id sid tid token lemma pos 37809 1 1 UNIVERSITY UNIVERSITY NNP 37809 1 2 OF of IN 37809 1 3 KANSAS KANSAS NNP 37809 1 4 MUSEUM MUSEUM NNP 37809 1 5 OF of IN 37809 1 6 NATURAL NATURAL NNP 37809 1 7 HISTORY HISTORY NNP 37809 1 8 Vol Vol NNP 37809 1 9 . . . 37809 2 1 20 20 CD 37809 2 2 , , , 37809 2 3 No no UH 37809 2 4 . . . 37809 3 1 1 1 CD 37809 3 2 , , , 37809 3 3 pp pp NNP 37809 3 4 . . . 37809 4 1 1 1 CD 37809 4 2 - - SYM 37809 4 3 45 45 CD 37809 4 4 , , , 37809 4 5 22 22 CD 37809 4 6 figs fig NNS 37809 4 7 . . . 37809 5 1 February February NNP 37809 5 2 20 20 CD 37809 5 3 , , , 37809 5 4 1970 1970 CD 37809 5 5 A a DT 37809 5 6 Taxonomic Taxonomic NNP 37809 5 7 Revision Revision NNP 37809 5 8 of of IN 37809 5 9 the the DT 37809 5 10 Leptodactylid leptodactylid JJ 37809 5 11 Frog Frog NNP 37809 5 12 Genus Genus NNP 37809 5 13 Syrrhophus Syrrhophus NNP 37809 5 14 Cope Cope NNP 37809 5 15 BY by IN 37809 5 16 JOHN JOHN NNP 37809 5 17 D. D. NNP 37809 5 18 LYNCH LYNCH NNP 37809 5 19 UNIVERSITY UNIVERSITY NNP 37809 5 20 OF of IN 37809 5 21 KANSAS KANSAS NNP 37809 5 22 LAWRENCE LAWRENCE NNP 37809 5 23 1970 1970 CD 37809 5 24 UNIVERSITY UNIVERSITY NNP 37809 5 25 OF of IN 37809 5 26 KANSAS KANSAS NNP 37809 5 27 PUBLICATIONS PUBLICATIONS NNP 37809 5 28 , , , 37809 5 29 MUSEUM MUSEUM NNP 37809 5 30 OF of IN 37809 5 31 NATURAL natural JJ 37809 5 32 HISTORY HISTORY NNP 37809 5 33 Editors Editors NNPS 37809 5 34 of of IN 37809 5 35 this this DT 37809 5 36 number number NN 37809 5 37 : : : 37809 5 38 Frank Frank NNP 37809 5 39 B. B. NNP 37809 6 1 Cross Cross NNP 37809 6 2 , , , 37809 6 3 Philip Philip NNP 37809 6 4 S. S. NNP 37809 6 5 Humphrey Humphrey NNP 37809 6 6 , , , 37809 6 7 William William NNP 37809 6 8 E. E. NNP 37809 6 9 Duellman Duellman NNP 37809 6 10 Volume Volume NNP 37809 6 11 20 20 CD 37809 6 12 , , , 37809 6 13 No no UH 37809 6 14 . . . 37809 7 1 1 1 CD 37809 7 2 , , , 37809 7 3 pp pp NNP 37809 7 4 . . . 37809 8 1 1 1 CD 37809 8 2 - - SYM 37809 8 3 45 45 CD 37809 8 4 , , , 37809 8 5 22 22 CD 37809 8 6 figs fig NNS 37809 8 7 . . . 37809 9 1 Published Published NNP 37809 9 2 February February NNP 37809 9 3 20 20 CD 37809 9 4 , , , 37809 9 5 1970 1970 CD 37809 9 6 UNIVERSITY UNIVERSITY NNP 37809 9 7 OF of IN 37809 9 8 KANSAS KANSAS NNP 37809 9 9 Lawrence Lawrence NNP 37809 9 10 , , , 37809 9 11 Kansas Kansas NNP 37809 9 12 PRINTED print VBD 37809 9 13 BY by IN 37809 9 14 THE the DT 37809 9 15 UNIVERSITY UNIVERSITY NNP 37809 9 16 OF of IN 37809 9 17 KANSAS KANSAS NNP 37809 9 18 PRINTING PRINTING NNP 37809 9 19 SERVICE SERVICE NNP 37809 9 20 LAWRENCE LAWRENCE NNP 37809 9 21 , , , 37809 9 22 KANSAS KANSAS NNP 37809 9 23 1970 1970 CD 37809 9 24 A A NNP 37809 9 25 Taxonomic Taxonomic NNP 37809 9 26 Revision Revision NNP 37809 9 27 of of IN 37809 9 28 the the DT 37809 9 29 Leptodactylid leptodactylid JJ 37809 9 30 Frog Frog NNP 37809 9 31 Genus Genus NNP 37809 9 32 Syrrhophus Syrrhophus NNP 37809 9 33 Cope Cope NNP 37809 9 34 BY by IN 37809 9 35 JOHN JOHN NNP 37809 9 36 D. D. NNP 37809 9 37 LYNCH LYNCH NNP 37809 9 38 INTRODUCTION INTRODUCTION NNP 37809 9 39 Cope Cope NNP 37809 9 40 ( ( -LRB- 37809 9 41 1878 1878 CD 37809 9 42 ) ) -RRB- 37809 9 43 proposed propose VBD 37809 9 44 the the DT 37809 9 45 genus genus NNP 37809 9 46 _ _ NNP 37809 9 47 Syrrhophus Syrrhophus NNP 37809 9 48 _ _ NNP 37809 9 49 for for IN 37809 9 50 a a DT 37809 9 51 medium medium JJ 37809 9 52 - - HYPH 37809 9 53 sized sized JJ 37809 9 54 leptodactylid leptodactylid JJ 37809 9 55 frog frog NN 37809 9 56 from from IN 37809 9 57 central central JJ 37809 9 58 Texas Texas NNP 37809 9 59 ; ; : 37809 9 60 in in IN 37809 9 61 the the DT 37809 9 62 ensuing ensue VBG 37809 9 63 75 75 CD 37809 9 64 years year NNS 37809 9 65 the the DT 37809 9 66 genus genus NN 37809 9 67 was be VBD 37809 9 68 expanded expand VBN 37809 9 69 to to TO 37809 9 70 include include VB 37809 9 71 a a DT 37809 9 72 heterogeneous heterogeneous JJ 37809 9 73 group group NN 37809 9 74 of of IN 37809 9 75 frogs frog NNS 37809 9 76 ranging range VBG 37809 9 77 from from IN 37809 9 78 Texas Texas NNP 37809 9 79 to to IN 37809 9 80 Peru Peru NNP 37809 9 81 . . . 37809 10 1 Taylor Taylor NNP 37809 10 2 ( ( -LRB- 37809 10 3 1952 1952 CD 37809 10 4 ) ) -RRB- 37809 10 5 and and CC 37809 10 6 Firschein Firschein NNP 37809 10 7 ( ( -LRB- 37809 10 8 1954 1954 CD 37809 10 9 ) ) -RRB- 37809 10 10 limited limit VBD 37809 10 11 the the DT 37809 10 12 genus genus NN 37809 10 13 to to IN 37809 10 14 several several JJ 37809 10 15 species specie NNS 37809 10 16 of of IN 37809 10 17 frogs frog NNS 37809 10 18 occurring occur VBG 37809 10 19 in in IN 37809 10 20 Guatemala Guatemala NNP 37809 10 21 , , , 37809 10 22 México México NNP 37809 10 23 , , , 37809 10 24 and and CC 37809 10 25 Texas Texas NNP 37809 10 26 . . . 37809 11 1 Lynch Lynch NNP 37809 11 2 ( ( -LRB- 37809 11 3 1968 1968 CD 37809 11 4 ) ) -RRB- 37809 11 5 provided provide VBD 37809 11 6 a a DT 37809 11 7 definition definition NN 37809 11 8 of of IN 37809 11 9 the the DT 37809 11 10 previously previously RB 37809 11 11 loosely loosely RB 37809 11 12 - - HYPH 37809 11 13 defined define VBN 37809 11 14 genus genus NN 37809 11 15 . . . 37809 12 1 With with IN 37809 12 2 the the DT 37809 12 3 exception exception NN 37809 12 4 of of IN 37809 12 5 Taylor Taylor NNP 37809 12 6 ( ( -LRB- 37809 12 7 1952 1952 CD 37809 12 8 ) ) -RRB- 37809 12 9 , , , 37809 12 10 who who WP 37809 12 11 treated treat VBD 37809 12 12 the the DT 37809 12 13 Costa costa JJ 37809 12 14 Rican rican JJ 37809 12 15 species specie NNS 37809 12 16 , , , 37809 12 17 none none NN 37809 12 18 of of IN 37809 12 19 these these DT 37809 12 20 authors author NNS 37809 12 21 dealt deal VBD 37809 12 22 with with IN 37809 12 23 the the DT 37809 12 24 present present JJ 37809 12 25 status status NN 37809 12 26 of of IN 37809 12 27 the the DT 37809 12 28 nineteen nineteen CD 37809 12 29 species specie NNS 37809 12 30 erroneously erroneously RB 37809 12 31 assigned assign VBN 37809 12 32 to to IN 37809 12 33 _ _ NNP 37809 12 34 Syrrhophus Syrrhophus NNP 37809 12 35 _ _ NNP 37809 12 36 . . . 37809 13 1 These these DT 37809 13 2 species specie NNS 37809 13 3 are be VBP 37809 13 4 listed list VBN 37809 13 5 in in IN 37809 13 6 Tables Tables NNP 37809 13 7 1 1 CD 37809 13 8 and and CC 37809 13 9 2 2 CD 37809 13 10 with with IN 37809 13 11 the the DT 37809 13 12 name name NN 37809 13 13 currently currently RB 37809 13 14 applied apply VBD 37809 13 15 . . . 37809 14 1 Some some DT 37809 14 2 of of IN 37809 14 3 them -PRON- PRP 37809 14 4 are be VBP 37809 14 5 new new JJ 37809 14 6 combinations combination NNS 37809 14 7 and and CC 37809 14 8 their -PRON- PRP$ 37809 14 9 justifications justification NNS 37809 14 10 will will MD 37809 14 11 be be VB 37809 14 12 published publish VBN 37809 14 13 elsewhere elsewhere RB 37809 14 14 . . . 37809 15 1 Gorham Gorham NNP 37809 15 2 ( ( -LRB- 37809 15 3 1966 1966 CD 37809 15 4 ) ) -RRB- 37809 15 5 is be VBZ 37809 15 6 the the DT 37809 15 7 most most RBS 37809 15 8 recent recent JJ 37809 15 9 author author NN 37809 15 10 to to TO 37809 15 11 include include VB 37809 15 12 South south JJ 37809 15 13 American american JJ 37809 15 14 species specie NNS 37809 15 15 in in IN 37809 15 16 the the DT 37809 15 17 genus genus NN 37809 15 18 _ _ NNP 37809 15 19 Syrrhophus Syrrhophus NNP 37809 15 20 _ _ NNP 37809 15 21 . . . 37809 16 1 Smith Smith NNP 37809 16 2 and and CC 37809 16 3 Taylor Taylor NNP 37809 16 4 ( ( -LRB- 37809 16 5 1948 1948 CD 37809 16 6 ) ) -RRB- 37809 16 7 recognized recognize VBD 37809 16 8 two two CD 37809 16 9 species specie NNS 37809 16 10 groups group NNS 37809 16 11 of of IN 37809 16 12 the the DT 37809 16 13 genus genus NN 37809 16 14 in in IN 37809 16 15 México México NNP 37809 16 16 , , , 37809 16 17 an an DT 37809 16 18 eastern eastern JJ 37809 16 19 and and CC 37809 16 20 a a DT 37809 16 21 western western JJ 37809 16 22 group group NN 37809 16 23 ( ( -LRB- 37809 16 24 here here RB 37809 16 25 termed term VBN 37809 16 26 complexes complex NNS 37809 16 27 for for IN 37809 16 28 purposes purpose NNS 37809 16 29 of of IN 37809 16 30 discussion discussion NN 37809 16 31 ) ) -RRB- 37809 16 32 , , , 37809 16 33 separated separate VBN 37809 16 34 on on IN 37809 16 35 the the DT 37809 16 36 basis basis NN 37809 16 37 of of IN 37809 16 38 the the DT 37809 16 39 number number NN 37809 16 40 of of IN 37809 16 41 palmar palmar NNP 37809 16 42 ( ( -LRB- 37809 16 43 metacarpal metacarpal NNP 37809 16 44 ) ) -RRB- 37809 16 45 tubercles tubercle NNS 37809 16 46 ( ( -LRB- 37809 16 47 three three CD 37809 16 48 palmar palmar NNP 37809 16 49 tubercles tubercle NNS 37809 16 50 in in IN 37809 16 51 the the DT 37809 16 52 members member NNS 37809 16 53 of of IN 37809 16 54 the the DT 37809 16 55 eastern eastern JJ 37809 16 56 complex complex NN 37809 16 57 and and CC 37809 16 58 two two CD 37809 16 59 in in IN 37809 16 60 those those DT 37809 16 61 of of IN 37809 16 62 the the DT 37809 16 63 western western JJ 37809 16 64 complex complex NN 37809 16 65 ) ) -RRB- 37809 16 66 . . . 37809 17 1 Duellman Duellman NNP 37809 17 2 ( ( -LRB- 37809 17 3 1958 1958 CD 37809 17 4 ) ) -RRB- 37809 17 5 reviewed review VBD 37809 17 6 the the DT 37809 17 7 species specie NNS 37809 17 8 of of IN 37809 17 9 the the DT 37809 17 10 genus genus NN 37809 17 11 occurring occur VBG 37809 17 12 in in IN 37809 17 13 western western JJ 37809 17 14 México México NNP 37809 17 15 and and CC 37809 17 16 concluded conclude VBD 37809 17 17 that that IN 37809 17 18 there there EX 37809 17 19 were be VBD 37809 17 20 five five CD 37809 17 21 species specie NNS 37809 17 22 ( ( -LRB- 37809 17 23 two two CD 37809 17 24 polytypic polytypic NN 37809 17 25 ) ) -RRB- 37809 17 26 . . . 37809 18 1 Dixon Dixon NNP 37809 18 2 and and CC 37809 18 3 Webb Webb NNP 37809 18 4 ( ( -LRB- 37809 18 5 1966 1966 CD 37809 18 6 ) ) -RRB- 37809 18 7 described describe VBD 37809 18 8 an an DT 37809 18 9 additional additional JJ 37809 18 10 species species NN 37809 18 11 from from IN 37809 18 12 Jalisco Jalisco NNP 37809 18 13 , , , 37809 18 14 México México NNP 37809 18 15 . . . 37809 19 1 The the DT 37809 19 2 distributions distribution NNS 37809 19 3 of of IN 37809 19 4 some some DT 37809 19 5 species specie NNS 37809 19 6 have have VBP 37809 19 7 been be VBN 37809 19 8 extended extend VBN 37809 19 9 , , , 37809 19 10 but but CC 37809 19 11 otherwise otherwise RB 37809 19 12 the the DT 37809 19 13 western western JJ 37809 19 14 complex complex NN 37809 19 15 of of IN 37809 19 16 species specie NNS 37809 19 17 remains remain VBZ 37809 19 18 unchanged unchanged JJ 37809 19 19 since since IN 37809 19 20 Duellman Duellman NNP 37809 19 21 's 's POS 37809 19 22 review review NN 37809 19 23 . . . 37809 20 1 Smith Smith NNP 37809 20 2 and and CC 37809 20 3 Taylor Taylor NNP 37809 20 4 ( ( -LRB- 37809 20 5 1948 1948 CD 37809 20 6 ) ) -RRB- 37809 20 7 recognized recognize VBD 37809 20 8 seven seven CD 37809 20 9 species specie NNS 37809 20 10 of of IN 37809 20 11 the the DT 37809 20 12 genus genus NN 37809 20 13 in in IN 37809 20 14 eastern eastern JJ 37809 20 15 México México NNP 37809 20 16 . . . 37809 21 1 Firschein Firschein NNP 37809 21 2 revised revise VBD 37809 21 3 the the DT 37809 21 4 eastern eastern JJ 37809 21 5 complex complex NN 37809 21 6 ( ( -LRB- 37809 21 7 as as IN 37809 21 8 then then RB 37809 21 9 understood understand VBN 37809 21 10 ) ) -RRB- 37809 21 11 , , , 37809 21 12 and and CC 37809 21 13 in in IN 37809 21 14 so so RB 37809 21 15 doing do VBG 37809 21 16 added add VBN 37809 21 17 one one CD 37809 21 18 new new JJ 37809 21 19 species specie NNS 37809 21 20 and and CC 37809 21 21 treated treat VBN 37809 21 22 _ _ NNP 37809 21 23 Syrrhophus Syrrhophus NNP 37809 21 24 verruculatus verruculatus NNP 37809 21 25 _ _ NNP 37809 21 26 as as IN 37809 21 27 a a DT 37809 21 28 _ _ NNP 37809 21 29 nomen nomen NNP 37809 21 30 dubium dubium NN 37809 21 31 _ _ NNP 37809 21 32 . . . 37809 22 1 Dixon Dixon NNP 37809 22 2 ( ( -LRB- 37809 22 3 1957 1957 CD 37809 22 4 ) ) -RRB- 37809 22 5 redefined redefine VBD 37809 22 6 the the DT 37809 22 7 related related JJ 37809 22 8 genus genus NN 37809 22 9 _ _ NNP 37809 22 10 Tomodactylus Tomodactylus NNP 37809 22 11 _ _ NNP 37809 22 12 and and CC 37809 22 13 transferred transfer VBN 37809 22 14 _ _ NNP 37809 22 15 T. T. NNP 37809 23 1 macrotympanum macrotympanum NNP 37809 23 2 _ _ NNP 37809 23 3 Taylor Taylor NNP 37809 23 4 to to IN 37809 23 5 the the DT 37809 23 6 genus genus JJ 37809 23 7 _ _ NNP 37809 23 8 Syrrhophus Syrrhophus NNP 37809 23 9 _ _ NNP 37809 23 10 . . . 37809 24 1 Neill Neill NNP 37809 24 2 ( ( -LRB- 37809 24 3 1965 1965 CD 37809 24 4 ) ) -RRB- 37809 24 5 described describe VBD 37809 24 6 a a DT 37809 24 7 new new JJ 37809 24 8 subspecies subspecie NNS 37809 24 9 of of IN 37809 24 10 _ _ NNP 37809 24 11 S. S. NNP 37809 25 1 leprus leprus NNP 37809 25 2 _ _ NNP 37809 25 3 from from IN 37809 25 4 British British NNP 37809 25 5 Honduras hondura NNS 37809 25 6 . . . 37809 26 1 Two two CD 37809 26 2 species specie NNS 37809 26 3 ( ( -LRB- 37809 26 4 _ _ NNP 37809 26 5 S. S. NNP 37809 27 1 gaigeae gaigeae NNP 37809 27 2 _ _ NNP 37809 27 3 and and CC 37809 27 4 _ _ NNP 37809 27 5 S. S. NNP 37809 28 1 marnockii marnockii NNP 37809 28 2 _ _ NNP 37809 28 3 ) ) -RRB- 37809 28 4 were be VBD 37809 28 5 recognized recognize VBN 37809 28 6 in in IN 37809 28 7 Texas Texas NNP 37809 28 8 until until IN 37809 28 9 Milstead Milstead NNP 37809 28 10 , , , 37809 28 11 Mecham Mecham NNP 37809 28 12 , , , 37809 28 13 and and CC 37809 28 14 McClintock McClintock NNP 37809 28 15 ( ( -LRB- 37809 28 16 1950 1950 CD 37809 28 17 ) ) -RRB- 37809 28 18 synonymized synonymize VBD 37809 28 19 _ _ NNP 37809 28 20 S. S. NNP 37809 29 1 gaigeae gaigeae NNP 37809 29 2 _ _ NNP 37809 29 3 with with IN 37809 29 4 _ _ NNP 37809 29 5 S. S. NNP 37809 30 1 marnockii marnockii NNP 37809 30 2 _ _ NNP 37809 30 3 . . . 37809 31 1 Thus thus RB 37809 31 2 , , , 37809 31 3 at at IN 37809 31 4 present present JJ 37809 31 5 , , , 37809 31 6 nine nine CD 37809 31 7 species specie NNS 37809 31 8 ( ( -LRB- 37809 31 9 one one CD 37809 31 10 polytypic polytypic NN 37809 31 11 ) ) -RRB- 37809 31 12 are be VBP 37809 31 13 recognized recognize VBN 37809 31 14 on on IN 37809 31 15 the the DT 37809 31 16 eastern eastern JJ 37809 31 17 slopes slope NNS 37809 31 18 and and CC 37809 31 19 lowlands lowland NNS 37809 31 20 from from IN 37809 31 21 central central JJ 37809 31 22 Texas Texas NNP 37809 31 23 to to IN 37809 31 24 British british JJ 37809 31 25 Honduras Honduras NNP 37809 31 26 . . . 37809 32 1 These these DT 37809 32 2 are be VBP 37809 32 3 currently currently RB 37809 32 4 placed place VBN 37809 32 5 on on IN 37809 32 6 one one CD 37809 32 7 species species NN 37809 32 8 group group NN 37809 32 9 equivalent equivalent JJ 37809 32 10 to to IN 37809 32 11 the the DT 37809 32 12 western western JJ 37809 32 13 complex complex NN 37809 32 14 reviewed review VBN 37809 32 15 by by IN 37809 32 16 Duellman Duellman NNP 37809 32 17 ( ( -LRB- 37809 32 18 1958 1958 CD 37809 32 19 ) ) -RRB- 37809 32 20 . . . 37809 33 1 TABLE TABLE NNP 37809 33 2 1 1 CD 37809 33 3 - - HYPH 37809 33 4 -Species -specie NNS 37809 33 5 Described describe VBN 37809 33 6 as as IN 37809 33 7 Members member NNS 37809 33 8 of of IN 37809 33 9 the the DT 37809 33 10 Genus Genus NNP 37809 33 11 _ _ NNP 37809 33 12 Syrrhophus Syrrhophus NNP 37809 33 13 _ _ NNP 37809 33 14 but but CC 37809 33 15 Now now RB 37809 33 16 Placed place VBN 37809 33 17 in in IN 37809 33 18 Other other JJ 37809 33 19 Genera Genera NNP 37809 33 20 . . . 37809 34 1 = = NFP 37809 34 2 = = NFP 37809 34 3 = = SYM 37809 34 4 = = SYM 37809 34 5 = = SYM 37809 34 6 = = SYM 37809 34 7 = = SYM 37809 34 8 = = SYM 37809 34 9 = = SYM 37809 34 10 = = SYM 37809 34 11 = = SYM 37809 34 12 = = SYM 37809 34 13 = = SYM 37809 34 14 = = SYM 37809 34 15 = = SYM 37809 34 16 = = SYM 37809 34 17 = = SYM 37809 34 18 = = SYM 37809 34 19 = = SYM 37809 34 20 = = SYM 37809 34 21 = = SYM 37809 34 22 = = SYM 37809 34 23 = = SYM 37809 34 24 = = SYM 37809 34 25 = = SYM 37809 34 26 = = SYM 37809 34 27 = = SYM 37809 34 28 = = SYM 37809 34 29 = = SYM 37809 34 30 = = SYM 37809 34 31 = = SYM 37809 34 32 = = SYM 37809 34 33 = = SYM 37809 34 34 = = SYM 37809 34 35 = = SYM 37809 34 36 = = SYM 37809 34 37 = = SYM 37809 34 38 = = SYM 37809 34 39 = = SYM 37809 34 40 = = SYM 37809 34 41 = = SYM 37809 34 42 = = SYM 37809 34 43 = = SYM 37809 34 44 = = SYM 37809 34 45 = = SYM 37809 34 46 = = SYM 37809 34 47 = = SYM 37809 34 48 = = SYM 37809 34 49 = = SYM 37809 34 50 = = SYM 37809 34 51 = = SYM 37809 34 52 = = SYM 37809 34 53 = = SYM 37809 34 54 = = SYM 37809 34 55 = = SYM 37809 34 56 = = SYM 37809 34 57 = = SYM 37809 34 58 = = SYM 37809 34 59 = = SYM 37809 34 60 = = SYM 37809 34 61 = = SYM 37809 34 62 = = SYM 37809 34 63 = = SYM 37809 34 64 = = SYM 37809 34 65 = = SYM 37809 34 66 = = SYM 37809 34 67 = = SYM 37809 34 68 = = SYM 37809 34 69 = = SYM 37809 34 70 = = NFP 37809 34 71 = = NFP 37809 34 72 Trivial trivial JJ 37809 34 73 name name NN 37809 34 74 and and CC 37809 34 75 author author VB 37809 34 76 Current current JJ 37809 34 77 combination combination NN 37809 34 78 ----------------------------------------------------------------------- ----------------------------------------------------------------------- : 37809 34 79 _ _ NNP 37809 34 80 areolatus areolatus NN 37809 34 81 _ _ NNP 37809 34 82 Boulenger Boulenger NNP 37809 34 83 , , , 37809 34 84 1898 1898 CD 37809 34 85 _ _ NNP 37809 34 86 Eleutherodactylus Eleutherodactylus NNP 37809 34 87 areolatus areolatus NN 37809 34 88 _ _ NNP 37809 34 89 _ _ NNP 37809 34 90 calcaratus calcaratus NN 37809 34 91 _ _ NNP 37809 34 92 Andersson Andersson NNP 37809 34 93 , , , 37809 34 94 1945 1945 CD 37809 34 95 _ _ NNP 37809 34 96 Eleutherodactylus Eleutherodactylus NNP 37809 34 97 anderssoni anderssoni NN 37809 34 98 _ _ NNP 37809 34 99 _ _ NNP 37809 34 100 caryophyllaceus caryophyllaceus NNP 37809 34 101 _ _ NNP 37809 34 102 Barbour Barbour NNP 37809 34 103 , , , 37809 34 104 1928 1928 CD 37809 34 105 _ _ NNP 37809 34 106 Eleutherodactylus Eleutherodactylus NNP 37809 34 107 caryophyllaceus caryophyllaceu NNS 37809 34 108 _ _ NNP 37809 34 109 _ _ NNP 37809 34 110 coeruleus coeruleus NN 37809 34 111 _ _ NNP 37809 34 112 Andersson Andersson NNP 37809 34 113 , , , 37809 34 114 1945 1945 CD 37809 34 115 _ _ NNP 37809 34 116 Eleutherodactylus Eleutherodactylus NNP 37809 34 117 coeruleus coeruleus NN 37809 34 118 _ _ NNP 37809 34 119 _ _ NNP 37809 34 120 ineptus ineptus NN 37809 34 121 _ _ NNP 37809 34 122 Barbour Barbour NNP 37809 34 123 , , , 37809 34 124 1928 1928 CD 37809 34 125 _ _ NNP 37809 34 126 Eleutherodactylus Eleutherodactylus NNP 37809 34 127 diastema diastema NN 37809 34 128 _ _ NNP 37809 34 129 _ _ NNP 37809 34 130 juninensis juninensis NN 37809 34 131 _ _ NNP 37809 34 132 Shreve Shreve NNP 37809 34 133 , , , 37809 34 134 1938 1938 CD 37809 34 135 _ _ NNP 37809 34 136 Eupsophus Eupsophus NNP 37809 34 137 juninensis juninensis NN 37809 34 138 _ _ NNP 37809 34 139 _ _ NNP 37809 34 140 lutosus lutosus NN 37809 34 141 _ _ NNP 37809 34 142 Barbour Barbour NNP 37809 34 143 and and CC 37809 34 144 Dunn Dunn NNP 37809 34 145 , , , 37809 34 146 1921 1921 CD 37809 34 147 _ _ NNP 37809 34 148 Eleutherodactylus Eleutherodactylus NNP 37809 34 149 lutosus lutosus NN 37809 34 150 _ _ NNP 37809 34 151 _ _ NNP 37809 34 152 molinoi molinoi NN 37809 34 153 _ _ NNP 37809 34 154 Barbour Barbour NNP 37809 34 155 , , , 37809 34 156 1928 1928 CD 37809 34 157 _ _ NNP 37809 34 158 Eleutherodactylus Eleutherodactylus NNP 37809 34 159 molinoi molinoi NN 37809 34 160 _ _ NNP 37809 34 161 _ _ NNP 37809 34 162 montium montium NN 37809 34 163 _ _ NNP 37809 34 164 Shreve Shreve NNP 37809 34 165 , , , 37809 34 166 1938 1938 CD 37809 34 167 _ _ NNP 37809 34 168 Niceforonia Niceforonia NNP 37809 34 169 montia montia NN 37809 34 170 _ _ NNP 37809 34 171 _ _ NNP 37809 34 172 mystaceus mystaceus NN 37809 34 173 _ _ NNP 37809 34 174 Barbour Barbour NNP 37809 34 175 , , , 37809 34 176 1922 1922 CD 37809 34 177 _ _ NNP 37809 34 178 Eleutherodactylus Eleutherodactylus NNP 37809 34 179 rhodopis rhodopis NN 37809 34 180 _ _ NNP 37809 34 181 _ _ NNP 37809 34 182 obesus obesus NN 37809 34 183 _ _ NNP 37809 34 184 Barbour Barbour NNP 37809 34 185 , , , 37809 34 186 1928 1928 CD 37809 34 187 _ _ NNP 37809 34 188 Eleutherodactylus Eleutherodactylus NNP 37809 34 189 punctariolus punctariolus NN 37809 34 190 _ _ NNP 37809 34 191 _ _ NNP 37809 34 192 omiltemanus omiltemanus NN 37809 34 193 _ _ NNP 37809 34 194 Gunther Gunther NNP 37809 34 195 , , , 37809 34 196 1900 1900 CD 37809 34 197 _ _ NNP 37809 34 198 Eleutherodactylus Eleutherodactylus NNP 37809 34 199 omiltemanus_[1 omiltemanus_[1 NNP 37809 34 200 ] ] -RRB- 37809 34 201 _ _ NNP 37809 34 202 pardalis pardali NNS 37809 34 203 _ _ NNP 37809 34 204 Barbour Barbour NNP 37809 34 205 , , , 37809 34 206 1928 1928 CD 37809 34 207 _ _ NNP 37809 34 208 Eleutherodactylus Eleutherodactylus NNP 37809 34 209 pardalis pardali NNS 37809 34 210 _ _ NNP 37809 34 211 = = SYM 37809 34 212 = = SYM 37809 34 213 = = SYM 37809 34 214 = = SYM 37809 34 215 = = SYM 37809 34 216 = = SYM 37809 34 217 = = SYM 37809 34 218 = = SYM 37809 34 219 = = SYM 37809 34 220 = = SYM 37809 34 221 = = SYM 37809 34 222 = = SYM 37809 34 223 = = SYM 37809 34 224 = = SYM 37809 34 225 = = SYM 37809 34 226 = = SYM 37809 34 227 = = SYM 37809 34 228 = = SYM 37809 34 229 = = SYM 37809 34 230 = = SYM 37809 34 231 = = SYM 37809 34 232 = = SYM 37809 34 233 = = SYM 37809 34 234 = = SYM 37809 34 235 = = SYM 37809 34 236 = = SYM 37809 34 237 = = SYM 37809 34 238 = = SYM 37809 34 239 = = SYM 37809 34 240 = = SYM 37809 34 241 = = SYM 37809 34 242 = = SYM 37809 34 243 = = SYM 37809 34 244 = = SYM 37809 34 245 = = SYM 37809 34 246 = = SYM 37809 34 247 = = SYM 37809 34 248 = = SYM 37809 34 249 = = SYM 37809 34 250 = = SYM 37809 34 251 = = SYM 37809 34 252 = = SYM 37809 34 253 = = SYM 37809 34 254 = = SYM 37809 34 255 = = SYM 37809 34 256 = = SYM 37809 34 257 = = SYM 37809 34 258 = = SYM 37809 34 259 = = SYM 37809 34 260 = = SYM 37809 34 261 = = SYM 37809 34 262 = = SYM 37809 34 263 = = SYM 37809 34 264 = = SYM 37809 34 265 = = SYM 37809 34 266 = = SYM 37809 34 267 = = SYM 37809 34 268 = = SYM 37809 34 269 = = SYM 37809 34 270 = = SYM 37809 34 271 = = SYM 37809 34 272 = = SYM 37809 34 273 = = SYM 37809 34 274 = = SYM 37809 34 275 = = SYM 37809 34 276 = = SYM 37809 34 277 = = SYM 37809 34 278 = = SYM 37809 34 279 = = NFP 37809 34 280 = = NFP 37809 34 281 = = NFP 37809 34 282 [ [ -LRB- 37809 34 283 1 1 CD 37809 34 284 ] ] -RRB- 37809 34 285 New new JJ 37809 34 286 combination combination NN 37809 34 287 . . . 37809 35 1 TABLE TABLE NNP 37809 35 2 2 2 CD 37809 35 3 - - : 37809 35 4 -Species -species NN 37809 35 5 Incorrectly incorrectly RB 37809 35 6 Regarded regard VBN 37809 35 7 as as IN 37809 35 8 Members member NNS 37809 35 9 of of IN 37809 35 10 the the DT 37809 35 11 Genus Genus NNP 37809 35 12 _ _ NNP 37809 35 13 Syrrhophus Syrrhophus NNP 37809 35 14 _ _ NNP 37809 35 15 but but CC 37809 35 16 Described describe VBN 37809 35 17 as as IN 37809 35 18 Members member NNS 37809 35 19 of of IN 37809 35 20 Other Other NNP 37809 35 21 Genera Genera NNP 37809 35 22 . . . 37809 36 1 = = NFP 37809 36 2 = = NFP 37809 36 3 = = SYM 37809 36 4 = = SYM 37809 36 5 = = SYM 37809 36 6 = = SYM 37809 36 7 = = SYM 37809 36 8 = = SYM 37809 36 9 = = SYM 37809 36 10 = = SYM 37809 36 11 = = SYM 37809 36 12 = = SYM 37809 36 13 = = SYM 37809 36 14 = = SYM 37809 36 15 = = SYM 37809 36 16 = = SYM 37809 36 17 = = SYM 37809 36 18 = = SYM 37809 36 19 = = SYM 37809 36 20 = = SYM 37809 36 21 = = SYM 37809 36 22 = = SYM 37809 36 23 = = SYM 37809 36 24 = = SYM 37809 36 25 = = SYM 37809 36 26 = = SYM 37809 36 27 = = SYM 37809 36 28 = = SYM 37809 36 29 = = SYM 37809 36 30 = = SYM 37809 36 31 = = SYM 37809 36 32 = = SYM 37809 36 33 = = SYM 37809 36 34 = = SYM 37809 36 35 = = SYM 37809 36 36 = = SYM 37809 36 37 = = SYM 37809 36 38 = = SYM 37809 36 39 = = SYM 37809 36 40 = = SYM 37809 36 41 = = SYM 37809 36 42 = = SYM 37809 36 43 = = SYM 37809 36 44 = = SYM 37809 36 45 = = SYM 37809 36 46 = = SYM 37809 36 47 = = SYM 37809 36 48 = = SYM 37809 36 49 = = SYM 37809 36 50 = = SYM 37809 36 51 = = SYM 37809 36 52 = = SYM 37809 36 53 = = SYM 37809 36 54 = = SYM 37809 36 55 = = SYM 37809 36 56 = = SYM 37809 36 57 = = SYM 37809 36 58 = = SYM 37809 36 59 = = SYM 37809 36 60 = = SYM 37809 36 61 = = SYM 37809 36 62 = = SYM 37809 36 63 = = SYM 37809 36 64 = = SYM 37809 36 65 = = SYM 37809 36 66 = = SYM 37809 36 67 = = SYM 37809 36 68 = = SYM 37809 36 69 = = SYM 37809 36 70 = = SYM 37809 36 71 = = SYM 37809 36 72 = = SYM 37809 36 73 = = NFP 37809 36 74 = = NFP 37809 36 75 Trivial trivial JJ 37809 36 76 name name NN 37809 36 77 , , , 37809 36 78 original original JJ 37809 36 79 generic generic JJ 37809 36 80 assignment assignment NN 37809 36 81 , , , 37809 36 82 and and CC 37809 36 83 author author VB 37809 36 84 Current current JJ 37809 36 85 combination combination NN 37809 36 86 -------------------------------------------------------------------------- -------------------------------------------------------------------------- : 37809 36 87 _ _ NNP 37809 36 88 chalceus chalceus NNP 37809 36 89 _ _ NNP 37809 36 90 ( ( -LRB- 37809 36 91 _ _ NNP 37809 36 92 Phyllobates Phyllobates NNPS 37809 36 93 _ _ NNP 37809 36 94 ) ) -RRB- 37809 36 95 Peters Peters NNP 37809 36 96 , , , 37809 36 97 1873 1873 CD 37809 36 98 _ _ NNP 37809 36 99 Eleutherodactylus Eleutherodactylus NNP 37809 36 100 chalceus chalceus NN 37809 36 101 _ _ NNP 37809 36 102 _ _ NNP 37809 36 103 festae festae VBD 37809 36 104 _ _ NNP 37809 36 105 ( ( -LRB- 37809 36 106 _ _ NNP 37809 36 107 Paludicola Paludicola NNP 37809 36 108 _ _ NNP 37809 36 109 ) ) -RRB- 37809 36 110 Peracca Peracca NNP 37809 36 111 , , , 37809 36 112 1904 1904 CD 37809 36 113 _ _ NNP 37809 36 114 Niceforonia Niceforonia NNP 37809 36 115 festae festae VBD 37809 36 116 _ _ NNP 37809 36 117 _ _ NNP 37809 36 118 hylaeformis hylaeformis NNP 37809 36 119 _ _ NNP 37809 36 120 ( ( -LRB- 37809 36 121 _ _ NNP 37809 36 122 Phyllobates Phyllobates NNPS 37809 36 123 _ _ NNP 37809 36 124 ) ) -RRB- 37809 36 125 Cope Cope NNP 37809 36 126 , , , 37809 36 127 1875 1875 CD 37809 36 128 _ _ NNP 37809 36 129 Eleutherodactylus Eleutherodactylus NNP 37809 36 130 hylaeformis hylaeformis NN 37809 36 131 _ _ NNP 37809 36 132 _ _ NNP 37809 36 133 palmatus palmatus NN 37809 36 134 _ _ NNP 37809 36 135 ( ( -LRB- 37809 36 136 _ _ NNP 37809 36 137 Phyllobates Phyllobates NNPS 37809 36 138 _ _ NNP 37809 36 139 ) ) -RRB- 37809 36 140 Werner Werner NNP 37809 36 141 , , , 37809 36 142 1899 1899 CD 37809 36 143 _ _ NNP 37809 36 144 Colostethus Colostethus NNP 37809 36 145 palmatus palmatus NN 37809 36 146 _ _ NNP 37809 36 147 _ _ NNP 37809 36 148 ridens ridens NNP 37809 36 149 _ _ NNP 37809 36 150 ( ( -LRB- 37809 36 151 _ _ NNP 37809 36 152 Phyllobates Phyllobates NNPS 37809 36 153 _ _ NNP 37809 36 154 ) ) -RRB- 37809 36 155 Cope Cope NNP 37809 36 156 , , , 37809 36 157 1866 1866 CD 37809 36 158 _ _ NNP 37809 36 159 Eleutherodactylus Eleutherodactylus NNP 37809 36 160 ridens riden NNS 37809 36 161 _ _ NNP 37809 36 162 _ _ NNP 37809 36 163 simonsii simonsii NNS 37809 36 164 _ _ NNP 37809 36 165 ( ( -LRB- 37809 36 166 _ _ NNP 37809 36 167 Paludicola Paludicola NNP 37809 36 168 _ _ NNP 37809 36 169 ) ) -RRB- 37809 36 170 Boulenger Boulenger NNP 37809 36 171 , , , 37809 36 172 1900 1900 CD 37809 36 173 _ _ NNP 37809 36 174 Niceforonia Niceforonia NNP 37809 36 175 simonsii simonsii VBZ 37809 36 176 _ _ NNP 37809 36 177 = = SYM 37809 36 178 = = SYM 37809 36 179 = = SYM 37809 36 180 = = SYM 37809 36 181 = = SYM 37809 36 182 = = SYM 37809 36 183 = = SYM 37809 36 184 = = SYM 37809 36 185 = = SYM 37809 36 186 = = SYM 37809 36 187 = = SYM 37809 36 188 = = SYM 37809 36 189 = = SYM 37809 36 190 = = SYM 37809 36 191 = = SYM 37809 36 192 = = SYM 37809 36 193 = = SYM 37809 36 194 = = SYM 37809 36 195 = = SYM 37809 36 196 = = SYM 37809 36 197 = = SYM 37809 36 198 = = SYM 37809 36 199 = = SYM 37809 36 200 = = SYM 37809 36 201 = = SYM 37809 36 202 = = SYM 37809 36 203 = = SYM 37809 36 204 = = SYM 37809 36 205 = = SYM 37809 36 206 = = SYM 37809 36 207 = = SYM 37809 36 208 = = SYM 37809 36 209 = = SYM 37809 36 210 = = SYM 37809 36 211 = = SYM 37809 36 212 = = SYM 37809 36 213 = = SYM 37809 36 214 = = SYM 37809 36 215 = = SYM 37809 36 216 = = SYM 37809 36 217 = = SYM 37809 36 218 = = SYM 37809 36 219 = = SYM 37809 36 220 = = SYM 37809 36 221 = = SYM 37809 36 222 = = SYM 37809 36 223 = = SYM 37809 36 224 = = SYM 37809 36 225 = = SYM 37809 36 226 = = SYM 37809 36 227 = = SYM 37809 36 228 = = SYM 37809 36 229 = = SYM 37809 36 230 = = SYM 37809 36 231 = = SYM 37809 36 232 = = SYM 37809 36 233 = = SYM 37809 36 234 = = SYM 37809 36 235 = = SYM 37809 36 236 = = SYM 37809 36 237 = = SYM 37809 36 238 = = SYM 37809 36 239 = = SYM 37809 36 240 = = SYM 37809 36 241 = = SYM 37809 36 242 = = SYM 37809 36 243 = = SYM 37809 36 244 = = SYM 37809 36 245 = = SYM 37809 36 246 = = SYM 37809 36 247 = = SYM 37809 36 248 = = SYM 37809 36 249 = = NFP 37809 36 250 = = NFP 37809 36 251 TABLE TABLE NNP 37809 36 252 3 3 CD 37809 36 253 - - HYPH 37809 36 254 -Nominal -Nominal NNP 37809 36 255 Species Species NNPS 37809 36 256 of of IN 37809 36 257 _ _ NNP 37809 36 258 Syrrhophus Syrrhophus NNP 37809 36 259 _ _ NNP 37809 36 260 ( ( -LRB- 37809 36 261 _ _ NNP 37809 36 262 sensu sensu NN 37809 36 263 strictu strictu NNP 37809 36 264 _ _ NNP 37809 36 265 ) ) -RRB- 37809 36 266 and and CC 37809 36 267 the the DT 37809 36 268 Name Name NNP 37809 36 269 Used use VBD 37809 36 270 Herein Herein NNP 37809 36 271 . . . 37809 37 1 = = NFP 37809 37 2 = = NFP 37809 37 3 = = SYM 37809 37 4 = = SYM 37809 37 5 = = SYM 37809 37 6 = = SYM 37809 37 7 = = SYM 37809 37 8 = = SYM 37809 37 9 = = SYM 37809 37 10 = = SYM 37809 37 11 = = SYM 37809 37 12 = = SYM 37809 37 13 = = SYM 37809 37 14 = = SYM 37809 37 15 = = SYM 37809 37 16 = = SYM 37809 37 17 = = SYM 37809 37 18 = = SYM 37809 37 19 = = SYM 37809 37 20 = = SYM 37809 37 21 = = SYM 37809 37 22 = = SYM 37809 37 23 = = SYM 37809 37 24 = = SYM 37809 37 25 = = SYM 37809 37 26 = = SYM 37809 37 27 = = SYM 37809 37 28 = = SYM 37809 37 29 = = SYM 37809 37 30 = = SYM 37809 37 31 = = SYM 37809 37 32 = = SYM 37809 37 33 = = SYM 37809 37 34 = = SYM 37809 37 35 = = SYM 37809 37 36 = = SYM 37809 37 37 = = SYM 37809 37 38 = = SYM 37809 37 39 = = SYM 37809 37 40 = = SYM 37809 37 41 = = SYM 37809 37 42 = = SYM 37809 37 43 = = SYM 37809 37 44 = = SYM 37809 37 45 = = SYM 37809 37 46 = = SYM 37809 37 47 = = SYM 37809 37 48 = = SYM 37809 37 49 = = SYM 37809 37 50 = = SYM 37809 37 51 = = SYM 37809 37 52 = = SYM 37809 37 53 = = SYM 37809 37 54 = = SYM 37809 37 55 = = SYM 37809 37 56 = = SYM 37809 37 57 = = SYM 37809 37 58 = = SYM 37809 37 59 = = SYM 37809 37 60 = = SYM 37809 37 61 = = SYM 37809 37 62 = = SYM 37809 37 63 = = SYM 37809 37 64 = = SYM 37809 37 65 = = SYM 37809 37 66 = = SYM 37809 37 67 = = SYM 37809 37 68 = = SYM 37809 37 69 = = SYM 37809 37 70 = = NFP 37809 37 71 = = SYM 37809 37 72 Original original JJ 37809 37 73 combination combination NN 37809 37 74 Current current JJ 37809 37 75 combination combination NN 37809 37 76 ----------------------------------------------------------------------- ----------------------------------------------------------------------- : 37809 37 77 _ _ NNP 37809 37 78 campi campi NNP 37809 37 79 _ _ NNP 37809 37 80 , , , 37809 37 81 _ _ NNP 37809 37 82 Syrrhophus Syrrhophus NNP 37809 37 83 _ _ NNP 37809 37 84 _ _ NNP 37809 37 85 cystignathoides cystignathoide VBZ 37809 37 86 campi campi NNP 37809 37 87 _ _ NNP 37809 37 88 _ _ NNP 37809 37 89 cholorum cholorum NN 37809 37 90 _ _ NNP 37809 37 91 , , , 37809 37 92 _ _ NNP 37809 37 93 Syrrhophus Syrrhophus NNP 37809 37 94 leprus leprus NNP 37809 37 95 _ _ NNP 37809 37 96 _ _ NNP 37809 37 97 leprus leprus NNP 37809 37 98 _ _ NNP 37809 37 99 _ _ NNP 37809 37 100 cystigathoides cystigathoide VBZ 37809 37 101 _ _ NNP 37809 37 102 , , , 37809 37 103 _ _ NNP 37809 37 104 Phyllobates Phyllobates NNPS 37809 37 105 _ _ NNP 37809 37 106 _ _ NNP 37809 37 107 cystignathoides cystignathoide VBZ 37809 37 108 cystignathoides cystignathoide VBZ 37809 37 109 _ _ NNP 37809 37 110 _ _ NNP 37809 37 111 dennisi dennisi NNP 37809 37 112 _ _ NNP 37809 37 113 , , , 37809 37 114 _ _ NNP 37809 37 115 Syrrhophus Syrrhophus NNP 37809 37 116 _ _ NNP 37809 37 117 _ _ NNP 37809 37 118 dennisi dennisi NNP 37809 37 119 _ _ NNP 37809 37 120 new new JJ 37809 37 121 species specie NNS 37809 37 122 _ _ NNP 37809 37 123 gaigeae gaigeae NN 37809 37 124 _ _ NNP 37809 37 125 , , , 37809 37 126 _ _ NNP 37809 37 127 Syrrhophus Syrrhophus NNP 37809 37 128 _ _ NNP 37809 37 129 _ _ NNP 37809 37 130 guttilatus guttilatus NN 37809 37 131 _ _ NNP 37809 37 132 _ _ NNP 37809 37 133 guttilatus guttilatus NN 37809 37 134 _ _ NNP 37809 37 135 , , , 37809 37 136 _ _ NNP 37809 37 137 Malachylodes Malachylodes NNP 37809 37 138 _ _ NNP 37809 37 139 _ _ NNP 37809 37 140 guttilatus guttilatus NN 37809 37 141 _ _ NNP 37809 37 142 _ _ NNP 37809 37 143 interorbitalis interorbitalis NN 37809 37 144 _ _ NNP 37809 37 145 , , , 37809 37 146 _ _ NNP 37809 37 147 Syrrhophus Syrrhophus NNP 37809 37 148 _ _ NNP 37809 37 149 _ _ NNP 37809 37 150 interorbitalis interorbitalis NN 37809 37 151 _ _ NNP 37809 37 152 _ _ NNP 37809 37 153 latodactylus latodactylus NN 37809 37 154 _ _ NNP 37809 37 155 , , , 37809 37 156 _ _ NNP 37809 37 157 Syrrhophus Syrrhophus NNP 37809 37 158 _ _ NNP 37809 37 159 _ _ NNP 37809 37 160 longipes longipe VBZ 37809 37 161 _ _ NNP 37809 37 162 _ _ NNP 37809 37 163 leprus leprus NNP 37809 37 164 _ _ NNP 37809 37 165 , , , 37809 37 166 _ _ NNP 37809 37 167 Syrrhophus Syrrhophus NNP 37809 37 168 _ _ NNP 37809 37 169 _ _ NNP 37809 37 170 leprus leprus NNP 37809 37 171 _ _ NNP 37809 37 172 _ _ NNP 37809 37 173 longipes longipe VBZ 37809 37 174 _ _ NNP 37809 37 175 , , , 37809 37 176 _ _ NNP 37809 37 177 Batrachyla Batrachyla NNP 37809 37 178 _ _ NNP 37809 37 179 _ _ NNP 37809 37 180 longipes longipe VBZ 37809 37 181 _ _ NNP 37809 37 182 _ _ NNP 37809 37 183 macrotympanum macrotympanum NNP 37809 37 184 _ _ NNP 37809 37 185 , , , 37809 37 186 _ _ NNP 37809 37 187 Tomodactylus Tomodactylus NNP 37809 37 188 _ _ NNP 37809 37 189 _ _ NNP 37809 37 190 verrucipes verrucipe NNS 37809 37 191 _ _ NNP 37809 37 192 _ _ NNP 37809 37 193 marnockii marnockii NNS 37809 37 194 _ _ NNP 37809 37 195 , , , 37809 37 196 _ _ NNP 37809 37 197 Syrrhophus Syrrhophus NNP 37809 37 198 _ _ NNP 37809 37 199 _ _ NNP 37809 37 200 marnockii marnockii NNS 37809 37 201 _ _ NNP 37809 37 202 _ _ NNP 37809 37 203 modestus modestus NN 37809 37 204 _ _ NNP 37809 37 205 , , , 37809 37 206 _ _ NNP 37809 37 207 Syrrhophus Syrrhophus NNP 37809 37 208 _ _ NNP 37809 37 209 _ _ NNP 37809 37 210 modestus modestus NN 37809 37 211 _ _ NNP 37809 37 212 _ _ NNP 37809 37 213 nebulosus nebulosus NN 37809 37 214 _ _ NNP 37809 37 215 , , , 37809 37 216 _ _ NNP 37809 37 217 Syrrhophus Syrrhophus NNP 37809 37 218 _ _ NNP 37809 37 219 _ _ NNP 37809 37 220 pipilans pipilans NNPS 37809 37 221 nebulosus nebulosus NN 37809 37 222 _ _ NNP 37809 37 223 _ _ NNP 37809 37 224 nivocolimae nivocolimae NN 37809 37 225 _ _ NNP 37809 37 226 , , , 37809 37 227 _ _ NNP 37809 37 228 Syrrhophus Syrrhophus NNP 37809 37 229 _ _ NNP 37809 37 230 _ _ NNP 37809 37 231 nivocolimae nivocolimae NN 37809 37 232 _ _ NNP 37809 37 233 _ _ NNP 37809 37 234 pallidus pallidus NNP 37809 37 235 _ _ NNP 37809 37 236 , , , 37809 37 237 _ _ NNP 37809 37 238 Syrrhophus Syrrhophus NNP 37809 37 239 modestus modestus NN 37809 37 240 _ _ NNP 37809 37 241 _ _ NNP 37809 37 242 pallidus pallidus NNP 37809 37 243 _ _ NNP 37809 37 244 _ _ NNP 37809 37 245 petrophilus petrophilus NN 37809 37 246 _ _ NNP 37809 37 247 , , , 37809 37 248 _ _ NNP 37809 37 249 Syrrhophus Syrrhophus NNP 37809 37 250 _ _ NNP 37809 37 251 _ _ NNP 37809 37 252 guttilatus guttilatus NN 37809 37 253 _ _ NNP 37809 37 254 _ _ NNP 37809 37 255 pipilans pipilans NNPS 37809 37 256 _ _ NNP 37809 37 257 , , , 37809 37 258 _ _ NNP 37809 37 259 Syrrhophus Syrrhophus NNP 37809 37 260 _ _ NNP 37809 37 261 _ _ NNP 37809 37 262 pipilans pipilans NNPS 37809 37 263 pipilans pipilans NNPS 37809 37 264 _ _ NNP 37809 37 265 _ _ NNP 37809 37 266 rubrimaculatus rubrimaculatus NN 37809 37 267 _ _ NNP 37809 37 268 , , , 37809 37 269 _ _ NNP 37809 37 270 Syrrhophus Syrrhophus NNP 37809 37 271 _ _ NNP 37809 37 272 _ _ NNP 37809 37 273 rubrimaculatus rubrimaculatus NN 37809 37 274 _ _ NNP 37809 37 275 _ _ NNP 37809 37 276 smithi smithi NN 37809 37 277 _ _ NNP 37809 37 278 , , , 37809 37 279 _ _ NNP 37809 37 280 Syrrhophus Syrrhophus NNP 37809 37 281 _ _ NNP 37809 37 282 _ _ NNP 37809 37 283 guttilatus guttilatus NN 37809 37 284 _ _ NNP 37809 37 285 _ _ NNP 37809 37 286 teretistes teretiste VBZ 37809 37 287 _ _ NNP 37809 37 288 , , , 37809 37 289 _ _ NNP 37809 37 290 Syrrhophus Syrrhophus NNP 37809 37 291 _ _ NNP 37809 37 292 _ _ NNP 37809 37 293 teretistes teretiste VBZ 37809 37 294 _ _ NNP 37809 37 295 _ _ NNP 37809 37 296 verrucipes verrucipe VBZ 37809 37 297 _ _ NNP 37809 37 298 , , , 37809 37 299 _ _ NNP 37809 37 300 Syrrhophus Syrrhophus NNP 37809 37 301 _ _ NNP 37809 37 302 _ _ NNP 37809 37 303 verrucipes verrucipe VBZ 37809 37 304 _ _ NNP 37809 37 305 _ _ NNP 37809 37 306 verruculatus verruculatus NNP 37809 37 307 _ _ NNP 37809 37 308 , , , 37809 37 309 _ _ NNP 37809 37 310 Phyllobates Phyllobates NNPS 37809 37 311 _ _ NNP 37809 37 312 _ _ NNP 37809 37 313 Nomen Nomen NNP 37809 37 314 dubium dubium NN 37809 37 315 _ _ NNP 37809 37 316 = = SYM 37809 37 317 = = SYM 37809 37 318 = = SYM 37809 37 319 = = SYM 37809 37 320 = = SYM 37809 37 321 = = SYM 37809 37 322 = = SYM 37809 37 323 = = SYM 37809 37 324 = = SYM 37809 37 325 = = SYM 37809 37 326 = = SYM 37809 37 327 = = SYM 37809 37 328 = = SYM 37809 37 329 = = SYM 37809 37 330 = = SYM 37809 37 331 = = SYM 37809 37 332 = = SYM 37809 37 333 = = SYM 37809 37 334 = = SYM 37809 37 335 = = SYM 37809 37 336 = = SYM 37809 37 337 = = SYM 37809 37 338 = = SYM 37809 37 339 = = SYM 37809 37 340 = = SYM 37809 37 341 = = SYM 37809 37 342 = = SYM 37809 37 343 = = SYM 37809 37 344 = = SYM 37809 37 345 = = SYM 37809 37 346 = = SYM 37809 37 347 = = SYM 37809 37 348 = = SYM 37809 37 349 = = SYM 37809 37 350 = = SYM 37809 37 351 = = SYM 37809 37 352 = = SYM 37809 37 353 = = SYM 37809 37 354 = = SYM 37809 37 355 = = SYM 37809 37 356 = = SYM 37809 37 357 = = SYM 37809 37 358 = = SYM 37809 37 359 = = SYM 37809 37 360 = = SYM 37809 37 361 = = SYM 37809 37 362 = = SYM 37809 37 363 = = SYM 37809 37 364 = = SYM 37809 37 365 = = SYM 37809 37 366 = = SYM 37809 37 367 = = SYM 37809 37 368 = = SYM 37809 37 369 = = SYM 37809 37 370 = = SYM 37809 37 371 = = SYM 37809 37 372 = = SYM 37809 37 373 = = SYM 37809 37 374 = = SYM 37809 37 375 = = SYM 37809 37 376 = = SYM 37809 37 377 = = SYM 37809 37 378 = = SYM 37809 37 379 = = SYM 37809 37 380 = = SYM 37809 37 381 = = SYM 37809 37 382 = = SYM 37809 37 383 = = SYM 37809 37 384 = = SYM 37809 37 385 = = NFP 37809 37 386 = = NFP 37809 37 387 In in IN 37809 37 388 the the DT 37809 37 389 course course NN 37809 37 390 of of IN 37809 37 391 preparing prepare VBG 37809 37 392 an an DT 37809 37 393 account account NN 37809 37 394 of of IN 37809 37 395 the the DT 37809 37 396 species specie NNS 37809 37 397 of of IN 37809 37 398 _ _ NNP 37809 37 399 Eleutherodactylus Eleutherodactylus NNP 37809 37 400 _ _ NNP 37809 37 401 occurring occur VBG 37809 37 402 in in IN 37809 37 403 México México NNP 37809 37 404 and and CC 37809 37 405 northern northern JJ 37809 37 406 Central Central NNP 37809 37 407 America America NNP 37809 37 408 , , , 37809 37 409 it -PRON- PRP 37809 37 410 became become VBD 37809 37 411 necessary necessary JJ 37809 37 412 to to TO 37809 37 413 reëxamine reëxamine VB 37809 37 414 the the DT 37809 37 415 status status NN 37809 37 416 of of IN 37809 37 417 the the DT 37809 37 418 genus genus NN 37809 37 419 _ _ NNP 37809 37 420 Syrrhophus Syrrhophus NNP 37809 37 421 _ _ NNP 37809 37 422 and and CC 37809 37 423 its -PRON- PRP$ 37809 37 424 nominal nominal JJ 37809 37 425 species specie NNS 37809 37 426 . . . 37809 38 1 It -PRON- PRP 37809 38 2 soon soon RB 37809 38 3 became become VBD 37809 38 4 evident evident JJ 37809 38 5 that that IN 37809 38 6 there there EX 37809 38 7 were be VBD 37809 38 8 more more JJR 37809 38 9 names name NNS 37809 38 10 than than IN 37809 38 11 species specie NNS 37809 38 12 , , , 37809 38 13 that that IN 37809 38 14 some some DT 37809 38 15 previously previously RB 37809 38 16 regarded regard VBN 37809 38 17 species specie NNS 37809 38 18 were be VBD 37809 38 19 geographic geographic JJ 37809 38 20 variants variant NNS 37809 38 21 , , , 37809 38 22 and and CC 37809 38 23 that that IN 37809 38 24 the the DT 37809 38 25 eastern eastern JJ 37809 38 26 and and CC 37809 38 27 western western JJ 37809 38 28 groups group NNS 37809 38 29 ( ( -LRB- 37809 38 30 complexes complex NNS 37809 38 31 here here RB 37809 38 32 ) ) -RRB- 37809 38 33 were be VBD 37809 38 34 artificial artificial JJ 37809 38 35 divisions division NNS 37809 38 36 of of IN 37809 38 37 the the DT 37809 38 38 genus genus NN 37809 38 39 . . . 37809 39 1 I -PRON- PRP 37809 39 2 conclude conclude VBP 37809 39 3 that that IN 37809 39 4 there there EX 37809 39 5 are be VBP 37809 39 6 seven seven CD 37809 39 7 species specie NNS 37809 39 8 ( ( -LRB- 37809 39 9 one one CD 37809 39 10 polytypic polytypic NN 37809 39 11 ) ) -RRB- 37809 39 12 of of IN 37809 39 13 _ _ NNP 37809 39 14 Syrrhophus Syrrhophus NNP 37809 39 15 _ _ NNP 37809 39 16 in in IN 37809 39 17 eastern eastern JJ 37809 39 18 México México NNP 37809 39 19 , , , 37809 39 20 Texas Texas NNP 37809 39 21 , , , 37809 39 22 and and CC 37809 39 23 El El NNP 37809 39 24 Petén Petén NNP 37809 39 25 of of IN 37809 39 26 Guatemala Guatemala NNP 37809 39 27 , , , 37809 39 28 and and CC 37809 39 29 seven seven CD 37809 39 30 species specie NNS 37809 39 31 ( ( -LRB- 37809 39 32 one one CD 37809 39 33 polytypic polytypic NN 37809 39 34 ) ) -RRB- 37809 39 35 in in IN 37809 39 36 western western JJ 37809 39 37 México México NNP 37809 39 38 . . . 37809 40 1 The the DT 37809 40 2 current current JJ 37809 40 3 status status NN 37809 40 4 of of IN 37809 40 5 each each DT 37809 40 6 of of IN 37809 40 7 the the DT 37809 40 8 23 23 CD 37809 40 9 names name NNS 37809 40 10 correctly correctly RB 37809 40 11 assigned assign VBN 37809 40 12 to to IN 37809 40 13 the the DT 37809 40 14 genus genus NN 37809 40 15 is be VBZ 37809 40 16 presented present VBN 37809 40 17 in in IN 37809 40 18 Table table NN 37809 40 19 3 3 CD 37809 40 20 . . . 37809 41 1 The the DT 37809 41 2 fourteen fourteen CD 37809 41 3 species specie NNS 37809 41 4 recognized recognize VBN 37809 41 5 by by IN 37809 41 6 me -PRON- PRP 37809 41 7 are be VBP 37809 41 8 placed place VBN 37809 41 9 in in IN 37809 41 10 five five CD 37809 41 11 species specie NNS 37809 41 12 groups group NNS 37809 41 13 . . . 37809 42 1 Two two CD 37809 42 2 of of IN 37809 42 3 these these DT 37809 42 4 groups group NNS 37809 42 5 are be VBP 37809 42 6 presently presently RB 37809 42 7 placed place VBN 37809 42 8 in in IN 37809 42 9 the the DT 37809 42 10 western western JJ 37809 42 11 complex complex NN 37809 42 12 ( ( -LRB- 37809 42 13 _ _ NNP 37809 42 14 modestus modestus NN 37809 42 15 _ _ NNP 37809 42 16 and and CC 37809 42 17 _ _ NNP 37809 42 18 pipilans pipilans NNPS 37809 42 19 _ _ NNP 37809 42 20 groups group NNS 37809 42 21 ) ) -RRB- 37809 42 22 and and CC 37809 42 23 three three CD 37809 42 24 in in IN 37809 42 25 the the DT 37809 42 26 eastern eastern JJ 37809 42 27 complex complex NN 37809 42 28 ( ( -LRB- 37809 42 29 _ _ NNP 37809 42 30 leprus leprus NNP 37809 42 31 _ _ NNP 37809 42 32 , , , 37809 42 33 _ _ NNP 37809 42 34 longipes longipe VBZ 37809 42 35 _ _ NNP 37809 42 36 and and CC 37809 42 37 _ _ NNP 37809 42 38 marnockii marnockii NNS 37809 42 39 _ _ NNP 37809 42 40 groups group NNS 37809 42 41 ) ) -RRB- 37809 42 42 . . . 37809 43 1 The the DT 37809 43 2 two two CD 37809 43 3 complexes complex NNS 37809 43 4 do do VBP 37809 43 5 not not RB 37809 43 6 correspond correspond VB 37809 43 7 exactly exactly RB 37809 43 8 with with IN 37809 43 9 the the DT 37809 43 10 eastern eastern JJ 37809 43 11 and and CC 37809 43 12 western western JJ 37809 43 13 groups group NNS 37809 43 14 of of IN 37809 43 15 Smith Smith NNP 37809 43 16 and and CC 37809 43 17 Taylor Taylor NNP 37809 43 18 ( ( -LRB- 37809 43 19 1948 1948 CD 37809 43 20 ) ) -RRB- 37809 43 21 , , , 37809 43 22 Firschein Firschein NNP 37809 43 23 ( ( -LRB- 37809 43 24 1954 1954 CD 37809 43 25 ) ) -RRB- 37809 43 26 , , , 37809 43 27 and and CC 37809 43 28 Duellman Duellman NNP 37809 43 29 ( ( -LRB- 37809 43 30 1958 1958 CD 37809 43 31 ) ) -RRB- 37809 43 32 since since IN 37809 43 33 _ _ NNP 37809 43 34 S. S. NNP 37809 44 1 rubrimaculatus rubrimaculatus NNP 37809 44 2 _ _ NNP 37809 44 3 is be VBZ 37809 44 4 now now RB 37809 44 5 associated associate VBN 37809 44 6 with with IN 37809 44 7 the the DT 37809 44 8 eastern eastern JJ 37809 44 9 _ _ NNP 37809 44 10 leprus leprus NNP 37809 44 11 _ _ NNP 37809 44 12 group group NN 37809 44 13 . . . 37809 45 1 The the DT 37809 45 2 definitions definition NNS 37809 45 3 and and CC 37809 45 4 contents content NNS 37809 45 5 of of IN 37809 45 6 the the DT 37809 45 7 five five CD 37809 45 8 species specie NNS 37809 45 9 groups group NNS 37809 45 10 are be VBP 37809 45 11 as as IN 37809 45 12 follows follow VBZ 37809 45 13 : : : 37809 45 14 _ _ NNP 37809 45 15 leprus leprus NNP 37809 45 16 _ _ NNP 37809 45 17 group group NN 37809 45 18 : : : 37809 45 19 digital digital JJ 37809 45 20 pads pad NNS 37809 45 21 not not RB 37809 45 22 or or CC 37809 45 23 only only RB 37809 45 24 slightly slightly RB 37809 45 25 expanded expand VBN 37809 45 26 , , , 37809 45 27 rounded round VBN 37809 45 28 in in IN 37809 45 29 outline outline NN 37809 45 30 ; ; : 37809 45 31 first first JJ 37809 45 32 finger finger NN 37809 45 33 longer long RBR 37809 45 34 or or CC 37809 45 35 shorter short JJR 37809 45 36 than than IN 37809 45 37 second second JJ 37809 45 38 ; ; : 37809 45 39 snout snout NN 37809 45 40 acuminate acuminate NN 37809 45 41 or or CC 37809 45 42 subacuminate subacuminate VB 37809 45 43 , , , 37809 45 44 not not RB 37809 45 45 rounded rounded JJ 37809 45 46 ; ; : 37809 45 47 outer outer JJ 37809 45 48 metatarsal metatarsal NNP 37809 45 49 tubercle tubercle NN 37809 45 50 conical conical NN 37809 45 51 ; ; : 37809 45 52 digits digit NNS 37809 45 53 lacking lack VBG 37809 45 54 distinct distinct JJ 37809 45 55 lateral lateral JJ 37809 45 56 fringes fringe NNS 37809 45 57 . . . 37809 46 1 content content NN 37809 46 2 : : : 37809 46 3 _ _ NNP 37809 46 4 cystignathoides cystignathoide VBZ 37809 46 5 _ _ NNP 37809 46 6 , , , 37809 46 7 _ _ NNP 37809 46 8 leprus leprus NNP 37809 46 9 _ _ NNP 37809 46 10 and and CC 37809 46 11 _ _ NNP 37809 46 12 rubrimaculatus rubrimaculatus NN 37809 46 13 _ _ NNP 37809 46 14 . . . 37809 47 1 _ _ NNP 37809 47 2 longipes longipes NNP 37809 47 3 _ _ NNP 37809 47 4 group group NN 37809 47 5 : : : 37809 47 6 digital digital NNP 37809 47 7 pads pad NNS 37809 47 8 widely widely RB 37809 47 9 expanded expand VBN 37809 47 10 , , , 37809 47 11 triangular triangular JJ 37809 47 12 in in IN 37809 47 13 outline outline NN 37809 47 14 ; ; : 37809 47 15 first first RB 37809 47 16 finger finger NN 37809 47 17 shorter short JJR 37809 47 18 than than IN 37809 47 19 second second JJ 37809 47 20 ; ; : 37809 47 21 snout snout NN 37809 47 22 acuminate acuminate NN 37809 47 23 ; ; : 37809 47 24 outer outer JJ 37809 47 25 metatarsal metatarsal NN 37809 47 26 tubercle tubercle NN 37809 47 27 not not RB 37809 47 28 conical conical JJ 37809 47 29 ; ; : 37809 47 30 digits digit VBZ 37809 47 31 bearing bearing NN 37809 47 32 lateral lateral JJ 37809 47 33 fringes fringe NNS 37809 47 34 . . . 37809 48 1 content content NN 37809 48 2 : : : 37809 48 3 _ _ NNP 37809 48 4 dennisi dennisi NNP 37809 48 5 _ _ NNP 37809 48 6 and and CC 37809 48 7 _ _ NNP 37809 48 8 longipes longipe VBZ 37809 48 9 _ _ NNP 37809 48 10 . . . 37809 49 1 _ _ NNP 37809 49 2 marnockii marnockii NNP 37809 49 3 _ _ NNP 37809 49 4 group group NN 37809 49 5 : : : 37809 49 6 digital digital NNP 37809 49 7 pads pad NNS 37809 49 8 expanded expand VBD 37809 49 9 , , , 37809 49 10 rounded round VBD 37809 49 11 to to TO 37809 49 12 truncate truncate VB 37809 49 13 in in IN 37809 49 14 outline outline NN 37809 49 15 ; ; : 37809 49 16 first first RB 37809 49 17 finger finger NN 37809 49 18 equal equal JJ 37809 49 19 in in IN 37809 49 20 length length NN 37809 49 21 to to TO 37809 49 22 second second JJ 37809 49 23 or or CC 37809 49 24 slightly slightly RB 37809 49 25 shorter short JJR 37809 49 26 ; ; : 37809 49 27 snout snout NN 37809 49 28 rounded round VBD 37809 49 29 ; ; : 37809 49 30 outer outer JJ 37809 49 31 metatarsal metatarsal NN 37809 49 32 tubercle tubercle NN 37809 49 33 not not RB 37809 49 34 conical conical JJ 37809 49 35 ; ; : 37809 49 36 digits digit NNS 37809 49 37 lacking lack VBG 37809 49 38 lateral lateral JJ 37809 49 39 fringes fringe NNS 37809 49 40 ; ; : 37809 49 41 generally generally RB 37809 49 42 stout stout JJ 37809 49 43 - - HYPH 37809 49 44 bodied bodied JJ 37809 49 45 frogs frog NNS 37809 49 46 . . . 37809 50 1 content content NN 37809 50 2 : : : 37809 50 3 _ _ NNP 37809 50 4 guttilatus guttilatus NNP 37809 50 5 _ _ NNP 37809 50 6 , , , 37809 50 7 _ _ NNP 37809 50 8 marnockii marnockii NNS 37809 50 9 _ _ NNP 37809 50 10 , , , 37809 50 11 and and CC 37809 50 12 _ _ NNP 37809 50 13 verrucipes verrucipe VBZ 37809 50 14 _ _ NNP 37809 50 15 . . . 37809 51 1 _ _ NNP 37809 51 2 modestus modestus NNP 37809 51 3 _ _ NNP 37809 51 4 group group NN 37809 51 5 : : : 37809 51 6 digital digital NNP 37809 51 7 pads pad NNS 37809 51 8 expanded expand VBD 37809 51 9 , , , 37809 51 10 truncate truncate JJ 37809 51 11 in in IN 37809 51 12 outline outline NN 37809 51 13 ; ; : 37809 51 14 first first JJ 37809 51 15 and and CC 37809 51 16 second second JJ 37809 51 17 fingers finger NNS 37809 51 18 subequal subequal JJ 37809 51 19 in in IN 37809 51 20 length length NN 37809 51 21 , , , 37809 51 22 first first RB 37809 51 23 usually usually RB 37809 51 24 slightly slightly RB 37809 51 25 shorter short JJR 37809 51 26 than than IN 37809 51 27 second second JJ 37809 51 28 ; ; : 37809 51 29 snout snout NN 37809 51 30 subacuminate subacuminate NN 37809 51 31 ; ; : 37809 51 32 inner inner JJ 37809 51 33 metatarsal metatarsal NN 37809 51 34 tubercle tubercle NN 37809 51 35 twice twice RB 37809 51 36 as as RB 37809 51 37 large large JJ 37809 51 38 ( ( -LRB- 37809 51 39 or or CC 37809 51 40 larger large JJR 37809 51 41 ) ) -RRB- 37809 51 42 as as IN 37809 51 43 outer outer JJ 37809 51 44 metatarsal metatarsal NN 37809 51 45 tubercle tubercle NN 37809 51 46 ; ; : 37809 51 47 digits digit NNS 37809 51 48 bearing bearing NN 37809 51 49 poorly poorly RB 37809 51 50 - - HYPH 37809 51 51 defined define VBN 37809 51 52 lateral lateral JJ 37809 51 53 fringes fringe NNS 37809 51 54 . . . 37809 52 1 content content NN 37809 52 2 : : : 37809 52 3 _ _ NNP 37809 52 4 interorbitalis interorbitalis NNP 37809 52 5 _ _ NNP 37809 52 6 , , , 37809 52 7 _ _ NNP 37809 52 8 modestus modestus NN 37809 52 9 _ _ NNP 37809 52 10 , , , 37809 52 11 _ _ NNP 37809 52 12 nivocolimae nivocolimae NN 37809 52 13 _ _ NNP 37809 52 14 , , , 37809 52 15 _ _ NNP 37809 52 16 pallidus pallidus NNP 37809 52 17 _ _ NNP 37809 52 18 , , , 37809 52 19 and and CC 37809 52 20 _ _ NNP 37809 52 21 teretistes teretistes NNPS 37809 52 22 _ _ NNP 37809 52 23 . . . 37809 53 1 _ _ NNP 37809 53 2 pipilans pipilans NNPS 37809 53 3 _ _ NNP 37809 53 4 group group NN 37809 53 5 : : : 37809 53 6 digital digital JJ 37809 53 7 pads pad NNS 37809 53 8 not not RB 37809 53 9 or or CC 37809 53 10 only only RB 37809 53 11 slightly slightly RB 37809 53 12 expanded expand VBN 37809 53 13 , , , 37809 53 14 truncate truncate JJ 37809 53 15 in in IN 37809 53 16 outline outline NN 37809 53 17 ; ; : 37809 53 18 first first RB 37809 53 19 finger finger NN 37809 53 20 equal equal JJ 37809 53 21 in in IN 37809 53 22 length length NN 37809 53 23 to to TO 37809 53 24 second second JJ 37809 53 25 ; ; : 37809 53 26 snout snout NN 37809 53 27 subacuminate subacuminate NN 37809 53 28 ; ; : 37809 53 29 metatarsal metatarsal NN 37809 53 30 tubercles tubercle NNS 37809 53 31 subequal subequal JJ 37809 53 32 in in IN 37809 53 33 size size NN 37809 53 34 ; ; , 37809 53 35 digits digit NNS 37809 53 36 lacking lack VBG 37809 53 37 lateral lateral JJ 37809 53 38 fringes fringe NNS 37809 53 39 . . . 37809 54 1 content content NN 37809 54 2 : : : 37809 54 3 _ _ NNP 37809 54 4 pipilans pipilans NNPS 37809 54 5 _ _ NNP 37809 54 6 . . . 37809 55 1 _ _ NNP 37809 55 2 Acknowledgments._--For acknowledgments._--for CD 37809 55 3 loan loan NN 37809 55 4 of of IN 37809 55 5 specimens specimen NNS 37809 55 6 , , , 37809 55 7 I -PRON- PRP 37809 55 8 am be VBP 37809 55 9 indebted indebted JJ 37809 55 10 to to IN 37809 55 11 Richard Richard NNP 37809 55 12 J. J. NNP 37809 55 13 Baldauf Baldauf NNP 37809 55 14 , , , 37809 55 15 Texas Texas NNP 37809 55 16 A A NNP 37809 55 17 & & CC 37809 55 18 M M NNP 37809 55 19 University University NNP 37809 55 20 ( ( -LRB- 37809 55 21 TCWC TCWC NNP 37809 55 22 ) ) -RRB- 37809 55 23 ; ; : 37809 55 24 W. W. NNP 37809 55 25 Frank Frank NNP 37809 55 26 Blair Blair NNP 37809 55 27 , , , 37809 55 28 University University NNP 37809 55 29 of of IN 37809 55 30 Texas Texas NNP 37809 55 31 ( ( -LRB- 37809 55 32 TNHC TNHC NNP 37809 55 33 ) ) -RRB- 37809 55 34 ; ; : 37809 55 35 Charles Charles NNP 37809 55 36 M. M. NNP 37809 55 37 Bogert Bogert NNP 37809 55 38 and and CC 37809 55 39 Richard Richard NNP 37809 55 40 G. G. NNP 37809 55 41 Zweifel Zweifel NNP 37809 55 42 , , , 37809 55 43 American American NNP 37809 55 44 Museum Museum NNP 37809 55 45 of of IN 37809 55 46 Natural Natural NNP 37809 55 47 History History NNP 37809 55 48 ( ( -LRB- 37809 55 49 AMNH AMNH NNP 37809 55 50 ) ) -RRB- 37809 55 51 ; ; : 37809 55 52 James James NNP 37809 55 53 E. E. NNP 37809 55 54 Böhlke Böhlke NNP 37809 55 55 and and CC 37809 55 56 Edmond Edmond NNP 37809 55 57 V. V. NNP 37809 55 58 Malnate Malnate NNP 37809 55 59 , , , 37809 55 60 Academy Academy NNP 37809 55 61 of of IN 37809 55 62 Natural Natural NNP 37809 55 63 Sciences Sciences NNPS 37809 55 64 of of IN 37809 55 65 Philadelphia Philadelphia NNP 37809 55 66 ( ( -LRB- 37809 55 67 ANSP ANSP NNP 37809 55 68 ) ) -RRB- 37809 55 69 ; ; : 37809 55 70 Robert Robert NNP 37809 55 71 F. F. NNP 37809 55 72 Inger Inger NNP 37809 55 73 and and CC 37809 55 74 Hymen Hymen NNP 37809 55 75 Marx Marx NNP 37809 55 76 , , , 37809 55 77 Field Field NNP 37809 55 78 Museum Museum NNP 37809 55 79 of of IN 37809 55 80 Natural Natural NNP 37809 55 81 History History NNP 37809 55 82 ( ( -LRB- 37809 55 83 FMNH FMNH NNP 37809 55 84 ) ) -RRB- 37809 55 85 ; ; : 37809 55 86 Ernest Ernest NNP 37809 55 87 A. a. NN 37809 55 88 Liner Liner NNP 37809 55 89 ( ( -LRB- 37809 55 90 EAL EAL NNP 37809 55 91 ) ) -RRB- 37809 55 92 ; ; : 37809 55 93 Michael Michael NNP 37809 55 94 Ovchynnyk Ovchynnyk NNP 37809 55 95 , , , 37809 55 96 Michigan Michigan NNP 37809 55 97 State State NNP 37809 55 98 University University NNP 37809 55 99 collection collection NN 37809 55 100 ( ( -LRB- 37809 55 101 MSU MSU NNP 37809 55 102 ) ) -RRB- 37809 55 103 ; ; : 37809 55 104 James James NNP 37809 55 105 A. A. NNP 37809 55 106 Peters Peters NNP 37809 55 107 , , , 37809 55 108 United United NNP 37809 55 109 States States NNP 37809 55 110 National National NNP 37809 55 111 Museum Museum NNP 37809 55 112 ( ( -LRB- 37809 55 113 USNM USNM NNP 37809 55 114 ) ) -RRB- 37809 55 115 ; ; : 37809 55 116 Douglas Douglas NNP 37809 55 117 A. A. NNP 37809 55 118 Rossman Rossman NNP 37809 55 119 , , , 37809 55 120 Louisiana Louisiana NNP 37809 55 121 State State NNP 37809 55 122 University University NNP 37809 55 123 Museum Museum NNP 37809 55 124 of of IN 37809 55 125 Zoology Zoology NNP 37809 55 126 ( ( -LRB- 37809 55 127 LSUMZ LSUMZ NNP 37809 55 128 ) ) -RRB- 37809 55 129 ; ; : 37809 55 130 Hobart Hobart NNP 37809 55 131 M. M. NNP 37809 55 132 Smith Smith NNP 37809 55 133 , , , 37809 55 134 University University NNP 37809 55 135 of of IN 37809 55 136 Illinois Illinois NNP 37809 55 137 Museum Museum NNP 37809 55 138 of of IN 37809 55 139 Natural Natural NNP 37809 55 140 History History NNP 37809 55 141 ( ( -LRB- 37809 55 142 UIMNH UIMNH NNP 37809 55 143 ) ) -RRB- 37809 55 144 ; ; : 37809 55 145 Charles Charles NNP 37809 55 146 F. F. NNP 37809 55 147 Walker Walker NNP 37809 55 148 , , , 37809 55 149 University University NNP 37809 55 150 of of IN 37809 55 151 Michigan Michigan NNP 37809 55 152 Museum Museum NNP 37809 55 153 of of IN 37809 55 154 Zoology Zoology NNP 37809 55 155 ( ( -LRB- 37809 55 156 UMMZ UMMZ NNP 37809 55 157 ) ) -RRB- 37809 55 158 ; ; : 37809 55 159 and and CC 37809 55 160 John John NNP 37809 55 161 W. W. NNP 37809 55 162 Wright Wright NNP 37809 55 163 , , , 37809 55 164 Los Los NNP 37809 55 165 Angeles Angeles NNP 37809 55 166 County County NNP 37809 55 167 Museum Museum NNP 37809 55 168 ( ( -LRB- 37809 55 169 LACM LACM NNP 37809 55 170 ) ) -RRB- 37809 55 171 . . . 37809 56 1 Specimens specimen NNS 37809 56 2 in in IN 37809 56 3 the the DT 37809 56 4 collection collection NN 37809 56 5 at at IN 37809 56 6 the the DT 37809 56 7 University University NNP 37809 56 8 of of IN 37809 56 9 Kansas Kansas NNP 37809 56 10 Museum Museum NNP 37809 56 11 of of IN 37809 56 12 Natural Natural NNP 37809 56 13 History History NNP 37809 56 14 are be VBP 37809 56 15 identified identify VBN 37809 56 16 as as IN 37809 56 17 KU KU NNP 37809 56 18 . . . 37809 57 1 The the DT 37809 57 2 abbreviations abbreviation NNS 37809 57 3 EHT EHT NNP 37809 57 4 - - HYPH 37809 57 5 HMS HMS NNP 37809 57 6 refer refer VB 37809 57 7 to to IN 37809 57 8 the the DT 37809 57 9 Edward Edward NNP 37809 57 10 H. H. NNP 37809 57 11 Taylor Taylor NNP 37809 57 12 - - HYPH 37809 57 13 Hobart Hobart NNP 37809 57 14 M. M. NNP 37809 57 15 Smith Smith NNP 37809 57 16 collection collection NN 37809 57 17 and and CC 37809 57 18 FAS FAS NNP 37809 57 19 to to IN 37809 57 20 the the DT 37809 57 21 Frederick Frederick NNP 37809 57 22 A. A. NNP 37809 57 23 Shannon Shannon NNP 37809 57 24 collection collection NN 37809 57 25 . . . 37809 58 1 The the DT 37809 58 2 type type NN 37809 58 3 - - HYPH 37809 58 4 specimens specimen NNS 37809 58 5 from from IN 37809 58 6 these these DT 37809 58 7 collections collection NNS 37809 58 8 are be VBP 37809 58 9 now now RB 37809 58 10 in in IN 37809 58 11 the the DT 37809 58 12 Field Field NNP 37809 58 13 Museum Museum NNP 37809 58 14 of of IN 37809 58 15 Natural Natural NNP 37809 58 16 History History NNP 37809 58 17 and and CC 37809 58 18 the the DT 37809 58 19 University University NNP 37809 58 20 of of IN 37809 58 21 Illinois Illinois NNP 37809 58 22 Museum Museum NNP 37809 58 23 of of IN 37809 58 24 Natural Natural NNP 37809 58 25 History History NNP 37809 58 26 . . . 37809 59 1 I -PRON- PRP 37809 59 2 have have VBP 37809 59 3 profited profit VBN 37809 59 4 from from IN 37809 59 5 discussions discussion NNS 37809 59 6 concerning concern VBG 37809 59 7 this this DT 37809 59 8 problem problem NN 37809 59 9 with with IN 37809 59 10 several several JJ 37809 59 11 persons person NNS 37809 59 12 , , , 37809 59 13 most most RBS 37809 59 14 notably notably RB 37809 59 15 William William NNP 37809 59 16 E. E. NNP 37809 59 17 Duellman Duellman NNP 37809 59 18 , , , 37809 59 19 Hobart Hobart NNP 37809 59 20 M. M. NNP 37809 59 21 Smith Smith NNP 37809 59 22 , , , 37809 59 23 Edward Edward NNP 37809 59 24 H. H. NNP 37809 59 25 Taylor Taylor NNP 37809 59 26 and and CC 37809 59 27 Charles Charles NNP 37809 59 28 F. F. NNP 37809 59 29 Walker Walker NNP 37809 59 30 . . . 37809 60 1 Nevertheless nevertheless RB 37809 60 2 , , , 37809 60 3 the the DT 37809 60 4 ideas idea NNS 37809 60 5 and and CC 37809 60 6 conclusions conclusion NNS 37809 60 7 presented present VBN 37809 60 8 here here RB 37809 60 9 should should MD 37809 60 10 not not RB 37809 60 11 be be VB 37809 60 12 construed construe VBN 37809 60 13 as as IN 37809 60 14 necessarily necessarily RB 37809 60 15 reflecting reflect VBG 37809 60 16 their -PRON- PRP$ 37809 60 17 opinions opinion NNS 37809 60 18 . . . 37809 61 1 David David NNP 37809 61 2 M. M. NNP 37809 61 3 Dennis Dennis NNP 37809 61 4 executed execute VBD 37809 61 5 all all DT 37809 61 6 of of IN 37809 61 7 the the DT 37809 61 8 figures figure NNS 37809 61 9 , , , 37809 61 10 and and CC 37809 61 11 my -PRON- PRP$ 37809 61 12 wife wife NN 37809 61 13 , , , 37809 61 14 Marsha Marsha NNP 37809 61 15 , , , 37809 61 16 typed type VBD 37809 61 17 the the DT 37809 61 18 manuscript manuscript NN 37809 61 19 . . . 37809 62 1 _ _ NNP 37809 62 2 Materials Materials NNP 37809 62 3 and and CC 37809 62 4 Methods._--In methods._--in NN 37809 62 5 the the DT 37809 62 6 course course NN 37809 62 7 of of IN 37809 62 8 this this DT 37809 62 9 study study NN 37809 62 10 , , , 37809 62 11 1003 1003 CD 37809 62 12 specimens specimen NNS 37809 62 13 of of IN 37809 62 14 the the DT 37809 62 15 genus genus NNPS 37809 62 16 were be VBD 37809 62 17 examined examine VBN 37809 62 18 . . . 37809 63 1 The the DT 37809 63 2 holotypes holotype NNS 37809 63 3 of of IN 37809 63 4 21 21 CD 37809 63 5 of of IN 37809 63 6 the the DT 37809 63 7 23 23 CD 37809 63 8 nominal nominal JJ 37809 63 9 species specie NNS 37809 63 10 are be VBP 37809 63 11 extant extant JJ 37809 63 12 ; ; : 37809 63 13 I -PRON- PRP 37809 63 14 have have VBP 37809 63 15 examined examine VBN 37809 63 16 19 19 CD 37809 63 17 of of IN 37809 63 18 these these DT 37809 63 19 . . . 37809 64 1 Nine nine CD 37809 64 2 measurements measurement NNS 37809 64 3 were be VBD 37809 64 4 taken take VBN 37809 64 5 , , , 37809 64 6 and and CC 37809 64 7 five five CD 37809 64 8 ratios ratio NNS 37809 64 9 computed compute VBN 37809 64 10 for for IN 37809 64 11 each each DT 37809 64 12 of of IN 37809 64 13 338 338 CD 37809 64 14 specimens specimen NNS 37809 64 15 . . . 37809 65 1 Females female NNS 37809 65 2 are be VBP 37809 65 3 available available JJ 37809 65 4 for for IN 37809 65 5 all all DT 37809 65 6 species specie NNS 37809 65 7 but but CC 37809 65 8 one one CD 37809 65 9 ; ; : 37809 65 10 thus thus RB 37809 65 11 , , , 37809 65 12 measurements measurement NNS 37809 65 13 were be VBD 37809 65 14 taken take VBN 37809 65 15 on on IN 37809 65 16 individuals individual NNS 37809 65 17 of of IN 37809 65 18 both both DT 37809 65 19 sexes sex NNS 37809 65 20 . . . 37809 66 1 ANALYSIS ANALYSIS NNP 37809 66 2 OF of IN 37809 66 3 CHARACTERS character NNS 37809 66 4 _ _ NNP 37809 66 5 Size Size NNP 37809 66 6 and and CC 37809 66 7 proportions._--Frogs proportions._--frog NNS 37809 66 8 of of IN 37809 66 9 this this DT 37809 66 10 genus genus JJ 37809 66 11 range range NN 37809 66 12 in in IN 37809 66 13 size size NN 37809 66 14 from from IN 37809 66 15 16 16 CD 37809 66 16 to to TO 37809 66 17 40 40 CD 37809 66 18 mm mm NNS 37809 66 19 . . . 37809 67 1 in in IN 37809 67 2 snout snout NN 37809 67 3 - - HYPH 37809 67 4 vent vent NN 37809 67 5 length length NN 37809 67 6 . . . 37809 68 1 Five five CD 37809 68 2 species specie NNS 37809 68 3 are be VBP 37809 68 4 relatively relatively RB 37809 68 5 small small JJ 37809 68 6 : : : 37809 68 7 _ _ NNP 37809 68 8 S. S. NNP 37809 69 1 cystignathoides cystignathoide VBZ 37809 69 2 _ _ NNP 37809 69 3 , , , 37809 69 4 _ _ NNP 37809 69 5 modestus modestus NN 37809 69 6 _ _ NNP 37809 69 7 , , , 37809 69 8 _ _ NNP 37809 69 9 nivocolimae nivocolimae NN 37809 69 10 _ _ NNP 37809 69 11 , , , 37809 69 12 _ _ NNP 37809 69 13 pallidus pallidus NNP 37809 69 14 _ _ NNP 37809 69 15 and and CC 37809 69 16 _ _ NNP 37809 69 17 rubrimaculatus rubrimaculatus NNP 37809 69 18 _ _ NNP 37809 69 19 ; ; : 37809 69 20 one one CD 37809 69 21 , , , 37809 69 22 _ _ NNP 37809 69 23 S. S. NNP 37809 70 1 longipes longipes NNPS 37809 70 2 _ _ NNP 37809 70 3 , , , 37809 70 4 is be VBZ 37809 70 5 relatively relatively RB 37809 70 6 large large JJ 37809 70 7 , , , 37809 70 8 and and CC 37809 70 9 the the DT 37809 70 10 remaining remain VBG 37809 70 11 eight eight CD 37809 70 12 species specie NNS 37809 70 13 are be VBP 37809 70 14 intermediate intermediate JJ 37809 70 15 in in IN 37809 70 16 size size NN 37809 70 17 ( ( -LRB- 37809 70 18 22 22 CD 37809 70 19 - - SYM 37809 70 20 30 30 CD 37809 70 21 mm mm NNP 37809 70 22 . . . 37809 70 23 ) ) -RRB- 37809 70 24 . . . 37809 71 1 Males male NNS 37809 71 2 are be VBP 37809 71 3 generally generally RB 37809 71 4 smaller small JJR 37809 71 5 than than IN 37809 71 6 females female NNS 37809 71 7 and and CC 37809 71 8 have have VBP 37809 71 9 proportionately proportionately RB 37809 71 10 longer long JJR 37809 71 11 heads head NNS 37809 71 12 and and CC 37809 71 13 usually usually RB 37809 71 14 larger large JJR 37809 71 15 tympani tympani NNS 37809 71 16 . . . 37809 72 1 No no DT 37809 72 2 significant significant JJ 37809 72 3 differences difference NNS 37809 72 4 were be VBD 37809 72 5 found find VBN 37809 72 6 among among IN 37809 72 7 proportions proportion NNS 37809 72 8 , , , 37809 72 9 except except IN 37809 72 10 that that IN 37809 72 11 _ _ NNP 37809 72 12 S. S. NNP 37809 73 1 longipes longipes NNP 37809 73 2 _ _ NNP 37809 73 3 has have VBZ 37809 73 4 a a DT 37809 73 5 larger large JJR 37809 73 6 tympanum tympanum NN 37809 73 7 / / SYM 37809 73 8 eye eye NN 37809 73 9 ratio ratio NN 37809 73 10 than than IN 37809 73 11 any any DT 37809 73 12 other other JJ 37809 73 13 species specie NNS 37809 73 14 . . . 37809 74 1 Frogs frog NNS 37809 74 2 in in IN 37809 74 3 the the DT 37809 74 4 _ _ NNP 37809 74 5 Syrrhophus Syrrhophus NNP 37809 74 6 marnockii marnockii NNS 37809 74 7 _ _ NNP 37809 74 8 group group NN 37809 74 9 tend tend VBP 37809 74 10 to to TO 37809 74 11 have have VB 37809 74 12 shorter short JJR 37809 74 13 shanks shank NNS 37809 74 14 and and CC 37809 74 15 feet foot NNS 37809 74 16 , , , 37809 74 17 thereby thereby RB 37809 74 18 giving give VBG 37809 74 19 those those DT 37809 74 20 species specie NNS 37809 74 21 a a DT 37809 74 22 more more RBR 37809 74 23 stocky stocky JJ 37809 74 24 appearance appearance NN 37809 74 25 . . . 37809 75 1 However however RB 37809 75 2 , , , 37809 75 3 the the DT 37809 75 4 differences difference NNS 37809 75 5 are be VBP 37809 75 6 not not RB 37809 75 7 significant significant JJ 37809 75 8 . . . 