 37602 807 10 Cuyabeno Cuyabeno NNP 37602 807 11 , , , 37602 807 12 UIMNH UIMNH NNP 37602 807 13 63158 63158 CD 37602 807 14 , , , 37602 807 15 90111 90111 CD 37602 807 16 ; ; : 37602 807 17 Lago Lago NNP 37602 807 18 Agrio Agrio NNP 37602 807 19 , , , 37602 807 20 KU KU NNP 37602 807 21 126647 126647 CD 37602 807 22 - - SYM 37602 807 23 55 55 CD 37602 807 24 ; ; : 37602 807 25 Limón Limón NNP 37602 807 26 Cocha Cocha NNP 37602 807 27 , , , 37602 807 28 Río Río NNP 37602 807 29 Napo Napo NNP 37602 807 30 , , , 37602 807 31 AUM AUM NNP 37602 807 32 8132 8132 CD 37602 807 33 - - SYM 37602 807 34 4 4 CD 37602 807 35 , , , 37602 807 36 KU KU NNP 37602 807 37 99207 99207 CD 37602 807 38 - - SYM 37602 807 39 8 8 CD 37602 807 40 , , , 37602 807 41 99421 99421 CD 37602 807 42 - - SYM 37602 807 43 3 3 CD 37602 807 44 , , , 37602 807 45 99424 99424 CD 37602 807 46 ( ( -LRB- 37602 807 47 skeleton skeleton NNP 37602 807 48 ) ) -RRB- 37602 807 49 , , , 37602 807 50 99425 99425 CD 37602 807 51 , , , 37602 807 52 UIMNH UIMNH NNP 37602 807 53 64801 64801 CD 37602 807 54 , , , 37602 807 55 87798 87798 CD 37602 807 56 , , , 37602 807 57 87800 87800 CD 37602 807 58 , , , 37602 807 59 88032 88032 CD 37602 807 60 - - SYM 37602 807 61 5 5 CD 37602 807 62 , , , 37602 807 63 88576 88576 CD 37602 807 64 , , , 37602 807 65 90066 90066 CD 37602 807 66 , , , 37602 807 67 90082 90082 CD 37602 807 68 , , , 37602 807 69 90102 90102 CD 37602 807 70 , , , 37602 807 71 90104 90104 CD 37602 807 72 , , , 37602 807 73 90314 90314 CD 37602 807 74 , , , 37602 807 75 90984 90984 CD 37602 807 76 ; ; : 37602 807 77 Loreto Loreto NNP 37602 807 78 , , , 37602 807 79 WCAB WCAB NNP 37602 807 80 35352 35352 CD 37602 807 81 ; ; : 37602 807 82 Río Río NNP 37602 807 83 Cotapino Cotapino NNP 37602 807 84 , , , 37602 807 85 UMMZ UMMZ NNP 37602 807 86 92080 92080 CD 37602 807 87 ; ; : 37602 807 88 Río Río NNP 37602 807 89 Napo Napo NNP 37602 807 90 , , , 37602 807 91 UMMZ UMMZ NNP 37602 807 92 84120 84120 CD 37602 807 93 ; ; : 37602 807 94 San San NNP 37602 807 95 José José NNP 37602 807 96 Abajo Abajo NNP 37602 807 97 , , , 37602 807 98 AMNH AMNH NNP 37602 807 99 1295 1295 CD 37602 807 100 , , , 37602 807 101 1449 1449 CD 37602 807 102 , , , 37602 807 103 22180 22180 CD 37602 807 104 , , , 37602 807 105 79990 79990 CD 37602 807 106 ; ; : 37602 807 107 Santa Santa NNP 37602 807 108 Cecilia Cecilia NNP 37602 807 109 , , , 37602 807 110 AUM AUM NNP 37602 807 111 8117 8117 CD 37602 807 112 , , , 37602 807 113 8150 8150 CD 37602 807 114 , , , 37602 807 115 KU KU NNP 37602 807 116 105230 105230 CD 37602 807 117 - - SYM 37602 807 118 3 3 CD 37602 807 119 ; ; : 37602 807 120 south south JJ 37602 807 121 slope slope NN 37602 807 122 Volcán Volcán NNP 37602 807 123 Sumaco Sumaco NNP 37602 807 124 , , , 37602 807 125 USNM USNM NNP 37602 807 126 166570 166570 CD 37602 807 127 . . . 37602 808 1 _ _ NNP 37602 808 2 Pastaza Pastaza NNP 37602 808 3 _ _ NNP 37602 808 4 : : : 37602 808 5 No no DT 37602 808 6 specific specific JJ 37602 808 7 locality locality NN 37602 808 8 , , , 37602 808 9 ZSM ZSM NNP 37602 808 10 31/1956 31/1956 LS 37602 808 11 ; ; : 37602 808 12 Arajuno Arajuno NNP 37602 808 13 , , , 37602 808 14 USNM USNM NNP 37602 808 15 165995 165995 CD 37602 808 16 ; ; : 37602 808 17 Bufeo Bufeo NNP 37602 808 18 , , , 37602 808 19 lower low JJR 37602 808 20 Río Río NNP 37602 808 21 Bobonaza Bobonaza NNP 37602 808 22 , , , 37602 808 23 USNM USNM NNP 37602 808 24 166046 166046 CD 37602 808 25 - - SYM 37602 808 26 8 8 CD 37602 808 27 ; ; : 37602 808 28 Canelos Canelos NNP 37602 808 29 , , , 37602 808 30 BMNH BMNH NNP 37602 808 31 80.12.5.179 80.12.5.179 CD 37602 808 32 , , , 37602 808 33 1947.2.13.48 1947.2.13.48 CD 37602 808 34 , , , 37602 808 35 UMMZ UMMZ NNP 37602 808 36 89066 89066 CD 37602 808 37 ; ; : 37602 808 38 Don Don NNP 37602 808 39 Tomás Tomás NNP 37602 808 40 , , , 37602 808 41 Río Río NNP 37602 808 42 Bobonaza Bobonaza NNP 37602 808 43 , , , 37602 808 44 USNM USNM NNP 37602 808 45 166049 166049 CD 37602 808 46 - - SYM 37602 808 47 50 50 CD 37602 808 48 ; ; : 37602 808 49 Montalvo Montalvo NNP 37602 808 50 , , , 37602 808 51 CAS CAS NNP 37602 808 52 - - HYPH 37602 808 53 SU SU NNP 37602 808 54 10320 10320 CD 37602 808 55 , , , 37602 808 56 USNM USNM NNP 37602 808 57 165987 165987 CD 37602 808 58 - - SYM 37602 808 59 9 9 CD 37602 808 60 , , , 37602 808 61 166058 166058 CD 37602 808 62 , , , 37602 808 63 166566 166566 CD 37602 808 64 ; ; : 37602 808 65 2.5 2.5 CD 37602 808 66 km km NN 37602 808 67 SE SE NNP 37602 808 68 Puyo Puyo NNP 37602 808 69 , , , 37602 808 70 USNM USNM NNP 37602 808 71 166051 166051 CD 37602 808 72 ; ; : 37602 808 73 Río Río NNP 37602 808 74 Arajuno Arajuno NNP 37602 808 75 , , , 37602 808 76 USNM USNM NNP 37602 808 77 166043 166043 CD 37602 808 78 - - SYM 37602 808 79 5 5 CD 37602 808 80 ; ; : 37602 808 81 Río Río NNP 37602 808 82 Arajuno Arajuno NNP 37602 808 83 ( ( -LRB- 37602 808 84 headwaters headwater NNS 37602 808 85 ) ) -RRB- 37602 808 86 , , , 37602 808 87 USNM USNM NNP 37602 808 88 166053 166053 CD 37602 808 89 ; ; : 37602 808 90 Río Río NNP 37602 808 91 Bobonaza Bobonaza NNP 37602 808 92 , , , 37602 808 93 WCAB WCAB NNP 37602 808 94 3613 3613 CD 37602 808 95 - - SYM 37602 808 96 4 4 CD 37602 808 97 , , , 37602 808 98 35504 35504 CD 37602 808 99 ; ; : 37602 808 100 Río Río NNP 37602 808 101 Capahuari Capahuari NNP 37602 808 102 , , , 37602 808 103 USNM USNM NNP 37602 808 104 165990 165990 CD 37602 808 105 , , , 37602 808 106 166555 166555 CD 37602 808 107 - - SYM 37602 808 108 6 6 CD 37602 808 109 ; ; : 37602 808 110 Río Río NNP 37602 808 111 Capahuari Capahuari NNP 37602 808 112 ( ( -LRB- 37602 808 113 headwaters headwater NNS 37602 808 114 ) ) -RRB- 37602 808 115 , , , 37602 808 116 USNM USNM NNP 37602 808 117 166057 166057 CD 37602 808 118 ; ; : 37602 808 119 Río Río NNP 37602 808 120 Conambo Conambo NNP 37602 808 121 , , , 37602 808 122 USNM USNM NNP 37602 808 123 166569 166569 CD 37602 808 124 , , , 37602 808 125 ZSM ZSM NNP 37602 808 126 28/1962 28/1962 CD 37602 808 127 , , , 37602 808 128 35/1962 35/1962 CD 37602 808 129 ; ; : 37602 808 130 Río Río NNP 37602 808 131 Conambo Conambo NNP 37602 808 132 at at IN 37602 808 133 Río Río NNP 37602 808 134 Ollaguanga Ollaguanga NNP 37602 808 135 , , , 37602 808 136 USNM USNM NNP 37602 808 137 166568 166568 CD 37602 808 138 ; ; : 37602 808 139 Río Río NNP 37602 808 140 Conambo Conambo NNP 37602 808 141 at at IN 37602 808 142 Río Río NNP 37602 808 143 Shiona Shiona NNP 37602 808 144 - - HYPH 37602 808 145 yacu yacu NNP 37602 808 146 , , , 37602 808 147 USNM USNM NNP 37602 808 148 166019 166019 CD 37602 808 149 , , , 37602 808 150 166563 166563 CD 37602 808 151 - - SYM 37602 808 152 5 5 CD 37602 808 153 ; ; : 37602 808 154 Río Río NNP 37602 808 155 Corrientes