id	sid	tid	token	lemma	pos
cord-005550-qrrdi667	1	1	key	key	NN
cord-005550-qrrdi667	1	2	:	:	:
cord-005550-qrrdi667	2	1	cord-005550-qrrdi667	cord-005550-qrrdi667	NNP
cord-005550-qrrdi667	2	2	authors	author	NNS
cord-005550-qrrdi667	2	3	:	:	:
cord-005550-qrrdi667	2	4	Mayer	Mayer	NNP
cord-005550-qrrdi667	2	5	,	,	,
cord-005550-qrrdi667	2	6	F	F	NNP
cord-005550-qrrdi667	2	7	;	;	:
cord-005550-qrrdi667	2	8	Brunner	Brunner	NNP
cord-005550-qrrdi667	2	9	,	,	,
cord-005550-qrrdi667	2	10	A	a	DT
cord-005550-qrrdi667	2	11	title	title	NN
cord-005550-qrrdi667	2	12	:	:	:
cord-005550-qrrdi667	2	13	Non	non	JJ
cord-005550-qrrdi667	2	14	-	-	JJ
cord-005550-qrrdi667	2	15	neutral	neutral	JJ
cord-005550-qrrdi667	2	16	evolution	evolution	NN
cord-005550-qrrdi667	2	17	of	of	IN
cord-005550-qrrdi667	2	18	the	the	DT
cord-005550-qrrdi667	2	19	major	major	JJ
cord-005550-qrrdi667	2	20	histocompatibility	histocompatibility	NN
cord-005550-qrrdi667	2	21	complex	complex	JJ
cord-005550-qrrdi667	2	22	class	class	NN
cord-005550-qrrdi667	2	23	II	II	NNP
cord-005550-qrrdi667	2	24	gene	gene	NN
cord-005550-qrrdi667	2	25	DRB1	DRB1	NNP
cord-005550-qrrdi667	2	26	in	in	IN
cord-005550-qrrdi667	2	27	the	the	DT
cord-005550-qrrdi667	2	28	sac	sac	NN
cord-005550-qrrdi667	2	29	-	-	HYPH
cord-005550-qrrdi667	2	30	winged	winged	JJ
cord-005550-qrrdi667	2	31	bat	bat	NN
cord-005550-qrrdi667	2	32	Saccopteryx	Saccopteryx	NNP
cord-005550-qrrdi667	2	33	bilineata	bilineata	NN
cord-005550-qrrdi667	2	34	date	date	NN
cord-005550-qrrdi667	2	35	:	:	:
cord-005550-qrrdi667	2	36	2007	2007	CD
cord-005550-qrrdi667	2	37	-	-	SYM
cord-005550-qrrdi667	2	38	05	05	CD
cord-005550-qrrdi667	2	39	-	-	HYPH
cord-005550-qrrdi667	2	40	23	23	CD
cord-005550-qrrdi667	2	41	journal	journal	NN
cord-005550-qrrdi667	2	42	:	:	:
cord-005550-qrrdi667	2	43	Heredity	heredity	NN
cord-005550-qrrdi667	2	44	(	(	-LRB-
cord-005550-qrrdi667	2	45	Edinb	Edinb	NNP
cord-005550-qrrdi667	2	46	)	)	-RRB-
cord-005550-qrrdi667	2	47	DOI	DOI	NNP
cord-005550-qrrdi667	2	48	:	:	:
cord-005550-qrrdi667	2	49	10.1038/	10.1038/	CD
cord-005550-qrrdi667	3	1	sj.hdy.6800989	sj.hdy.6800989	NNP
cord-005550-qrrdi667	3	2	sha	sha	NNP
cord-005550-qrrdi667	3	3	:	:	:
cord-005550-qrrdi667	3	4	c5ba328b74a58b850dd2224e4cc1ba1bcd58838a	c5ba328b74a58b850dd2224e4cc1ba1bcd58838a	NNP
cord-005550-qrrdi667	3	5	doc_id	doc_id	NNP
cord-005550-qrrdi667	3	6	:	:	:
cord-005550-qrrdi667	3	7	5550	5550	CD
cord-005550-qrrdi667	3	8	cord_uid	cord_uid	FW
cord-005550-qrrdi667	3	9	:	:	:
cord-005550-qrrdi667	4	1	qrrdi667	qrrdi667	NNP
cord-005550-qrrdi667	5	1	The	the	DT
cord-005550-qrrdi667	5	2	immune	immune	JJ
cord-005550-qrrdi667	5	3	genes	gene	NNS
cord-005550-qrrdi667	5	4	of	of	IN
cord-005550-qrrdi667	5	5	the	the	DT
cord-005550-qrrdi667	5	6	major	major	JJ
cord-005550-qrrdi667	5	7	histocompatibility	histocompatibility	NN
cord-005550-qrrdi667	5	8	complex	complex	NN
cord-005550-qrrdi667	5	9	(	(	-LRB-
cord-005550-qrrdi667	5	10	MHC	MHC	NNP
cord-005550-qrrdi667	5	11	)	)	-RRB-
cord-005550-qrrdi667	5	12	are	be	VBP
cord-005550-qrrdi667	5	13	classical	classical	JJ
cord-005550-qrrdi667	5	14	examples	example	NNS
cord-005550-qrrdi667	5	15	for	for	IN
cord-005550-qrrdi667	5	16	high	high	JJ
cord-005550-qrrdi667	5	17	levels	level	NNS
cord-005550-qrrdi667	5	18	of	of	IN
cord-005550-qrrdi667	5	19	genetic	genetic	JJ
cord-005550-qrrdi667	5	20	diversity	diversity	NN
cord-005550-qrrdi667	5	21	and	and	CC
cord-005550-qrrdi667	5	22	non	non	JJ
cord-005550-qrrdi667	5	23	-	-	JJ
cord-005550-qrrdi667	5	24	neutral	neutral	JJ
cord-005550-qrrdi667	5	25	evolution	evolution	NN
cord-005550-qrrdi667	5	26	.	.	.
cord-005550-qrrdi667	6	1	This	this	DT
cord-005550-qrrdi667	6	2	is	be	VBZ
cord-005550-qrrdi667	6	3	particularly	particularly	RB
cord-005550-qrrdi667	6	4	true	true	JJ
cord-005550-qrrdi667	6	5	for	for	IN
cord-005550-qrrdi667	6	6	the	the	DT
cord-005550-qrrdi667	6	7	regions	region	NNS
cord-005550-qrrdi667	6	8	containing	contain	VBG
cord-005550-qrrdi667	6	9	the	the	DT
cord-005550-qrrdi667	6	10	antigen	antigen	NN
cord-005550-qrrdi667	6	11	-	-	HYPH
cord-005550-qrrdi667	6	12	binding	bind	VBG
cord-005550-qrrdi667	6	13	sites	site	NNS
cord-005550-qrrdi667	6	14	as	as	IN
cord-005550-qrrdi667	6	15	,	,	,
cord-005550-qrrdi667	6	16	for	for	IN
cord-005550-qrrdi667	6	17	instance	instance	NN
cord-005550-qrrdi667	6	18	,	,	,
cord-005550-qrrdi667	6	19	in	in	IN
cord-005550-qrrdi667	6	20	the	the	DT
cord-005550-qrrdi667	6	21	exon	exon	NN
cord-005550-qrrdi667	6	22	2	2	CD
cord-005550-qrrdi667	6	23	of	of	IN
cord-005550-qrrdi667	6	24	the	the	DT
cord-005550-qrrdi667	6	25	MHC	MHC	NNP
cord-005550-qrrdi667	6	26	class	class	NN
cord-005550-qrrdi667	6	27	II	II	NNP
cord-005550-qrrdi667	6	28	gene	gene	NN
cord-005550-qrrdi667	6	29	DRB	DRB	NNP
cord-005550-qrrdi667	6	30	.	.	.
cord-005550-qrrdi667	7	1	We	-PRON-	PRP
cord-005550-qrrdi667	7	2	surveyed	survey	VBD
cord-005550-qrrdi667	7	3	,	,	,
cord-005550-qrrdi667	7	4	for	for	IN
cord-005550-qrrdi667	7	5	the	the	DT
cord-005550-qrrdi667	7	6	first	first	JJ
cord-005550-qrrdi667	7	7	time	time	NN
cord-005550-qrrdi667	7	8	in	in	IN
cord-005550-qrrdi667	7	9	the	the	DT
cord-005550-qrrdi667	7	10	order	order	NN
cord-005550-qrrdi667	7	11	Chiroptera	Chiroptera	NNP
cord-005550-qrrdi667	7	12	,	,	,
cord-005550-qrrdi667	7	13	the	the	DT
cord-005550-qrrdi667	7	14	genetic	genetic	JJ
cord-005550-qrrdi667	7	15	diversity	diversity	NN
cord-005550-qrrdi667	7	16	within	within	IN
cord-005550-qrrdi667	7	17	this	this	DT
cord-005550-qrrdi667	7	18	exon	exon	NN
cord-005550-qrrdi667	7	19	in	in	IN
cord-005550-qrrdi667	7	20	the	the	DT
cord-005550-qrrdi667	7	21	sac	sac	NN
cord-005550-qrrdi667	7	22	-	-	HYPH
cord-005550-qrrdi667	7	23	winged	winged	JJ
cord-005550-qrrdi667	7	24	bat	bat	NN
cord-005550-qrrdi667	7	25	Saccopteryx	Saccopteryx	NNP
cord-005550-qrrdi667	7	26	bilineata	bilineata	NN
cord-005550-qrrdi667	7	27	.	.	.
cord-005550-qrrdi667	8	1	We	-PRON-	PRP
cord-005550-qrrdi667	8	2	detected	detect	VBD
cord-005550-qrrdi667	8	3	11	11	CD
cord-005550-qrrdi667	8	4	alleles	allele	NNS
cord-005550-qrrdi667	8	5	among	among	IN
cord-005550-qrrdi667	8	6	85	85	CD
cord-005550-qrrdi667	8	7	bats	bat	NNS
cord-005550-qrrdi667	8	8	,	,	,
cord-005550-qrrdi667	8	9	of	of	IN
cord-005550-qrrdi667	8	10	which	which	WDT
cord-005550-qrrdi667	8	11	79	79	CD
cord-005550-qrrdi667	8	12	were	be	VBD
cord-005550-qrrdi667	8	13	sampled	sample	VBN
cord-005550-qrrdi667	8	14	in	in	IN
cord-005550-qrrdi667	8	15	one	one	CD
cord-005550-qrrdi667	8	16	population	population	NN
cord-005550-qrrdi667	8	17	.	.	.
cord-005550-qrrdi667	9	1	Pairwise	pairwise	JJ
cord-005550-qrrdi667	9	2	comparisons	comparison	NNS
cord-005550-qrrdi667	9	3	revealed	reveal	VBD
cord-005550-qrrdi667	9	4	that	that	IN
cord-005550-qrrdi667	9	5	interallelic	interallelic	NNP
cord-005550-qrrdi667	9	6	sequence	sequence	NN
cord-005550-qrrdi667	9	7	differences	difference	NNS
cord-005550-qrrdi667	9	8	ranged	range	VBD
cord-005550-qrrdi667	9	9	between	between	IN
cord-005550-qrrdi667	9	10	3	3	CD
cord-005550-qrrdi667	9	11	and	and	CC
cord-005550-qrrdi667	9	12	22	22	CD
cord-005550-qrrdi667	9	13	%	%	NN
cord-005550-qrrdi667	9	14	,	,	,
cord-005550-qrrdi667	9	15	although	although	IN
cord-005550-qrrdi667	9	16	nucleotide	nucleotide	JJ
cord-005550-qrrdi667	9	17	substitutions	substitution	NNS
cord-005550-qrrdi667	9	18	were	be	VBD
cord-005550-qrrdi667	9	19	not	not	RB
cord-005550-qrrdi667	9	20	evenly	evenly	RB
cord-005550-qrrdi667	9	21	distributed	distribute	VBN
cord-005550-qrrdi667	9	22	along	along	IN
cord-005550-qrrdi667	9	23	the	the	DT
cord-005550-qrrdi667	9	24	exon	exon	NN
cord-005550-qrrdi667	9	25	sequence	sequence	NN
cord-005550-qrrdi667	9	26	.	.	.
cord-005550-qrrdi667	10	1	This	this	DT
cord-005550-qrrdi667	10	2	was	be	VBD
cord-005550-qrrdi667	10	3	most	most	RBS
cord-005550-qrrdi667	10	4	probably	probably	RB
cord-005550-qrrdi667	10	5	the	the	DT
cord-005550-qrrdi667	10	6	result	result	NN
cord-005550-qrrdi667	10	7	of	of	IN
cord-005550-qrrdi667	10	8	intragenic	intragenic	JJ
cord-005550-qrrdi667	10	9	recombination	recombination	NN
cord-005550-qrrdi667	10	10	.	.	.
cord-005550-qrrdi667	11	1	High	high	JJ
cord-005550-qrrdi667	11	2	levels	level	NNS
cord-005550-qrrdi667	11	3	of	of	IN
cord-005550-qrrdi667	11	4	sequence	sequence	NN
cord-005550-qrrdi667	11	5	divergence	divergence	NN
cord-005550-qrrdi667	11	6	and	and	CC
cord-005550-qrrdi667	11	7	significantly	significantly	RB
cord-005550-qrrdi667	11	8	more	more	RBR
cord-005550-qrrdi667	11	9	nonsynonymous	nonsynonymous	JJ
cord-005550-qrrdi667	11	10	than	than	IN
cord-005550-qrrdi667	11	11	synonymous	synonymous	JJ
cord-005550-qrrdi667	11	12	substitutions	substitution	NNS
cord-005550-qrrdi667	11	13	(	(	-LRB-
cord-005550-qrrdi667	11	14	d(N)/d(S)>1	d(n)/d(s)>1	VB
cord-005550-qrrdi667	11	15	)	)	-RRB-
cord-005550-qrrdi667	11	16	suggest	suggest	VBP
cord-005550-qrrdi667	11	17	long	long	JJ
cord-005550-qrrdi667	11	18	-	-	HYPH
cord-005550-qrrdi667	11	19	term	term	NN
cord-005550-qrrdi667	11	20	balancing	balancing	NN
cord-005550-qrrdi667	11	21	selection	selection	NN
cord-005550-qrrdi667	11	22	.	.	.
cord-005550-qrrdi667	12	1	Thus	thus	RB
cord-005550-qrrdi667	12	2	,	,	,
cord-005550-qrrdi667	12	3	the	the	DT
cord-005550-qrrdi667	12	4	data	datum	NNS
cord-005550-qrrdi667	12	5	are	be	VBP
cord-005550-qrrdi667	12	6	consistent	consistent	JJ
cord-005550-qrrdi667	12	7	with	with	IN
cord-005550-qrrdi667	12	8	the	the	DT
cord-005550-qrrdi667	12	9	hypothesis	hypothesis	NN
cord-005550-qrrdi667	12	10	that	that	IN
cord-005550-qrrdi667	12	11	recombination	recombination	NN
cord-005550-qrrdi667	12	12	gives	give	VBZ
cord-005550-qrrdi667	12	13	rise	rise	NN
cord-005550-qrrdi667	12	14	to	to	IN
cord-005550-qrrdi667	12	15	new	new	JJ
cord-005550-qrrdi667	12	16	alleles	allele	NNS
cord-005550-qrrdi667	12	17	at	at	IN
cord-005550-qrrdi667	12	18	the	the	DT
cord-005550-qrrdi667	12	19	DRB	DRB	NNP
cord-005550-qrrdi667	12	20	locus	locus	NN
cord-005550-qrrdi667	12	21	of	of	IN
cord-005550-qrrdi667	12	22	the	the	DT
cord-005550-qrrdi667	12	23	sac	sac	NN
cord-005550-qrrdi667	12	24	-	-	HYPH
cord-005550-qrrdi667	12	25	winged	winged	JJ
cord-005550-qrrdi667	12	26	bat	bat	NN
cord-005550-qrrdi667	12	27	,	,	,
cord-005550-qrrdi667	12	28	and	and	CC
cord-005550-qrrdi667	12	29	these	these	DT
cord-005550-qrrdi667	12	30	are	be	VBP
cord-005550-qrrdi667	12	31	maintained	maintain	VBN
cord-005550-qrrdi667	12	32	in	in	IN
cord-005550-qrrdi667	12	33	the	the	DT
cord-005550-qrrdi667	12	34	population	population	NN
cord-005550-qrrdi667	12	35	through	through	IN
cord-005550-qrrdi667	12	36	balancing	balance	VBG
cord-005550-qrrdi667	12	37	selection	selection	NN
cord-005550-qrrdi667	12	38	.	.	.
cord-005550-qrrdi667	13	1	In	in	IN
cord-005550-qrrdi667	13	2	this	this	DT
cord-005550-qrrdi667	13	3	respect	respect	NN
cord-005550-qrrdi667	13	4	,	,	,
cord-005550-qrrdi667	13	5	the	the	DT
cord-005550-qrrdi667	13	6	sac	sac	NN
cord-005550-qrrdi667	13	7	-	-	HYPH
cord-005550-qrrdi667	13	8	winged	winged	JJ
cord-005550-qrrdi667	13	9	bat	bat	NN
cord-005550-qrrdi667	13	10	closely	closely	RB
cord-005550-qrrdi667	13	11	resembles	resemble	VBZ
cord-005550-qrrdi667	13	12	other	other	JJ
cord-005550-qrrdi667	13	13	mammalian	mammalian	JJ
cord-005550-qrrdi667	13	14	species	specie	NNS
cord-005550-qrrdi667	13	15	.	.	.
cord-005550-qrrdi667	14	1	The	the	DT
cord-005550-qrrdi667	14	2	major	major	JJ
cord-005550-qrrdi667	14	3	histocompatibility	histocompatibility	NN
cord-005550-qrrdi667	14	4	complex	complex	NN
cord-005550-qrrdi667	14	5	(	(	-LRB-
cord-005550-qrrdi667	14	6	MHC	MHC	NNP
cord-005550-qrrdi667	14	7	)	)	-RRB-
cord-005550-qrrdi667	14	8	plays	play	VBZ
cord-005550-qrrdi667	14	9	a	a	DT
cord-005550-qrrdi667	14	10	central	central	JJ
cord-005550-qrrdi667	14	11	role	role	NN
cord-005550-qrrdi667	14	12	in	in	IN
cord-005550-qrrdi667	14	13	the	the	DT
cord-005550-qrrdi667	14	14	vertebrate	vertebrate	JJ
cord-005550-qrrdi667	14	15	immune	immune	JJ
cord-005550-qrrdi667	14	16	system	system	NN
cord-005550-qrrdi667	14	17	.	.	.
cord-005550-qrrdi667	15	1	MHC	MHC	NNP
cord-005550-qrrdi667	15	2	genes	gene	NNS
cord-005550-qrrdi667	15	3	encode	encode	VBP
cord-005550-qrrdi667	15	4	two	two	CD
cord-005550-qrrdi667	15	5	major	major	JJ
cord-005550-qrrdi667	15	6	classes	class	NNS
cord-005550-qrrdi667	15	7	of	of	IN
cord-005550-qrrdi667	15	8	transmembrane	transmembrane	JJ
cord-005550-qrrdi667	15	9	glycoproteins	glycoprotein	NNS
cord-005550-qrrdi667	15	10	consisting	consist	VBG
cord-005550-qrrdi667	15	11	of	of	IN
cord-005550-qrrdi667	15	12	two	two	CD
cord-005550-qrrdi667	15	13	chains	chain	NNS
cord-005550-qrrdi667	15	14	,	,	,
cord-005550-qrrdi667	15	15	a	a	NN
cord-005550-qrrdi667	15	16	and	and	CC
cord-005550-qrrdi667	15	17	b.	b.	NNP
cord-005550-qrrdi667	15	18	MHC	MHC	NNP
cord-005550-qrrdi667	15	19	class	class	NN
cord-005550-qrrdi667	15	20	I	-PRON-	PRP
cord-005550-qrrdi667	15	21	genes	gene	NNS
cord-005550-qrrdi667	15	22	are	be	VBP
cord-005550-qrrdi667	15	23	expressed	express	VBN
cord-005550-qrrdi667	15	24	in	in	IN
cord-005550-qrrdi667	15	25	almost	almost	RB
cord-005550-qrrdi667	15	26	all	all	DT
cord-005550-qrrdi667	15	27	nucleated	nucleated	JJ
cord-005550-qrrdi667	15	28	somatic	somatic	JJ
cord-005550-qrrdi667	15	29	cells	cell	NNS
cord-005550-qrrdi667	15	30	,	,	,
cord-005550-qrrdi667	15	31	whereas	whereas	IN
cord-005550-qrrdi667	15	32	MHC	MHC	NNP
cord-005550-qrrdi667	15	33	class	class	NN
cord-005550-qrrdi667	15	34	II	II	NNP
cord-005550-qrrdi667	15	35	genes	gene	NNS
cord-005550-qrrdi667	15	36	are	be	VBP
cord-005550-qrrdi667	15	37	confined	confine	VBN
cord-005550-qrrdi667	15	38	to	to	IN
cord-005550-qrrdi667	15	39	antigen	antigen	NN
cord-005550-qrrdi667	15	40	-	-	HYPH
cord-005550-qrrdi667	15	41	presenting	present	VBG
cord-005550-qrrdi667	15	42	cells	cell	NNS
cord-005550-qrrdi667	15	43	of	of	IN
cord-005550-qrrdi667	15	44	the	the	DT
cord-005550-qrrdi667	15	45	immune	immune	JJ
cord-005550-qrrdi667	15	46	system	system	NN
cord-005550-qrrdi667	15	47	(	(	-LRB-
cord-005550-qrrdi667	15	48	Klein	Klein	NNP
cord-005550-qrrdi667	15	49	,	,	,
cord-005550-qrrdi667	15	50	1986	1986	CD
cord-005550-qrrdi667	15	51	)	)	-RRB-
cord-005550-qrrdi667	15	52	.	.	.
cord-005550-qrrdi667	16	1	Although	although	IN
cord-005550-qrrdi667	16	2	this	this	DT
cord-005550-qrrdi667	16	3	general	general	JJ
cord-005550-qrrdi667	16	4	function	function	NN
cord-005550-qrrdi667	16	5	of	of	IN
cord-005550-qrrdi667	16	6	the	the	DT
cord-005550-qrrdi667	16	7	MHC	MHC	NNP
cord-005550-qrrdi667	16	8	is	be	VBZ
cord-005550-qrrdi667	16	9	conserved	conserve	VBN
cord-005550-qrrdi667	16	10	among	among	IN
cord-005550-qrrdi667	16	11	vertebrates	vertebrate	NNS
cord-005550-qrrdi667	16	12	,	,	,
cord-005550-qrrdi667	16	13	the	the	DT
cord-005550-qrrdi667	16	14	genetic	genetic	JJ
cord-005550-qrrdi667	16	15	architecture	architecture	NN
cord-005550-qrrdi667	16	16	of	of	IN
cord-005550-qrrdi667	16	17	the	the	DT
cord-005550-qrrdi667	16	18	MHC	MHC	NNP
cord-005550-qrrdi667	16	19	can	can	MD
cord-005550-qrrdi667	16	20	differ	differ	VB
cord-005550-qrrdi667	16	21	within	within	IN
cord-005550-qrrdi667	16	22	and	and	CC
cord-005550-qrrdi667	16	23	between	between	IN
cord-005550-qrrdi667	16	24	vertebrate	vertebrate	JJ
cord-005550-qrrdi667	16	25	classes	class	NNS
cord-005550-qrrdi667	16	26	(	(	-LRB-
cord-005550-qrrdi667	16	27	recently	recently	RB
cord-005550-qrrdi667	16	28	reviewed	review	VBN
cord-005550-qrrdi667	16	29	in	in	IN
cord-005550-qrrdi667	16	30	Kelley	Kelley	NNP
cord-005550-qrrdi667	16	31	et	et	NNP
cord-005550-qrrdi667	16	32	al	al	NNP
cord-005550-qrrdi667	16	33	.	.	NNP
cord-005550-qrrdi667	16	34	,	,	,
cord-005550-qrrdi667	16	35	2005	2005	CD
cord-005550-qrrdi667	16	36	)	)	-RRB-
cord-005550-qrrdi667	16	37	.	.	.
cord-005550-qrrdi667	17	1	Gene	gene	NN
cord-005550-qrrdi667	17	2	duplication	duplication	NN
cord-005550-qrrdi667	17	3	occurred	occur	VBD
cord-005550-qrrdi667	17	4	frequently	frequently	RB
cord-005550-qrrdi667	17	5	,	,	,
cord-005550-qrrdi667	17	6	and	and	CC
cord-005550-qrrdi667	17	7	thus	thus	RB
cord-005550-qrrdi667	17	8	the	the	DT
cord-005550-qrrdi667	17	9	number	number	NN
cord-005550-qrrdi667	17	10	of	of	IN
cord-005550-qrrdi667	17	11	classes	class	NNS
cord-005550-qrrdi667	17	12	I	-PRON-	PRP
cord-005550-qrrdi667	17	13	and	and	CC
cord-005550-qrrdi667	17	14	II	II	NNP
cord-005550-qrrdi667	17	15	loci	locus	NNS
cord-005550-qrrdi667	17	16	can	can	MD
cord-005550-qrrdi667	17	17	substantially	substantially	RB
cord-005550-qrrdi667	17	18	vary	vary	VB
cord-005550-qrrdi667	17	19	between	between	IN
cord-005550-qrrdi667	17	20	species	specie	NNS
cord-005550-qrrdi667	17	21	(	(	-LRB-
cord-005550-qrrdi667	17	22	Kelley	Kelley	NNP
cord-005550-qrrdi667	17	23	et	et	NNP
cord-005550-qrrdi667	17	24	al	al	NNP
cord-005550-qrrdi667	17	25	.	.	NNP
cord-005550-qrrdi667	17	26	,	,	,
cord-005550-qrrdi667	17	27	2005	2005	CD
cord-005550-qrrdi667	17	28	;	;	:
cord-005550-qrrdi667	17	29	Nei	Nei	NNP
cord-005550-qrrdi667	17	30	and	and	CC
cord-005550-qrrdi667	17	31	Rooney	Rooney	NNP
cord-005550-qrrdi667	17	32	,	,	,
cord-005550-qrrdi667	17	33	2005	2005	CD
cord-005550-qrrdi667	17	34	)	)	-RRB-
cord-005550-qrrdi667	17	35	.	.	.
cord-005550-qrrdi667	18	1	In	in	IN
cord-005550-qrrdi667	18	2	mammals	mammal	NNS
cord-005550-qrrdi667	18	3	,	,	,
cord-005550-qrrdi667	18	4	the	the	DT
cord-005550-qrrdi667	18	5	genetic	genetic	JJ
cord-005550-qrrdi667	18	6	diversity	diversity	NN
cord-005550-qrrdi667	18	7	at	at	IN
cord-005550-qrrdi667	18	8	various	various	JJ
cord-005550-qrrdi667	18	9	MHC	MHC	NNP
cord-005550-qrrdi667	18	10	loci	locus	NNS
cord-005550-qrrdi667	18	11	has	have	VBZ
cord-005550-qrrdi667	18	12	been	be	VBN
cord-005550-qrrdi667	18	13	studied	study	VBN
cord-005550-qrrdi667	18	14	in	in	IN
cord-005550-qrrdi667	18	15	several	several	JJ
cord-005550-qrrdi667	18	16	species	specie	NNS
cord-005550-qrrdi667	18	17	of	of	IN
cord-005550-qrrdi667	18	18	all	all	DT
cord-005550-qrrdi667	18	19	major	major	JJ
cord-005550-qrrdi667	18	20	mammalian	mammalian	JJ
cord-005550-qrrdi667	18	21	orders	order	NNS
cord-005550-qrrdi667	18	22	except	except	IN
cord-005550-qrrdi667	18	23	for	for	IN
cord-005550-qrrdi667	18	24	the	the	DT
cord-005550-qrrdi667	18	25	second	second	RB
cord-005550-qrrdi667	18	26	largest	large	JJS
cord-005550-qrrdi667	18	27	order	order	NN
cord-005550-qrrdi667	18	28	Chiroptera	Chiroptera	NNP
cord-005550-qrrdi667	18	29	that	that	WDT
cord-005550-qrrdi667	18	30	comprise	comprise	VBP
cord-005550-qrrdi667	18	31	more	more	JJR
cord-005550-qrrdi667	18	32	than	than	IN
cord-005550-qrrdi667	18	33	1000	1000	CD
cord-005550-qrrdi667	18	34	species	specie	NNS
cord-005550-qrrdi667	18	35	worldwide	worldwide	RB
cord-005550-qrrdi667	18	36	(	(	-LRB-
cord-005550-qrrdi667	18	37	Simmons	Simmons	NNP
cord-005550-qrrdi667	18	38	,	,	,
cord-005550-qrrdi667	18	39	2006	2006	CD
cord-005550-qrrdi667	18	40	)	)	-RRB-
cord-005550-qrrdi667	18	41	.	.	.
cord-005550-qrrdi667	19	1	Bats	bat	NNS
cord-005550-qrrdi667	19	2	and	and	CC
cord-005550-qrrdi667	19	3	flying	fly	VBG
cord-005550-qrrdi667	19	4	foxes	fox	NNS
cord-005550-qrrdi667	19	5	are	be	VBP
cord-005550-qrrdi667	19	6	of	of	IN
cord-005550-qrrdi667	19	7	particular	particular	JJ
cord-005550-qrrdi667	19	8	immunological	immunological	JJ
cord-005550-qrrdi667	19	9	interest	interest	NN
cord-005550-qrrdi667	19	10	,	,	,
cord-005550-qrrdi667	19	11	because	because	IN
cord-005550-qrrdi667	19	12	they	-PRON-	PRP
cord-005550-qrrdi667	19	13	are	be	VBP
cord-005550-qrrdi667	19	14	hosts	host	NNS
cord-005550-qrrdi667	19	15	of	of	IN
cord-005550-qrrdi667	19	16	viruses	virus	NNS
cord-005550-qrrdi667	19	17	pathogenic	pathogenic	JJ
cord-005550-qrrdi667	19	18	to	to	IN
cord-005550-qrrdi667	19	19	humans	human	NNS
cord-005550-qrrdi667	19	20	including	include	VBG
cord-005550-qrrdi667	19	21	rabies	rabie	NNS
cord-005550-qrrdi667	20	1	(	(	-LRB-
cord-005550-qrrdi667	20	2	Fooks	Fooks	NNP
cord-005550-qrrdi667	20	3	et	et	NNP
cord-005550-qrrdi667	20	4	al	al	NNP
cord-005550-qrrdi667	20	5	.	.	NNP
cord-005550-qrrdi667	20	6	,	,	,
cord-005550-qrrdi667	20	7	2003	2003	CD
cord-005550-qrrdi667	20	8	;	;	:
cord-005550-qrrdi667	20	9	Cliquet	Cliquet	NNP
cord-005550-qrrdi667	20	10	and	and	CC
cord-005550-qrrdi667	20	11	Picard	Picard	NNP
cord-005550-qrrdi667	20	12	-	-	HYPH
cord-005550-qrrdi667	20	13	Meyer	Meyer	NNP
cord-005550-qrrdi667	20	14	,	,	,
cord-005550-qrrdi667	20	15	2004	2004	CD
cord-005550-qrrdi667	20	16	)	)	-RRB-
cord-005550-qrrdi667	20	17	,	,	,
cord-005550-qrrdi667	20	18	Ebola	Ebola	NNP
cord-005550-qrrdi667	20	19	(	(	-LRB-
cord-005550-qrrdi667	20	20	Leroy	Leroy	NNP
cord-005550-qrrdi667	20	21	et	et	NNP
cord-005550-qrrdi667	20	22	al	al	NNP
cord-005550-qrrdi667	20	23	.	.	NNP
cord-005550-qrrdi667	20	24	,	,	,
cord-005550-qrrdi667	20	25	2005	2005	CD
cord-005550-qrrdi667	20	26	)	)	-RRB-
cord-005550-qrrdi667	20	27	,	,	,
cord-005550-qrrdi667	20	28	SARS	SARS	NNP
cord-005550-qrrdi667	20	29	-	-	HYPH
cord-005550-qrrdi667	20	30	like	like	JJ
cord-005550-qrrdi667	20	31	viruses	virus	NNS
cord-005550-qrrdi667	21	1	(	(	-LRB-
cord-005550-qrrdi667	21	2	Li	Li	NNP
cord-005550-qrrdi667	21	3	et	et	NNP
cord-005550-qrrdi667	21	4	al	al	NNP
cord-005550-qrrdi667	21	5	.	.	NNP
cord-005550-qrrdi667	21	6	,	,	,
cord-005550-qrrdi667	21	7	2005	2005	CD
cord-005550-qrrdi667	21	8	)	)	-RRB-
cord-005550-qrrdi667	21	9	,	,	,
cord-005550-qrrdi667	21	10	and	and	CC
cord-005550-qrrdi667	21	11	Nipah	Nipah	NNP
cord-005550-qrrdi667	21	12	and	and	CC
cord-005550-qrrdi667	21	13	Hendra	Hendra	NNP
cord-005550-qrrdi667	21	14	viruses	virus	NNS
cord-005550-qrrdi667	22	1	(	(	-LRB-
cord-005550-qrrdi667	22	2	Wong	Wong	NNP
cord-005550-qrrdi667	22	3	et	et	NNP
cord-005550-qrrdi667	22	4	al	al	NNP
cord-005550-qrrdi667	22	5	.	.	NNP
cord-005550-qrrdi667	22	6	,	,	,
cord-005550-qrrdi667	22	7	2007	2007	CD
cord-005550-qrrdi667	22	8	)	)	-RRB-
cord-005550-qrrdi667	22	9	.	.	.
cord-005550-qrrdi667	23	1	MHC	MHC	NNP
cord-005550-qrrdi667	23	2	genes	gene	NNS
cord-005550-qrrdi667	23	3	are	be	VBP
cord-005550-qrrdi667	23	4	the	the	DT
cord-005550-qrrdi667	23	5	most	most	RBS
cord-005550-qrrdi667	23	6	variable	variable	JJ
cord-005550-qrrdi667	23	7	genes	gene	NNS
cord-005550-qrrdi667	23	8	described	describe	VBN
cord-005550-qrrdi667	23	9	in	in	IN
cord-005550-qrrdi667	23	10	vertebrates	vertebrate	NNS
cord-005550-qrrdi667	23	11	.	.	.
cord-005550-qrrdi667	24	1	Up	up	IN
cord-005550-qrrdi667	24	2	to	to	TO
cord-005550-qrrdi667	24	3	499	499	CD
cord-005550-qrrdi667	24	4	alleles	allele	NNS
cord-005550-qrrdi667	24	5	were	be	VBD
cord-005550-qrrdi667	24	6	described	describe	VBN
cord-005550-qrrdi667	24	7	at	at	IN
cord-005550-qrrdi667	24	8	the	the	DT
cord-005550-qrrdi667	24	9	human	human	JJ
cord-005550-qrrdi667	24	10	MHC	MHC	NNP
cord-005550-qrrdi667	24	11	locus	locus	NN
cord-005550-qrrdi667	24	12	HLA	HLA	NNP
cord-005550-qrrdi667	24	13	-	-	HYPH
cord-005550-qrrdi667	24	14	B	B	NNP
cord-005550-qrrdi667	24	15	(	(	-LRB-
cord-005550-qrrdi667	24	16	Garrigan	Garrigan	NNP
cord-005550-qrrdi667	24	17	and	and	CC
cord-005550-qrrdi667	24	18	Hedrick	Hedrick	NNP
cord-005550-qrrdi667	24	19	,	,	,
cord-005550-qrrdi667	24	20	2003	2003	CD
cord-005550-qrrdi667	24	21	)	)	-RRB-
cord-005550-qrrdi667	24	22	.	.	.
cord-005550-qrrdi667	25	1	Highest	high	JJS
cord-005550-qrrdi667	25	2	levels	level	NNS
cord-005550-qrrdi667	25	3	of	of	IN
cord-005550-qrrdi667	25	4	nucleotide	nucleotide	JJ
cord-005550-qrrdi667	25	5	diversity	diversity	NN
cord-005550-qrrdi667	25	6	are	be	VBP
cord-005550-qrrdi667	25	7	found	find	VBN
cord-005550-qrrdi667	25	8	in	in	IN
cord-005550-qrrdi667	25	9	regions	region	NNS
cord-005550-qrrdi667	25	10	containing	contain	VBG
cord-005550-qrrdi667	25	11	antigen	antigen	NN
cord-005550-qrrdi667	25	12	-	-	HYPH
cord-005550-qrrdi667	25	13	binding	bind	VBG
cord-005550-qrrdi667	25	14	sites	site	NNS
cord-005550-qrrdi667	25	15	(	(	-LRB-
cord-005550-qrrdi667	25	16	ABS	abs	NN
cord-005550-qrrdi667	25	17	)	)	-RRB-
cord-005550-qrrdi667	25	18	and	and	CC
cord-005550-qrrdi667	25	19	patterns	pattern	NNS
cord-005550-qrrdi667	25	20	of	of	IN
cord-005550-qrrdi667	25	21	nucleotide	nucleotide	JJ
cord-005550-qrrdi667	25	22	substitutions	substitution	NNS
cord-005550-qrrdi667	25	23	deviate	deviate	VBP
cord-005550-qrrdi667	25	24	from	from	IN
cord-005550-qrrdi667	25	25	neutral	neutral	JJ
cord-005550-qrrdi667	25	26	evolution	evolution	NN
cord-005550-qrrdi667	25	27	expectation	expectation	NN
cord-005550-qrrdi667	25	28	in	in	IN
cord-005550-qrrdi667	25	29	almost	almost	RB
cord-005550-qrrdi667	25	30	all	all	DT
cord-005550-qrrdi667	25	31	studies	study	NNS
cord-005550-qrrdi667	25	32	.	.	.
cord-005550-qrrdi667	26	1	Some	some	DT
cord-005550-qrrdi667	26	2	form	form	NN
cord-005550-qrrdi667	26	3	of	of	IN
cord-005550-qrrdi667	26	4	balancing	balance	VBG
cord-005550-qrrdi667	26	5	selection	selection	NN
cord-005550-qrrdi667	26	6	best	good	JJS
cord-005550-qrrdi667	26	7	explains	explain	VBZ
cord-005550-qrrdi667	26	8	the	the	DT
cord-005550-qrrdi667	26	9	enormous	enormous	JJ
cord-005550-qrrdi667	26	10	diversity	diversity	NN
cord-005550-qrrdi667	26	11	of	of	IN
cord-005550-qrrdi667	26	12	certain	certain	JJ
cord-005550-qrrdi667	26	13	MHC	MHC	NNP
cord-005550-qrrdi667	26	14	genes	gene	NNS
cord-005550-qrrdi667	26	15	(	(	-LRB-
cord-005550-qrrdi667	26	16	Hedrick	Hedrick	NNP
cord-005550-qrrdi667	26	17	and	and	CC
cord-005550-qrrdi667	26	18	Thomson	Thomson	NNP
cord-005550-qrrdi667	26	19	,	,	,
cord-005550-qrrdi667	26	20	1983	1983	CD
cord-005550-qrrdi667	26	21	)	)	-RRB-
cord-005550-qrrdi667	26	22	,	,	,
cord-005550-qrrdi667	26	23	and	and	CC
cord-005550-qrrdi667	26	24	two	two	CD
cord-005550-qrrdi667	26	25	major	major	JJ
cord-005550-qrrdi667	26	26	mutually	mutually	RB
cord-005550-qrrdi667	26	27	nonexclusive	nonexclusive	JJ
cord-005550-qrrdi667	26	28	mechanisms	mechanism	NNS
cord-005550-qrrdi667	26	29	of	of	IN
cord-005550-qrrdi667	26	30	pathogen	pathogen	NN
cord-005550-qrrdi667	26	31	-	-	HYPH
cord-005550-qrrdi667	26	32	driven	drive	VBN
cord-005550-qrrdi667	26	33	selection	selection	NN
cord-005550-qrrdi667	26	34	have	have	VBP
cord-005550-qrrdi667	26	35	been	be	VBN
cord-005550-qrrdi667	26	36	proposed	propose	VBN
cord-005550-qrrdi667	26	37	to	to	TO
cord-005550-qrrdi667	26	38	explain	explain	VB
cord-005550-qrrdi667	26	39	this	this	DT
cord-005550-qrrdi667	26	40	pattern	pattern	NN
cord-005550-qrrdi667	26	41	.	.	.
cord-005550-qrrdi667	27	1	First	first	RB
cord-005550-qrrdi667	27	2	,	,	,
cord-005550-qrrdi667	27	3	the	the	DT
cord-005550-qrrdi667	27	4	frequency	frequency	NN
cord-005550-qrrdi667	27	5	-	-	HYPH
cord-005550-qrrdi667	27	6	dependent	dependent	JJ
cord-005550-qrrdi667	27	7	selection	selection	NN
cord-005550-qrrdi667	27	8	hypothesis	hypothesis	NN
cord-005550-qrrdi667	27	9	or	or	CC
cord-005550-qrrdi667	27	10	the	the	DT
cord-005550-qrrdi667	27	11	rare	rare	JJ
cord-005550-qrrdi667	27	12	allele	allele	NN
cord-005550-qrrdi667	27	13	advantage	advantage	NN
cord-005550-qrrdi667	27	14	(	(	-LRB-
cord-005550-qrrdi667	27	15	Clarke	Clarke	NNP
cord-005550-qrrdi667	27	16	and	and	CC
cord-005550-qrrdi667	27	17	Kirby	Kirby	NNP
cord-005550-qrrdi667	27	18	,	,	,
cord-005550-qrrdi667	27	19	1966	1966	CD
cord-005550-qrrdi667	27	20	)	)	-RRB-
cord-005550-qrrdi667	27	21	predicts	predict	VBZ
cord-005550-qrrdi667	27	22	that	that	IN
cord-005550-qrrdi667	27	23	rare	rare	JJ
cord-005550-qrrdi667	27	24	alleles	allele	NNS
cord-005550-qrrdi667	27	25	can	can	MD
cord-005550-qrrdi667	27	26	have	have	VB
cord-005550-qrrdi667	27	27	a	a	DT
cord-005550-qrrdi667	27	28	selective	selective	JJ
cord-005550-qrrdi667	27	29	advantage	advantage	NN
cord-005550-qrrdi667	27	30	as	as	IN
cord-005550-qrrdi667	27	31	parasites	parasite	NNS
cord-005550-qrrdi667	27	32	will	will	MD
cord-005550-qrrdi667	27	33	adapt	adapt	VB
cord-005550-qrrdi667	27	34	to	to	IN
cord-005550-qrrdi667	27	35	common	common	JJ
cord-005550-qrrdi667	27	36	host	host	NN
cord-005550-qrrdi667	27	37	alleles	allele	NNS
cord-005550-qrrdi667	27	38	.	.	.
cord-005550-qrrdi667	28	1	Since	since	IN
cord-005550-qrrdi667	28	2	an	an	DT
cord-005550-qrrdi667	28	3	advantageous	advantageous	JJ
cord-005550-qrrdi667	28	4	allele	allele	NN
cord-005550-qrrdi667	28	5	will	will	MD
cord-005550-qrrdi667	28	6	increase	increase	VB
cord-005550-qrrdi667	28	7	in	in	IN
cord-005550-qrrdi667	28	8	frequency	frequency	NN
cord-005550-qrrdi667	28	9	,	,	,
cord-005550-qrrdi667	28	10	parasites	parasite	NNS
cord-005550-qrrdi667	28	11	resistant	resistant	JJ
cord-005550-qrrdi667	28	12	to	to	IN
cord-005550-qrrdi667	28	13	this	this	DT
cord-005550-qrrdi667	28	14	allele	allele	NN
cord-005550-qrrdi667	28	15	will	will	MD
cord-005550-qrrdi667	28	16	be	be	VB
cord-005550-qrrdi667	28	17	favoured	favour	VBN
cord-005550-qrrdi667	28	18	.	.	.
cord-005550-qrrdi667	29	1	Negative	negative	JJ
cord-005550-qrrdi667	29	2	frequency	frequency	NN
cord-005550-qrrdi667	29	3	-	-	HYPH
cord-005550-qrrdi667	29	4	dependent	dependent	JJ
cord-005550-qrrdi667	29	5	selection	selection	NN
cord-005550-qrrdi667	29	6	models	model	NNS
cord-005550-qrrdi667	29	7	of	of	IN
cord-005550-qrrdi667	29	8	host	host	NN
cord-005550-qrrdi667	29	9	-	-	HYPH
cord-005550-qrrdi667	29	10	parasite	parasite	NN
cord-005550-qrrdi667	29	11	coevolution	coevolution	NN
cord-005550-qrrdi667	29	12	showed	show	VBD
cord-005550-qrrdi667	29	13	that	that	IN
cord-005550-qrrdi667	29	14	a	a	DT
cord-005550-qrrdi667	29	15	large	large	JJ
cord-005550-qrrdi667	29	16	number	number	NN
cord-005550-qrrdi667	29	17	of	of	IN
cord-005550-qrrdi667	29	18	alleles	allele	NNS
cord-005550-qrrdi667	29	19	can	can	MD
cord-005550-qrrdi667	29	20	be	be	VB
cord-005550-qrrdi667	29	21	retained	retain	VBN
cord-005550-qrrdi667	29	22	(	(	-LRB-
cord-005550-qrrdi667	29	23	Borghans	Borghans	NNP
cord-005550-qrrdi667	29	24	et	et	FW
cord-005550-qrrdi667	29	25	al	al	NNP
cord-005550-qrrdi667	29	26	.	.	NNP
cord-005550-qrrdi667	29	27	,	,	,
cord-005550-qrrdi667	29	28	2004	2004	CD
cord-005550-qrrdi667	29	29	)	)	-RRB-
cord-005550-qrrdi667	29	30	.	.	.
cord-005550-qrrdi667	30	1	The	the	DT
cord-005550-qrrdi667	30	2	spatial	spatial	JJ
cord-005550-qrrdi667	30	3	and	and	CC
cord-005550-qrrdi667	30	4	rapid	rapid	JJ
cord-005550-qrrdi667	30	5	temporal	temporal	JJ
cord-005550-qrrdi667	30	6	variation	variation	NN
cord-005550-qrrdi667	30	7	in	in	IN
cord-005550-qrrdi667	30	8	frequency	frequency	NN
cord-005550-qrrdi667	30	9	of	of	IN
cord-005550-qrrdi667	30	10	MHC	MHC	NNP
cord-005550-qrrdi667	30	11	alleles	allele	NNS
cord-005550-qrrdi667	30	12	and	and	CC
cord-005550-qrrdi667	30	13	resistance	resistance	NN
cord-005550-qrrdi667	30	14	provides	provide	VBZ
cord-005550-qrrdi667	30	15	evidence	evidence	NN
cord-005550-qrrdi667	30	16	for	for	IN
cord-005550-qrrdi667	30	17	frequency	frequency	NN
cord-005550-qrrdi667	30	18	-	-	HYPH
cord-005550-qrrdi667	30	19	dependent	dependent	JJ
cord-005550-qrrdi667	30	20	selection	selection	NN
cord-005550-qrrdi667	30	21	(	(	-LRB-
cord-005550-qrrdi667	30	22	Hill	Hill	NNP
cord-005550-qrrdi667	30	23	et	et	FW
cord-005550-qrrdi667	30	24	al	al	NNP
cord-005550-qrrdi667	30	25	.	.	.