37809 76 1 A a DT 37809 76 2 summary summary NN 37809 76 3 of of IN 37809 76 4 the the DT 37809 76 5 data datum NNS 37809 76 6 on on IN 37809 76 7 size size NN 37809 76 8 and and CC 37809 76 9 proportions proportion NNS 37809 76 10 for for IN 37809 76 11 the the DT 37809 76 12 frogs frog NNS 37809 76 13 of of IN 37809 76 14 the the DT 37809 76 15 genus genus NNP 37809 76 16 _ _ NNP 37809 76 17 Syrrhophus Syrrhophus NNP 37809 76 18 _ _ NNP 37809 76 19 is be VBZ 37809 76 20 given give VBN 37809 76 21 in in IN 37809 76 22 Tables Tables NNP 37809 76 23 4 4 CD 37809 76 24 , , , 37809 76 25 5 5 CD 37809 76 26 , , , 37809 76 27 and and CC 37809 76 28 6 6 CD 37809 76 29 . . . 37809 77 1 _ _ NNP 37809 77 2 Hands Hands NNP 37809 77 3 and and CC 37809 77 4 Feet._--Taylor Feet._--Taylor NNP 37809 77 5 and and CC 37809 77 6 Smith Smith NNP 37809 77 7 ( ( -LRB- 37809 77 8 1945 1945 CD 37809 77 9 ) ) -RRB- 37809 77 10 , , , 37809 77 11 Smith Smith NNP 37809 77 12 and and CC 37809 77 13 Taylor Taylor NNP 37809 77 14 ( ( -LRB- 37809 77 15 1948 1948 CD 37809 77 16 ) ) -RRB- 37809 77 17 , , , 37809 77 18 Firschein Firschein NNP 37809 77 19 ( ( -LRB- 37809 77 20 1954 1954 CD 37809 77 21 ) ) -RRB- 37809 77 22 and and CC 37809 77 23 Duellman Duellman NNP 37809 77 24 ( ( -LRB- 37809 77 25 1958 1958 CD 37809 77 26 ) ) -RRB- 37809 77 27 discussed discuss VBD 37809 77 28 the the DT 37809 77 29 value value NN 37809 77 30 of of IN 37809 77 31 the the DT 37809 77 32 palmar palmar NNP 37809 77 33 tubercles tubercle NNS 37809 77 34 in in IN 37809 77 35 identifying identify VBG 37809 77 36 frogs frog NNS 37809 77 37 of of IN 37809 77 38 this this DT 37809 77 39 genus genus NN 37809 77 40 . . . 37809 78 1 The the DT 37809 78 2 eastern eastern JJ 37809 78 3 complex complex NN 37809 78 4 in in IN 37809 78 5 general general JJ 37809 78 6 has have VBZ 37809 78 7 a a DT 37809 78 8 well well RB 37809 78 9 - - HYPH 37809 78 10 developed develop VBN 37809 78 11 outer outer JJ 37809 78 12 palmar palmar NNP 37809 78 13 tubercle tubercle NN 37809 78 14 ( ( -LRB- 37809 78 15 Fig fig NN 37809 78 16 . . . 37809 79 1 1 1 LS 37809 79 2 ) ) -RRB- 37809 79 3 in in IN 37809 79 4 distinction distinction NN 37809 79 5 to to IN 37809 79 6 the the DT 37809 79 7 western western JJ 37809 79 8 complex complex NN 37809 79 9 in in IN 37809 79 10 which which WDT 37809 79 11 the the DT 37809 79 12 outer outer JJ 37809 79 13 palmar palmar NNP 37809 79 14 tubercle tubercle NN 37809 79 15 is be VBZ 37809 79 16 reduced reduce VBN 37809 79 17 or or CC 37809 79 18 absent absent JJ 37809 79 19 ( ( -LRB- 37809 79 20 Fig fig NN 37809 79 21 . . . 37809 80 1 2 2 LS 37809 80 2 ) ) -RRB- 37809 80 3 . . . 37809 81 1 Dixon Dixon NNP 37809 81 2 and and CC 37809 81 3 Webb Webb NNP 37809 81 4 ( ( -LRB- 37809 81 5 1966 1966 CD 37809 81 6 ) ) -RRB- 37809 81 7 imply imply VBP 37809 81 8 that that IN 37809 81 9 the the DT 37809 81 10 outer outer JJ 37809 81 11 palmar palmar NNP 37809 81 12 tubercle tubercle NN 37809 81 13 is be VBZ 37809 81 14 rarely rarely RB 37809 81 15 absent absent JJ 37809 81 16 but but CC 37809 81 17 is be VBZ 37809 81 18 usually usually RB 37809 81 19 smaller small JJR 37809 81 20 than than IN 37809 81 21 the the DT 37809 81 22 first first JJ 37809 81 23 supernumerary supernumerary JJ 37809 81 24 tubercle tubercle NN 37809 81 25 of of IN 37809 81 26 the the DT 37809 81 27 fourth fourth JJ 37809 81 28 finger finger NN 37809 81 29 . . . 37809 82 1 My -PRON- PRP$ 37809 82 2 study study NN 37809 82 3 of of IN 37809 82 4 the the DT 37809 82 5 western western JJ 37809 82 6 species species NN 37809 82 7 demonstrates demonstrate VBZ 37809 82 8 that that IN 37809 82 9 the the DT 37809 82 10 outer outer JJ 37809 82 11 palmar palmar NNP 37809 82 12 tubercle tubercle NN 37809 82 13 is be VBZ 37809 82 14 indeed indeed RB 37809 82 15 usually usually RB 37809 82 16 present present JJ 37809 82 17 and and CC 37809 82 18 smaller small JJR 37809 82 19 than than IN 37809 82 20 the the DT 37809 82 21 first first JJ 37809 82 22 supernumerary supernumerary JJ 37809 82 23 tubercle tubercle NN 37809 82 24 . . . 37809 83 1 Differences difference NNS 37809 83 2 in in IN 37809 83 3 interpretation interpretation NN 37809 83 4 of of IN 37809 83 5 the the DT 37809 83 6 terms term NNS 37809 83 7 " " `` 37809 83 8 unexpanded unexpanded JJ 37809 83 9 " " '' 37809 83 10 and and CC 37809 83 11 " " `` 37809 83 12 narrow narrow JJ 37809 83 13 , , , 37809 83 14 " " '' 37809 83 15 as as RB 37809 83 16 well well RB 37809 83 17 as as IN 37809 83 18 differences difference NNS 37809 83 19 in in IN 37809 83 20 techniques technique NNS 37809 83 21 of of IN 37809 83 22 preservation preservation NN 37809 83 23 , , , 37809 83 24 have have VBP 37809 83 25 led lead VBN 37809 83 26 to to IN 37809 83 27 confusion confusion NN 37809 83 28 of of IN 37809 83 29 the the DT 37809 83 30 reported report VBN 37809 83 31 digital digital JJ 37809 83 32 shapes shape NNS 37809 83 33 in in IN 37809 83 34 various various JJ 37809 83 35 species specie NNS 37809 83 36 . . . 37809 84 1 Constant constant JJ 37809 84 2 specific specific JJ 37809 84 3 differences difference NNS 37809 84 4 are be VBP 37809 84 5 evident evident JJ 37809 84 6 in in IN 37809 84 7 the the DT 37809 84 8 hands hand NNS 37809 84 9 ( ( -LRB- 37809 84 10 Fig fig NN 37809 84 11 . . . 37809 85 1 1 1 LS 37809 85 2 ) ) -RRB- 37809 85 3 . . . 37809 86 1 Except except IN 37809 86 2 in in IN 37809 86 3 the the DT 37809 86 4 cases case NNS 37809 86 5 of of IN 37809 86 6 excessive excessive JJ 37809 86 7 uptake uptake NN 37809 86 8 of of IN 37809 86 9 fluids fluid NNS 37809 86 10 , , , 37809 86 11 all all DT 37809 86 12 species specie NNS 37809 86 13 have have VBP 37809 86 14 a a DT 37809 86 15 terminal terminal JJ 37809 86 16 transverse transverse JJ 37809 86 17 groove groove NN 37809 86 18 at at IN 37809 86 19 the the DT 37809 86 20 tip tip NN 37809 86 21 of of IN 37809 86 22 each each DT 37809 86 23 digit digit NN 37809 86 24 . . . 37809 87 1 Taylor Taylor NNP 37809 87 2 ( ( -LRB- 37809 87 3 1940b 1940b CD 37809 87 4 ) ) -RRB- 37809 87 5 stated state VBD 37809 87 6 that that IN 37809 87 7 _ _ NNP 37809 87 8 S. S. NNP 37809 88 1 smithi smithi NNP 37809 88 2 _ _ NNP 37809 88 3 lacked lack VBD 37809 88 4 grooves groove NNS 37809 88 5 , , , 37809 88 6 but but CC 37809 88 7 examination examination NN 37809 88 8 of of IN 37809 88 9 the the DT 37809 88 10 holotype holotype NN 37809 88 11 reveals reveal VBZ 37809 88 12 faint faint JJ 37809 88 13 grooves groove NNS 37809 88 14 at at IN 37809 88 15 the the DT 37809 88 16 tops top NNS 37809 88 17 of of IN 37809 88 18 the the DT 37809 88 19 digits digit NNS 37809 88 20 . . . 37809 89 1 _ _ NNP 37809 89 2 Syrrhophus Syrrhophus NNP 37809 89 3 guttilatus guttilatus NN 37809 89 4 _ _ NNP 37809 89 5 , , , 37809 89 6 _ _ NNP 37809 89 7 leprus leprus NNP 37809 89 8 _ _ NNP 37809 89 9 , , , 37809 89 10 _ _ NNP 37809 89 11 pipilans pipilans NNPS 37809 89 12 _ _ NNP 37809 89 13 , , , 37809 89 14 and and CC 37809 89 15 _ _ NNP 37809 89 16 verrucipes verrucipe NNS 37809 89 17 _ _ NNP 37809 89 18 lack lack VBP 37809 89 19 lateral lateral JJ 37809 89 20 fringes fringe NNS 37809 89 21 on on IN 37809 89 22 the the DT 37809 89 23 fingers finger NNS 37809 89 24 . . . 37809 90 1 Lateral lateral JJ 37809 90 2 fringes fringe NNS 37809 90 3 are be VBP 37809 90 4 well well RB 37809 90 5 developed develop VBN 37809 90 6 in in IN 37809 90 7 the the DT 37809 90 8 _ _ NNP 37809 90 9 longipes longipe NNS 37809 90 10 _ _ NNP 37809 90 11 and and CC 37809 90 12 _ _ NNP 37809 90 13 modestus modestus NN 37809 90 14 _ _ NNP 37809 90 15 groups group NNS 37809 90 16 but but CC 37809 90 17 poorly poorly RB 37809 90 18 defined define VBN 37809 90 19 or or CC 37809 90 20 absent absent JJ 37809 90 21 in in IN 37809 90 22 the the DT 37809 90 23 other other JJ 37809 90 24 members member NNS 37809 90 25 of of IN 37809 90 26 the the DT 37809 90 27 genus genus NN 37809 90 28 . . . 37809 91 1 The the DT 37809 91 2 digital digital JJ 37809 91 3 pads pad NNS 37809 91 4 of of IN 37809 91 5 the the DT 37809 91 6 frogs frog NNS 37809 91 7 of of IN 37809 91 8 the the DT 37809 91 9 _ _ NNP 37809 91 10 longipes longipes NNPS 37809 91 11 _ _ NNP 37809 91 12 group group NN 37809 91 13 are be VBP 37809 91 14 much much RB 37809 91 15 broader broad JJR 37809 91 16 than than IN 37809 91 17 those those DT 37809 91 18 of of IN 37809 91 19 the the DT 37809 91 20 other other JJ 37809 91 21 species specie NNS 37809 91 22 and and CC 37809 91 23 are be VBP 37809 91 24 narrowest narrow JJS 37809 91 25 in in IN 37809 91 26 the the DT 37809 91 27 frogs frog NNS 37809 91 28 of of IN 37809 91 29 the the DT 37809 91 30 _ _ NNP 37809 91 31 leprus leprus NNP 37809 91 32 _ _ NNP 37809 91 33 group group NN 37809 91 34 . . . 37809 92 1 Supernumerary supernumerary JJ 37809 92 2 tubercles tubercle NNS 37809 92 3 are be VBP 37809 92 4 present present JJ 37809 92 5 on on IN 37809 92 6 the the DT 37809 92 7 palmar palmar NN 37809 92 8 surfaces surface NNS 37809 92 9 of of IN 37809 92 10 all all DT 37809 92 11 species specie NNS 37809 92 12 of of IN 37809 92 13 the the DT 37809 92 14 genus genus NN 37809 92 15 . . . 37809 93 1 TABLE TABLE NNP 37809 93 2 4 4 CD 37809 93 3 - - HYPH 37809 93 4 -Size -size CD 37809 93 5 and and CC 37809 93 6 Proportions Proportions NNPS 37809 93 7 in in IN 37809 93 8 the the DT 37809 93 9 Frogs Frogs NNP 37809 93 10 of of IN 37809 93 11 the the DT 37809 93 12 _ _ NNP 37809 93 13 Syrrhophus Syrrhophus NNP 37809 93 14 leprus leprus NN 37809 93 15 _ _ NNP 37809 93 16 Group Group NNP 37809 93 17 . . . 37809 94 1 A a DT 37809 94 2 : : : 37809 94 3 _ _ NNP 37809 94 4 cystignathoides cystignathoide VBZ 37809 94 5 campi campi NNP 37809 94 6 _ _ NNP 37809 94 7 B B NNP 37809 94 8 : : : 37809 94 9 _ _ NNP 37809 94 10 c. c. NN 37809 95 1 cystignathoides cystignathoide NNS 37809 95 2 _ _ NNP 37809 95 3 C C NNP 37809 95 4 : : : 37809 95 5 _ _ NNP 37809 95 6 leprus leprus NNP 37809 95 7 _ _ NNP 37809 95 8 D D NNP 37809 95 9 : : : 37809 95 10 _ _ NNP 37809 95 11 rubrimaculatus rubrimaculatus NNP 37809 95 12 _ _ NNP 37809 95 13 = = SYM 37809 95 14 = = SYM 37809 95 15 = = SYM 37809 95 16 = = SYM 37809 95 17 = = SYM 37809 95 18 = = SYM 37809 95 19 = = SYM 37809 95 20 = = SYM 37809 95 21 = = SYM 37809 95 22 = = SYM 37809 95 23 = = SYM 37809 95 24 = = SYM 37809 95 25 = = SYM 37809 95 26 = = SYM 37809 95 27 = = SYM 37809 95 28 = = SYM 37809 95 29 = = SYM 37809 95 30 = = SYM 37809 95 31 = = SYM 37809 95 32 = = SYM 37809 95 33 = = SYM 37809 95 34 = = SYM 37809 95 35 = = SYM 37809 95 36 = = SYM 37809 95 37 = = SYM 37809 95 38 = = SYM 37809 95 39 = = SYM 37809 95 40 = = SYM 37809 95 41 = = SYM 37809 95 42 = = SYM 37809 95 43 = = SYM 37809 95 44 = = SYM 37809 95 45 = = SYM 37809 95 46 = = SYM 37809 95 47 = = SYM 37809 95 48 = = SYM 37809 95 49 = = SYM 37809 95 50 = = SYM 37809 95 51 = = SYM 37809 95 52 = = SYM 37809 95 53 = = SYM 37809 95 54 = = SYM 37809 95 55 = = SYM 37809 95 56 = = SYM 37809 95 57 = = SYM 37809 95 58 = = SYM 37809 95 59 = = SYM 37809 95 60 = = SYM 37809 95 61 = = SYM 37809 95 62 = = SYM 37809 95 63 = = SYM 37809 95 64 = = SYM 37809 95 65 = = SYM 37809 95 66 = = SYM 37809 95 67 = = SYM 37809 95 68 = = SYM 37809 95 69 = = SYM 37809 95 70 = = SYM 37809 95 71 = = SYM 37809 95 72 = = SYM 37809 95 73 = = SYM 37809 95 74 = = SYM 37809 95 75 = = SYM 37809 95 76 = = SYM 37809 95 77 = = SYM 37809 95 78 = = SYM 37809 95 79 = = SYM 37809 95 80 = = SYM 37809 95 81 = = SYM 37809 95 82 = = SYM 37809 95 83 = = SYM 37809 95 84 = = SYM 37809 95 85 = = NFP 37809 95 86 = = SYM 37809 95 87 Snout snout NN 37809 95 88 - - HYPH 37809 95 89 vent vent NN 37809 95 90 Tibia Tibia NNP 37809 95 91 Head Head NNP 37809 95 92 Tympanum/ tympanum/ PRP 37809 95 93 Eyelid/ Eyelid/ -LRB- 37809 95 94 length length NN 37809 95 95 length/ length/ NN 37809 95 96 width/ width/ NN 37809 95 97 Eye Eye NNP 37809 95 98 Interorbital Interorbital NNP 37809 95 99 Species Species NNP 37809 95 100 Sex Sex NNP 37809 95 101 N N NNP 37809 95 102 ( ( -LRB- 37809 95 103 SVL SVL NNP 37809 95 104 ) ) -RRB- 37809 95 105 SVL SVL NNP 37809 95 106 SVL SVL NNP 37809 95 107 -------------------------------------------------------------------------- -------------------------------------------------------------------------- NFP 37809 95 108 A a DT 37809 95 109 [ [ -LRB- 37809 95 110 M m NN 37809 95 111 ] ] -RRB- 37809 95 112 33 33 CD 37809 95 113 16.3 16.3 CD 37809 95 114 - - SYM 37809 95 115 23.5 23.5 CD 37809 95 116 41.3 41.3 CD 37809 95 117 - - SYM 37809 95 118 49.6 49.6 CD 37809 95 119 34.0 34.0 CD 37809 95 120 - - SYM 37809 95 121 40.1 40.1 CD 37809 95 122 43.7 43.7 CD 37809 95 123 - - CD 37809 95 124 66.5 66.5 CD 37809 95 125 43.2 43.2 CD 37809 95 126 - - HYPH 37809 95 127 89.6 89.6 CD 37809 95 128 ( ( -LRB- 37809 95 129 45.8 45.8 CD 37809 95 130 ) ) -RRB- 37809 95 131 ( ( -LRB- 37809 95 132 37.0 37.0 CD 37809 95 133 ) ) -RRB- 37809 95 134 ( ( -LRB- 37809 95 135 56.2 56.2 CD 37809 95 136 ) ) -RRB- 37809 95 137 ( ( -LRB- 37809 95 138 61.5 61.5 CD 37809 95 139 ) ) -RRB- 37809 95 140 [ [ -LRB- 37809 95 141 F f NN 37809 95 142 ] ] -RRB- 37809 95 143 12 12 CD 37809 95 144 16.0 16.0 CD 37809 95 145 - - SYM 37809 95 146 25.8 25.8 CD 37809 95 147 41.5 41.5 CD 37809 95 148 - - SYM 37809 95 149 51.0 51.0 CD 37809 95 150 33.0 33.0 CD 37809 95 151 - - HYPH 37809 95 152 38.0 38.0 CD 37809 95 153 42.8 42.8 CD 37809 95 154 - - SYM 37809 95 155 60.0 60.0 CD 37809 95 156 48.2 48.2 CD 37809 95 157 - - HYPH 37809 95 158 69.2 69.2 CD 37809 95 159 ( ( -LRB- 37809 95 160 45.8 45.8 CD 37809 95 161 ) ) -RRB- 37809 95 162 ( ( -LRB- 37809 95 163 35.0 35.0 CD 37809 95 164 ) ) -RRB- 37809 95 165 ( ( -LRB- 37809 95 166 51.2 51.2 CD 37809 95 167 ) ) -RRB- 37809 95 168 ( ( -LRB- 37809 95 169 60.1 60.1 CD 37809 95 170 ) ) -RRB- 37809 95 171 B b NN 37809 95 172 [ [ -LRB- 37809 95 173 M M NNP 37809 95 174 ] ] -RRB- 37809 95 175 15 15 CD 37809 95 176 16.8 16.8 CD 37809 95 177 - - HYPH 37809 95 178 22.1 22.1 CD 37809 95 179 45.1 45.1 CD 37809 95 180 - - HYPH 37809 95 181 50.4 50.4 CD 37809 95 182 33.2 33.2 CD 37809 95 183 - - HYPH 37809 95 184 40.7 40.7 CD 37809 95 185 44.3 44.3 CD 37809 95 186 - - HYPH 37809 95 187 68.7 68.7 CD 37809 95 188 44.6 44.6 CD 37809 95 189 - - SYM 37809 95 190 65.4 65.4 CD 37809 95 191 ( ( -LRB- 37809 95 192 47.3 47.3 CD 37809 95 193 ) ) -RRB- 37809 95 194 ( ( -LRB- 37809 95 195 37.8 37.8 CD 37809 95 196 ) ) -RRB- 37809 95 197 ( ( -LRB- 37809 95 198 54.8 54.8 CD 37809 95 199 ) ) -RRB- 37809 95 200 ( ( -LRB- 37809 95 201 60.0 60.0 CD 37809 95 202 ) ) -RRB- 37809 95 203 [ [ -LRB- 37809 95 204 F f NN 37809 95 205 ] ] -RRB- 37809 95 206 6 6 CD 37809 95 207 19.6 19.6 CD 37809 95 208 - - HYPH 37809 95 209 24.2 24.2 CD 37809 95 210 46.4 46.4 CD 37809 95 211 - - HYPH 37809 95 212 50.0 50.0 CD 37809 95 213 34.1 34.1 CD 37809 95 214 - - SYM 37809 95 215 38.1 38.1 CD 37809 95 216 43.3 43.3 CD 37809 95 217 - - HYPH 37809 95 218 56.5 56.5 CD 37809 95 219 53.2 53.2 CD 37809 95 220 - - HYPH 37809 95 221 65.4 65.4 CD 37809 95 222 ( ( -LRB- 37809 95 223 47.6 47.6 CD 37809 95 224 ) ) -RRB- 37809 95 225 ( ( -LRB- 37809 95 226 36.2 36.2 CD 37809 95 227 ) ) -RRB- 37809 95 228 ( ( -LRB- 37809 95 229 46.9 46.9 CD 37809 95 230 ) ) -RRB- 37809 95 231 ( ( -LRB- 37809 95 232 59.2 59.2 CD 37809 95 233 ) ) -RRB- 37809 95 234 C C NNP 37809 95 235 [ [ -LRB- 37809 95 236 M m NN 37809 95 237 ] ] -RRB- 37809 95 238 14 14 CD 37809 95 239 20.6 20.6 CD 37809 95 240 - - SYM 37809 95 241 26.4 26.4 CD 37809 95 242 42.3 42.3 CD 37809 95 243 - - SYM 37809 95 244 52.3 52.3 CD 37809 95 245 35.0 35.0 CD 37809 95 246 - - HYPH 37809 95 247 40.3 40.3 CD 37809 95 248 47.5 47.5 CD 37809 95 249 - - SYM 37809 95 250 62.5 62.5 CD 37809 95 251 58.2 58.2 CD 37809 95 252 - - HYPH 37809 95 253 72.5 72.5 CD 37809 95 254 ( ( -LRB- 37809 95 255 46.8 46.8 CD 37809 95 256 ) ) -RRB- 37809 95 257 ( ( -LRB- 37809 95 258 37.4 37.4 CD 37809 95 259 ) ) -RRB- 37809 95 260 ( ( -LRB- 37809 95 261 56.5 56.5 CD 37809 95 262 ) ) -RRB- 37809 95 263 ( ( -LRB- 37809 95 264 67.3 67.3 CD 37809 95 265 ) ) -RRB- 37809 95 266 [ [ -LRB- 37809 95 267 F f NN 37809 95 268 ] ] -RRB- 37809 95 269 15 15 CD 37809 95 270 22.1 22.1 CD 37809 95 271 - - HYPH 37809 95 272 29.2 29.2 CD 37809 95 273 43.4 43.4 CD 37809 95 274 - - SYM 37809 95 275 53.3 53.3 CD 37809 95 276 32.6 32.6 CD 37809 95 277 - - HYPH 37809 95 278 38.9 38.9 CD 37809 95 279 38.6 38.6 CD 37809 95 280 - - SYM 37809 95 281 57.9 57.9 CD 37809 95 282 50.2 50.2 CD 37809 95 283 - - HYPH 37809 95 284 86.9 86.9 CD 37809 95 285 ( ( -LRB- 37809 95 286 47.1 47.1 CD 37809 95 287 ) ) -RRB- 37809 95 288 ( ( -LRB- 37809 95 289 35.8 35.8 CD 37809 95 290 ) ) -RRB- 37809 95 291 ( ( -LRB- 37809 95 292 47.1 47.1 CD 37809 95 293 ) ) -RRB- 37809 95 294 ( ( -LRB- 37809 95 295 68.1 68.1 CD 37809 95 296 ) ) -RRB- 37809 95 297 D D NNP 37809 95 298 [ [ -LRB- 37809 95 299 M m NN 37809 95 300 ] ] -RRB- 37809 95 301 12 12 CD 37809 95 302 18.2 18.2 CD 37809 95 303 - - SYM 37809 95 304 23.5 23.5 CD 37809 95 305 40.4 40.4 CD 37809 95 306 - - SYM 37809 95 307 46.2 46.2 CD 37809 95 308 31.8 31.8 CD 37809 95 309 - - SYM 37809 95 310 35.5 35.5 CD 37809 95 311 35.5 35.5 CD 37809 95 312 - - HYPH 37809 95 313 46.5 46.5 CD 37809 95 314 65.1 65.1 CD 37809 95 315 - - HYPH 37809 95 316 78.5 78.5 CD 37809 95 317 ( ( -LRB- 37809 95 318 43.4 43.4 CD 37809 95 319 ) ) -RRB- 37809 95 320 ( ( -LRB- 37809 95 321 33.8 33.8 CD 37809 95 322 ) ) -RRB- 37809 95 323 ( ( -LRB- 37809 95 324 41.7 41.7 CD 37809 95 325 ) ) -RRB- 37809 95 326 ( ( -LRB- 37809 95 327 71.7 71.7 CD 37809 95 328 ) ) -RRB- 37809 95 329 = = NFP 37809 95 330 = = NFP 37809 95 331 = = SYM 37809 95 332 = = SYM 37809 95 333 = = SYM 37809 95 334 = = SYM 37809 95 335 = = SYM 37809 95 336 = = SYM 37809 95 337 = = SYM 37809 95 338 = = SYM 37809 95 339 = = SYM 37809 95 340 = = SYM 37809 95 341 = = SYM 37809 95 342 = = SYM 37809 95 343 = = SYM 37809 95 344 = = SYM 37809 95 345 = = SYM 37809 95 346 = = SYM 37809 95 347 = = SYM 37809 95 348 = = SYM 37809 95 349 = = SYM 37809 95 350 = = SYM 37809 95 351 = = SYM 37809 95 352 = = SYM 37809 95 353 = = SYM 37809 95 354 = = SYM 37809 95 355 = = SYM 37809 95 356 = = SYM 37809 95 357 = = SYM 37809 95 358 = = SYM 37809 95 359 = = SYM 37809 95 360 = = SYM 37809 95 361 = = SYM 37809 95 362 = = SYM 37809 95 363 = = SYM 37809 95 364 = = SYM 37809 95 365 = = SYM 37809 95 366 = = SYM 37809 95 367 = = SYM 37809 95 368 = = SYM 37809 95 369 = = SYM 37809 95 370 = = SYM 37809 95 371 = = SYM 37809 95 372 = = SYM 37809 95 373 = = SYM 37809 95 374 = = SYM 37809 95 375 = = SYM 37809 95 376 = = SYM 37809 95 377 = = SYM 37809 95 378 = = SYM 37809 95 379 = = SYM 37809 95 380 = = SYM 37809 95 381 = = SYM 37809 95 382 = = SYM 37809 95 383 = = SYM 37809 95 384 = = SYM 37809 95 385 = = SYM 37809 95 386 = = SYM 37809 95 387 = = SYM 37809 95 388 = = SYM 37809 95 389 = = SYM 37809 95 390 = = SYM 37809 95 391 = = SYM 37809 95 392 = = SYM 37809 95 393 = = SYM 37809 95 394 = = SYM 37809 95 395 = = SYM 37809 95 396 = = SYM 37809 95 397 = = SYM 37809 95 398 = = SYM 37809 95 399 = = SYM 37809 95 400 = = SYM 37809 95 401 = = NFP 37809 95 402 = = NFP 37809 95 403 TABLE TABLE NNP 37809 95 404 5 5 CD 37809 95 405 - - HYPH 37809 95 406 -Size -size CD 37809 95 407 and and CC 37809 95 408 Proportions Proportions NNPS 37809 95 409 in in IN 37809 95 410 the the DT 37809 95 411 Frogs Frogs NNP 37809 95 412 of of IN 37809 95 413 the the DT 37809 95 414 _ _ NNP 37809 95 415 Syrrhophus Syrrhophus NNP 37809 95 416 longipes longipe VBZ 37809 95 417 _ _ NNP 37809 95 418 and and CC 37809 95 419 _ _ NNP 37809 95 420 S. S. NNP 37809 96 1 marnockii marnockii NNP 37809 96 2 _ _ NNP 37809 96 3 Groups Groups NNPS 37809 96 4 . . . 37809 97 1 A a DT 37809 97 2 : : : 37809 97 3 _ _ NNP 37809 97 4 dennisi dennisi NNP 37809 97 5 _ _ NNP 37809 97 6 B B NNP 37809 97 7 : : : 37809 97 8 _ _ NNP 37809 97 9 longipes longipe NNS 37809 97 10 _ _ NNP 37809 97 11 C C NNP 37809 97 12 : : : 37809 97 13 _ _ NNP 37809 97 14 guttilatus guttilatus NN 37809 97 15 _ _ NNP 37809 97 16 D D NNP 37809 97 17 : : : 37809 97 18 _ _ NNP 37809 97 19 marnockii marnockii NNS 37809 97 20 _ _ NNP 37809 97 21 E E NNP 37809 97 22 : : : 37809 97 23 _ _ NNP 37809 97 24 verrucipes verrucipes NNP 37809 97 25 _ _ NNP 37809 97 26 = = SYM 37809 97 27 = = SYM 37809 97 28 = = SYM 37809 97 29 = = SYM 37809 97 30 = = SYM 37809 97 31 = = SYM 37809 97 32 = = SYM 37809 97 33 = = SYM 37809 97 34 = = SYM 37809 97 35 = = SYM 37809 97 36 = = SYM 37809 97 37 = = SYM 37809 97 38 = = SYM 37809 97 39 = = SYM 37809 97 40 = = SYM 37809 97 41 = = SYM 37809 97 42 = = SYM 37809 97 43 = = SYM 37809 97 44 = = SYM 37809 97 45 = = SYM 37809 97 46 = = SYM 37809 97 47 = = SYM 37809 97 48 = = SYM 37809 97 49 = = SYM 37809 97 50 = = SYM 37809 97 51 = = SYM 37809 97 52 = = SYM 37809 97 53 = = SYM 37809 97 54 = = SYM 37809 97 55 = = SYM 37809 97 56 = = SYM 37809 97 57 = = SYM 37809 97 58 = = SYM 37809 97 59 = = SYM 37809 97 60 = = SYM 37809 97 61 = = SYM 37809 97 62 = = SYM 37809 97 63 = = SYM 37809 97 64 = = SYM 37809 97 65 = = SYM 37809 97 66 = = SYM 37809 97 67 = = SYM 37809 97 68 = = SYM 37809 97 69 = = SYM 37809 97 70 = = SYM 37809 97 71 = = SYM 37809 97 72 = = SYM 37809 97 73 = = SYM 37809 97 74 = = SYM 37809 97 75 = = SYM 37809 97 76 = = SYM 37809 97 77 = = SYM 37809 97 78 = = SYM 37809 97 79 = = SYM 37809 97 80 = = SYM 37809 97 81 = = SYM 37809 97 82 = = SYM 37809 97 83 = = SYM 37809 97 84 = = SYM 37809 97 85 = = SYM 37809 97 86 = = SYM 37809 97 87 = = SYM 37809 97 88 = = SYM 37809 97 89 = = SYM 37809 97 90 = = SYM 37809 97 91 = = SYM 37809 97 92 = = SYM 37809 97 93 = = SYM 37809 97 94 = = SYM 37809 97 95 = = SYM 37809 97 96 = = SYM 37809 97 97 = = SYM 37809 97 98 = = NFP 37809 97 99 = = SYM 37809 97 100 Snout snout NN 37809 97 101 - - HYPH 37809 97 102 vent vent NN 37809 97 103 Tibia Tibia NNP 37809 97 104 Head Head NNP 37809 97 105 Tympanum/ tympanum/ PRP 37809 97 106 Eyelid/ Eyelid/ -LRB- 37809 97 107 length length NN 37809 97 108 length/ length/ NN 37809 97 109 width/ width/ NN 37809 97 110 Eye Eye NNP 37809 97 111 Interorbital Interorbital NNP 37809 97 112 Species Species NNP 37809 97 113 Sex Sex NNP 37809 97 114 N N NNP 37809 97 115 ( ( -LRB- 37809 97 116 SVL SVL NNP 37809 97 117 ) ) -RRB- 37809 97 118 SVL SVL NNP 37809 97 119 SVL SVL NNP 37809 97 120 -------------------------------------------------------------------------- -------------------------------------------------------------------------- NFP 37809 97 121 A a DT 37809 97 122 [ [ -LRB- 37809 97 123 M m NN 37809 97 124 ] ] -RRB- 37809 97 125 16 16 CD 37809 97 126 22.8 22.8 CD 37809 97 127 - - SYM 37809 97 128 28.4 28.4 CD 37809 97 129 43.9 43.9 CD 37809 97 130 - - CD 37809 97 131 49.7 49.7 CD 37809 97 132 35.3 35.3 CD 37809 97 133 - - SYM 37809 97 134 41.2 41.2 CD 37809 97 135 53.9 53.9 CD 37809 97 136 - - HYPH 37809 97 137 64.2 64.2 CD 37809 97 138 55.3 55.3 CD 37809 97 139 - - HYPH 37809 97 140 74.0 74.0 CD 37809 97 141 ( ( -LRB- 37809 97 142 47.4 47.4 CD 37809 97 143 ) ) -RRB- 37809 97 144 ( ( -LRB- 37809 97 145 38.8 38.8 CD 37809 97 146 ) ) -RRB- 37809 97 147 ( ( -LRB- 37809 97 148 58.9 58.9 CD 37809 97 149 ) ) -RRB- 37809 97 150 ( ( -LRB- 37809 97 151 65.1 65.1 CD 37809 97 152 ) ) -RRB- 37809 97 153 [ [ -LRB- 37809 97 154 F f NN 37809 97 155 ] ] -RRB- 37809 97 156 10 10 CD 37809 97 157 25.9 25.9 CD 37809 97 158 - - HYPH 37809 97 159 32.0 32.0 CD 37809 97 160 46.3 46.3 CD 37809 97 161 - - HYPH 37809 97 162 50.8 50.8 CD 37809 97 163 35.6 35.6 CD 37809 97 164 - - HYPH 37809 97 165 40.3 40.3 CD 37809 97 166 50.6 50.6 CD 37809 97 167 - - HYPH 37809 97 168 58.7 58.7 CD 37809 97 169 58.1 58.1 CD 37809 97 170 - - HYPH 37809 97 171 70.9 70.9 CD 37809 97 172 ( ( -LRB- 37809 97 173 48.2 48.2 CD 37809 97 174 ) ) -RRB- 37809 97 175 ( ( -LRB- 37809 97 176 37.7 37.7 CD 37809 97 177 ) ) -RRB- 37809 97 178 ( ( -LRB- 37809 97 179 54.9 54.9 CD 37809 97 180 ) ) -RRB- 37809 97 181 ( ( -LRB- 37809 97 182 63.6 63.6 CD 37809 97 183 ) ) -RRB- 37809 97 184 B b NN 37809 97 185 [ [ -LRB- 37809 97 186 M m NN 37809 97 187 ] ] -RRB- 37809 97 188 22 22 CD 37809 97 189 22.1 22.1 CD 37809 97 190 - - SYM 37809 97 191 33.2 33.2 CD 37809 97 192 45.8 45.8 CD 37809 97 193 - - HYPH 37809 97 194 51.7 51.7 CD 37809 97 195 38.7 38.7 CD 37809 97 196 - - HYPH 37809 97 197 44.4 44.4 CD 37809 97 198 61.1 61.1 CD 37809 97 199 - - SYM 37809 97 200 87.2 87.2 CD 37809 97 201 61.5 61.5 CD 37809 97 202 - - HYPH 37809 97 203 83.0 83.0 CD 37809 97 204 ( ( -LRB- 37809 97 205 48.4 48.4 CD 37809 97 206 ) ) -RRB- 37809 97 207 ( ( -LRB- 37809 97 208 41.8 41.8 CD 37809 97 209 ) ) -RRB- 37809 97 210 ( ( -LRB- 37809 97 211 72.0 72.0 CD 37809 97 212 ) ) -RRB- 37809 97 213 ( ( -LRB- 37809 97 214 72.0 72.0 CD 37809 97 215 ) ) -RRB- 37809 97 216 [ [ -LRB- 37809 97 217 F f NN 37809 97 218 ] ] -RRB- 37809 97 219 19 19 CD 37809 97 220 26.8 26.8 CD 37809 97 221 - - SYM 37809 97 222 39.6 39.6 CD 37809 97 223 44.3 44.3 CD 37809 97 224 - - HYPH 37809 97 225 51.0 51.0 CD 37809 97 226 36.3 36.3 CD 37809 97 227 - - HYPH 37809 97 228 40.8 40.8 CD 37809 97 229 49.5 49.5 CD 37809 97 230 - - HYPH 37809 97 231 72.1 72.1 CD 37809 97 232 55.3 55.3 CD 37809 97 233 - - HYPH 37809 97 234 85.9 85.9 CD 37809 97 235 ( ( -LRB- 37809 97 236 47.2 47.2 CD 37809 97 237 ) ) -RRB- 37809 97 238 ( ( -LRB- 37809 97 239 39.1 39.1 CD 37809 97 240 ) ) -RRB- 37809 97 241 ( ( -LRB- 37809 97 242 59.5 59.5 CD 37809 97 243 ) ) -RRB- 37809 97 244 ( ( -LRB- 37809 97 245 67.9 67.9 CD 37809 97 246 ) ) -RRB- 37809 97 247 C C NNP 37809 97 248 [ [ -LRB- 37809 97 249 M m NN 37809 97 250 ] ] -RRB- 37809 97 251 19 19 CD 37809 97 252 20.6 20.6 CD 37809 97 253 - - SYM 37809 97 254 29.0 29.0 CD 37809 97 255 41.2 41.2 CD 37809 97 256 - - SYM 37809 97 257 48.1 48.1 CD 37809 97 258 36.9 36.9 CD 37809 97 259 - - HYPH 37809 97 260 44.9 44.9 CD 37809 97 261 55.1 55.1 CD 37809 97 262 - - HYPH 37809 97 263 75.7 75.7 CD 37809 97 264 53.3 53.3 CD 37809 97 265 - - HYPH 37809 97 266 79.5 79.5 CD 37809 97 267 ( ( -LRB- 37809 97 268 44.5 44.5 CD 37809 97 269 ) ) -RRB- 37809 97 270 ( ( -LRB- 37809 97 271 40.6 40.6 CD 37809 97 272 ) ) -RRB- 37809 97 273 ( ( -LRB- 37809 97 274 64.1 64.1 CD 37809 97 275 ) ) -RRB- 37809 97 276 ( ( -LRB- 37809 97 277 66.0 66.0 CD 37809 97 278 ) ) -RRB- 37809 97 279 [ [ -LRB- 37809 97 280 F f NN 37809 97 281 ] ] -RRB- 37809 97 282 5 5 CD 37809 97 283 25.7 25.7 CD 37809 97 284 - - SYM 37809 97 285 31.0 31.0 CD 37809 97 286 41.4 41.4 CD 37809 97 287 - - HYPH 37809 97 288 46.8 46.8 CD 37809 97 289 35.9 35.9 CD 37809 97 290 - - SYM 37809 97 291 42.3 42.3 CD 37809 97 292 47.6 47.6 CD 37809 97 293 - - HYPH 37809 97 294 61.7 61.7 CD 37809 97 295 62.3 62.3 CD 37809 97 296 - - HYPH 37809 97 297 79.8 79.8 CD 37809 97 298 ( ( -LRB- 37809 97 299 43.6 43.6 CD 37809 97 300 ) ) -RRB- 37809 97 301 ( ( -LRB- 37809 97 302 38.5 38.5 CD 37809 97 303 ) ) -RRB- 37809 97 304 ( ( -LRB- 37809 97 305 54.0 54.0 CD 37809 97 306 ) ) -RRB- 37809 97 307 ( ( -LRB- 37809 97 308 72.9 72.9 CD 37809 97 309 ) ) -RRB- 37809 97 310 D D NNP 37809 97 311 [ [ -LRB- 37809 97 312 M m NN 37809 97 313 ] ] -RRB- 37809 97 314 14 14 CD 37809 97 315 18.4 18.4 CD 37809 97 316 - - SYM 37809 97 317 28.9 28.9 CD 37809 97 318 42.3 42.3 CD 37809 97 319 - - SYM 37809 97 320 47.2 47.2 CD 37809 97 321 36.1 36.1 CD 37809 97 322 - - SYM 37809 97 323 43.0 43.0 CD 37809 97 324 47.2 47.2 CD 37809 97 325 - - SYM 37809 97 326 68.3 68.3 CD 37809 97 327 51.6 51.6 CD 37809 97 328 - - SYM 37809 97 329 74.4 74.4 CD 37809 97 330 ( ( -LRB- 37809 97 331 44.1 44.1 CD 37809 97 332 ) ) -RRB- 37809 97 333 ( ( -LRB- 37809 97 334 39.6 39.6 CD 37809 97 335 ) ) -RRB- 37809 97 336 ( ( -LRB- 37809 97 337 61.2 61.2 CD 37809 97 338 ) ) -RRB- 37809 97 339 ( ( -LRB- 37809 97 340 66.3 66.3 CD 37809 97 341 ) ) -RRB- 37809 97 342 [ [ -LRB- 37809 97 343 F f NN 37809 97 344 ] ] -RRB- 37809 97 345 29 29 CD 37809 97 346 20.4 20.4 CD 37809 97 347 - - SYM 37809 97 348 35.4 35.4 CD 37809 97 349 38.7 38.7 CD 37809 97 350 - - HYPH 37809 97 351 46.4 46.4 CD 37809 97 352 35.9 35.9 CD 37809 97 353 - - SYM 37809 97 354 41.3 41.3 CD 37809 97 355 45.8 45.8 CD 37809 97 356 - - SYM 37809 97 357 73.3 73.3 CD 37809 97 358 52.1 52.1 CD 37809 97 359 - - HYPH 37809 97 360 70.5 70.5 CD 37809 97 361 ( ( -LRB- 37809 97 362 42.7 42.7 CD 37809 97 363 ) ) -RRB- 37809 97 364 ( ( -LRB- 37809 97 365 38.2 38.2 CD 37809 97 366 ) ) -RRB- 37809 97 367 ( ( -LRB- 37809 97 368 60.3 60.3 CD 37809 97 369 ) ) -RRB- 37809 97 370 ( ( -LRB- 37809 97 371 60.7 60.7 CD 37809 97 372 ) ) -RRB- 37809 97 373 E e NN 37809 97 374 [ [ -LRB- 37809 97 375 M m NN 37809 97 376 ] ] -RRB- 37809 97 377 29 29 CD 37809 97 378 17.5 17.5 CD 37809 97 379 - - SYM 37809 97 380 29.2 29.2 CD 37809 97 381 42.7 42.7 CD 37809 97 382 - - HYPH 37809 97 383 49.5 49.5 CD 37809 97 384 36.2 36.2 CD 37809 97 385 - - SYM 37809 97 386 42.4 42.4 CD 37809 97 387 56.1 56.1 CD 37809 97 388 - - HYPH 37809 97 389 82.2 82.2 CD 37809 97 390 56.8 56.8 CD 37809 97 391 - - HYPH 37809 97 392 82.8 82.8 CD 37809 97 393 ( ( -LRB- 37809 97 394 46.3 46.3 CD 37809 97 395 ) ) -RRB- 37809 97 396 ( ( -LRB- 37809 97 397 39.1 39.1 CD 37809 97 398 ) ) -RRB- 37809 97 399 ( ( -LRB- 37809 97 400 67.8 67.8 CD 37809 97 401 ) ) -RRB- 37809 97 402 ( ( -LRB- 37809 97 403 70.4 70.4 CD 37809 97 404 ) ) -RRB- 37809 97 405 [ [ -LRB- 37809 97 406 F f NN 37809 97 407 ] ] -RRB- 37809 97 408 6 6 CD 37809 97 409 26.5 26.5 CD 37809 97 410 - - SYM 37809 97 411 31.7 31.7 CD 37809 97 412 42.4 42.4 CD 37809 97 413 - - SYM 37809 97 414 47.7 47.7 CD 37809 97 415 36.0 36.0 CD 37809 97 416 - - SYM 37809 97 417 38.1 38.1 CD 37809 97 418 45.8 45.8 CD 37809 97 419 - - SYM 37809 97 420 57.8 57.8 CD 37809 97 421 61.0 61.0 CD 37809 97 422 - - HYPH 37809 97 423 77.9 77.9 CD 37809 97 424 ( ( -LRB- 37809 97 425 44.6 44.6 CD 37809 97 426 ) ) -RRB- 37809 97 427 ( ( -LRB- 37809 97 428 37.0 37.0 CD 37809 97 429 ) ) -RRB- 37809 97 430 ( ( -LRB- 37809 97 431 53.9 53.9 CD 37809 97 432 ) ) -RRB- 37809 97 433 ( ( -LRB- 37809 97 434 69.0 69.0 CD 37809 97 435 ) ) -RRB- 37809 97 436 = = NFP 37809 97 437 = = NFP 37809 97 438 = = SYM 37809 97 439 = = SYM 37809 97 440 = = SYM 37809 97 441 = = SYM 37809 97 442 = = SYM 37809 97 443 = = SYM 37809 97 444 = = SYM 37809 97 445 = = SYM 37809 97 446 = = SYM 37809 97 447 = = SYM 37809 97 448 = = SYM 37809 97 449 = = SYM 37809 97 450 = = SYM 37809 97 451 = = SYM 37809 97 452 = = SYM 37809 97 453 = = SYM 37809 97 454 = = SYM 37809 97 455 = = SYM 37809 97 456 = = SYM 37809 97 457 = = SYM 37809 97 458 = = SYM 37809 97 459 = = SYM 37809 97 460 = = SYM 37809 97 461 = = SYM 37809 97 462 = = SYM 37809 97 463 = = SYM 37809 97 464 = = SYM 37809 97 465 = = SYM 37809 97 466 = = SYM 37809 97 467 = = SYM 37809 97 468 = = SYM 37809 97 469 = = SYM 37809 97 470 = = SYM 37809 97 471 = = SYM 37809 97 472 = = SYM 37809 97 473 = = SYM 37809 97 474 = = SYM 37809 97 475 = = SYM 37809 97 476 = = SYM 37809 97 477 = = SYM 37809 97 478 = = SYM 37809 97 479 = = SYM 37809 97 480 = = SYM 37809 97 481 = = SYM 37809 97 482 = = SYM 37809 97 483 = = SYM 37809 97 484 = = SYM 37809 97 485 = = SYM 37809 97 486 = = SYM 37809 97 487 = = SYM 37809 97 488 = = SYM 37809 97 489 = = SYM 37809 97 490 = = SYM 37809 97 491 = = SYM 37809 97 492 = = SYM 37809 97 493 = = SYM 37809 97 494 = = SYM 37809 97 495 = = SYM 37809 97 496 = = SYM 37809 97 497 = = SYM 37809 97 498 = = SYM 37809 97 499 = = SYM 37809 97 500 = = SYM 37809 97 501 = = SYM 37809 97 502 = = SYM 37809 97 503 = = SYM 37809 97 504 = = SYM 37809 97 505 = = SYM 37809 97 506 = = SYM 37809 97 507 = = SYM 37809 97 508 = = NFP 37809 97 509 = = NFP 37809 97 510 TABLE TABLE NNP 37809 97 511 6 6 CD 37809 97 512 - - HYPH 37809 97 513 -Size -size CD 37809 97 514 and and CC 37809 97 515 Proportions Proportions NNPS 37809 97 516 in in IN 37809 97 517 the the DT 37809 97 518 Frogs Frogs NNP 37809 97 519 of of IN 37809 97 520 the the DT 37809 97 521 _ _ NNP 37809 97 522 Syrrhophus Syrrhophus NNP 37809 97 523 pipilans pipilan NNS 37809 97 524 _ _ NNP 37809 97 525 and and CC 37809 97 526 _ _ NNP 37809 97 527 S. S. NNP 37809 98 1 modestus modestus NNP 37809 98 2 _ _ NNP 37809 98 3 Groups Groups NNPS 37809 98 4 . . . 37809 99 1 A a DT 37809 99 2 : : : 37809 99 3 _ _ NNP 37809 99 4 pipilans pipilans NNPS 37809 99 5 nebulosus nebulosus NN 37809 99 6 _ _ NNP 37809 99 7 B B NNP 37809 99 8 : : : 37809 99 9 _ _ NNP 37809 99 10 pipilans pipilans NNPS 37809 99 11 pipilans pipilans NNPS 37809 99 12 _ _ NNP 37809 99 13 C C NNP 37809 99 14 : : : 37809 99 15 _ _ NNP 37809 99 16 modestus modestus NN 37809 99 17 _ _ NNP 37809 99 18 D D NNP 37809 99 19 : : : 37809 99 20 _ _ NNP 37809 99 21 pallidus pallidus NNP 37809 99 22 _ _ NNP 37809 99 23 E E NNP 37809 99 24 : : : 37809 99 25 _ _ NNP 37809 99 26 teretistes teretiste NNS 37809 99 27 _ _ NNP 37809 99 28 F F NNP 37809 99 29 : : : 37809 99 30 _ _ NNP 37809 99 31 nivocolimae nivocolimae NN 37809 99 32 _ _ NNP 37809 99 33 G G NNP 37809 99 34 : : : 37809 99 35 _ _ NNP 37809 99 36 interorbitalis interorbitalis NNP 37809 99 37 _ _ NNP 37809 99 38 = = SYM 37809 99 39 = = SYM 37809 99 40 = = SYM 37809 99 41 = = SYM 37809 99 42 = = SYM 37809 99 43 = = SYM 37809 99 44 = = SYM 37809 99 45 = = SYM 37809 99 46 = = SYM 37809 99 47 = = SYM 37809 99 48 = = SYM 37809 99 49 = = SYM 37809 99 50 = = SYM 37809 99 51 = = SYM 37809 99 52 = = SYM 37809 99 53 = = SYM 37809 99 54 = = SYM 37809 99 55 = = SYM 37809 99 56 = = SYM 37809 99 57 = = SYM 37809 99 58 = = SYM 37809 99 59 = = SYM 37809 99 60 = = SYM 37809 99 61 = = SYM 37809 99 62 = = SYM 37809 99 63 = = SYM 37809 99 64 = = SYM 37809 99 65 = = SYM 37809 99 66 = = SYM 37809 99 67 = = SYM 37809 99 68 = = SYM 37809 99 69 = = SYM 37809 99 70 = = SYM 37809 99 71 = = SYM 37809 99 72 = = SYM 37809 99 73 = = SYM 37809 99 74 = = SYM 37809 99 75 = = SYM 37809 99 76 = = SYM 37809 99 77 = = SYM 37809 99 78 = = SYM 37809 99 79 = = SYM 37809 99 80 = = SYM 37809 99 81 = = SYM 37809 99 82 = = SYM 37809 99 83 = = SYM 37809 99 84 = = SYM 37809 99 85 = = SYM 37809 99 86 = = SYM 37809 99 87 = = SYM 37809 99 88 = = SYM 37809 99 89 = = SYM 37809 99 90 = = SYM 37809 99 91 = = SYM 37809 99 92 = = SYM 37809 99 93 = = SYM 37809 99 94 = = SYM 37809 99 95 = = SYM 37809 99 96 = = SYM 37809 99 97 = = SYM 37809 99 98 = = SYM 37809 99 99 = = SYM 37809 99 100 = = SYM 37809 99 101 = = SYM 37809 99 102 = = SYM 37809 99 103 = = SYM 37809 99 104 = = SYM 37809 99 105 = = SYM 37809 99 106 = = SYM 37809 99 107 = = SYM 37809 99 108 = = SYM 37809 99 109 = = SYM 37809 99 110 = = NFP 37809 99 111 = = SYM 37809 99 112 Snout snout NN 37809 99 113 - - HYPH 37809 99 114 vent vent NN 37809 99 115 Tibia Tibia NNP 37809 99 116 Head Head NNP 37809 99 117 Tympanum/ tympanum/ PRP 37809 99 118 Eyelid/ Eyelid/ -LRB- 37809 99 119 length length NN 37809 99 120 length/ length/ NN 37809 99 121 width/ width/ NN 37809 99 122 Eye Eye NNP 37809 99 123 Interorbital Interorbital NNP 37809 99 124 Species Species NNP 37809 99 125 Sex Sex NNP 37809 99 126 N N NNP 37809 99 127 ( ( -LRB- 37809 99 128 SVL SVL NNP 37809 99 129 ) ) -RRB- 37809 99 130 SVL SVL NNP 37809 99 131 SVL SVL NNP 37809 99 132 -------------------------------------------------------------------------- -------------------------------------------------------------------------- NFP 37809 99 133 A a DT 37809 99 134 [ [ -LRB- 37809 99 135 M m NN 37809 99 136 ] ] -RRB- 37809 99 137 17 17 CD 37809 99 138 22.9 22.9 CD 37809 99 139 - - HYPH 37809 99 140 28.5 28.5 CD 37809 99 141 38.1 38.1 CD 37809 99 142 - - SYM 37809 99 143 42.0 42.0 CD 37809 99 144 34.4 34.4 CD 37809 99 145 - - HYPH 37809 99 146 37.2 37.2 CD 37809 99 147 36.6 36.6 CD 37809 99 148 - - HYPH 37809 99 149 47.8 47.8 CD 37809 99 150 56.1 56.1 CD 37809 99 151 - - HYPH 37809 99 152 82.4 82.4 CD 37809 99 153 ( ( -LRB- 37809 99 154 40.0 40.0 CD 37809 99 155 ) ) -RRB- 37809 99 156 ( ( -LRB- 37809 99 157 35.4 35.4 CD 37809 99 158 ) ) -RRB- 37809 99 159 ( ( -LRB- 37809 99 160 43.6 43.6 CD 37809 99 161 ) ) -RRB- 37809 99 162 ( ( -LRB- 37809 99 163 68.2 68.2 CD 37809 99 164 ) ) -RRB- 37809 99 165 [ [ -LRB- 37809 99 166 F f NN 37809 99 167 ] ] -RRB- 37809 99 168 3 3 CD 37809 99 169 21.1 21.1 CD 37809 99 170 - - SYM 37809 99 171 22.7 22.7 CD 37809 99 172 42.1 42.1 CD 37809 99 173 - - CD 37809 99 174 44.5 44.5 CD 37809 99 175 33.2 33.2 CD 37809 99 176 - - SYM 37809 99 177 35.8 35.8 CD 37809 99 178 36.6 36.6 CD 37809 99 179 - - HYPH 37809 99 180 47.6 47.6 CD 37809 99 181 64.3 64.3 CD 37809 99 182 - - HYPH 37809 99 183 65.4 65.4 CD 37809 99 184 B b NN 37809 99 185 [ [ -LRB- 37809 99 186 M M NNP 37809 99 187 ] ] -RRB- 37809 99 188 18 18 CD 37809 99 189 22.6 22.6 CD 37809 99 190 - - HYPH 37809 99 191 27.8 27.8 CD 37809 99 192 37.9 37.9 CD 37809 99 193 - - SYM 37809 99 194 44.0 44.0 CD 37809 99 195 32.2 32.2 CD 37809 99 196 - - SYM 37809 99 197 36.5 36.5 CD 37809 99 198 38.0 38.0 CD 37809 99 199 - - HYPH 37809 99 200 54.0 54.0 CD 37809 99 201 56.1 56.1 CD 37809 99 202 - - HYPH 37809 99 203 79.5 79.5 CD 37809 99 204 ( ( -LRB- 37809 99 205 41.4 41.4 CD 37809 99 206 ) ) -RRB- 37809 99 207 ( ( -LRB- 37809 99 208 33.0 33.0 CD 37809 99 209 ) ) -RRB- 37809 99 210 ( ( -LRB- 37809 99 211 46.2 46.2 CD 37809 99 212 ) ) -RRB- 37809 99 213 ( ( -LRB- 37809 99 214 67.3 67.3 CD 37809 99 215 ) ) -RRB- 37809 99 216 [ [ -LRB- 37809 99 217 F f NN 37809 99 218 ] ] -RRB- 37809 99 219 1 1 CD 37809 99 220 29.4 29.4 CD 37809 99 221 38.4 38.4 CD 37809 99 222 32.5 32.5 CD 37809 99 223 44.6 44.6 CD 37809 99 224 55.0 55.0 CD 37809 99 225 C C NNP 37809 99 226 [ [ -LRB- 37809 99 227 M m NN 37809 99 228 ] ] -RRB- 37809 99 229 8 8 CD 37809 99 230 15.8 15.8 CD 37809 99 231 - - SYM 37809 99 232 20.1 20.1 CD 37809 99 233 38.5 38.5 CD 37809 99 234 - - HYPH 37809 99 235 42.6 42.6 CD 37809 99 236 32.1 32.1 CD 37809 99 237 - - SYM 37809 99 238 38.1 38.1 CD 37809 99 239 26.8 26.8 CD 37809 99 240 - - HYPH 37809 99 241 39.3 39.3 CD 37809 99 242 57.0 57.0 CD 37809 99 243 - - HYPH 37809 99 244 86.9 86.9 CD 37809 99 245 ( ( -LRB- 37809 99 246 40.6 40.6 CD 37809 99 247 ) ) -RRB- 37809 99 248 ( ( -LRB- 37809 99 249 34.2 34.2 CD 37809 99 250 ) ) -RRB- 37809 99 251 ( ( -LRB- 37809 99 252 31.5 31.5 CD 37809 99 253 ) ) -RRB- 37809 99 254 ( ( -LRB- 37809 99 255 69.1 69.1 CD 37809 99 256 ) ) -RRB- 37809 99 257 [ [ -LRB- 37809 99 258 F f NN 37809 99 259 ] ] -RRB- 37809 99 260 1 1 CD 37809 99 261 18.5 18.5 CD 37809 99 262 44.2 44.2 CD 37809 99 263 36.0 36.0 CD 37809 99 264 24.0 24.0 CD 37809 99 265 52.1 52.1 CD 37809 99 266 D d NN 37809 99 267 [ [ -LRB- 37809 99 268 M m NN 37809 99 269 ] ] -RRB- 37809 99 270 6 6 CD 37809 99 271 17.9 17.9 CD 37809 99 272 - - SYM 37809 99 273 19.3 19.3 CD 37809 99 274 41.0 41.0 CD 37809 99 275 - - SYM 37809 99 276 44.9 44.9 CD 37809 99 277 32.6 32.6 CD 37809 99 278 - - SYM 37809 99 279 36.2 36.2 CD 37809 99 280 27.0 27.0 CD 37809 99 281 - - SYM 37809 99 282 35.6 35.6 CD 37809 99 283 59.4 59.4 CD 37809 99 284 - - HYPH 37809 99 285 67.7 67.7 CD 37809 99 286 ( ( -LRB- 37809 99 287 43.4 43.4 CD 37809 99 288 ) ) -RRB- 37809 99 289 ( ( -LRB- 37809 99 290 35.2 35.2 CD 37809 99 291 ) ) -RRB- 37809 99 292 ( ( -LRB- 37809 99 293 30.9 30.9 CD 37809 99 294 ) ) -RRB- 37809 99 295 ( ( -LRB- 37809 99 296 65.2 65.2 CD 37809 99 297 ) ) -RRB- 37809 99 298 E e NN 37809 99 299 [ [ -LRB- 37809 99 300 M m NN 37809 99 301 ] ] -RRB- 37809 99 302 18 18 CD 37809 99 303 19.2 19.2 CD 37809 99 304 - - HYPH 37809 99 305 23.2 23.2 CD 37809 99 306 41.5 41.5 CD 37809 99 307 - - SYM 37809 99 308 45.3 45.3 CD 37809 99 309 32.5 32.5 CD 37809 99 310 - - SYM 37809 99 311 36.4 36.4 CD 37809 99 312 28.6 28.6 CD 37809 99 313 - - HYPH 37809 99 314 43.8 43.8 CD 37809 99 315 51.2 51.2 CD 37809 99 316 - - HYPH 37809 99 317 75.0 75.0 CD 37809 99 318 ( ( -LRB- 37809 99 319 43.7 43.7 CD 37809 99 320 ) ) -RRB- 37809 99 321 ( ( -LRB- 37809 99 322 34.0 34.0 CD 37809 99 323 ) ) -RRB- 37809 99 324 ( ( -LRB- 37809 99 325 33.7 33.7 CD 37809 99 326 ) ) -RRB- 37809 99 327 ( ( -LRB- 37809 99 328 62.2 62.2 CD 37809 99 329 ) ) -RRB- 37809 99 330 [ [ -LRB- 37809 99 331 F f NN 37809 99 332 ] ] -RRB- 37809 99 333 1 1 CD 37809 99 334 24.8 24.8 CD 37809 99 335 41.8 41.8 CD 37809 99 336 30.8 30.8 CD 37809 99 337 37.9 37.9 CD 37809 99 338 60.5 60.5 CD 37809 99 339 F F NNP 37809 99 340 [ [ -LRB- 37809 99 341 M M NNP 37809 99 342 ] ] -RRB- 37809 99 343 15 15 CD 37809 99 344 18.9 18.9 CD 37809 99 345 - - SYM 37809 99 346 21.1 21.1 CD 37809 99 347 42.2 42.2 CD 37809 99 348 - - SYM 37809 99 349 48.6 48.6 CD 37809 99 350 30.9 30.9 CD 37809 99 351 - - HYPH 37809 99 352 37.1 37.1 CD 37809 99 353 30.0 30.0 CD 37809 99 354 - - SYM 37809 99 355 39.3 39.3 CD 37809 99 356 42.6 42.6 CD 37809 99 357 - - HYPH 37809 99 358 69.1 69.1 CD 37809 99 359 ( ( -LRB- 37809 99 360 45.0 45.0 CD 37809 99 361 ) ) -RRB- 37809 99 362 ( ( -LRB- 37809 99 363 33.7 33.7 CD 37809 99 364 ) ) -RRB- 37809 99 365 ( ( -LRB- 37809 99 366 34.7 34.7 CD 37809 99 367 ) ) -RRB- 37809 99 368 ( ( -LRB- 37809 99 369 55.0 55.0 CD 37809 99 370 ) ) -RRB- 37809 99 371 [ [ -LRB- 37809 99 372 F f NN 37809 99 373 ] ] -RRB- 37809 99 374 1 1 CD 37809 99 375 24.1 24.1 CD 37809 99 376 40.9 40.9 CD 37809 99 377 33.5 33.5 CD 37809 99 378 27.6 27.6 CD 37809 99 379 56.5 56.5 CD 37809 99 380 G g NN 37809 99 381 [ [ -LRB- 37809 99 382 M m NN 37809 99 383 ] ] -RRB- 37809 99 384 1 1 CD 37809 99 385 25.6 25.6 CD 37809 99 386 43.0 43.0 CD 37809 99 387 ---- ---- CD 37809 99 388 39.4 39.4 CD 37809 99 389 57.6 57.6 CD 37809 99 390 [ [ -LRB- 37809 99 391 F f NN 37809 99 392 ] ] -RRB- 37809 99 393 9 9 CD 37809 99 394 20.2 20.2 CD 37809 99 395 - - HYPH 37809 99 396 26.7 26.7 CD 37809 99 397 39.9 39.9 CD 37809 99 398 - - SYM 37809 99 399 47.1 47.1 CD 37809 99 400 32.6 32.6 CD 37809 99 401 - - SYM 37809 99 402 39.3 39.3 CD 37809 99 403 29.1 29.1 CD 37809 99 404 - - SYM 37809 99 405 41.2 41.2 CD 37809 99 406 58.2 58.2 CD 37809 99 407 - - HYPH 37809 99 408 76.9 76.9 CD 37809 99 409 ( ( -LRB- 37809 99 410 43.2 43.2 CD 37809 99 411 ) ) -RRB- 37809 99 412 ( ( -LRB- 37809 99 413 35.8 35.8 CD 37809 99 414 ) ) -RRB- 37809 99 415 ( ( -LRB- 37809 99 416 36.4 36.4 CD 37809 99 417 ) ) -RRB- 37809 99 418 ( ( -LRB- 37809 99 419 69.2 69.2 CD 37809 99 420 ) ) -RRB- 37809 99 421 = = NFP 37809 99 422 = = NFP 37809 99 423 = = SYM 37809 99 424 = = SYM 37809 99 425 = = SYM 37809 99 426 = = SYM 37809 99 427 = = SYM 37809 99 428 = = SYM 37809 99 429 = = SYM 37809 99 430 = = SYM 37809 99 431 = = SYM 37809 99 432 = = SYM 37809 99 433 = = SYM 37809 99 434 = = SYM 37809 99 435 = = SYM 37809 99 436 = = SYM 37809 99 437 = = SYM 37809 99 438 = = SYM 37809 99 439 = = SYM 37809 99 440 = = SYM 37809 99 441 = = SYM 37809 99 442 = = SYM 37809 99 443 = = SYM 37809 99 444 = = SYM 37809 99 445 = = SYM 37809 99 446 = = SYM 37809 99 447 = = SYM 37809 99 448 = = SYM 37809 99 449 = = SYM 37809 99 450 = = SYM 37809 99 451 = = SYM 37809 99 452 = = SYM 37809 99 453 = = SYM 37809 99 454 = = SYM 37809 99 455 = = SYM 37809 99 456 = = SYM 37809 99 457 = = SYM 37809 99 458 = = SYM 37809 99 459 = = SYM 37809 99 460 = = SYM 37809 99 461 = = SYM 37809 99 462 = = SYM 37809 99 463 = = SYM 37809 99 464 = = SYM 37809 99 465 = = SYM 37809 99 466 = = SYM 37809 99 467 = = SYM 37809 99 468 = = SYM 37809 99 469 = = SYM 37809 99 470 = = SYM 37809 99 471 = = SYM 37809 99 472 = = SYM 37809 99 473 = = SYM 37809 99 474 = = SYM 37809 99 475 = = SYM 37809 99 476 = = SYM 37809 99 477 = = SYM 37809 99 478 = = SYM 37809 99 479 = = SYM 37809 99 480 = = SYM 37809 99 481 = = SYM 37809 99 482 = = SYM 37809 99 483 = = SYM 37809 99 484 = = SYM 37809 99 485 = = SYM 37809 99 486 = = SYM 37809 99 487 = = SYM 37809 99 488 = = SYM 37809 99 489 = = SYM 37809 99 490 = = SYM 37809 99 491 = = SYM 37809 99 492 = = NFP 37809 99 493 = = NFP 37809 99 494 = = NFP 37809 99 495 [ [ -LRB- 37809 99 496 Illustration illustration NN 37809 99 497 : : : 37809 99 498 FIG FIG NNP 37809 99 499 . . . 37809 100 1 1 1 LS 37809 100 2 : : : 37809 100 3 Palmar Palmar NNP 37809 100 4 views view NNS 37809 100 5 of of IN 37809 100 6 hands hand NNS 37809 100 7 of of IN 37809 100 8 six six CD 37809 100 9 species specie NNS 37809 100 10 of of IN 37809 100 11 the the DT 37809 100 12 eastern eastern JJ 37809 100 13 complex complex NN 37809 100 14 of of IN 37809 100 15 _ _ NNP 37809 100 16 Syrrhophus Syrrhophus NNP 37809 100 17 _ _ NNP 37809 100 18 . . . 37809 101 1 ( ( -LRB- 37809 101 2 A a DT 37809 101 3 ) ) -RRB- 37809 101 4 _ _ NNP 37809 101 5 verrucipes verrucipe NNS 37809 101 6 _ _ NNP 37809 101 7 ( ( -LRB- 37809 101 8 UIMNH UIMNH NNP 37809 101 9 15995 15995 CD 37809 101 10 ) ) -RRB- 37809 101 11 , , , 37809 101 12 ( ( -LRB- 37809 101 13 B b NN 37809 101 14 ) ) -RRB- 37809 101 15 _ _ NNP 37809 101 16 rubrimaculatus rubrimaculatus NN 37809 101 17 _ _ NNP 37809 101 18 ( ( -LRB- 37809 101 19 KU KU NNP 37809 101 20 58911 58911 CD 37809 101 21 ) ) -RRB- 37809 101 22 , , , 37809 101 23 ( ( -LRB- 37809 101 24 C C NNP 37809 101 25 ) ) -RRB- 37809 101 26 _ _ NNP 37809 101 27 dennisi dennisi NNP 37809 101 28 _ _ NNP 37809 101 29 sp sp NNP 37809 101 30 . . . 37809 102 1 nov nov NNP 37809 102 2 . . NNP 37809 102 3 ( ( -LRB- 37809 102 4 holotype holotype NN 37809 102 5 , , , 37809 102 6 UMMZ UMMZ NNP 37809 102 7 101121 101121 CD 37809 102 8 ) ) -RRB- 37809 102 9 , , , 37809 102 10 ( ( -LRB- 37809 102 11 D D NNP 37809 102 12 ) ) -RRB- 37809 102 13 _ _ NNP 37809 102 14 guttilatus guttilatus NN 37809 102 15 _ _ NNP 37809 102 16 ( ( -LRB- 37809 102 17 UIMNH UIMNH NNP 37809 102 18 55520 55520 CD 37809 102 19 ) ) -RRB- 37809 102 20 , , , 37809 102 21 ( ( -LRB- 37809 102 22 E e NN 37809 102 23 ) ) -RRB- 37809 102 24 _ _ NNP 37809 102 25 marnockii marnockii NNS 37809 102 26 _ _ NNP 37809 102 27 ( ( -LRB- 37809 102 28 TCWC TCWC NNP 37809 102 29 4782 4782 CD 37809 102 30 ) ) -RRB- 37809 102 31 , , , 37809 102 32 and and CC 37809 102 33 ( ( -LRB- 37809 102 34 F F NNP 37809 102 35 ) ) -RRB- 37809 102 36 _ _ NNP 37809 102 37 longipes longipe NNS 37809 102 38 _ _ NNP 37809 102 39 ( ( -LRB- 37809 102 40 TCWC TCWC NNP 37809 102 41 12179 12179 CD 37809 102 42 ) ) -RRB- 37809 102 43 . . . 37809 103 1 All all DT 37809 103 2 ×6.5 ×6.5 NNP 37809 103 3 . . . 37809 103 4 ] ] -RRB- 37809 104 1 [ [ -LRB- 37809 104 2 Illustration illustration NN 37809 104 3 : : : 37809 104 4 FIG FIG NNP 37809 104 5 . . . 37809 105 1 2 2 LS 37809 105 2 : : : 37809 105 3 Palmar Palmar NNP 37809 105 4 views view NNS 37809 105 5 of of IN 37809 105 6 hands hand NNS 37809 105 7 of of IN 37809 105 8 two two CD 37809 105 9 species specie NNS 37809 105 10 of of IN 37809 105 11 the the DT 37809 105 12 western western JJ 37809 105 13 complex complex NN 37809 105 14 of of IN 37809 105 15 _ _ NNP 37809 105 16 Syrrhophus Syrrhophus NNP 37809 105 17 _ _ NNP 37809 105 18 . . . 37809 106 1 _ _ NNP 37809 106 2 pipilans pipilans NNPS 37809 106 3 _ _ NNP 37809 106 4 ( ( -LRB- 37809 106 5 left leave VBN 37809 106 6 , , , 37809 106 7 KU KU NNP 37809 106 8 58908 58908 CD 37809 106 9 , , , 37809 106 10 ×6 ×6 XX 37809 106 11 ) ) -RRB- 37809 106 12 and and CC 37809 106 13 _ _ NNP 37809 106 14 teretistes teretistes NNPS 37809 106 15 _ _ NNP 37809 106 16 ( ( -LRB- 37809 106 17 center center NN 37809 106 18 , , , 37809 106 19 KU KU NNP 37809 106 20 75269 75269 CD 37809 106 21 , , , 37809 106 22 and and CC 37809 106 23 right right UH 37809 106 24 , , , 37809 106 25 KU KU NNP 37809 106 26 75263 75263 CD 37809 106 27 , , , 37809 106 28 respectively respectively RB 37809 106 29 , , , 37809 106 30 ×9 ×9 LS 37809 106 31 ) ) -RRB- 37809 106 32 . . . 37809 106 33 ] ] -RRB- 37809 107 1 In in IN 37809 107 2 _ _ NNP 37809 107 3 S. S. NNP 37809 108 1 cystignathoides cystignathoide VBZ 37809 108 2 _ _ NNP 37809 108 3 and and CC 37809 108 4 _ _ NNP 37809 108 5 leprus leprus NNP 37809 108 6 _ _ NNP 37809 108 7 , , , 37809 108 8 the the DT 37809 108 9 first first JJ 37809 108 10 finger finger NN 37809 108 11 is be VBZ 37809 108 12 longer long JJR 37809 108 13 than than IN 37809 108 14 the the DT 37809 108 15 second second JJ 37809 108 16 , , , 37809 108 17 and and CC 37809 108 18 the the DT 37809 108 19 first first JJ 37809 108 20 two two CD 37809 108 21 fingers finger NNS 37809 108 22 are be VBP 37809 108 23 equal equal JJ 37809 108 24 in in IN 37809 108 25 length length NN 37809 108 26 in in IN 37809 108 27 _ _ NNP 37809 108 28 guttilatus guttilatus NN 37809 108 29 _ _ NNP 37809 108 30 and and CC 37809 108 31 _ _ NNP 37809 108 32 marnockii marnockii NNS 37809 108 33 _ _ NNP 37809 108 34 . . . 37809 109 1 In in IN 37809 109 2 the the DT 37809 109 3 other other JJ 37809 109 4 species specie NNS 37809 109 5 the the DT 37809 109 6 first first JJ 37809 109 7 finger finger NN 37809 109 8 is be VBZ 37809 109 9 shorter short JJR 37809 109 10 than than IN 37809 109 11 the the DT 37809 109 12 second second JJ 37809 109 13 . . . 37809 110 1 Supernumerary supernumerary JJ 37809 110 2 tubercles tubercle NNS 37809 110 3 are be VBP 37809 110 4 well well RB 37809 110 5 developed developed JJ 37809 110 6 on on IN 37809 110 7 the the DT 37809 110 8 plantar plantar NN 37809 110 9 surfaces surface NNS 37809 110 10 in in IN 37809 110 11 all all DT 37809 110 12 species specie NNS 37809 110 13 , , , 37809 110 14 except except IN 37809 110 15 _ _ NNP 37809 110 16 S. S. NNP 37809 111 1 guttilatus guttilatus NNP 37809 111 2 _ _ NNP 37809 111 3 , , , 37809 111 4 in in IN 37809 111 5 which which WDT 37809 111 6 they -PRON- PRP 37809 111 7 are be VBP 37809 111 8 poorly poorly RB 37809 111 9 defined define VBN 37809 111 10 ( ( -LRB- 37809 111 11 Fig fig NN 37809 111 12 . . . 37809 112 1 3 3 LS 37809 112 2 ) ) -RRB- 37809 112 3 . . . 37809 113 1 The the DT 37809 113 2 relative relative JJ 37809 113 3 sizes size NNS 37809 113 4 of of IN 37809 113 5 the the DT 37809 113 6 metatarsal metatarsal NN 37809 113 7 tubercles tubercle NNS 37809 113 8 has have VBZ 37809 113 9 been be VBN 37809 113 10 used use VBN 37809 113 11 in in IN 37809 113 12 the the DT 37809 113 13 classification classification NN 37809 113 14 of of IN 37809 113 15 the the DT 37809 113 16 species specie NNS 37809 113 17 and and CC 37809 113 18 species specie NNS 37809 113 19 groups group NNS 37809 113 20 of of IN 37809 113 21 _ _ NNP 37809 113 22 Syrrhophus Syrrhophus NNP 37809 113 23 _ _ NNP 37809 113 24 . . . 37809 114 1 The the DT 37809 114 2 metatarsal metatarsal NN 37809 114 3 tubercles tubercle NNS 37809 114 4 are be VBP 37809 114 5 similar similar JJ 37809 114 6 in in IN 37809 114 7 all all DT 37809 114 8 species specie NNS 37809 114 9 of of IN 37809 114 10 the the DT 37809 114 11 eastern eastern JJ 37809 114 12 complex complex NN 37809 114 13 ( ( -LRB- 37809 114 14 including include VBG 37809 114 15 _ _ NNP 37809 114 16 rubrimaculatus rubrimaculatus NN 37809 114 17 _ _ NNP 37809 114 18 ) ) -RRB- 37809 114 19 ; ; : 37809 114 20 the the DT 37809 114 21 outer outer JJ 37809 114 22 tubercle tubercle NN 37809 114 23 is be VBZ 37809 114 24 always always RB 37809 114 25 about about RB 37809 114 26 one one CD 37809 114 27 - - HYPH 37809 114 28 half half NN 37809 114 29 the the DT 37809 114 30 size size NN 37809 114 31 of of IN 37809 114 32 the the DT 37809 114 33 ovoid ovoid NNP 37809 114 34 inner inner JJ 37809 114 35 metatarsal metatarsal NN 37809 114 36 tubercle tubercle NN 37809 114 37 . . . 37809 115 1 In in IN 37809 115 2 the the DT 37809 115 3 _ _ NNP 37809 115 4 leprus leprus NNP 37809 115 5 _ _ NNP 37809 115 6 group group NN 37809 115 7 the the DT 37809 115 8 outer outer JJ 37809 115 9 tubercle tubercle NN 37809 115 10 is be VBZ 37809 115 11 conical conical JJ 37809 115 12 and and CC 37809 115 13 compressed compressed JJ 37809 115 14 . . . 37809 116 1 The the DT 37809 116 2 metatarsal metatarsal NN 37809 116 3 tubercles tubercle NNS 37809 116 4 of of IN 37809 116 5 _ _ NNP 37809 116 6 pipilans pipilans NNPS 37809 116 7 _ _ NNP 37809 116 8 are be VBP 37809 116 9 about about IN 37809 116 10 the the DT 37809 116 11 same same JJ 37809 116 12 size size NN 37809 116 13 , , , 37809 116 14 or or CC 37809 116 15 the the DT 37809 116 16 outer outer NN 37809 116 17 is be VBZ 37809 116 18 slightly slightly RB 37809 116 19 smaller small JJR 37809 116 20 than than IN 37809 116 21 the the DT 37809 116 22 inner inner JJ 37809 116 23 . . . 37809 117 1 In in IN 37809 117 2 the the DT 37809 117 3 _ _ NNP 37809 117 4 modestus modestus NN 37809 117 5 _ _ NNP 37809 117 6 group group NN 37809 117 7 the the DT 37809 117 8 outer outer JJ 37809 117 9 metatarsal metatarsal NN 37809 117 10 tubercle tubercle NN 37809 117 11 is be VBZ 37809 117 12 about about RB 37809 117 13 one one CD 37809 117 14 - - HYPH 37809 117 15 third third NN 37809 117 16 the the DT 37809 117 17 size size NN 37809 117 18 of of IN 37809 117 19 the the DT 37809 117 20 inner inner JJ 37809 117 21 . . . 37809 118 1 All all DT 37809 118 2 species specie NNS 37809 118 3 , , , 37809 118 4 except except IN 37809 118 5 _ _ NNP 37809 118 6 guttilatus guttilatus NNP 37809 118 7 _ _ NNP 37809 118 8 , , , 37809 118 9 have have VBP 37809 118 10 well well RB 37809 118 11 - - HYPH 37809 118 12 defined define VBN 37809 118 13 to to IN 37809 118 14 poorly poorly RB 37809 118 15 defined define VBN 37809 118 16 lateral lateral JJ 37809 118 17 fringes fringe NNS 37809 118 18 on on IN 37809 118 19 the the DT 37809 118 20 toes toe NNS 37809 118 21 . . . 37809 119 1 All all DT 37809 119 2 species specie NNS 37809 119 3 have have VBP 37809 119 4 expanded expand VBN 37809 119 5 toe toe NN 37809 119 6 pads pad NNS 37809 119 7 . . . 37809 120 1 The the DT 37809 120 2 fifth fifth JJ 37809 120 3 toe toe NN 37809 120 4 is be VBZ 37809 120 5 usually usually RB 37809 120 6 shorter short JJR 37809 120 7 than than IN 37809 120 8 the the DT 37809 120 9 third third JJ 37809 120 10 , , , 37809 120 11 but but CC 37809 120 12 the the DT 37809 120 13 second second JJ 37809 120 14 is be VBZ 37809 120 15 equal equal JJ 37809 120 16 in in IN 37809 120 17 length length NN 37809 120 18 to to IN 37809 120 19 the the DT 37809 120 20 fifth fifth NN 37809 120 21 in in IN 37809 120 22 some some DT 37809 120 23 specimens specimen NNS 37809 120 24 of of IN 37809 120 25 _ _ NNP 37809 120 26 S. S. NNP 37809 121 1 cystignathoides cystignathoide VBZ 37809 121 2 _ _ NNP 37809 121 3 and and CC 37809 121 4 _ _ NNP 37809 121 5 S. S. NNP 37809 122 1 marnockii marnockii NNP 37809 122 2 _ _ NNP 37809 122 3 . . . 37809 123 1 _ _ NNP 37809 123 2 Syrrhophus Syrrhophus NNP 37809 123 3 nivocolimae nivocolimae NN 37809 123 4 _ _ NNP 37809 123 5 is be VBZ 37809 123 6 the the DT 37809 123 7 only only JJ 37809 123 8 species specie NNS 37809 123 9 with with IN 37809 123 10 tubercles tubercle NNS 37809 123 11 along along IN 37809 123 12 the the DT 37809 123 13 outer outer JJ 37809 123 14 edge edge NN 37809 123 15 of of IN 37809 123 16 the the DT 37809 123 17 tarsus tarsus NN 37809 123 18 ; ; : 37809 123 19 this this DT 37809 123 20 is be VBZ 37809 123 21 merely merely RB 37809 123 22 a a DT 37809 123 23 reflection reflection NN 37809 123 24 of of IN 37809 123 25 the the DT 37809 123 26 highly highly RB 37809 123 27 tuberculate tuberculate JJ 37809 123 28 nature nature NN 37809 123 29 of of IN 37809 123 30 the the DT 37809 123 31 skin skin NN 37809 123 32 in in IN 37809 123 33 this this DT 37809 123 34 species specie NNS 37809 123 35 . . . 37809 124 1 _ _ NNP 37809 124 2 Skin skin NN 37809 124 3 texture._--The texture._--the RB 37809 124 4 skin skin NN 37809 124 5 of of IN 37809 124 6 the the DT 37809 124 7 dorsum dorsum NN 37809 124 8 is be VBZ 37809 124 9 smooth smooth JJ 37809 124 10 or or CC 37809 124 11 very very RB 37809 124 12 weakly weakly RB 37809 124 13 pustular pustular NN 37809 124 14 in in IN 37809 124 15 all all DT 37809 124 16 species specie NNS 37809 124 17 of of IN 37809 124 18 the the DT 37809 124 19 genus genus NN 37809 124 20 except except IN 37809 124 21 _ _ NNP 37809 124 22 nivocolimae nivocolimae NN 37809 124 23 _ _ NNP 37809 124 24 and and CC 37809 124 25 _ _ NNP 37809 124 26 verrucipes verrucipe NNS 37809 124 27 _ _ NNP 37809 124 28 . . . 37809 125 1 The the DT 37809 125 2 dorsal dorsal NN 37809 125 3 surfaces surface NNS 37809 125 4 of of IN 37809 125 5 _ _ NNP 37809 125 6 nivocolimae nivocolimae NN 37809 125 7 _ _ NNP 37809 125 8 are be VBP 37809 125 9 warty warty JJ 37809 125 10 ; ; : 37809 125 11 in in IN 37809 125 12 _ _ NNP 37809 125 13 verrucipes verrucipe NNS 37809 125 14 _ _ NNP 37809 125 15 the the DT 37809 125 16 skin skin NN 37809 125 17 is be VBZ 37809 125 18 pustular pustular JJ 37809 125 19 . . . 37809 126 1 The the DT 37809 126 2 skin skin NN 37809 126 3 of of IN 37809 126 4 the the DT 37809 126 5 venter venter NN 37809 126 6 is be VBZ 37809 126 7 areolate areolate NN 37809 126 8 in in IN 37809 126 9 _ _ NNP 37809 126 10 cystignathoides cystignathoides NNP 37809 126 11 cystignathoides cystignathoide VBZ 37809 126 12 _ _ NNP 37809 126 13 , , , 37809 126 14 _ _ NNP 37809 126 15 dennisi dennisi NNP 37809 126 16 _ _ NNP 37809 126 17 and and CC 37809 126 18 _ _ NNP 37809 126 19 verrucipes verrucipe NNS 37809 126 20 _ _ NNP 37809 126 21 but but CC 37809 126 22 is be VBZ 37809 126 23 smooth smooth JJ 37809 126 24 in in IN 37809 126 25 all all DT 37809 126 26 other other JJ 37809 126 27 species specie NNS 37809 126 28 of of IN 37809 126 29 the the DT 37809 126 30 genus genus NN 37809 126 31 . . . 37809 127 1 [ [ -LRB- 37809 127 2 Illustration illustration NN 37809 127 3 : : : 37809 127 4 FIG FIG NNP 37809 127 5 . . . 37809 128 1 3 3 LS 37809 128 2 : : : 37809 128 3 Plantar Plantar NNP 37809 128 4 views view NNS 37809 128 5 of of IN 37809 128 6 feet foot NNS 37809 128 7 of of IN 37809 128 8 four four CD 37809 128 9 species specie NNS 37809 128 10 of of IN 37809 128 11 the the DT 37809 128 12 eastern eastern JJ 37809 128 13 complex complex NN 37809 128 14 of of IN 37809 128 15 _ _ NNP 37809 128 16 Syrrhophus Syrrhophus NNP 37809 128 17 _ _ NNP 37809 128 18 . . . 37809 129 1 ( ( -LRB- 37809 129 2 A a DT 37809 129 3 ) ) -RRB- 37809 129 4 _ _ NNP 37809 129 5 guttilatus guttilatus NN 37809 129 6 _ _ NNP 37809 129 7 ( ( -LRB- 37809 129 8 UIMNH UIMNH NNP 37809 129 9 55519 55519 CD 37809 129 10 , , , 37809 129 11 ×6 ×6 JJ 37809 129 12 ) ) -RRB- 37809 129 13 , , , 37809 129 14 ( ( -LRB- 37809 129 15 B B NNP 37809 129 16 ) ) -RRB- 37809 129 17 _ _ NNP 37809 129 18 leprus leprus NNP 37809 129 19 _ _ NNP 37809 129 20 ( ( -LRB- 37809 129 21 UIMNH UIMNH NNP 37809 129 22 42726 42726 CD 37809 129 23 , , , 37809 129 24 ×6 ×6 JJ 37809 129 25 ) ) -RRB- 37809 129 26 , , , 37809 129 27 ( ( -LRB- 37809 129 28 C C NNP 37809 129 29 ) ) -RRB- 37809 129 30 _ _ NNP 37809 129 31 verrucipes verrucipes NNP 37809 129 32 _ _ NNP 37809 129 33 ( ( -LRB- 37809 129 34 UIMNH UIMNH NNP 37809 129 35 15995 15995 CD 37809 129 36 , , , 37809 129 37 ×6 ×6 JJ 37809 129 38 ) ) -RRB- 37809 129 39 , , , 37809 129 40 and and CC 37809 129 41 ( ( -LRB- 37809 129 42 D D NNP 37809 129 43 ) ) -RRB- 37809 129 44 _ _ NNP 37809 129 45 longipes longipe NNS 37809 129 46 _ _ NNP 37809 129 47 ( ( -LRB- 37809 129 48 TCWC TCWC NNP 37809 129 49 12179 12179 CD 37809 129 50 , , , 37809 129 51 ×4.6 ×4.6 NNP 37809 129 52 ) ) -RRB- 37809 129 53 . . . 37809 129 54 ] ] -RRB- 37809 130 1 _ _ NNP 37809 130 2 Color Color NNP 37809 130 3 pattern._--As pattern._--As NNP 37809 130 4 is be VBZ 37809 130 5 evident evident JJ 37809 130 6 in in IN 37809 130 7 the the DT 37809 130 8 diagnoses diagnosis NNS 37809 130 9 , , , 37809 130 10 the the DT 37809 130 11 color color NN 37809 130 12 patterns pattern NNS 37809 130 13 of of IN 37809 130 14 given give VBN 37809 130 15 populations population NNS 37809 130 16 have have VBP 37809 130 17 been be VBN 37809 130 18 regarded regard VBN 37809 130 19 as as IN 37809 130 20 useful useful JJ 37809 130 21 in in IN 37809 130 22 separating separate VBG 37809 130 23 the the DT 37809 130 24 species specie NNS 37809 130 25 and and CC 37809 130 26 subspecies subspecie NNS 37809 130 27 . . . 37809 131 1 Duellman Duellman NNP 37809 131 2 ( ( -LRB- 37809 131 3 1958 1958 CD 37809 131 4 ) ) -RRB- 37809 131 5 suggested suggest VBD 37809 131 6 that that IN 37809 131 7 the the DT 37809 131 8 coloration coloration NN 37809 131 9 , , , 37809 131 10 with with IN 37809 131 11 the the DT 37809 131 12 exception exception NN 37809 131 13 of of IN 37809 131 14 _ _ NNP 37809 131 15 modestus modestus NN 37809 131 16 _ _ NNP 37809 131 17 , , , 37809 131 18 was be VBD 37809 131 19 a a DT 37809 131 20 dark dark JJ 37809 131 21 ground ground NN 37809 131 22 color color NN 37809 131 23 with with IN 37809 131 24 pale pale JJ 37809 131 25 markings marking NNS 37809 131 26 . . . 37809 132 1 It -PRON- PRP 37809 132 2 is be VBZ 37809 132 3 a a DT 37809 132 4 moot moot JJ 37809 132 5 point point NN 37809 132 6 whether whether IN 37809 132 7 the the DT 37809 132 8 frogs frog NNS 37809 132 9 have have VBP 37809 132 10 light light JJ 37809 132 11 spots spot NNS 37809 132 12 on on IN 37809 132 13 a a DT 37809 132 14 dark dark JJ 37809 132 15 background background NN 37809 132 16 or or CC 37809 132 17 have have VB 37809 132 18 a a DT 37809 132 19 light light JJ 37809 132 20 background background NN 37809 132 21 with with IN 37809 132 22 an an DT 37809 132 23 extensive extensive JJ 37809 132 24 reticulate reticulate JJ 37809 132 25 dark dark JJ 37809 132 26 pattern pattern NN 37809 132 27 . . . 37809 133 1 The the DT 37809 133 2 venters venter NNS 37809 133 3 are be VBP 37809 133 4 gray gray JJ 37809 133 5 or or CC 37809 133 6 white white JJ 37809 133 7 , , , 37809 133 8 and and CC 37809 133 9 the the DT 37809 133 10 vocal vocal JJ 37809 133 11 sac sac NN 37809 133 12 is be VBZ 37809 133 13 nearly nearly RB 37809 133 14 black black JJ 37809 133 15 in in IN 37809 133 16 some some DT 37809 133 17 species specie NNS 37809 133 18 . . . 37809 134 1 Interorbital interorbital JJ 37809 134 2 dark dark JJ 37809 134 3 bars bar NNS 37809 134 4 or or CC 37809 134 5 triangles triangle NNS 37809 134 6 are be VBP 37809 134 7 absent absent JJ 37809 134 8 in in IN 37809 134 9 only only RB 37809 134 10 two two CD 37809 134 11 species specie NNS 37809 134 12 of of IN 37809 134 13 the the DT 37809 134 14 eastern eastern JJ 37809 134 15 complex complex NN 37809 134 16 , , , 37809 134 17 _ _ NNP 37809 134 18 cystignathoides cystignathoide VBZ 37809 134 19 campi campi NNP 37809 134 20 _ _ NNP 37809 134 21 and and CC 37809 134 22 _ _ NNP 37809 134 23 marnockii marnockii NNS 37809 134 24 _ _ NNP 37809 134 25 ; ; : 37809 134 26 the the DT 37809 134 27 latter latter NN 37809 134 28 lacks lack VBZ 37809 134 29 a a DT 37809 134 30 supratympanic supratympanic JJ 37809 134 31 stripe stripe NN 37809 134 32 , , , 37809 134 33 which which WDT 37809 134 34 is be VBZ 37809 134 35 present present JJ 37809 134 36 in in IN 37809 134 37 the the DT 37809 134 38 other other JJ 37809 134 39 members member NNS 37809 134 40 of of IN 37809 134 41 the the DT 37809 134 42 eastern eastern JJ 37809 134 43 complex complex NN 37809 134 44 . . . 37809 135 1 _ _ NNP 37809 135 2 Syrrhophus Syrrhophus NNP 37809 135 3 interorbitalis interorbitalis NN 37809 135 4 _ _ NNP 37809 135 5 and and CC 37809 135 6 _ _ NNP 37809 135 7 nivocolimae nivocolimae NN 37809 135 8 _ _ NNP 37809 135 9 have have VBP 37809 135 10 light light JJ 37809 135 11 interorbital interorbital NN 37809 135 12 bars bar NNS 37809 135 13 ; ; : 37809 135 14 these these DT 37809 135 15 bars bar NNS 37809 135 16 occur occur VBP 37809 135 17 in in IN 37809 135 18 only only RB 37809 135 19 one one CD 37809 135 20 other other JJ 37809 135 21 population population NN 37809 135 22 of of IN 37809 135 23 the the DT 37809 135 24 genus genus NNP 37809 135 25 ( ( -LRB- 37809 135 26 _ _ NNP 37809 135 27 S. S. NNP 37809 136 1 c. c. NNP 37809 136 2 cystignathoides cystignathoides NNP 37809 136 3 _ _ NNP 37809 136 4 ) ) -RRB- 37809 136 5 . . . 37809 137 1 Bars bar NNS 37809 137 2 on on IN 37809 137 3 the the DT 37809 137 4 thighs thigh NNS 37809 137 5 are be VBP 37809 137 6 ill ill RB 37809 137 7 defined define VBN 37809 137 8 or or CC 37809 137 9 absent absent JJ 37809 137 10 in in IN 37809 137 11 the the DT 37809 137 12 members member NNS 37809 137 13 of of IN 37809 137 14 the the DT 37809 137 15 _ _ NNP 37809 137 16 marnockii marnockii NNS 37809 137 17 _ _ NNP 37809 137 18 and and CC 37809 137 19 part part NN 37809 137 20 of of IN 37809 137 21 the the DT 37809 137 22 _ _ NNP 37809 137 23 modestus modestus NN 37809 137 24 _ _ NNP 37809 137 25 groups group NNS 37809 137 26 . . . 37809 138 1 The the DT 37809 138 2 color color NN 37809 138 3 in in IN 37809 138 4 life life NN 37809 138 5 is be VBZ 37809 138 6 noted note VBN 37809 138 7 in in IN 37809 138 8 the the DT 37809 138 9 species species NN 37809 138 10 accounts account NNS 37809 138 11 . . . 37809 139 1 _ _ NNP 37809 139 2 Voice._--The Voice._--The NNP 37809 139 3 voices voice NNS 37809 139 4 of of IN 37809 139 5 all all DT 37809 139 6 _ _ NNP 37809 139 7 Syrrhophus Syrrhophus NNP 37809 139 8 _ _ NNP 37809 139 9 can can MD 37809 139 10 be be VB 37809 139 11 described describe VBN 37809 139 12 as as IN 37809 139 13 a a DT 37809 139 14 single single JJ 37809 139 15 short short JJ 37809 139 16 chirp chirp NN 37809 139 17 or or CC 37809 139 18 peep peep NNP 37809 139 19 ; ; : 37809 139 20 without without IN 37809 139 21 audiospectrographic audiospectrographic JJ 37809 139 22 analyses analyse VBZ 37809 139 23 the the DT 37809 139 24 significance significance NN 37809 139 25 of of IN 37809 139 26 the the DT 37809 139 27 differences difference NNS 37809 139 28 between between IN 37809 139 29 a a DT 37809 139 30 chirp chirp NN 37809 139 31 , , , 37809 139 32 peep peep NN 37809 139 33 , , , 37809 139 34 or or CC 37809 139 35 short short JJ 37809 139 36 whistle whistle NN 37809 139 37 can can MD 37809 139 38 not not RB 37809 139 39 be be VB 37809 139 40 appreciated appreciate VBN 37809 139 41 . . . 37809 140 1 Martin Martin NNP 37809 140 2 ( ( -LRB- 37809 140 3 1958 1958 CD 37809 140 4 ) ) -RRB- 37809 140 5 and and CC 37809 140 6 Wright Wright NNP 37809 140 7 and and CC 37809 140 8 Wright Wright NNP 37809 140 9 ( ( -LRB- 37809 140 10 1949 1949 CD 37809 140 11 ) ) -RRB- 37809 140 12 reported report VBD 37809 140 13 multi multi JJ 37809 140 14 - - JJ 37809 140 15 noted noted JJ 37809 140 16 calls call NNS 37809 140 17 , , , 37809 140 18 and and CC 37809 140 19 one one CD 37809 140 20 collector collector NN 37809 140 21 of of IN 37809 140 22 _ _ NNP 37809 140 23 S. S. NNP 37809 141 1 verrucipes verrucipes NNP 37809 141 2 _ _ NNP 37809 141 3 noted note VBD 37809 141 4 the the DT 37809 141 5 frog frog NN 37809 141 6 " " `` 37809 141 7 trilled trill VBN 37809 141 8 . . . 37809 141 9 " " '' 37809 142 1 Fouquette Fouquette NNP 37809 142 2 ( ( -LRB- 37809 142 3 1960 1960 CD 37809 142 4 ) ) -RRB- 37809 142 5 presented present VBD 37809 142 6 analyses analysis NNS 37809 142 7 of of IN 37809 142 8 two two CD 37809 142 9 species specie NNS 37809 142 10 ( ( -LRB- 37809 142 11 _ _ NNP 37809 142 12 marnockii marnockii NNS 37809 142 13 _ _ NNP 37809 142 14 and and CC 37809 142 15 _ _ NNP 37809 142 16 pipilans pipilans NNPS 37809 142 17 nebulosus nebulosus NNP 37809 142 18 _ _ NNP 37809 142 19 ) ) -RRB- 37809 142 20 . . . 37809 143 1 The the DT 37809 143 2 voices voice NNS 37809 143 3 were be VBD 37809 143 4 very very RB 37809 143 5 similar similar JJ 37809 143 6 ; ; : 37809 143 7 both both DT 37809 143 8 frogs frog NNS 37809 143 9 were be VBD 37809 143 10 reported report VBN 37809 143 11 to to IN 37809 143 12 " " `` 37809 143 13 trill trill NN 37809 143 14 " " '' 37809 143 15 and and CC 37809 143 16 " " `` 37809 143 17 chirp chirp NNP 37809 143 18 . . . 