Corrientes NNP 37602 808 156 , , , 37602 808 157 USNM USNM NNP 37602 808 158 195994 195994 CD 37602 808 159 , , , 37602 808 160 166020 166020 CD 37602 808 161 - - SYM 37602 808 162 38 38 CD 37602 808 163 , , , 37602 808 164 WCAB WCAB NNP 37602 808 165 3841 3841 CD 37602 808 166 - - SYM 37602 808 167 2 2 CD 37602 808 168 ; ; : 37602 808 169 Río Río NNP 37602 808 170 Huiyo Huiyo NNP 37602 808 171 - - HYPH 37602 808 172 yacu yacu NNP 37602 808 173 , , , 37602 808 174 Pico Pico NNP 37602 808 175 de de NNP 37602 808 176 Conambo Conambo NNP 37602 808 177 , , , 37602 808 178 USNM USNM NNP 37602 808 179 166052 166052 CD 37602 808 180 ; ; : 37602 808 181 Río Río NNP 37602 808 182 Pastaza Pastaza NNP 37602 808 183 , , , 37602 808 184 MCZ MCZ NNP 37602 808 185 19697 19697 CD 37602 808 186 ; ; : 37602 808 187 Río Río NNP 37602 808 188 Pastaza Pastaza NNP 37602 808 189 ( ( -LRB- 37602 808 190 drainage drainage NN 37602 808 191 ) ) -RRB- 37602 808 192 , , , 37602 808 193 NHRM nhrm NN 37602 808 194 1966 1966 CD 37602 808 195 , , , 37602 808 196 USNM USNM NNP 37602 808 197 165996 165996 CD 37602 808 198 ; ; : 37602 808 199 Río Río NNP 37602 808 200 Pindo Pindo NNP 37602 808 201 , , , 37602 808 202 USNM USNM NNP 37602 808 203 166039 166039 CD 37602 808 204 - - SYM 37602 808 205 41 41 CD 37602 808 206 ; ; : 37602 808 207 Río Río NNP 37602 808 208 Pindo Pindo NNP 37602 808 209 at at IN 37602 808 210 Río Río NNP 37602 808 211 Tigre Tigre NNP 37602 808 212 ( ( -LRB- 37602 808 213 village village NN 37602 808 214 ) ) -RRB- 37602 808 215 , , , 37602 808 216 USNM USNM NNP 37602 808 217 165992 165992 CD 37602 808 218 - - SYM 37602 808 219 3 3 CD 37602 808 220 , , , 37602 808 221 166042 166042 CD 37602 808 222 ; ; : 37602 808 223 Río Río NNP 37602 808 224 Pucayacu Pucayacu NNP 37602 808 225 , , , 37602 808 226 USNM USNM NNP 37602 808 227 166054 166054 CD 37602 808 228 , , , 37602 808 229 166056 166056 CD 37602 808 230 ; ; : 37602 808 231 Río Río NNP 37602 808 232 Rutuno Rutuno NNP 37602 808 233 , , , 37602 808 234 USNM USNM NNP 37602 808 235 166055 166055 CD 37602 808 236 ; ; : 37602 808 237 Río Río NNP 37602 808 238 Solis Solis NNP 37602 808 239 , , , 37602 808 240 upper upper JJ 37602 808 241 Río Río NNP 37602 808 242 Bobonaza Bobonaza NNP 37602 808 243 , , , 37602 808 244 WCAB WCAB NNP 37602 808 245 39914 39914 CD 37602 808 246 ; ; : 37602 808 247 Río Río NNP 37602 808 248 Villano Villano NNP 37602 808 249 , , , 37602 808 250 USNM USNM NNP 37602 808 251 165991 165991 CD 37602 808 252 , , , 37602 808 253 166567 166567 CD 37602 808 254 ; ; : 37602 808 255 Sarayacu Sarayacu NNP 37602 808 256 , , , 37602 808 257 BMNH BMNH NNP 37602 808 258 80.12.5.213 80.12.5.213 CD 37602 808 259 , , , 37602 808 260 80.12.5.239 80.12.5.239 NNP 37602 808 261 - - SYM 37602 808 262 40 40 CD 37602 808 263 , , , 37602 808 264 MZUSP MZUSP NNP 37602 808 265 323 323 CD 37602 808 266 ; ; : 37602 808 267 Shell Shell NNP 37602 808 268 Mera Mera NNP 37602 808 269 , , , 37602 808 270 KU KU NNP 37602 808 271 99420 99420 CD 37602 808 272 . . . 37602 809 1 _ _ NNP 37602 809 2 Zamora Zamora NNP 37602 809 3 - - HYPH 37602 809 4 Chinchipe Chinchipe NNP 37602 809 5 _ _ NNP 37602 809 6 : : : 37602 809 7 " " `` 37602 809 8 Yani Yani NNP 37602 809 9 - - HYPH 37602 809 10 Inzari Inzari NNP 37602 809 11 , , , 37602 809 12 " " '' 37602 809 13 AMNH AMNH NNP 37602 809 14 43259 43259 CD 37602 809 15 , , , 37602 809 16 43394 43394 CD 37602 809 17 ; ; : 37602 809 18 Zamora Zamora NNP 37602 809 19 , , , 37602 809 20 AMNH AMNH NNP 37602 809 21 78928 78928 CD 37602 809 22 . . . 37602 810 1 FRENCH french JJ 37602 810 2 GUIANA guiana NN 37602 810 3 : : : 37602 810 4 _ _ NNP 37602 810 5 Cayenne Cayenne NNP 37602 810 6 _ _ NNP 37602 810 7 : : : 37602 810 8 Crique Crique NNP 37602 810 9 Grégoire Grégoire NNP 37602 810 10 , , , 37602 810 11 UP up RB 37602 810 12 40 40 CD 37602 810 13 ; ; : 37602 810 14 Maripa Maripa NNP 37602 810 15 , , , 37602 810 16 Oyapok Oyapok NNP 37602 810 17 River River NNP 37602 810 18 , , , 37602 810 19 UP up RB 37602 810 20 72 72 CD 37602 810 21 ; ; : 37602 810 22 Oyapok Oyapok NNP 37602 810 23 River River NNP 37602 810 24 , , , 37602 810 25 UZM UZM NNP 37602 810 26 1473 1473 CD 37602 810 27 . . . 37602 811 1 _ _ NNP 37602 811 2 Inini Inini NNP 37602 811 3 _ _ NNP 37602 811 4 : : : 37602 811 5 Crique Crique NNP 37602 811 6 Gabrielle Gabrielle NNP 37602 811 7 , , , 37602 811 8 UP up RB 37602 811 9 118 118 CD 37602 811 10 - - SYM 37602 811 11 20 20 CD 37602 811 12 . . . 37602 812 1 GUYANA guyana NN 37602 812 2 : : : 37602 812 3 No no DT 37602 812 4 specific specific JJ 37602 812 5 locality locality NN 37602 812 6 : : : 37602 812 7 RMNH RMNH NNP 37602 812 8 1873(3 1873(3 CD 37602 812 9 ) ) -RRB- 37602 812 10 , , , 37602 812 11 ZMB ZMB NNP 37602 812 12 3102(2 3102(2 CD 37602 812 13 ) ) -RRB- 37602 812 14 . . . 37602 813 1 _ _ NNP 37602 813 2 East East NNP 37602 813 3 Demerara Demerara NNP 37602 813 4 _ _ NNP 37602 813 5 : : : 37602 813 6 Atkinson Atkinson NNP 37602 813 7 Field Field NNP 37602 813 8 , , , 37602 813 9 ASU ASU NNP 37602 813 10 11622 11622 CD 37602 813 11 . . . 37602 814 1 _ _ NNP 37602 814 2 Mazaruni Mazaruni NNP 37602 814 3 - - HYPH 37602 814 4 Potaro Potaro NNP 37602 814 5 _ _ NNP 37602 814 6 : : : 37602 814 7 Chinapora Chinapora NNP 37602 814 8 River River NNP 37602 814 9 , , , 37602 814 10 upper upper JJ 37602 814 11 Potaro Potaro NNP 37602 814 12 River River NNP 37602 814 13 , , , 37602 814 14 BMNH BMNH VBZ 37602 814 15 1905.11.1.20 1905.11.1.20 CD 37602 814 16 - - SYM 37602 814 17 1 1 CD 37602 814 18 ; ; : 37602 814 19 Kamakusa Kamakusa NNP 37602 814 20 , , , 37602 814 21 AMNH AMNH NNP 37602 814 22 21416 21416 CD 37602 814 23 , , , 37602 814 24 21418 21418 CD 37602 814 25 - - SYM 37602 814 26 9 9 CD 37602 814 27 , , , 37602 814 28 21422 21422 CD 37602 814 29 ; ; : 37602 814 30 Kartabo Kartabo NNP 37602 814 31 , , , 37602 814 32 AMNH AMNH NNP 37602 814 33 11689 11689 CD 37602 814 34 , , , 37602 814 35 11691 11691 CD 37602 814 36 , , , 37602 814 37 11697 11697 CD 37602 814 38 - - SYM 37602 814 39 9 9 CD 37602 814 40 , , , 37602 814 41 11703 11703 CD 37602 814 42 , , , 37602 814 43 11706 11706 CD 37602 814 44 - - SYM 37602 814 45 8 8 CD 37602 814 46 , , , 37602 814 47 23107 23107 CD 37602 814 48 , , , 37602 814 49 39730 39730 CD 37602 814 50 , , , 37602 814 51 70966 70966 CD 37602 814 52 , , , 37602 814 53 70969 70969 CD 37602 814 54 - - SYM 37602 814 55 70 70 CD 37602 814 56 , , , 37602 814 57 70973 70973 CD 37602 814 58 - - SYM 37602 814 59 5 5 CD 37602 814 60 , , , 37602 814 61 USNM USNM NNP 37602 814 62 118057 118057 CD 37602 814 63 ; ; : 37602 814 64 Moraballi Moraballi NNP 37602 814 65 Creek Creek NNP 37602 814 66 , , , 37602 814 67 Essequibo Essequibo NNP 37602 814 68 River River NNP 37602 814 69 , , , 37602 814 70 BMNH BMNH NNP 37602 814 71 1930.10.10.47 1930.10.10.47 CD 37602 814 72 - - HYPH 37602 814 73 51 51 CD 37602 814 74 ; ; : 37602 814 75 Oko Oko NNP 37602 814 76 Mountains Mountains NNPS 37602 814 77 , , , 37602 814 78 FMNH FMNH NNP 37602 814 79 26692 26692 CD 37602 814 80 - - SYM 37602 814 81 705 705 CD 37602 814 82 ; ; : 37602 814 83 upper upper JJ 37602 814 84 Potaro Potaro NNP 37602 814 85 River River NNP 37602 814 86 , , , 37602 814 87 Tung Tung NNP 37602 814 88 District District NNP 37602 814 89 , , , 37602 814 90 BMNH BMNH VBZ 37602 814 91 1905.11.1.40 1905.11.1.40 CD 37602 814 92 ; ; : 37602 814 93 Rockstone Rockstone NNP 37602 814 94 , , , 37602 814 95 FMNH FMNH NNP 37602 814 96 26591 26591 CD 37602 814 97 . . . 