cord-005550-qrrdi667	31	1	,	,	,
cord-005550-qrrdi667	31	2	1994	1994	CD
cord-005550-qrrdi667	31	3	;	;	:
cord-005550-qrrdi667	31	4	Westerdahl	Westerdahl	NNP
cord-005550-qrrdi667	31	5	et	et	NNP
cord-005550-qrrdi667	31	6	al	al	NNP
cord-005550-qrrdi667	31	7	.	.	NNP
cord-005550-qrrdi667	31	8	,	,	,
cord-005550-qrrdi667	31	9	2004	2004	CD
cord-005550-qrrdi667	31	10	;	;	:
cord-005550-qrrdi667	31	11	Charbonnel	Charbonnel	NNP
cord-005550-qrrdi667	31	12	and	and	CC
cord-005550-qrrdi667	31	13	Pemberton	Pemberton	NNP
cord-005550-qrrdi667	31	14	,	,	,
cord-005550-qrrdi667	31	15	2005	2005	CD
cord-005550-qrrdi667	31	16	)	)	-RRB-
cord-005550-qrrdi667	31	17	.	.	.
cord-005550-qrrdi667	32	1	Second	second	RB
cord-005550-qrrdi667	32	2	,	,	,
cord-005550-qrrdi667	32	3	the	the	DT
cord-005550-qrrdi667	32	4	overdominance	overdominance	NN
cord-005550-qrrdi667	32	5	or	or	CC
cord-005550-qrrdi667	32	6	heterozygote	heterozygote	NN
cord-005550-qrrdi667	32	7	advantage	advantage	NN
cord-005550-qrrdi667	32	8	hypothesis	hypothesis	NN
cord-005550-qrrdi667	32	9	(	(	-LRB-
cord-005550-qrrdi667	32	10	Doherty	Doherty	NNP
cord-005550-qrrdi667	32	11	and	and	CC
cord-005550-qrrdi667	32	12	Zinkernagel	Zinkernagel	NNP
cord-005550-qrrdi667	32	13	,	,	,
cord-005550-qrrdi667	32	14	1975	1975	CD
cord-005550-qrrdi667	32	15	)	)	-RRB-
cord-005550-qrrdi667	32	16	assumes	assume	VBZ
cord-005550-qrrdi667	32	17	that	that	IN
cord-005550-qrrdi667	32	18	heterozygous	heterozygous	JJ
cord-005550-qrrdi667	32	19	individuals	individual	NNS
cord-005550-qrrdi667	32	20	have	have	VBP
cord-005550-qrrdi667	32	21	a	a	DT
cord-005550-qrrdi667	32	22	higher	high	JJR
cord-005550-qrrdi667	32	23	average	average	JJ
cord-005550-qrrdi667	32	24	fitness	fitness	NN
cord-005550-qrrdi667	32	25	,	,	,
cord-005550-qrrdi667	32	26	because	because	IN
cord-005550-qrrdi667	32	27	individuals	individual	NNS
cord-005550-qrrdi667	32	28	being	be	VBG
cord-005550-qrrdi667	32	29	heterozygous	heterozygous	JJ
cord-005550-qrrdi667	32	30	at	at	IN
cord-005550-qrrdi667	32	31	the	the	DT
cord-005550-qrrdi667	32	32	antigen	antigen	NN
cord-005550-qrrdi667	32	33	-	-	HYPH
cord-005550-qrrdi667	32	34	binding	bind	VBG
cord-005550-qrrdi667	32	35	region	region	NN
cord-005550-qrrdi667	32	36	can	can	MD
cord-005550-qrrdi667	32	37	present	present	VB
cord-005550-qrrdi667	32	38	a	a	DT
cord-005550-qrrdi667	32	39	wider	wide	JJR
cord-005550-qrrdi667	32	40	array	array	NN
cord-005550-qrrdi667	32	41	of	of	IN
cord-005550-qrrdi667	32	42	antigens	antigen	NNS
cord-005550-qrrdi667	32	43	than	than	IN
cord-005550-qrrdi667	32	44	homozygous	homozygous	JJ
cord-005550-qrrdi667	32	45	individuals	individual	NNS
cord-005550-qrrdi667	32	46	.	.	.
cord-005550-qrrdi667	33	1	Therefore	therefore	RB
cord-005550-qrrdi667	33	2	,	,	,
cord-005550-qrrdi667	33	3	heterozyotes	heterozyote	NNS
cord-005550-qrrdi667	33	4	are	be	VBP
cord-005550-qrrdi667	33	5	expected	expect	VBN
cord-005550-qrrdi667	33	6	to	to	TO
cord-005550-qrrdi667	33	7	be	be	VB
cord-005550-qrrdi667	33	8	more	more	RBR
cord-005550-qrrdi667	33	9	resistant	resistant	JJ
cord-005550-qrrdi667	33	10	against	against	IN
cord-005550-qrrdi667	33	11	pathogens	pathogen	NNS
cord-005550-qrrdi667	33	12	than	than	IN
cord-005550-qrrdi667	33	13	homozygotes	homozygote	NNS
cord-005550-qrrdi667	33	14	.	.	.
cord-005550-qrrdi667	34	1	An	an	DT
cord-005550-qrrdi667	34	2	overdominance	overdominance	NN
cord-005550-qrrdi667	34	3	effect	effect	NN
cord-005550-qrrdi667	34	4	was	be	VBD
cord-005550-qrrdi667	34	5	shown	show	VBN
cord-005550-qrrdi667	34	6	in	in	IN
cord-005550-qrrdi667	34	7	mice	mouse	NNS
cord-005550-qrrdi667	34	8	that	that	WDT
cord-005550-qrrdi667	34	9	were	be	VBD
cord-005550-qrrdi667	34	10	coinfected	coinfecte	VBN
cord-005550-qrrdi667	34	11	with	with	IN
cord-005550-qrrdi667	34	12	Salmonella	Salmonella	NNP
cord-005550-qrrdi667	34	13	enterica	enterica	NN
cord-005550-qrrdi667	34	14	and	and	CC
cord-005550-qrrdi667	34	15	Theiler	Theiler	NNP
cord-005550-qrrdi667	34	16	's	's	POS
cord-005550-qrrdi667	34	17	murine	murine	JJ
cord-005550-qrrdi667	34	18	encephalomyelitis	encephalomyelitis	NN
cord-005550-qrrdi667	34	19	virus	virus	NN
cord-005550-qrrdi667	34	20	(	(	-LRB-
cord-005550-qrrdi667	34	21	McClelland	McClelland	NNP
cord-005550-qrrdi667	34	22	et	et	NNP
cord-005550-qrrdi667	34	23	al	al	NNP
cord-005550-qrrdi667	34	24	.	.	NNP
cord-005550-qrrdi667	34	25	,	,	,
cord-005550-qrrdi667	34	26	2003	2003	CD
cord-005550-qrrdi667	34	27	)	)	-RRB-
cord-005550-qrrdi667	34	28	.	.	.
cord-005550-qrrdi667	35	1	Distinguishing	distinguish	VBG
cord-005550-qrrdi667	35	2	between	between	IN
cord-005550-qrrdi667	35	3	the	the	DT
cord-005550-qrrdi667	35	4	frequencydependent	frequencydependent	NN
cord-005550-qrrdi667	35	5	selection	selection	NN
cord-005550-qrrdi667	35	6	and	and	CC
cord-005550-qrrdi667	35	7	the	the	DT
cord-005550-qrrdi667	35	8	heterozygote	heterozygote	NN
cord-005550-qrrdi667	35	9	advantage	advantage	NN
cord-005550-qrrdi667	35	10	hypothesis	hypothesis	NN
cord-005550-qrrdi667	35	11	is	be	VBZ
cord-005550-qrrdi667	35	12	difficult	difficult	JJ
cord-005550-qrrdi667	35	13	,	,	,
cord-005550-qrrdi667	35	14	because	because	IN
cord-005550-qrrdi667	35	15	both	both	DT
cord-005550-qrrdi667	35	16	models	model	NNS
cord-005550-qrrdi667	35	17	require	require	VBP
cord-005550-qrrdi667	35	18	a	a	DT
cord-005550-qrrdi667	35	19	correlation	correlation	NN
cord-005550-qrrdi667	35	20	between	between	IN
cord-005550-qrrdi667	35	21	parasites	parasite	NNS
cord-005550-qrrdi667	35	22	and	and	CC
cord-005550-qrrdi667	35	23	particular	particular	JJ
cord-005550-qrrdi667	35	24	MHC	MHC	NNP
cord-005550-qrrdi667	35	25	alleles	allele	NNS
cord-005550-qrrdi667	35	26	.	.	.
cord-005550-qrrdi667	36	1	Meanwhile	meanwhile	RB
cord-005550-qrrdi667	36	2	,	,	,
cord-005550-qrrdi667	36	3	such	such	JJ
cord-005550-qrrdi667	36	4	relationships	relationship	NNS
cord-005550-qrrdi667	36	5	were	be	VBD
cord-005550-qrrdi667	36	6	found	find	VBN
cord-005550-qrrdi667	36	7	in	in	IN
cord-005550-qrrdi667	36	8	several	several	JJ
cord-005550-qrrdi667	36	9	species	specie	NNS
cord-005550-qrrdi667	36	10	including	include	VBG
cord-005550-qrrdi667	36	11	many	many	JJ
cord-005550-qrrdi667	36	12	non	non	JJ
cord-005550-qrrdi667	36	13	-	-	JJ
cord-005550-qrrdi667	36	14	model	model	JJ
cord-005550-qrrdi667	36	15	organisms	organism	NNS
cord-005550-qrrdi667	36	16	(	(	-LRB-
cord-005550-qrrdi667	36	17	Table	table	NN
cord-005550-qrrdi667	36	18	1	1	CD
cord-005550-qrrdi667	36	19	in	in	IN
cord-005550-qrrdi667	36	20	Sommer	Sommer	NNP
cord-005550-qrrdi667	36	21	,	,	,
cord-005550-qrrdi667	36	22	2005	2005	CD
cord-005550-qrrdi667	36	23	)	)	-RRB-
cord-005550-qrrdi667	36	24	.	.	.
cord-005550-qrrdi667	37	1	The	the	DT
cord-005550-qrrdi667	37	2	evolutionary	evolutionary	JJ
cord-005550-qrrdi667	37	3	ecology	ecology	NN
cord-005550-qrrdi667	37	4	of	of	IN
cord-005550-qrrdi667	37	5	the	the	DT
cord-005550-qrrdi667	37	6	MHC	MHC	NNP
cord-005550-qrrdi667	37	7	was	be	VBD
cord-005550-qrrdi667	37	8	recently	recently	RB
cord-005550-qrrdi667	37	9	summarized	summarize	VBN
cord-005550-qrrdi667	37	10	in	in	IN
cord-005550-qrrdi667	37	11	several	several	JJ
cord-005550-qrrdi667	37	12	reviews	review	NNS
cord-005550-qrrdi667	37	13	(	(	-LRB-
cord-005550-qrrdi667	37	14	Bernatchez	Bernatchez	NNP
cord-005550-qrrdi667	37	15	and	and	CC
cord-005550-qrrdi667	37	16	Landry	Landry	NNP
cord-005550-qrrdi667	37	17	,	,	,
cord-005550-qrrdi667	37	18	2003	2003	CD
cord-005550-qrrdi667	37	19	;	;	:
cord-005550-qrrdi667	37	20	Garrigan	Garrigan	NNP
cord-005550-qrrdi667	37	21	and	and	CC
cord-005550-qrrdi667	37	22	Hedrick	Hedrick	NNP
cord-005550-qrrdi667	37	23	,	,	,
cord-005550-qrrdi667	37	24	2003	2003	CD
cord-005550-qrrdi667	37	25	;	;	:
cord-005550-qrrdi667	37	26	Sommer	Sommer	NNP
cord-005550-qrrdi667	37	27	,	,	,
cord-005550-qrrdi667	37	28	2005	2005	CD
cord-005550-qrrdi667	37	29	;	;	:
cord-005550-qrrdi667	37	30	Piertney	Piertney	NNP
cord-005550-qrrdi667	37	31	and	and	CC
cord-005550-qrrdi667	37	32	Oliver	Oliver	NNP
cord-005550-qrrdi667	37	33	,	,	,
cord-005550-qrrdi667	37	34	2006	2006	CD
cord-005550-qrrdi667	37	35	)	)	-RRB-
cord-005550-qrrdi667	37	36	.	.	.
cord-005550-qrrdi667	38	1	We	-PRON-	PRP
cord-005550-qrrdi667	38	2	analysed	analyse	VBD
cord-005550-qrrdi667	38	3	the	the	DT
cord-005550-qrrdi667	38	4	antigen	antigen	NN
cord-005550-qrrdi667	38	5	-	-	HYPH
cord-005550-qrrdi667	38	6	binding	bind	VBG
cord-005550-qrrdi667	38	7	region	region	NN
cord-005550-qrrdi667	38	8	of	of	IN
cord-005550-qrrdi667	38	9	the	the	DT
cord-005550-qrrdi667	38	10	MHC	MHC	NNP
cord-005550-qrrdi667	38	11	class	class	NN
cord-005550-qrrdi667	38	12	II	II	NNP
cord-005550-qrrdi667	38	13	gene	gene	NN
cord-005550-qrrdi667	38	14	DRB	DRB	NNP
cord-005550-qrrdi667	38	15	of	of	IN
cord-005550-qrrdi667	38	16	the	the	DT
cord-005550-qrrdi667	38	17	sac	sac	NN
cord-005550-qrrdi667	38	18	-	-	HYPH
cord-005550-qrrdi667	38	19	winged	winged	JJ
cord-005550-qrrdi667	38	20	bat	bat	NN
cord-005550-qrrdi667	38	21	Saccopteryx	Saccopteryx	NNP
cord-005550-qrrdi667	38	22	bilineata	bilineata	NN
cord-005550-qrrdi667	38	23	by	by	IN
cord-005550-qrrdi667	38	24	sequencing	sequence	VBG
cord-005550-qrrdi667	38	25	cloned	clone	VBN
cord-005550-qrrdi667	38	26	PCR	PCR	NNP
cord-005550-qrrdi667	38	27	products	product	NNS
cord-005550-qrrdi667	38	28	.	.	.
cord-005550-qrrdi667	39	1	We	-PRON-	PRP
cord-005550-qrrdi667	39	2	selected	select	VBD
cord-005550-qrrdi667	39	3	this	this	DT
cord-005550-qrrdi667	39	4	neotropical	neotropical	JJ
cord-005550-qrrdi667	39	5	,	,	,
cord-005550-qrrdi667	39	6	insectivorous	insectivorous	JJ
cord-005550-qrrdi667	39	7	bat	bat	NN
cord-005550-qrrdi667	39	8	species	specie	NNS
cord-005550-qrrdi667	39	9	,	,	,
cord-005550-qrrdi667	39	10	because	because	IN
cord-005550-qrrdi667	39	11	completely	completely	RB
cord-005550-qrrdi667	39	12	sampled	sample	VBN
cord-005550-qrrdi667	39	13	families	family	NNS
cord-005550-qrrdi667	39	14	consisting	consist	VBG
cord-005550-qrrdi667	39	15	of	of	IN
cord-005550-qrrdi667	39	16	an	an	DT
cord-005550-qrrdi667	39	17	offspring	offspring	NN
cord-005550-qrrdi667	39	18	and	and	CC
cord-005550-qrrdi667	39	19	both	both	DT
cord-005550-qrrdi667	39	20	parents	parent	NNS
cord-005550-qrrdi667	39	21	were	be	VBD
cord-005550-qrrdi667	39	22	available	available	JJ
cord-005550-qrrdi667	39	23	(	(	-LRB-
cord-005550-qrrdi667	39	24	Heckel	Heckel	NNP
cord-005550-qrrdi667	39	25	and	and	CC
cord-005550-qrrdi667	39	26	von	von	NNP
cord-005550-qrrdi667	39	27	Helversen	Helversen	NNP
cord-005550-qrrdi667	39	28	,	,	,
cord-005550-qrrdi667	39	29	2003	2003	CD
cord-005550-qrrdi667	39	30	)	)	-RRB-
cord-005550-qrrdi667	39	31	that	that	WDT
cord-005550-qrrdi667	39	32	allowed	allow	VBD
cord-005550-qrrdi667	39	33	testing	test	VBG
cord-005550-qrrdi667	39	34	Mendelian	mendelian	JJ
cord-005550-qrrdi667	39	35	inheritance	inheritance	NN
cord-005550-qrrdi667	39	36	of	of	IN
cord-005550-qrrdi667	39	37	haplotypes	haplotype	NNS
cord-005550-qrrdi667	39	38	.	.	.
cord-005550-qrrdi667	40	1	In	in	IN
cord-005550-qrrdi667	40	2	this	this	DT
cord-005550-qrrdi667	40	3	study	study	NN
cord-005550-qrrdi667	40	4	,	,	,
cord-005550-qrrdi667	40	5	we	-PRON-	PRP
cord-005550-qrrdi667	40	6	describe	describe	VBP
cord-005550-qrrdi667	40	7	the	the	DT
cord-005550-qrrdi667	40	8	genetic	genetic	JJ
cord-005550-qrrdi667	40	9	diversity	diversity	NN
cord-005550-qrrdi667	40	10	within	within	IN
cord-005550-qrrdi667	40	11	exon	exon	NN
cord-005550-qrrdi667	40	12	2	2	CD
cord-005550-qrrdi667	40	13	of	of	IN
cord-005550-qrrdi667	40	14	the	the	DT
cord-005550-qrrdi667	40	15	MHC	MHC	NNP
cord-005550-qrrdi667	40	16	class	class	NN
cord-005550-qrrdi667	40	17	II	II	NNP
cord-005550-qrrdi667	40	18	gene	gene	NN
cord-005550-qrrdi667	40	19	DRB	DRB	NNP
cord-005550-qrrdi667	40	20	,	,	,
cord-005550-qrrdi667	40	21	and	and	CC
cord-005550-qrrdi667	40	22	discuss	discuss	VB
cord-005550-qrrdi667	40	23	the	the	DT
cord-005550-qrrdi667	40	24	role	role	NN
cord-005550-qrrdi667	40	25	of	of	IN
cord-005550-qrrdi667	40	26	recombination	recombination	NN
cord-005550-qrrdi667	40	27	and	and	CC
cord-005550-qrrdi667	40	28	selection	selection	NN
cord-005550-qrrdi667	40	29	in	in	IN
cord-005550-qrrdi667	40	30	generating	generate	VBG
cord-005550-qrrdi667	40	31	and	and	CC
cord-005550-qrrdi667	40	32	maintaining	maintain	VBG
cord-005550-qrrdi667	40	33	high	high	JJ
cord-005550-qrrdi667	40	34	levels	level	NNS
cord-005550-qrrdi667	40	35	of	of	IN
cord-005550-qrrdi667	40	36	genetic	genetic	JJ
cord-005550-qrrdi667	40	37	diversity	diversity	NN
cord-005550-qrrdi667	40	38	in	in	IN
cord-005550-qrrdi667	40	39	colonies	colony	NNS
cord-005550-qrrdi667	40	40	of	of	IN
cord-005550-qrrdi667	40	41	the	the	DT
cord-005550-qrrdi667	40	42	sac	sac	NN
cord-005550-qrrdi667	40	43	-	-	HYPH
cord-005550-qrrdi667	40	44	winged	winged	JJ
cord-005550-qrrdi667	40	45	bat	bat	NN
cord-005550-qrrdi667	40	46	.	.	.
cord-005550-qrrdi667	41	1	Bats	bat	NNS
cord-005550-qrrdi667	41	2	were	be	VBD
cord-005550-qrrdi667	41	3	caught	catch	VBN
cord-005550-qrrdi667	41	4	by	by	IN
cord-005550-qrrdi667	41	5	mist	mist	NN
cord-005550-qrrdi667	41	6	netting	netting	NN
cord-005550-qrrdi667	41	7	at	at	IN
cord-005550-qrrdi667	41	8	the	the	DT
cord-005550-qrrdi667	41	9	biological	biological	JJ
cord-005550-qrrdi667	41	10	station	station	NN
cord-005550-qrrdi667	41	11	La	La	NNP
cord-005550-qrrdi667	41	12	Selva	Selva	NNP
cord-005550-qrrdi667	41	13	in	in	IN
cord-005550-qrrdi667	41	14	Costa	Costa	NNP
cord-005550-qrrdi667	41	15	Rica	Rica	NNP
cord-005550-qrrdi667	41	16	(	(	-LRB-
cord-005550-qrrdi667	41	17	10120	10120	CD
cord-005550-qrrdi667	41	18	0	0	CD
cord-005550-qrrdi667	41	19	N	n	CD
cord-005550-qrrdi667	41	20	,	,	,
cord-005550-qrrdi667	41	21	84110	84110	CD
cord-005550-qrrdi667	41	22	0	0	CD
cord-005550-qrrdi667	41	23	W	W	NNP
cord-005550-qrrdi667	41	24	,	,	,
cord-005550-qrrdi667	41	25	N	n	CD
cord-005550-qrrdi667	41	26	¼	¼	CD
cord-005550-qrrdi667	41	27	79	79	CD
cord-005550-qrrdi667	41	28	)	)	-RRB-
cord-005550-qrrdi667	41	29	,	,	,
cord-005550-qrrdi667	41	30	on	on	IN
cord-005550-qrrdi667	41	31	Barro	Barro	NNP
cord-005550-qrrdi667	41	32	Colorado	Colorado	NNP
cord-005550-qrrdi667	41	33	Island	Island	NNP
cord-005550-qrrdi667	41	34	(	(	-LRB-
cord-005550-qrrdi667	41	35	BCI	BCI	NNP
cord-005550-qrrdi667	41	36	)	)	-RRB-
cord-005550-qrrdi667	41	37	in	in	IN
cord-005550-qrrdi667	41	38	Panama	Panama	NNP
cord-005550-qrrdi667	41	39	(	(	-LRB-
cord-005550-qrrdi667	41	40	9109	9109	CD
cord-005550-qrrdi667	41	41	0	0	CD
cord-005550-qrrdi667	41	42	N	n	CD
cord-005550-qrrdi667	41	43	,	,	,
cord-005550-qrrdi667	41	44	79151	79151	CD
cord-005550-qrrdi667	41	45	0	0	CD
cord-005550-qrrdi667	41	46	W	W	NNP
cord-005550-qrrdi667	41	47	,	,	,
cord-005550-qrrdi667	41	48	N	n	NN
cord-005550-qrrdi667	41	49	¼	¼	NN
cord-005550-qrrdi667	41	50	3	3	CD
cord-005550-qrrdi667	41	51	)	)	-RRB-
cord-005550-qrrdi667	41	52	and	and	CC
cord-005550-qrrdi667	41	53	at	at	IN
cord-005550-qrrdi667	41	54	the	the	DT
cord-005550-qrrdi667	41	55	Tiputini	Tiputini	NNP
cord-005550-qrrdi667	41	56	Biodiversity	Biodiversity	NNP
cord-005550-qrrdi667	41	57	Station	Station	NNP
cord-005550-qrrdi667	41	58	(	(	-LRB-
cord-005550-qrrdi667	41	59	TBS	TBS	NNP
cord-005550-qrrdi667	41	60	)	)	-RRB-
cord-005550-qrrdi667	41	61	in	in	IN
cord-005550-qrrdi667	41	62	the	the	DT
cord-005550-qrrdi667	41	63	Amazonian	amazonian	JJ
cord-005550-qrrdi667	41	64	rainforest	rainforest	NN
cord-005550-qrrdi667	41	65	of	of	IN
cord-005550-qrrdi667	41	66	Ecuador	Ecuador	NNP
cord-005550-qrrdi667	41	67	(	(	-LRB-
cord-005550-qrrdi667	41	68	B0138	B0138	NNP
cord-005550-qrrdi667	41	69	0	0	NNP
cord-005550-qrrdi667	41	70	S	S	NNP
cord-005550-qrrdi667	41	71	,	,	,
cord-005550-qrrdi667	41	72	76108	76108	CD
cord-005550-qrrdi667	41	73	0	0	CD
cord-005550-qrrdi667	41	74	W	W	NNP
cord-005550-qrrdi667	41	75	,	,	,
cord-005550-qrrdi667	41	76	N	n	NN
cord-005550-qrrdi667	41	77	¼	¼	NN
cord-005550-qrrdi667	41	78	3	3	CD
cord-005550-qrrdi667	41	79	)	)	-RRB-
cord-005550-qrrdi667	41	80	between	between	IN
cord-005550-qrrdi667	41	81	1998	1998	CD
cord-005550-qrrdi667	41	82	and	and	CC
cord-005550-qrrdi667	41	83	2001	2001	CD
cord-005550-qrrdi667	41	84	.	.	.
cord-005550-qrrdi667	42	1	Animals	animal	NNS
cord-005550-qrrdi667	42	2	were	be	VBD
cord-005550-qrrdi667	42	3	sexed	sex	VBN
cord-005550-qrrdi667	42	4	,	,	,
cord-005550-qrrdi667	42	5	aged	aged	JJ
cord-005550-qrrdi667	42	6	and	and	CC
cord-005550-qrrdi667	42	7	measured	measure	VBN
cord-005550-qrrdi667	42	8	before	before	IN
cord-005550-qrrdi667	42	9	a	a	DT
cord-005550-qrrdi667	42	10	circular	circular	JJ
cord-005550-qrrdi667	42	11	wing	wing	NN
cord-005550-qrrdi667	42	12	biopsy	biopsy	NN
cord-005550-qrrdi667	42	13	with	with	IN
cord-005550-qrrdi667	42	14	a	a	DT
cord-005550-qrrdi667	42	15	diameter	diameter	NN
cord-005550-qrrdi667	42	16	of	of	IN
cord-005550-qrrdi667	42	17	4	4	CD
cord-005550-qrrdi667	42	18	mm	mm	NN
cord-005550-qrrdi667	42	19	was	be	VBD
cord-005550-qrrdi667	42	20	taken	take	VBN
cord-005550-qrrdi667	42	21	from	from	IN
cord-005550-qrrdi667	42	22	the	the	DT
cord-005550-qrrdi667	42	23	proximal	proximal	JJ
cord-005550-qrrdi667	42	24	plagiopatagium	plagiopatagium	NN
cord-005550-qrrdi667	42	25	:	:	:
cord-005550-qrrdi667	43	1	biopsies	biopsy	NNS
cord-005550-qrrdi667	43	2	were	be	VBD
cord-005550-qrrdi667	43	3	stored	store	VBN
cord-005550-qrrdi667	43	4	in	in	IN
cord-005550-qrrdi667	43	5	80	80	CD
cord-005550-qrrdi667	43	6	%	%	NN
cord-005550-qrrdi667	43	7	ethanol	ethanol	NN
cord-005550-qrrdi667	43	8	at	at	IN
cord-005550-qrrdi667	43	9	room	room	NN
cord-005550-qrrdi667	43	10	temperature	temperature	NN
cord-005550-qrrdi667	43	11	until	until	IN
cord-005550-qrrdi667	43	12	DNA	dna	NN
cord-005550-qrrdi667	43	13	isolation	isolation	NN
cord-005550-qrrdi667	43	14	.	.	.
cord-005550-qrrdi667	44	1	Genetic	genetic	JJ
cord-005550-qrrdi667	44	2	analysis	analysis	NN
cord-005550-qrrdi667	44	3	DNA	dna	NN
cord-005550-qrrdi667	44	4	was	be	VBD
cord-005550-qrrdi667	44	5	isolated	isolate	VBN
cord-005550-qrrdi667	44	6	by	by	IN
cord-005550-qrrdi667	44	7	a	a	DT
cord-005550-qrrdi667	44	8	salting	salting	NN
cord-005550-qrrdi667	44	9	out	out	RP
cord-005550-qrrdi667	44	10	procedure	procedure	NN
cord-005550-qrrdi667	44	11	(	(	-LRB-
cord-005550-qrrdi667	44	12	Mü	Mü	NNP
cord-005550-qrrdi667	44	13	llenbach	llenbach	NNP
cord-005550-qrrdi667	44	14	et	et	NNP
cord-005550-qrrdi667	44	15	al	al	NNP
cord-005550-qrrdi667	44	16	.	.	NNP
cord-005550-qrrdi667	44	17	,	,	,
cord-005550-qrrdi667	44	18	1989	1989	CD
cord-005550-qrrdi667	44	19	)	)	-RRB-
cord-005550-qrrdi667	44	20	,	,	,
cord-005550-qrrdi667	44	21	which	which	WDT
cord-005550-qrrdi667	44	22	yielded	yield	VBD
cord-005550-qrrdi667	44	23	about	about	RB
cord-005550-qrrdi667	44	24	0.5	0.5	CD
cord-005550-qrrdi667	44	25	mg	mg	CD
cord-005550-qrrdi667	44	26	of	of	IN
cord-005550-qrrdi667	44	27	total	total	JJ
cord-005550-qrrdi667	44	28	genomic	genomic	JJ
cord-005550-qrrdi667	44	29	DNA	dna	NN
cord-005550-qrrdi667	44	30	.	.	.
cord-005550-qrrdi667	45	1	The	the	DT
cord-005550-qrrdi667	45	2	exon	exon	NN
cord-005550-qrrdi667	45	3	2	2	CD
cord-005550-qrrdi667	45	4	of	of	IN
cord-005550-qrrdi667	45	5	the	the	DT
cord-005550-qrrdi667	45	6	class	class	NN
cord-005550-qrrdi667	45	7	II	II	NNP
cord-005550-qrrdi667	45	8	MHC	MHC	NNP
cord-005550-qrrdi667	45	9	gene	gene	NN
cord-005550-qrrdi667	45	10	DRB	DRB	NNP
cord-005550-qrrdi667	45	11	was	be	VBD
cord-005550-qrrdi667	45	12	initially	initially	RB
cord-005550-qrrdi667	45	13	amplified	amplify	VBN
cord-005550-qrrdi667	45	14	by	by	IN
cord-005550-qrrdi667	45	15	PCR	PCR	NNP
cord-005550-qrrdi667	45	16	with	with	IN
cord-005550-qrrdi667	45	17	the	the	DT
cord-005550-qrrdi667	45	18	primers	primer	NNS
cord-005550-qrrdi667	45	19	GH46	GH46	NNP
cord-005550-qrrdi667	45	20	and	and	CC
cord-005550-qrrdi667	45	21	GH50	GH50	NNP
cord-005550-qrrdi667	45	22	(	(	-LRB-
cord-005550-qrrdi667	45	23	Scharf	Scharf	NNP
cord-005550-qrrdi667	45	24	et	et	NNP
cord-005550-qrrdi667	45	25	al	al	NNP
cord-005550-qrrdi667	45	26	.	.	.
cord-005550-qrrdi667	46	1	,	,	,
cord-005550-qrrdi667	46	2	1988	1988	CD
cord-005550-qrrdi667	46	3	)	)	-RRB-
cord-005550-qrrdi667	46	4	but	but	CC
cord-005550-qrrdi667	46	5	only	only	RB
cord-005550-qrrdi667	46	6	small	small	JJ
cord-005550-qrrdi667	46	7	amounts	amount	NNS
cord-005550-qrrdi667	46	8	of	of	IN
cord-005550-qrrdi667	46	9	DNA	DNA	NNP
cord-005550-qrrdi667	46	10	were	be	VBD
cord-005550-qrrdi667	46	11	obtained	obtain	VBN
cord-005550-qrrdi667	46	12	.	.	.
cord-005550-qrrdi667	47	1	The	the	DT
cord-005550-qrrdi667	47	2	design	design	NN
cord-005550-qrrdi667	47	3	of	of	IN
cord-005550-qrrdi667	47	4	two	two	CD
cord-005550-qrrdi667	47	5	shorter	short	JJR
cord-005550-qrrdi667	47	6	primers	primer	NNS
cord-005550-qrrdi667	47	7	ER407B	ER407B	NNP
cord-005550-qrrdi667	47	8	(	(	-LRB-
cord-005550-qrrdi667	47	9	5	5	CD
cord-005550-qrrdi667	47	10	0	0	CD
cord-005550-qrrdi667	47	11	-AACCCCGTAGTTGTGTCTGCA-3	-aaccccgtagttgtgtctgca-3	NN
cord-005550-qrrdi667	47	12	0	0	CD
cord-005550-qrrdi667	47	13	)	)	-RRB-
cord-005550-qrrdi667	47	14	and	and	CC
cord-005550-qrrdi667	47	15	ER408	ER408	NNP
cord-005550-qrrdi667	47	16	(	(	-LRB-
cord-005550-qrrdi667	47	17	5	5	CD
cord-005550-qrrdi667	47	18	0	0	CD
cord-005550-qrrdi667	47	19	-GGATCCTTCGTGTCCCCA-3	-GGATCCTTCGTGTCCCCA-3	NNP
cord-005550-qrrdi667	47	20	0	0	CD
cord-005550-qrrdi667	47	21	)	)	-RRB-
cord-005550-qrrdi667	47	22	,	,	,
cord-005550-qrrdi667	47	23	which	which	WDT
cord-005550-qrrdi667	47	24	have	have	VBP
cord-005550-qrrdi667	47	25	a	a	DT
cord-005550-qrrdi667	47	26	similar	similar	JJ
cord-005550-qrrdi667	47	27	T	t	NN
cord-005550-qrrdi667	47	28	m	m	NN
cord-005550-qrrdi667	47	29	value	value	NN
cord-005550-qrrdi667	47	30	and	and	CC
cord-005550-qrrdi667	47	31	contain	contain	VBP
cord-005550-qrrdi667	47	32	only	only	RB
cord-005550-qrrdi667	47	33	the	the	DT
cord-005550-qrrdi667	47	34	core	core	NN
cord-005550-qrrdi667	47	35	sequences	sequence	NNS
cord-005550-qrrdi667	47	36	of	of	IN
cord-005550-qrrdi667	47	37	the	the	DT
cord-005550-qrrdi667	47	38	primers	primer	NNS
cord-005550-qrrdi667	47	39	GH46	GH46	NNP
cord-005550-qrrdi667	47	40	and	and	CC
cord-005550-qrrdi667	47	41	GH50	GH50	NNP
cord-005550-qrrdi667	47	42	,	,	,
cord-005550-qrrdi667	47	43	improved	improve	VBD
cord-005550-qrrdi667	47	44	the	the	DT
cord-005550-qrrdi667	47	45	PCR	PCR	NNP
cord-005550-qrrdi667	47	46	substantially	substantially	RB
cord-005550-qrrdi667	47	47	.	.	.
cord-005550-qrrdi667	48	1	The	the	DT
cord-005550-qrrdi667	48	2	reaction	reaction	NN
cord-005550-qrrdi667	48	3	was	be	VBD
cord-005550-qrrdi667	48	4	performed	perform	VBN
cord-005550-qrrdi667	48	5	in	in	IN
cord-005550-qrrdi667	48	6	a	a	DT
cord-005550-qrrdi667	48	7	total	total	JJ
cord-005550-qrrdi667	48	8	volume	volume	NN
cord-005550-qrrdi667	48	9	of	of	IN
cord-005550-qrrdi667	48	10	25	25	CD
cord-005550-qrrdi667	48	11	ml	ml	NNS
cord-005550-qrrdi667	48	12	,	,	,
cord-005550-qrrdi667	48	13	which	which	WDT
cord-005550-qrrdi667	48	14	contained	contain	VBD
cord-005550-qrrdi667	48	15	about	about	RB
cord-005550-qrrdi667	48	16	20	20	CD
cord-005550-qrrdi667	48	17	ng	ng	NNP
cord-005550-qrrdi667	48	18	total	total	JJ
cord-005550-qrrdi667	48	19	genomic	genomic	JJ
cord-005550-qrrdi667	48	20	DNA	dna	NN
cord-005550-qrrdi667	48	21	,	,	,
cord-005550-qrrdi667	48	22	0.4	0.4	CD
cord-005550-qrrdi667	48	23	%	%	NN
cord-005550-qrrdi667	48	24	bovine	bovine	JJ
cord-005550-qrrdi667	48	25	serum	serum	NN
cord-005550-qrrdi667	48	26	albumin	albumin	NN
cord-005550-qrrdi667	48	27	,	,	,
cord-005550-qrrdi667	48	28	1	1	CD
cord-005550-qrrdi667	48	29	mM	mm	CD
cord-005550-qrrdi667	48	30	of	of	IN
cord-005550-qrrdi667	48	31	each	each	DT
cord-005550-qrrdi667	48	32	primer	primer	NN
cord-005550-qrrdi667	48	33	(	(	-LRB-
cord-005550-qrrdi667	48	34	ER407B	er407b	NN
cord-005550-qrrdi667	48	35	and	and	CC
cord-005550-qrrdi667	48	36	ER408	ER408	NNP
cord-005550-qrrdi667	48	37	)	)	-RRB-
cord-005550-qrrdi667	48	38	,	,	,
cord-005550-qrrdi667	48	39	0.1	0.1	CD
cord-005550-qrrdi667	48	40	mM	mm	CD
cord-005550-qrrdi667	48	41	of	of	IN
cord-005550-qrrdi667	48	42	each	each	DT
cord-005550-qrrdi667	48	43	dNTP	dntp	NN
cord-005550-qrrdi667	48	44	,	,	,
cord-005550-qrrdi667	48	45	2	2	CD
cord-005550-qrrdi667	48	46	mM	mm	CD
cord-005550-qrrdi667	48	47	MgCl	MgCl	NNP
cord-005550-qrrdi667	48	48	2	2	CD
cord-005550-qrrdi667	48	49	,	,	,
cord-005550-qrrdi667	48	50	0.5	0.5	CD
cord-005550-qrrdi667	48	51	U	u	NN
cord-005550-qrrdi667	48	52	Taq	taq	NN
cord-005550-qrrdi667	48	53	-	-	HYPH
cord-005550-qrrdi667	48	54	DNA	dna	NN
cord-005550-qrrdi667	48	55	-	-	HYPH
cord-005550-qrrdi667	48	56	polymerase	polymerase	NN
cord-005550-qrrdi667	48	57	(	(	-LRB-
cord-005550-qrrdi667	48	58	Promega	Promega	NNP
cord-005550-qrrdi667	48	59	Corporation	Corporation	NNP
cord-005550-qrrdi667	48	60	,	,	,
cord-005550-qrrdi667	48	61	Madison	Madison	NNP
cord-005550-qrrdi667	48	62	,	,	,
cord-005550-qrrdi667	48	63	WI	WI	NNP
cord-005550-qrrdi667	48	64	,	,	,
cord-005550-qrrdi667	48	65	USA	USA	NNP
cord-005550-qrrdi667	48	66	,	,	,
cord-005550-qrrdi667	48	67	no	no	UH
cord-005550-qrrdi667	48	68	.	.	.
cord-005550-qrrdi667	49	1	M1665	M1665	NNP
cord-005550-qrrdi667	49	2	)	)	-RRB-
cord-005550-qrrdi667	50	1	and	and	CC
cord-005550-qrrdi667	50	2	1	1	CD
cord-005550-qrrdi667	50	3	Â	â	NN
cord-005550-qrrdi667	50	4	Taq	taq	NN
cord-005550-qrrdi667	50	5	-	-	HYPH
cord-005550-qrrdi667	50	6	DNA	dna	NN
cord-005550-qrrdi667	50	7	-	-	HYPH
cord-005550-qrrdi667	50	8	polymerase	polymerase	NN
cord-005550-qrrdi667	50	9	buffer	buffer	NN
cord-005550-qrrdi667	50	10	(	(	-LRB-
cord-005550-qrrdi667	50	11	supplied	supply	VBN
cord-005550-qrrdi667	50	12	with	with	IN
cord-005550-qrrdi667	50	13	the	the	DT
cord-005550-qrrdi667	50	14	enzyme	enzyme	NN
cord-005550-qrrdi667	50	15	)	)	-RRB-
cord-005550-qrrdi667	50	16	.	.	.
cord-005550-qrrdi667	51	1	A	a	DT
cord-005550-qrrdi667	51	2	PCR	PCR	NNP
cord-005550-qrrdi667	51	3	was	be	VBD
cord-005550-qrrdi667	51	4	started	start	VBN
cord-005550-qrrdi667	51	5	with	with	IN
cord-005550-qrrdi667	51	6	incubation	incubation	NN
cord-005550-qrrdi667	51	7	at	at	IN
cord-005550-qrrdi667	51	8	951C	951c	CD
cord-005550-qrrdi667	51	9	for	for	IN
cord-005550-qrrdi667	51	10	10	10	CD
cord-005550-qrrdi667	51	11	min	min	NN
cord-005550-qrrdi667	51	12	followed	follow	VBN
cord-005550-qrrdi667	51	13	by	by	IN
cord-005550-qrrdi667	51	14	30	30	CD
cord-005550-qrrdi667	51	15	cycles	cycle	NNS
cord-005550-qrrdi667	51	16	of	of	IN
cord-005550-qrrdi667	51	17	951C	951c	CD
cord-005550-qrrdi667	51	18	for	for	IN
cord-005550-qrrdi667	51	19	30	30	CD
cord-005550-qrrdi667	51	20	s	s	NN
cord-005550-qrrdi667	51	21	,	,	,
cord-005550-qrrdi667	51	22	601C	601c	CD
cord-005550-qrrdi667	51	23	for	for	IN
cord-005550-qrrdi667	51	24	30	30	CD
cord-005550-qrrdi667	51	25	s	s	NN
cord-005550-qrrdi667	51	26	,	,	,
cord-005550-qrrdi667	51	27	721C	721c	CD
cord-005550-qrrdi667	51	28	for	for	IN
cord-005550-qrrdi667	51	29	1.5	1.5	CD
cord-005550-qrrdi667	51	30	min	min	NN
cord-005550-qrrdi667	51	31	and	and	CC
cord-005550-qrrdi667	51	32	finished	finish	VBD
cord-005550-qrrdi667	51	33	by	by	IN
cord-005550-qrrdi667	51	34	10	10	CD
cord-005550-qrrdi667	51	35	min	min	NN
cord-005550-qrrdi667	51	36	period	period	NN
cord-005550-qrrdi667	51	37	at	at	IN
cord-005550-qrrdi667	52	1	721C.	721C.	NNP
cord-005550-qrrdi667	52	2	Amplified	amplify	VBN
cord-005550-qrrdi667	52	3	DNA	dna	NN
cord-005550-qrrdi667	52	4	fragments	fragment	NNS
cord-005550-qrrdi667	52	5	were	be	VBD
cord-005550-qrrdi667	52	6	cleaned	clean	VBN
cord-005550-qrrdi667	52	7	with	with	IN
cord-005550-qrrdi667	52	8	QIAquick	QIAquick	NNP
cord-005550-qrrdi667	52	9	PCR	PCR	NNP
cord-005550-qrrdi667	52	10	Purification	Purification	NNP
cord-005550-qrrdi667	52	11	columns	column	NNS
cord-005550-qrrdi667	52	12	(	(	-LRB-
cord-005550-qrrdi667	52	13	QIAGEN	QIAGEN	NNP
cord-005550-qrrdi667	52	14	,	,	,
cord-005550-qrrdi667	52	15	GmbH	GmbH	NNP
cord-005550-qrrdi667	52	16	,	,	,
cord-005550-qrrdi667	52	17	Hilden	Hilden	NNP
cord-005550-qrrdi667	52	18	,	,	,
cord-005550-qrrdi667	52	19	Germany	Germany	NNP
cord-005550-qrrdi667	52	20	)	)	-RRB-
cord-005550-qrrdi667	52	21	and	and	CC
cord-005550-qrrdi667	52	22	reconstituted	reconstitute	VBD
cord-005550-qrrdi667	52	23	in	in	IN
cord-005550-qrrdi667	52	24	50	50	CD
cord-005550-qrrdi667	52	25	ml	ml	NN
cord-005550-qrrdi667	52	26	Tris	Tris	NNP
cord-005550-qrrdi667	52	27	-	-	HYPH
cord-005550-qrrdi667	52	28	Cl	Cl	NNP
cord-005550-qrrdi667	52	29	buffer	buffer	NN
cord-005550-qrrdi667	52	30	(	(	-LRB-
cord-005550-qrrdi667	52	31	10	10	CD
cord-005550-qrrdi667	52	32	mM	mm	NN
cord-005550-qrrdi667	52	33	,	,	,
cord-005550-qrrdi667	52	34	pH	pH	NNP
cord-005550-qrrdi667	52	35	8.5	8.5	CD
cord-005550-qrrdi667	52	36	)	)	-RRB-
cord-005550-qrrdi667	52	37	.	.	.
cord-005550-qrrdi667	53	1	The	the	DT
cord-005550-qrrdi667	53	2	amplified	amplify	VBN
cord-005550-qrrdi667	53	3	DNA	dna	NN
cord-005550-qrrdi667	53	4	of	of	IN
cord-005550-qrrdi667	53	5	each	each	DT
cord-005550-qrrdi667	53	6	individual	individual	NN
cord-005550-qrrdi667	53	7	investigated	investigate	VBD
cord-005550-qrrdi667	53	8	was	be	VBD
cord-005550-qrrdi667	53	9	cloned	clone	VBN
cord-005550-qrrdi667	53	10	using	use	VBG
cord-005550-qrrdi667	53	11	the	the	DT
cord-005550-qrrdi667	53	12	pGEM	pGEM	NNS
cord-005550-qrrdi667	53	13	-	-	HYPH
cord-005550-qrrdi667	53	14	T	T	NNP
cord-005550-qrrdi667	53	15	Easy	Easy	NNP
cord-005550-qrrdi667	53	16	Kit	kit	NN
cord-005550-qrrdi667	53	17	(	(	-LRB-
cord-005550-qrrdi667	53	18	Promega	Promega	NNP
cord-005550-qrrdi667	53	19	)	)	-RRB-
cord-005550-qrrdi667	53	20	and	and	CC
cord-005550-qrrdi667	53	21	following	follow	VBG
cord-005550-qrrdi667	53	22	the	the	DT
cord-005550-qrrdi667	53	23	manufacturer	manufacturer	NN
cord-005550-qrrdi667	53	24	's	's	POS
cord-005550-qrrdi667	53	25	protocol	protocol	NN
cord-005550-qrrdi667	53	26	.	.	.
cord-005550-qrrdi667	54	1	TAM1-F	TAM1-F	NNP
cord-005550-qrrdi667	54	2	'	'	``
cord-005550-qrrdi667	54	3	Extra	extra	JJ
cord-005550-qrrdi667	54	4	competent	competent	JJ
cord-005550-qrrdi667	54	5	cells	cell	NNS
cord-005550-qrrdi667	54	6	(	(	-LRB-
cord-005550-qrrdi667	54	7	Active	active	JJ
cord-005550-qrrdi667	54	8	Motif	motif	NN
cord-005550-qrrdi667	54	9	,	,	,
cord-005550-qrrdi667	54	10	Rixensart	Rixensart	NNP
cord-005550-qrrdi667	54	11	,	,	,
cord-005550-qrrdi667	54	12	Belgium	Belgium	NNP
cord-005550-qrrdi667	54	13	)	)	-RRB-
cord-005550-qrrdi667	54	14	were	be	VBD
cord-005550-qrrdi667	54	15	used	use	VBN
cord-005550-qrrdi667	54	16	in	in	IN
cord-005550-qrrdi667	54	17	transformation	transformation	NN
cord-005550-qrrdi667	54	18	.	.	.