37809 143 19 " " '' 37809 144 1 SYSTEMATIC systematic JJ 37809 144 2 ACCOUNT account NN 37809 144 3 The the DT 37809 144 4 genus genus NN 37809 144 5 _ _ NNP 37809 144 6 Syrrhophus Syrrhophus NNP 37809 144 7 _ _ NNP 37809 144 8 has have VBZ 37809 144 9 been be VBN 37809 144 10 defined define VBN 37809 144 11 ( ( -LRB- 37809 144 12 Lynch Lynch NNP 37809 144 13 , , , 37809 144 14 1968 1968 CD 37809 144 15 ) ) -RRB- 37809 144 16 and and CC 37809 144 17 limited limit VBN 37809 144 18 to to IN 37809 144 19 the the DT 37809 144 20 group group NN 37809 144 21 of of IN 37809 144 22 species specie NNS 37809 144 23 occurring occur VBG 37809 144 24 in in IN 37809 144 25 Guatemala Guatemala NNP 37809 144 26 , , , 37809 144 27 México México NNP 37809 144 28 and and CC 37809 144 29 the the DT 37809 144 30 United United NNP 37809 144 31 States States NNP 37809 144 32 . . . 37809 145 1 The the DT 37809 145 2 closest close JJS 37809 145 3 relatives relative NNS 37809 145 4 of of IN 37809 145 5 _ _ NNP 37809 145 6 Syrrhophus Syrrhophus NNP 37809 145 7 _ _ NNP 37809 145 8 are be VBP 37809 145 9 the the DT 37809 145 10 frogs frog NNS 37809 145 11 of of IN 37809 145 12 the the DT 37809 145 13 genus genus NNP 37809 145 14 _ _ NNP 37809 145 15 Tomodactylus Tomodactylus NNP 37809 145 16 _ _ NNP 37809 145 17 ( ( -LRB- 37809 145 18 Dixon Dixon NNP 37809 145 19 , , , 37809 145 20 1957 1957 CD 37809 145 21 ; ; : 37809 145 22 Firschein Firschein NNP 37809 145 23 , , , 37809 145 24 1954 1954 CD 37809 145 25 ) ) -RRB- 37809 145 26 . . . 37809 146 1 Lynch Lynch NNP 37809 146 2 ( ( -LRB- 37809 146 3 1968 1968 CD 37809 146 4 ) ) -RRB- 37809 146 5 implied imply VBN 37809 146 6 there there EX 37809 146 7 were be VBD 37809 146 8 no no DT 37809 146 9 osteological osteological JJ 37809 146 10 bases basis NNS 37809 146 11 for for IN 37809 146 12 the the DT 37809 146 13 separation separation NN 37809 146 14 of of IN 37809 146 15 _ _ NNP 37809 146 16 Eleutherodactylus Eleutherodactylus NNP 37809 146 17 _ _ NNP 37809 146 18 , , , 37809 146 19 _ _ NNP 37809 146 20 Syrrhophus Syrrhophus NNP 37809 146 21 _ _ NNP 37809 146 22 , , , 37809 146 23 and and CC 37809 146 24 _ _ NNP 37809 146 25 Tomodactylus Tomodactylus NNP 37809 146 26 _ _ NNP 37809 146 27 . . . 37809 147 1 At at IN 37809 147 2 that that DT 37809 147 3 time time NN 37809 147 4 , , , 37809 147 5 I -PRON- PRP 37809 147 6 believed believe VBD 37809 147 7 such such JJ 37809 147 8 to to TO 37809 147 9 be be VB 37809 147 10 the the DT 37809 147 11 case case NN 37809 147 12 and and CC 37809 147 13 derived derive VBN 37809 147 14 _ _ NNP 37809 147 15 Syrrhophus Syrrhophus NNP 37809 147 16 _ _ NNP 37809 147 17 and and CC 37809 147 18 _ _ NNP 37809 147 19 Tomodactylus Tomodactylus NNP 37809 147 20 _ _ NNP 37809 147 21 from from IN 37809 147 22 the the DT 37809 147 23 _ _ NNP 37809 147 24 rhodopis rhodopis NN 37809 147 25 _ _ NNP 37809 147 26 complex complex NN 37809 147 27 of of IN 37809 147 28 _ _ NNP 37809 147 29 Eleutherodactylus Eleutherodactylus NNP 37809 147 30 _ _ NNP 37809 147 31 , , , 37809 147 32 with with IN 37809 147 33 which which WDT 37809 147 34 they -PRON- PRP 37809 147 35 share share VBP 37809 147 36 terrestrial terrestrial JJ 37809 147 37 habits habit NNS 37809 147 38 and and CC 37809 147 39 relatively relatively RB 37809 147 40 short short JJ 37809 147 41 limbs limb NNS 37809 147 42 . . . 37809 148 1 In in IN 37809 148 2 the the DT 37809 148 3 _ _ NNP 37809 148 4 rhodopis rhodopis NN 37809 148 5 _ _ NNP 37809 148 6 complex complex NN 37809 148 7 there there EX 37809 148 8 is be VBZ 37809 148 9 a a DT 37809 148 10 tendency tendency NN 37809 148 11 for for IN 37809 148 12 the the DT 37809 148 13 loss loss NN 37809 148 14 of of IN 37809 148 15 the the DT 37809 148 16 outer outer JJ 37809 148 17 palmar palmar NNP 37809 148 18 tubercle tubercle NN 37809 148 19 , , , 37809 148 20 a a DT 37809 148 21 not not RB 37809 148 22 uncommon uncommon JJ 37809 148 23 condition condition NN 37809 148 24 in in IN 37809 148 25 _ _ NNP 37809 148 26 Syrrhophus Syrrhophus NNP 37809 148 27 _ _ NNP 37809 148 28 and and CC 37809 148 29 _ _ NNP 37809 148 30 Tomodactylus Tomodactylus NNP 37809 148 31 _ _ NNP 37809 148 32 . . . 37809 149 1 However however RB 37809 149 2 , , , 37809 149 3 the the DT 37809 149 4 skulls skull NNS 37809 149 5 of of IN 37809 149 6 _ _ NNP 37809 149 7 Syrrhophus Syrrhophus NNP 37809 149 8 _ _ NNP 37809 149 9 and and CC 37809 149 10 _ _ NNP 37809 149 11 Tomodactylus Tomodactylus NNP 37809 149 12 _ _ NNP 37809 149 13 show show NN 37809 149 14 departures departure NNS 37809 149 15 from from IN 37809 149 16 the the DT 37809 149 17 pattern pattern NN 37809 149 18 observed observe VBN 37809 149 19 in in IN 37809 149 20 the the DT 37809 149 21 Middle Middle NNP 37809 149 22 American American NNP 37809 149 23 _ _ NNP 37809 149 24 Eleutherodactylus Eleutherodactylus NNP 37809 149 25 _ _ NNP 37809 149 26 , , , 37809 149 27 as as RB 37809 149 28 well well RB 37809 149 29 as as IN 37809 149 30 many many JJ 37809 149 31 of of IN 37809 149 32 those those DT 37809 149 33 species specie NNS 37809 149 34 in in IN 37809 149 35 western western JJ 37809 149 36 South South NNP 37809 149 37 America America NNP 37809 149 38 . . . 37809 150 1 Baldauf Baldauf NNP 37809 150 2 and and CC 37809 150 3 Tanzer Tanzer NNP 37809 150 4 ( ( -LRB- 37809 150 5 1965 1965 CD 37809 150 6 ) ) -RRB- 37809 150 7 reported report VBD 37809 150 8 that that IN 37809 150 9 the the DT 37809 150 10 frontoparietals frontoparietal NNS 37809 150 11 and and CC 37809 150 12 prootics prootic NNS 37809 150 13 were be VBD 37809 150 14 fused fuse VBN 37809 150 15 in in IN 37809 150 16 _ _ NNP 37809 150 17 Syrrhophus Syrrhophus NNP 37809 150 18 marnockii marnockii NNS 37809 150 19 _ _ NNP 37809 150 20 and and CC 37809 150 21 that that IN 37809 150 22 the the DT 37809 150 23 prootics prootic NNS 37809 150 24 and and CC 37809 150 25 exoccipitals exoccipital NNS 37809 150 26 appeared appear VBD 37809 150 27 to to TO 37809 150 28 be be VB 37809 150 29 one one CD 37809 150 30 bone bone NN 37809 150 31 ( ( -LRB- 37809 150 32 otoccipital otoccipital JJ 37809 150 33 ) ) -RRB- 37809 150 34 . . . 37809 151 1 The the DT 37809 151 2 otoccipital otoccipital NN 37809 151 3 is be VBZ 37809 151 4 not not RB 37809 151 5 uncommon uncommon JJ 37809 151 6 in in IN 37809 151 7 eleutherodactyline eleutherodactyline JJ 37809 151 8 frogs frog NNS 37809 151 9 , , , 37809 151 10 but but CC 37809 151 11 the the DT 37809 151 12 fusion fusion NN 37809 151 13 of of IN 37809 151 14 the the DT 37809 151 15 frontoparietals frontoparietal NNS 37809 151 16 with with IN 37809 151 17 the the DT 37809 151 18 prootics prootic NNS 37809 151 19 ( ( -LRB- 37809 151 20 regardless regardless RB 37809 151 21 of of IN 37809 151 22 the the DT 37809 151 23 fusion fusion NN 37809 151 24 of of IN 37809 151 25 the the DT 37809 151 26 latter latter JJ 37809 151 27 with with IN 37809 151 28 the the DT 37809 151 29 exoccipital exoccipital NN 37809 151 30 ) ) -RRB- 37809 151 31 is be VBZ 37809 151 32 uncommon uncommon JJ 37809 151 33 in in IN 37809 151 34 the the DT 37809 151 35 family family NN 37809 151 36 . . . 37809 152 1 I -PRON- PRP 37809 152 2 have have VBP 37809 152 3 found find VBN 37809 152 4 the the DT 37809 152 5 frontoparietal frontoparietal JJ 37809 152 6 - - HYPH 37809 152 7 prootic prootic JJ 37809 152 8 fusion fusion NN 37809 152 9 only only RB 37809 152 10 in in IN 37809 152 11 _ _ NNP 37809 152 12 Syrrhophus Syrrhophus NNP 37809 152 13 _ _ NNP 37809 152 14 ( ( -LRB- 37809 152 15 all all DT 37809 152 16 species specie NNS 37809 152 17 ) ) -RRB- 37809 152 18 , , , 37809 152 19 _ _ NNP 37809 152 20 Tomodactylus Tomodactylus NNP 37809 152 21 _ _ NNP 37809 152 22 ( ( -LRB- 37809 152 23 all all DT 37809 152 24 species specie NNS 37809 152 25 ) ) -RRB- 37809 152 26 , , , 37809 152 27 and and CC 37809 152 28 _ _ NNP 37809 152 29 Eleutherodactylus Eleutherodactylus NNP 37809 152 30 _ _ NNP 37809 152 31 ( ( -LRB- 37809 152 32 West West NNP 37809 152 33 Indies Indies NNP 37809 152 34 species specie NNS 37809 152 35 ) ) -RRB- 37809 152 36 . . . 37809 153 1 None none NN 37809 153 2 of of IN 37809 153 3 the the DT 37809 153 4 Middle Middle NNP 37809 153 5 American American NNP 37809 153 6 _ _ NNP 37809 153 7 Eleutherodactylus Eleutherodactylus NNP 37809 153 8 _ _ NNP 37809 153 9 has have VBZ 37809 153 10 the the DT 37809 153 11 two two CD 37809 153 12 bones bone NNS 37809 153 13 fused fuse VBN 37809 153 14 . . . 37809 154 1 Examination examination NN 37809 154 2 of of IN 37809 154 3 the the DT 37809 154 4 character character NN 37809 154 5 is be VBZ 37809 154 6 difficult difficult JJ 37809 154 7 in in IN 37809 154 8 dried dry VBN 37809 154 9 skeletal skeletal JJ 37809 154 10 preparations preparation NNS 37809 154 11 . . . 37809 155 1 Cleared clear VBN 37809 155 2 and and CC 37809 155 3 stained stain VBN 37809 155 4 or or CC 37809 155 5 macerated macerate VBN 37809 155 6 preparations preparation NNS 37809 155 7 are be VBP 37809 155 8 satisfactory satisfactory JJ 37809 155 9 for for IN 37809 155 10 checking check VBG 37809 155 11 this this DT 37809 155 12 character character NN 37809 155 13 . . . 37809 156 1 Thus thus RB 37809 156 2 , , , 37809 156 3 in in IN 37809 156 4 addition addition NN 37809 156 5 to to IN 37809 156 6 the the DT 37809 156 7 presence presence NN 37809 156 8 of of IN 37809 156 9 numerous numerous JJ 37809 156 10 plantar plantar NNP 37809 156 11 supernumerary supernumerary NNP 37809 156 12 tubercles tubercle NNS 37809 156 13 in in IN 37809 156 14 the the DT 37809 156 15 frogs frog NNS 37809 156 16 of of IN 37809 156 17 the the DT 37809 156 18 genera genera NN 37809 156 19 _ _ NNP 37809 156 20 Syrrhophus Syrrhophus NNP 37809 156 21 _ _ NNP 37809 156 22 and and CC 37809 156 23 _ _ NNP 37809 156 24 Tomodactylus Tomodactylus NNP 37809 156 25 _ _ NNP 37809 156 26 , , , 37809 156 27 these these DT 37809 156 28 two two CD 37809 156 29 genera genera NN 37809 156 30 can can MD 37809 156 31 be be VB 37809 156 32 separated separate VBN 37809 156 33 from from IN 37809 156 34 other other JJ 37809 156 35 Middle middle JJ 37809 156 36 American american JJ 37809 156 37 eleutherodactylines eleutherodactyline NNS 37809 156 38 by by IN 37809 156 39 the the DT 37809 156 40 fusion fusion NN 37809 156 41 of of IN 37809 156 42 the the DT 37809 156 43 frontoparietals frontoparietal NNS 37809 156 44 and and CC 37809 156 45 prootics prootic NNS 37809 156 46 . . . 37809 157 1 This this DT 37809 157 2 character character NN 37809 157 3 not not RB 37809 157 4 only only RB 37809 157 5 further far RBR 37809 157 6 strengthens strengthen VBZ 37809 157 7 the the DT 37809 157 8 argument argument NN 37809 157 9 that that IN 37809 157 10 the the DT 37809 157 11 two two CD 37809 157 12 genera genera NN 37809 157 13 are be VBP 37809 157 14 closely closely RB 37809 157 15 related related JJ 37809 157 16 but but CC 37809 157 17 poses pose VBZ 37809 157 18 a a DT 37809 157 19 problem problem NN 37809 157 20 of of IN 37809 157 21 zoogeographic zoogeographic JJ 37809 157 22 analysis analysis NN 37809 157 23 of of IN 37809 157 24 the the DT 37809 157 25 distribution distribution NN 37809 157 26 of of IN 37809 157 27 the the DT 37809 157 28 character character NN 37809 157 29 , , , 37809 157 30 which which WDT 37809 157 31 will will MD 37809 157 32 be be VB 37809 157 33 discussed discuss VBN 37809 157 34 fully fully RB 37809 157 35 elsewhere elsewhere RB 37809 157 36 . . . 37809 158 1 Key key JJ 37809 158 2 to to IN 37809 158 3 the the DT 37809 158 4 Species Species NNPS 37809 158 5 of of IN 37809 158 6 the the DT 37809 158 7 Frog Frog NNP 37809 158 8 Genus Genus NNP 37809 158 9 _ _ NNP 37809 158 10 Syrrhophus Syrrhophus NNP 37809 158 11 _ _ NNP 37809 158 12 1 1 CD 37809 158 13 . . . 37809 159 1 Three three CD 37809 159 2 large large JJ 37809 159 3 , , , 37809 159 4 well well RB 37809 159 5 - - HYPH 37809 159 6 developed develop VBN 37809 159 7 palmar palmar NN 37809 159 8 tubercles tubercle NNS 37809 159 9 2 2 CD 37809 159 10 Two two CD 37809 159 11 large large JJ 37809 159 12 palmar palmar NN 37809 159 13 tubercles tubercle NNS 37809 159 14 ; ; , 37809 159 15 outer outer NNP 37809 159 16 ( ( -LRB- 37809 159 17 third third JJ 37809 159 18 ) ) -RRB- 37809 159 19 palmar palmar NNP 37809 159 20 tubercle tubercle NN 37809 159 21 reduced reduce VBN 37809 159 22 in in IN 37809 159 23 size size NN 37809 159 24 or or CC 37809 159 25 absent absent JJ 37809 159 26 9 9 CD 37809 159 27 2 2 CD 37809 159 28 . . . 37809 160 1 Digital Digital NNP 37809 160 2 pads pad NNS 37809 160 3 more more RBR 37809 160 4 than than IN 37809 160 5 twice twice RB 37809 160 6 ( ( -LRB- 37809 160 7 usually usually RB 37809 160 8 three three CD 37809 160 9 or or CC 37809 160 10 more more JJR 37809 160 11 ) ) -RRB- 37809 160 12 times time NNS 37809 160 13 width width NN 37809 160 14 of of IN 37809 160 15 digit digit NN 37809 160 16 3 3 CD 37809 160 17 Digital Digital NNP 37809 160 18 pads pad NNS 37809 160 19 less less RBR 37809 160 20 than than IN 37809 160 21 twice twice RB 37809 160 22 width width NN 37809 160 23 of of IN 37809 160 24 digit digit NN 37809 160 25 4 4 CD 37809 160 26 3 3 CD 37809 160 27 . . . 37809 161 1 Males male NNS 37809 161 2 having have VBG 37809 161 3 vocal vocal JJ 37809 161 4 slits slit NNS 37809 161 5 ; ; , 37809 161 6 dorsum dorsum NN 37809 161 7 vermiculate vermiculate NN 37809 161 8 ; ; : 37809 161 9 diameter diameter NN 37809 161 10 of of IN 37809 161 11 tympanum tympanum NN 37809 161 12 in in IN 37809 161 13 males male NNS 37809 161 14 about about IN 37809 161 15 one one CD 37809 161 16 - - HYPH 37809 161 17 half half NN 37809 161 18 diameter diameter NN 37809 161 19 of of IN 37809 161 20 eye eye NN 37809 161 21 _ _ NNP 37809 161 22 S. S. NNP 37809 162 1 dennisi dennisi NNP 37809 162 2 _ _ NNP 37809 162 3 Males Males NNP 37809 162 4 lacking lack VBG 37809 162 5 vocal vocal JJ 37809 162 6 slits slit NNS 37809 162 7 ; ; , 37809 162 8 dorsum dorsum NN 37809 162 9 flecked fleck VBN 37809 162 10 , , , 37809 162 11 spotted spot VBD 37809 162 12 , , , 37809 162 13 or or CC 37809 162 14 blotched blotch VBN 37809 162 15 ; ; : 37809 162 16 diameter diameter NN 37809 162 17 of of IN 37809 162 18 tympanum tympanum NN 37809 162 19 in in IN 37809 162 20 male male NN 37809 162 21 about about RB 37809 162 22 three three CD 37809 162 23 - - HYPH 37809 162 24 fourths fourth NNS 37809 162 25 that that DT 37809 162 26 of of IN 37809 162 27 eye eye NN 37809 162 28 _ _ NNP 37809 162 29 S. S. NNP 37809 163 1 longipes longipes NNP 37809 163 2 _ _ NNP 37809 163 3 4 4 CD 37809 163 4 . . . 37809 164 1 First first JJ 37809 164 2 finger finger NN 37809 164 3 longer long RBR 37809 164 4 than than IN 37809 164 5 second second JJ 37809 164 6 5 5 CD 37809 164 7 First first JJ 37809 164 8 finger finger NN 37809 164 9 shorter short JJR 37809 164 10 than than IN 37809 164 11 or or CC 37809 164 12 equal equal JJ 37809 164 13 to to IN 37809 164 14 second second JJ 37809 164 15 7 7 CD 37809 164 16 5 5 CD 37809 164 17 . . . 37809 165 1 Venter venter VB 37809 165 2 smooth smooth JJ 37809 165 3 ; ; : 37809 165 4 dorsum dorsum NN 37809 165 5 spotted spot VBN 37809 165 6 or or CC 37809 165 7 vermiculate vermiculate JJ 37809 165 8 _ _ NNP 37809 165 9 S. S. NNP 37809 166 1 leprus leprus NNP 37809 166 2 _ _ NNP 37809 166 3 Venter Venter NNP 37809 166 4 areolate areolate NN 37809 166 5 , , , 37809 166 6 or or CC 37809 166 7 if if IN 37809 166 8 smooth smooth JJ 37809 166 9 , , , 37809 166 10 dorsum dorsum NN 37809 166 11 flecked fleck VBN 37809 166 12 and and CC 37809 166 13 interorbital interorbital NNP 37809 166 14 bar bar NN 37809 166 15 lacking lack VBG 37809 166 16 6 6 CD 37809 166 17 6 6 CD 37809 166 18 . . . 37809 167 1 Venter venter JJ 37809 167 2 areolate areolate NN 37809 167 3 ; ; : 37809 167 4 interorbital interorbital NNP 37809 167 5 bar bar NN 37809 167 6 present present NN 37809 167 7 ; ; : 37809 167 8 ground ground NN 37809 167 9 color color NN 37809 167 10 yellowish yellowish JJ 37809 167 11 _ _ NNP 37809 167 12 S. S. NNP 37809 168 1 cystignathoides cystignathoide NNS 37809 168 2 cystignathoides cystignathoide NNS 37809 168 3 _ _ NNP 37809 168 4 Venter Venter NNP 37809 168 5 smooth smooth JJ 37809 168 6 ; ; : 37809 168 7 interorbital interorbital NNP 37809 168 8 bar bar NN 37809 168 9 absent absent NN 37809 168 10 ; ; : 37809 168 11 ground ground NN 37809 168 12 color color NN 37809 168 13 brown brown NNP 37809 168 14 _ _ NNP 37809 168 15 S. S. NNP 37809 169 1 cystignathoides cystignathoide NNS 37809 169 2 campi campi NNP 37809 169 3 _ _ NNP 37809 169 4 7 7 CD 37809 169 5 . . . 37809 170 1 First first JJ 37809 170 2 finger finger NN 37809 170 3 shorter short JJR 37809 170 4 than than IN 37809 170 5 second second JJ 37809 170 6 ; ; : 37809 170 7 digital digital JJ 37809 170 8 tips tip NNS 37809 170 9 only only RB 37809 170 10 slightly slightly RB 37809 170 11 dilated dilate VBN 37809 170 12 ; ; : 37809 170 13 green green JJ 37809 170 14 in in IN 37809 170 15 life life NN 37809 170 16 with with IN 37809 170 17 darker dark JJR 37809 170 18 green green JJ 37809 170 19 spots spot NNS 37809 170 20 _ _ NNP 37809 170 21 S. S. NNP 37809 171 1 verrucipes verrucipes NNP 37809 171 2 _ _ NNP 37809 171 3 First First NNP 37809 171 4 finger finger NN 37809 171 5 equal equal JJ 37809 171 6 to to IN 37809 171 7 second second JJ 37809 171 8 ; ; : 37809 171 9 digital digital JJ 37809 171 10 tips tip NNS 37809 171 11 slightly slightly RB 37809 171 12 to to IN 37809 171 13 moderately moderately RB 37809 171 14 expanded expand VBN 37809 171 15 8 8 CD 37809 171 16 8 8 CD 37809 171 17 . . . 37809 172 1 Dorsum dorsum NN 37809 172 2 vermiculate vermiculate NN 37809 172 3 ; ; : 37809 172 4 interorbital interorbital NNP 37809 172 5 bar bar NN 37809 172 6 present present NN 37809 172 7 ; ; : 37809 172 8 ground ground NN 37809 172 9 color color NN 37809 172 10 cream cream NN 37809 172 11 to to TO 37809 172 12 brown brown VB 37809 172 13 in in IN 37809 172 14 life life NN 37809 172 15 _ _ NNP 37809 172 16 S. S. NNP 37809 173 1 guttilatus guttilatus NNP 37809 173 2 _ _ NNP 37809 173 3 Dorsum Dorsum NNP 37809 173 4 punctate punctate NN 37809 173 5 or or CC 37809 173 6 flecked flecked JJ 37809 173 7 ; ; : 37809 173 8 interorbital interorbital NNP 37809 173 9 bar bar NNP 37809 173 10 absent absent NN 37809 173 11 ; ; : 37809 173 12 ground ground NN 37809 173 13 color color NN 37809 173 14 green green JJ 37809 173 15 in in IN 37809 173 16 life life NN 37809 173 17 _ _ NNP 37809 173 18 S. S. NNP 37809 174 1 marnockii marnockii NNP 37809 174 2 _ _ NNP 37809 174 3 9 9 CD 37809 174 4 . . . 37809 175 1 Dorsum dorsum NN 37809 175 2 dark dark NN 37809 175 3 with with IN 37809 175 4 pale pale JJ 37809 175 5 ( ( -LRB- 37809 175 6 red red JJ 37809 175 7 in in IN 37809 175 8 life life NN 37809 175 9 ) ) -RRB- 37809 175 10 spots spot NNS 37809 175 11 ; ; , 37809 175 12 digital digital JJ 37809 175 13 pads pad NNS 37809 175 14 not not RB 37809 175 15 expanded expand VBN 37809 175 16 _ _ NNP 37809 175 17 S. S. NNP 37809 176 1 rubrimaculatus rubrimaculatus NNP 37809 176 2 _ _ NNP 37809 176 3 Dorsum Dorsum NNP 37809 176 4 pale pale NN 37809 176 5 with with IN 37809 176 6 dark dark JJ 37809 176 7 markings marking NNS 37809 176 8 and and CC 37809 176 9 digital digital JJ 37809 176 10 pads pad NNS 37809 176 11 slightly slightly RB 37809 176 12 to to IN 37809 176 13 widely widely RB 37809 176 14 expanded expand VBN 37809 176 15 10 10 CD 37809 176 16 10 10 CD 37809 176 17 . . . 37809 177 1 Digital digital JJ 37809 177 2 tips tip NNS 37809 177 3 not not RB 37809 177 4 widely widely RB 37809 177 5 expanded expand VBN 37809 177 6 ; ; : 37809 177 7 tympanum tympanum PRP 37809 177 8 well well RB 37809 177 9 - - HYPH 37809 177 10 defined define VBN 37809 177 11 ; ; : 37809 177 12 outer outer JJ 37809 177 13 metatarsal metatarsal NN 37809 177 14 tubercle tubercle NN 37809 177 15 more more JJR 37809 177 16 than than IN 37809 177 17 one one CD 37809 177 18 - - HYPH 37809 177 19 half half NN 37809 177 20 size size NN 37809 177 21 of of IN 37809 177 22 inner inner JJ 37809 177 23 11 11 CD 37809 177 24 Digital Digital NNP 37809 177 25 tips tip NNS 37809 177 26 widely widely RB 37809 177 27 expanded expand VBN 37809 177 28 , , , 37809 177 29 truncate truncate JJ 37809 177 30 in in IN 37809 177 31 outline outline NN 37809 177 32 ; ; : 37809 177 33 tympanum tympanum NN 37809 177 34 poorly poorly RB 37809 177 35 defined define VBN 37809 177 36 ; ; : 37809 177 37 outer outer JJ 37809 177 38 metatarsal metatarsal NN 37809 177 39 tubercle tubercle NN 37809 177 40 less less JJR 37809 177 41 than than IN 37809 177 42 one one CD 37809 177 43 - - HYPH 37809 177 44 half half NN 37809 177 45 size size NN 37809 177 46 of of IN 37809 177 47 inner inner JJ 37809 177 48 12 12 CD 37809 177 49 11 11 CD 37809 177 50 . . . 37809 178 1 Dorsum dorsum NN 37809 178 2 dark dark JJ 37809 178 3 brown brown NN 37809 178 4 with with IN 37809 178 5 large large JJ 37809 178 6 light light JJ 37809 178 7 spots spot NNS 37809 178 8 or or CC 37809 178 9 blotches blotch NNS 37809 178 10 ; ; , 37809 178 11 tympanum tympanum NN 37809 178 12 / / SYM 37809 178 13 eye eye NN 37809 178 14 ratio ratio NN 37809 178 15 usually usually RB 37809 178 16 greater great JJR 37809 178 17 than than IN 37809 178 18 43 43 CD 37809 178 19 percent percent NN 37809 178 20 _ _ NNP 37809 178 21 S. S. NNP 37809 179 1 pipilans pipilans NNPS 37809 179 2 pipilans pipilans NNPS 37809 179 3 _ _ NNP 37809 179 4 Dorsum Dorsum NNP 37809 179 5 dark dark JJ 37809 179 6 brown brown NN 37809 179 7 with with IN 37809 179 8 small small JJ 37809 179 9 light light JJ 37809 179 10 spots spot NNS 37809 179 11 ; ; : 37809 179 12 tympanum tympanum NN 37809 179 13 / / SYM 37809 179 14 eye eye NN 37809 179 15 ratio ratio NN 37809 179 16 less less JJR 37809 179 17 than than IN 37809 179 18 48 48 CD 37809 179 19 percent percent NN 37809 179 20 _ _ NNP 37809 179 21 S. S. NNP 37809 180 1 pipilans pipilans NNPS 37809 180 2 nebulosus nebulosus NNP 37809 180 3 _ _ NNP 37809 180 4 12 12 CD 37809 180 5 . . . 37809 181 1 Light light JJ 37809 181 2 interorbital interorbital NN 37809 181 3 bar bar NN 37809 181 4 present present JJ 37809 181 5 13 13 CD 37809 181 6 Light Light NNP 37809 181 7 interorbital interorbital NN 37809 181 8 bar bar NN 37809 181 9 absent absent NN 37809 181 10 14 14 CD 37809 181 11 13 13 CD 37809 181 12 . . . 37809 182 1 Adults adult NNS 37809 182 2 small small JJ 37809 182 3 , , , 37809 182 4 less less JJR 37809 182 5 than than IN 37809 182 6 22 22 CD 37809 182 7 mm mm NNS 37809 182 8 . . . 37809 183 1 snout snout NN 37809 183 2 - - HYPH 37809 183 3 vent vent NN 37809 183 4 length length NN 37809 183 5 with with IN 37809 183 6 a a DT 37809 183 7 broad broad JJ 37809 183 8 mid mid JJ 37809 183 9 - - JJ 37809 183 10 dorsal dorsal JJ 37809 183 11 stripe stripe NN 37809 183 12 ; ; : 37809 183 13 dark dark JJ 37809 183 14 bands band NNS 37809 183 15 on on IN 37809 183 16 shank shank JJ 37809 183 17 narrower narrow JJR 37809 183 18 than than IN 37809 183 19 light light JJ 37809 183 20 interspaces interspace NNS 37809 183 21 _ _ NNP 37809 183 22 S. S. NNP 37809 184 1 nivocolimae nivocolimae NNP 37809 184 2 _ _ NNP 37809 184 3 Adults Adults NNP 37809 184 4 larger large JJR 37809 184 5 , , , 37809 184 6 more more JJR 37809 184 7 than than IN 37809 184 8 22 22 CD 37809 184 9 mm mm NNS 37809 184 10 . . . 37809 185 1 snout snout NN 37809 185 2 - - HYPH 37809 185 3 vent vent NN 37809 185 4 length length NN 37809 185 5 ; ; : 37809 185 6 dorsum dorsum NN 37809 185 7 vermiculate vermiculate NN 37809 185 8 ; ; : 37809 185 9 dark dark JJ 37809 185 10 bands band NNS 37809 185 11 on on IN 37809 185 12 shank shank JJ 37809 185 13 broader broad JJR 37809 185 14 than than IN 37809 185 15 light light JJ 37809 185 16 interspaces interspace NNS 37809 185 17 _ _ NNP 37809 185 18 S. S. NNP 37809 186 1 interorbitalis interorbitalis NNP 37809 186 2 _ _ NNP 37809 186 3 14 14 CD 37809 186 4 . . . 37809 187 1 Dorsum dorsum NN 37809 187 2 spotted spot VBN 37809 187 3 with with IN 37809 187 4 discrete discrete JJ 37809 187 5 black black JJ 37809 187 6 spots spot NNS 37809 187 7 ; ; : 37809 187 8 pattern pattern NN 37809 187 9 definite definite NN 37809 187 10 _ _ NNP 37809 187 11 S. S. NNP 37809 188 1 modestus modestus NNP 37809 188 2 _ _ NNP 37809 188 3 Dorsum Dorsum NNP 37809 188 4 reticulate reticulate NN 37809 188 5 or or CC 37809 188 6 vermiculate vermiculate NN 37809 188 7 , , , 37809 188 8 pattern pattern NN 37809 188 9 poorly poorly RB 37809 188 10 defined define VBN 37809 188 11 15 15 CD 37809 188 12 15 15 CD 37809 188 13 . . . 37809 189 1 Adults adult NNS 37809 189 2 small small JJ 37809 189 3 , , , 37809 189 4 less less JJR 37809 189 5 than than IN 37809 189 6 21 21 CD 37809 189 7 mm mm NNS 37809 189 8 . . . 37809 190 1 snout snout NN 37809 190 2 - - HYPH 37809 190 3 vent vent NN 37809 190 4 length length NN 37809 190 5 ; ; : 37809 190 6 upper upper JJ 37809 190 7 arm arm NN 37809 190 8 not not RB 37809 190 9 banded band VBN 37809 190 10 _ _ NNP 37809 190 11 S. S. NNP 37809 191 1 pallidus pallidus NNP 37809 191 2 _ _ NNP 37809 191 3 Adults Adults NNP 37809 191 4 larger large JJR 37809 191 5 , , , 37809 191 6 usually usually RB 37809 191 7 greater great JJR 37809 191 8 than than IN 37809 191 9 21 21 CD 37809 191 10 mm mm CD 37809 191 11 . . . 37809 192 1 snout snout NN 37809 192 2 - - HYPH 37809 192 3 vent vent NN 37809 192 4 length length NN 37809 192 5 ; ; : 37809 192 6 upper upper JJ 37809 192 7 arm arm NN 37809 192 8 banded band VBD 37809 192 9 _ _ NNP 37809 192 10 S. S. NNP 37809 193 1 teretistes teretistes NNPS 37809 193 2 _ _ NNP 37809 193 3 SPECIES specie NNS 37809 193 4 ACCOUNTS ACCOUNTS NNP 37809 193 5 The the DT 37809 193 6 following follow VBG 37809 193 7 accounts account NNS 37809 193 8 do do VBP 37809 193 9 not not RB 37809 193 10 include include VB 37809 193 11 complete complete JJ 37809 193 12 descriptions description NNS 37809 193 13 of of IN 37809 193 14 each each DT 37809 193 15 taxon taxon NN 37809 193 16 , , , 37809 193 17 because because IN 37809 193 18 a a DT 37809 193 19 more more JJR 37809 193 20 than than IN 37809 193 21 adequate adequate JJ 37809 193 22 number number NN 37809 193 23 of of IN 37809 193 24 descriptions description NNS 37809 193 25 is be VBZ 37809 193 26 available available JJ 37809 193 27 in in IN 37809 193 28 the the DT 37809 193 29 recent recent JJ 37809 193 30 ( ( -LRB- 37809 193 31 1940 1940 CD 37809 193 32 - - SYM 37809 193 33 1966 1966 CD 37809 193 34 ) ) -RRB- 37809 193 35 literature literature NN 37809 193 36 . . . 37809 194 1 An an DT 37809 194 2 abbreviated abbreviate VBN 37809 194 3 synonymy synonymy NN 37809 194 4 , , , 37809 194 5 in in IN 37809 194 6 which which WDT 37809 194 7 are be VBP 37809 194 8 listed list VBN 37809 194 9 all all DT 37809 194 10 combinations combination NNS 37809 194 11 and and CC 37809 194 12 emendations emendation NNS 37809 194 13 of of IN 37809 194 14 names name NNS 37809 194 15 and and CC 37809 194 16 significant significant JJ 37809 194 17 contributions contribution NNS 37809 194 18 to to IN 37809 194 19 our -PRON- PRP$ 37809 194 20 knowledge knowledge NN 37809 194 21 of of IN 37809 194 22 the the DT 37809 194 23 taxon taxon NN 37809 194 24 , , , 37809 194 25 is be VBZ 37809 194 26 given give VBN 37809 194 27 for for IN 37809 194 28 each each DT 37809 194 29 . . . 37809 195 1 For for IN 37809 195 2 each each DT 37809 195 3 species specie NNS 37809 195 4 and and CC 37809 195 5 subspecies subspecie NNS 37809 195 6 the the DT 37809 195 7 following follow VBG 37809 195 8 are be VBP 37809 195 9 given give VBN 37809 195 10 : : : 37809 195 11 descriptive descriptive JJ 37809 195 12 diagnosis diagnosis NN 37809 195 13 , , , 37809 195 14 statement statement NN 37809 195 15 of of IN 37809 195 16 range range NNP 37809 195 17 , , , 37809 195 18 remarks remark NNS 37809 195 19 on on IN 37809 195 20 taxonomy taxonomy NN 37809 195 21 , , , 37809 195 22 list list NN 37809 195 23 of of IN 37809 195 24 specimens specimen NNS 37809 195 25 examined examine VBN 37809 195 26 , , , 37809 195 27 illustration illustration NN 37809 195 28 of of IN 37809 195 29 color color NN 37809 195 30 pattern pattern NN 37809 195 31 , , , 37809 195 32 and and CC 37809 195 33 distribution distribution NN 37809 195 34 map map NN 37809 195 35 . . . 37809 196 1 = = NFP 37809 196 2 Syrrhophus Syrrhophus NNP 37809 196 3 cystignathoides= cystignathoides= NN 37809 196 4 ( ( -LRB- 37809 196 5 Cope Cope NNP 37809 196 6 ) ) -RRB- 37809 196 7 _ _ NNP 37809 196 8 Phyllobates Phyllobates NNPS 37809 196 9 cystignathoides cystignathoide VBZ 37809 196 10 _ _ NNP 37809 196 11 Cope Cope NNP 37809 196 12 , , , 37809 196 13 1877:89 1877:89 CD 37809 196 14 - - SYM 37809 196 15 90 90 CD 37809 196 16 [ [ -LRB- 37809 196 17 Syntypes.--Originally syntypes.--originally CD 37809 196 18 USNM USNM NNP 37809 196 19 32402 32402 CD 37809 196 20 - - SYM 37809 196 21 32409 32409 CD 37809 196 22 , , , 37809 196 23 ( ( -LRB- 37809 196 24 32405 32405 CD 37809 196 25 now now RB 37809 196 26 in in IN 37809 196 27 MCZ MCZ NNP 37809 196 28 ) ) -RRB- 37809 196 29 from from IN 37809 196 30 Potrero Potrero NNP 37809 196 31 , , , 37809 196 32 near near IN 37809 196 33 Córdoba Córdoba NNP 37809 196 34 , , , 37809 196 35 Veracruz Veracruz NNP 37809 196 36 , , , 37809 196 37 México México NNP 37809 196 38 , , , 37809 196 39 Francis Francis NNP 37809 196 40 Sumichrast Sumichrast NNP 37809 196 41 collector collector NN 37809 196 42 . . . 37809 196 43 ] ] -RRB- 37809 197 1 _ _ NNP 37809 197 2 Diagnosis._--Adults Diagnosis._--Adults NNP 37809 197 3 small small JJ 37809 197 4 , , , 37809 197 5 males male VBZ 37809 197 6 16.0 16.0 CD 37809 197 7 to to IN 37809 197 8 23.5 23.5 CD 37809 197 9 mm mm NNS 37809 197 10 . . . 37809 198 1 in in IN 37809 198 2 snout snout NN 37809 198 3 - - HYPH 37809 198 4 vent vent NN 37809 198 5 length length NN 37809 198 6 , , , 37809 198 7 females female VBZ 37809 198 8 16.0 16.0 CD 37809 198 9 - - SYM 37809 198 10 25.8 25.8 CD 37809 198 11 mm mm NNP 37809 198 12 . . . 37809 199 1 in in IN 37809 199 2 snout snout NN 37809 199 3 - - HYPH 37809 199 4 vent vent NN 37809 199 5 length length NN 37809 199 6 ; ; : 37809 199 7 vocal vocal NN 37809 199 8 slits slit NNS 37809 199 9 present present JJ 37809 199 10 in in IN 37809 199 11 males male NNS 37809 199 12 ; ; : 37809 199 13 finger finger NN 37809 199 14 tips tip NNS 37809 199 15 slightly slightly RB 37809 199 16 expanded expand VBN 37809 199 17 ; ; : 37809 199 18 first first RB 37809 199 19 finger finger NN 37809 199 20 longer long RBR 37809 199 21 than than IN 37809 199 22 second second JJ 37809 199 23 ; ; : 37809 199 24 outer outer JJ 37809 199 25 metatarsal metatarsal NN 37809 199 26 tubercle tubercle NN 37809 199 27 one one CD 37809 199 28 - - HYPH 37809 199 29 half half NN 37809 199 30 size size NN 37809 199 31 of of IN 37809 199 32 inner inner JJ 37809 199 33 , , , 37809 199 34 conical conical JJ 37809 199 35 , , , 37809 199 36 compressed compress VBN 37809 199 37 ; ; : 37809 199 38 skin skin NN 37809 199 39 of of IN 37809 199 40 dorsum dorsum NN 37809 199 41 weakly weakly RB 37809 199 42 pustular pustular NN 37809 199 43 , , , 37809 199 44 that that DT 37809 199 45 of of IN 37809 199 46 venter venter NN 37809 199 47 smooth smooth JJ 37809 199 48 to to TO 37809 199 49 areolate areolate VB 37809 199 50 ; ; : 37809 199 51 tympanum tympanum PRP 37809 199 52 44 44 CD 37809 199 53 to to TO 37809 199 54 69 69 CD 37809 199 55 per per NN 37809 199 56 cent cent NN 37809 199 57 diameter diameter NN 37809 199 58 of of IN 37809 199 59 eye eye NN 37809 199 60 ( ( -LRB- 37809 199 61 mean mean NNP 37809 199 62 55.5 55.5 CD 37809 199 63 per per IN 37809 199 64 cent cent NN 37809 199 65 ) ) -RRB- 37809 199 66 ; ; : 37809 199 67 ground ground NN 37809 199 68 color color NN 37809 199 69 yellow yellow NNP 37809 199 70 to to TO 37809 199 71 brown brown VB 37809 199 72 in in IN 37809 199 73 life life NN 37809 199 74 with with IN 37809 199 75 brown brown JJ 37809 199 76 to to IN 37809 199 77 black black JJ 37809 199 78 fleckings flecking NNS 37809 199 79 on on IN 37809 199 80 dorsum dorsum NN 37809 199 81 and and CC 37809 199 82 flanks flank NNS 37809 199 83 ; ; : 37809 199 84 limbs limb NNS 37809 199 85 banded band VBD 37809 199 86 ; ; : 37809 199 87 interorbital interorbital NNP 37809 199 88 bar bar NN 37809 199 89 present present JJ 37809 199 90 or or CC 37809 199 91 not not RB 37809 199 92 . . . 37809 200 1 _ _ NNP 37809 200 2 Remarks._--Two Remarks._--Two NNP 37809 200 3 geographic geographic JJ 37809 200 4 races race NNS 37809 200 5 ( ( -LRB- 37809 200 6 subspecies subspecie NNS 37809 200 7 ) ) -RRB- 37809 200 8 are be VBP 37809 200 9 herein herein NNP 37809 200 10 recognized recognize VBN 37809 200 11 ; ; : 37809 200 12 previously previously RB 37809 200 13 these these DT 37809 200 14 were be VBD 37809 200 15 held hold VBN 37809 200 16 by by IN 37809 200 17 various various JJ 37809 200 18 authors author NNS 37809 200 19 to to TO 37809 200 20 be be VB 37809 200 21 species specie NNS 37809 200 22 ( ( -LRB- 37809 200 23 _ _ NNP 37809 200 24 campi campi NNP 37809 200 25 _ _ NNP 37809 200 26 and and CC 37809 200 27 _ _ NNP 37809 200 28 cystignathoides cystignathoide VBZ 37809 200 29 _ _ NNP 37809 200 30 ) ) -RRB- 37809 200 31 . . . 37809 201 1 Intergradation intergradation NN 37809 201 2 occurs occur VBZ 37809 201 3 in in IN 37809 201 4 southern southern JJ 37809 201 5 Tamaulipas Tamaulipas NNP 37809 201 6 and and CC 37809 201 7 eastern eastern JJ 37809 201 8 San San NNP 37809 201 9 Luis Luis NNP 37809 201 10 Potosí Potosí NNP 37809 201 11 , , , 37809 201 12 México México NNP 37809 201 13 . . . 37809 202 1 The the DT 37809 202 2 two two CD 37809 202 3 subspecies subspecie NNS 37809 202 4 can can MD 37809 202 5 be be VB 37809 202 6 distinguished distinguish VBN 37809 202 7 on on IN 37809 202 8 the the DT 37809 202 9 basis basis NN 37809 202 10 of of IN 37809 202 11 color color NN 37809 202 12 pattern pattern NN 37809 202 13 and and CC 37809 202 14 the the DT 37809 202 15 condition condition NN 37809 202 16 of of IN 37809 202 17 the the DT 37809 202 18 skin skin NN 37809 202 19 of of IN 37809 202 20 the the DT 37809 202 21 venter venter NN 37809 202 22 . . . 37809 203 1 _ _ NNP 37809 203 2 Distribution._--Low Distribution._--Low NNP 37809 203 3 to to TO 37809 203 4 moderate moderate VB 37809 203 5 elevations elevation NNS 37809 203 6 from from IN 37809 203 7 the the DT 37809 203 8 Río Río NNP 37809 203 9 Grande Grande NNP 37809 203 10 embayment embayment NN 37809 203 11 to to IN 37809 203 12 central central JJ 37809 203 13 Veracruz Veracruz NNP 37809 203 14 , , , 37809 203 15 México México NNP 37809 203 16 ( ( -LRB- 37809 203 17 Fig Fig NNP 37809 203 18 . . . 37809 204 1 5 5 LS 37809 204 2 ) ) -RRB- 37809 204 3 . . . 37809 205 1 = = NFP 37809 205 2 Syrrhophus Syrrhophus NNP 37809 205 3 cystignathoides cystignathoide VBZ 37809 205 4 campi= campi= NN 37809 205 5 Stejneger Stejneger NNP 37809 205 6 , , , 37809 205 7 New new JJ 37809 205 8 combination combination NN 37809 205 9 _ _ NNP 37809 205 10 Syrrhophus Syrrhophus NNP 37809 205 11 campi campi NN 37809 205 12 _ _ NNP 37809 205 13 Stejneger Stejneger NNP 37809 205 14 , , , 37809 205 15 1915:131 1915:131 CD 37809 205 16 - - SYM 37809 205 17 32 32 CD 37809 205 18 . . . 37809 206 1 [ [ -LRB- 37809 206 2 Holotype.--USNM Holotype.--USNM NNP 37809 206 3 52290 52290 CD 37809 206 4 , , , 37809 206 5 from from IN 37809 206 6 Brownsville Brownsville NNP 37809 206 7 , , , 37809 206 8 Cameron Cameron NNP 37809 206 9 Co. Co. NNP 37809 206 10 , , , 37809 206 11 Texas Texas NNP 37809 206 12 ; ; : 37809 206 13 R. R. NNP 37809 206 14 D. D. NNP 37809 206 15 Camp Camp NNP 37809 206 16 collector collector NN 37809 206 17 , , , 37809 206 18 March March NNP 37809 206 19 31 31 CD 37809 206 20 , , , 37809 206 21 1915 1915 CD 37809 206 22 ] ] -RRB- 37809 206 23 . . . 37809 207 1 Smith Smith NNP 37809 207 2 and and CC 37809 207 3 Taylor Taylor NNP 37809 207 4 , , , 37809 207 5 1948:52 1948:52 CD 37809 207 6 . . . 37809 208 1 Martin Martin NNP 37809 208 2 , , , 37809 208 3 1958:50 1958:50 CD 37809 208 4 . . . 37809 209 1 _ _ NNP 37809 209 2 Diagnosis._--Venter Diagnosis._--Venter NNP 37809 209 3 smooth smooth JJ 37809 209 4 ; ; : 37809 209 5 usually usually RB 37809 209 6 no no DT 37809 209 7 interorbital interorbital JJ 37809 209 8 light light NN 37809 209 9 and and CC 37809 209 10 dark dark JJ 37809 209 11 bars bar NNS 37809 209 12 present present JJ 37809 209 13 ; ; : 37809 209 14 ground ground NN 37809 209 15 color color NN 37809 209 16 brown brown NNP 37809 209 17 in in IN 37809 209 18 life life NN 37809 209 19 ( ( -LRB- 37809 209 20 Fig fig NN 37809 209 21 . . . 37809 210 1 4a 4a LS 37809 210 2 ) ) -RRB- 37809 210 3 . . . 37809 211 1 _ _ NNP 37809 211 2 Remarks._--Martin Remarks._--Martin NNP 37809 211 3 ( ( -LRB- 37809 211 4 1958 1958 CD 37809 211 5 ) ) -RRB- 37809 211 6 was be VBD 37809 211 7 the the DT 37809 211 8 first first JJ 37809 211 9 author author NN 37809 211 10 to to TO 37809 211 11 point point VB 37809 211 12 out out RP 37809 211 13 that that IN 37809 211 14 _ _ NNP 37809 211 15 S. S. NNP 37809 212 1 campi campi NNP 37809 212 2 _ _ NNP 37809 212 3 was be VBD 37809 212 4 probably probably RB 37809 212 5 a a DT 37809 212 6 subspecies subspecie NNS 37809 212 7 of of IN 37809 212 8 the the DT 37809 212 9 more more RBR 37809 212 10 southern southern JJ 37809 212 11 _ _ NNP 37809 212 12 S. S. NNP 37809 213 1 cystignathoides cystignathoide VBZ 37809 213 2 _ _ NNP 37809 213 3 . . . 37809 214 1 Various various JJ 37809 214 2 references reference NNS 37809 214 3 in in IN 37809 214 4 the the DT 37809 214 5 literature literature NN 37809 214 6 might may MD 37809 214 7 lead lead VB 37809 214 8 one one PRP 37809 214 9 to to TO 37809 214 10 believe believe VB 37809 214 11 that that IN 37809 214 12 the the DT 37809 214 13 two two CD 37809 214 14 were be VBD 37809 214 15 sympatric sympatric JJ 37809 214 16 over over IN 37809 214 17 much much JJ 37809 214 18 of of IN 37809 214 19 northeastern northeastern JJ 37809 214 20 México México NNP 37809 214 21 ; ; : 37809 214 22 this this DT 37809 214 23 error error NN 37809 214 24 was be VBD 37809 214 25 created create VBN 37809 214 26 by by IN 37809 214 27 the the DT 37809 214 28 use use NN 37809 214 29 of of IN 37809 214 30 a a DT 37809 214 31 single single JJ 37809 214 32 character character NN 37809 214 33 ( ( -LRB- 37809 214 34 condition condition NN 37809 214 35 of of IN 37809 214 36 the the DT 37809 214 37 skin skin NN 37809 214 38 of of IN 37809 214 39 the the DT 37809 214 40 venter venter NN 37809 214 41 ) ) -RRB- 37809 214 42 to to TO 37809 214 43 characterize characterize VB 37809 214 44 the the DT 37809 214 45 two two CD 37809 214 46 populations population NNS 37809 214 47 . . . 37809 215 1 Specimens specimen NNS 37809 215 2 from from IN 37809 215 3 southern southern JJ 37809 215 4 Texas Texas NNP 37809 215 5 have have VBP 37809 215 6 a a DT 37809 215 7 smooth smooth JJ 37809 215 8 venter venter NN 37809 215 9 , , , 37809 215 10 lack lack NN 37809 215 11 interorbital interorbital JJ 37809 215 12 bars bar NNS 37809 215 13 and and CC 37809 215 14 have have VBP 37809 215 15 , , , 37809 215 16 in in IN 37809 215 17 general general JJ 37809 215 18 , , , 37809 215 19 a a DT 37809 215 20 brown brown JJ 37809 215 21 ground ground NN 37809 215 22 color color NN 37809 215 23 , , , 37809 215 24 whereas whereas IN 37809 215 25 specimens specimen NNS 37809 215 26 from from IN 37809 215 27 central central JJ 37809 215 28 Veracruz Veracruz NNP 37809 215 29 have have VBP 37809 215 30 an an DT 37809 215 31 areolate areolate NN 37809 215 32 venter venter NN 37809 215 33 , , , 37809 215 34 interorbital interorbital NNP 37809 215 35 light light NN 37809 215 36 and and CC 37809 215 37 dark dark JJ 37809 215 38 bars bar NNS 37809 215 39 and and CC 37809 215 40 a a DT 37809 215 41 yellow yellow JJ 37809 215 42 ground ground NN 37809 215 43 color color NN 37809 215 44 . . . 37809 216 1 In in IN 37809 216 2 southern southern JJ 37809 216 3 Tamaulipas Tamaulipas NNP 37809 216 4 and and CC 37809 216 5 eastern eastern JJ 37809 216 6 San San NNP 37809 216 7 Luis Luis NNP 37809 216 8 Potosí Potosí NNP 37809 216 9 , , , 37809 216 10 these these DT 37809 216 11 characters character NNS 37809 216 12 vary vary VBP 37809 216 13 discordantly discordantly RB 37809 216 14 , , , 37809 216 15 thereby thereby RB 37809 216 16 strongly strongly RB 37809 216 17 suggesting suggest VBG 37809 216 18 that that IN 37809 216 19 the the DT 37809 216 20 two two CD 37809 216 21 populations population NNS 37809 216 22 intergrade intergrade NN 37809 216 23 . . . 37809 217 1 Both both DT 37809 217 2 populations population NNS 37809 217 3 agree agree VBP 37809 217 4 in in IN 37809 217 5 other other JJ 37809 217 6 morphological morphological JJ 37809 217 7 characters character NNS 37809 217 8 ; ; : 37809 217 9 therefore therefore RB 37809 217 10 , , , 37809 217 11 they -PRON- PRP 37809 217 12 are be VBP 37809 217 13 here here RB 37809 217 14 treated treat VBN 37809 217 15 as as IN 37809 217 16 geographic geographic JJ 37809 217 17 variants variant NNS 37809 217 18 . . . 37809 218 1 _ _ NNP 37809 218 2 Etymology._--Named Etymology._--Named NNP 37809 218 3 for for IN 37809 218 4 the the DT 37809 218 5 collector collector NN 37809 218 6 of of IN 37809 218 7 the the DT 37809 218 8 type type NN 37809 218 9 specimens specimen NNS 37809 218 10 , , , 37809 218 11 Mr. Mr. NNP 37809 218 12 R. R. NNP 37809 218 13 D. D. NNP 37809 218 14 Camp Camp NNP 37809 218 15 of of IN 37809 218 16 Brownsville Brownsville NNP 37809 218 17 , , , 37809 218 18 Texas Texas NNP 37809 218 19 . . . 37809 219 1 _ _ NNP 37809 219 2 Distribution._--Lower Distribution._--Lower NNP 37809 219 3 Río Río NNP 37809 219 4 Grande Grande NNP 37809 219 5 embayment embayment NN 37809 219 6 in in IN 37809 219 7 Texas Texas NNP 37809 219 8 to to IN 37809 219 9 central central JJ 37809 219 10 Nuevo Nuevo NNP 37809 219 11 León León NNP 37809 219 12 and and CC 37809 219 13 Tamaulipas Tamaulipas NNP 37809 219 14 , , , 37809 219 15 México México NNP 37809 219 16 . . . 37809 220 1 Intergrades intergrade NNS 37809 220 2 are be VBP 37809 220 3 known know VBN 37809 220 4 from from IN 37809 220 5 southern southern JJ 37809 220 6 Tamaulipas Tamaulipas NNP 37809 220 7 and and CC 37809 220 8 adjacent adjacent JJ 37809 220 9 San San NNP 37809 220 10 Luis Luis NNP 37809 220 11 Potosí Potosí NNP 37809 220 12 , , , 37809 220 13 México México NNP 37809 220 14 ( ( -LRB- 37809 220 15 Fig Fig NNP 37809 220 16 . . . 37809 221 1 5 5 LS 37809 221 2 ) ) -RRB- 37809 221 3 . . . 37809 222 1 _ _ NNP 37809 222 2 Specimens Specimens NNP 37809 222 3 examined._--(113 examined._--(113 NNP 37809 222 4 ) ) -RRB- 37809 222 5 TEXAS TEXAS NNP 37809 222 6 , , , 37809 222 7 Cameron Cameron NNP 37809 222 8 Co. Co. NNP 37809 222 9 : : : 37809 222 10 MCZ MCZ NNP 37809 222 11 10277 10277 CD 37809 222 12 - - SYM 37809 222 13 85 85 CD 37809 222 14 , , , 37809 222 15 10286 10286 CD 37809 222 16 ( ( -LRB- 37809 222 17 10 10 CD 37809 222 18 ) ) -RRB- 37809 222 19 ; ; : 37809 222 20 Brownsville Brownsville NNP 37809 222 21 , , , 37809 222 22 AMNH AMNH NNP 37809 222 23 3215 3215 CD 37809 222 24 , , , 37809 222 25 3218 3218 CD 37809 222 26 - - SYM 37809 222 27 20 20 CD 37809 222 28 , , , 37809 222 29 3221 3221 CD 37809 222 30 ( ( -LRB- 37809 222 31 3 3 CD 37809 222 32 ) ) -RRB- 37809 222 33 , , , 37809 222 34 5376 5376 CD 37809 222 35 , , , 37809 222 36 62117 62117 CD 37809 222 37 , , , 37809 222 38 FMNH FMNH NNP 37809 222 39 105336 105336 CD 37809 222 40 , , , 37809 222 41 KU KU NNP 37809 222 42 8135 8135 CD 37809 222 43 - - SYM 37809 222 44 39 39 CD 37809 222 45 , , , 37809 222 46 MCZ MCZ NNP 37809 222 47 3738 3738 CD 37809 222 48 - - SYM 37809 222 49 42 42 CD 37809 222 50 , , , 37809 222 51 3743 3743 CD 37809 222 52 ( ( -LRB- 37809 222 53 10 10 CD 37809 222 54 ) ) -RRB- 37809 222 55 , , , 37809 222 56 TCWC TCWC NNP 37809 222 57 5908 5908 CD 37809 222 58 , , , 37809 222 59 7139 7139 CD 37809 222 60 , , , 37809 222 61 TNHC TNHC NNP 37809 222 62 92 92 CD 37809 222 63 - - SYM 37809 222 64 94 94 CD 37809 222 65 , , , 37809 222 66 20909 20909 CD 37809 222 67 , , , 37809 222 68 UMMZ UMMZ NNP 37809 222 69 51760 51760 CD 37809 222 70 , , , 37809 222 71 54031 54031 CD 37809 222 72 ( ( -LRB- 37809 222 73 5 5 CD 37809 222 74 ) ) -RRB- 37809 222 75 , , , 37809 222 76 USNM USNM NNP 37809 222 77 52290 52290 CD 37809 222 78 ( ( -LRB- 37809 222 79 holotype holotype NN 37809 222 80 ) ) -RRB- 37809 222 81 ; ; : 37809 222 82 22 22 CD 37809 222 83 mi mi JJ 37809 222 84 . . . 37809 223 1 SE SE NNP 37809 223 2 Brownsville Brownsville NNP 37809 223 3 , , , 37809 223 4 TNMC TNMC NNP 37809 223 5 14223 14223 CD 37809 223 6 ; ; : 37809 223 7 8 8 CD 37809 223 8 mi mi NNS 37809 223 9 . . . 37809 224 1 SW SW NNP 37809 224 2 Brownsville Brownsville NNP 37809 224 3 , , , 37809 224 4 UMMZ UMMZ NNP 37809 224 5 101127 101127 CD 37809 224 6 ( ( -LRB- 37809 224 7 3 3 CD 37809 224 8 ) ) -RRB- 37809 224 9 ; ; : 37809 224 10 Harlingen Harlingen NNP 37809 224 11 , , , 37809 224 12 AMNH AMNH NNP 37809 224 13 62118 62118 CD 37809 224 14 , , , 37809 224 15 UMMZ UMMZ NNP 37809 224 16 105200 105200 CD 37809 224 17 - - SYM 37809 224 18 205 205 CD 37809 224 19 , , , 37809 224 20 105206 105206 CD 37809 224 21 ( ( -LRB- 37809 224 22 5 5 CD 37809 224 23 ) ) -RRB- 37809 224 24 , , , 37809 224 25 105207 105207 CD 37809 224 26 ( ( -LRB- 37809 224 27 4 4 CD 37809 224 28 ) ) -RRB- 37809 224 29 . . . 37809 225 1 _ _ NNP 37809 225 2 Hidalgo Hidalgo NNP 37809 225 3 Co. Co. NNP 37809 225 4 _ _ NNP 37809 225 5 : : : 37809 225 6 Bentsen Bentsen NNP 37809 225 7 - - HYPH 37809 225 8 Río Río NNP 37809 225 9 Grande Grande NNP 37809 225 10 State State NNP 37809 225 11 Park Park NNP 37809 225 12 , , , 37809 225 13 UMMZ UMMZ NNP 37809 225 14 114378 114378 CD 37809 225 15 ; ; : 37809 225 16 6 6 CD 37809 225 17 mi mi NNS 37809 225 18 . . . 37809 226 1 S S NNP 37809 226 2 McAllen McAllen NNP 37809 226 3 , , , 37809 226 4 TNHC TNHC NNP 37809 226 5 7136 7136 CD 37809 226 6 - - SYM 37809 226 7 39 39 CD 37809 226 8 ; ; : 37809 226 9 Santa Santa NNP 37809 226 10 Ana Ana NNP 37809 226 11 Refuge Refuge NNP 37809 226 12 , , , 37809 226 13 TCWC TCWC NNP 37809 226 14 13495 13495 CD 37809 226 15 - - SYM 37809 226 16 96 96 CD 37809 226 17 ; ; : 37809 226 18 Weslaco Weslaco NNP 37809 226 19 , , , 37809 226 20 TCWC TCWC NNP 37809 226 21 17658 17658 CD 37809 226 22 - - SYM 37809 226 23 60 60 CD 37809 226 24 . . . 37809 227 1 MEXICO MEXICO NNP 37809 227 2 , , , 37809 227 3 _ _ NNP 37809 227 4 Nuevo Nuevo NNP 37809 227 5 León León NNP 37809 227 6 _ _ NNP 37809 227 7 : : : 37809 227 8 Salto Salto NNP 37809 227 9 Cola Cola NNP 37809 227 10 de de NNP 37809 227 11 Caballo Caballo NNP 37809 227 12 , , , 37809 227 13 AMNH AMNH NNP 37809 227 14 57953 57953 CD 37809 227 15 - - SYM 37809 227 16 54 54 CD 37809 227 17 , , , 37809 227 18 FMNH FMNH NNP 37809 227 19 30644 30644 CD 37809 227 20 - - SYM 37809 227 21 45 45 CD 37809 227 22 , , , 37809 227 23 37169 37169 CD 37809 227 24 - - SYM 37809 227 25 70 70 CD 37809 227 26 ; ; : 37809 227 27 Monterrey Monterrey NNP 37809 227 28 , , , 37809 227 29 UIMNH UIMNH NNP 37809 227 30 13324 13324 CD 37809 227 31 ; ; : 37809 227 32 40 40 CD 37809 227 33 km km NN 37809 227 34 . . . 37809 228 1 SE SE NNP 37809 228 2 Monterrey Monterrey NNP 37809 228 3 , , , 37809 228 4 UIMNH UIMNH NNP 37809 228 5 3686 3686 CD 37809 228 6 . . . 37809 229 1 _ _ NNP 37809 229 2 Tamaulipas Tamaulipas NNP 37809 229 3 _ _ NNP 37809 229 4 : : : 37809 229 5 80 80 CD 37809 229 6 km km NN 37809 229 7 . . . 37809 230 1 Matamoros Matamoros NNP 37809 230 2 , , , 37809 230 3 FMNH FMNH NNP 37809 230 4 27150 27150 CD 37809 230 5 ( ( -LRB- 37809 230 6 13 13 CD 37809 230 7 ) ) -RRB- 37809 230 8 . . . 37809 231 1 Intergrades Intergrades NNP 37809 231 2 [ [ -LRB- 37809 231 3 _ _ NNP 37809 231 4 S. S. NNP 37809 232 1 c. c. NNP 37809 232 2 cystignathoides cystignathoides NNP 37809 232 3 _ _ NNP 37809 232 4 × × NN 37809 232 5 _ _ NNP 37809 232 6 S. S. NNP 37809 233 1 c. c. NNP 37809 233 2 campi campi NNP 37809 233 3 _ _ NNP 37809 233 4 ( ( -LRB- 37809 233 5 88 88 CD 37809 233 6 ) ) -RRB- 37809 233 7 ] ] -RRB- 37809 233 8 MÉXICO MÉXICO NNP 37809 233 9 , , , 37809 233 10 _ _ NNP 37809 233 11 San San NNP 37809 233 12 Luis Luis NNP 37809 233 13 Potosí Potosí NNP 37809 233 14 _ _ NNP 37809 233 15 : : : 37809 233 16 5 5 CD 37809 233 17 km km NN 37809 233 18 . . . 37809 234 1 E E NNP 37809 234 2 Ciudad Ciudad NNP 37809 234 3 del del NNP 37809 234 4 Maiz Maiz NNP 37809 234 5 , , , 37809 234 6 UMMZ UMMZ NNP 37809 234 7 106435 106435 CD 37809 234 8 ; ; : 37809 234 9 16 16 CD 37809 234 10 km km NNS 37809 234 11 . . . 37809 235 1 W W NNP 37809 235 2 Naranjo Naranjo NNP 37809 235 3 , , , 37809 235 4 FMNH FMNH NNP 37809 235 5 104584 104584 CD 37809 235 6 ; ; : 37809 235 7 Salto Salto NNP 37809 235 8 de de NNP 37809 235 9 Agua Agua NNP 37809 235 10 , , , 37809 235 11 34 34 CD 37809 235 12 km km NN 37809 235 13 . . . 37809 236 1 WSW WSW NNP 37809 236 2 Antigua Antigua NNP 37809 236 3 Morelos Morelos NNP 37809 236 4 , , , 37809 236 5 TCWC TCWC NNP 37809 236 6 6980 6980 CD 37809 236 7 . . . 37809 237 1 _ _ NNP 37809 237 2 Tamaulipas Tamaulipas NNP 37809 237 3 _ _ NNP 37809 237 4 : : : 37809 237 5 5 5 CD 37809 237 6 km km NN 37809 237 7 . . . 37809 238 1 W W NNP 37809 238 2 Acuña Acuña NNP 37809 238 3 , , , 37809 238 4 1060 1060 CD 37809 238 5 m. m. NN 37809 238 6 , , , 37809 238 7 UMMZ UMMZ NNP 37809 238 8 101172 101172 CD 37809 238 9 , , , 37809 238 10 101173 101173 CD 37809 238 11 ( ( -LRB- 37809 238 12 16 16 CD 37809 238 13 ) ) -RRB- 37809 238 14 , , , 37809 238 15 101174 101174 CD 37809 238 16 - - SYM 37809 238 17 76 76 CD 37809 238 18 , , , 37809 238 19 101177 101177 CD 37809 238 20 ( ( -LRB- 37809 238 21 6 6 CD 37809 238 22 ) ) -RRB- 37809 238 23 ; ; : 37809 238 24 14.5 14.5 CD 37809 238 25 km km NNS 37809 238 26 . . . 37809 239 1 NNW NNW NNP 37809 239 2 Chamal Chamal NNP 37809 239 3 , , , 37809 239 4 430 430 CD 37809 239 5 m. m. NN 37809 239 6 , , , 37809 239 7 UMMZ UMMZ NNP 37809 239 8 111337 111337 CD 37809 239 9 ( ( -LRB- 37809 239 10 2 2 CD 37809 239 11 ) ) -RRB- 37809 239 12 ; ; : 37809 239 13 20 20 CD 37809 239 14 km km NN 37809 239 15 . . . 37809 240 1 NNW NNW NNP 37809 240 2 Chamal Chamal NNP 37809 240 3 , , , 37809 240 4 700 700 CD 37809 240 5 m. m. NN 37809 240 6 , , , 37809 240 7 UMMZ UMMZ NNP 37809 240 8 111338 111338 CD 37809 240 9 ( ( -LRB- 37809 240 10 11 11 CD 37809 240 11 ) ) -RRB- 37809 240 12 ; ; : 37809 240 13 8 8 CD 37809 240 14 km km NN 37809 240 15 . . . 37809 241 1 N n CD 37809 241 2 Gómez Gómez NNP 37809 241 3 Farías Farías NNP 37809 241 4 , , , 37809 241 5 450 450 CD 37809 241 6 m. m. NN 37809 241 7 , , , 37809 241 8 UMMZ UMMZ NNP 37809 241 9 101165 101165 CD 37809 241 10 ; ; : 37809 241 11 8 8 CD 37809 241 12 km km NN 37809 241 13 . . . 37809 242 1 NE NE NNP 37809 242 2 Gómez Gómez NNP 37809 242 3 Farías Farías NNP 37809 242 4 , , , 37809 242 5 Pano Pano NNP 37809 242 6 Ayuctle Ayuctle NNP 37809 242 7 , , , 37809 242 8 UMMZ UMMZ NNP 37809 242 9 102264 102264 CD 37809 242 10 , , , 37809 242 11 102924 102924 CD 37809 242 12 ( ( -LRB- 37809 242 13 6 6 CD 37809 242 14 ) ) -RRB- 37809 242 15 ; ; : 37809 242 16 8 8 CD 37809 242 17 km km NN 37809 242 18 . . . 37809 243 1 NW NW NNP 37809 243 2 Gómez Gómez NNP 37809 243 3 Farías Farías NNP 37809 243 4 , , , 37809 243 5 1060 1060 CD 37809 243 6 m. m. NN 37809 243 7 , , , 37809 243 8 LSUMZ LSUMZ NNP 37809 243 9 11084 11084 CD 37809 243 10 , , , 37809 243 11 UMMZ UMMZ NNP 37809 243 12 101199 101199 CD 37809 243 13 , , , 37809 243 14 102928 102928 CD 37809 243 15 ( ( -LRB- 37809 243 16 5 5 CD 37809 243 17 ) ) -RRB- 37809 243 18 , , , 37809 243 19 102929 102929 CD 37809 243 20 - - SYM 37809 243 21 32 32 CD 37809 243 22 , , , 37809 243 23 110124 110124 CD 37809 243 24 ( ( -LRB- 37809 243 25 3 3 CD 37809 243 26 ) ) -RRB- 37809 243 27 ; ; : 37809 243 28 Río Río NNP 37809 243 29 Guayala Guayala NNP 37809 243 30 , , , 37809 243 31 near near IN 37809 243 32 Magiscatzin Magiscatzin NNP 37809 243 33 , , , 37809 243 34 MCZ MCZ NNP 37809 243 35 24138 24138 CD 37809 243 36 - - SYM 37809 243 37 42 42 CD 37809 243 38 , , , 37809 243 39 85071 85071 CD 37809 243 40 - - SYM 37809 243 41 81 81 CD 37809 243 42 , , , 37809 243 43 UMMZ UMMZ NNP 37809 243 44 88242 88242 CD 37809 243 45 ( ( -LRB- 37809 243 46 2 2 CD 37809 243 47 ) ) -RRB- 37809 243 48 ; ; : 37809 243 49 Magiscatzin Magiscatzin NNP 37809 243 50 , , , 37809 243 51 TCWC TCWC VBD 37809 243 52 6981 6981 CD 37809 243 53 ; ; : 37809 243 54 Las Las NNP 37809 243 55 Yucas Yucas NNP 37809 243 56 , , , 37809 243 57 north north RB 37809 243 58 of of IN 37809 243 59 Aldama Aldama NNP 37809 243 60 , , , 37809 243 61 MCZ MCZ NNP 37809 243 62 29665 29665 CD 37809 243 63 - - SYM 37809 243 64 68 68 CD 37809 243 65 ; ; : 37809 243 66 16 16 CD 37809 243 67 km km NN 37809 243 68 . . . 37809 244 1 NE NE NNP 37809 244 2 Zamorina Zamorina NNP 37809 244 3 , , , 37809 244 4 UMMZ UMMZ NNP 37809 244 5 101124 101124 CD 37809 244 6 . . . 37809 245 1 [ [ -LRB- 37809 245 2 Illustration illustration NN 37809 245 3 : : : 37809 245 4 FIG FIG NNP 37809 245 5 . . . 37809 246 1 4 4 LS 37809 246 2 : : : 37809 246 3 _ _ NNP 37809 246 4 Syrrhophus Syrrhophus NNP 37809 246 5 cystignathoides cystignathoide VBZ 37809 246 6 campi campi NNP 37809 246 7 _ _ NNP 37809 246 8 ( ( -LRB- 37809 246 9 left leave VBN 37809 246 10 , , , 37809 246 11 TCWC TCWC NNP 37809 246 12 13490 13490 CD 37809 246 13 ) ) -RRB- 37809 246 14 and and CC 37809 246 15 _ _ NNP 37809 246 16 S. S. NNP 37809 247 1 c. c. NNP 37809 247 2 cystignathoides cystignathoides NNP 37809 247 3 _ _ NNP 37809 247 4 ( ( -LRB- 37809 247 5 right right UH 37809 247 6 , , , 37809 247 7 KU KU NNP 37809 247 8 105500 105500 CD 37809 247 9 ) ) -RRB- 37809 247 10 . . . 37809 248 1 Dorsal dorsal NN 37809 248 2 views view NNS 37809 248 3 ×2 ×2 NNS 37809 248 4 , , , 37809 248 5 sides side NNS 37809 248 6 of of IN 37809 248 7 heads head NNS 37809 248 8 ×3 ×3 NNP 37809 248 9 . . . 37809 248 10 ] ] -RRB- 37809 249 1 = = NFP 37809 249 2 Syrrhophus Syrrhophus NNP 37809 249 3 cystignathoides cystignathoide VBZ 37809 249 4 cystignathoides= cystignathoides= JJ 37809 249 5 ( ( -LRB- 37809 249 6 Cope Cope NNP 37809 249 7 ) ) -RRB- 37809 249 8 , , , 37809 249 9 New new JJ 37809 249 10 combination combination NN 37809 249 11 _ _ NNP 37809 249 12 Phyllobates Phyllobates NNPS 37809 249 13 cystignathoides cystignathoide VBZ 37809 249 14 _ _ NNP 37809 249 15 Cope Cope NNP 37809 249 16 , , , 37809 249 17 1877:89 1877:89 CD 37809 249 18 - - SYM 37809 249 19 90 90 CD 37809 249 20 [ [ -LRB- 37809 249 21 Syntypes.--USNM Syntypes.--USNM NNP 37809 249 22 32402 32402 CD 37809 249 23 - - SYM 37809 249 24 32409 32409 CD 37809 249 25 , , , 37809 249 26 from from IN 37809 249 27 Potrero Potrero NNP 37809 249 28 , , , 37809 249 29 near near IN 37809 249 30 Córdoba Córdoba NNP 37809 249 31 , , , 37809 249 32 Veracruz Veracruz NNP 37809 249 33 , , , 37809 249 34 México México NNP 37809 249 35 , , , 37809 249 36 collected collect VBN 37809 249 37 by by IN 37809 249 38 Francis Francis NNP 37809 249 39 Sumichrast Sumichrast NNP 37809 249 40 ] ] -RRB- 37809 249 41 . . . 37809 250 1 Boulenger boulenger NN 37809 250 2 , , , 37809 250 3 1882:196 1882:196 CD 37809 250 4 . . . 37809 251 1 _ _ NNP 37809 251 2 Syrrhophus Syrrhophus NNP 37809 251 3 cystignathoides cystignathoide VBZ 37809 251 4 _ _ NNP 37809 251 5 : : : 37809 251 6 Cope cope VB 37809 251 7 , , , 37809 251 8 1879:268 1879:268 CD 37809 251 9 . . . 37809 252 1 Kellogg Kellogg NNP 37809 252 2 , , , 37809 252 3 1932 1932 CD 37809 252 4 : : : 37809 252 5 126 126 CD 37809 252 6 - - SYM 37809 252 7 27 27 CD 37809 252 8 . . . 37809 253 1 Taylor Taylor NNP 37809 253 2 and and CC 37809 253 3 Smith Smith NNP 37809 253 4 , , , 37809 253 5 1945 1945 CD 37809 253 6 : : : 37809 253 7 582 582 CD 37809 253 8 - - SYM 37809 253 9 83 83 CD 37809 253 10 . . . 37809 254 1 Smith Smith NNP 37809 254 2 and and CC 37809 254 3 Taylor Taylor NNP 37809 254 4 , , , 37809 254 5 1948:50 1948:50 CD 37809 254 6 . . . 37809 255 1 Martin Martin NNP 37809 255 2 , , , 37809 255 3 1958:49 1958:49 CD 37809 255 4 . . . 37809 256 1 _ _ NNP 37809 256 2 Syrrhaphus Syrrhaphus NNP 37809 256 3 cystignathoides cystignathoide VBZ 37809 256 4 _ _ NNP 37809 256 5 : : : 37809 256 6 Günther Günther NNP 37809 256 7 , , , 37809 256 8 1900:218 1900:218 CD 37809 256 9 . . . 37809 257 1 _ _ NNP 37809 257 2 Syrraphus Syrraphus NNP 37809 257 3 cystignathoides cystignathoide VBZ 37809 257 4 _ _ NNP 37809 257 5 : : : 37809 257 6 Díaz Díaz NNP 37809 257 7 de de FW 37809 257 8 León León NNP 37809 257 9 , , , 37809 257 10 1904:10 1904:10 CD 37809 257 11 . . . 37809 258 1 _ _ NNP 37809 258 2 Syrrhopus Syrrhopus NNP 37809 258 3 cystignathoides cystignathoide VBZ 37809 258 4 _ _ NNP 37809 258 5 : : : 37809 258 6 Barbour Barbour NNP 37809 258 7 and and CC 37809 258 8 Loveridge Loveridge NNP 37809 258 9 , , , 37809 258 10 1946:170 1946:170 CD 37809 258 11 . . . 37809 259 1 [ [ -LRB- 37809 259 2 Illustration illustration NN 37809 259 3 : : : 37809 259 4 FIG FIG NNP 37809 259 5 . . . 37809 260 1 5 5 LS 37809 260 2 : : : 37809 260 3 Distribution distribution NN 37809 260 4 of of IN 37809 260 5 _ _ NNP 37809 260 6 Syrrhophus Syrrhophus NNP 37809 260 7 cystignathoides cystignathoide VBZ 37809 260 8 campi campi NNP 37809 260 9 _ _ NNP 37809 260 10 ( ( -LRB- 37809 260 11 solid solid JJ 37809 260 12 symbols symbol NNS 37809 260 13 ) ) -RRB- 37809 260 14 and and CC 37809 260 15 the the DT 37809 260 16 nominate nominate NN 37809 260 17 subspecies subspecie NNS 37809 260 18 ( ( -LRB- 37809 260 19 open open JJ 37809 260 20 symbols symbol NNS 37809 260 21 ) ) -RRB- 37809 260 22 . . . 37809 260 23 ] ] -RRB- 37809 261 1 _ _ NNP 37809 261 2 Diagnosis._--Venter diagnosis._--venter CD 37809 261 3 areolate areolate NN 37809 261 4 ; ; : 37809 261 5 interorbital interorbital JJ 37809 261 6 light light NN 37809 261 7 and and CC 37809 261 8 dark dark JJ 37809 261 9 bars bar NNS 37809 261 10 present present JJ 37809 261 11 ; ; : 37809 261 12 ground ground NN 37809 261 13 color color NN 37809 261 14 yellow yellow NNP 37809 261 15 to to IN 37809 261 16 brownish brownish NN 37809 261 17 - - HYPH 37809 261 18 yellow yellow NN 37809 261 19 in in IN 37809 261 20 life life NN 37809 261 21 ( ( -LRB- 37809 261 22 Fig fig NN 37809 261 23 . . . 37809 262 1 4b 4b LS 37809 262 2 ) ) -RRB- 37809 262 3 . . . 37809 263 1 _ _ NNP 37809 263 2 Remarks._--Firschein Remarks._--Firschein NNP 37809 263 3 ( ( -LRB- 37809 263 4 1954 1954 CD 37809 263 5 ) ) -RRB- 37809 263 6 briefly briefly RB 37809 263 7 considered consider VBD 37809 263 8 the the DT 37809 263 9 status status NN 37809 263 10 of of IN 37809 263 11 Peters Peters NNP 37809 263 12 ' ' POS 37809 263 13 ( ( -LRB- 37809 263 14 1871 1871 CD 37809 263 15 ) ) -RRB- 37809 263 16 _ _ NNP 37809 263 17 Phyllobates Phyllobates NNPS 37809 263 18 verruculatus verruculatus VBP 37809 263 19 _ _ NNP 37809 263 20 and and CC 37809 263 21 noted note VBD 37809 263 22 that that IN 37809 263 23 if if IN 37809 263 24 it -PRON- PRP 37809 263 25 was be VBD 37809 263 26 a a DT 37809 263 27 _ _ NNP 37809 263 28 Syrrhophus Syrrhophus NNP 37809 263 29 _ _ NNP 37809 263 30 it -PRON- PRP 37809 263 31 would would MD 37809 263 32 probably probably RB 37809 263 33 be be VB 37809 263 34 referrable referrable JJ 37809 263 35 to to IN 37809 263 36 _ _ NNP 37809 263 37 S. S. NNP 37809 264 1 cystignathoides cystignathoide VBZ 37809 264 2 _ _ NNP 37809 264 3 . . . 37809 265 1 Peters Peters NNP 37809 265 2 ' ' POS 37809 265 3 ( ( -LRB- 37809 265 4 1871 1871 CD 37809 265 5 ) ) -RRB- 37809 265 6 original original JJ 37809 265 7 description description NN 37809 265 8 corresponds correspond VBZ 37809 265 9 well well RB 37809 265 10 with with IN 37809 265 11 _ _ NNP 37809 265 12 S. S. NNP 37809 266 1 cystignathoides cystignathoide VBZ 37809 266 2 _ _ NNP 37809 266 3 , , , 37809 266 4 and and CC 37809 266 5 the the DT 37809 266 6 type type NN 37809 266 7 - - HYPH 37809 266 8 locality locality NN 37809 266 9 ( ( -LRB- 37809 266 10 " " `` 37809 266 11 Huanusco Huanusco NNP 37809 266 12 " " '' 37809 266 13 = = SYM 37809 266 14 Huatusco Huatusco NNP 37809 266 15 ) ) -RRB- 37809 266 16 is be VBZ 37809 266 17 within within IN 37809 266 18 the the DT 37809 266 19 range range NN 37809 266 20 of of IN 37809 266 21 that that DT 37809 266 22 species specie NNS 37809 266 23 . . . 37809 267 1 Firschein Firschein NNP 37809 267 2 ( ( -LRB- 37809 267 3 1954 1954 CD 37809 267 4 ) ) -RRB- 37809 267 5 expressed express VBD 37809 267 6 doubt doubt NN 37809 267 7 that that IN 37809 267 8 _ _ NNP 37809 267 9 verruculatus verruculatus NNP 37809 267 10 _ _ NNP 37809 267 11 was be VBD 37809 267 12 a a DT 37809 267 13 _ _ NNP 37809 267 14 Syrrhophus Syrrhophus NNP 37809 267 15 _ _ NNP 37809 267 16 , , , 37809 267 17 because because IN 37809 267 18 Peters Peters NNP 37809 267 19 placed place VBD 37809 267 20 it -PRON- PRP 37809 267 21 in in IN 37809 267 22 another another DT 37809 267 23 genus genus NN 37809 267 24 . . . 37809 268 1 However however RB 37809 268 2 , , , 37809 268 3 Peters Peters NNP 37809 268 4 described describe VBD 37809 268 5 _ _ NNP 37809 268 6 verruculatus verruculatus NNP 37809 268 7 _ _ NNP 37809 268 8 a a DT 37809 268 9 decade decade NN 37809 268 10 before before IN 37809 268 11 Cope Cope NNP 37809 268 12 diagnosed diagnose VBD 37809 268 13 the the DT 37809 268 14 genus genus JJ 37809 268 15 Syrrhophus Syrrhophus NNP 37809 268 16 . . . 37809 269 1 Most Most JJS 37809 269 2 frogs frog NNS 37809 269 3 now now RB 37809 269 4 called call VBN 37809 269 5 _ _ NNP 37809 269 6 Syrrhophus Syrrhophus NNP 37809 269 7 _ _ NNP 37809 269 8 , , , 37809 269 9 plus plus CC 37809 269 10 a a DT 37809 269 11 number number NN 37809 269 12 of of IN 37809 269 13 lower low JJR 37809 269 14 Central central JJ 37809 269 15 American american JJ 37809 269 16 frogs frog NNS 37809 269 17 now now RB 37809 269 18 placed place VBN 37809 269 19 in in IN 37809 269 20 a a DT 37809 269 21 variety variety NN 37809 269 22 of of IN 37809 269 23 genera genera NN 37809 269 24 were be VBD 37809 269 25 placed place VBN 37809 269 26 in in IN 37809 269 27 _ _ NNP 37809 269 28 Phyllobates Phyllobates NNPS 37809 269 29 _ _ NNP 37809 269 30 by by IN 37809 269 31 Boulenger Boulenger NNP 37809 269 32 , , , 37809 269 33 Cope Cope NNP 37809 269 34 , , , 37809 269 35 and and CC 37809 269 36 Peters Peters NNP 37809 269 37 . . . 37809 270 1 The the DT 37809 270 2 types type NNS 37809 270 3 of of IN 37809 270 4 _ _ NNP 37809 270 5 Phyllobates Phyllobates NNPS 37809 270 6 verruculatus verruculatus NNP 37809 270 7 _ _ NNP 37809 270 8 were be VBD 37809 270 9 destroyed destroy VBN 37809 270 10 during during IN 37809 270 11 World World NNP 37809 270 12 War War NNP 37809 270 13 II II NNP 37809 270 14 ( ( -LRB- 37809 270 15 Günther Günther NNP 37809 270 16 Peters Peters NNP 37809 270 17 , , , 37809 270 18 _ _ NNP 37809 270 19 in in IN 37809 270 20 litt litt NNP 37809 270 21 . . . 37809 270 22 _ _ NNP 37809 270 23 ) ) -RRB- 37809 270 24 ; ; : 37809 270 25 the the DT 37809 270 26 specimens specimen NNS 37809 270 27 subsequently subsequently RB 37809 270 28 assigned assign VBN 37809 270 29 to to IN 37809 270 30 the the DT 37809 270 31 taxon taxon NN 37809 270 32 by by IN 37809 270 33 Kellogg Kellogg NNP 37809 270 34 ( ( -LRB- 37809 270 35 1932 1932 CD 37809 270 36 ) ) -RRB- 37809 270 37 are be VBP 37809 270 38 _ _ NNP 37809 270 39 Syrrhophus Syrrhophus NNP 37809 270 40 cystignathoides cystignathoide VBZ 37809 270 41 _ _ NNP 37809 270 42 . . . 37809 271 1 Because because IN 37809 271 2 the the DT 37809 271 3 type type NN 37809 271 4 specimens specimen NNS 37809 271 5 are be VBP 37809 271 6 lost lose VBN 37809 271 7 and and CC 37809 271 8 because because IN 37809 271 9 the the DT 37809 271 10 name name NN 37809 271 11 antedates antedate VBZ 37809 271 12 the the DT 37809 271 13 more more RBR 37809 271 14 established established JJ 37809 271 15 name name NN 37809 271 16 , , , 37809 271 17 _ _ NNP 37809 271 18 cystignathoides cystignathoide VBZ 37809 271 19 _ _ NNP 37809 271 20 , , , 37809 271 21 I -PRON- PRP 37809 271 22 favor favor VBP 37809 271 23 retaining retain VBG 37809 271 24 _ _ NNP 37809 271 25 Phyllobates Phyllobates NNPS 37809 271 26 verruculatus verruculatus NNP 37809 271 27 _ _ NNP 37809 271 28 Peters Peters NNP 37809 271 29 as as IN 37809 271 30 a a DT 37809 271 31 _ _ NNP 37809 271 32 nomen nomen NNP 37809 271 33 dubium dubium NN 37809 271 34 _ _ NNP 37809 271 35 . . . 37809 272 1 Smith Smith NNP 37809 272 2 and and CC 37809 272 3 Taylor Taylor NNP 37809 272 4 ( ( -LRB- 37809 272 5 1948 1948 CD 37809 272 6 ) ) -RRB- 37809 272 7 reported report VBD 37809 272 8 _ _ NNP 37809 272 9 S. S. NNP 37809 273 1 verruculatus verruculatus NNP 37809 273 2 _ _ NNP 37809 273 3 from from IN 37809 273 4 Tianguistengo Tianguistengo NNP 37809 273 5 , , , 37809 273 6 Hidalgo Hidalgo NNP 37809 273 7 , , , 37809 273 8 México México NNP 37809 273 9 . . . 37809 274 1 These these DT 37809 274 2 specimens specimen NNS 37809 274 3 are be VBP 37809 274 4 examples example NNS 37809 274 5 of of IN 37809 274 6 _ _ NNP 37809 274 7 verrucipes verrucipes NNP 37809 274 8 _ _ NNP 37809 274 9 . . . 37809 275 1 Smith Smith NNP 37809 275 2 ( ( -LRB- 37809 275 3 1947 1947 CD 37809 275 4 ) ) -RRB- 37809 275 5 reported report VBD 37809 275 6 a a DT 37809 275 7 specimen speciman NNS 37809 275 8 of of IN 37809 275 9 _ _ NNP 37809 275 10 verruculatus verruculatus NNP 37809 275 11 _ _ NNP 37809 275 12 from from IN 37809 275 13 San San NNP 37809 275 14 Lorenzo Lorenzo NNP 37809 275 15 , , , 37809 275 16 Veracruz Veracruz NNP 37809 275 17 . . . 37809 276 1 Firschein Firschein NNP 37809 276 2 ( ( -LRB- 37809 276 3 1954 1954 CD 37809 276 4 ) ) -RRB- 37809 276 5 referred refer VBD 37809 276 6 it -PRON- PRP 37809 276 7 to to IN 37809 276 8 _ _ NNP 37809 276 9 cystignathoides cystignathoides NNP 37809 276 10 _ _ NNP 37809 276 11 , , , 37809 276 12 and and CC 37809 276 13 Duellman Duellman NNP 37809 276 14 ( ( -LRB- 37809 276 15 1960 1960 CD 37809 276 16 ) ) -RRB- 37809 276 17 concluded conclude VBD 37809 276 18 that that IN 37809 276 19 both both DT 37809 276 20 authors author NNS 37809 276 21 were be VBD 37809 276 22 in in IN 37809 276 23 error error NN 37809 276 24 and and CC 37809 276 25 that that IN 37809 276 26 the the DT 37809 276 27 specimen speciman NNS 37809 276 28 ( ( -LRB- 37809 276 29 USNM USNM NNP 37809 276 30 123530 123530 CD 37809 276 31 ) ) -RRB- 37809 276 32 was be VBD 37809 276 33 a a DT 37809 276 34 _ _ NNP 37809 276 35 leprus leprus NNP 37809 276 36 _ _ NNP 37809 276 37 . . . 