37602 815 1 _ _ NNP 37602 815 2 North North NNP 37602 815 3 West West NNP 37602 815 4 _ _ NNP 37602 815 5 : : : 37602 815 6 Amakura Amakura NNP 37602 815 7 River River NNP 37602 815 8 , , , 37602 815 9 Haulover Haulover NNP 37602 815 10 , , , 37602 815 11 UMMZ UMMZ NNP 37602 815 12 83735 83735 CD 37602 815 13 . . . 37602 816 1 _ _ NNP 37602 816 2 Rupununi Rupununi NNP 37602 816 3 _ _ NNP 37602 816 4 : : : 37602 816 5 north north NN 37602 816 6 of of IN 37602 816 7 Acaray Acaray NNP 37602 816 8 River River NNP 37602 816 9 , , , 37602 816 10 west west NN 37602 816 11 of of IN 37602 816 12 New New NNP 37602 816 13 River River NNP 37602 816 14 , , , 37602 816 15 KU KU NNP 37602 816 16 69747 69747 CD 37602 816 17 - - SYM 37602 816 18 8 8 CD 37602 816 19 ; ; : 37602 816 20 Kuyuwini Kuyuwini NNP 37602 816 21 Landing Landing NNP 37602 816 22 , , , 37602 816 23 AMNH AMNH NNP 37602 816 24 46283 46283 CD 37602 816 25 ; ; : 37602 816 26 Pakaraima Pakaraima NNP 37602 816 27 Mountains Mountains NNPS 37602 816 28 , , , 37602 816 29 BMNH BMNH NNS 37602 816 30 1933.6.19.49 1933.6.19.49 CD 37602 816 31 ; ; : 37602 816 32 Shudi Shudi NNP 37602 816 33 - - HYPH 37602 816 34 kar kar NNP 37602 816 35 - - HYPH 37602 816 36 wau wau NNP 37602 816 37 , , , 37602 816 38 AMNH AMNH NNP 37602 816 39 10665 10665 CD 37602 816 40 , , , 37602 816 41 39637 39637 CD 37602 816 42 , , , 37602 816 43 49256(2 49256(2 CD 37602 816 44 ) ) -RRB- 37602 816 45 . . . 37602 817 1 _ _ NNP 37602 817 2 West West NNP 37602 817 3 Demerara Demerara NNP 37602 817 4 _ _ NNP 37602 817 5 : : : 37602 817 6 Demerara Demerara NNP 37602 817 7 , , , 37602 817 8 CAS CAS NNP 37602 817 9 54773 54773 CD 37602 817 10 - - SYM 37602 817 11 4 4 CD 37602 817 12 ; ; : 37602 817 13 Demerara Demerara NNP 37602 817 14 Falls Falls NNP 37602 817 15 , , , 37602 817 16 BMNH BMNH VBZ 37602 817 17 72.10.16.16 72.10.16.16 CD 37602 817 18 - - SYM 37602 817 19 22 22 CD 37602 817 20 , , , 37602 817 21 72.10.16.25 72.10.16.25 CD 37602 817 22 - - SYM 37602 817 23 32 32 CD 37602 817 24 ; ; : 37602 817 25 Dunoon Dunoon NNP 37602 817 26 , , , 37602 817 27 MCZ MCZ NNP 37602 817 28 4834 4834 CD 37602 817 29 , , , 37602 817 30 UMMZ UMMZ NNP 37602 817 31 46736 46736 CD 37602 817 32 , , , 37602 817 33 52493 52493 CD 37602 817 34 - - SYM 37602 817 35 4 4 CD 37602 817 36 , , , 37602 817 37 52502 52502 CD 37602 817 38 , , , 37602 817 39 52504 52504 CD 37602 817 40 - - SYM 37602 817 41 5 5 CD 37602 817 42 , , , 37602 817 43 57271 57271 CD 37602 817 44 ; ; : 37602 817 45 Vryheid Vryheid NNP 37602 817 46 , , , 37602 817 47 BMNH BMNH NNP 37602 817 48 78.12.13.18 78.12.13.18 CD 37602 817 49 . . . 37602 818 1 PERÚ PERÚ NNP 37602 818 2 : : : 37602 818 3 _ _ NNP 37602 818 4 Amazonas Amazonas NNP 37602 818 5 _ _ NNP 37602 818 6 : : : 37602 818 7 Río Río NNP 37602 818 8 Cenepa Cenepa NNP 37602 818 9 , , , 37602 818 10 AMNH AMNH NNP 37602 818 11 43400 43400 CD 37602 818 12 . . . 37602 819 1 _ _ NNP 37602 819 2 Huanuco Huanuco NNP 37602 819 3 _ _ NNP 37602 819 4 : : : 37602 819 5 Monte Monte NNP 37602 819 6 Alegre Alegre NNP 37602 819 7 , , , 37602 819 8 Río Río NNP 37602 819 9 Pachitea Pachitea NNP 37602 819 10 , , , 37602 819 11 AMNH AMNH NNP 37602 819 12 43014 43014 CD 37602 819 13 , , , 37602 819 14 43019 43019 CD 37602 819 15 . . . 37602 820 1 _ _ NNP 37602 820 2 Loreto Loreto NNP 37602 820 3 _ _ NNP 37602 820 4 : : : 37602 820 5 Achinamisa Achinamisa NNP 37602 820 6 , , , 37602 820 7 Río Río NNP 37602 820 8 Huallaga Huallaga NNP 37602 820 9 , , , 37602 820 10 AMNH AMNH NNP 37602 820 11 42178 42178 CD 37602 820 12 , , , 37602 820 13 42502 42502 CD 37602 820 14 ; ; : 37602 820 15 Andoas Andoas NNP 37602 820 16 , , , 37602 820 17 Río Río NNP 37602 820 18 Pastaza Pastaza NNP 37602 820 19 , , , 37602 820 20 AMNH AMNH NNP 37602 820 21 79991 79991 CD 37602 820 22 ; ; : 37602 820 23 Cashiboya Cashiboya NNP 37602 820 24 , , , 37602 820 25 AMNH AMNH NNP 37602 820 26 43388 43388 CD 37602 820 27 , , , 37602 820 28 43453 43453 CD 37602 820 29 ; ; : 37602 820 30 Estirón Estirón NNP 37602 820 31 , , , 37602 820 32 Río Río NNP 37602 820 33 Ampiyacu Ampiyacu NNP 37602 820 34 , , , 37602 820 35 CAS CAS NNP 37602 820 36 93264 93264 CD 37602 820 37 - - SYM 37602 820 38 74 74 CD 37602 820 39 , , , 37602 820 40 93276 93276 CD 37602 820 41 , , , 37602 820 42 93278 93278 CD 37602 820 43 - - SYM 37602 820 44 9 9 CD 37602 820 45 , , , 37602 820 46 93281 93281 CD 37602 820 47 , , , 37602 820 48 93283 93283 CD 37602 820 49 - - SYM 37602 820 50 6 6 CD 37602 820 51 , , , 37602 820 52 93289 93289 CD 37602 820 53 , , , 37602 820 54 93311 93311 CD 37602 820 55 , , , 37602 820 56 93327 93327 CD 37602 820 57 ; ; : 37602 820 58 Igarapé Igarapé NNP 37602 820 59 Champuia Champuia NNP 37602 820 60 , , , 37602 820 61 upper upper JJ 37602 820 62 Río Río NNP 37602 820 63 Curiuja Curiuja NNP 37602 820 64 , , , 37602 820 65 MZUSP MZUSP NNP 37602 820 66 10339 10339 CD 37602 820 67 ; ; : 37602 820 68 Iquitos Iquitos NNP 37602 820 69 , , , 37602 820 70 AMNH AMNH NNP 37602 820 71 42204 42204 CD 37602 820 72 , , , 37602 820 73 42442 42442 CD 37602 820 74 , , , 37602 820 75 43468 43468 CD 37602 820 76 , , , 37602 820 77 NHMW NHMW NNP 37602 820 78 6118 6118 CD 37602 820 79 ; ; : 37602 820 80 Lago Lago NNP 37602 820 81 de de NNP 37602 820 82 Miraño Miraño NNP 37602 820 83 , , , 37602 820 84 mouth mouth NN 37602 820 85 of of IN 37602 820 86 Río Río NNP 37602 820 87 Napo Napo NNP 37602 820 88 , , , 37602 820 89 AMNH AMNH NNP 37602 820 90 42712 42712 CD 37602 820 91 , , , 37602 820 92 43186 43186 CD 37602 820 93 ; ; : 37602 820 94 Nauta Nauta NNP 37602 820 95 , , , 37602 820 96 ANSP ANSP NNP 37602 820 97 11399 11399 CD 37602 820 98 ; ; : 37602 820 99 Ollanta Ollanta NNP 37602 820 100 , , , 37602 820 101 AMNH AMNH NNP 37602 820 102 42865 42865 CD 37602 820 103 ; ; : 37602 820 104 Pampa Pampa NNP 37602 820 105 Hermosa Hermosa NNP 37602 820 106 , , , 37602 820 107 Río Río NNP 37602 820 108 Cushabatay Cushabatay NNP 37602 820 109 , , , 37602 820 110 AMNH AMNH NNP 37602 820 111 43124 43124 CD 37602 820 112 , , , 37602 820 113 43146 43146 