cord-005550-qrrdi667	55	1	Positive	positive	JJ
cord-005550-qrrdi667	55	2	clones	clone	NNS
cord-005550-qrrdi667	55	3	were	be	VBD
cord-005550-qrrdi667	55	4	picked	pick	VBN
cord-005550-qrrdi667	55	5	and	and	CC
cord-005550-qrrdi667	55	6	transferred	transfer	VBN
cord-005550-qrrdi667	55	7	into	into	IN
cord-005550-qrrdi667	55	8	a	a	DT
cord-005550-qrrdi667	55	9	10	10	CD
cord-005550-qrrdi667	55	10	ml	ml	NN
cord-005550-qrrdi667	55	11	PCR	PCR	NNP
cord-005550-qrrdi667	55	12	mix	mix	NN
cord-005550-qrrdi667	55	13	as	as	IN
cord-005550-qrrdi667	55	14	described	describe	VBN
cord-005550-qrrdi667	55	15	above	above	RB
cord-005550-qrrdi667	55	16	.	.	.
cord-005550-qrrdi667	56	1	If	if	IN
cord-005550-qrrdi667	56	2	a	a	DT
cord-005550-qrrdi667	56	3	DNA	dna	NN
cord-005550-qrrdi667	56	4	fragment	fragment	NN
cord-005550-qrrdi667	56	5	of	of	IN
cord-005550-qrrdi667	56	6	the	the	DT
cord-005550-qrrdi667	56	7	expected	expect	VBN
cord-005550-qrrdi667	56	8	length	length	NN
cord-005550-qrrdi667	56	9	was	be	VBD
cord-005550-qrrdi667	56	10	amplified	amplify	VBN
cord-005550-qrrdi667	56	11	,	,	,
cord-005550-qrrdi667	56	12	the	the	DT
cord-005550-qrrdi667	56	13	same	same	JJ
cord-005550-qrrdi667	56	14	clone	clone	NN
cord-005550-qrrdi667	56	15	was	be	VBD
cord-005550-qrrdi667	56	16	picked	pick	VBN
cord-005550-qrrdi667	56	17	again	again	RB
cord-005550-qrrdi667	56	18	,	,	,
cord-005550-qrrdi667	56	19	transferred	transfer	VBN
cord-005550-qrrdi667	56	20	into	into	IN
cord-005550-qrrdi667	56	21	3	3	CD
cord-005550-qrrdi667	56	22	ml	ml	NN
cord-005550-qrrdi667	56	23	LB	lb	NN
cord-005550-qrrdi667	56	24	medium	medium	NN
cord-005550-qrrdi667	56	25	containing	contain	VBG
cord-005550-qrrdi667	56	26	100	100	CD
cord-005550-qrrdi667	56	27	mg	mg	NNP
cord-005550-qrrdi667	56	28	/	/	SYM
cord-005550-qrrdi667	56	29	ml	ml	NNS
cord-005550-qrrdi667	56	30	ampicillin	ampicillin	NN
cord-005550-qrrdi667	56	31	and	and	CC
cord-005550-qrrdi667	56	32	incubated	incubate	VBN
cord-005550-qrrdi667	56	33	at	at	IN
cord-005550-qrrdi667	56	34	371C	371c	CD
cord-005550-qrrdi667	56	35	for	for	IN
cord-005550-qrrdi667	56	36	14	14	CD
cord-005550-qrrdi667	56	37	h.	h.	NN
cord-005550-qrrdi667	56	38	DNA	DNA	NNP
cord-005550-qrrdi667	56	39	was	be	VBD
cord-005550-qrrdi667	56	40	extracted	extract	VBN
cord-005550-qrrdi667	56	41	using	use	VBG
cord-005550-qrrdi667	56	42	the	the	DT
cord-005550-qrrdi667	56	43	Qiaprep	Qiaprep	NNP
cord-005550-qrrdi667	56	44	Miniprep	Miniprep	NNP
cord-005550-qrrdi667	56	45	Kit	Kit	NNP
cord-005550-qrrdi667	56	46	(	(	-LRB-
cord-005550-qrrdi667	56	47	Qiagen	Qiagen	NNP
cord-005550-qrrdi667	56	48	)	)	-RRB-
cord-005550-qrrdi667	56	49	following	follow	VBG
cord-005550-qrrdi667	56	50	the	the	DT
cord-005550-qrrdi667	56	51	recommendations	recommendation	NNS
cord-005550-qrrdi667	56	52	of	of	IN
cord-005550-qrrdi667	56	53	the	the	DT
cord-005550-qrrdi667	56	54	manufacturer	manufacturer	NN
cord-005550-qrrdi667	56	55	.	.	.
cord-005550-qrrdi667	57	1	Six	six	CD
cord-005550-qrrdi667	57	2	to	to	TO
cord-005550-qrrdi667	57	3	18	18	CD
cord-005550-qrrdi667	57	4	clones	clone	NNS
cord-005550-qrrdi667	57	5	with	with	IN
cord-005550-qrrdi667	57	6	an	an	DT
cord-005550-qrrdi667	57	7	insert	insert	NN
cord-005550-qrrdi667	57	8	of	of	IN
cord-005550-qrrdi667	57	9	the	the	DT
cord-005550-qrrdi667	57	10	DRB	DRB	NNP
cord-005550-qrrdi667	57	11	fragment	fragment	NN
cord-005550-qrrdi667	57	12	were	be	VBD
cord-005550-qrrdi667	57	13	sequenced	sequence	VBN
cord-005550-qrrdi667	57	14	from	from	IN
cord-005550-qrrdi667	57	15	each	each	DT
cord-005550-qrrdi667	57	16	individual	individual	NN
cord-005550-qrrdi667	57	17	on	on	IN
cord-005550-qrrdi667	57	18	a	a	DT
cord-005550-qrrdi667	57	19	DNA	dna	NN
cord-005550-qrrdi667	57	20	sequencer	sequencer	NN
cord-005550-qrrdi667	57	21	4200	4200	CD
cord-005550-qrrdi667	57	22	(	(	-LRB-
cord-005550-qrrdi667	57	23	LI	LI	NNP
cord-005550-qrrdi667	57	24	-	-	HYPH
cord-005550-qrrdi667	57	25	COR	COR	NNP
cord-005550-qrrdi667	57	26	)	)	-RRB-
cord-005550-qrrdi667	57	27	using	use	VBG
cord-005550-qrrdi667	57	28	the	the	DT
cord-005550-qrrdi667	57	29	ThermoSequenase	ThermoSequenase	NNP
cord-005550-qrrdi667	57	30	DYEnamic	DYEnamic	NNP
cord-005550-qrrdi667	57	31	Direct	Direct	NNP
cord-005550-qrrdi667	57	32	Sequencing	Sequencing	NNP
cord-005550-qrrdi667	57	33	Kit	kit	NN
cord-005550-qrrdi667	57	34	(	(	-LRB-
cord-005550-qrrdi667	57	35	Amersham	Amersham	NNP
cord-005550-qrrdi667	57	36	Biosciences	Biosciences	NNP
cord-005550-qrrdi667	57	37	,	,	,
cord-005550-qrrdi667	57	38	Piscataway	Piscataway	NNP
cord-005550-qrrdi667	57	39	,	,	,
cord-005550-qrrdi667	57	40	NJ	NJ	NNP
cord-005550-qrrdi667	57	41	,	,	,
cord-005550-qrrdi667	57	42	USA	USA	NNP
cord-005550-qrrdi667	57	43	)	)	-RRB-
cord-005550-qrrdi667	57	44	.	.	.
cord-005550-qrrdi667	58	1	The	the	DT
cord-005550-qrrdi667	58	2	probability	probability	NN
cord-005550-qrrdi667	58	3	of	of	IN
cord-005550-qrrdi667	58	4	missing	miss	VBG
cord-005550-qrrdi667	58	5	one	one	CD
cord-005550-qrrdi667	58	6	of	of	IN
cord-005550-qrrdi667	58	7	two	two	CD
cord-005550-qrrdi667	58	8	gene	gene	NN
cord-005550-qrrdi667	58	9	copies	copy	NNS
cord-005550-qrrdi667	58	10	among	among	IN
cord-005550-qrrdi667	58	11	six	six	CD
cord-005550-qrrdi667	58	12	clones	clone	NNS
cord-005550-qrrdi667	58	13	is	be	VBZ
cord-005550-qrrdi667	58	14	0.5	0.5	CD
cord-005550-qrrdi667	58	15	5	5	CD
cord-005550-qrrdi667	58	16	¼	¼	CD
cord-005550-qrrdi667	58	17	0.031	0.031	CD
cord-005550-qrrdi667	58	18	and	and	CC
cord-005550-qrrdi667	58	19	decreases	decrease	VBZ
cord-005550-qrrdi667	58	20	to	to	IN
cord-005550-qrrdi667	58	21	Po0.001	Po0.001	NNP
cord-005550-qrrdi667	58	22	with	with	IN
cord-005550-qrrdi667	58	23	11	11	CD
cord-005550-qrrdi667	58	24	or	or	CC
cord-005550-qrrdi667	58	25	more	more	JJR
cord-005550-qrrdi667	58	26	clones	clone	NNS
cord-005550-qrrdi667	58	27	.	.	.
cord-005550-qrrdi667	59	1	We	-PRON-	PRP
cord-005550-qrrdi667	59	2	excluded	exclude	VBD
cord-005550-qrrdi667	59	3	sequence	sequence	NN
cord-005550-qrrdi667	59	4	types	type	NNS
cord-005550-qrrdi667	59	5	from	from	IN
cord-005550-qrrdi667	59	6	the	the	DT
cord-005550-qrrdi667	59	7	analysis	analysis	NN
cord-005550-qrrdi667	59	8	,	,	,
cord-005550-qrrdi667	59	9	if	if	IN
cord-005550-qrrdi667	59	10	they	-PRON-	PRP
cord-005550-qrrdi667	59	11	were	be	VBD
cord-005550-qrrdi667	59	12	found	find	VBN
cord-005550-qrrdi667	59	13	only	only	RB
cord-005550-qrrdi667	59	14	in	in	IN
cord-005550-qrrdi667	59	15	one	one	CD
cord-005550-qrrdi667	59	16	clone	clone	NN
cord-005550-qrrdi667	59	17	to	to	TO
cord-005550-qrrdi667	59	18	account	account	VB
cord-005550-qrrdi667	59	19	for	for	IN
cord-005550-qrrdi667	59	20	nucleotide	nucleotide	JJ
cord-005550-qrrdi667	59	21	substitutions	substitution	NNS
cord-005550-qrrdi667	59	22	or	or	CC
cord-005550-qrrdi667	59	23	recombination	recombination	NN
cord-005550-qrrdi667	59	24	during	during	IN
cord-005550-qrrdi667	59	25	the	the	DT
cord-005550-qrrdi667	59	26	PCR	PCR	NNP
cord-005550-qrrdi667	59	27	or	or	CC
cord-005550-qrrdi667	59	28	cloning	cloning	NN
cord-005550-qrrdi667	59	29	.	.	.
cord-005550-qrrdi667	60	1	The	the	DT
cord-005550-qrrdi667	60	2	inheritance	inheritance	NN
cord-005550-qrrdi667	60	3	of	of	IN
cord-005550-qrrdi667	60	4	sequence	sequence	NN
cord-005550-qrrdi667	60	5	types	type	NNS
cord-005550-qrrdi667	60	6	was	be	VBD
cord-005550-qrrdi667	60	7	studied	study	VBN
cord-005550-qrrdi667	60	8	in	in	IN
cord-005550-qrrdi667	60	9	29	29	CD
cord-005550-qrrdi667	60	10	juvenile	juvenile	JJ
cord-005550-qrrdi667	60	11	bats	bat	NNS
cord-005550-qrrdi667	60	12	with	with	IN
cord-005550-qrrdi667	60	13	known	know	VBN
cord-005550-qrrdi667	60	14	mothers	mother	NNS
cord-005550-qrrdi667	60	15	and/or	and/or	CC
cord-005550-qrrdi667	60	16	fathers	father	NNS
cord-005550-qrrdi667	60	17	from	from	IN
cord-005550-qrrdi667	60	18	a	a	DT
cord-005550-qrrdi667	60	19	large	large	JJ
cord-005550-qrrdi667	60	20	colony	colony	NN
cord-005550-qrrdi667	60	21	at	at	IN
cord-005550-qrrdi667	60	22	the	the	DT
cord-005550-qrrdi667	60	23	biological	biological	JJ
cord-005550-qrrdi667	60	24	station	station	NN
cord-005550-qrrdi667	60	25	La	La	NNP
cord-005550-qrrdi667	60	26	Selva	Selva	NNP
cord-005550-qrrdi667	60	27	.	.	.
cord-005550-qrrdi667	61	1	They	-PRON-	PRP
cord-005550-qrrdi667	61	2	were	be	VBD
cord-005550-qrrdi667	61	3	assigned	assign	VBN
cord-005550-qrrdi667	61	4	to	to	IN
cord-005550-qrrdi667	61	5	their	-PRON-	PRP$
cord-005550-qrrdi667	61	6	parents	parent	NNS
cord-005550-qrrdi667	61	7	by	by	IN
cord-005550-qrrdi667	61	8	behavioural	behavioural	JJ
cord-005550-qrrdi667	61	9	observations	observation	NNS
cord-005550-qrrdi667	61	10	and	and	CC
cord-005550-qrrdi667	61	11	genotyping	genotyping	NN
cord-005550-qrrdi667	61	12	of	of	IN
cord-005550-qrrdi667	61	13	the	the	DT
cord-005550-qrrdi667	61	14	juveniles	juvenile	NNS
cord-005550-qrrdi667	61	15	and	and	CC
cord-005550-qrrdi667	61	16	their	-PRON-	PRP$
cord-005550-qrrdi667	61	17	putative	putative	JJ
cord-005550-qrrdi667	61	18	parents	parent	NNS
cord-005550-qrrdi667	61	19	at	at	IN
cord-005550-qrrdi667	61	20	11	11	CD
cord-005550-qrrdi667	61	21	microsatellite	microsatellite	JJ
cord-005550-qrrdi667	61	22	loci	locus	NNS
cord-005550-qrrdi667	61	23	as	as	IN
cord-005550-qrrdi667	61	24	described	describe	VBN
cord-005550-qrrdi667	61	25	in	in	IN
cord-005550-qrrdi667	61	26	Heckel	Heckel	NNP
cord-005550-qrrdi667	61	27	and	and	CC
cord-005550-qrrdi667	61	28	von	von	NNP
cord-005550-qrrdi667	61	29	Helversen	Helversen	NNP
cord-005550-qrrdi667	61	30	(	(	-LRB-
cord-005550-qrrdi667	61	31	2003	2003	CD
cord-005550-qrrdi667	61	32	)	)	-RRB-
cord-005550-qrrdi667	61	33	.	.	.
cord-005550-qrrdi667	62	1	Allele	allele	NN
cord-005550-qrrdi667	62	2	frequencies	frequency	NNS
cord-005550-qrrdi667	62	3	were	be	VBD
cord-005550-qrrdi667	62	4	calculated	calculate	VBN
cord-005550-qrrdi667	62	5	only	only	RB
cord-005550-qrrdi667	62	6	from	from	IN
cord-005550-qrrdi667	62	7	adult	adult	NN
cord-005550-qrrdi667	62	8	males	male	NNS
cord-005550-qrrdi667	62	9	and	and	CC
cord-005550-qrrdi667	62	10	females	female	NNS
cord-005550-qrrdi667	62	11	,	,	,
cord-005550-qrrdi667	62	12	which	which	WDT
cord-005550-qrrdi667	62	13	were	be	VBD
cord-005550-qrrdi667	62	14	sampled	sample	VBN
cord-005550-qrrdi667	62	15	in	in	IN
cord-005550-qrrdi667	62	16	Costa	Costa	NNP
cord-005550-qrrdi667	62	17	Rica	Rica	NNP
cord-005550-qrrdi667	62	18	.	.	.
cord-005550-qrrdi667	63	1	Using	use	VBG
cord-005550-qrrdi667	63	2	these	these	DT
cord-005550-qrrdi667	63	3	allele	allele	NN
cord-005550-qrrdi667	63	4	frequencies	frequency	NNS
cord-005550-qrrdi667	63	5	,	,	,
cord-005550-qrrdi667	63	6	the	the	DT
cord-005550-qrrdi667	63	7	heterozygosity	heterozygosity	NN
cord-005550-qrrdi667	63	8	at	at	IN
cord-005550-qrrdi667	63	9	a	a	DT
cord-005550-qrrdi667	63	10	given	give	VBN
cord-005550-qrrdi667	63	11	amino	amino	NN
cord-005550-qrrdi667	63	12	-	-	HYPH
cord-005550-qrrdi667	63	13	acid	acid	NN
cord-005550-qrrdi667	63	14	site	site	NN
cord-005550-qrrdi667	63	15	was	be	VBD
cord-005550-qrrdi667	63	16	calculated	calculate	VBN
cord-005550-qrrdi667	63	17	(	(	-LRB-
cord-005550-qrrdi667	63	18	Hedrick	Hedrick	NNP
cord-005550-qrrdi667	63	19	et	et	NNP
cord-005550-qrrdi667	63	20	al	al	NNP
cord-005550-qrrdi667	63	21	.	.	NNP
cord-005550-qrrdi667	63	22	,	,	,
cord-005550-qrrdi667	63	23	1991	1991	CD
cord-005550-qrrdi667	63	24	)	)	-RRB-
cord-005550-qrrdi667	63	25	.	.	.
cord-005550-qrrdi667	64	1	Hardy	Hardy	NNP
cord-005550-qrrdi667	64	2	-	-	HYPH
cord-005550-qrrdi667	64	3	Weinberg	Weinberg	NNP
cord-005550-qrrdi667	64	4	Exact	Exact	NNP
cord-005550-qrrdi667	64	5	Tests	Tests	NNPS
cord-005550-qrrdi667	64	6	were	be	VBD
cord-005550-qrrdi667	64	7	performed	perform	VBN
cord-005550-qrrdi667	64	8	with	with	IN
cord-005550-qrrdi667	64	9	the	the	DT
cord-005550-qrrdi667	64	10	program	program	NN
cord-005550-qrrdi667	64	11	GENEPOP	GENEPOP	NNP
cord-005550-qrrdi667	64	12	(	(	-LRB-
cord-005550-qrrdi667	64	13	web	web	NN
cord-005550-qrrdi667	64	14	version	version	NN
cord-005550-qrrdi667	64	15	3.4	3.4	CD
cord-005550-qrrdi667	64	16	)	)	-RRB-
cord-005550-qrrdi667	65	1	(	(	-LRB-
cord-005550-qrrdi667	65	2	Raymond	Raymond	NNP
cord-005550-qrrdi667	65	3	and	and	CC
cord-005550-qrrdi667	65	4	Rousset	Rousset	NNP
cord-005550-qrrdi667	65	5	,	,	,
cord-005550-qrrdi667	65	6	1995	1995	CD
cord-005550-qrrdi667	65	7	)	)	-RRB-
cord-005550-qrrdi667	65	8	using	use	VBG
cord-005550-qrrdi667	65	9	the	the	DT
cord-005550-qrrdi667	65	10	default	default	NN
cord-005550-qrrdi667	65	11	values	value	NNS
cord-005550-qrrdi667	65	12	(	(	-LRB-
cord-005550-qrrdi667	65	13	Markov	Markov	NNP
cord-005550-qrrdi667	65	14	chain	chain	NN
cord-005550-qrrdi667	65	15	:	:	:
cord-005550-qrrdi667	65	16	100	100	CD
cord-005550-qrrdi667	65	17	batches	batch	NNS
cord-005550-qrrdi667	65	18	and	and	CC
cord-005550-qrrdi667	65	19	1000	1000	CD
cord-005550-qrrdi667	65	20	iterations	iteration	NNS
cord-005550-qrrdi667	65	21	per	per	IN
cord-005550-qrrdi667	65	22	batch	batch	NN
cord-005550-qrrdi667	65	23	)	)	-RRB-
cord-005550-qrrdi667	65	24	.	.	.
cord-005550-qrrdi667	66	1	The	the	DT
cord-005550-qrrdi667	66	2	software	software	NN
cord-005550-qrrdi667	66	3	package	package	NN
cord-005550-qrrdi667	66	4	MEGA3	MEGA3	.
cord-005550-qrrdi667	67	1	(	(	-LRB-
cord-005550-qrrdi667	67	2	Kumar	Kumar	NNP
cord-005550-qrrdi667	67	3	et	et	NNP
cord-005550-qrrdi667	67	4	al	al	NNP
cord-005550-qrrdi667	67	5	.	.	NNP
cord-005550-qrrdi667	67	6	,	,	,
cord-005550-qrrdi667	67	7	2004	2004	CD
cord-005550-qrrdi667	67	8	)	)	-RRB-
cord-005550-qrrdi667	67	9	allowed	allow	VBD
cord-005550-qrrdi667	67	10	the	the	DT
cord-005550-qrrdi667	67	11	calculation	calculation	NN
cord-005550-qrrdi667	67	12	of	of	IN
cord-005550-qrrdi667	67	13	the	the	DT
cord-005550-qrrdi667	67	14	relative	relative	JJ
cord-005550-qrrdi667	67	15	rate	rate	NN
cord-005550-qrrdi667	67	16	of	of	IN
cord-005550-qrrdi667	67	17	nonsynonymous	nonsynonymous	JJ
cord-005550-qrrdi667	67	18	and	and	CC
cord-005550-qrrdi667	67	19	synonymous	synonymous	JJ
cord-005550-qrrdi667	67	20	substitutions	substitution	NNS
cord-005550-qrrdi667	67	21	according	accord	VBG
cord-005550-qrrdi667	67	22	to	to	IN
cord-005550-qrrdi667	67	23	Nei	Nei	NNP
cord-005550-qrrdi667	67	24	and	and	CC
cord-005550-qrrdi667	67	25	Gojobori	Gojobori	NNP
cord-005550-qrrdi667	67	26	(	(	-LRB-
cord-005550-qrrdi667	67	27	1986	1986	CD
cord-005550-qrrdi667	67	28	)	)	-RRB-
cord-005550-qrrdi667	67	29	by	by	IN
cord-005550-qrrdi667	67	30	applying	apply	VBG
cord-005550-qrrdi667	67	31	the	the	DT
cord-005550-qrrdi667	67	32	correction	correction	NN
cord-005550-qrrdi667	67	33	of	of	IN
cord-005550-qrrdi667	67	34	Jukes	Jukes	NNP
cord-005550-qrrdi667	67	35	and	and	CC
cord-005550-qrrdi667	67	36	Cantor	Cantor	NNP
cord-005550-qrrdi667	67	37	(	(	-LRB-
cord-005550-qrrdi667	67	38	1969	1969	CD
cord-005550-qrrdi667	67	39	)	)	-RRB-
cord-005550-qrrdi667	67	40	for	for	IN
cord-005550-qrrdi667	67	41	multiple	multiple	JJ
cord-005550-qrrdi667	67	42	hits	hit	NNS
cord-005550-qrrdi667	67	43	.	.	.
cord-005550-qrrdi667	68	1	The	the	DT
cord-005550-qrrdi667	68	2	program	program	NN
cord-005550-qrrdi667	68	3	Arelequin	Arelequin	NNP
cord-005550-qrrdi667	68	4	2.000	2.000	CD
cord-005550-qrrdi667	68	5	(	(	-LRB-
cord-005550-qrrdi667	68	6	Schneider	Schneider	NNP
cord-005550-qrrdi667	68	7	et	et	NNP
cord-005550-qrrdi667	68	8	al	al	NNP
cord-005550-qrrdi667	68	9	.	.	NNP
cord-005550-qrrdi667	68	10	,	,	,
cord-005550-qrrdi667	68	11	2000	2000	CD
cord-005550-qrrdi667	68	12	)	)	-RRB-
cord-005550-qrrdi667	68	13	was	be	VBD
cord-005550-qrrdi667	68	14	used	use	VBN
cord-005550-qrrdi667	68	15	to	to	TO
cord-005550-qrrdi667	68	16	perform	perform	VB
cord-005550-qrrdi667	68	17	the	the	DT
cord-005550-qrrdi667	68	18	Ewens	Ewens	NNP
cord-005550-qrrdi667	68	19	-	-	HYPH
cord-005550-qrrdi667	68	20	Watterson	Watterson	NNP
cord-005550-qrrdi667	68	21	homozygosity	homozygosity	NN
cord-005550-qrrdi667	68	22	test	test	NN
cord-005550-qrrdi667	68	23	of	of	IN
cord-005550-qrrdi667	68	24	neutrality	neutrality	NN
cord-005550-qrrdi667	68	25	.	.	.
cord-005550-qrrdi667	69	1	The	the	DT
cord-005550-qrrdi667	69	2	computer	computer	NN
cord-005550-qrrdi667	69	3	program	program	NN
cord-005550-qrrdi667	69	4	GENECONV	GENECONV	NNP
cord-005550-qrrdi667	69	5	(	(	-LRB-
cord-005550-qrrdi667	69	6	Sawyer	Sawyer	NNP
cord-005550-qrrdi667	69	7	,	,	,
cord-005550-qrrdi667	69	8	1999	1999	CD
cord-005550-qrrdi667	69	9	)	)	-RRB-
cord-005550-qrrdi667	69	10	was	be	VBD
cord-005550-qrrdi667	69	11	used	use	VBN
cord-005550-qrrdi667	69	12	to	to	TO
cord-005550-qrrdi667	69	13	test	test	VB
cord-005550-qrrdi667	69	14	for	for	IN
cord-005550-qrrdi667	69	15	statistical	statistical	JJ
cord-005550-qrrdi667	69	16	evidence	evidence	NN
cord-005550-qrrdi667	69	17	of	of	IN
cord-005550-qrrdi667	69	18	gene	gene	NN
cord-005550-qrrdi667	69	19	conversion	conversion	NN
cord-005550-qrrdi667	69	20	(	(	-LRB-
cord-005550-qrrdi667	69	21	Sawyer	Sawyer	NNP
cord-005550-qrrdi667	69	22	,	,	,
cord-005550-qrrdi667	69	23	1989	1989	CD
cord-005550-qrrdi667	69	24	)	)	-RRB-
cord-005550-qrrdi667	69	25	.	.	.
cord-005550-qrrdi667	70	1	We	-PRON-	PRP
cord-005550-qrrdi667	70	2	used	use	VBD
cord-005550-qrrdi667	70	3	the	the	DT
cord-005550-qrrdi667	70	4	default	default	NN
cord-005550-qrrdi667	70	5	settings	setting	NNS
cord-005550-qrrdi667	70	6	that	that	WDT
cord-005550-qrrdi667	70	7	do	do	VBP
cord-005550-qrrdi667	70	8	not	not	RB
cord-005550-qrrdi667	70	9	allow	allow	VB
cord-005550-qrrdi667	70	10	mismatches	mismatch	NNS
cord-005550-qrrdi667	70	11	.	.	.
cord-005550-qrrdi667	71	1	Global	global	JJ
cord-005550-qrrdi667	71	2	and	and	CC
cord-005550-qrrdi667	71	3	pairwise	pairwise	JJ
cord-005550-qrrdi667	71	4	comparisons	comparison	NNS
cord-005550-qrrdi667	71	5	were	be	VBD
cord-005550-qrrdi667	71	6	statistically	statistically	RB
cord-005550-qrrdi667	71	7	evaluated	evaluate	VBN
cord-005550-qrrdi667	71	8	with	with	IN
cord-005550-qrrdi667	71	9	10	10	CD
cord-005550-qrrdi667	71	10	000	000	CD
cord-005550-qrrdi667	71	11	permutation	permutation	NN
cord-005550-qrrdi667	71	12	runs	run	NNS
cord-005550-qrrdi667	71	13	.	.	.
cord-005550-qrrdi667	72	1	P	p	NN
cord-005550-qrrdi667	72	2	-	-	HYPH
cord-005550-qrrdi667	72	3	values	value	NNS
cord-005550-qrrdi667	72	4	were	be	VBD
cord-005550-qrrdi667	72	5	corrected	correct	VBN
cord-005550-qrrdi667	72	6	for	for	IN
cord-005550-qrrdi667	72	7	multiple	multiple	JJ
cord-005550-qrrdi667	72	8	comparisons	comparison	NNS
cord-005550-qrrdi667	72	9	.	.	.
cord-005550-qrrdi667	73	1	All	all	DT
cord-005550-qrrdi667	73	2	other	other	JJ
cord-005550-qrrdi667	73	3	statistical	statistical	JJ
cord-005550-qrrdi667	73	4	tests	test	NNS
cord-005550-qrrdi667	73	5	were	be	VBD
cord-005550-qrrdi667	73	6	performed	perform	VBN
cord-005550-qrrdi667	73	7	with	with	IN
cord-005550-qrrdi667	73	8	SPSS	SPSS	NNP
cord-005550-qrrdi667	73	9	11.0	11.0	CD
cord-005550-qrrdi667	73	10	for	for	IN
cord-005550-qrrdi667	73	11	Mac	Mac	NNP
cord-005550-qrrdi667	73	12	OSX	OSX	NNP
cord-005550-qrrdi667	73	13	.	.	.
cord-005550-qrrdi667	74	1	We	-PRON-	PRP
cord-005550-qrrdi667	74	2	detected	detect	VBD
cord-005550-qrrdi667	74	3	13	13	CD
cord-005550-qrrdi667	74	4	sequence	sequence	NN
cord-005550-qrrdi667	74	5	types	type	NNS
cord-005550-qrrdi667	74	6	(	(	-LRB-
cord-005550-qrrdi667	74	7	Figure	figure	NN
cord-005550-qrrdi667	74	8	1	1	CD
cord-005550-qrrdi667	74	9	)	)	-RRB-
cord-005550-qrrdi667	74	10	by	by	IN
cord-005550-qrrdi667	74	11	sequencing	sequence	VBG
cord-005550-qrrdi667	74	12	6	6	CD
cord-005550-qrrdi667	74	13	-	-	SYM
cord-005550-qrrdi667	74	14	18	18	CD
cord-005550-qrrdi667	74	15	clones	clone	NNS
cord-005550-qrrdi667	74	16	from	from	IN
cord-005550-qrrdi667	74	17	each	each	DT
cord-005550-qrrdi667	74	18	of	of	IN
cord-005550-qrrdi667	74	19	the	the	DT
cord-005550-qrrdi667	74	20	85	85	CD
cord-005550-qrrdi667	74	21	individuals	individual	NNS
cord-005550-qrrdi667	74	22	studied	study	VBD
cord-005550-qrrdi667	74	23	.	.	.
cord-005550-qrrdi667	75	1	Except	except	IN
cord-005550-qrrdi667	75	2	for	for	IN
cord-005550-qrrdi667	75	3	two	two	CD
cord-005550-qrrdi667	75	4	individuals	individual	NNS
cord-005550-qrrdi667	75	5	that	that	WDT
cord-005550-qrrdi667	75	6	had	have	VBD
cord-005550-qrrdi667	75	7	three	three	CD
cord-005550-qrrdi667	75	8	different	different	JJ
cord-005550-qrrdi667	75	9	sequence	sequence	NN
cord-005550-qrrdi667	75	10	types	type	NNS
cord-005550-qrrdi667	75	11	,	,	,
cord-005550-qrrdi667	75	12	only	only	RB
cord-005550-qrrdi667	75	13	one	one	CD
cord-005550-qrrdi667	75	14	or	or	CC
cord-005550-qrrdi667	75	15	two	two	CD
cord-005550-qrrdi667	75	16	sequence	sequence	NN
cord-005550-qrrdi667	75	17	types	type	NNS
cord-005550-qrrdi667	75	18	were	be	VBD
cord-005550-qrrdi667	75	19	found	find	VBN
cord-005550-qrrdi667	75	20	per	per	IN
cord-005550-qrrdi667	75	21	individual	individual	NN
cord-005550-qrrdi667	75	22	.	.	.
cord-005550-qrrdi667	76	1	The	the	DT
cord-005550-qrrdi667	76	2	sequence	sequence	NN
cord-005550-qrrdi667	76	3	type	type	NN
cord-005550-qrrdi667	76	4	B1	b1	NN
cord-005550-qrrdi667	76	5	was	be	VBD
cord-005550-qrrdi667	76	6	detected	detect	VBN
cord-005550-qrrdi667	76	7	only	only	RB
cord-005550-qrrdi667	76	8	in	in	IN
cord-005550-qrrdi667	76	9	one	one	CD
cord-005550-qrrdi667	76	10	and	and	CC
cord-005550-qrrdi667	76	11	B2	b2	NN
cord-005550-qrrdi667	76	12	in	in	IN
cord-005550-qrrdi667	76	13	another	another	DT
cord-005550-qrrdi667	76	14	individual	individual	JJ
cord-005550-qrrdi667	76	15	MHC	MHC	NNP
cord-005550-qrrdi667	76	16	diversity	diversity	NN
cord-005550-qrrdi667	76	17	in	in	IN
cord-005550-qrrdi667	76	18	the	the	DT
cord-005550-qrrdi667	76	19	sac	sac	NN
cord-005550-qrrdi667	76	20	-	-	HYPH
cord-005550-qrrdi667	76	21	winged	winged	JJ
cord-005550-qrrdi667	76	22	bat	bat	NN
cord-005550-qrrdi667	76	23	F	F	NNP
cord-005550-qrrdi667	76	24	Mayer	Mayer	NNP
cord-005550-qrrdi667	76	25	and	and	CC
cord-005550-qrrdi667	76	26	A	a	DT
cord-005550-qrrdi667	76	27	Brunner	Brunner	NNP
cord-005550-qrrdi667	76	28	(	(	-LRB-
cord-005550-qrrdi667	76	29	two	two	CD
cord-005550-qrrdi667	76	30	out	out	IN
cord-005550-qrrdi667	76	31	of	of	IN
cord-005550-qrrdi667	76	32	nine	nine	CD
cord-005550-qrrdi667	76	33	clones	clone	NNS
cord-005550-qrrdi667	76	34	in	in	IN
cord-005550-qrrdi667	76	35	each	each	DT
cord-005550-qrrdi667	76	36	individual	individual	NN
cord-005550-qrrdi667	76	37	)	)	-RRB-
cord-005550-qrrdi667	76	38	,	,	,
cord-005550-qrrdi667	76	39	although	although	IN
cord-005550-qrrdi667	76	40	two	two	CD
cord-005550-qrrdi667	76	41	different	different	JJ
cord-005550-qrrdi667	76	42	A	a	DT
cord-005550-qrrdi667	76	43	sequence	sequence	NN
cord-005550-qrrdi667	76	44	types	type	NNS
cord-005550-qrrdi667	76	45	were	be	VBD
cord-005550-qrrdi667	76	46	found	find	VBN
cord-005550-qrrdi667	76	47	in	in	IN
cord-005550-qrrdi667	76	48	both	both	DT
cord-005550-qrrdi667	76	49	individuals	individual	NNS
cord-005550-qrrdi667	76	50	.	.	.
cord-005550-qrrdi667	77	1	The	the	DT
cord-005550-qrrdi667	77	2	three	three	CD
cord-005550-qrrdi667	77	3	sequence	sequence	NN
cord-005550-qrrdi667	77	4	types	type	NNS
cord-005550-qrrdi667	77	5	indicate	indicate	VBP
cord-005550-qrrdi667	77	6	the	the	DT
cord-005550-qrrdi667	77	7	existence	existence	NN
cord-005550-qrrdi667	77	8	of	of	IN
cord-005550-qrrdi667	77	9	at	at	RB
cord-005550-qrrdi667	77	10	least	least	RBS
cord-005550-qrrdi667	77	11	two	two	CD
cord-005550-qrrdi667	77	12	DRB	DRB	NNP
cord-005550-qrrdi667	77	13	loci	locus	NNS
cord-005550-qrrdi667	77	14	in	in	IN
cord-005550-qrrdi667	77	15	the	the	DT
cord-005550-qrrdi667	77	16	genome	genome	NN
cord-005550-qrrdi667	77	17	of	of	IN
cord-005550-qrrdi667	77	18	the	the	DT
cord-005550-qrrdi667	77	19	sac	sac	NN
cord-005550-qrrdi667	77	20	-	-	HYPH
cord-005550-qrrdi667	77	21	winged	winged	JJ
cord-005550-qrrdi667	77	22	bat	bat	NN
cord-005550-qrrdi667	77	23	.	.	.
cord-005550-qrrdi667	78	1	A	a	NN
cord-005550-qrrdi667	78	2	and	and	CC
cord-005550-qrrdi667	78	3	B	b	NN
cord-005550-qrrdi667	78	4	sequence	sequence	NN
cord-005550-qrrdi667	78	5	types	type	NNS
cord-005550-qrrdi667	78	6	differ	differ	VBP
cord-005550-qrrdi667	78	7	in	in	IN
cord-005550-qrrdi667	78	8	length	length	NN
cord-005550-qrrdi667	78	9	by	by	IN
cord-005550-qrrdi667	78	10	75	75	CD
cord-005550-qrrdi667	78	11	bp	bp	NNS
cord-005550-qrrdi667	78	12	.	.	.
cord-005550-qrrdi667	79	1	All	all	DT
cord-005550-qrrdi667	79	2	sequence	sequence	NN
cord-005550-qrrdi667	79	3	types	type	NNS
cord-005550-qrrdi667	79	4	contained	contain	VBD
cord-005550-qrrdi667	79	5	an	an	DT
cord-005550-qrrdi667	79	6	open	open	JJ
cord-005550-qrrdi667	79	7	reading	reading	NN
cord-005550-qrrdi667	79	8	frame	frame	NN
cord-005550-qrrdi667	79	9	without	without	IN
cord-005550-qrrdi667	79	10	stop	stop	NN
cord-005550-qrrdi667	79	11	codons	codon	NNS
cord-005550-qrrdi667	79	12	(	(	-LRB-
cord-005550-qrrdi667	79	13	Figure	figure	NN
cord-005550-qrrdi667	79	14	2	2	CD
cord-005550-qrrdi667	79	15	)	)	-RRB-
cord-005550-qrrdi667	79	16	and	and	CC
cord-005550-qrrdi667	79	17	showed	show	VBD
cord-005550-qrrdi667	79	18	high	high	JJ
cord-005550-qrrdi667	79	19	similarity	similarity	NN
cord-005550-qrrdi667	79	20	to	to	IN
cord-005550-qrrdi667	79	21	the	the	DT
cord-005550-qrrdi667	79	22	exon	exon	NN
cord-005550-qrrdi667	79	23	2	2	CD
cord-005550-qrrdi667	79	24	amino	amino	NN
cord-005550-qrrdi667	79	25	-	-	HYPH
cord-005550-qrrdi667	79	26	acid	acid	NN
cord-005550-qrrdi667	79	27	sequence	sequence	NN
cord-005550-qrrdi667	79	28	of	of	IN
cord-005550-qrrdi667	79	29	the	the	DT
cord-005550-qrrdi667	79	30	mammalian	mammalian	JJ
cord-005550-qrrdi667	79	31	DRB	DRB	NNP
cord-005550-qrrdi667	79	32	locus	locus	NN
cord-005550-qrrdi667	79	33	.	.	.
cord-005550-qrrdi667	80	1	Inheritance	inheritance	NN
cord-005550-qrrdi667	80	2	of	of	IN
cord-005550-qrrdi667	80	3	DRB	DRB	NNP
cord-005550-qrrdi667	80	4	sequence	sequence	NN
cord-005550-qrrdi667	80	5	types	type	NNS
cord-005550-qrrdi667	80	6	was	be	VBD
cord-005550-qrrdi667	80	7	studied	study	VBN
cord-005550-qrrdi667	80	8	in	in	IN
cord-005550-qrrdi667	80	9	12	12	CD
cord-005550-qrrdi667	80	10	single	single	JJ
cord-005550-qrrdi667	80	11	parent	parent	NN
cord-005550-qrrdi667	80	12	-	-	HYPH
cord-005550-qrrdi667	80	13	offspring	offspre	VBG
cord-005550-qrrdi667	80	14	pairs	pair	NNS
cord-005550-qrrdi667	80	15	and	and	CC
cord-005550-qrrdi667	80	16	17	17	CD
cord-005550-qrrdi667	80	17	complete	complete	JJ
cord-005550-qrrdi667	80	18	families	family	NNS
cord-005550-qrrdi667	80	19	that	that	WDT
cord-005550-qrrdi667	80	20	consisted	consist	VBD
cord-005550-qrrdi667	80	21	of	of	IN
cord-005550-qrrdi667	80	22	a	a	DT
cord-005550-qrrdi667	80	23	juvenile	juvenile	NN
cord-005550-qrrdi667	80	24	and	and	CC
cord-005550-qrrdi667	80	25	both	both	DT
cord-005550-qrrdi667	80	26	parents	parent	NNS
cord-005550-qrrdi667	80	27	.	.	.
cord-005550-qrrdi667	81	1	Among	among	IN
cord-005550-qrrdi667	81	2	all	all	DT
cord-005550-qrrdi667	81	3	individuals	individual	NNS
cord-005550-qrrdi667	81	4	,	,	,
cord-005550-qrrdi667	81	5	only	only	RB
cord-005550-qrrdi667	81	6	A	a	DT
cord-005550-qrrdi667	81	7	sequence	sequence	NN
cord-005550-qrrdi667	81	8	types	type	NNS
cord-005550-qrrdi667	81	9	were	be	VBD
cord-005550-qrrdi667	81	10	found	find	VBN
cord-005550-qrrdi667	81	11	.	.	.
cord-005550-qrrdi667	82	1	The	the	DT
cord-005550-qrrdi667	82	2	genotypes	genotype	NNS
cord-005550-qrrdi667	82	3	of	of	IN
cord-005550-qrrdi667	82	4	all	all	DT
cord-005550-qrrdi667	82	5	juveniles	juvenile	NNS
cord-005550-qrrdi667	82	6	matched	match	VBN
cord-005550-qrrdi667	82	7	to	to	IN
cord-005550-qrrdi667	82	8	the	the	DT
cord-005550-qrrdi667	82	9	genotypes	genotype	NNS
cord-005550-qrrdi667	82	10	of	of	IN
cord-005550-qrrdi667	82	11	their	-PRON-	PRP$
cord-005550-qrrdi667	82	12	parents	parent	NNS
cord-005550-qrrdi667	82	13	as	as	IN
cord-005550-qrrdi667	82	14	expected	expect	VBN
cord-005550-qrrdi667	82	15	under	under	IN
cord-005550-qrrdi667	82	16	Mendelian	mendelian	JJ
cord-005550-qrrdi667	82	17	inheritance	inheritance	NN
cord-005550-qrrdi667	82	18	.	.	.
cord-005550-qrrdi667	83	1	According	accord	VBG
cord-005550-qrrdi667	83	2	to	to	IN
cord-005550-qrrdi667	83	3	the	the	DT
cord-005550-qrrdi667	83	4	differences	difference	NNS
cord-005550-qrrdi667	83	5	in	in	IN
cord-005550-qrrdi667	83	6	length	length	NN
cord-005550-qrrdi667	83	7	between	between	IN
cord-005550-qrrdi667	83	8	A	a	NN
cord-005550-qrrdi667	83	9	and	and	CC
cord-005550-qrrdi667	83	10	B	b	NN
cord-005550-qrrdi667	83	11	sequence	sequence	NN
cord-005550-qrrdi667	83	12	types	type	NNS
cord-005550-qrrdi667	83	13	and	and	CC
cord-005550-qrrdi667	83	14	the	the	DT
cord-005550-qrrdi667	83	15	Mendelian	mendelian	JJ
cord-005550-qrrdi667	83	16	inheritance	inheritance	NN
cord-005550-qrrdi667	83	17	of	of	IN
cord-005550-qrrdi667	83	18	A	a	DT
cord-005550-qrrdi667	83	19	sequence	sequence	NN
cord-005550-qrrdi667	83	20	types	type	NNS
cord-005550-qrrdi667	83	21	,	,	,
cord-005550-qrrdi667	83	22	we	-PRON-	PRP
cord-005550-qrrdi667	83	23	assume	assume	VBP
cord-005550-qrrdi667	83	24	that	that	IN
cord-005550-qrrdi667	83	25	all	all	PDT
cord-005550-qrrdi667	83	26	A	a	DT
cord-005550-qrrdi667	83	27	sequence	sequence	NN
cord-005550-qrrdi667	83	28	types	type	NNS
cord-005550-qrrdi667	83	29	are	be	VBP
cord-005550-qrrdi667	83	30	coded	code	VBN
cord-005550-qrrdi667	83	31	by	by	IN
cord-005550-qrrdi667	83	32	a	a	DT
cord-005550-qrrdi667	83	33	single	single	JJ
cord-005550-qrrdi667	83	34	locus	locus	NN
cord-005550-qrrdi667	83	35	and	and	CC
cord-005550-qrrdi667	83	36	refer	refer	VB
cord-005550-qrrdi667	83	37	to	to	IN
cord-005550-qrrdi667	83	38	them	-PRON-	PRP
cord-005550-qrrdi667	83	39	as	as	IN
cord-005550-qrrdi667	83	40	alleles	allele	NNS
cord-005550-qrrdi667	83	41	of	of	IN
cord-005550-qrrdi667	83	42	the	the	DT
cord-005550-qrrdi667	83	43	DRB1	DRB1	NNP
cord-005550-qrrdi667	83	44	locus	locus	NN
cord-005550-qrrdi667	83	45	of	of	IN
cord-005550-qrrdi667	83	46	S.	S.	NNP
cord-005550-qrrdi667	83	47	bilineata	bilineata	NN
cord-005550-qrrdi667	83	48	.	.	.
cord-005550-qrrdi667	84	1	The	the	DT
cord-005550-qrrdi667	84	2	genotypes	genotype	NNS
cord-005550-qrrdi667	84	3	of	of	IN
cord-005550-qrrdi667	84	4	79	79	CD
cord-005550-qrrdi667	84	5	adult	adult	NN
cord-005550-qrrdi667	84	6	individuals	individual	NNS
cord-005550-qrrdi667	84	7	from	from	IN
cord-005550-qrrdi667	84	8	the	the	DT
cord-005550-qrrdi667	84	9	La	La	NNP
cord-005550-qrrdi667	84	10	Selva	Selva	NNP
cord-005550-qrrdi667	84	11	population	population	NN
cord-005550-qrrdi667	84	12	did	do	VBD
cord-005550-qrrdi667	84	13	not	not	RB
cord-005550-qrrdi667	84	14	deviate	deviate	VB
cord-005550-qrrdi667	84	15	from	from	IN
cord-005550-qrrdi667	84	16	Hardy	Hardy	NNP
cord-005550-qrrdi667	84	17	-	-	HYPH
cord-005550-qrrdi667	84	18	Weinberg	Weinberg	NNP
cord-005550-qrrdi667	84	19	expectation	expectation	NN
cord-005550-qrrdi667	84	20	(	(	-LRB-
cord-005550-qrrdi667	84	21	P	p	NN
cord-005550-qrrdi667	84	22	¼	¼	CD
cord-005550-qrrdi667	84	23	0.13	0.13	CD
cord-005550-qrrdi667	84	24	)	)	-RRB-
cord-005550-qrrdi667	84	25	.	.	.