37809 277 1 _ _ NNP 37809 277 2 Etymology._--The Etymology._--The NNP 37809 277 3 trivial trivial JJ 37809 277 4 name name NN 37809 277 5 is be VBZ 37809 277 6 the the DT 37809 277 7 diminutive diminutive NN 37809 277 8 of of IN 37809 277 9 _ _ NNP 37809 277 10 Cystignathus Cystignathus NNP 37809 277 11 _ _ NNP 37809 277 12 , , , 37809 277 13 a a DT 37809 277 14 once once RB 37809 277 15 - - HYPH 37809 277 16 used use VBN 37809 277 17 generic generic JJ 37809 277 18 name name NN 37809 277 19 for for IN 37809 277 20 several several JJ 37809 277 21 leptodactylid leptodactylid JJ 37809 277 22 frogs frog NNS 37809 277 23 . . . 37809 278 1 _ _ NNP 37809 278 2 Distribution._--Low Distribution._--Low NNP 37809 278 3 and and CC 37809 278 4 moderate moderate JJ 37809 278 5 elevations elevation NNS 37809 278 6 in in IN 37809 278 7 the the DT 37809 278 8 foothills foothill NNS 37809 278 9 along along IN 37809 278 10 the the DT 37809 278 11 Sierra Sierra NNP 37809 278 12 Madre Madre NNP 37809 278 13 Oriental Oriental NNP 37809 278 14 from from IN 37809 278 15 eastern eastern JJ 37809 278 16 San San NNP 37809 278 17 Luis Luis NNP 37809 278 18 Potosí Potosí NNP 37809 278 19 to to IN 37809 278 20 Central Central NNP 37809 278 21 Veracruz Veracruz NNP 37809 278 22 , , , 37809 278 23 México México NNP 37809 278 24 ( ( -LRB- 37809 278 25 Fig Fig NNP 37809 278 26 . . . 37809 279 1 5 5 LS 37809 279 2 ) ) -RRB- 37809 279 3 . . . 37809 280 1 _ _ NNP 37809 280 2 Specimens Specimens NNP 37809 280 3 examined._--(130 examined._--(130 NNP 37809 280 4 ) ) -RRB- 37809 280 5 , , , 37809 280 6 MÉXICO MÉXICO NNP 37809 280 7 , , , 37809 280 8 _ _ NNP 37809 280 9 Puebla Puebla NNP 37809 280 10 _ _ NNP 37809 280 11 : : : 37809 280 12 Necaxa Necaxa NNP 37809 280 13 , , , 37809 280 14 UMMZ UMMZ NNP 37809 280 15 69519 69519 CD 37809 280 16 - - SYM 37809 280 17 20 20 CD 37809 280 18 . . . 37809 281 1 _ _ NNP 37809 281 2 San San NNP 37809 281 3 Luis Luis NNP 37809 281 4 Potosí Potosí NNP 37809 281 5 _ _ NNP 37809 281 6 : : : 37809 281 7 5 5 CD 37809 281 8 km km NN 37809 281 9 . . . 37809 282 1 W W NNP 37809 282 2 Aguismón Aguismón NNP 37809 282 3 , , , 37809 282 4 LSUMZ LSUMZ NNP 37809 282 5 4962 4962 CD 37809 282 6 - - SYM 37809 282 7 63 63 CD 37809 282 8 ; ; : 37809 282 9 along along IN 37809 282 10 Río Río NNP 37809 282 11 Axtla Axtla NNP 37809 282 12 , , , 37809 282 13 road road NN 37809 282 14 to to IN 37809 282 15 Xilitla Xilitla NNP 37809 282 16 , , , 37809 282 17 UMMZ UMMZ NNP 37809 282 18 105500 105500 CD 37809 282 19 ; ; : 37809 282 20 Tamazunchale Tamazunchale NNP 37809 282 21 , , , 37809 282 22 UIMNH UIMNH NNP 37809 282 23 3199 3199 CD 37809 282 24 ; ; : 37809 282 25 6.5 6.5 CD 37809 282 26 km km NN 37809 282 27 . . . 37809 283 1 N N NNP 37809 283 2 Tamazunchale Tamazunchale NNP 37809 283 3 , , , 37809 283 4 UMMZ UMMZ NNP 37809 283 5 104039 104039 CD 37809 283 6 ; ; : 37809 283 7 8 8 CD 37809 283 8 km km NN 37809 283 9 . . . 37809 284 1 N N NNP 37809 284 2 Tamazunchale Tamazunchale NNP 37809 284 3 , , , 37809 284 4 UMMZ UMMZ NNP 37809 284 5 119490 119490 CD 37809 284 6 . . . 37809 285 1 _ _ NNP 37809 285 2 Veracruz Veracruz NNP 37809 285 3 _ _ NNP 37809 285 4 : : : 37809 285 5 Coatepec Coatepec NNP 37809 285 6 , , , 37809 285 7 1210 1210 CD 37809 285 8 m. m. NN 37809 285 9 , , , 37809 285 10 FMNH FMNH NNP 37809 285 11 704966 704966 CD 37809 285 12 - - SYM 37809 285 13 67 67 CD 37809 285 14 ; ; : 37809 285 15 11 11 CD 37809 285 16 km km NN 37809 285 17 . . . 37809 286 1 SE SE NNP 37809 286 2 Coatepec Coatepec NNP 37809 286 3 , , , 37809 286 4 850 850 CD 37809 286 5 m. m. NN 37809 286 6 , , , 37809 286 7 FMNH FMNH NNP 37809 286 8 70468 70468 CD 37809 286 9 - - SYM 37809 286 10 70 70 CD 37809 286 11 ; ; : 37809 286 12 below below IN 37809 286 13 Córdoba Córdoba NNP 37809 286 14 , , , 37809 286 15 FMNH FMNH NNP 37809 286 16 104588 104588 CD 37809 286 17 , , , 37809 286 18 UIMNH UIMNH NNP 37809 286 19 13321 13321 CD 37809 286 20 ; ; : 37809 286 21 Cuautlapam Cuautlapam NNP 37809 286 22 , , , 37809 286 23 1000 1000 CD 37809 286 24 m. m. NN 37809 286 25 , , , 37809 286 26 FMNH FMNH NNP 37809 286 27 106477 106477 CD 37809 286 28 - - SYM 37809 286 29 80 80 CD 37809 286 30 , , , 37809 286 31 KU KU NNP 37809 286 32 100364 100364 CD 37809 286 33 , , , 37809 286 34 UIMNH UIMNH NNP 37809 286 35 58200 58200 CD 37809 286 36 - - SYM 37809 286 37 03 03 CD 37809 286 38 , , , 37809 286 39 UMMZ UMMZ NNP 37809 286 40 105392 105392 CD 37809 286 41 ; ; : 37809 286 42 Fortín Fortín NNP 37809 286 43 de de IN 37809 286 44 las las NNP 37809 286 45 Flores Flores NNP 37809 286 46 , , , 37809 286 47 UIMNH UIMNH NNP 37809 286 48 13322 13322 CD 37809 286 49 , , , 37809 286 50 13339 13339 CD 37809 286 51 ; ; : 37809 286 52 1.6 1.6 CD 37809 286 53 km km NNS 37809 286 54 . . . 37809 287 1 N N NNP 37809 287 2 Fortín Fortín NNP 37809 287 3 de de NN 37809 287 4 las las NNP 37809 287 5 Flores Flores NNP 37809 287 6 , , , 37809 287 7 UIMNH UIMNH NNP 37809 287 8 42799 42799 CD 37809 287 9 - - SYM 37809 287 10 808 808 CD 37809 287 11 , , , 37809 287 12 UMMZ UMMZ NNP 37809 287 13 105389 105389 CD 37809 287 14 ; ; : 37809 287 15 3.2 3.2 CD 37809 287 16 km km NN 37809 287 17 . . . 37809 288 1 N N NNP 37809 288 2 Fortín Fortín NNP 37809 288 3 de de NN 37809 288 4 las las NNP 37809 288 5 Flores Flores NNP 37809 288 6 , , , 37809 288 7 UIMNH UIMNH NNP 37809 288 8 26633 26633 CD 37809 288 9 - - SYM 37809 288 10 35 35 CD 37809 288 11 ; ; : 37809 288 12 4.8 4.8 CD 37809 288 13 km km NNS 37809 288 14 . . . 37809 289 1 N N NNP 37809 289 2 Fortín Fortín NNP 37809 289 3 de de NN 37809 289 4 las las NNP 37809 289 5 Flores Flores NNP 37809 289 6 , , , 37809 289 7 UIMNH UIMNH NNP 37809 289 8 71967 71967 CD 37809 289 9 - - SYM 37809 289 10 68 68 CD 37809 289 11 ; ; : 37809 289 12 3.2 3.2 CD 37809 289 13 km km NN 37809 289 14 . . . 37809 290 1 W W NNP 37809 290 2 Fortín Fortín NNP 37809 290 3 de de NN 37809 290 4 las las NNP 37809 290 5 Flores Flores NNP 37809 290 6 ( ( -LRB- 37809 290 7 Barranca Barranca NNP 37809 290 8 Metlac Metlac NNP 37809 290 9 ) ) -RRB- 37809 290 10 , , , 37809 290 11 910 910 CD 37809 290 12 m. m. NN 37809 290 13 , , , 37809 290 14 UIMNH UIMNH NNP 37809 290 15 49294 49294 CD 37809 290 16 - - SYM 37809 290 17 95 95 CD 37809 290 18 , , , 37809 290 19 UMMZ UMMZ NNP 37809 290 20 115444 115444 CD 37809 290 21 - - SYM 37809 290 22 46 46 CD 37809 290 23 , , , 37809 290 24 118221 118221 CD 37809 290 25 , , , 37809 290 26 119893 119893 CD 37809 290 27 ( ( -LRB- 37809 290 28 2 2 CD 37809 290 29 ) ) -RRB- 37809 290 30 ; ; : 37809 290 31 Huatusco Huatusco NNP 37809 290 32 , , , 37809 290 33 KU KU NNP 37809 290 34 100363 100363 CD 37809 290 35 ; ; : 37809 290 36 Jalapa Jalapa NNP 37809 290 37 , , , 37809 290 38 1400 1400 CD 37809 290 39 m. m. NN 37809 290 40 , , , 37809 290 41 FMNH FMNH NNP 37809 290 42 70440 70440 CD 37809 290 43 , , , 37809 290 44 70443 70443 CD 37809 290 45 - - SYM 37809 290 46 51 51 CD 37809 290 47 , , , 37809 290 48 70454 70454 CD 37809 290 49 - - SYM 37809 290 50 65 65 CD 37809 290 51 ; ; : 37809 290 52 16 16 CD 37809 290 53 km km NN 37809 290 54 . . . 37809 291 1 NE NE NNP 37809 291 2 Jalapa Jalapa NNP 37809 291 3 , , , 37809 291 4 1300 1300 CD 37809 291 5 m. m. NN 37809 291 6 , , , 37809 291 7 FMNH FMNH NNP 37809 291 8 70452 70452 CD 37809 291 9 - - SYM 37809 291 10 53 53 CD 37809 291 11 ; ; : 37809 291 12 8 8 CD 37809 291 13 km km NN 37809 291 14 . . . 37809 292 1 E E NNP 37809 292 2 Jalapa Jalapa NNP 37809 292 3 , , , 37809 292 4 UIMNH UIMNH NNP 37809 292 5 13338 13338 CD 37809 292 6 ; ; : 37809 292 7 9.5 9.5 CD 37809 292 8 km km NN 37809 292 9 . . . 37809 293 1 S S NNP 37809 293 2 Jalapa Jalapa NNP 37809 293 3 , , , 37809 293 4 UMMZ UMMZ NNP 37809 293 5 122083 122083 CD 37809 293 6 ( ( -LRB- 37809 293 7 2 2 CD 37809 293 8 ) ) -RRB- 37809 293 9 ; ; : 37809 293 10 Mirador Mirador NNP 37809 293 11 , , , 37809 293 12 KU KU NNP 37809 293 13 23967 23967 CD 37809 293 14 ; ; : 37809 293 15 Paraja Paraja NNP 37809 293 16 Nuevo Nuevo NNP 37809 293 17 , , , 37809 293 18 El El NNP 37809 293 19 Suchil Suchil NNP 37809 293 20 , , , 37809 293 21 UMMZ UMMZ NNP 37809 293 22 85490 85490 CD 37809 293 23 ( ( -LRB- 37809 293 24 7 7 CD 37809 293 25 ) ) -RRB- 37809 293 26 , , , 37809 293 27 85491 85491 CD 37809 293 28 ( ( -LRB- 37809 293 29 2 2 CD 37809 293 30 ) ) -RRB- 37809 293 31 , , , 37809 293 32 90315 90315 CD 37809 293 33 ; ; : 37809 293 34 La La NNP 37809 293 35 Passa Passa NNP 37809 293 36 , , , 37809 293 37 UIMNH UIMNH NNP 37809 293 38 49293 49293 CD 37809 293 39 , , , 37809 293 40 49297 49297 CD 37809 293 41 ; ; : 37809 293 42 1 1 CD 37809 293 43 km km NN 37809 293 44 . . . 37809 294 1 E E NNP 37809 294 2 Plan Plan NNP 37809 294 3 del del NNP 37809 294 4 Río Río NNP 37809 294 5 , , , 37809 294 6 240 240 CD 37809 294 7 m. m. NN 37809 294 8 , , , 37809 294 9 UMMZ UMMZ NNP 37809 294 10 102067 102067 CD 37809 294 11 ( ( -LRB- 37809 294 12 2 2 CD 37809 294 13 ) ) -RRB- 37809 294 14 ; ; : 37809 294 15 Potrero Potrero NNP 37809 294 16 Viejo Viejo NNP 37809 294 17 , , , 37809 294 18 FMNH FMNH NNP 37809 294 19 104583 104583 CD 37809 294 20 , , , 37809 294 21 104586 104586 CD 37809 294 22 , , , 37809 294 23 105326 105326 CD 37809 294 24 - - SYM 37809 294 25 27 27 CD 37809 294 26 , , , 37809 294 27 KU KU NNP 37809 294 28 26789 26789 CD 37809 294 29 , , , 37809 294 30 100357 100357 CD 37809 294 31 - - SYM 37809 294 32 62 62 CD 37809 294 33 , , , 37809 294 34 UIMNH UIMNH NNP 37809 294 35 13323 13323 CD 37809 294 36 , , , 37809 294 37 13340 13340 CD 37809 294 38 - - SYM 37809 294 39 43 43 CD 37809 294 40 ; ; : 37809 294 41 USNM USNM NNP 37809 294 42 32402 32402 CD 37809 294 43 ( ( -LRB- 37809 294 44 lectotype lectotype NNP 37809 294 45 ) ) -RRB- 37809 294 46 , , , 37809 294 47 32403 32403 CD 37809 294 48 - - SYM 37809 294 49 04 04 CD 37809 294 50 , , , 37809 294 51 32406 32406 CD 37809 294 52 - - SYM 37809 294 53 09 09 CD 37809 294 54 ; ; : 37809 294 55 9.6 9.6 CD 37809 294 56 km km NN 37809 294 57 . . . 37809 295 1 S S NNP 37809 295 2 Santa Santa NNP 37809 295 3 Rosa Rosa NNP 37809 295 4 , , , 37809 295 5 TCWC TCWC NNP 37809 295 6 12785 12785 CD 37809 295 7 ; ; : 37809 295 8 24 24 CD 37809 295 9 km km NNS 37809 295 10 . . . 37809 296 1 NE NE NNP 37809 296 2 Tezuitlán Tezuitlán NNP 37809 296 3 ( ( -LRB- 37809 296 4 Puebla Puebla NNP 37809 296 5 ) ) -RRB- 37809 296 6 , , , 37809 296 7 UMMZ UMMZ NNP 37809 296 8 105388 105388 CD 37809 296 9 ; ; : 37809 296 10 Teocelo Teocelo NNP 37809 296 11 , , , 37809 296 12 FMNH FMNH NNP 37809 296 13 70437 70437 CD 37809 296 14 - - SYM 37809 296 15 38 38 CD 37809 296 16 , , , 37809 296 17 KU KU NNP 37809 296 18 26080 26080 CD 37809 296 19 , , , 37809 296 20 26790 26790 CD 37809 296 21 ; ; : 37809 296 22 3.2 3.2 CD 37809 296 23 km km NN 37809 296 24 . . . 37809 297 1 N N NNP 37809 297 2 Teocelo Teocelo NNP 37809 297 3 , , , 37809 297 4 FMNH FMNH NNP 37809 297 5 70439 70439 CD 37809 297 6 , , , 37809 297 7 70441 70441 CD 37809 297 8 - - SYM 37809 297 9 42 42 CD 37809 297 10 ; ; : 37809 297 11 9.6 9.6 CD 37809 297 12 km km NN 37809 297 13 . . . 37809 298 1 NW NW NNP 37809 298 2 Tihuatlán Tihuatlán NNP 37809 298 3 , , , 37809 298 4 UIMNH UIMNH NNP 37809 298 5 3684 3684 CD 37809 298 6 - - SYM 37809 298 7 85 85 CD 37809 298 8 ; ; : 37809 298 9 15 15 CD 37809 298 10 km km NN 37809 298 11 . . . 37809 299 1 ENE ENE NNP 37809 299 2 Tlacotepec Tlacotepec NNP 37809 299 3 , , , 37809 299 4 KU KU NNP 37809 299 5 23966 23966 CD 37809 299 6 ; ; : 37809 299 7 26 26 CD 37809 299 8 km km NN 37809 299 9 . . . 37809 300 1 NW NW NNP 37809 300 2 Tuxpan Tuxpan NNP 37809 300 3 , , , 37809 300 4 UMMZ UMMZ NNP 37809 300 5 126419 126419 CD 37809 300 6 . . . 37809 301 1 = = NFP 37809 301 2 Syrrhophus Syrrhophus NNP 37809 301 3 leprus= leprus= VBZ 37809 301 4 Cope Cope NNP 37809 301 5 _ _ NNP 37809 301 6 Syrrhophus Syrrhophus NNP 37809 301 7 leprus leprus NN 37809 301 8 _ _ NNP 37809 301 9 Cope Cope NNP 37809 301 10 , , , 37809 301 11 1879:268 1879:268 CD 37809 301 12 - - SYM 37809 301 13 69 69 CD 37809 301 14 [ [ -LRB- 37809 301 15 Holotype.--USNM holotype.--usnm NN 37809 301 16 10040 10040 CD 37809 301 17 , , , 37809 301 18 from from IN 37809 301 19 Santa Santa NNP 37809 301 20 Efigena Efigena NNP 37809 301 21 , , , 37809 301 22 Oaxaca Oaxaca NNP 37809 301 23 , , , 37809 301 24 México México NNP 37809 301 25 , , , 37809 301 26 Francis Francis NNP 37809 301 27 Sumichrast Sumichrast NNP 37809 301 28 collector collector NN 37809 301 29 ] ] -RRB- 37809 301 30 . . . 37809 302 1 Kellogg Kellogg NNP 37809 302 2 , , , 37809 302 3 1932:124 1932:124 CD 37809 302 4 - - SYM 37809 302 5 5 5 CD 37809 302 6 , , , 37809 302 7 128 128 CD 37809 302 8 . . . 37809 303 1 Taylor Taylor NNP 37809 303 2 and and CC 37809 303 3 Smith Smith NNP 37809 303 4 , , , 37809 303 5 1945:582 1945:582 CD 37809 303 6 . . . 37809 304 1 Smith Smith NNP 37809 304 2 and and CC 37809 304 3 Taylor Taylor NNP 37809 304 4 , , , 37809 304 5 1948:50 1948:50 CD 37809 304 6 - - SYM 37809 304 7 51 51 CD 37809 304 8 . . . 37809 305 1 Duellman Duellman NNP 37809 305 2 , , , 37809 305 3 1958:8 1958:8 CD 37809 305 4 , , , 37809 305 5 pl pl NNP 37809 305 6 . . . 37809 306 1 1 1 LS 37809 306 2 , , , 37809 306 3 Fig Fig NNP 37809 306 4 . . . 37809 307 1 2 2 LS 37809 307 2 ; ; : 37809 307 3 1960:56 1960:56 CD 37809 307 4 - - SYM 37809 307 5 57 57 CD 37809 307 6 . . . 37809 308 1 Gorham Gorham NNP 37809 308 2 , , , 37809 308 3 1966:165 1966:165 CD 37809 308 4 . . . 37809 309 1 _ _ NNP 37809 309 2 Syrrhaphus Syrrhaphus NNP 37809 309 3 leprus leprus NNP 37809 309 4 _ _ NNP 37809 309 5 : : : 37809 309 6 Günther Günther NNP 37809 309 7 , , , 37809 309 8 1900:217 1900:217 CD 37809 309 9 . . . 37809 310 1 _ _ NNP 37809 310 2 Syrrhophus Syrrhophus NNP 37809 310 3 leprus leprus NNP 37809 310 4 leprus leprus NNP 37809 310 5 _ _ NNP 37809 310 6 : : : 37809 310 7 Neill Neill NNP 37809 310 8 , , , 37809 310 9 1965:85 1965:85 NNP 37809 310 10 - - HYPH 37809 310 11 86 86 CD 37809 310 12 . . . 37809 311 1 _ _ NNP 37809 311 2 Syrrhophus Syrrhophus NNP 37809 311 3 leprus leprus NNP 37809 311 4 cholorum cholorum NN 37809 311 5 _ _ NNP 37809 311 6 Neill Neill NNP 37809 311 7 , , , 37809 311 8 1965:85 1965:85 NNP 37809 311 9 - - HYPH 37809 311 10 86 86 CD 37809 311 11 [ [ -LRB- 37809 311 12 Holotype.--Wilfred Holotype.--Wilfred NNP 37809 311 13 T. T. NNP 37809 311 14 Neill Neill NNP 37809 311 15 collection collection NN 37809 311 16 1525 1525 CD 37809 311 17 , , , 37809 311 18 from from IN 37809 311 19 3.9 3.9 CD 37809 311 20 mi mi NNP 37809 311 21 . . . 37809 312 1 N N NNP 37809 312 2 San San NNP 37809 312 3 Antonio Antonio NNP 37809 312 4 , , , 37809 312 5 Toledo Toledo NNP 37809 312 6 District District NNP 37809 312 7 , , , 37809 312 8 British British NNP 37809 312 9 Honduras Honduras NNP 37809 312 10 , , , 37809 312 11 collected collect VBN 37809 312 12 October October NNP 37809 312 13 28 28 CD 37809 312 14 , , , 37809 312 15 1959 1959 CD 37809 312 16 , , , 37809 312 17 by by IN 37809 312 18 R. R. NNP 37809 312 19 A. A. NNP 37809 312 20 Allen Allen NNP 37809 312 21 , , , 37809 312 22 T. T. NNP 37809 312 23 C. C. NNP 37809 312 24 Allen Allen NNP 37809 312 25 , , , 37809 312 26 and and CC 37809 312 27 W. W. NNP 37809 312 28 T. T. NNP 37809 312 29 Neill Neill NNP 37809 312 30 ] ] -RRB- 37809 312 31 . . . 37809 313 1 _ _ NNP 37809 313 2 Diagnosis._--Medium Diagnosis._--Medium NNP 37809 313 3 - - HYPH 37809 313 4 sized sized JJ 37809 313 5 frogs frog NNS 37809 313 6 , , , 37809 313 7 males males FW 37809 313 8 20.5 20.5 CD 37809 313 9 - - HYPH 37809 313 10 26.5 26.5 CD 37809 313 11 mm mm NNP 37809 313 12 . . . 37809 314 1 in in IN 37809 314 2 snout snout NN 37809 314 3 - - HYPH 37809 314 4 vent vent NN 37809 314 5 , , , 37809 314 6 females female VBZ 37809 314 7 22.0 22.0 CD 37809 314 8 - - SYM 37809 314 9 29.3 29.3 CD 37809 314 10 mm mm NN 37809 314 11 . . . 37809 315 1 in in IN 37809 315 2 snout snout NN 37809 315 3 - - HYPH 37809 315 4 vent vent NN 37809 315 5 length length NN 37809 315 6 ; ; : 37809 315 7 vocal vocal NN 37809 315 8 slits slit NNS 37809 315 9 present present JJ 37809 315 10 in in IN 37809 315 11 males male NNS 37809 315 12 ; ; : 37809 315 13 tips tip NNS 37809 315 14 of of IN 37809 315 15 fingers finger NNS 37809 315 16 dilated dilate VBN 37809 315 17 slightly slightly RB 37809 315 18 ; ; : 37809 315 19 first first JJ 37809 315 20 finger finger NN 37809 315 21 longer long RBR 37809 315 22 than than IN 37809 315 23 second second JJ 37809 315 24 ; ; : 37809 315 25 inner inner JJ 37809 315 26 metatarsal metatarsal NN 37809 315 27 tubercle tubercle NN 37809 315 28 twice twice RB 37809 315 29 size size NN 37809 315 30 of of IN 37809 315 31 small small JJ 37809 315 32 , , , 37809 315 33 conical conical JJ 37809 315 34 outer outer JJ 37809 315 35 metatarsal metatarsal NN 37809 315 36 tubercle tubercle NN 37809 315 37 ; ; : 37809 315 38 skin skin NN 37809 315 39 of of IN 37809 315 40 dorsum dorsum NN 37809 315 41 pustular pustular NN 37809 315 42 , , , 37809 315 43 that that DT 37809 315 44 of of IN 37809 315 45 venter venter NN 37809 315 46 smooth smooth JJ 37809 315 47 ; ; : 37809 315 48 snout snout NN 37809 315 49 subacuminate subacuminate NN 37809 315 50 ; ; : 37809 315 51 diameter diameter NN 37809 315 52 of of IN 37809 315 53 tympanum tympanum CD 37809 315 54 47.5 47.5 CD 37809 315 55 - - SYM 37809 315 56 62.5 62.5 CD 37809 315 57 per per IN 37809 315 58 cent cent NN 37809 315 59 of of IN 37809 315 60 eye eye NN 37809 315 61 in in IN 37809 315 62 males male NNS 37809 315 63 , , , 37809 315 64 38.6 38.6 CD 37809 315 65 - - SYM 37809 315 66 57.9 57.9 CD 37809 315 67 per per IN 37809 315 68 cent cent NN 37809 315 69 in in IN 37809 315 70 females female NNS 37809 315 71 ; ; : 37809 315 72 dorsum dorsum NN 37809 315 73 yellowish yellowish JJ 37809 315 74 - - HYPH 37809 315 75 green green JJ 37809 315 76 with with IN 37809 315 77 chocolate chocolate NN 37809 315 78 brown brown JJ 37809 315 79 blotches blotch NNS 37809 315 80 or or CC 37809 315 81 spots spot NNS 37809 315 82 forming form VBG 37809 315 83 reticulations reticulation NNS 37809 315 84 in in IN 37809 315 85 most most JJS 37809 315 86 specimens specimen NNS 37809 315 87 ; ; : 37809 315 88 venter venter NN 37809 315 89 white white JJ 37809 315 90 to to TO 37809 315 91 gray gray VB 37809 315 92 ; ; : 37809 315 93 flanks flanks NNP 37809 315 94 brown brown NNP 37809 315 95 , , , 37809 315 96 spotted spot VBD 37809 315 97 with with IN 37809 315 98 white white JJ 37809 315 99 or or CC 37809 315 100 not not RB 37809 315 101 ; ; : 37809 315 102 limbs limb NNS 37809 315 103 banded band VBD 37809 315 104 ; ; : 37809 315 105 interorbital interorbital NNP 37809 315 106 bar bar NN 37809 315 107 obscured obscure VBN 37809 315 108 by by IN 37809 315 109 dorsal dorsal NN 37809 315 110 pattern pattern NN 37809 315 111 . . . 37809 316 1 [ [ -LRB- 37809 316 2 Illustration illustration NN 37809 316 3 : : : 37809 316 4 FIG FIG NNP 37809 316 5 . . . 37809 317 1 6 6 CD 37809 317 2 : : : 37809 317 3 Dorsal dorsal NN 37809 317 4 views view NNS 37809 317 5 of of IN 37809 317 6 _ _ NNP 37809 317 7 Syrrhophus Syrrhophus NNP 37809 317 8 leprus leprus NN 37809 317 9 _ _ NNP 37809 317 10 showing show VBG 37809 317 11 variation variation NN 37809 317 12 in in IN 37809 317 13 dorsal dorsal NN 37809 317 14 pattern pattern NN 37809 317 15 ( ( -LRB- 37809 317 16 left leave VBN 37809 317 17 , , , 37809 317 18 UMMZ UMMZ NNP 37809 317 19 121244 121244 CD 37809 317 20 , , , 37809 317 21 ×2 ×2 NNS 37809 317 22 ; ; : 37809 317 23 right right UH 37809 317 24 , , , 37809 317 25 KU KU NNP 37809 317 26 26106 26106 CD 37809 317 27 , , , 37809 317 28 ×1.7 ×1.7 NNP 37809 317 29 ) ) -RRB- 37809 317 30 . . . 37809 318 1 Side side NN 37809 318 2 of of IN 37809 318 3 head head NN 37809 318 4 ( ( -LRB- 37809 318 5 UIMNH UIMNH NNP 37809 318 6 42726 42726 CD 37809 318 7 , , , 37809 318 8 ×7 ×7 ADD 37809 318 9 ) ) -RRB- 37809 318 10 . . . 37809 318 11 ] ] -RRB- 37809 319 1 [ [ -LRB- 37809 319 2 Illustration illustration NN 37809 319 3 : : : 37809 319 4 FIG FIG NNP 37809 319 5 . . . 37809 320 1 7 7 LS 37809 320 2 : : : 37809 320 3 Distribution distribution NN 37809 320 4 of of IN 37809 320 5 three three CD 37809 320 6 species specie NNS 37809 320 7 of of IN 37809 320 8 eastern eastern JJ 37809 320 9 complex complex NN 37809 320 10 _ _ NNP 37809 320 11 Syrrhophus Syrrhophus NNP 37809 320 12 _ _ NNP 37809 320 13 : : : 37809 320 14 _ _ NNP 37809 320 15 leprus leprus NNP 37809 320 16 _ _ NNP 37809 320 17 ( ( -LRB- 37809 320 18 circles circles NNP 37809 320 19 ) ) -RRB- 37809 320 20 , , , 37809 320 21 _ _ NNP 37809 320 22 rubrimaculatus rubrimaculatus NN 37809 320 23 _ _ NNP 37809 320 24 ( ( -LRB- 37809 320 25 triangles triangle NNS 37809 320 26 ) ) -RRB- 37809 320 27 , , , 37809 320 28 and and CC 37809 320 29 _ _ NNP 37809 320 30 verrucipes verrucipe NNS 37809 320 31 _ _ NNP 37809 320 32 ( ( -LRB- 37809 320 33 squares square NNS 37809 320 34 ) ) -RRB- 37809 320 35 . . . 37809 320 36 ] ] -RRB- 37809 321 1 _ _ NNP 37809 321 2 Remarks._--My remarks._--my NN 37809 321 3 distribution distribution NN 37809 321 4 map map NN 37809 321 5 ( ( -LRB- 37809 321 6 Fig fig NN 37809 321 7 . . . 37809 322 1 7 7 LS 37809 322 2 ) ) -RRB- 37809 322 3 differs differ VBZ 37809 322 4 somewhat somewhat RB 37809 322 5 from from IN 37809 322 6 that that DT 37809 322 7 of of IN 37809 322 8 Duellman Duellman NNP 37809 322 9 ( ( -LRB- 37809 322 10 1958 1958 CD 37809 322 11 ) ) -RRB- 37809 322 12 , , , 37809 322 13 who who WP 37809 322 14 was be VBD 37809 322 15 unaware unaware JJ 37809 322 16 of of IN 37809 322 17 specimens specimen NNS 37809 322 18 reported report VBN 37809 322 19 by by IN 37809 322 20 Taylor Taylor NNP 37809 322 21 and and CC 37809 322 22 Smith Smith NNP 37809 322 23 ( ( -LRB- 37809 322 24 1945 1945 CD 37809 322 25 ) ) -RRB- 37809 322 26 from from IN 37809 322 27 central central JJ 37809 322 28 Veracruz Veracruz NNP 37809 322 29 , , , 37809 322 30 México México NNP 37809 322 31 . . . 37809 323 1 Duellman Duellman NNP 37809 323 2 ( ( -LRB- 37809 323 3 1958 1958 CD 37809 323 4 , , , 37809 323 5 1960 1960 CD 37809 323 6 ) ) -RRB- 37809 323 7 regarded regard VBD 37809 323 8 _ _ NNP 37809 323 9 S. S. NNP 37809 324 1 leprus leprus NNP 37809 324 2 _ _ NNP 37809 324 3 as as IN 37809 324 4 having have VBG 37809 324 5 a a DT 37809 324 6 gray gray JJ 37809 324 7 venter venter NN 37809 324 8 . . . 37809 325 1 Neill Neill NNP 37809 325 2 ( ( -LRB- 37809 325 3 1965 1965 CD 37809 325 4 ) ) -RRB- 37809 325 5 characterized characterize VBD 37809 325 6 his -PRON- PRP$ 37809 325 7 new new JJ 37809 325 8 subspecies subspecie NNS 37809 325 9 on on IN 37809 325 10 the the DT 37809 325 11 basis basis NN 37809 325 12 of of IN 37809 325 13 white white JJ 37809 325 14 venter venter NN 37809 325 15 and and CC 37809 325 16 spots spot NNS 37809 325 17 on on IN 37809 325 18 the the DT 37809 325 19 dorsum dorsum NN 37809 325 20 . . . 37809 326 1 Some some DT 37809 326 2 specimens specimen NNS 37809 326 3 from from IN 37809 326 4 throughout throughout IN 37809 326 5 the the DT 37809 326 6 range range NN 37809 326 7 have have VBP 37809 326 8 only only JJ 37809 326 9 small small JJ 37809 326 10 round round JJ 37809 326 11 spots spot NNS 37809 326 12 , , , 37809 326 13 instead instead RB 37809 326 14 of of IN 37809 326 15 vermiculations vermiculation NNS 37809 326 16 ( ( -LRB- 37809 326 17 Fig fig NN 37809 326 18 . . . 37809 327 1 6 6 LS 37809 327 2 ) ) -RRB- 37809 327 3 . . . 37809 328 1 The the DT 37809 328 2 gray gray JJ 37809 328 3 ventral ventral JJ 37809 328 4 coloration coloration NN 37809 328 5 is be VBZ 37809 328 6 largely largely RB 37809 328 7 restricted restrict VBN 37809 328 8 to to IN 37809 328 9 the the DT 37809 328 10 population population NN 37809 328 11 in in IN 37809 328 12 Los Los NNP 37809 328 13 Tuxtlas Tuxtlas NNP 37809 328 14 , , , 37809 328 15 Veracruz Veracruz NNP 37809 328 16 , , , 37809 328 17 but but CC 37809 328 18 only only RB 37809 328 19 about about RB 37809 328 20 80 80 CD 37809 328 21 per per NN 37809 328 22 cent cent NN 37809 328 23 of of IN 37809 328 24 the the DT 37809 328 25 specimens specimen NNS 37809 328 26 from from IN 37809 328 27 the the DT 37809 328 28 Los Los NNP 37809 328 29 Tuxtlas Tuxtlas NNP 37809 328 30 have have VBP 37809 328 31 gray gray JJ 37809 328 32 venters venter NNS 37809 328 33 , , , 37809 328 34 whereas whereas IN 37809 328 35 specimens specimen NNS 37809 328 36 from from IN 37809 328 37 Guatemala Guatemala NNP 37809 328 38 , , , 37809 328 39 Oaxaca Oaxaca NNP 37809 328 40 , , , 37809 328 41 Tabasco Tabasco NNP 37809 328 42 , , , 37809 328 43 and and CC 37809 328 44 central central JJ 37809 328 45 Veracruz Veracruz NNP 37809 328 46 , , , 37809 328 47 México México NNP 37809 328 48 , , , 37809 328 49 have have VBP 37809 328 50 white white JJ 37809 328 51 venters venter NNS 37809 328 52 ( ( -LRB- 37809 328 53 rarely rarely RB 37809 328 54 gray gray JJ 37809 328 55 ) ) -RRB- 37809 328 56 . . . 37809 329 1 Since since IN 37809 329 2 the the DT 37809 329 3 specimens specimen NNS 37809 329 4 from from IN 37809 329 5 British British NNP 37809 329 6 Honduras Honduras NNPS 37809 329 7 are be VBP 37809 329 8 not not RB 37809 329 9 distinct distinct JJ 37809 329 10 from from IN 37809 329 11 specimens specimen NNS 37809 329 12 throughout throughout IN 37809 329 13 most most JJS 37809 329 14 of of IN 37809 329 15 the the DT 37809 329 16 range range NN 37809 329 17 , , , 37809 329 18 there there EX 37809 329 19 is be VBZ 37809 329 20 no no DT 37809 329 21 reason reason NN 37809 329 22 to to TO 37809 329 23 recognize recognize VB 37809 329 24 them -PRON- PRP 37809 329 25 as as IN 37809 329 26 a a DT 37809 329 27 subspecies subspecie NNS 37809 329 28 . . . 37809 330 1 _ _ NNP 37809 330 2 Etymology._--Greek Etymology._--Greek NNP 37809 330 3 , , , 37809 330 4 _ _ NNP 37809 330 5 lepra lepra NNP 37809 330 6 _ _ NNP 37809 330 7 , , , 37809 330 8 leprosy leprosy NN 37809 330 9 , , , 37809 330 10 in in IN 37809 330 11 reference reference NN 37809 330 12 to to IN 37809 330 13 the the DT 37809 330 14 mottled mottle VBN 37809 330 15 color color NN 37809 330 16 pattern pattern NN 37809 330 17 . . . 37809 331 1 _ _ NNP 37809 331 2 Distribution._--Discontinuous distribution._--discontinuous CD 37809 331 3 ; ; : 37809 331 4 central central JJ 37809 331 5 Veracruz Veracruz NNP 37809 331 6 to to IN 37809 331 7 British British NNP 37809 331 8 Honduras Honduras NNP 37809 331 9 to to IN 37809 331 10 low low JJ 37809 331 11 elevations elevation NNS 37809 331 12 in in IN 37809 331 13 the the DT 37809 331 14 foothills foothill NNS 37809 331 15 of of IN 37809 331 16 the the DT 37809 331 17 Sierra Sierra NNP 37809 331 18 Madre Madre NNP 37809 331 19 Oriental Oriental NNP 37809 331 20 , , , 37809 331 21 Los Los NNP 37809 331 22 Tuxtlas Tuxtlas NNP 37809 331 23 , , , 37809 331 24 Sierra Sierra NNP 37809 331 25 Madre Madre NNP 37809 331 26 de de NNP 37809 331 27 Chiapas Chiapas NNP 37809 331 28 ( ( -LRB- 37809 331 29 Isthmus Isthmus NNP 37809 331 30 of of IN 37809 331 31 Tehuantepec Tehuantepec NNP 37809 331 32 ( ( -LRB- 37809 331 33 Fig Fig NNP 37809 331 34 . . . 37809 332 1 7 7 LS 37809 332 2 ) ) -RRB- 37809 332 3 ) ) -RRB- 37809 332 4 . . . 37809 333 1 _ _ NNP 37809 333 2 Specimens Specimens NNP 37809 333 3 examined._--(84 examined._--(84 NNP 37809 333 4 ) ) -RRB- 37809 333 5 . . . 37809 334 1 GUATEMALA GUATEMALA NNP 37809 334 2 , , , 37809 334 3 _ _ NNP 37809 334 4 Alta Alta NNP 37809 334 5 Verapaz Verapaz NNP 37809 334 6 _ _ NNP 37809 334 7 : : : 37809 334 8 Chinajá Chinajá NNP 37809 334 9 , , , 37809 334 10 KU KU NNP 37809 334 11 55961 55961 CD 37809 334 12 - - SYM 37809 334 13 62 62 CD 37809 334 14 . . . 37809 335 1 _ _ NNP 37809 335 2 El El NNP 37809 335 3 Petén Petén NNP 37809 335 4 _ _ NNP 37809 335 5 : : : 37809 335 6 15 15 CD 37809 335 7 km km NN 37809 335 8 . . . 37809 336 1 NW NW NNP 37809 336 2 Chinajá Chinajá NNP 37809 336 3 , , , 37809 336 4 KU KU NNP 37809 336 5 55963 55963 CD 37809 336 6 ; ; : 37809 336 7 Piedras Piedras NNP 37809 336 8 Negras Negras NNP 37809 336 9 , , , 37809 336 10 USNM USNM NNP 37809 336 11 114085 114085 CD 37809 336 12 - - SYM 37809 336 13 92 92 CD 37809 336 14 ; ; : 37809 336 15 Tikal Tikal NNP 37809 336 16 , , , 37809 336 17 UMMZ UMMZ NNP 37809 336 18 117035 117035 CD 37809 336 19 ; ; : 37809 336 20 Uaxactún Uaxactún NNP 37809 336 21 , , , 37809 336 22 AMNH AMNH NNP 37809 336 23 55121 55121 CD 37809 336 24 - - SYM 37809 336 25 22 22 CD 37809 336 26 . . . 37809 337 1 MÉXICO MÉXICO NNP 37809 337 2 , , , 37809 337 3 _ _ NNP 37809 337 4 Oaxaca Oaxaca NNP 37809 337 5 _ _ NNP 37809 337 6 : : : 37809 337 7 Cerro Cerro NNP 37809 337 8 San San NNP 37809 337 9 Pedro Pedro NNP 37809 337 10 del del NNP 37809 337 11 Isthmo Isthmo NNP 37809 337 12 , , , 37809 337 13 UIMNH UIMNH NNP 37809 337 14 35510 35510 CD 37809 337 15 ; ; : 37809 337 16 Finca Finca NNP 37809 337 17 La La NNP 37809 337 18 Gloria Gloria NNP 37809 337 19 , , , 37809 337 20 USNM USNM NNP 37809 337 21 114093 114093 CD 37809 337 22 ; ; : 37809 337 23 30.5 30.5 CD 37809 337 24 km km NNS 37809 337 25 . . . 37809 338 1 N N NNP 37809 338 2 Matías Matías NNP 37809 338 3 Romero Romero NNP 37809 338 4 , , , 37809 338 5 UIMNH UIMNH NNP 37809 338 6 39459 39459 CD 37809 338 7 , , , 37809 338 8 71969 71969 CD 37809 338 9 ; ; : 37809 338 10 Santa Santa NNP 37809 338 11 Efigenia Efigenia NNP 37809 338 12 , , , 37809 338 13 USNM USNM NNP 37809 338 14 10040 10040 CD 37809 338 15 ( ( -LRB- 37809 338 16 holotype holotype NN 37809 338 17 ) ) -RRB- 37809 338 18 . . . 37809 339 1 _ _ NNP 37809 339 2 Tabasco Tabasco NNP 37809 339 3 _ _ NNP 37809 339 4 : : : 37809 339 5 Teapa Teapa NNP 37809 339 6 , , , 37809 339 7 UMMZ UMMZ NNP 37809 339 8 113799 113799 CD 37809 339 9 - - SYM 37809 339 10 800 800 CD 37809 339 11 ; ; : 37809 339 12 13.5 13.5 CD 37809 339 13 km km NNS 37809 339 14 . . . 37809 340 1 W W NNP 37809 340 2 Teapa Teapa NNP 37809 340 3 , , , 37809 340 4 UMMZ UMMZ NNP 37809 340 5 120253 120253 CD 37809 340 6 . . . 37809 341 1 _ _ NNP 37809 341 2 Veracruz Veracruz NNP 37809 341 3 _ _ NNP 37809 341 4 : : : 37809 341 5 27.5 27.5 CD 37809 341 6 km km NNS 37809 341 7 . . . 37809 342 1 N N NNP 37809 342 2 Acayucan Acayucan NNP 37809 342 3 , , , 37809 342 4 UIMNH UIMNH NNP 37809 342 5 42726 42726 CD 37809 342 6 ; ; : 37809 342 7 Atoyac Atoyac NNP 37809 342 8 , , , 37809 342 9 UIMNH UIMNH NNP 37809 342 10 13331 13331 CD 37809 342 11 , , , 37809 342 12 49296 49296 CD 37809 342 13 ; ; : 37809 342 14 3.2 3.2 CD 37809 342 15 km km NN 37809 342 16 . . . 37809 343 1 N N NNP 37809 343 2 Catemaco Catemaco NNP 37809 343 3 , , , 37809 343 4 UIMNH UIMNH NNP 37809 343 5 71976 71976 CD 37809 343 6 - - SYM 37809 343 7 77 77 CD 37809 343 8 ; ; : 37809 343 9 Coyame coyame NN 37809 343 10 , , , 37809 343 11 UIMNH UIMNH NNP 37809 343 12 38995 38995 CD 37809 343 13 , , , 37809 343 14 38998 38998 CD 37809 343 15 , , , 37809 343 16 40342 40342 CD 37809 343 17 ; ; : 37809 343 18 Dos Dos NNP 37809 343 19 Amates Amates NNP 37809 343 20 , , , 37809 343 21 TCWC tcwc NN 37809 343 22 21211 21211 CD 37809 343 23 ; ; : 37809 343 24 Fortín Fortín NNP 37809 343 25 de de IN 37809 343 26 Las Las NNP 37809 343 27 Flores Flores NNP 37809 343 28 , , , 37809 343 29 FMNH FMNH NNP 37809 343 30 113751 113751 CD 37809 343 31 , , , 37809 343 32 113753 113753 CD 37809 343 33 ; ; : 37809 343 34 Paraja Paraja NNP 37809 343 35 Nuevo Nuevo NNP 37809 343 36 , , , 37809 343 37 El El NNP 37809 343 38 Suchil Suchil NNP 37809 343 39 , , , 37809 343 40 UMMZ UMMZ NNP 37809 343 41 90315 90315 CD 37809 343 42 ; ; : 37809 343 43 Potrero Potrero NNP 37809 343 44 Viejo Viejo NNP 37809 343 45 , , , 37809 343 46 FMNH FMNH NNP 37809 343 47 113743 113743 CD 37809 343 48 - - SYM 37809 343 49 50 50 CD 37809 343 50 , , , 37809 343 51 126114 126114 CD 37809 343 52 - - SYM 37809 343 53 18 18 CD 37809 343 54 , , , 37809 343 55 KU KU NNP 37809 343 56 26104 26104 CD 37809 343 57 - - SYM 37809 343 58 06 06 CD 37809 343 59 , , , 37809 343 60 UIMNH UIMNH NNP 37809 343 61 13332 13332 CD 37809 343 62 - - SYM 37809 343 63 37 37 CD 37809 343 64 , , , 37809 343 65 UMMZ UMMZ NNP 37809 343 66 88837 88837 CD 37809 343 67 ; ; : 37809 343 68 San San NNP 37809 343 69 Andrés Andrés NNP 37809 343 70 Tuxtla Tuxtla NNP 37809 343 71 , , , 37809 343 72 UIMNH UIMNH NNP 37809 343 73 27123 27123 CD 37809 343 74 - - SYM 37809 343 75 31 31 CD 37809 343 76 , , , 37809 343 77 28611 28611 CD 37809 343 78 , , , 37809 343 79 71975 71975 CD 37809 343 80 , , , 37809 343 81 UMMZ UMMZ NNP 37809 343 82 115450 115450 CD 37809 343 83 ( ( -LRB- 37809 343 84 5 5 CD 37809 343 85 ) ) -RRB- 37809 343 86 ; ; : 37809 343 87 San San NNP 37809 343 88 Lorenzo Lorenzo NNP 37809 343 89 , , , 37809 343 90 USNM USNM NNP 37809 343 91 123530 123530 CD 37809 343 92 ; ; : 37809 343 93 4.5 4.5 CD 37809 343 94 km km NNS 37809 343 95 . . . 37809 344 1 NW NW NNP 37809 344 2 Santiago Santiago NNP 37809 344 3 Tuxtla Tuxtla NNP 37809 344 4 , , , 37809 344 5 JDL JDL NNP 37809 344 6 992 992 CD 37809 344 7 ( ( -LRB- 37809 344 8 skeleton skeleton NN 37809 344 9 ) ) -RRB- 37809 344 10 , , , 37809 344 11 UIMNH UIMNH NNP 37809 344 12 27122 27122 CD 37809 344 13 ; ; : 37809 344 14 32 32 CD 37809 344 15 km km NNS 37809 344 16 . . . 37809 345 1 S S NNP 37809 345 2 Sayula Sayula NNP 37809 345 3 , , , 37809 345 4 EAL EAL NNP 37809 345 5 1696 1696 CD 37809 345 6 ; ; : 37809 345 7 Tepalapan Tepalapan NNP 37809 345 8 , , , 37809 345 9 1.6 1.6 CD 37809 345 10 km km NN 37809 345 11 . . . 37809 346 1 S S NNP 37809 346 2 Catemaco Catemaco NNP 37809 346 3 , , , 37809 346 4 UMMZ UMMZ NNP 37809 346 5 118222 118222 CD 37809 346 6 ( ( -LRB- 37809 346 7 2 2 CD 37809 346 8 ) ) -RRB- 37809 346 9 ; ; : 37809 346 10 Volcán Volcán NNP 37809 346 11 San San NNP 37809 346 12 Martín Martín NNP 37809 346 13 , , , 37809 346 14 south south JJ 37809 346 15 slope slope NN 37809 346 16 , , , 37809 346 17 UMMZ UMMZ NNP 37809 346 18 118223 118223 CD 37809 346 19 ; ; : 37809 346 20 Volcán Volcán NNP 37809 346 21 San San NNP 37809 346 22 Martín Martín NNP 37809 346 23 , , , 37809 346 24 Rancho Rancho NNP 37809 346 25 El El NNP 37809 346 26 Tular Tular NNP 37809 346 27 , , , 37809 346 28 UIMNH UIMNH NNP 37809 346 29 35399 35399 CD 37809 346 30 - - SYM 37809 346 31 400 400 CD 37809 346 32 , , , 37809 346 33 40340 40340 CD 37809 346 34 - - SYM 37809 346 35 41 41 CD 37809 346 36 . . . 37809 347 1 = = NFP 37809 347 2 Syrrhophus Syrrhophus NNP 37809 347 3 rubrimaculatus= rubrimaculatus= VBZ 37809 347 4 Taylor Taylor NNP 37809 347 5 and and CC 37809 347 6 Smith Smith NNP 37809 347 7 _ _ NNP 37809 347 8 Syrrhophus Syrrhophus NNP 37809 347 9 rubrimaculatus rubrimaculatus NN 37809 347 10 _ _ NNP 37809 347 11 Taylor Taylor NNP 37809 347 12 and and CC 37809 347 13 Smith Smith NNP 37809 347 14 , , , 37809 347 15 1945:583 1945:583 CD 37809 347 16 - - SYM 37809 347 17 85 85 CD 37809 347 18 [ [ -LRB- 37809 347 19 Holotype.--USNM Holotype.--USNM NNP 37809 347 20 114070 114070 CD 37809 347 21 , , , 37809 347 22 from from IN 37809 347 23 La La NNP 37809 347 24 Esperanza Esperanza NNP 37809 347 25 , , , 37809 347 26 near near IN 37809 347 27 Escuintla Escuintla NNP 37809 347 28 , , , 37809 347 29 Chiapas Chiapas NNP 37809 347 30 , , , 37809 347 31 México México NNP 37809 347 32 , , , 37809 347 33 collected collect VBD 37809 347 34 May May NNP 37809 347 35 13 13 CD 37809 347 36 , , , 37809 347 37 1940 1940 CD 37809 347 38 , , , 37809 347 39 by by IN 37809 347 40 H. H. NNP 37809 347 41 M. M. NNP 37809 347 42 and and CC 37809 347 43 R. R. NNP 37809 347 44 Smith Smith NNP 37809 347 45 ] ] -RRB- 37809 347 46 . . . 37809 348 1 Duellman duellman NN 37809 348 2 , , , 37809 348 3 1958:1 1958:1 CD 37809 348 4 - - SYM 37809 348 5 4 4 CD 37809 348 6 , , , 37809 348 7 7 7 CD 37809 348 8 , , , 37809 348 9 12 12 CD 37809 348 10 , , , 37809 348 11 14 14 CD 37809 348 12 . . . 37809 349 1 Gorham Gorham NNP 37809 349 2 , , , 37809 349 3 1966:167 1966:167 CD 37809 349 4 . . . 37809 350 1 _ _ NNP 37809 350 2 Syrrhophus Syrrhophus NNP 37809 350 3 rubrimaculata rubrimaculata NN 37809 350 4 _ _ NNP 37809 350 5 : : : 37809 350 6 Smith Smith NNP 37809 350 7 and and CC 37809 350 8 Taylor Taylor NNP 37809 350 9 , , , 37809 350 10 1948:48 1948:48 NNP 37809 350 11 - - HYPH 37809 350 12 49 49 CD 37809 350 13 . . . 37809 351 1 [ [ -LRB- 37809 351 2 Illustration illustration NN 37809 351 3 : : : 37809 351 4 FIG FIG NNP 37809 351 5 . . . 37809 352 1 8 8 LS 37809 352 2 : : : 37809 352 3 _ _ NNP 37809 352 4 Syrrhophus Syrrhophus NNP 37809 352 5 rubrimaculatus rubrimaculatus NN 37809 352 6 _ _ NNP 37809 352 7 ( ( -LRB- 37809 352 8 upper upper JJ 37809 352 9 right right UH 37809 352 10 , , , 37809 352 11 KU KU NNP 37809 352 12 58911 58911 CD 37809 352 13 , , , 37809 352 14 ×1.6 ×1.6 NNP 37809 352 15 ; ; : 37809 352 16 lower lower RBR 37809 352 17 right right UH 37809 352 18 , , , 37809 352 19 KU KU NNP 37809 352 20 58910 58910 CD 37809 352 21 , , , 37809 352 22 ×4 ×4 LS 37809 352 23 ) ) -RRB- 37809 352 24 and and CC 37809 352 25 _ _ NNP 37809 352 26 S. S. NNP 37809 353 1 verrucipes verrucipes NNP 37809 353 2 _ _ NNP 37809 353 3 ( ( -LRB- 37809 353 4 upper upper JJ 37809 353 5 left left NN 37809 353 6 , , , 37809 353 7 UIMNH UIMNH NNP 37809 353 8 15995 15995 CD 37809 353 9 , , , 37809 353 10 ×1.6 ×1.6 NNP 37809 353 11 ; ; : 37809 353 12 lower low JJR 37809 353 13 left leave VBN 37809 353 14 , , , 37809 353 15 UIMNH UIMNH NNP 37809 353 16 15989 15989 CD 37809 353 17 , , , 37809 353 18 ×3.7 ×3.7 NNP 37809 353 19 ) ) -RRB- 37809 353 20 . . . 37809 353 21 ] ] -RRB- 37809 354 1 _ _ NNP 37809 354 2 Diagnosis._--Small Diagnosis._--Small NNP 37809 354 3 frogs frog NNS 37809 354 4 , , , 37809 354 5 males male VBZ 37809 354 6 18.2 18.2 CD 37809 354 7 - - SYM 37809 354 8 23.5 23.5 CD 37809 354 9 mm mm NN 37809 354 10 . . . 37809 355 1 snout snout NN 37809 355 2 - - HYPH 37809 355 3 vent vent NN 37809 355 4 , , , 37809 355 5 females female VBZ 37809 355 6 19.0 19.0 CD 37809 355 7 - - : 37809 355 8 22.5 22.5 CD 37809 355 9 mm mm NNP 37809 355 10 . . . 37809 356 1 snout snout NN 37809 356 2 - - HYPH 37809 356 3 vent vent NN 37809 356 4 length length NN 37809 356 5 ( ( -LRB- 37809 356 6 small small JJ 37809 356 7 sample sample NN 37809 356 8 ) ) -RRB- 37809 356 9 ; ; : 37809 356 10 vocal vocal NN 37809 356 11 slits slit NNS 37809 356 12 in in IN 37809 356 13 males male NNS 37809 356 14 ; ; , 37809 356 15 digital digital JJ 37809 356 16 tips tip NNS 37809 356 17 scarcely scarcely RB 37809 356 18 expanded expand VBN 37809 356 19 ( ( -LRB- 37809 356 20 Fig fig NN 37809 356 21 . . . 37809 357 1 1 1 LS 37809 357 2 ) ) -RRB- 37809 357 3 ; ; : 37809 357 4 first first RB 37809 357 5 finger finger NN 37809 357 6 shorter short JJR 37809 357 7 than than IN 37809 357 8 second second JJ 37809 357 9 ; ; : 37809 357 10 outer outer JJ 37809 357 11 palmar palmar NNP 37809 357 12 tubercle tubercle NN 37809 357 13 reduced reduce VBN 37809 357 14 in in IN 37809 357 15 size size NN 37809 357 16 ; ; , 37809 357 17 inner inner JJ 37809 357 18 metatarsal metatarsal NN 37809 357 19 tubercle tubercle NN 37809 357 20 elongate elongate NN 37809 357 21 , , , 37809 357 22 twice twice PDT 37809 357 23 the the DT 37809 357 24 size size NN 37809 357 25 of of IN 37809 357 26 small small JJ 37809 357 27 , , , 37809 357 28 conical conical JJ 37809 357 29 outer outer JJ 37809 357 30 metatarsal metatarsal NN 37809 357 31 tubercle tubercle NN 37809 357 32 ; ; : 37809 357 33 diameter diameter NN 37809 357 34 of of IN 37809 357 35 tympanum tympanum NN 37809 357 36 35.5 35.5 CD 37809 357 37 - - SYM 37809 357 38 46.5 46.5 CD 37809 357 39 per per IN 37809 357 40 cent cent NN 37809 357 41 that that DT 37809 357 42 of of IN 37809 357 43 eye eye NN 37809 357 44 in in IN 37809 357 45 both both DT 37809 357 46 sexes sex NNS 37809 357 47 ; ; , 37809 357 48 dorsum dorsum NN 37809 357 49 brown brown NNP 37809 357 50 with with IN 37809 357 51 small small JJ 37809 357 52 pale pale JJ 37809 357 53 spots spot NNS 37809 357 54 ( ( -LRB- 37809 357 55 red red NN 37809 357 56 in in IN 37809 357 57 life life NN 37809 357 58 ) ) -RRB- 37809 357 59 ; ; : 37809 357 60 venter venter NN 37809 357 61 gray gray JJ 37809 357 62 . . . 37809 358 1 _ _ NNP 37809 358 2 Remarks._--Previous remarks._--previous JJ 37809 358 3 authors author NNS 37809 358 4 who who WP 37809 358 5 treated treat VBD 37809 358 6 _ _ NNP 37809 358 7 Syrrhophus Syrrhophus NNP 37809 358 8 _ _ NNP 37809 358 9 placed place VBD 37809 358 10 this this DT 37809 358 11 species specie NNS 37809 358 12 in in IN 37809 358 13 the the DT 37809 358 14 western western JJ 37809 358 15 complex complex NN 37809 358 16 , , , 37809 358 17 because because IN 37809 358 18 it -PRON- PRP 37809 358 19 occurs occur VBZ 37809 358 20 on on IN 37809 358 21 the the DT 37809 358 22 Pacific Pacific NNP 37809 358 23 versant versant NN 37809 358 24 and and CC 37809 358 25 has have VBZ 37809 358 26 a a DT 37809 358 27 reduced reduced JJ 37809 358 28 outer outer JJ 37809 358 29 palmar palmar NNP 37809 358 30 tubercle tubercle NN 37809 358 31 . . . 37809 359 1 Duellman Duellman NNP 37809 359 2 ( ( -LRB- 37809 359 3 1958 1958 CD 37809 359 4 ) ) -RRB- 37809 359 5 placed place VBN 37809 359 6 _ _ NNP 37809 359 7 rubrimaculatus rubrimaculatus NN 37809 359 8 _ _ NNP 37809 359 9 apart apart RB 37809 359 10 from from IN 37809 359 11 the the DT 37809 359 12 other other JJ 37809 359 13 western western JJ 37809 359 14 species specie NNS 37809 359 15 , , , 37809 359 16 because because IN 37809 359 17 of of IN 37809 359 18 its -PRON- PRP$ 37809 359 19 relatively relatively RB 37809 359 20 unexpanded unexpanded JJ 37809 359 21 digital digital JJ 37809 359 22 tips tip NNS 37809 359 23 and and CC 37809 359 24 coloration coloration NN 37809 359 25 . . . 37809 360 1 The the DT 37809 360 2 digital digital JJ 37809 360 3 tips tip NNS 37809 360 4 are be VBP 37809 360 5 like like IN 37809 360 6 those those DT 37809 360 7 in in IN 37809 360 8 _ _ NNP 37809 360 9 leprus leprus NNP 37809 360 10 _ _ NNP 37809 360 11 , , , 37809 360 12 which which WDT 37809 360 13 _ _ NNP 37809 360 14 rubrimaculatus rubrimaculatus NN 37809 360 15 _ _ NNP 37809 360 16 resembles resemble VBZ 37809 360 17 . . . 37809 361 1 Except except IN 37809 361 2 for for IN 37809 361 3 the the DT 37809 361 4 reduction reduction NN 37809 361 5 of of IN 37809 361 6 the the DT 37809 361 7 outer outer JJ 37809 361 8 palmar palmar NNP 37809 361 9 tubercle tubercle NN 37809 361 10 , , , 37809 361 11 _ _ NNP 37809 361 12 rubrimaculatus rubrimaculatus NN 37809 361 13 _ _ NNP 37809 361 14 could could MD 37809 361 15 be be VB 37809 361 16 a a DT 37809 361 17 member member NN 37809 361 18 of of IN 37809 361 19 the the DT 37809 361 20 _ _ NNP 37809 361 21 leprus leprus NNP 37809 361 22 _ _ NNP 37809 361 23 group group NN 37809 361 24 . . . 37809 362 1 _ _ NNP 37809 362 2 Syrrhophus Syrrhophus NNP 37809 362 3 rubrimaculatus rubrimaculatus NN 37809 362 4 _ _ NNP 37809 362 5 is be VBZ 37809 362 6 probably probably RB 37809 362 7 best well RBS 37809 362 8 treated treat VBN 37809 362 9 as as IN 37809 362 10 a a DT 37809 362 11 Pacific Pacific NNP 37809 362 12 derivative derivative NN 37809 362 13 of of IN 37809 362 14 the the DT 37809 362 15 _ _ NNP 37809 362 16 leprus leprus NNP 37809 362 17 _ _ NNP 37809 362 18 group group NN 37809 362 19 , , , 37809 362 20 even even RB 37809 362 21 though though IN 37809 362 22 the the DT 37809 362 23 palmar palmar NNP 37809 362 24 tubercles tubercle NNS 37809 362 25 do do VBP 37809 362 26 not not RB 37809 362 27 agree agree VB 37809 362 28 . . . 37809 363 1 The the DT 37809 363 2 removal removal NN 37809 363 3 of of IN 37809 363 4 _ _ NNP 37809 363 5 rubrimaculatus rubrimaculatus NN 37809 363 6 _ _ NNP 37809 363 7 from from IN 37809 363 8 the the DT 37809 363 9 western western JJ 37809 363 10 complex complex JJ 37809 363 11 results result NNS 37809 363 12 in in IN 37809 363 13 a a DT 37809 363 14 more more RBR 37809 363 15 homogeneous homogeneous JJ 37809 363 16 remainder remainder NN 37809 363 17 and and CC 37809 363 18 does do VBZ 37809 363 19 not not RB 37809 363 20 greatly greatly RB 37809 363 21 increase increase VB 37809 363 22 the the DT 37809 363 23 heterogeneity heterogeneity NN 37809 363 24 of of IN 37809 363 25 the the DT 37809 363 26 eastern eastern JJ 37809 363 27 complex complex NN 37809 363 28 . . . 37809 364 1 _ _ NNP 37809 364 2 Etymology._--Latin Etymology._--Latin NNP 37809 364 3 , , , 37809 364 4 meaning mean VBG 37809 364 5 spotted spot VBN 37809 364 6 with with IN 37809 364 7 red red JJ 37809 364 8 ; ; : 37809 364 9 in in IN 37809 364 10 reference reference NN 37809 364 11 to to IN 37809 364 12 the the DT 37809 364 13 colors color NNS 37809 364 14 in in IN 37809 364 15 life life NN 37809 364 16 . . . 37809 365 1 _ _ NNP 37809 365 2 Distribution._--Low Distribution._--Low NNP 37809 365 3 to to TO 37809 365 4 moderate moderate VB 37809 365 5 elevations elevation NNS 37809 365 6 on on IN 37809 365 7 the the DT 37809 365 8 Pacific Pacific NNP 37809 365 9 versant versant NN 37809 365 10 of of IN 37809 365 11 southeastern southeastern JJ 37809 365 12 Chiapas Chiapas NNP 37809 365 13 , , , 37809 365 14 México México NNP 37809 365 15 ( ( -LRB- 37809 365 16 Fig Fig NNP 37809 365 17 . . . 37809 366 1 7 7 LS 37809 366 2 ) ) -RRB- 37809 366 3 ; ; : 37809 366 4 probably probably RB 37809 366 5 extending extend VBG 37809 366 6 into into IN 37809 366 7 adjacent adjacent JJ 37809 366 8 Guatemala Guatemala NNP 37809 366 9 . . . 37809 367 1 _ _ NNP 37809 367 2 Specimens Specimens NNP 37809 367 3 examined._--(48 examined._--(48 NNP 37809 367 4 ) ) -RRB- 37809 367 5 MÉXICO MÉXICO NNP 37809 367 6 , , , 37809 367 7 _ _ NNP 37809 367 8 Chiapas Chiapas NNP 37809 367 9 _ _ NNP 37809 367 10 : : : 37809 367 11 Escuintla Escuintla NNP 37809 367 12 , , , 37809 367 13 UMMZ UMMZ NNP 37809 367 14 88283 88283 CD 37809 367 15 ; ; : 37809 367 16 6 6 CD 37809 367 17 km km NN 37809 367 18 . . . 37809 368 1 NE NE NNP 37809 368 2 Escuintla Escuintla NNP 37809 368 3 , , , 37809 368 4 UMMZ UMMZ NNP 37809 368 5 87876 87876 CD 37809 368 6 - - SYM 37809 368 7 80 80 CD 37809 368 8 ; ; : 37809 368 9 La La NNP 37809 368 10 Esperanza Esperanza NNP 37809 368 11 , , , 37809 368 12 UIMNH UIMNH NNP 37809 368 13 13285 13285 CD 37809 368 14 , , , 37809 368 15 UMMZ UMMZ NNP 37809 368 16 88496 88496 CD 37809 368 17 - - SYM 37809 368 18 97 97 CD 37809 368 19 , , , 37809 368 20 USNM USNM NNP 37809 368 21 114070 114070 CD 37809 368 22 ( ( -LRB- 37809 368 23 holotype holotype NNP 37809 368 24 ) ) -RRB- 37809 368 25 , , , 37809 368 26 114054 114054 CD 37809 368 27 - - SYM 37809 368 28 69 69 CD 37809 368 29 , , , 37809 368 30 114072 114072 CD 37809 368 31 ; ; : 37809 368 32 Monte Monte NNP 37809 368 33 Cristo Cristo NNP 37809 368 34 , , , 37809 368 35 UMMZ UMMZ NNP 37809 368 36 88353 88353 CD 37809 368 37 ; ; : 37809 368 38 1.3 1.3 CD 37809 368 39 km km NNS 37809 368 40 . . . 37809 369 1 N N NNP 37809 369 2 Puerto Puerto NNP 37809 369 3 Madero Madero NNP 37809 369 4 , , , 37809 369 5 KU KU NNP 37809 369 6 58910 58910 CD 37809 369 7 - - SYM 37809 369 8 11 11 CD 37809 369 9 ; ; : 37809 369 10 Finca Finca NNP 37809 369 11 San San NNP 37809 369 12 Jerónimo Jerónimo NNP 37809 369 13 , , , 37809 369 14 600 600 CD 37809 369 15 - - SYM 37809 369 16 650 650 CD 37809 369 17 m. m. NN 37809 369 18 , , , 37809 369 19 UIMNH UIMNH NNP 37809 369 20 55299 55299 CD 37809 369 21 - - SYM 37809 369 22 312 312 CD 37809 369 23 , , , 37809 369 24 55313 55313 CD 37809 369 25 - - SYM 37809 369 26 16 16 CD 37809 369 27 ( ( -LRB- 37809 369 28 cleared clear VBN 37809 369 29 and and CC 37809 369 30 stained stain VBN 37809 369 31 ) ) -RRB- 37809 369 32 . . . 37809 370 1 = = NFP 37809 370 2 Syrrhophus Syrrhophus NNP 37809 370 3 guttilatus= guttilatus= NN 37809 370 4 ( ( -LRB- 37809 370 5 Cope Cope NNP 37809 370 6 ) ) -RRB- 37809 370 7 _ _ NNP 37809 370 8 Malachylodes Malachylodes NNP 37809 370 9 guttilatus guttilatus NN 37809 370 10 _ _ NNP 37809 370 11 Cope Cope NNP 37809 370 12 , , , 37809 370 13 1879:264 1879:264 CD 37809 370 14 [ [ -LRB- 37809 370 15 Holotype.--USNM Holotype.--USNM NNP 37809 370 16 9888 9888 CD 37809 370 17 , , , 37809 370 18 from from IN 37809 370 19 Guanajuato Guanajuato NNP 37809 370 20 , , , 37809 370 21 Guanajuato Guanajuato NNP 37809 370 22 , , , 37809 370 23 México México NNP 37809 370 24 ; ; , 37809 370 25 collected collect VBN 37809 370 26 in in IN 37809 370 27 1877 1877 CD 37809 370 28 by by IN 37809 370 29 Alfredo Alfredo NNP 37809 370 30 Duges Duges NNPS 37809 370 31 ] ] -RRB- 37809 370 32 . . . 37809 371 1 _ _ NNP 37809 371 2 Syrrhopus Syrrhopus NNP 37809 371 3 guttulatus guttulatus NN 37809 371 4 _ _ NNP 37809 371 5 : : : 37809 371 6 Boulenger boulenger NN 37809 371 7 , , , 37809 371 8 1888:204 1888:204 CD 37809 371 9 - - SYM 37809 371 10 06 06 CD 37809 371 11 . . . 37809 372 1 _ _ NNP 37809 372 2 Syrrhaphus Syrrhaphus NNP 37809 372 3 guttulatus guttulatus NNP 37809 372 4 _ _ NNP 37809 372 5 : : : 37809 372 6 Günther Günther NNP 37809 372 7 , , , 37809 372 8 1900:317 1900:317 CD 37809 372 9 . . . 37809 373 1 _ _ NNP 37809 373 2 Syrraphus Syrraphus NNP 37809 373 3 guttulatus guttulatus NNP 37809 373 4 _ _ NNP 37809 373 5 : : : 37809 373 6 Díaz Díaz NNP 37809 373 7 de de FW 37809 373 8 León León NNP 37809 373 9 , , , 37809 373 10 1904:11 1904:11 CD 37809 373 11 . . . 37809 374 1 _ _ NNP 37809 374 2 Syrrhophus Syrrhophus NNP 37809 374 3 guttilatus guttilatus NN 37809 374 4 _ _ NNP 37809 374 5 : : : 37809 374 6 Nieden Nieden NNP 37809 374 7 , , , 37809 374 8 1923:399 1923:399 CD 37809 374 9 - - SYM 37809 374 10 400 400 CD 37809 374 11 . . . 37809 375 1 Kellogg Kellogg NNP 37809 375 2 , , , 37809 375 3 1932:125 1932:125 CD 37809 375 4 , , , 37809 375 5 127 127 CD 37809 375 6 - - SYM 37809 375 7 28 28 CD 37809 375 8 . . . 37809 376 1 Smith Smith NNP 37809 376 2 and and CC 37809 376 3 Taylor Taylor NNP 37809 376 4 , , , 37809 376 5 1948:49 1948:49 CD 37809 376 6 , , , 37809 376 7 51 51 CD 37809 376 8 . . . 37809 377 1 Firschein Firschein NNP 37809 377 2 , , , 37809 377 3 1954:52 1954:52 CD 37809 377 4 - - SYM 37809 377 5 54 54 CD 37809 377 6 . . . 37809 378 1 Gorham Gorham NNP 37809 378 2 , , , 37809 378 3 1966:164 1966:164 CD 37809 378 4 . . . 37809 379 1 _ _ NNP 37809 379 2 Syrrhophus Syrrhophus NNP 37809 379 3 smithi smithi NN 37809 379 4 _ _ NNP 37809 379 5 Taylor Taylor NNP 37809 379 6 , , , 37809 379 7 1940b:43 1940b:43 CD 37809 379 8 - - SYM 37809 379 9 45 45 CD 37809 379 10 , , , 37809 379 11 pl pl NNP 37809 379 12 . . . 37809 380 1 1 1 CD 37809 380 2 [ [ -LRB- 37809 380 3 Holotype.--USNM Holotype.--USNM NNP 37809 380 4 108594 108594 CD 37809 380 5 , , , 37809 380 6 from from IN 37809 380 7 15 15 CD 37809 380 8 mi mi NN 37809 380 9 . . . 37809 381 1 SW SW NNP 37809 381 2 Galeana Galeana NNP 37809 381 3 , , , 37809 381 4 Nuevo Nuevo NNP 37809 381 5 León León NNP 37809 381 6 , , , 37809 381 7 México México NNP 37809 381 8 , , , 37809 381 9 1575 1575 CD 37809 381 10 m. m. NN 37809 381 11 ; ; , 37809 381 12 collected collect VBN 37809 381 13 on on IN 37809 381 14 October October NNP 37809 381 15 13 13 CD 37809 381 16 , , , 37809 381 17 1939 1939 CD 37809 381 18 , , , 37809 381 19 by by IN 37809 381 20 Hobart Hobart NNP 37809 381 21 M. M. NNP 37809 381 22 Smith Smith NNP 37809 381 23 ] ] -RRB- 37809 381 24 . . . 37809 382 1 Smith Smith NNP 37809 382 2 and and CC 37809 382 3 Taylor Taylor NNP 37809 382 4 , , , 37809 382 5 1948:49 1948:49 CD 37809 382 6 , , , 37809 382 7 51 51 CD 37809 382 8 . . . 37809 383 1 Firschein Firschein NNP 37809 383 2 , , , 37809 383 3 1954:54 1954:54 CD 37809 383 4 - - SYM 37809 383 5 55 55 CD 37809 383 6 . . . 37809 384 1 Martin Martin NNP 37809 384 2 , , , 37809 384 3 1958:50 1958:50 CD 37809 384 4 . . . 37809 385 1 Gorham Gorham NNP 37809 385 2 , , , 37809 385 3 1966:167 1966:167 CD 37809 385 4 . . . 37809 386 1 _ _ NNP 37809 386 2 Syrrhophus Syrrhophus NNP 37809 386 3 gaigeae gaigeae NN 37809 386 4 _ _ NNP 37809 386 5 Schmidt Schmidt NNP 37809 386 6 and and CC 37809 386 7 Smith Smith NNP 37809 386 8 , , , 37809 386 9 1944:80 1944:80 CD 37809 386 10 [ [ -LRB- 37809 386 11 Holotype.--FMNH holotype.--fmnh PRP$ 37809 386 12 27361 27361 CD 37809 386 13 , , , 37809 386 14 from from IN 37809 386 15 the the DT 37809 386 16 Basin Basin NNP 37809 386 17 , , , 37809 386 18 Chisos Chisos NNP 37809 386 19 Mountains Mountains NNPS 37809 386 20 , , , 37809 386 21 Brewster Brewster NNP 37809 386 22 Co. Co. NNP 37809 386 23 , , , 37809 386 24 Texas Texas NNP 37809 386 25 ; ; : 37809 386 26 collected collect VBN 37809 386 27 on on IN 37809 386 28 July July NNP 37809 386 29 24 24 CD 37809 386 30 , , , 37809 386 31 1937 1937 CD 37809 386 32 , , , 37809 386 33 by by IN 37809 386 34 Walter Walter NNP 37809 386 35 L. L. NNP 37809 386 36 Necker Necker NNP 37809 386 37 ] ] -RRB- 37809 386 38 . . . 37809 387 1 _ _ NNP 37809 387 2 Syrrhophus Syrrhophus NNP 37809 387 3 petrophilus petrophilus NN 37809 387 4 _ _ NNP 37809 387 5 Firschein Firschein NNP 37809 387 6 , , , 37809 387 7 1954:50 1954:50 CD 37809 387 8 - - SYM 37809 387 9 52 52 CD 37809 387 10 [ [ -LRB- 37809 387 11 Holotype.--UIMNH Holotype.--UIMNH NNP 37809 387 12 7807 7807 CD 37809 387 13 , , , 37809 387 14 from from IN 37809 387 15 5 5 CD 37809 387 16 km km NNS 37809 387 17 . . . 37809 388 1 SW SW NNP 37809 388 2 San San NNP 37809 388 3 Luis Luis NNP 37809 388 4 Potosí Potosí NNP 37809 388 5 , , , 37809 388 6 San San NNP 37809 388 7 Luis Luis NNP 37809 388 8 Potosí Potosí NNP 37809 388 9 , , , 37809 388 10 México México NNP 37809 388 11 ; ; , 37809 388 12 collected collect VBN 37809 388 13 on on IN 37809 388 14 July July NNP 37809 388 15 18 18 CD 37809 388 16 , , , 37809 388 17 1949 1949 CD 37809 388 18 , , , 37809 388 19 by by IN 37809 388 20 David David NNP 37809 388 21 Langebartel Langebartel NNP 37809 388 22 ] ] -RRB- 37809 388 23 . . . 37809 389 1 Gorham Gorham NNP 37809 389 2 , , , 37809 389 3 1966:166 1966:166 CD 37809 389 4 . . . 37809 390 1 _ _ NNP 37809 390 2 Syrrhophus Syrrhophus NNP 37809 390 3 marnocki marnocki NN 37809 390 4 _ _ NNP 37809 390 5 : : : 37809 390 6 Milstead Milstead NNP 37809 390 7 , , , 37809 390 8 Mecham Mecham NNP 37809 390 9 , , , 37809 390 10 and and CC 37809 390 11 McClintock McClintock NNP 37809 390 12 , , , 37809 390 13 1950:548 1950:548 CD 37809 390 14 ( ( -LRB- 37809 390 15 in in IN 37809 390 16 part part NN 37809 390 17 ) ) -RRB- 37809 390 18 . . . 37809 391 1 _ _ NNP 37809 391 2 Diagnosis._--Medium Diagnosis._--Medium NNP 37809 391 3 - - HYPH 37809 391 4 sized sized JJ 37809 391 5 frogs frog NNS 37809 391 6 , , , 37809 391 7 males male VBZ 37809 391 8 20.6 20.6 CD 37809 391 9 - - SYM 37809 391 10 29.0 29.0 CD 37809 391 11 mm mm NN 37809 391 12 . . . 37809 392 1 snout snout NN 37809 392 2 - - HYPH 37809 392 3 vent vent NN 37809 392 4 , , , 37809 392 5 females female VBZ 37809 392 6 25.7 25.7 CD 37809 392 7 - - HYPH 37809 392 8 31.0 31.0 CD 37809 392 9 mm mm NN 37809 392 10 . . . 37809 393 1 snout snout NN 37809 393 2 - - HYPH 37809 393 3 vent vent NN 37809 393 4 length length NN 37809 393 5 ; ; : 37809 393 6 vocal vocal NN 37809 393 7 slits slit NNS 37809 393 8 in in IN 37809 393 9 males male NNS 37809 393 10 ; ; , 37809 393 11 digital digital JJ 37809 393 12 tips tip NNS 37809 393 13 slightly slightly RB 37809 393 14 expanded expand VBN 37809 393 15 ( ( -LRB- 37809 393 16 Fig fig NN 37809 393 17 . . . 37809 394 1 1 1 LS 37809 394 2 ) ) -RRB- 37809 394 3 ; ; : 37809 394 4 first first JJ 37809 394 5 and and CC 37809 394 6 second second JJ 37809 394 7 fingers finger NNS 37809 394 8 equal equal JJ 37809 394 9 ; ; : 37809 394 10 skin skin NN 37809 394 11 of of IN 37809 394 12 dorsum dorsum NN 37809 394 13 smooth smooth JJ 37809 394 14 to to IN 37809 394 15 moderately moderately RB 37809 394 16 pustular pustular VB 37809 394 17 , , , 37809 394 18 that that DT 37809 394 19 of of IN 37809 394 20 venter venter NN 37809 394 21 smooth smooth JJ 37809 394 22 ; ; : 37809 394 23 snout snout NN 37809 394 24 blunt blunt JJ 37809 394 25 ; ; : 37809 394 26 diameter diameter NN 37809 394 27 of of IN 37809 394 28 tympanum tympanum CD 37809 394 29 55.1 55.1 CD 37809 394 30 - - SYM 37809 394 31 75.7 75.7 CD 37809 394 32 per per IN 37809 394 33 cent cent NN 37809 394 34 that that DT 37809 394 35 of of IN 37809 394 36 eye eye NN 37809 394 37 in in IN 37809 394 38 males male NNS 37809 394 39 , , , 37809 394 40 47.6 47.6 CD 37809 394 41 - - SYM 37809 394 42 61.7 61.7 CD 37809 394 43 in in IN 37809 394 44 females female NNS 37809 394 45 ; ; : 37809 394 46 dorsum dorsum NN 37809 394 47 and and CC 37809 394 48 flanks flank NNS 37809 394 49 cream cream NN 37809 394 50 to to TO 37809 394 51 gray gray VB 37809 394 52 with with IN 37809 394 53 light light JJ 37809 394 54 brown brown NNP 37809 394 55 to to IN 37809 394 56 black black JJ 37809 394 57 flecking flecking NN 37809 394 58 and and CC 37809 394 59 vermiculations vermiculation NNS 37809 394 60 ; ; , 37809 394 61 thighs thigh NNS 37809 394 62 usually usually RB 37809 394 63 not not RB 37809 394 64 banded band VBN 37809 394 65 ; ; : 37809 394 66 interorbital interorbital NNP 37809 394 67 bar bar NN 37809 394 68 present present NN 37809 394 69 ( ( -LRB- 37809 394 70 Fig fig NN 37809 394 71 . . . 37809 395 1 8) 8) CD 37809 395 2 . . . 37809 396 1 [ [ -LRB- 37809 396 2 Illustration illustration NN 37809 396 3 : : : 37809 396 4 FIG FIG NNP 37809 396 5 . . . 37809 397 1 9 9 LS 37809 397 2 : : : 37809 397 3 _ _ NNP 37809 397 4 Syrrhophus Syrrhophus NNP 37809 397 5 guttilatus guttilatus NN 37809 397 6 _ _ NNP 37809 397 7 ( ( -LRB- 37809 397 8 upper upper JJ 37809 397 9 left left NN 37809 397 10 , , , 37809 397 11 UIMNH UIMNH NNP 37809 397 12 55519 55519 CD 37809 397 13 , , , 37809 397 14 ×1.4 ×1.4 XX 37809 397 15 ; ; : 37809 397 16 lower low JJR 37809 397 17 left leave VBN 37809 397 18 , , , 37809 397 19 UIMNH UIMNH NNP 37809 397 20 55519 55519 CD 37809 397 21 , , , 37809 397 22 ×2.3 ×2.3 LS 37809 397 23 ) ) -RRB- 37809 397 24 and and CC 37809 397 25 _ _ NNP 37809 397 26 S. S. NNP 37809 398 1 marnockii marnockii NNP 37809 398 2 _ _ NNP 37809 398 3 ( ( -LRB- 37809 398 4 upper upper JJ 37809 398 5 right right NN 37809 398 6 , , , 37809 398 7 TCWC TCWC NNP 37809 398 8 9317 9317 CD 37809 398 9 , , , 37809 398 10 ×1.4 ×1.4 DT 37809 398 11 ; ; : 37809 398 12 lower lower RBR 37809 398 13 right right RB 37809 398 14 , , , 37809 398 15 TCWC TCWC NNP 37809 398 16 13510 13510 CD 37809 398 17 , , , 37809 398 18 ×2.1 ×2.1 LS 37809 398 19 ) ) -RRB- 37809 398 20 . . . 37809 398 21 ] ] -RRB- 37809 399 1 _ _ NNP 37809 399 2 Remarks._--Cope Remarks._--Cope NNP 37809 399 3 ( ( -LRB- 37809 399 4 1879 1879 CD 37809 399 5 ) ) -RRB- 37809 399 6 distinguished distinguish VBD 37809 399 7 _ _ NNP 37809 399 8 Malachylodes Malachylodes NNP 37809 399 9 _ _ NNP 37809 399 10 from from IN 37809 399 11 _ _ NNP 37809 399 12 Syrrhophus Syrrhophus NNP 37809 399 13 _ _ NNP 37809 399 14 on on IN 37809 399 15 the the DT 37809 399 16 basis basis NN 37809 399 17 of of IN 37809 399 18 the the DT 37809 399 19 presence presence NN 37809 399 20 of of IN 37809 399 21 a a DT 37809 399 22 frontoparietal frontoparietal JJ 37809 399 23 fontanelle fontanelle NN 37809 399 24 in in IN 37809 399 25 the the DT 37809 399 26 holotype holotype NN 37809 399 27 of of IN 37809 399 28 _ _ NNP 37809 399 29 guttilatus guttilatus NNP 37809 399 30 _ _ NNP 37809 399 31 . . . 37809 400 1 The the DT 37809 400 2 holotype holotype NN 37809 400 3 is be VBZ 37809 400 4 a a DT 37809 400 5 juvenile juvenile JJ 37809 400 6 female female NN 37809 400 7 and and CC 37809 400 8 as as IN 37809 400 9 is be VBZ 37809 400 10 the the DT 37809 400 11 case case NN 37809 400 12 in in IN 37809 400 13 the the DT 37809 400 14 juveniles juvenile NNS 37809 400 15 of of IN 37809 400 16 nearly nearly RB 37809 400 17 all all DT 37809 400 18 leptodactylids leptodactylid NNS 37809 400 19 , , , 37809 400 20 a a DT 37809 400 21 frontoparietal frontoparietal JJ 37809 400 22 fontanelle fontanelle NN 37809 400 23 is be VBZ 37809 400 24 present present JJ 37809 400 25 . . . 37809 401 1 Firschein Firschein NNP 37809 401 2 ( ( -LRB- 37809 401 3 1954 1954 CD 37809 401 4 ) ) -RRB- 37809 401 5 used use VBD 37809 401 6 the the DT 37809 401 7 presence presence NN 37809 401 8 of of IN 37809 401 9 the the DT 37809 401 10 fontanelle fontanelle NN 37809 401 11 to to TO 37809 401 12 distinguish distinguish VB 37809 401 13 _ _ NNP 37809 401 14 guttilatus guttilatus NN 37809 401 15 _ _ NNP 37809 401 16 from from IN 37809 401 17 his -PRON- PRP$ 37809 401 18 _ _ NNP 37809 401 19 petrophilus petrophilus NN 37809 401 20 _ _ NNP 37809 401 21 . . . 37809 402 1 As as IN 37809 402 2 is be VBZ 37809 402 3 clearly clearly RB 37809 402 4 evident evident JJ 37809 402 5 from from IN 37809 402 6 the the DT 37809 402 7 length length NN 37809 402 8 of of IN 37809 402 9 the the DT 37809 402 10 synonymy synonymy NN 37809 402 11 , , , 37809 402 12 I -PRON- PRP 37809 402 13 consider consider VBP 37809 402 14 a a DT 37809 402 15 number number NN 37809 402 16 of of IN 37809 402 17 currently currently RB 37809 402 18 used use VBN 37809 402 19 names name NNS 37809 402 20 to to TO 37809 402 21 be be VB 37809 402 22 synonymous synonymous JJ 37809 402 23 with with IN 37809 402 24 _ _ NNP 37809 402 25 guttilatus guttilatus NNP 37809 402 26 _ _ NNP 37809 402 27 . . . 37809 403 1 I -PRON- PRP 37809 403 2 have have VBP 37809 403 3 seen see VBN 37809 403 4 the the DT 37809 403 5 holotypes holotype NNS 37809 403 6 of of IN 37809 403 7 all all DT 37809 403 8 four four CD 37809 403 9 names name NNS 37809 403 10 and and CC 37809 403 11 am be VBP 37809 403 12 unable unable JJ 37809 403 13 to to TO 37809 403 14 recognize recognize VB 37809 403 15 more more JJR 37809 403 16 than than IN 37809 403 17 a a DT 37809 403 18 single single JJ 37809 403 19 species specie NNS 37809 403 20 . . . 37809 404 1 The the DT 37809 404 2 holotype holotype NN 37809 404 3 of of IN 37809 404 4 _ _ NNP 37809 404 5 petrophilus petrophilus NN 37809 404 6 _ _ NNP 37809 404 7 is be VBZ 37809 404 8 a a DT 37809 404 9 male male NN 37809 404 10 , , , 37809 404 11 whereas whereas IN 37809 404 12 that that DT 37809 404 13 of of IN 37809 404 14 _ _ NNP 37809 404 15 smithi smithi NN 37809 404 16 _ _ NNP 37809 404 17 is be VBZ 37809 404 18 a a DT 37809 404 19 female female NN 37809 404 20 . . . 37809 405 1 The the DT 37809 405 2 supposed suppose VBN 37809 405 3 differences difference NNS 37809 405 4 are be VBP 37809 405 5 a a DT 37809 405 6 reflection reflection NN 37809 405 7 of of IN 37809 405 8 sexual sexual JJ 37809 405 9 dimorphism dimorphism NN 37809 405 10 in in IN 37809 405 11 the the DT 37809 405 12 size size NN 37809 405 13 of of IN 37809 405 14 the the DT 37809 405 15 eye eye NN 37809 405 16 ( ( -LRB- 37809 405 17 Table table NN 37809 405 18 5 5 CD 37809 405 19 ) ) -RRB- 37809 405 20 . . . 37809 406 1 The the DT 37809 406 2 two two CD 37809 406 3 holotypes holotype NNS 37809 406 4 , , , 37809 406 5 as as RB 37809 406 6 well well RB 37809 406 7 as as IN 37809 406 8 those those DT 37809 406 9 of of IN 37809 406 10 _ _ NNP 37809 406 11 gaigeae gaigeae NN 37809 406 12 _ _ NNP 37809 406 13 and and CC 37809 406 14 _ _ NNP 37809 406 15 Malachylodes Malachylodes NNP 37809 406 16 guttilatus guttilatus VBP 37809 406 17 _ _ NNP 37809 406 18 agree agree VBP 37809 406 19 in in IN 37809 406 20 color color NN 37809 406 21 pattern pattern NN 37809 406 22 . . . 37809 407 1 Schmidt Schmidt NNP 37809 407 2 and and CC 37809 407 3 Smith Smith NNP 37809 407 4 ( ( -LRB- 37809 407 5 1944 1944 CD 37809 407 6 ) ) -RRB- 37809 407 7 named name VBN 37809 407 8 _ _ NNP 37809 407 9 Syrrhophus Syrrhophus NNP 37809 407 10 gaigeae gaigeae NN 37809 407 11 _ _ NNP 37809 407 12 from from IN 37809 407 13 the the DT 37809 407 14 Chisos Chisos NNP 37809 407 15 Mountains Mountains NNPS 37809 407 16 of of IN 37809 407 17 the the DT 37809 407 18 Big Big NNP 37809 407 19 Bend Bend NNP 37809 407 20 region region NN 37809 407 21 of of IN 37809 407 22 Texas Texas NNP 37809 407 23 and and CC 37809 407 24 compared compare VBD 37809 407 25 it -PRON- PRP 37809 407 26 only only RB 37809 407 27 with with IN 37809 407 28 _ _ NNP 37809 407 29 S. S. NNP 37809 408 1 marnockii marnockii NNP 37809 408 2 _ _ NNP 37809 408 3 . . . 37809 409 1 Milstead Milstead NNP 37809 409 2 , , , 37809 409 3 Mecham Mecham NNP 37809 409 4 and and CC 37809 409 5 McClintock McClintock NNP 37809 409 6 ( ( -LRB- 37809 409 7 1950 1950 CD 37809 409 8 ) ) -RRB- 37809 409 9 synonymized synonymize VBD 37809 409 10 _ _ NNP 37809 409 11 gaigeae gaigeae NN 37809 409 12 _ _ NNP 37809 409 13 and and CC 37809 409 14 _ _ NNP 37809 409 15 marnockii marnockii NNS 37809 409 16 _ _ NNP 37809 409 17 because because IN 37809 409 18 they -PRON- PRP 37809 409 19 were be VBD 37809 409 20 unable unable JJ 37809 409 21 to to TO 37809 409 22 verify verify VB 37809 409 23 the the DT 37809 409 24 characters character NNS 37809 409 25 Wright Wright NNP 37809 409 26 and and CC 37809 409 27 Wright Wright NNP 37809 409 28 ( ( -LRB- 37809 409 29 1949 1949 CD 37809 409 30 ) ) -RRB- 37809 409 31 used use VBD 37809 409 32 to to TO 37809 409 33 separate separate VB 37809 409 34 them -PRON- PRP 37809 409 35 . . . 37809 410 1 Specimens specimen NNS 37809 410 2 from from IN 37809 410 3 the the DT 37809 410 4 Big Big NNP 37809 410 5 Bend Bend NNP 37809 410 6 region region NN 37809 410 7 differ differ VBP 37809 410 8 from from IN 37809 410 9 those those DT 37809 410 10 of of IN 37809 410 11 the the DT 37809 410 12 Edward Edward NNP 37809 410 13 and and CC 37809 410 14 Stockton Stockton NNP 37809 410 15 Plateaus Plateaus NNP 37809 410 16 in in IN 37809 410 17 having have VBG 37809 410 18 a a DT 37809 410 19 vermiculate vermiculate JJ 37809 410 20 pattern pattern NN 37809 410 21 , , , 37809 410 22 an an DT 37809 410 23 interorbital interorbital JJ 37809 410 24 bar bar NN 37809 410 25 , , , 37809 410 26 and and CC 37809 410 27 a a DT 37809 410 28 supratympanic supratympanic JJ 37809 410 29 stripe stripe NN 37809 410 30 . . . 37809 411 1 In in IN 37809 411 2 these these DT 37809 411 3 respects respect NNS 37809 411 4 they -PRON- PRP 37809 411 5 agree agree VBP 37809 411 6 with with IN 37809 411 7 specimens specimen NNS 37809 411 8 from from IN 37809 411 9 northern northern JJ 37809 411 10 México México NNP 37809 411 11 . . . 37809 412 1 Based base VBN 37809 412 2 on on IN 37809 412 3 limited limited JJ 37809 412 4 observations observation NNS 37809 412 5 , , , 37809 412 6 the the DT 37809 412 7 Mexican mexican JJ 37809 412 8 population population NN 37809 412 9 is be VBZ 37809 412 10 yellowish yellowish JJ 37809 412 11 to to TO 37809 412 12 brownish brownish VB 37809 412 13 in in IN 37809 412 14 life life NN 37809 412 15 whereas whereas IN 37809 412 16 the the DT 37809 412 17 central central JJ 37809 412 18 Texas Texas NNP 37809 412 19 population population NN 37809 412 20 is be VBZ 37809 412 21 green green JJ 37809 412 22 in in IN 37809 412 23 life life NN 37809 412 24 . . . 