CD 37602 820 114 ; ; : 37602 820 115 Pebas Pebas NNP 37602 820 116 , , , 37602 820 117 CAS CAS NNP 37602 820 118 - - HYPH 37602 820 119 SU SU NNP 37602 820 120 6375 6375 CD 37602 820 121 ; ; : 37602 820 122 Pucallpa Pucallpa NNP 37602 820 123 , , , 37602 820 124 MJP MJP NNP 37602 820 125 101(2 101(2 CD 37602 820 126 ) ) -RRB- 37602 820 127 , , , 37602 820 128 140(3 140(3 CD 37602 820 129 ) ) -RRB- 37602 820 130 ; ; : 37602 820 131 Punga Punga NNP 37602 820 132 , , , 37602 820 133 Río Río NNP 37602 820 134 Tapiche Tapiche NNP 37602 820 135 , , , 37602 820 136 AMNH AMNH NNP 37602 820 137 43194 43194 CD 37602 820 138 ; ; : 37602 820 139 " " `` 37602 820 140 Rancho Rancho NNP 37602 820 141 de de IN 37602 820 142 Indiana Indiana NNP 37602 820 143 , , , 37602 820 144 Iquitos Iquitos NNP 37602 820 145 District District NNP 37602 820 146 , , , 37602 820 147 " " '' 37602 820 148 MVZ MVZ NNP 37602 820 149 16890 16890 CD 37602 820 150 ; ; : 37602 820 151 upper upper JJ 37602 820 152 Río Río NNP 37602 820 153 Abujao Abujao NNP 37602 820 154 , , , 37602 820 155 AMNH AMNH NNP 37602 820 156 42908 42908 CD 37602 820 157 ; ; : 37602 820 158 Río Río NNP 37602 820 159 Itaya Itaya NNP 37602 820 160 , , , 37602 820 161 AMNH AMNH NNP 37602 820 162 42755 42755 CD 37602 820 163 ; ; : 37602 820 164 upper upper JJ 37602 820 165 Río Río NNP 37602 820 166 Pisqui Pisqui NNP 37602 820 167 , , , 37602 820 168 AMNH AMNH NNP 37602 820 169 43536 43536 CD 37602 820 170 ; ; : 37602 820 171 Río Río NNP 37602 820 172 Tapiche Tapiche NNP 37602 820 173 at at IN 37602 820 174 Río Río NNP 37602 820 175 Contaya Contaya NNP 37602 820 176 , , , 37602 820 177 AMNH AMNH NNP 37602 820 178 42983 42983 CD 37602 820 179 ; ; : 37602 820 180 Río Río NNP 37602 820 181 Utoquinia Utoquinia NNP 37602 820 182 at at IN 37602 820 183 Brasilian Brasilian NNP 37602 820 184 frontier frontier NN 37602 820 185 , , , 37602 820 186 AMNH AMNH NNP 37602 820 187 43137 43137 CD 37602 820 188 ; ; : 37602 820 189 Sobral Sobral NNP 37602 820 190 , , , 37602 820 191 Río Río NNP 37602 820 192 Tamaya Tamaya NNP 37602 820 193 , , , 37602 820 194 AMNH AMNH NNP 37602 820 195 43242 43242 CD 37602 820 196 ; ; : 37602 820 197 Yurimaguas yurimaguas UH 37602 820 198 , , , 37602 820 199 BMNH BMNH NNP 37602 820 200 84.2.18.50 84.2.18.50 CD 37602 820 201 . . . 37602 821 1 _ _ NNP 37602 821 2 San San NNP 37602 821 3 Martín Martín NNP 37602 821 4 _ _ NNP 37602 821 5 : : : 37602 821 6 Cainarachi Cainarachi NNP 37602 821 7 , , , 37602 821 8 AMNH AMNH NNP 37602 821 9 42763 42763 CD 37602 821 10 ; ; : 37602 821 11 Moyobamba Moyobamba NNP 37602 821 12 , , , 37602 821 13 ZSM ZSM NNP 37602 821 14 19/1914 19/1914 CD 37602 821 15 . . . 37602 822 1 SURINAM SURINAM NNP 37602 822 2 : : : 37602 822 3 No no DT 37602 822 4 specific specific JJ 37602 822 5 locality locality NN 37602 822 6 , , , 37602 822 7 BMNH BMNH NNP 37602 822 8 70.3.10.67 70.3.10.67 CD 37602 822 9 , , , 37602 822 10 NHMW NHMW NNP 37602 822 11 18433.3 18433.3 CD 37602 822 12 . . . 37602 823 1 _ _ NNP 37602 823 2 Brokopondo Brokopondo NNP 37602 823 3 _ _ NNP 37602 823 4 : : : 37602 823 5 Afobaka Afobaka NNP 37602 823 6 , , , 37602 823 7 RMNH RMNH NNP 37602 823 8 16536 16536 CD 37602 823 9 ; ; : 37602 823 10 Brownsweg Brownsweg NNP 37602 823 11 , , , 37602 823 12 RMNH RMNH NNP 37602 823 13 16537 16537 CD 37602 823 14 ; ; : 37602 823 15 Railway Railway NNP 37602 823 16 km km NNP 37602 823 17 . . . 37602 824 1 121 121 CD 37602 824 2 , , , 37602 824 3 RMNH RMNH NNP 37602 824 4 16534 16534 CD 37602 824 5 . . . 37602 825 1 _ _ NNP 37602 825 2 Marowijne Marowijne NNP 37602 825 3 _ _ NNP 37602 825 4 : : : 37602 825 5 Djai Djai NNP 37602 825 6 Creek Creek NNP 37602 825 7 , , , 37602 825 8 RMNH RMNH NNP 37602 825 9 16513 16513 CD 37602 825 10 - - SYM 37602 825 11 4 4 CD 37602 825 12 ; ; : 37602 825 13 Maroni Maroni NNP 37602 825 14 River River NNP 37602 825 15 , , , 37602 825 16 ZMB ZMB NNP 37602 825 17 8240 8240 CD 37602 825 18 , , , 37602 825 19 8531 8531 CD 37602 825 20 ; ; : 37602 825 21 Nassaugebergte Nassaugebergte NNP 37602 825 22 , , , 37602 825 23 RMNH RMNH NNP 37602 825 24 16517 16517 CD 37602 825 25 - - SYM 37602 825 26 33 33 CD 37602 825 27 ; ; : 37602 825 28 Paloemeu Paloemeu NNP 37602 825 29 , , , 37602 825 30 USNM USNM NNP 37602 825 31 159025 159025 CD 37602 825 32 ; ; : 37602 825 33 Swamp Swamp NNP 37602 825 34 Camp Camp NNP 37602 825 35 , , , 37602 825 36 RMNH RMNH NNP 37602 825 37 16515 16515 CD 37602 825 38 . . . 37602 826 1 _ _ NNP 37602 826 2 Nickerie Nickerie NNP 37602 826 3 _ _ NNP 37602 826 4 : : : 37602 826 5 Sipaliwini Sipaliwini NNP 37602 826 6 , , , 37602 826 7 RMNH RMNH NNP 37602 826 8 16538 16538 CD 37602 826 9 . . . 37602 827 1 _ _ NNP 37602 827 2 Saramacca Saramacca NNP 37602 827 3 _ _ NNP 37602 827 4 : : : 37602 827 5 Left Left NNP 37602 827 6 Coppename Coppename NNP 37602 827 7 River River NNP 37602 827 8 , , , 37602 827 9 RMNH RMNH NNP 37602 827 10 16535 16535 CD 37602 827 11 ; ; : 37602 827 12 Tibiti Tibiti NNP 37602 827 13 , , , 37602 827 14 RMNH RMNH NNP 37602 827 15 16516 16516 CD 37602 827 16 . . . 37602 828 1 _ _ NNP 37602 828 2 Suriname Suriname NNP 37602 828 3 _ _ NNP 37602 828 4 : : : 37602 828 5 Berlijn Berlijn NNP 37602 828 6 , , , 37602 828 7 RMNH RMNH NNP 37602 828 8 15064 15064 CD 37602 828 9 ; ; : 37602 828 10 Powakka Powakka NNP 37602 828 11 , , , 37602 828 12 CM CM NNP 37602 828 13 44226 44226 CD 37602 828 14 ; ; : 37602 828 15 Zanderij Zanderij NNP 37602 828 16 , , , 37602 828 17 CM CM NNP 37602 828 18 50568 50568 CD 37602 828 19 . . . 37602 829 1 VENEZUELA VENEZUELA NNP 37602 829 2 : : : 37602 829 3 _ _ NNP 37602 829 4 Amazonas Amazonas NNP 37602 829 5 _ _ NNP 37602 829 6 : : : 37602 829 7 Cerro Cerro NNP 37602 829 8 Duida Duida NNP 37602 829 9 , , , 37602 829 10 UPR UPR NNP 37602 829 11 - - HYPH 37602 829 12 M M NNP 37602 829 13 2875 2875 CD 37602 829 14 ; ; : 37602 829 15 Cerro Cerro NNP 37602 829 16 Marahuaca Marahuaca NNP 37602 829 17 , , , 37602 829 18 UPR UPR NNP 37602 829 19 - - HYPH 37602 829 20 M M NNP 37602 829 21 114 114 CD 37602 829 22 - - SYM 37602 829 23 5 5 CD 37602 829 24 ; ; : 37602 829 25 Esmeralda Esmeralda NNP 37602 829 26 , , , 37602 829 27 AMNH AMNH NNP 37602 829 28 23174 23174 CD 37602 829 29 ; ; : 37602 829 30 Iniridi Iniridi NNP 37602 829 31 , , , 37602 829 32 SMF SMF NNP 37602 829 33 2640 2640 CD 37602 829 34 ; ; : 37602 829 35 La La NNP 37602 829 36 Culebra Culebra NNP 37602 829 37 , , , 37602 829 38 MCZ MCZ NNP 37602 829 39 28572 28572 CD 37602 829 40 , , , 37602 829 41 UPR UPR NNP 37602 829 42 - - HYPH 37602 829 43 M M NNP 37602 829 44 117 117 CD 37602 829 45 ; ; : 37602 829 46 Laguna Laguna NNP 37602 829 47 , , , 37602 829 48 between between IN 37602 829 49 Tama Tama NNP 37602 829 50 Tama Tama NNP 37602 829 51 and and CC 37602 829 52 Esmeralda Esmeralda NNP 37602 829 53 , , , 37602 829 54 UPR UPR NNP 37602 829 55 - - HYPH 37602 829 56 M M NNP 37602 829 57 2760 2760 CD 37602 829 58 ; ; : 37602 829 59 Río Río NNP 37602 829 60 Pescado Pescado NNP 37602 829 61 , , , 37602 829 62 AMNH AMNH NNP 37602 829 63 23177 23177 CD 37602 829 64 ; ; : 37602 829 65 Tapara Tapara NNP 37602 829 66 , , , 37602 829 67 UPR UPR NNP 37602 829 68 - - HYPH 37602 829 69 M M NNP 37602 829 70 113 113 CD 37602 829 71 . . . 