cord-005550-qrrdi667	85	1	It	-PRON-	PRP
cord-005550-qrrdi667	85	2	remains	remain	VBZ
cord-005550-qrrdi667	85	3	unclear	unclear	JJ
cord-005550-qrrdi667	85	4	,	,	,
cord-005550-qrrdi667	85	5	whether	whether	IN
cord-005550-qrrdi667	85	6	the	the	DT
cord-005550-qrrdi667	85	7	two	two	CD
cord-005550-qrrdi667	85	8	B	b	NN
cord-005550-qrrdi667	85	9	sequence	sequence	NN
cord-005550-qrrdi667	85	10	types	type	NNS
cord-005550-qrrdi667	85	11	originate	originate	VBP
cord-005550-qrrdi667	85	12	from	from	IN
cord-005550-qrrdi667	85	13	one	one	CD
cord-005550-qrrdi667	85	14	or	or	CC
cord-005550-qrrdi667	85	15	two	two	CD
cord-005550-qrrdi667	85	16	loci	locus	NNS
cord-005550-qrrdi667	85	17	.	.	.
cord-005550-qrrdi667	86	1	Alignment	alignment	NN
cord-005550-qrrdi667	86	2	of	of	IN
cord-005550-qrrdi667	86	3	all	all	DT
cord-005550-qrrdi667	86	4	91	91	CD
cord-005550-qrrdi667	86	5	polymorphic	polymorphic	JJ
cord-005550-qrrdi667	86	6	nucleotide	nucleotide	JJ
cord-005550-qrrdi667	86	7	sites	site	NNS
cord-005550-qrrdi667	86	8	among	among	IN
cord-005550-qrrdi667	86	9	all	all	DT
cord-005550-qrrdi667	86	10	sequence	sequence	NN
cord-005550-qrrdi667	86	11	types	type	NNS
cord-005550-qrrdi667	86	12	found	find	VBD
cord-005550-qrrdi667	86	13	.	.	.
cord-005550-qrrdi667	87	1	Boxes	box	NNS
cord-005550-qrrdi667	87	2	mark	mark	VBP
cord-005550-qrrdi667	87	3	identical	identical	JJ
cord-005550-qrrdi667	87	4	sequence	sequence	NN
cord-005550-qrrdi667	87	5	fragments	fragment	NNS
cord-005550-qrrdi667	87	6	among	among	IN
cord-005550-qrrdi667	87	7	DRB1	DRB1	NNP
cord-005550-qrrdi667	87	8	alleles	allele	NNS
cord-005550-qrrdi667	87	9	that	that	WDT
cord-005550-qrrdi667	87	10	are	be	VBP
cord-005550-qrrdi667	87	11	statistically	statistically	RB
cord-005550-qrrdi667	87	12	supported	support	VBN
cord-005550-qrrdi667	87	13	by	by	IN
cord-005550-qrrdi667	87	14	permutation	permutation	NN
cord-005550-qrrdi667	87	15	tests	test	NNS
cord-005550-qrrdi667	87	16	implemented	implement	VBN
cord-005550-qrrdi667	87	17	in	in	IN
cord-005550-qrrdi667	87	18	the	the	DT
cord-005550-qrrdi667	87	19	software	software	NN
cord-005550-qrrdi667	87	20	GENECONV	GENECONV	NNP
cord-005550-qrrdi667	87	21	(	(	-LRB-
cord-005550-qrrdi667	87	22	Sawyer	Sawyer	NNP
cord-005550-qrrdi667	87	23	,	,	,
cord-005550-qrrdi667	87	24	1999	1999	CD
cord-005550-qrrdi667	87	25	)	)	-RRB-
cord-005550-qrrdi667	87	26	.	.	.
cord-005550-qrrdi667	88	1	A	a	DT
cord-005550-qrrdi667	88	2	shared	share	VBN
cord-005550-qrrdi667	88	3	fragment	fragment	NN
cord-005550-qrrdi667	88	4	between	between	IN
cord-005550-qrrdi667	88	5	the	the	DT
cord-005550-qrrdi667	88	6	sequence	sequence	NN
cord-005550-qrrdi667	88	7	types	type	VBZ
cord-005550-qrrdi667	88	8	A4	A4	NNP
cord-005550-qrrdi667	88	9	and	and	CC
cord-005550-qrrdi667	88	10	B2	B2	NNP
cord-005550-qrrdi667	88	11	is	be	VBZ
cord-005550-qrrdi667	88	12	underlined	underline	VBN
cord-005550-qrrdi667	88	13	.	.	.
cord-005550-qrrdi667	89	1	Only	only	RB
cord-005550-qrrdi667	89	2	significantly	significantly	RB
cord-005550-qrrdi667	89	3	shared	share	VBN
cord-005550-qrrdi667	89	4	fragments	fragment	NNS
cord-005550-qrrdi667	89	5	of	of	IN
cord-005550-qrrdi667	89	6	global	global	JJ
cord-005550-qrrdi667	89	7	comparisons	comparison	NNS
cord-005550-qrrdi667	89	8	are	be	VBP
cord-005550-qrrdi667	89	9	shown	show	VBN
cord-005550-qrrdi667	89	10	.	.	.
cord-005550-qrrdi667	90	1	B	b	NN
cord-005550-qrrdi667	90	2	sequence	sequence	NN
cord-005550-qrrdi667	90	3	types	type	NNS
cord-005550-qrrdi667	90	4	are	be	VBP
cord-005550-qrrdi667	90	5	characterized	characterize	VBN
cord-005550-qrrdi667	90	6	by	by	IN
cord-005550-qrrdi667	90	7	a	a	DT
cord-005550-qrrdi667	90	8	75	75	CD
cord-005550-qrrdi667	90	9	bp	bp	NNP
cord-005550-qrrdi667	90	10	fragment	fragment	NN
cord-005550-qrrdi667	90	11	that	that	WDT
cord-005550-qrrdi667	90	12	lacks	lack	VBZ
cord-005550-qrrdi667	90	13	in	in	IN
cord-005550-qrrdi667	90	14	A	a	DT
cord-005550-qrrdi667	90	15	sequence	sequence	NN
cord-005550-qrrdi667	90	16	types	type	NNS
cord-005550-qrrdi667	90	17	.	.	.
cord-005550-qrrdi667	91	1	The	the	DT
cord-005550-qrrdi667	91	2	complete	complete	JJ
cord-005550-qrrdi667	91	3	DNA	dna	NN
cord-005550-qrrdi667	91	4	sequences	sequence	NNS
cord-005550-qrrdi667	91	5	are	be	VBP
cord-005550-qrrdi667	91	6	available	available	JJ
cord-005550-qrrdi667	91	7	at	at	IN
cord-005550-qrrdi667	91	8	GenBank	GenBank	NNP
cord-005550-qrrdi667	91	9	(	(	-LRB-
cord-005550-qrrdi667	91	10	accession	accession	NN
cord-005550-qrrdi667	91	11	nos	nos	NNP
cord-005550-qrrdi667	91	12	.	.	.
cord-005550-qrrdi667	91	13	EF533888-EF53390	EF533888-EF53390	NNP
cord-005550-qrrdi667	91	14	)	)	-RRB-
cord-005550-qrrdi667	91	15	.	.	.
cord-005550-qrrdi667	92	1	,	,	,
cord-005550-qrrdi667	92	2	11	11	CD
cord-005550-qrrdi667	92	3	,	,	,
cord-005550-qrrdi667	92	4	13	13	CD
cord-005550-qrrdi667	92	5	,	,	,
cord-005550-qrrdi667	92	6	28	28	CD
cord-005550-qrrdi667	92	7	,	,	,
cord-005550-qrrdi667	92	8	30	30	CD
cord-005550-qrrdi667	92	9	,	,	,
cord-005550-qrrdi667	92	10	32	32	CD
cord-005550-qrrdi667	92	11	,	,	,
cord-005550-qrrdi667	92	12	37	37	CD
cord-005550-qrrdi667	92	13	,	,	,
cord-005550-qrrdi667	92	14	38	38	CD
cord-005550-qrrdi667	92	15	,	,	,
cord-005550-qrrdi667	92	16	47	47	CD
cord-005550-qrrdi667	92	17	,	,	,
cord-005550-qrrdi667	92	18	56	56	CD
cord-005550-qrrdi667	92	19	,	,	,
cord-005550-qrrdi667	92	20	60	60	CD
cord-005550-qrrdi667	92	21	,	,	,
cord-005550-qrrdi667	92	22	61	61	CD
cord-005550-qrrdi667	92	23	,	,	,
cord-005550-qrrdi667	92	24	65	65	CD
cord-005550-qrrdi667	92	25	,	,	,
cord-005550-qrrdi667	92	26	68	68	CD
cord-005550-qrrdi667	92	27	,	,	,
cord-005550-qrrdi667	92	28	70	70	CD
cord-005550-qrrdi667	92	29	,	,	,
cord-005550-qrrdi667	92	30	71	71	CD
cord-005550-qrrdi667	92	31	,	,	,
cord-005550-qrrdi667	92	32	74	74	CD
cord-005550-qrrdi667	92	33	and	and	CC
cord-005550-qrrdi667	92	34	78	78	CD
cord-005550-qrrdi667	92	35	are	be	VBP
cord-005550-qrrdi667	92	36	assumed	assume	VBN
cord-005550-qrrdi667	92	37	to	to	TO
cord-005550-qrrdi667	92	38	be	be	VB
cord-005550-qrrdi667	92	39	antigen	antigen	NN
cord-005550-qrrdi667	92	40	-	-	HYPH
cord-005550-qrrdi667	92	41	binding	bind	VBG
cord-005550-qrrdi667	92	42	sites	site	NNS
cord-005550-qrrdi667	92	43	(	(	-LRB-
cord-005550-qrrdi667	92	44	Brown	Brown	NNP
cord-005550-qrrdi667	92	45	et	et	NNP
cord-005550-qrrdi667	92	46	al	al	NNP
cord-005550-qrrdi667	92	47	.	.	NNP
cord-005550-qrrdi667	92	48	,	,	,
cord-005550-qrrdi667	92	49	1993	1993	CD
cord-005550-qrrdi667	92	50	)	)	-RRB-
cord-005550-qrrdi667	92	51	and	and	CC
cord-005550-qrrdi667	92	52	are	be	VBP
cord-005550-qrrdi667	92	53	labelled	label	VBN
cord-005550-qrrdi667	92	54	with	with	IN
cord-005550-qrrdi667	92	55	an	an	DT
cord-005550-qrrdi667	92	56	asterisk	asterisk	NN
cord-005550-qrrdi667	92	57	.	.	.
cord-005550-qrrdi667	93	1	MHC	MHC	NNP
cord-005550-qrrdi667	93	2	diversity	diversity	NN
cord-005550-qrrdi667	93	3	in	in	IN
cord-005550-qrrdi667	93	4	the	the	DT
cord-005550-qrrdi667	93	5	sac	sac	NN
cord-005550-qrrdi667	93	6	-	-	HYPH
cord-005550-qrrdi667	93	7	winged	winged	JJ
cord-005550-qrrdi667	93	8	bat	bat	NN
cord-005550-qrrdi667	93	9	Variation	variation	NN
cord-005550-qrrdi667	93	10	at	at	IN
cord-005550-qrrdi667	93	11	the	the	DT
cord-005550-qrrdi667	93	12	DRB1	DRB1	NNP
cord-005550-qrrdi667	93	13	locus	locus	NN
cord-005550-qrrdi667	93	14	;	;	:
cord-005550-qrrdi667	93	15	evidence	evidence	NN
cord-005550-qrrdi667	93	16	of	of	IN
cord-005550-qrrdi667	93	17	recombination/	recombination/	NNS
cord-005550-qrrdi667	93	18	gene	gene	NN
cord-005550-qrrdi667	93	19	conversion	conversion	NN
cord-005550-qrrdi667	94	1	Ten	ten	CD
cord-005550-qrrdi667	94	2	alleles	allele	NNS
cord-005550-qrrdi667	94	3	were	be	VBD
cord-005550-qrrdi667	94	4	detected	detect	VBN
cord-005550-qrrdi667	94	5	within	within	IN
cord-005550-qrrdi667	94	6	the	the	DT
cord-005550-qrrdi667	94	7	main	main	JJ
cord-005550-qrrdi667	94	8	sample	sample	NN
cord-005550-qrrdi667	94	9	of	of	IN
cord-005550-qrrdi667	94	10	79	79	CD
cord-005550-qrrdi667	94	11	individuals	individual	NNS
cord-005550-qrrdi667	94	12	from	from	IN
cord-005550-qrrdi667	94	13	the	the	DT
cord-005550-qrrdi667	94	14	La	La	NNP
cord-005550-qrrdi667	94	15	Selva	Selva	NNP
cord-005550-qrrdi667	94	16	population	population	NN
cord-005550-qrrdi667	94	17	.	.	.
cord-005550-qrrdi667	95	1	Allele	Allele	NNP
cord-005550-qrrdi667	95	2	A1	A1	NNP
cord-005550-qrrdi667	95	3	was	be	VBD
cord-005550-qrrdi667	95	4	most	most	RBS
cord-005550-qrrdi667	95	5	frequent	frequent	JJ
cord-005550-qrrdi667	95	6	and	and	CC
cord-005550-qrrdi667	95	7	occurred	occur	VBD
cord-005550-qrrdi667	95	8	at	at	IN
cord-005550-qrrdi667	95	9	a	a	DT
cord-005550-qrrdi667	95	10	frequency	frequency	NN
cord-005550-qrrdi667	95	11	of	of	IN
cord-005550-qrrdi667	95	12	80.4	80.4	CD
cord-005550-qrrdi667	95	13	%	%	NN
cord-005550-qrrdi667	95	14	among	among	IN
cord-005550-qrrdi667	95	15	69	69	CD
cord-005550-qrrdi667	95	16	adult	adult	NN
cord-005550-qrrdi667	95	17	bats	bat	NNS
cord-005550-qrrdi667	95	18	(	(	-LRB-
cord-005550-qrrdi667	95	19	Figure	figure	NN
cord-005550-qrrdi667	95	20	3	3	CD
cord-005550-qrrdi667	95	21	)	)	-RRB-
cord-005550-qrrdi667	95	22	.	.	.
cord-005550-qrrdi667	96	1	The	the	DT
cord-005550-qrrdi667	96	2	alleles	allele	NNS
cord-005550-qrrdi667	96	3	A5	A5	NNP
cord-005550-qrrdi667	96	4	,	,	,
cord-005550-qrrdi667	96	5	A8	A8	NNP
cord-005550-qrrdi667	96	6	,	,	,
cord-005550-qrrdi667	96	7	A10	A10	NNP
cord-005550-qrrdi667	96	8	and	and	CC
cord-005550-qrrdi667	96	9	A11	A11	NNP
cord-005550-qrrdi667	96	10	were	be	VBD
cord-005550-qrrdi667	96	11	detected	detect	VBN
cord-005550-qrrdi667	96	12	in	in	IN
cord-005550-qrrdi667	96	13	only	only	RB
cord-005550-qrrdi667	96	14	one	one	CD
cord-005550-qrrdi667	96	15	individual	individual	NN
cord-005550-qrrdi667	96	16	.	.	.
cord-005550-qrrdi667	97	1	Observed	observed	JJ
cord-005550-qrrdi667	97	2	heterozygosity	heterozygosity	NN
cord-005550-qrrdi667	97	3	(	(	-LRB-
cord-005550-qrrdi667	97	4	0.30	0.30	CD
cord-005550-qrrdi667	97	5	)	)	-RRB-
cord-005550-qrrdi667	97	6	was	be	VBD
cord-005550-qrrdi667	97	7	lower	low	JJR
cord-005550-qrrdi667	97	8	than	than	IN
cord-005550-qrrdi667	97	9	expected	expect	VBN
cord-005550-qrrdi667	97	10	heterozygosity	heterozygosity	NN
cord-005550-qrrdi667	97	11	(	(	-LRB-
cord-005550-qrrdi667	97	12	0.35	0.35	CD
cord-005550-qrrdi667	97	13	)	)	-RRB-
cord-005550-qrrdi667	97	14	.	.	.
cord-005550-qrrdi667	98	1	The	the	DT
cord-005550-qrrdi667	98	2	Hardy	Hardy	NNP
cord-005550-qrrdi667	98	3	-	-	HYPH
cord-005550-qrrdi667	98	4	Weinberg	Weinberg	NNP
cord-005550-qrrdi667	98	5	Exact	Exact	NNP
cord-005550-qrrdi667	98	6	Test	Test	NNP
cord-005550-qrrdi667	98	7	on	on	IN
cord-005550-qrrdi667	98	8	heterozygote	heterozygote	NN
cord-005550-qrrdi667	98	9	deficiency	deficiency	NN
cord-005550-qrrdi667	98	10	(	(	-LRB-
cord-005550-qrrdi667	98	11	GENEPOP	GENEPOP	NNP
cord-005550-qrrdi667	98	12	)	)	-RRB-
cord-005550-qrrdi667	98	13	revealed	reveal	VBD
cord-005550-qrrdi667	98	14	an	an	DT
cord-005550-qrrdi667	98	15	almost	almost	RB
cord-005550-qrrdi667	98	16	significant	significant	JJ
cord-005550-qrrdi667	98	17	result	result	NN
cord-005550-qrrdi667	98	18	(	(	-LRB-
cord-005550-qrrdi667	98	19	P	p	NN
cord-005550-qrrdi667	98	20	¼	¼	NN
cord-005550-qrrdi667	98	21	0.0575	0.0575	CD
cord-005550-qrrdi667	98	22	)	)	-RRB-
cord-005550-qrrdi667	98	23	.	.	.
cord-005550-qrrdi667	99	1	Within	within	IN
cord-005550-qrrdi667	99	2	the	the	DT
cord-005550-qrrdi667	99	3	Panama	Panama	NNP
cord-005550-qrrdi667	99	4	sample	sample	NN
cord-005550-qrrdi667	99	5	of	of	IN
cord-005550-qrrdi667	99	6	three	three	CD
cord-005550-qrrdi667	99	7	individuals	individual	NNS
cord-005550-qrrdi667	99	8	,	,	,
cord-005550-qrrdi667	99	9	allele	allele	NN
cord-005550-qrrdi667	99	10	A1	a1	NN
cord-005550-qrrdi667	99	11	was	be	VBD
cord-005550-qrrdi667	99	12	also	also	RB
cord-005550-qrrdi667	99	13	most	most	RBS
cord-005550-qrrdi667	99	14	common	common	JJ
cord-005550-qrrdi667	99	15	(	(	-LRB-
cord-005550-qrrdi667	99	16	four	four	CD
cord-005550-qrrdi667	99	17	copies	copy	NNS
cord-005550-qrrdi667	99	18	)	)	-RRB-
cord-005550-qrrdi667	99	19	and	and	CC
cord-005550-qrrdi667	99	20	a	a	DT
cord-005550-qrrdi667	99	21	single	single	JJ
cord-005550-qrrdi667	99	22	copy	copy	NN
cord-005550-qrrdi667	99	23	of	of	IN
cord-005550-qrrdi667	99	24	allele	allele	NN
cord-005550-qrrdi667	99	25	A2	a2	NN
cord-005550-qrrdi667	99	26	and	and	CC
cord-005550-qrrdi667	99	27	a	a	DT
cord-005550-qrrdi667	99	28	new	new	JJ
cord-005550-qrrdi667	99	29	allele	allele	NN
cord-005550-qrrdi667	100	1	A11	A11	NNP
cord-005550-qrrdi667	100	2	were	be	VBD
cord-005550-qrrdi667	100	3	found	find	VBN
cord-005550-qrrdi667	100	4	.	.	.
cord-005550-qrrdi667	101	1	In	in	IN
cord-005550-qrrdi667	101	2	contrast	contrast	NN
cord-005550-qrrdi667	101	3	,	,	,
cord-005550-qrrdi667	101	4	allele	allele	NN
cord-005550-qrrdi667	101	5	A6	A6	NNP
cord-005550-qrrdi667	101	6	was	be	VBD
cord-005550-qrrdi667	101	7	most	most	RBS
cord-005550-qrrdi667	101	8	common	common	JJ
cord-005550-qrrdi667	101	9	in	in	IN
cord-005550-qrrdi667	101	10	three	three	CD
cord-005550-qrrdi667	101	11	animals	animal	NNS
cord-005550-qrrdi667	101	12	from	from	IN
cord-005550-qrrdi667	101	13	Ecuador	Ecuador	NNP
cord-005550-qrrdi667	101	14	(	(	-LRB-
cord-005550-qrrdi667	101	15	three	three	CD
cord-005550-qrrdi667	101	16	copies	copy	NNS
cord-005550-qrrdi667	101	17	)	)	-RRB-
cord-005550-qrrdi667	101	18	.	.	.
cord-005550-qrrdi667	102	1	The	the	DT
cord-005550-qrrdi667	102	2	remaining	remain	VBG
cord-005550-qrrdi667	102	3	alleles	allele	NNS
cord-005550-qrrdi667	102	4	were	be	VBD
cord-005550-qrrdi667	102	5	A1	A1	NNP
cord-005550-qrrdi667	102	6	(	(	-LRB-
cord-005550-qrrdi667	102	7	two	two	CD
cord-005550-qrrdi667	102	8	copies	copy	NNS
cord-005550-qrrdi667	102	9	)	)	-RRB-
cord-005550-qrrdi667	102	10	and	and	CC
cord-005550-qrrdi667	102	11	A7	A7	NNP
cord-005550-qrrdi667	102	12	(	(	-LRB-
cord-005550-qrrdi667	102	13	one	one	CD
cord-005550-qrrdi667	102	14	copy	copy	NN
cord-005550-qrrdi667	102	15	)	)	-RRB-
cord-005550-qrrdi667	102	16	.	.	.
cord-005550-qrrdi667	103	1	The	the	DT
cord-005550-qrrdi667	103	2	DNA	dna	NN
cord-005550-qrrdi667	103	3	sequence	sequence	NN
cord-005550-qrrdi667	103	4	consists	consist	VBZ
cord-005550-qrrdi667	103	5	of	of	IN
cord-005550-qrrdi667	103	6	224	224	CD
cord-005550-qrrdi667	103	7	base	base	NN
cord-005550-qrrdi667	103	8	pairs	pair	NNS
cord-005550-qrrdi667	103	9	of	of	IN
cord-005550-qrrdi667	103	10	which	which	WDT
cord-005550-qrrdi667	103	11	80	80	CD
cord-005550-qrrdi667	103	12	(	(	-LRB-
cord-005550-qrrdi667	103	13	35.7	35.7	CD
cord-005550-qrrdi667	103	14	%	%	NN
cord-005550-qrrdi667	103	15	)	)	-RRB-
cord-005550-qrrdi667	103	16	were	be	VBD
cord-005550-qrrdi667	103	17	polymorphic	polymorphic	JJ
cord-005550-qrrdi667	103	18	.	.	.
cord-005550-qrrdi667	104	1	On	on	IN
cord-005550-qrrdi667	104	2	average	average	JJ
cord-005550-qrrdi667	104	3	,	,	,
cord-005550-qrrdi667	104	4	alleles	allele	NNS
cord-005550-qrrdi667	104	5	differed	differ	VBN
cord-005550-qrrdi667	104	6	by	by	IN
cord-005550-qrrdi667	104	7	31.9	31.9	CD
cord-005550-qrrdi667	104	8	nucleotides	nucleotide	NNS
cord-005550-qrrdi667	104	9	(	(	-LRB-
cord-005550-qrrdi667	104	10	14.2	14.2	CD
cord-005550-qrrdi667	104	11	%	%	NN
cord-005550-qrrdi667	104	12	)	)	-RRB-
cord-005550-qrrdi667	104	13	.	.	.
cord-005550-qrrdi667	105	1	Alleles	Alleles	NNP
cord-005550-qrrdi667	105	2	A1	A1	NNP
cord-005550-qrrdi667	105	3	and	and	CC
cord-005550-qrrdi667	105	4	A7	A7	NNP
cord-005550-qrrdi667	105	5	differed	differ	VBD
cord-005550-qrrdi667	105	6	by	by	IN
cord-005550-qrrdi667	105	7	only	only	RB
cord-005550-qrrdi667	105	8	six	six	CD
cord-005550-qrrdi667	105	9	nucleotides	nucleotide	NNS
cord-005550-qrrdi667	105	10	(	(	-LRB-
cord-005550-qrrdi667	105	11	2.7	2.7	CD
cord-005550-qrrdi667	105	12	%	%	NN
cord-005550-qrrdi667	105	13	)	)	-RRB-
cord-005550-qrrdi667	105	14	.	.	.
cord-005550-qrrdi667	106	1	The	the	DT
cord-005550-qrrdi667	106	2	most	most	RBS
cord-005550-qrrdi667	106	3	distinct	distinct	JJ
cord-005550-qrrdi667	106	4	alleles	allele	NNS
cord-005550-qrrdi667	106	5	were	be	VBD
cord-005550-qrrdi667	106	6	A9	A9	NNP
cord-005550-qrrdi667	106	7	and	and	CC
cord-005550-qrrdi667	106	8	A11	A11	NNP
cord-005550-qrrdi667	106	9	,	,	,
cord-005550-qrrdi667	106	10	which	which	WDT
cord-005550-qrrdi667	106	11	differed	differ	VBD
cord-005550-qrrdi667	106	12	by	by	IN
cord-005550-qrrdi667	106	13	49	49	CD
cord-005550-qrrdi667	106	14	nucleotides	nucleotide	NNS
cord-005550-qrrdi667	106	15	(	(	-LRB-
cord-005550-qrrdi667	106	16	21.9	21.9	CD
cord-005550-qrrdi667	106	17	%	%	NN
cord-005550-qrrdi667	106	18	)	)	-RRB-
cord-005550-qrrdi667	106	19	.	.	.
cord-005550-qrrdi667	107	1	Pairwise	pairwise	JJ
cord-005550-qrrdi667	107	2	DNA	dna	NN
cord-005550-qrrdi667	107	3	sequence	sequence	NN
cord-005550-qrrdi667	107	4	differences	difference	NNS
cord-005550-qrrdi667	107	5	between	between	IN
cord-005550-qrrdi667	107	6	the	the	DT
cord-005550-qrrdi667	107	7	alleles	allele	NNS
cord-005550-qrrdi667	107	8	are	be	VBP
cord-005550-qrrdi667	107	9	given	give	VBN
cord-005550-qrrdi667	107	10	in	in	IN
cord-005550-qrrdi667	107	11	Table	Table	NNP
cord-005550-qrrdi667	107	12	1	1	CD
cord-005550-qrrdi667	107	13	.	.	.
cord-005550-qrrdi667	108	1	Allele	Allele	NNP
cord-005550-qrrdi667	108	2	A11	A11	NNP
cord-005550-qrrdi667	108	3	,	,	,
cord-005550-qrrdi667	108	4	which	which	WDT
cord-005550-qrrdi667	108	5	was	be	VBD
cord-005550-qrrdi667	108	6	only	only	RB
cord-005550-qrrdi667	108	7	found	find	VBN
cord-005550-qrrdi667	108	8	in	in	IN
cord-005550-qrrdi667	108	9	individuals	individual	NNS
cord-005550-qrrdi667	108	10	from	from	IN
cord-005550-qrrdi667	108	11	Panama	Panama	NNP
cord-005550-qrrdi667	108	12	,	,	,
cord-005550-qrrdi667	108	13	was	be	VBD
cord-005550-qrrdi667	108	14	not	not	RB
cord-005550-qrrdi667	108	15	more	more	RBR
cord-005550-qrrdi667	108	16	distinct	distinct	JJ
cord-005550-qrrdi667	108	17	to	to	IN
cord-005550-qrrdi667	108	18	alleles	allele	NNS
cord-005550-qrrdi667	108	19	from	from	IN
cord-005550-qrrdi667	108	20	Costa	Costa	NNP
cord-005550-qrrdi667	108	21	Rica	Rica	NNP
cord-005550-qrrdi667	108	22	than	than	IN
cord-005550-qrrdi667	108	23	alleles	allele	NNS
cord-005550-qrrdi667	108	24	from	from	IN
cord-005550-qrrdi667	108	25	Costa	Costa	NNP
cord-005550-qrrdi667	108	26	Rica	Rica	NNP
cord-005550-qrrdi667	108	27	to	to	IN
cord-005550-qrrdi667	108	28	each	each	DT
cord-005550-qrrdi667	108	29	other	other	JJ
cord-005550-qrrdi667	108	30	.	.	.
cord-005550-qrrdi667	109	1	Several	several	JJ
cord-005550-qrrdi667	109	2	alleles	allele	NNS
cord-005550-qrrdi667	109	3	share	share	VBP
cord-005550-qrrdi667	109	4	different	different	JJ
cord-005550-qrrdi667	109	5	sequence	sequence	NN
cord-005550-qrrdi667	109	6	fragments	fragment	NNS
cord-005550-qrrdi667	109	7	of	of	IN
cord-005550-qrrdi667	109	8	identical	identical	JJ
cord-005550-qrrdi667	109	9	sequence	sequence	NN
cord-005550-qrrdi667	109	10	(	(	-LRB-
cord-005550-qrrdi667	109	11	Figure	figure	NN
cord-005550-qrrdi667	109	12	1	1	CD
cord-005550-qrrdi667	109	13	)	)	-RRB-
cord-005550-qrrdi667	109	14	.	.	.
cord-005550-qrrdi667	110	1	These	these	DT
cord-005550-qrrdi667	110	2	shared	share	VBN
cord-005550-qrrdi667	110	3	fragments	fragment	NNS
cord-005550-qrrdi667	110	4	were	be	VBD
cord-005550-qrrdi667	110	5	significantly	significantly	RB
cord-005550-qrrdi667	110	6	similar	similar	JJ
cord-005550-qrrdi667	110	7	when	when	WRB
cord-005550-qrrdi667	110	8	the	the	DT
cord-005550-qrrdi667	110	9	global	global	JJ
cord-005550-qrrdi667	110	10	comparison	comparison	NN
cord-005550-qrrdi667	110	11	procedures	procedure	NNS
cord-005550-qrrdi667	110	12	and	and	CC
cord-005550-qrrdi667	110	13	permutation	permutation	NN
cord-005550-qrrdi667	110	14	test	test	NN
cord-005550-qrrdi667	110	15	implemented	implement	VBN
cord-005550-qrrdi667	110	16	in	in	IN
cord-005550-qrrdi667	110	17	the	the	DT
cord-005550-qrrdi667	110	18	software	software	NN
cord-005550-qrrdi667	110	19	GENECONV	GENECONV	NNP
cord-005550-qrrdi667	110	20	(	(	-LRB-
cord-005550-qrrdi667	110	21	Sawyer	Sawyer	NNP
cord-005550-qrrdi667	110	22	,	,	,
cord-005550-qrrdi667	110	23	1999	1999	CD
cord-005550-qrrdi667	110	24	)	)	-RRB-
cord-005550-qrrdi667	110	25	was	be	VBD
cord-005550-qrrdi667	110	26	applied	apply	VBN
cord-005550-qrrdi667	110	27	.	.	.
cord-005550-qrrdi667	111	1	Thus	thus	RB
cord-005550-qrrdi667	111	2	,	,	,
cord-005550-qrrdi667	111	3	some	some	DT
cord-005550-qrrdi667	111	4	alleles	allele	NNS
cord-005550-qrrdi667	111	5	are	be	VBP
cord-005550-qrrdi667	111	6	partly	partly	RB
cord-005550-qrrdi667	111	7	characterized	characterize	VBN
cord-005550-qrrdi667	111	8	by	by	IN
cord-005550-qrrdi667	111	9	different	different	JJ
cord-005550-qrrdi667	111	10	arrangements	arrangement	NNS
cord-005550-qrrdi667	111	11	of	of	IN
cord-005550-qrrdi667	111	12	these	these	DT
cord-005550-qrrdi667	111	13	fragments	fragment	NNS
cord-005550-qrrdi667	111	14	that	that	WDT
cord-005550-qrrdi667	111	15	is	be	VBZ
cord-005550-qrrdi667	111	16	likely	likely	JJ
cord-005550-qrrdi667	111	17	due	due	IN
cord-005550-qrrdi667	111	18	to	to	IN
cord-005550-qrrdi667	111	19	gene	gene	NN
cord-005550-qrrdi667	111	20	conversion	conversion	NN
cord-005550-qrrdi667	111	21	.	.	.
cord-005550-qrrdi667	112	1	The	the	DT
cord-005550-qrrdi667	112	2	two	two	CD
cord-005550-qrrdi667	112	3	most	most	RBS
cord-005550-qrrdi667	112	4	common	common	JJ
cord-005550-qrrdi667	112	5	alleles	allele	NNS
cord-005550-qrrdi667	112	6	(	(	-LRB-
cord-005550-qrrdi667	112	7	A1	A1	NNP
cord-005550-qrrdi667	112	8	and	and	CC
cord-005550-qrrdi667	112	9	A6	A6	NNP
cord-005550-qrrdi667	112	10	)	)	-RRB-
cord-005550-qrrdi667	112	11	are	be	VBP
cord-005550-qrrdi667	112	12	distinct	distinct	JJ
cord-005550-qrrdi667	112	13	at	at	IN
cord-005550-qrrdi667	112	14	15.5	15.5	CD
cord-005550-qrrdi667	112	15	%	%	NN
cord-005550-qrrdi667	112	16	of	of	IN
cord-005550-qrrdi667	112	17	all	all	DT
cord-005550-qrrdi667	112	18	nucleotide	nucleotide	JJ
cord-005550-qrrdi667	112	19	sites	site	NNS
cord-005550-qrrdi667	112	20	and	and	CC
cord-005550-qrrdi667	112	21	do	do	VBP
cord-005550-qrrdi667	112	22	not	not	RB
cord-005550-qrrdi667	112	23	share	share	VB
cord-005550-qrrdi667	112	24	an	an	DT
cord-005550-qrrdi667	112	25	obvious	obvious	JJ
cord-005550-qrrdi667	112	26	sequence	sequence	NN
cord-005550-qrrdi667	112	27	fragment	fragment	NN
cord-005550-qrrdi667	112	28	.	.	.
cord-005550-qrrdi667	113	1	The	the	DT
cord-005550-qrrdi667	113	2	two	two	CD
cord-005550-qrrdi667	113	3	B	b	NN
cord-005550-qrrdi667	113	4	sequence	sequence	NN
cord-005550-qrrdi667	113	5	types	type	NNS
cord-005550-qrrdi667	113	6	do	do	VBP
cord-005550-qrrdi667	113	7	not	not	RB
cord-005550-qrrdi667	113	8	share	share	VB
cord-005550-qrrdi667	113	9	a	a	DT
cord-005550-qrrdi667	113	10	sequence	sequence	NN
cord-005550-qrrdi667	113	11	fragment	fragment	NN
cord-005550-qrrdi667	113	12	with	with	IN
cord-005550-qrrdi667	113	13	each	each	DT
cord-005550-qrrdi667	113	14	other	other	JJ
cord-005550-qrrdi667	113	15	.	.	.
cord-005550-qrrdi667	114	1	Only	only	RB
cord-005550-qrrdi667	114	2	the	the	DT
cord-005550-qrrdi667	114	3	sequence	sequence	NN
cord-005550-qrrdi667	114	4	type	type	NN
cord-005550-qrrdi667	114	5	B2	b2	NN
cord-005550-qrrdi667	114	6	has	have	VBZ
cord-005550-qrrdi667	114	7	a	a	DT
cord-005550-qrrdi667	114	8	short	short	JJ
cord-005550-qrrdi667	114	9	sequence	sequence	NN
cord-005550-qrrdi667	114	10	fragment	fragment	NN
cord-005550-qrrdi667	114	11	in	in	IN
cord-005550-qrrdi667	114	12	common	common	JJ
cord-005550-qrrdi667	114	13	with	with	IN
cord-005550-qrrdi667	114	14	the	the	DT
cord-005550-qrrdi667	114	15	A4	A4	NNP
cord-005550-qrrdi667	114	16	allele	allele	NN
cord-005550-qrrdi667	114	17	.	.	.
cord-005550-qrrdi667	115	1	In	in	IN
cord-005550-qrrdi667	115	2	contrast	contrast	NN
cord-005550-qrrdi667	115	3	,	,	,
cord-005550-qrrdi667	115	4	most	most	JJS
cord-005550-qrrdi667	115	5	DRB1	DRB1	NNP
cord-005550-qrrdi667	115	6	alleles	allele	NNS
cord-005550-qrrdi667	115	7	(	(	-LRB-
cord-005550-qrrdi667	115	8	that	that	DT
cord-005550-qrrdi667	115	9	is	be	VBZ
cord-005550-qrrdi667	115	10	A	a	DT
cord-005550-qrrdi667	115	11	sequence	sequence	NN
cord-005550-qrrdi667	115	12	types	type	NNS
cord-005550-qrrdi667	115	13	)	)	-RRB-
cord-005550-qrrdi667	115	14	are	be	VBP
cord-005550-qrrdi667	115	15	linked	link	VBN
cord-005550-qrrdi667	115	16	by	by	IN
cord-005550-qrrdi667	115	17	shared	share	VBN
cord-005550-qrrdi667	115	18	sequence	sequence	NN
cord-005550-qrrdi667	115	19	fragments	fragment	NNS
cord-005550-qrrdi667	115	20	.	.	.
cord-005550-qrrdi667	116	1	This	this	DT
cord-005550-qrrdi667	116	2	corroborates	corroborate	VBZ
cord-005550-qrrdi667	116	3	our	-PRON-	PRP$
cord-005550-qrrdi667	116	4	view	view	NN
cord-005550-qrrdi667	116	5	that	that	IN
cord-005550-qrrdi667	116	6	B	b	NN
cord-005550-qrrdi667	116	7	sequence	sequence	NN
cord-005550-qrrdi667	116	8	types	type	NNS
cord-005550-qrrdi667	116	9	are	be	VBP
cord-005550-qrrdi667	116	10	not	not	RB
cord-005550-qrrdi667	116	11	alleles	allele	NNS
cord-005550-qrrdi667	116	12	of	of	IN
cord-005550-qrrdi667	116	13	the	the	DT
cord-005550-qrrdi667	116	14	DRB1	DRB1	NNP
cord-005550-qrrdi667	116	15	locus	locus	NN
cord-005550-qrrdi667	116	16	.	.	.
cord-005550-qrrdi667	117	1	Evidence	evidence	NN
cord-005550-qrrdi667	117	2	for	for	IN
cord-005550-qrrdi667	117	3	balancing	balance	VBG
cord-005550-qrrdi667	117	4	selection	selection	NN
cord-005550-qrrdi667	117	5	DRB1	DRB1	NNP
cord-005550-qrrdi667	117	6	alleles	allele	NNS
cord-005550-qrrdi667	117	7	differed	differ	VBD
cord-005550-qrrdi667	117	8	in	in	IN
cord-005550-qrrdi667	117	9	3	3	CD
cord-005550-qrrdi667	117	10	(	(	-LRB-
cord-005550-qrrdi667	117	11	4	4	CD
cord-005550-qrrdi667	117	12	%	%	NN
cord-005550-qrrdi667	117	13	)	)	-RRB-
cord-005550-qrrdi667	117	14	to	to	IN
cord-005550-qrrdi667	117	15	31	31	CD
cord-005550-qrrdi667	117	16	(	(	-LRB-
cord-005550-qrrdi667	117	17	42	42	CD
cord-005550-qrrdi667	117	18	%	%	NN
cord-005550-qrrdi667	117	19	)	)	-RRB-
cord-005550-qrrdi667	117	20	of	of	IN
cord-005550-qrrdi667	117	21	all	all	DT
cord-005550-qrrdi667	117	22	74	74	CD
cord-005550-qrrdi667	117	23	amino	amino	NN
cord-005550-qrrdi667	117	24	-	-	HYPH
cord-005550-qrrdi667	117	25	acid	acid	NN
cord-005550-qrrdi667	117	26	sites	site	NNS
cord-005550-qrrdi667	117	27	studied	study	VBD
cord-005550-qrrdi667	117	28	.	.	.
cord-005550-qrrdi667	118	1	Up	up	IN
cord-005550-qrrdi667	118	2	to	to	TO
cord-005550-qrrdi667	118	3	five	five	CD
cord-005550-qrrdi667	118	4	different	different	JJ
cord-005550-qrrdi667	118	5	amino	amino	NN
cord-005550-qrrdi667	118	6	acids	acid	NNS
cord-005550-qrrdi667	118	7	were	be	VBD
cord-005550-qrrdi667	118	8	found	find	VBN
cord-005550-qrrdi667	118	9	at	at	IN
cord-005550-qrrdi667	118	10	a	a	DT
cord-005550-qrrdi667	118	11	single	single	JJ
cord-005550-qrrdi667	118	12	codon	codon	NN
cord-005550-qrrdi667	118	13	position	position	NN
cord-005550-qrrdi667	118	14	and	and	CC
cord-005550-qrrdi667	118	15	44	44	CD
cord-005550-qrrdi667	118	16	(	(	-LRB-
cord-005550-qrrdi667	118	17	59.5	59.5	CD
cord-005550-qrrdi667	118	18	%	%	NN
cord-005550-qrrdi667	118	19	)	)	-RRB-
cord-005550-qrrdi667	119	1	amino	amino	NN
cord-005550-qrrdi667	119	2	-	-	HYPH
cord-005550-qrrdi667	119	3	acid	acid	NN
cord-005550-qrrdi667	119	4	sites	site	NNS
cord-005550-qrrdi667	119	5	were	be	VBD
cord-005550-qrrdi667	119	6	variable	variable	JJ
cord-005550-qrrdi667	119	7	(	(	-LRB-
cord-005550-qrrdi667	119	8	Figure	figure	NN
cord-005550-qrrdi667	119	9	2	2	CD
cord-005550-qrrdi667	119	10	)	)	-RRB-
cord-005550-qrrdi667	119	11	.	.	.
cord-005550-qrrdi667	120	1	In	in	IN
cord-005550-qrrdi667	120	2	humans	human	NNS
cord-005550-qrrdi667	120	3	,	,	,
cord-005550-qrrdi667	120	4	18	18	CD
cord-005550-qrrdi667	120	5	amino	amino	NN
cord-005550-qrrdi667	120	6	-	-	HYPH
cord-005550-qrrdi667	120	7	acid	acid	NN
cord-005550-qrrdi667	120	8	positions	position	NNS
cord-005550-qrrdi667	120	9	,	,	,
cord-005550-qrrdi667	120	10	located	locate	VBN
cord-005550-qrrdi667	120	11	within	within	IN
cord-005550-qrrdi667	120	12	the	the	DT
cord-005550-qrrdi667	120	13	investigated	investigate	VBN
cord-005550-qrrdi667	120	14	sequence	sequence	NN
cord-005550-qrrdi667	120	15	region	region	NN
cord-005550-qrrdi667	120	16	,	,	,
cord-005550-qrrdi667	120	17	are	be	VBP
cord-005550-qrrdi667	120	18	documented	document	VBN
cord-005550-qrrdi667	120	19	to	to	TO
cord-005550-qrrdi667	120	20	be	be	VB
cord-005550-qrrdi667	120	21	important	important	JJ
cord-005550-qrrdi667	120	22	in	in	IN
cord-005550-qrrdi667	120	23	the	the	DT
cord-005550-qrrdi667	120	24	ABS	abs	NN
cord-005550-qrrdi667	120	25	using	use	VBG
cord-005550-qrrdi667	120	26	X	x	NN
cord-005550-qrrdi667	120	27	-	-	NN
cord-005550-qrrdi667	120	28	ray	ray	NN
cord-005550-qrrdi667	120	29	crystallography	crystallography	NN
cord-005550-qrrdi667	120	30	(	(	-LRB-
cord-005550-qrrdi667	120	31	Brown	Brown	NNP
cord-005550-qrrdi667	120	32	et	et	NNP
cord-005550-qrrdi667	120	33	al	al	NNP
cord-005550-qrrdi667	120	34	.	.	NNP
cord-005550-qrrdi667	120	35	,	,	,
cord-005550-qrrdi667	120	36	1993	1993	CD
cord-005550-qrrdi667	120	37	)	)	-RRB-
cord-005550-qrrdi667	120	38	.	.	.
cord-005550-qrrdi667	121	1	Most	Most	JJS
cord-005550-qrrdi667	121	2	variation	variation	NN
cord-005550-qrrdi667	121	3	was	be	VBD
cord-005550-qrrdi667	121	4	concentrated	concentrate	VBN
cord-005550-qrrdi667	121	5	at	at	IN
cord-005550-qrrdi667	121	6	these	these	DT
cord-005550-qrrdi667	121	7	ABSs	abs	NNS
cord-005550-qrrdi667	121	8	.	.	.
cord-005550-qrrdi667	122	1	The	the	DT
cord-005550-qrrdi667	122	2	proportion	proportion	NN
cord-005550-qrrdi667	122	3	of	of	IN
cord-005550-qrrdi667	122	4	variable	variable	JJ
cord-005550-qrrdi667	122	5	positions	position	NNS
cord-005550-qrrdi667	122	6	was	be	VBD
cord-005550-qrrdi667	122	7	significantly	significantly	RB
cord-005550-qrrdi667	122	8	higher	high	JJR
cord-005550-qrrdi667	122	9	at	at	IN
cord-005550-qrrdi667	122	10	ABS	abs	NN
cord-005550-qrrdi667	122	11	than	than	IN
cord-005550-qrrdi667	122	12	at	at	IN
cord-005550-qrrdi667	122	13	nonantigen	nonantigen	NN
cord-005550-qrrdi667	122	14	-	-	HYPH
cord-005550-qrrdi667	122	15	binding	bind	VBG
cord-005550-qrrdi667	122	16	sites	site	NNS
cord-005550-qrrdi667	122	17	(	(	-LRB-
cord-005550-qrrdi667	122	18	non	non	JJ
cord-005550-qrrdi667	122	19	-	-	JJ
cord-005550-qrrdi667	122	20	ABSs	abss	JJ
cord-005550-qrrdi667	122	21	)	)	-RRB-
cord-005550-qrrdi667	122	22	(	(	-LRB-
cord-005550-qrrdi667	122	23	15	15	CD
cord-005550-qrrdi667	122	24	of	of	IN
cord-005550-qrrdi667	122	25	18	18	CD
cord-005550-qrrdi667	122	26	(	(	-LRB-
cord-005550-qrrdi667	122	27	83	83	CD
cord-005550-qrrdi667	122	28	%	%	NN
cord-005550-qrrdi667	122	29	)	)	-RRB-
cord-005550-qrrdi667	122	30	and	and	CC
cord-005550-qrrdi667	122	31	29	29	CD
cord-005550-qrrdi667	122	32	of	of	IN
cord-005550-qrrdi667	122	33	56	56	CD
cord-005550-qrrdi667	122	34	(	(	-LRB-
cord-005550-qrrdi667	122	35	52	52	CD
cord-005550-qrrdi667	122	36	%	%	NN
cord-005550-qrrdi667	122	37	)	)	-RRB-
cord-005550-qrrdi667	122	38	,	,	,
cord-005550-qrrdi667	122	39	respectively	respectively	RB
cord-005550-qrrdi667	122	40	,	,	,
cord-005550-qrrdi667	122	41	w	w	NN
cord-005550-qrrdi667	122	42	2	2	CD
cord-005550-qrrdi667	122	43	¼	¼	CD
cord-005550-qrrdi667	122	44	5.6	5.6	CD
cord-005550-qrrdi667	122	45	,	,	,
cord-005550-qrrdi667	122	46	Po0.05	Po0.05	NNP
cord-005550-qrrdi667	122	47	)	)	-RRB-
cord-005550-qrrdi667	122	48	.	.	.