37809 413 1 Lacking lack VBG 37809 413 2 evidence evidence NN 37809 413 3 of of IN 37809 413 4 genetic genetic JJ 37809 413 5 exchange exchange NN 37809 413 6 , , , 37809 413 7 the the DT 37809 413 8 two two CD 37809 413 9 are be VBP 37809 413 10 held hold VBN 37809 413 11 to to TO 37809 413 12 be be VB 37809 413 13 specifically specifically RB 37809 413 14 distinct distinct JJ 37809 413 15 . . . 37809 414 1 Nearly nearly RB 37809 414 2 every every DT 37809 414 3 specimen speciman NNS 37809 414 4 examined examine VBN 37809 414 5 was be VBD 37809 414 6 infested infest VBN 37809 414 7 with with IN 37809 414 8 chiggers chigger NNS 37809 414 9 of of IN 37809 414 10 the the DT 37809 414 11 genus genus NN 37809 414 12 _ _ NNP 37809 414 13 Hannemania Hannemania NNP 37809 414 14 _ _ NNP 37809 414 15 . . . 37809 415 1 The the DT 37809 415 2 greatest great JJS 37809 415 3 concentrations concentration NNS 37809 415 4 are be VBP 37809 415 5 on on IN 37809 415 6 the the DT 37809 415 7 venter venter NN 37809 415 8 , , , 37809 415 9 in in IN 37809 415 10 the the DT 37809 415 11 groin groin NN 37809 415 12 , , , 37809 415 13 and and CC 37809 415 14 on on IN 37809 415 15 the the DT 37809 415 16 thighs thigh NNS 37809 415 17 . . . 37809 416 1 Many many JJ 37809 416 2 specimens specimen NNS 37809 416 3 have have VBP 37809 416 4 chiggers chigger NNS 37809 416 5 on on IN 37809 416 6 the the DT 37809 416 7 digits digit NNS 37809 416 8 and and CC 37809 416 9 tarsi tarsi NNP 37809 416 10 . . . 37809 417 1 The the DT 37809 417 2 same same JJ 37809 417 3 , , , 37809 417 4 or or CC 37809 417 5 a a DT 37809 417 6 related related JJ 37809 417 7 , , , 37809 417 8 chigger chigger NNP 37809 417 9 was be VBD 37809 417 10 found find VBN 37809 417 11 on on IN 37809 417 12 many many JJ 37809 417 13 specimens specimen NNS 37809 417 14 of of IN 37809 417 15 _ _ NNP 37809 417 16 Syrrhophus Syrrhophus NNP 37809 417 17 marnockii marnockii NNS 37809 417 18 _ _ NNP 37809 417 19 and and CC 37809 417 20 a a DT 37809 417 21 few few JJ 37809 417 22 _ _ NNP 37809 417 23 S. S. NNP 37809 418 1 verrucipes verrucipes NNP 37809 418 2 _ _ NNP 37809 418 3 , , , 37809 418 4 but but CC 37809 418 5 on on IN 37809 418 6 no no DT 37809 418 7 other other JJ 37809 418 8 species specie NNS 37809 418 9 of of IN 37809 418 10 the the DT 37809 418 11 genus genus NN 37809 418 12 . . . 37809 419 1 Mr. Mr. NNP 37809 419 2 Willy Willy NNP 37809 419 3 Wrenn Wrenn NNP 37809 419 4 told tell VBD 37809 419 5 me -PRON- PRP 37809 419 6 that that IN 37809 419 7 he -PRON- PRP 37809 419 8 has have VBZ 37809 419 9 seen see VBN 37809 419 10 heavy heavy JJ 37809 419 11 infestations infestation NNS 37809 419 12 of of IN 37809 419 13 _ _ NNP 37809 419 14 Hannemania Hannemania NNP 37809 419 15 _ _ NNP 37809 419 16 on on IN 37809 419 17 _ _ NNP 37809 419 18 Syrrhophus Syrrhophus NNP 37809 419 19 pallidus pallidus NNP 37809 419 20 _ _ NNP 37809 419 21 . . . 37809 420 1 Infestation infestation NN 37809 420 2 by by IN 37809 420 3 _ _ NNP 37809 420 4 Hannemania Hannemania NNP 37809 420 5 _ _ NNP 37809 420 6 probably probably RB 37809 420 7 reflects reflect VBZ 37809 420 8 similar similar JJ 37809 420 9 ecologies ecology NNS 37809 420 10 rather rather RB 37809 420 11 than than IN 37809 420 12 close close JJ 37809 420 13 relationships relationship NNS 37809 420 14 . . . 37809 421 1 [ [ -LRB- 37809 421 2 Illustration illustration NN 37809 421 3 : : : 37809 421 4 FIG FIG NNP 37809 421 5 . . . 37809 422 1 10 10 LS 37809 422 2 : : : 37809 422 3 Distribution distribution NN 37809 422 4 of of IN 37809 422 5 _ _ NNP 37809 422 6 Syrrhophus Syrrhophus NNP 37809 422 7 guttilatus guttilatus NN 37809 422 8 _ _ NNP 37809 422 9 . . . 37809 422 10 ] ] -RRB- 37809 423 1 _ _ NNP 37809 423 2 Etymology._--Latin Etymology._--Latin NNP 37809 423 3 , , , 37809 423 4 _ _ NNP 37809 423 5 guttula guttula NNP 37809 423 6 _ _ NNP 37809 423 7 , , , 37809 423 8 meaning mean VBG 37809 423 9 spotting spot VBG 37809 423 10 or or CC 37809 423 11 flecking fleck VBG 37809 423 12 , , , 37809 423 13 in in IN 37809 423 14 reference reference NN 37809 423 15 to to IN 37809 423 16 the the DT 37809 423 17 color color NN 37809 423 18 pattern pattern NN 37809 423 19 . . . 37809 424 1 _ _ NNP 37809 424 2 Distribution._--Moderate distribution._--moderate CD 37809 424 3 to to TO 37809 424 4 intermediate intermediate VB 37809 424 5 elevations elevation NNS 37809 424 6 ( ( -LRB- 37809 424 7 600 600 CD 37809 424 8 to to TO 37809 424 9 2000 2000 CD 37809 424 10 m. m. NN 37809 424 11 ) ) -RRB- 37809 424 12 along along IN 37809 424 13 the the DT 37809 424 14 Sierra Sierra NNP 37809 424 15 Madre Madre NNP 37809 424 16 Oriental Oriental NNP 37809 424 17 from from IN 37809 424 18 the the DT 37809 424 19 Big Big NNP 37809 424 20 Bend Bend NNP 37809 424 21 Region Region NNP 37809 424 22 of of IN 37809 424 23 Texas Texas NNP 37809 424 24 to to TO 37809 424 25 Guanajuato guanajuato VB 37809 424 26 , , , 37809 424 27 México México NNP 37809 424 28 ( ( -LRB- 37809 424 29 Fig Fig NNP 37809 424 30 . . . 37809 425 1 10 10 LS 37809 425 2 ) ) -RRB- 37809 425 3 . . . 37809 426 1 _ _ NNP 37809 426 2 Specimens Specimens NNP 37809 426 3 examined._--(32 examined._--(32 NNP 37809 426 4 ) ) -RRB- 37809 426 5 TEXAS TEXAS NNP 37809 426 6 , , , 37809 426 7 _ _ NNP 37809 426 8 Brewster Brewster NNP 37809 426 9 Co. Co. NNP 37809 426 10 _ _ NNP 37809 426 11 : : : 37809 426 12 Juniper Juniper NNP 37809 426 13 Canyon Canyon NNP 37809 426 14 , , , 37809 426 15 Chisos Chisos NNP 37809 426 16 Mts Mts NNP 37809 426 17 . . NNP 37809 426 18 , , , 37809 426 19 FMNH FMNH NNP 37809 426 20 27361 27361 CD 37809 426 21 ( ( -LRB- 37809 426 22 holotype holotype NN 37809 426 23 of of IN 37809 426 24 _ _ NNP 37809 426 25 S. S. NNP 37809 427 1 gaigeae gaigeae NNP 37809 427 2 _ _ NNP 37809 427 3 ) ) -RRB- 37809 427 4 , , , 37809 427 5 27360 27360 CD 37809 427 6 , , , 37809 427 7 27362 27362 CD 37809 427 8 - - SYM 37809 427 9 63 63 CD 37809 427 10 , , , 37809 427 11 MCZ MCZ NNP 37809 427 12 15346 15346 CD 37809 427 13 , , , 37809 427 14 27801 27801 CD 37809 427 15 , , , 37809 427 16 UMMZ UMMZ NNP 37809 427 17 66080 66080 CD 37809 427 18 , , , 37809 427 19 66082 66082 CD 37809 427 20 , , , 37809 427 21 66085 66085 CD 37809 427 22 - - SYM 37809 427 23 91 91 CD 37809 427 24 , , , 37809 427 25 USNM USNM NNP 37809 427 26 76876 76876 CD 37809 427 27 ; ; : 37809 427 28 Upper Upper NNP 37809 427 29 Green Green NNP 37809 427 30 Gulch Gulch NNP 37809 427 31 , , , 37809 427 32 TCWC TCWC NNP 37809 427 33 15943 15943 CD 37809 427 34 . . . 37809 428 1 MÉXICO MÉXICO NNP 37809 428 2 : : : 37809 428 3 _ _ NNP 37809 428 4 Coahuila Coahuila NNP 37809 428 5 _ _ NNP 37809 428 6 : : : 37809 428 7 8 8 CD 37809 428 8 km km NN 37809 428 9 . . . 37809 429 1 S S NNP 37809 429 2 Saltillo Saltillo NNP 37809 429 3 , , , 37809 429 4 UIMNH UIMNH NNP 37809 429 5 55518 55518 CD 37809 429 6 - - SYM 37809 429 7 21 21 CD 37809 429 8 . . . 37809 430 1 _ _ NNP 37809 430 2 Guanajuato Guanajuato NNP 37809 430 3 _ _ NNP 37809 430 4 : : : 37809 430 5 Guanajuato Guanajuato NNP 37809 430 6 , , , 37809 430 7 USNM USNM NNP 37809 430 8 9888 9888 CD 37809 430 9 ( ( -LRB- 37809 430 10 holotype holotype NN 37809 430 11 of of IN 37809 430 12 _ _ NNP 37809 430 13 Malachulodes Malachulodes NNPS 37809 430 14 guttilatus guttilatus NN 37809 430 15 _ _ NNP 37809 430 16 ) ) -RRB- 37809 430 17 ; ; : 37809 430 18 8 8 CD 37809 430 19 km km NN 37809 430 20 . . . 37809 431 1 E E NNP 37809 431 2 Guanajuato Guanajuato NNP 37809 431 3 , , , 37809 431 4 AMNH AMNH NNP 37809 431 5 73425 73425 CD 37809 431 6 ; ; : 37809 431 7 Cerro Cerro NNP 37809 431 8 Cubilete Cubilete NNP 37809 431 9 , , , 37809 431 10 AMNH AMNH NNP 37809 431 11 73424 73424 CD 37809 431 12 . . . 37809 432 1 _ _ NNP 37809 432 2 Nuevo Nuevo NNP 37809 432 3 León León NNP 37809 432 4 _ _ NNP 37809 432 5 : : : 37809 432 6 3 3 CD 37809 432 7 km km NN 37809 432 8 . . . 37809 433 1 S S NNP 37809 433 2 Galeana Galeana NNP 37809 433 3 , , , 37809 433 4 JDL JDL NNP 37809 433 5 1215 1215 CD 37809 433 6 ( ( -LRB- 37809 433 7 skeleton skeleton NNP 37809 433 8 ) ) -RRB- 37809 433 9 , , , 37809 433 10 UIMNH UIMNH NNP 37809 433 11 58204 58204 CD 37809 433 12 ; ; : 37809 433 13 24 24 CD 37809 433 14 km km NN 37809 433 15 . . . 37809 434 1 SW SW NNP 37809 434 2 Galeana Galeana NNP 37809 434 3 . . . 37809 435 1 1575 1575 CD 37809 435 2 m. m. NN 37809 435 3 , , , 37809 435 4 USNM USNM NNP 37809 435 5 108594 108594 CD 37809 435 6 ( ( -LRB- 37809 435 7 holotype holotype NN 37809 435 8 of of IN 37809 435 9 _ _ NNP 37809 435 10 Syrrhophus Syrrhophus NNP 37809 435 11 smithi smithi NN 37809 435 12 _ _ NNP 37809 435 13 ) ) -RRB- 37809 435 14 . . . 37809 436 1 _ _ NNP 37809 436 2 San San NNP 37809 436 3 Luis Luis NNP 37809 436 4 Potosí Potosí NNP 37809 436 5 _ _ NNP 37809 436 6 : : : 37809 436 7 5 5 CD 37809 436 8 km km NN 37809 436 9 . . . 37809 437 1 SW SW NNP 37809 437 2 San San NNP 37809 437 3 Luis Luis NNP 37809 437 4 Potosí Potosí NNP 37809 437 5 , , , 37809 437 6 UIMNH UIMNH NNP 37809 437 7 7807 7807 CD 37809 437 8 ( ( -LRB- 37809 437 9 holotype holotype NN 37809 437 10 of of IN 37809 437 11 _ _ NNP 37809 437 12 S. S. NNP 37809 438 1 petrophilus petrophilus NNP 37809 438 2 _ _ NNP 37809 438 3 ) ) -RRB- 37809 438 4 . . . 37809 439 1 _ _ NNP 37809 439 2 Tamaulipas Tamaulipas NNP 37809 439 3 _ _ NNP 37809 439 4 : : : 37809 439 5 1.6 1.6 CD 37809 439 6 km km NNS 37809 439 7 . . . 37809 440 1 NW NW NNP 37809 440 2 La La NNP 37809 440 3 Joya Joya NNP 37809 440 4 de de FW 37809 440 5 Salas Salas NNP 37809 440 6 , , , 37809 440 7 1530 1530 CD 37809 440 8 m. m. NN 37809 440 9 , , , 37809 440 10 UMMZ UMMZ NNP 37809 440 11 110736 110736 CD 37809 440 12 ( ( -LRB- 37809 440 13 4 4 CD 37809 440 14 ) ) -RRB- 37809 440 15 . . . 37809 441 1 = = NFP 37809 441 2 Syrrhophus Syrrhophus NNP 37809 441 3 marnockii= marnockii= NN 37809 441 4 Cope Cope NNP 37809 441 5 _ _ NNP 37809 441 6 Syrrhophus Syrrhophus NNP 37809 441 7 marnockii marnockii NNS 37809 441 8 _ _ NNP 37809 441 9 Cope Cope NNP 37809 441 10 , , , 37809 441 11 1878:253 1878:253 CD 37809 441 12 [ [ -LRB- 37809 441 13 Syntypes.--ANSP Syntypes.--ANSP NNP 37809 441 14 10765 10765 CD 37809 441 15 - - SYM 37809 441 16 68 68 CD 37809 441 17 , , , 37809 441 18 from from IN 37809 441 19 " " `` 37809 441 20 near near IN 37809 441 21 San San NNP 37809 441 22 Antonio Antonio NNP 37809 441 23 , , , 37809 441 24 " " `` 37809 441 25 Bexar Bexar NNP 37809 441 26 Co. Co. NNP 37809 441 27 , , , 37809 441 28 Texas Texas NNP 37809 441 29 ; ; : 37809 441 30 collected collect VBN 37809 441 31 by by IN 37809 441 32 G. G. NNP 37809 441 33 W. W. NNP 37809 441 34 Marnock Marnock NNP 37809 441 35 ] ] -RRB- 37809 441 36 . . . 37809 442 1 _ _ NNP 37809 442 2 Syrrhophus Syrrhophus NNP 37809 442 3 marnocki marnocki NN 37809 442 4 _ _ NNP 37809 442 5 : : : 37809 442 6 Yarrow Yarrow NNP 37809 442 7 , , , 37809 442 8 1882:24 1882:24 CD 37809 442 9 , , , 37809 442 10 193 193 CD 37809 442 11 . . . 37809 443 1 Milstead Milstead NNP 37809 443 2 , , , 37809 443 3 Mecham Mecham NNP 37809 443 4 , , , 37809 443 5 and and CC 37809 443 6 McClintock McClintock NNP 37809 443 7 , , , 37809 443 8 1950:550 1950:550 CD 37809 443 9 . . . 37809 444 1 _ _ NNP 37809 444 2 Diagnosis._--Medium Diagnosis._--Medium NNP 37809 444 3 - - HYPH 37809 444 4 sized sized JJ 37809 444 5 frogs frog NNS 37809 444 6 , , , 37809 444 7 males male VBZ 37809 444 8 18.4 18.4 CD 37809 444 9 - - SYM 37809 444 10 28.9 28.9 CD 37809 444 11 mm mm NN 37809 444 12 . . . 37809 445 1 snout snout NN 37809 445 2 - - HYPH 37809 445 3 vent vent NN 37809 445 4 , , , 37809 445 5 females female VBZ 37809 445 6 20.4 20.4 CD 37809 445 7 - - SYM 37809 445 8 35.4 35.4 CD 37809 445 9 mm mm NN 37809 445 10 . . . 37809 446 1 snout snout NN 37809 446 2 - - HYPH 37809 446 3 vent vent NN 37809 446 4 length length NN 37809 446 5 ; ; : 37809 446 6 vocal vocal NN 37809 446 7 slits slit NNS 37809 446 8 in in IN 37809 446 9 males male NNS 37809 446 10 ; ; , 37809 446 11 digital digital JJ 37809 446 12 tips tip NNS 37809 446 13 widened widen VBD 37809 446 14 ( ( -LRB- 37809 446 15 Fig fig NN 37809 446 16 . . . 37809 447 1 1 1 LS 37809 447 2 ) ) -RRB- 37809 447 3 ; ; : 37809 447 4 first first JJ 37809 447 5 and and CC 37809 447 6 second second JJ 37809 447 7 fingers finger NNS 37809 447 8 equal equal JJ 37809 447 9 ; ; : 37809 447 10 skin skin NN 37809 447 11 of of IN 37809 447 12 dorsum dorsum NN 37809 447 13 smooth smooth JJ 37809 447 14 to to IN 37809 447 15 weakly weakly NNP 37809 447 16 pustular pustular NNP 37809 447 17 , , , 37809 447 18 that that DT 37809 447 19 of of IN 37809 447 20 venter venter NN 37809 447 21 smooth smooth JJ 37809 447 22 ; ; : 37809 447 23 snout snout NNP 37809 447 24 blunt blunt NNP 37809 447 25 , , , 37809 447 26 rounded round VBD 37809 447 27 ; ; : 37809 447 28 diameter diameter NN 37809 447 29 of of IN 37809 447 30 tympanum tympanum NNP 37809 447 31 47.2 47.2 CD 37809 447 32 - - SYM 37809 447 33 68.3 68.3 CD 37809 447 34 per per IN 37809 447 35 cent cent NN 37809 447 36 that that DT 37809 447 37 of of IN 37809 447 38 eye eye NN 37809 447 39 in in IN 37809 447 40 males male NNS 37809 447 41 , , , 37809 447 42 45.8 45.8 CD 37809 447 43 - - SYM 37809 447 44 73.3 73.3 CD 37809 447 45 in in IN 37809 447 46 females female NNS 37809 447 47 ; ; : 37809 447 48 dorsum dorsum NN 37809 447 49 tan tan NN 37809 447 50 to to IN 37809 447 51 light light NN 37809 447 52 brown brown NNP 37809 447 53 in in IN 37809 447 54 preservative preservative NN 37809 447 55 with with IN 37809 447 56 rusty rusty JJ 37809 447 57 - - HYPH 37809 447 58 brown brown JJ 37809 447 59 flecks fleck NNS 37809 447 60 , , , 37809 447 61 venter venter NN 37809 447 62 white white JJ 37809 447 63 ; ; : 37809 447 64 ground ground NN 37809 447 65 color color NN 37809 447 66 green green JJ 37809 447 67 in in IN 37809 447 68 life life NN 37809 447 69 ; ; : 37809 447 70 thighs thigh NNS 37809 447 71 banded band VBD 37809 447 72 ; ; : 37809 447 73 interorbital interorbital NNP 37809 447 74 bar bar NN 37809 447 75 absent absent NN 37809 447 76 . . . 37809 448 1 _ _ NNP 37809 448 2 Remarks._--Specimens Remarks._--Specimens NNPS 37809 448 3 from from IN 37809 448 4 the the DT 37809 448 5 southern southern JJ 37809 448 6 edge edge NN 37809 448 7 of of IN 37809 448 8 the the DT 37809 448 9 Edwards Edwards NNP 37809 448 10 Plateau Plateau NNP 37809 448 11 and and CC 37809 448 12 the the DT 37809 448 13 eastern eastern JJ 37809 448 14 edge edge NN 37809 448 15 of of IN 37809 448 16 the the DT 37809 448 17 Stockton Stockton NNP 37809 448 18 Plateau Plateau NNP 37809 448 19 have have VBP 37809 448 20 larger large JJR 37809 448 21 flecks fleck NNS 37809 448 22 on on IN 37809 448 23 the the DT 37809 448 24 back back NN 37809 448 25 that that WDT 37809 448 26 tend tend VBP 37809 448 27 to to TO 37809 448 28 form form VB 37809 448 29 a a DT 37809 448 30 vermiculate vermiculate JJ 37809 448 31 pattern pattern NN 37809 448 32 like like IN 37809 448 33 that that DT 37809 448 34 of of IN 37809 448 35 _ _ NNP 37809 448 36 S. S. NNP 37809 449 1 guttilatus guttilatus NNP 37809 449 2 _ _ NNP 37809 449 3 . . . 37809 450 1 The the DT 37809 450 2 vermiculation vermiculation NN 37809 450 3 is be VBZ 37809 450 4 never never RB 37809 450 5 well well RB 37809 450 6 developed developed JJ 37809 450 7 ( ( -LRB- 37809 450 8 see see VB 37809 450 9 plate plate NN 37809 450 10 38 38 CD 37809 450 11 in in IN 37809 450 12 Conant Conant NNP 37809 450 13 , , , 37809 450 14 1958 1958 CD 37809 450 15 ) ) -RRB- 37809 450 16 . . . 37809 451 1 Most Most JJS 37809 451 2 of of IN 37809 451 3 the the DT 37809 451 4 specimens specimen NNS 37809 451 5 from from IN 37809 451 6 the the DT 37809 451 7 Edwards Edwards NNP 37809 451 8 Plateau Plateau NNP 37809 451 9 have have VBP 37809 451 10 a a DT 37809 451 11 punctate punctate NN 37809 451 12 pattern pattern NN 37809 451 13 ( ( -LRB- 37809 451 14 Fig fig NN 37809 451 15 . . . 37809 452 1 9 9 LS 37809 452 2 ) ) -RRB- 37809 452 3 . . . 37809 453 1 Fossils fossil NNS 37809 453 2 are be VBP 37809 453 3 known know VBN 37809 453 4 from from IN 37809 453 5 the the DT 37809 453 6 Sangamon Sangamon NNP 37809 453 7 interglacial interglacial JJ 37809 453 8 deposits deposit NNS 37809 453 9 in in IN 37809 453 10 Foard Foard NNP 37809 453 11 and and CC 37809 453 12 Knox Knox NNP 37809 453 13 Counties Counties NNPS 37809 453 14 , , , 37809 453 15 Texas Texas NNP 37809 453 16 ( ( -LRB- 37809 453 17 Lynch Lynch NNP 37809 453 18 , , , 37809 453 19 1964 1964 CD 37809 453 20 ; ; : 37809 453 21 Tihen Tihen NNP 37809 453 22 , , , 37809 453 23 1960 1960 CD 37809 453 24 ) ) -RRB- 37809 453 25 . . . 37809 454 1 _ _ NNP 37809 454 2 Etymology._--A Etymology._--A NNP 37809 454 3 patronym patronym NN 37809 454 4 for for IN 37809 454 5 the the DT 37809 454 6 collector collector NN 37809 454 7 of of IN 37809 454 8 the the DT 37809 454 9 type type NN 37809 454 10 specimens specimen NNS 37809 454 11 . . . 37809 455 1 _ _ NNP 37809 455 2 Distribution._--The Distribution._--The NNP 37809 455 3 Edwards Edwards NNP 37809 455 4 Plateau Plateau NNP 37809 455 5 and and CC 37809 455 6 the the DT 37809 455 7 extreme extreme JJ 37809 455 8 eastern eastern JJ 37809 455 9 edge edge NN 37809 455 10 of of IN 37809 455 11 the the DT 37809 455 12 Stockton Stockton NNP 37809 455 13 Plateau Plateau NNP 37809 455 14 in in IN 37809 455 15 Texas Texas NNP 37809 455 16 ( ( -LRB- 37809 455 17 Fig Fig NNP 37809 455 18 . . . 37809 456 1 11 11 CD 37809 456 2 ) ) -RRB- 37809 456 3 . . . 37809 457 1 The the DT 37809 457 2 fossil fossil NN 37809 457 3 records record NNS 37809 457 4 lie lie VBP 37809 457 5 some some DT 37809 457 6 200 200 CD 37809 457 7 miles mile NNS 37809 457 8 to to IN 37809 457 9 the the DT 37809 457 10 north north NN 37809 457 11 . . . 37809 458 1 Two two CD 37809 458 2 specimens specimen NNS 37809 458 3 ( ( -LRB- 37809 458 4 FMNH FMNH NNP 37809 458 5 103216 103216 CD 37809 458 6 - - SYM 37809 458 7 17 17 CD 37809 458 8 ) ) -RRB- 37809 458 9 from from IN 37809 458 10 Brownsville Brownsville NNP 37809 458 11 , , , 37809 458 12 Cameron Cameron NNP 37809 458 13 Co. Co. NNP 37809 458 14 , , , 37809 458 15 Texas Texas NNP 37809 458 16 , , , 37809 458 17 were be VBD 37809 458 18 formerly formerly RB 37809 458 19 in in IN 37809 458 20 the the DT 37809 458 21 EHT EHT NNP 37809 458 22 - - HYPH 37809 458 23 HMS HMS NNP 37809 458 24 collection collection NN 37809 458 25 ( ( -LRB- 37809 458 26 nos nos NNP 37809 458 27 . . . 37809 459 1 31348 31348 CD 37809 459 2 - - SYM 37809 459 3 49 49 CD 37809 459 4 ) ) -RRB- 37809 459 5 . . . 37809 460 1 Data datum NNS 37809 460 2 given give VBN 37809 460 3 in in IN 37809 460 4 Taylor Taylor NNP 37809 460 5 's 's POS 37809 460 6 field field NN 37809 460 7 catalogue catalogue NN 37809 460 8 ( ( -LRB- 37809 460 9 housed house VBN 37809 460 10 in in IN 37809 460 11 the the DT 37809 460 12 Division Division NNP 37809 460 13 of of IN 37809 460 14 Reptiles Reptiles NNP 37809 460 15 , , , 37809 460 16 Field Field NNP 37809 460 17 Museum Museum NNP 37809 460 18 ) ) -RRB- 37809 460 19 are be VBP 37809 460 20 " " `` 37809 460 21 Brownsville Brownsville NNP 37809 460 22 , , , 37809 460 23 A. A. NNP 37809 460 24 J. J. NNP 37809 460 25 Kirn Kirn NNP 37809 460 26 collector collector NN 37809 460 27 , , , 37809 460 28 April April NNP 37809 460 29 15 15 CD 37809 460 30 , , , 37809 460 31 1934 1934 CD 37809 460 32 . . . 37809 460 33 " " '' 37809 461 1 Until until IN 37809 461 2 verification verification NN 37809 461 3 by by IN 37809 461 4 recently recently RB 37809 461 5 collected collect VBN 37809 461 6 material material NN 37809 461 7 is be VBZ 37809 461 8 available available JJ 37809 461 9 , , , 37809 461 10 this this DT 37809 461 11 record record NN 37809 461 12 must must MD 37809 461 13 be be VB 37809 461 14 disregarded disregard VBN 37809 461 15 . . . 37809 462 1 _ _ NNP 37809 462 2 Specimens Specimens NNP 37809 462 3 examined._--(103 examined._--(103 NNP 37809 462 4 ) ) -RRB- 37809 462 5 TEXAS TEXAS NNP 37809 462 6 , , , 37809 462 7 _ _ NNP 37809 462 8 Bandera Bandera NNP 37809 462 9 Co. Co. NNP 37809 462 10 _ _ NNP 37809 462 11 : : : 37809 462 12 10 10 CD 37809 462 13 mi mi JJ 37809 462 14 . . . 37809 463 1 SW SW NNP 37809 463 2 Medina Medina NNP 37809 463 3 , , , 37809 463 4 TCWC TCWC NNP 37809 463 5 13508 13508 CD 37809 463 6 - - SYM 37809 463 7 10 10 CD 37809 463 8 ; ; : 37809 463 9 8 8 CD 37809 463 10 mi mi NNS 37809 463 11 . . . 37809 464 1 W W NNP 37809 464 2 Medina Medina NNP 37809 464 3 , , , 37809 464 4 KU KU NNP 37809 464 5 60243 60243 CD 37809 464 6 ; ; : 37809 464 7 13 13 CD 37809 464 8 mi mi JJ 37809 464 9 . . . 37809 465 1 W W NNP 37809 465 2 Medina Medina NNP 37809 465 3 , , , 37809 465 4 KU KU NNP 37809 465 5 60242 60242 CD 37809 465 6 , , , 37809 465 7 TCWC TCWC NNP 37809 465 8 13506 13506 CD 37809 465 9 - - SYM 37809 465 10 07 07 CD 37809 465 11 . . . 37809 466 1 _ _ NNP 37809 466 2 Bexar Bexar NNP 37809 466 3 Co. Co. NNP 37809 466 4 _ _ XX 37809 466 5 : : : 37809 466 6 UIMNH UIMNH NNP 37809 466 7 34694 34694 CD 37809 466 8 ; ; : 37809 466 9 Classen Classen NNP 37809 466 10 ranch ranch NN 37809 466 11 , , , 37809 466 12 near near IN 37809 466 13 San San NNP 37809 466 14 Antonio Antonio NNP 37809 466 15 . . . 37809 467 1 UMMZ UMMZ NNP 37809 467 2 98891 98891 CD 37809 467 3 ; ; : 37809 467 4 Helotes Helotes NNP 37809 467 5 , , , 37809 467 6 EAL EAL NNP 37809 467 7 1560 1560 CD 37809 467 8 , , , 37809 467 9 MCZ MCZ NNP 37809 467 10 11837 11837 CD 37809 467 11 ( ( -LRB- 37809 467 12 2 2 CD 37809 467 13 ) ) -RRB- 37809 467 14 , , , 37809 467 15 UMMZ UMMZ NNP 37809 467 16 64045 64045 CD 37809 467 17 , , , 37809 467 18 USNM USNM NNP 37809 467 19 13635 13635 CD 37809 467 20 ; ; : 37809 467 21 2 2 CD 37809 467 22 mi mi NNP 37809 467 23 . . . 37809 468 1 N N NNP 37809 468 2 Helotes Helotes NNP 37809 468 3 , , , 37809 468 4 TCWC TCWC VBD 37809 468 5 9234 9234 CD 37809 468 6 - - SYM 37809 468 7 35 35 CD 37809 468 8 ; ; : 37809 468 9 3.5 3.5 CD 37809 468 10 mi mi JJ 37809 468 11 . . . 37809 469 1 N N NNP 37809 469 2 Helotes Helotes NNP 37809 469 3 , , , 37809 469 4 LSUMZ LSUMZ NNP 37809 469 5 10363 10363 CD 37809 469 6 ; ; : 37809 469 7 8 8 CD 37809 469 8 mi mi NNS 37809 469 9 . . . 37809 470 1 N N NNP 37809 470 2 Helotes Helotes NNP 37809 470 3 , , , 37809 470 4 TCWC tcwc NN 37809 470 5 1549 1549 CD 37809 470 6 , , , 37809 470 7 4364 4364 CD 37809 470 8 ; ; : 37809 470 9 San San NNP 37809 470 10 Antonio Antonio NNP 37809 470 11 , , , 37809 470 12 FMNH FMNH NNP 37809 470 13 15553 15553 CD 37809 470 14 - - SYM 37809 470 15 56 56 CD 37809 470 16 , , , 37809 470 17 TCWC TCWC NNP 37809 470 18 13497 13497 CD 37809 470 19 - - SYM 37809 470 20 99 99 CD 37809 470 21 . . . 37809 471 1 _ _ NNP 37809 471 2 Blanco Blanco NNP 37809 471 3 Co. Co. NNP 37809 471 4 _ _ NNP 37809 471 5 : : : 37809 471 6 8 8 CD 37809 471 7 mi mi NNP 37809 471 8 . . . 37809 472 1 NE NE NNP 37809 472 2 Blanco Blanco NNP 37809 472 3 , , , 37809 472 4 TCWC TCWC NNP 37809 472 5 4782 4782 CD 37809 472 6 . . . 37809 473 1 _ _ NNP 37809 473 2 Comal Comal NNP 37809 473 3 Co. Co. NNP 37809 473 4 _ _ NNP 37809 473 5 : : : 37809 473 6 New New NNP 37809 473 7 Braunfels braunfel NNS 37809 473 8 , , , 37809 473 9 TCWC TCWC NNP 37809 473 10 13500 13500 CD 37809 473 11 - - SYM 37809 473 12 05 05 CD 37809 473 13 ; ; : 37809 473 14 5 5 CD 37809 473 15 mi mi JJ 37809 473 16 . . . 37809 474 1 NE NE NNP 37809 474 2 New New NNP 37809 474 3 Braunfels Braunfels NNP 37809 474 4 , , , 37809 474 5 UMMZ UMMZ NNP 37809 474 6 71016 71016 CD 37809 474 7 ( ( -LRB- 37809 474 8 10 10 CD 37809 474 9 ) ) -RRB- 37809 474 10 . . . 37809 475 1 _ _ NNP 37809 475 2 Hays Hays NNP 37809 475 3 Co. Co. NNP 37809 475 4 _ _ NNP 37809 475 5 : : : 37809 475 6 San San NNP 37809 475 7 Marcos Marcos NNP 37809 475 8 , , , 37809 475 9 AMNH AMNH NNP 37809 475 10 22661 22661 CD 37809 475 11 - - SYM 37809 475 12 64 64 CD 37809 475 13 , , , 37809 475 14 32700 32700 CD 37809 475 15 , , , 37809 475 16 FMNH FMNH NNP 37809 475 17 15245 15245 CD 37809 475 18 - - SYM 37809 475 19 46 46 CD 37809 475 20 , , , 37809 475 21 26250 26250 CD 37809 475 22 , , , 37809 475 23 26253 26253 CD 37809 475 24 - - SYM 37809 475 25 57 57 CD 37809 475 26 , , , 37809 475 27 37617 37617 CD 37809 475 28 , , , 37809 475 29 37665 37665 CD 37809 475 30 , , , 37809 475 31 MCZ MCZ NNP 37809 475 32 15649 15649 CD 37809 475 33 - - SYM 37809 475 34 50 50 CD 37809 475 35 , , , 37809 475 36 23268 23268 CD 37809 475 37 - - SYM 37809 475 38 69 69 CD 37809 475 39 ; ; : 37809 475 40 6 6 CD 37809 475 41 mi mi NNS 37809 475 42 . . . 37809 476 1 SW SW NNP 37809 476 2 San San NNP 37809 476 3 Marcos Marcos NNP 37809 476 4 , , , 37809 476 5 TCWC TCWC NNP 37809 476 6 5070 5070 CD 37809 476 7 - - SYM 37809 476 8 71 71 CD 37809 476 9 , , , 37809 476 10 7140 7140 CD 37809 476 11 , , , 37809 476 12 9232 9232 CD 37809 476 13 - - SYM 37809 476 14 33 33 CD 37809 476 15 , , , 37809 476 16 9236 9236 CD 37809 476 17 , , , 37809 476 18 9316 9316 CD 37809 476 19 - - SYM 37809 476 20 17 17 CD 37809 476 21 , , , 37809 476 22 9320 9320 CD 37809 476 23 . . . 37809 477 1 _ _ NNP 37809 477 2 Kendall Kendall NNP 37809 477 3 Co. Co. NNP 37809 477 4 _ _ NNP 37809 477 5 : : : 37809 477 6 11 11 CD 37809 477 7 mi mi NN 37809 477 8 . . . 37809 478 1 E E NNP 37809 478 2 Boerne Boerne NNP 37809 478 3 , , , 37809 478 4 AMNH AMNH NNP 37809 478 5 54660 54660 CD 37809 478 6 - - SYM 37809 478 7 61 61 CD 37809 478 8 , , , 37809 478 9 54662 54662 CD 37809 478 10 ( ( -LRB- 37809 478 11 2 2 CD 37809 478 12 ) ) -RRB- 37809 478 13 ; ; : 37809 478 14 10 10 CD 37809 478 15 mi mi JJ 37809 478 16 . . . 37809 479 1 W W NNP 37809 479 2 Boerne Boerne NNP 37809 479 3 , , , 37809 479 4 KU KU NNP 37809 479 5 18441 18441 CD 37809 479 6 ; ; : 37809 479 7 Kendalia Kendalia NNP 37809 479 8 , , , 37809 479 9 UIMNH UIMNH NNP 37809 479 10 21434 21434 CD 37809 479 11 . . . 37809 480 1 _ _ NNP 37809 480 2 Kerr Kerr NNP 37809 480 3 Co. Co. NNP 37809 480 4 _ _ NNP 37809 480 5 : : : 37809 480 6 Kerr Kerr NNP 37809 480 7 W. W. NNP 37809 480 8 M. M. NNP 37809 480 9 Area Area NNP 37809 480 10 , , , 37809 480 11 TCWC TCWC NNP 37809 480 12 15859 15859 CD 37809 480 13 ; ; : 37809 480 14 40 40 CD 37809 480 15 mi mi JJ 37809 480 16 . . . 37809 481 1 NW NW NNP 37809 481 2 Kerrville Kerrville NNP 37809 481 3 , , , 37809 481 4 TCWC TCWC NNP 37809 481 5 6555 6555 CD 37809 481 6 . . . 37809 482 1 _ _ NNP 37809 482 2 Medina Medina NNP 37809 482 3 Co. Co. NNP 37809 482 4 _ _ NNP 37809 482 5 : : : 37809 482 6 UIMNH UIMNH NNP 37809 482 7 13287 13287 CD 37809 482 8 - - SYM 37809 482 9 88 88 CD 37809 482 10 ; ; : 37809 482 11 12 12 CD 37809 482 12 mi mi NNS 37809 482 13 . . . 37809 483 1 N N NNP 37809 483 2 Castroville Castroville NNP 37809 483 3 , , , 37809 483 4 UIMNH UIMNH NNP 37809 483 5 21423 21423 CD 37809 483 6 ; ; : 37809 483 7 14 14 CD 37809 483 8 mi mi NNP 37809 483 9 . . . 37809 484 1 N N NNP 37809 484 2 Castroville Castroville NNP 37809 484 3 , , , 37809 484 4 UIMNH UIMNH NNP 37809 484 5 21424 21424 CD 37809 484 6 - - SYM 37809 484 7 25 25 CD 37809 484 8 ; ; : 37809 484 9 16 16 CD 37809 484 10 mi mi JJ 37809 484 11 . . . 37809 485 1 N N NNP 37809 485 2 Castroville Castroville NNP 37809 485 3 , , , 37809 485 4 UIMNH UIMNH NNP 37809 485 5 21421 21421 CD 37809 485 6 - - SYM 37809 485 7 22 22 CD 37809 485 8 ; ; : 37809 485 9 17 17 CD 37809 485 10 mi mi JJ 37809 485 11 . . . 37809 486 1 N N NNP 37809 486 2 Castroville Castroville NNP 37809 486 3 , , , 37809 486 4 UIMNH UIMNH NNP 37809 486 5 21428 21428 CD 37809 486 6 - - SYM 37809 486 7 29 29 CD 37809 486 8 ; ; : 37809 486 9 18 18 CD 37809 486 10 mi mi JJ 37809 486 11 . . . 37809 487 1 N N NNP 37809 487 2 Castroville Castroville NNP 37809 487 3 , , , 37809 487 4 UIMNH UIMNH NNP 37809 487 5 21426 21426 CD 37809 487 6 - - SYM 37809 487 7 27 27 CD 37809 487 8 , , , 37809 487 9 21430 21430 CD 37809 487 10 - - SYM 37809 487 11 33 33 CD 37809 487 12 ; ; : 37809 487 13 6.5 6.5 CD 37809 487 14 mi mi UH 37809 487 15 . . . 37809 488 1 NW NW NNP 37809 488 2 Rio Rio NNP 37809 488 3 Medina Medina NNP 37809 488 4 , , , 37809 488 5 KU KU NNP 37809 488 6 18440 18440 CD 37809 488 7 . . . 37809 489 1 _ _ NNP 37809 489 2 Real Real NNP 37809 489 3 Co. Co. NNP 37809 489 4 _ _ XX 37809 489 5 : : : 37809 489 6 Rio Rio NNP 37809 489 7 Frio Frio NNP 37809 489 8 , , , 37809 489 9 FMNH FMNH NNP 37809 489 10 55156 55156 CD 37809 489 11 - - SYM 37809 489 12 57 57 CD 37809 489 13 . . . 37809 490 1 _ _ NNP 37809 490 2 Travis Travis NNP 37809 490 3 Co. Co. NNP 37809 490 4 _ _ NNP 37809 490 5 : : : 37809 490 6 Austin Austin NNP 37809 490 7 , , , 37809 490 8 AMNH AMNH NNP 37809 490 9 44221 44221 CD 37809 490 10 - - SYM 37809 490 11 22 22 CD 37809 490 12 ; ; : 37809 490 13 Mount Mount NNP 37809 490 14 Bonnell Bonnell NNP 37809 490 15 , , , 37809 490 16 5 5 CD 37809 490 17 mi mi JJ 37809 490 18 . . . 37809 491 1 S S NNP 37809 491 2 Austin Austin NNP 37809 491 3 , , , 37809 491 4 UMMZ UMMZ NNP 37809 491 5 101453 101453 CD 37809 491 6 ( ( -LRB- 37809 491 7 10 10 CD 37809 491 8 ) ) -RRB- 37809 491 9 . . . 37809 492 1 _ _ NNP 37809 492 2 Uvalde Uvalde NNP 37809 492 3 Co. Co. NNP 37809 492 4 _ _ NNP 37809 492 5 : : : 37809 492 6 13 13 CD 37809 492 7 mi mi JJ 37809 492 8 . . . 37809 493 1 from from IN 37809 493 2 Uvalde Uvalde NNP 37809 493 3 , , , 37809 493 4 UIMNH UIMNH NNP 37809 493 5 62322 62322 CD 37809 493 6 . . . 37809 494 1 _ _ NNP 37809 494 2 Val Val NNP 37809 494 3 - - HYPH 37809 494 4 Verde Verde NNP 37809 494 5 Co. Co. NNP 37809 494 6 _ _ NNP 37809 494 7 : : : 37809 494 8 40 40 CD 37809 494 9 mi mi JJ 37809 494 10 . . . 37809 495 1 N N NNP 37809 495 2 Del Del NNP 37809 495 3 Rio Rio NNP 37809 495 4 , , , 37809 495 5 JDL JDL NNP 37809 495 6 214 214 CD 37809 495 7 ( ( -LRB- 37809 495 8 skeleton skeleton NNP 37809 495 9 ) ) -RRB- 37809 495 10 . . . 37809 496 1 [ [ -LRB- 37809 496 2 Illustration illustration NN 37809 496 3 : : : 37809 496 4 FIG FIG NNP 37809 496 5 . . . 37809 497 1 11 11 LS 37809 497 2 : : : 37809 497 3 Distribution distribution NN 37809 497 4 of of IN 37809 497 5 _ _ NNP 37809 497 6 Syrrhophus Syrrhophus NNP 37809 497 7 marnockii marnockii NNS 37809 497 8 _ _ NNP 37809 497 9 ( ( -LRB- 37809 497 10 circles circles NNP 37809 497 11 ) ) -RRB- 37809 497 12 . . . 37809 498 1 Starred starred JJ 37809 498 2 localities locality NNS 37809 498 3 are be VBP 37809 498 4 late late JJ 37809 498 5 Pleistocene Pleistocene NNP 37809 498 6 records record NNS 37809 498 7 . . . 37809 498 8 ] ] -RRB- 37809 499 1 = = NFP 37809 499 2 Syrrhophus Syrrhophus NNP 37809 499 3 verrucipes= verrucipes= NN 37809 499 4 Cope Cope NNP 37809 499 5 _ _ NNP 37809 499 6 Syrrhophus Syrrhophus NNP 37809 499 7 verrucipes verrucipe VBZ 37809 499 8 _ _ NNP 37809 499 9 Cope Cope NNP 37809 499 10 , , , 37809 499 11 1885:383 1885:383 CD 37809 499 12 [ [ -LRB- 37809 499 13 Holotype.--ANSP Holotype.--ANSP NNP 37809 499 14 11325 11325 CD 37809 499 15 , , , 37809 499 16 from from IN 37809 499 17 near near IN 37809 499 18 Zacualtipán Zacualtipán NNP 37809 499 19 , , , 37809 499 20 Hidalgo Hidalgo NNP 37809 499 21 , , , 37809 499 22 México México NNP 37809 499 23 ( ( -LRB- 37809 499 24 1800 1800 CD 37809 499 25 feet foot NNS 37809 499 26 lower low JJR 37809 499 27 in in IN 37809 499 28 a a DT 37809 499 29 rocky rocky JJ 37809 499 30 gorge gorge NN 37809 499 31 of of IN 37809 499 32 a a DT 37809 499 33 stream stream NN 37809 499 34 near near IN 37809 499 35 its -PRON- PRP$ 37809 499 36 junction junction NN 37809 499 37 with with IN 37809 499 38 the the DT 37809 499 39 Río Río NNP 37809 499 40 San San NNP 37809 499 41 Miguel Miguel NNP 37809 499 42 ) ) -RRB- 37809 499 43 , , , 37809 499 44 collected collect VBN 37809 499 45 by by IN 37809 499 46 Dr. Dr. NNP 37809 499 47 Santiago Santiago NNP 37809 499 48 Bernard Bernard NNP 37809 499 49 ] ] -RRB- 37809 499 50 . . . 37809 500 1 Kellogg Kellogg NNP 37809 500 2 , , , 37809 500 3 1932:126 1932:126 CD 37809 500 4 - - SYM 37809 500 5 29 29 CD 37809 500 6 . . . 37809 501 1 Smith Smith NNP 37809 501 2 and and CC 37809 501 3 Taylor Taylor NNP 37809 501 4 , , , 37809 501 5 1948:52 1948:52 CD 37809 501 6 - - SYM 37809 501 7 53 53 CD 37809 501 8 . . . 37809 502 1 Firschein Firschein NNP 37809 502 2 , , , 37809 502 3 1954:55 1954:55 CD 37809 502 4 - - SYM 37809 502 5 57 57 CD 37809 502 6 . . . 37809 503 1 Gorham Gorham NNP 37809 503 2 , , , 37809 503 3 1966:167 1966:167 CD 37809 503 4 . . . 37809 504 1 _ _ NNP 37809 504 2 Syrrhaphus Syrrhaphus NNP 37809 504 3 verrucipes verrucipe VBZ 37809 504 4 _ _ NNP 37809 504 5 : : : 37809 504 6 Günther Günther NNP 37809 504 7 , , , 37809 504 8 1900:216 1900:216 CD 37809 504 9 - - SYM 37809 504 10 17 17 CD 37809 504 11 . . . 37809 505 1 _ _ NNP 37809 505 2 Tomodactylus Tomodactylus NNP 37809 505 3 macrotympanum macrotympanum NN 37809 505 4 _ _ NNP 37809 505 5 Taylor Taylor NNP 37809 505 6 , , , 37809 505 7 1940e:496 1940e:496 CD 37809 505 8 - - SYM 37809 505 9 99 99 CD 37809 505 10 , , , 37809 505 11 pl pl NN 37809 505 12 . . . 37809 506 1 55 55 CD 37809 506 2 , , , 37809 506 3 figs fig NNS 37809 506 4 . . . 37809 507 1 2a 2a CD 37809 507 2 - - HYPH 37809 507 3 b b NN 37809 507 4 . . . 37809 508 1 [ [ -LRB- 37809 508 2 Holotype.--FMNH holotype.--fmnh NN 37809 508 3 100049 100049 CD 37809 508 4 ( ( -LRB- 37809 508 5 formerly formerly RB 37809 508 6 EHT EHT NNP 37809 508 7 - - HYPH 37809 508 8 HMS HMS NNP 37809 508 9 6838 6838 CD 37809 508 10 ) ) -RRB- 37809 508 11 , , , 37809 508 12 from from IN 37809 508 13 La La NNP 37809 508 14 Placita Placita NNP 37809 508 15 , , , 37809 508 16 8 8 CD 37809 508 17 km km NN 37809 508 18 . . . 37809 509 1 S S NNP 37809 509 2 Jacala Jacala NNP 37809 509 3 , , , 37809 509 4 Hidalgo Hidalgo NNP 37809 509 5 , , , 37809 509 6 México México NNP 37809 509 7 , , , 37809 509 8 1850 1850 CD 37809 509 9 m. m. NN 37809 509 10 ; ; , 37809 509 11 collected collect VBN 37809 509 12 on on IN 37809 509 13 July July NNP 37809 509 14 2 2 CD 37809 509 15 , , , 37809 509 16 1936 1936 CD 37809 509 17 , , , 37809 509 18 by by IN 37809 509 19 Edward Edward NNP 37809 509 20 H. H. NNP 37809 509 21 Taylor Taylor NNP 37809 509 22 ] ] -RRB- 37809 509 23 . . . 37809 510 1 Smith Smith NNP 37809 510 2 and and CC 37809 510 3 Taylor Taylor NNP 37809 510 4 , , , 37809 510 5 1948:47 1948:47 CD 37809 510 6 - - SYM 37809 510 7 48 48 CD 37809 510 8 . . . 37809 511 1 _ _ NNP 37809 511 2 Syrrhophus Syrrhophus NNP 37809 511 3 macrotympanum macrotympanum NNP 37809 511 4 _ _ NNP 37809 511 5 : : : 37809 511 6 Dixon Dixon NNP 37809 511 7 , , , 37809 511 8 1957:384 1957:384 CD 37809 511 9 . . . 37809 512 1 Gorham Gorham NNP 37809 512 2 , , , 37809 512 3 1966:165 1966:165 CD 37809 512 4 . . . 37809 513 1 _ _ NNP 37809 513 2 Diagnosis._--Medium Diagnosis._--Medium NNP 37809 513 3 - - HYPH 37809 513 4 sized sized JJ 37809 513 5 frogs frog NNS 37809 513 6 , , , 37809 513 7 males male VBZ 37809 513 8 17.5 17.5 CD 37809 513 9 - - SYM 37809 513 10 26.1 26.1 CD 37809 513 11 mm mm CD 37809 513 12 . . . 37809 514 1 snout snout NN 37809 514 2 - - HYPH 37809 514 3 vent vent NN 37809 514 4 , , , 37809 514 5 females female VBZ 37809 514 6 28.0 28.0 CD 37809 514 7 - - HYPH 37809 514 8 31.7 31.7 CD 37809 514 9 mm mm NNP 37809 514 10 . . . 37809 515 1 snout snout NN 37809 515 2 - - HYPH 37809 515 3 vent vent NN 37809 515 4 length length NN 37809 515 5 ; ; : 37809 515 6 vocal vocal NN 37809 515 7 slits slit NNS 37809 515 8 in in IN 37809 515 9 males male NNS 37809 515 10 ; ; , 37809 515 11 digital digital JJ 37809 515 12 tips tip NNS 37809 515 13 slightly slightly RB 37809 515 14 expanded expand VBN 37809 515 15 ; ; : 37809 515 16 first first RB 37809 515 17 finger finger NN 37809 515 18 shorter short JJR 37809 515 19 than than IN 37809 515 20 second second JJ 37809 515 21 ; ; : 37809 515 22 skin skin NN 37809 515 23 of of IN 37809 515 24 dorsum dorsum NN 37809 515 25 pustular pustular NN 37809 515 26 , , , 37809 515 27 that that DT 37809 515 28 of of IN 37809 515 29 venter venter NN 37809 515 30 areolate areolate NN 37809 515 31 ; ; : 37809 515 32 snout snout NNP 37809 515 33 elongate elongate NN 37809 515 34 , , , 37809 515 35 subacuminate subacuminate VBP 37809 515 36 ; ; : 37809 515 37 diameter diameter NN 37809 515 38 of of IN 37809 515 39 tympanum tympanum CD 37809 515 40 56.1 56.1 CD 37809 515 41 - - SYM 37809 515 42 76.7 76.7 CD 37809 515 43 per per IN 37809 515 44 cent cent NN 37809 515 45 that that DT 37809 515 46 of of IN 37809 515 47 eye eye NN 37809 515 48 in in IN 37809 515 49 males male NNS 37809 515 50 , , , 37809 515 51 54.3 54.3 CD 37809 515 52 - - SYM 37809 515 53 56.8 56.8 CD 37809 515 54 in in IN 37809 515 55 females female NNS 37809 515 56 ; ; : 37809 515 57 in in IN 37809 515 58 preservative preservative JJ 37809 515 59 , , , 37809 515 60 dorsum dorsum NN 37809 515 61 reddish reddish JJ 37809 515 62 brown brown JJ 37809 515 63 with with IN 37809 515 64 numerous numerous JJ 37809 515 65 small small JJ 37809 515 66 black black JJ 37809 515 67 or or CC 37809 515 68 dark dark JJ 37809 515 69 brown brown JJ 37809 515 70 spots spot NNS 37809 515 71 ( ( -LRB- 37809 515 72 Fig fig NN 37809 515 73 . . . 37809 516 1 8) 8) CD 37809 516 2 ; ; : 37809 516 3 venter venter NN 37809 516 4 white white JJ 37809 516 5 to to TO 37809 516 6 cream cream VB 37809 516 7 ; ; : 37809 516 8 in in IN 37809 516 9 life life NN 37809 516 10 dorsum dorsum NN 37809 516 11 green green JJ 37809 516 12 with with IN 37809 516 13 darker dark JJR 37809 516 14 green green JJ 37809 516 15 spots spot NNS 37809 516 16 , , , 37809 516 17 belly belly NN 37809 516 18 white white JJ 37809 516 19 ; ; : 37809 516 20 iris iris JJ 37809 516 21 gold gold NN 37809 516 22 above above RB 37809 516 23 , , , 37809 516 24 bronze bronze NN 37809 516 25 below below RB 37809 516 26 . . . 37809 517 1 _ _ NNP 37809 517 2 Remarks._--Cope Remarks._--Cope NNP 37809 517 3 's 's POS 37809 517 4 ( ( -LRB- 37809 517 5 1885 1885 CD 37809 517 6 ) ) -RRB- 37809 517 7 original original JJ 37809 517 8 description description NN 37809 517 9 was be VBD 37809 517 10 not not RB 37809 517 11 sufficiently sufficiently RB 37809 517 12 clear clear JJ 37809 517 13 to to TO 37809 517 14 enable enable VB 37809 517 15 subsequent subsequent JJ 37809 517 16 authors author NNS 37809 517 17 to to TO 37809 517 18 recognize recognize VB 37809 517 19 this this DT 37809 517 20 species specie NNS 37809 517 21 . . . 37809 518 1 Taylor Taylor NNP 37809 518 2 ( ( -LRB- 37809 518 3 1940e 1940e CD 37809 518 4 ) ) -RRB- 37809 518 5 described describe VBD 37809 518 6 it -PRON- PRP 37809 518 7 as as IN 37809 518 8 a a DT 37809 518 9 _ _ NNP 37809 518 10 Tomodactylus Tomodactylus NNP 37809 518 11 _ _ NNP 37809 518 12 , , , 37809 518 13 but but CC 37809 518 14 Dixon Dixon NNP 37809 518 15 ( ( -LRB- 37809 518 16 1957 1957 CD 37809 518 17 ) ) -RRB- 37809 518 18 pointed point VBD 37809 518 19 out out RP 37809 518 20 that that IN 37809 518 21 _ _ NNP 37809 518 22 T. T. NNP 37809 519 1 macrotympanum macrotympanum NNP 37809 519 2 _ _ NNP 37809 519 3 differed differ VBD 37809 519 4 from from IN 37809 519 5 the the DT 37809 519 6 other other JJ 37809 519 7 species specie NNS 37809 519 8 of of IN 37809 519 9 the the DT 37809 519 10 genus genus NN 37809 519 11 in in IN 37809 519 12 having have VBG 37809 519 13 a a DT 37809 519 14 poorly poorly RB 37809 519 15 developed develop VBN 37809 519 16 lumbo lumbo JJ 37809 519 17 - - HYPH 37809 519 18 inguinal inguinal NN 37809 519 19 ( ( -LRB- 37809 519 20 inguinal inguinal JJ 37809 519 21 ) ) -RRB- 37809 519 22 gland gland NNP 37809 519 23 , , , 37809 519 24 and and CC 37809 519 25 placed place VBD 37809 519 26 the the DT 37809 519 27 species specie NNS 37809 519 28 in in IN 37809 519 29 the the DT 37809 519 30 genus genus NN 37809 519 31 _ _ NNP 37809 519 32 Syrrhophus Syrrhophus NNP 37809 519 33 _ _ NNP 37809 519 34 . . . 37809 520 1 Comparison comparison NN 37809 520 2 of of IN 37809 520 3 the the DT 37809 520 4 holotypes holotype NNS 37809 520 5 of of IN 37809 520 6 _ _ NNP 37809 520 7 S. S. NNP 37809 521 1 verrucipes verrucipes NNP 37809 521 2 _ _ NNP 37809 521 3 and and CC 37809 521 4 _ _ NNP 37809 521 5 T. T. NNP 37809 522 1 macrotympanum macrotympanum NNP 37809 522 2 _ _ NNP 37809 522 3 leaves leave VBZ 37809 522 4 no no DT 37809 522 5 doubt doubt NN 37809 522 6 in in IN 37809 522 7 my -PRON- PRP$ 37809 522 8 mind mind NN 37809 522 9 that that IN 37809 522 10 a a DT 37809 522 11 single single JJ 37809 522 12 species species NN 37809 522 13 is be VBZ 37809 522 14 involved involve VBN 37809 522 15 . . . 37809 523 1 This this DT 37809 523 2 same same JJ 37809 523 3 species specie NNS 37809 523 4 was be VBD 37809 523 5 reported report VBN 37809 523 6 by by IN 37809 523 7 Smith Smith NNP 37809 523 8 and and CC 37809 523 9 Taylor Taylor NNP 37809 523 10 ( ( -LRB- 37809 523 11 1948 1948 CD 37809 523 12 ) ) -RRB- 37809 523 13 as as IN 37809 523 14 _ _ NNP 37809 523 15 S. S. NNP 37809 524 1 verruculatus verruculatus NNP 37809 524 2 _ _ NNP 37809 524 3 . . . 37809 525 1 _ _ NNP 37809 525 2 Syrrhophus Syrrhophus NNP 37809 525 3 verrucipes verrucipe VBZ 37809 525 4 _ _ NNP 37809 525 5 bears bear VBZ 37809 525 6 resemblance resemblance NN 37809 525 7 to to IN 37809 525 8 members member NNS 37809 525 9 of of IN 37809 525 10 both both CC 37809 525 11 the the DT 37809 525 12 _ _ NNP 37809 525 13 leprus leprus NNP 37809 525 14 _ _ NNP 37809 525 15 and and CC 37809 525 16 _ _ NNP 37809 525 17 marnockii marnockii NNS 37809 525 18 _ _ NNP 37809 525 19 groups group NNS 37809 525 20 . . . 37809 526 1 In in IN 37809 526 2 snout snout NN 37809 526 3 shape shape NN 37809 526 4 it -PRON- PRP 37809 526 5 is be VBZ 37809 526 6 closer close JJR 37809 526 7 to to IN 37809 526 8 the the DT 37809 526 9 _ _ NNP 37809 526 10 leprus leprus NNP 37809 526 11 _ _ NNP 37809 526 12 group group NN 37809 526 13 , , , 37809 526 14 whereas whereas IN 37809 526 15 in in IN 37809 526 16 digital digital NNP 37809 526 17 pad pad NNP 37809 526 18 , , , 37809 526 19 the the DT 37809 526 20 shape shape NN 37809 526 21 of of IN 37809 526 22 the the DT 37809 526 23 general general JJ 37809 526 24 body body NN 37809 526 25 form form NN 37809 526 26 , , , 37809 526 27 and and CC 37809 526 28 contiguity contiguity NN 37809 526 29 of of IN 37809 526 30 habitat habitat NN 37809 526 31 it -PRON- PRP 37809 526 32 is be VBZ 37809 526 33 most most RBS 37809 526 34 similar similar JJ 37809 526 35 to to IN 37809 526 36 the the DT 37809 526 37 _ _ NNP 37809 526 38 marnockii marnockii NNS 37809 526 39 _ _ NNP 37809 526 40 group group NN 37809 526 41 ( ( -LRB- 37809 526 42 _ _ NNP 37809 526 43 S. S. NNP 37809 527 1 guttilatus guttilatus NNP 37809 527 2 _ _ NNP 37809 527 3 ) ) -RRB- 37809 527 4 . . . 37809 528 1 _ _ NNP 37809 528 2 Etymology._--Latin Etymology._--Latin NNP 37809 528 3 , , , 37809 528 4 meaning mean VBG 37809 528 5 warty warty JJ 37809 528 6 foot foot NN 37809 528 7 , , , 37809 528 8 probably probably RB 37809 528 9 in in IN 37809 528 10 reference reference NN 37809 528 11 to to IN 37809 528 12 the the DT 37809 528 13 numerous numerous JJ 37809 528 14 plantar plantar NNP 37809 528 15 supernumerary supernumerary NNP 37809 528 16 tubercles tubercle NNS 37809 528 17 . . . 37809 529 1 _ _ NNP 37809 529 2 Distribution._--Moderate distribution._--moderate JJ 37809 529 3 elevations elevation NNS 37809 529 4 in in IN 37809 529 5 southeastern southeastern JJ 37809 529 6 San San NNP 37809 529 7 Luis Luis NNP 37809 529 8 Potosí Potosí NNP 37809 529 9 , , , 37809 529 10 Queretaro Queretaro NNP 37809 529 11 , , , 37809 529 12 and and CC 37809 529 13 northwestern northwestern JJ 37809 529 14 Hidalgo Hidalgo NNP 37809 529 15 , , , 37809 529 16 México México NNP 37809 529 17 ( ( -LRB- 37809 529 18 Fig Fig NNP 37809 529 19 . . . 37809 530 1 7 7 LS 37809 530 2 ) ) -RRB- 37809 530 3 . . . 37809 531 1 _ _ NNP 37809 531 2 Specimens Specimens NNP 37809 531 3 examined_--(43 examined_--(43 NNS 37809 531 4 ) ) -RRB- 37809 531 5 MÉXICO MÉXICO NNP 37809 531 6 , , , 37809 531 7 _ _ NNP 37809 531 8 Hidalgo Hidalgo NNP 37809 531 9 _ _ NNP 37809 531 10 : : : 37809 531 11 Jacala Jacala NNP 37809 531 12 , , , 37809 531 13 UMMZ UMMZ NNP 37809 531 14 106434 106434 CD 37809 531 15 ; ; : 37809 531 16 9.6 9.6 CD 37809 531 17 km km NN 37809 531 18 . . . 37809 532 1 NE NE NNP 37809 532 2 Jacala Jacala NNP 37809 532 3 , , , 37809 532 4 Puerto Puerto NNP 37809 532 5 de de NNP 37809 532 6 la la NNP 37809 532 7 Zorra Zorra NNP 37809 532 8 , , , 37809 532 9 1820 1820 CD 37809 532 10 m. m. NN 37809 532 11 , , , 37809 532 12 KU KU NNP 37809 532 13 60240 60240 CD 37809 532 14 - - SYM 37809 532 15 41 41 CD 37809 532 16 , , , 37809 532 17 TCWC TCWC NNP 37809 532 18 11090 11090 CD 37809 532 19 , , , 37809 532 20 11147 11147 CD 37809 532 21 ; ; : 37809 532 22 8 8 CD 37809 532 23 km km NN 37809 532 24 . . . 37809 533 1 S S NNP 37809 533 2 Jacala Jacala NNP 37809 533 3 , , , 37809 533 4 La La NNP 37809 533 5 Placita Placita NNP 37809 533 6 , , , 37809 533 7 1850 1850 CD 37809 533 8 m. m. NN 37809 533 9 , , , 37809 533 10 FMNH FMNH NNP 37809 533 11 100049 100049 CD 37809 533 12 ( ( -LRB- 37809 533 13 holotype holotype NN 37809 533 14 of of IN 37809 533 15 _ _ NNP 37809 533 16 Tomodactylus Tomodactylus NNP 37809 533 17 macrotympanum macrotympanum NN 37809 533 18 _ _ NNP 37809 533 19 ) ) -RRB- 37809 533 20 , , , 37809 533 21 100791 100791 CD 37809 533 22 - - SYM 37809 533 23 803 803 CD 37809 533 24 , , , 37809 533 25 105334 105334 CD 37809 533 26 - - SYM 37809 533 27 35 35 CD 37809 533 28 , , , 37809 533 29 114287 114287 CD 37809 533 30 , , , 37809 533 31 UIMNH UIMNH NNP 37809 533 32 15989 15989 CD 37809 533 33 - - SYM 37809 533 34 92 92 CD 37809 533 35 , , , 37809 533 36 15995 15995 CD 37809 533 37 - - SYM 37809 533 38 96 96 CD 37809 533 39 , , , 37809 533 40 UMMZ UMMZ NNP 37809 533 41 117252 117252 CD 37809 533 42 , , , 37809 533 43 USNM USNM NNP 37809 533 44 137202 137202 CD 37809 533 45 ; ; : 37809 533 46 Tianguistengo Tianguistengo NNP 37809 533 47 , , , 37809 533 48 FMNH FMNH NNP 37809 533 49 113705 113705 CD 37809 533 50 - - SYM 37809 533 51 09 09 CD 37809 533 52 , , , 37809 533 53 UIMNH UIMNH NNP 37809 533 54 13328 13328 CD 37809 533 55 - - SYM 37809 533 56 30 30 CD 37809 533 57 ; ; : 37809 533 58 near near IN 37809 533 59 Zacualtipán Zacualtipán NNP 37809 533 60 , , , 37809 533 61 ANSP ANSP NNP 37809 533 62 11325 11325 CD 37809 533 63 ( ( -LRB- 37809 533 64 holotype holotype NN 37809 533 65 of of IN 37809 533 66 _ _ NNP 37809 533 67 Syrrhophus Syrrhophus NNP 37809 533 68 verrucipes verrucipe VBZ 37809 533 69 _ _ NNP 37809 533 70 ) ) -RRB- 37809 533 71 . . . 37809 534 1 _ _ NNP 37809 534 2 Queretaro Queretaro NNP 37809 534 3 _ _ NNP 37809 534 4 : : : 37809 534 5 3.5 3.5 CD 37809 534 6 km km NN 37809 534 7 . . . 37809 535 1 S S NNP 37809 535 2 San San NNP 37809 535 3 Juan Juan NNP 37809 535 4 del del NNP 37809 535 5 Río Río NNP 37809 535 6 , , , 37809 535 7 EAL EAL NNP 37809 535 8 1343 1343 CD 37809 535 9 . . . 37809 536 1 _ _ NNP 37809 536 2 San San NNP 37809 536 3 Luis Luis NNP 37809 536 4 Potosí Potosí NNP 37809 536 5 _ _ NNP 37809 536 6 : : : 37809 536 7 9.6 9.6 CD 37809 536 8 km km NN 37809 536 9 . . . 37809 537 1 W W NNP 37809 537 2 Ahuacatlán Ahuacatlán NNP 37809 537 3 , , , 37809 537 4 LSUMZ LSUMZ NNP 37809 537 5 4968 4968 CD 37809 537 6 - - SYM 37809 537 7 70 70 CD 37809 537 8 . . . 37809 538 1 = = NFP 37809 538 2 Syrrhophus Syrrhophus NNP 37809 538 3 dennisi= dennisi= NN 37809 538 4 new new JJ 37809 538 5 species specie NNS 37809 538 6 _ _ NNP 37809 538 7 Syrrhophus Syrrhophus NNP 37809 538 8 latodactylus latodactylus NN 37809 538 9 _ _ NNP 37809 538 10 : : : 37809 538 11 Martin Martin NNP 37809 538 12 , , , 37809 538 13 1958:49 1958:49 CD 37809 538 14 ( ( -LRB- 37809 538 15 in in IN 37809 538 16 part part NN 37809 538 17 ) ) -RRB- 37809 538 18 . . . 37809 539 1 _ _ NNP 37809 539 2 Holotype._--UMMZ Holotype._--UMMZ NNP 37809 539 3 101121 101121 CD 37809 539 4 , , , 37809 539 5 adult adult NN 37809 539 6 male male NN 37809 539 7 from from IN 37809 539 8 a a DT 37809 539 9 cave cave NN 37809 539 10 near near IN 37809 539 11 El El NNP 37809 539 12 Pachón Pachón NNP 37809 539 13 , , , 37809 539 14 8 8 CD 37809 539 15 km km NN 37809 539 16 . . . 37809 540 1 N n IN 37809 540 2 Antiguo Antiguo NNP 37809 540 3 Morelos Morelos NNP 37809 540 4 , , , 37809 540 5 Tamaulipas Tamaulipas NNP 37809 540 6 , , , 37809 540 7 México México NNP 37809 540 8 , , , 37809 540 9 250 250 CD 37809 540 10 m. m. NN 37809 540 11 , , , 37809 540 12 collected collect VBN 37809 540 13 on on IN 37809 540 14 March March NNP 37809 540 15 13 13 CD 37809 540 16 , , , 37809 540 17 1949 1949 CD 37809 540 18 , , , 37809 540 19 by by IN 37809 540 20 Paul Paul NNP 37809 540 21 S. S. NNP 37809 540 22 Martin Martin NNP 37809 540 23 . . . 37809 541 1 _ _ NNP 37809 541 2 Paratopotypes._--(26 Paratopotypes._--(26 NNP 37809 541 3 ) ) -RRB- 37809 541 4 . . . 37809 542 1 UMMZ UMMZ NNP 37809 542 2 101122 101122 CD 37809 542 3 ( ( -LRB- 37809 542 4 10 10 CD 37809 542 5 ) ) -RRB- 37809 542 6 , , , 37809 542 7 101123 101123 CD 37809 542 8 ( ( -LRB- 37809 542 9 2 2 CD 37809 542 10 ) ) -RRB- 37809 542 11 , , , 37809 542 12 101126 101126 CD 37809 542 13 , , , 37809 542 14 126993 126993 CD 37809 542 15 ( ( -LRB- 37809 542 16 12 12 CD 37809 542 17 ) ) -RRB- 37809 542 18 . . . 37809 543 1 _ _ NNP 37809 543 2 Diagnosis._--Medium Diagnosis._--Medium NNP 37809 543 3 - - HYPH 37809 543 4 sized sized JJ 37809 543 5 frogs frog NNS 37809 543 6 , , , 37809 543 7 males male VBZ 37809 543 8 22.8 22.8 CD 37809 543 9 - - SYM 37809 543 10 28.4 28.4 CD 37809 543 11 mm mm NNP 37809 543 12 . . . 37809 544 1 snout snout NN 37809 544 2 - - HYPH 37809 544 3 vent vent NN 37809 544 4 , , , 37809 544 5 females female VBZ 37809 544 6 25.9 25.9 CD 37809 544 7 - - HYPH 37809 544 8 32.0 32.0 CD 37809 544 9 mm mm NN 37809 544 10 . . . 37809 545 1 snout snout NN 37809 545 2 - - HYPH 37809 545 3 vent vent NN 37809 545 4 ; ; : 37809 545 5 vocal vocal NN 37809 545 6 slits slit NNS 37809 545 7 in in IN 37809 545 8 males male NNS 37809 545 9 ; ; , 37809 545 10 digital digital JJ 37809 545 11 tips tip NNS 37809 545 12 greatly greatly RB 37809 545 13 expanded expand VBN 37809 545 14 , , , 37809 545 15 more more JJR 37809 545 16 than than IN 37809 545 17 twice twice RB 37809 545 18 width width NN 37809 545 19 of of IN 37809 545 20 digit digit NN 37809 545 21 ; ; : 37809 545 22 first first RB 37809 545 23 finger finger NN 37809 545 24 shorter short JJR 37809 545 25 than than IN 37809 545 26 second second JJ 37809 545 27 ; ; : 37809 545 28 skin skin NN 37809 545 29 of of IN 37809 545 30 dorsum dorsum NN 37809 545 31 shagreened shagreene VBN 37809 545 32 to to IN 37809 545 33 pustular pustular NNP 37809 545 34 , , , 37809 545 35 that that DT 37809 545 36 of of IN 37809 545 37 venter venter NN 37809 545 38 weakly weakly RB 37809 545 39 to to TO 37809 545 40 moderately moderately RB 37809 545 41 areolate areolate VB 37809 545 42 ; ; : 37809 545 43 toes toe NNS 37809 545 44 webbed web VBD 37809 545 45 basally basally RB 37809 545 46 ; ; : 37809 545 47 dorsum dorsum NN 37809 545 48 light light NN 37809 545 49 brown brown NNP 37809 545 50 to to IN 37809 545 51 tan tan NNP 37809 545 52 with with IN 37809 545 53 brown brown JJ 37809 545 54 vermiculations vermiculation NNS 37809 545 55 ; ; : 37809 545 56 venter venter NN 37809 545 57 white white JJ 37809 545 58 ; ; : 37809 545 59 diameter diameter NN 37809 545 60 of of IN 37809 545 61 tympanum tympanum NN 37809 545 62 53.9 53.9 CD 37809 545 63 to to IN 37809 545 64 64.2 64.2 CD 37809 545 65 per per IN 37809 545 66 cent cent NN 37809 545 67 that that DT 37809 545 68 of of IN 37809 545 69 eye eye NN 37809 545 70 in in IN 37809 545 71 males male NNS 37809 545 72 , , , 37809 545 73 50.6 50.6 CD 37809 545 74 to to IN 37809 545 75 58.7 58.7 CD 37809 545 76 per per IN 37809 545 77 cent cent NN 37809 545 78 in in IN 37809 545 79 females female NNS 37809 545 80 . . . 37809 546 1 _ _ NNP 37809 546 2 Description Description NNP 37809 546 3 and and CC 37809 546 4 variation._--(Fig variation._--(fig NN 37809 546 5 . . . 37809 547 1 12 12 CD 37809 547 2 ) ) -RRB- 37809 547 3 . . . 37809 548 1 Head head VB 37809 548 2 wider wider RBR 37809 548 3 than than IN 37809 548 4 body body NN 37809 548 5 ; ; : 37809 548 6 head head VB 37809 548 7 as as RB 37809 548 8 wide wide JJ 37809 548 9 or or CC 37809 548 10 wider wide JJR 37809 548 11 than than IN 37809 548 12 long long RB 37809 548 13 in in IN 37809 548 14 males male NNS 37809 548 15 , , , 37809 548 16 sometimes sometimes RB 37809 548 17 longer long JJR 37809 548 18 than than IN 37809 548 19 wide wide JJ 37809 548 20 in in IN 37809 548 21 females female NNS 37809 548 22 ; ; : 37809 548 23 snout snout NN 37809 548 24 acuminate acuminate NN 37809 548 25 in in IN 37809 548 26 dorsal dorsal NN 37809 548 27 view view NN 37809 548 28 , , , 37809 548 29 elongate elongate VB 37809 548 30 and and CC 37809 548 31 rounded round VBD 37809 548 32 in in IN 37809 548 33 lateral lateral JJ 37809 548 34 profile profile NN 37809 548 35 ; ; : 37809 548 36 canthus canthus NNP 37809 548 37 rostralis rostralis NNP 37809 548 38 rounded round VBD 37809 548 39 but but CC 37809 548 40 distinct distinct JJ 37809 548 41 ; ; : 37809 548 42 loreal loreal NN 37809 548 43 region region NN 37809 548 44 slightly slightly RB 37809 548 45 concave concave NNP 37809 548 46 , , , 37809 548 47 sloping slope VBG 37809 548 48 abruptly abruptly RB 37809 548 49 to to IN 37809 548 50 lip lip NN 37809 548 51 ; ; : 37809 548 52 lips lip NNS 37809 548 53 not not RB 37809 548 54 flared flare VBN 37809 548 55 ; ; : 37809 548 56 eyelid eyelid JJR 37809 548 57 about about RB 37809 548 58 two two CD 37809 548 59 - - HYPH 37809 548 60 thirds third NNS 37809 548 61 interorbital interorbital JJ 37809 548 62 distance distance NN 37809 548 63 ; ; : 37809 548 64 length length NN 37809 548 65 of of IN 37809 548 66 eye eye NN 37809 548 67 less less JJR 37809 548 68 than than IN 37809 548 69 distance distance NN 37809 548 70 between between IN 37809 548 71 eye eye NN 37809 548 72 and and CC 37809 548 73 nostril nostril NNP 37809 548 74 ; ; : 37809 548 75 diameter diameter NN 37809 548 76 of of IN 37809 548 77 tympanum tympanum NN 37809 548 78 53.9 53.9 CD 37809 548 79 to to IN 37809 548 80 64.2 64.2 CD 37809 548 81 per per IN 37809 548 82 cent cent NN 37809 548 83 that that DT 37809 548 84 of of IN 37809 548 85 eye eye NN 37809 548 86 in in IN 37809 548 87 males male NNS 37809 548 88 , , , 37809 548 89 50.6 50.6 CD 37809 548 90 to to IN 37809 548 91 58.7 58.7 CD 37809 548 92 per per IN 37809 548 93 cent cent NN 37809 548 94 in in IN 37809 548 95 females female NNS 37809 548 96 ; ; , 37809 548 97 tympanum tympanum VB 37809 548 98 round round NN 37809 548 99 and and CC 37809 548 100 distinct distinct JJ 37809 548 101 in in IN 37809 548 102 both both DT 37809 548 103 sexes sex NNS 37809 548 104 ; ; : 37809 548 105 supratympanic supratympanic NNP 37809 548 106 fold fold VBP 37809 548 107 moderately moderately RB 37809 548 108 distinct distinct JJ 37809 548 109 ; ; : 37809 548 110 choanae choanae NN 37809 548 111 within within IN 37809 548 112 border border NN 37809 548 113 of of IN 37809 548 114 jaws jaw NNS 37809 548 115 , , , 37809 548 116 completely completely RB 37809 548 117 visible visible JJ 37809 548 118 from from IN 37809 548 119 directly directly RB 37809 548 120 below below RB 37809 548 121 , , , 37809 548 122 rounded round VBD 37809 548 123 to to IN 37809 548 124 slightly slightly RB 37809 548 125 oval oval VB 37809 548 126 ; ; , 37809 548 127 dentigerous dentigerous JJ 37809 548 128 processes process NNS 37809 548 129 of of IN 37809 548 130 prevomers prevomer NNS 37809 548 131 and and CC 37809 548 132 teeth tooth NNS 37809 548 133 absent absent JJ 37809 548 134 ; ; : 37809 548 135 tongue tongue NN 37809 548 136 free free JJ 37809 548 137 for for IN 37809 548 138 posterior posterior RB 37809 548 139 one one CD 37809 548 140 - - HYPH 37809 548 141 half half NN 37809 548 142 , , , 37809 548 143 generally generally RB 37809 548 144 oval oval JJ 37809 548 145 in in IN 37809 548 146 outline outline NN 37809 548 147 ; ; : 37809 548 148 vocal vocal JJ 37809 548 149 slits slit NNS 37809 548 150 present present JJ 37809 548 151 in in IN 37809 548 152 males male NNS 37809 548 153 . . . 37809 549 1 Many many JJ 37809 549 2 scattered scatter VBN 37809 549 3 pustules pustule NNS 37809 549 4 on on IN 37809 549 5 dorsum dorsum NN 37809 549 6 ; ; : 37809 549 7 flanks flank NNS 37809 549 8 areolate areolate VBP 37809 549 9 ; ; : 37809 549 10 skin skin NN 37809 549 11 of of IN 37809 549 12 venter venter NN 37809 549 13 areolate areolate NN 37809 549 14 or or CC 37809 549 15 not not RB 37809 549 16 ( ( -LRB- 37809 549 17 variability variability NN 37809 549 18 may may MD 37809 549 19 be be VB 37809 549 20 due due JJ 37809 549 21 to to IN 37809 549 22 differences difference NNS 37809 549 23 in in IN 37809 549 24 preservation preservation NN 37809 549 25 ) ) -RRB- 37809 549 26 ; ; : 37809 549 27 ventral ventral JJ 37809 549 28 disc disc NN 37809 549 29 distinct distinct JJ 37809 549 30 on on IN 37809 549 31 chest chest NN 37809 549 32 and and CC 37809 549 33 lower low JJR 37809 549 34 abdomen abdoman NNS 37809 549 35 ; ; : 37809 549 36 inguinal inguinal NNP 37809 549 37 gland gland NNP 37809 549 38 present present JJ 37809 549 39 or or CC 37809 549 40 not not RB 37809 549 41 , , , 37809 549 42 when when WRB 37809 549 43 present present JJ 37809 549 44 varying varying NN 37809 549 45 from from IN 37809 549 46 very very RB 37809 549 47 large large JJ 37809 549 48 and and CC 37809 549 49 distinct distinct JJ 37809 549 50 to to TO 37809 549 51 poorly poorly RB 37809 549 52 defined define VBN 37809 549 53 ; ; : 37809 549 54 axillary axillary JJ 37809 549 55 gland gland NN 37809 549 56 absent absent NN 37809 549 57 . . . 37809 550 1 First first JJ 37809 550 2 finger finger NN 37809 550 3 shorter short JJR 37809 550 4 than than IN 37809 550 5 second second RB 37809 550 6 ; ; : 37809 550 7 all all DT 37809 550 8 fingers finger NNS 37809 550 9 bearing bear VBG 37809 550 10 truncate truncate JJ 37809 550 11 tips tip NNS 37809 550 12 with with IN 37809 550 13 pads pad NNS 37809 550 14 , , , 37809 550 15 each each DT 37809 550 16 pad pad NN 37809 550 17 having have VBG 37809 550 18 a a DT 37809 550 19 terminal terminal JJ 37809 550 20 groove groove NN 37809 550 21 ; ; : 37809 550 22 fingers finger NNS 37809 550 23 fringed fringe VBN 37809 550 24 ; ; : 37809 550 25 fingers finger NNS 37809 550 26 three three CD 37809 550 27 and and CC 37809 550 28 four four CD 37809 550 29 having have VBG 37809 550 30 dilated dilate VBN 37809 550 31 pads pad NNS 37809 550 32 two two CD 37809 550 33 to to IN 37809 550 34 three three CD 37809 550 35 times time NNS 37809 550 36 width width NN 37809 550 37 of of IN 37809 550 38 digit digit NN 37809 550 39 ; ; , 37809 550 40 subarticular subarticular JJ 37809 550 41 tubercles tubercle NNS 37809 550 42 large large JJ 37809 550 43 , , , 37809 550 44 conical conical JJ 37809 550 45 , , , 37809 550 46 rounded rounded JJ 37809 550 47 , , , 37809 550 48 simple simple JJ 37809 550 49 ; ; : 37809 550 50 supernumerary supernumerary JJ 37809 550 51 tubercles tubercle NNS 37809 550 52 numerous numerous JJ 37809 550 53 on on IN 37809 550 54 thenar thenar NN 37809 550 55 surface surface NN 37809 550 56 , , , 37809 550 57 none none NN 37809 550 58 on on IN 37809 550 59 digits digit NNS 37809 550 60 ; ; : 37809 550 61 three three CD 37809 550 62 palmar palmar NNP 37809 550 63 tubercles tubercle NNS 37809 550 64 , , , 37809 550 65 outer outer VBP 37809 550 66 slightly slightly RB 37809 550 67 smaller small JJR 37809 550 68 than than IN 37809 550 69 largest large JJS 37809 550 70 supernumerary supernumerary JJ 37809 550 71 tubercles tubercle NNS 37809 550 72 ; ; : 37809 550 73 row row NN 37809 550 74 of of IN 37809 550 75 tubercles tubercle NNS 37809 550 76 on on IN 37809 550 77 outer outer JJ 37809 550 78 edge edge NN 37809 550 79 of of IN 37809 550 80 forearm forearm NN 37809 550 81 variable variable NN 37809 550 82 , , , 37809 550 83 weak weak JJ 37809 550 84 to to IN 37809 550 85 very very RB 37809 550 86 distinct distinct JJ 37809 550 87 ; ; : 37809 550 88 tips tip NNS 37809 550 89 of of IN 37809 550 90 toes toe NNS 37809 550 91 wider wide JJR 37809 550 92 than than IN 37809 550 93 digits digit NNS 37809 550 94 , , , 37809 550 95 rounded round VBD 37809 550 96 to to TO 37809 550 97 truncate truncate VB 37809 550 98 at at IN 37809 550 99 tips tip NNS 37809 550 100 , , , 37809 550 101 each each DT 37809 550 102 pad pad NN 37809 550 103 having have VBG 37809 550 104 terminal terminal JJ 37809 550 105 groove groove NN 37809 550 106 ; ; : 37809 550 107 toes toe NNS 37809 550 108 having have VBG 37809 550 109 lateral lateral JJ 37809 550 110 fringes fringe NNS 37809 550 111 , , , 37809 550 112 bases basis NNS 37809 550 113 of of IN 37809 550 114 toes toe NNS 37809 550 115 united unite VBN 37809 550 116 by by IN 37809 550 117 web web NN 37809 550 118 , , , 37809 550 119 web web VBD 37809 550 120 not not RB 37809 550 121 extending extend VBG 37809 550 122 to to TO 37809 550 123 basal basal VB 37809 550 124 subarticular subarticular JJ 37809 550 125 tubercle tubercle NN 37809 550 126 ; ; : 37809 550 127 subarticular subarticular JJ 37809 550 128 tubercles tubercle NNS 37809 550 129 smaller small JJR 37809 550 130 than than IN 37809 550 131 those those DT 37809 550 132 of of IN 37809 550 133 hand hand NN 37809 550 134 , , , 37809 550 135 round round NN 37809 550 136 , , , 37809 550 137 conical conical JJ 37809 550 138 , , , 37809 550 139 simple simple JJ 37809 550 140 ; ; : 37809 550 141 supernumerary supernumerary JJ 37809 550 142 tubercles tubercle NNS 37809 550 143 numerous numerous JJ 37809 550 144 on on IN 37809 550 145 plantar plantar NNP 37809 550 146 surfaces surface NNS 37809 550 147 , , , 37809 550 148 extending extend VBG 37809 550 149 between between IN 37809 550 150 metatarsal metatarsal NN 37809 550 151 tubercles tubercle NNS 37809 550 152 , , , 37809 550 153 present present JJ 37809 550 154 on on IN 37809 550 155 toes toe NNS 37809 550 156 between between IN 37809 550 157 basal basal NN 37809 550 158 two two CD 37809 550 159 subarticular subarticular JJ 37809 550 160 tubercles tubercle NNS 37809 550 161 in in IN 37809 550 162 some some DT 37809 550 163 specimens specimen NNS 37809 550 164 ; ; : 37809 550 165 outer outer JJ 37809 550 166 metatarsal metatarsal NN 37809 550 167 tubercle tubercle NN 37809 550 168 round round NN 37809 550 169 , , , 37809 550 170 conical conical JJ 37809 550 171 , , , 37809 550 172 one one CD 37809 550 173 - - HYPH 37809 550 174 half half NN 37809 550 175 as as RB 37809 550 176 large large JJ 37809 550 177 as as IN 37809 550 178 ovoid ovoid JJ 37809 550 179 , , , 37809 550 180 non non JJ 37809 550 181 - - JJ 37809 550 182 compressed compressed JJ 37809 550 183 inner inner JJ 37809 550 184 metatarsal metatarsal NN 37809 550 185 tubercle tubercle NN 37809 550 186 ; ; : 37809 550 187 tarsal tarsal NN 37809 550 188 tubercles tubercle NNS 37809 550 189 or or CC 37809 550 190 folds fold NNS 37809 550 191 absent absent JJ 37809 550 192 . . . 37809 551 1 Ground ground NN 37809 551 2 color color NN 37809 551 3 pale pale JJ 37809 551 4 reddish reddish JJ 37809 551 5 - - HYPH 37809 551 6 brown brown NN 37809 551 7 to to IN 37809 551 8 tan tan NNP 37809 551 9 dorsally dorsally RB 37809 551 10 , , , 37809 551 11 creamy creamy JJ 37809 551 12 on on IN 37809 551 13 flanks flank NNS 37809 551 14 ; ; : 37809 551 15 dorsal dorsal NN 37809 551 16 pattern pattern NN 37809 551 17 consisting consist VBG 37809 551 18 of of IN 37809 551 19 reddish reddish JJ 37809 551 20 - - HYPH 37809 551 21 brown brown NN 37809 551 22 to to IN 37809 551 23 brown brown JJ 37809 551 24 vermiculations vermiculation NNS 37809 551 25 extending extend VBG 37809 551 26 onto onto IN 37809 551 27 flanks flank NNS 37809 551 28 ; ; : 37809 551 29 distinct distinct JJ 37809 551 30 interorbital interorbital NNP 37809 551 31 light light NNP 37809 551 32 bar bar NN 37809 551 33 present present JJ 37809 551 34 ; ; : 37809 551 35 loreal loreal NN 37809 551 36 region region NN 37809 551 37 darker dark JJR 37809 551 38 than than IN 37809 551 39 snout snout NN 37809 551 40 , , , 37809 551 41 reddish reddish JJ 37809 551 42 - - HYPH 37809 551 43 brown brown NN 37809 551 44 compared compare VBN 37809 551 45 to to IN 37809 551 46 tan tan NN 37809 551 47 or or CC 37809 551 48 pale pale JJ 37809 551 49 reddish reddish JJ 37809 551 50 - - HYPH 37809 551 51 brown brown NN 37809 551 52 ; ; : 37809 551 53 arms arm NNS 37809 551 54 colored color VBN 37809 551 55 like like IN 37809 551 56 dorsum dorsum NN 37809 551 57 ; ; : 37809 551 58 thighs thigh NNS 37809 551 59 banded band VBD 37809 551 60 , , , 37809 551 61 unicolor unicolor NNP 37809 551 62 brown brown JJ 37809 551 63 on on IN 37809 551 64 posterior posterior NNP 37809 551 65 surfaces surface NNS 37809 551 66 ; ; : 37809 551 67 shanks shank NNS 37809 551 68 and and CC 37809 551 69 tarsi tarsi NNP 37809 551 70 banded banded JJ 37809 551 71 ; ; : 37809 551 72 venter venter NN 37809 551 73 white white JJ 37809 551 74 to to TO 37809 551 75 cream cream VB 37809 551 76 punctated punctate VBN 37809 551 77 with with IN 37809 551 78 brown brown NNP 37809 551 79 in in IN 37809 551 80 some some DT 37809 551 81 specimens specimen NNS 37809 551 82 . . . 37809 552 1 The the DT 37809 552 2 variation variation NN 37809 552 3 in in IN 37809 552 4 proportions proportion NNS 37809 552 5 is be VBZ 37809 552 6 summarized summarize VBN 37809 552 7 in in IN 37809 552 8 Table table NN 37809 552 9 5 5 CD 37809 552 10 . . . 37809 553 1 _ _ NNP 37809 553 2 Remarks._--Martin Remarks._--Martin NNP 37809 553 3 ( ( -LRB- 37809 553 4 1958 1958 CD 37809 553 5 ) ) -RRB- 37809 553 6 expressed express VBD 37809 553 7 some some DT 37809 553 8 doubt doubt NN 37809 553 9 that that IN 37809 553 10 this this DT 37809 553 11 series series NN 37809 553 12 of of IN 37809 553 13 26 26 CD 37809 553 14 specimens specimen NNS 37809 553 15 was be VBD 37809 553 16 identical identical JJ 37809 553 17 with with IN 37809 553 18 " " `` 37809 553 19 _ _ NNP 37809 553 20 S. S. NNP 37809 554 1 latodactylus latodactylus NNP 37809 554 2 _ _ NNP 37809 554 3 . . . 37809 554 4 " " '' 37809 555 1 My -PRON- PRP$ 37809 555 2 study study NN 37809 555 3 indicates indicate VBZ 37809 555 4 that that IN 37809 555 5 the the DT 37809 555 6 specimens specimen NNS 37809 555 7 from from IN 37809 555 8 El El NNP 37809 555 9 Pachón Pachón NNP 37809 555 10 represent represent VBP 37809 555 11 a a DT 37809 555 12 distinctive distinctive JJ 37809 555 13 but but CC 37809 555 14 allied allied JJ 37809 555 15 species specie NNS 37809 555 16 . . . 