id author title date pages extension mime words sentences flesch summary cache txt work_jajnfdt3vzhsla3qd3sexbff5u Marshall Abrams The Unity of Fitness 2009 11 .pdf application/pdf 5224 350 56 number of offspring; different mathematical functions are needed to define fitness in Though in such cases fitness must be defined in terms of probabilities of reproductive effects over several generations, I argue that it nevertheless has to do with Notice that the worry that prompted Beatty and Finsen and Brandon to revise the original expectation-based definition of fitness was that expected number of offspring sometimes didn't correspond to probable long-term success. notice that an apparently long-term measure like geometric mean in fact just captures a mathematical fact about a short-term probability distribution over numbers Case 3: Ariew and Lewontin (2004) point out that when generations overlap, probabilities of long-term success can depend on whether the population size is increasing cases fitness cannot be defined by short-term probabilities, but instead must be derived from actual events over a relatively long period of time. ./cache/work_jajnfdt3vzhsla3qd3sexbff5u.pdf ./txt/work_jajnfdt3vzhsla3qd3sexbff5u.txt