cord-005550-qrrdi667	123	1	Using	use	VBG
cord-005550-qrrdi667	123	2	the	the	DT
cord-005550-qrrdi667	123	3	observed	observed	JJ
cord-005550-qrrdi667	123	4	allele	allele	NN
cord-005550-qrrdi667	123	5	frequencies	frequency	NNS
cord-005550-qrrdi667	123	6	in	in	IN
cord-005550-qrrdi667	123	7	the	the	DT
cord-005550-qrrdi667	123	8	Costa	Costa	NNP
cord-005550-qrrdi667	123	9	Rica	Rica	NNP
cord-005550-qrrdi667	123	10	sample	sample	NN
cord-005550-qrrdi667	123	11	,	,	,
cord-005550-qrrdi667	123	12	we	-PRON-	PRP
cord-005550-qrrdi667	123	13	calculated	calculate	VBD
cord-005550-qrrdi667	123	14	the	the	DT
cord-005550-qrrdi667	123	15	heterozygosity	heterozygosity	NN
cord-005550-qrrdi667	123	16	for	for	IN
cord-005550-qrrdi667	123	17	each	each	DT
cord-005550-qrrdi667	123	18	amino	amino	NN
cord-005550-qrrdi667	123	19	-	-	HYPH
cord-005550-qrrdi667	123	20	acid	acid	NN
cord-005550-qrrdi667	123	21	position	position	NN
cord-005550-qrrdi667	123	22	(	(	-LRB-
cord-005550-qrrdi667	123	23	Figure	figure	NN
cord-005550-qrrdi667	123	24	4	4	CD
cord-005550-qrrdi667	123	25	)	)	-RRB-
cord-005550-qrrdi667	123	26	.	.	.
cord-005550-qrrdi667	124	1	Figure	figure	NN
cord-005550-qrrdi667	124	2	4	4	CD
cord-005550-qrrdi667	124	3	Average	average	JJ
cord-005550-qrrdi667	124	4	heterozygosity	heterozygosity	NN
cord-005550-qrrdi667	124	5	for	for	IN
cord-005550-qrrdi667	124	6	individual	individual	JJ
cord-005550-qrrdi667	124	7	amino	amino	NN
cord-005550-qrrdi667	124	8	-	-	HYPH
cord-005550-qrrdi667	124	9	acid	acid	NN
cord-005550-qrrdi667	124	10	positions	position	NNS
cord-005550-qrrdi667	124	11	calculated	calculate	VBN
cord-005550-qrrdi667	124	12	from	from	IN
cord-005550-qrrdi667	124	13	the	the	DT
cord-005550-qrrdi667	124	14	allele	allele	NN
cord-005550-qrrdi667	124	15	frequencies	frequency	NNS
cord-005550-qrrdi667	124	16	in	in	IN
cord-005550-qrrdi667	124	17	the	the	DT
cord-005550-qrrdi667	124	18	Costa	Costa	NNP
cord-005550-qrrdi667	124	19	Rica	Rica	NNP
cord-005550-qrrdi667	124	20	sample	sample	NN
cord-005550-qrrdi667	124	21	.	.	.
cord-005550-qrrdi667	125	1	Asterisks	asterisk	NNS
cord-005550-qrrdi667	125	2	indicate	indicate	VBP
cord-005550-qrrdi667	125	3	positions	position	NNS
cord-005550-qrrdi667	125	4	,	,	,
cord-005550-qrrdi667	125	5	which	which	WDT
cord-005550-qrrdi667	125	6	are	be	VBP
cord-005550-qrrdi667	125	7	assumed	assume	VBN
cord-005550-qrrdi667	125	8	to	to	TO
cord-005550-qrrdi667	125	9	be	be	VB
cord-005550-qrrdi667	125	10	antigenbinding	antigenbinde	VBG
cord-005550-qrrdi667	125	11	sites	site	NNS
cord-005550-qrrdi667	125	12	in	in	IN
cord-005550-qrrdi667	125	13	humans	human	NNS
cord-005550-qrrdi667	125	14	(	(	-LRB-
cord-005550-qrrdi667	125	15	according	accord	VBG
cord-005550-qrrdi667	125	16	to	to	IN
cord-005550-qrrdi667	125	17	Brown	Brown	NNP
cord-005550-qrrdi667	125	18	et	et	NNP
cord-005550-qrrdi667	125	19	al	al	NNP
cord-005550-qrrdi667	125	20	.	.	NNP
cord-005550-qrrdi667	125	21	,	,	,
cord-005550-qrrdi667	125	22	1993	1993	CD
cord-005550-qrrdi667	125	23	)	)	-RRB-
cord-005550-qrrdi667	125	24	.	.	.
cord-005550-qrrdi667	126	1	MHC	MHC	NNP
cord-005550-qrrdi667	126	2	diversity	diversity	NN
cord-005550-qrrdi667	126	3	in	in	IN
cord-005550-qrrdi667	126	4	the	the	DT
cord-005550-qrrdi667	126	5	sac	sac	NN
cord-005550-qrrdi667	126	6	-	-	HYPH
cord-005550-qrrdi667	126	7	winged	winged	JJ
cord-005550-qrrdi667	126	8	bat	bat	NN
cord-005550-qrrdi667	127	1	The	the	DT
cord-005550-qrrdi667	127	2	highest	high	JJS
cord-005550-qrrdi667	127	3	heterozygosity	heterozygosity	NN
cord-005550-qrrdi667	127	4	was	be	VBD
cord-005550-qrrdi667	127	5	found	find	VBN
cord-005550-qrrdi667	127	6	at	at	IN
cord-005550-qrrdi667	127	7	position	position	NN
cord-005550-qrrdi667	127	8	74	74	CD
cord-005550-qrrdi667	127	9	(	(	-LRB-
cord-005550-qrrdi667	127	10	0.319	0.319	CD
cord-005550-qrrdi667	127	11	)	)	-RRB-
cord-005550-qrrdi667	127	12	.	.	.
cord-005550-qrrdi667	128	1	An	an	DT
cord-005550-qrrdi667	128	2	average	average	JJ
cord-005550-qrrdi667	128	3	heterozygosity	heterozygosity	NN
cord-005550-qrrdi667	128	4	of	of	IN
cord-005550-qrrdi667	128	5	0.153	0.153	CD
cord-005550-qrrdi667	128	6	was	be	VBD
cord-005550-qrrdi667	128	7	observed	observe	VBN
cord-005550-qrrdi667	128	8	at	at	IN
cord-005550-qrrdi667	128	9	ABS	abs	NN
cord-005550-qrrdi667	128	10	,	,	,
cord-005550-qrrdi667	128	11	which	which	WDT
cord-005550-qrrdi667	128	12	was	be	VBD
cord-005550-qrrdi667	128	13	significantly	significantly	RB
cord-005550-qrrdi667	128	14	higher	high	JJR
cord-005550-qrrdi667	128	15	than	than	IN
cord-005550-qrrdi667	128	16	the	the	DT
cord-005550-qrrdi667	128	17	average	average	JJ
cord-005550-qrrdi667	128	18	heterozygosity	heterozygosity	NN
cord-005550-qrrdi667	128	19	at	at	IN
cord-005550-qrrdi667	128	20	non	non	JJ
cord-005550-qrrdi667	128	21	-	-	JJ
cord-005550-qrrdi667	128	22	ABSs	abss	JJ
cord-005550-qrrdi667	128	23	(	(	-LRB-
cord-005550-qrrdi667	128	24	0.066	0.066	CD
cord-005550-qrrdi667	128	25	,	,	,
cord-005550-qrrdi667	128	26	U	U	NNP
cord-005550-qrrdi667	128	27	¼	¼	NNP
cord-005550-qrrdi667	128	28	266	266	CD
cord-005550-qrrdi667	128	29	,	,	,
cord-005550-qrrdi667	128	30	P	p	NN
cord-005550-qrrdi667	128	31	¼	¼	CD
cord-005550-qrrdi667	128	32	0.002	0.002	CD
cord-005550-qrrdi667	128	33	)	)	-RRB-
cord-005550-qrrdi667	128	34	.	.	.
cord-005550-qrrdi667	129	1	The	the	DT
cord-005550-qrrdi667	129	2	maximum	maximum	NN
cord-005550-qrrdi667	129	3	of	of	IN
cord-005550-qrrdi667	129	4	five	five	CD
cord-005550-qrrdi667	129	5	different	different	JJ
cord-005550-qrrdi667	129	6	amino	amino	NN
cord-005550-qrrdi667	129	7	-	-	HYPH
cord-005550-qrrdi667	129	8	acid	acid	NN
cord-005550-qrrdi667	129	9	substitutions	substitution	NNS
cord-005550-qrrdi667	129	10	was	be	VBD
cord-005550-qrrdi667	129	11	only	only	RB
cord-005550-qrrdi667	129	12	found	find	VBN
cord-005550-qrrdi667	129	13	at	at	IN
cord-005550-qrrdi667	129	14	two	two	CD
cord-005550-qrrdi667	129	15	ABS	abs	NN
cord-005550-qrrdi667	129	16	(	(	-LRB-
cord-005550-qrrdi667	129	17	positions	position	NNS
cord-005550-qrrdi667	129	18	71	71	CD
cord-005550-qrrdi667	129	19	and	and	CC
cord-005550-qrrdi667	129	20	74	74	CD
cord-005550-qrrdi667	129	21	,	,	,
cord-005550-qrrdi667	129	22	Figure	figure	NN
cord-005550-qrrdi667	129	23	2	2	CD
cord-005550-qrrdi667	129	24	)	)	-RRB-
cord-005550-qrrdi667	129	25	.	.	.
cord-005550-qrrdi667	130	1	Higher	high	JJR
cord-005550-qrrdi667	130	2	divergence	divergence	NN
cord-005550-qrrdi667	130	3	among	among	IN
cord-005550-qrrdi667	130	4	amino	amino	NN
cord-005550-qrrdi667	130	5	-	-	HYPH
cord-005550-qrrdi667	130	6	acid	acid	NN
cord-005550-qrrdi667	130	7	sequences	sequence	NNS
cord-005550-qrrdi667	130	8	than	than	IN
cord-005550-qrrdi667	130	9	among	among	IN
cord-005550-qrrdi667	130	10	DNA	dna	NN
cord-005550-qrrdi667	130	11	sequences	sequence	NNS
cord-005550-qrrdi667	130	12	,	,	,
cord-005550-qrrdi667	130	13	and	and	CC
cord-005550-qrrdi667	130	14	the	the	DT
cord-005550-qrrdi667	130	15	concentration	concentration	NN
cord-005550-qrrdi667	130	16	of	of	IN
cord-005550-qrrdi667	130	17	aminoacid	aminoacid	JJ
cord-005550-qrrdi667	130	18	variation	variation	NN
cord-005550-qrrdi667	130	19	at	at	IN
cord-005550-qrrdi667	130	20	ABS	abs	NN
cord-005550-qrrdi667	130	21	indicate	indicate	VBP
cord-005550-qrrdi667	130	22	the	the	DT
cord-005550-qrrdi667	130	23	existence	existence	NN
cord-005550-qrrdi667	130	24	of	of	IN
cord-005550-qrrdi667	130	25	balancing	balance	VBG
cord-005550-qrrdi667	130	26	selection	selection	NN
cord-005550-qrrdi667	130	27	.	.	.
cord-005550-qrrdi667	131	1	Additional	additional	JJ
cord-005550-qrrdi667	131	2	evidence	evidence	NN
cord-005550-qrrdi667	131	3	comes	come	VBZ
cord-005550-qrrdi667	131	4	from	from	IN
cord-005550-qrrdi667	131	5	the	the	DT
cord-005550-qrrdi667	131	6	comparison	comparison	NN
cord-005550-qrrdi667	131	7	of	of	IN
cord-005550-qrrdi667	131	8	the	the	DT
cord-005550-qrrdi667	131	9	relative	relative	JJ
cord-005550-qrrdi667	131	10	rate	rate	NN
cord-005550-qrrdi667	131	11	of	of	IN
cord-005550-qrrdi667	131	12	nonsynonymous	nonsynonymous	NNP
cord-005550-qrrdi667	131	13	(	(	-LRB-
cord-005550-qrrdi667	131	14	d	d	NNP
cord-005550-qrrdi667	131	15	N	N	NNP
cord-005550-qrrdi667	131	16	)	)	-RRB-
cord-005550-qrrdi667	131	17	and	and	CC
cord-005550-qrrdi667	131	18	synonymous	synonymous	JJ
cord-005550-qrrdi667	131	19	(	(	-LRB-
cord-005550-qrrdi667	131	20	d	d	NNP
cord-005550-qrrdi667	131	21	S	S	NNP
cord-005550-qrrdi667	131	22	)	)	-RRB-
cord-005550-qrrdi667	131	23	substitutions	substitution	NNS
cord-005550-qrrdi667	131	24	at	at	IN
cord-005550-qrrdi667	131	25	the	the	DT
cord-005550-qrrdi667	131	26	18	18	CD
cord-005550-qrrdi667	131	27	antigen	antigen	NN
cord-005550-qrrdi667	131	28	(	(	-LRB-
cord-005550-qrrdi667	131	29	ABS	abs	NN
cord-005550-qrrdi667	131	30	)	)	-RRB-
cord-005550-qrrdi667	131	31	and	and	CC
cord-005550-qrrdi667	131	32	the	the	DT
cord-005550-qrrdi667	131	33	56	56	CD
cord-005550-qrrdi667	131	34	nonantigen	nonantigen	NN
cord-005550-qrrdi667	131	35	-	-	HYPH
cord-005550-qrrdi667	131	36	binding	bind	VBG
cord-005550-qrrdi667	131	37	amino	amino	NN
cord-005550-qrrdi667	131	38	-	-	HYPH
cord-005550-qrrdi667	131	39	acid	acid	NN
cord-005550-qrrdi667	131	40	sites	site	NNS
cord-005550-qrrdi667	131	41	.	.	.
cord-005550-qrrdi667	132	1	At	at	IN
cord-005550-qrrdi667	132	2	ABS	abs	NN
cord-005550-qrrdi667	132	3	,	,	,
cord-005550-qrrdi667	132	4	nonsynonymous	nonsynonymous	JJ
cord-005550-qrrdi667	132	5	substitutions	substitution	NNS
cord-005550-qrrdi667	132	6	(	(	-LRB-
cord-005550-qrrdi667	132	7	d	d	NNP
cord-005550-qrrdi667	132	8	N	N	NNP
cord-005550-qrrdi667	132	9	¼	¼	NNP
cord-005550-qrrdi667	132	10	0.37970.180	0.37970.180	CD
cord-005550-qrrdi667	132	11	)	)	-RRB-
cord-005550-qrrdi667	132	12	were	be	VBD
cord-005550-qrrdi667	132	13	on	on	IN
cord-005550-qrrdi667	132	14	average	average	JJ
cord-005550-qrrdi667	132	15	2.4	2.4	CD
cord-005550-qrrdi667	132	16	times	time	NNS
cord-005550-qrrdi667	132	17	more	more	RBR
cord-005550-qrrdi667	132	18	frequent	frequent	JJ
cord-005550-qrrdi667	132	19	(	(	-LRB-
cord-005550-qrrdi667	132	20	Z	Z	NNP
cord-005550-qrrdi667	132	21	¼	¼	NNP
cord-005550-qrrdi667	132	22	À6.33	À6.33	NNP
cord-005550-qrrdi667	132	23	,	,	,
cord-005550-qrrdi667	132	24	N	N	NNP
cord-005550-qrrdi667	132	25	¼	¼	CD
cord-005550-qrrdi667	132	26	55	55	CD
cord-005550-qrrdi667	132	27	,	,	,
cord-005550-qrrdi667	132	28	Po0.001	Po0.001	NNP
cord-005550-qrrdi667	132	29	)	)	-RRB-
cord-005550-qrrdi667	132	30	than	than	IN
cord-005550-qrrdi667	132	31	synonymous	synonymous	JJ
cord-005550-qrrdi667	132	32	substitutions	substitution	NNS
cord-005550-qrrdi667	132	33	(	(	-LRB-
cord-005550-qrrdi667	132	34	d	d	NNP
cord-005550-qrrdi667	132	35	S	S	NNP
cord-005550-qrrdi667	132	36	¼	¼	CD
cord-005550-qrrdi667	132	37	0.16070.093	0.16070.093	CD
cord-005550-qrrdi667	132	38	)	)	-RRB-
cord-005550-qrrdi667	132	39	.	.	.
cord-005550-qrrdi667	133	1	The	the	DT
cord-005550-qrrdi667	133	2	corresponding	corresponding	JJ
cord-005550-qrrdi667	133	3	ratio	ratio	NN
cord-005550-qrrdi667	133	4	d	d	NN
cord-005550-qrrdi667	133	5	N	N	NNP
cord-005550-qrrdi667	133	6	/d	/d	NNP
cord-005550-qrrdi667	133	7	S	S	NNP
cord-005550-qrrdi667	133	8	at	at	IN
cord-005550-qrrdi667	133	9	non	non	JJ
cord-005550-qrrdi667	133	10	-	-	JJ
cord-005550-qrrdi667	133	11	ABSs	abss	JJ
cord-005550-qrrdi667	133	12	was	be	VBD
cord-005550-qrrdi667	133	13	reverse	reverse	JJ
cord-005550-qrrdi667	133	14	(	(	-LRB-
cord-005550-qrrdi667	133	15	0.65	0.65	CD
cord-005550-qrrdi667	133	16	)	)	-RRB-
cord-005550-qrrdi667	133	17	and	and	CC
cord-005550-qrrdi667	133	18	also	also	RB
cord-005550-qrrdi667	133	19	both	both	DT
cord-005550-qrrdi667	133	20	substitution	substitution	NN
cord-005550-qrrdi667	134	1	According	accord	VBG
cord-005550-qrrdi667	134	2	to	to	IN
cord-005550-qrrdi667	134	3	the	the	DT
cord-005550-qrrdi667	134	4	overdominance	overdominance	NN
cord-005550-qrrdi667	134	5	hypothesis	hypothesis	NN
cord-005550-qrrdi667	134	6	and	and	CC
cord-005550-qrrdi667	134	7	the	the	DT
cord-005550-qrrdi667	134	8	negative	negative	JJ
cord-005550-qrrdi667	134	9	frequency	frequency	NN
cord-005550-qrrdi667	134	10	-	-	HYPH
cord-005550-qrrdi667	134	11	dependent	dependent	JJ
cord-005550-qrrdi667	134	12	selection	selection	NN
cord-005550-qrrdi667	134	13	hypothesis	hypothesis	NN
cord-005550-qrrdi667	134	14	,	,	,
cord-005550-qrrdi667	134	15	balancing	balance	VBG
cord-005550-qrrdi667	134	16	selection	selection	NN
cord-005550-qrrdi667	134	17	should	should	MD
cord-005550-qrrdi667	134	18	lead	lead	VB
cord-005550-qrrdi667	134	19	to	to	IN
cord-005550-qrrdi667	134	20	a	a	DT
cord-005550-qrrdi667	134	21	more	more	RBR
cord-005550-qrrdi667	134	22	even	even	JJ
cord-005550-qrrdi667	134	23	distribution	distribution	NN
cord-005550-qrrdi667	134	24	of	of	IN
cord-005550-qrrdi667	134	25	allele	allele	NN
cord-005550-qrrdi667	134	26	frequencies	frequency	NNS
cord-005550-qrrdi667	134	27	than	than	IN
cord-005550-qrrdi667	134	28	under	under	IN
cord-005550-qrrdi667	134	29	neutral	neutral	JJ
cord-005550-qrrdi667	134	30	expectation	expectation	NN
cord-005550-qrrdi667	134	31	,	,	,
cord-005550-qrrdi667	134	32	which	which	WDT
cord-005550-qrrdi667	134	33	can	can	MD
cord-005550-qrrdi667	134	34	be	be	VB
cord-005550-qrrdi667	134	35	tested	test	VBN
cord-005550-qrrdi667	134	36	by	by	IN
cord-005550-qrrdi667	134	37	the	the	DT
cord-005550-qrrdi667	134	38	commonly	commonly	RB
cord-005550-qrrdi667	134	39	used	use	VBN
cord-005550-qrrdi667	134	40	Ewens	Ewens	NNP
cord-005550-qrrdi667	134	41	-	-	HYPH
cord-005550-qrrdi667	134	42	Watterson	Watterson	NNP
cord-005550-qrrdi667	134	43	homozygosity	homozygosity	NN
cord-005550-qrrdi667	134	44	test	test	NN
cord-005550-qrrdi667	134	45	of	of	IN
cord-005550-qrrdi667	134	46	neutrality	neutrality	NN
cord-005550-qrrdi667	134	47	.	.	.
cord-005550-qrrdi667	135	1	We	-PRON-	PRP
cord-005550-qrrdi667	135	2	did	do	VBD
cord-005550-qrrdi667	135	3	not	not	RB
cord-005550-qrrdi667	135	4	find	find	VB
cord-005550-qrrdi667	135	5	a	a	DT
cord-005550-qrrdi667	135	6	significant	significant	JJ
cord-005550-qrrdi667	135	7	deviation	deviation	NN
cord-005550-qrrdi667	135	8	from	from	IN
cord-005550-qrrdi667	135	9	neutral	neutral	JJ
cord-005550-qrrdi667	135	10	expectation	expectation	NN
cord-005550-qrrdi667	135	11	(	(	-LRB-
cord-005550-qrrdi667	135	12	P	p	NN
cord-005550-qrrdi667	135	13	¼	¼	CD
cord-005550-qrrdi667	135	14	0.99	0.99	CD
cord-005550-qrrdi667	135	15	)	)	-RRB-
cord-005550-qrrdi667	135	16	by	by	IN
cord-005550-qrrdi667	135	17	applying	apply	VBG
cord-005550-qrrdi667	135	18	this	this	DT
cord-005550-qrrdi667	135	19	test	test	NN
cord-005550-qrrdi667	135	20	to	to	IN
cord-005550-qrrdi667	135	21	our	-PRON-	PRP$
cord-005550-qrrdi667	135	22	Costa	Costa	NNP
cord-005550-qrrdi667	135	23	Rica	Rica	NNP
cord-005550-qrrdi667	135	24	sample	sample	NN
cord-005550-qrrdi667	135	25	.	.	.
cord-005550-qrrdi667	136	1	The	the	DT
cord-005550-qrrdi667	136	2	first	first	JJ
cord-005550-qrrdi667	136	3	analysis	analysis	NN
cord-005550-qrrdi667	136	4	of	of	IN
cord-005550-qrrdi667	136	5	a	a	DT
cord-005550-qrrdi667	136	6	MHC	MHC	NNP
cord-005550-qrrdi667	136	7	gene	gene	NN
cord-005550-qrrdi667	136	8	in	in	IN
cord-005550-qrrdi667	136	9	a	a	DT
cord-005550-qrrdi667	136	10	bat	bat	NN
cord-005550-qrrdi667	136	11	species	specie	NNS
cord-005550-qrrdi667	136	12	revealed	reveal	VBD
cord-005550-qrrdi667	136	13	concordant	concordant	JJ
cord-005550-qrrdi667	136	14	diversity	diversity	NN
cord-005550-qrrdi667	136	15	patterns	pattern	NNS
cord-005550-qrrdi667	136	16	as	as	IN
cord-005550-qrrdi667	136	17	in	in	IN
cord-005550-qrrdi667	136	18	species	specie	NNS
cord-005550-qrrdi667	136	19	of	of	IN
cord-005550-qrrdi667	136	20	other	other	JJ
cord-005550-qrrdi667	136	21	mammalian	mammalian	JJ
cord-005550-qrrdi667	136	22	orders	order	NNS
cord-005550-qrrdi667	136	23	.	.	.
cord-005550-qrrdi667	137	1	In	in	IN
cord-005550-qrrdi667	137	2	the	the	DT
cord-005550-qrrdi667	137	3	genome	genome	NN
cord-005550-qrrdi667	137	4	of	of	IN
cord-005550-qrrdi667	137	5	the	the	DT
cord-005550-qrrdi667	137	6	sac	sac	NN
cord-005550-qrrdi667	137	7	-	-	HYPH
cord-005550-qrrdi667	137	8	winged	winged	JJ
cord-005550-qrrdi667	137	9	bat	bat	NN
cord-005550-qrrdi667	137	10	S.	S.	NNP
cord-005550-qrrdi667	137	11	bilineata	bilineata	NNP
cord-005550-qrrdi667	137	12	,	,	,
cord-005550-qrrdi667	137	13	we	-PRON-	PRP
cord-005550-qrrdi667	137	14	found	find	VBD
cord-005550-qrrdi667	137	15	evidence	evidence	NN
cord-005550-qrrdi667	137	16	for	for	IN
cord-005550-qrrdi667	137	17	two	two	CD
cord-005550-qrrdi667	137	18	gene	gene	NN
cord-005550-qrrdi667	137	19	copies	copy	NNS
cord-005550-qrrdi667	137	20	of	of	IN
cord-005550-qrrdi667	137	21	the	the	DT
cord-005550-qrrdi667	137	22	MHC	MHC	NNP
cord-005550-qrrdi667	137	23	class	class	NN
cord-005550-qrrdi667	137	24	II	II	NNP
cord-005550-qrrdi667	137	25	gene	gene	NN
cord-005550-qrrdi667	137	26	DRB	DRB	NNP
cord-005550-qrrdi667	137	27	,	,	,
cord-005550-qrrdi667	137	28	which	which	WDT
cord-005550-qrrdi667	137	29	is	be	VBZ
cord-005550-qrrdi667	137	30	involved	involve	VBN
cord-005550-qrrdi667	137	31	in	in	IN
cord-005550-qrrdi667	137	32	antigen	antigen	NN
cord-005550-qrrdi667	137	33	presentation	presentation	NN
cord-005550-qrrdi667	137	34	.	.	.
cord-005550-qrrdi667	138	1	Cloning	clone	VBG
cord-005550-qrrdi667	138	2	and	and	CC
cord-005550-qrrdi667	138	3	sequencing	sequence	VBG
cord-005550-qrrdi667	138	4	PCR	pcr	NN
cord-005550-qrrdi667	138	5	products	product	NNS
cord-005550-qrrdi667	138	6	amplified	amplify	VBN
cord-005550-qrrdi667	138	7	by	by	IN
cord-005550-qrrdi667	138	8	MHC	MHC	NNP
cord-005550-qrrdi667	138	9	class	class	NN
cord-005550-qrrdi667	138	10	II	II	NNP
cord-005550-qrrdi667	138	11	b	b	NN
cord-005550-qrrdi667	138	12	-	-	HYPH
cord-005550-qrrdi667	138	13	specific	specific	JJ
cord-005550-qrrdi667	138	14	primers	primer	NNS
cord-005550-qrrdi667	138	15	produced	produce	VBD
cord-005550-qrrdi667	138	16	a	a	DT
cord-005550-qrrdi667	138	17	maximum	maximum	NN
cord-005550-qrrdi667	138	18	of	of	IN
cord-005550-qrrdi667	138	19	three	three	CD
cord-005550-qrrdi667	138	20	haplotypes	haplotype	NNS
cord-005550-qrrdi667	138	21	per	per	IN
cord-005550-qrrdi667	138	22	individual	individual	NN
cord-005550-qrrdi667	138	23	.	.	.
cord-005550-qrrdi667	139	1	Three	three	CD
cord-005550-qrrdi667	139	2	haplotypes	haplotype	NNS
cord-005550-qrrdi667	139	3	were	be	VBD
cord-005550-qrrdi667	139	4	found	find	VBN
cord-005550-qrrdi667	139	5	only	only	RB
cord-005550-qrrdi667	139	6	in	in	IN
cord-005550-qrrdi667	139	7	two	two	CD
cord-005550-qrrdi667	139	8	out	out	IN
cord-005550-qrrdi667	139	9	of	of	IN
cord-005550-qrrdi667	139	10	85	85	CD
cord-005550-qrrdi667	139	11	individuals	individual	NNS
cord-005550-qrrdi667	139	12	(	(	-LRB-
cord-005550-qrrdi667	139	13	2	2	CD
cord-005550-qrrdi667	139	14	%	%	NN
cord-005550-qrrdi667	139	15	)	)	-RRB-
cord-005550-qrrdi667	139	16	.	.	.
cord-005550-qrrdi667	140	1	Only	only	RB
cord-005550-qrrdi667	140	2	in	in	IN
cord-005550-qrrdi667	140	3	these	these	DT
cord-005550-qrrdi667	140	4	two	two	CD
cord-005550-qrrdi667	140	5	individuals	individual	NNS
cord-005550-qrrdi667	140	6	,	,	,
cord-005550-qrrdi667	140	7	one	one	CD
cord-005550-qrrdi667	140	8	haplotype	haplotype	NN
cord-005550-qrrdi667	140	9	differed	differ	VBD
cord-005550-qrrdi667	140	10	from	from	IN
cord-005550-qrrdi667	140	11	all	all	PDT
cord-005550-qrrdi667	140	12	the	the	DT
cord-005550-qrrdi667	140	13	other	other	JJ
cord-005550-qrrdi667	140	14	haplotypes	haplotype	NNS
cord-005550-qrrdi667	140	15	detected	detect	VBN
cord-005550-qrrdi667	140	16	by	by	IN
cord-005550-qrrdi667	140	17	a	a	DT
cord-005550-qrrdi667	140	18	75	75	CD
cord-005550-qrrdi667	140	19	bp	bp	NNP
cord-005550-qrrdi667	140	20	insertion	insertion	NN
cord-005550-qrrdi667	140	21	and	and	CC
cord-005550-qrrdi667	140	22	thus	thus	RB
cord-005550-qrrdi667	140	23	likely	likely	RB
cord-005550-qrrdi667	140	24	originated	originate	VBD
cord-005550-qrrdi667	140	25	from	from	IN
cord-005550-qrrdi667	140	26	another	another	DT
cord-005550-qrrdi667	140	27	(	(	-LRB-
cord-005550-qrrdi667	140	28	second	second	JJ
cord-005550-qrrdi667	140	29	)	)	-RRB-
cord-005550-qrrdi667	140	30	locus	locus	NN
cord-005550-qrrdi667	140	31	.	.	.
cord-005550-qrrdi667	141	1	The	the	DT
cord-005550-qrrdi667	141	2	occasional	occasional	JJ
cord-005550-qrrdi667	141	3	occurrence	occurrence	NN
cord-005550-qrrdi667	141	4	of	of	IN
cord-005550-qrrdi667	141	5	three	three	CD
cord-005550-qrrdi667	141	6	haplotypes	haplotype	NNS
cord-005550-qrrdi667	141	7	can	can	MD
cord-005550-qrrdi667	141	8	have	have	VB
cord-005550-qrrdi667	141	9	two	two	CD
cord-005550-qrrdi667	141	10	reasons	reason	NNS
cord-005550-qrrdi667	141	11	.	.	.
cord-005550-qrrdi667	142	1	Either	either	CC
cord-005550-qrrdi667	142	2	duplicated	duplicate	VBN
cord-005550-qrrdi667	142	3	DRB	DRB	NNP
cord-005550-qrrdi667	142	4	loci	locus	NNS
cord-005550-qrrdi667	142	5	exist	exist	VBP
cord-005550-qrrdi667	142	6	only	only	RB
cord-005550-qrrdi667	142	7	in	in	IN
cord-005550-qrrdi667	142	8	some	some	DT
cord-005550-qrrdi667	142	9	individuals	individual	NNS
cord-005550-qrrdi667	142	10	or	or	CC
cord-005550-qrrdi667	142	11	the	the	DT
cord-005550-qrrdi667	142	12	primers	primer	NNS
cord-005550-qrrdi667	142	13	we	-PRON-	PRP
cord-005550-qrrdi667	142	14	used	use	VBD
cord-005550-qrrdi667	142	15	amplify	amplify	VBP
cord-005550-qrrdi667	142	16	preferably	preferably	RB
cord-005550-qrrdi667	142	17	only	only	RB
cord-005550-qrrdi667	142	18	one	one	CD
cord-005550-qrrdi667	142	19	locus	locus	NN
cord-005550-qrrdi667	142	20	.	.	.
cord-005550-qrrdi667	143	1	At	at	IN
cord-005550-qrrdi667	143	2	the	the	DT
cord-005550-qrrdi667	143	3	moment	moment	NN
cord-005550-qrrdi667	143	4	,	,	,
cord-005550-qrrdi667	143	5	we	-PRON-	PRP
cord-005550-qrrdi667	143	6	can	can	MD
cord-005550-qrrdi667	143	7	not	not	RB
cord-005550-qrrdi667	143	8	distinguish	distinguish	VB
cord-005550-qrrdi667	143	9	between	between	IN
cord-005550-qrrdi667	143	10	both	both	DT
cord-005550-qrrdi667	143	11	hypotheses	hypothesis	NNS
cord-005550-qrrdi667	143	12	.	.	.
cord-005550-qrrdi667	144	1	Although	although	IN
cord-005550-qrrdi667	144	2	we	-PRON-	PRP
cord-005550-qrrdi667	144	3	used	use	VBD
cord-005550-qrrdi667	144	4	DRB	DRB	NNP
cord-005550-qrrdi667	144	5	exon	exon	NN
cord-005550-qrrdi667	144	6	2-specific	2-specific	CD
cord-005550-qrrdi667	144	7	primers	primer	NNS
cord-005550-qrrdi667	144	8	,	,	,
cord-005550-qrrdi667	144	9	which	which	WDT
cord-005550-qrrdi667	144	10	were	be	VBD
cord-005550-qrrdi667	144	11	also	also	RB
cord-005550-qrrdi667	144	12	successfully	successfully	RB
cord-005550-qrrdi667	144	13	applied	apply	VBN
cord-005550-qrrdi667	144	14	in	in	IN
cord-005550-qrrdi667	144	15	other	other	JJ
cord-005550-qrrdi667	144	16	mammalian	mammalian	JJ
cord-005550-qrrdi667	144	17	orders	order	NNS
cord-005550-qrrdi667	144	18	,	,	,
cord-005550-qrrdi667	144	19	including	include	VBG
cord-005550-qrrdi667	144	20	primates	primate	NNS
cord-005550-qrrdi667	144	21	and	and	CC
cord-005550-qrrdi667	144	22	rodents	rodent	NNS
cord-005550-qrrdi667	144	23	(	(	-LRB-
cord-005550-qrrdi667	144	24	Gyllensten	Gyllensten	NNP
cord-005550-qrrdi667	144	25	et	et	NNP
cord-005550-qrrdi667	144	26	al	al	NNP
cord-005550-qrrdi667	144	27	.	.	NNP
cord-005550-qrrdi667	144	28	,	,	,
cord-005550-qrrdi667	144	29	1991	1991	CD
cord-005550-qrrdi667	144	30	;	;	:
cord-005550-qrrdi667	144	31	Schad	Schad	NNP
cord-005550-qrrdi667	144	32	et	et	FW
cord-005550-qrrdi667	144	33	al	al	NNP
cord-005550-qrrdi667	144	34	.	.	.
cord-005550-qrrdi667	145	1	,	,	,
cord-005550-qrrdi667	145	2	2004	2004	CD
cord-005550-qrrdi667	145	3	;	;	:
cord-005550-qrrdi667	145	4	Froeschke	Froeschke	NNP
cord-005550-qrrdi667	145	5	and	and	CC
cord-005550-qrrdi667	145	6	Sommer	Sommer	NNP
cord-005550-qrrdi667	145	7	,	,	,
cord-005550-qrrdi667	145	8	2005	2005	CD
cord-005550-qrrdi667	145	9	)	)	-RRB-
cord-005550-qrrdi667	145	10	,	,	,
cord-005550-qrrdi667	145	11	we	-PRON-	PRP
cord-005550-qrrdi667	145	12	can	can	MD
cord-005550-qrrdi667	145	13	not	not	RB
cord-005550-qrrdi667	145	14	rule	rule	VB
cord-005550-qrrdi667	145	15	out	out	RP
cord-005550-qrrdi667	145	16	the	the	DT
cord-005550-qrrdi667	145	17	amplification	amplification	NN
cord-005550-qrrdi667	145	18	failure	failure	NN
cord-005550-qrrdi667	145	19	of	of	IN
cord-005550-qrrdi667	145	20	a	a	DT
cord-005550-qrrdi667	145	21	locus	locus	NN
cord-005550-qrrdi667	145	22	due	due	JJ
cord-005550-qrrdi667	145	23	to	to	IN
cord-005550-qrrdi667	145	24	mutations	mutation	NNS
cord-005550-qrrdi667	145	25	in	in	IN
cord-005550-qrrdi667	145	26	a	a	DT
cord-005550-qrrdi667	145	27	primer	primer	NN
cord-005550-qrrdi667	145	28	-	-	HYPH
cord-005550-qrrdi667	145	29	binding	bind	VBG
cord-005550-qrrdi667	145	30	site	site	NN
cord-005550-qrrdi667	145	31	.	.	.
cord-005550-qrrdi667	146	1	In	in	IN
cord-005550-qrrdi667	146	2	98	98	CD
cord-005550-qrrdi667	146	3	%	%	NN
cord-005550-qrrdi667	146	4	of	of	IN
cord-005550-qrrdi667	146	5	all	all	DT
cord-005550-qrrdi667	146	6	bats	bat	NNS
cord-005550-qrrdi667	146	7	studied	study	VBD
cord-005550-qrrdi667	146	8	,	,	,
cord-005550-qrrdi667	146	9	we	-PRON-	PRP
cord-005550-qrrdi667	146	10	amplified	amplify	VBD
cord-005550-qrrdi667	146	11	one	one	CD
cord-005550-qrrdi667	146	12	or	or	CC
cord-005550-qrrdi667	146	13	two	two	CD
cord-005550-qrrdi667	146	14	haplotypes	haplotype	NNS
cord-005550-qrrdi667	146	15	.	.	.
cord-005550-qrrdi667	147	1	We	-PRON-	PRP
cord-005550-qrrdi667	147	2	regarded	regard	VBD
cord-005550-qrrdi667	147	3	them	-PRON-	PRP
cord-005550-qrrdi667	147	4	as	as	IN
cord-005550-qrrdi667	147	5	alleles	allele	NNS
cord-005550-qrrdi667	147	6	from	from	IN
cord-005550-qrrdi667	147	7	one	one	CD
cord-005550-qrrdi667	147	8	locus	locus	NN
cord-005550-qrrdi667	147	9	(	(	-LRB-
cord-005550-qrrdi667	147	10	DRB1	DRB1	NNP
cord-005550-qrrdi667	147	11	)	)	-RRB-
cord-005550-qrrdi667	147	12	.	.	.
cord-005550-qrrdi667	148	1	This	this	DT
cord-005550-qrrdi667	148	2	assumption	assumption	NN
cord-005550-qrrdi667	148	3	is	be	VBZ
cord-005550-qrrdi667	148	4	supported	support	VBN
cord-005550-qrrdi667	148	5	by	by	IN
cord-005550-qrrdi667	148	6	the	the	DT
cord-005550-qrrdi667	148	7	Mendelian	mendelian	JJ
cord-005550-qrrdi667	148	8	inheritance	inheritance	NN
cord-005550-qrrdi667	148	9	of	of	IN
cord-005550-qrrdi667	148	10	alleles	allele	NNS
cord-005550-qrrdi667	148	11	that	that	WDT
cord-005550-qrrdi667	148	12	was	be	VBD
cord-005550-qrrdi667	148	13	tested	test	VBN
cord-005550-qrrdi667	148	14	in	in	IN
cord-005550-qrrdi667	148	15	46	46	CD
cord-005550-qrrdi667	148	16	parentoffspring	parentoffspring	NN
cord-005550-qrrdi667	148	17	pairs	pair	NNS
cord-005550-qrrdi667	148	18	and	and	CC
cord-005550-qrrdi667	148	19	by	by	IN
cord-005550-qrrdi667	148	20	the	the	DT
cord-005550-qrrdi667	148	21	observation	observation	NN
cord-005550-qrrdi667	148	22	that	that	IN
cord-005550-qrrdi667	148	23	genotype	genotype	NN
cord-005550-qrrdi667	148	24	frequencies	frequency	NNS
cord-005550-qrrdi667	148	25	matched	match	VBD
cord-005550-qrrdi667	148	26	Hardy	Hardy	NNP
cord-005550-qrrdi667	148	27	-	-	HYPH
cord-005550-qrrdi667	148	28	Weinberg	Weinberg	NNP
cord-005550-qrrdi667	148	29	expectations	expectation	NNS
cord-005550-qrrdi667	148	30	.	.	.
cord-005550-qrrdi667	149	1	Allelic	allelic	JJ
cord-005550-qrrdi667	149	2	diversity	diversity	NN
cord-005550-qrrdi667	150	1	A	a	DT
cord-005550-qrrdi667	150	2	total	total	NN
cord-005550-qrrdi667	150	3	of	of	IN
cord-005550-qrrdi667	150	4	10	10	CD
cord-005550-qrrdi667	150	5	alleles	allele	NNS
cord-005550-qrrdi667	150	6	was	be	VBD
cord-005550-qrrdi667	150	7	found	find	VBN
cord-005550-qrrdi667	150	8	within	within	IN
cord-005550-qrrdi667	150	9	a	a	DT
cord-005550-qrrdi667	150	10	small	small	JJ
cord-005550-qrrdi667	150	11	geographic	geographic	JJ
cord-005550-qrrdi667	150	12	area	area	NN
cord-005550-qrrdi667	150	13	of	of	IN
cord-005550-qrrdi667	150	14	the	the	DT
cord-005550-qrrdi667	150	15	biological	biological	JJ
cord-005550-qrrdi667	150	16	station	station	NN
cord-005550-qrrdi667	150	17	La	La	NNP
cord-005550-qrrdi667	150	18	Selva	Selva	NNP
cord-005550-qrrdi667	150	19	in	in	IN
cord-005550-qrrdi667	150	20	Costa	Costa	NNP
cord-005550-qrrdi667	150	21	Rica	Rica	NNP
cord-005550-qrrdi667	150	22	.	.	.
cord-005550-qrrdi667	151	1	Alleles	allele	NNS
cord-005550-qrrdi667	151	2	differed	differ	VBD
cord-005550-qrrdi667	151	3	substantially	substantially	RB
cord-005550-qrrdi667	151	4	in	in	IN
cord-005550-qrrdi667	151	5	frequency	frequency	NN
cord-005550-qrrdi667	151	6	.	.	.
cord-005550-qrrdi667	152	1	Although	although	IN
cord-005550-qrrdi667	152	2	allele	allele	NN
cord-005550-qrrdi667	152	3	A1	A1	NNP
cord-005550-qrrdi667	152	4	occurred	occur	VBD
cord-005550-qrrdi667	152	5	at	at	IN
cord-005550-qrrdi667	152	6	a	a	DT
cord-005550-qrrdi667	152	7	frequency	frequency	NN
cord-005550-qrrdi667	152	8	of	of	IN
cord-005550-qrrdi667	152	9	80	80	CD
cord-005550-qrrdi667	152	10	%	%	NN
cord-005550-qrrdi667	152	11	,	,	,
cord-005550-qrrdi667	152	12	the	the	DT
cord-005550-qrrdi667	152	13	frequencies	frequency	NNS
cord-005550-qrrdi667	152	14	of	of	IN
cord-005550-qrrdi667	152	15	the	the	DT
cord-005550-qrrdi667	152	16	remaining	remain	VBG
cord-005550-qrrdi667	152	17	alleles	allele	NNS
cord-005550-qrrdi667	152	18	were	be	VBD
cord-005550-qrrdi667	152	19	5	5	CD
cord-005550-qrrdi667	152	20	%	%	NN
cord-005550-qrrdi667	152	21	or	or	CC
cord-005550-qrrdi667	152	22	less	less	JJR
cord-005550-qrrdi667	152	23	.	.	.
cord-005550-qrrdi667	153	1	We	-PRON-	PRP
cord-005550-qrrdi667	153	2	did	do	VBD
cord-005550-qrrdi667	153	3	not	not	RB
cord-005550-qrrdi667	153	4	find	find	VB
cord-005550-qrrdi667	153	5	evidence	evidence	NN
cord-005550-qrrdi667	153	6	of	of	IN
cord-005550-qrrdi667	153	7	an	an	DT
cord-005550-qrrdi667	153	8	amplification	amplification	NN
cord-005550-qrrdi667	153	9	bias	bias	NN
cord-005550-qrrdi667	153	10	among	among	IN
cord-005550-qrrdi667	153	11	alleles	allele	NNS
cord-005550-qrrdi667	153	12	,	,	,
cord-005550-qrrdi667	153	13	which	which	WDT
cord-005550-qrrdi667	153	14	could	could	MD
cord-005550-qrrdi667	153	15	explain	explain	VB
cord-005550-qrrdi667	153	16	such	such	JJ
cord-005550-qrrdi667	153	17	frequency	frequency	NN
cord-005550-qrrdi667	153	18	differences	difference	NNS
cord-005550-qrrdi667	153	19	.	.	.
cord-005550-qrrdi667	154	1	Among	among	IN
cord-005550-qrrdi667	154	2	25	25	CD
cord-005550-qrrdi667	154	3	heterozygous	heterozygous	JJ
cord-005550-qrrdi667	154	4	individuals	individual	NNS
cord-005550-qrrdi667	154	5	,	,	,
cord-005550-qrrdi667	154	6	clones	clone	NNS
cord-005550-qrrdi667	154	7	containing	contain	VBG
cord-005550-qrrdi667	154	8	the	the	DT
cord-005550-qrrdi667	154	9	allele	allele	NN
cord-005550-qrrdi667	154	10	A1	A1	NNP
cord-005550-qrrdi667	154	11	were	be	VBD
cord-005550-qrrdi667	154	12	not	not	RB
cord-005550-qrrdi667	154	13	more	more	RBR
cord-005550-qrrdi667	154	14	frequent	frequent	JJ
cord-005550-qrrdi667	154	15	than	than	IN
cord-005550-qrrdi667	154	16	clones	clone	NNS
cord-005550-qrrdi667	154	17	containing	contain	VBG
cord-005550-qrrdi667	154	18	another	another	DT
cord-005550-qrrdi667	154	19	allele	allele	NN
cord-005550-qrrdi667	154	20	(	(	-LRB-
cord-005550-qrrdi667	154	21	Wilcoxon	Wilcoxon	NNP
cord-005550-qrrdi667	154	22	signed	sign	VBN
cord-005550-qrrdi667	154	23	-	-	HYPH
cord-005550-qrrdi667	154	24	rank	rank	NN
cord-005550-qrrdi667	154	25	test	test	NN
cord-005550-qrrdi667	154	26	,	,	,
cord-005550-qrrdi667	154	27	Z	z	NN
cord-005550-qrrdi667	154	28	¼	¼	CD
cord-005550-qrrdi667	154	29	À0.07	À0.07	NNP
cord-005550-qrrdi667	154	30	,	,	,
cord-005550-qrrdi667	154	31	P	p	NN
cord-005550-qrrdi667	154	32	¼	¼	CD
cord-005550-qrrdi667	154	33	0.94	0.94	CD
cord-005550-qrrdi667	154	34	)	)	-RRB-
cord-005550-qrrdi667	154	35	.	.	.