37602 830 1 _ _ NNP 37602 830 2 Distrito Distrito NNP 37602 830 3 Federal Federal NNP 37602 830 4 _ _ NNP 37602 830 5 : : : 37602 830 6 Caracas Caracas NNP 37602 830 7 , , , 37602 830 8 BMNH BMNH NNP 37602 830 9 51.7.17.182 51.7.17.182 CD 37602 830 10 . . . 37602 831 1 _ _ NNP 37602 831 2 Osteocephalus Osteocephalus NNP 37602 831 3 verrucigerus verrucigerus NN 37602 831 4 _ _ NNP 37602 831 5 COLOMBIA COLOMBIA NNP 37602 831 6 : : : 37602 831 7 _ _ NNP 37602 831 8 Huila Huila NNP 37602 831 9 _ _ NNP 37602 831 10 : : : 37602 831 11 Acevedo Acevedo NNP 37602 831 12 , , , 37602 831 13 Río Río NNP 37602 831 14 Suaza Suaza NNP 37602 831 15 , , , 37602 831 16 FMNH FMNH NNP 37602 831 17 69709 69709 CD 37602 831 18 - - SYM 37602 831 19 10 10 CD 37602 831 20 . . . 37602 832 1 ECUADOR ECUADOR NNP 37602 832 2 : : : 37602 832 3 No no DT 37602 832 4 specific specific JJ 37602 832 5 locality locality NN 37602 832 6 , , , 37602 832 7 ZMB ZMB NNP 37602 832 8 16589 16589 CD 37602 832 9 . . . 37602 833 1 _ _ NNP 37602 833 2 Napo Napo NNP 37602 833 3 _ _ NNP 37602 833 4 : : : 37602 833 5 Avila Avila NNP 37602 833 6 , , , 37602 833 7 UMMZ UMMZ NNP 37602 833 8 90413 90413 CD 37602 833 9 ; ; : 37602 833 10 south south JJ 37602 833 11 slope slope NN 37602 833 12 Cordillera Cordillera NNP 37602 833 13 del del NNP 37602 833 14 Dué Dué NNP 37602 833 15 , , , 37602 833 16 KU KU NNP 37602 833 17 123176 123176 CD 37602 833 18 - - SYM 37602 833 19 88 88 CD 37602 833 20 , , , 37602 833 21 123189 123189 CD 37602 833 22 ( ( -LRB- 37602 833 23 skeleton skeleton NNP 37602 833 24 ) ) -RRB- 37602 833 25 , , , 37602 833 26 124208 124208 CD 37602 833 27 ( ( -LRB- 37602 833 28 eggs egg NNS 37602 833 29 ) ) -RRB- 37602 833 30 , , , 37602 833 31 124209 124209 CD 37602 833 32 - - SYM 37602 833 33 11 11 CD 37602 833 34 ( ( -LRB- 37602 833 35 tadpoles tadpole NNS 37602 833 36 ) ) -RRB- 37602 833 37 ; ; : 37602 833 38 L'Alegria L'Alegria NNP 37602 833 39 , , , 37602 833 40 USNM USNM NNP 37602 833 41 167472 167472 CD 37602 833 42 - - SYM 37602 833 43 3 3 CD 37602 833 44 ; ; : 37602 833 45 Río Río NNP 37602 833 46 Pacayacu Pacayacu NNP 37602 833 47 , , , 37602 833 48 tributary tributary NN 37602 833 49 of of IN 37602 833 50 Río Río NNP 37602 833 51 Cotapino Cotapino NNP 37602 833 52 , , , 37602 833 53 CAS CAS NNP 37602 833 54 - - HYPH 37602 833 55 SU SU NNP 37602 833 56 13150 13150 CD 37602 833 57 ; ; : 37602 833 58 southeast southeast JJ 37602 833 59 slope slope NN 37602 833 60 Volcán Volcán NNP 37602 833 61 Sumaco Sumaco NNP 37602 833 62 , , , 37602 833 63 CAS CAS NNP 37602 833 64 - - HYPH 37602 833 65 SU SU NNP 37602 833 66 11442 11442 CD 37602 833 67 . . . 37602 834 1 _ _ NNP 37602 834 2 Pastaza Pastaza NNP 37602 834 3 _ _ NNP 37602 834 4 : : : 37602 834 5 Abitagua Abitagua NNP 37602 834 6 , , , 37602 834 7 CAS CAS NNP 37602 834 8 - - HYPH 37602 834 9 SU SU NNP 37602 834 10 5067 5067 CD 37602 834 11 , , , 37602 834 12 FMNH FMNH NNP 37602 834 13 25791 25791 CD 37602 834 14 , , , 37602 834 15 27619 27619 CD 37602 834 16 , , , 37602 834 17 UMMZ UMMZ NNP 37602 834 18 90414 90414 CD 37602 834 19 , , , 37602 834 20 92092 92092 CD 37602 834 21 ; ; : 37602 834 22 Alpayaca Alpayaca NNP 37602 834 23 , , , 37602 834 24 Río Río NNP 37602 834 25 Pastaza Pastaza NNP 37602 834 26 , , , 37602 834 27 BMNH BMNH VBZ 37602 834 28 1912.11.1.64 1912.11.1.64 CD 37602 834 29 ; ; : 37602 834 30 Mera Mera NNP 37602 834 31 , , , 37602 834 32 UMMZ UMMZ NNP 37602 834 33 90412(4 90412(4 CD 37602 834 34 ) ) -RRB- 37602 834 35 . . . 37602 835 1 _ _ NNP 37602 835 2 Tungurahua Tungurahua NNP 37602 835 3 _ _ NNP 37602 835 4 : : : 37602 835 5 Baños Baños NNP 37602 835 6 , , , 37602 835 7 NHRM nhrm NN 37602 835 8 1960 1960 CD 37602 835 9 . . . 37602 836 1 PERÚ PERÚ NNP 37602 836 2 : : : 37602 836 3 _ _ NNP 37602 836 4 Ayacucho Ayacucho NNP 37602 836 5 _ _ NNP 37602 836 6 : : : 37602 836 7 La La NNP 37602 836 8 Mar Mar NNP 37602 836 9 , , , 37602 836 10 Sivia Sivia NNP 37602 836 11 , , , 37602 836 12 Río Río NNP 37602 836 13 Apurimac Apurimac NNP 37602 836 14 , , , 37602 836 15 FMNH FMNH NNP 37602 836 16 39853 39853 CD 37602 836 17 . . . 37602 837 1 _ _ NNP 37602 837 2 Huanuco Huanuco NNP 37602 837 3 _ _ NNP 37602 837 4 : : : 37602 837 5 Río Río NNP 37602 837 6 Pachitea Pachitea NNP 37602 837 7 , , , 37602 837 8 midway midway RB 37602 837 9 between between IN 37602 837 10 Puerto Puerto NNP 37602 837 11 Victoria Victoria NNP 37602 837 12 and and CC 37602 837 13 Puerto Puerto NNP 37602 837 14 Inca Inca NNP 37602 837 15 , , , 37602 837 16 CAS CAS NNP 37602 837 17 - - HYPH 37602 837 18 SU SU NNP 37602 837 19 17745 17745 CD 37602 837 20 . . . 37602 838 1 _ _ NNP 37602 838 2 Junín Junín NNP 37602 838 3 _ _ NNP 37602 838 4 : : : 37602 838 5 Satipo Satipo NNP 37602 838 6 , , , 37602 838 7 MJP MJP NNP 37602 838 8 38 38 CD 37602 838 9 . . . 37602 839 1 LITERATURE LITERATURE NNP 37602 839 2 CITED cite VBD 37602 839 3 ANDERSSON ANDERSSON NNP 37602 839 4 , , , 37602 839 5 L. L. NNP 37602 839 6 G. G. NNP 37602 839 7 1945 1945 CD 37602 839 8 . . . 37602 840 1 Batrachians batrachian NNS 37602 840 2 from from IN 37602 840 3 east east JJ 37602 840 4 Ecuador Ecuador NNP 37602 840 5 collected collect VBD 37602 840 6 1937 1937 CD 37602 840 7 , , , 37602 840 8 1938 1938 CD 37602 840 9 by by IN 37602 840 10 Win Win NNP 37602 840 11 . . . 37602 841 1 Clarke Clarke NNP 37602 841 2 - - HYPH 37602 841 3 MacIntyre MacIntyre NNP 37602 841 4 and and CC 37602 841 5 Rolf Rolf NNP 37602 841 6 Blomberg Blomberg NNP 37602 841 7 . . . 37602 842 1 Arkiv Arkiv NNP 37602 842 2 Zool Zool NNP 37602 842 3 . . NNP 37602 842 4 , , , 37602 842 5 37A(2):1 37a(2):1 CD 37602 842 6 - - SYM 37602 842 7 88 88 CD 37602 842 8 . . . 37602 843 1 BOKERMANN BOKERMANN NNP 37602 843 2 , , , 37602 843 3 W. W. NNP 37602 843 4 C. C. NNP 37602 843 5 A. a. NN 37602 844 1 1964 1964 CD 37602 844 2 . . . 37602 845 1 Field field NN 37602 845 2 observations observation NNS 37602 845 3 on on IN 37602 845 4 the the DT 37602 845 5 hylid hylid JJ 37602 845 6 frog frog NN 37602 845 7 _ _ NNP 37602 845 8 Osteocephalus osteocephalus JJ 37602 845 9 taurinus taurinus NN 37602 845 10 _ _ NNP 37602 845 11 Fitz Fitz NNP 37602 845 12 . . . 37602 846 1 Herpetologica Herpetologica NNP 37602 846 2 , , , 37602 846 3 20:252 20:252 CD 37602 846 4 - - SYM 37602 846 5 255 255 CD 37602 846 6 . . . 