37809 556 1 Males male NNS 37809 556 2 of of IN 37809 556 3 the the DT 37809 556 4 two two CD 37809 556 5 species specie NNS 37809 556 6 can can MD 37809 556 7 be be VB 37809 556 8 readily readily RB 37809 556 9 separated separate VBN 37809 556 10 by by IN 37809 556 11 the the DT 37809 556 12 relative relative JJ 37809 556 13 sizes size NNS 37809 556 14 of of IN 37809 556 15 the the DT 37809 556 16 tympani tympani NNS 37809 556 17 , , , 37809 556 18 presence presence NN 37809 556 19 or or CC 37809 556 20 absence absence NN 37809 556 21 of of IN 37809 556 22 vocal vocal JJ 37809 556 23 slits slit NNS 37809 556 24 , , , 37809 556 25 and and CC 37809 556 26 color color NN 37809 556 27 pattern pattern NN 37809 556 28 . . . 37809 557 1 Females female NNS 37809 557 2 of of IN 37809 557 3 the the DT 37809 557 4 two two CD 37809 557 5 species specie NNS 37809 557 6 can can MD 37809 557 7 be be VB 37809 557 8 separated separate VBN 37809 557 9 by by IN 37809 557 10 color color NN 37809 557 11 pattern pattern NN 37809 557 12 . . . 37809 558 1 Within within IN 37809 558 2 the the DT 37809 558 3 type type NN 37809 558 4 - - HYPH 37809 558 5 series series NN 37809 558 6 , , , 37809 558 7 the the DT 37809 558 8 pattern pattern NN 37809 558 9 varies vary VBZ 37809 558 10 from from IN 37809 558 11 weakly weakly RB 37809 558 12 to to TO 37809 558 13 strongly strongly RB 37809 558 14 vermiculate vermiculate VB 37809 558 15 but but CC 37809 558 16 is be VBZ 37809 558 17 always always RB 37809 558 18 recognizable recognizable JJ 37809 558 19 as as IN 37809 558 20 vermiculate vermiculate JJ 37809 558 21 rather rather RB 37809 558 22 than than IN 37809 558 23 spotted spot VBN 37809 558 24 as as IN 37809 558 25 in in IN 37809 558 26 _ _ NNP 37809 558 27 S. S. NNP 37809 559 1 longipes longipe NNS 37809 559 2 _ _ NNP 37809 559 3 (= (= NNP 37809 559 4 _ _ NNP 37809 559 5 S. S. NNP 37809 560 1 latodactylus latodactylus NNP 37809 560 2 _ _ NNP 37809 560 3 of of IN 37809 560 4 Taylor Taylor NNP 37809 560 5 and and CC 37809 560 6 Martin Martin NNP 37809 560 7 ) ) -RRB- 37809 560 8 . . . 37809 561 1 [ [ -LRB- 37809 561 2 Illustration illustration NN 37809 561 3 : : : 37809 561 4 FIG FIG NNP 37809 561 5 . . . 37809 562 1 12 12 CD 37809 562 2 : : : 37809 562 3 _ _ NNP 37809 562 4 Syrrhophus Syrrhophus NNP 37809 562 5 dennisi dennisi RB 37809 562 6 _ _ NNP 37809 562 7 sp sp NNP 37809 562 8 . . . 37809 563 1 nov nov NNP 37809 563 2 . . NNP 37809 563 3 , , , 37809 563 4 holotype holotype NN 37809 563 5 , , , 37809 563 6 UMMZ UMMZ NNP 37809 563 7 101121 101121 CD 37809 563 8 ( ( -LRB- 37809 563 9 dorsum dorsum NN 37809 563 10 ×1.8 ×1.8 UH 37809 563 11 , , , 37809 563 12 side side NN 37809 563 13 of of IN 37809 563 14 head head NN 37809 563 15 ×6.1 ×6.1 NFP 37809 563 16 ) ) -RRB- 37809 563 17 . . . 37809 563 18 ] ] -RRB- 37809 564 1 _ _ NNP 37809 564 2 Etymology._--The Etymology._--The NNP 37809 564 3 specific specific JJ 37809 564 4 name name NN 37809 564 5 is be VBZ 37809 564 6 a a DT 37809 564 7 patronym patronym NN 37809 564 8 for for IN 37809 564 9 David David NNP 37809 564 10 M. M. NNP 37809 564 11 Dennis Dennis NNP 37809 564 12 , , , 37809 564 13 whose whose WP$ 37809 564 14 drawings drawing NNS 37809 564 15 greatly greatly RB 37809 564 16 enhance enhance VBP 37809 564 17 the the DT 37809 564 18 worth worth NN 37809 564 19 of of IN 37809 564 20 this this DT 37809 564 21 paper paper NN 37809 564 22 . . . 37809 565 1 _ _ NNP 37809 565 2 Distribution._--Known distribution._--known XX 37809 565 3 only only RB 37809 565 4 from from IN 37809 565 5 the the DT 37809 565 6 type type NN 37809 565 7 series series NN 37809 565 8 . . . 37809 566 1 = = NFP 37809 566 2 Syrrhophus Syrrhophus NNP 37809 566 3 longipes= longipes= NN 37809 566 4 ( ( -LRB- 37809 566 5 Baird Baird NNP 37809 566 6 ) ) -RRB- 37809 566 7 , , , 37809 566 8 New new JJ 37809 566 9 combination combination NN 37809 566 10 _ _ NNP 37809 566 11 Batrachyla Batrachyla NNP 37809 566 12 longipes longipe VBZ 37809 566 13 _ _ NNP 37809 566 14 Baird Baird NNP 37809 566 15 , , , 37809 566 16 1859:35 1859:35 NNP 37809 566 17 , , , 37809 566 18 pl pl NN 37809 566 19 . . . 37809 567 1 37 37 CD 37809 567 2 , , , 37809 567 3 fig fig NN 37809 567 4 . . . 37809 568 1 1 1 LS 37809 568 2 - - SYM 37809 568 3 3 3 CD 37809 568 4 [ [ -LRB- 37809 568 5 Holotype.--apparently holotype.--apparently RB 37809 568 6 USNM USNM NNP 37809 568 7 3237 3237 CD 37809 568 8 ( ( -LRB- 37809 568 9 cited cite VBN 37809 568 10 as as IN 37809 568 11 3207 3207 CD 37809 568 12 by by IN 37809 568 13 Cope Cope NNP 37809 568 14 , , , 37809 568 15 1887:16 1887:16 CD 37809 568 16 ) ) -RRB- 37809 568 17 , , , 37809 568 18 now now RB 37809 568 19 lost lose VBN 37809 568 20 , , , 37809 568 21 from from IN 37809 568 22 40 40 CD 37809 568 23 Leagues league NNS 37809 568 24 from from IN 37809 568 25 ( ( -LRB- 37809 568 26 probably probably RB 37809 568 27 north north NN 37809 568 28 ) ) -RRB- 37809 568 29 México México NNP 37809 568 30 City City NNP 37809 568 31 ; ; , 37809 568 32 collected collect VBN 37809 568 33 by by IN 37809 568 34 John John NNP 37809 568 35 Potts Potts NNP 37809 568 36 ] ] -RRB- 37809 568 37 . . . 37809 569 1 Kellogg Kellogg NNP 37809 569 2 , , , 37809 569 3 1932:107 1932:107 CD 37809 569 4 . . . 37809 570 1 _ _ NNP 37809 570 2 Epirhexis Epirhexis NNP 37809 570 3 longipes longipe VBZ 37809 570 4 _ _ NNP 37809 570 5 : : : 37809 570 6 Cope cope VB 37809 570 7 , , , 37809 570 8 1866:96 1866:96 CD 37809 570 9 . . . 37809 571 1 _ _ NNP 37809 571 2 Eleutherodactylus Eleutherodactylus NNP 37809 571 3 longipes longipe VBZ 37809 571 4 _ _ NNP 37809 571 5 : : : 37809 571 6 Kellogg Kellogg NNP 37809 571 7 , , , 37809 571 8 1932:107 1932:107 CD 37809 571 9 ( ( -LRB- 37809 571 10 part part NN 37809 571 11 ) ) -RRB- 37809 571 12 . . . 37809 572 1 Smith Smith NNP 37809 572 2 and and CC 37809 572 3 Taylor Taylor NNP 37809 572 4 , , , 37809 572 5 1948:61 1948:61 CD 37809 572 6 . . . 37809 573 1 Lynch Lynch NNP 37809 573 2 , , , 37809 573 3 1963:580 1963:580 CD 37809 573 4 - - SYM 37809 573 5 581 581 CD 37809 573 6 . . . 37809 574 1 Gorham Gorham NNP 37809 574 2 , , , 37809 574 3 1966:82 1966:82 CD 37809 574 4 . . . 37809 575 1 _ _ NNP 37809 575 2 Syrrhophus Syrrhophus NNP 37809 575 3 latodactylus latodactylus NN 37809 575 4 _ _ NNP 37809 575 5 Taylor Taylor NNP 37809 575 6 , , , 37809 575 7 1940d:396 1940d:396 CD 37809 575 8 - - SYM 37809 575 9 401 401 CD 37809 575 10 , , , 37809 575 11 pl pl NNP 37809 575 12 . . . 37809 576 1 43 43 CD 37809 576 2 , , , 37809 576 3 figs fig NNS 37809 576 4 . . . 37809 577 1 A A NNP 37809 577 2 - - HYPH 37809 577 3 F F NNP 37809 577 4 , , , 37809 577 5 text text NN 37809 577 6 fig fig NN 37809 577 7 . . . 37809 578 1 7 7 LS 37809 578 2 [ [ -LRB- 37809 578 3 Holotype.--FMNH holotype.--fmnh NN 37809 578 4 100063 100063 CD 37809 578 5 ( ( -LRB- 37809 578 6 formerly formerly RB 37809 578 7 EHT EHT NNP 37809 578 8 - - HYPH 37809 578 9 HMS HMS NNP 37809 578 10 6807 6807 CD 37809 578 11 ) ) -RRB- 37809 578 12 , , , 37809 578 13 from from IN 37809 578 14 Huasteca Huasteca NNP 37809 578 15 Canyon Canyon NNP 37809 578 16 , , , 37809 578 17 15 15 CD 37809 578 18 km km NN 37809 578 19 . . . 37809 579 1 W W NNP 37809 579 2 Monterrey Monterrey NNP 37809 579 3 , , , 37809 579 4 Nuevo Nuevo NNP 37809 579 5 León León NNP 37809 579 6 , , , 37809 579 7 México México NNP 37809 579 8 , , , 37809 579 9 680 680 CD 37809 579 10 m. m. NN 37809 579 11 ; ; , 37809 579 12 collected collect VBN 37809 579 13 on on IN 37809 579 14 June June NNP 37809 579 15 20 20 CD 37809 579 16 , , , 37809 579 17 1936 1936 CD 37809 579 18 , , , 37809 579 19 by by IN 37809 579 20 Edward Edward NNP 37809 579 21 H. H. NNP 37809 579 22 Taylor Taylor NNP 37809 579 23 ] ] -RRB- 37809 579 24 . . . 37809 580 1 Smith Smith NNP 37809 580 2 and and CC 37809 580 3 Taylor Taylor NNP 37809 580 4 , , , 37809 580 5 1948:50 1948:50 CD 37809 580 6 - - SYM 37809 580 7 52 52 CD 37809 580 8 . . . 37809 581 1 Martin Martin NNP 37809 581 2 , , , 37809 581 3 1958:48 1958:48 CD 37809 581 4 - - SYM 37809 581 5 50 50 CD 37809 581 6 . . . 37809 582 1 Gorham Gorham NNP 37809 582 2 , , , 37809 582 3 1966:165 1966:165 CD 37809 582 4 . . . 37809 583 1 _ _ NNP 37809 583 2 Diagnosis._--Large diagnosis._--large CD 37809 583 3 frogs frog NNS 37809 583 4 , , , 37809 583 5 males male VBZ 37809 583 6 22.1 22.1 CD 37809 583 7 - - SYM 37809 583 8 33.2 33.2 CD 37809 583 9 mm mm NNP 37809 583 10 . . . 37809 584 1 snout snout NN 37809 584 2 - - HYPH 37809 584 3 vent vent NN 37809 584 4 , , , 37809 584 5 females female VBZ 37809 584 6 26.8 26.8 CD 37809 584 7 - - SYM 37809 584 8 39.6 39.6 CD 37809 584 9 mm mm NNP 37809 584 10 . . . 37809 585 1 snout snout NN 37809 585 2 - - HYPH 37809 585 3 vent vent NN 37809 585 4 length length NN 37809 585 5 ; ; : 37809 585 6 vocal vocal NN 37809 585 7 slits slit NNS 37809 585 8 lacking lack VBG 37809 585 9 in in IN 37809 585 10 males male NNS 37809 585 11 ; ; , 37809 585 12 digital digital JJ 37809 585 13 tips tip NNS 37809 585 14 greatly greatly RB 37809 585 15 expanded expand VBN 37809 585 16 ( ( -LRB- 37809 585 17 more more RBR 37809 585 18 than than IN 37809 585 19 twice twice PDT 37809 585 20 the the DT 37809 585 21 width width NN 37809 585 22 of of IN 37809 585 23 digit digit NN 37809 585 24 ) ) -RRB- 37809 585 25 ; ; : 37809 585 26 first first RB 37809 585 27 finger finger NN 37809 585 28 shorter short JJR 37809 585 29 than than IN 37809 585 30 second second JJ 37809 585 31 ; ; : 37809 585 32 skin skin NN 37809 585 33 of of IN 37809 585 34 dorsum dorsum NN 37809 585 35 pustular pustular NN 37809 585 36 , , , 37809 585 37 that that DT 37809 585 38 of of IN 37809 585 39 venter venter NN 37809 585 40 smooth smooth JJ 37809 585 41 ; ; : 37809 585 42 diameter diameter NN 37809 585 43 of of IN 37809 585 44 tympanum tympanum NN 37809 585 45 in in IN 37809 585 46 males male NNS 37809 585 47 61.1 61.1 CD 37809 585 48 - - SYM 37809 585 49 87.2 87.2 CD 37809 585 50 per per IN 37809 585 51 cent cent NN 37809 585 52 that that DT 37809 585 53 of of IN 37809 585 54 eye eye NN 37809 585 55 , , , 37809 585 56 49.5 49.5 CD 37809 585 57 - - SYM 37809 585 58 72.1 72.1 CD 37809 585 59 per per IN 37809 585 60 cent cent NN 37809 585 61 in in IN 37809 585 62 females female NNS 37809 585 63 ; ; : 37809 585 64 dorsum dorsum NN 37809 585 65 tan tan NN 37809 585 66 with with IN 37809 585 67 large large JJ 37809 585 68 or or CC 37809 585 69 small small JJ 37809 585 70 spots spot NNS 37809 585 71 and and CC 37809 585 72 blotches blotch NNS 37809 585 73 ; ; : 37809 585 74 limbs limb NNS 37809 585 75 banded band VBD 37809 585 76 ; ; : 37809 585 77 interorbital interorbital NNP 37809 585 78 bar bar NN 37809 585 79 or or CC 37809 585 80 triangle triangle VB 37809 585 81 present present JJ 37809 585 82 . . . 37809 586 1 _ _ NNP 37809 586 2 Remarks._--I Remarks._--I NNP 37809 586 3 have have VBP 37809 586 4 applied apply VBN 37809 586 5 Baird Baird NNP 37809 586 6 's 's POS 37809 586 7 _ _ NNP 37809 586 8 Batrachyla Batrachyla NNP 37809 586 9 longipes longipe VBZ 37809 586 10 _ _ IN 37809 586 11 to to IN 37809 586 12 the the DT 37809 586 13 frog frog NN 37809 586 14 Taylor Taylor NNP 37809 586 15 ( ( -LRB- 37809 586 16 1940d 1940d CD 37809 586 17 ) ) -RRB- 37809 586 18 called call VBN 37809 586 19 _ _ NNP 37809 586 20 Syrrhophus Syrrhophus NNP 37809 586 21 latodactylus latodactylus NN 37809 586 22 _ _ NNP 37809 586 23 because because IN 37809 586 24 the the DT 37809 586 25 color color NN 37809 586 26 pattern pattern NN 37809 586 27 ( ( -LRB- 37809 586 28 Fig fig NN 37809 586 29 . . . 37809 587 1 13 13 LS 37809 587 2 ) ) -RRB- 37809 587 3 predominant predominant NN 37809 587 4 in in IN 37809 587 5 the the DT 37809 587 6 southern southern JJ 37809 587 7 part part NN 37809 587 8 of of IN 37809 587 9 the the DT 37809 587 10 range range NN 37809 587 11 agrees agree VBZ 37809 587 12 with with IN 37809 587 13 that that DT 37809 587 14 described describe VBN 37809 587 15 ( ( -LRB- 37809 587 16 figured figure VBN 37809 587 17 ) ) -RRB- 37809 587 18 for for IN 37809 587 19 _ _ NNP 37809 587 20 Batrachyla Batrachyla NNP 37809 587 21 longipes longipe VBZ 37809 587 22 _ _ NNP 37809 587 23 . . . 37809 588 1 The the DT 37809 588 2 color color NN 37809 588 3 pattern pattern NN 37809 588 4 of of IN 37809 588 5 individuals individual NNS 37809 588 6 in in IN 37809 588 7 the the DT 37809 588 8 southern southern JJ 37809 588 9 part part NN 37809 588 10 of of IN 37809 588 11 the the DT 37809 588 12 range range NN 37809 588 13 of of IN 37809 588 14 this this DT 37809 588 15 species species NN 37809 588 16 consists consist VBZ 37809 588 17 of of IN 37809 588 18 large large JJ 37809 588 19 spots spot NNS 37809 588 20 or or CC 37809 588 21 blotches blotch NNS 37809 588 22 , , , 37809 588 23 whereas whereas IN 37809 588 24 in in IN 37809 588 25 the the DT 37809 588 26 northwestern northwestern JJ 37809 588 27 part part NN 37809 588 28 the the DT 37809 588 29 pattern pattern NN 37809 588 30 is be VBZ 37809 588 31 made make VBN 37809 588 32 up up RP 37809 588 33 of of IN 37809 588 34 smaller small JJR 37809 588 35 spots spot NNS 37809 588 36 . . . 37809 589 1 In in IN 37809 589 2 the the DT 37809 589 3 northeastern northeastern JJ 37809 589 4 part part NN 37809 589 5 of of IN 37809 589 6 the the DT 37809 589 7 range range NN 37809 589 8 , , , 37809 589 9 the the DT 37809 589 10 pattern pattern NN 37809 589 11 is be VBZ 37809 589 12 more more RBR 37809 589 13 reduced reduce VBN 37809 589 14 and and CC 37809 589 15 tends tend VBZ 37809 589 16 to to TO 37809 589 17 consist consist VB 37809 589 18 of of IN 37809 589 19 heavy heavy JJ 37809 589 20 flecking flecking NN 37809 589 21 . . . 37809 590 1 The the DT 37809 590 2 interorbital interorbital JJ 37809 590 3 bar bar NN 37809 590 4 is be VBZ 37809 590 5 narrower narrow JJR 37809 590 6 in in IN 37809 590 7 specimens specimen NNS 37809 590 8 from from IN 37809 590 9 Nuevo Nuevo NNP 37809 590 10 León León NNP 37809 590 11 and and CC 37809 590 12 Tamaulipas Tamaulipas NNP 37809 590 13 and and CC 37809 590 14 is be VBZ 37809 590 15 triangular triangular JJ 37809 590 16 in in IN 37809 590 17 specimens specimen NNS 37809 590 18 from from IN 37809 590 19 Hidalgo Hidalgo NNP 37809 590 20 and and CC 37809 590 21 Queretaro Queretaro NNP 37809 590 22 . . . 37809 591 1 The the DT 37809 591 2 status status NN 37809 591 3 of of IN 37809 591 4 the the DT 37809 591 5 name name NN 37809 591 6 _ _ NNP 37809 591 7 Batrachyla Batrachyla NNP 37809 591 8 longipes longipe VBZ 37809 591 9 _ _ NNP 37809 591 10 is be VBZ 37809 591 11 currently currently RB 37809 591 12 that that DT 37809 591 13 of of IN 37809 591 14 a a DT 37809 591 15 _ _ NNP 37809 591 16 nomen nomen NNP 37809 591 17 dubium dubium NN 37809 591 18 _ _ NNP 37809 591 19 ( ( -LRB- 37809 591 20 Lynch Lynch NNP 37809 591 21 , , , 37809 591 22 1963 1963 CD 37809 591 23 ) ) -RRB- 37809 591 24 . . . 37809 592 1 At at IN 37809 592 2 that that DT 37809 592 3 time time NN 37809 592 4 , , , 37809 592 5 I -PRON- PRP 37809 592 6 was be VBD 37809 592 7 unaware unaware JJ 37809 592 8 of of IN 37809 592 9 the the DT 37809 592 10 geographic geographic JJ 37809 592 11 variation variation NN 37809 592 12 in in IN 37809 592 13 color color NN 37809 592 14 pattern pattern NN 37809 592 15 in in IN 37809 592 16 _ _ NNP 37809 592 17 Syrrhophus Syrrhophus NNP 37809 592 18 latodactylus latodactylus NN 37809 592 19 _ _ NNP 37809 592 20 . . . 37809 593 1 The the DT 37809 593 2 exact exact JJ 37809 593 3 type type NN 37809 593 4 - - HYPH 37809 593 5 locality locality NN 37809 593 6 of of IN 37809 593 7 _ _ NNP 37809 593 8 Batrachyla Batrachyla NNP 37809 593 9 longipes longipe VBZ 37809 593 10 _ _ NNP 37809 593 11 is be VBZ 37809 593 12 not not RB 37809 593 13 known know VBN 37809 593 14 . . . 37809 594 1 If if IN 37809 594 2 it -PRON- PRP 37809 594 3 is be VBZ 37809 594 4 40 40 CD 37809 594 5 Leagues Leagues NNPS 37809 594 6 north north RB 37809 594 7 of of IN 37809 594 8 México México NNP 37809 594 9 City City NNP 37809 594 10 , , , 37809 594 11 the the DT 37809 594 12 locality locality NN 37809 594 13 would would MD 37809 594 14 be be VB 37809 594 15 in in IN 37809 594 16 an an DT 37809 594 17 area area NN 37809 594 18 where where WRB 37809 594 19 the the DT 37809 594 20 species specie NNS 37809 594 21 has have VBZ 37809 594 22 a a DT 37809 594 23 blotched blotched JJ 37809 594 24 instead instead RB 37809 594 25 of of IN 37809 594 26 a a DT 37809 594 27 flecked flecked JJ 37809 594 28 or or CC 37809 594 29 spotted spotted JJ 37809 594 30 pattern pattern NN 37809 594 31 . . . 37809 595 1 No no DT 37809 595 2 justifiable justifiable JJ 37809 595 3 evidence evidence NN 37809 595 4 was be VBD 37809 595 5 presented present VBN 37809 595 6 to to TO 37809 595 7 place place VB 37809 595 8 _ _ NNP 37809 595 9 Batrachyla Batrachyla NNP 37809 595 10 longipes longipe VBZ 37809 595 11 _ _ NNP 37809 595 12 in in IN 37809 595 13 _ _ NNP 37809 595 14 Eleutherodactylus Eleutherodactylus NNP 37809 595 15 _ _ NNP 37809 595 16 instead instead RB 37809 595 17 of of IN 37809 595 18 _ _ NNP 37809 595 19 Syrrhophus Syrrhophus NNP 37809 595 20 _ _ NNP 37809 595 21 . . . 37809 596 1 Barbour Barbour NNP 37809 596 2 ( ( -LRB- 37809 596 3 1923 1923 CD 37809 596 4 ) ) -RRB- 37809 596 5 and and CC 37809 596 6 Kellogg Kellogg NNP 37809 596 7 ( ( -LRB- 37809 596 8 1932 1932 CD 37809 596 9 ) ) -RRB- 37809 596 10 associated associate VBD 37809 596 11 another another DT 37809 596 12 species specie NNS 37809 596 13 ( ( -LRB- 37809 596 14 _ _ NNP 37809 596 15 E. E. NNP 37809 597 1 batrachylus batrachylus NNP 37809 597 2 _ _ NNP 37809 597 3 ) ) -RRB- 37809 597 4 with with IN 37809 597 5 _ _ NNP 37809 597 6 longipes longipes NNPS 37809 597 7 _ _ NNP 37809 597 8 . . . 37809 598 1 Taylor Taylor NNP 37809 598 2 ( ( -LRB- 37809 598 3 1940a 1940a CD 37809 598 4 ) ) -RRB- 37809 598 5 noted note VBD 37809 598 6 this this DT 37809 598 7 as as IN 37809 598 8 a a DT 37809 598 9 case case NN 37809 598 10 of of IN 37809 598 11 misidentification misidentification NN 37809 598 12 and and CC 37809 598 13 corrected correct VBD 37809 598 14 the the DT 37809 598 15 error error NN 37809 598 16 but but CC 37809 598 17 left leave VBD 37809 598 18 _ _ NNP 37809 598 19 longipes longipe NNS 37809 598 20 _ _ NNP 37809 598 21 in in IN 37809 598 22 the the DT 37809 598 23 genus genus NNP 37809 598 24 _ _ NNP 37809 598 25 Eleutherodactylus Eleutherodactylus NNP 37809 598 26 _ _ NNP 37809 598 27 . . . 37809 599 1 Lynch Lynch NNP 37809 599 2 ( ( -LRB- 37809 599 3 1963 1963 CD 37809 599 4 ) ) -RRB- 37809 599 5 noted note VBD 37809 599 6 several several JJ 37809 599 7 points point NNS 37809 599 8 of of IN 37809 599 9 morphological morphological JJ 37809 599 10 agreement agreement NN 37809 599 11 between between IN 37809 599 12 _ _ NNP 37809 599 13 Syrrhophus Syrrhophus NNP 37809 599 14 _ _ NNP 37809 599 15 and and CC 37809 599 16 _ _ NNP 37809 599 17 B. B. NNP 37809 600 1 longipes longipes NNPS 37809 600 2 _ _ NNP 37809 600 3 but but CC 37809 600 4 did do VBD 37809 600 5 not not RB 37809 600 6 place place VB 37809 600 7 _ _ NNP 37809 600 8 longipes longipe NNS 37809 600 9 _ _ NNP 37809 600 10 in in IN 37809 600 11 _ _ NNP 37809 600 12 Syrrhophus Syrrhophus NNP 37809 600 13 _ _ NNP 37809 600 14 . . . 37809 601 1 Baird Baird NNP 37809 601 2 's 's POS 37809 601 3 ( ( -LRB- 37809 601 4 1859 1859 CD 37809 601 5 ) ) -RRB- 37809 601 6 figures figure NNS 37809 601 7 of of IN 37809 601 8 the the DT 37809 601 9 holotype holotype NN 37809 601 10 do do VBP 37809 601 11 not not RB 37809 601 12 illustrate illustrate VB 37809 601 13 prevomerine prevomerine JJ 37809 601 14 teeth tooth NNS 37809 601 15 , , , 37809 601 16 but but CC 37809 601 17 according accord VBG 37809 601 18 to to IN 37809 601 19 Cope Cope NNP 37809 601 20 ( ( -LRB- 37809 601 21 1866 1866 CD 37809 601 22 ) ) -RRB- 37809 601 23 they -PRON- PRP 37809 601 24 were be VBD 37809 601 25 present present JJ 37809 601 26 in in IN 37809 601 27 the the DT 37809 601 28 holotype holotype NN 37809 601 29 . . . 37809 602 1 The the DT 37809 602 2 digital digital JJ 37809 602 3 tips tip NNS 37809 602 4 of of IN 37809 602 5 the the DT 37809 602 6 frog frog NN 37809 602 7 in in IN 37809 602 8 the the DT 37809 602 9 figure figure NN 37809 602 10 are be VBP 37809 602 11 somewhat somewhat RB 37809 602 12 narrower narrow JJR 37809 602 13 than than IN 37809 602 14 those those DT 37809 602 15 typically typically RB 37809 602 16 seen see VBN 37809 602 17 in in IN 37809 602 18 _ _ NNP 37809 602 19 S. S. NNP 37809 603 1 latodactylus latodactylus NNP 37809 603 2 _ _ NNP 37809 603 3 . . . 37809 604 1 If if IN 37809 604 2 the the DT 37809 604 3 specimen speciman NNS 37809 604 4 was be VBD 37809 604 5 slightly slightly RB 37809 604 6 desiccated desiccate VBN 37809 604 7 , , , 37809 604 8 as as IN 37809 604 9 possibly possibly RB 37809 604 10 was be VBD 37809 604 11 the the DT 37809 604 12 case case NN 37809 604 13 , , , 37809 604 14 the the DT 37809 604 15 digits digit NNS 37809 604 16 would would MD 37809 604 17 appear appear VB 37809 604 18 narrower narrow JJR 37809 604 19 . . . 37809 605 1 There there EX 37809 605 2 is be VBZ 37809 605 3 no no DT 37809 605 4 evidence evidence NN 37809 605 5 contrary contrary JJ 37809 605 6 to to IN 37809 605 7 placing place VBG 37809 605 8 _ _ NNP 37809 605 9 Syrrhophus Syrrhophus NNP 37809 605 10 latodactylus latodactylus NN 37809 605 11 _ _ NNP 37809 605 12 in in IN 37809 605 13 the the DT 37809 605 14 synonymy synonymy NN 37809 605 15 of of IN 37809 605 16 _ _ NNP 37809 605 17 Batrachyla Batrachyla NNP 37809 605 18 longipes longipe VBZ 37809 605 19 _ _ NNP 37809 605 20 . . . 37809 606 1 [ [ -LRB- 37809 606 2 Illustration illustration NN 37809 606 3 : : : 37809 606 4 FIG FIG NNP 37809 606 5 . . . 37809 607 1 13 13 CD 37809 607 2 : : : 37809 607 3 Dorsal dorsal NN 37809 607 4 views view NNS 37809 607 5 of of IN 37809 607 6 _ _ NNP 37809 607 7 Syrrhophus Syrrhophus NNP 37809 607 8 longipes longipe VBZ 37809 607 9 _ _ NNP 37809 607 10 illustrating illustrate VBG 37809 607 11 geographic geographic JJ 37809 607 12 variation variation NN 37809 607 13 in in IN 37809 607 14 pattern pattern NN 37809 607 15 ( ( -LRB- 37809 607 16 left leave VBN 37809 607 17 , , , 37809 607 18 TCWC TCWC NNP 37809 607 19 12179 12179 CD 37809 607 20 , , , 37809 607 21 ×1.5 ×1.5 NN 37809 607 22 ; ; : 37809 607 23 right right UH 37809 607 24 , , , 37809 607 25 KU KU NNP 37809 607 26 92572 92572 CD 37809 607 27 , , , 37809 607 28 ×1.8 ×1.8 XX 37809 607 29 ) ) -RRB- 37809 607 30 ; ; : 37809 607 31 side side NN 37809 607 32 of of IN 37809 607 33 head head NN 37809 607 34 ( ( -LRB- 37809 607 35 TCWC TCWC NNP 37809 607 36 10966 10966 CD 37809 607 37 , , , 37809 607 38 ×6 ×6 XX 37809 607 39 ) ) -RRB- 37809 607 40 . . . 37809 607 41 ] ] -RRB- 37809 608 1 Application application NN 37809 608 2 of of IN 37809 608 3 Baird Baird NNP 37809 608 4 's 's POS 37809 608 5 name name NN 37809 608 6 _ _ NNP 37809 608 7 Batrachyla Batrachyla NNP 37809 608 8 longipes longipe VBZ 37809 608 9 _ _ IN 37809 608 10 to to IN 37809 608 11 the the DT 37809 608 12 species species NN 37809 608 13 of of IN 37809 608 14 frog frog NNP 37809 608 15 heretofore heretofore RB 37809 608 16 called call VBN 37809 608 17 _ _ NNP 37809 608 18 Syrrhophus Syrrhophus NNP 37809 608 19 latodactylus latodactylus NN 37809 608 20 _ _ NNP 37809 608 21 poses pose VBZ 37809 608 22 one one CD 37809 608 23 serious serious JJ 37809 608 24 problem problem NN 37809 608 25 . . . 37809 609 1 _ _ NNP 37809 609 2 Batrachyla Batrachyla NNP 37809 609 3 longipes longipe VBZ 37809 609 4 _ _ NNP 37809 609 5 is be VBZ 37809 609 6 the the DT 37809 609 7 type type NN 37809 609 8 - - HYPH 37809 609 9 species species NN 37809 609 10 ( ( -LRB- 37809 609 11 by by IN 37809 609 12 original original JJ 37809 609 13 designation designation NN 37809 609 14 ) ) -RRB- 37809 609 15 of of IN 37809 609 16 the the DT 37809 609 17 genus genus NNP 37809 609 18 _ _ NNP 37809 609 19 Epirhexis Epirhexis NNP 37809 609 20 _ _ NNP 37809 609 21 Cope Cope NNP 37809 609 22 , , , 37809 609 23 1866 1866 CD 37809 609 24 , , , 37809 609 25 which which WDT 37809 609 26 has have VBZ 37809 609 27 priority priority NN 37809 609 28 over over IN 37809 609 29 _ _ NNP 37809 609 30 Syrrhophus Syrrhophus NNP 37809 609 31 _ _ NNP 37809 609 32 Cope Cope NNP 37809 609 33 , , , 37809 609 34 1878 1878 CD 37809 609 35 . . . 37809 610 1 If if IN 37809 610 2 _ _ NNP 37809 610 3 Batrachyla Batrachyla NNP 37809 610 4 longipes longipe VBZ 37809 610 5 _ _ NNP 37809 610 6 is be VBZ 37809 610 7 left leave VBN 37809 610 8 in in IN 37809 610 9 the the DT 37809 610 10 status status NN 37809 610 11 of of IN 37809 610 12 a a DT 37809 610 13 _ _ NNP 37809 610 14 nomen nomen NNP 37809 610 15 dubium dubium NN 37809 610 16 _ _ NNP 37809 610 17 , , , 37809 610 18 _ _ NNP 37809 610 19 Epirhexis Epirhexis NNP 37809 610 20 _ _ NNP 37809 610 21 can can MD 37809 610 22 be be VB 37809 610 23 forgotten forget VBN 37809 610 24 , , , 37809 610 25 for for IN 37809 610 26 the the DT 37809 610 27 two two CD 37809 610 28 names name NNS 37809 610 29 are be VBP 37809 610 30 tied tie VBN 37809 610 31 together together RB 37809 610 32 . . . 37809 611 1 However however RB 37809 611 2 , , , 37809 611 3 since since IN 37809 611 4 it -PRON- PRP 37809 611 5 seems seem VBZ 37809 611 6 almost almost RB 37809 611 7 certain certain JJ 37809 611 8 that that IN 37809 611 9 _ _ NNP 37809 611 10 Batrachyla Batrachyla NNP 37809 611 11 longipes longipe VBZ 37809 611 12 _ _ NNP 37809 611 13 and and CC 37809 611 14 _ _ NNP 37809 611 15 Syrrhophus Syrrhophus NNP 37809 611 16 latodactylus latodactylus NN 37809 611 17 _ _ NNP 37809 611 18 are be VBP 37809 611 19 conspecific conspecific NN 37809 611 20 , , , 37809 611 21 the the DT 37809 611 22 former former JJ 37809 611 23 name name NN 37809 611 24 should should MD 37809 611 25 not not RB 37809 611 26 be be VB 37809 611 27 left leave VBN 37809 611 28 as as IN 37809 611 29 a a DT 37809 611 30 _ _ NNP 37809 611 31 nomen nomen NNP 37809 611 32 dubium dubium NN 37809 611 33 _ _ NNP 37809 611 34 . . . 37809 612 1 _ _ NNP 37809 612 2 Epirhexis Epirhexis NNP 37809 612 3 _ _ NNP 37809 612 4 never never RB 37809 612 5 came come VBD 37809 612 6 into into IN 37809 612 7 general general JJ 37809 612 8 usage usage NN 37809 612 9 ( ( -LRB- 37809 612 10 Cope Cope NNP 37809 612 11 cited cite VBD 37809 612 12 the the DT 37809 612 13 name name NN 37809 612 14 four four CD 37809 612 15 times time NNS 37809 612 16 , , , 37809 612 17 but but CC 37809 612 18 no no DT 37809 612 19 one one NN 37809 612 20 else else RB 37809 612 21 has have VBZ 37809 612 22 used use VBN 37809 612 23 it -PRON- PRP 37809 612 24 ) ) -RRB- 37809 612 25 , , , 37809 612 26 whereas whereas IN 37809 612 27 _ _ NNP 37809 612 28 Syrrhophus Syrrhophus NNP 37809 612 29 _ _ NNP 37809 612 30 is be VBZ 37809 612 31 well well RB 37809 612 32 established establish VBN 37809 612 33 in in IN 37809 612 34 the the DT 37809 612 35 zoological zoological JJ 37809 612 36 literature literature NN 37809 612 37 . . . 37809 613 1 It -PRON- PRP 37809 613 2 would would MD 37809 613 3 serve serve VB 37809 613 4 only only RB 37809 613 5 to to TO 37809 613 6 confuse confuse VB 37809 613 7 the the DT 37809 613 8 literature literature NN 37809 613 9 to to TO 37809 613 10 adhere adhere VB 37809 613 11 strictly strictly RB 37809 613 12 to to IN 37809 613 13 the the DT 37809 613 14 Law Law NNP 37809 613 15 of of IN 37809 613 16 Priority Priority NNP 37809 613 17 and and CC 37809 613 18 replace replace VB 37809 613 19 _ _ NNP 37809 613 20 Syrrhophus Syrrhophus NNP 37809 613 21 _ _ NNP 37809 613 22 with with IN 37809 613 23 _ _ NNP 37809 613 24 Epirhexis Epirhexis NNP 37809 613 25 _ _ NNP 37809 613 26 . . . 37809 614 1 Therefore therefore RB 37809 614 2 , , , 37809 614 3 _ _ NNP 37809 614 4 Syrrhophus Syrrhophus NNP 37809 614 5 _ _ NNP 37809 614 6 is be VBZ 37809 614 7 used use VBN 37809 614 8 in in IN 37809 614 9 this this DT 37809 614 10 paper paper NN 37809 614 11 , , , 37809 614 12 even even RB 37809 614 13 though though IN 37809 614 14 _ _ NNP 37809 614 15 Epirhexis Epirhexis NNP 37809 614 16 _ _ NNP 37809 614 17 has have VBZ 37809 614 18 priority priority NN 37809 614 19 . . . 37809 615 1 A a DT 37809 615 2 request request NN 37809 615 3 for for IN 37809 615 4 the the DT 37809 615 5 suppression suppression NN 37809 615 6 of of IN 37809 615 7 _ _ NNP 37809 615 8 Epirhexis Epirhexis NNP 37809 615 9 _ _ NNP 37809 615 10 Cope Cope NNP 37809 615 11 , , , 37809 615 12 1866 1866 CD 37809 615 13 , , , 37809 615 14 has have VBZ 37809 615 15 been be VBN 37809 615 16 submitted submit VBN 37809 615 17 to to IN 37809 615 18 the the DT 37809 615 19 International International NNP 37809 615 20 Commission Commission NNP 37809 615 21 of of IN 37809 615 22 Zoological Zoological NNP 37809 615 23 Nomenclature Nomenclature NNP 37809 615 24 ( ( -LRB- 37809 615 25 Lynch Lynch NNP 37809 615 26 , , , 37809 615 27 1967 1967 CD 37809 615 28 ) ) -RRB- 37809 615 29 . . . 37809 616 1 _ _ NNP 37809 616 2 Etymology._--Latin Etymology._--Latin NNP 37809 616 3 , , , 37809 616 4 meaning mean VBG 37809 616 5 long long RB 37809 616 6 - - HYPH 37809 616 7 footed footed JJ 37809 616 8 ; ; : 37809 616 9 Taylor Taylor NNP 37809 616 10 's 's POS 37809 616 11 _ _ NNP 37809 616 12 latodactylus latodactylus NN 37809 616 13 _ _ NNP 37809 616 14 refers refer VBZ 37809 616 15 to to IN 37809 616 16 the the DT 37809 616 17 wide wide JJ 37809 616 18 digital digital JJ 37809 616 19 pads pad NNS 37809 616 20 . . . 37809 617 1 [ [ -LRB- 37809 617 2 Illustration illustration NN 37809 617 3 : : : 37809 617 4 FIG FIG NNP 37809 617 5 . . . 37809 618 1 14 14 CD 37809 618 2 : : : 37809 618 3 Distribution distribution NN 37809 618 4 of of IN 37809 618 5 _ _ NNP 37809 618 6 Syrrhophus Syrrhophus NNP 37809 618 7 dennisi dennisi RB 37809 618 8 _ _ NNP 37809 618 9 ( ( -LRB- 37809 618 10 triangle triangle NNP 37809 618 11 ) ) -RRB- 37809 618 12 and and CC 37809 618 13 _ _ NNP 37809 618 14 S. S. NNP 37809 619 1 longipes longipes NNPS 37809 619 2 _ _ NNP 37809 619 3 ( ( -LRB- 37809 619 4 circles circles NNP 37809 619 5 ) ) -RRB- 37809 619 6 . . . 37809 619 7 ] ] -RRB- 37809 620 1 _ _ NNP 37809 620 2 Distribution._--Moderate distribution._--moderate JJ 37809 620 3 elevations elevation NNS 37809 620 4 ( ( -LRB- 37809 620 5 650 650 CD 37809 620 6 to to TO 37809 620 7 2000 2000 CD 37809 620 8 meters meter NNS 37809 620 9 ) ) -RRB- 37809 620 10 along along IN 37809 620 11 the the DT 37809 620 12 Sierra Sierra NNP 37809 620 13 Madre Madre NNP 37809 620 14 Oriental Oriental NNP 37809 620 15 from from IN 37809 620 16 central central JJ 37809 620 17 Nuevo Nuevo NNP 37809 620 18 León León NNP 37809 620 19 to to IN 37809 620 20 northern northern JJ 37809 620 21 Hidalgo Hidalgo NNP 37809 620 22 , , , 37809 620 23 México México NNP 37809 620 24 ( ( -LRB- 37809 620 25 Fig Fig NNP 37809 620 26 . . . 37809 621 1 14 14 CD 37809 621 2 ) ) -RRB- 37809 621 3 . . . 37809 622 1 _ _ NNP 37809 622 2 Specimens Specimens NNP 37809 622 3 examined._--(122 examined._--(122 NNP 37809 622 4 ) ) -RRB- 37809 622 5 MÉXICO MÉXICO NNP 37809 622 6 , , , 37809 622 7 _ _ NNP 37809 622 8 Hidalgo Hidalgo NNP 37809 622 9 _ _ NNP 37809 622 10 : : : 37809 622 11 3 3 CD 37809 622 12 km km NN 37809 622 13 . . . 37809 623 1 NE NE NNP 37809 623 2 Jacala Jacala NNP 37809 623 3 , , , 37809 623 4 AMNH AMNH NNP 37809 623 5 52977 52977 CD 37809 623 6 ; ; : 37809 623 7 9.6 9.6 CD 37809 623 8 km km NN 37809 623 9 . . . 37809 624 1 NE NE NNP 37809 624 2 Jacala Jacala NNP 37809 624 3 , , , 37809 624 4 1800 1800 CD 37809 624 5 m. m. NN 37809 624 6 , , , 37809 624 7 TCWC TCWC NNP 37809 624 8 10966 10966 CD 37809 624 9 - - SYM 37809 624 10 70 70 CD 37809 624 11 , , , 37809 624 12 12179 12179 CD 37809 624 13 ; ; : 37809 624 14 8 8 CD 37809 624 15 km km NN 37809 624 16 . . . 37809 625 1 S S NNP 37809 625 2 Jacala Jacala NNP 37809 625 3 , , , 37809 625 4 La La NNP 37809 625 5 Placita Placita NNP 37809 625 6 , , , 37809 625 7 1850 1850 CD 37809 625 8 m. m. NN 37809 625 9 , , , 37809 625 10 FMNH FMNH NNP 37809 625 11 100266 100266 CD 37809 625 12 - - SYM 37809 625 13 68 68 CD 37809 625 14 , , , 37809 625 15 103244 103244 CD 37809 625 16 , , , 37809 625 17 UIMNH UIMNH NNP 37809 625 18 13291 13291 CD 37809 625 19 , , , 37809 625 20 13327 13327 CD 37809 625 21 . . . 37809 626 1 _ _ NNP 37809 626 2 Nuevo Nuevo NNP 37809 626 3 León León NNP 37809 626 4 _ _ NNP 37809 626 5 : : : 37809 626 6 Salto Salto NNP 37809 626 7 Cola Cola NNP 37809 626 8 de de NNP 37809 626 9 Caballo Caballo NNP 37809 626 10 , , , 37809 626 11 KU KU NNP 37809 626 12 92572 92572 CD 37809 626 13 ; ; : 37809 626 14 Huasteca Huasteca NNP 37809 626 15 Canyon Canyon NNP 37809 626 16 , , , 37809 626 17 15 15 CD 37809 626 18 km km NN 37809 626 19 . . . 37809 627 1 W W NNP 37809 627 2 Monterrey Monterrey NNP 37809 627 3 , , , 37809 627 4 680 680 CD 37809 627 5 m. m. NN 37809 627 6 , , , 37809 627 7 FMNH FMNH NNP 37809 627 8 100063 100063 CD 37809 627 9 ( ( -LRB- 37809 627 10 holotype holotype NN 37809 627 11 of of IN 37809 627 12 _ _ NNP 37809 627 13 S. S. NNP 37809 628 1 latodactylus latodactylus NNP 37809 628 2 _ _ NNP 37809 628 3 ) ) -RRB- 37809 628 4 , , , 37809 628 5 UIMNH UIMNH NNP 37809 628 6 13290 13290 CD 37809 628 7 ; ; : 37809 628 8 6.5 6.5 CD 37809 628 9 km km NN 37809 628 10 . . . 37809 629 1 N n CD 37809 629 2 Pablillo Pablillo NNP 37809 629 3 , , , 37809 629 4 EAL EAL NNP 37809 629 5 1319 1319 CD 37809 629 6 ; ; : 37809 629 7 Sabinas Sabinas NNP 37809 629 8 Hidalgo Hidalgo NNP 37809 629 9 , , , 37809 629 10 USNM USNM NNP 37809 629 11 139728 139728 CD 37809 629 12 . . . 37809 630 1 _ _ NNP 37809 630 2 Queretaro Queretaro NNP 37809 630 3 _ _ NNP 37809 630 4 : : : 37809 630 5 Cueva Cueva NNP 37809 630 6 de de NNP 37809 630 7 los los NNP 37809 630 8 Riscos Riscos NNP 37809 630 9 , , , 37809 630 10 8 8 CD 37809 630 11 km km NN 37809 630 12 . . . 37809 631 1 SW SW NNP 37809 631 2 Jalpan Jalpan NNP 37809 631 3 , , , 37809 631 4 KU KU NNP 37809 631 5 106300 106300 CD 37809 631 6 . . . 37809 632 1 _ _ NNP 37809 632 2 San San NNP 37809 632 3 Luis Luis NNP 37809 632 4 Potosí Potosí NNP 37809 632 5 _ _ NNP 37809 632 6 : : : 37809 632 7 13 13 CD 37809 632 8 km km NN 37809 632 9 . . . 37809 633 1 E E NNP 37809 633 2 Santa Santa NNP 37809 633 3 Barberita Barberita NNP 37809 633 4 , , , 37809 633 5 LSUMZ LSUMZ NNP 37809 633 6 2295 2295 CD 37809 633 7 ; ; : 37809 633 8 second second JJ 37809 633 9 camp camp NN 37809 633 10 , , , 37809 633 11 San San NNP 37809 633 12 Luis Luis NNP 37809 633 13 Potosí Potosí NNP 37809 633 14 road road NN 37809 633 15 , , , 37809 633 16 UIMNH UIMNH NNP 37809 633 17 13326 13326 CD 37809 633 18 ; ; : 37809 633 19 Xilitla Xilitla NNP 37809 633 20 , , , 37809 633 21 Cueva Cueva NNP 37809 633 22 sin sin NN 37809 633 23 nombre nombre NNP 37809 633 24 , , , 37809 633 25 UMMZ UMMZ NNP 37809 633 26 125892 125892 CD 37809 633 27 . . . 37809 634 1 _ _ NNP 37809 634 2 Tamaulipas Tamaulipas NNP 37809 634 3 _ _ NNP 37809 634 4 : : : 37809 634 5 4 4 CD 37809 634 6 km km NN 37809 634 7 . . . 37809 635 1 W W NNP 37809 635 2 El El NNP 37809 635 3 Carrizo Carrizo NNP 37809 635 4 , , , 37809 635 5 500 500 CD 37809 635 6 m. m. NN 37809 635 7 , , , 37809 635 8 UMMZ UMMZ NNP 37809 635 9 111343 111343 CD 37809 635 10 ( ( -LRB- 37809 635 11 31 31 CD 37809 635 12 ) ) -RRB- 37809 635 13 ; ; : 37809 635 14 8 8 CD 37809 635 15 km km NN 37809 635 16 . . . 37809 636 1 N N NNP 37809 636 2 Chamal Chamal NNP 37809 636 3 , , , 37809 636 4 Bee Bee NNP 37809 636 5 Cave Cave NNP 37809 636 6 , , , 37809 636 7 KU KU NNP 37809 636 8 106299 106299 CD 37809 636 9 ; ; : 37809 636 10 14.5 14.5 CD 37809 636 11 km km NNS 37809 636 12 . . . 37809 637 1 NNW NNW NNP 37809 637 2 Chamal Chamal NNP 37809 637 3 , , , 37809 637 4 420 420 CD 37809 637 5 m. m. NN 37809 637 6 , , , 37809 637 7 UMMZ UMMZ NNP 37809 637 8 111339 111339 CD 37809 637 9 - - SYM 37809 637 10 40 40 CD 37809 637 11 , , , 37809 637 12 111342 111342 CD 37809 637 13 ( ( -LRB- 37809 637 14 4 4 CD 37809 637 15 ) ) -RRB- 37809 637 16 , , , 37809 637 17 111344 111344 CD 37809 637 18 ( ( -LRB- 37809 637 19 11 11 CD 37809 637 20 ) ) -RRB- 37809 637 21 ; ; : 37809 637 22 19 19 CD 37809 637 23 km km NN 37809 637 24 . . . 37809 638 1 NNW NNW NNP 37809 638 2 Chamal Chamal NNP 37809 638 3 , , , 37809 638 4 700 700 CD 37809 638 5 m. m. NN 37809 638 6 , , , 37809 638 7 UMMZ UMMZ NNP 37809 638 8 111341 111341 CD 37809 638 9 ( ( -LRB- 37809 638 10 3 3 CD 37809 638 11 ) ) -RRB- 37809 638 12 ; ; : 37809 638 13 El El NNP 37809 638 14 Chihue Chihue NNP 37809 638 15 , , , 37809 638 16 1880 1880 CD 37809 638 17 m. m. NN 37809 638 18 , , , 37809 638 19 UMMZ UMMZ NNP 37809 638 20 111289 111289 CD 37809 638 21 ( ( -LRB- 37809 638 22 4 4 CD 37809 638 23 ) ) -RRB- 37809 638 24 ; ; : 37809 638 25 11 11 CD 37809 638 26 km km NN 37809 638 27 . . . 37809 639 1 N n CD 37809 639 2 Gómez Gómez NNP 37809 639 3 Farías Farías NNP 37809 639 4 , , , 37809 639 5 1060 1060 CD 37809 639 6 m. m. NN 37809 639 7 , , , 37809 639 8 UMMZ UMMZ NNP 37809 639 9 101166 101166 CD 37809 639 10 ; ; : 37809 639 11 11 11 CD 37809 639 12 km km NN 37809 639 13 . . . 37809 640 1 WNW WNW NNP 37809 640 2 Gómez Gómez NNP 37809 640 3 Farías Farías NNP 37809 640 4 , , , 37809 640 5 1800 1800 CD 37809 640 6 m. m. NN 37809 640 7 , , , 37809 640 8 UMMZ UMMZ NNP 37809 640 9 108507 108507 CD 37809 640 10 ( ( -LRB- 37809 640 11 3 3 CD 37809 640 12 ) ) -RRB- 37809 640 13 ; ; : 37809 640 14 8 8 CD 37809 640 15 km km NN 37809 640 16 . . . 37809 641 1 NW NW NNP 37809 641 2 Gómez Gómez NNP 37809 641 3 Farías Farías NNP 37809 641 4 , , , 37809 641 5 1060 1060 CD 37809 641 6 - - SYM 37809 641 7 1400 1400 CD 37809 641 8 m. m. NN 37809 641 9 , , , 37809 641 10 LSUMZ LSUMZ NNP 37809 641 11 11085 11085 CD 37809 641 12 , , , 37809 641 13 UMMZ UMMZ NNP 37809 641 14 101167 101167 CD 37809 641 15 ( ( -LRB- 37809 641 16 3 3 CD 37809 641 17 ) ) -RRB- 37809 641 18 , , , 37809 641 19 101168 101168 CD 37809 641 20 ( ( -LRB- 37809 641 21 4 4 CD 37809 641 22 ) ) -RRB- 37809 641 23 , , , 37809 641 24 101169 101169 CD 37809 641 25 ( ( -LRB- 37809 641 26 2 2 CD 37809 641 27 ) ) -RRB- 37809 641 28 , , , 37809 641 29 101170 101170 CD 37809 641 30 ( ( -LRB- 37809 641 31 3 3 CD 37809 641 32 ) ) -RRB- 37809 641 33 , , , 37809 641 34 101171 101171 CD 37809 641 35 ( ( -LRB- 37809 641 36 2 2 CD 37809 641 37 ) ) -RRB- 37809 641 38 , , , 37809 641 39 101360 101360 CD 37809 641 40 - - SYM 37809 641 41 61 61 CD 37809 641 42 , , , 37809 641 43 102860 102860 CD 37809 641 44 , , , 37809 641 45 102933 102933 CD 37809 641 46 ( ( -LRB- 37809 641 47 4 4 CD 37809 641 48 ) ) -RRB- 37809 641 49 , , , 37809 641 50 102934 102934 CD 37809 641 51 ( ( -LRB- 37809 641 52 2 2 CD 37809 641 53 ) ) -RRB- 37809 641 54 , , , 37809 641 55 102935 102935 CD 37809 641 56 - - SYM 37809 641 57 38 38 CD 37809 641 58 , , , 37809 641 59 102939 102939 CD 37809 641 60 ( ( -LRB- 37809 641 61 2 2 CD 37809 641 62 ) ) -RRB- 37809 641 63 , , , 37809 641 64 102940 102940 CD 37809 641 65 - - SYM 37809 641 66 43 43 CD 37809 641 67 , , , 37809 641 68 108800 108800 CD 37809 641 69 ( ( -LRB- 37809 641 70 3 3 CD 37809 641 71 ) ) -RRB- 37809 641 72 , , , 37809 641 73 110735 110735 CD 37809 641 74 , , , 37809 641 75 111345 111345 CD 37809 641 76 - - SYM 37809 641 77 46 46 CD 37809 641 78 . . . 37809 642 1 = = NFP 37809 642 2 Syrrhophus Syrrhophus NNP 37809 642 3 pipilans= pipilans= NN 37809 642 4 Taylor Taylor NNP 37809 642 5 _ _ NNP 37809 642 6 Syrrhophus Syrrhophus NNP 37809 642 7 pipilans pipilan NNS 37809 642 8 _ _ NNP 37809 642 9 Taylor Taylor NNP 37809 642 10 , , , 37809 642 11 1940c:95 1940c:95 CD 37809 642 12 - - SYM 37809 642 13 97 97 CD 37809 642 14 , , , 37809 642 15 pl pl NNP 37809 642 16 . . . 37809 643 1 1 1 LS 37809 643 2 [ [ -LRB- 37809 643 3 Holotype.--FMNH holotype.--fmnh NN 37809 643 4 100072 100072 CD 37809 643 5 ( ( -LRB- 37809 643 6 formerly formerly RB 37809 643 7 EHT EHT NNP 37809 643 8 - - HYPH 37809 643 9 HMS HMS NNP 37809 643 10 6843 6843 CD 37809 643 11 ) ) -RRB- 37809 643 12 , , , 37809 643 13 14.6 14.6 CD 37809 643 14 km km NNS 37809 643 15 . . . 37809 644 1 S S NNP 37809 644 2 Mazatlán Mazatlán NNP 37809 644 3 , , , 37809 644 4 Guerrero Guerrero NNP 37809 644 5 , , , 37809 644 6 México México NNP 37809 644 7 ; ; , 37809 644 8 collected collect VBN 37809 644 9 on on IN 37809 644 10 July July NNP 37809 644 11 22 22 CD 37809 644 12 , , , 37809 644 13 1936 1936 CD 37809 644 14 , , , 37809 644 15 by by IN 37809 644 16 Edward Edward NNP 37809 644 17 H. H. NNP 37809 644 18 Taylor Taylor NNP 37809 644 19 ] ] -RRB- 37809 644 20 . . . 37809 645 1 _ _ NNP 37809 645 2 Diagnosis._--Medium Diagnosis._--Medium NNP 37809 645 3 sized sized JJ 37809 645 4 frogs frog NNS 37809 645 5 , , , 37809 645 6 males male VBZ 37809 645 7 22.6 22.6 CD 37809 645 8 - - HYPH 37809 645 9 28.5 28.5 CD 37809 645 10 mm mm NNP 37809 645 11 . . . 37809 646 1 snout snout NN 37809 646 2 - - HYPH 37809 646 3 vent vent NN 37809 646 4 , , , 37809 646 5 females female VBZ 37809 646 6 21.1 21.1 CD 37809 646 7 - - SYM 37809 646 8 29.4 29.4 CD 37809 646 9 mm mm CD 37809 646 10 . . . 37809 647 1 snout snout NN 37809 647 2 - - HYPH 37809 647 3 vent vent NN 37809 647 4 length length NN 37809 647 5 ; ; : 37809 647 6 vocal vocal NN 37809 647 7 slits slit NNS 37809 647 8 present present JJ 37809 647 9 in in IN 37809 647 10 males male NNS 37809 647 11 ; ; : 37809 647 12 finger finger NN 37809 647 13 tips tip NNS 37809 647 14 slightly slightly RB 37809 647 15 expanded expand VBN 37809 647 16 , , , 37809 647 17 truncate truncate JJ 37809 647 18 in in IN 37809 647 19 outline outline NN 37809 647 20 ; ; , 37809 647 21 inner inner JJ 37809 647 22 metatarsal metatarsal NN 37809 647 23 tubercle tubercle NN 37809 647 24 less less RBR 37809 647 25 than than IN 37809 647 26 twice twice PDT 37809 647 27 the the DT 37809 647 28 size size NN 37809 647 29 of of IN 37809 647 30 outer outer NN 37809 647 31 ; ; : 37809 647 32 skin skin NN 37809 647 33 of of IN 37809 647 34 dorsum dorsum NN 37809 647 35 smooth smooth JJ 37809 647 36 to to IN 37809 647 37 shagreened shagreene VBN 37809 647 38 , , , 37809 647 39 that that DT 37809 647 40 of of IN 37809 647 41 venter venter NN 37809 647 42 smooth smooth JJ 37809 647 43 ; ; : 37809 647 44 tympanum tympanum CD 37809 647 45 36.5 36.5 CD 37809 647 46 - - SYM 37809 647 47 54.0 54.0 CD 37809 647 48 per per IN 37809 647 49 cent cent NN 37809 647 50 diameter diameter NN 37809 647 51 of of IN 37809 647 52 eye eye NN 37809 647 53 ; ; : 37809 647 54 dorsum dorsum NN 37809 647 55 dark dark JJ 37809 647 56 brown brown NN 37809 647 57 with with IN 37809 647 58 large large JJ 37809 647 59 or or CC 37809 647 60 small small JJ 37809 647 61 light light NN 37809 647 62 brown brown NNP 37809 647 63 , , , 37809 647 64 orange orange NNP 37809 647 65 - - HYPH 37809 647 66 brown brown JJ 37809 647 67 , , , 37809 647 68 or or CC 37809 647 69 yellowish yellowish JJ 37809 647 70 spots spot NNS 37809 647 71 or or CC 37809 647 72 blotches blotch NNS 37809 647 73 ; ; : 37809 647 74 limbs limb NNS 37809 647 75 banded band VBD 37809 647 76 ; ; : 37809 647 77 interorbital interorbital NNP 37809 647 78 bar bar NN 37809 647 79 absent absent NN 37809 647 80 . . . 37809 648 1 [ [ -LRB- 37809 648 2 Illustration illustration NN 37809 648 3 : : : 37809 648 4 FIG FIG NNP 37809 648 5 . . . 37809 649 1 15 15 CD 37809 649 2 : : : 37809 649 3 Dicegrams dicegram NNS 37809 649 4 of of IN 37809 649 5 ear ear NN 37809 649 6 size size NN 37809 649 7 relative relative JJ 37809 649 8 to to IN 37809 649 9 eye eye NN 37809 649 10 diameter diameter NN 37809 649 11 in in IN 37809 649 12 the the DT 37809 649 13 two two CD 37809 649 14 subspecies subspecie NNS 37809 649 15 of of IN 37809 649 16 _ _ NNP 37809 649 17 Syrrhophus Syrrhophus NNP 37809 649 18 pipilans pipilans NNPS 37809 649 19 _ _ NNP 37809 649 20 . . . 37809 650 1 N n NN 37809 650 2 = = SYM 37809 650 3 17 17 CD 37809 650 4 in in IN 37809 650 5 _ _ NNP 37809 650 6 nebulosus nebulosus NN 37809 650 7 _ _ NNP 37809 650 8 , , , 37809 650 9 18 18 CD 37809 650 10 in in IN 37809 650 11 _ _ NNP 37809 650 12 pipilans pipilans NNPS 37809 650 13 _ _ NNP 37809 650 14 . . . 37809 650 15 ] ] -RRB- 37809 651 1 _ _ NNP 37809 651 2 Remarks._--Two remarks._--two CD 37809 651 3 subspecies subspecie NNS 37809 651 4 were be VBD 37809 651 5 recognized recognize VBN 37809 651 6 by by IN 37809 651 7 Duellman Duellman NNP 37809 651 8 ( ( -LRB- 37809 651 9 1958 1958 CD 37809 651 10 ) ) -RRB- 37809 651 11 . . . 37809 652 1 Previously previously RB 37809 652 2 both both DT 37809 652 3 had have VBD 37809 652 4 been be VBN 37809 652 5 treated treat VBN 37809 652 6 as as IN 37809 652 7 species specie NNS 37809 652 8 . . . 37809 653 1 The the DT 37809 653 2 two two CD 37809 653 3 populations population NNS 37809 653 4 were be VBD 37809 653 5 distinguished distinguish VBN 37809 653 6 on on IN 37809 653 7 the the DT 37809 653 8 basis basis NN 37809 653 9 of of IN 37809 653 10 color color NN 37809 653 11 pattern pattern NN 37809 653 12 and and CC 37809 653 13 the the DT 37809 653 14 size size NN 37809 653 15 of of IN 37809 653 16 the the DT 37809 653 17 tympanum tympanum NN 37809 653 18 . . . 37809 654 1 Measurements measurement NNS 37809 654 2 of of IN 37809 654 3 17 17 CD 37809 654 4 males male NNS 37809 654 5 of of IN 37809 654 6 _ _ NNP 37809 654 7 S. S. NNP 37809 655 1 p. p. NNP 37809 655 2 nebulosus nebulosus NNP 37809 655 3 _ _ NNP 37809 655 4 from from IN 37809 655 5 central central JJ 37809 655 6 Chiapas Chiapas NNPS 37809 655 7 and and CC 37809 655 8 18 18 CD 37809 655 9 males male NNS 37809 655 10 of of IN 37809 655 11 _ _ NNP 37809 655 12 S. S. NNP 37809 656 1 p. p. NN 37809 656 2 pipilans pipilans NNPS 37809 656 3 _ _ NNP 37809 656 4 from from IN 37809 656 5 southcentral southcentral JJ 37809 656 6 Oaxaca Oaxaca NNP 37809 656 7 and and CC 37809 656 8 Guerrero Guerrero NNP 37809 656 9 , , , 37809 656 10 México México NNP 37809 656 11 , , , 37809 656 12 demonstrates demonstrate VBZ 37809 656 13 that that IN 37809 656 14 the the DT 37809 656 15 supposed suppose VBN 37809 656 16 difference difference NN 37809 656 17 in in IN 37809 656 18 tympanum tympanum NN 37809 656 19 size size NN 37809 656 20 is be VBZ 37809 656 21 not not RB 37809 656 22 significant significant JJ 37809 656 23 ( ( -LRB- 37809 656 24 Fig fig NN 37809 656 25 . . . 37809 657 1 15 15 CD 37809 657 2 ) ) -RRB- 37809 657 3 . . . 37809 658 1 There there EX 37809 658 2 is be VBZ 37809 658 3 , , , 37809 658 4 however however RB 37809 658 5 , , , 37809 658 6 a a DT 37809 658 7 tendency tendency NN 37809 658 8 for for IN 37809 658 9 the the DT 37809 658 10 western western JJ 37809 658 11 population population NN 37809 658 12 of of IN 37809 658 13 _ _ NNP 37809 658 14 S. S. NNP 37809 659 1 pipilans pipilans NNPS 37809 659 2 _ _ NNP 37809 659 3 to to TO 37809 659 4 have have VB 37809 659 5 larger large JJR 37809 659 6 tympani tympani NNS 37809 659 7 . . . 37809 660 1 Based base VBN 37809 660 2 on on IN 37809 660 3 the the DT 37809 660 4 present present JJ 37809 660 5 examination examination NN 37809 660 6 of of IN 37809 660 7 112 112 CD 37809 660 8 specimens specimen NNS 37809 660 9 of of IN 37809 660 10 this this DT 37809 660 11 species specie NNS 37809 660 12 the the DT 37809 660 13 two two CD 37809 660 14 populations population NNS 37809 660 15 are be VBP 37809 660 16 held hold VBN 37809 660 17 to to TO 37809 660 18 be be VB 37809 660 19 sufficiently sufficiently RB 37809 660 20 distinct distinct JJ 37809 660 21 to to TO 37809 660 22 warrant warrant VB 37809 660 23 taxonomic taxonomic JJ 37809 660 24 recognition recognition NN 37809 660 25 as as IN 37809 660 26 subspecies subspecie NNS 37809 660 27 ( ( -LRB- 37809 660 28 Fig fig NN 37809 660 29 . . . 37809 661 1 16 16 CD 37809 661 2 ) ) -RRB- 37809 661 3 . . . 37809 662 1 [ [ -LRB- 37809 662 2 Illustration illustration NN 37809 662 3 : : : 37809 662 4 FIG FIG NNP 37809 662 5 . . . 37809 663 1 16 16 CD 37809 663 2 : : : 37809 663 3 _ _ NNP 37809 663 4 Syrrhophus Syrrhophus NNP 37809 663 5 pipilans pipilans NNPS 37809 663 6 nebulosus nebulosus NNP 37809 663 7 _ _ NNP 37809 663 8 ( ( -LRB- 37809 663 9 left leave VBN 37809 663 10 , , , 37809 663 11 KU KU NNP 37809 663 12 58908 58908 CD 37809 663 13 ) ) -RRB- 37809 663 14 and and CC 37809 663 15 _ _ NNP 37809 663 16 S. S. NNP 37809 664 1 p. p. NN 37809 664 2 pipilans pipilans NNPS 37809 664 3 _ _ NNP 37809 664 4 ( ( -LRB- 37809 664 5 right right UH 37809 664 6 , , , 37809 664 7 KU KU NNP 37809 664 8 86885 86885 CD 37809 664 9 ) ) -RRB- 37809 664 10 . . . 37809 665 1 ×2.7 ×2.7 UH 37809 665 2 . . . 37809 665 3 ] ] -RRB- 37809 666 1 The the DT 37809 666 2 parotoid parotoid NN 37809 666 3 glands gland NNS 37809 666 4 attributed attribute VBN 37809 666 5 to to IN 37809 666 6 this this DT 37809 666 7 species species NN 37809 666 8 by by IN 37809 666 9 Taylor Taylor NNP 37809 666 10 ( ( -LRB- 37809 666 11 1940c:95 1940c:95 CD 37809 666 12 ) ) -RRB- 37809 666 13 are be VBP 37809 666 14 merely merely RB 37809 666 15 the the DT 37809 666 16 superficial superficial JJ 37809 666 17 expression expression NN 37809 666 18 of of IN 37809 666 19 the the DT 37809 666 20 _ _ NNP 37809 666 21 m. m. NN 37809 667 1 depressor depressor NNP 37809 667 2 mandibulae mandibulae NNP 37809 667 3 _ _ NNP 37809 667 4 and and CC 37809 667 5 scapula scapula NNP 37809 667 6 . . . 37809 668 1 No no DT 37809 668 2 true true JJ 37809 668 3 glands gland NNS 37809 668 4 are be VBP 37809 668 5 present present JJ 37809 668 6 in in IN 37809 668 7 the the DT 37809 668 8 parotoid parotoid NN 37809 668 9 region region NN 37809 668 10 . . . 37809 669 1 = = NFP 37809 669 2 Syrrhophus Syrrhophus NNP 37809 669 3 pipilans pipilan VBZ 37809 669 4 nebulosus= nebulosus= VBP 37809 669 5 Taylor Taylor NNP 37809 669 6 _ _ NNP 37809 669 7 Syrrhophus Syrrhophus NNP 37809 669 8 nebulosus nebulosus NN 37809 669 9 _ _ NNP 37809 669 10 Taylor Taylor NNP 37809 669 11 , , , 37809 669 12 1943:353 1943:353 CD 37809 669 13 - - SYM 37809 669 14 55 55 CD 37809 669 15 , , , 37809 669 16 pl pl NNP 37809 669 17 . . . 37809 670 1 27 27 CD 37809 670 2 , , , 37809 670 3 figs fig NNS 37809 670 4 . . . 37809 671 1 3 3 LS 37809 671 2 - - SYM 37809 671 3 5 5 CD 37809 671 4 [ [ -LRB- 37809 671 5 Holotype.--FMNH holotype.--fmnh NN 37809 671 6 100095 100095 CD 37809 671 7 ( ( -LRB- 37809 671 8 formerly formerly RB 37809 671 9 EHT EHT NNP 37809 671 10 - - HYPH 37809 671 11 HMS HMS NNP 37809 671 12 3774 3774 CD 37809 671 13 ) ) -RRB- 37809 671 14 , , , 37809 671 15 near near IN 37809 671 16 Tonolá Tonolá NNP 37809 671 17 , , , 37809 671 18 Chiapas Chiapas NNP 37809 671 19 , , , 37809 671 20 México México NNP 37809 671 21 ; ; , 37809 671 22 collected collect VBN 37809 671 23 on on IN 37809 671 24 August August NNP 37809 671 25 27 27 CD 37809 671 26 , , , 37809 671 27 1935 1935 CD 37809 671 28 , , , 37809 671 29 by by IN 37809 671 30 Hobart Hobart NNP 37809 671 31 M. M. NNP 37809 671 32 Smith Smith NNP 37809 671 33 and and CC 37809 671 34 Edward Edward NNP 37809 671 35 H. H. NNP 37809 671 36 Taylor Taylor NNP 37809 671 37 ] ] -RRB- 37809 671 38 . . . 37809 672 1 Smith Smith NNP 37809 672 2 and and CC 37809 672 3 Taylor Taylor NNP 37809 672 4 , , , 37809 672 5 1948:49 1948:49 CD 37809 672 6 , , , 37809 672 7 51 51 CD 37809 672 8 . . . 37809 673 1 _ _ NNP 37809 673 2 Syrrhophus Syrrhophus NNP 37809 673 3 pipilans pipilans NNPS 37809 673 4 nebulosus nebulosus NNP 37809 673 5 _ _ NNP 37809 673 6 : : : 37809 673 7 Duellman Duellman NNP 37809 673 8 , , , 37809 673 9 1958:2 1958:2 CD 37809 673 10 - - SYM 37809 673 11 4 4 CD 37809 673 12 , , , 37809 673 13 9 9 CD 37809 673 14 , , , 37809 673 15 12 12 CD 37809 673 16 , , , 37809 673 17 14 14 CD 37809 673 18 . . . 37809 674 1 Stuart Stuart NNP 37809 674 2 , , , 37809 674 3 1963:32 1963:32 CD 37809 674 4 - - SYM 37809 674 5 33 33 CD 37809 674 6 . . . 37809 675 1 Gorham Gorham NNP 37809 675 2 , , , 37809 675 3 1966:166 1966:166 CD 37809 675 4 - - SYM 37809 675 5 67 67 CD 37809 675 6 . . . 37809 676 1 _ _ NNP 37809 676 2 Diagnosis._--Diameter Diagnosis._--Diameter NNP 37809 676 3 of of IN 37809 676 4 tympanum tympanum CD 37809 676 5 36.6 36.6 CD 37809 676 6 - - HYPH 37809 676 7 47.8 47.8 CD 37809 676 8 per per IN 37809 676 9 cent cent NN 37809 676 10 that that DT 37809 676 11 of of IN 37809 676 12 eye eye NN 37809 676 13 ; ; : 37809 676 14 dorsum dorsum NN 37809 676 15 dark dark JJ 37809 676 16 brown brown JJ 37809 676 17 with with IN 37809 676 18 numerous numerous JJ 37809 676 19 small small JJ 37809 676 20 light light NN 37809 676 21 brown brown JJ 37809 676 22 to to IN 37809 676 23 yellowish yellowish JJ 37809 676 24 spots spot NNS 37809 676 25 . . . 37809 677 1 _ _ NNP 37809 677 2 Remarks._--The remarks._--the CD 37809 677 3 distribution distribution NN 37809 677 4 of of IN 37809 677 5 this this DT 37809 677 6 subspecies subspecie NNS 37809 677 7 is be VBZ 37809 677 8 adequately adequately RB 37809 677 9 described describe VBN 37809 677 10 by by IN 37809 677 11 Duellman Duellman NNP 37809 677 12 ( ( -LRB- 37809 677 13 1958 1958 CD 37809 677 14 ) ) -RRB- 37809 677 15 . . . 37809 678 1 Fouquette Fouquette NNP 37809 678 2 ( ( -LRB- 37809 678 3 1960 1960 CD 37809 678 4 ) ) -RRB- 37809 678 5 described describe VBD 37809 678 6 the the DT 37809 678 7 vocalization vocalization NN 37809 678 8 of of IN 37809 678 9 this this DT 37809 678 10 frog frog NN 37809 678 11 . . . 37809 679 1 _ _ NNP 37809 679 2 Etymology._--Latin Etymology._--Latin NNP 37809 679 3 , , , 37809 679 4 _ _ NNP 37809 679 5 nebula nebula NNP 37809 679 6 _ _ NNP 37809 679 7 , , , 37809 679 8 in in IN 37809 679 9 reference reference NN 37809 679 10 to to IN 37809 679 11 the the DT 37809 679 12 clouded clouded JJ 37809 679 13 dorsal dorsal NN 37809 679 14 pattern pattern NN 37809 679 15 . . . 37809 680 1 _ _ NNP 37809 680 2 Distribution._--Low Distribution._--Low NNP 37809 680 3 to to TO 37809 680 4 moderate moderate VB 37809 680 5 elevations elevation NNS 37809 680 6 along along IN 37809 680 7 the the DT 37809 680 8 Pacific Pacific NNP 37809 680 9 versant versant NN 37809 680 10 of of IN 37809 680 11 Chiapas Chiapas NNPS 37809 680 12 and and CC 37809 680 13 in in IN 37809 680 14 the the DT 37809 680 15 Grijalva Grijalva NNP 37809 680 16 valley valley NN 37809 680 17 of of IN 37809 680 18 Chiapas Chiapas NNP 37809 680 19 and and CC 37809 680 20 Guatemala Guatemala NNP 37809 680 21 ( ( -LRB- 37809 680 22 Fig Fig NNP 37809 680 23 . . . 37809 681 1 17 17 CD 37809 681 2 ) ) -RRB- 37809 681 3 . . . 37809 682 1 _ _ NNP 37809 682 2 Specimens Specimens NNP 37809 682 3 examined._--(54 examined._--(54 NNP 37809 682 4 ) ) -RRB- 37809 682 5 GUATEMALA GUATEMALA NNP 37809 682 6 , , , 37809 682 7 _ _ NNP 37809 682 8 Huehuetenango Huehuetenango NNP 37809 682 9 _ _ NNP 37809 682 10 : : : 37809 682 11 Jacaltenango Jacaltenango NNP 37809 682 12 , , , 37809 682 13 UMMZ UMMZ NNP 37809 682 14 117036 117036 CD 37809 682 15 ; ; : 37809 682 16 35 35 CD 37809 682 17 km km NN 37809 682 18 . . . 37809 683 1 SE SE NNP 37809 683 2 La La NNP 37809 683 3 Mesilla Mesilla NNP 37809 683 4 , , , 37809 683 5 TNHC TNHC NNP 37809 683 6 29652 29652 CD 37809 683 7 . . . 37809 684 1 MÉXICO MÉXICO NNP 37809 684 2 , , , 37809 684 3 _ _ NNP 37809 684 4 Chiapas Chiapas NNP 37809 684 5 _ _ NNP 37809 684 6 : : : 37809 684 7 11.2 11.2 CD 37809 684 8 km km NN 37809 684 9 . . . 37809 685 1 N N NNP 37809 685 2 Arriaga Arriaga NNP 37809 685 3 , , , 37809 685 4 300 300 CD 37809 685 5 m. m. NN 37809 685 6 , , , 37809 685 7 UMMZ UMMZ NNP 37809 685 8 125891 125891 CD 37809 685 9 ; ; : 37809 685 10 11.8 11.8 CD 37809 685 11 km km NNS 37809 685 12 . . . 37809 686 1 N N NNP 37809 686 2 Arriaga Arriaga NNPS 37809 686 3 , , , 37809 686 4 UMMZ UMMZ NNP 37809 686 5 117279 117279 CD 37809 686 6 ; ; : 37809 686 7 12.8 12.8 CD 37809 686 8 km km NNS 37809 686 9 . . . 37809 687 1 N N NNP 37809 687 2 Arriaga Arriaga NNPS 37809 687 3 , , , 37809 687 4 UMMZ UMMZ NNP 37809 687 5 117280 117280 CD 37809 687 6 ; ; : 37809 687 7 17.5 17.5 CD 37809 687 8 km km NNS 37809 687 9 . . . 37809 688 1 S S NNP 37809 688 2 Arriaga Arriaga NNP 37809 688 3 , , , 37809 688 4 UIMNH UIMNH NNP 37809 688 5 57108 57108 CD 37809 688 6 - - SYM 37809 688 7 109 109 CD 37809 688 8 ; ; : 37809 688 9 1.5 1.5 CD 37809 688 10 km km NN 37809 688 11 . . . 37809 689 1 S S NNP 37809 689 2 Bochil Bochil NNP 37809 689 3 , , , 37809 689 4 1250 1250 CD 37809 689 5 m. m. NN 37809 689 6 , , , 37809 689 7 KU KU NNP 37809 689 8 58898 58898 CD 37809 689 9 - - SYM 37809 689 10 908 908 CD 37809 689 11 ; ; : 37809 689 12 Cerro Cerro NNP 37809 689 13 Hueco Hueco NNP 37809 689 14 , , , 37809 689 15 7 7 CD 37809 689 16 km km NN 37809 689 17 . . . 37809 690 1 S S NNP 37809 690 2 Tuxtla Tuxtla NNP 37809 690 3 Gutierrez Gutierrez NNP 37809 690 4 , , , 37809 690 5 UMMZ UMMZ NNP 37809 690 6 123007 123007 CD 37809 690 7 ; ; : 37809 690 8 3.2 3.2 CD 37809 690 9 km km NN 37809 690 10 . . . 37809 691 1 S S NNP 37809 691 2 Ixtapa Ixtapa NNP 37809 691 3 , , , 37809 691 4 UMMZ UMMZ NNP 37809 691 5 124000 124000 CD 37809 691 6 ; ; : 37809 691 7 Linda Linda NNP 37809 691 8 Vista Vista NNP 37809 691 9 , , , 37809 691 10 ca ca MD 37809 691 11 . . . 37809 692 1 2 2 CD 37809 692 2 km km NN 37809 692 3 . . . 37809 693 1 NW NW NNP 37809 693 2 Pueblo Pueblo NNP 37809 693 3 Nuevo Nuevo NNP 37809 693 4 Solistahuacán Solistahuacán NNP 37809 693 5 , , , 37809 693 6 KU KU NNP 37809 693 7 58897 58897 CD 37809 693 8 ; ; : 37809 693 9 Hda Hda NNP 37809 693 10 . . . 37809 694 1 Monserrate Monserrate NNP 37809 694 2 , , , 37809 694 3 40 40 CD 37809 694 4 km km NN 37809 694 5 . . . 37809 695 1 NW NW NNP 37809 695 2 Arriaga Arriaga NNP 37809 695 3 , , , 37809 695 4 UMMZ UMMZ NNP 37809 695 5 102258 102258 CD 37809 695 6 ; ; : 37809 695 7 near near IN 37809 695 8 San San NNP 37809 695 9 Ricardo Ricardo NNP 37809 695 10 , , , 37809 695 11 FMNH FMNH NNP 37809 695 12 100720 100720 CD 37809 695 13 ; ; : 37809 695 14 Tapachula Tapachula NNP 37809 695 15 , , , 37809 695 16 FMNH FMNH NNP 37809 695 17 75792 75792 CD 37809 695 18 , , , 37809 695 19 103242 103242 CD 37809 695 20 , , , 37809 695 21 100695 100695 CD 37809 695 22 - - SYM 37809 695 23 96 96 CD 37809 695 24 , , , 37809 695 25 UIMNH UIMNH NNP 37809 695 26 13292 13292 CD 37809 695 27 ; ; : 37809 695 28 56 56 CD 37809 695 29 km km NNS 37809 695 30 . . . 37809 696 1 E E NNP 37809 696 2 Tapanatepec Tapanatepec NNP 37809 696 3 , , , 37809 696 4 Oaxaca Oaxaca NNP 37809 696 5 , , , 37809 696 6 TNHC TNHC NNP 37809 696 7 26942 26942 CD 37809 696 8 , , , 37809 696 9 Tonolá Tonolá NNP 37809 696 10 , , , 37809 696 11 FMNH FMNH NNP 37809 696 12 100095 100095 CD 37809 696 13 ( ( -LRB- 37809 696 14 holotype holotype NN 37809 696 15 ) ) -RRB- 37809 696 16 , , , 37809 696 17 100686 100686 CD 37809 696 18 - - SYM 37809 696 19 92 92 CD 37809 696 20 , , , 37809 696 21 UIMNH UIMNH NNP 37809 696 22 13293 13293 CD 37809 696 23 - - SYM 37809 696 24 95 95 CD 37809 696 25 ; ; : 37809 696 26 Tuxtla Tuxtla NNP 37809 696 27 Gutierrez Gutierrez NNP 37809 696 28 , , , 37809 696 29 FMNH FMNH NNP 37809 696 30 100693 100693 CD 37809 696 31 - - SYM 37809 696 32 94 94 CD 37809 696 33 , , , 37809 696 34 UIMNH UIMNH NNP 37809 696 35 13297 13297 CD 37809 696 36 ; ; : 37809 696 37 19 19 CD 37809 696 38 km km NN 37809 696 39 . . . 37809 697 1 N N NNP 37809 697 2 Tuxtla Tuxtla NNP 37809 697 3 Gutierrez Gutierrez NNP 37809 697 4 , , , 37809 697 5 TNHC TNHC NNP 37809 697 6 25229 25229 CD 37809 697 7 - - SYM 37809 697 8 30 30 CD 37809 697 9 ; ; : 37809 697 10 15.5 15.5 CD 37809 697 11 km km NNS 37809 697 12 . . . 37809 698 1 NE NE NNP 37809 698 2 Tuxtla Tuxtla NNP 37809 698 3 Gutierrez Gutierrez NNP 37809 698 4 , , , 37809 698 5 UMMZ UMMZ NNP 37809 698 6 119892 119892 CD 37809 698 7 ( ( -LRB- 37809 698 8 3 3 CD 37809 698 9 ) ) -RRB- 37809 698 10 ; ; : 37809 698 11 19 19 CD 37809 698 12 km km NN 37809 698 13 . . . 37809 699 1 NE NE NNP 37809 699 2 Tuxtla Tuxtla NNP 37809 699 3 Gutierrez Gutierrez NNP 37809 699 4 , , , 37809 699 5 UMMZ UMMZ NNP 37809 699 6 119891 119891 CD 37809 699 7 ( ( -LRB- 37809 699 8 3 3 CD 37809 699 9 ) ) -RRB- 37809 699 10 ; ; : 37809 699 11 8 8 CD 37809 699 12 km km NN 37809 699 13 . . . 37809 700 1 NNW NNW NNP 37809 700 2 Tuxtla Tuxtla NNP 37809 700 3 Gutierrez Gutierrez NNP 37809 700 4 , , , 37809 700 5 KU KU NNP 37809 700 6 37809 37809 CD 37809 700 7 ; ; : 37809 700 8 Unión Unión NNP 37809 700 9 de de NNP 37809 700 10 Juarez Juarez NNP 37809 700 11 , , , 37809 700 12 FMNH FMNH NNP 37809 700 13 105294 105294 CD 37809 700 14 . . . 37809 701 1 = = NFP 37809 701 2 Syrrhophus Syrrhophus NNP 37809 701 3 pipilans pipilan NNS 37809 701 4 pipilans= pipilans= VBP 37809 701 5 Taylor Taylor NNP 37809 701 6 _ _ NNP 37809 701 7 ? ? . 37809 701 8 Syrrhopus Syrrhopus NNP 37809 701 9 verruculatus verruculatus NNP 37809 701 10 _ _ NNP 37809 701 11 : : : 37809 701 12 Gadow Gadow NNP 37809 701 13 , , , 37809 701 14 1905:194 1905:194 CD 37809 701 15 . . . 37809 702 1 _ _ NNP 37809 702 2 Syrrhophus Syrrhophus NNP 37809 702 3 pipilans pipilan NNS 37809 702 4 _ _ NNP 37809 702 5 Taylor Taylor NNP 37809 702 6 , , , 37809 702 7 1940c:95 1940c:95 CD 37809 702 8 - - SYM 37809 702 9 97 97 CD 37809 702 10 , , , 37809 702 11 pl pl NNP 37809 702 12 . . . 37809 703 1 1 1 LS 37809 703 2 [ [ -LRB- 37809 703 3 Holotype.--FMNH holotype.--fmnh NN 37809 703 4 100072 100072 CD 37809 703 5 ( ( -LRB- 37809 703 6 formerly formerly RB 37809 703 7 EHT EHT NNP 37809 703 8 - - HYPH 37809 703 9 HMS HMS NNP 37809 703 10 6843 6843 CD 37809 703 11 ) ) -RRB- 37809 703 12 , , , 37809 703 13 from from IN 37809 703 14 14.6 14.6 CD 37809 703 15 km km NNS 37809 703 16 . . . 37809 704 1 S S NNP 37809 704 2 Mazatlán Mazatlán NNP 37809 704 3 , , , 37809 704 4 Guerrero Guerrero NNP 37809 704 5 , , , 37809 704 6 México México NNP 37809 704 7 ; ; , 37809 704 8 collected collect VBN 37809 704 9 on on IN 37809 704 10 July July NNP 37809 704 11 22 22 CD 37809 704 12 , , , 37809 704 13 1936 1936 CD 37809 704 14 , , , 37809 704 15 by by IN 37809 704 16 Edward Edward NNP 37809 704 17 H. H. NNP 37809 704 18 Taylor Taylor NNP 37809 704 19 ] ] -RRB- 37809 704 20 . . . 37809 705 1 Taylor Taylor NNP 37809 705 2 and and CC 37809 705 3 Smith Smith NNP 37809 705 4 , , , 37809 705 5 1945:581 1945:581 CD 37809 705 6 - - SYM 37809 705 7 82 82 CD 37809 705 8 . . . 37809 706 1 Smith Smith NNP 37809 706 2 and and CC 37809 706 3 Taylor Taylor NNP 37809 706 4 , , , 37809 706 5 1948:49 1948:49 CD 37809 706 6 , , , 37809 706 7 50 50 CD 37809 706 8 - - SYM 37809 706 9 51 51 CD 37809 706 10 . . . 37809 707 1 _ _ NNP 37809 707 2 Syrrhophus Syrrhophus NNP 37809 707 3 pipilans pipilan NNS 37809 707 4 pipilans pipilans NNPS 37809 707 5 _ _ NNP 37809 707 6 : : : 37809 707 7 Duellman Duellman NNP 37809 707 8 , , , 37809 707 9 1958:1 1958:1 CD 37809 707 10 - - SYM 37809 707 11 4 4 CD 37809 707 12 , , , 37809 707 13 8 8 CD 37809 707 14 - - SYM 37809 707 15 9 9 CD 37809 707 16 , , , 37809 707 17 13 13 CD 37809 707 18 - - SYM 37809 707 19 14 14 CD 37809 707 20 , , , 37809 707 21 pl pl NNP 37809 707 22 . . . 37809 708 1 2 2 LS 37809 708 2 , , , 37809 708 3 fig fig NN 37809 708 4 . . . 37809 709 1 1 1 LS 37809 709 2 . . . 37809 710 1 Gorham Gorham NNP 37809 710 2 , , , 37809 710 3 1966:166 1966:166 CD 37809 710 4 . . . 37809 711 1 _ _ NNP 37809 711 2 Diagnosis._--Diameter Diagnosis._--Diameter NNP 37809 711 3 of of IN 37809 711 4 tympanum tympanum CD 37809 711 5 40.6 40.6 CD 37809 711 6 - - SYM 37809 711 7 54.0 54.0 CD 37809 711 8 per per IN 37809 711 9 cent cent NN 37809 711 10 that that DT 37809 711 11 of of IN 37809 711 12 eye eye NN 37809 711 13 ; ; : 37809 711 14 dorsum dorsum NN 37809 711 15 dark dark JJ 37809 711 16 brown brown NN 37809 711 17 with with IN 37809 711 18 large large JJ 37809 711 19 light light JJ 37809 711 20 spots spot NNS 37809 711 21 or or CC 37809 711 22 blotches blotch NNS 37809 711 23 . . . 37809 712 1 _ _ NNP 37809 712 2 Remarks._--Duellman Remarks._--Duellman NNP 37809 712 3 's 's POS 37809 712 4 ( ( -LRB- 37809 712 5 1958 1958 CD 37809 712 6 ) ) -RRB- 37809 712 7 synopsis synopsis NN 37809 712 8 of of IN 37809 712 9 this this DT 37809 712 10 subspecies subspecie NNS 37809 712 11 is be VBZ 37809 712 12 adequate adequate JJ 37809 712 13 ; ; : 37809 712 14 the the DT 37809 712 15 distribution distribution NN 37809 712 16 has have VBZ 37809 712 17 not not RB 37809 712 18 been be VBN 37809 712 19 extended extend VBN 37809 712 20 , , , 37809 712 21 but but CC 37809 712 22 several several JJ 37809 712 23 records record NNS 37809 712 24 are be VBP 37809 712 25 now now RB 37809 712 26 available available JJ 37809 712 27 which which WDT 37809 712 28 fill fill VBP 37809 712 29 in in IN 37809 712 30 gaps gap NNS 37809 712 31 . . . 37809 713 1 [ [ -LRB- 37809 713 2 Illustration illustration NN 37809 713 3 : : : 37809 713 4 FIG FIG NNP 37809 713 5 . . . 37809 714 1 17 17 LS 37809 714 2 : : : 37809 714 3 Distribution distribution NN 37809 714 4 of of IN 37809 714 5 _ _ NNP 37809 714 6 Syrrhophus Syrrhophus NNP 37809 714 7 pipilans pipilans NNPS 37809 714 8 _ _ NNP 37809 714 9 : : : 37809 714 10 _ _ NNP 37809 714 11 nebulosus nebulosus NN 37809 714 12 _ _ NNP 37809 714 13 ( ( -LRB- 37809 714 14 open open JJ 37809 714 15 circles circle NNS 37809 714 16 ) ) -RRB- 37809 714 17 and and CC 37809 714 18 _ _ NNP 37809 714 19 pipilans pipilans NNPS 37809 714 20 _ _ NNP 37809 714 21 ( ( -LRB- 37809 714 22 solid solid JJ 37809 714 23 circles circle NNS 37809 714 24 ) ) -RRB- 37809 714 25 . . . 37809 714 26 ] ] -RRB- 37809 715 1 Gadow Gadow NNP 37809 715 2 's 's POS 37809 715 3 ( ( -LRB- 37809 715 4 1905 1905 CD 37809 715 5 ) ) -RRB- 37809 715 6 record record NN 37809 715 7 of of IN 37809 715 8 _ _ NNP 37809 715 9 S. S. NNP 37809 716 1 verruculatus verruculatus NNP 37809 716 2 _ _ NNP 37809 716 3 from from IN 37809 716 4 " " `` 37809 716 5 Buena Buena NNP 37809 716 6 Vista Vista NNP 37809 716 7 , , , 37809 716 8 S. S. NNP 37809 716 9 Guerrero Guerrero NNP 37809 716 10 " " `` 37809 716 11 is be VBZ 37809 716 12 most most RBS 37809 716 13 likely likely RB 37809 716 14 applicable applicable JJ 37809 716 15 to to IN 37809 716 16 this this DT 37809 716 17 species species NN 37809 716 18 . . . 37809 717 1 Gadow Gadow NNP 37809 717 2 simply simply RB 37809 717 3 included include VBD 37809 717 4 the the DT 37809 717 5 name name NN 37809 717 6 in in IN 37809 717 7 a a DT 37809 717 8 list list NN 37809 717 9 of of IN 37809 717 10 the the DT 37809 717 11 species specie NNS 37809 717 12 he -PRON- PRP 37809 717 13 had have VBD 37809 717 14 collected collect VBN 37809 717 15 during during IN 37809 717 16 his -PRON- PRP$ 37809 717 17 trip trip NN 37809 717 18 in in IN 37809 717 19 México México NNP 37809 717 20 ( ( -LRB- 37809 717 21 1902 1902 CD 37809 717 22 - - SYM 37809 717 23 04 04 CD 37809 717 24 ) ) -RRB- 37809 717 25 ; ; : 37809 717 26 no no DT 37809 717 27 further further JJ 37809 717 28 comment comment NN 37809 717 29 was be VBD 37809 717 30 made make VBN 37809 717 31 on on IN 37809 717 32 this this DT 37809 717 33 species species NN 37809 717 34 although although IN 37809 717 35 references reference NNS 37809 717 36 to to IN 37809 717 37 _ _ NNP 37809 717 38 Syrrhopus Syrrhopus NNP 37809 717 39 _ _ NNP 37809 717 40 ( ( -LRB- 37809 717 41 sic sic NN 37809 717 42 ) ) -RRB- 37809 717 43 appear appear VBP 37809 717 44 in in IN 37809 717 45 several several JJ 37809 717 46 places place NNS 37809 717 47 in in IN 37809 717 48 the the DT 37809 717 49 paper paper NN 37809 717 50 and and CC 37809 717 51 would would MD 37809 717 52 appear appear VB 37809 717 53 to to TO 37809 717 54 apply apply VB 37809 717 55 to to IN 37809 717 56 the the DT 37809 717 57 species specie NNS 37809 717 58 he -PRON- PRP 37809 717 59 had have VBD 37809 717 60 . . . 