cord-005550-qrrdi667	155	1	The	the	DT
cord-005550-qrrdi667	155	2	number	number	NN
cord-005550-qrrdi667	155	3	of	of	IN
cord-005550-qrrdi667	155	4	DRB	DRB	NNP
cord-005550-qrrdi667	155	5	alleles	allele	NNS
cord-005550-qrrdi667	155	6	varies	vary	VBZ
cord-005550-qrrdi667	155	7	substantially	substantially	RB
cord-005550-qrrdi667	155	8	among	among	IN
cord-005550-qrrdi667	155	9	mammalian	mammalian	JJ
cord-005550-qrrdi667	155	10	species	specie	NNS
cord-005550-qrrdi667	155	11	ranging	range	VBG
cord-005550-qrrdi667	155	12	from	from	IN
cord-005550-qrrdi667	155	13	one	one	CD
cord-005550-qrrdi667	155	14	or	or	CC
cord-005550-qrrdi667	155	15	two	two	CD
cord-005550-qrrdi667	155	16	in	in	IN
cord-005550-qrrdi667	155	17	some	some	DT
cord-005550-qrrdi667	155	18	small	small	JJ
cord-005550-qrrdi667	155	19	and	and	CC
cord-005550-qrrdi667	155	20	inbred	inbred	JJ
cord-005550-qrrdi667	155	21	populations	population	NNS
cord-005550-qrrdi667	155	22	(	(	-LRB-
cord-005550-qrrdi667	155	23	ruminants	ruminant	NNS
cord-005550-qrrdi667	155	24	:	:	:
cord-005550-qrrdi667	155	25	Mikko	Mikko	NNP
cord-005550-qrrdi667	155	26	et	et	NNP
cord-005550-qrrdi667	155	27	al	al	NNP
cord-005550-qrrdi667	155	28	.	.	NNP
cord-005550-qrrdi667	155	29	,	,	,
cord-005550-qrrdi667	155	30	1999	1999	CD
cord-005550-qrrdi667	155	31	;	;	:
cord-005550-qrrdi667	155	32	canids	canid	NNS
cord-005550-qrrdi667	155	33	:	:	:
cord-005550-qrrdi667	155	34	Aguilar	Aguilar	NNP
cord-005550-qrrdi667	155	35	et	et	NNP
cord-005550-qrrdi667	155	36	al	al	NNP
cord-005550-qrrdi667	155	37	.	.	NNP
cord-005550-qrrdi667	155	38	,	,	,
cord-005550-qrrdi667	155	39	2004	2004	CD
cord-005550-qrrdi667	155	40	;	;	:
cord-005550-qrrdi667	155	41	rodents	rodent	NNS
cord-005550-qrrdi667	155	42	:	:	:
cord-005550-qrrdi667	155	43	Sommer	Sommer	NNP
cord-005550-qrrdi667	155	44	and	and	CC
cord-005550-qrrdi667	155	45	Tichy	Tichy	NNP
cord-005550-qrrdi667	155	46	,	,	,
cord-005550-qrrdi667	155	47	1999	1999	CD
cord-005550-qrrdi667	155	48	;	;	:
cord-005550-qrrdi667	155	49	pinnipeds	pinniped	NNS
cord-005550-qrrdi667	155	50	:	:	:
cord-005550-qrrdi667	155	51	Weber	Weber	NNP
cord-005550-qrrdi667	155	52	et	et	NNP
cord-005550-qrrdi667	155	53	al	al	NNP
cord-005550-qrrdi667	155	54	.	.	NNP
cord-005550-qrrdi667	155	55	,	,	,
cord-005550-qrrdi667	155	56	2004	2004	CD
cord-005550-qrrdi667	155	57	)	)	-RRB-
cord-005550-qrrdi667	155	58	to	to	IN
cord-005550-qrrdi667	155	59	321	321	CD
cord-005550-qrrdi667	155	60	alleles	allele	NNS
cord-005550-qrrdi667	155	61	at	at	IN
cord-005550-qrrdi667	155	62	the	the	DT
cord-005550-qrrdi667	155	63	human	human	JJ
cord-005550-qrrdi667	155	64	HLA	HLA	NNP
cord-005550-qrrdi667	155	65	-	-	HYPH
cord-005550-qrrdi667	155	66	DRB1	DRB1	NNP
cord-005550-qrrdi667	155	67	locus	locus	NN
cord-005550-qrrdi667	155	68	(	(	-LRB-
cord-005550-qrrdi667	155	69	Garrigan	Garrigan	NNP
cord-005550-qrrdi667	155	70	and	and	CC
cord-005550-qrrdi667	155	71	Hedrick	Hedrick	NNP
cord-005550-qrrdi667	155	72	,	,	,
cord-005550-qrrdi667	155	73	2003	2003	CD
cord-005550-qrrdi667	155	74	)	)	-RRB-
cord-005550-qrrdi667	155	75	.	.	.
cord-005550-qrrdi667	156	1	This	this	DT
cord-005550-qrrdi667	156	2	enormous	enormous	JJ
cord-005550-qrrdi667	156	3	variation	variation	NN
cord-005550-qrrdi667	156	4	has	have	VBZ
cord-005550-qrrdi667	156	5	to	to	TO
cord-005550-qrrdi667	156	6	be	be	VB
cord-005550-qrrdi667	156	7	treated	treat	VBN
cord-005550-qrrdi667	156	8	with	with	IN
cord-005550-qrrdi667	156	9	caution	caution	NN
cord-005550-qrrdi667	156	10	because	because	IN
cord-005550-qrrdi667	156	11	the	the	DT
cord-005550-qrrdi667	156	12	number	number	NN
cord-005550-qrrdi667	156	13	of	of	IN
cord-005550-qrrdi667	156	14	alleles	allele	NNS
cord-005550-qrrdi667	156	15	detected	detect	VBN
cord-005550-qrrdi667	156	16	also	also	RB
cord-005550-qrrdi667	156	17	depends	depend	VBZ
cord-005550-qrrdi667	156	18	on	on	IN
cord-005550-qrrdi667	156	19	the	the	DT
cord-005550-qrrdi667	156	20	number	number	NN
cord-005550-qrrdi667	156	21	of	of	IN
cord-005550-qrrdi667	156	22	individuals	individual	NNS
cord-005550-qrrdi667	156	23	sampled	sample	VBN
cord-005550-qrrdi667	156	24	and	and	CC
cord-005550-qrrdi667	156	25	population	population	NN
cord-005550-qrrdi667	156	26	coverage	coverage	NN
cord-005550-qrrdi667	156	27	.	.	.
cord-005550-qrrdi667	157	1	The	the	DT
cord-005550-qrrdi667	157	2	highest	high	JJS
cord-005550-qrrdi667	157	3	numbers	number	NNS
cord-005550-qrrdi667	157	4	of	of	IN
cord-005550-qrrdi667	157	5	DRB	DRB	NNP
cord-005550-qrrdi667	157	6	alleles	allele	NNS
cord-005550-qrrdi667	157	7	are	be	VBP
cord-005550-qrrdi667	157	8	described	describe	VBN
cord-005550-qrrdi667	157	9	from	from	IN
cord-005550-qrrdi667	157	10	intensively	intensively	RB
cord-005550-qrrdi667	157	11	studied	study	VBN
cord-005550-qrrdi667	157	12	species	specie	NNS
cord-005550-qrrdi667	157	13	like	like	IN
cord-005550-qrrdi667	157	14	humans	human	NNS
cord-005550-qrrdi667	157	15	(	(	-LRB-
cord-005550-qrrdi667	157	16	321	321	CD
cord-005550-qrrdi667	157	17	alleles	allele	NNS
cord-005550-qrrdi667	157	18	,	,	,
cord-005550-qrrdi667	157	19	Garrigan	Garrigan	NNP
cord-005550-qrrdi667	157	20	and	and	CC
cord-005550-qrrdi667	157	21	Hedrick	Hedrick	NNP
cord-005550-qrrdi667	157	22	,	,	,
cord-005550-qrrdi667	157	23	2003	2003	CD
cord-005550-qrrdi667	157	24	)	)	-RRB-
cord-005550-qrrdi667	157	25	or	or	CC
cord-005550-qrrdi667	157	26	sheep	sheep	NN
cord-005550-qrrdi667	157	27	(	(	-LRB-
cord-005550-qrrdi667	157	28	106	106	CD
cord-005550-qrrdi667	157	29	alleles	allele	NNS
cord-005550-qrrdi667	157	30	,	,	,
cord-005550-qrrdi667	157	31	Konnai	Konnai	NNP
cord-005550-qrrdi667	157	32	et	et	NNP
cord-005550-qrrdi667	157	33	al	al	NNP
cord-005550-qrrdi667	157	34	.	.	NNP
cord-005550-qrrdi667	157	35	,	,	,
cord-005550-qrrdi667	157	36	2003	2003	CD
cord-005550-qrrdi667	157	37	)	)	-RRB-
cord-005550-qrrdi667	157	38	.	.	.
cord-005550-qrrdi667	158	1	In	in	IN
cord-005550-qrrdi667	158	2	the	the	DT
cord-005550-qrrdi667	158	3	sac	sac	NN
cord-005550-qrrdi667	158	4	-	-	HYPH
cord-005550-qrrdi667	158	5	winged	winged	JJ
cord-005550-qrrdi667	158	6	bat	bat	NN
cord-005550-qrrdi667	158	7	,	,	,
cord-005550-qrrdi667	158	8	the	the	DT
cord-005550-qrrdi667	158	9	number	number	NN
cord-005550-qrrdi667	158	10	of	of	IN
cord-005550-qrrdi667	158	11	DRB1	DRB1	NNP
cord-005550-qrrdi667	158	12	alleles	allele	NNS
cord-005550-qrrdi667	158	13	increased	increase	VBD
cord-005550-qrrdi667	158	14	to	to	IN
cord-005550-qrrdi667	158	15	11	11	CD
cord-005550-qrrdi667	158	16	by	by	IN
cord-005550-qrrdi667	158	17	analysing	analyse	VBG
cord-005550-qrrdi667	158	18	additional	additional	JJ
cord-005550-qrrdi667	158	19	six	six	CD
cord-005550-qrrdi667	158	20	individuals	individual	NNS
cord-005550-qrrdi667	158	21	from	from	IN
cord-005550-qrrdi667	158	22	Panama	Panama	NNP
cord-005550-qrrdi667	158	23	and	and	CC
cord-005550-qrrdi667	158	24	Ecuador	Ecuador	NNP
cord-005550-qrrdi667	158	25	.	.	.
cord-005550-qrrdi667	159	1	Therefore	therefore	RB
cord-005550-qrrdi667	159	2	,	,	,
cord-005550-qrrdi667	159	3	the	the	DT
cord-005550-qrrdi667	159	4	allele	allele	NN
cord-005550-qrrdi667	159	5	number	number	NN
cord-005550-qrrdi667	159	6	found	find	VBN
cord-005550-qrrdi667	159	7	in	in	IN
cord-005550-qrrdi667	159	8	the	the	DT
cord-005550-qrrdi667	159	9	sacwinged	sacwinge	VBN
cord-005550-qrrdi667	159	10	bat	bat	NN
cord-005550-qrrdi667	159	11	is	be	VBZ
cord-005550-qrrdi667	159	12	within	within	IN
cord-005550-qrrdi667	159	13	the	the	DT
cord-005550-qrrdi667	159	14	range	range	NN
cord-005550-qrrdi667	159	15	of	of	IN
cord-005550-qrrdi667	159	16	other	other	JJ
cord-005550-qrrdi667	159	17	mammalian	mammalian	JJ
cord-005550-qrrdi667	159	18	species	specie	NNS
cord-005550-qrrdi667	159	19	,	,	,
cord-005550-qrrdi667	159	20	although	although	IN
cord-005550-qrrdi667	159	21	it	-PRON-	PRP
cord-005550-qrrdi667	159	22	ranges	range	VBZ
cord-005550-qrrdi667	159	23	at	at	IN
cord-005550-qrrdi667	159	24	the	the	DT
cord-005550-qrrdi667	159	25	lower	low	JJR
cord-005550-qrrdi667	159	26	end	end	NN
cord-005550-qrrdi667	159	27	of	of	IN
cord-005550-qrrdi667	159	28	an	an	DT
cord-005550-qrrdi667	159	29	outbred	outbred	JJ
cord-005550-qrrdi667	159	30	population	population	NN
cord-005550-qrrdi667	159	31	(	(	-LRB-
cord-005550-qrrdi667	159	32	for	for	IN
cord-005550-qrrdi667	159	33	example	example	NN
cord-005550-qrrdi667	159	34	,	,	,
cord-005550-qrrdi667	159	35	Mikko	Mikko	NNP
cord-005550-qrrdi667	159	36	et	et	NNP
cord-005550-qrrdi667	159	37	al	al	NNP
cord-005550-qrrdi667	159	38	.	.	NNP
cord-005550-qrrdi667	159	39	,	,	,
cord-005550-qrrdi667	159	40	1999	1999	CD
cord-005550-qrrdi667	159	41	)	)	-RRB-
cord-005550-qrrdi667	159	42	.	.	.
cord-005550-qrrdi667	160	1	Recombination	recombination	NN
cord-005550-qrrdi667	160	2	between	between	IN
cord-005550-qrrdi667	160	3	alleles	allele	NNS
cord-005550-qrrdi667	160	4	(	(	-LRB-
cord-005550-qrrdi667	160	5	intragenic	intragenic	NNP
cord-005550-qrrdi667	160	6	recombination	recombination	NN
cord-005550-qrrdi667	160	7	)	)	-RRB-
cord-005550-qrrdi667	160	8	or	or	CC
cord-005550-qrrdi667	160	9	between	between	IN
cord-005550-qrrdi667	160	10	loci	locus	NNS
cord-005550-qrrdi667	160	11	(	(	-LRB-
cord-005550-qrrdi667	160	12	intergenic	intergenic	JJ
cord-005550-qrrdi667	160	13	recombination	recombination	NN
cord-005550-qrrdi667	160	14	)	)	-RRB-
cord-005550-qrrdi667	160	15	can	can	MD
cord-005550-qrrdi667	160	16	rapidly	rapidly	RB
cord-005550-qrrdi667	160	17	generate	generate	VB
cord-005550-qrrdi667	160	18	allelic	allelic	JJ
cord-005550-qrrdi667	160	19	diversity	diversity	NN
cord-005550-qrrdi667	160	20	at	at	IN
cord-005550-qrrdi667	160	21	MHC	MHC	NNP
cord-005550-qrrdi667	160	22	class	class	NN
cord-005550-qrrdi667	160	23	II	II	NNP
cord-005550-qrrdi667	160	24	genes	gene	NNS
cord-005550-qrrdi667	160	25	(	(	-LRB-
cord-005550-qrrdi667	160	26	Andersson	Andersson	NNP
cord-005550-qrrdi667	160	27	and	and	CC
cord-005550-qrrdi667	160	28	Mikko	Mikko	NNP
cord-005550-qrrdi667	160	29	,	,	,
cord-005550-qrrdi667	160	30	1995	1995	CD
cord-005550-qrrdi667	160	31	)	)	-RRB-
cord-005550-qrrdi667	160	32	.	.	.
cord-005550-qrrdi667	161	1	Our	-PRON-	PRP$
cord-005550-qrrdi667	161	2	study	study	NN
cord-005550-qrrdi667	161	3	provides	provide	VBZ
cord-005550-qrrdi667	161	4	further	further	JJ
cord-005550-qrrdi667	161	5	evidence	evidence	NN
cord-005550-qrrdi667	161	6	for	for	IN
cord-005550-qrrdi667	161	7	the	the	DT
cord-005550-qrrdi667	161	8	significance	significance	NN
cord-005550-qrrdi667	161	9	of	of	IN
cord-005550-qrrdi667	161	10	recombination	recombination	NN
cord-005550-qrrdi667	161	11	in	in	IN
cord-005550-qrrdi667	161	12	generating	generate	VBG
cord-005550-qrrdi667	161	13	allelic	allelic	JJ
cord-005550-qrrdi667	161	14	diversity	diversity	NN
cord-005550-qrrdi667	161	15	of	of	IN
cord-005550-qrrdi667	161	16	MHC	MHC	NNP
cord-005550-qrrdi667	161	17	class	class	NN
cord-005550-qrrdi667	161	18	II	II	NNP
cord-005550-qrrdi667	161	19	genes	gene	NNS
cord-005550-qrrdi667	161	20	.	.	.
cord-005550-qrrdi667	162	1	Several	several	JJ
cord-005550-qrrdi667	162	2	DRB1	DRB1	NNP
cord-005550-qrrdi667	162	3	alleles	allele	NNS
cord-005550-qrrdi667	162	4	of	of	IN
cord-005550-qrrdi667	162	5	the	the	DT
cord-005550-qrrdi667	162	6	sac	sac	NN
cord-005550-qrrdi667	162	7	-	-	HYPH
cord-005550-qrrdi667	162	8	winged	winged	JJ
cord-005550-qrrdi667	162	9	bat	bat	NN
cord-005550-qrrdi667	162	10	shared	share	VBD
cord-005550-qrrdi667	162	11	different	different	JJ
cord-005550-qrrdi667	162	12	sequence	sequence	NN
cord-005550-qrrdi667	162	13	motifs	motif	NNS
cord-005550-qrrdi667	162	14	with	with	IN
cord-005550-qrrdi667	162	15	different	different	JJ
cord-005550-qrrdi667	162	16	alleles	allele	NNS
cord-005550-qrrdi667	162	17	(	(	-LRB-
cord-005550-qrrdi667	162	18	Figure	figure	NN
cord-005550-qrrdi667	162	19	1	1	CD
cord-005550-qrrdi667	162	20	)	)	-RRB-
cord-005550-qrrdi667	162	21	.	.	.
cord-005550-qrrdi667	163	1	In	in	IN
cord-005550-qrrdi667	163	2	contrast	contrast	NN
cord-005550-qrrdi667	163	3	,	,	,
cord-005550-qrrdi667	163	4	the	the	DT
cord-005550-qrrdi667	163	5	two	two	CD
cord-005550-qrrdi667	163	6	B	b	NN
cord-005550-qrrdi667	163	7	sequence	sequence	NN
cord-005550-qrrdi667	163	8	types	type	NNS
cord-005550-qrrdi667	163	9	,	,	,
cord-005550-qrrdi667	163	10	which	which	WDT
cord-005550-qrrdi667	163	11	are	be	VBP
cord-005550-qrrdi667	163	12	likely	likely	RB
cord-005550-qrrdi667	163	13	encoded	encode	VBN
cord-005550-qrrdi667	163	14	by	by	IN
cord-005550-qrrdi667	163	15	another	another	DT
cord-005550-qrrdi667	163	16	locus	locus	NN
cord-005550-qrrdi667	163	17	,	,	,
cord-005550-qrrdi667	163	18	did	do	VBD
cord-005550-qrrdi667	163	19	not	not	RB
cord-005550-qrrdi667	163	20	share	share	VB
cord-005550-qrrdi667	163	21	long	long	JJ
cord-005550-qrrdi667	163	22	-	-	HYPH
cord-005550-qrrdi667	163	23	sequence	sequence	NN
cord-005550-qrrdi667	163	24	fragments	fragment	NNS
cord-005550-qrrdi667	163	25	with	with	IN
cord-005550-qrrdi667	163	26	DRB1	DRB1	NNP
cord-005550-qrrdi667	163	27	alleles	allele	NNS
cord-005550-qrrdi667	163	28	.	.	.
cord-005550-qrrdi667	164	1	This	this	DT
cord-005550-qrrdi667	164	2	suggests	suggest	VBZ
cord-005550-qrrdi667	164	3	that	that	IN
cord-005550-qrrdi667	164	4	intragenic	intragenic	JJ
cord-005550-qrrdi667	164	5	recombination	recombination	NN
cord-005550-qrrdi667	164	6	of	of	IN
cord-005550-qrrdi667	164	7	the	the	DT
cord-005550-qrrdi667	164	8	DRB1	DRB1	NNP
cord-005550-qrrdi667	164	9	locus	locus	NN
cord-005550-qrrdi667	164	10	and	and	CC
cord-005550-qrrdi667	164	11	less	less	RBR
cord-005550-qrrdi667	164	12	intergenic	intergenic	JJ
cord-005550-qrrdi667	164	13	recombination	recombination	NN
cord-005550-qrrdi667	164	14	plays	play	VBZ
cord-005550-qrrdi667	164	15	an	an	DT
cord-005550-qrrdi667	164	16	important	important	JJ
cord-005550-qrrdi667	164	17	role	role	NN
cord-005550-qrrdi667	164	18	in	in	IN
cord-005550-qrrdi667	164	19	generating	generate	VBG
cord-005550-qrrdi667	164	20	allelic	allelic	JJ
cord-005550-qrrdi667	164	21	diversity	diversity	NN
cord-005550-qrrdi667	164	22	at	at	IN
cord-005550-qrrdi667	164	23	the	the	DT
cord-005550-qrrdi667	164	24	DRB1	DRB1	NNP
cord-005550-qrrdi667	164	25	locus	locus	NN
cord-005550-qrrdi667	164	26	of	of	IN
cord-005550-qrrdi667	164	27	the	the	DT
cord-005550-qrrdi667	164	28	sac	sac	NN
cord-005550-qrrdi667	164	29	-	-	HYPH
cord-005550-qrrdi667	164	30	winged	winged	JJ
cord-005550-qrrdi667	164	31	bat	bat	NN
cord-005550-qrrdi667	164	32	.	.	.
cord-005550-qrrdi667	165	1	Despite	despite	IN
cord-005550-qrrdi667	165	2	the	the	DT
cord-005550-qrrdi667	165	3	lack	lack	NN
cord-005550-qrrdi667	165	4	of	of	IN
cord-005550-qrrdi667	165	5	clear	clear	JJ
cord-005550-qrrdi667	165	6	evidence	evidence	NN
cord-005550-qrrdi667	165	7	for	for	IN
cord-005550-qrrdi667	165	8	intergenic	intergenic	JJ
cord-005550-qrrdi667	165	9	recombination	recombination	NN
cord-005550-qrrdi667	165	10	we	-PRON-	PRP
cord-005550-qrrdi667	165	11	are	be	VBP
cord-005550-qrrdi667	165	12	not	not	RB
cord-005550-qrrdi667	165	13	able	able	JJ
cord-005550-qrrdi667	165	14	to	to	TO
cord-005550-qrrdi667	165	15	rule	rule	VB
cord-005550-qrrdi667	165	16	it	-PRON-	PRP
cord-005550-qrrdi667	165	17	out	out	RP
cord-005550-qrrdi667	165	18	,	,	,
cord-005550-qrrdi667	165	19	because	because	IN
cord-005550-qrrdi667	165	20	we	-PRON-	PRP
cord-005550-qrrdi667	165	21	might	may	MD
cord-005550-qrrdi667	165	22	have	have	VB
cord-005550-qrrdi667	165	23	missed	miss	VBN
cord-005550-qrrdi667	165	24	to	to	TO
cord-005550-qrrdi667	165	25	amplify	amplify	VB
cord-005550-qrrdi667	165	26	paralog	paralog	NN
cord-005550-qrrdi667	165	27	DRB	DRB	NNP
cord-005550-qrrdi667	165	28	loci	locus	NNS
cord-005550-qrrdi667	165	29	possibly	possibly	RB
cord-005550-qrrdi667	165	30	involved	involve	VBN
cord-005550-qrrdi667	165	31	in	in	IN
cord-005550-qrrdi667	165	32	intergenic	intergenic	JJ
cord-005550-qrrdi667	165	33	recombination	recombination	NN
cord-005550-qrrdi667	165	34	.	.	.
cord-005550-qrrdi667	166	1	Methodological	methodological	JJ
cord-005550-qrrdi667	166	2	artefacts	artefact	NNS
cord-005550-qrrdi667	166	3	,	,	,
cord-005550-qrrdi667	166	4	like	like	IN
cord-005550-qrrdi667	166	5	recombination	recombination	NN
cord-005550-qrrdi667	166	6	during	during	IN
cord-005550-qrrdi667	166	7	PCR	PCR	NNP
cord-005550-qrrdi667	166	8	,	,	,
cord-005550-qrrdi667	166	9	can	can	MD
cord-005550-qrrdi667	166	10	be	be	VB
cord-005550-qrrdi667	166	11	ruled	rule	VBN
cord-005550-qrrdi667	166	12	out	out	RP
cord-005550-qrrdi667	166	13	because	because	IN
cord-005550-qrrdi667	166	14	recombined	recombine	VBN
cord-005550-qrrdi667	166	15	alleles	allele	NNS
cord-005550-qrrdi667	166	16	were	be	VBD
cord-005550-qrrdi667	166	17	independently	independently	RB
cord-005550-qrrdi667	166	18	detected	detect	VBN
cord-005550-qrrdi667	166	19	in	in	IN
cord-005550-qrrdi667	166	20	several	several	JJ
cord-005550-qrrdi667	166	21	individuals	individual	NNS
cord-005550-qrrdi667	166	22	.	.	.
cord-005550-qrrdi667	167	1	Several	several	JJ
cord-005550-qrrdi667	167	2	recent	recent	JJ
cord-005550-qrrdi667	167	3	studies	study	NNS
cord-005550-qrrdi667	167	4	on	on	IN
cord-005550-qrrdi667	167	5	genetic	genetic	JJ
cord-005550-qrrdi667	167	6	diversity	diversity	NN
cord-005550-qrrdi667	167	7	of	of	IN
cord-005550-qrrdi667	167	8	MHC	MHC	NNP
cord-005550-qrrdi667	167	9	class	class	NN
cord-005550-qrrdi667	167	10	II	II	NNP
cord-005550-qrrdi667	167	11	genes	gene	NNS
cord-005550-qrrdi667	167	12	documented	document	VBD
cord-005550-qrrdi667	167	13	intragenic	intragenic	JJ
cord-005550-qrrdi667	167	14	recombination	recombination	NN
cord-005550-qrrdi667	167	15	(	(	-LRB-
cord-005550-qrrdi667	167	16	human	human	JJ
cord-005550-qrrdi667	167	17	HLA	HLA	NNP
cord-005550-qrrdi667	167	18	-	-	HYPH
cord-005550-qrrdi667	167	19	DPB1	DPB1	NNP
cord-005550-qrrdi667	167	20	:	:	:
cord-005550-qrrdi667	168	1	Zangenberg	Zangenberg	NNP
cord-005550-qrrdi667	168	2	et	et	FW
cord-005550-qrrdi667	168	3	al	al	NNP
cord-005550-qrrdi667	168	4	.	.	.
cord-005550-qrrdi667	169	1	,	,	,
cord-005550-qrrdi667	169	2	1995	1995	CD
cord-005550-qrrdi667	169	3	;	;	:
cord-005550-qrrdi667	169	4	deer	deer	NN
cord-005550-qrrdi667	169	5	mouse	mouse	NN
cord-005550-qrrdi667	169	6	EB	EB	NNP
cord-005550-qrrdi667	169	7	:	:	:
cord-005550-qrrdi667	169	8	Richman	Richman	NNP
cord-005550-qrrdi667	169	9	et	et	NNP
cord-005550-qrrdi667	169	10	al	al	NNP
cord-005550-qrrdi667	169	11	.	.	NNP
cord-005550-qrrdi667	169	12	,	,	,
cord-005550-qrrdi667	169	13	2003	2003	CD
cord-005550-qrrdi667	169	14	;	;	:
cord-005550-qrrdi667	169	15	chamois	chamois	NNP
cord-005550-qrrdi667	169	16	DRB	DRB	NNP
cord-005550-qrrdi667	169	17	:	:	:
cord-005550-qrrdi667	170	1	Schaschl	Schaschl	NNP
cord-005550-qrrdi667	170	2	et	et	FW
cord-005550-qrrdi667	170	3	al	al	NNP
cord-005550-qrrdi667	170	4	.	.	.
cord-005550-qrrdi667	171	1	,	,	,
cord-005550-qrrdi667	171	2	2005	2005	CD
cord-005550-qrrdi667	171	3	;	;	:
cord-005550-qrrdi667	171	4	stickleback	stickleback	NN
cord-005550-qrrdi667	171	5	MHC	MHC	NNP
cord-005550-qrrdi667	171	6	class	class	NN
cord-005550-qrrdi667	171	7	IIB	IIB	NNP
cord-005550-qrrdi667	171	8	genes	gene	NNS
cord-005550-qrrdi667	171	9	:	:	:
cord-005550-qrrdi667	172	1	Reusch	Reusch	NNP
cord-005550-qrrdi667	172	2	and	and	CC
cord-005550-qrrdi667	172	3	Langefors	Langefors	NNP
cord-005550-qrrdi667	172	4	,	,	,
cord-005550-qrrdi667	172	5	2005	2005	CD
cord-005550-qrrdi667	172	6	)	)	-RRB-
cord-005550-qrrdi667	172	7	.	.	.
cord-005550-qrrdi667	173	1	Usually	usually	RB
cord-005550-qrrdi667	173	2	only	only	RB
cord-005550-qrrdi667	173	3	short	short	JJ
cord-005550-qrrdi667	173	4	-	-	HYPH
cord-005550-qrrdi667	173	5	sequence	sequence	NN
cord-005550-qrrdi667	173	6	motifs	motif	NNS
cord-005550-qrrdi667	173	7	are	be	VBP
cord-005550-qrrdi667	173	8	shared	share	VBN
cord-005550-qrrdi667	173	9	among	among	IN
cord-005550-qrrdi667	173	10	alleles	allele	NNS
cord-005550-qrrdi667	173	11	.	.	.
cord-005550-qrrdi667	174	1	In	in	IN
cord-005550-qrrdi667	174	2	contrast	contrast	NN
cord-005550-qrrdi667	174	3	,	,	,
cord-005550-qrrdi667	174	4	shared	share	VBN
cord-005550-qrrdi667	174	5	sequence	sequence	NN
cord-005550-qrrdi667	174	6	fragments	fragment	NNS
cord-005550-qrrdi667	174	7	are	be	VBP
cord-005550-qrrdi667	174	8	rather	rather	RB
cord-005550-qrrdi667	174	9	long	long	RB
cord-005550-qrrdi667	174	10	in	in	IN
cord-005550-qrrdi667	174	11	the	the	DT
cord-005550-qrrdi667	174	12	sac	sac	NN
cord-005550-qrrdi667	174	13	-	-	HYPH
cord-005550-qrrdi667	174	14	winged	winged	JJ
cord-005550-qrrdi667	174	15	bat	bat	NN
cord-005550-qrrdi667	174	16	.	.	.
cord-005550-qrrdi667	175	1	A	a	DT
cord-005550-qrrdi667	175	2	similar	similar	JJ
cord-005550-qrrdi667	175	3	pattern	pattern	NN
cord-005550-qrrdi667	175	4	of	of	IN
cord-005550-qrrdi667	175	5	shared	share	VBN
cord-005550-qrrdi667	175	6	sequence	sequence	NN
cord-005550-qrrdi667	175	7	fragments	fragment	NNS
cord-005550-qrrdi667	175	8	among	among	IN
cord-005550-qrrdi667	175	9	alleles	allele	NNS
cord-005550-qrrdi667	175	10	was	be	VBD
cord-005550-qrrdi667	175	11	observed	observe	VBN
cord-005550-qrrdi667	175	12	by	by	IN
cord-005550-qrrdi667	175	13	Zangenberg	Zangenberg	NNP
cord-005550-qrrdi667	175	14	et	et	NNP
cord-005550-qrrdi667	175	15	al	al	NNP
cord-005550-qrrdi667	175	16	.	.	.
cord-005550-qrrdi667	176	1	(	(	-LRB-
cord-005550-qrrdi667	176	2	1995	1995	CD
cord-005550-qrrdi667	176	3	)	)	-RRB-
cord-005550-qrrdi667	176	4	,	,	,
cord-005550-qrrdi667	176	5	who	who	WP
cord-005550-qrrdi667	176	6	studied	study	VBD
cord-005550-qrrdi667	176	7	the	the	DT
cord-005550-qrrdi667	176	8	rate	rate	NN
cord-005550-qrrdi667	176	9	of	of	IN
cord-005550-qrrdi667	176	10	evolution	evolution	NN
cord-005550-qrrdi667	176	11	within	within	IN
cord-005550-qrrdi667	176	12	the	the	DT
cord-005550-qrrdi667	176	13	exon	exon	NN
cord-005550-qrrdi667	176	14	2	2	CD
cord-005550-qrrdi667	176	15	of	of	IN
cord-005550-qrrdi667	176	16	the	the	DT
cord-005550-qrrdi667	176	17	human	human	JJ
cord-005550-qrrdi667	176	18	HLA	HLA	NNP
cord-005550-qrrdi667	176	19	-	-	HYPH
cord-005550-qrrdi667	176	20	DPB1	DPB1	NNP
cord-005550-qrrdi667	176	21	locus	locus	NN
cord-005550-qrrdi667	176	22	by	by	IN
cord-005550-qrrdi667	176	23	screening	screen	VBG
cord-005550-qrrdi667	176	24	pools	pool	NNS
cord-005550-qrrdi667	176	25	of	of	IN
cord-005550-qrrdi667	176	26	sperm	sperm	NN
cord-005550-qrrdi667	176	27	.	.	.
cord-005550-qrrdi667	177	1	The	the	DT
cord-005550-qrrdi667	177	2	similarity	similarity	NN
cord-005550-qrrdi667	177	3	between	between	IN
cord-005550-qrrdi667	177	4	this	this	DT
cord-005550-qrrdi667	177	5	sperm	sperm	NN
cord-005550-qrrdi667	177	6	analysis	analysis	NN
cord-005550-qrrdi667	177	7	study	study	NN
cord-005550-qrrdi667	177	8	and	and	CC
cord-005550-qrrdi667	177	9	our	-PRON-	PRP$
cord-005550-qrrdi667	177	10	study	study	NN
cord-005550-qrrdi667	177	11	suggests	suggest	VBZ
cord-005550-qrrdi667	177	12	that	that	IN
cord-005550-qrrdi667	177	13	some	some	DT
cord-005550-qrrdi667	177	14	recombination	recombination	NN
cord-005550-qrrdi667	177	15	events	event	NNS
cord-005550-qrrdi667	177	16	among	among	IN
cord-005550-qrrdi667	177	17	DRB1	DRB1	NNP
cord-005550-qrrdi667	177	18	alleles	allele	NNS
cord-005550-qrrdi667	177	19	in	in	IN
cord-005550-qrrdi667	177	20	the	the	DT
cord-005550-qrrdi667	177	21	sac	sac	NN
cord-005550-qrrdi667	177	22	-	-	HYPH
cord-005550-qrrdi667	177	23	winged	winged	JJ
cord-005550-qrrdi667	177	24	bat	bat	NN
cord-005550-qrrdi667	177	25	occurred	occur	VBD
cord-005550-qrrdi667	177	26	rather	rather	RB
cord-005550-qrrdi667	177	27	recently	recently	RB
cord-005550-qrrdi667	177	28	.	.	.
cord-005550-qrrdi667	178	1	This	this	DT
cord-005550-qrrdi667	178	2	is	be	VBZ
cord-005550-qrrdi667	178	3	further	further	RB
cord-005550-qrrdi667	178	4	supported	support	VBN
cord-005550-qrrdi667	178	5	by	by	IN
cord-005550-qrrdi667	178	6	the	the	DT
cord-005550-qrrdi667	178	7	lack	lack	NN
cord-005550-qrrdi667	178	8	of	of	IN
cord-005550-qrrdi667	178	9	nucleotide	nucleotide	JJ
cord-005550-qrrdi667	178	10	substitutions	substitution	NNS
cord-005550-qrrdi667	178	11	between	between	IN
cord-005550-qrrdi667	178	12	some	some	DT
cord-005550-qrrdi667	178	13	DNA	dna	NN
cord-005550-qrrdi667	178	14	sequence	sequence	NN
cord-005550-qrrdi667	178	15	fragments	fragment	NNS
cord-005550-qrrdi667	178	16	that	that	WDT
cord-005550-qrrdi667	178	17	are	be	VBP
cord-005550-qrrdi667	178	18	shared	share	VBN
cord-005550-qrrdi667	178	19	by	by	IN
cord-005550-qrrdi667	178	20	two	two	CD
cord-005550-qrrdi667	178	21	alleles	allele	NNS
cord-005550-qrrdi667	178	22	(	(	-LRB-
cord-005550-qrrdi667	178	23	Figure	figure	NN
cord-005550-qrrdi667	178	24	1	1	CD
cord-005550-qrrdi667	178	25	)	)	-RRB-
cord-005550-qrrdi667	178	26	.	.	.
cord-005550-qrrdi667	179	1	One	one	CD
cord-005550-qrrdi667	179	2	of	of	IN
cord-005550-qrrdi667	179	3	the	the	DT
cord-005550-qrrdi667	179	4	strongest	strong	JJS
cord-005550-qrrdi667	179	5	arguments	argument	NNS
cord-005550-qrrdi667	179	6	for	for	IN
cord-005550-qrrdi667	179	7	balancing	balance	VBG
cord-005550-qrrdi667	179	8	selection	selection	NN
cord-005550-qrrdi667	179	9	comes	come	VBZ
cord-005550-qrrdi667	179	10	from	from	IN
cord-005550-qrrdi667	179	11	a	a	DT
cord-005550-qrrdi667	179	12	higher	high	JJR
cord-005550-qrrdi667	179	13	rate	rate	NN
cord-005550-qrrdi667	179	14	of	of	IN
cord-005550-qrrdi667	179	15	nonsynonymous	nonsynonymous	JJ
cord-005550-qrrdi667	179	16	nucleotide	nucleotide	JJ
cord-005550-qrrdi667	179	17	substitutions	substitution	NNS
cord-005550-qrrdi667	179	18	than	than	IN
cord-005550-qrrdi667	179	19	synonymous	synonymous	JJ
cord-005550-qrrdi667	179	20	nucleotide	nucleotide	JJ
cord-005550-qrrdi667	179	21	substitutions	substitution	NNS
cord-005550-qrrdi667	179	22	(	(	-LRB-
cord-005550-qrrdi667	179	23	Garrigan	Garrigan	NNP
cord-005550-qrrdi667	179	24	and	and	CC
cord-005550-qrrdi667	179	25	Hedrick	Hedrick	NNP
cord-005550-qrrdi667	179	26	,	,	,
cord-005550-qrrdi667	179	27	2003	2003	CD
cord-005550-qrrdi667	179	28	)	)	-RRB-
cord-005550-qrrdi667	179	29	.	.	.
cord-005550-qrrdi667	180	1	The	the	DT
cord-005550-qrrdi667	180	2	comparison	comparison	NN
cord-005550-qrrdi667	180	3	of	of	IN
cord-005550-qrrdi667	180	4	nonsynonymous	nonsynonymous	NNP
cord-005550-qrrdi667	180	5	to	to	TO
cord-005550-qrrdi667	180	6	synonymous	synonymous	JJ
cord-005550-qrrdi667	180	7	nucleotide	nucleotide	JJ
cord-005550-qrrdi667	180	8	substitutions	substitution	NNS
cord-005550-qrrdi667	180	9	(	(	-LRB-
cord-005550-qrrdi667	180	10	d	d	NNP
cord-005550-qrrdi667	180	11	N	N	NNP
cord-005550-qrrdi667	180	12	/d	/d	NNP
cord-005550-qrrdi667	180	13	S	S	NNP
cord-005550-qrrdi667	180	14	)	)	-RRB-
cord-005550-qrrdi667	180	15	revealed	reveal	VBD
cord-005550-qrrdi667	180	16	contrasting	contrast	VBG
cord-005550-qrrdi667	180	17	results	result	NNS
cord-005550-qrrdi667	180	18	between	between	IN
cord-005550-qrrdi667	180	19	sites	site	NNS
cord-005550-qrrdi667	180	20	involved	involve	VBN
cord-005550-qrrdi667	180	21	in	in	IN
cord-005550-qrrdi667	180	22	ABS	abs	NN
cord-005550-qrrdi667	180	23	and	and	CC
cord-005550-qrrdi667	180	24	non	non	JJ
cord-005550-qrrdi667	180	25	-	-	JJ
cord-005550-qrrdi667	180	26	ABS	abs	NN
cord-005550-qrrdi667	180	27	.	.	.
cord-005550-qrrdi667	181	1	At	at	IN
cord-005550-qrrdi667	181	2	ABS	abs	NN
cord-005550-qrrdi667	181	3	,	,	,
cord-005550-qrrdi667	181	4	nonsynonymous	nonsynonymous	JJ
cord-005550-qrrdi667	181	5	mutations	mutation	NNS
cord-005550-qrrdi667	181	6	were	be	VBD
cord-005550-qrrdi667	181	7	2.4	2.4	CD
cord-005550-qrrdi667	181	8	times	time	NNS
cord-005550-qrrdi667	181	9	more	more	RBR
cord-005550-qrrdi667	181	10	frequent	frequent	JJ
cord-005550-qrrdi667	181	11	than	than	IN
cord-005550-qrrdi667	181	12	synonymous	synonymous	JJ
cord-005550-qrrdi667	181	13	mutations	mutation	NNS
cord-005550-qrrdi667	181	14	,	,	,
cord-005550-qrrdi667	181	15	whereas	whereas	IN
cord-005550-qrrdi667	181	16	this	this	DT
cord-005550-qrrdi667	181	17	was	be	VBD
cord-005550-qrrdi667	181	18	not	not	RB
cord-005550-qrrdi667	181	19	the	the	DT
cord-005550-qrrdi667	181	20	case	case	NN
cord-005550-qrrdi667	181	21	for	for	IN
cord-005550-qrrdi667	181	22	non	non	JJ
cord-005550-qrrdi667	181	23	-	-	JJ
cord-005550-qrrdi667	181	24	ABS	abs	NN
cord-005550-qrrdi667	181	25	.	.	.
cord-005550-qrrdi667	182	1	In	in	IN
cord-005550-qrrdi667	182	2	addition	addition	NN
cord-005550-qrrdi667	182	3	,	,	,
cord-005550-qrrdi667	182	4	ABSs	abs	NNS
cord-005550-qrrdi667	182	5	were	be	VBD
cord-005550-qrrdi667	182	6	more	more	RBR
cord-005550-qrrdi667	182	7	variable	variable	JJ
cord-005550-qrrdi667	182	8	than	than	IN
cord-005550-qrrdi667	182	9	non	non	JJ
cord-005550-qrrdi667	182	10	-	-	JJ
cord-005550-qrrdi667	182	11	ABSs	abss	JJ
cord-005550-qrrdi667	182	12	.	.	.
cord-005550-qrrdi667	183	1	The	the	DT
cord-005550-qrrdi667	183	2	proportion	proportion	NN
cord-005550-qrrdi667	183	3	of	of	IN
cord-005550-qrrdi667	183	4	polymorph	polymorph	NN
cord-005550-qrrdi667	183	5	ABS	abs	NN
cord-005550-qrrdi667	183	6	was	be	VBD
cord-005550-qrrdi667	183	7	significantly	significantly	RB
cord-005550-qrrdi667	183	8	higher	high	JJR
cord-005550-qrrdi667	183	9	than	than	IN
cord-005550-qrrdi667	183	10	that	that	DT
cord-005550-qrrdi667	183	11	of	of	IN
cord-005550-qrrdi667	183	12	non	non	JJ
cord-005550-qrrdi667	183	13	-	-	JJ
cord-005550-qrrdi667	183	14	ABS	abs	JJ
cord-005550-qrrdi667	183	15	;	;	:
cord-005550-qrrdi667	183	16	and	and	CC
cord-005550-qrrdi667	183	17	at	at	IN
cord-005550-qrrdi667	183	18	the	the	DT
cord-005550-qrrdi667	183	19	protein	protein	NN
cord-005550-qrrdi667	183	20	level	level	NN
cord-005550-qrrdi667	183	21	,	,	,
cord-005550-qrrdi667	183	22	amino	amino	NN
cord-005550-qrrdi667	183	23	-	-	HYPH
cord-005550-qrrdi667	183	24	acid	acid	NN
cord-005550-qrrdi667	183	25	sites	site	NNS
cord-005550-qrrdi667	183	26	involved	involve	VBN
cord-005550-qrrdi667	183	27	in	in	IN
cord-005550-qrrdi667	183	28	antigen	antigen	NN
cord-005550-qrrdi667	183	29	binding	binding	NN
cord-005550-qrrdi667	183	30	showed	show	VBD
cord-005550-qrrdi667	183	31	higher	high	JJR
cord-005550-qrrdi667	183	32	levels	level	NNS
cord-005550-qrrdi667	183	33	of	of	IN
cord-005550-qrrdi667	183	34	heterozygosity	heterozygosity	NN
cord-005550-qrrdi667	183	35	than	than	IN
cord-005550-qrrdi667	183	36	nonantigen	nonantigen	NN
cord-005550-qrrdi667	183	37	-	-	HYPH
cord-005550-qrrdi667	183	38	binding	bind	VBG
cord-005550-qrrdi667	183	39	amino	amino	NN
cord-005550-qrrdi667	183	40	acid	acid	NN
cord-005550-qrrdi667	183	41	sites	site	NNS
cord-005550-qrrdi667	183	42	.	.	.
cord-005550-qrrdi667	184	1	The	the	DT
cord-005550-qrrdi667	184	2	d	d	NN
cord-005550-qrrdi667	184	3	N	N	NNP
cord-005550-qrrdi667	184	4	/d	/d	NNP
cord-005550-qrrdi667	184	5	S	S	NNP
cord-005550-qrrdi667	184	6	test	test	NN
cord-005550-qrrdi667	184	7	is	be	VBZ
cord-005550-qrrdi667	184	8	most	most	RBS
cord-005550-qrrdi667	184	9	commonly	commonly	RB
cord-005550-qrrdi667	184	10	used	use	VBN
cord-005550-qrrdi667	184	11	in	in	IN
cord-005550-qrrdi667	184	12	MHC	MHC	NNP
cord-005550-qrrdi667	184	13	studies	study	NNS
cord-005550-qrrdi667	184	14	to	to	TO
cord-005550-qrrdi667	184	15	infer	infer	VB
cord-005550-qrrdi667	184	16	a	a	DT
cord-005550-qrrdi667	184	17	possible	possible	JJ
cord-005550-qrrdi667	184	18	role	role	NN
cord-005550-qrrdi667	184	19	of	of	IN
cord-005550-qrrdi667	184	20	balancing	balance	VBG
cord-005550-qrrdi667	184	21	selection	selection	NN
cord-005550-qrrdi667	184	22	.	.	.