37602 847 1 1966 1966 CD 37602 847 2 . . . 37602 848 1 Lista Lista NNP 37602 848 2 anotada anotada VBP 37602 848 3 das das NNP 37602 848 4 localidades localidade NNS 37602 848 5 tipo tipo NNP 37602 848 6 de de FW 37602 848 7 anfíbios anfíbios NNP 37602 848 8 Brasileiros Brasileiros NNP 37602 848 9 . . . 37602 849 1 São São NNP 37602 849 2 Paulo Paulo NNP 37602 849 3 , , , 37602 849 4 183 183 CD 37602 849 5 pp pp NNS 37602 849 6 . . . 37602 850 1 BOULENGER BOULENGER NNP 37602 850 2 , , , 37602 850 3 G. G. NNP 37602 850 4 A. A. NNP 37602 851 1 1882 1882 CD 37602 851 2 . . . 37602 852 1 Catalogue catalogue NN 37602 852 2 of of IN 37602 852 3 the the DT 37602 852 4 Batrachia Batrachia NNP 37602 852 5 Salientia Salientia NNP 37602 852 6 s. s. NNP 37602 852 7 Ecaudata Ecaudata NNP 37602 852 8 in in IN 37602 852 9 the the DT 37602 852 10 collection collection NN 37602 852 11 of of IN 37602 852 12 the the DT 37602 852 13 British British NNP 37602 852 14 Museum Museum NNP 37602 852 15 , , , 37602 852 16 ed ed NNP 37602 852 17 . . . 37602 853 1 2 2 LS 37602 853 2 , , , 37602 853 3 London London NNP 37602 853 4 , , , 37602 853 5 xvi+503 xvi+503 NNP 37602 853 6 pp pp NNP 37602 853 7 . . . 37602 854 1 COCHRAN COCHRAN NNP 37602 854 2 , , , 37602 854 3 D. D. NNP 37602 854 4 M. M. NNP 37602 854 5 and and CC 37602 854 6 C. C. NNP 37602 854 7 J. J. NNP 37602 854 8 GOIN GOIN NNP 37602 854 9 1970 1970 CD 37602 854 10 . . . 37602 855 1 Frogs Frogs NNP 37602 855 2 of of IN 37602 855 3 Colombia Colombia NNP 37602 855 4 . . . 37602 856 1 Bull Bull NNP 37602 856 2 . . . 37602 857 1 U.S. U.S. NNP 37602 857 2 Natl Natl NNP 37602 857 3 . . . 37602 858 1 Mus Mus NNP 37602 858 2 . . NNP 37602 858 3 , , , 37602 858 4 288 288 CD 37602 858 5 : : : 37602 858 6 xii+655 xii+655 NNP 37602 858 7 pp pp NNP 37602 858 8 . . . 37602 859 1 COPE COPE NNP 37602 859 2 , , , 37602 859 3 E. E. NNP 37602 859 4 D. D. NNP 37602 859 5 1867 1867 CD 37602 859 6 . . . 37602 860 1 On on IN 37602 860 2 the the DT 37602 860 3 families family NNS 37602 860 4 of of IN 37602 860 5 the the DT 37602 860 6 raniform raniform NN 37602 860 7 Anura Anura NNP 37602 860 8 . . . 37602 861 1 Jour Jour NNP 37602 861 2 . . . 37602 862 1 Acad Acad NNS 37602 862 2 . . . 37602 863 1 Nat Nat NNP 37602 863 2 . . . 37602 864 1 Sci Sci NNP 37602 864 2 . . . 37602 865 1 Philadelphia Philadelphia NNP 37602 865 2 , , , 37602 865 3 2:189 2:189 CD 37602 865 4 - - SYM 37602 865 5 206 206 CD 37602 865 6 . . . 37602 866 1 1874 1874 CD 37602 866 2 . . . 37602 867 1 On on IN 37602 867 2 some some DT 37602 867 3 Batrachia Batrachia NNP 37602 867 4 and and CC 37602 867 5 Nematognathi Nematognathi NNP 37602 867 6 brought bring VBD 37602 867 7 from from IN 37602 867 8 the the DT 37602 867 9 upper upper JJ 37602 867 10 Amazon Amazon NNP 37602 867 11 by by IN 37602 867 12 Prof. Prof. NNP 37602 867 13 Orton Orton NNP 37602 867 14 . . . 37602 868 1 Proc Proc NNP 37602 868 2 . . . 37602 869 1 Acad Acad NNS 37602 869 2 . . . 37602 870 1 Nat Nat NNP 37602 870 2 . . . 37602 871 1 Sci Sci NNP 37602 871 2 . . . 37602 872 1 Philadelphia Philadelphia NNP 37602 872 2 , , , 37602 872 3 25:120 25:120 CD 37602 872 4 - - SYM 37602 872 5 137 137 CD 37602 872 6 . . . 37602 873 1 DUELLMAN DUELLMAN NNP 37602 873 2 , , , 37602 873 3 W. W. NNP 37602 873 4 E. E. NNP 37602 873 5 1970a 1970a CD 37602 873 6 . . . 37602 874 1 Identity identity NN 37602 874 2 of of IN 37602 874 3 the the DT 37602 874 4 South south JJ 37602 874 5 American american JJ 37602 874 6 hylid hylid NNP 37602 874 7 frog frog NN 37602 874 8 _ _ NNP 37602 874 9 Garbeana Garbeana NNP 37602 874 10 garbei garbei NNP 37602 874 11 _ _ NNP 37602 874 12 . . . 37602 875 1 Copeia Copeia NNP 37602 875 2 , , , 37602 875 3 ( ( -LRB- 37602 875 4 3):534 3):534 JJ 37602 875 5 - - SYM 37602 875 6 538 538 CD 37602 875 7 . . . 37602 876 1 1970b 1970b CD 37602 876 2 . . . 37602 877 1 The the DT 37602 877 2 hylid hylid JJ 37602 877 3 frogs frog NNS 37602 877 4 of of IN 37602 877 5 Middle Middle NNP 37602 877 6 America America NNP 37602 877 7 . . . 37602 878 1 Monog Monog NNP 37602 878 2 . . . 37602 879 1 Mus mus NN 37602 879 2 . . . 37602 880 1 Nat Nat NNP 37602 880 2 . . . 37602 881 1 Hist Hist NNP 37602 881 2 . . . 37602 881 3 , , , 37602 881 4 Univ Univ NNP 37602 881 5 . . . 37602 882 1 Kansas Kansas NNP 37602 882 2 , , , 37602 882 3 1 1 CD 37602 882 4 : : : 37602 882 5 xi+753 xi+753 NNP 37602 882 6 pp pp NNP 37602 882 7 . . . 37602 883 1 DUMÉRIL DUMÉRIL NNP 37602 883 2 , , , 37602 883 3 A. a. NN 37602 883 4 M. M. NNP 37602 883 5 C. C. NNP 37602 883 6 and and CC 37602 883 7 G. G. NNP 37602 883 8 BIBRON BIBRON NNP 37602 883 9 1841 1841 CD 37602 883 10 . . . 37602 884 1 Erpétologie Erpétologie VBG 37602 884 2 générale générale JJ 37602 884 3 ou ou NNP 37602 884 4 histoire histoire NNP 37602 884 5 naturelle naturelle NNP 37602 884 6 compléte compléte NNP 37602 884 7 des des NNP 37602 884 8 reptiles reptiles NNP 37602 884 9 , , , 37602 884 10 vol vol NNP 37602 884 11 . . . 37602 885 1 8 8 LS 37602 885 2 . . . 37602 886 1 Paris Paris NNP 37602 886 2 , , , 37602 886 3 792 792 CD 37602 886 4 pp pp NNP 37602 886 5 . . . 37602 887 1 FITZINGER FITZINGER NNP 37602 887 2 , , , 37602 887 3 L. L. NNP 37602 887 4 1843 1843 CD 37602 887 5 . . . 37602 888 1 Systema systema JJ 37602 888 2 reptilium reptilium NN 37602 888 3 . . . 37602 889 1 Vienna Vienna NNP 37602 889 2 , , , 37602 889 3 ix+106 ix+106 XX 37602 889 4 pp pp NNP 37602 889 5 . . . 37602 890 1 FUNKHOUSER FUNKHOUSER NNP 37602 890 2 , , , 37602 890 3 J. J. NNP 37602 891 1 1956 1956 CD 37602 891 2 . . . 37602 892 1 New new JJ 37602 892 2 frogs frog NNS 37602 892 3 from from IN 37602 892 4 Ecuador Ecuador NNP 37602 892 5 and and CC 37602 892 6 southwestern southwestern JJ 37602 892 7 Colombia Colombia NNP 37602 892 8 . . . 37602 893 1 Zoologica Zoologica NNP 37602 893 2 , , , 37602 893 3 91:73 91:73 CD 37602 893 4 - - SYM 37602 893 5 80 80 CD 37602 893 6 . . . 37602 894 1 GAIGE GAIGE NNP 37602 894 2 , , , 37602 894 3 H. H. NNP 37602 894 4 T. T. NNP 37602 894 5 1929 1929 CD 37602 894 6 . . . 37602 895 1 Three three CD 37602 895 2 new new JJ 37602 895 3 tree tree NN 37602 895 4 - - HYPH 37602 895 5 frogs frog NNS 37602 895 6 from from IN 37602 895 7 Panama Panama NNP 37602 895 8 and and CC 37602 895 9 Bolivia Bolivia NNP 37602 895 10 . . . 37602 896 1 Occas occas JJ 37602 896 2 . . . 37602 897 1 Papers Papers NNPS 37602 897 2 Mus Mus NNP 37602 897 3 . . . 37602 898 1 Zool Zool NNP 37602 898 2 . . . 37602 899 1 Univ Univ NNP 37602 899 2 . . . 37602 900 1 Michigan Michigan NNP 37602 900 2 , , , 37602 900 3 207:1 207:1 CD 37602 900 4 - - SYM 37602 900 5 6 6 CD 37602 900 6 . . . 37602 901 1 GOIN GOIN NNP 37602 901 2 , , , 37602 901 3 C. C. NNP 37602 901 4 J. J. NNP 37602 902 1 1961 1961 CD 37602 902 2 . . . 