37809 718 1 _ _ NNP 37809 718 2 Etymology._--Latin Etymology._--Latin NNP 37809 718 3 , , , 37809 718 4 _ _ NNP 37809 718 5 pipilo pipilo VBD 37809 718 6 _ _ NNP 37809 718 7 , , , 37809 718 8 chirping chirp VBG 37809 718 9 , , , 37809 718 10 peeping peeping NN 37809 718 11 , , , 37809 718 12 in in IN 37809 718 13 reference reference NN 37809 718 14 to to IN 37809 718 15 the the DT 37809 718 16 call call NN 37809 718 17 of of IN 37809 718 18 the the DT 37809 718 19 male male NN 37809 718 20 . . . 37809 719 1 _ _ NNP 37809 719 2 Distribution._--Sea Distribution._--Sea NNP 37809 719 3 level level NN 37809 719 4 to to IN 37809 719 5 about about RB 37809 719 6 1800 1800 CD 37809 719 7 meters meter NNS 37809 719 8 along along IN 37809 719 9 the the DT 37809 719 10 Pacific Pacific NNP 37809 719 11 versant versant NN 37809 719 12 of of IN 37809 719 13 western western JJ 37809 719 14 México México NNP 37809 719 15 from from IN 37809 719 16 central central JJ 37809 719 17 Guerrero Guerrero NNP 37809 719 18 to to IN 37809 719 19 the the DT 37809 719 20 Isthmus Isthmus NNP 37809 719 21 of of IN 37809 719 22 Tehuantepec Tehuantepec NNP 37809 719 23 ( ( -LRB- 37809 719 24 Fig Fig NNP 37809 719 25 . . . 37809 720 1 17 17 CD 37809 720 2 ) ) -RRB- 37809 720 3 . . . 37809 721 1 _ _ NNP 37809 721 2 Specimens Specimens NNP 37809 721 3 examined._--(62 examined._--(62 NNP 37809 721 4 ) ) -RRB- 37809 721 5 . . . 37809 722 1 MÉXICO MÉXICO NNP 37809 722 2 , , , 37809 722 3 _ _ NNP 37809 722 4 Guerrero Guerrero NNP 37809 722 5 _ _ NNP 37809 722 6 : : : 37809 722 7 Acapulco Acapulco NNP 37809 722 8 , , , 37809 722 9 UMMZ UMMZ NNP 37809 722 10 110125 110125 CD 37809 722 11 ; ; : 37809 722 12 6.4 6.4 CD 37809 722 13 km km NN 37809 722 14 . . . 37809 723 1 N n IN 37809 723 2 Acapulco Acapulco NNP 37809 723 3 , , , 37809 723 4 FMNH FMNH NNP 37809 723 5 100389 100389 CD 37809 723 6 , , , 37809 723 7 100525 100525 CD 37809 723 8 ; ; : 37809 723 9 Agua Agua NNP 37809 723 10 del del NNP 37809 723 11 Obispo Obispo NNP 37809 723 12 , , , 37809 723 13 980 980 CD 37809 723 14 - - SYM 37809 723 15 1000 1000 CD 37809 723 16 m. m. NN 37809 723 17 , , , 37809 723 18 FMNH FMNH NNP 37809 723 19 75791 75791 CD 37809 723 20 , , , 37809 723 21 100518 100518 CD 37809 723 22 - - SYM 37809 723 23 21 21 CD 37809 723 24 , , , 37809 723 25 100526 100526 CD 37809 723 26 , , , 37809 723 27 KU KU NNP 37809 723 28 86884 86884 CD 37809 723 29 - - SYM 37809 723 30 86 86 CD 37809 723 31 , , , 37809 723 32 UIMNH UIMNH NNP 37809 723 33 13315 13315 CD 37809 723 34 , , , 37809 723 35 UMMZ UMMZ NNP 37809 723 36 119152 119152 CD 37809 723 37 , , , 37809 723 38 125890 125890 CD 37809 723 39 ( ( -LRB- 37809 723 40 4 4 CD 37809 723 41 ) ) -RRB- 37809 723 42 ; ; : 37809 723 43 13.3 13.3 CD 37809 723 44 km km NNS 37809 723 45 . . . 37809 724 1 NW NW NNP 37809 724 2 Coyuca Coyuca NNP 37809 724 3 , , , 37809 724 4 UIMNH UIMNH NNP 37809 724 5 38367 38367 CD 37809 724 6 , , , 37809 724 7 71982 71982 CD 37809 724 8 - - SYM 37809 724 9 83 83 CD 37809 724 10 ; ; : 37809 724 11 14.5 14.5 CD 37809 724 12 km km NNS 37809 724 13 . . . 37809 725 1 S S NNP 37809 725 2 Mazatlán Mazatlán NNP 37809 725 3 , , , 37809 725 4 FMNH FMNH NNP 37809 725 5 100072 100072 CD 37809 725 6 ( ( -LRB- 37809 725 7 holotype holotype NNP 37809 725 8 ) ) -RRB- 37809 725 9 , , , 37809 725 10 100408 100408 CD 37809 725 11 , , , 37809 725 12 100511 100511 CD 37809 725 13 - - SYM 37809 725 14 17 17 CD 37809 725 15 , , , 37809 725 16 UIMNH UIMNH NNP 37809 725 17 13302 13302 CD 37809 725 18 - - SYM 37809 725 19 309 309 CD 37809 725 20 ; ; : 37809 725 21 Tierra Tierra NNP 37809 725 22 Colorado Colorado NNP 37809 725 23 , , , 37809 725 24 300 300 CD 37809 725 25 m. m. NN 37809 725 26 , , , 37809 725 27 KU KU NNP 37809 725 28 67961 67961 CD 37809 725 29 , , , 37809 725 30 UIMNH UIMNH NNP 37809 725 31 13313 13313 CD 37809 725 32 - - SYM 37809 725 33 14 14 CD 37809 725 34 ; ; : 37809 725 35 near near IN 37809 725 36 El El NNP 37809 725 37 Treinte Treinte NNP 37809 725 38 , , , 37809 725 39 FMNH FMNH NNP 37809 725 40 126639 126639 CD 37809 725 41 ; ; : 37809 725 42 Xaltinanguis Xaltinanguis NNP 37809 725 43 , , , 37809 725 44 FMNH FMNH NNP 37809 725 45 100522 100522 CD 37809 725 46 - - SYM 37809 725 47 24 24 CD 37809 725 48 , , , 37809 725 49 126640 126640 CD 37809 725 50 . . . 37809 726 1 _ _ NNP 37809 726 2 Oaxaca Oaxaca NNP 37809 726 3 _ _ NNP 37809 726 4 : : : 37809 726 5 Cacahuatepec Cacahuatepec NNP 37809 726 6 , , , 37809 726 7 UIMNH UIMNH NNP 37809 726 8 52853 52853 CD 37809 726 9 ; ; : 37809 726 10 8 8 CD 37809 726 11 km km NN 37809 726 12 . . . 37809 727 1 NW NW NNP 37809 727 2 Río Río NNP 37809 727 3 Canoa Canoa NNP 37809 727 4 , , , 37809 727 5 53 53 CD 37809 727 6 km km NN 37809 727 7 . . . 37809 728 1 ESE ESE NNP 37809 728 2 Cuajinicuilapa Cuajinicuilapa NNP 37809 728 3 , , , 37809 728 4 UIMNH UIMNH NNP 37809 728 5 52852 52852 CD 37809 728 6 ; ; : 37809 728 7 6.4 6.4 CD 37809 728 8 km km NN 37809 728 9 . . . 37809 729 1 N N NNP 37809 729 2 El El NNP 37809 729 3 Candelaria Candelaria NNP 37809 729 4 , , , 37809 729 5 UIMNH UIMNH NNP 37809 729 6 9501 9501 CD 37809 729 7 ; ; : 37809 729 8 11.2 11.2 CD 37809 729 9 km km NNS 37809 729 10 . . . 37809 730 1 S S NNP 37809 730 2 El El NNP 37809 730 3 Candelaria Candelaria NNP 37809 730 4 , , , 37809 730 5 UIMNH UIMNH NNP 37809 730 6 9502 9502 CD 37809 730 7 ; ; : 37809 730 8 17 17 CD 37809 730 9 km km NN 37809 730 10 . . . 37809 731 1 NE NE NNP 37809 731 2 Juchatengo Juchatengo NNP 37809 731 3 , , , 37809 731 4 1600 1600 CD 37809 731 5 m. m. NN 37809 731 6 , , , 37809 731 7 KU KU NNP 37809 731 8 86887 86887 CD 37809 731 9 ; ; : 37809 731 10 31.5 31.5 CD 37809 731 11 km km NNS 37809 731 12 . . . 37809 732 1 N n IN 37809 732 2 Pochutla Pochutla NNP 37809 732 3 , , , 37809 732 4 UMMZ UMMZ NNP 37809 732 5 123999 123999 CD 37809 732 6 ( ( -LRB- 37809 732 7 2 2 CD 37809 732 8 ) ) -RRB- 37809 732 9 ; ; : 37809 732 10 32.9 32.9 CD 37809 732 11 km km NN 37809 732 12 . . . 37809 733 1 N N NNP 37809 733 2 Pochutla Pochutla NNP 37809 733 3 , , , 37809 733 4 850 850 CD 37809 733 5 m. m. NN 37809 733 6 , , , 37809 733 7 UMMZ UMMZ NNP 37809 733 8 123996 123996 CD 37809 733 9 ; ; : 37809 733 10 37.1 37.1 CD 37809 733 11 km km NNS 37809 733 12 . . . 37809 734 1 N n IN 37809 734 2 Pochutla Pochutla NNP 37809 734 3 , , , 37809 734 4 UMMZ UMMZ NNP 37809 734 5 123998 123998 CD 37809 734 6 ( ( -LRB- 37809 734 7 2 2 CD 37809 734 8 ) ) -RRB- 37809 734 9 ; ; : 37809 734 10 41.4 41.4 CD 37809 734 11 km km NNS 37809 734 12 . . . 37809 735 1 N n IN 37809 735 2 Pochutla Pochutla NNP 37809 735 3 , , , 37809 735 4 UMMZ UMMZ NNP 37809 735 5 123997 123997 CD 37809 735 6 ( ( -LRB- 37809 735 7 2 2 CD 37809 735 8 ) ) -RRB- 37809 735 9 ; ; : 37809 735 10 Cerro Cerro NNP 37809 735 11 Quiengola Quiengola NNP 37809 735 12 , , , 37809 735 13 FMNH FMNH NNP 37809 735 14 105653 105653 CD 37809 735 15 ; ; : 37809 735 16 3.8 3.8 CD 37809 735 17 km km NN 37809 735 18 . . . 37809 736 1 N N NNP 37809 736 2 Santiago Santiago NNP 37809 736 3 Chivela Chivela NNP 37809 736 4 , , , 37809 736 5 UMMZ UMMZ NNP 37809 736 6 115449 115449 CD 37809 736 7 ; ; : 37809 736 8 14.5 14.5 CD 37809 736 9 km km NNS 37809 736 10 . . . 37809 737 1 W W NNP 37809 737 2 Tehuantepec Tehuantepec NNP 37809 737 3 , , , 37809 737 4 UMMZ UMMZ NNP 37809 737 5 115448 115448 CD 37809 737 6 ( ( -LRB- 37809 737 7 2 2 CD 37809 737 8 ) ) -RRB- 37809 737 9 . . . 37809 738 1 = = NFP 37809 738 2 Syrrhophus Syrrhophus NNP 37809 738 3 interorbitalis= interorbitalis= NN 37809 738 4 Langebartel Langebartel NNP 37809 738 5 and and CC 37809 738 6 Shannon Shannon NNP 37809 738 7 _ _ NNP 37809 738 8 Syrrhophus Syrrhophus NNP 37809 738 9 interorbitalis interorbitalis NN 37809 738 10 _ _ NNP 37809 738 11 Langebartel Langebartel NNP 37809 738 12 and and CC 37809 738 13 Shannon Shannon NNP 37809 738 14 , , , 37809 738 15 1956 1956 CD 37809 738 16 : : : 37809 738 17 161 161 CD 37809 738 18 - - SYM 37809 738 19 65 65 CD 37809 738 20 , , , 37809 738 21 figs fig NNS 37809 738 22 . . . 37809 739 1 1 1 LS 37809 739 2 - - SYM 37809 739 3 2 2 CD 37809 739 4 [ [ -LRB- 37809 739 5 Holotype.--UIMNH Holotype.--UIMNH NNP 37809 739 6 67061 67061 CD 37809 739 7 ( ( -LRB- 37809 739 8 formerly formerly RB 37809 739 9 FAS FAS NNP 37809 739 10 9378 9378 CD 37809 739 11 ) ) -RRB- 37809 739 12 , , , 37809 739 13 36 36 CD 37809 739 14 mi mi JJ 37809 739 15 . . . 37809 740 1 N N NNP 37809 740 2 Mazatlán Mazatlán NNP 37809 740 3 , , , 37809 740 4 Sinaloa Sinaloa NNP 37809 740 5 , , , 37809 740 6 México México NNP 37809 740 7 , , , 37809 740 8 collected collect VBD 37809 740 9 on on IN 37809 740 10 November November NNP 37809 740 11 17 17 CD 37809 740 12 , , , 37809 740 13 1955 1955 CD 37809 740 14 , , , 37809 740 15 by by IN 37809 740 16 E. E. NNP 37809 740 17 C. C. NNP 37809 740 18 Bay Bay NNP 37809 740 19 , , , 37809 740 20 J. J. NNP 37809 740 21 C. C. NNP 37809 740 22 Schaffner Schaffner NNP 37809 740 23 , , , 37809 740 24 and and CC 37809 740 25 D. D. NNP 37809 740 26 A. a. NN 37809 740 27 Langebartel Langebartel NNP 37809 740 28 ] ] -RRB- 37809 740 29 . . . 37809 741 1 Duellman duellman NN 37809 741 2 , , , 37809 741 3 1958:1 1958:1 CD 37809 741 4 - - SYM 37809 741 5 4 4 CD 37809 741 6 , , , 37809 741 7 10 10 CD 37809 741 8 , , , 37809 741 9 12 12 CD 37809 741 10 , , , 37809 741 11 14 14 CD 37809 741 12 . . . 37809 742 1 Gorham Gorham NNP 37809 742 2 , , , 37809 742 3 1966:164 1966:164 CD 37809 742 4 - - SYM 37809 742 5 65 65 CD 37809 742 6 . . . 37809 743 1 _ _ NNP 37809 743 2 Syrrhophis Syrrhophis NNP 37809 743 3 interorbitalis interorbitalis NN 37809 743 4 _ _ NNP 37809 743 5 : : : 37809 743 6 Campbell Campbell NNP 37809 743 7 and and CC 37809 743 8 Simmons Simmons NNP 37809 743 9 , , , 37809 743 10 1962:194 1962:194 CD 37809 743 11 , , , 37809 743 12 fig fig NN 37809 743 13 . . . 37809 744 1 1 1 LS 37809 744 2 . . . 37809 745 1 [ [ -LRB- 37809 745 2 Illustration illustration NN 37809 745 3 : : : 37809 745 4 FIG FIG NNP 37809 745 5 . . . 37809 746 1 18 18 CD 37809 746 2 : : : 37809 746 3 Left leave VBN 37809 746 4 to to IN 37809 746 5 right right NN 37809 746 6 . . . 37809 747 1 _ _ NNP 37809 747 2 Syrrhophus Syrrhophus NNP 37809 747 3 interorbitalis interorbitalis NN 37809 747 4 _ _ NNP 37809 747 5 ( ( -LRB- 37809 747 6 UIMNH UIMNH NNP 37809 747 7 38095 38095 CD 37809 747 8 , , , 37809 747 9 ×1.5 ×1.5 NN 37809 747 10 ) ) -RRB- 37809 747 11 , , , 37809 747 12 _ _ NNP 37809 747 13 S. S. NNP 37809 748 1 nivocolimae nivocolimae NNP 37809 748 2 _ _ NNP 37809 748 3 ( ( -LRB- 37809 748 4 LACM LACM NNP 37809 748 5 3203 3203 CD 37809 748 6 , , , 37809 748 7 ×1.3 ×1.3 LS 37809 748 8 ) ) -RRB- 37809 748 9 , , , 37809 748 10 and and CC 37809 748 11 _ _ NNP 37809 748 12 S. S. NNP 37809 749 1 teretistes teretistes NNPS 37809 749 2 _ _ NNP 37809 749 3 ( ( -LRB- 37809 749 4 KU KU NNP 37809 749 5 75263 75263 CD 37809 749 6 , , , 37809 749 7 ×1.5 ×1.5 LS 37809 749 8 ) ) -RRB- 37809 749 9 . . . 37809 749 10 ] ] -RRB- 37809 750 1 _ _ NNP 37809 750 2 Diagnosis._--Medium Diagnosis._--Medium NNP 37809 750 3 sized sized JJ 37809 750 4 frogs frog NNS 37809 750 5 , , , 37809 750 6 only only RB 37809 750 7 known know VBN 37809 750 8 male male NNP 37809 750 9 25.6 25.6 CD 37809 750 10 mm mm NNS 37809 750 11 . . . 37809 751 1 snout snout NN 37809 751 2 - - HYPH 37809 751 3 vent vent NN 37809 751 4 , , , 37809 751 5 females female VBZ 37809 751 6 20.0 20.0 CD 37809 751 7 - - HYPH 37809 751 8 26.7 26.7 CD 37809 751 9 mm mm NNS 37809 751 10 . . . 37809 752 1 snout snout NN 37809 752 2 - - HYPH 37809 752 3 vent vent NN 37809 752 4 length length NN 37809 752 5 ( ( -LRB- 37809 752 6 small small JJ 37809 752 7 sample sample NN 37809 752 8 ) ) -RRB- 37809 752 9 ; ; : 37809 752 10 vocal vocal NN 37809 752 11 slits slit NNS 37809 752 12 in in IN 37809 752 13 males male NNS 37809 752 14 ; ; : 37809 752 15 finger finger NN 37809 752 16 tips tip NNS 37809 752 17 expanded expand VBD 37809 752 18 ; ; : 37809 752 19 first first RB 37809 752 20 finger finger NN 37809 752 21 shorter short JJR 37809 752 22 than than IN 37809 752 23 second second JJ 37809 752 24 ; ; : 37809 752 25 outer outer JJ 37809 752 26 metatarsal metatarsal NN 37809 752 27 tubercle tubercle NN 37809 752 28 one one CD 37809 752 29 - - HYPH 37809 752 30 third third NN 37809 752 31 size size NN 37809 752 32 of of IN 37809 752 33 inner inner JJ 37809 752 34 ; ; : 37809 752 35 skin skin NN 37809 752 36 of of IN 37809 752 37 dorsum dorsum NN 37809 752 38 shagreened shagreene VBN 37809 752 39 , , , 37809 752 40 that that DT 37809 752 41 of of IN 37809 752 42 venter venter NN 37809 752 43 smooth smooth JJ 37809 752 44 ; ; : 37809 752 45 diameter diameter NN 37809 752 46 of of IN 37809 752 47 tympanum tympanum NN 37809 752 48 37.7 37.7 CD 37809 752 49 - - HYPH 37809 752 50 42.4 42.4 CD 37809 752 51 per per IN 37809 752 52 cent cent NN 37809 752 53 that that DT 37809 752 54 of of IN 37809 752 55 eye eye NN 37809 752 56 in in IN 37809 752 57 both both DT 37809 752 58 sexes sex NNS 37809 752 59 ; ; , 37809 752 60 pale pale JJ 37809 752 61 yellow yellow JJ 37809 752 62 - - HYPH 37809 752 63 brown brown JJ 37809 752 64 ground ground NN 37809 752 65 color color NN 37809 752 66 mottled mottle VBN 37809 752 67 with with IN 37809 752 68 brown brown JJ 37809 752 69 ; ; : 37809 752 70 limb limb NN 37809 752 71 bands band NNS 37809 752 72 broad broad RB 37809 752 73 , , , 37809 752 74 much much RB 37809 752 75 wider wide JJR 37809 752 76 than than IN 37809 752 77 narrow narrow JJ 37809 752 78 light light JJ 37809 752 79 interspaces interspace NNS 37809 752 80 ; ; : 37809 752 81 interorbital interorbital NNP 37809 752 82 bar bar NN 37809 752 83 very very RB 37809 752 84 long long RB 37809 752 85 , , , 37809 752 86 edged edge VBD 37809 752 87 with with IN 37809 752 88 dark dark JJ 37809 752 89 brown brown NNP 37809 752 90 to to IN 37809 752 91 black black NN 37809 752 92 ( ( -LRB- 37809 752 93 Fig fig NN 37809 752 94 . . . 37809 753 1 18 18 CD 37809 753 2 ) ) -RRB- 37809 753 3 . . . 37809 754 1 _ _ NNP 37809 754 2 Remarks._--Duellman Remarks._--Duellman NNP 37809 754 3 's 's POS 37809 754 4 ( ( -LRB- 37809 754 5 1958 1958 CD 37809 754 6 ) ) -RRB- 37809 754 7 measurements measurement NNS 37809 754 8 and and CC 37809 754 9 proportions proportion NNS 37809 754 10 of of IN 37809 754 11 _ _ NNP 37809 754 12 S. S. NNP 37809 755 1 interorbitalis interorbitalis NNP 37809 755 2 _ _ NNP 37809 755 3 were be VBD 37809 755 4 based base VBN 37809 755 5 exclusively exclusively RB 37809 755 6 on on IN 37809 755 7 the the DT 37809 755 8 type type NN 37809 755 9 series series NN 37809 755 10 , , , 37809 755 11 which which WDT 37809 755 12 is be VBZ 37809 755 13 composed compose VBN 37809 755 14 of of IN 37809 755 15 only only JJ 37809 755 16 females female NNS 37809 755 17 ; ; : 37809 755 18 therefore therefore RB 37809 755 19 his -PRON- PRP$ 37809 755 20 _ _ NNP 37809 755 21 interorbitalis interorbitalis NN 37809 755 22 _ _ NNP 37809 755 23 data datum NNS 37809 755 24 are be VBP 37809 755 25 not not RB 37809 755 26 comparable comparable JJ 37809 755 27 with with IN 37809 755 28 the the DT 37809 755 29 data datum NNS 37809 755 30 for for IN 37809 755 31 the the DT 37809 755 32 other other JJ 37809 755 33 species specie NNS 37809 755 34 in in IN 37809 755 35 his -PRON- PRP$ 37809 755 36 table table NN 37809 755 37 . . . 37809 756 1 Campbell Campbell NNP 37809 756 2 and and CC 37809 756 3 Simmons Simmons NNP 37809 756 4 ( ( -LRB- 37809 756 5 1962 1962 CD 37809 756 6 ) ) -RRB- 37809 756 7 collected collect VBD 37809 756 8 the the DT 37809 756 9 only only JJ 37809 756 10 known know VBN 37809 756 11 male male NN 37809 756 12 . . . 37809 757 1 The the DT 37809 757 2 type type NN 37809 757 3 series series NN 37809 757 4 was be VBD 37809 757 5 collected collect VBN 37809 757 6 beneath beneath IN 37809 757 7 rocks rock NNS 37809 757 8 in in IN 37809 757 9 a a DT 37809 757 10 stream stream NN 37809 757 11 bed bed NN 37809 757 12 ; ; : 37809 757 13 the the DT 37809 757 14 collectors collector NNS 37809 757 15 heard hear VBD 37809 757 16 calling call VBG 37809 757 17 frogs frog NNS 37809 757 18 in in IN 37809 757 19 the the DT 37809 757 20 bushes bush NNS 37809 757 21 but but CC 37809 757 22 were be VBD 37809 757 23 unable unable JJ 37809 757 24 to to TO 37809 757 25 obtain obtain VB 37809 757 26 specimens specimen NNS 37809 757 27 ( ( -LRB- 37809 757 28 Langebartel Langebartel NNP 37809 757 29 and and CC 37809 757 30 Shannon Shannon NNP 37809 757 31 , , , 37809 757 32 1956 1956 CD 37809 757 33 ) ) -RRB- 37809 757 34 . . . 37809 758 1 Campbell Campbell NNP 37809 758 2 and and CC 37809 758 3 Simmons Simmons NNP 37809 758 4 ( ( -LRB- 37809 758 5 1962 1962 CD 37809 758 6 ) ) -RRB- 37809 758 7 reported report VBD 37809 758 8 that that IN 37809 758 9 their -PRON- PRP$ 37809 758 10 specimen speciman NNS 37809 758 11 had have VBD 37809 758 12 a a DT 37809 758 13 poorly poorly RB 37809 758 14 developed develop VBN 37809 758 15 interorbital interorbital JJ 37809 758 16 bar bar NN 37809 758 17 in in IN 37809 758 18 life life NN 37809 758 19 ; ; : 37809 758 20 in in IN 37809 758 21 preservative preservative JJ 37809 758 22 the the DT 37809 758 23 bar bar NN 37809 758 24 compares compare VBZ 37809 758 25 favorably favorably RB 37809 758 26 with with IN 37809 758 27 the the DT 37809 758 28 bar bar NN 37809 758 29 in in IN 37809 758 30 the the DT 37809 758 31 female female NN 37809 758 32 ( ( -LRB- 37809 758 33 Fig fig NN 37809 758 34 . . . 37809 759 1 18 18 CD 37809 759 2 ) ) -RRB- 37809 759 3 . . . 37809 760 1 _ _ NNP 37809 760 2 Etymology._--Latin Etymology._--Latin NNP 37809 760 3 , , , 37809 760 4 in in IN 37809 760 5 reference reference NN 37809 760 6 to to IN 37809 760 7 the the DT 37809 760 8 pale pale JJ 37809 760 9 interocular interocular JJ 37809 760 10 band band NN 37809 760 11 . . . 37809 761 1 _ _ NNP 37809 761 2 Distribution._--Pacific distribution._--pacific JJ 37809 761 3 lowlands lowland NNS 37809 761 4 of of IN 37809 761 5 Sinaloa Sinaloa NNP 37809 761 6 , , , 37809 761 7 México México NNP 37809 761 8 ( ( -LRB- 37809 761 9 Fig Fig NNP 37809 761 10 . . . 37809 762 1 20 20 CD 37809 762 2 ) ) -RRB- 37809 762 3 . . . 37809 763 1 _ _ NNP 37809 763 2 Specimens Specimens NNP 37809 763 3 examined._--(10 examined._--(10 NNP 37809 763 4 ) ) -RRB- 37809 763 5 . . . 37809 764 1 MÉXICO MÉXICO NNP 37809 764 2 , , , 37809 764 3 _ _ NNP 37809 764 4 Sinaloa Sinaloa NNP 37809 764 5 _ _ NNP 37809 764 6 : : : 37809 764 7 36 36 CD 37809 764 8 mi mi JJ 37809 764 9 . . . 37809 765 1 N N NNP 37809 765 2 Mazatlán Mazatlán NNP 37809 765 3 , , , 37809 765 4 UIMNH UIMNH NNP 37809 765 5 38094 38094 CD 37809 765 6 - - SYM 37809 765 7 96 96 CD 37809 765 8 , , , 37809 765 9 67061 67061 CD 37809 765 10 ( ( -LRB- 37809 765 11 holotype holotype NNP 37809 765 12 ) ) -RRB- 37809 765 13 , , , 37809 765 14 71970 71970 CD 37809 765 15 - - SYM 37809 765 16 74 74 CD 37809 765 17 ; ; : 37809 765 18 65 65 CD 37809 765 19 mi mi UH 37809 765 20 . . . 37809 766 1 N N NNP 37809 766 2 Mazatlán Mazatlán NNP 37809 766 3 , , , 37809 766 4 LACM LACM NNP 37809 766 5 13773 13773 CD 37809 766 6 . . . 37809 767 1 = = NFP 37809 767 2 Syrrhophus Syrrhophus NNP 37809 767 3 modestus= modestus= NN 37809 767 4 Taylor Taylor NNP 37809 767 5 _ _ NNP 37809 767 6 Syrrhophus Syrrhophus NNP 37809 767 7 modestus modestus NN 37809 767 8 _ _ NNP 37809 767 9 Taylor Taylor NNP 37809 767 10 , , , 37809 767 11 1942:304 1942:304 CD 37809 767 12 - - SYM 37809 767 13 06 06 CD 37809 767 14 , , , 37809 767 15 pl pl NNP 37809 767 16 . . . 37809 768 1 29 29 CD 37809 768 2 [ [ -LRB- 37809 768 3 Holotype.--FMNH holotype.--fmnh NN 37809 768 4 100048 100048 CD 37809 768 5 ( ( -LRB- 37809 768 6 formerly formerly RB 37809 768 7 EHT EHT NNP 37809 768 8 - - HYPH 37809 768 9 HMS HMS NNP 37809 768 10 3756 3756 CD 37809 768 11 ) ) -RRB- 37809 768 12 , , , 37809 768 13 from from IN 37809 768 14 Hacienda Hacienda NNP 37809 768 15 Paso Paso NNP 37809 768 16 del del NNP 37809 768 17 Río Río NNP 37809 768 18 , , , 37809 768 19 Colima Colima NNP 37809 768 20 , , , 37809 768 21 México México NNP 37809 768 22 ; ; , 37809 768 23 collected collect VBN 37809 768 24 on on IN 37809 768 25 July July NNP 37809 768 26 8 8 CD 37809 768 27 , , , 37809 768 28 1935 1935 CD 37809 768 29 , , , 37809 768 30 by by IN 37809 768 31 Hobart Hobart NNP 37809 768 32 M. M. NNP 37809 768 33 Smith Smith NNP 37809 768 34 ] ] -RRB- 37809 768 35 . . . 37809 769 1 Smith Smith NNP 37809 769 2 and and CC 37809 769 3 Taylor Taylor NNP 37809 769 4 , , , 37809 769 5 1948:49 1948:49 CD 37809 769 6 - - SYM 37809 769 7 50 50 CD 37809 769 8 . . . 37809 770 1 _ _ NNP 37809 770 2 Syrrhophus Syrrhophus NNP 37809 770 3 modestus modestus NN 37809 770 4 modestus modestus NN 37809 770 5 _ _ NNP 37809 770 6 : : : 37809 770 7 Duellman Duellman NNP 37809 770 8 , , , 37809 770 9 1958:2 1958:2 CD 37809 770 10 - - SYM 37809 770 11 5 5 CD 37809 770 12 , , , 37809 770 13 7 7 CD 37809 770 14 , , , 37809 770 15 14 14 CD 37809 770 16 , , , 37809 770 17 pl pl NNP 37809 770 18 . . . 37809 771 1 1 1 CD 37809 771 2 , , , 37809 771 3 fig fig VB 37809 771 4 . . . 37809 772 1 1 1 LS 37809 772 2 . . . 37809 773 1 Gorham Gorham NNP 37809 773 2 , , , 37809 773 3 1966:166 1966:166 CD 37809 773 4 . . . 37809 774 1 _ _ NNP 37809 774 2 Diagnosis._--Small Diagnosis._--Small NNP 37809 774 3 frogs frog NNS 37809 774 4 , , , 37809 774 5 males male VBZ 37809 774 6 15.8 15.8 CD 37809 774 7 - - SYM 37809 774 8 20.1 20.1 CD 37809 774 9 mm mm NNS 37809 774 10 . . . 37809 775 1 snout snout NN 37809 775 2 - - HYPH 37809 775 3 vent vent NN 37809 775 4 length length NN 37809 775 5 , , , 37809 775 6 single single JJ 37809 775 7 female female JJ 37809 775 8 18.5 18.5 CD 37809 775 9 mm mm NNS 37809 775 10 . . . 37809 776 1 ; ; : 37809 776 2 vocal vocal JJ 37809 776 3 slits slit NNS 37809 776 4 present present JJ 37809 776 5 in in IN 37809 776 6 males male NNS 37809 776 7 ; ; : 37809 776 8 finger finger NN 37809 776 9 tips tip NNS 37809 776 10 widely widely RB 37809 776 11 expanded expand VBN 37809 776 12 ; ; : 37809 776 13 first first RB 37809 776 14 finger finger NN 37809 776 15 shorter short JJR 37809 776 16 than than IN 37809 776 17 second second JJ 37809 776 18 ; ; : 37809 776 19 inner inner JJ 37809 776 20 metatarsal metatarsal NN 37809 776 21 tubercle tubercle NN 37809 776 22 about about IN 37809 776 23 three three CD 37809 776 24 times time NNS 37809 776 25 size size NN 37809 776 26 of of IN 37809 776 27 outer outer NN 37809 776 28 ; ; : 37809 776 29 skin skin NN 37809 776 30 of of IN 37809 776 31 dorsum dorsum NN 37809 776 32 shagreened shagreene VBN 37809 776 33 , , , 37809 776 34 that that DT 37809 776 35 of of IN 37809 776 36 venter venter NN 37809 776 37 smooth smooth JJ 37809 776 38 ; ; : 37809 776 39 tympanum tympanum NNP 37809 776 40 concealed conceal VBD 37809 776 41 ; ; : 37809 776 42 pale pale JJ 37809 776 43 cream cream NN 37809 776 44 in in IN 37809 776 45 preservative preservative JJ 37809 776 46 with with IN 37809 776 47 dark dark JJ 37809 776 48 brown brown JJ 37809 776 49 spots spot NNS 37809 776 50 ; ; : 37809 776 51 limbs limb NNS 37809 776 52 banded band VBD 37809 776 53 ; ; : 37809 776 54 bands band NNS 37809 776 55 on on IN 37809 776 56 forearm forearm NN 37809 776 57 and and CC 37809 776 58 thigh thigh NN 37809 776 59 poorly poorly RB 37809 776 60 developed develop VBN 37809 776 61 or or CC 37809 776 62 absent absent JJ 37809 776 63 ; ; : 37809 776 64 interorbital interorbital NNP 37809 776 65 bar bar NN 37809 776 66 absent absent NN 37809 776 67 . . . 37809 777 1 _ _ NNP 37809 777 2 Remarks._--The Remarks._--The NNP 37809 777 3 tympanum tympanum NN 37809 777 4 is be VBZ 37809 777 5 concealed conceal VBN 37809 777 6 in in IN 37809 777 7 _ _ NNP 37809 777 8 S. S. NNP 37809 778 1 modestus modestus NNP 37809 778 2 _ _ NNP 37809 778 3 , , , 37809 778 4 _ _ NNP 37809 778 5 S. S. NNP 37809 779 1 nivocolimae nivocolimae NNP 37809 779 2 _ _ NNP 37809 779 3 , , , 37809 779 4 _ _ NNP 37809 779 5 S. S. NNP 37809 780 1 pallidus pallidus NNP 37809 780 2 _ _ NNP 37809 780 3 , , , 37809 780 4 _ _ NNP 37809 780 5 S. S. NNP 37809 781 1 teretistes teretistes NNPS 37809 781 2 _ _ NNP 37809 781 3 , , , 37809 781 4 and and CC 37809 781 5 to to IN 37809 781 6 a a DT 37809 781 7 lesser less JJR 37809 781 8 degree degree NN 37809 781 9 in in IN 37809 781 10 _ _ NNP 37809 781 11 S. S. NNP 37809 782 1 interorbitalis interorbitalis NNP 37809 782 2 _ _ NNP 37809 782 3 . . . 37809 783 1 However however RB 37809 783 2 , , , 37809 783 3 if if IN 37809 783 4 the the DT 37809 783 5 specimen speciman NNS 37809 783 6 is be VBZ 37809 783 7 permitted permit VBN 37809 783 8 to to TO 37809 783 9 dry dry VB 37809 783 10 slightly slightly RB 37809 783 11 , , , 37809 783 12 the the DT 37809 783 13 annulus annulus JJ 37809 783 14 tympanicus tympanicus NN 37809 783 15 becomes become VBZ 37809 783 16 visible visible JJ 37809 783 17 through through IN 37809 783 18 the the DT 37809 783 19 skin skin NN 37809 783 20 and and CC 37809 783 21 a a DT 37809 783 22 tympanum tympanum NN 37809 783 23 / / SYM 37809 783 24 eye eye NN 37809 783 25 ratio ratio NN 37809 783 26 can can MD 37809 783 27 be be VB 37809 783 28 computed compute VBN 37809 783 29 . . . 37809 784 1 One one CD 37809 784 2 of of IN 37809 784 3 the the DT 37809 784 4 few few JJ 37809 784 5 cases case NNS 37809 784 6 of of IN 37809 784 7 sympatry sympatry NN 37809 784 8 within within IN 37809 784 9 the the DT 37809 784 10 genus genus NNP 37809 784 11 _ _ NNP 37809 784 12 Syrrhophus Syrrhophus NNP 37809 784 13 _ _ NNP 37809 784 14 involves involve VBZ 37809 784 15 this this DT 37809 784 16 species species NN 37809 784 17 ; ; : 37809 784 18 _ _ NNP 37809 784 19 modestus modestus NN 37809 784 20 _ _ NNP 37809 784 21 and and CC 37809 784 22 _ _ NNP 37809 784 23 nivocolimae nivocolimae NN 37809 784 24 _ _ NNP 37809 784 25 are be VBP 37809 784 26 known know VBN 37809 784 27 to to TO 37809 784 28 be be VB 37809 784 29 sympatric sympatric JJ 37809 784 30 at at IN 37809 784 31 one one CD 37809 784 32 locality locality NN 37809 784 33 in in IN 37809 784 34 southwestern southwestern JJ 37809 784 35 Jalisco Jalisco NNP 37809 784 36 , , , 37809 784 37 México México NNP 37809 784 38 . . . 37809 785 1 Duellman Duellman NNP 37809 785 2 ( ( -LRB- 37809 785 3 1958 1958 CD 37809 785 4 ) ) -RRB- 37809 785 5 used use VBD 37809 785 6 the the DT 37809 785 7 trinomial trinomial NN 37809 785 8 for for IN 37809 785 9 this this DT 37809 785 10 population population NN 37809 785 11 and and CC 37809 785 12 named name VBD 37809 785 13 a a DT 37809 785 14 new new JJ 37809 785 15 subspecies subspecie NNS 37809 785 16 , , , 37809 785 17 _ _ NNP 37809 785 18 pallidus pallidus NNP 37809 785 19 _ _ NNP 37809 785 20 , , , 37809 785 21 from from IN 37809 785 22 Nayarit Nayarit NNP 37809 785 23 . . . 37809 786 1 I -PRON- PRP 37809 786 2 consider consider VBP 37809 786 3 _ _ NNP 37809 786 4 pallidus pallidus NNP 37809 786 5 _ _ NNP 37809 786 6 to to TO 37809 786 7 be be VB 37809 786 8 specifically specifically RB 37809 786 9 distinct distinct JJ 37809 786 10 from from IN 37809 786 11 _ _ NNP 37809 786 12 modestus modestus NN 37809 786 13 _ _ NNP 37809 786 14 because because IN 37809 786 15 there there EX 37809 786 16 is be VBZ 37809 786 17 no no DT 37809 786 18 evidence evidence NN 37809 786 19 of of IN 37809 786 20 genetic genetic JJ 37809 786 21 exchange exchange NN 37809 786 22 , , , 37809 786 23 and and CC 37809 786 24 there there EX 37809 786 25 is be VBZ 37809 786 26 no no DT 37809 786 27 overlap overlap NN 37809 786 28 in in IN 37809 786 29 the the DT 37809 786 30 distinguishing distinguish VBG 37809 786 31 morphological morphological JJ 37809 786 32 features feature NNS 37809 786 33 . . . 37809 787 1 I -PRON- PRP 37809 787 2 do do VBP 37809 787 3 consider consider VB 37809 787 4 the the DT 37809 787 5 two two CD 37809 787 6 populations population NNS 37809 787 7 to to TO 37809 787 8 be be VB 37809 787 9 closely closely RB 37809 787 10 related related JJ 37809 787 11 but but CC 37809 787 12 feel feel VBP 37809 787 13 the the DT 37809 787 14 interrelationships interrelationship NNS 37809 787 15 between between IN 37809 787 16 _ _ NNP 37809 787 17 modestus modestus NNP 37809 787 18 _ _ NNP 37809 787 19 , , , 37809 787 20 _ _ NNP 37809 787 21 pallidus pallidus NNP 37809 787 22 _ _ NNP 37809 787 23 , , , 37809 787 24 _ _ NNP 37809 787 25 nivocolimae nivocolimae NN 37809 787 26 _ _ NNP 37809 787 27 , , , 37809 787 28 and and CC 37809 787 29 _ _ NNP 37809 787 30 teretistes teretistes NNPS 37809 787 31 _ _ NNP 37809 787 32 are be VBP 37809 787 33 more more RBR 37809 787 34 complex complex JJ 37809 787 35 than than IN 37809 787 36 would would MD 37809 787 37 be be VB 37809 787 38 indicated indicate VBN 37809 787 39 by by IN 37809 787 40 the the DT 37809 787 41 use use NN 37809 787 42 of of IN 37809 787 43 trinomials trinomial NNS 37809 787 44 . . . 37809 788 1 The the DT 37809 788 2 sympatric sympatric JJ 37809 788 3 occurrence occurrence NN 37809 788 4 of of IN 37809 788 5 _ _ NNP 37809 788 6 modestus modestus NN 37809 788 7 _ _ NNP 37809 788 8 and and CC 37809 788 9 _ _ NNP 37809 788 10 nivocolimae nivocolimae NN 37809 788 11 _ _ NNP 37809 788 12 is be VBZ 37809 788 13 significant significant JJ 37809 788 14 ; ; : 37809 788 15 morphologically morphologically RB 37809 788 16 , , , 37809 788 17 they -PRON- PRP 37809 788 18 might may MD 37809 788 19 otherwise otherwise RB 37809 788 20 be be VB 37809 788 21 regarded regard VBN 37809 788 22 as as IN 37809 788 23 subspecies subspecie NNS 37809 788 24 . . . 37809 789 1 Although although IN 37809 789 2 allopatric allopatric JJ 37809 789 3 , , , 37809 789 4 similar similar JJ 37809 789 5 arguments argument NNS 37809 789 6 could could MD 37809 789 7 be be VB 37809 789 8 advanced advance VBN 37809 789 9 for for IN 37809 789 10 the the DT 37809 789 11 morphologically morphologically RB 37809 789 12 similar similar JJ 37809 789 13 _ _ NNP 37809 789 14 pallidus pallidus NNP 37809 789 15 _ _ NNP 37809 789 16 and and CC 37809 789 17 _ _ NNP 37809 789 18 teretistes teretistes NNPS 37809 789 19 _ _ NNP 37809 789 20 . . . 37809 790 1 The the DT 37809 790 2 four four CD 37809 790 3 are be VBP 37809 790 4 here here RB 37809 790 5 afforded afford VBN 37809 790 6 species specie NNS 37809 790 7 rank rank NN 37809 790 8 since since IN 37809 790 9 morphological morphological JJ 37809 790 10 similarity similarity NN 37809 790 11 and and CC 37809 790 12 allopatry allopatry NN 37809 790 13 are be VBP 37809 790 14 not not RB 37809 790 15 sufficient sufficient JJ 37809 790 16 grounds ground NNS 37809 790 17 for for IN 37809 790 18 the the DT 37809 790 19 assumption assumption NN 37809 790 20 of of IN 37809 790 21 genetic genetic JJ 37809 790 22 exchange exchange NN 37809 790 23 . . . 37809 791 1 [ [ -LRB- 37809 791 2 Illustration illustration NN 37809 791 3 : : : 37809 791 4 FIG FIG NNP 37809 791 5 . . . 37809 792 1 19 19 CD 37809 792 2 : : : 37809 792 3 _ _ NNP 37809 792 4 Syrrhophus Syrrhophus NNP 37809 792 5 modestus modestus NN 37809 792 6 _ _ NNP 37809 792 7 [ [ -LRB- 37809 792 8 left leave VBN 37809 792 9 , , , 37809 792 10 UMMZ UMMZ NNP 37809 792 11 115447 115447 CD 37809 792 12 ( ( -LRB- 37809 792 13 WED WED NNP 37809 792 14 11155 11155 CD 37809 792 15 ) ) -RRB- 37809 792 16 ] ] -RRB- 37809 792 17 and and CC 37809 792 18 _ _ NNP 37809 792 19 S. S. NNP 37809 793 1 pallidus pallidus NNP 37809 793 2 _ _ NNP 37809 793 3 ( ( -LRB- 37809 793 4 right right UH 37809 793 5 , , , 37809 793 6 UMMZ UMMZ NNP 37809 793 7 115453 115453 CD 37809 793 8 ) ) -RRB- 37809 793 9 . . . 37809 794 1 ×2.2 ×2.2 UH 37809 794 2 . . . 37809 794 3 ] ] -RRB- 37809 795 1 _ _ NNP 37809 795 2 Etymology._--Latin Etymology._--Latin NNP 37809 795 3 , , , 37809 795 4 meaning mean VBG 37809 795 5 unassuming unassuming NN 37809 795 6 , , , 37809 795 7 modest modest JJ 37809 795 8 , , , 37809 795 9 in in IN 37809 795 10 reference reference NN 37809 795 11 to to IN 37809 795 12 the the DT 37809 795 13 small small JJ 37809 795 14 size size NN 37809 795 15 of of IN 37809 795 16 the the DT 37809 795 17 species specie NNS 37809 795 18 . . . 37809 796 1 _ _ NNP 37809 796 2 Distribution._--Low Distribution._--Low NNP 37809 796 3 elevations elevation NNS 37809 796 4 ( ( -LRB- 37809 796 5 up up IN 37809 796 6 to to TO 37809 796 7 700 700 CD 37809 796 8 meters meter NNS 37809 796 9 ) ) -RRB- 37809 796 10 in in IN 37809 796 11 the the DT 37809 796 12 lowlands lowland NNS 37809 796 13 and and CC 37809 796 14 foothills foothill NNS 37809 796 15 of of IN 37809 796 16 Colima Colima NNP 37809 796 17 and and CC 37809 796 18 southwestern southwestern JJ 37809 796 19 Jalisco Jalisco NNP 37809 796 20 , , , 37809 796 21 México México NNP 37809 796 22 ( ( -LRB- 37809 796 23 Fig Fig NNP 37809 796 24 . . . 37809 797 1 20 20 CD 37809 797 2 ) ) -RRB- 37809 797 3 . . . 37809 798 1 _ _ NNP 37809 798 2 Specimens Specimens NNP 37809 798 3 examined._--(14 examined._--(14 NNP 37809 798 4 ) ) -RRB- 37809 798 5 . . . 37809 799 1 MÉXICO MÉXICO NNP 37809 799 2 , , , 37809 799 3 _ _ NNP 37809 799 4 Colima Colima NNP 37809 799 5 _ _ NNP 37809 799 6 : : : 37809 799 7 Hda Hda NNP 37809 799 8 . . . 37809 800 1 Paso Paso NNP 37809 800 2 del del NNP 37809 800 3 Río Río NNP 37809 800 4 , , , 37809 800 5 FMNH FMNH NNP 37809 800 6 100048 100048 CD 37809 800 7 ( ( -LRB- 37809 800 8 holotype holotype NNP 37809 800 9 ) ) -RRB- 37809 800 10 , , , 37809 800 11 100167 100167 CD 37809 800 12 , , , 37809 800 13 100299 100299 CD 37809 800 14 , , , 37809 800 15 UIMNH UIMNH NNP 37809 800 16 13300 13300 CD 37809 800 17 , , , 37809 800 18 UMMZ UMMZ NNP 37809 800 19 110877 110877 CD 37809 800 20 ( ( -LRB- 37809 800 21 2 2 CD 37809 800 22 ) ) -RRB- 37809 800 23 , , , 37809 800 24 USNM USNM NNP 37809 800 25 139729 139729 CD 37809 800 26 ; ; : 37809 800 27 7.2 7.2 CD 37809 800 28 km km NN 37809 800 29 . . . 37809 801 1 SW SW NNP 37809 801 2 Tecolapa Tecolapa NNP 37809 801 3 , , , 37809 801 4 UMMZ UMMZ NNP 37809 801 5 115477 115477 CD 37809 801 6 ( ( -LRB- 37809 801 7 4 4 CD 37809 801 8 ) ) -RRB- 37809 801 9 ; ; : 37809 801 10 _ _ NNP 37809 801 11 Jalisco Jalisco NNP 37809 801 12 _ _ NNP 37809 801 13 : : : 37809 801 14 17.6 17.6 CD 37809 801 15 km km NNS 37809 801 16 . . . 37809 802 1 SW SW NNP 37809 802 2 Autlan Autlan NNP 37809 802 3 , , , 37809 802 4 606 606 CD 37809 802 5 m. m. NN 37809 802 6 , , , 37809 802 7 KU KU NNP 37809 802 8 102627 102627 CD 37809 802 9 ; ; : 37809 802 10 3.2 3.2 CD 37809 802 11 km km NN 37809 802 12 . . . 37809 803 1 N N NNP 37809 803 2 La La NNP 37809 803 3 Resolana Resolana NNP 37809 803 4 , , , 37809 803 5 UMMZ UMMZ NNP 37809 803 6 102100 102100 CD 37809 803 7 ; ; : 37809 803 8 Bahía Bahía NNP 37809 803 9 Tenacatita Tenacatita NNP 37809 803 10 , , , 37809 803 11 UMMZ UMMZ NNP 37809 803 12 84264 84264 CD 37809 803 13 . . . 37809 804 1 = = NFP 37809 804 2 Syrrhophus Syrrhophus NNP 37809 804 3 nivocolimae= nivocolimae= NN 37809 804 4 Dixon Dixon NNP 37809 804 5 and and CC 37809 804 6 Webb Webb NNP 37809 804 7 _ _ NNP 37809 804 8 Syrrhophus Syrrhophus NNP 37809 804 9 nivocolimae nivocolimae NN 37809 804 10 _ _ NNP 37809 804 11 Dixon Dixon NNP 37809 804 12 and and CC 37809 804 13 Webb Webb NNP 37809 804 14 , , , 37809 804 15 1966:1 1966:1 CD 37809 804 16 - - SYM 37809 804 17 4 4 CD 37809 804 18 , , , 37809 804 19 Fig Fig NNP 37809 804 20 . . . 37809 805 1 1 1 LS 37809 805 2 [ [ -LRB- 37809 805 3 Holotype.--LACM Holotype.--LACM NNP 37809 805 4 3200 3200 CD 37809 805 5 , , , 37809 805 6 from from IN 37809 805 7 Nevado Nevado NNP 37809 805 8 de de FW 37809 805 9 Colima Colima NNP 37809 805 10 ( ( -LRB- 37809 805 11 6 6 CD 37809 805 12 airline airline NN 37809 805 13 miles mile NNS 37809 805 14 west west RB 37809 805 15 of of IN 37809 805 16 Atenquique Atenquique NNP 37809 805 17 ) ) -RRB- 37809 805 18 , , , 37809 805 19 Jalisco Jalisco NNP 37809 805 20 , , , 37809 805 21 México México NNP 37809 805 22 , , , 37809 805 23 7800 7800 CD 37809 805 24 feet foot NNS 37809 805 25 ; ; , 37809 805 26 collected collect VBN 37809 805 27 on on IN 37809 805 28 July July NNP 37809 805 29 20 20 CD 37809 805 30 , , , 37809 805 31 1964 1964 CD 37809 805 32 , , , 37809 805 33 by by IN 37809 805 34 Robert Robert NNP 37809 805 35 G. G. NNP 37809 805 36 Webb Webb NNP 37809 805 37 ] ] -RRB- 37809 805 38 . . . 37809 806 1 _ _ NNP 37809 806 2 Diagnosis._--Small Diagnosis._--Small NNP 37809 806 3 frogs frog NNS 37809 806 4 , , , 37809 806 5 males male VBZ 37809 806 6 18.5 18.5 CD 37809 806 7 - - SYM 37809 806 8 21.1 21.1 CD 37809 806 9 mm mm CD 37809 806 10 . . . 37809 807 1 snout snout NN 37809 807 2 - - HYPH 37809 807 3 vent vent NN 37809 807 4 length length NN 37809 807 5 , , , 37809 807 6 only only RB 37809 807 7 known know VBN 37809 807 8 female female JJ 37809 807 9 24.1 24.1 CD 37809 807 10 mm mm NNS 37809 807 11 . . . 37809 808 1 snout snout NN 37809 808 2 - - HYPH 37809 808 3 vent vent NN 37809 808 4 ; ; : 37809 808 5 vocal vocal JJ 37809 808 6 slits slit NNS 37809 808 7 present present JJ 37809 808 8 in in IN 37809 808 9 males male NNS 37809 808 10 ; ; : 37809 808 11 finger finger NN 37809 808 12 tips tip NNS 37809 808 13 widely widely RB 37809 808 14 expanded expand VBN 37809 808 15 ; ; : 37809 808 16 first first RB 37809 808 17 finger finger NN 37809 808 18 shorter short JJR 37809 808 19 than than IN 37809 808 20 second second JJ 37809 808 21 ; ; : 37809 808 22 inner inner JJ 37809 808 23 metatarsal metatarsal NN 37809 808 24 tubercle tubercle NN 37809 808 25 about about IN 37809 808 26 three three CD 37809 808 27 times time NNS 37809 808 28 size size NN 37809 808 29 of of IN 37809 808 30 outer outer NN 37809 808 31 ; ; : 37809 808 32 skin skin NN 37809 808 33 of of IN 37809 808 34 dorsum dorsum NN 37809 808 35 warty warty NN 37809 808 36 , , , 37809 808 37 that that DT 37809 808 38 of of IN 37809 808 39 venter venter NN 37809 808 40 smooth smooth JJ 37809 808 41 ; ; : 37809 808 42 tympanum tympanum NNP 37809 808 43 concealed conceal VBD 37809 808 44 , , , 37809 808 45 its -PRON- PRP$ 37809 808 46 diameter diameter NN 37809 808 47 30.0 30.0 CD 37809 808 48 - - HYPH 37809 808 49 39.3 39.3 CD 37809 808 50 per per IN 37809 808 51 cent cent NN 37809 808 52 that that DT 37809 808 53 of of IN 37809 808 54 eye eye NN 37809 808 55 in in IN 37809 808 56 males male NNS 37809 808 57 ; ; : 37809 808 58 mid mid JJ 37809 808 59 - - JJ 37809 808 60 dorsal dorsal JJ 37809 808 61 brown brown JJ 37809 808 62 band band NN 37809 808 63 from from IN 37809 808 64 interorbital interorbital NNP 37809 808 65 bar bar NN 37809 808 66 to to IN 37809 808 67 anus anus NN 37809 808 68 ; ; : 37809 808 69 bands band NNS 37809 808 70 on on IN 37809 808 71 limbs limb NNS 37809 808 72 narrow narrow JJ 37809 808 73 , , , 37809 808 74 dark dark JJ 37809 808 75 bands band NNS 37809 808 76 less less JJR 37809 808 77 than than IN 37809 808 78 one one CD 37809 808 79 - - HYPH 37809 808 80 half half NN 37809 808 81 width width NN 37809 808 82 of of IN 37809 808 83 light light JJ 37809 808 84 bands band NNS 37809 808 85 , , , 37809 808 86 upper upper JJ 37809 808 87 arm arm NN 37809 808 88 not not RB 37809 808 89 banded band VBN 37809 808 90 ; ; : 37809 808 91 narrow narrow JJ 37809 808 92 interorbital interorbital NNP 37809 808 93 light light NNP 37809 808 94 bar bar NN 37809 808 95 . . . 37809 809 1 _ _ NNP 37809 809 2 Remarks._--This remarks._--this CD 37809 809 3 species species NN 37809 809 4 is be VBZ 37809 809 5 closely closely RB 37809 809 6 related relate VBN 37809 809 7 to to IN 37809 809 8 _ _ NNP 37809 809 9 S. S. NNP 37809 810 1 modestus modestus NNP 37809 810 2 _ _ NNP 37809 810 3 and and CC 37809 810 4 differs differ VBZ 37809 810 5 in in IN 37809 810 6 color color NN 37809 810 7 pattern pattern NN 37809 810 8 and and CC 37809 810 9 degree degree NN 37809 810 10 of of IN 37809 810 11 wartiness wartiness NN 37809 810 12 of of IN 37809 810 13 the the DT 37809 810 14 skin skin NN 37809 810 15 . . . 37809 811 1 Dixon Dixon NNP 37809 811 2 and and CC 37809 811 3 Webb Webb NNP 37809 811 4 ( ( -LRB- 37809 811 5 1966 1966 CD 37809 811 6 ) ) -RRB- 37809 811 7 held hold VBD 37809 811 8 that that IN 37809 811 9 _ _ NNP 37809 811 10 nivocolimae nivocolimae NN 37809 811 11 _ _ NNP 37809 811 12 had have VBD 37809 811 13 no no DT 37809 811 14 close close JJ 37809 811 15 relatives relative NNS 37809 811 16 , , , 37809 811 17 but but CC 37809 811 18 the the DT 37809 811 19 condition condition NN 37809 811 20 of of IN 37809 811 21 the the DT 37809 811 22 tympanum tympanum NN 37809 811 23 , , , 37809 811 24 size size NN 37809 811 25 , , , 37809 811 26 nature nature NN 37809 811 27 of of IN 37809 811 28 the the DT 37809 811 29 outer outer JJ 37809 811 30 palmar palmar NNP 37809 811 31 tubercle tubercle NN 37809 811 32 , , , 37809 811 33 relative relative JJ 37809 811 34 sizes size NNS 37809 811 35 of of IN 37809 811 36 the the DT 37809 811 37 metatarsal metatarsal NN 37809 811 38 tubercles tubercle NNS 37809 811 39 , , , 37809 811 40 and and CC 37809 811 41 shape shape NN 37809 811 42 and and CC 37809 811 43 size size NN 37809 811 44 of of IN 37809 811 45 the the DT 37809 811 46 digital digital JJ 37809 811 47 pads pad NNS 37809 811 48 all all DT 37809 811 49 point point VBP 37809 811 50 to to IN 37809 811 51 a a DT 37809 811 52 close close JJ 37809 811 53 relationship relationship NN 37809 811 54 between between IN 37809 811 55 _ _ NNP 37809 811 56 S. S. NNP 37809 812 1 modestus modestus NNP 37809 812 2 _ _ NNP 37809 812 3 , , , 37809 812 4 _ _ NNP 37809 812 5 S. S. NNP 37809 813 1 nivocolimae nivocolimae NNP 37809 813 2 _ _ NNP 37809 813 3 , , , 37809 813 4 and and CC 37809 813 5 _ _ NNP 37809 813 6 S. S. NNP 37809 814 1 pallidus pallidus NNP 37809 814 2 _ _ NNP 37809 814 3 . . . 37809 815 1 [ [ -LRB- 37809 815 2 Illustration illustration NN 37809 815 3 : : : 37809 815 4 FIG FIG NNP 37809 815 5 . . . 37809 816 1 20 20 CD 37809 816 2 : : : 37809 816 3 Distribution distribution NN 37809 816 4 of of IN 37809 816 5 the the DT 37809 816 6 species specie NNS 37809 816 7 of of IN 37809 816 8 the the DT 37809 816 9 _ _ NNP 37809 816 10 modestus modestus NN 37809 816 11 _ _ NNP 37809 816 12 group group NN 37809 816 13 : : : 37809 816 14 _ _ NNP 37809 816 15 interorbitalis interorbitalis NNP 37809 816 16 _ _ NNP 37809 816 17 ( ( -LRB- 37809 816 18 open open JJ 37809 816 19 circles circle NNS 37809 816 20 ) ) -RRB- 37809 816 21 , , , 37809 816 22 _ _ NNP 37809 816 23 teretistes teretiste NNS 37809 816 24 _ _ NNP 37809 816 25 ( ( -LRB- 37809 816 26 solid solid JJ 37809 816 27 circles circle NNS 37809 816 28 ) ) -RRB- 37809 816 29 , , , 37809 816 30 _ _ NNP 37809 816 31 modestus modestus NN 37809 816 32 _ _ NNP 37809 816 33 ( ( -LRB- 37809 816 34 open open JJ 37809 816 35 triangles triangle NNS 37809 816 36 ) ) -RRB- 37809 816 37 , , , 37809 816 38 _ _ NNP 37809 816 39 pallidus pallidus NNP 37809 816 40 _ _ NNP 37809 816 41 ( ( -LRB- 37809 816 42 solid solid JJ 37809 816 43 triangles triangle NNS 37809 816 44 ) ) -RRB- 37809 816 45 and and CC 37809 816 46 _ _ NNP 37809 816 47 nivocolimae nivocolimae NN 37809 816 48 _ _ NNP 37809 816 49 ( ( -LRB- 37809 816 50 square square NNP 37809 816 51 ) ) -RRB- 37809 816 52 . . . 37809 817 1 Arrow Arrow NNP 37809 817 2 indicates indicate VBZ 37809 817 3 locality locality NN 37809 817 4 of of IN 37809 817 5 sympatry sympatry NN 37809 817 6 between between IN 37809 817 7 _ _ NNP 37809 817 8 modestus modestus NNP 37809 817 9 _ _ NNP 37809 817 10 and and CC 37809 817 11 _ _ NNP 37809 817 12 nivocolimae nivocolimae NN 37809 817 13 _ _ NNP 37809 817 14 . . . 37809 818 1 Solid solid JJ 37809 818 2 line line NN 37809 818 3 about about IN 37809 818 4 the the DT 37809 818 5 localities locality NNS 37809 818 6 for for IN 37809 818 7 _ _ NNP 37809 818 8 interorbitalis interorbitalis NNP 37809 818 9 _ _ NNP 37809 818 10 is be VBZ 37809 818 11 a a DT 37809 818 12 range range NN 37809 818 13 estimate estimate NN 37809 818 14 based base VBN 37809 818 15 on on IN 37809 818 16 call call NN 37809 818 17 records record NNS 37809 818 18 and and CC 37809 818 19 specimens specimen NNS 37809 818 20 examined examine VBN 37809 818 21 . . . 37809 818 22 ] ] -RRB- 37809 819 1 Dixon Dixon NNP 37809 819 2 and and CC 37809 819 3 Webb Webb NNP 37809 819 4 ( ( -LRB- 37809 819 5 1966 1966 CD 37809 819 6 ) ) -RRB- 37809 819 7 reported report VBD 37809 819 8 that that IN 37809 819 9 _ _ NNP 37809 819 10 S. S. NNP 37809 820 1 nivocolimae nivocolimae NNP 37809 820 2 _ _ NNP 37809 820 3 has have VBZ 37809 820 4 a a DT 37809 820 5 large large JJ 37809 820 6 tympanum tympanum NN 37809 820 7 ( ( -LRB- 37809 820 8 50.0 50.0 CD 37809 820 9 - - HYPH 37809 820 10 59.0 59.0 CD 37809 820 11 per per IN 37809 820 12 cent cent NN 37809 820 13 diameter diameter NN 37809 820 14 of of IN 37809 820 15 eye eye NN 37809 820 16 ) ) -RRB- 37809 820 17 . . . 37809 821 1 However however RB 37809 821 2 , , , 37809 821 3 my -PRON- PRP$ 37809 821 4 examination examination NN 37809 821 5 of of IN 37809 821 6 the the DT 37809 821 7 type type NN 37809 821 8 series series NN 37809 821 9 and and CC 37809 821 10 several several JJ 37809 821 11 other other JJ 37809 821 12 specimens specimen NNS 37809 821 13 from from IN 37809 821 14 Jalisco Jalisco NNP 37809 821 15 reveals reveal VBZ 37809 821 16 that that IN 37809 821 17 the the DT 37809 821 18 largest large JJS 37809 821 19 tympanum tympanum NN 37809 821 20 / / SYM 37809 821 21 eye eye NN 37809 821 22 ratio ratio NN 37809 821 23 is be VBZ 37809 821 24 39.3 39.3 CD 37809 821 25 per per IN 37809 821 26 cent cent NN 37809 821 27 . . . 37809 822 1 Therefore therefore RB 37809 822 2 , , , 37809 822 3 the the DT 37809 822 4 tympanum tympanum NN 37809 822 5 / / SYM 37809 822 6 eye eye NN 37809 822 7 ratio ratio NN 37809 822 8 in in IN 37809 822 9 _ _ NNP 37809 822 10 S. S. NNP 37809 823 1 nivocolimae nivocolimae NNP 37809 823 2 _ _ NNP 37809 823 3 is be VBZ 37809 823 4 in in IN 37809 823 5 agreement agreement NN 37809 823 6 with with IN 37809 823 7 those those DT 37809 823 8 for for IN 37809 823 9 _ _ NNP 37809 823 10 S. S. NNP 37809 824 1 modestus modestus NNP 37809 824 2 _ _ NNP 37809 824 3 , , , 37809 824 4 _ _ NNP 37809 824 5 S. S. NNP 37809 825 1 pallidus pallidus NNP 37809 825 2 _ _ NNP 37809 825 3 , , , 37809 825 4 and and CC 37809 825 5 _ _ NNP 37809 825 6 S. S. NNP 37809 826 1 teretistes teretistes NNPS 37809 826 2 _ _ NNP 37809 826 3 ( ( -LRB- 37809 826 4 Table Table NNP 37809 826 5 6 6 CD 37809 826 6 ) ) -RRB- 37809 826 7 . . . 37809 827 1 _ _ NNP 37809 827 2 Etymology._--_niv Etymology._--_niv NNP 37809 827 3 _ _ NNP 37809 827 4 , , , 37809 827 5 Latin Latin NNP 37809 827 6 , , , 37809 827 7 and and CC 37809 827 8 Colima Colima NNP 37809 827 9 ( ( -LRB- 37809 827 10 Nevado Nevado NNP 37809 827 11 de de NNP 37809 827 12 ) ) -RRB- 37809 827 13 , , , 37809 827 14 meaning mean VBG 37809 827 15 high high RB 37809 827 16 on on IN 37809 827 17 the the DT 37809 827 18 volcano volcano NN 37809 827 19 , , , 37809 827 20 in in IN 37809 827 21 reference reference NN 37809 827 22 to to IN 37809 827 23 the the DT 37809 827 24 higher high JJR 37809 827 25 distribution distribution NN 37809 827 26 of of IN 37809 827 27 this this DT 37809 827 28 species specie NNS 37809 827 29 ( ( -LRB- 37809 827 30 around around RB 37809 827 31 2000 2000 CD 37809 827 32 meters meter NNS 37809 827 33 ) ) -RRB- 37809 827 34 than than IN 37809 827 35 other other JJ 37809 827 36 members member NNS 37809 827 37 of of IN 37809 827 38 the the DT 37809 827 39 group group NN 37809 827 40 . . . 37809 828 1 _ _ NNP 37809 828 2 Distribution._--Known Distribution._--Known NNP 37809 828 3 from from IN 37809 828 4 southwestern southwestern JJ 37809 828 5 Jalisco Jalisco NNP 37809 828 6 , , , 37809 828 7 México México NNP 37809 828 8 , , , 37809 828 9 at at IN 37809 828 10 moderate moderate JJ 37809 828 11 to to IN 37809 828 12 high high JJ 37809 828 13 elevations elevation NNS 37809 828 14 ( ( -LRB- 37809 828 15 600 600 CD 37809 828 16 - - SYM 37809 828 17 2400 2400 CD 37809 828 18 meters meter NNS 37809 828 19 ) ) -RRB- 37809 828 20 . . . 37809 829 1 _ _ NNP 37809 829 2 Specimens Specimens NNP 37809 829 3 examined._--(48 examined._--(48 NNP 37809 829 4 ) ) -RRB- 37809 829 5 MÉXICO MÉXICO NNP 37809 829 6 , , , 37809 829 7 _ _ NNP 37809 829 8 Jalisco Jalisco NNP 37809 829 9 _ _ NNP 37809 829 10 : : : 37809 829 11 17.6 17.6 CD 37809 829 12 km km NNS 37809 829 13 . . . 37809 830 1 SW SW NNP 37809 830 2 Autlán Autlán NNP 37809 830 3 , , , 37809 830 4 606 606 CD 37809 830 5 m. m. NN 37809 830 6 , , , 37809 830 7 KU KU NNP 37809 830 8 102626 102626 CD 37809 830 9 , , , 37809 830 10 102631 102631 CD 37809 830 11 ; ; : 37809 830 12 6.4 6.4 CD 37809 830 13 km km NN 37809 830 14 . . . 37809 831 1 W W NNP 37809 831 2 Atenquique Atenquique NNP 37809 831 3 , , , 37809 831 4 2060 2060 CD 37809 831 5 m. m. NN 37809 831 6 , , , 37809 831 7 KU KU NNP 37809 831 8 102628 102628 CD 37809 831 9 - - SYM 37809 831 10 30 30 CD 37809 831 11 , , , 37809 831 12 102632 102632 CD 37809 831 13 ; ; : 37809 831 14 8 8 CD 37809 831 15 km km NN 37809 831 16 . . . 37809 832 1 W W NNP 37809 832 2 Atenquique Atenquique NNP 37809 832 3 , , , 37809 832 4 1970 1970 CD 37809 832 5 m. m. NN 37809 832 6 , , , 37809 832 7 LACM LACM NNP 37809 832 8 3210 3210 CD 37809 832 9 - - SYM 37809 832 10 12 12 CD 37809 832 11 ; ; : 37809 832 12 9.6 9.6 CD 37809 832 13 km km NN 37809 832 14 . . . 37809 833 1 W W NNP 37809 833 2 Atenquique Atenquique NNP 37809 833 3 , , , 37809 833 4 2360 2360 CD 37809 833 5 m. m. NN 37809 833 6 , , , 37809 833 7 LACM LACM NNP 37809 833 8 3200 3200 CD 37809 833 9 ( ( -LRB- 37809 833 10 holotype holotype NNP 37809 833 11 ) ) -RRB- 37809 833 12 , , , 37809 833 13 3201 3201 CD 37809 833 14 - - SYM 37809 833 15 09 09 CD 37809 833 16 ; ; : 37809 833 17 14.5 14.5 CD 37809 833 18 km km NNS 37809 833 19 . . . 37809 834 1 W W NNP 37809 834 2 Atenquique Atenquique NNP 37809 834 3 , , , 37809 834 4 2000 2000 CD 37809 834 5 m. m. NN 37809 834 6 , , , 37809 834 7 LACM LACM NNP 37809 834 8 25424 25424 CD 37809 834 9 - - SYM 37809 834 10 36 36 CD 37809 834 11 , , , 37809 834 12 25439 25439 CD 37809 834 13 - - SYM 37809 834 14 41 41 CD 37809 834 15 , , , 37809 834 16 25446 25446 CD 37809 834 17 ; ; : 37809 834 18 15 15 CD 37809 834 19 km km NN 37809 834 20 . . . 37809 835 1 W W NNP 37809 835 2 Atenquique Atenquique NNP 37809 835 3 , , , 37809 835 4 LACM LACM NNP 37809 835 5 37044 37044 CD 37809 835 6 - - SYM 37809 835 7 46 46 CD 37809 835 8 , , , 37809 835 9 37244 37244 CD 37809 835 10 - - SYM 37809 835 11 47 47 CD 37809 835 12 ; ; : 37809 835 13 16 16 CD 37809 835 14 km km NN 37809 835 15 . . . 37809 836 1 W W NNP 37809 836 2 Atenquique Atenquique NNP 37809 836 3 , , , 37809 836 4 2105 2105 CD 37809 836 5 m. m. NN 37809 836 6 , , , 37809 836 7 LACM LACM NNP 37809 836 8 25443 25443 CD 37809 836 9 - - SYM 37809 836 10 45 45 CD 37809 836 11 ; ; : 37809 836 12 17 17 CD 37809 836 13 km km NN 37809 836 14 . . . 37809 837 1 W W NNP 37809 837 2 Atenquique Atenquique NNP 37809 837 3 , , , 37809 837 4 2180 2180 CD 37809 837 5 m. m. NN 37809 837 6 , , , 37809 837 7 LACM LACM NNP 37809 837 8 25442 25442 CD 37809 837 9 . . . 37809 838 1 = = NFP 37809 838 2 Syrrhophus Syrrhophus NNP 37809 838 3 pallidus= pallidus= NN 37809 838 4 Duellman Duellman NNP 37809 838 5 , , , 37809 838 6 New new JJ 37809 838 7 combination combination NN 37809 838 8 _ _ NNP 37809 838 9 Syrrhophus Syrrhophus NNP 37809 838 10 modestus modestus NN 37809 838 11 _ _ NNP 37809 838 12 : : : 37809 838 13 Davis Davis NNP 37809 838 14 and and CC 37809 838 15 Dixon Dixon NNP 37809 838 16 , , , 37809 838 17 1957:146 1957:146 CD 37809 838 18 . . . 37809 839 1 _ _ NNP 37809 839 2 Syrrhophus Syrrhophus NNP 37809 839 3 modestus modestus NN 37809 839 4 pallidus pallidus NNP 37809 839 5 _ _ NNP 37809 839 6 Duellman Duellman NNP 37809 839 7 , , , 37809 839 8 1958:2 1958:2 CD 37809 839 9 - - SYM 37809 839 10 3 3 CD 37809 839 11 , , , 37809 839 12 5 5 CD 37809 839 13 - - SYM 37809 839 14 7 7 CD 37809 839 15 , , , 37809 839 16 14 14 CD 37809 839 17 , , , 37809 839 18 pl pl NNP 37809 839 19 . . . 37809 840 1 3 3 LS 37809 840 2 [ [ -LRB- 37809 840 3 Holotype.--UMMZ holotype.--ummz NN 37809 840 4 115452 115452 CD 37809 840 5 , , , 37809 840 6 from from IN 37809 840 7 San San NNP 37809 840 8 Blas Blas NNP 37809 840 9 , , , 37809 840 10 Nayarit Nayarit NNP 37809 840 11 , , , 37809 840 12 México México NNP 37809 840 13 , , , 37809 840 14 sea sea NN 37809 840 15 level level NN 37809 840 16 ; ; , 37809 840 17 collected collect VBN 37809 840 18 on on IN 37809 840 19 August August NNP 37809 840 20 13 13 CD 37809 840 21 , , , 37809 840 22 1956 1956 CD 37809 840 23 , , , 37809 840 24 by by IN 37809 840 25 William William NNP 37809 840 26 E. E. NNP 37809 840 27 and and CC 37809 840 28 Ann Ann NNP 37809 840 29 S. S. NNP 37809 840 30 Duellman Duellman NNP 37809 840 31 ] ] -RRB- 37809 840 32 . . . 37809 841 1 Zweifel Zweifel NNP 37809 841 2 , , , 37809 841 3 1960:86 1960:86 NNP 37809 841 4 - - SYM 37809 841 5 88 88 CD 37809 841 6 , , , 37809 841 7 91 91 CD 37809 841 8 , , , 37809 841 9 93 93 CD 37809 841 10 - - SYM 37809 841 11 94 94 CD 37809 841 12 , , , 37809 841 13 118 118 CD 37809 841 14 , , , 37809 841 15 120 120 CD 37809 841 16 - - SYM 37809 841 17 22 22 CD 37809 841 18 . . . 37809 842 1 Gorham Gorham NNP 37809 842 2 , , , 37809 842 3 1966:166 1966:166 CD 37809 842 4 . . . 37809 843 1 _ _ NNP 37809 843 2 Syrrhophis Syrrhophis NNP 37809 843 3 modestus modestus NN 37809 843 4 pallidus pallidus NN 37809 843 5 _ _ NNP 37809 843 6 : : : 37809 843 7 Campbell Campbell NNP 37809 843 8 and and CC 37809 843 9 Simmons Simmons NNP 37809 843 10 , , , 37809 843 11 1962:194 1962:194 CD 37809 843 12 . . . 37809 844 1 _ _ NNP 37809 844 2 Diagnosis._--Small Diagnosis._--Small NNP 37809 844 3 frogs frog NNS 37809 844 4 , , , 37809 844 5 males male VBZ 37809 844 6 17.9 17.9 CD 37809 844 7 - - SYM 37809 844 8 19.3 19.3 CD 37809 844 9 mm mm NN 37809 844 10 . . . 37809 845 1 snout snout NN 37809 845 2 - - HYPH 37809 845 3 vent vent NN 37809 845 4 length length NN 37809 845 5 ; ; : 37809 845 6 vocal vocal NN 37809 845 7 slits slit NNS 37809 845 8 in in IN 37809 845 9 males male NNS 37809 845 10 ; ; : 37809 845 11 finger finger NN 37809 845 12 tips tip NNS 37809 845 13 widely widely RB 37809 845 14 expanded expand VBN 37809 845 15 ; ; : 37809 845 16 first first RB 37809 845 17 finger finger NN 37809 845 18 shorter short JJR 37809 845 19 than than IN 37809 845 20 second second JJ 37809 845 21 ; ; : 37809 845 22 inner inner JJ 37809 845 23 metatarsal metatarsal NN 37809 845 24 tubercle tubercle NN 37809 845 25 about about IN 37809 845 26 three three CD 37809 845 27 times time NNS 37809 845 28 size size NN 37809 845 29 of of IN 37809 845 30 outer outer NN 37809 845 31 ; ; : 37809 845 32 skin skin NN 37809 845 33 of of IN 37809 845 34 dorsum dorsum NN 37809 845 35 shagreened shagreene VBN 37809 845 36 , , , 37809 845 37 that that DT 37809 845 38 of of IN 37809 845 39 venter venter NN 37809 845 40 smooth smooth JJ 37809 845 41 ; ; : 37809 845 42 tympanum tympanum NNP 37809 845 43 concealed conceal VBD 37809 845 44 , , , 37809 845 45 its -PRON- PRP$ 37809 845 46 diameter diameter NN 37809 845 47 27.0 27.0 CD 37809 845 48 - - HYPH 37809 845 49 35.6 35.6 CD 37809 845 50 per per IN 37809 845 51 cent cent NN 37809 845 52 of of IN 37809 845 53 eye eye NN 37809 845 54 in in IN 37809 845 55 males male NNS 37809 845 56 ; ; , 37809 845 57 ground ground NN 37809 845 58 color color NN 37809 845 59 cream cream NN 37809 845 60 vermiculated vermiculate VBN 37809 845 61 with with IN 37809 845 62 brown brown JJ 37809 845 63 , , , 37809 845 64 upper upper JJ 37809 845 65 arm arm NN 37809 845 66 and and CC 37809 845 67 thigh thigh NN 37809 845 68 lacking lacking NN 37809 845 69 , , , 37809 845 70 or or CC 37809 845 71 with with IN 37809 845 72 few few JJ 37809 845 73 , , , 37809 845 74 indistinct indistinct JJ 37809 845 75 , , , 37809 845 76 bands band NNS 37809 845 77 ; ; : 37809 845 78 interorbital interorbital NNP 37809 845 79 bar bar NN 37809 845 80 absent absent NN 37809 845 81 . . . 37809 846 1 _ _ NNP 37809 846 2 Remarks._--Considerable remarks._--considerable JJ 37809 846 3 debate debate NN 37809 846 4 has have VBZ 37809 846 5 been be VBN 37809 846 6 waged wage VBN 37809 846 7 relative relative JJ 37809 846 8 to to IN 37809 846 9 the the DT 37809 846 10 value value NN 37809 846 11 of of IN 37809 846 12 subspecies subspecie NNS 37809 846 13 and and CC 37809 846 14 to to IN 37809 846 15 the the DT 37809 846 16 reasons reason NNS 37809 846 17 for for IN 37809 846 18 recognizing recognize VBG 37809 846 19 distinct distinct JJ 37809 846 20 disjunct disjunct JJ 37809 846 21 populations population NNS 37809 846 22 as as IN 37809 846 23 species specie NNS 37809 846 24 versus versus IN 37809 846 25 subspecies subspecie NNS 37809 846 26 . . . 37809 847 1 Lacking lack VBG 37809 847 2 evidence evidence NN 37809 847 3 of of IN 37809 847 4 genetic genetic JJ 37809 847 5 exchange exchange NN 37809 847 6 , , , 37809 847 7 I -PRON- PRP 37809 847 8 prefer prefer VBP 37809 847 9 to to TO 37809 847 10 retain retain VB 37809 847 11 disjunct disjunct JJ 37809 847 12 populations population NNS 37809 847 13 that that WDT 37809 847 14 are be VBP 37809 847 15 distinctive distinctive JJ 37809 847 16 as as IN 37809 847 17 species specie NNS 37809 847 18 . . . 37809 848 1 All all DT 37809 848 2 known know VBN 37809 848 3 specimens specimen NNS 37809 848 4 of of IN 37809 848 5 _ _ NNP 37809 848 6 pallidus pallidus NNP 37809 848 7 _ _ NNP 37809 848 8 can can MD 37809 848 9 be be VB 37809 848 10 separated separate VBN 37809 848 11 from from IN 37809 848 12 those those DT 37809 848 13 of of IN 37809 848 14 _ _ NNP 37809 848 15 modestus modestus NN 37809 848 16 _ _ NNP 37809 848 17 by by IN 37809 848 18 color color NN 37809 848 19 pattern pattern NN 37809 848 20 . . . 37809 849 1 The the DT 37809 849 2 two two CD 37809 849 3 nominal nominal JJ 37809 849 4 species specie NNS 37809 849 5 exhibit exhibit NN 37809 849 6 overlap overlap VBP 37809 849 7 in in IN 37809 849 8 proportions proportion NNS 37809 849 9 but but CC 37809 849 10 the the DT 37809 849 11 same same JJ 37809 849 12 can can MD 37809 849 13 be be VB 37809 849 14 said say VBN 37809 849 15 about about IN 37809 849 16 nearly nearly RB 37809 849 17 every every DT 37809 849 18 species species NN 37809 849 19 of of IN 37809 849 20 _ _ NNP 37809 849 21 Syrrhophus Syrrhophus NNP 37809 849 22 _ _ NNP 37809 849 23 ; ; : 37809 849 24 therefore therefore RB 37809 849 25 , , , 37809 849 26 overlap overlap VBP 37809 849 27 in in IN 37809 849 28 proportions proportion NNS 37809 849 29 can can MD 37809 849 30 be be VB 37809 849 31 disregarded disregard VBN 37809 849 32 in in IN 37809 849 33 assessing assess VBG 37809 849 34 specific specific JJ 37809 849 35 versus versus IN 37809 849 36 subspecific subspecific NN 37809 849 37 rank rank NN 37809 849 38 . . . 37809 850 1 Until until IN 37809 850 2 contrary contrary JJ 37809 850 3 evidence evidence NN 37809 850 4 is be VBZ 37809 850 5 forthcoming forthcoming JJ 37809 850 6 , , , 37809 850 7 I -PRON- PRP 37809 850 8 consider consider VBP 37809 850 9 the the DT 37809 850 10 disjunct disjunct JJ 37809 850 11 populations population NNS 37809 850 12 heretofore heretofore RB 37809 850 13 held hold VBN 37809 850 14 to to TO 37809 850 15 be be VB 37809 850 16 subspecies subspecie NNS 37809 850 17 of of IN 37809 850 18 _ _ NNP 37809 850 19 modestus modestus NN 37809 850 20 _ _ NNP 37809 850 21 to to TO 37809 850 22 be be VB 37809 850 23 specifically specifically RB 37809 850 24 distinct distinct JJ 37809 850 25 . . . 37809 851 1 The the DT 37809 851 2 specimens specimen NNS 37809 851 3 of of IN 37809 851 4 the the DT 37809 851 5 disjunct disjunct JJ 37809 851 6 population population NN 37809 851 7 of of IN 37809 851 8 _ _ NNP 37809 851 9 pallidus pallidus NNP 37809 851 10 _ _ NNP 37809 851 11 on on IN 37809 851 12 the the DT 37809 851 13 Tres Tres NNP 37809 851 14 Marias Marias NNP 37809 851 15 do do VBP 37809 851 16 not not RB 37809 851 17 differ differ VB 37809 851 18 from from IN 37809 851 19 the the DT 37809 851 20 mainland mainland NN 37809 851 21 population population NN 37809 851 22 in in IN 37809 851 23 Nayarit Nayarit NNP 37809 851 24 . . . 37809 852 1 This this DT 37809 852 2 evidence evidence NN 37809 852 3 , , , 37809 852 4 though though IN 37809 852 5 perhaps perhaps RB 37809 852 6 secondary secondary JJ 37809 852 7 , , , 37809 852 8 supports support VBZ 37809 852 9 my -PRON- PRP$ 37809 852 10 contention contention NN 37809 852 11 that that IN 37809 852 12 two two CD 37809 852 13 species specie NNS 37809 852 14 should should MD 37809 852 15 be be VB 37809 852 16 recognized recognize VBN 37809 852 17 . . . 37809 853 1 _ _ NNP 37809 853 2 Etymology._--Latin Etymology._--Latin NNP 37809 853 3 , , , 37809 853 4 in in IN 37809 853 5 reference reference NN 37809 853 6 to to IN 37809 853 7 the the DT 37809 853 8 pale pale JJ 37809 853 9 ground ground NN 37809 853 10 color color NN 37809 853 11 in in IN 37809 853 12 comparison comparison NN 37809 853 13 with with IN 37809 853 14 that that DT 37809 853 15 of of IN 37809 853 16 _ _ NNP 37809 853 17 S. S. NNP 37809 854 1 modestus modestus NNP 37809 854 2 _ _ NNP 37809 854 3 . . . 37809 855 1 _ _ NNP 37809 855 2 Distribution._--Low Distribution._--Low NNP 37809 855 3 elevations elevation NNS 37809 855 4 in in IN 37809 855 5 coastal coastal JJ 37809 855 6 Nayarit Nayarit NNP 37809 855 7 and and CC 37809 855 8 on on IN 37809 855 9 Islas Islas NNP 37809 855 10 Tres Tres NNP 37809 855 11 Marias Marias NNP 37809 855 12 ( ( -LRB- 37809 855 13 Fig fig NN 37809 855 14 . . . 37809 856 1 20 20 CD 37809 856 2 ) ) -RRB- 37809 856 3 . . . 37809 857 1 _ _ NNP 37809 857 2 Specimens Specimens NNP 37809 857 3 examined._--(12 examined._--(12 NNP 37809 857 4 ) ) -RRB- 37809 857 5 MÉXICO MÉXICO NNP 37809 857 6 , , , 37809 857 7 _ _ NNP 37809 857 8 Nayarit Nayarit NNP 37809 857 9 _ _ NNP 37809 857 10 : : : 37809 857 11 18.8 18.8 CD 37809 857 12 mi mi NNP 37809 857 13 . . . 37809 858 1 NW NW NNP 37809 858 2 Ahuacatlán Ahuacatlán NNP 37809 858 3 , , , 37809 858 4 UIMNH UIMNH NNP 37809 858 5 7808 7808 CD 37809 858 6 ; ; : 37809 858 7 San San NNP 37809 858 8 Blas Blas NNP 37809 858 9 , , , 37809 858 10 UMMZ UMMZ NNP 37809 858 11 115452 115452 CD 37809 858 12 ( ( -LRB- 37809 858 13 holotype holotype NN 37809 858 14 ) ) -RRB- 37809 858 15 , , , 37809 858 16 115453 115453 CD 37809 858 17 - - SYM 37809 858 18 57 57 CD 37809 858 19 ; ; : 37809 858 20 17 17 CD 37809 858 21 km km NN 37809 858 22 . . . 37809 859 1 NE NE NNP 37809 859 2 San San NNP 37809 859 3 Blas Blas NNP 37809 859 4 , , , 37809 859 5 150 150 CD 37809 859 6 m. m. NN 37809 859 7 , , , 37809 859 8 MSU MSU NNP 37809 859 9 5085 5085 CD 37809 859 10 ; ; : 37809 859 11 12.8 12.8 CD 37809 859 12 km km NNS 37809 859 13 . . . 37809 860 1 E E NNP 37809 860 2 San San NNP 37809 860 3 Blas Blas NNP 37809 860 4 , , , 37809 860 5 UIMNH UIMNH NNP 37809 860 6 71979 71979 CD 37809 860 7 ; ; : 37809 860 8 31 31 CD 37809 860 9 km km NN 37809 860 10 . . . 37809 861 1 E E NNP 37809 861 2 San San NNP 37809 861 3 Blas Blas NNP 37809 861 4 , , , 37809 861 5 UIMNH UIMNH NNP 37809 861 6 71978 71978 CD 37809 861 7 ; ; : 37809 861 8 13.5 13.5 CD 37809 861 9 km km NNS 37809 861 10 . . . 37809 862 1 N N NNP 37809 862 2 Tepic Tepic NNP 37809 862 3 , , , 37809 862 4 UIMNH UIMNH NNP 37809 862 5 71980 71980 CD 37809 862 6 - - SYM 37809 862 7 81 81 CD 37809 862 8 . . . 37809 863 1 = = NFP 37809 863 2 Syrrhophus Syrrhophus NNP 37809 863 3 teretistes= teretistes= NN 37809 863 4 Duellman Duellman NNP 37809 863 5 _ _ NNP 37809 863 6 Syrrhophus Syrrhophus NNP 37809 863 7 teretistes teretiste VBZ 37809 863 8 _ _ NNP 37809 863 9 Duellman Duellman NNP 37809 863 10 , , , 37809 863 11 1958:2 1958:2 CD 37809 863 12 - - SYM 37809 863 13 3 3 CD 37809 863 14 , , , 37809 863 15 10 10 CD 37809 863 16 - - SYM 37809 863 17 14 14 CD 37809 863 18 , , , 37809 863 19 pl pl NNP 37809 863 20 . . . 37809 864 1 2 2 LS 37809 864 2 , , , 37809 864 3 fig fig NN 37809 864 4 . . . 37809 865 1 2 2 LS 37809 865 2 [ [ -LRB- 37809 865 3 Holotype.--UMMZ holotype.--ummz NN 37809 865 4 115451 115451 CD 37809 865 5 , , , 37809 865 6 from from IN 37809 865 7 4.8 4.8 CD 37809 865 8 km km NNS 37809 865 9 . . . 37809 866 1 NW NW NNP 37809 866 2 Tepic Tepic NNP 37809 866 3 , , , 37809 866 4 Nayarit Nayarit NNP 37809 866 5 , , , 37809 866 6 México México NNP 37809 866 7 , , , 37809 866 8 840 840 CD 37809 866 9 m. m. NN 37809 866 10 ; ; , 37809 866 11 collected collect VBN 37809 866 12 on on IN 37809 866 13 August August NNP 37809 866 14 12 12 CD 37809 866 15 , , , 37809 866 16 1956 1956 CD 37809 866 17 , , , 37809 866 18 by by IN 37809 866 19 William William NNP 37809 866 20 E. E. NNP 37809 866 21 Duellman Duellman NNP 37809 866 22 ] ] -RRB- 37809 866 23 . . . 37809 867 1 Gorham Gorham NNP 37809 867 2 , , , 37809 867 3 1966:167 1966:167 CD 37809 867 4 . . . 37809 868 1 _ _ NNP 37809 868 2 Diagnosis._--Medium Diagnosis._--Medium NNP 37809 868 3 - - HYPH 37809 868 4 sized sized JJ 37809 868 5 frogs frog NNS 37809 868 6 , , , 37809 868 7 males male VBZ 37809 868 8 19.2 19.2 CD 37809 868 9 - - HYPH 37809 868 10 23.2 23.2 CD 37809 868 11 mm mm NNP 37809 868 12 . . . 37809 869 1 snout snout NN 37809 869 2 - - HYPH 37809 869 3 vent vent NN 37809 869 4 length length NN 37809 869 5 , , , 37809 869 6 single single JJ 37809 869 7 known know VBN 37809 869 8 female female JJ 37809 869 9 24.8 24.8 CD 37809 869 10 mm mm NNS 37809 869 11 . . . 