cord-005550-qrrdi667	185	1	In	in	IN
cord-005550-qrrdi667	185	2	47	47	CD
cord-005550-qrrdi667	185	3	of	of	IN
cord-005550-qrrdi667	185	4	48	48	CD
cord-005550-qrrdi667	185	5	studies	study	NNS
cord-005550-qrrdi667	185	6	,	,	,
cord-005550-qrrdi667	185	7	the	the	DT
cord-005550-qrrdi667	185	8	rate	rate	NN
cord-005550-qrrdi667	185	9	of	of	IN
cord-005550-qrrdi667	185	10	nonsynonymous	nonsynonymous	JJ
cord-005550-qrrdi667	185	11	substitutions	substitution	NNS
cord-005550-qrrdi667	185	12	exceeded	exceed	VBD
cord-005550-qrrdi667	185	13	the	the	DT
cord-005550-qrrdi667	185	14	rate	rate	NN
cord-005550-qrrdi667	185	15	of	of	IN
cord-005550-qrrdi667	185	16	synonymous	synonymous	JJ
cord-005550-qrrdi667	185	17	nucleotide	nucleotide	JJ
cord-005550-qrrdi667	185	18	substitutions	substitution	NNS
cord-005550-qrrdi667	185	19	(	(	-LRB-
cord-005550-qrrdi667	185	20	d	d	NNP
cord-005550-qrrdi667	185	21	N	N	NNP
cord-005550-qrrdi667	185	22	/d	/d	NNP
cord-005550-qrrdi667	185	23	S	S	NNP
cord-005550-qrrdi667	185	24	41	41	CD
cord-005550-qrrdi667	185	25	)	)	-RRB-
cord-005550-qrrdi667	185	26	(	(	-LRB-
cord-005550-qrrdi667	185	27	Bernatchez	Bernatchez	NNP
cord-005550-qrrdi667	185	28	and	and	CC
cord-005550-qrrdi667	185	29	Landry	Landry	NNP
cord-005550-qrrdi667	185	30	,	,	,
cord-005550-qrrdi667	185	31	2003	2003	CD
cord-005550-qrrdi667	185	32	)	)	-RRB-
cord-005550-qrrdi667	185	33	,	,	,
cord-005550-qrrdi667	185	34	and	and	CC
cord-005550-qrrdi667	185	35	thus	thus	RB
cord-005550-qrrdi667	185	36	our	-PRON-	PRP$
cord-005550-qrrdi667	185	37	study	study	NN
cord-005550-qrrdi667	185	38	on	on	IN
cord-005550-qrrdi667	185	39	the	the	DT
cord-005550-qrrdi667	185	40	sac	sac	NN
cord-005550-qrrdi667	185	41	-	-	HYPH
cord-005550-qrrdi667	185	42	winged	winged	JJ
cord-005550-qrrdi667	185	43	bat	bat	NN
cord-005550-qrrdi667	185	44	is	be	VBZ
cord-005550-qrrdi667	185	45	in	in	IN
cord-005550-qrrdi667	185	46	line	line	NN
cord-005550-qrrdi667	185	47	with	with	IN
cord-005550-qrrdi667	185	48	these	these	DT
cord-005550-qrrdi667	185	49	studies	study	NNS
cord-005550-qrrdi667	185	50	.	.	.
cord-005550-qrrdi667	186	1	Whether	whether	IN
cord-005550-qrrdi667	186	2	balancing	balance	VBG
cord-005550-qrrdi667	186	3	selection	selection	NN
cord-005550-qrrdi667	186	4	occurred	occur	VBD
cord-005550-qrrdi667	186	5	over	over	IN
cord-005550-qrrdi667	186	6	the	the	DT
cord-005550-qrrdi667	186	7	history	history	NN
cord-005550-qrrdi667	186	8	of	of	IN
cord-005550-qrrdi667	186	9	the	the	DT
cord-005550-qrrdi667	186	10	species	specie	NNS
cord-005550-qrrdi667	186	11	and	and	CC
cord-005550-qrrdi667	186	12	whether	whether	IN
cord-005550-qrrdi667	186	13	it	-PRON-	PRP
cord-005550-qrrdi667	186	14	still	still	RB
cord-005550-qrrdi667	186	15	acts	act	VBZ
cord-005550-qrrdi667	186	16	in	in	IN
cord-005550-qrrdi667	186	17	contemporary	contemporary	JJ
cord-005550-qrrdi667	186	18	populations	population	NNS
cord-005550-qrrdi667	186	19	can	can	MD
cord-005550-qrrdi667	186	20	be	be	VB
cord-005550-qrrdi667	186	21	tested	test	VBN
cord-005550-qrrdi667	186	22	by	by	IN
cord-005550-qrrdi667	186	23	different	different	JJ
cord-005550-qrrdi667	186	24	approaches	approach	NNS
cord-005550-qrrdi667	186	25	(	(	-LRB-
cord-005550-qrrdi667	186	26	Garrigan	Garrigan	NNP
cord-005550-qrrdi667	186	27	and	and	CC
cord-005550-qrrdi667	186	28	Hedrick	Hedrick	NNP
cord-005550-qrrdi667	186	29	,	,	,
cord-005550-qrrdi667	186	30	2003	2003	CD
cord-005550-qrrdi667	186	31	)	)	-RRB-
cord-005550-qrrdi667	186	32	.	.	.
cord-005550-qrrdi667	187	1	The	the	DT
cord-005550-qrrdi667	187	2	nucleotide	nucleotide	JJ
cord-005550-qrrdi667	187	3	sequence	sequence	NN
cord-005550-qrrdi667	187	4	of	of	IN
cord-005550-qrrdi667	187	5	the	the	DT
cord-005550-qrrdi667	187	6	11	11	CD
cord-005550-qrrdi667	187	7	alleles	allele	NNS
cord-005550-qrrdi667	187	8	differed	differ	VBD
cord-005550-qrrdi667	187	9	substantially	substantially	RB
cord-005550-qrrdi667	187	10	(	(	-LRB-
cord-005550-qrrdi667	187	11	mean	mean	VB
cord-005550-qrrdi667	187	12	14.2	14.2	CD
cord-005550-qrrdi667	187	13	%	%	NN
cord-005550-qrrdi667	187	14	)	)	-RRB-
cord-005550-qrrdi667	187	15	.	.	.
cord-005550-qrrdi667	188	1	This	this	DT
cord-005550-qrrdi667	188	2	is	be	VBZ
cord-005550-qrrdi667	188	3	in	in	IN
cord-005550-qrrdi667	188	4	accordance	accordance	NN
cord-005550-qrrdi667	188	5	with	with	IN
cord-005550-qrrdi667	188	6	balancing	balance	VBG
cord-005550-qrrdi667	188	7	selection	selection	NN
cord-005550-qrrdi667	188	8	in	in	IN
cord-005550-qrrdi667	188	9	the	the	DT
cord-005550-qrrdi667	188	10	past	past	NN
cord-005550-qrrdi667	188	11	,	,	,
cord-005550-qrrdi667	188	12	which	which	WDT
cord-005550-qrrdi667	188	13	can	can	MD
cord-005550-qrrdi667	188	14	maintain	maintain	VB
cord-005550-qrrdi667	188	15	particular	particular	JJ
cord-005550-qrrdi667	188	16	haplotype	haplotype	NN
cord-005550-qrrdi667	188	17	lineages	lineage	NNS
cord-005550-qrrdi667	188	18	for	for	IN
cord-005550-qrrdi667	188	19	a	a	DT
cord-005550-qrrdi667	188	20	long	long	JJ
cord-005550-qrrdi667	188	21	evolutionary	evolutionary	JJ
cord-005550-qrrdi667	188	22	period	period	NN
cord-005550-qrrdi667	188	23	.	.	.
cord-005550-qrrdi667	189	1	Therefore	therefore	RB
cord-005550-qrrdi667	189	2	,	,	,
cord-005550-qrrdi667	189	3	despite	despite	IN
cord-005550-qrrdi667	189	4	the	the	DT
cord-005550-qrrdi667	189	5	important	important	JJ
cord-005550-qrrdi667	189	6	role	role	NN
cord-005550-qrrdi667	189	7	of	of	IN
cord-005550-qrrdi667	189	8	balancing	balance	VBG
cord-005550-qrrdi667	189	9	selection	selection	NN
cord-005550-qrrdi667	189	10	for	for	IN
cord-005550-qrrdi667	189	11	maintaining	maintain	VBG
cord-005550-qrrdi667	189	12	allelic	allelic	JJ
cord-005550-qrrdi667	189	13	diversity	diversity	NN
cord-005550-qrrdi667	189	14	for	for	IN
cord-005550-qrrdi667	189	15	a	a	DT
cord-005550-qrrdi667	189	16	long	long	JJ
cord-005550-qrrdi667	189	17	evolutionary	evolutionary	JJ
cord-005550-qrrdi667	189	18	time	time	NN
cord-005550-qrrdi667	189	19	,	,	,
cord-005550-qrrdi667	189	20	its	-PRON-	PRP$
cord-005550-qrrdi667	189	21	contemporary	contemporary	JJ
cord-005550-qrrdi667	189	22	role	role	NN
cord-005550-qrrdi667	189	23	remains	remain	VBZ
cord-005550-qrrdi667	189	24	unclear	unclear	JJ
cord-005550-qrrdi667	189	25	.	.	.
cord-005550-qrrdi667	190	1	Our	-PRON-	PRP$
cord-005550-qrrdi667	190	2	data	datum	NNS
cord-005550-qrrdi667	190	3	do	do	VBP
cord-005550-qrrdi667	190	4	not	not	RB
cord-005550-qrrdi667	190	5	support	support	VB
cord-005550-qrrdi667	190	6	ongoing	ongoing	JJ
cord-005550-qrrdi667	190	7	balancing	balancing	NN
cord-005550-qrrdi667	190	8	selection	selection	NN
cord-005550-qrrdi667	190	9	at	at	IN
cord-005550-qrrdi667	190	10	least	least	JJS
cord-005550-qrrdi667	190	11	not	not	RB
cord-005550-qrrdi667	190	12	within	within	IN
cord-005550-qrrdi667	190	13	our	-PRON-	PRP$
cord-005550-qrrdi667	190	14	local	local	JJ
cord-005550-qrrdi667	190	15	population	population	NN
cord-005550-qrrdi667	190	16	of	of	IN
cord-005550-qrrdi667	190	17	sac	sac	NN
cord-005550-qrrdi667	190	18	-	-	HYPH
cord-005550-qrrdi667	190	19	winged	winged	JJ
cord-005550-qrrdi667	190	20	bats	bat	NNS
cord-005550-qrrdi667	190	21	from	from	IN
cord-005550-qrrdi667	190	22	the	the	DT
cord-005550-qrrdi667	190	23	biological	biological	JJ
cord-005550-qrrdi667	190	24	station	station	NN
cord-005550-qrrdi667	190	25	La	La	NNP
cord-005550-qrrdi667	190	26	Selva	Selva	NNP
cord-005550-qrrdi667	190	27	.	.	.
cord-005550-qrrdi667	191	1	We	-PRON-	PRP
cord-005550-qrrdi667	191	2	neither	neither	CC
cord-005550-qrrdi667	191	3	found	find	VBD
cord-005550-qrrdi667	191	4	evidence	evidence	NN
cord-005550-qrrdi667	191	5	for	for	IN
cord-005550-qrrdi667	191	6	heterozygote	heterozygote	NN
cord-005550-qrrdi667	191	7	excess	excess	NN
cord-005550-qrrdi667	191	8	nor	nor	CC
cord-005550-qrrdi667	191	9	did	do	VBD
cord-005550-qrrdi667	191	10	the	the	DT
cord-005550-qrrdi667	191	11	Ewens	Ewens	NNP
cord-005550-qrrdi667	191	12	-	-	HYPH
cord-005550-qrrdi667	191	13	Watterson	Watterson	NNP
cord-005550-qrrdi667	191	14	test	test	VBP
cord-005550-qrrdi667	191	15	reveal	reveal	VBP
cord-005550-qrrdi667	191	16	a	a	DT
cord-005550-qrrdi667	191	17	deviation	deviation	NN
cord-005550-qrrdi667	191	18	of	of	IN
cord-005550-qrrdi667	191	19	heterozygosity	heterozygosity	NN
cord-005550-qrrdi667	191	20	from	from	IN
cord-005550-qrrdi667	191	21	a	a	DT
cord-005550-qrrdi667	191	22	neutral	neutral	JJ
cord-005550-qrrdi667	191	23	equilibrium	equilibrium	NN
cord-005550-qrrdi667	191	24	distribution	distribution	NN
cord-005550-qrrdi667	191	25	.	.	.
cord-005550-qrrdi667	192	1	Instead	instead	RB
cord-005550-qrrdi667	192	2	the	the	DT
cord-005550-qrrdi667	192	3	Hardy	Hardy	NNP
cord-005550-qrrdi667	192	4	-	-	HYPH
cord-005550-qrrdi667	192	5	Weinberg	Weinberg	NNP
cord-005550-qrrdi667	192	6	-	-	HYPH
cord-005550-qrrdi667	192	7	Exact	Exact	NNP
cord-005550-qrrdi667	192	8	Test	Test	NNP
cord-005550-qrrdi667	192	9	on	on	IN
cord-005550-qrrdi667	192	10	heterozygote	heterozygote	NN
cord-005550-qrrdi667	192	11	deficiency	deficiency	NN
cord-005550-qrrdi667	192	12	revealed	reveal	VBD
cord-005550-qrrdi667	192	13	an	an	DT
cord-005550-qrrdi667	192	14	almost	almost	RB
cord-005550-qrrdi667	192	15	significant	significant	JJ
cord-005550-qrrdi667	192	16	result	result	NN
cord-005550-qrrdi667	192	17	due	due	IN
cord-005550-qrrdi667	192	18	to	to	IN
cord-005550-qrrdi667	192	19	an	an	DT
cord-005550-qrrdi667	192	20	increased	increase	VBN
cord-005550-qrrdi667	192	21	proportion	proportion	NN
cord-005550-qrrdi667	192	22	of	of	IN
cord-005550-qrrdi667	192	23	homozygotes	homozygote	NNS
cord-005550-qrrdi667	192	24	of	of	IN
cord-005550-qrrdi667	192	25	the	the	DT
cord-005550-qrrdi667	192	26	most	most	RBS
cord-005550-qrrdi667	192	27	common	common	JJ
cord-005550-qrrdi667	192	28	allele	allele	NN
cord-005550-qrrdi667	192	29	.	.	.
cord-005550-qrrdi667	193	1	This	this	DT
cord-005550-qrrdi667	193	2	trend	trend	NN
cord-005550-qrrdi667	193	3	is	be	VBZ
cord-005550-qrrdi667	193	4	in	in	IN
cord-005550-qrrdi667	193	5	contrast	contrast	NN
cord-005550-qrrdi667	193	6	to	to	IN
cord-005550-qrrdi667	193	7	the	the	DT
cord-005550-qrrdi667	193	8	expectation	expectation	NN
cord-005550-qrrdi667	193	9	of	of	IN
cord-005550-qrrdi667	193	10	ongoing	ongoing	JJ
cord-005550-qrrdi667	193	11	balancing	balancing	NN
cord-005550-qrrdi667	193	12	selection	selection	NN
cord-005550-qrrdi667	193	13	due	due	IN
cord-005550-qrrdi667	193	14	to	to	IN
cord-005550-qrrdi667	193	15	a	a	DT
cord-005550-qrrdi667	193	16	heterozygote	heterozygote	NN
cord-005550-qrrdi667	193	17	advantage	advantage	NN
cord-005550-qrrdi667	193	18	(	(	-LRB-
cord-005550-qrrdi667	193	19	Doherty	Doherty	NNP
cord-005550-qrrdi667	193	20	and	and	CC
cord-005550-qrrdi667	193	21	Zinkernagel	Zinkernagel	NNP
cord-005550-qrrdi667	193	22	,	,	,
cord-005550-qrrdi667	193	23	1975	1975	CD
cord-005550-qrrdi667	193	24	)	)	-RRB-
cord-005550-qrrdi667	193	25	.	.	.
cord-005550-qrrdi667	194	1	However	however	RB
cord-005550-qrrdi667	194	2	,	,	,
cord-005550-qrrdi667	194	3	detection	detection	NN
cord-005550-qrrdi667	194	4	of	of	IN
cord-005550-qrrdi667	194	5	selection	selection	NN
cord-005550-qrrdi667	194	6	from	from	IN
cord-005550-qrrdi667	194	7	heterozygote	heterozygote	NN
cord-005550-qrrdi667	194	8	excess	excess	NN
cord-005550-qrrdi667	194	9	has	have	VBZ
cord-005550-qrrdi667	194	10	low	low	JJ
cord-005550-qrrdi667	194	11	power	power	NN
cord-005550-qrrdi667	194	12	.	.	.
cord-005550-qrrdi667	195	1	Small	small	JJ
cord-005550-qrrdi667	195	2	deme	deme	NNP
cord-005550-qrrdi667	195	3	size	size	NN
cord-005550-qrrdi667	195	4	itself	-PRON-	PRP
cord-005550-qrrdi667	195	5	leads	lead	VBZ
cord-005550-qrrdi667	195	6	to	to	TO
cord-005550-qrrdi667	195	7	heterozygote	heterozygote	VB
cord-005550-qrrdi667	195	8	excess	excess	JJ
cord-005550-qrrdi667	195	9	(	(	-LRB-
cord-005550-qrrdi667	195	10	Pudovkin	Pudovkin	NNP
cord-005550-qrrdi667	195	11	et	et	NNP
cord-005550-qrrdi667	195	12	al	al	NNP
cord-005550-qrrdi667	195	13	.	.	NNP
cord-005550-qrrdi667	195	14	,	,	,
cord-005550-qrrdi667	195	15	1996	1996	CD
cord-005550-qrrdi667	195	16	;	;	:
cord-005550-qrrdi667	195	17	Luikart	Luikart	NNP
cord-005550-qrrdi667	195	18	and	and	CC
cord-005550-qrrdi667	195	19	Cornuet	Cornuet	NNP
cord-005550-qrrdi667	195	20	,	,	,
cord-005550-qrrdi667	195	21	1999	1999	CD
cord-005550-qrrdi667	195	22	;	;	:
cord-005550-qrrdi667	195	23	Balloux	Balloux	NNP
cord-005550-qrrdi667	195	24	,	,	,
cord-005550-qrrdi667	195	25	2004	2004	CD
cord-005550-qrrdi667	195	26	)	)	-RRB-
cord-005550-qrrdi667	195	27	and	and	CC
cord-005550-qrrdi667	195	28	cryptic	cryptic	JJ
cord-005550-qrrdi667	195	29	population	population	NN
cord-005550-qrrdi667	195	30	structure	structure	NN
cord-005550-qrrdi667	195	31	could	could	MD
cord-005550-qrrdi667	195	32	cause	cause	VB
cord-005550-qrrdi667	195	33	homozygote	homozygote	NN
cord-005550-qrrdi667	195	34	excess	excess	NN
cord-005550-qrrdi667	195	35	.	.	.
cord-005550-qrrdi667	196	1	Thus	thus	RB
cord-005550-qrrdi667	196	2	,	,	,
cord-005550-qrrdi667	196	3	one	one	PRP
cord-005550-qrrdi667	196	4	would	would	MD
cord-005550-qrrdi667	196	5	need	need	VB
cord-005550-qrrdi667	196	6	a	a	DT
cord-005550-qrrdi667	196	7	comparative	comparative	JJ
cord-005550-qrrdi667	196	8	study	study	NN
cord-005550-qrrdi667	196	9	with	with	IN
cord-005550-qrrdi667	196	10	other	other	JJ
cord-005550-qrrdi667	196	11	markers	marker	NNS
cord-005550-qrrdi667	196	12	to	to	TO
cord-005550-qrrdi667	196	13	see	see	VB
cord-005550-qrrdi667	196	14	if	if	IN
cord-005550-qrrdi667	196	15	observed	observed	JJ
cord-005550-qrrdi667	196	16	excess	excess	NN
cord-005550-qrrdi667	196	17	was	be	VBD
cord-005550-qrrdi667	196	18	itself	-PRON-	PRP
cord-005550-qrrdi667	196	19	in	in	IN
cord-005550-qrrdi667	196	20	excess	excess	NN
cord-005550-qrrdi667	196	21	of	of	IN
cord-005550-qrrdi667	196	22	a	a	DT
cord-005550-qrrdi667	196	23	heterozygote	heterozygote	NN
cord-005550-qrrdi667	196	24	or	or	CC
cord-005550-qrrdi667	196	25	homozygote	homozygote	NN
cord-005550-qrrdi667	196	26	excess	excess	NN
cord-005550-qrrdi667	196	27	at	at	IN
cord-005550-qrrdi667	196	28	other	other	JJ
cord-005550-qrrdi667	196	29	loci	locus	NNS
cord-005550-qrrdi667	196	30	.	.	.
cord-005550-qrrdi667	197	1	Despite	despite	IN
cord-005550-qrrdi667	197	2	the	the	DT
cord-005550-qrrdi667	197	3	diversity	diversity	NN
cord-005550-qrrdi667	197	4	of	of	IN
cord-005550-qrrdi667	197	5	11	11	CD
cord-005550-qrrdi667	197	6	divergent	divergent	JJ
cord-005550-qrrdi667	197	7	alleles	allele	NNS
cord-005550-qrrdi667	197	8	,	,	,
cord-005550-qrrdi667	197	9	the	the	DT
cord-005550-qrrdi667	197	10	observed	observed	JJ
cord-005550-qrrdi667	197	11	and	and	CC
cord-005550-qrrdi667	197	12	expected	expected	JJ
cord-005550-qrrdi667	197	13	heterozygosity	heterozygosity	NN
cord-005550-qrrdi667	197	14	was	be	VBD
cord-005550-qrrdi667	197	15	very	very	RB
cord-005550-qrrdi667	197	16	low	low	JJ
cord-005550-qrrdi667	197	17	(	(	-LRB-
cord-005550-qrrdi667	197	18	0.30	0.30	CD
cord-005550-qrrdi667	197	19	and	and	CC
cord-005550-qrrdi667	197	20	0.35	0.35	CD
cord-005550-qrrdi667	197	21	,	,	,
cord-005550-qrrdi667	197	22	respectively	respectively	RB
cord-005550-qrrdi667	197	23	)	)	-RRB-
cord-005550-qrrdi667	197	24	.	.	.
cord-005550-qrrdi667	198	1	Such	such	JJ
cord-005550-qrrdi667	198	2	low	low	JJ
cord-005550-qrrdi667	198	3	levels	level	NNS
cord-005550-qrrdi667	198	4	of	of	IN
cord-005550-qrrdi667	198	5	heterozygosity	heterozygosity	NN
cord-005550-qrrdi667	198	6	are	be	VBP
cord-005550-qrrdi667	198	7	known	know	VBN
cord-005550-qrrdi667	198	8	only	only	RB
cord-005550-qrrdi667	198	9	from	from	IN
cord-005550-qrrdi667	198	10	populations	population	NNS
cord-005550-qrrdi667	198	11	of	of	IN
cord-005550-qrrdi667	198	12	small	small	JJ
cord-005550-qrrdi667	198	13	size	size	NN
cord-005550-qrrdi667	198	14	or	or	CC
cord-005550-qrrdi667	198	15	with	with	IN
cord-005550-qrrdi667	198	16	a	a	DT
cord-005550-qrrdi667	198	17	bottleneck	bottleneck	NN
cord-005550-qrrdi667	198	18	in	in	IN
cord-005550-qrrdi667	198	19	the	the	DT
cord-005550-qrrdi667	198	20	past	past	NN
cord-005550-qrrdi667	198	21	.	.	.
cord-005550-qrrdi667	199	1	They	-PRON-	PRP
cord-005550-qrrdi667	199	2	are	be	VBP
cord-005550-qrrdi667	199	3	characterized	characterize	VBN
cord-005550-qrrdi667	199	4	by	by	IN
cord-005550-qrrdi667	199	5	a	a	DT
cord-005550-qrrdi667	199	6	small	small	JJ
cord-005550-qrrdi667	199	7	number	number	NN
cord-005550-qrrdi667	199	8	of	of	IN
cord-005550-qrrdi667	199	9	alleles	allele	NNS
cord-005550-qrrdi667	199	10	with	with	IN
cord-005550-qrrdi667	199	11	allele	allele	NN
cord-005550-qrrdi667	199	12	frequencies	frequency	NNS
cord-005550-qrrdi667	199	13	not	not	RB
cord-005550-qrrdi667	199	14	exceeding	exceed	VBG
cord-005550-qrrdi667	199	15	50	50	CD
cord-005550-qrrdi667	199	16	%	%	NN
cord-005550-qrrdi667	199	17	in	in	IN
cord-005550-qrrdi667	199	18	polymorphic	polymorphic	JJ
cord-005550-qrrdi667	199	19	populations	population	NNS
cord-005550-qrrdi667	199	20	(	(	-LRB-
cord-005550-qrrdi667	199	21	Sena	Sena	NNP
cord-005550-qrrdi667	199	22	et	et	NNP
cord-005550-qrrdi667	199	23	al	al	NNP
cord-005550-qrrdi667	199	24	.	.	NNP
cord-005550-qrrdi667	199	25	,	,	,
cord-005550-qrrdi667	199	26	2003	2003	CD
cord-005550-qrrdi667	199	27	;	;	:
cord-005550-qrrdi667	199	28	Aguilar	Aguilar	NNP
cord-005550-qrrdi667	199	29	et	et	NNP
cord-005550-qrrdi667	199	30	al	al	NNP
cord-005550-qrrdi667	199	31	.	.	NNP
cord-005550-qrrdi667	199	32	,	,	,
cord-005550-qrrdi667	199	33	2004	2004	CD
cord-005550-qrrdi667	199	34	;	;	:
cord-005550-qrrdi667	199	35	Amills	Amills	NNP
cord-005550-qrrdi667	199	36	et	et	NNP
cord-005550-qrrdi667	199	37	al	al	NNP
cord-005550-qrrdi667	199	38	.	.	NNP
cord-005550-qrrdi667	199	39	,	,	,
cord-005550-qrrdi667	199	40	2004	2004	CD
cord-005550-qrrdi667	199	41	;	;	:
cord-005550-qrrdi667	199	42	Weber	Weber	NNP
cord-005550-qrrdi667	199	43	et	et	NNP
cord-005550-qrrdi667	199	44	al	al	NNP
cord-005550-qrrdi667	199	45	.	.	NNP
cord-005550-qrrdi667	199	46	,	,	,
cord-005550-qrrdi667	199	47	2004	2004	CD
cord-005550-qrrdi667	199	48	)	)	-RRB-
cord-005550-qrrdi667	199	49	.	.	.
cord-005550-qrrdi667	200	1	In	in	IN
cord-005550-qrrdi667	200	2	contrast	contrast	NN
cord-005550-qrrdi667	200	3	,	,	,
cord-005550-qrrdi667	200	4	in	in	IN
cord-005550-qrrdi667	200	5	the	the	DT
cord-005550-qrrdi667	200	6	sac	sac	NN
cord-005550-qrrdi667	200	7	-	-	HYPH
cord-005550-qrrdi667	200	8	winged	winged	JJ
cord-005550-qrrdi667	200	9	bat	bat	NN
cord-005550-qrrdi667	200	10	,	,	,
cord-005550-qrrdi667	200	11	a	a	DT
cord-005550-qrrdi667	200	12	singlefrequent	singlefrequent	NN
cord-005550-qrrdi667	200	13	allele	allele	NN
cord-005550-qrrdi667	200	14	(	(	-LRB-
cord-005550-qrrdi667	200	15	A1	A1	NNP
cord-005550-qrrdi667	200	16	)	)	-RRB-
cord-005550-qrrdi667	200	17	occurred	occur	VBD
cord-005550-qrrdi667	200	18	at	at	IN
cord-005550-qrrdi667	200	19	a	a	DT
cord-005550-qrrdi667	200	20	frequency	frequency	NN
cord-005550-qrrdi667	200	21	of	of	IN
cord-005550-qrrdi667	200	22	80	80	CD
cord-005550-qrrdi667	200	23	%	%	NN
cord-005550-qrrdi667	200	24	that	that	WDT
cord-005550-qrrdi667	200	25	caused	cause	VBD
cord-005550-qrrdi667	200	26	the	the	DT
cord-005550-qrrdi667	200	27	low	low	JJ
cord-005550-qrrdi667	200	28	heterozygosity	heterozygosity	NN
cord-005550-qrrdi667	200	29	.	.	.
cord-005550-qrrdi667	201	1	The	the	DT
cord-005550-qrrdi667	201	2	nine	nine	CD
cord-005550-qrrdi667	201	3	remaining	remain	VBG
cord-005550-qrrdi667	201	4	alleles	allele	NNS
cord-005550-qrrdi667	201	5	had	have	VBD
cord-005550-qrrdi667	201	6	a	a	DT
cord-005550-qrrdi667	201	7	frequency	frequency	NN
cord-005550-qrrdi667	201	8	of	of	IN
cord-005550-qrrdi667	201	9	5	5	CD
cord-005550-qrrdi667	201	10	%	%	NN
cord-005550-qrrdi667	201	11	or	or	CC
cord-005550-qrrdi667	201	12	less	less	JJR
cord-005550-qrrdi667	201	13	.	.	.
cord-005550-qrrdi667	202	1	Such	such	PDT
cord-005550-qrrdi667	202	2	a	a	DT
cord-005550-qrrdi667	202	3	high	high	JJ
cord-005550-qrrdi667	202	4	frequency	frequency	NN
cord-005550-qrrdi667	202	5	of	of	IN
cord-005550-qrrdi667	202	6	a	a	DT
cord-005550-qrrdi667	202	7	single	single	JJ
cord-005550-qrrdi667	202	8	allele	allele	NN
cord-005550-qrrdi667	202	9	can	can	MD
cord-005550-qrrdi667	202	10	be	be	VB
cord-005550-qrrdi667	202	11	the	the	DT
cord-005550-qrrdi667	202	12	result	result	NN
cord-005550-qrrdi667	202	13	of	of	IN
cord-005550-qrrdi667	202	14	genetic	genetic	JJ
cord-005550-qrrdi667	202	15	drift	drift	NN
cord-005550-qrrdi667	202	16	and	and	CC
cord-005550-qrrdi667	202	17	is	be	VBZ
cord-005550-qrrdi667	202	18	unexpected	unexpected	JJ
cord-005550-qrrdi667	202	19	if	if	IN
cord-005550-qrrdi667	202	20	heterozygotes	heterozygote	NNS
cord-005550-qrrdi667	202	21	(	(	-LRB-
cord-005550-qrrdi667	202	22	Doherty	Doherty	NNP
cord-005550-qrrdi667	202	23	and	and	CC
cord-005550-qrrdi667	202	24	Zinkernagel	Zinkernagel	NNP
cord-005550-qrrdi667	202	25	,	,	,
cord-005550-qrrdi667	202	26	1975	1975	CD
cord-005550-qrrdi667	202	27	)	)	-RRB-
cord-005550-qrrdi667	202	28	or	or	CC
cord-005550-qrrdi667	202	29	rare	rare	JJ
cord-005550-qrrdi667	202	30	alleles	allele	NNS
cord-005550-qrrdi667	202	31	(	(	-LRB-
cord-005550-qrrdi667	202	32	Clarke	Clarke	NNP
cord-005550-qrrdi667	202	33	and	and	CC
cord-005550-qrrdi667	202	34	Kirby	Kirby	NNP
cord-005550-qrrdi667	202	35	,	,	,
cord-005550-qrrdi667	202	36	1966	1966	CD
cord-005550-qrrdi667	202	37	)	)	-RRB-
cord-005550-qrrdi667	202	38	have	have	VBP
cord-005550-qrrdi667	202	39	an	an	DT
cord-005550-qrrdi667	202	40	advantage	advantage	NN
cord-005550-qrrdi667	202	41	.	.	.
cord-005550-qrrdi667	203	1	We	-PRON-	PRP
cord-005550-qrrdi667	203	2	observed	observe	VBD
cord-005550-qrrdi667	203	3	more	more	JJR
cord-005550-qrrdi667	203	4	homozygotes	homozygote	NNS
cord-005550-qrrdi667	203	5	for	for	IN
cord-005550-qrrdi667	203	6	the	the	DT
cord-005550-qrrdi667	203	7	common	common	JJ
cord-005550-qrrdi667	203	8	allele	allele	NN
cord-005550-qrrdi667	203	9	than	than	IN
cord-005550-qrrdi667	203	10	expected	expect	VBN
cord-005550-qrrdi667	203	11	,	,	,
cord-005550-qrrdi667	203	12	although	although	IN
cord-005550-qrrdi667	203	13	this	this	DT
cord-005550-qrrdi667	203	14	result	result	NN
cord-005550-qrrdi667	203	15	was	be	VBD
cord-005550-qrrdi667	203	16	not	not	RB
cord-005550-qrrdi667	203	17	significant	significant	JJ
cord-005550-qrrdi667	203	18	.	.	.
cord-005550-qrrdi667	204	1	An	an	DT
cord-005550-qrrdi667	204	2	excess	excess	NN
cord-005550-qrrdi667	204	3	of	of	IN
cord-005550-qrrdi667	204	4	a	a	DT
cord-005550-qrrdi667	204	5	particular	particular	JJ
cord-005550-qrrdi667	204	6	homozygous	homozygous	JJ
cord-005550-qrrdi667	204	7	genotype	genotype	NN
cord-005550-qrrdi667	204	8	is	be	VBZ
cord-005550-qrrdi667	204	9	expected	expect	VBN
cord-005550-qrrdi667	204	10	,	,	,
cord-005550-qrrdi667	204	11	if	if	IN
cord-005550-qrrdi667	204	12	individuals	individual	NNS
cord-005550-qrrdi667	204	13	with	with	IN
cord-005550-qrrdi667	204	14	this	this	DT
cord-005550-qrrdi667	204	15	genotype	genotype	NN
cord-005550-qrrdi667	204	16	have	have	VBP
cord-005550-qrrdi667	204	17	higher	high	JJR
cord-005550-qrrdi667	204	18	fitness	fitness	NN
cord-005550-qrrdi667	204	19	,	,	,
cord-005550-qrrdi667	204	20	for	for	IN
cord-005550-qrrdi667	204	21	example	example	NN
cord-005550-qrrdi667	204	22	,	,	,
cord-005550-qrrdi667	204	23	by	by	IN
cord-005550-qrrdi667	204	24	having	have	VBG
cord-005550-qrrdi667	204	25	a	a	DT
cord-005550-qrrdi667	204	26	higher	high	JJR
cord-005550-qrrdi667	204	27	chance	chance	NN
cord-005550-qrrdi667	204	28	of	of	IN
cord-005550-qrrdi667	204	29	survival	survival	NN
cord-005550-qrrdi667	204	30	.	.	.
cord-005550-qrrdi667	205	1	One	one	CD
cord-005550-qrrdi667	205	2	possibility	possibility	NN
cord-005550-qrrdi667	205	3	is	be	VBZ
cord-005550-qrrdi667	205	4	pathogen	pathogen	NN
cord-005550-qrrdi667	205	5	-	-	HYPH
cord-005550-qrrdi667	205	6	driven	drive	VBN
cord-005550-qrrdi667	205	7	selection	selection	NN
cord-005550-qrrdi667	205	8	that	that	WDT
cord-005550-qrrdi667	205	9	was	be	VBD
cord-005550-qrrdi667	205	10	recently	recently	RB
cord-005550-qrrdi667	205	11	shown	show	VBN
cord-005550-qrrdi667	205	12	for	for	IN
cord-005550-qrrdi667	205	13	a	a	DT
cord-005550-qrrdi667	205	14	number	number	NN
cord-005550-qrrdi667	205	15	of	of	IN
cord-005550-qrrdi667	205	16	wild	wild	JJ
cord-005550-qrrdi667	205	17	populations	population	NNS
cord-005550-qrrdi667	205	18	of	of	IN
cord-005550-qrrdi667	205	19	several	several	JJ
cord-005550-qrrdi667	205	20	species	specie	NNS
cord-005550-qrrdi667	205	21	.	.	.
cord-005550-qrrdi667	206	1	Particular	particular	JJ
cord-005550-qrrdi667	206	2	alleles	allele	NNS
cord-005550-qrrdi667	206	3	were	be	VBD
cord-005550-qrrdi667	206	4	significantly	significantly	RB
cord-005550-qrrdi667	206	5	increased	increase	VBN
cord-005550-qrrdi667	206	6	in	in	IN
cord-005550-qrrdi667	206	7	frequency	frequency	NN
cord-005550-qrrdi667	206	8	in	in	IN
cord-005550-qrrdi667	206	9	individuals	individual	NNS
cord-005550-qrrdi667	206	10	,	,	,
cord-005550-qrrdi667	206	11	infected	infect	VBN
cord-005550-qrrdi667	206	12	by	by	IN
cord-005550-qrrdi667	206	13	a	a	DT
cord-005550-qrrdi667	206	14	particular	particular	JJ
cord-005550-qrrdi667	206	15	pathogen	pathogen	NN
cord-005550-qrrdi667	206	16	compared	compare	VBN
cord-005550-qrrdi667	206	17	to	to	IN
cord-005550-qrrdi667	206	18	uninfected	uninfected	JJ
cord-005550-qrrdi667	206	19	individuals	individual	NNS
cord-005550-qrrdi667	206	20	(	(	-LRB-
cord-005550-qrrdi667	206	21	reviewed	review	VBN
cord-005550-qrrdi667	206	22	in	in	IN
cord-005550-qrrdi667	206	23	Bernatchez	Bernatchez	NNP
cord-005550-qrrdi667	206	24	and	and	CC
cord-005550-qrrdi667	206	25	Landry	Landry	NNP
cord-005550-qrrdi667	206	26	,	,	,
cord-005550-qrrdi667	206	27	2003	2003	CD
cord-005550-qrrdi667	206	28	;	;	:
cord-005550-qrrdi667	206	29	Piertney	Piertney	NNP
cord-005550-qrrdi667	206	30	and	and	CC
cord-005550-qrrdi667	206	31	Oliver	Oliver	NNP
cord-005550-qrrdi667	206	32	,	,	,
cord-005550-qrrdi667	206	33	2006	2006	CD
cord-005550-qrrdi667	206	34	)	)	-RRB-
cord-005550-qrrdi667	206	35	.	.	.
cord-005550-qrrdi667	207	1	For	for	IN
cord-005550-qrrdi667	207	2	example	example	NN
cord-005550-qrrdi667	207	3	,	,	,
cord-005550-qrrdi667	207	4	the	the	DT
cord-005550-qrrdi667	207	5	frequency	frequency	NN
cord-005550-qrrdi667	207	6	of	of	IN
cord-005550-qrrdi667	207	7	one	one	CD
cord-005550-qrrdi667	207	8	allele	allele	NN
cord-005550-qrrdi667	207	9	differed	differ	VBN
cord-005550-qrrdi667	207	10	by	by	IN
cord-005550-qrrdi667	207	11	the	the	DT
cord-005550-qrrdi667	207	12	factor	factor	NN
cord-005550-qrrdi667	207	13	of	of	IN
cord-005550-qrrdi667	207	14	more	more	JJR
cord-005550-qrrdi667	207	15	than	than	IN
cord-005550-qrrdi667	207	16	two	two	CD
cord-005550-qrrdi667	207	17	between	between	IN
cord-005550-qrrdi667	207	18	nematode	nematode	NN
cord-005550-qrrdi667	207	19	infected	infect	VBN
cord-005550-qrrdi667	207	20	and	and	CC
cord-005550-qrrdi667	207	21	uninfected	uninfected	JJ
cord-005550-qrrdi667	207	22	striped	striped	JJ
cord-005550-qrrdi667	207	23	mice	mouse	NNS
cord-005550-qrrdi667	208	1	Rhabdomys	Rhabdomys	NNP
cord-005550-qrrdi667	208	2	pumilio	pumilio	NNP
cord-005550-qrrdi667	208	3	(	(	-LRB-
cord-005550-qrrdi667	208	4	Froeschke	Froeschke	NNP
cord-005550-qrrdi667	208	5	and	and	CC
cord-005550-qrrdi667	208	6	Sommer	Sommer	NNP
cord-005550-qrrdi667	208	7	,	,	,
cord-005550-qrrdi667	208	8	2005	2005	CD
cord-005550-qrrdi667	208	9	)	)	-RRB-
cord-005550-qrrdi667	208	10	.	.	.
cord-005550-qrrdi667	209	1	The	the	DT
cord-005550-qrrdi667	209	2	high	high	JJ
cord-005550-qrrdi667	209	3	frequency	frequency	NN
cord-005550-qrrdi667	209	4	of	of	IN
cord-005550-qrrdi667	209	5	the	the	DT
cord-005550-qrrdi667	209	6	A1	A1	NNP
cord-005550-qrrdi667	209	7	allele	allele	NN
cord-005550-qrrdi667	209	8	in	in	IN
cord-005550-qrrdi667	209	9	the	the	DT
cord-005550-qrrdi667	209	10	sac	sac	NN
cord-005550-qrrdi667	209	11	-	-	HYPH
cord-005550-qrrdi667	209	12	winged	winged	JJ
cord-005550-qrrdi667	209	13	bat	bat	NN
cord-005550-qrrdi667	209	14	could	could	MD
cord-005550-qrrdi667	209	15	be	be	VB
cord-005550-qrrdi667	209	16	a	a	DT
cord-005550-qrrdi667	209	17	result	result	NN
cord-005550-qrrdi667	209	18	of	of	IN
cord-005550-qrrdi667	209	19	positive	positive	JJ
cord-005550-qrrdi667	209	20	selection	selection	NN
cord-005550-qrrdi667	209	21	by	by	IN
cord-005550-qrrdi667	209	22	an	an	DT
cord-005550-qrrdi667	209	23	unknown	unknown	JJ
cord-005550-qrrdi667	209	24	pathogen	pathogen	NN
cord-005550-qrrdi667	209	25	that	that	WDT
cord-005550-qrrdi667	209	26	invaded	invade	VBD
cord-005550-qrrdi667	209	27	our	-PRON-	PRP$
cord-005550-qrrdi667	209	28	study	study	NN
cord-005550-qrrdi667	209	29	population	population	NN
cord-005550-qrrdi667	209	30	.	.	.
cord-005550-qrrdi667	210	1	This	this	DT
cord-005550-qrrdi667	210	2	remains	remain	VBZ
cord-005550-qrrdi667	210	3	purely	purely	RB
cord-005550-qrrdi667	210	4	speculative	speculative	JJ
cord-005550-qrrdi667	210	5	at	at	IN
cord-005550-qrrdi667	210	6	the	the	DT
cord-005550-qrrdi667	210	7	moment	moment	NN
cord-005550-qrrdi667	210	8	,	,	,
cord-005550-qrrdi667	210	9	as	as	IN
cord-005550-qrrdi667	210	10	we	-PRON-	PRP
cord-005550-qrrdi667	210	11	do	do	VBP
cord-005550-qrrdi667	210	12	not	not	RB
cord-005550-qrrdi667	210	13	have	have	VB
cord-005550-qrrdi667	210	14	information	information	NN
cord-005550-qrrdi667	210	15	on	on	IN
cord-005550-qrrdi667	210	16	the	the	DT
cord-005550-qrrdi667	210	17	infection	infection	NN
cord-005550-qrrdi667	210	18	of	of	IN
cord-005550-qrrdi667	210	19	sac	sac	NN
cord-005550-qrrdi667	210	20	-	-	HYPH
cord-005550-qrrdi667	210	21	winged	winged	JJ
cord-005550-qrrdi667	210	22	bats	bat	NNS
cord-005550-qrrdi667	210	23	by	by	IN
cord-005550-qrrdi667	210	24	parasites	parasite	NNS
cord-005550-qrrdi667	210	25	.	.	.
cord-005550-qrrdi667	211	1	Even	even	RB
cord-005550-qrrdi667	211	2	when	when	WRB
cord-005550-qrrdi667	211	3	positive	positive	JJ
cord-005550-qrrdi667	211	4	selection	selection	NN
cord-005550-qrrdi667	211	5	occurs	occur	VBZ
cord-005550-qrrdi667	211	6	,	,	,
cord-005550-qrrdi667	211	7	allelic	allelic	JJ
cord-005550-qrrdi667	211	8	diversity	diversity	NN
cord-005550-qrrdi667	211	9	can	can	MD
cord-005550-qrrdi667	211	10	be	be	VB
cord-005550-qrrdi667	211	11	maintained	maintain	VBN
cord-005550-qrrdi667	211	12	due	due	IN
cord-005550-qrrdi667	211	13	to	to	IN
cord-005550-qrrdi667	211	14	immigrant	immigrant	JJ
cord-005550-qrrdi667	211	15	individuals	individual	NNS
cord-005550-qrrdi667	211	16	from	from	IN
cord-005550-qrrdi667	211	17	other	other	JJ
cord-005550-qrrdi667	211	18	populations	population	NNS
cord-005550-qrrdi667	211	19	,	,	,
cord-005550-qrrdi667	211	20	if	if	IN
cord-005550-qrrdi667	211	21	these	these	DT
cord-005550-qrrdi667	211	22	populations	population	NNS
cord-005550-qrrdi667	211	23	experience	experience	VBP
cord-005550-qrrdi667	211	24	different	different	JJ
cord-005550-qrrdi667	211	25	selection	selection	NN
cord-005550-qrrdi667	211	26	pressures	pressure	NNS
cord-005550-qrrdi667	211	27	.	.	.