37602 903 1 Synopsis synopsis NN 37602 903 2 of of IN 37602 903 3 the the DT 37602 903 4 genera genera NN 37602 903 5 of of IN 37602 903 6 hylid hylid NNP 37602 903 7 frogs frog NNS 37602 903 8 . . . 37602 904 1 Ann Ann NNP 37602 904 2 . . . 37602 905 1 Carnegie Carnegie NNP 37602 905 2 Mus Mus NNP 37602 905 3 . . NNP 37602 905 4 , , , 37602 905 5 36:5 36:5 CD 37602 905 6 - - SYM 37602 905 7 18 18 CD 37602 905 8 . . . 37602 906 1 MELIN MELIN NNP 37602 906 2 , , , 37602 906 3 D. D. NNP 37602 906 4 1941 1941 CD 37602 906 5 . . . 37602 907 1 Contribution contribution NN 37602 907 2 to to IN 37602 907 3 the the DT 37602 907 4 knowledge knowledge NN 37602 907 5 of of IN 37602 907 6 Amphibia Amphibia NNP 37602 907 7 of of IN 37602 907 8 South South NNP 37602 907 9 America America NNP 37602 907 10 . . . 37602 908 1 Göteborgs Göteborgs NNP 37602 908 2 Kungl Kungl NNP 37602 908 3 . . . 37602 909 1 Vetensk.-och Vetensk.-och NNP 37602 909 2 Vitterh.-Sam Vitterh.-Sam NNP 37602 909 3 . . . 37602 910 1 Handl handl RB 37602 910 2 . . . 37602 910 3 , , , 37602 910 4 Ser Ser NNP 37602 910 5 . . . 37602 911 1 B b NN 37602 911 2 , , , 37602 911 3 1(4):1 1(4):1 NNPS 37602 911 4 - - SYM 37602 911 5 71 71 CD 37602 911 6 . . . 37602 912 1 PERACCA PERACCA NNP 37602 912 2 , , , 37602 912 3 M. M. NNP 37602 912 4 G. G. NNP 37602 912 5 1904 1904 CD 37602 912 6 . . . 37602 913 1 Viaggio Viaggio NNP 37602 913 2 del del NNP 37602 913 3 Dr. Dr. NNP 37602 913 4 Enrico Enrico NNP 37602 913 5 Festa Festa NNP 37602 913 6 nell nell RB 37602 913 7 ' ' `` 37602 913 8 Ecuador Ecuador NNP 37602 913 9 e e NNP 37602 913 10 regioni regioni NN 37602 913 11 vicine vicine NN 37602 913 12 . . . 37602 914 1 Reptile reptile NN 37602 914 2 ed ed NN 37602 914 3 amfibii amfibii NN 37602 914 4 . . . 37602 915 1 Boll Boll NNP 37602 915 2 . . . 37602 916 1 Mus mus NN 37602 916 2 . . . 37602 917 1 Zool Zool NNP 37602 917 2 . . . 37602 918 1 Anat Anat NNP 37602 918 2 . . . 37602 919 1 Comp Comp NNP 37602 919 2 . . . 37602 919 3 , , , 37602 919 4 Univ Univ NNP 37602 919 5 . . . 37602 920 1 Torino Torino NNP 37602 920 2 , , , 37602 920 3 19:1 19:1 CD 37602 920 4 - - SYM 37602 920 5 41 41 CD 37602 920 6 . . . 37602 921 1 STEINDACHNER STEINDACHNER NNP 37602 921 2 , , , 37602 921 3 F. F. NNP 37602 921 4 1862 1862 CD 37602 921 5 . . . 37602 922 1 Über Über NNP 37602 922 2 zwei zwei NNP 37602 922 3 noch noch NNP 37602 922 4 unbeschriebene unbeschriebene NN 37602 922 5 Batrachier Batrachier NNP 37602 922 6 . . . 37602 923 1 Arch arch UH 37602 923 2 . . . 37602 924 1 Zool Zool NNP 37602 924 2 . . . 37602 925 1 Anat Anat NNP 37602 925 2 . . . 37602 926 1 Fisiol Fisiol NNP 37602 926 2 . . . 37602 926 3 , , , 37602 926 4 2:77 2:77 CD 37602 926 5 - - SYM 37602 926 6 82 82 CD 37602 926 7 . . . 37602 927 1 1867 1867 CD 37602 927 2 . . . 37602 928 1 Amphibien Amphibien NNP 37602 928 2 . . . 37602 929 1 Novara Novara NNP 37602 929 2 Expedition Expedition NNP 37602 929 3 . . . 37602 930 1 Zool Zool NNP 37602 930 2 . . . 37602 931 1 Theil Theil NNP 37602 931 2 , , , 37602 931 3 I -PRON- PRP 37602 931 4 , , , 37602 931 5 Vienna Vienna NNP 37602 931 6 , , , 37602 931 7 70 70 CD 37602 931 8 pp pp NN 37602 931 9 . . . 37602 932 1 TRUEB TRUEB NNP 37602 932 2 , , , 37602 932 3 L. L. NNP 37602 932 4 1970a 1970a CD 37602 932 5 . . . 37602 933 1 The the DT 37602 933 2 evolutionary evolutionary JJ 37602 933 3 relationships relationship NNS 37602 933 4 of of IN 37602 933 5 casque casque NN 37602 933 6 - - HYPH 37602 933 7 headed head VBN 37602 933 8 treefrogs treefrog NNS 37602 933 9 with with IN 37602 933 10 co co JJ 37602 933 11 - - JJ 37602 933 12 ossified ossified JJ 37602 933 13 skulls skull NNS 37602 933 14 ( ( -LRB- 37602 933 15 family family NN 37602 933 16 Hylidae Hylidae NNP 37602 933 17 ) ) -RRB- 37602 933 18 . . . 37602 934 1 Univ Univ NNP 37602 934 2 . . . 37602 935 1 Kansas Kansas NNP 37602 935 2 Publ Publ NNP 37602 935 3 . . . 37602 936 1 Mus mus NN 37602 936 2 . . . 37602 937 1 Nat Nat NNP 37602 937 2 . . . 37602 938 1 Hist Hist NNP 37602 938 2 . . NNP 37602 938 3 , , , 37602 938 4 18:547 18:547 CD 37602 938 5 - - SYM 37602 938 6 716 716 CD 37602 938 7 . . . 37602 939 1 1970b 1970b CD 37602 939 2 . . . 37602 940 1 The the DT 37602 940 2 generic generic JJ 37602 940 3 status status NN 37602 940 4 of of IN 37602 940 5 _ _ NNP 37602 940 6 Hyla Hyla NNP 37602 940 7 siemersi siemersi NN 37602 940 8 _ _ NNP 37602 940 9 Mertens Mertens NNP 37602 940 10 . . . 37602 941 1 Herpetologica Herpetologica NNP 37602 941 2 , , , 37602 941 3 26:254 26:254 CD 37602 941 4 - - SYM 37602 941 5 267 267 CD 37602 941 6 . . . 37602 942 1 TRUEB TRUEB NNP 37602 942 2 , , , 37602 942 3 L. L. NNP 37602 942 4 and and CC 37602 942 5 W. W. NNP 37602 942 6 E. E. NNP 37602 942 7 DUELLMAN DUELLMAN NNP 37602 942 8 1970 1970 CD 37602 942 9 . . . 37602 943 1 The the DT 37602 943 2 systematic systematic JJ 37602 943 3 status status NN 37602 943 4 and and CC 37602 943 5 life life NN 37602 943 6 history history NN 37602 943 7 of of IN 37602 943 8 _ _ NNP 37602 943 9 Hyla Hyla NNP 37602 943 10 verrucigera verrucigera NN 37602 943 11 _ _ NNP 37602 943 12 Werner Werner NNP 37602 943 13 . . . 37602 944 1 Copeia Copeia NNP 37602 944 2 ( ( -LRB- 37602 944 3 4):601 4):601 NN 37602 944 4 - - SYM 37602 944 5 610 610 CD 37602 944 6 . . . 37602 945 1 TYLER TYLER NNP 37602 945 2 , , , 37602 945 3 M. M. NNP 37602 945 4 1971 1971 CD 37602 945 5 . . . 37602 946 1 The the DT 37602 946 2 phylogenetic phylogenetic JJ 37602 946 3 significance significance NN 37602 946 4 of of IN 37602 946 5 vocal vocal JJ 37602 946 6 sac sac NN 37602 946 7 structure structure NN 37602 946 8 in in IN 37602 946 9 hylid hylid JJ 37602 946 10 frogs frog NNS 37602 946 11 . . . 37602 947 1 Univ Univ NNP 37602 947 2 . . . 37602 948 1 Kansas Kansas NNP 37602 948 2 Publ Publ NNP 37602 948 3 . . . 37602 949 1 Mus mus NN 37602 949 2 . . . 37602 950 1 Nat Nat NNP 37602 950 2 . . . 37602 951 1 Hist Hist NNP 37602 951 2 . . . 37602 951 3 , , , 37602 951 4 19:319 19:319 CD 37602 951 5 - - SYM 37602 951 6 360 360 CD 37602 951 7 . . . 37602 952 1 WERNER WERNER NNP 37602 952 2 , , , 37602 952 3 F. F. NNP 37602 952 4 1901 1901 CD 37602 952 5 . . . 37602 953 1 Ueber Ueber NNP 37602 953 2 Reptilien Reptilien NNP 37602 953 3 und und NN 37602 953 4 Batrachier Batrachier NNP 37602 953 5 aus au VBD 37602 953 6 Ecuador Ecuador NNP 37602 953 7 und und VB 37602 953 8 Neu Neu NNP 37602 953 9 - - HYPH 37602 953 10 Guinea Guinea NNP 37602 953 11 . . . 37602 954 1 Verh Verh NNP 37602 954 2 . . . 37602 955 1 Zool.-Bot Zool.-Bot NNP 37602 955 2 . . . 37602 956 1 Gesell gesell NN 37602 956 2 . . . 37602 957 1 Wien Wien NNP 37602 957 2 , , , 37602 957 3 50:593 50:593 CD 37602 957 4 - - SYM 37602 957 5 614 614 CD 37602 957 6 . . . 