37809 870 1 snout snout NN 37809 870 2 - - HYPH 37809 870 3 vent vent NN 37809 870 4 ; ; : 37809 870 5 vocal vocal NN 37809 870 6 slits slit NNS 37809 870 7 in in IN 37809 870 8 males male NNS 37809 870 9 ; ; : 37809 870 10 finger finger NN 37809 870 11 tips tip NNS 37809 870 12 widely widely RB 37809 870 13 expanded expand VBN 37809 870 14 ; ; : 37809 870 15 first first RB 37809 870 16 finger finger NN 37809 870 17 shorter short JJR 37809 870 18 than than IN 37809 870 19 second second JJ 37809 870 20 ; ; : 37809 870 21 inner inner JJ 37809 870 22 metatarsal metatarsal NN 37809 870 23 tubercle tubercle NN 37809 870 24 about about IN 37809 870 25 three three CD 37809 870 26 times time NNS 37809 870 27 size size NN 37809 870 28 of of IN 37809 870 29 outer outer NN 37809 870 30 ; ; : 37809 870 31 skin skin NN 37809 870 32 of of IN 37809 870 33 dorsum dorsum NN 37809 870 34 shagreened shagreene VBN 37809 870 35 , , , 37809 870 36 that that DT 37809 870 37 of of IN 37809 870 38 venter venter NN 37809 870 39 smooth smooth JJ 37809 870 40 ; ; : 37809 870 41 tympanum tympanum FW 37809 870 42 partially partially RB 37809 870 43 concealed conceal VBD 37809 870 44 , , , 37809 870 45 its -PRON- PRP$ 37809 870 46 diameter diameter NN 37809 870 47 28.6 28.6 CD 37809 870 48 - - HYPH 37809 870 49 43.8 43.8 CD 37809 870 50 per per IN 37809 870 51 cent cent NN 37809 870 52 of of IN 37809 870 53 eye eye NN 37809 870 54 in in IN 37809 870 55 males male NNS 37809 870 56 ; ; , 37809 870 57 ground ground NN 37809 870 58 color color NN 37809 870 59 brown brown NNP 37809 870 60 vermiculated vermiculate VBD 37809 870 61 with with IN 37809 870 62 dark dark JJ 37809 870 63 brown brown NNP 37809 870 64 to to IN 37809 870 65 nearly nearly RB 37809 870 66 black black JJ 37809 870 67 ; ; : 37809 870 68 upper upper JJ 37809 870 69 arm arm NN 37809 870 70 and and CC 37809 870 71 thigh thigh NN 37809 870 72 banded band VBD 37809 870 73 ; ; : 37809 870 74 interorbital interorbital NNP 37809 870 75 light light NNP 37809 870 76 bar bar NN 37809 870 77 absent absent NN 37809 870 78 . . . 37809 871 1 _ _ NNP 37809 871 2 Remarks._--_S. remarks._--_s. VBP 37809 872 1 teretistes teretistes NNPS 37809 872 2 _ _ NNP 37809 872 3 appears appear VBZ 37809 872 4 to to TO 37809 872 5 be be VB 37809 872 6 most most RBS 37809 872 7 closely closely RB 37809 872 8 related related JJ 37809 872 9 to to IN 37809 872 10 _ _ NNP 37809 872 11 S. S. NNP 37809 873 1 pallidus pallidus NNP 37809 873 2 _ _ NNP 37809 873 3 ; ; : 37809 873 4 I -PRON- PRP 37809 873 5 consider consider VBP 37809 873 6 it -PRON- PRP 37809 873 7 to to TO 37809 873 8 be be VB 37809 873 9 an an DT 37809 873 10 upland upland JJ 37809 873 11 derivative derivative NN 37809 873 12 of of IN 37809 873 13 _ _ NNP 37809 873 14 pallidus pallidus NNP 37809 873 15 _ _ NNP 37809 873 16 . . . 37809 874 1 Morphologically morphologically RB 37809 874 2 , , , 37809 874 3 the the DT 37809 874 4 differences difference NNS 37809 874 5 between between IN 37809 874 6 the the DT 37809 874 7 two two CD 37809 874 8 are be VBP 37809 874 9 few few JJ 37809 874 10 , , , 37809 874 11 but but CC 37809 874 12 lacking lack VBG 37809 874 13 evidence evidence NN 37809 874 14 of of IN 37809 874 15 genetic genetic JJ 37809 874 16 exchange exchange NN 37809 874 17 they -PRON- PRP 37809 874 18 are be VBP 37809 874 19 retained retain VBN 37809 874 20 as as IN 37809 874 21 species specie NNS 37809 874 22 . . . 37809 875 1 _ _ NNP 37809 875 2 Etymology._--Greek Etymology._--Greek NNP 37809 875 3 , , , 37809 875 4 in in IN 37809 875 5 reference reference NN 37809 875 6 to to IN 37809 875 7 the the DT 37809 875 8 whistle whistle NN 37809 875 9 - - HYPH 37809 875 10 like like JJ 37809 875 11 nature nature NN 37809 875 12 of of IN 37809 875 13 the the DT 37809 875 14 call call NN 37809 875 15 . . . 37809 876 1 _ _ NNP 37809 876 2 Distribution._--Moderate distribution._--moderate JJ 37809 876 3 elevations elevation NNS 37809 876 4 ( ( -LRB- 37809 876 5 840 840 CD 37809 876 6 - - SYM 37809 876 7 1200 1200 CD 37809 876 8 meters meter NNS 37809 876 9 ) ) -RRB- 37809 876 10 in in IN 37809 876 11 the the DT 37809 876 12 Sierra Sierra NNP 37809 876 13 Occidental Occidental NNP 37809 876 14 of of IN 37809 876 15 Nayarit Nayarit NNP 37809 876 16 , , , 37809 876 17 Sinaloa Sinaloa NNP 37809 876 18 , , , 37809 876 19 and and CC 37809 876 20 Durango Durango NNP 37809 876 21 , , , 37809 876 22 México México NNP 37809 876 23 ( ( -LRB- 37809 876 24 Fig Fig NNP 37809 876 25 . . . 37809 877 1 20 20 CD 37809 877 2 ) ) -RRB- 37809 877 3 . . . 37809 878 1 _ _ NNP 37809 878 2 Specimens Specimens NNP 37809 878 3 examined._--(13 examined._--(13 NNP 37809 878 4 ) ) -RRB- 37809 878 5 MÉXICO MÉXICO NNP 37809 878 6 , , , 37809 878 7 _ _ NNP 37809 878 8 Nayarit Nayarit NNP 37809 878 9 _ _ NNP 37809 878 10 : : : 37809 878 11 4.8 4.8 CD 37809 878 12 km km NNS 37809 878 13 . . . 37809 879 1 NW NW NNP 37809 879 2 Tepic Tepic NNP 37809 879 3 , , , 37809 879 4 840 840 CD 37809 879 5 m. m. NN 37809 879 6 , , , 37809 879 7 UMMZ UMMZ NNP 37809 879 8 115451 115451 CD 37809 879 9 ( ( -LRB- 37809 879 10 holotype holotype NN 37809 879 11 ) ) -RRB- 37809 879 12 . . . 37809 880 1 _ _ NNP 37809 880 2 Sinaloa Sinaloa NNP 37809 880 3 _ _ NNP 37809 880 4 : : : 37809 880 5 Santa Santa NNP 37809 880 6 Lucía Lucía NNP 37809 880 7 , , , 37809 880 8 1090 1090 CD 37809 880 9 m. m. NN 37809 880 10 , , , 37809 880 11 KU KU NNP 37809 880 12 75263 75263 CD 37809 880 13 - - SYM 37809 880 14 72 72 CD 37809 880 15 ; ; : 37809 880 16 1 1 CD 37809 880 17 km km NN 37809 880 18 . . . 37809 881 1 NE NE NNP 37809 881 2 Santa Santa NNP 37809 881 3 Lucía Lucía NNP 37809 881 4 , , , 37809 881 5 1156 1156 CD 37809 881 6 m. m. NN 37809 881 7 , , , 37809 881 8 KU KU NNP 37809 881 9 78257 78257 CD 37809 881 10 ; ; : 37809 881 11 2.2 2.2 CD 37809 881 12 km km NN 37809 881 13 . . . 37809 882 1 NE NE NNP 37809 882 2 Santa Santa NNP 37809 882 3 Lucía Lucía NNP 37809 882 4 , , , 37809 882 5 1156 1156 CD 37809 882 6 m. m. NN 37809 882 7 , , , 37809 882 8 KU KU NNP 37809 882 9 78258 78258 CD 37809 882 10 . . . 37809 883 1 DISCUSSION DISCUSSION NNP 37809 883 2 There there EX 37809 883 3 are be VBP 37809 883 4 relatively relatively RB 37809 883 5 few few JJ 37809 883 6 clear clear RB 37809 883 7 - - HYPH 37809 883 8 cut cut VBN 37809 883 9 morphological morphological JJ 37809 883 10 differences difference NNS 37809 883 11 among among IN 37809 883 12 the the DT 37809 883 13 fourteen fourteen CD 37809 883 14 species specie NNS 37809 883 15 now now RB 37809 883 16 assigned assign VBN 37809 883 17 to to IN 37809 883 18 _ _ NNP 37809 883 19 Syrrhophus Syrrhophus NNP 37809 883 20 _ _ NNP 37809 883 21 . . . 37809 884 1 The the DT 37809 884 2 majority majority NN 37809 884 3 of of IN 37809 884 4 the the DT 37809 884 5 species specie NNS 37809 884 6 are be VBP 37809 884 7 allopatric allopatric JJ 37809 884 8 and and CC 37809 884 9 differ differ VBP 37809 884 10 primarily primarily RB 37809 884 11 in in IN 37809 884 12 color color NN 37809 884 13 patterns pattern NNS 37809 884 14 . . . 37809 885 1 Sympatric sympatric JJ 37809 885 2 occurrence occurrence NN 37809 885 3 serves serve VBZ 37809 885 4 as as IN 37809 885 5 an an DT 37809 885 6 indicator indicator NN 37809 885 7 of of IN 37809 885 8 specific specific JJ 37809 885 9 distinctness distinctness NN 37809 885 10 and and CC 37809 885 11 is be VBZ 37809 885 12 one one CD 37809 885 13 of of IN 37809 885 14 the the DT 37809 885 15 more more RBR 37809 885 16 practical practical JJ 37809 885 17 tests test NNS 37809 885 18 of of IN 37809 885 19 species specie NNS 37809 885 20 validity validity NN 37809 885 21 when when WRB 37809 885 22 cross cross NN 37809 885 23 - - JJ 37809 885 24 breeding breeding JJ 37809 885 25 experiments experiment NNS 37809 885 26 are be VBP 37809 885 27 not not RB 37809 885 28 possible possible JJ 37809 885 29 . . . 37809 886 1 Two two CD 37809 886 2 cases case NNS 37809 886 3 of of IN 37809 886 4 sympatric sympatric JJ 37809 886 5 occurrence occurrence NN 37809 886 6 are be VBP 37809 886 7 known know VBN 37809 886 8 for for IN 37809 886 9 the the DT 37809 886 10 species specie NNS 37809 886 11 of of IN 37809 886 12 _ _ NNP 37809 886 13 Syrrhophus Syrrhophus NNP 37809 886 14 _ _ NNP 37809 886 15 in in IN 37809 886 16 western western JJ 37809 886 17 México México NNP 37809 886 18 : : : 37809 886 19 _ _ NNP 37809 886 20 modestus modestus NN 37809 886 21 _ _ NNP 37809 886 22 and and CC 37809 886 23 _ _ NNP 37809 886 24 nivocolimae nivocolimae NN 37809 886 25 _ _ NNP 37809 886 26 are be VBP 37809 886 27 sympatric sympatric JJ 37809 886 28 in in IN 37809 886 29 southern southern JJ 37809 886 30 Jalisco Jalisco NNP 37809 886 31 and and CC 37809 886 32 _ _ NNP 37809 886 33 pipilans pipilans NNPS 37809 886 34 nebulosus nebulosus NNP 37809 886 35 _ _ NNP 37809 886 36 and and CC 37809 886 37 _ _ NNP 37809 886 38 rubrimaculatus rubrimaculatus NN 37809 886 39 _ _ NNP 37809 886 40 are be VBP 37809 886 41 sympatric sympatric JJ 37809 886 42 in in IN 37809 886 43 southeastern southeastern JJ 37809 886 44 Chiapas chiapa NNS 37809 886 45 . . . 37809 887 1 In in IN 37809 887 2 eastern eastern JJ 37809 887 3 México México NNP 37809 887 4 , , , 37809 887 5 _ _ NNP 37809 887 6 longipes longipe VBZ 37809 887 7 _ _ NNP 37809 887 8 and and CC 37809 887 9 _ _ NNP 37809 887 10 verrucipes verrucipe NNS 37809 887 11 _ _ NNP 37809 887 12 are be VBP 37809 887 13 sympatric sympatric JJ 37809 887 14 in in IN 37809 887 15 southern southern JJ 37809 887 16 Hidalgo Hidalgo NNP 37809 887 17 , , , 37809 887 18 and and CC 37809 887 19 _ _ NNP 37809 887 20 longipes longipe VBZ 37809 887 21 _ _ NNP 37809 887 22 is be VBZ 37809 887 23 sympatric sympatric NN 37809 887 24 with with IN 37809 887 25 _ _ NNP 37809 887 26 cystignathoides cystignathoides NNP 37809 887 27 _ _ NNP 37809 887 28 , , , 37809 887 29 _ _ NNP 37809 887 30 dennisi dennisi NNP 37809 887 31 _ _ NNP 37809 887 32 , , , 37809 887 33 and and CC 37809 887 34 _ _ NNP 37809 887 35 guttilatus guttilatus NN 37809 887 36 _ _ NNP 37809 887 37 in in IN 37809 887 38 southern southern JJ 37809 887 39 Tamaulipas Tamaulipas NNP 37809 887 40 . . . 37809 888 1 _ _ NNP 37809 888 2 Syrrhophus Syrrhophus NNP 37809 888 3 cystignathoides cystignathoide VBZ 37809 888 4 _ _ NNP 37809 888 5 and and CC 37809 888 6 _ _ NNP 37809 888 7 leprus leprus NNP 37809 888 8 _ _ NNP 37809 888 9 are be VBP 37809 888 10 apparently apparently RB 37809 888 11 sympatric sympatric JJ 37809 888 12 in in IN 37809 888 13 central central JJ 37809 888 14 Veracruz Veracruz NNP 37809 888 15 . . . 37809 889 1 Subspecific Subspecific NNP 37809 889 2 assignments assignment NNS 37809 889 3 have have VBP 37809 889 4 been be VBN 37809 889 5 made make VBN 37809 889 6 only only RB 37809 889 7 when when WRB 37809 889 8 there there EX 37809 889 9 is be VBZ 37809 889 10 evidence evidence NN 37809 889 11 of of IN 37809 889 12 intergradation intergradation NN 37809 889 13 . . . 37809 890 1 The the DT 37809 890 2 sympatric sympatric JJ 37809 890 3 occurrence occurrence NN 37809 890 4 of of IN 37809 890 5 morphologically morphologically RB 37809 890 6 similar similar JJ 37809 890 7 species specie NNS 37809 890 8 in in IN 37809 890 9 this this DT 37809 890 10 genus genus NN 37809 890 11 has have VBZ 37809 890 12 led lead VBN 37809 890 13 me -PRON- PRP 37809 890 14 to to TO 37809 890 15 adopt adopt VB 37809 890 16 a a DT 37809 890 17 conservative conservative JJ 37809 890 18 approach approach NN 37809 890 19 to to IN 37809 890 20 the the DT 37809 890 21 degree degree NN 37809 890 22 of of IN 37809 890 23 difference difference NN 37809 890 24 philosophy philosophy NN 37809 890 25 . . . 37809 891 1 I -PRON- PRP 37809 891 2 have have VBP 37809 891 3 therefore therefore RB 37809 891 4 recognized recognize VBN 37809 891 5 all all DT 37809 891 6 morphologically morphologically RB 37809 891 7 distinct distinct JJ 37809 891 8 allopatric allopatric NN 37809 891 9 populations population NNS 37809 891 10 as as IN 37809 891 11 species specie NNS 37809 891 12 . . . 37809 892 1 [ [ -LRB- 37809 892 2 Illustration illustration NN 37809 892 3 : : : 37809 892 4 FIG FIG NNP 37809 892 5 . . . 37809 893 1 21 21 LS 37809 893 2 : : : 37809 893 3 Generic generic JJ 37809 893 4 distributions distribution NNS 37809 893 5 of of IN 37809 893 6 _ _ NNP 37809 893 7 Syrrhophus Syrrhophus NNP 37809 893 8 _ _ NNP 37809 893 9 ( ( -LRB- 37809 893 10 stipple stipple NN 37809 893 11 ) ) -RRB- 37809 893 12 and and CC 37809 893 13 _ _ NNP 37809 893 14 Tomodactylus Tomodactylus NNP 37809 893 15 _ _ NNP 37809 893 16 ( ( -LRB- 37809 893 17 hatching hatching NN 37809 893 18 ) ) -RRB- 37809 893 19 . . . 37809 894 1 Black black JJ 37809 894 2 areas area NNS 37809 894 3 are be VBP 37809 894 4 zones zone NNS 37809 894 5 of of IN 37809 894 6 intergeneric intergeneric NNP 37809 894 7 sympatry sympatry NNP 37809 894 8 . . . 37809 894 9 ] ] -RRB- 37809 895 1 _ _ NNP 37809 895 2 Syrrhophus Syrrhophus NNP 37809 895 3 _ _ NNP 37809 895 4 is be VBZ 37809 895 5 closely closely RB 37809 895 6 allied ally VBN 37809 895 7 to to IN 37809 895 8 another another DT 37809 895 9 Mexican mexican JJ 37809 895 10 leptodactylid leptodactylid JJ 37809 895 11 genus genus NN 37809 895 12 , , , 37809 895 13 _ _ NNP 37809 895 14 Tomodactylus Tomodactylus NNP 37809 895 15 _ _ NNP 37809 895 16 , , , 37809 895 17 which which WDT 37809 895 18 was be VBD 37809 895 19 revised revise VBN 37809 895 20 by by IN 37809 895 21 Dixon Dixon NNP 37809 895 22 ( ( -LRB- 37809 895 23 1957 1957 CD 37809 895 24 ) ) -RRB- 37809 895 25 , , , 37809 895 26 who who WP 37809 895 27 along along IN 37809 895 28 with with IN 37809 895 29 numerous numerous JJ 37809 895 30 other other JJ 37809 895 31 authors author NNS 37809 895 32 noted note VBD 37809 895 33 the the DT 37809 895 34 close close JJ 37809 895 35 relationship relationship NN 37809 895 36 between between IN 37809 895 37 the the DT 37809 895 38 two two CD 37809 895 39 genera genera NN 37809 895 40 . . . 37809 896 1 There there EX 37809 896 2 is be VBZ 37809 896 3 an an DT 37809 896 4 almost almost RB 37809 896 5 complete complete JJ 37809 896 6 lack lack NN 37809 896 7 of of IN 37809 896 8 sympatry sympatry NN 37809 896 9 between between IN 37809 896 10 the the DT 37809 896 11 two two CD 37809 896 12 genera genera NN 37809 896 13 ; ; : 37809 896 14 in in IN 37809 896 15 very very RB 37809 896 16 few few JJ 37809 896 17 places place NNS 37809 896 18 in in IN 37809 896 19 México México NNP 37809 896 20 do do VBP 37809 896 21 they -PRON- PRP 37809 896 22 coexist coexist VB 37809 896 23 ( ( -LRB- 37809 896 24 Fig fig NN 37809 896 25 . . . 37809 897 1 21 21 CD 37809 897 2 ) ) -RRB- 37809 897 3 . . . 37809 898 1 _ _ NNP 37809 898 2 Tomodactylus Tomodactylus NNP 37809 898 3 _ _ NNP 37809 898 4 has have VBZ 37809 898 5 its -PRON- PRP$ 37809 898 6 greatest great JJS 37809 898 7 diversity diversity NN 37809 898 8 in in IN 37809 898 9 the the DT 37809 898 10 Cordillera Cordillera NNP 37809 898 11 Volcánica Volcánica NNP 37809 898 12 and and CC 37809 898 13 Sierra Sierra NNP 37809 898 14 Madre Madre NNP 37809 898 15 del del NNP 37809 898 16 Sur Sur NNP 37809 898 17 , , , 37809 898 18 whereas whereas IN 37809 898 19 _ _ NNP 37809 898 20 Syrrhophus Syrrhophus NNP 37809 898 21 _ _ NNP 37809 898 22 reaches reach VBZ 37809 898 23 its -PRON- PRP$ 37809 898 24 greatest great JJS 37809 898 25 diversity diversity NN 37809 898 26 in in IN 37809 898 27 the the DT 37809 898 28 Sierra Sierra NNP 37809 898 29 Madre Madre NNP 37809 898 30 Oriental Oriental NNP 37809 898 31 and and CC 37809 898 32 eastern eastern JJ 37809 898 33 foothills foothill NNS 37809 898 34 . . . 37809 899 1 The the DT 37809 899 2 species specie NNS 37809 899 3 of of IN 37809 899 4 both both DT 37809 899 5 genera genera NN 37809 899 6 are be VBP 37809 899 7 about about IN 37809 899 8 the the DT 37809 899 9 same same JJ 37809 899 10 size size NN 37809 899 11 and and CC 37809 899 12 presumably presumably RB 37809 899 13 have have VBP 37809 899 14 similar similar JJ 37809 899 15 requirements requirement NNS 37809 899 16 insofar insofar RB 37809 899 17 as as IN 37809 899 18 food food NN 37809 899 19 , , , 37809 899 20 breeding breed VBG 37809 899 21 sites site NNS 37809 899 22 , , , 37809 899 23 and and CC 37809 899 24 habitat habitat NN 37809 899 25 selection selection NN 37809 899 26 . . . 37809 900 1 Four four CD 37809 900 2 cases case NNS 37809 900 3 of of IN 37809 900 4 intergeneric intergeneric JJ 37809 900 5 sympatry sympatry NN 37809 900 6 are be VBP 37809 900 7 known know VBN 37809 900 8 for for IN 37809 900 9 the the DT 37809 900 10 two two CD 37809 900 11 genera genera NN 37809 900 12 : : : 37809 900 13 1 1 LS 37809 900 14 ) ) -RRB- 37809 900 15 the the DT 37809 900 16 Chilpancingo Chilpancingo NNP 37809 900 17 region region NN 37809 900 18 of of IN 37809 900 19 Guerrero Guerrero NNP 37809 900 20 , , , 37809 900 21 2 2 LS 37809 900 22 ) ) -RRB- 37809 900 23 the the DT 37809 900 24 lowlands lowland NNS 37809 900 25 of of IN 37809 900 26 Colima Colima NNP 37809 900 27 and and CC 37809 900 28 the the DT 37809 900 29 mountains mountain NNS 37809 900 30 just just RB 37809 900 31 inland inland RB 37809 900 32 in in IN 37809 900 33 Jalisco Jalisco NNP 37809 900 34 , , , 37809 900 35 3 3 LS 37809 900 36 ) ) -RRB- 37809 900 37 the the DT 37809 900 38 lowlands lowland NNS 37809 900 39 of of IN 37809 900 40 central central JJ 37809 900 41 Nayarit Nayarit NNP 37809 900 42 , , , 37809 900 43 and and CC 37809 900 44 4 4 LS 37809 900 45 ) ) -RRB- 37809 900 46 the the DT 37809 900 47 Sierra Sierra NNP 37809 900 48 Madre Madre NNP 37809 900 49 Occidental Occidental NNP 37809 900 50 on on IN 37809 900 51 the the DT 37809 900 52 Durango Durango NNP 37809 900 53 - - HYPH 37809 900 54 Sinaloan Sinaloan NNP 37809 900 55 border border NN 37809 900 56 . . . 37809 901 1 The the DT 37809 901 2 apparent apparent JJ 37809 901 3 sympatry sympatry NN 37809 901 4 in in IN 37809 901 5 the the DT 37809 901 6 Chilpancingo Chilpancingo NNP 37809 901 7 region region NN 37809 901 8 involves involve VBZ 37809 901 9 four four CD 37809 901 10 species specie NNS 37809 901 11 : : : 37809 901 12 _ _ NNP 37809 901 13 S. S. NNP 37809 902 1 pipilans pipilans NNPS 37809 902 2 _ _ NNP 37809 902 3 , , , 37809 902 4 _ _ NNP 37809 902 5 T. T. NNP 37809 903 1 albolabris albolabris NNP 37809 903 2 _ _ NNP 37809 903 3 , , , 37809 903 4 _ _ NNP 37809 903 5 T. T. NNP 37809 904 1 dilatus dilatus NNP 37809 904 2 _ _ NNP 37809 904 3 , , , 37809 904 4 and and CC 37809 904 5 _ _ NNP 37809 904 6 T. T. NNP 37809 905 1 nitidus nitidus NNP 37809 905 2 _ _ NNP 37809 905 3 . . . 37809 906 1 Of of IN 37809 906 2 the the DT 37809 906 3 four four CD 37809 906 4 , , , 37809 906 5 _ _ NNP 37809 906 6 T. T. NNP 37809 907 1 dilatus dilatus NNP 37809 907 2 _ _ NNP 37809 907 3 appears appear VBZ 37809 907 4 to to TO 37809 907 5 be be VB 37809 907 6 completely completely RB 37809 907 7 allopatric allopatric JJ 37809 907 8 in in IN 37809 907 9 that that IN 37809 907 10 it -PRON- PRP 37809 907 11 occurs occur VBZ 37809 907 12 at at IN 37809 907 13 higher high JJR 37809 907 14 altitudes altitude NNS 37809 907 15 ( ( -LRB- 37809 907 16 above above IN 37809 907 17 2000 2000 CD 37809 907 18 meters meter NNS 37809 907 19 ) ) -RRB- 37809 907 20 , , , 37809 907 21 whereas whereas IN 37809 907 22 the the DT 37809 907 23 other other JJ 37809 907 24 three three CD 37809 907 25 occur occur VBP 37809 907 26 below below IN 37809 907 27 1800 1800 CD 37809 907 28 meters meter NNS 37809 907 29 in in IN 37809 907 30 the the DT 37809 907 31 region region NN 37809 907 32 ( ( -LRB- 37809 907 33 Davis Davis NNP 37809 907 34 and and CC 37809 907 35 Dixon Dixon NNP 37809 907 36 , , , 37809 907 37 1965 1965 CD 37809 907 38 ) ) -RRB- 37809 907 39 . . . 37809 908 1 In in IN 37809 908 2 the the DT 37809 908 3 Colima Colima NNP 37809 908 4 - - HYPH 37809 908 5 Jalisco Jalisco NNP 37809 908 6 region region NN 37809 908 7 , , , 37809 908 8 _ _ NNP 37809 908 9 Tomodactylus Tomodactylus NNP 37809 908 10 _ _ NNP 37809 908 11 tends tend VBZ 37809 908 12 to to TO 37809 908 13 occur occur VB 37809 908 14 higher high JJR 37809 908 15 ( ( -LRB- 37809 908 16 Dixon Dixon NNP 37809 908 17 and and CC 37809 908 18 Webb Webb NNP 37809 908 19 , , , 37809 908 20 1966 1966 CD 37809 908 21 ) ) -RRB- 37809 908 22 than than IN 37809 908 23 some some DT 37809 908 24 of of IN 37809 908 25 the the DT 37809 908 26 _ _ NNP 37809 908 27 Syrrhophus Syrrhophus NNP 37809 908 28 _ _ NNP 37809 908 29 , , , 37809 908 30 but but CC 37809 908 31 one one CD 37809 908 32 subspecies subspecie NNS 37809 908 33 of of IN 37809 908 34 _ _ NNP 37809 908 35 Tomodactylus Tomodactylus NNP 37809 908 36 nitidus nitidus NN 37809 908 37 _ _ NNP 37809 908 38 is be VBZ 37809 908 39 a a DT 37809 908 40 lowland lowland NN 37809 908 41 frog frog NN 37809 908 42 , , , 37809 908 43 occurring occur VBG 37809 908 44 sympatrically sympatrically RB 37809 908 45 with with IN 37809 908 46 the the DT 37809 908 47 lowland lowland NN 37809 908 48 _ _ NNP 37809 908 49 Syrrhophus Syrrhophus NNP 37809 908 50 modestus modestus NN 37809 908 51 _ _ NNP 37809 908 52 . . . 37809 909 1 A a DT 37809 909 2 similar similar JJ 37809 909 3 situation situation NN 37809 909 4 is be VBZ 37809 909 5 observed observe VBN 37809 909 6 in in IN 37809 909 7 Nayarit Nayarit NNP 37809 909 8 ; ; : 37809 909 9 the the DT 37809 909 10 lowland lowland NN 37809 909 11 _ _ NNP 37809 909 12 Tomodactylus Tomodactylus NNP 37809 909 13 _ _ NNP 37809 909 14 occurs occur VBZ 37809 909 15 sympatrically sympatrically RB 37809 909 16 with with IN 37809 909 17 the the DT 37809 909 18 small small JJ 37809 909 19 _ _ NNP 37809 909 20 Syrrhophus Syrrhophus NNP 37809 909 21 pallidus pallidus NNP 37809 909 22 _ _ NNP 37809 909 23 . . . 37809 910 1 In in IN 37809 910 2 both both DT 37809 910 3 cases case NNS 37809 910 4 the the DT 37809 910 5 _ _ NNP 37809 910 6 Syrrhophus Syrrhophus NNP 37809 910 7 _ _ NNP 37809 910 8 is be VBZ 37809 910 9 smaller small JJR 37809 910 10 than than IN 37809 910 11 the the DT 37809 910 12 _ _ NNP 37809 910 13 Tomodactylus Tomodactylus NNP 37809 910 14 _ _ NNP 37809 910 15 . . . 37809 911 1 [ [ -LRB- 37809 911 2 Illustration illustration NN 37809 911 3 : : : 37809 911 4 FIG FIG NNP 37809 911 5 . . . 37809 912 1 22 22 CD 37809 912 2 : : : 37809 912 3 Altitudinal altitudinal JJ 37809 912 4 distributions distribution NNS 37809 912 5 of of IN 37809 912 6 _ _ NNP 37809 912 7 Syrrhophus Syrrhophus NNP 37809 912 8 _ _ NNP 37809 912 9 and and CC 37809 912 10 _ _ NNP 37809 912 11 Tomodactylus Tomodactylus NNP 37809 912 12 _ _ NNP 37809 912 13 . . . 37809 913 1 Widths width NNS 37809 913 2 of of IN 37809 913 3 the the DT 37809 913 4 columns column NNS 37809 913 5 are be VBP 37809 913 6 proportional proportional JJ 37809 913 7 to to IN 37809 913 8 the the DT 37809 913 9 numbers number NNS 37809 913 10 of of IN 37809 913 11 species specie NNS 37809 913 12 at at IN 37809 913 13 a a DT 37809 913 14 given give VBN 37809 913 15 altitude altitude NN 37809 913 16 ; ; : 37809 913 17 narrowest narrow JJS 37809 913 18 width width NN 37809 913 19 equals equal VBZ 37809 913 20 one one CD 37809 913 21 species species NN 37809 913 22 . . . 37809 913 23 ] ] -RRB- 37809 914 1 Frogs frog NNS 37809 914 2 of of IN 37809 914 3 the the DT 37809 914 4 genus genus NN 37809 914 5 _ _ NNP 37809 914 6 Syrrhophus Syrrhophus NNP 37809 914 7 _ _ NNP 37809 914 8 tend tend VB 37809 914 9 to to TO 37809 914 10 occur occur VB 37809 914 11 at at IN 37809 914 12 lower low JJR 37809 914 13 elevations elevation NNS 37809 914 14 than than IN 37809 914 15 do do VBP 37809 914 16 their -PRON- PRP$ 37809 914 17 close close JJ 37809 914 18 relatives relative NNS 37809 914 19 of of IN 37809 914 20 the the DT 37809 914 21 genus genus NNP 37809 914 22 _ _ NNP 37809 914 23 Tomodactylus Tomodactylus NNP 37809 914 24 _ _ NNP 37809 914 25 ( ( -LRB- 37809 914 26 Fig Fig NNP 37809 914 27 . . . 37809 915 1 22 22 CD 37809 915 2 ) ) -RRB- 37809 915 3 . . . 37809 916 1 This this DT 37809 916 2 generalization generalization NN 37809 916 3 is be VBZ 37809 916 4 complicated complicate VBN 37809 916 5 by by IN 37809 916 6 the the DT 37809 916 7 occurrence occurrence NN 37809 916 8 in in IN 37809 916 9 the the DT 37809 916 10 Sierra Sierra NNP 37809 916 11 Madre Madre NNP 37809 916 12 Oriental Oriental NNP 37809 916 13 in in IN 37809 916 14 relatively relatively RB 37809 916 15 high high JJ 37809 916 16 altitude altitude NN 37809 916 17 _ _ NNP 37809 916 18 Syrrhophus Syrrhophus NNP 37809 916 19 _ _ NNP 37809 916 20 ( ( -LRB- 37809 916 21 up up IN 37809 916 22 to to TO 37809 916 23 2000 2000 CD 37809 916 24 m. m. NN 37809 916 25 ) ) -RRB- 37809 916 26 and and CC 37809 916 27 the the DT 37809 916 28 occurrence occurrence NN 37809 916 29 in in IN 37809 916 30 Michoacán Michoacán NNP 37809 916 31 of of IN 37809 916 32 low low JJ 37809 916 33 altitude altitude NN 37809 916 34 _ _ NNP 37809 916 35 Tomodactylus Tomodactylus NNP 37809 916 36 _ _ NNP 37809 916 37 ( ( -LRB- 37809 916 38 to to IN 37809 916 39 sea sea NN 37809 916 40 level level NN 37809 916 41 ) ) -RRB- 37809 916 42 . . . 37809 917 1 There there EX 37809 917 2 are be VBP 37809 917 3 no no DT 37809 917 4 _ _ NNP 37809 917 5 Tomodactylus Tomodactylus NNP 37809 917 6 _ _ NNP 37809 917 7 in in IN 37809 917 8 the the DT 37809 917 9 Sierra Sierra NNP 37809 917 10 Madre Madre NNP 37809 917 11 Oriental Oriental NNP 37809 917 12 , , , 37809 917 13 whereas whereas IN 37809 917 14 the the DT 37809 917 15 genus genus NN 37809 917 16 _ _ NNP 37809 917 17 Syrrhophus Syrrhophus NNP 37809 917 18 _ _ NNP 37809 917 19 is be VBZ 37809 917 20 represented represent VBN 37809 917 21 in in IN 37809 917 22 the the DT 37809 917 23 lowlands lowland NNS 37809 917 24 of of IN 37809 917 25 western western JJ 37809 917 26 México México NNP 37809 917 27 ( ( -LRB- 37809 917 28 _ _ NNP 37809 917 29 modestus modestus NN 37809 917 30 _ _ NNP 37809 917 31 group group NN 37809 917 32 ) ) -RRB- 37809 917 33 . . . 37809 918 1 _ _ NNP 37809 918 2 Syrrhophus Syrrhophus NNP 37809 918 3 _ _ NNP 37809 918 4 and and CC 37809 918 5 _ _ NNP 37809 918 6 Tomodactylus Tomodactylus NNP 37809 918 7 _ _ NNP 37809 918 8 exhibit exhibit NN 37809 918 9 essentially essentially RB 37809 918 10 parapatric parapatric VBP 37809 918 11 distributions distribution NNS 37809 918 12 . . . 37809 919 1 The the DT 37809 919 2 two two CD 37809 919 3 genera genera NN 37809 919 4 as as RB 37809 919 5 now now RB 37809 919 6 composed compose VBN 37809 919 7 can can MD 37809 919 8 be be VB 37809 919 9 characterized characterize VBN 37809 919 10 as as RB 37809 919 11 low low JJ 37809 919 12 to to IN 37809 919 13 moderate moderate JJ 37809 919 14 elevation elevation NN 37809 919 15 frogs frog NNS 37809 919 16 ( ( -LRB- 37809 919 17 _ _ NNP 37809 919 18 Syrrhophus Syrrhophus NNP 37809 919 19 _ _ NNP 37809 919 20 ) ) -RRB- 37809 919 21 and and CC 37809 919 22 moderate moderate JJ 37809 919 23 to to TO 37809 919 24 intermediate intermediate VB 37809 919 25 elevation elevation NN 37809 919 26 frogs frog NNS 37809 919 27 ( ( -LRB- 37809 919 28 _ _ NNP 37809 919 29 Tomodactylus Tomodactylus NNP 37809 919 30 _ _ NNP 37809 919 31 ) ) -RRB- 37809 919 32 . . . 37809 920 1 LITERATURE LITERATURE NNP 37809 920 2 CITED cite VBD 37809 920 3 BAIRD BAIRD NNP 37809 920 4 , , , 37809 920 5 S. S. NNP 37809 920 6 F. F. NNP 37809 920 7 1859 1859 CD 37809 920 8 . . . 37809 921 1 Reptiles reptile NNS 37809 921 2 of of IN 37809 921 3 the the DT 37809 921 4 Boundary Boundary NNP 37809 921 5 . . . 37809 922 1 United United NNP 37809 922 2 States States NNP 37809 922 3 and and CC 37809 922 4 Mexican Mexican NNP 37809 922 5 Boundary Boundary NNP 37809 922 6 Survey Survey NNP 37809 922 7 , , , 37809 922 8 pp pp NNP 37809 922 9 . . . 37809 923 1 1 1 CD 37809 923 2 - - SYM 37809 923 3 35 35 CD 37809 923 4 , , , 37809 923 5 pls pls UH 37809 923 6 . . . 37809 924 1 1 1 CD 37809 924 2 - - SYM 37809 924 3 41 41 CD 37809 924 4 . . . 37809 925 1 BARBOUR BARBOUR NNP 37809 925 2 , , , 37809 925 3 T. T. NNP 37809 925 4 1923 1923 CD 37809 925 5 . . . 37809 926 1 The the DT 37809 926 2 reappearance reappearance NN 37809 926 3 of of IN 37809 926 4 Batrachyla Batrachyla NNP 37809 926 5 longipes longipe NNS 37809 926 6 . . . 37809 927 1 Proc Proc NNP 37809 927 2 . . . 37809 928 1 New New NNP 37809 928 2 England England NNP 37809 928 3 Zool Zool NNP 37809 928 4 . . . 37809 929 1 Club Club NNP 37809 929 2 , , , 37809 929 3 8:81 8:81 CD 37809 929 4 - - SYM 37809 929 5 83 83 CD 37809 929 6 . . . 37809 930 1 BARBOUR BARBOUR NNP 37809 930 2 , , , 37809 930 3 T. T. NNP 37809 930 4 , , , 37809 930 5 and and CC 37809 930 6 A. A. NNP 37809 930 7 LOVERIDGE LOVERIDGE NNP 37809 930 8 1946 1946 CD 37809 930 9 . . . 37809 931 1 Typical typical JJ 37809 931 2 reptiles reptile NNS 37809 931 3 and and CC 37809 931 4 amphibians amphibian NNS 37809 931 5 ; ; , 37809 931 6 supplement supplement VB 37809 931 7 . . . 37809 932 1 Bull Bull NNP 37809 932 2 . . . 37809 933 1 Mus mus NN 37809 933 2 . . . 37809 934 1 Comp Comp NNP 37809 934 2 . . . 37809 935 1 Zool Zool NNP 37809 935 2 . . . 37809 935 3 , , , 37809 935 4 96:59 96:59 CD 37809 935 5 - - HYPH 37809 935 6 214 214 CD 37809 935 7 . . . 37809 936 1 BOULENGER BOULENGER NNP 37809 936 2 , , , 37809 936 3 G. G. NNP 37809 936 4 A. A. NNP 37809 937 1 1882 1882 CD 37809 937 2 . . . 37809 938 1 Catalogue catalogue NN 37809 938 2 of of IN 37809 938 3 the the DT 37809 938 4 Batrachia Batrachia NNP 37809 938 5 Salientia Salientia NNP 37809 938 6 ... ... : 37809 938 7 British British NNP 37809 938 8 Museum Museum NNP 37809 938 9 . . NNP 37809 938 10 , , , 37809 938 11 2nd 2nd NNP 37809 938 12 ed ed NNP 37809 938 13 . . . 37809 939 1 1888 1888 CD 37809 939 2 . . . 37809 940 1 Note note VB 37809 940 2 on on IN 37809 940 3 the the DT 37809 940 4 classification classification NN 37809 940 5 of of IN 37809 940 6 the the DT 37809 940 7 Ranidae Ranidae NNP 37809 940 8 . . . 37809 941 1 Proc Proc NNP 37809 941 2 . . . 37809 942 1 Zool Zool NNP 37809 942 2 . . . 37809 943 1 Soc Soc NNP 37809 943 2 . . . 37809 944 1 London London NNP 37809 944 2 , , , 37809 944 3 1888 1888 CD 37809 944 4 , , , 37809 944 5 pt pt UH 37809 944 6 . . . 37809 945 1 2:204 2:204 CD 37809 945 2 - - SYM 37809 945 3 06 06 CD 37809 945 4 . . . 37809 946 1 CAMPBELL CAMPBELL NNP 37809 946 2 , , , 37809 946 3 H. H. NNP 37809 946 4 W. W. NNP 37809 946 5 , , , 37809 946 6 and and CC 37809 946 7 R. R. NNP 37809 946 8 S. S. NNP 37809 946 9 SIMMONS SIMMONS NNP 37809 946 10 1962 1962 CD 37809 946 11 . . . 37809 947 1 Notes note NNS 37809 947 2 on on IN 37809 947 3 some some DT 37809 947 4 reptiles reptile NNS 37809 947 5 and and CC 37809 947 6 amphibians amphibian NNS 37809 947 7 from from IN 37809 947 8 western western JJ 37809 947 9 Mexico Mexico NNP 37809 947 10 . . . 37809 948 1 Bull Bull NNP 37809 948 2 . . . 37809 949 1 So so RB 37809 949 2 . . . 37809 950 1 California California NNP 37809 950 2 Acad Acad NNP 37809 950 3 . . . 37809 951 1 Sci Sci NNP 37809 951 2 . . NNP 37809 951 3 , , , 37809 951 4 61:193 61:193 CD 37809 951 5 - - SYM 37809 951 6 203 203 CD 37809 951 7 . . . 37809 952 1 CONANT CONANT NNP 37809 952 2 , , , 37809 952 3 R. R. NNP 37809 952 4 1958 1958 CD 37809 952 5 . . . 37809 953 1 A a DT 37809 953 2 field field NN 37809 953 3 guide guide NN 37809 953 4 to to IN 37809 953 5 reptiles reptile NNS 37809 953 6 and and CC 37809 953 7 amphibians amphibian NNS 37809 953 8 . . . 37809 954 1 Houghton Houghton NNP 37809 954 2 - - HYPH 37809 954 3 Mifflin Mifflin NNP 37809 954 4 Co. Co. NNP 37809 954 5 Boston Boston NNP 37809 954 6 . . . 37809 955 1 366 366 CD 37809 955 2 pp pp NNP 37809 955 3 . . . 37809 956 1 COPE COPE NNP 37809 956 2 , , , 37809 956 3 E. E. NNP 37809 956 4 D. D. NNP 37809 956 5 1866 1866 CD 37809 956 6 . . . 37809 957 1 On on IN 37809 957 2 the the DT 37809 957 3 structures structure NNS 37809 957 4 and and CC 37809 957 5 distribution distribution NN 37809 957 6 of of IN 37809 957 7 the the DT 37809 957 8 genera genera NN 37809 957 9 of of IN 37809 957 10 the the DT 37809 957 11 arciferous arciferous JJ 37809 957 12 Anura Anura NNP 37809 957 13 . . . 37809 958 1 J. J. NNP 37809 958 2 Acad Acad NNP 37809 958 3 . . . 37809 959 1 Nat Nat NNP 37809 959 2 . . . 37809 960 1 Sci Sci NNP 37809 960 2 . . . 37809 961 1 Philadelphia Philadelphia NNP 37809 961 2 , , , 37809 961 3 n. n. NN 37809 962 1 ser ser NNP 37809 962 2 . . . 37809 962 3 , , , 37809 962 4 6:67 6:67 CD 37809 962 5 - - SYM 37809 962 6 112 112 CD 37809 962 7 . . . 37809 963 1 1877 1877 CD 37809 963 2 . . . 37809 964 1 Tenth tenth JJ 37809 964 2 contribution contribution NN 37809 964 3 to to IN 37809 964 4 the the DT 37809 964 5 herpetology herpetology NN 37809 964 6 of of IN 37809 964 7 tropical tropical JJ 37809 964 8 America America NNP 37809 964 9 . . . 37809 965 1 Proc Proc NNP 37809 965 2 . . . 37809 966 1 Amer amer NN 37809 966 2 . . . 37809 967 1 Philos Philos NNP 37809 967 2 . . . 37809 968 1 Soc Soc NNP 37809 968 2 . . . 37809 968 3 , , , 37809 968 4 17:85 17:85 CD 37809 968 5 - - SYM 37809 968 6 98 98 CD 37809 968 7 . . . 37809 969 1 1878 1878 CD 37809 969 2 . . . 37809 970 1 New new JJ 37809 970 2 genus genus NN 37809 970 3 of of IN 37809 970 4 Cystignathidae Cystignathidae NNP 37809 970 5 from from IN 37809 970 6 Texas Texas NNP 37809 970 7 . . . 37809 971 1 Amer amer NN 37809 971 2 . . . 37809 972 1 Nat Nat NNP 37809 972 2 . . NNP 37809 972 3 , , , 37809 972 4 12:252 12:252 CD 37809 972 5 - - SYM 37809 972 6 53 53 CD 37809 972 7 . . . 37809 973 1 1879 1879 CD 37809 973 2 . . . 37809 974 1 Eleventh eleventh JJ 37809 974 2 contribution contribution NN 37809 974 3 to to IN 37809 974 4 the the DT 37809 974 5 herpetology herpetology NN 37809 974 6 of of IN 37809 974 7 tropical tropical JJ 37809 974 8 America America NNP 37809 974 9 . . . 37809 975 1 Proc Proc NNP 37809 975 2 . . . 37809 976 1 Amer amer NN 37809 976 2 . . . 37809 977 1 Philos Philos NNP 37809 977 2 . . . 37809 978 1 Soc Soc NNP 37809 978 2 . . NNP 37809 978 3 , , , 37809 978 4 18:261 18:261 CD 37809 978 5 - - SYM 37809 978 6 77 77 CD 37809 978 7 . . . 37809 979 1 1885 1885 CD 37809 979 2 . . . 37809 980 1 A a DT 37809 980 2 contribution contribution NN 37809 980 3 to to IN 37809 980 4 the the DT 37809 980 5 herpetology herpetology NN 37809 980 6 of of IN 37809 980 7 Mexico Mexico NNP 37809 980 8 . . . 37809 981 1 _ _ NNP 37809 981 2 Ibid Ibid NNP 37809 981 3 . . . 37809 981 4 _ _ NNP 37809 981 5 , , , 37809 981 6 22:379 22:379 CD 37809 981 7 - - SYM 37809 981 8 404 404 CD 37809 981 9 . . . 37809 982 1 DAVIS DAVIS NNP 37809 982 2 , , , 37809 982 3 W. W. NNP 37809 982 4 B. B. NNP 37809 982 5 , , , 37809 982 6 and and CC 37809 982 7 J. J. NNP 37809 982 8 R. R. NNP 37809 982 9 DIXON DIXON NNP 37809 982 10 1957 1957 CD 37809 982 11 . . . 37809 983 1 Notes note NNS 37809 983 2 on on IN 37809 983 3 Mexican mexican JJ 37809 983 4 amphibians amphibian NNS 37809 983 5 , , , 37809 983 6 with with IN 37809 983 7 description description NN 37809 983 8 of of IN 37809 983 9 a a DT 37809 983 10 new new JJ 37809 983 11 _ _ NNP 37809 983 12 Microbatrachylus Microbatrachylus NNP 37809 983 13 _ _ NNP 37809 983 14 . . . 37809 984 1 Herpetologica Herpetologica NNP 37809 984 2 , , , 37809 984 3 13:145 13:145 CD 37809 984 4 - - SYM 37809 984 5 47 47 CD 37809 984 6 . . . 37809 985 1 1965 1965 CD 37809 985 2 . . . 37809 986 1 Amphibians amphibian NNS 37809 986 2 of of IN 37809 986 3 the the DT 37809 986 4 Chilpancingo Chilpancingo NNP 37809 986 5 Region Region NNP 37809 986 6 , , , 37809 986 7 Mexico Mexico NNP 37809 986 8 . . . 37809 987 1 _ _ NNP 37809 987 2 Ibid Ibid NNP 37809 987 3 . . . 37809 987 4 _ _ NNP 37809 987 5 , , , 37809 987 6 20:225 20:225 CD 37809 987 7 - - SYM 37809 987 8 33 33 CD 37809 987 9 . . . 37809 988 1 DÍAZ DÍAZ NNP 37809 988 2 DE DE NNP 37809 988 3 LEÓN LEÓN NNP 37809 988 4 , , , 37809 988 5 J. J. NNP 37809 989 1 1904 1904 CD 37809 989 2 . . . 37809 990 1 Indice Indice NNP 37809 990 2 de de NNP 37809 990 3 los los NNP 37809 990 4 Batracios Batracios NNP 37809 990 5 que que NNP 37809 990 6 se se NNP 37809 990 7 enquentran enquentran VBP 37809 990 8 en en FW 37809 990 9 la la FW 37809 990 10 Republica Republica NNP 37809 990 11 Méxicana Méxicana NNP 37809 990 12 . . . 37809 991 1 Imprenta Imprenta NNP 37809 991 2 de de NNP 37809 991 3 Ricardo Ricardo NNP 37809 991 4 Rodriquez Rodriquez NNP 37809 991 5 Romo Romo NNP 37809 991 6 . . . 37809 992 1 Aguascalientes aguascaliente NNS 37809 992 2 . . . 37809 993 1 40 40 CD 37809 993 2 pp pp NN 37809 993 3 . . . 37809 994 1 DIXON DIXON NNP 37809 994 2 , , , 37809 994 3 J. J. NNP 37809 994 4 R. R. NNP 37809 994 5 1957 1957 CD 37809 994 6 . . . 37809 995 1 Geographic geographic JJ 37809 995 2 variation variation NN 37809 995 3 and and CC 37809 995 4 distribution distribution NN 37809 995 5 of of IN 37809 995 6 the the DT 37809 995 7 genus genus NNP 37809 995 8 Tomodactylus Tomodactylus NNP 37809 995 9 in in IN 37809 995 10 Mexico Mexico NNP 37809 995 11 . . . 37809 996 1 Texas Texas NNP 37809 996 2 J. J. NNP 37809 997 1 Sci Sci NNP 37809 997 2 . . NNP 37809 997 3 , , , 37809 997 4 9:379 9:379 CD 37809 997 5 - - HYPH 37809 997 6 409 409 CD 37809 997 7 . . . 37809 998 1 DIXON DIXON NNP 37809 998 2 , , , 37809 998 3 J. J. NNP 37809 998 4 R. R. NNP 37809 998 5 , , , 37809 998 6 and and CC 37809 998 7 R. R. NNP 37809 998 8 G. G. NNP 37809 998 9 WEBB WEBB NNP 37809 998 10 1966 1966 CD 37809 998 11 . . . 37809 999 1 A a DT 37809 999 2 new new JJ 37809 999 3 _ _ NNP 37809 999 4 Syrrhophus Syrrhophus NNP 37809 999 5 _ _ NNP 37809 999 6 from from IN 37809 999 7 Mexico Mexico NNP 37809 999 8 ( ( -LRB- 37809 999 9 Amphibia Amphibia NNP 37809 999 10 : : : 37809 999 11 Leptodactylidae Leptodactylidae NNP 37809 999 12 ) ) -RRB- 37809 999 13 . . . 37809 1000 1 Cont Cont NNP 37809 1000 2 . . . 37809 1001 1 Sc sc IN 37809 1001 2 . . . 37809 1001 3 , , , 37809 1001 4 Los Los NNP 37809 1001 5 Angeles Angeles NNP 37809 1001 6 Co. Co. NNP 37809 1002 1 Mus Mus NNP 37809 1002 2 . . NNP 37809 1002 3 , , , 37809 1002 4 102:1 102:1 CD 37809 1002 5 - - SYM 37809 1002 6 5 5 CD 37809 1002 7 . . . 37809 1003 1 DUELLMAN DUELLMAN NNP 37809 1003 2 , , , 37809 1003 3 W. W. NNP 37809 1003 4 E. E. NNP 37809 1003 5 1958 1958 CD 37809 1003 6 . . . 37809 1004 1 A a DT 37809 1004 2 review review NN 37809 1004 3 of of IN 37809 1004 4 the the DT 37809 1004 5 frogs frog NNS 37809 1004 6 of of IN 37809 1004 7 the the DT 37809 1004 8 genus genus NNP 37809 1004 9 _ _ NNP 37809 1004 10 Syrrhophus Syrrhophus NNP 37809 1004 11 _ _ NNP 37809 1004 12 in in IN 37809 1004 13 western western JJ 37809 1004 14 Mexico Mexico NNP 37809 1004 15 . . . 37809 1005 1 Occ occ UH 37809 1005 2 . . . 37809 1006 1 Pap Pap NNP 37809 1006 2 . . . 37809 1007 1 Mus mus NN 37809 1007 2 . . . 37809 1008 1 Zool Zool NNP 37809 1008 2 . . . 37809 1009 1 Univ Univ NNP 37809 1009 2 . . . 37809 1010 1 Michigan Michigan NNP 37809 1010 2 , , , 37809 1010 3 594:1 594:1 CD 37809 1010 4 - - SYM 37809 1010 5 15 15 CD 37809 1010 6 . . . 37809 1011 1 1960 1960 CD 37809 1011 2 . . . 37809 1012 1 A a DT 37809 1012 2 distributional distributional JJ 37809 1012 3 study study NN 37809 1012 4 of of IN 37809 1012 5 the the DT 37809 1012 6 amphibians amphibian NNS 37809 1012 7 of of IN 37809 1012 8 the the DT 37809 1012 9 Isthmus Isthmus NNP 37809 1012 10 of of IN 37809 1012 11 Tehuantepec Tehuantepec NNP 37809 1012 12 , , , 37809 1012 13 Mexico Mexico NNP 37809 1012 14 . . . 37809 1013 1 Univ Univ NNP 37809 1013 2 . . . 37809 1014 1 Kansas Kansas NNP 37809 1014 2 Publs Publs NNP 37809 1014 3 . . . 37809 1015 1 Mus mus NN 37809 1015 2 . . . 37809 1016 1 Nat Nat NNP 37809 1016 2 . . . 37809 1017 1 Hist Hist NNP 37809 1017 2 . . . 37809 1017 3 , , , 37809 1017 4 13:19 13:19 CD 37809 1017 5 - - SYM 37809 1017 6 72 72 CD 37809 1017 7 . . . 37809 1018 1 FIRSCHEIN FIRSCHEIN NNP 37809 1018 2 , , , 37809 1018 3 I. I. NNP 37809 1018 4 L. L. NNP 37809 1018 5 1954 1954 CD 37809 1018 6 . . . 37809 1019 1 Definition definition NN 37809 1019 2 of of IN 37809 1019 3 some some DT 37809 1019 4 little little RB 37809 1019 5 - - HYPH 37809 1019 6 understood understand VBN 37809 1019 7 members member NNS 37809 1019 8 of of IN 37809 1019 9 the the DT 37809 1019 10 leptodactylid leptodactylid JJ 37809 1019 11 genus genus NNP 37809 1019 12 _ _ NNP 37809 1019 13 Syrrhophus Syrrhophus NNP 37809 1019 14 _ _ NNP 37809 1019 15 , , , 37809 1019 16 with with IN 37809 1019 17 a a DT 37809 1019 18 description description NN 37809 1019 19 of of IN 37809 1019 20 a a DT 37809 1019 21 new new JJ 37809 1019 22 species specie NNS 37809 1019 23 . . . 37809 1020 1 Copeia Copeia NNP 37809 1020 2 , , , 37809 1020 3 ( ( -LRB- 37809 1020 4 1):48 1):48 CD 37809 1020 5 - - SYM 37809 1020 6 58 58 CD 37809 1020 7 . . . 37809 1021 1 FOUQUETTE FOUQUETTE NNP 37809 1021 2 , , , 37809 1021 3 M. M. NNP 37809 1021 4 J. J. NNP 37809 1022 1 1960 1960 CD 37809 1022 2 . . . 37809 1023 1 Call call VB 37809 1023 2 structure structure NN 37809 1023 3 in in IN 37809 1023 4 frogs frog NNS 37809 1023 5 of of IN 37809 1023 6 the the DT 37809 1023 7 family family NN 37809 1023 8 Leptodactylidae Leptodactylidae NNP 37809 1023 9 . . . 37809 1024 1 Texas Texas NNP 37809 1024 2 J. J. NNP 37809 1025 1 Sci Sci NNP 37809 1025 2 . . NNP 37809 1025 3 , , , 37809 1025 4 12:201 12:201 CD 37809 1025 5 - - SYM 37809 1025 6 15 15 CD 37809 1025 7 . . . 37809 1026 1 GADOW GADOW NNP 37809 1026 2 , , , 37809 1026 3 H. H. NNP 37809 1026 4 1905 1905 CD 37809 1026 5 . . . 37809 1027 1 The the DT 37809 1027 2 distribution distribution NN 37809 1027 3 of of IN 37809 1027 4 Mexican mexican JJ 37809 1027 5 amphibians amphibian NNS 37809 1027 6 and and CC 37809 1027 7 reptiles reptile NNS 37809 1027 8 . . . 37809 1028 1 Proc Proc NNP 37809 1028 2 . . . 37809 1029 1 Zool Zool NNP 37809 1029 2 . . . 37809 1030 1 Soc Soc NNP 37809 1030 2 . . . 37809 1031 1 London London NNP 37809 1031 2 , , , 37809 1031 3 1905 1905 CD 37809 1031 4 , , , 37809 1031 5 pt pt UH 37809 1031 6 . . . 37809 1032 1 2:191 2:191 CD 37809 1032 2 - - SYM 37809 1032 3 244 244 CD 37809 1032 4 . . . 37809 1033 1 GORHAM GORHAM NNP 37809 1033 2 , , , 37809 1033 3 S. S. NNP 37809 1033 4 W. W. NNP 37809 1033 5 1966 1966 CD 37809 1033 6 . . . 37809 1034 1 Liste liste VB 37809 1034 2 der der RB 37809 1034 3 rezenten rezenten VBN 37809 1034 4 Amphibien Amphibien NNP 37809 1034 5 und und NN 37809 1034 6 Reptilien Reptilien NNP 37809 1034 7 .... .... . 37809 1034 8 Das Das NNP 37809 1034 9 Tierreich Tierreich NNP 37809 1034 10 . . . 37809 1035 1 Lief Lief NNP 37809 1035 2 , , , 37809 1035 3 85:1 85:1 CD 37809 1035 4 - - SYM 37809 1035 5 222 222 CD 37809 1035 6 . . . 37809 1036 1 GÜNTHER GÜNTHER NNP 37809 1036 2 , , , 37809 1036 3 A. a. NN 37809 1036 4 C. C. NNP 37809 1036 5 L. L. NNP 37809 1036 6 G. G. NNP 37809 1036 7 1885 1885 CD 37809 1036 8 - - SYM 37809 1036 9 1902 1902 CD 37809 1036 10 . . . 37809 1037 1 Biologia Biologia NNP 37809 1037 2 Centrali Centrali NNP 37809 1037 3 - - HYPH 37809 1037 4 Americana Americana NNP 37809 1037 5 . . . 37809 1038 1 Reptilia Reptilia NNP 37809 1038 2 and and CC 37809 1038 3 Batrachia Batrachia NNP 37809 1038 4 . . . 37809 1039 1 326 326 CD 37809 1039 2 pp pp NNP 37809 1039 3 . . NNP 37809 1039 4 , , , 37809 1039 5 76 76 CD 37809 1039 6 pls pls UH 37809 1039 7 . . . 37809 1040 1 Syrrhophus Syrrhophus NNP 37809 1040 2 section section NN 37809 1040 3 dated date VBD 37809 1040 4 1900 1900 CD 37809 1040 5 . . . 37809 1041 1 KELLOGG KELLOGG NNP 37809 1041 2 , , , 37809 1041 3 R. R. NNP 37809 1041 4 1932 1932 CD 37809 1041 5 . . . 37809 1042 1 Mexican mexican JJ 37809 1042 2 tailless tailless NN 37809 1042 3 amphibians amphibian NNS 37809 1042 4 in in IN 37809 1042 5 the the DT 37809 1042 6 United United NNP 37809 1042 7 States States NNP 37809 1042 8 National National NNP 37809 1042 9 Museum Museum NNP 37809 1042 10 . . . 37809 1043 1 Bull Bull NNP 37809 1043 2 . . . 37809 1044 1 U.S. U.S. NNP 37809 1044 2 Natl Natl NNP 37809 1044 3 . . . 37809 1045 1 Mus Mus NNP 37809 1045 2 . . NNP 37809 1045 3 , , , 37809 1045 4 160 160 CD 37809 1045 5 . . . 37809 1046 1 LANGEBARTEL LANGEBARTEL NNP 37809 1046 2 , , , 37809 1046 3 D. D. NNP 37809 1046 4 A. A. NNP 37809 1046 5 , , , 37809 1046 6 and and CC 37809 1046 7 F. F. NNP 37809 1046 8 A. A. NNP 37809 1046 9 SHANNON SHANNON NNP 37809 1046 10 1956 1956 CD 37809 1046 11 . . . 37809 1047 1 A a DT 37809 1047 2 new new JJ 37809 1047 3 frog frog NN 37809 1047 4 ( ( -LRB- 37809 1047 5 Syrrhophus Syrrhophus NNP 37809 1047 6 ) ) -RRB- 37809 1047 7 from from IN 37809 1047 8 the the DT 37809 1047 9 Sinoloan Sinoloan NNP 37809 1047 10 lowlands lowland NNS 37809 1047 11 of of IN 37809 1047 12 Mexico Mexico NNP 37809 1047 13 . . . 37809 1048 1 Herpetologica Herpetologica NNP 37809 1048 2 , , , 37809 1048 3 12:161 12:161 CD 37809 1048 4 - - SYM 37809 1048 5 65 65 CD 37809 1048 6 . . . 37809 1049 1 LYNCH LYNCH NNP 37809 1049 2 , , , 37809 1049 3 J. J. NNP 37809 1049 4 D. D. NNP 37809 1049 5 1963 1963 CD 37809 1049 6 . . . 37809 1050 1 The the DT 37809 1050 2 status status NN 37809 1050 3 of of IN 37809 1050 4 _ _ NNP 37809 1050 5 Eleutherodactylus Eleutherodactylus NNP 37809 1050 6 longipes longipe NNS 37809 1050 7 _ _ NNP 37809 1050 8 ( ( -LRB- 37809 1050 9 Baird Baird NNP 37809 1050 10 ) ) -RRB- 37809 1050 11 of of IN 37809 1050 12 Mexico Mexico NNP 37809 1050 13 ( ( -LRB- 37809 1050 14 Amphibia Amphibia NNP 37809 1050 15 : : : 37809 1050 16 Leptodactylidae Leptodactylidae NNP 37809 1050 17 ) ) -RRB- 37809 1050 18 . . . 37809 1051 1 Copeia Copeia NNP 37809 1051 2 , , , 37809 1051 3 ( ( -LRB- 37809 1051 4 3):580 3):580 NNP 37809 1051 5 - - HYPH 37809 1051 6 81 81 CD 37809 1051 7 . . . 37809 1052 1 1964 1964 CD 37809 1052 2 . . . 37809 1053 1 Additional additional JJ 37809 1053 2 hylid hylid NN 37809 1053 3 and and CC 37809 1053 4 leptodactylid leptodactylid JJ 37809 1053 5 remains remain NNS 37809 1053 6 from from IN 37809 1053 7 the the DT 37809 1053 8 Pleistocene Pleistocene NNP 37809 1053 9 of of IN 37809 1053 10 Texas Texas NNP 37809 1053 11 and and CC 37809 1053 12 Florida Florida NNP 37809 1053 13 Herpetologica Herpetologica NNP 37809 1053 14 . . . 37809 1054 1 20:141 20:141 CD 37809 1054 2 - - SYM 37809 1054 3 42 42 CD 37809 1054 4 . . . 37809 1055 1 1967 1967 CD 37809 1055 2 . . . 37809 1056 1 _ _ NNP 37809 1056 2 Epirhexis Epirhexis NNP 37809 1056 3 _ _ NNP 37809 1056 4 Cope Cope NNP 37809 1056 5 , , , 37809 1056 6 1866 1866 CD 37809 1056 7 ( ( -LRB- 37809 1056 8 Amphibia Amphibia NNP 37809 1056 9 : : : 37809 1056 10 Salientia Salientia NNP 37809 1056 11 ) ) -RRB- 37809 1056 12 : : : 37809 1056 13 request request NN 37809 1056 14 for for IN 37809 1056 15 suppression suppression NN 37809 1056 16 under under IN 37809 1056 17 the the DT 37809 1056 18 plenary plenary JJ 37809 1056 19 powers power NNS 37809 1056 20 . . . 37809 1057 1 I. I. NNP 37809 1057 2 N. N. NNP 37809 1057 3 ( ( -LRB- 37809 1057 4 S S NNP 37809 1057 5 ) ) -RRB- 37809 1057 6 . . . 37809 1058 1 Bull Bull NNP 37809 1058 2 . . . 37809 1059 1 Zool Zool NNP 37809 1059 2 . . . 37809 1060 1 Nomencl Nomencl NNP 37809 1060 2 . . NNP 37809 1060 3 , , , 37809 1060 4 24:313 24:313 CD 37809 1060 5 - - SYM 37809 1060 6 15 15 CD 37809 1060 7 . . . 37809 1061 1 1968 1968 CD 37809 1061 2 . . . 37809 1062 1 Genera Genera NNP 37809 1062 2 of of IN 37809 1062 3 leptodactylid leptodactylid JJ 37809 1062 4 frogs frog NNS 37809 1062 5 in in IN 37809 1062 6 México México NNP 37809 1062 7 . . . 37809 1063 1 Univ Univ NNP 37809 1063 2 . . . 37809 1064 1 Kansas Kansas NNP 37809 1064 2 Publs Publs NNP 37809 1064 3 . . . 37809 1064 4 , , , 37809 1064 5 Mus mus NN 37809 1064 6 . . . 37809 1065 1 Nat Nat NNP 37809 1065 2 . . . 37809 1066 1 Hist Hist NNP 37809 1066 2 . . . 37809 1066 3 , , , 37809 1066 4 17:503 17:503 CD 37809 1066 5 - - SYM 37809 1066 6 15 15 CD 37809 1066 7 . . . 37809 1067 1 MARTIN MARTIN NNP 37809 1067 2 , , , 37809 1067 3 P. P. NNP 37809 1067 4 S. S. NNP 37809 1067 5 1958 1958 CD 37809 1067 6 . . . 37809 1068 1 A a DT 37809 1068 2 biogeography biogeography NN 37809 1068 3 of of IN 37809 1068 4 reptiles reptile NNS 37809 1068 5 and and CC 37809 1068 6 amphibians amphibian NNS 37809 1068 7 in in IN 37809 1068 8 the the DT 37809 1068 9 Gomez Gomez NNP 37809 1068 10 Farias Farias NNP 37809 1068 11 region region NN 37809 1068 12 , , , 37809 1068 13 Tamaulipas Tamaulipas NNP 37809 1068 14 , , , 37809 1068 15 Mexico Mexico NNP 37809 1068 16 . . . 37809 1069 1 Misc Misc NNP 37809 1069 2 . . . 37809 1070 1 Publs publs RB 37809 1070 2 . . . 37809 1071 1 Mus mus NN 37809 1071 2 . . . 37809 1072 1 Zool Zool NNP 37809 1072 2 . . . 37809 1073 1 Univ Univ NNP 37809 1073 2 . . . 37809 1074 1 Michigan Michigan NNP 37809 1074 2 , , , 37809 1074 3 101:1 101:1 CD 37809 1074 4 - - SYM 37809 1074 5 102 102 CD 37809 1074 6 . . . 37809 1075 1 MILSTEAD MILSTEAD NNP 37809 1075 2 , , , 37809 1075 3 W. W. NNP 37809 1075 4 M. M. NNP 37809 1075 5 , , , 37809 1075 6 J. J. NNP 37809 1075 7 S. S. NNP 37809 1075 8 MECHAM MECHAM NNP 37809 1075 9 , , , 37809 1075 10 and and CC 37809 1075 11 H. H. NNP 37809 1075 12 MCCLINTOCK MCCLINTOCK NNP 37809 1075 13 1950 1950 CD 37809 1075 14 . . . 37809 1076 1 The the DT 37809 1076 2 amphibians amphibian NNS 37809 1076 3 and and CC 37809 1076 4 reptiles reptile NNS 37809 1076 5 of of IN 37809 1076 6 the the DT 37809 1076 7 Stockton Stockton NNP 37809 1076 8 Plateau Plateau NNP 37809 1076 9 in in IN 37809 1076 10 northern northern JJ 37809 1076 11 Terrell Terrell NNP 37809 1076 12 County County NNP 37809 1076 13 , , , 37809 1076 14 Texas Texas NNP 37809 1076 15 . . . 37809 1077 1 Texas Texas NNP 37809 1077 2 J. J. NNP 37809 1078 1 Sci Sci NNP 37809 1078 2 . . NNP 37809 1078 3 , , , 37809 1078 4 2:543 2:543 CD 37809 1078 5 - - SYM 37809 1078 6 62 62 CD 37809 1078 7 . . . 37809 1079 1 NEILL NEILL NNP 37809 1079 2 , , , 37809 1079 3 W. W. NNP 37809 1079 4 T. T. NNP 37809 1079 5 1965 1965 CD 37809 1079 6 . . . 37809 1080 1 New new JJ 37809 1080 2 and and CC 37809 1080 3 noteworthy noteworthy JJ 37809 1080 4 amphibians amphibian NNS 37809 1080 5 and and CC 37809 1080 6 reptiles reptile NNS 37809 1080 7 from from IN 37809 1080 8 British british JJ 37809 1080 9 Honduras hondura NNS 37809 1080 10 . . . 37809 1081 1 Bull Bull NNP 37809 1081 2 . . . 37809 1082 1 Florida Florida NNP 37809 1082 2 State State NNP 37809 1082 3 Mus Mus NNP 37809 1082 4 . . NNP 37809 1082 5 , , , 37809 1082 6 9:77 9:77 CD 37809 1082 7 - - SYM 37809 1082 8 130 130 CD 37809 1082 9 . . . 37809 1083 1 NIEDEN NIEDEN NNP 37809 1083 2 , , , 37809 1083 3 F. F. NNP 37809 1083 4 1923 1923 CD 37809 1083 5 . . . 37809 1084 1 Anura anura UH 37809 1084 2 I -PRON- PRP 37809 1084 3 ... ... . 37809 1084 4 Das Das NNP 37809 1084 5 Tierreich Tierreich NNP 37809 1084 6 . . . 37809 1085 1 Lief Lief NNP 37809 1085 2 . . . 37809 1085 3 , , , 37809 1085 4 46:1 46:1 CD 37809 1085 5 - - SYM 37809 1085 6 584 584 CD 37809 1085 7 . . . 37809 1086 1 PETERS PETERS NNP 37809 1086 2 , , , 37809 1086 3 W. W. NNP 37809 1086 4 1871 1871 CD 37809 1086 5 . . . 37809 1087 1 Über Über NNP 37809 1087 2 neue neue NN 37809 1087 3 Amphibien Amphibien NNP 37809 1087 4 ... ... : 37809 1087 5 des des NNP 37809 1087 6 Konigl Konigl NNP 37809 1087 7 . . . 37809 1088 1 Zoologischen Zoologischen NNP 37809 1088 2 Museums Museums NNP 37809 1088 3 . . . 37809 1089 1 Monatsb Monatsb NNP 37809 1089 2 . . . 37809 1090 1 k. k. NNP 37809 1090 2 k. k. NNP 37809 1090 3 Preuss Preuss NNP 37809 1090 4 . . . 37809 1091 1 Akad Akad NNP 37809 1091 2 . . . 37809 1092 1 Wiss Wiss NNP 37809 1092 2 . . . 37809 1093 1 Berlin Berlin NNP 37809 1093 2 , , , 37809 1093 3 1870:641 1870:641 CD 37809 1093 4 - - SYM 37809 1093 5 52 52 CD 37809 1093 6 . . . 37809 1094 1 SCHMIDT schmidt NN 37809 1094 2 , , , 37809 1094 3 K. K. NNP 37809 1094 4 P. P. NNP 37809 1094 5 , , , 37809 1094 6 and and CC 37809 1094 7 T. T. NNP 37809 1094 8 F. F. NNP 37809 1094 9 SMITH SMITH NNP 37809 1094 10 1944 1944 CD 37809 1094 11 . . . 37809 1095 1 Amphibians amphibian NNS 37809 1095 2 and and CC 37809 1095 3 reptiles reptile NNS 37809 1095 4 of of IN 37809 1095 5 the the DT 37809 1095 6 Big Big NNP 37809 1095 7 Bend Bend NNP 37809 1095 8 Region Region NNP 37809 1095 9 of of IN 37809 1095 10 Texas Texas NNP 37809 1095 11 . . . 37809 1096 1 Field field NN 37809 1096 2 Mus Mus NNP 37809 1096 3 . . . 37809 1097 1 Nat Nat NNP 37809 1097 2 . . . 37809 1098 1 Hist Hist NNP 37809 1098 2 . . NNP 37809 1098 3 , , , 37809 1098 4 zool zool NNP 37809 1098 5 . . . 37809 1099 1 ser ser NNP 37809 1099 2 . . . 37809 1099 3 , , , 37809 1099 4 29:75 29:75 CD 37809 1099 5 - - SYM 37809 1099 6 96 96 CD 37809 1099 7 . . . 37809 1100 1 SMITH SMITH NNP 37809 1100 2 , , , 37809 1100 3 H. H. NNP 37809 1100 4 M. M. NNP 37809 1100 5 1947 1947 CD 37809 1100 6 . . . 37809 1101 1 Notes note NNS 37809 1101 2 on on IN 37809 1101 3 Mexican mexican JJ 37809 1101 4 amphibians amphibian NNS 37809 1101 5 and and CC 37809 1101 6 reptiles reptile NNS 37809 1101 7 . . . 37809 1102 1 J. J. NNP 37809 1102 2 Washington Washington NNP 37809 1102 3 Acad Acad NNP 37809 1102 4 . . . 37809 1103 1 Sci Sci NNP 37809 1103 2 . . NNP 37809 1103 3 , , , 37809 1103 4 37:408 37:408 CD 37809 1103 5 - - SYM 37809 1103 6 12 12 CD 37809 1103 7 . . . 37809 1104 1 SMITH SMITH NNP 37809 1104 2 , , , 37809 1104 3 H. H. NNP 37809 1104 4 M. M. NNP 37809 1104 5 , , , 37809 1104 6 and and CC 37809 1104 7 E. E. NNP 37809 1104 8 H. H. NNP 37809 1104 9 TAYLOR TAYLOR NNP 37809 1104 10 1948 1948 CD 37809 1104 11 . . . 37809 1105 1 An an DT 37809 1105 2 annotated annotate VBN 37809 1105 3 checklist checklist NN 37809 1105 4 and and CC 37809 1105 5 key key NN 37809 1105 6 to to IN 37809 1105 7 the the DT 37809 1105 8 Amphibia Amphibia NNP 37809 1105 9 of of IN 37809 1105 10 Mexico Mexico NNP 37809 1105 11 . . . 37809 1106 1 Bull Bull NNP 37809 1106 2 . . . 37809 1107 1 U.S. U.S. NNP 37809 1107 2 Natl Natl NNP 37809 1107 3 . . . 37809 1108 1 Mus Mus NNP 37809 1108 2 . . NNP 37809 1108 3 , , , 37809 1108 4 194:1 194:1 CD 37809 1108 5 - - SYM 37809 1108 6 118 118 CD 37809 1108 7 . . . 37809 1109 1 STEJNEGER STEJNEGER NNP 37809 1109 2 , , , 37809 1109 3 L. L. NNP 37809 1109 4 1915 1915 CD 37809 1109 5 . . . 37809 1110 1 A a DT 37809 1110 2 new new JJ 37809 1110 3 species specie NNS 37809 1110 4 of of IN 37809 1110 5 tailless tailless NN 37809 1110 6 batrachian batrachian NN 37809 1110 7 from from IN 37809 1110 8 North North NNP 37809 1110 9 America America NNP 37809 1110 10 . . . 37809 1111 1 Proc Proc NNP 37809 1111 2 . . . 37809 1112 1 Biol Biol NNP 37809 1112 2 . . . 37809 1113 1 Soc Soc NNP 37809 1113 2 . . . 37809 1114 1 Washington Washington NNP 37809 1114 2 , , , 37809 1114 3 28:131 28:131 CD 37809 1114 4 - - SYM 37809 1114 5 32 32 CD 37809 1114 6 . . . 37809 1115 1 TAYLOR TAYLOR NNP 37809 1115 2 , , , 37809 1115 3 E. E. NNP 37809 1115 4 H. H. NNP 37809 1115 5 1940a 1940a CD 37809 1115 6 . . . 37809 1116 1 A a DT 37809 1116 2 new new JJ 37809 1116 3 eleutherodactylid eleutherodactylid JJ 37809 1116 4 frog frog NN 37809 1116 5 from from IN 37809 1116 6 Mexico Mexico NNP 37809 1116 7 . . . 37809 1117 1 Proc Proc NNP 37809 1117 2 . . . 37809 1118 1 New New NNP 37809 1118 2 England England NNP 37809 1118 3 Zool Zool NNP 37809 1118 4 . . . 37809 1119 1 Club Club NNP 37809 1119 2 , , , 37809 1119 3 18:13 18:13 CD 37809 1119 4 - - SYM 37809 1119 5 16 16 CD 37809 1119 6 . . . 37809 1120 1 1940b 1940b CD 37809 1120 2 . . . 37809 1121 1 Two two CD 37809 1121 2 new new JJ 37809 1121 3 anuran anuran NN 37809 1121 4 amphibians amphibian NNS 37809 1121 5 from from IN 37809 1121 6 Mexico Mexico NNP 37809 1121 7 . . . 37809 1122 1 Proc Proc NNP 37809 1122 2 . . . 37809 1123 1 U.S. U.S. NNP 37809 1123 2 Natl Natl NNP 37809 1123 3 . . . 37809 1124 1 Mus Mus NNP 37809 1124 2 . . . 37809 1124 3 , , , 37809 1124 4 89:43 89:43 CD 37809 1124 5 - - SYM 37809 1124 6 47 47 CD 37809 1124 7 , , , 37809 1124 8 1 1 CD 37809 1124 9 pl pl NN 37809 1124 10 . . . 37809 1125 1 1940c 1940c CD 37809 1125 2 . . . 37809 1126 1 A a DT 37809 1126 2 new new JJ 37809 1126 3 Syrrhophus Syrrhophus NNP 37809 1126 4 from from IN 37809 1126 5 Guerrero Guerrero NNP 37809 1126 6 , , , 37809 1126 7 Mexico Mexico NNP 37809 1126 8 . . . 37809 1127 1 Proc Proc NNP 37809 1127 2 . . . 37809 1128 1 Biol Biol NNP 37809 1128 2 . . . 37809 1129 1 Soc Soc NNP 37809 1129 2 . . . 37809 1130 1 Washington Washington NNP 37809 1130 2 , , , 37809 1130 3 53:95 53:95 CD 37809 1130 4 - - SYM 37809 1130 5 98 98 CD 37809 1130 6 , , , 37809 1130 7 1 1 CD 37809 1130 8 pl pl NN 37809 1130 9 . . . 37809 1131 1 1940d 1940d CD 37809 1131 2 . . . 37809 1132 1 New new JJ 37809 1132 2 species specie NNS 37809 1132 3 of of IN 37809 1132 4 Mexican Mexican NNP 37809 1132 5 Anura Anura NNP 37809 1132 6 . . . 37809 1133 1 Univ Univ NNP 37809 1133 2 . . . 37809 1134 1 Kansas Kansas NNP 37809 1134 2 Sci Sci NNP 37809 1134 3 . . . 37809 1135 1 Bull Bull NNP 37809 1135 2 . . . 37809 1135 3 , , , 37809 1135 4 26:385 26:385 CD 37809 1135 5 - - SYM 37809 1135 6 405 405 CD 37809 1135 7 . . . 37809 1136 1 1940e 1940e CD 37809 1136 2 . . . 37809 1137 1 Herpetological herpetological JJ 37809 1137 2 miscellany miscellany JJ 37809 1137 3 no no UH 37809 1137 4 . . . 37809 1138 1 I. I. NNP 37809 1139 1 _ _ NNP 37809 1139 2 Ibid Ibid NNP 37809 1139 3 . . . 37809 1139 4 _ _ NNP 37809 1139 5 , , , 37809 1139 6 26:489 26:489 CD 37809 1139 7 - - SYM 37809 1139 8 571 571 CD 37809 1139 9 . . . 37809 1140 1 1942 1942 CD 37809 1140 2 . . . 37809 1141 1 New New NNP 37809 1141 2 Caudata Caudata NNP 37809 1141 3 and and CC 37809 1141 4 Salientia Salientia NNP 37809 1141 5 from from IN 37809 1141 6 México México NNP 37809 1141 7 . . . 37809 1142 1 _ _ NNP 37809 1142 2 Ibid Ibid NNP 37809 1142 3 . . . 37809 1142 4 _ _ NNP 37809 1142 5 , , , 37809 1142 6 28:295 28:295 CD 37809 1142 7 - - SYM 37809 1142 8 323 323 CD 37809 1142 9 . . . 37809 1143 1 1943 1943 CD 37809 1143 2 . . . 37809 1144 1 Herpetological herpetological JJ 37809 1144 2 novelties novelty NNS 37809 1144 3 from from IN 37809 1144 4 Mexico Mexico NNP 37809 1144 5 . . . 37809 1145 1 _ _ NNP 37809 1145 2 Ibid Ibid NNP 37809 1145 3 . . . 37809 1145 4 _ _ NNP 37809 1145 5 , , , 37809 1145 6 29:343 29:343 CD 37809 1145 7 - - SYM 37809 1145 8 61 61 CD 37809 1145 9 . . . 37809 1146 1 1952 1952 CD 37809 1146 2 . . . 37809 1147 1 A a DT 37809 1147 2 review review NN 37809 1147 3 of of IN 37809 1147 4 the the DT 37809 1147 5 frogs frog NNS 37809 1147 6 and and CC 37809 1147 7 toads toad NNS 37809 1147 8 of of IN 37809 1147 9 Costa Costa NNP 37809 1147 10 Rica Rica NNP 37809 1147 11 . . . 37809 1148 1 _ _ NNP 37809 1148 2 Ibid Ibid NNP 37809 1148 3 . . . 37809 1148 4 _ _ NNP 37809 1148 5 , , , 37809 1148 6 35:577 35:577 CD 37809 1148 7 - - SYM 37809 1148 8 942 942 CD 37809 1148 9 . . . 37809 1149 1 TAYLOR TAYLOR NNP 37809 1149 2 , , , 37809 1149 3 E. E. NNP 37809 1149 4 H. H. NNP 37809 1149 5 , , , 37809 1149 6 and and CC 37809 1149 7 H. H. NNP 37809 1149 8 M. M. NNP 37809 1149 9 SMITH SMITH NNP 37809 1149 10 1945 1945 CD 37809 1149 11 . . . 37809 1150 1 Summary summary NN 37809 1150 2 of of IN 37809 1150 3 the the DT 37809 1150 4 collections collection NNS 37809 1150 5 of of IN 37809 1150 6 amphibians amphibian NNS 37809 1150 7 made make VBN 37809 1150 8 in in IN 37809 1150 9 Mexico Mexico NNP 37809 1150 10 under under IN 37809 1150 11 the the DT 37809 1150 12 Walter Walter NNP 37809 1150 13 Rathbone Rathbone NNP 37809 1150 14 Bacon Bacon NNP 37809 1150 15 Traveling Traveling NNP 37809 1150 16 Scholarship Scholarship NNP 37809 1150 17 . . . 37809 1151 1 Proc Proc NNP 37809 1151 2 . . . 37809 1152 1 U.S. U.S. NNP 37809 1152 2 Natl Natl NNP 37809 1152 3 . . . 37809 1153 1 Mus Mus NNP 37809 1153 2 . . NNP 37809 1153 3 , , , 37809 1153 4 95:521 95:521 CD 37809 1153 5 - - SYM 37809 1153 6 613 613 CD 37809 1153 7 . . . 37809 1154 1 TIHEN TIHEN NNP 37809 1154 2 , , , 37809 1154 3 J. J. NNP 37809 1155 1 A. A. NNP 37809 1156 1 1960 1960 CD 37809 1156 2 . . . 37809 1157 1 Notes note NNS 37809 1157 2 on on IN 37809 1157 3 Late Late NNP 37809 1157 4 Cenozoic Cenozoic NNP 37809 1157 5 hylid hylid NN 37809 1157 6 and and CC 37809 1157 7 leptodactylid leptodactylid JJ 37809 1157 8 frogs frog NNS 37809 1157 9 from from IN 37809 1157 10 Kansas Kansas NNP 37809 1157 11 , , , 37809 1157 12 Oklahoma Oklahoma NNP 37809 1157 13 and and CC 37809 1157 14 Texas Texas NNP 37809 1157 15 . . . 37809 1158 1 Southwest southwest UH 37809 1158 2 . . . 37809 1159 1 Nat Nat NNP 37809 1159 2 . . NNP 37809 1159 3 , , , 37809 1159 4 5:66 5:66 CD 37809 1159 5 - - SYM 37809 1159 6 70 70 CD 37809 1159 7 . . . 37809 1160 1 WRIGHT WRIGHT NNP 37809 1160 2 , , , 37809 1160 3 A. a. NN 37809 1160 4 H. H. NNP 37809 1160 5 , , , 37809 1160 6 and and CC 37809 1160 7 A. a. NN 37809 1161 1 A. A. NNP 37809 1161 2 WRIGHT WRIGHT NNP 37809 1161 3 1949 1949 CD 37809 1161 4 . . . 37809 1162 1 Handbook handbook NN 37809 1162 2 of of IN 37809 1162 3 frogs frog NNS 37809 1162 4 and and CC 37809 1162 5 toads toad NNS 37809 1162 6 . . . 37809 1163 1 3rd 3rd NNP 37809 1163 2 ed ed NNP 37809 1163 3 . . . 37809 1164 1 Comstock comstock NN 37809 1164 2 . . . 37809 1165 1 640 640 CD 37809 1165 2 pp pp NNP 37809 1165 3 . . . 37809 1166 1 YARROW YARROW NNP 37809 1166 2 , , , 37809 1166 3 H. H. NNP 37809 1166 4 C. C. NNP 37809 1166 5 1882 1882 CD 37809 1166 6 . . . 37809 1167 1 Checklist checklist NN 37809 1167 2 of of IN 37809 1167 3 North North NNP 37809 1167 4 American American NNP 37809 1167 5 Reptilia Reptilia NNP 37809 1167 6 and and CC 37809 1167 7 Batrachia Batrachia NNP 37809 1167 8 , , , 37809 1167 9 with with IN 37809 1167 10 catalogue catalogue NN 37809 1167 11 of of IN 37809 1167 12 specimens specimen NNS 37809 1167 13 in in IN 37809 1167 14 U.S. U.S. NNP 37809 1167 15 National National NNP 37809 1167 16 Museum Museum NNP 37809 1167 17 . . . 37809 1168 1 Bull Bull NNP 37809 1168 2 . . . 37809 1169 1 U.S. U.S. NNP 37809 1169 2 Natl Natl NNP 37809 1169 3 . . . 37809 1170 1 Mus Mus NNP 37809 1170 2 . . NNP 37809 1170 3 , , , 37809 1170 4 24:1 24:1 CD 37809 1170 5 - - SYM 37809 1170 6 249 249 CD 37809 1170 7 . . . 37809 1171 1 ZWEIFEL ZWEIFEL NNP 37809 1171 2 , , , 37809 1171 3 R. R. NNP 37809 1171 4 G. G. NNP 37809 1171 5 1960 1960 CD 37809 1171 6 . . . 37809 1172 1 Results result NNS 37809 1172 2 of of IN 37809 1172 3 the the DT 37809 1172 4 Puritan Puritan NNP 37809 1172 5 - - HYPH 37809 1172 6 American American NNP 37809 1172 7 Museum Museum NNP 37809 1172 8 of of IN 37809 1172 9 Natural Natural NNP 37809 1172 10 History History NNP 37809 1172 11 Expedition Expedition NNP 37809 1172 12 to to IN 37809 1172 13 Western Western NNP 37809 1172 14 Mexico Mexico NNP 37809 1172 15 . . . 37809 1173 1 9 9 CD 37809 1173 2 . . . 37809 1174 1 Herpetology herpetology NN 37809 1174 2 of of IN 37809 1174 3 the the DT 37809 1174 4 Tres Tres NNP 37809 1174 5 Marias Marias NNP 37809 1174 6 Islands Islands NNP 37809 1174 7 . . . 37809 1175 1 Bull Bull NNP 37809 1175 2 . . . 37809 1176 1 Amer amer NN 37809 1176 2 . . . 37809 1177 1 Mus mus NN 37809 1177 2 . . . 37809 1178 1 Nat Nat NNP 37809 1178 2 . . . 37809 1179 1 Hist Hist NNP 37809 1179 2 . . . 37809 1179 3 , , , 37809 1179 4 119:81 119:81 CD 37809 1179 5 - - SYM 37809 1179 6 128 128 CD 37809 1179 7 . . . 37809 1180 1 TRANSCRIBER TRANSCRIBER NNP 37809 1180 2 'S be VBZ 37809 1180 3 NOTES note NNS 37809 1180 4 Although although IN 37809 1180 5 _ _ NNP 37809 1180 6 Syrrhophus Syrrhophus NNP 37809 1180 7 marnocki marnocki NN 37809 1180 8 _ _ NNP 37809 1180 9 and and CC 37809 1180 10 _ _ NNP 37809 1180 11 Syrrhophus Syrrhophus NNP 37809 1180 12 marnockii marnockii NNS 37809 1180 13 _ _ IN 37809 1180 14 both both DT 37809 1180 15 appear appear VBP 37809 1180 16 in in IN 37809 1180 17 this this DT 37809 1180 18 text text NN 37809 1180 19 , , , 37809 1180 20 a a DT 37809 1180 21 literature literature NN 37809 1180 22 search search NN 37809 1180 23 shows show VBZ 37809 1180 24 that that IN 37809 1180 25 both both DT 37809 1180 26 spellings spelling NNS 37809 1180 27 have have VBP 37809 1180 28 been be VBN 37809 1180 29 used use VBN 37809 1180 30 and and CC 37809 1180 31 the the DT 37809 1180 32 two two CD 37809 1180 33 instances instance NNS 37809 1180 34 where where WRB 37809 1180 35 there there EX 37809 1180 36 is be VBZ 37809 1180 37 only only RB 37809 1180 38 one one CD 37809 1180 39 " " `` 37809 1180 40 i i PRP 37809 1180 41 " " '' 37809 1180 42 at at IN 37809 1180 43 the the DT 37809 1180 44 end end NN 37809 1180 45 are be VBP 37809 1180 46 in in IN 37809 1180 47 reference reference NN 37809 1180 48 to to TO 37809 1180 49 priviously priviously RB 37809 1180 50 published publish VBN 37809 1180 51 names name NNS 37809 1180 52 . . . 37809 1181 1 Therefore therefore RB 37809 1181 2 , , , 37809 1181 3 they -PRON- PRP 37809 1181 4 were be VBD 37809 1181 5 left leave VBN 37809 1181 6 as as IN 37809 1181 7 is be VBZ 37809 1181 8 . . . 37809 1182 1 With with IN 37809 1182 2 the the DT 37809 1182 3 exception exception NN 37809 1182 4 of of IN 37809 1182 5 the the DT 37809 1182 6 list list NN 37809 1182 7 below below RB 37809 1182 8 and and CC 37809 1182 9 a a DT 37809 1182 10 number number NN 37809 1182 11 of of IN 37809 1182 12 silent silent JJ 37809 1182 13 corrections correction NNS 37809 1182 14 , , , 37809 1182 15 the the DT 37809 1182 16 text text NN 37809 1182 17 presented present VBD 37809 1182 18 is be VBZ 37809 1182 19 that that DT 37809 1182 20 of of IN 37809 1182 21 the the DT 37809 1182 22 original original JJ 37809 1182 23 printed print VBN 37809 1182 24 version version NN 37809 1182 25 . . . 37809 1183 1 Typographical Typographical NNP 37809 1183 2 Corrections Corrections NNPS 37809 1183 3 Page Page NNP 37809 1183 4 Correction Correction NNP 37809 1183 5 = = SYM 37809 1183 6 = = SYM 37809 1183 7 = = SYM 37809 1183 8 = = SYM 37809 1183 9 = = SYM 37809 1183 10 = = SYM 37809 1183 11 = = SYM 37809 1183 12 = = SYM 37809 1183 13 = = SYM 37809 1183 14 = = SYM 37809 1183 15 = = SYM 37809 1183 16 = = SYM 37809 1183 17 = = SYM 37809 1183 18 = = SYM 37809 1183 19 = = SYM 37809 1183 20 = = SYM 37809 1183 21 = = SYM 37809 1183 22 = = SYM 37809 1183 23 = = SYM 37809 1183 24 = = SYM 37809 1183 25 = = SYM 37809 1183 26 = = SYM 37809 1183 27 = = SYM 37809 1183 28 = = SYM 37809 1183 29 = = NNS 37809 1183 30 = = SYM 37809 1183 31 3 3 CD 37809 1183 32 otherwse otherwse NN 37809 1183 33 = = SYM 37809 1183 34 > > XX 37809 1183 35 otherwise otherwise RB 37809 1183 36 5 5 CD 37809 1183 37 poltypic poltypic NN 37809 1183 38 = = NFP 37809 1183 39 > > XX 37809 1183 40 polytypic polytypic NNP 37809 1183 41 12 12 CD 37809 1183 42 interorbtal interorbtal NNP 37809 1183 43 = = SYM 37809 1183 44 > > XX 37809 1183 45 interorbital interorbital NNP 37809 1183 46 14 14 CD 37809 1183 47 neublosus neublosus NNP 37809 1183 48 = = SYM 37809 1183 49 > > XX 37809 1183 50 nebulosus nebulosus NN 37809 1183 51 16 16 CD 37809 1183 52 Cuidad Cuidad NNP 37809 1183 53 = = SYM 37809 1183 54 > > XX 37809 1183 55 Ciudad Ciudad NNP 37809 1183 56 16 16 CD 37809 1183 57 1946 1946 CD 37809 1183 58 - - SYM 37809 1183 59 170 170 CD 37809 1183 60 = = SYM 37809 1183 61 > > XX 37809 1183 62 1946:170 1946:170 CD 37809 1183 63 22 22 CD 37809 1183 64 rubrimacultaus rubrimacultau NNS 37809 1183 65 = = SYM 37809 1183 66 > > XX 37809 1183 67 rubrimaculatus rubrimaculatus NNP 37809 1183 68 27 27 CD 37809 1183 69 resemblence resemblence NN 37809 1183 70 = = NFP 37809 1183 71 > > XX 37809 1183 72 resemblance resemblance NN 37809 1183 73 Text Text NNP 37809 1183 74 Emphasis Emphasis NNP 37809 1183 75 _ _ NNP 37809 1183 76 Text Text NNP 37809 1183 77 _ _ NNP 37809 1183 78 - - HYPH 37809 1183 79 Italics Italics NNP 37809 1183 80 = = SYM 37809 1183 81 Text= text= NN 37809 1183 82 - - HYPH 37809 1183 83 Bold bold NN