cord-005550-qrrdi667	212	1	The	the	DT
cord-005550-qrrdi667	212	2	analysis	analysis	NN
cord-005550-qrrdi667	212	3	of	of	IN
cord-005550-qrrdi667	212	4	bats	bat	NNS
cord-005550-qrrdi667	212	5	from	from	IN
cord-005550-qrrdi667	212	6	other	other	JJ
cord-005550-qrrdi667	212	7	localities	locality	NNS
cord-005550-qrrdi667	212	8	and	and	CC
cord-005550-qrrdi667	212	9	of	of	IN
cord-005550-qrrdi667	212	10	neutrally	neutrally	RB
cord-005550-qrrdi667	212	11	evolving	evolve	VBG
cord-005550-qrrdi667	212	12	sequence	sequence	NN
cord-005550-qrrdi667	212	13	regions	region	NNS
cord-005550-qrrdi667	212	14	close	close	JJ
cord-005550-qrrdi667	212	15	to	to	IN
cord-005550-qrrdi667	212	16	the	the	DT
cord-005550-qrrdi667	212	17	DRB	DRB	NNP
cord-005550-qrrdi667	212	18	locus	locus	NN
cord-005550-qrrdi667	212	19	will	will	MD
cord-005550-qrrdi667	212	20	give	give	VB
cord-005550-qrrdi667	212	21	some	some	DT
cord-005550-qrrdi667	212	22	further	further	JJ
cord-005550-qrrdi667	212	23	insights	insight	NNS
cord-005550-qrrdi667	212	24	(	(	-LRB-
cord-005550-qrrdi667	212	25	Garrigan	Garrigan	NNP
cord-005550-qrrdi667	212	26	and	and	CC
cord-005550-qrrdi667	212	27	Hedrick	Hedrick	NNP
cord-005550-qrrdi667	212	28	,	,	,
cord-005550-qrrdi667	212	29	2003	2003	CD
cord-005550-qrrdi667	212	30	;	;	:
cord-005550-qrrdi667	212	31	Charlesworth	Charlesworth	NNP
cord-005550-qrrdi667	212	32	,	,	,
cord-005550-qrrdi667	212	33	2006	2006	CD
cord-005550-qrrdi667	212	34	Huang	Huang	NNP
cord-005550-qrrdi667	212	35	MM	MM	NNP
cord-005550-qrrdi667	212	36	,	,	,
cord-005550-qrrdi667	212	37	Arnheim	Arnheim	NNP
cord-005550-qrrdi667	212	38	N	N	NNP
cord-005550-qrrdi667	212	39	,	,	,
cord-005550-qrrdi667	212	40	Erlich	Erlich	NNP
cord-005550-qrrdi667	212	41	H	H	NNP
cord-005550-qrrdi667	212	42	(	(	-LRB-
cord-005550-qrrdi667	212	43	1995	1995	CD
cord-005550-qrrdi667	212	44	)	)	-RRB-
cord-005550-qrrdi667	212	45	.	.	.
cord-005550-qrrdi667	213	1	New	New	NNP
cord-005550-qrrdi667	213	2	HLA	HLA	NNP
cord-005550-qrrdi667	213	3	-	-	HYPH
cord-005550-qrrdi667	213	4	DPB1	DPB1	NNP
cord-005550-qrrdi667	213	5	alleles	allele	NNS
cord-005550-qrrdi667	213	6	generated	generate	VBN
cord-005550-qrrdi667	213	7	by	by	IN
cord-005550-qrrdi667	213	8	interallelic	interallelic	NNP
cord-005550-qrrdi667	213	9	gene	gene	NN
cord-005550-qrrdi667	213	10	conversion	conversion	NN
cord-005550-qrrdi667	213	11	detected	detect	VBN
cord-005550-qrrdi667	213	12	by	by	IN
cord-005550-qrrdi667	213	13	analysis	analysis	NN
cord-005550-qrrdi667	213	14	of	of	IN
cord-005550-qrrdi667	213	15	sperm	sperm	NN
cord-005550-qrrdi667	213	16	.	.	.
cord-005550-qrrdi667	214	1	Nat	Nat	NNP
cord-005550-qrrdi667	214	2	Genet	Genet	NNP
cord-005550-qrrdi667	214	3	10	10	CD
cord-005550-qrrdi667	214	4	:	:	SYM
cord-005550-qrrdi667	214	5	407	407	CD
cord-005550-qrrdi667	214	6	-	-	SYM
cord-005550-qrrdi667	214	7	414	414	CD
cord-005550-qrrdi667	214	8	.	.	.
cord-005550-qrrdi667	215	1	MHC	MHC	NNP
cord-005550-qrrdi667	215	2	diversity	diversity	NN
cord-005550-qrrdi667	215	3	in	in	IN
cord-005550-qrrdi667	215	4	the	the	DT
cord-005550-qrrdi667	215	5	sac	sac	NN
cord-005550-qrrdi667	215	6	-	-	HYPH
cord-005550-qrrdi667	215	7	winged	winged	JJ
cord-005550-qrrdi667	215	8	bat	bat	NN
cord-005550-qrrdi667	216	1	High	high	JJ
cord-005550-qrrdi667	216	2	MHC	MHC	NNP
cord-005550-qrrdi667	216	3	diversity	diversity	NN
cord-005550-qrrdi667	216	4	maintained	maintain	VBN
cord-005550-qrrdi667	216	5	by	by	IN
cord-005550-qrrdi667	216	6	balancing	balance	VBG
cord-005550-qrrdi667	216	7	selection	selection	NN
cord-005550-qrrdi667	216	8	in	in	IN
cord-005550-qrrdi667	216	9	an	an	DT
cord-005550-qrrdi667	216	10	otherwise	otherwise	RB
cord-005550-qrrdi667	216	11	genetically	genetically	RB
cord-005550-qrrdi667	216	12	monomorphic	monomorphic	JJ
cord-005550-qrrdi667	216	13	mammal	mammal	NN
cord-005550-qrrdi667	216	14	Low	low	JJ
cord-005550-qrrdi667	216	15	diversity	diversity	NN
cord-005550-qrrdi667	216	16	in	in	IN
cord-005550-qrrdi667	216	17	the	the	DT
cord-005550-qrrdi667	216	18	major	major	JJ
cord-005550-qrrdi667	216	19	histocompatibility	histocompatibility	NN
cord-005550-qrrdi667	216	20	complex	complex	JJ
cord-005550-qrrdi667	216	21	class	class	NN
cord-005550-qrrdi667	216	22	II	II	NNP
cord-005550-qrrdi667	216	23	DRB1	DRB1	NNP
cord-005550-qrrdi667	216	24	gene	gene	NN
cord-005550-qrrdi667	216	25	of	of	IN
cord-005550-qrrdi667	216	26	the	the	DT
cord-005550-qrrdi667	216	27	Spanish	spanish	JJ
cord-005550-qrrdi667	216	28	ibex	ibex	NN
cord-005550-qrrdi667	216	29	,	,	,
cord-005550-qrrdi667	216	30	Capra	Capra	NNP
cord-005550-qrrdi667	216	31	pyrenaica	pyrenaica	NNP
cord-005550-qrrdi667	216	32	Generation	Generation	NNP
cord-005550-qrrdi667	216	33	of	of	IN
cord-005550-qrrdi667	216	34	MHC	MHC	NNP
cord-005550-qrrdi667	216	35	class	class	NN
cord-005550-qrrdi667	216	36	II	II	NNP
cord-005550-qrrdi667	216	37	diversity	diversity	NN
cord-005550-qrrdi667	216	38	by	by	IN
cord-005550-qrrdi667	216	39	intra	intra	NNP
cord-005550-qrrdi667	216	40	-	-	JJ
cord-005550-qrrdi667	216	41	and	and	CC
cord-005550-qrrdi667	216	42	intergenic	intergenic	JJ
cord-005550-qrrdi667	216	43	recombination	recombination	NN
cord-005550-qrrdi667	217	1	Heterozygote	Heterozygote	NNP
cord-005550-qrrdi667	217	2	excess	excess	NN
cord-005550-qrrdi667	217	3	in	in	IN
cord-005550-qrrdi667	217	4	small	small	JJ
cord-005550-qrrdi667	217	5	populations	population	NNS
cord-005550-qrrdi667	217	6	and	and	CC
cord-005550-qrrdi667	217	7	the	the	DT
cord-005550-qrrdi667	217	8	heterozygote	heterozygote	NN
cord-005550-qrrdi667	217	9	-	-	HYPH
cord-005550-qrrdi667	217	10	excess	excess	JJ
cord-005550-qrrdi667	217	11	effective	effective	JJ
cord-005550-qrrdi667	217	12	population	population	NN
cord-005550-qrrdi667	217	13	size	size	NN
cord-005550-qrrdi667	217	14	MHC	MHC	NNP
cord-005550-qrrdi667	217	15	studies	study	NNS
cord-005550-qrrdi667	217	16	in	in	IN
cord-005550-qrrdi667	217	17	nonmodel	nonmodel	NNP
cord-005550-qrrdi667	217	18	vertebrates	vertebrate	NNS
cord-005550-qrrdi667	217	19	:	:	:
cord-005550-qrrdi667	217	20	what	what	WP
cord-005550-qrrdi667	217	21	have	have	VBP
cord-005550-qrrdi667	217	22	we	-PRON-	PRP
cord-005550-qrrdi667	217	23	learned	learn	VBN
cord-005550-qrrdi667	217	24	about	about	IN
cord-005550-qrrdi667	217	25	natural	natural	JJ
cord-005550-qrrdi667	217	26	selection	selection	NN
cord-005550-qrrdi667	217	27	in	in	IN
cord-005550-qrrdi667	217	28	15	15	CD
cord-005550-qrrdi667	217	29	years	year	NNS
cord-005550-qrrdi667	217	30	?	?	.
cord-005550-qrrdi667	218	1	MHC	MHC	NNP
cord-005550-qrrdi667	218	2	polymorphism	polymorphism	NN
cord-005550-qrrdi667	218	3	under	under	IN
cord-005550-qrrdi667	218	4	host	host	NN
cord-005550-qrrdi667	218	5	-	-	HYPH
cord-005550-qrrdi667	218	6	pathogen	pathogen	NN
cord-005550-qrrdi667	218	7	coevolution	coevolution	NN
cord-005550-qrrdi667	218	8	Three	three	CD
cord-005550-qrrdi667	218	9	-	-	HYPH
cord-005550-qrrdi667	218	10	dimensional	dimensional	JJ
cord-005550-qrrdi667	218	11	structure	structure	NN
cord-005550-qrrdi667	218	12	of	of	IN
cord-005550-qrrdi667	218	13	the	the	DT
cord-005550-qrrdi667	218	14	human	human	JJ
cord-005550-qrrdi667	218	15	class	class	NN
cord-005550-qrrdi667	218	16	II	II	NNP
cord-005550-qrrdi667	218	17	histocompatibility	histocompatibility	NN
cord-005550-qrrdi667	218	18	antigen	antigen	NN
cord-005550-qrrdi667	218	19	HLA	HLA	NNP
cord-005550-qrrdi667	218	20	-	-	HYPH
cord-005550-qrrdi667	218	21	DR1	DR1	NNP
cord-005550-qrrdi667	219	1	A	a	DT
cord-005550-qrrdi667	219	2	long	long	JJ
cord-005550-qrrdi667	219	3	-	-	HYPH
cord-005550-qrrdi667	219	4	term	term	NN
cord-005550-qrrdi667	219	5	genetic	genetic	JJ
cord-005550-qrrdi667	219	6	survey	survey	NN
cord-005550-qrrdi667	219	7	of	of	IN
cord-005550-qrrdi667	219	8	an	an	DT
cord-005550-qrrdi667	219	9	ungulate	ungulate	JJ
cord-005550-qrrdi667	219	10	population	population	NN
cord-005550-qrrdi667	219	11	reveals	reveal	VBZ
cord-005550-qrrdi667	219	12	balancing	balance	VBG
cord-005550-qrrdi667	219	13	selection	selection	NN
cord-005550-qrrdi667	219	14	acting	act	VBG
cord-005550-qrrdi667	219	15	on	on	IN
cord-005550-qrrdi667	219	16	MHC	MHC	NNP
cord-005550-qrrdi667	219	17	through	through	IN
cord-005550-qrrdi667	219	18	spatial	spatial	JJ
cord-005550-qrrdi667	219	19	and	and	CC
cord-005550-qrrdi667	219	20	temporal	temporal	JJ
cord-005550-qrrdi667	219	21	fluctuations	fluctuation	NNS
cord-005550-qrrdi667	219	22	in	in	IN
cord-005550-qrrdi667	219	23	selection	selection	NN
cord-005550-qrrdi667	219	24	Balancing	balancing	NN
cord-005550-qrrdi667	219	25	selection	selection	NN
cord-005550-qrrdi667	219	26	and	and	CC
cord-005550-qrrdi667	219	27	its	-PRON-	PRP$
cord-005550-qrrdi667	219	28	effects	effect	NNS
cord-005550-qrrdi667	219	29	on	on	IN
cord-005550-qrrdi667	219	30	sequences	sequence	NNS
cord-005550-qrrdi667	219	31	in	in	IN
cord-005550-qrrdi667	219	32	nearby	nearby	JJ
cord-005550-qrrdi667	219	33	genome	genome	NN
cord-005550-qrrdi667	219	34	regions	region	NNS
cord-005550-qrrdi667	219	35	Maintenance	maintenance	NN
cord-005550-qrrdi667	219	36	of	of	IN
cord-005550-qrrdi667	219	37	histocompatibility	histocompatibility	NN
cord-005550-qrrdi667	219	38	polymorphisms	polymorphism	NNS
cord-005550-qrrdi667	219	39	Rabies	rabie	NNS
cord-005550-qrrdi667	219	40	and	and	CC
cord-005550-qrrdi667	219	41	rabies	rabies	NN
cord-005550-qrrdi667	219	42	-	-	HYPH
cord-005550-qrrdi667	219	43	related	relate	VBN
cord-005550-qrrdi667	219	44	viruses	virus	NNS
cord-005550-qrrdi667	219	45	:	:	:
cord-005550-qrrdi667	219	46	a	a	DT
cord-005550-qrrdi667	219	47	modern	modern	JJ
cord-005550-qrrdi667	219	48	perspective	perspective	NN
cord-005550-qrrdi667	219	49	on	on	IN
cord-005550-qrrdi667	219	50	an	an	DT
cord-005550-qrrdi667	219	51	ancient	ancient	JJ
cord-005550-qrrdi667	219	52	disease	disease	NN
cord-005550-qrrdi667	220	1	Enhanced	enhanced	JJ
cord-005550-qrrdi667	220	2	immunological	immunological	JJ
cord-005550-qrrdi667	220	3	surveillance	surveillance	NN
cord-005550-qrrdi667	220	4	in	in	IN
cord-005550-qrrdi667	220	5	mice	mouse	NNS
cord-005550-qrrdi667	220	6	heterozygous	heterozygous	JJ
cord-005550-qrrdi667	220	7	at	at	IN
cord-005550-qrrdi667	220	8	the	the	DT
cord-005550-qrrdi667	220	9	H-2	H-2	NNP
cord-005550-qrrdi667	220	10	gene	gene	NN
cord-005550-qrrdi667	220	11	complex	complex	JJ
cord-005550-qrrdi667	220	12	European	european	JJ
cord-005550-qrrdi667	220	13	bat	bat	NN
cord-005550-qrrdi667	220	14	lyssaviruses	lyssavirus	NNS
cord-005550-qrrdi667	220	15	:	:	:
cord-005550-qrrdi667	221	1	an	an	DT
cord-005550-qrrdi667	221	2	emerging	emerge	VBG
cord-005550-qrrdi667	221	3	zoonosis	zoonosis	NN
cord-005550-qrrdi667	221	4	MHC	MHC	NNP
cord-005550-qrrdi667	221	5	class	class	NN
cord-005550-qrrdi667	221	6	II	II	NNP
cord-005550-qrrdi667	221	7	DRB	DRB	NNP
cord-005550-qrrdi667	221	8	variability	variability	NN
cord-005550-qrrdi667	221	9	and	and	CC
cord-005550-qrrdi667	221	10	parasite	parasite	NN
cord-005550-qrrdi667	221	11	load	load	NN
cord-005550-qrrdi667	221	12	in	in	IN
cord-005550-qrrdi667	221	13	the	the	DT
cord-005550-qrrdi667	221	14	striped	striped	JJ
cord-005550-qrrdi667	221	15	mouse	mouse	NN
cord-005550-qrrdi667	221	16	(	(	-LRB-
cord-005550-qrrdi667	221	17	Rhabdomys	Rhabdomys	NNP
cord-005550-qrrdi667	221	18	pumilio	pumilio	NNP
cord-005550-qrrdi667	221	19	)	)	-RRB-
cord-005550-qrrdi667	221	20	in	in	IN
cord-005550-qrrdi667	221	21	the	the	DT
cord-005550-qrrdi667	221	22	Southern	Southern	NNP
cord-005550-qrrdi667	221	23	Kalahari	Kalahari	NNP
cord-005550-qrrdi667	221	24	Perspective	perspective	NN
cord-005550-qrrdi667	221	25	:	:	:
cord-005550-qrrdi667	221	26	detecting	detect	VBG
cord-005550-qrrdi667	221	27	adaptive	adaptive	JJ
cord-005550-qrrdi667	221	28	molecular	molecular	JJ
cord-005550-qrrdi667	221	29	polymorphism	polymorphism	NN
cord-005550-qrrdi667	221	30	:	:	:
cord-005550-qrrdi667	221	31	lessons	lesson	NNS
cord-005550-qrrdi667	221	32	from	from	IN
cord-005550-qrrdi667	221	33	the	the	DT
cord-005550-qrrdi667	221	34	MHC	MHC	NNP
cord-005550-qrrdi667	221	35	Genetic	genetic	JJ
cord-005550-qrrdi667	221	36	diversity	diversity	NN
cord-005550-qrrdi667	221	37	at	at	IN
cord-005550-qrrdi667	221	38	class	class	NN
cord-005550-qrrdi667	221	39	II	II	NNP
cord-005550-qrrdi667	221	40	DRB	DRB	NNP
cord-005550-qrrdi667	221	41	loci	locus	NNS
cord-005550-qrrdi667	221	42	of	of	IN
cord-005550-qrrdi667	221	43	the	the	DT
cord-005550-qrrdi667	221	44	primate	primate	NN
cord-005550-qrrdi667	221	45	MHC	MHC	NNP
cord-005550-qrrdi667	221	46	Genetic	genetic	JJ
cord-005550-qrrdi667	221	47	mating	mating	NN
cord-005550-qrrdi667	221	48	system	system	NN
cord-005550-qrrdi667	221	49	and	and	CC
cord-005550-qrrdi667	221	50	the	the	DT
cord-005550-qrrdi667	221	51	significance	significance	NN
cord-005550-qrrdi667	221	52	of	of	IN
cord-005550-qrrdi667	221	53	harem	harem	JJ
cord-005550-qrrdi667	221	54	associations	association	NNS
cord-005550-qrrdi667	221	55	in	in	IN
cord-005550-qrrdi667	221	56	the	the	DT
cord-005550-qrrdi667	221	57	bat	bat	NN
cord-005550-qrrdi667	221	58	Saccopteryx	Saccopteryx	NNP
cord-005550-qrrdi667	221	59	bilineata	bilineata	NN
cord-005550-qrrdi667	221	60	Evidence	evidence	NN
cord-005550-qrrdi667	221	61	for	for	IN
cord-005550-qrrdi667	221	62	balancing	balance	VBG
cord-005550-qrrdi667	221	63	selection	selection	NN
cord-005550-qrrdi667	221	64	at	at	IN
cord-005550-qrrdi667	221	65	HLA	HLA	NNP
cord-005550-qrrdi667	221	66	Heterozygosity	Heterozygosity	NNP
cord-005550-qrrdi667	221	67	at	at	IN
cord-005550-qrrdi667	221	68	individual	individual	JJ
cord-005550-qrrdi667	221	69	amino	amino	NN
cord-005550-qrrdi667	221	70	acid	acid	NN
cord-005550-qrrdi667	221	71	sites	site	NNS
cord-005550-qrrdi667	221	72	:	:	:
cord-005550-qrrdi667	221	73	extremely	extremely	RB
cord-005550-qrrdi667	221	74	high	high	JJ
cord-005550-qrrdi667	221	75	levels	level	NNS
cord-005550-qrrdi667	221	76	for	for	IN
cord-005550-qrrdi667	221	77	HLA	HLA	NNP
cord-005550-qrrdi667	221	78	-	-	HYPH
cord-005550-qrrdi667	221	79	A	A	NNP
cord-005550-qrrdi667	221	80	and	and	CC
cord-005550-qrrdi667	221	81	-B	-B	NFP
cord-005550-qrrdi667	221	82	genes	gene	NNS
cord-005550-qrrdi667	221	83	Human	human	JJ
cord-005550-qrrdi667	221	84	leukocyte	leukocyte	NN
cord-005550-qrrdi667	221	85	antigens	antigen	NNS
cord-005550-qrrdi667	221	86	and	and	CC
cord-005550-qrrdi667	221	87	natural	natural	JJ
cord-005550-qrrdi667	221	88	selection	selection	NN
cord-005550-qrrdi667	221	89	by	by	IN
cord-005550-qrrdi667	221	90	malaria	malaria	NN
cord-005550-qrrdi667	221	91	Evolution	evolution	NN
cord-005550-qrrdi667	221	92	of	of	IN
cord-005550-qrrdi667	221	93	protein	protein	NN
cord-005550-qrrdi667	221	94	molecules	molecule	NNS
cord-005550-qrrdi667	222	1	Comparative	comparative	JJ
cord-005550-qrrdi667	222	2	genomics	genomic	NNS
cord-005550-qrrdi667	222	3	of	of	IN
cord-005550-qrrdi667	222	4	major	major	JJ
cord-005550-qrrdi667	222	5	histocompatibility	histocompatibility	NN
cord-005550-qrrdi667	222	6	complexes	complex	NNS
cord-005550-qrrdi667	223	1	The	the	DT
cord-005550-qrrdi667	223	2	Natural	natural	JJ
cord-005550-qrrdi667	223	3	History	history	NN
cord-005550-qrrdi667	223	4	of	of	IN
cord-005550-qrrdi667	223	5	the	the	DT
cord-005550-qrrdi667	223	6	Major	Major	NNP
cord-005550-qrrdi667	223	7	Histocompatibility	Histocompatibility	NNP
cord-005550-qrrdi667	223	8	Complex	Complex	NNP
cord-005550-qrrdi667	223	9	Sequences	Sequences	NNPS
cord-005550-qrrdi667	223	10	and	and	CC
cord-005550-qrrdi667	223	11	diversity	diversity	NN
cord-005550-qrrdi667	223	12	of	of	IN
cord-005550-qrrdi667	223	13	17	17	CD
cord-005550-qrrdi667	223	14	new	new	JJ
cord-005550-qrrdi667	223	15	Ovar	Ovar	NNP
cord-005550-qrrdi667	223	16	-	-	HYPH
cord-005550-qrrdi667	223	17	DRB1	DRB1	NNP
cord-005550-qrrdi667	223	18	alleles	allele	NNS
cord-005550-qrrdi667	223	19	from	from	IN
cord-005550-qrrdi667	223	20	three	three	CD
cord-005550-qrrdi667	223	21	breeds	breed	NNS
cord-005550-qrrdi667	223	22	of	of	IN
cord-005550-qrrdi667	223	23	sheep	sheep	NNS
cord-005550-qrrdi667	223	24	MEGA	MEGA	NNP
cord-005550-qrrdi667	223	25	3	3	CD
cord-005550-qrrdi667	223	26	:	:	SYM
cord-005550-qrrdi667	223	27	integrated	integrated	JJ
cord-005550-qrrdi667	223	28	software	software	NN
cord-005550-qrrdi667	223	29	for	for	IN
cord-005550-qrrdi667	223	30	molecular	molecular	JJ
cord-005550-qrrdi667	223	31	evolutionary	evolutionary	JJ
cord-005550-qrrdi667	223	32	genetics	genetics	NN
cord-005550-qrrdi667	223	33	analysis	analysis	NN
cord-005550-qrrdi667	223	34	and	and	CC
cord-005550-qrrdi667	223	35	sequence	sequence	NN
cord-005550-qrrdi667	223	36	alignment	alignment	NN
cord-005550-qrrdi667	223	37	Fruit	Fruit	NNP
cord-005550-qrrdi667	223	38	bats	bat	NNS
cord-005550-qrrdi667	223	39	as	as	IN
cord-005550-qrrdi667	223	40	reservoirs	reservoir	NNS
cord-005550-qrrdi667	223	41	of	of	IN
cord-005550-qrrdi667	223	42	Ebola	Ebola	NNP
cord-005550-qrrdi667	223	43	virus	virus	NN
cord-005550-qrrdi667	223	44	Bats	bat	NNS
cord-005550-qrrdi667	223	45	are	be	VBP
cord-005550-qrrdi667	223	46	natural	natural	JJ
cord-005550-qrrdi667	223	47	reservoirs	reservoir	NNS
cord-005550-qrrdi667	223	48	of	of	IN
cord-005550-qrrdi667	223	49	SARS	SARS	NNP
cord-005550-qrrdi667	223	50	-	-	HYPH
cord-005550-qrrdi667	223	51	like	like	JJ
cord-005550-qrrdi667	223	52	coronaviruses	coronaviruse	NNS
cord-005550-qrrdi667	223	53	Estimating	estimate	VBG
cord-005550-qrrdi667	223	54	the	the	DT
cord-005550-qrrdi667	223	55	effective	effective	JJ
cord-005550-qrrdi667	223	56	number	number	NN
cord-005550-qrrdi667	223	57	of	of	IN
cord-005550-qrrdi667	223	58	breeders	breeder	NNS
cord-005550-qrrdi667	223	59	from	from	IN
cord-005550-qrrdi667	223	60	heterozygote	heterozygote	NN
cord-005550-qrrdi667	223	61	excess	excess	NN
cord-005550-qrrdi667	223	62	in	in	IN
cord-005550-qrrdi667	223	63	progeny	progeny	JJ
cord-005550-qrrdi667	223	64	Major	major	JJ
cord-005550-qrrdi667	223	65	histocompatibility	histocompatibility	NN
cord-005550-qrrdi667	223	66	complex	complex	JJ
cord-005550-qrrdi667	223	67	heterozygote	heterozygote	NN
cord-005550-qrrdi667	223	68	superiority	superiority	NN
cord-005550-qrrdi667	223	69	during	during	IN
cord-005550-qrrdi667	223	70	coinfection	coinfection	NN
cord-005550-qrrdi667	223	71	Monomorphism	Monomorphism	NNP
cord-005550-qrrdi667	223	72	and	and	CC
cord-005550-qrrdi667	223	73	polymorphism	polymorphism	NN
cord-005550-qrrdi667	223	74	at	at	IN
cord-005550-qrrdi667	223	75	MHC	MHC	NNP
cord-005550-qrrdi667	223	76	DRB	DRB	NNP
cord-005550-qrrdi667	223	77	loci	locus	NNS
cord-005550-qrrdi667	223	78	in	in	IN
cord-005550-qrrdi667	223	79	domestic	domestic	JJ
cord-005550-qrrdi667	223	80	and	and	CC
cord-005550-qrrdi667	223	81	wild	wild	JJ
cord-005550-qrrdi667	223	82	ruminants	ruminant	NNS
cord-005550-qrrdi667	224	1	An	an	DT
cord-005550-qrrdi667	224	2	efficient	efficient	JJ
cord-005550-qrrdi667	224	3	saltchloroform	saltchloroform	NN
cord-005550-qrrdi667	224	4	extraction	extraction	NN
cord-005550-qrrdi667	224	5	of	of	IN
cord-005550-qrrdi667	224	6	DNA	DNA	NNP
cord-005550-qrrdi667	224	7	from	from	IN
cord-005550-qrrdi667	224	8	blood	blood	NN
cord-005550-qrrdi667	224	9	and	and	CC
cord-005550-qrrdi667	224	10	tissues	tissue	VBZ
cord-005550-qrrdi667	224	11	Simple	simple	JJ
cord-005550-qrrdi667	224	12	methods	method	NNS
cord-005550-qrrdi667	224	13	for	for	IN
cord-005550-qrrdi667	224	14	estimating	estimate	VBG
cord-005550-qrrdi667	224	15	the	the	DT
cord-005550-qrrdi667	224	16	numbers	number	NNS
cord-005550-qrrdi667	224	17	of	of	IN
cord-005550-qrrdi667	224	18	synonymous	synonymous	JJ
cord-005550-qrrdi667	224	19	and	and	CC
cord-005550-qrrdi667	224	20	nonsynonymous	nonsynonymous	JJ
cord-005550-qrrdi667	224	21	nucleotide	nucleotide	JJ
cord-005550-qrrdi667	224	22	substitutions	substitution	NNS
cord-005550-qrrdi667	224	23	Concerted	concerted	JJ
cord-005550-qrrdi667	224	24	and	and	CC
cord-005550-qrrdi667	224	25	birth	birth	NN
cord-005550-qrrdi667	224	26	-	-	HYPH
cord-005550-qrrdi667	224	27	and	and	CC
cord-005550-qrrdi667	224	28	-	-	HYPH
cord-005550-qrrdi667	224	29	death	death	NN
cord-005550-qrrdi667	224	30	evolution	evolution	NN
cord-005550-qrrdi667	224	31	of	of	IN
cord-005550-qrrdi667	224	32	multigene	multigene	JJ
cord-005550-qrrdi667	224	33	families	family	NNS
cord-005550-qrrdi667	225	1	The	the	DT
cord-005550-qrrdi667	225	2	evolutionary	evolutionary	JJ
cord-005550-qrrdi667	225	3	ecology	ecology	NN
cord-005550-qrrdi667	225	4	of	of	IN
cord-005550-qrrdi667	225	5	the	the	DT
cord-005550-qrrdi667	225	6	major	major	JJ
cord-005550-qrrdi667	225	7	histocompatibility	histocompatibility	NN
cord-005550-qrrdi667	225	8	complex	complex	NN
cord-005550-qrrdi667	226	1	On	on	IN
cord-005550-qrrdi667	226	2	the	the	DT
cord-005550-qrrdi667	226	3	potential	potential	NN
cord-005550-qrrdi667	226	4	for	for	IN
cord-005550-qrrdi667	226	5	estimating	estimate	VBG
cord-005550-qrrdi667	226	6	the	the	DT
cord-005550-qrrdi667	226	7	effective	effective	JJ
cord-005550-qrrdi667	226	8	number	number	NN
cord-005550-qrrdi667	226	9	of	of	IN
cord-005550-qrrdi667	226	10	breeders	breeder	NNS
cord-005550-qrrdi667	226	11	from	from	IN
cord-005550-qrrdi667	226	12	heterozygote	heterozygote	NN
cord-005550-qrrdi667	226	13	-	-	HYPH
cord-005550-qrrdi667	226	14	excess	excess	NN
cord-005550-qrrdi667	226	15	in	in	IN
cord-005550-qrrdi667	226	16	progeny	progeny	JJ
cord-005550-qrrdi667	226	17	GENEPOP	GENEPOP	NNP
cord-005550-qrrdi667	226	18	(	(	-LRB-
cord-005550-qrrdi667	226	19	version	version	NN
cord-005550-qrrdi667	226	20	1.2	1.2	CD
cord-005550-qrrdi667	226	21	)	)	-RRB-
cord-005550-qrrdi667	226	22	:	:	:
cord-005550-qrrdi667	226	23	population	population	NN
cord-005550-qrrdi667	226	24	genetics	genetics	NN
cord-005550-qrrdi667	226	25	software	software	NN
cord-005550-qrrdi667	226	26	for	for	IN
cord-005550-qrrdi667	226	27	exact	exact	JJ
cord-005550-qrrdi667	226	28	tests	test	NNS
cord-005550-qrrdi667	226	29	and	and	CC
cord-005550-qrrdi667	226	30	ecumenicism	ecumenicism	NN
cord-005550-qrrdi667	227	1	Inter	Inter	NNP
cord-005550-qrrdi667	227	2	-	-	:
cord-005550-qrrdi667	227	3	and	and	CC
cord-005550-qrrdi667	227	4	intralocus	intralocus	NNP
cord-005550-qrrdi667	227	5	recombination	recombination	NN
cord-005550-qrrdi667	227	6	drive	drive	NN
cord-005550-qrrdi667	227	7	MHC	MHC	NNP
cord-005550-qrrdi667	227	8	Class	Class	NNP
cord-005550-qrrdi667	227	9	IIB	IIB	NNP
cord-005550-qrrdi667	227	10	gene	gene	NN
cord-005550-qrrdi667	227	11	diversification	diversification	NN
cord-005550-qrrdi667	227	12	in	in	IN
cord-005550-qrrdi667	227	13	a	a	DT
cord-005550-qrrdi667	227	14	teleost	teleost	NN
cord-005550-qrrdi667	227	15	,	,	,
cord-005550-qrrdi667	227	16	the	the	DT
cord-005550-qrrdi667	227	17	three	three	CD
cord-005550-qrrdi667	227	18	-	-	HYPH
cord-005550-qrrdi667	227	19	spined	spined	JJ
cord-005550-qrrdi667	227	20	stickleback	stickleback	NN
cord-005550-qrrdi667	227	21	Gasterosteus	Gasterosteus	NNP
cord-005550-qrrdi667	227	22	aculeatus	aculeatus	NN
cord-005550-qrrdi667	227	23	Relative	relative	JJ
cord-005550-qrrdi667	227	24	roles	role	NNS
cord-005550-qrrdi667	227	25	of	of	IN
cord-005550-qrrdi667	227	26	mutation	mutation	NN
cord-005550-qrrdi667	227	27	and	and	CC
cord-005550-qrrdi667	227	28	recombination	recombination	NN
cord-005550-qrrdi667	227	29	in	in	IN
cord-005550-qrrdi667	227	30	generating	generate	VBG
cord-005550-qrrdi667	227	31	allelic	allelic	JJ
cord-005550-qrrdi667	227	32	polymorphism	polymorphism	NN
cord-005550-qrrdi667	227	33	at	at	IN
cord-005550-qrrdi667	227	34	an	an	DT
cord-005550-qrrdi667	227	35	MHC	MHC	NNP
cord-005550-qrrdi667	227	36	class	class	NN
cord-005550-qrrdi667	227	37	II	II	NNP
cord-005550-qrrdi667	227	38	locus	locus	NN
cord-005550-qrrdi667	227	39	in	in	IN
cord-005550-qrrdi667	227	40	Peromyscus	Peromyscus	NNP
cord-005550-qrrdi667	227	41	maniculatus	maniculatus	NNP
cord-005550-qrrdi667	227	42	Statistical	statistical	JJ
cord-005550-qrrdi667	227	43	tests	test	NNS
cord-005550-qrrdi667	227	44	for	for	IN
cord-005550-qrrdi667	227	45	detecting	detect	VBG
cord-005550-qrrdi667	227	46	gene	gene	NN
cord-005550-qrrdi667	227	47	conversion	conversion	NN
cord-005550-qrrdi667	227	48	MHC	MHC	NNP
cord-005550-qrrdi667	227	49	variability	variability	NN
cord-005550-qrrdi667	227	50	of	of	IN
cord-005550-qrrdi667	227	51	a	a	DT
cord-005550-qrrdi667	227	52	small	small	JJ
cord-005550-qrrdi667	227	53	lemur	lemur	NN
cord-005550-qrrdi667	227	54	in	in	IN
cord-005550-qrrdi667	227	55	the	the	DT
cord-005550-qrrdi667	227	56	littoral	littoral	JJ
cord-005550-qrrdi667	227	57	forest	forest	NN
cord-005550-qrrdi667	227	58	fragments	fragment	NNS
cord-005550-qrrdi667	227	59	of	of	IN
cord-005550-qrrdi667	227	60	southeastern	southeastern	JJ
cord-005550-qrrdi667	227	61	Madagascar	Madagascar	NNP
cord-005550-qrrdi667	227	62	Sequence	Sequence	NNP
cord-005550-qrrdi667	227	63	analysis	analysis	NN
cord-005550-qrrdi667	227	64	of	of	IN
cord-005550-qrrdi667	227	65	the	the	DT
cord-005550-qrrdi667	227	66	HLA	HLA	NNP
cord-005550-qrrdi667	227	67	-	-	HYPH
cord-005550-qrrdi667	227	68	DR	DR	NNP
cord-005550-qrrdi667	227	69	beta	beta	NN
cord-005550-qrrdi667	227	70	and	and	CC
cord-005550-qrrdi667	227	71	HLA	HLA	NNP
cord-005550-qrrdi667	227	72	-	-	HYPH
cord-005550-qrrdi667	227	73	DQ	dq	NN
cord-005550-qrrdi667	227	74	beta	beta	NN
cord-005550-qrrdi667	227	75	loci	locus	NNS
cord-005550-qrrdi667	227	76	from	from	IN
cord-005550-qrrdi667	227	77	three	three	CD
cord-005550-qrrdi667	227	78	Pemphigus	Pemphigus	NNP
cord-005550-qrrdi667	227	79	vulgaris	vulgaris	NN
cord-005550-qrrdi667	227	80	patients	patient	NNS
cord-005550-qrrdi667	227	81	Recombination	Recombination	NNP
cord-005550-qrrdi667	227	82	and	and	CC
cord-005550-qrrdi667	227	83	the	the	DT
cord-005550-qrrdi667	227	84	origin	origin	NN
cord-005550-qrrdi667	227	85	of	of	IN
cord-005550-qrrdi667	227	86	sequence	sequence	NN
cord-005550-qrrdi667	227	87	diversity	diversity	NN
cord-005550-qrrdi667	227	88	in	in	IN
cord-005550-qrrdi667	227	89	the	the	DT
cord-005550-qrrdi667	227	90	DRB	DRB	NNP
cord-005550-qrrdi667	227	91	MHC	MHC	NNP
cord-005550-qrrdi667	227	92	class	class	NN
cord-005550-qrrdi667	227	93	II	II	NNP
cord-005550-qrrdi667	227	94	locus	locus	NN
cord-005550-qrrdi667	227	95	in	in	IN
cord-005550-qrrdi667	227	96	chamois	chamois	NNP
cord-005550-qrrdi667	227	97	(	(	-LRB-
cord-005550-qrrdi667	227	98	Rupicapra	Rupicapra	NNP
cord-005550-qrrdi667	227	99	spp	spp	NNP
cord-005550-qrrdi667	227	100	)	)	-RRB-
cord-005550-qrrdi667	228	1	Arlequin	Arlequin	NNP
cord-005550-qrrdi667	228	2	:	:	:
cord-005550-qrrdi667	228	3	a	a	DT
cord-005550-qrrdi667	228	4	Software	Software	NNP
cord-005550-qrrdi667	228	5	for	for	IN
cord-005550-qrrdi667	228	6	Population	Population	NNP
cord-005550-qrrdi667	228	7	Genetics	Genetics	NNP
cord-005550-qrrdi667	228	8	Data	Data	NNP
cord-005550-qrrdi667	228	9	Analysis	Analysis	NNP
cord-005550-qrrdi667	228	10	.	.	.
cord-005550-qrrdi667	229	1	Genetics	Genetics	NNP
cord-005550-qrrdi667	229	2	and	and	CC
cord-005550-qrrdi667	229	3	Biometry	Biometry	NNP
cord-005550-qrrdi667	229	4	Lab	Lab	NNP
cord-005550-qrrdi667	229	5	Polymorphisms	Polymorphisms	NNPS
cord-005550-qrrdi667	229	6	in	in	IN
cord-005550-qrrdi667	229	7	MHC	MHC	NNP
cord-005550-qrrdi667	229	8	-	-	HYPH
cord-005550-qrrdi667	229	9	DRA	DRA	NNP
cord-005550-qrrdi667	229	10	and	and	CC
cord-005550-qrrdi667	229	11	-DRB	-drb	WDT
cord-005550-qrrdi667	229	12	alleles	allele	NNS
cord-005550-qrrdi667	229	13	of	of	IN
cord-005550-qrrdi667	229	14	water	water	NN
cord-005550-qrrdi667	229	15	buffalo	buffalo	NNP
cord-005550-qrrdi667	229	16	(	(	-LRB-
cord-005550-qrrdi667	229	17	Bubalus	Bubalus	NNP
cord-005550-qrrdi667	229	18	bubalis	bubalis	NNP
cord-005550-qrrdi667	229	19	)	)	-RRB-
cord-005550-qrrdi667	229	20	reveal	reveal	VBP
cord-005550-qrrdi667	229	21	different	different	JJ
cord-005550-qrrdi667	229	22	features	feature	NNS
cord-005550-qrrdi667	229	23	from	from	IN
cord-005550-qrrdi667	229	24	cattle	cattle	NNS
cord-005550-qrrdi667	229	25	DR	DR	NNP
cord-005550-qrrdi667	229	26	alleles	allele	NNS
cord-005550-qrrdi667	229	27	Mammal	Mammal	NNP
cord-005550-qrrdi667	229	28	Species	Species	NNPS
cord-005550-qrrdi667	229	29	of	of	IN
cord-005550-qrrdi667	229	30	the	the	DT
cord-005550-qrrdi667	229	31	World	world	NN
cord-005550-qrrdi667	229	32	:	:	:
cord-005550-qrrdi667	230	1	A	a	DT
cord-005550-qrrdi667	230	2	Taxonomic	Taxonomic	NNP
cord-005550-qrrdi667	230	3	and	and	CC
cord-005550-qrrdi667	230	4	Geographic	Geographic	NNP
cord-005550-qrrdi667	230	5	Reference	Reference	NNP
cord-005550-qrrdi667	231	1	The	the	DT
cord-005550-qrrdi667	231	2	importance	importance	NN
cord-005550-qrrdi667	231	3	of	of	IN
cord-005550-qrrdi667	231	4	immune	immune	JJ
cord-005550-qrrdi667	231	5	gene	gene	NN
cord-005550-qrrdi667	231	6	variability	variability	NN
cord-005550-qrrdi667	231	7	(	(	-LRB-
cord-005550-qrrdi667	231	8	MHC	MHC	NNP
cord-005550-qrrdi667	231	9	)	)	-RRB-
cord-005550-qrrdi667	231	10	in	in	IN
cord-005550-qrrdi667	231	11	evolutionary	evolutionary	JJ
cord-005550-qrrdi667	231	12	ecology	ecology	NN
cord-005550-qrrdi667	231	13	and	and	CC
cord-005550-qrrdi667	231	14	conservation	conservation	NN
cord-005550-qrrdi667	231	15	Major	major	JJ
cord-005550-qrrdi667	231	16	histocompatibility	histocompatibility	NN
cord-005550-qrrdi667	231	17	complex	complex	NN
cord-005550-qrrdi667	231	18	(	(	-LRB-
cord-005550-qrrdi667	231	19	MHC	MHC	NNP
cord-005550-qrrdi667	231	20	)	)	-RRB-
cord-005550-qrrdi667	231	21	class	class	NN
cord-005550-qrrdi667	231	22	II	II	NNP
cord-005550-qrrdi667	231	23	polymorphism	polymorphism	NN
cord-005550-qrrdi667	231	24	and	and	CC
cord-005550-qrrdi667	231	25	paternity	paternity	NN
cord-005550-qrrdi667	231	26	in	in	IN
cord-005550-qrrdi667	231	27	the	the	DT
cord-005550-qrrdi667	231	28	monogamous	monogamous	JJ
cord-005550-qrrdi667	231	29	Hypogeomys	Hypogeomys	NNP
cord-005550-qrrdi667	231	30	antimena	antimena	NNP
cord-005550-qrrdi667	231	31	,	,	,
cord-005550-qrrdi667	231	32	the	the	DT
cord-005550-qrrdi667	231	33	endangered	endanger	VBN
cord-005550-qrrdi667	231	34	,	,	,
cord-005550-qrrdi667	231	35	largest	large	JJS
cord-005550-qrrdi667	231	36	endemic	endemic	JJ
cord-005550-qrrdi667	231	37	Malagasy	Malagasy	NNP
cord-005550-qrrdi667	231	38	rodent	rodent	NN
cord-005550-qrrdi667	231	39	Christian	Christian	NNP
cord-005550-qrrdi667	231	40	Voigt	Voigt	NNP
cord-005550-qrrdi667	231	41	and	and	CC
cord-005550-qrrdi667	231	42	Gerald	Gerald	NNP
cord-005550-qrrdi667	231	43	Heckel	Heckel	NNP
cord-005550-qrrdi667	231	44	participated	participate	VBD
cord-005550-qrrdi667	231	45	in	in	IN
cord-005550-qrrdi667	231	46	field	field	NN
cord-005550-qrrdi667	231	47	work	work	NN
cord-005550-qrrdi667	231	48	.	.	.
cord-005550-qrrdi667	232	1	We	-PRON-	PRP
cord-005550-qrrdi667	232	2	are	be	VBP
cord-005550-qrrdi667	232	3	grateful	grateful	JJ
cord-005550-qrrdi667	232	4	for	for	IN
cord-005550-qrrdi667	232	5	discussions	discussion	NNS
cord-005550-qrrdi667	232	6	and	and	CC
cord-005550-qrrdi667	232	7	suggestions	suggestion	NNS
cord-005550-qrrdi667	232	8	by	by	IN
cord-005550-qrrdi667	232	9	Gerald	Gerald	NNP
cord-005550-qrrdi667	232	10	Heckel	Heckel	NNP
cord-005550-qrrdi667	232	11	,	,	,
cord-005550-qrrdi667	232	12	Otto	Otto	NNP
cord-005550-qrrdi667	232	13	von	von	NNP
cord-005550-qrrdi667	232	14	Helversen	Helversen	NNP
cord-005550-qrrdi667	232	15	,	,	,
cord-005550-qrrdi667	232	16	Martina	Martina	NNP
cord-005550-qrrdi667	232	17	Nagy	Nagy	NNP
cord-005550-qrrdi667	232	18	,	,	,
cord-005550-qrrdi667	232	19	Simone	Simone	NNP
cord-005550-qrrdi667	232	20	Sommer	Sommer	NNP
cord-005550-qrrdi667	232	21	,	,	,
cord-005550-qrrdi667	232	22	Jana	Jana	NNP
cord-005550-qrrdi667	232	23	Ustinova	Ustinova	NNP
cord-005550-qrrdi667	232	24	and	and	CC
cord-005550-qrrdi667	232	25	Christian	Christian	NNP
cord-005550-qrrdi667	232	26	Voigt	Voigt	NNP
cord-005550-qrrdi667	232	27	.	.	.
cord-005550-qrrdi667	233	1	Two	two	CD
cord-005550-qrrdi667	233	2	anonymous	anonymous	JJ
cord-005550-qrrdi667	233	3	referees	referee	NNS
cord-005550-qrrdi667	233	4	provided	provide	VBD
cord-005550-qrrdi667	233	5	important	important	JJ
cord-005550-qrrdi667	233	6	suggestions	suggestion	NNS
cord-005550-qrrdi667	233	7	.	.	.
cord-005550-qrrdi667	234	1	The	the	DT
cord-005550-qrrdi667	234	2	research	research	NN
cord-005550-qrrdi667	234	3	was	be	VBD
cord-005550-qrrdi667	234	4	financially	financially	RB
cord-005550-qrrdi667	234	5	supported	support	VBN
cord-005550-qrrdi667	234	6	by	by	IN
cord-005550-qrrdi667	234	7	the	the	DT
cord-005550-qrrdi667	234	8	Deutsche	Deutsche	NNP
cord-005550-qrrdi667	234	9	Forschungsgemeinschaft	Forschungsgemeinschaft	NNP
cord-005550-qrrdi667	234	10	(	(	-LRB-
cord-005550-qrrdi667	234	11	DFG	DFG	NNP
cord-005550-qrrdi667	234	12	)	)	-RRB-
cord-005550-qrrdi667	234	13	and	and	CC
cord-005550-qrrdi667	234	14	the	the	DT
cord-005550-qrrdi667	234	15	Deutscher	Deutscher	NNP
cord-005550-qrrdi667	234	16	Akademischer	Akademischer	NNP
cord-005550-qrrdi667	234	17	Austauschdienst	Austauschdienst	NNP
cord-005550-qrrdi667	234	18	(	(	-LRB-
cord-005550-qrrdi667	234	19	DAAD	DAAD	NNP
cord-005550-qrrdi667	234	20	)	)	-RRB-
cord-005550-qrrdi667	234	21	.	.	.