37602 958 1 UNIVERSITY UNIVERSITY NNP 37602 958 2 OF of IN 37602 958 3 KANSAS KANSAS NNP 37602 958 4 PUBLICATIONS PUBLICATIONS NNP 37602 958 5 MUSEUM MUSEUM NNP 37602 958 6 OF of IN 37602 958 7 NATURAL natural JJ 37602 958 8 HISTORY HISTORY NNP 37602 958 9 The the DT 37602 958 10 University University NNP 37602 958 11 of of IN 37602 958 12 Kansas Kansas NNP 37602 958 13 Publications Publications NNP 37602 958 14 , , , 37602 958 15 Museum Museum NNP 37602 958 16 of of IN 37602 958 17 Natural Natural NNP 37602 958 18 History History NNP 37602 958 19 , , , 37602 958 20 beginning begin VBG 37602 958 21 with with IN 37602 958 22 volume volume NN 37602 958 23 1 1 CD 37602 958 24 in in IN 37602 958 25 1946 1946 CD 37602 958 26 , , , 37602 958 27 was be VBD 37602 958 28 discontinued discontinue VBN 37602 958 29 with with IN 37602 958 30 volume volume NN 37602 958 31 20 20 CD 37602 958 32 in in IN 37602 958 33 1971 1971 CD 37602 958 34 . . . 37602 959 1 Shorter short JJR 37602 959 2 research research NN 37602 959 3 papers paper NNS 37602 959 4 formerly formerly RB 37602 959 5 published publish VBN 37602 959 6 in in IN 37602 959 7 the the DT 37602 959 8 above above JJ 37602 959 9 series series NN 37602 959 10 are be VBP 37602 959 11 now now RB 37602 959 12 published publish VBN 37602 959 13 as as IN 37602 959 14 Occasional Occasional NNP 37602 959 15 Papers Papers NNP 37602 959 16 , , , 37602 959 17 Museum Museum NNP 37602 959 18 of of IN 37602 959 19 Natural Natural NNP 37602 959 20 History History NNP 37602 959 21 . . . 37602 960 1 The the DT 37602 960 2 Miscellaneous Miscellaneous NNP 37602 960 3 Publications Publications NNPS 37602 960 4 , , , 37602 960 5 Museum Museum NNP 37602 960 6 of of IN 37602 960 7 Natural Natural NNP 37602 960 8 History History NNP 37602 960 9 , , , 37602 960 10 began begin VBD 37602 960 11 with with IN 37602 960 12 number number NN 37602 960 13 1 1 CD 37602 960 14 in in IN 37602 960 15 1946 1946 CD 37602 960 16 . . . 37602 961 1 Longer long JJR 37602 961 2 research research NN 37602 961 3 papers paper NNS 37602 961 4 are be VBP 37602 961 5 published publish VBN 37602 961 6 in in IN 37602 961 7 that that DT 37602 961 8 series series NN 37602 961 9 . . . 37602 962 1 Monographs Monographs NNP 37602 962 2 of of IN 37602 962 3 the the DT 37602 962 4 Museum Museum NNP 37602 962 5 of of IN 37602 962 6 Natural Natural NNP 37602 962 7 History History NNP 37602 962 8 were be VBD 37602 962 9 initiated initiate VBN 37602 962 10 in in IN 37602 962 11 1970 1970 CD 37602 962 12 . . . 37602 963 1 Institutional institutional JJ 37602 963 2 libraries library NNS 37602 963 3 interested interested JJ 37602 963 4 in in IN 37602 963 5 exchanging exchange VBG 37602 963 6 publications publication NNS 37602 963 7 may may MD 37602 963 8 obtain obtain VB 37602 963 9 the the DT 37602 963 10 Occasional Occasional NNP 37602 963 11 Papers Papers NNPS 37602 963 12 and and CC 37602 963 13 Miscellaneous Miscellaneous NNP 37602 963 14 Publications Publications NNP 37602 963 15 by by IN 37602 963 16 addressing address VBG 37602 963 17 the the DT 37602 963 18 Exchange Exchange NNP 37602 963 19 Librarian Librarian NNP 37602 963 20 , , , 37602 963 21 University University NNP 37602 963 22 of of IN 37602 963 23 Kansas Kansas NNP 37602 963 24 Library Library NNP 37602 963 25 , , , 37602 963 26 Lawrence Lawrence NNP 37602 963 27 , , , 37602 963 28 Kansas Kansas NNP 37602 963 29 66044 66044 CD 37602 963 30 . . . 37602 964 1 Individuals individual NNS 37602 964 2 may may MD 37602 964 3 purchase purchase VB 37602 964 4 separate separate JJ 37602 964 5 numbers number NNS 37602 964 6 of of IN 37602 964 7 all all DT 37602 964 8 series series NN 37602 964 9 . . . 37602 965 1 Prices price NNS 37602 965 2 may may MD 37602 965 3 be be VB 37602 965 4 obtained obtain VBN 37602 965 5 upon upon IN 37602 965 6 request request NN 37602 965 7 addressed address VBN 37602 965 8 to to IN 37602 965 9 Publications Publications NNP 37602 965 10 Secretary Secretary NNP 37602 965 11 , , , 37602 965 12 Museum Museum NNP 37602 965 13 of of IN 37602 965 14 Natural Natural NNP 37602 965 15 History History NNP 37602 965 16 , , , 37602 965 17 University University NNP 37602 965 18 of of IN 37602 965 19 Kansas Kansas NNP 37602 965 20 , , , 37602 965 21 Lawrence Lawrence NNP 37602 965 22 , , , 37602 965 23 Kansas Kansas NNP 37602 965 24 66044 66044 CD 37602 965 25 . . . 37602 966 1 Transcriber Transcriber NNP 37602 966 2 's 's POS 37602 966 3 Notes Notes NNPS 37602 966 4 Except except IN 37602 966 5 for for IN 37602 966 6 the the DT 37602 966 7 list list NN 37602 966 8 of of IN 37602 966 9 corrections correction NNS 37602 966 10 below below IN 37602 966 11 and and CC 37602 966 12 minor minor JJ 37602 966 13 corrections correction NNS 37602 966 14 not not RB 37602 966 15 listed list VBN 37602 966 16 , , , 37602 966 17 the the DT 37602 966 18 text text NN 37602 966 19 presented present VBN 37602 966 20 here here RB 37602 966 21 is be VBZ 37602 966 22 that that DT 37602 966 23 of of IN 37602 966 24 the the DT 37602 966 25 original original JJ 37602 966 26 printed print VBN 37602 966 27 version version NN 37602 966 28 . . . 37602 967 1 Typographical Typographical NNP 37602 967 2 Corrections Corrections NNPS 37602 967 3 Page Page NNP 37602 967 4 Correction Correction NNP 37602 967 5 = = SYM 37602 967 6 = = SYM 37602 967 7 = = SYM 37602 967 8 = = SYM 37602 967 9 = = SYM 37602 967 10 = = SYM 37602 967 11 = = SYM 37602 967 12 = = SYM 37602 967 13 = = SYM 37602 967 14 = = SYM 37602 967 15 = = SYM 37602 967 16 = = SYM 37602 967 17 = = SYM 37602 967 18 = = SYM 37602 967 19 = = SYM 37602 967 20 = = SYM 37602 967 21 = = SYM 37602 967 22 = = SYM 37602 967 23 = = SYM 37602 967 24 = = SYM 37602 967 25 5 5 CD 37602 967 26 is be VBZ 37602 967 27 = = SYM 37602 967 28 > > XX 37602 967 29 in in IN 37602 967 30 5 5 CD 37602 967 31 buckley buckley NNP 37602 967 32 = = SYM 37602 967 33 > > XX 37602 967 34 buckleyi buckleyi NNP 37602 967 35 17 17 CD 37602 967 36 scaral scaral JJ 37602 967 37 = = NFP 37602 967 38 > > XX 37602 967 39 sacral sacral JJ 37602 967 40 19 19 CD 37602 967 41 Provicia Provicia NNP 37602 967 42 = = NFP 37602 967 43 > > NFP 37602 967 44 Provincia Provincia NNP 37602 967 45 25 25 CD 37602 967 46 in in IN 37602 967 47 = = NFP 37602 967 48 > > XX 37602 967 49 is be VBZ 37602 967 50 25 25 CD 37602 967 51 metalic metalic JJ 37602 967 52 = = SYM 37602 967 53 > > XX 37602 967 54 metallic metallic NNP 37602 967 55 41 41 CD 37602 967 56 bromeiad bromeiad NNS 37602 967 57 = = SYM 37602 967 58 > > XX 37602 967 59 bromeliad bromeliad NNP 37602 967 60 Text Text NNP 37602 967 61 Emphasis Emphasis NNP 37602 967 62 _ _ NNP 37602 967 63 Text Text NNP 37602 967 64 _ _ NNP 37602 967 65 - - HYPH 37602 967 66 Italics Italics NNP 37602 967 67 = = SYM 37602 967 68 Text= text= NN 37602 967 69 - - HYPH 37602 967 70 Bold bold NN